Adenodolichos

Taxonomy

Adenodolichos H.A.T. Harms Bot. Jahrb. Syst. 33: 179. 18 Nov 1902.

Subfamily: Faboideae.
Phylogenetic Number: 3.10.83.
Tribe: Phaseoleae.
Subtribe: Cajaninae.
Species Studied - Species in Genus: 3 studied; 15 in genus.

Description

Fruit: A legume; unilocular; 3–5.2 cm long; 1.2–1.5 cm wide; 2–9 times longer than wide; with deciduous androecial sheath; with persistent corolla, or deciduous corolla; with keel petal; with persistent calyx, or deciduous calyx; with calyx shorter than fruit; without orifice formed by curving of fruit or fruit segments; curved; not plicate; not twisted; asymmetrical; C-shaped; with both sutures parallelly curved; not inflated; compressed; without beak; short tapered at apex; oblique with longitudinal axis of fruit; tapered at base; aligned with longitudinal axis of fruit, or right angled with longitudinal axis of fruit; with the apex and base uniform in texture; coriaceous; seed chambers externally invisible; margin not constricted; margin without sulcus; margin plain; wing(s) absent; nonstipitate; with all layers dehiscing; splitting along suture(s). Dehiscence of valves along both sutures; apical and down; active; with valves twisting. Replum invisible. Epicarp dull; monochrome; brown to reddish brown; with surface texture uniform; pubescent and indurate, or pubescent but soon deciduous; with hairs erect; with 1 type of pubescence, or 2 types of pubescence; pilose; with pubescence brown (reddish), or golden; with golden glandular hairs and short-pilose reddish-brown intermixed; with pubescence uniformly distributed; with simple hairs and glandular hairs (sometimes); pliable; with hair bases swollen and plain, or swollen, or plain; glandular; with glandular hairs; without spines; not smooth; with elevated features; veined; reticulately veined; not tuberculate; wrinkled; without cracks; without embedded tissue, much thicker than epicarp, running from base to apex. Mesocarp present; thin; surface not veined; 1-layered; without balsamic vesicles; without reniform canals; solid; coriaceous. Endocarp present; visible; dull and glossy (usually under seed); opaque; monochrome; tan; smooth; without adhering pieces of testa; subseptate; with septa thin (tissue paper-like), flexible; with septa eglandular; chartaceous; not exfoliating; remaining fused to mesocarp and epicarp; without wings; entire. Seed(s) 1, or 2; length parallel with fruit length, or transverse to fruit length; neither overlapping nor touching; in 1 series. Funiculus of 1 length only; flattened; straight. Aril present; dry; when dry rim-aril; entire; gray to tan.

Seed: 6.5–9.5 mm long; 5.5–8.7 mm wide; 2.6–4.1 mm thick; not overgrown; not angular; symmetrical; circular; compressed; with surface smooth; without visible radicle and cotyledon lobes; with shallow hilar sinus; without umbo on seed faces; without medial ridge on each face. Cuticle not exfoliating; not inflated; not wrinkled. Testa present; without pieces of adhering epicarp; not adhering to endocarp; free from endocarp; glossy; not modified by a bloom, or modified by a bloom; colored; monochrome; reddish to dark brown; glabrous; smooth, or not smooth; with recessed features; large depressions on each face; coriaceous. Pleurogram absent. Pseudopleurogram absent. Fracture lines absent. Rim absent. Wing(s) absent. Raphe not visible. Hilum present; partially concealed; concealed by funicular remnant; with faboid split; with the lips of the faboid split the same color as the rest of the hilum; larger than punctiform; 2 mm long; with straight outline; linear; apical according to radicle tip but marginal according to seed length; recessed; within rim; rim color of testa, or lighter than testa. Lens discernible; 0.5–1 mm long; with margins straight; linear; not in groove of raphe; confluent with hilum; mounded; same color as testa; not within corona, halo, or rim, or within rim; rim color of testa. Endosperm present; trace; not pluglike and not resembling tip of radicle; restricted to region of embryo; adnate to testa. Cotyledons smooth; both outer faces convex; both the same thickness; both more or less of equal length; not folded; margin entire 180 degrees from base of radicle; similar at apex; not concealing radicle; notched at radicle; without lobes; with the interface division terminating at base of radicle; without margins recessed; white to tan; inner face flat; glabrous on inner face. Embryonic axis oblique to right angled; oblique to length of seed, or perpendicular to length of seed; without a joint evident between the radicle and the cotyledons. Radicle differentiated from cotyledon; linear; lobe tip straight; oblique to cotyledons; centered between cotyledons; less than 1/2 length of cotyledons. Plumule moderately developed; glabrous.

Distribution

Tropical Africa.

Old World; Africa.

Tribal Notes

Tribe Phaseoleae

Bruneau et al. (1995) carried out cladistic analyses of tribe Phaseoleae using chloroplast DNA restriction site data. Their results indicated that the tribe is not monophyletic and that the tribal delimitations between Phaseoleae and Desmodieae (11) and between Phaseoleae and Millettieae (7) are problematic.