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Plant Diversity Comparison between natural Bamboo (Sinarundinaria alpina) and Exotic Tree Plantations (Cupressus lusitanica) in South Kinangop Area of the Aberdare Range Report for The Bamboo Trading Company www.thebambootradingcompany.com JUNE 2012 Impatiens fischeri W.R.Q. Luke & P.A. Luke P.O.Box 24133 Nairobi 00502 quentin.luke@swiftkenya.com Plant Diversity of Bamboo versus Cupressus Plantations in South Kinangop W.R.Q. Luke & P.A. Luke 2 1. Background The Bamboo Trading Company (BTC) requested a botanical study of the Southern Kinangop Forest Reserve as part of their feasibility study and Environmental and Social Impact Assessment (ESIA) for a proposed energy project. This study is restricted to a rapid comparison of the diversity of associated plant species found within the indigenous bamboo zone versus that found within the Kenya Forest Service (KFS) plantations. Bamboo and Erica arborea at GPS pt 035 The area of bamboo in the entire Aberdare range was reported as 647 km2 (Wimbush, 1945) or approximately 20% of the total reported for Kenya (O’Meara, 2012). No figure can be found for the area of the plantations within the Forest Reserve (FR) but a total for the whole country in 1991 was estimated as 1600 km2 (Wass, 1995) and more recently 1350 km2 (KFS call for tenders, Standard Newspaper, 14th June 2012). Due to a long period of closure of the plantations under a ‘logging ban’ the condition of these areas has deteriorated from a forestry perspective, but possibly improved in terms of natural biodiversity. The ban on harvesting was also applied to bamboo poles and it is the recent adoption of a new Bamboo Policy by KFS that has lead to the BTC proposal. Thus the bamboo areas have had little disturbance over several years prior to this study. 2. Methodology An exploratory visit was made to the south Kinangop on 1 st June with BTC’s Bamboo expert and the team setting out bamboo plots to measure biomass. Due to the limited time given for the diversity study (2 days) it was decided to adopt a simple walking line transect and to do three transects, one ‘bamboo’ below Sasumua Dam (pts 038-040, see Google Map), another ‘bamboo’ above the forest station at the end of the road (pts 041044) and one ‘plantation’ above the forest station (pts 045-047). The survey was carried out between 7th to 9th June by QL and PL using their own vehicle and camping at pt 035 for 2 nights. Plant Diversity of Bamboo versus Cupressus Plantations in South Kinangop W.R.Q. Luke & P.A. Luke 3 Tracks 1st June to 9th June inclusive No decision was made on the length of the transect until the recording began and it was found that by walking along the bamboo boundary with the road, the number of new species recorded dropped off after 200m. Thus for all 3 surveys a point was chosen on the road, marked by GPS, 200m was walked along the road recording all species encountered, another GPS point was recorded, a line was then walked at right angles into the bamboo/plantation for a further 200m recording addition species and then an end point was marked. No prior investigation of the composition was made, thus on the first survey the line into the bamboo moved into a patch of mixed montane forest and the third survey, the Cupressus plantation, found that after 160m the mature plantation ended and the walk at right angles after 200m (pt 046) was outside in new plantings of Cupressus lusitanica with open Pennisetum clandestinum grass in between. This grass was manifestly heavily grazed by livestock and was much more diverse than if the line had run within the completely shaded mature plantation. The plant species were recorded as “sight records” (SR) when there was no doubt over their identification. If the plant was known to QL, but needed confirmation, a temporary specimen was collected for later keying using UKWF2 (Agnew & Agnew, 1994). When the plant’s identity needed checking, a voucher specimen was collected (HS) for drying and later identification in the Nairobi Herbarium (Beentje, 1994; FTEA, 1952-; Ibrahim & Kabuye, 1988; Haines & Lye, 1983). 3. Results The list of plant species recorded is shown in the Appendix. A summary of the transects and the number of taxa recorded is as follows: Plant Diversity of Bamboo versus Cupressus Plantations in South Kinangop W.R.Q. Luke & P.A. Luke Survey No. GPS Points Direction Altitude (m) 1 Bamboo 1 Bamboo 038 - 039 039 - 040 SSE SW 2480 – 2475 2475 – 2460 No. of Plant Taxa 60 44 2 Bamboo 2 Bamboo 3 Plantation 3 Plantation 041 - 043 043 - 044 045 - 046 046 - 047 E N WSW NNW 2780 – 2770 2770 – 2800 2700 – 2680 2680 – 2700 57 8 71 19 Total per site 104 65 90 4 Comment Downhill, afternoon shade Downhill leading to montane forest, medium shade Almost level, much sun Uphill, full shade Downhill, little morning sun Uphill, young plantation full sun 4. Discussion The total number of plant taxa recorded over the 3 surveys is 160. Of these 22 are recorded from all 3 sites, 55 from 2 of the 3 sites and 84 from just one site. The expected large difference in diversity between the natural bamboo areas and the plantation was not found, in fact the plantation site proved to have more species than the second bamboo site. This is understandable when it is realised that the second ‘leg’ of plantation survey was not through mature plantation but through open grassland with young trees. The extra diversity encountered at the first site (104) can be explained by the fact that the second part of the second ‘leg’ ran into a patch of montane forest and thus a different group of species. A probable average number of taxa for pure bamboo sites would be around 80 taxa and that of mature, closely planted plantation around 70. This does not seem to be a huge difference and could be attributable to factors such as altitude, soil, sunlight (orientation), and predation. Only more complex ecological surveys would adjust for these factors. Studies on the ‘bamboo cycle’ (Wimbush, 1945; Agnew 1985) show that there is a succession process following the death of bamboo after flowering and thus the age of a bamboo area in terms of how far through its cycle will influence the composition of associated species. Asteraceae: Helichrysum argyranthum & Berkheya spekeana Species composition is not very surprising with a majority of ‘weedy’ or ruderal plants, particularly those in the ‘Daisy’ family Asteraceae with a total of 28 taxa noted. A number of sedges (Cyperaceae) and grasses (Poaceae) were recorded as well as several ferns. Also well represented were the families Acanthaceae and Lamiaceae. One way of interpreting the different compositions is to look at the rarity of individual species. Although obviously no figures exist for the overall total numbers of each taxon, Plant Diversity of Bamboo versus Cupressus Plantations in South Kinangop W.R.Q. Luke & P.A. Luke 5 their distributions are fairly well known. We then find that 3 of the plants recorded in the bamboo areas have very limited distributions: Impatiens hoehnelii is a Kenyan endemic; Impatiens fischeri (cover photo, not recorded within transects but at pt 039) is only known from the Aberdares and Mt Kenya; the large clumped grass (Eragrostis amanda) recorded in the first bamboo area is restricted to the Aberdares and Kieni (Ibrahim & Kabuye, 1988). Oreoschimperella aberdarensis is also endemic and only known from 4 areas in Kenya (FTEA, 1952-). This plant was not recorded during the survey but is known from the area. Only one plant found in the plantation area has similar limited distribution, Rubus friesiorum is restricted to the Aberdares and Mt Kenya (FTEA, 1952-). If a more extensive survey and comparison were made, it is most likely that the bamboo zone species would be found to be more restricted in range and could be considered of more ‘value’ than the more widely distributed plants growing in the plantations. Cyperaceae: Cyperus tomaiophyllus and Carex cf. conferta. Lamiaceae: Plectranthus alpinus & Stachys aculeolata Plant Diversity of Bamboo versus Cupressus Plantations in South Kinangop W.R.Q. Luke & P.A. Luke 6 5. Conclusion Although the number of different plants (diversity) found in the two habitats is not hugely different, perhaps the ‘quality’ of plants in the bamboo zone is higher due to their more restricted range. This study provides an initial assessment of the issue and it is recommended that more comprehensive sampling is undertaken to provide a clearer understanding of the issue. The presence of the noxious, yellow flowered weed Senecio madagascariensis (Fireweed) was noted throughout the area and every effort should be made to remove it. 6. References Agnew, A.D.Q. (1985) Cyclic sequences of vegetation in the plant communities of the Aberdare Mountains, Kenya. Journal of the East Africa Natural History Society and National Museum, 75, 1-12. Agnew, A.D.Q. & S. Agnew (1994). Upland Kenya Wild Flowers: a Flora of the Ferns and Herbaceous Flowering Plants of Upland Kenya. 2nd edition. East African Natural History Society, Nairobi. Beentje, H.J. (1994). Kenya Trees, Shrubs and Lianas. National Museums of Kenya, Nairobi. Flora of Tropical East Africa (FTEA). (1952–). Authors Various. Crown Agents for Overseas Governments and Administration, London; Balkema, Rotterdam & Royal Botanic Gardens, Kew. Haines, R.W. & K.A. Lye (1983). The Sedges and Rushes of East Africa. East Africa Natural History Society, Nairobi. Ibrahim, K.M. & C.H.S. Kabuye (1988). An Illustrated Manual of Kenya Grasses. FAO, Rome. O’Meara L. (2012). Energy Proposal. Unpublished Report, The Bamboo Trading Company. Nairobi Schmitt, K. (1991). The Vegetation of the Aberdare Nation Park Kenya. Univ.-Verl. Wagner. Innsbruck. Wass P (Ed.) (1995) Kenya’s Indigenous Forests: Status, Management and Conservation. IUCN, Gland, Switzerland, and Cambridge, UK. Wimbush, S.H. (1945). The African Alpine Bamboo. Empire Forestry Journal 24:33-39. Plant Diversity of Bamboo versus Cupressus Plantations in South Kinangop W.R.Q. Luke & P.A. Luke 7 Appendix cat. No. Bamboo Site 1 family species Comment Oldenlandia monanthos (A. Rich.) Hiern Clinopodium uhligii (Guerke) Ryding var. obtusifolium (Avetta) Ryding Platostoma africanum P. Beauv. Spermacoce princeae (K. Schum.) Verdc. var. princeae Selaginella kraussiana (Kunze) A. Braun Alchemilla cryptantha A. Rich. H H Geranium kilimandscharicum Engl. Pavonia urens Cav. var. urens H H Oenanthe procumbens (H. Wolff) Norman Acacia melanoxylon R. Br. H E Plectranthus laxiflorus Benth. Cyphostemma kilimandscharicum (Gilg) Wild & R.B. Drumm. var. kilimandscharicum Droguetia iners (Forssk.) Schweinf. ssp. iners Cyperus derreilema Steud. Oxalis corniculata L. H C SR SR 1 2 Rubiaceae Lamiaceae SR SR 3 4 Lamiaceae Rubiaceae SR SR 5 6 Selaginellaceae Rosaceae SR SR 7 8 Geraniaceae Malvaceae HS SR 15345 10 SR SR 11 12 Apiaceae Mimosaceae (Leguminosae) Lamiaceae Vitaceae SR SR SR 13 14 15 Urticaceae Cyperaceae Oxalidaceae SR 16 Tacazzea conferta N.E. Br. C SR SR SR SR 17 18 19 20 Asclepiadaceae (Apocynaceae) Violaceae Plantaginaceae Asteraceae Asteraceae Viola abyssinica Oliv. Plantago palmata Hook.f. Adenostemma mauritianum DC. Helichrysum globosum A. Rich. H H H H SR SR 21 22 Juniperus procera Endl. Desmodium repandum (Vahl) DC. T H SR 23 Cupressaceae Papilionaceae (Leguminosae) Rosaceae C SR SR 24 25 Scrophulariaceae Asteraceae Rubus steudneri Schweinf. var. dictyophyllus (Oliv.) R.A. Graham Halleria lucida L. Helichrysum meyeri-johannis Engl. SR SR SR SR SR SR 26 27 28 29 30 31 Ericaceae Cornaceae Polygalaceae Cyperaceae Iridaceae Lobeliaceae H T H H H H SR SR SR SR SR 32 33 34 35 36 Amaranthaceae Loganiaceae Poaceae Poaceae Asteraceae Erica silvatica (Engl.) Beentje Afrocrania volkensii (Harms) Hutch. Polygala sphenoptera Fresen. Carex chlorosaccus C.B. Clarke Aristea alata Baker Monopsis stellarioides (C. Presl) Urb. ssp. schimperiana (Urb.) Thulin Cyathula polycephala Baker Nuxia congesta Fresen. Panicum calvum Stapf? Eragrostis schweinfurthii Chiov.? Solanecio mannii (Hook.f.) C. Jeffrey SR SR 37 38 Ericaceae Asteraceae Agarista salicifolia (Lam.) G. Don Sonchus afromontanus R.E. Fr. T H H H F H H H H S H photo H T H H S Plant Diversity of Bamboo versus Cupressus Plantations in South Kinangop W.R.Q. Luke & P.A. Luke SR 39 Asteraceae SR 40 Convolvulaceae SR 41 Asteraceae SR SR SR SR SR SR 42 43 44 45 46 47 Commelinaceae Clusiaceae Cyperaceae Scrophulariaceae Rubiaceae Ranunculaceae HS 15346 SR SR SR SR SR 49 50 51 52 53 Rosaceae Acanthaceae Rubiaceae Berberidaceae Tiliaceae SR SR SR SR SR 54 55 56 57 58 SR SR SR SR 59 60 61 62 Cucurbitaceae Convolvulaceae Acanthaceae Adiantaceae Papilionaceae (Leguminosae) Asteraceae Asteraceae Aspleniaceae Rubiaceae SR SR 63 64 Podocarpaceae Ranunculaceae SR 65 Acanthaceae SR HS 66 15347 Urticaceae Cyperaceae SR SR SR SR SR SR SR SR 68 69 70 71 72 73 74 75 Melianthaceae Asteraceae Rubiaceae Acanthaceae Acanthaceae Euphorbiaceae Solanaceae Menispermaceae SR SR 76 77 SR 78 Salicaceae Papilionaceae (Leguminosae) Euphorbiaceae SR 79 Solanaceae Cyperaceae 8 Conyza subscaposa O. Hoffm. H Dichondra repens J.R. Forst. & G. Forst. var. repens Helichrysum forskahlii (J.F. Gmel.) Hilliard & B.L. Burtt var. forskahlii indet. Hypericum revolutum Vahl Pycreus nigricans (Steud.) C.B. Clarke Veronica abyssinica Fresen. Galium aparinoides Forssk. Clematis simensis Fresen. H Kyllinga odorata Vahl var. major (C.B. Clark) Chiov. Alchemilla fischeri Engl. Justicia diclipteroides Lindau Galiniera saxifraga (Hochst.) Bridson Berberis holstii Engl. Sparrmannia ricinocarpa (Eckl. & Zeyh.) Kuntze var. macrocarpa (Ulbr.) Weim. Peponium vogelii (Hook.f.) Engl. Convolvulus kilimandschari Engl. Mimulopsis alpina Chiov.? Pellaea quadripinnata (Forssk.) Prantl Indigofera atriceps Hook.f. ssp. atriceps H Laggera brevipes Oliv. & Hiern Conyza bonariensis (L.) Cronquist Asplenium buettneri Hieron. var. buettneri Rubia cordifolia L. ssp. conotricha (Gandoger) Verdc. Podocarpus latifolius (Thunb.) Mirb. Thalictrum rhynchocarpum Quart.-Dill. & A. Rich. Phaulopsis imbricata (Forssk.) Sweet ssp. imbricata Droguetia debilis Rendle Carex vallisrosetto K. Schum. H H F C Bersama abyssinica Fresen. ssp. abyssinica Senecio syringifolius O. Hoffm. Psychotria fractinervata E.M.A.Petit Acanthopale pubescens (Engl.) C.B. Clarke Isoglossa gregorii (S. Moore) Lindau Clutia robusta Pax Solanum nigriviolaceum Bitter Stephania abyssinica (Quart.-Dill. & A. Rich.) Walp. var. abyssinica Dovyalis abyssinica (A. Rich.) Warb. Parochetus africanus Polhill T C S S photo H H H C Euphorbia ugandensis Pax S Solanum terminale Forssk. C H H (sterile) S H H H C H H T T H C C S F H T H H H H T H Plant Diversity of Bamboo versus Cupressus Plantations in South Kinangop W.R.Q. Luke & P.A. Luke SR 80 SR 81 Papilionaceae (Leguminosae) Asteraceae SR 82 SR SR HS 83 84 15348 SR SR SR SR 9 Crotalaria axillaris Aiton H H Rhamnaceae Crassocephalum montuosum (S. Moore) Milne-Redh. Rhamnus prinoides L'Hér. Aspleniaceae Meliaceae Poaceae Asplenium friesiorum C. Chr. Lepidotrichilia volkensii (Guerke) J.-F.Leroy Eragrostis amanda Clayton 86 87 88 89 Rutaceae Lamiaceae Dennstaedtiaceae Celastraceae SR SR SR SR SR SR 90 91 92 93 94 95 Icacinaceae Meliaceae Oleaceae Apiaceae Apiaceae Polypodiaceae Toddalia asiatica (L.) Lam. Plectranthus melleri Baker Pteridium aquilinum (L.) Kuhn ssp. aquilinum Gymnosporia heterophylla (Eckl. & Zeyh.) Loes. Apodytes dimidiata Arn. var. dimidiata Ekebergia capensis Sparrm. Jasminum fluminense Vell.? Peucedanum aculeolatum Engl. Hydrocotyle mannii Hook.f. var. mannii Lepisorus excavatus (Willd.) Ching F T H (narrow endemic) C H F S SR SR SR 96 97 98 Asparagaceae Urticaceae Balsaminaceae Asparagus africanus Lam. Laportea alatipes Hook.f. Impatiens hoehnelii T.C.E. Fr. SR SR 99 100 Myrsinaceae Asteraceae SR 101 Lycopodiaceae Myrsine africana L. Dichrocephala integrifolia (L.f.) Kuntze ssp. integrifolia Lycopodium clavatum L. ssp. clavatum SR SR SR 102 103 104 Rhizophoraceae Cucurbitaceae Solanaceae Cassipourea malosana (Baker) Alston Momordica friesiorum (Harms) C. Jeffrey? Solanum nigrum L. s.l. T C H Cestrum elegans (Neumann) Schltdl. S (exotic) C S T T C H H F (lobed fronds) C H H (limited distribution) S H F Sasumua Dam HS 15349 Solanaceae Bamboo Site 2 SR 105 Rosaceae SR SR 106 107 Violaceae Menispermaceae SR SR 108 109 Cyperaceae Apiaceae Rubus steudneri Schweinf. var. dictyophyllus (Oliv.) R.A. Graham Viola abyssinica Oliv. Stephania abyssinica (Quart.-Dill. & A. Rich.) Walp. var. abyssinica Cyperus tomaiophyllus K. Schum. Agrocharis melanantha Hochst. SR SR 110 111 Acanthaceae Lamiaceae Mimulopsis alpina Chiov.? Plectranthus laxiflorus Benth. S H SR SR 112 113 Asteraceae Asteraceae Conyza bonariensis (L.) Cronquist Senecio syringifolius O. Hoffm. H C SR SR 114 115 Ericaceae Cyperaceae Erica arborea L. Carex vallisrosetto K. Schum. T H HS 15350 Asteraceae H HS 15351 Cyperaceae Helichrysum forskahlii (J.F. Gmel.) Hilliard & B.L. Burtt var. forskahlii Cyperus derreilema Steud. H photo C H photo H H photo Plant Diversity of Bamboo versus Cupressus Plantations in South Kinangop W.R.Q. Luke & P.A. Luke SR 118 Aspleniaceae SR SR 119 120 Lycopodiaceae Cyperaceae SR SR SR SR HS SR SR SR SR SR 121 122 123 124 15352 126 127 128 129 130 Berberidaceae Asteraceae Commelinaceae Balsaminaceae Dryopteridaceae Ranunculaceae Aspleniaceae Podocarpaceae Lamiaceae Asteraceae SR SR HS SR HS SR SR SR SR SR 131 132 15353 134 15354 136 137 138 139 140 Thelypteridaceae Pteridaceae Pteridaceae Boraginaceae Asteraceae Geraniaceae Rosaceae Cyperaceae Asteraceae Vitaceae SR 141 Tiliaceae SR SR SR SR 142 143 144 145 Rubiaceae Euphorbiaceae Loganiaceae Lamiaceae SR SR SR SR SR SR SR SR SR HS HS SR 146 147 148 149 150 151 152 153 154 15355 15356 157 Convolvulaceae Asteraceae Lobeliaceae Rubiaceae Iridaceae Clusiaceae Polygonaceae Solanaceae Asteraceae Asteraceae Asteraceae Rubiaceae SR 158 Selaginellaceae SR SR HS 159 160 15357 Lamiaceae Apiaceae Asteraceae 10 Asplenium buettneri Hieron. var. buettneri F Lycopodium clavatum L. ssp. clavatum Kyllinga odorata Vahl var. major (C.B. Clark) Chiov. Berberis holstii Engl. Berkheya spekeana Oliv. indet. Impatiens hoehnelii T.C.E. Fr. Dryopteris? Clematis simensis Fresen. Asplenium friesiorum C. Chr. Podocarpus latifolius (Thunb.) Mirb. Plectranthus alpinus (Vatke) Ryding Dichrocephala integrifolia (L.f.) Kuntze ssp. integrifolia indet. Pteris catoptera Kunze var. catoptera Pteris buchananii? Cynoglossum lanceolatum Forssk. Conyza newii Oliv. & Hiern Geranium kilimandscharicum Engl. Alchemilla fischeri Engl. Pycreus nigricans (Steud.) C.B. Clarke Carduus sp. Cyphostemma kilimandscharicum (Gilg) Wild & R.B. Drumm. var. kilimandscharicum Sparrmannia ricinocarpa (Eckl. & Zeyh.) Kuntze var. macrocarpa (Ulbr.) Weim. Galium aparinoides Forssk. Euphorbia ugandensis Pax Nuxia congesta Fresen. Clinopodium uhligii (Guerke) Ryding var. obtusifolium (Avetta) Ryding Convolvulus kilimandschari Engl. Senecio moorei R.E. Fr. Lobelia sp. nr minutula Oldenlandia monanthos (A. Rich.) Hiern Aristea alata Baker Hypericum revolutum Vahl Rumex ruwenzoriensis Chiov. Solanum phoxocarpum Voronts. Sonchus sp. Senecio madagascariensis Poir. Bothriocline fusca (S. Moore) M.G. Gilbert Spermacoce princeae (K. Schum.) Verdc. var. princeae Selaginella kraussiana (Kunze) A. Braun F H Stachys aculeolata Hook.f. var. aculeolata Pseudocarum eminii (Engl.) H. Wolff Crassocephalum montuosum (S. Moore) H photo C H T H H (sterile) H photo F C F T H photo H F (sterile) F F H H H H H H C H H S T H C H H H H S H T H H Invasive S photo H F Plant Diversity of Bamboo versus Cupressus Plantations in South Kinangop W.R.Q. Luke & P.A. Luke 11 Milne-Redh. SR SR 162 163 Polypodiaceae Aspleniaceae Lepisorus excavatus (Willd.) Ching Asplenium sp. F F SR SR SR SR 164 165 166 167 Hydrocotyle mannii Hook.f. var. mannii Afrocrania volkensii (Harms) Hutch. Lagenaria sphaerica (Sond.) Naudin Tacazzea conferta N.E. Br. H T C C SR SR 168 169 Apiaceae Cornaceae Cucurbitaceae Asclepiadaceae (Apocynaceae) Araceae Urticaceae Arisaema mildbraedii Engl. Laportea alatipes Hook.f. H H Dichondra repens J.R. Forst. & G. Forst. var. repens Cyathula polycephala Baker Dichrocephala integrifolia (L.f.) Kuntze ssp. integrifolia Asplenium buettneri Hieron. var. buettneri Lepisorus excavatus (Willd.) Ching Acacia melanoxylon R. Br. H Rubus steudneri Schweinf. var. dictyophyllus (Oliv.) R.A. Graham Parochetus africanus Polhill C Helichrysum schimperi (A. Rich.) Moeser Bothriocline fusca (S. Moore) M.G. Gilbert Agrocharis melanantha Hochst. Podocarpus latifolius (Thunb.) Mirb. Clematis simensis Fresen. Agarista salicifolia (Lam.) G. Don Dryopteris? Trifolium semipilosum Fresen. H S H T C T F H Monopsis stellarioides (C. Presl) Urb. ssp. schimperiana (Urb.) Thulin Viola abyssinica Oliv. Clinopodium uhligii (Guerke) Ryding var. obtusifolium (Avetta) Ryding Thalictrum rhynchocarpum Quart.-Dill. & A. Rich. Pteridium aquilinum (L.) Kuhn ssp. aquilinum Kalanchoe densiflora Rolfe var. densiflora Helichrysum forskahlii (J.F. Gmel.) Hilliard & B.L. Burtt var. forskahlii Launaea sp. Hypericum revolutum Vahl Justicia diclipteroides Lindau Nuxia congesta Fresen. Conyza bonariensis (L.) Cronquist Droguetia iners (Forssk.) Schweinf. ssp. iners Polystichum wilsonii H. Christ? Bersama abyssinica Fresen. ssp. abyssinica H Plantation Site SR 170 Convolvulaceae SR SR 171 172 Amaranthaceae Asteraceae SR SR SR 173 174 175 SR 176 Aspleniaceae Polypodiaceae Mimosaceae (Leguminosae) Rosaceae SR 177 SR SR SR SR SR SR SR SR 178 179 180 181 182 183 184 185 SR 186 Papilionaceae (Leguminosae) Asteraceae Asteraceae Apiaceae Podocarpaceae Ranunculaceae Ericaceae Dryopteridaceae Papilionaceae (Leguminosae) Lobeliaceae SR SR 187 188 Violaceae Lamiaceae SR 189 Ranunculaceae SR SR SR 190 191 192 Dennstaedtiaceae Crassulaceae Asteraceae SR SR SR SR SR SR SR SR 193 194 195 196 197 198 199 200 Asteraceae Clusiaceae Acanthaceae Loganiaceae Asteraceae Urticaceae Dryopteridaceae Melianthaceae H H F F T H H H H F H H H S H T H H F T Plant Diversity of Bamboo versus Cupressus Plantations in South Kinangop W.R.Q. Luke & P.A. Luke SR 201 Cornaceae SR SR 202 203 Myrsinaceae Asteraceae SR HS SR 204 15358 206 Rosaceae Boraginaceae Rosaceae SR 207 Tiliaceae SR SR SR SR 208 209 210 211 SR HS SR SR 212 15359 214 215 Lamiaceae Asteraceae Tiliaceae Papilionaceae (Leguminosae) Asteraceae Dryopteridaceae Malvaceae Vitaceae SR SR SR SR 216 217 218 219 HS SR SR SR 15360 221 222 223 Asteraceae Solanaceae Convolvulaceae Sterculiaceae SR SR SR 224 225 226 SR SR 227 228 Apiaceae Araceae Asclepiadaceae (Apocynaceae) Cucurbitaceae Asteraceae SR SR SR SR SR SR SR SR SR SR SR 229 230 231 232 233 234 235 236 237 238 239 HS SR SR 15361 241 242 Polygalaceae Asteraceae Poaceae Apiaceae Poaceae Asteraceae Balsaminaceae Solanaceae Rosaceae Pteridaceae Araliaceae Scrophulariaceae Plantaginaceae Solanaceae Papilionaceae (Leguminosae) Asteraceae Lamiaceae Cyperaceae 12 Afrocrania volkensii (Harms) Hutch. T Rapanea melanophloeos (L.) Mez Crassocephalum montuosum (S. Moore) Milne-Redh. Alchemilla fischeri Engl. Cynoglossum lanceolatum Forssk. Rubus friesiorum Gust. ssp. friesiorum T H Sparrmannia ricinocarpa (Eckl. & Zeyh.) Kuntze var. macrocarpa (Ulbr.) Weim. Pycnostachys meyeri Guerke Carduus sp. Triumfetta rhomboidea Jacq. Adenocarpus/Argyrolobium H H C (limited distribution) H H H H H Helichrysum stenopterum DC. Megalastrum? Pavonia urens Cav. var. urens Cyphostemma kilimandscharicum (Gilg) Wild & R.B. Drumm. var. kilimandscharicum Polygala sphenoptera Fresen. Helichrysum globosum A. Rich. Pennisetum clandestinum Chiov. Peucedanum aculeolatum Engl. H F H C Helichrysum foetidum (L.) Moench. Physalis peruviana L. Convolvulus kilimandschari Engl. Dombeya torrida (J.F. Gmel.) P.Bamps ssp. torrida Hydrocotyle mannii Hook.f. var. mannii Arisaema mildbraedii Engl. Tacazzea conferta N.E. Br. H photo H C T Zehneria scabra (L.f.) Sond. ssp. scabra Senecio syringifolius O. Hoffm. C C Panicum calvum Stapf? Conyza newii Oliv. & Hiern Impatiens pseudoviola Gilg Solanum phoxocarpum Voronts. Alchemilla cryptantha A. Rich. Pteris catoptera Kunze var. catoptera Schefflera volkensii (Engl.) Harms Veronica abyssinica Fresen. Plantago palmata Hook.f. Solanum nigriviolaceum Bitter Desmodium repandum (Vahl) DC. H H H photo T H F T H H H H Senecio moorei R.E. Fr. Leonotis nepetifolia (L.) R. Br. var. nepetifolia Isolepis sp. H photo H H H H H H H H C Plant Diversity of Bamboo versus Cupressus Plantations in South Kinangop W.R.Q. Luke & P.A. Luke SR 243 HS HS 13 Cyperaceae Cyperus rigidifolius Steud. H 15362 15363 Asteraceae Polygonaceae Conyza schimperi A. Rich. Rumex acetosella L. SR SR SR SR SR HS SR SR SR HS SR SR 246 247 248 249 250 15364 252 253 254 15365 256 257 Asteraceae Asteraceae Commelinaceae Oxalidaceae Ranunculaceae Asteraceae Geraniaceae Rosaceae Asteraceae Asteraceae Asteraceae Asteraceae H H Naturalised H H Invasive H H H H H C photo H photo H photo H photo H SR SR 258 259 Cyperaceae Meliaceae Laggera brevipes Oliv. & Hiern Senecio madagascariensis Poir. Commelina africana L. Oxalis corniculata L. Ranunculus multifidus Forssk. Conyza steudelii A. Rich. Geranium kilimandscharicum Engl. Rubus volkensii Engl. Berkheya spekeana Oliv. Helichrysum argyranthum O. Hoffm. Carduus schimperi Sch. Bip. Pseudognaphalium luteo-album (L.) Hilliard & B.L. Burtt Kyllinga alba Nees Ekebergia capensis Sparrm. Key: SR = Sight Record; HS = Herbarium Specimen F = Fern; H = Herb; S = Shrub; C = Climber; T = Tree H T