!"#$#%& !"#$%&'$()*'&+%'+,$%"$&"'($!"%)!*#,$"#-. /01($-#2-3-"%456789:2-.# STRUCTURE AND DIVERSITY OF OAK FORESTS IN THE EL TEPOZTECO NATIONAL PARK (MORELOS, MEXICO) SEBASTIÁN BLOCK AND JORGE A. MEAVE1 Departamento de Ecología y Recursos Naturales, Facultad de Ciencias, Universidad Nacional Autónoma de México, México 04510, D.F., México. 1 Author for correspondence. 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><9C=C?:;<VVW,<;;CAF:=J,>C,/KA:D>8=KGDP,!'$()5,L#R[[T,<=F, >C,>K8,'=J:CD@8A?,eKWVCJ8=W,4ACG@,))),L'e4,)))Q,#RRaT5 , `8, ;C=D>AG;>8F, D?CC>K, D@8;:8D, <;;G?GV<>:C=, ;GAH8D, X:>K,>K8,6<C,(<G,@AC;8FGA8,<=F,<;;G?GV<>:C=,;GAH8D,CB,>K8, 8BB8;>:H8,=G?E8A,CB,D@8;:8D,CB,*+,L2CD>Q,#RRlTd,*+,:D,$K<=I =C=,:=F89,<=F,X<D,;<V;GV<>8F,X:>K,>K8,BCVVCX:=J,BCA?GV<Y ,, , , , ,,, $ $ $ $$$$*^c,b, ,-.,V=,-. $ $ $ $ ∑ .$/$[ XK8A8,,,D><=FD,BCA,>C><V,D@8;:8D,A:;K=8DD,<=F,-.,BCA,>K8,A8V<I >:H8,BA8OG8=;W,CB,>K8,.I>K,D@8;:8D,L6<JGAA<=Q,#RR!T5,(C,8DI >:?<>8,>K8,>C><V,@C>8=>:<V,D:P8,CB,>K8,!,CA<,X8,GD8F,>K8,/K<C, #,<VJCA:>K?,L,%0(1$ 2d,/K<CQ,[am!Q,[amSTQ,XK:;K,:D,E<D8F,C=, !"# :=;:F8=;8, F<><, L@A8D8=;8n<ED8=;8, CB, D@8;:8D, :=, D<?@V:=J, @VC>DTY, , , , ,, , , ,,,,,,o,[# ,,/K<C,#,c,,134,p,bbbbQ ,,,,,, ,,,,,,,#o# XK8A8,,134,:D,>K8,CED8AH8F,=G?E8A,CB,D@8;:8DQ,5[,:D,>K8,=G?I E8A,CB,D@8;:8D,C;;GAA:=J,:=,MGD>,C=8,@VC>,<=F,5#,:D,>K8,=G?E8A, CB,D@8;:8D,C;;GAA:=J,:=,89<;>VW,>XC,@VC>D5,(K8D8,;<V;GV<>:C=D, 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CB,>K8?,Lal5S,qT,X:>K:=,>K8,D<?@V:=J,@VC>D5,`8,;CGVF,:F8=I >:BW,>C,D@8;:8D,V8H8V,<,V<AJ8,BA<;>:C=,CB,>K8D8,D@8;:?8=D,Lm"5a, qTQ, XK:V8, l5\, q, <=F, !5S, q, ;CGVF, E8, C=VW, F8>8A?:=8F, >C, J8=GD,<=F,B<?:VW,V8H8VDQ,A8D@8;>:H8VW5,(K8,A8?<:=:=J,D@8;:I ?8=D,L!5a,qT,;CGVF,=C>,E8,F8>8A?:=8F,>C,<=W,V8H8V5,'@@8=I F:9,#,:D,>K8,;K8;gV:D>,CB,><9<,:F8=>:",8F,<>,V8<D>,>C,J8=GD,V8H8VQ, :=;VGF:=J,mm,B<?:V:8DQ,#Rm,J8=8A<,<=F,"![,D@8;:8D5,'?C=J, >K8,V<>>8AQ,m\5l,q,X8A8,<=J:CD@8A?DQ,[#5[,q,X8A8,B8A=D,<=F, <VV:8DQ, <=F, #5", q, X8A8, JW?=CD@8A?D5,'D>8A<;8<8, X<D, >K8, A:;K8D>,B<?:VWQ,XK:;K,X:>K,\S,D@8;:8D,<=F,#a,J8=8A<Q,V<AJ8VW, Table 1. Number of species and genera of vascular plant families with more than five species, present in the oak forests from the El Tepozteco National Park, Mexico. Family Asteraceae Fabaceae Poaceae Pteridaceae Lamiaceae Solanaceae Polypodiaceae Rubiaceae Orchidaceae Asparagaceae Pinaceae Euphorbiaceae Fagaceae No. of species No. of genera 57 28 19 17 17 13 11 9 7 7 7 6 6 31 18 11 9 7 4 4 7 6 3 1 2 1 @1'(8.6()$,6.!86!4,a",L"TY,!#aI!lRQ,#R[\ #$%&'#()#'#&,-&).&')/#0')1#&-$',#-$.&/$(% 9:2&76C6>&6@26&6A5&UY&\&D;&9:2&46ADE?&4DW2@&M2@2&644DLA925& ;D@&7?&DA>?&GNZ&W2@?&>6@K2&9@22C&F%.(I8&]^_&≥&NP&4<HI&1DAQ W2@C2>?8& 52AC=9?& 4DA42A9@6925& 6<DAK& 9:2& =A5=W=5L6>C& =A& >DQ M2@&5=6<29@=4&4692KD@=2C8&C=A42&<D@2&9:6A&:6>;&D;&9:2&C92<C& M2@2&729M22A&NIS&6A5&GP&4<&=A&]^_I Table 2. Vascular plant genera with more than five species in the oak forests from the El Tepozteco National Park, Mexico. Genus No. of species 10 7 7 7 6 6 6 6 6 Salvia Pinus Polypodium Roldana Ageratina Cestrum Desmodium Quercus Stevia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≥&NIS&4<H8&5=C9@=7L925&6<DAK&SN&CE24=2CI&OD@2& 9:6A&6&XL6@92@&D;&9:2<&M2@2&<L>9=QC92<<258&9:LC&=A&9D96>&M2& <26CL@25&YGY&C92<C&M=9:&6&]^_&`&NIS&4<I&#W2@&YP&\&D;& &140"7%+1)/ 8(*=(()/ 9(14170'1#19%3"#/ <)%*+I& O26A& CE2Q 4=2C&@=4:A2CC&E2@&E>D9&W6@=25&C=KA="&46A9>?&729M22A&K2D<D@Q E:D>DK=46>&LA=9C&F>&a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b2A2@6>>?8& 9:2@2&M2@2&DA>?&DA2&D@&9MD&5D<=A6A9&CE24=2C&=A&264:&LA=9& Table 3. Structural variables assessed in oak forests occurring in six geomorphological units of the El Tepozteco National Park. Values represent means ± 1 S.E. Values between parentheses indicate the ranges for those variables. Values are for plants with DBH ≥ 2.5 cm, which correspond to the stratum named canopy in this study. Geomorphological unit Basal area (m2 ha-1) Cover (m2 ha-1) Height (m) Stem density (ind. ha-1) Chichinautzin 19.6 ± 2.8 (3.4-29.0) 10,424 ± 1,830 (1,743-18,577) 9.0 ± 0.7 (5.5-11.8) 610.0 ± 99.4 (100-1100) Suchiooc 42.0 ± 4.3 (20.3-66.5) 20,919 ± 2,315 (9,889-29,957) 14.4 ± 1.2 (11.0-24.0) 1,040.0 ± 172.0 (400-1,900) Upper Otates 46.4 ± 10.6 (10.3-120.9) 21,526 ± 3,995 (6,440-42,313) 16.7 ± 2.1 (7.2-25.5) 940.0 ± 131.8 (300-1,900) Lower Otates 42.3 ± 7.9 (0.2-81.3) 31,341 ± 6,135 (577-65,475) 15.2 ± 2.0 (2.9-23.2) 900.0 ± 182.0 (100-2,200) Oclayuca 60.5 ± 15.4 (6.5-145.9) 26,041 ± 3,868 (8,536- 44,260) 22.8 ± 4.0 (11.5-55.0) 990.0 ± 172.8 (300-2,000) El Tepozteco Range 32.7 ± 6.2 (0.5-56.1) 22,028 ± 2,903 (2,768-31,671) 13.3 ± 1.3 (7.6-20.0) 810.0 ± 79.5 (400-1,100) *1*")%3"#/!3%()3(+&Z"&F"HT&!NZQ!UP8&NPGS !"" $$%&$9BH.'%/$L&'&.N'O$PQ$'&D'8$&R$ !"#$%&'()&$%&&'()*'*+,)-.+'/'0112%230'+&4'12-,).'5&-'.6)1+).'-+1(4)..'011&-*+47''&''()'80&'902'6-&1)*2-)':1&4'+42&2.'3+4);'7-0/'0-)0'*)< 6+1'.'=#'?'1&4"'*)41)'+4')-,03.@'04*'5&-''()')55)1'+,)'42%A)-'&5'.6)1+).'&5'$(044&4'B4*)C':*0.()*'3+4)@'5&-')01('7)&%&-6(&3&7+103'24+'D A2''%04/'-0-)'.6)1+).'S+'('.+%+30-'BRB.D'9()'&43/')C1)6'+&4' '&''(+.'60'')-4'S0.'$130/210T'S()-)'5&2-'.6)1+).'(0*'(+7(' BRB.D'9()''S&'.6)1+).'(0,+47''()'(+7().''BRB.'+4''()'$3'9)< 6&U')1&'P047)'04*'+4''()'3&S)-'$'0').'5&-).'.T'!"#$%&'()* !*%&(+",')' 04*' -)./)0"#* .1")#"0)2"#T' -).6)1'+,)3/T' (0*' 30-7)'1&,)-',032).'+4',)-/'5)S'63&'.;'&4''()'1&4'-0-/T'&'()-' .6)1+).'3+V)'3%#)&4)*4'"0(.'04*'5./04,("#*#%,),264.'S)-)' ,)-/'5-)W2)4''+4'.&%)'24+'.'A2''(0*',)-/'3&S'1&,)-'*2)''&' '()+-'.%033'.+U)D ' X)'5&24*'1&4.+*)-0A3)'*+55)-)41).'+4''()'.'-21'2-03',0-+< 0A3).' 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People who helped in the identification of plant specimens, arranged by institution. Acronyms: FC-UNAM = Faculty of Sciences, Universidad Nacional Autónoma de México (UNAM); IB-UNAM = Institute of Biology, UNAM; FESI-UNAM = Faculty of Higher Studies at Iztacala, UNAM; FB-UAEM = Faculty of Biology, Universidad Autónoma del Estado de Morelos; CUCBAUdeG = Center for Biological Sciences, Agriculture and Livestock Industry, Universidad de Guadalajara. Institution and name FB-UAEM Rodrigo Hernández Cárdenas CUCBA-UdeG Aarón Rodríguez Contreras IB-UNAM Alfonso Delgado Salinas Eduardo Domínguez Licona Hilda Flores Olvera María del Rosario García Peña Oscar Hinojosa Espinosa Verónica Juárez Jaimes Esteban Martínez Salas Gilda Ortiz Calderón Jaime Pacheco Trejo Angélica Ramírez Roa Clara Hilda Ramos Álvarez Alberto Reyes García Gerardo Adolfo Salazar Chávez Jorge Sánchez Ken !!! IB-UNAM (continues) Rafael Torres Colín José Luis Villaseñor Ríos Leticia Torres Colín FC-UNAM Sebastián Block Munguía Ramiro Cruz Durán Nelly Diego Pérez Jaime Jiménez Ramírez Lucio Lozada Pérez Martha Martínez Gordillo Eduardo Alberto Pérez García Jorge Rojas Gutiérrez Jesús Ricardo de Santiago Pérez Susana Valencia Ávalos Ernesto Velázquez Montes FESI-UNAM Rafael Lira Saade !"#$%&'$()*'&+%'+,,89,:9;<,!=8>!"#?,=#@A $%&'($)*+,+'-.'*/',*0$1,*2$'.%,-$.%/'0%)& Appendix 2. Checklist of vascular plant taxa from the oak forests of El Tepozteco National Park (Morelos, Mexico). Reference vouchers were deposited at MEXU, at FCME [Herbarium of the Faculty of Sciences, Universidad Nacional Autónoma de México (UNAM) (indicated with the herbarium’s code next to voucher number)] or, for vouchers lacking reproductive structures or of few value for a collection, in the plant collection kept at the Plant Ecology and Diversity Laboratory of the Faculty of Sciences, UNAM (indicated with an asterisk). The geomorphological units where each taxon was recorded are also shown. Abbreviations for geomorphological units: CH = Chichinautzin, SU = Suchiooc, UO = upper Otates, LO = lower Otates, OC = Oclayuca, TE = El Tepozteco Range. LYCOPODIOPHYTA Selaginellaceae Willk. Selaginella pallescens (C.Presl) Spring SB 1238 Selaginella aff. porphyrospora A.Braun SB 859* POLYPODIOPHYTA Aspleniaceae Newman Asplenium castaneum Schltdl. & Cham. SB 1254 Asplenium hallbergii Mickel & Beitel SB 817 Asplenium monanthes L. SB 293 Asplenium praemorsum Sw. SB 14 Dennstaedtiaceae Lotsy Pteridium aquilinum (L.) Kuhn var. arachnoideum (Kaulf.) Brade SB 1262* Dryopteridaceae Herter Dryopteris cinnamomea (Cav.) C.Chr. SB 912 Dryopteris maxonii Underw. & C.Chr. SB 1273 Dryopteris patula (Sw.) Underw. SB 1119 Polystichum rachichlaena Fée SB 623 Ophioglossaceae Martinov Botrychium virginianum (L.) Sw. SB 834 Polypodiaceae J. Presl Pecluma ferruginea (M.Martens & Galeotti) M.G.Price SB 764 Phlebodium pseudoaureum (Cav.) Lellinger SB 353 Pleopeltis mexicana (Fée) Mickel & Beitel SB 587* Pleopeltis polylepis (Roemer ex Kunze) T.Moore SB 1358 Polypodium furfuraceum Schltdl. & Cham. SB 748 Polypodium hartwegianum Hook. SB 1044 Polypodium madrense J.Sm. SB 1140 !"#$%&'$()*'&+%'+,'#3'4356'!7#8!9:;'7:<" CH SU X X UO LO OC X TE X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X !!" $%&'$()*+,&-./0,'+1,2.34%,'5,6%'7 % Appendix 2. Continuation. CH Polypodium plesiosorum Kunze SB 29 Polypodium polypodioides Weath. SB 1297 Polypodium rosei Maxon SB 1118 Polypodium subpetiolatum Hook. SB 818 Pteridaceae E.D.M. Kirchn Adiantum andicola Liebm. SB 1215 Adiantum braunii Mett. ex Kuhn SB 761* Adiantum concinnum Humb. & Bonpl. ex Willd. SB 1280 Adiantum poiretii Wikstr. SB 1067 Adiantum raddianum C.Presl SB 533 Anogramma leptophylla (L.) Link SB 95 Bommeria pedata (Sw.) E.Fourn. SB 876* Cheilanthes bonariensis (Willd.) Proctor SB 334 Cheilanthes cuneata Kaulf. ex Link SB 1275 Cheilanthes farinosa (Forssk.) Kaulf. SB 1087 Cheilanthes kaulfussii Kunze SB 1314 Cheilanthes lendigera (Cav.) Sw. SB 332 Cheilanthes marginata Kunth SB 1313* Cheilanthes myriophylla Desv. SB 1256 Cheilanthes pyramidalis Fée SB 697* Notholaena ochracea (Hook.) Yatsk. & Arbeláez SB 1319 Pellaea ternifolia (Cav.) Link SB 331 Pteris quadriaurita Retz. SB 1262* Woodsiaceae Herter Woodsia mollis (Kaulf.) J.Sm. SB 931 GYMNOSPERMAE Cupressaceae Bartl. Juniperus flaccida Schltdl. SB 41* !!" SU UO LO OC TE X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X !"#$%&'$()*'&+%'+,,89,:9;<,!=8>!"#?,=#@A $%&'($)*+,+'-.'*/',*0$1,*2$'.%,-$.%/'0%)& Appendix 2. Continuation. CH Pinaceae Lindl. Pinus aff. ayacahuite C.Ehrenb. ex Schltdl. SB 1352* Pinus aff. hartwegii Lindl. SB 1020* Pinus aff. leiophylla Schiede ex Schltdl. & Cham. SB 514* Pinus aff. maximinoi H.E.Moore SB 248* Pinus aff. montezumae Lamb. SB 1054* Pinus aff. pseudostrobus Lindl. SB 1173* Pinus aff. teocote Schltdl. & Cham. SB 1327* ANGIOSPERMAE: MAGNOLIOPSIDA Actinidiaceae Gilg & Werderm. Saurauia sp. SB 325* Adoxaceae E. Mey. Sambucus sp. SB 1376* Amaranthaceae Juss. Iresine diffusa Humb. & Bonpl. ex Willd. SB 821 Iresine sp. SB 1164* Anacardiaceae R.Br. Toxicodendron radicans (L.) Kuntze SB 612* Annonaceae Juss. Annona cherimola Mill. SB 894* Apiaceae Lindl. Arracacia tolucensis (Kunth) Hemsl. SB 1022* Donnellsmithia mexicana (B.L.Rob.) Mathias & Constance SB 924 Eryngium cf. ghiesbreghtii Decne. SB 727 Eryngium cf. pectinatum C.Presl ex DC. SB 57 Apocynaceae Juss. Asclepias pringlei (Greenm.) Woodson SB 1086* Mandevilla sp. SB 731 Gonolobus uniflorus Kunth SB 1045 Araliaceae Juss. Oreopanax peltatus Linden SB 134* !"#$%&'$()*'&+%'+,'34'5467'!839!:;<'8;=" SU UO LO OC TE X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X !!# $%&'$()*+,&-./0,'+1,2.34%,'5,6%'7% Appendix 2. Continuation. CH Asteraceae Bercht. & J.Presl Acourtia sp1. SB 430 Acourtia sp2. SB 60* Ageratina areolaris (DC.) Gage ex B.L.Turner SB 837 Ageratina choricephala (B.L.Rob.) R.M.King & H.Rob. SB 1283 Ageratina glabrata (Kunth) R.M.King & H.Rob. SB 163 Ageratina mairetiana (DC.) R.M.King & H.Rob. SB 1010 Ageratina pazcuarensis (Kunth) R.M.King & H.Rob. SB 1093 Ageratina aff. petiolaris (Moc. ex DC.) R.M.King & H.Rob. SB 1059 Ageratina photina (B.L. Rob.) R.M.King & H.Rob. SB 520 Ageratum corymbosum Zuccagni SB 525 Alloispermum scabrum (Lag.) H.Rob. SB 1304 Archibaccharis serratifolia (Kunth) S.F.Blake SB 1136 Baccharis conferta Kunth SB 1208 Bidens ocellatus (Greenm.) Melchert SB 196 Bidens sp1. SB 745 Brickellia scoparia (DC.) A.Gray var. scoparia SB 1317 Coreopsis rhyacophila Greenm. SB 598 Cosmos crithmifolius Kunth SB 1346 Critoniopsis salicifolia (DC.) H.Rob. SB 1300 Dahlia coccinea Cav. SB 595 Dahlia merckii Lehm. SB 1008 Dahlia rudis Sørensen SB 566a Eupatorium collinum DC. SB 878 Hieracium sp1. SB 1267* Jaegeria hirta (Lag.) Less. SB 578 !!# SU UO X X LO OC TE X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X !"#$%&'$()*'&+%'+,,89,:9;<,!=8>!?@A,=@B" $%&'($)*+,+'-.'*/',*0$1,*2$'.%,-$.%/'0%)& Appendix 2. Continuation. CH Lagascea rigida (Cav.) Stuessy SB 1311 Melampodium montanum Benth var. viridulum Stuessy SB 1138 Montanoa frutescens (Mairet ex DC.) Hemsl. SB 218 Psacalium peltatum (Kunth) Cass. var. peltatum SB 836 Pseudognaphalium attenuatum var. silvicola Anderb. SB 1007 Pseudognaphalium liebmannii var. monticola (Sch.Bip. ex Klatt) Anderb. SB 1012 Pseudognaphalium oxyphyllum (DC.) Kirp. SB 541 Pseudognaphalium roseum (Kunth) Anderb. SB 599 Roldana albonervia (Greenm.) H.Rob. & Brettell SB 277 Roldana angulifolia (DC.) H.Rob. & Brettell SB 1091 Roldana chapalensis (S.Watson) H.Rob. & Brettell SB 644 Roldana lineolata (DC.) H.Rob. & Brettell SB 477 Roldana lobata La Llave SB 544 Roldana suffulta (Greenm.) H.Rob. & Brettell SB 1341 Roldana sp1. SB 559* Rumfordia floribunda DC. var. floribunda SB 1359 Smallanthus maculatus (Cav.) H.Rob. SB 641 Stevia aschenborniana Sch.Bip. SB 1269 Stevia monardifolia Kunth SB 1196 Stevia origanoides Kunth SB 879 Stevia ovata Willd. SB 391 Stevia suaveolens Lag. SB 1212 Stevia subpubescens Lag. var. subpubescens SB 1018 Tagetes filifolia Lag. SB 402 Tagetes lucida Cav. SB 105 Tagetes lunulata Ortega SB 404 !"#$%&'$()*'&+%'+,'#3'4356'!7#8!9:;'7:<" SU UO LO OC TE X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X !!# $%&'$()*+,&-./0,'+1,2.34%,'5,6%'7 % Appendix 2. Continuation. CH Tridax brachylepis Hemsl. SB 1312 Trigonospermum melampodioides DC. SB 1344 Verbesina fastigiata B.L.Rob. & Greenm. SB 1263 Verbesina oncophora B.L.Rob. & Seaton var. oncophora SB 1103 Verbesina virgata Cav. var. virgata SB 219 Viguiera dentata (Cav.) Spreng. SB 1279 Begoniaceae C. Agardh Begonia cf. balmisiana Balmis SB 604* Begonia gracilis Kunth SB 10 Betulaceae Gray Alnus acuminata Kunth SB 617* Alnus jorullensis Kunth SB 1066* Bignoniaceae Juss. Tecoma stans (L.) Juss. ex Kunth SB 45* Boraginaceae Juss. Lithospermum trinervium (Lehm.) J.Cohen SB 1229 Nama origanifolia Kunth SB 1259 Tournefortia cf. acutiflora M.Martens & Galeotti SB 739 Wigandia urens (Ruiz & Pav.) Kunth SB 1143* Cactaceae Juss. Heliocereus speciosus (Cav.) Britton & Rose subsp. speciosus SB 547 Campanulaceae Juss. Lobelia laxiflora Kunth SB 683* Lobelia schmitzii E.Wimm. SB 26a Caprifoliaceae Juss. Symphoricarpos microphyllus Kunth SB 1234* Valeriana robertianifolia Briq. SB 483 Valeriana sorbifolia var. mexicana (DC.) F.G.Mey. SB 503 Valeriana urticifolia Kunth SB 129 !"# SU UO LO OC TE X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X !"#$%&'$()*'&+%'+,,89,:9;<,!=8>!"#?,=#@A $%&'($)*+,+'-.'*/',*0$1,*2$'.%,-$.%/'0%)& Appendix 2. Continuation. Caryophyllaceae Juss. Arenaria lanuginosa (Michx.) Rohrb. SB 856 Drymaria glandulosa Bartl. SB 762 Drymaria villosa Schltdl. & Cham. SB 1025 Minuartia moehringioides (Moc. & Sessé ex DC.) Mattf. SB 1001 Stellaria cuspidata Willd. ex Schltdl. SB 1230 Celastraceae R. Br. Celastrus pringlei Rose SB 305* Clethraceae Klotzsch Clethra mexicana DC. SB 1000 Convolvulaceae Juss. Ipomoea aff. caudata Fernald SB 654 Ipomoea aff. murucoides Roem. & Schult. SB 890* Ipomoea purpurea (L.) Roth SB 113 Crassulaceae J. St.-Hil. Echeveria gibbiflora Moc. & Sessé ex DC. (No specimens of this species were collected) Sedum frutescens Rose SB 6* Sedum jaliscanum S.Watson SB 899 Sedum oxypetalum Kunth SB 289* Cucurbitaceae Juss. Cyclanthera integrifoliola Cogn. SB 1165 Sechiopsis triqueter (Ser.) Naudin SB 1282 Ericaceae Juss. Arbutus xalapensis Kunth SB 154* Comarostaphylis glaucescens (Kunth) Zucc. ex Klotzsch SB 387 Euphorbiaceae Juss. Acalypha langiana Müll.Arg. SB 52 Acalypha aff. triloba Müll.Arg. SB 118 Acalypha sp. SB 246* Euphorbia ariensis Kunth SB 1251 !"#$%&'$()*'&+%'+,'#3'4356'!7#8!9:;'7:<" CH SU UO X X X LO OC TE X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X !"< $%&'$()*+,&-./0,'+1,2.34%,'5,6%'7 % Appendix 2. Continuation. CH Euphorbia sonorae Rose SB 1287a [FCME] Euphorbia subreniformis S.Watson SB 874 Fabaceae Lindl. Acacia pennatula (Schltdl. & Cham.) Benth. SB 101* Acaciella angustissima (Mill.) Britton & Rose var. angustissima SB 674 Acaciella painteri Britton & Rose var. houghii L.Rico SB 456* Astragalus guatemalensis Hemsl. var. brevidentatus Barneby SB 1340 Brongniartia lupinoides (Kunth) Taub. SB 1285 Calliandra grandiflora (L’Hér.) Benth. SB 1247 Canavalia villosa Benth. SB 104 Cologania broussonetii (Balb.) DC. SB 677 Crotalaria vitellina KerGawl. SB 568 Dalea reclinata (Cav.) Willd. SB 109 Desmodium bellum (S.F.Blake) B.G.Schub. SB 1240 Desmodium aff. cinereum (Kunth) DC. SB 1332* Desmodium aff. densiflorum Hemsl. SB 729* Desmodium intortum (Mill.) Urb. SB 111 Desmodium aff. jaliscanum S.Watson SB 467* Desmodium sp. SB 372* Erythrina breviflora Sessé & Moc. ex DC. SB 567 Eysenhardtia orthocarpa (A.Gray) S.Watson SB 1330* Indigofera thibaudiana DC. SB 106 Lupinus aff. elegans Kunth SB 1207 Lysiloma divaricatum (Jacq.) J.F.Macbr. SB 778* Mimosa albida Humb. & Bonpl. ex Willd. SB 871 Mimosa caerulea Rose SB 1135* !"# SU UO LO X OC TE X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X !"#$%&'$()*'&+%'+,,89,:9;<,!=8>!"#?,=#@A $%&'($)*+,+'-.'*/',*0$1,*2$'.%,-$.%/'0%)& Appendix 2. Continuation. Phaseolus coccineus L. SB 930 Phaseolus leptostachyus Benth. SB 884 Phaseolus vulgaris L. SB 820 Trifolium amabile Kunth SB 126 Vachellia campechiana (Mill.) Seigler & Ebinger SB 1290* Fagaceae Dumort. Quercus castanea Née SB 226 Quercus glabrescens Benth. SB 91 [FCME] Quercus glaucoides M.Martens & Galeotti SB 718 [FCME] Quercus laurina Bonpl. SB 561 Quercus obtusata Bonpl. SB 892 Quercus rugosa Née SB 1 Garryaceae Lindl. Garrya laurifolia Hartw. ex Benth. SB 386* Geraniaceae Juss. Geranium seemannii Peyr. SB 1046 Gesneriaceae Rich. & Juss. Achimenes heterophylla (Mart.) DC. SB 565a Grossulariaceae DC. Ribes sp. SB 1028* Hypericaceae Juss. Hypericum philonotis Schltdl. & Cham. SB 535 Lamiaceae Martinov Asterohyptis stellulata (Benth.) Epling SB 116 [FCME] Cunila lythrifolia Benth. SB 1201 Hyptis mutabilis (Rich.) Briq. SB 107 [FCME] Lepechinia caulescens (Ortega) Epling SB 1199 [FCME] Lepechinia sp1. 1210* Salvia gesneraeflora Lindl. & Paxton SB 1183 !"#$%&'$()*'&+%'+,'#3'4356'!7#8!9:;'7:<" CH SU X X UO LO OC TE X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X !"3 $%&'$()*+,&-./0,'+1,2.34%,'5,6%'7 % Appendix 2. Continuation. CH Salvia iodantha Fernald SB 1334 Salvia lavanduloides Kunth SB 542 Salvia longistyla Benth. SB 1342 [FCME] Salvia mexicana L. SB 187 Salvia polystachia Cav. SB 526 Salvia purpurea Cav. SB 772 [FCME] Salvia rhyacophila (Fernald) Epling SB 1286 [FCME] Salvia sessei Benth. SB 358 Salvia setulosa Fernald SB 838 Salvia aff. tiliifolia Vahl SB 237* Scutellaria dumetorum Schltdl. SB 550 Stachys coccinea Ortega SB 1222 Lauraceae Juss. Litsea glaucescens Kunth SB 740* Persea americana Mill. SB 1380* Linaceae DC. ex Perleb Linum orizabae Planch. SB 1192 Lythraceae J. St.-Hil. Cuphea wrightii A.Gray SB 87 Malvaceae Juss. Sida rhombifolia L. SB 110 Meliaceae Juss. Trichilia sp. SB 31* Moraceae Gaudich. Trophis mexicana (Liebm.) Bureau SB 835* Nyctaginaceae Juss. Mirabilis longiflora L. SB 1166* Oleaceae Hoffmanns. & Link Fraxinus uhdei (Wenz.) Lingelsh. SB 147* !"! SU UO LO OC X X TE X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X !"#$%&'$()*'&+%'+,,89,:9;<,!=8>!"#?,=#@A $%&'($)*+,+'-.'*/',*0$1,*2$'.%,-$.%/'0%)& Appendix 2. Continuation. Onagraceae Juss. Lopezia racemosa Cav. SB 566 Fuchsia thymifolia Kunth subsp. thymifolia SB 1169 Fuchsia sp1. SB 614* Oxalidaceae R.Br. Oxalis aff. hernandezii DC. SB 796* Oxalis aff. jacquiniana Kunth SB 500* Oxalis aff. tetraphylla Cav. SB 652* Orobanchaceae Vent. Castilleja aff. stipifolia G.L.Nesom SB 400* Castilleja tenuiflora Benth. SB 605 Papaveraceae Juss. Bocconia aff. arborea S.Watson SB 888* Passifloraceae Juss. ex Roussel Passiflora cf. exsudans Zucc. SB 465* Pentaphylacaceae Engl. Ternstroemia lineata DC. subsp. lineata SB 1080 Phytolaccaceae R.Br. Phytolacca rivinoides Kunth & C.D.Bouché SB 549 Piperaceae Giseke Peperomia bracteata A.W.Hill SB 428* Peperomia galioides Kunth SB 34 Peperomia hispidula (Sw.) A.Dietr. SB 291 Plantaginaceae Juss. Lophospermum scandens D.Don SB 574 Penstemon roseus (Cerv. ex Sweet) G.Don SB 158 Russelia coccinea (L.) Wettst. SB 732 Polygalaceae Hoffmanns. & Link Monnina ciliolata Sessé & Moc. ex DC. SB 532 Primulaceae Batsch ex Borkh. Anagallis arvensis L. SB 648* !"#$%&'$()*'&+%'+,'#3'4356'!7#8!9:;'7:<" CH SU UO LO OC TE X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X !"" $%&'$()*+,&-./0,'+1,2.34%,'5,6%'7 % Appendix 2. Continuation. Ranunculaceae Juss. Clematis dioica L. SB 702 Delphinium pedatisectum Hemsl. SB 649 Thalictrum gibbosum Lecoy. SB 274* Thalictrum grandifolium S.Watson SB 553 Rosaceae Juss. Alchemilla procumbens Rose SB 1048 Rubus cf. cymosus Rydb. SB 1197* Rubus pringlei Rydb. SB 308 Prunus aff. serotina Ehrh. SB 728* Rubiaceae Juss. Borreria remota (Lam.) Bacigalupo & E.L.Cabral SB 54 Bouvardia ternifolia (Cav.) Schltdl. SB 1307 Crusea coccinea DC. var. coccínea SB 266* Crusea longiflora (Willd. ex Roem. & Schult.) W.R.Anderson SB 811 Crusea sp1. SB 1071* Didymaea mexicana Hook.f. SB 507* Galium aschenbornii Nees & S. Schauer SB 1006 Hoffmannia conzattii B.L.Rob. SB 632* Mitracarpus sp. SB 1182* Sabiaceae Blume Meliosma dentata (Liebm.) Urb. SB 1186* Salicaceae Mirb. Xylosma flexuosa (Kunth) Hemsl. SB 61* Sapindaceae Juss. Dodonaea viscosa Jacq. SB 46 Serjania triquetra Radlk. SB 872* Scrophulariaceae Juss. Buddleja cordata Kunth SB 1046a !"" CH SU UO LO X X X X OC TE X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X !"#$%&'$()*'&+%'+,,89,:9;<,!=8>!"#?,=#@A $%&'($)*+,+'-.'*/',*0$1,*2$'.%,-$.%/'0%)& Appendix 2. Continuation. CH Buddleja parviflora Kunth SB 1100 Buddleja cf. sessiliflora Kunth SB 1097 Solanaceae Juss. Cestrum anagyris Dunal SB 1090* Cestrum nitidum M.Martens & Galeotti SB 1205* Cestrum nocturnum L. SB 638* Cestrum oblongifolium Schltdl. SB 564* Cestrum thyrsoideum Kunth SB 1343* Cestrum tomentosum L.f. SB 317* Jaltomata procumbens (Cav.) J.L.Gentry SB 1065* Lycianthes lenta (Cav.) Bitter SB 462* Solanum aligerum Schltdl. SB 1178* Solanum appendiculatum Dunal SB 634* Solanum nigrescens M.Martens & Galeotti SB 551* Solanum nigricans M.Martens & Galeotti SB 1189* Solanum stoloniferum Schltdl. & Bouché SB 943* Staphyleaceae Martinov Turpinia aff. paniculata Vent. SB 1376* Styracaceae DC. & Spreng. Styrax ramirezii Greenm. SB 135* Symplocaceae Desf. Symplocos citrea Lex. ex La Llave & Lex. SB 252* Verbenaceae J.St.-Hil. Lantana camara L. SB 873 Lantana velutina M.Martens & Galeotti SB 548 Lippia mexicana Grieve SB 678 Violaceae Batsch Viola aff. guatemalensis W. Becker SB 1194 !"#$%&'$()*'&+%'+,'34'5467'!839!:;<'8;=" SU UO LO OC TE X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X !"# $%&'$()*+,&-./0,'+1,2.34%,'5,6%'7% Appendix 2. Continuation. CH Vitaceae Juss. Vitis aff. tiliifolia Humb. & Bonpl. ex Roem & Schult. SB 620* ANGIOSPERMAE: LILIOPSIDA Alstroemeriaceae Dumort. Bomarea edulis (Tussac) Herb. SB 307 Amaryllidaceae J.St.-Hil. Hymenocallis aff. glauca (Zucc.) M.Roem. SB 724* Manfreda pringlei Rose SB 885* Manfreda pubescens (Regel & Ortgies) Verh.-Will. ex Pina SB 1284* Araceae Juss. Syngonium neglectum Schott SB 758 Asparagaceae Juss. Agave aff. angustifolia Haw. SB 341 Agave horrida Lem. ex Jacobi SB 347* Agave aff. inaequidens K.Koch SB 1017 Echeandia mexicana Cruden SB 920 Echeandia sp1. SB 659* Bromeliaceae Juss. Hechtia podantha Mez SB 354* Pitcairnia cf. heterophylla (Lindl.) Beer SB 352* Tillandsia cf. bourgaei Baker SB 336* Commelinaceae Mirb. Commelina cf. leiocarpa Benth. SB 712 Commelina tuberosa L. SB 686 Tinantia erecta (Jacq.) Schltdl. SB 765 Tradescantia poelliae D.R.Hunt SB 651 Cyperaceae Juss. Cyperus seslerioides Kunth SB 660 Cyperus hermaphroditus (Jacq.) Standl. SB 1260 Cyperus sp1. SB 497* !"# X SU X UO X LO OC TE X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X !"#$%&'$()*'&+%'+,,89,:9;<,!=8>!?@A,=@B" $%&'($)*+,+'-.'*/',*0$1,*2$'.%,-$.%/'0%)& Appendix 2. Continuation. CH Dioscoreaceae R.Br. Dioscorea galeottiana Kunth SB 752 Dioscorea cf. urceolata Uline SB 1031 Hypoxidaceae R.Br. Hypoxis sp1. SB 349* Iridaceae Juss. Sisyrinchium angustissimum (B.L.Rob. & Greenm.) Greenm. & C.H.Thomps. SB 388 Orchidaceae Juss. Aulosepalum aff. pyramidale (Lindl.) M.A.Dix & M.W.Dix SB 245* Bletia sp1. SB 1029* Epidendrum matudae L.O.Williams SB 330 Govenia sp1. SB 260* Malaxis fastigiata (Rchb.f.) Kuntze SB 936 Malaxis rosilloi R.González & E.W.Greenw. SB 768 Sarcoglottis sp1. SB 139 Poaceae Banhart Brachypodium mexicanum (Roem. & Schult.) Link SB 1200 Briza minor L. SB 395 Bromus carinatus Hook. & Arn. SB 448 Bromus dolichocarpus Wagnon SB 1051 Lasiacis nigra Davidse SB 1276 Lasiacis procerrima (Hack.) Hitchc. SB 1278 Melinis repens (Willd.) Zizka SB 1291a Muhlenbergia cenchroides (Humb. & Bonpl. ex Willd.) P.M.Peterson SB 1014 Muhlenbergia diversiglumis Trin. SB 35 Muhlenbergia robusta (E.Fourn.) Hitchc. SB 1291 Muhlenbergia versicolor Swallen SB 1325 Muhlenbergia virletii (E.Fourn.) Soderstr. SB 1015 !"#$%&'$()*'&+%'+,'#3'4356'!7#8!9:;'7:<" SU UO LO OC TE X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X X !"# $%&'$()*+,&-./0,'+1,2.34%,'5,6%'7 % Appendix 2. Continuation. CH Oplismenus burmannii (Retz.) P.Beauv. var. burmannii SB 62 Oplismenus hirtellus (L.) P.Beauv. SB 893 Paspalum prostratum Scribn. & Merr. SB 27* Paspalum squamulatum E.Fourn. SB 119 Piptochaetium virescens (Kunth) Parodi SB 447 Vulpia myuros (L.) C.C.Gmel. SB 407 Zuloagaea bulbosa (Kunth) Bess SB 1145* Smilacaceae Vent. Smilax aff. moranensis M.Martens & Galeotti SB 619* !"# SU UO LO OC X TE X X X X X X X X X X X !"#$%&'$()*'&+%'+,,89,:9;<,!=8>!"#?,=#@A