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WIKTROP - Weed Identification and Knowledge in the Tropical and Mediterranean areas
WIKTROP - Weed Identification and Knowledge in the Tropical and Mediterranean areas
SpeciesMapsDocumentsIDAO

Amaranthus viridis L.

Accepted
Amaranthus viridis L.
Amaranthus viridis L.
Amaranthus viridis L.
Amaranthus viridis L.
Amaranthus viridis L.
Amaranthus viridis L.
Amaranthus viridis L.
Amaranthus viridis L.
Amaranthus viridis L.
Amaranthus viridis L.
Amaranthus viridis L.
Amaranthus viridis L.
Amaranthus viridis L.
Amaranthus viridis L.
Amaranthus viridis L.
Amaranthus viridis L.
Amaranthus viridis L.
Amaranthus viridis L.
Amaranthus viridis L.
Amaranthus viridis L.
Amaranthus viridis L.
Amaranthus viridis L.
Amaranthus viridis L.
Amaranthus viridis L.
Amaranthus viridis L.
Amaranthus viridis L.
Amaranthus viridis L.
Amaranthus viridis L.
Amaranthus viridis L.
Amaranthus viridis L.
Amaranthus viridis L.
Amaranthus viridis L.
Amaranthus viridis L.
Amaranthus viridis L.
Amaranthus viridis L.
Amaranthus viridis L.
Amaranthus viridis L.
Amaranthus viridis L.
Amaranthus viridis L.
Amaranthus viridis L.
Amaranthus viridis L.
Amaranthus viridis L.
Amaranthus viridis L.
Amaranthus viridis L.
Amaranthus viridis L.
Amaranthus viridis L.
Amaranthus viridis L.
Amaranthus viridis L.
Amaranthus viridis L.
Amaranthus viridis L.
Amaranthus viridis L.
Amaranthus viridis L.
Amaranthus viridis L.
Amaranthus viridis L.
Amaranthus viridis L.
Amaranthus viridis L.
Amaranthus viridis L.
Amaranthus viridis L.
Amaranthus viridis L.
Amaranthus viridis L.
Amaranthus viridis L.
Amaranthus viridis L.
Amaranthus viridis L.
Amaranthus viridis L.
Amaranthus viridis L.
Amaranthus viridis L.
Amaranthus viridis L.
Amaranthus viridis L.
Amaranthus viridis L.
Amaranthus viridis L.
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Amaranthus viridis L.
/Amaranthus viridis/198.jpg
Amaranthus viridis L.
Amaranthus viridis L.
Amaranthus viridis L.
🗒 Synonyms
synonymAlbersia caudata (Jacq.) Boiss.
synonymAlbersia emarginata Asch.
synonymAlbersia gracilis (Desf.) Webb & Berthel.
synonymAlbersia polystachya (Willd.) Kunth
synonymAmaranthus acutilobus Uline & W.L.Bray
synonymAmaranthus emarginatus Salzm. ex Moq.
synonymAmaranthus fasciatus Roxb.
synonymAmaranthus flexuosus Ambrosi
synonymAmaranthus gracilis Desf.
synonymAmaranthus gracilis Desf. ex Poir.
synonymAmaranthus littoralis Bernh. ex Moq.
synonymAmaranthus major Salzm.
synonymAmaranthus polystachyus Buch.-Ham. ex Wall.
synonymAmaranthus polystachyus Willd.
synonymAmaranthus viridus L.
synonymBlitum viride (L.) Moench
synonymChenopodium caudatum Jacq.
synonymEuxolus caudatus var. gracilis Moq.
synonymEuxolus caudatus var. maximus Moq.
synonymEuxolus polystachyus (Willd.) Miq.
synonymEuxolus viridis (L.) Moq.
synonymGalliaria adscendens Bubani
synonymGlomeraria viridis (L.) Cav.
synonymPyxidium viride (L.) Moq.
🗒 Common Names
Anglais / English
  • Slender amaranth
  • Pig weed
Bengali
  • Shak natey
  • Marissag
Chinese
  • 皱果苋, Zhòu guǒ xiàn
Comorian
  • Debere
Creoles and pidgins; French-based
  • Zépina ranpan (Antilles)
Créole Maurice
  • Brède malabar
Créole Réunion
  • Brède pariétaire
  • Pariétaire
  • Paillatère
  • Amarante
Créole Seychelles
  • Brède malabar
  • Brède pariétaire
  • Bred pariater
English
  • Waterleaf, Green amaranth, Slender amaranth
French
  • Amarante verte, Amarante grêle,
Hindi
  • Jangali chaulai
Italian
  • Amaranto verde
Malgache
  • Anapatsa
Other
  • Niewe bole (Shimaore, Mayotte)
  • Hanga tanana be (Kibushi, Mayotte)
Portuguese
  • Bredo verde,
  • Caruru da mancha, Caruru comun, Caruru de porco, Caruru de soldado, Bredo (Brazil)
Spanish; Castilian
  • Bledo blanco, Caruru, Chichimeca, Citaco (Argentina)
  • Bledo manso, Bledo verde (Colombia)
📚 Overview
Overview
Brief
Code
AMAVI
Growth form
Broadleaf
Biological cycle
annual
Habitat
terrestrial

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    Diagnostic Keys
    Description
    Global description
    Amaranthus viridis is an erect plant or sometimes spread out at the base. It has fairly deep taproots. The stem is green, often becoming red. It is rather slender and highly branched. It is not hairy. The leaves are alternately arranged along the stem. They are oval and are held by long petioles. The apex of the leaf blade is notched and ends with a short tip. The flowers are small, green to reddish in color. They are grouped in small dense balls spread along slender spikes at the base of the leaves and the top of the stems. The fruit is a small capsule which does not open at maturity. It contains a single shiny seed, dark brown to black in colour.

    Cotyledons

    Cotyledons are linear to lanceolate and petiolate. Lamina 18 mm long and 3 mm large, smooth, without apparent nerves. The lower surface can be tainted purple.

    First leaves

    First leaves are simple and alternate, with a long petiole. The lamina is elliptical and then becomes oval. The apex of lamina is deeply notched. The extremity of the central nerve forms a mucro. The lower side is generally reddish purple, at least at its base.

    General habit

    Plant with erect growth habit or somewhat ascending, reaching up to 75 cm high.

    Underground system

    Root system consists of one or more strong branched taproots.

    Stem

    Erect stem and they are generally rounded, may have some ridges, and glabrous (without hairs).

    Leaf

    The leaves are simple and alternate, long-stalked (10 cm). The lamina is triangular to oval, 2 to 7 cm long and 1.5 to 5.5 cm wide. Wedged base, almost perpendicular to the petiole. The apex deeply notched, carrying a mucro. Margin full. 7 to 8 visible side ribs, slightly arched. Upper and lower surfaces glabrous, bearing many small translucent spots.

    Inflorescence

    The inflorescence consists of very small green to red flowers, assembled in sessile glomerules or grouped in axillary and terminal spikes, more or less branched.

    Flower

    The flower has 3 scarious sepals, 1 mm long, linear to lanceolate, acute apex. Female flowers at the base of the spikes, male flowers at the top have 5 stamens. 

    Fruit

    The fruit is an ovoid capsule, indehiscent, measuring 1.2 mm high and 1 mm wide, containing a single seed. Integument with numerous longitudinal ridges. Summit end with a short trifid stigma.

    Seed

    The seed is lenticular, measuring about 1 to 1.25 mm in diameter, smooth and shiny. Dark brown to black in color.
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      No Data
      📚 Natural History
      Life Cycle

      Life cycle

      Annual
      Annual

      Brazil: In the south-eastern region Amaranthus viridis germinates mainly in spring and summer. The cycle from germination to seed production lasts 80-90 days. Under certain conditions, the plant can produce seeds in 4 weeks, allowing 3 to 4 generations per growing season.
      China
      : Amaranthus viridis flowers from June to August and fruits from August to October.
      Mayotte
      : Amaranthus viridis flowers and fruits all year round.
      Morocco: Amaranthus viridis flowers and fruits from May to October.
      Nicaragua: Amaranthus viridis flowers and fruits all year round.
      West Indies: Amaranthus viridis flowers and fruits almost all year round, but mainly from March to June.

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        Cyclicity

        Northern Cameroon: Amaranthus viridis is present throughout the rainy season. In cultivated plot, it is virtually absent at the end of the crop cycle because it is eliminated by weeding. In long days (June-August), the duration of the development cycle, from emergence to seed production, is two months. The plants are then very developed. However, at the end of the rainy season when the days get shorter, the duration of this cycle can be reduced to three weeks. Flowering occurs then in very underdeveloped plants.

         

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          Reproduction

          Amaranthus viridis is annual a species which reproduces only from seeds. The plant can produce 7,000 up to 10 000 seeds that are dispersed by water or wind. They produce a large amount of seeds that have a long viability.

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            Morphology

            Type of prefoliation

            Leaf ratio medium
            Leaf ratio medium
            Broad leaves
            Broad leaves

            Latex

            Without latex
            Without latex

            Root type

            Taproot
            Taproot

            Stipule type

            No stipule
            No stipule

            Fruit type

            Capsule splitting horizontally
            Capsule splitting horizontally

            Lamina base

            rounded
            rounded
            attenuate
            attenuate

            Lamina apex

            mucronate
            mucronate
            emarginate
            emarginate

            Simple leaf type

            Lamina elliptic
            Lamina elliptic

            Lamina section

            flat
            flat
            embossed
            embossed

            Lamina Veination

            in arc
            in arc
            pennate
            pennate

            Flower color

            Red flowers
            Red flowers
            Green
            Green

            Inflorescence type

            Spike
            Spike
            Raceme with alternate sessile flowers
            Raceme with alternate sessile flowers

            Life form

            Broadleaf plant
            Broadleaf plant
            Look Alikes

            Key for Amaranthus

            Prostrate Growth habit A. blitum
            Erect Growth Habit a pair of spines on the leaf axils A. spinosus
            No spine Leaves 10 to 20 cm long A. hybridus
             Leaves 10 cm long Flowers with 3 tepals A. viridis
            Flowers with 5 tepals A. dubius

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              Ecology

              Amaranthus viridis is widespread in tropical and subtropical regions and in the rice fields in altitude. It is found on well-drained soils in open areas and cultures, preferably on sandy soil or humus-rich and rather humid. The plant grows best on land where water is not stagnant. It is very common in Java between altitude of 300 and 600 m.

              Brazil:
              Amaranthus viridis can occur in all regions of Brazil, especially in coffee crops or in plots previously cultivated with coffee. It is a common ruderal species in cities and in disturbed environments, especially on ploughed soils provided they are not excessively wet.
              Northern Cameroon
              : A. viridis is primarily a ruderal species that grows on roadsides, near the villages, especially surrounding the livestock-rearing areas and on the discharge areas of household waste. It is a weed of very infrequent cultures. It is very nitrophilous thus its development is more important when the soil is rich in organic matter and nitrogen.
              China : A ruderal species and a crop weed.
              Comoros: The species grows in cultivated or abandoned land and in the courtyards of houses. It is present at low altitude in all parts of Grande Comore.
              French Guiana: Very common plant on the Mana polder and its surroundings, among the ruderal vegetation of the non-flooded environments. It is a nitrophilic weed common in vegetable crops.
              Madagascar: A species found mainly at medium altitude on  organic soils rich enough or on semi humid soil of terraces with slopes used in the growing of vegetables.
              Mauritius: Common species of cultivated fields, vacant abandoned land, roadsides. It occurs primarily in the humid and dry part of the island.
              Mayotte: Species naturalized in degraded environments. As a weed it is frequent in market gardening, mainly in the north of the island but also in the south.
              Morocco: Amaranthus viridis is a nitrophilous species that thrives on wasteland, ruderalized areas, rubble, crops and wadi beds.
              Nicaragua: An occasional species, in disturbed sites and brackish marshes throughout the country; from 0 to 400 m altitude.
              Reunion: The species is found on well-drained soils. The plant has an average water need. It grows best in soil rich in organic matter and nitrogen. It occurs mainly in low and medium altitude, but very little on the east and south coasts of the island. It is much more common in the West.
              Seychelles: The species grows well on red soil rich in organic matter of high altitude regions and on very sandy soils.
              West Indies: Amaranthus viridis is a common species in all wastelands, stream banks and cultivated fields, especially in irrigated market gardening, from 0 to 200 m (500 m) altitude.

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                No Data
                📚 Habitat and Distribution
                Description

                Geographical distibution

                Madagascar
                Madagascar
                Reunion Island
                Reunion Island
                Comoros
                Comoros
                Mauritius
                Mauritius
                Seychelles
                Seychelles

                Origin

                Amaranthus viridis is native to Central and South America.

                Worldwide distribution

                Amaranthus viridis has become a pantropical species, widespread in all tropical or subtropical regions up to high altitude.

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                  No Data
                  📚 Occurrence
                  No Data
                  📚 Demography and Conservation
                  Population Biology
                  Global Weediness

                  A weed of minor importance.

                  Local Weediness

                  Bangladesh: can cause yield reduction up to 80%.
                  Kenya: Frequent but not abundant.
                  India: The slender amaranthus is considered as one of the most harmful weeds which may lead to heavy losses in crop yield. Its weedy status is more important than its culinary value.
                  Tanzania: Frequent but not abundant.
                  Uganda: Frequent but not abundant.

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                    Risk Statement

                    Global harmfulness

                    Amaranthus viridis is a weed of minor importance.
                     
                    Local harmfulness

                    Burkina Faso: rare and scarce.
                    Chad: frequent and scarce.
                    Comoros
                    : A. viridis is a ruderal species and common weed of vegetable crops.
                    Ivory Coast: frequent and scarce.
                    Ghana: rare and scarce.
                    Kenya: common and scarce.
                    Madagascar: Weed generally infrequent and sparse.
                    Mali: frequent and scarce.
                    Mauritius: A weed with a strong harmfulness in vegetables and sugar cane cultivation, when it grows in large numbers.
                    Mayotte: Amaranthus viridis is more abundant in the north of the island, but also present in the south.
                    Nigeria: rare and scarce.
                    Uganda: common and generally abundant.
                    Reunion: It is a weed that has average ecological amplitude in 23% of the plots. It is uncommon in sugarcane, and found primarily on vegetable crops (Fr> 25%). However, its harmfulness is low, with an overlap of 7 to 15%. Rather it is a ruderal species.
                    Senegal: frequent and scarce.
                    Seychelles: Very common species that can be a serious weed in all cultures.
                    West Indies: Amaranthus viridis is not very present in sugarcane fields. It is particularly widespread in vegetable crops, but has a lesser impact on them.

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                      📚 Uses and Management
                      Uses

                      Food: Young tips eaten as vegetable.
                      Livestock Feeding: It is also given to the pigs.
                      Medicinal: The whole plant extracts of Amaranthus viridis have antidiabetic, antihyperlipidemic and antioxidant effects. An infusion of the entire plant is used in Indonesia as a remedy against intestinal cramps, diarrhea and dysentery, and externally as a cooling agent to treat fever. In Malaysia, used against inflammation and fever, and externally to treat wounds. A. viridis is used in local medicine in Taiwan, Often in mixtures with other medicinal plants, to treat hepatitis, tight chest, bronchitis, asthma, and lung troubles, to stop bleeding and as a hair tonic. It is used locally in India against dysentery, as a cholagogue, abortifacient and to treat snake bite, in flamed wounds and boils, and in Thailand and Sri Lanka as galactagogue.

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                        Management

                        Global control

                        Cultivation favours emergence of A. viridis, and is a useful mean of killing seedlings before sowing rice.

                        Biological controlHypolixus trunculatus whose larve form tunnels and galls in the stems of A. viridis , is known in Pakistan, India and Thailand and feeds on A. viridis. But its suitability as a control agent is hampered by its relatively long life cycle and low reproductive capacity. The caterpillars of Hymenia recurvalis in large number defoliate the weed and, where infestation is severe, practically every plant is infested. But H. recurvalis has a wide host range, is a part of several vegetable crops, which makes it unsuitable for biological control of A. viridis.

                        Chemical: Pre-emergence application of Buttachlor at 2 to 2.5 kg ha-1 gives excellent control. Post-emergence application of 2 4-D at 500 g/ha.

                        Management recommandations for annual broad-leaved weeds in rice fields: http://portal.wiktrop.org/document/show/20

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                          No Data
                          📚 Information Listing
                          References
                          1. Marnotte, P. and A. Carrara. (2007). "Plantes des rizières de Guyane." from http://plantes-rizieres-guyane.cirad.fr/.
                          2. Berton, A. (2020). Flore spontanée des cultures maraichères et fruitières de Guyane. Guide de reconnaissance des 140 adventices les plus communes des parcelles cultivées. Cayenne, Guyane, FREDON Guyane: 186.https://portal.wiktrop.org/document/show/173
                          3. Grard, P., T. Le Bourgeois, J. Rodenburg, P. Marnotte, A. Carrara, R. Irakiza, D. Makokha, G. kyalo, K. Aloys, K. Iswaria, N. Nguyen and G. Tzelepoglou (2012). AFROweeds V.1.0: African weeds of rice. Cédérom. Montpellier, France & Cotonou, Bénin, Cirad-AfricaRice eds.
                          4. https://www.cabi.org/isc/datasheet/4654
                          5. Barthelat, F. 2019. La Flore illustrée de Mayotte. Meze, Paris, France, Collection Inventaires et Biodiversité, Biotope – Muséum national d’Histoire naturelle. 487 p.
                          6. Grossard, F., Le Bourgeois, T., Dumbardon-Martial, E. & Gervais, L. 2013. Adventilles - Guadeloupe & Martinique - Les adventices des Antilles françaises. Abymes, Guadeloupe, France, Les éditions du CTCS Guadeloupe.
                          7. Plants of the World Online https://powo.science.kew.org/taxon/urn:lsid:ipni.org:names:316349-2
                          8. The World Flora Online http://www.worldfloraonline.org/taxon/wfo-0000530552
                          9. Flora of China http://www.efloras.org/florataxon.aspx?flora_id=2&taxon_id=200006991
                          10. Fournet, J. 2002. Flore illustrée des phanérogames de Guadeloupe et de Martinique. Montpellier, France, Cirad, Gondwana éditions.
                          11. Huat, J., Nagy, M., Carpente, A., Schwartz, M., Le Bourgeois, T. & Marnotte, P. 2021. Guide de la flore spontannée des agrosystèmes de Mayotte. Montpellier, Cirad. 150 p.
                          12. Kissmann, K.G. & Groth, D. 1992. Plantas Infestantes e Nocivas. Sao Paulo, Brasil.
                          13. Flora Maroccana (Dobignard A.) https://www.floramaroccana.fr/amaranthus-viridis.html
                          1. Galinato M., Moody K., Piggin C.M. 1999. Upland rice weeds of South and Southeast Asia. IRRI.Philippines.
                          1. Le Bourgeois, T., Carrara, A., Dodet, M., Dogley, W., Gaungoo, A., Grard, P., Ibrahim, Y., Jeuffrault, E., Lebreton, G., Poilecot, P., Prosperi, J., Randriamampianina, J.A., Andrianaivo, A.P., Théveny, F. 2008. Advent-OI : Principales adventices des îles du sud-ouest de l'Océan Indien.V.1.0. In Cirad [ed.]. Cirad, Montpellier, France. Cdrom.
                          1. Bari, M.N. 1997. Major rice weeds in Bangladesh. Department of Agronomy. BSMR Agricultural University, Gazipur, Bangladesh.
                          1. Koch W., 1981. Mauvaises herbes des cultures tropicales. In Kranz J., Schmutterer H. & Koch W. : Maladies, ravageurs et mauvaises herbes des cultures tropicales. Verlag Paul Parey, Berlin, Hambourg : 587-665.
                          1. Le Bourgeois Th., 1993. Les mauvaises herbes dans la rotation cotonnière au Nord-Cameroun (Afrique) - Amplitude d'habitat et degré d'infestation - Cycle de développement. Thèse USTL Montpellier II, Montpellier, France, 241 p.
                          1. Merlier H. & Montégut J., 1982. Adventices tropicales. ORSTOM-GERDAT-ENSH éd., Montpellier, France, 490 p.
                          1. Grard, P., Homsombath, K., Kessler, P., Khuon, E., Le Bourgeois, T., Prospéri, J., Risdale, C. 2006. Oswald V.1.0: A multimedia identification system of the major weeds of rice paddy fields of Cambodia and Lao P.D.R. In Cirad [ed.]. Cirad, Montpellier, France. Cdrom. ISBN 978-2-87614-653-2.
                          1. Berhaut J., 1967. Flore du Sénégal. 2ème éd. Clairafrique éd., Dakar, Sénégal, 485 p.
                          1. Berhaut J., 1971. Flore illustrée du Sénégal. Tome 1. Clairafrique éd., Dakar, Sénégal, 626 p.
                          1. Cavaco A., 1974. Flore du Cameroun. 17 - Amaranthacées. Muséum National d'Histoire Naturelle éd., Paris, France, 65p.
                          1. Hutchinson J., Dalziel J. M., Keay R. W. J. & Hepper F. N., 1954. Flora of West Tropical Africa. Vol. I part. 1. 2ème éd. The Whitefriars Press ed., London & Tonbridge, 295 p.
                          1. Ivens G. W., 1989. East African Weeds and Their Control. Oxford University Press, Nairobi, Kenya, 289 p.
                          1. Ivens G. W., Moody K. & Egunjobi J. K., 1978. West African Weeds. Oxford University Press, Ibadan, Nigeria, 255 p.
                          1. Le Bourgeois T., Jeuffrault E., Grard P., Carrara A. 2001. AdvenRun V.1.0. Les principales mauvaises herbes de La Réunion. CD-ROM. Cirad, SPV. France.
                          2. Nayyar M. M., Ashiq M., Ahmad J. 2001. Manual on Punjab weeds (Part I). Directorate of Agronomy. Ayub Agricultural Research Institute, Faisalabad Pakistan.
                          1. Holm, Leroy/Doll, Jerry/Holm, Eric/Pancho, Jaun/Herberger, James. 1997. World weeds: natural histories and distribution. John Wiley & Sons. 1129 pp.
                          2. Waterhouse D. F. 1994. Biological control of weeds: Southeast Asian prospects. ACIAR Monograph No. 26, 302 pp.
                          Information Listing > References
                          1. Marnotte, P. and A. Carrara. (2007). "Plantes des rizières de Guyane." from http://plantes-rizieres-guyane.cirad.fr/.
                          2. Berton, A. (2020). Flore spontanée des cultures maraichères et fruitières de Guyane. Guide de reconnaissance des 140 adventices les plus communes des parcelles cultivées. Cayenne, Guyane, FREDON Guyane: 186.https://portal.wiktrop.org/document/show/173
                          3. Grard, P., T. Le Bourgeois, J. Rodenburg, P. Marnotte, A. Carrara, R. Irakiza, D. Makokha, G. kyalo, K. Aloys, K. Iswaria, N. Nguyen and G. Tzelepoglou (2012). AFROweeds V.1.0: African weeds of rice. Cédérom. Montpellier, France & Cotonou, Bénin, Cirad-AfricaRice eds.
                          4. https://www.cabi.org/isc/datasheet/4654
                          5. Barthelat, F. 2019. La Flore illustrée de Mayotte. Meze, Paris, France, Collection Inventaires et Biodiversité, Biotope – Muséum national d’Histoire naturelle. 487 p.
                          6. Grossard, F., Le Bourgeois, T., Dumbardon-Martial, E. & Gervais, L. 2013. Adventilles - Guadeloupe & Martinique - Les adventices des Antilles françaises. Abymes, Guadeloupe, France, Les éditions du CTCS Guadeloupe.
                          7. Plants of the World Online https://powo.science.kew.org/taxon/urn:lsid:ipni.org:names:316349-2
                          8. The World Flora Online http://www.worldfloraonline.org/taxon/wfo-0000530552
                          9. Flora of China http://www.efloras.org/florataxon.aspx?flora_id=2&taxon_id=200006991
                          10. Fournet, J. 2002. Flore illustrée des phanérogames de Guadeloupe et de Martinique. Montpellier, France, Cirad, Gondwana éditions.
                          11. Huat, J., Nagy, M., Carpente, A., Schwartz, M., Le Bourgeois, T. & Marnotte, P. 2021. Guide de la flore spontannée des agrosystèmes de Mayotte. Montpellier, Cirad. 150 p.
                          12. Kissmann, K.G. & Groth, D. 1992. Plantas Infestantes e Nocivas. Sao Paulo, Brasil.
                          13. Flora Maroccana (Dobignard A.) https://www.floramaroccana.fr/amaranthus-viridis.html
                          14. Galinato M., Moody K., Piggin C.M. 1999. Upland rice weeds of South and Southeast Asia. IRRI.Philippines.
                          15. Le Bourgeois, T., Carrara, A., Dodet, M., Dogley, W., Gaungoo, A., Grard, P., Ibrahim, Y., Jeuffrault, E., Lebreton, G., Poilecot, P., Prosperi, J., Randriamampianina, J.A., Andrianaivo, A.P., Théveny, F. 2008. Advent-OI : Principales adventices des îles du sud-ouest de l'Océan Indien.V.1.0. In Cirad [ed.]. Cirad, Montpellier, France. Cdrom.
                          16. Bari, M.N. 1997. Major rice weeds in Bangladesh. Department of Agronomy. BSMR Agricultural University, Gazipur, Bangladesh.
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                          18. Le Bourgeois Th., 1993. Les mauvaises herbes dans la rotation cotonnière au Nord-Cameroun (Afrique) - Amplitude d'habitat et degré d'infestation - Cycle de développement. Thèse USTL Montpellier II, Montpellier, France, 241 p.
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                          20. Grard, P., Homsombath, K., Kessler, P., Khuon, E., Le Bourgeois, T., Prospéri, J., Risdale, C. 2006. Oswald V.1.0: A multimedia identification system of the major weeds of rice paddy fields of Cambodia and Lao P.D.R. In Cirad [ed.]. Cirad, Montpellier, France. Cdrom. ISBN 978-2-87614-653-2.
                          21. Berhaut J., 1967. Flore du Sénégal. 2ème éd. Clairafrique éd., Dakar, Sénégal, 485 p.
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                          24. Hutchinson J., Dalziel J. M., Keay R. W. J. & Hepper F. N., 1954. Flora of West Tropical Africa. Vol. I part. 1. 2ème éd. The Whitefriars Press ed., London & Tonbridge, 295 p.
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                          28. Nayyar M. M., Ashiq M., Ahmad J. 2001. Manual on Punjab weeds (Part I). Directorate of Agronomy. Ayub Agricultural Research Institute, Faisalabad Pakistan.
                          29. Holm, Leroy/Doll, Jerry/Holm, Eric/Pancho, Jaun/Herberger, James. 1997. World weeds: natural histories and distribution. John Wiley & Sons. 1129 pp.
                          30. Waterhouse D. F. 1994. Biological control of weeds: Southeast Asian prospects. ACIAR Monograph No. 26, 302 pp.

                          Etude floristique et phytoécologique des adventices des complexes sucriers de Ferké 1 et 2, de Borotou-Koro et de Zuenoula, en Côte d'Ivoire

                          Thomas Le Bourgeois
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                          Thomas Le Bourgeois
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                            WIKTROP - Weed Identification and Knowledge in the Tropical and Mediterranean areasWIKTROP - Weed Identification and Knowledge in the Tropical and Mediterranean areas
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