PhytoKeys 147: 1–191 (2020)
doi: 10.3897/phytokeys.147.49602
Checklist of coastal forests of Kenya
A peer-reviewed open-access journal
1
CHECKLIST
http://phytokeys.pensoft.net
Launched to accelerate biodiversity research
An annotated checklist of the coastal forests
of Kenya, East Africa
Veronicah Mutele Ngumbau1,2,3,4, Quentin Luke4, Mwadime Nyange4,
Vincent Okelo Wanga1,2,3, Benjamin Muema Watuma1,2,3,
Yuvenalis Morara Mbuni1,2,3,4, Jacinta Ndunge Munyao1,2,3,
Millicent Akinyi Oulo1,2,3, Elijah Mbandi Mkala1,2,3,
Solomon Kipkoech1,2,3, Malombe Itambo4,
Guang-Wan Hu1,2, Qing-Feng Wang1,2
1 CAS Key Laboratory of Plant Germplasm Enhancement and Specialty Agriculture, Wuhan Botanical Garden, Chinese Academy of Sciences, Wuhan 430074, Hubei, China 2 Sino-Africa Joint Research Center (SAJOREC), Chinese Academy of Sciences, Wuhan 430074, Hubei, China 3 University of Chinese Academy of
Sciences, Beijing 100049, China 4 East African Herbarium, National Museums of Kenya, P. O. Box 45166
00100, Nairobi, Kenya
Corresponding author: Guang-Wan Hu (guangwanhu@wbgcas.cn)
Academic editor: P. Herendeen | Received 23 December 2019 | Accepted 17 March 2020 | Published 12 May 2020
Citation: Ngumbau VM, Luke Q, Nyange M, Wanga VO, Watuma BM, Mbuni YuM, Munyao JN, Oulo MA, Mkala
EM, Kipkoech S, Itambo M, Hu G-W, Wang Q-F (2020) An annotated checklist of the coastal forests of Kenya, East
Africa. PhytoKeys 147: 1–191. https://doi.org/10.3897/phytokeys.147.49602
Abstract
The inadequacy of information impedes society’s competence to find out the cause or degree of a problem or even to avoid further losses in an ecosystem. It becomes even harder to identify all the biological resources at risk because there is no exhaustive inventory of either fauna or flora of a particular region. Coastal forests of Kenya are located in the southeast part of Kenya and are distributed mainly in
four counties: Kwale, Kilifi, Lamu, and Tana River County. They are a stretch of fragmented forests ca.
30−120 km away from the Indian Ocean, and they have existed for millions of years. Diversity of both
fauna and flora is very high in these relicts and the coastal forests of Eastern Africa, extending along the
coast from Somalia through Kenya and Tanzania to Mozambique, are ranked among the priority biodiversity hotspot in the world. In spite of the high plant species richness and their importance towards
supporting the livelihoods of the communities that live around them, floristic studies in these forests have
remained poorly investigated. Hence, based on numerous field investigations, plant lists from published
monograph/literature, and data from BRAHMS (Botanical Records and Herbarium Management System) database at East African herbarium (EA), we present a detailed checklist of vascular plants recorded
in this region. Our results show that Kenyan coastal forests play an essential role in the flora of Kenya and
Copyright Veronicah M. Ngumbau et al. This is an open access article distributed under the terms of the Creative Commons Attribution License
(CC BY 4.0), which permits unrestricted use, distribution, and reproduction in any medium, provided the original author and source are credited.
2
Veronicah M. Ngumbau et al. / PhytoKeys 147: 1–191 (2020)
the plant diversity of the coastal forests of East Africa. The checklist represents 176 families, 981 genera,
2489 species, 100 infraspecific taxa, 90 endemic plants species, 72 exotic species, and 120 species that are
included in the current IUCN Red List of Threatened Species as species of major concern. We also discovered three new species to the world from these relicts. Thus, Kenyan coastal forests present a remarkable
and significant center of plant diversity.
Keywords
Biodiversity, checklist, coastal forests, endemic, threatened species
Table of contents
Introduction ............................................................................................................. 2
Material and methods............................................................................................... 4
Study area .......................................................................................................... 4
Plant collection and nomenclature ..................................................................... 4
Results and conclusion ............................................................................................. 6
Species diversity ................................................................................................. 6
Growth forms .................................................................................................... 6
Species in need of conservation attention ........................................................... 7
Endemic species of Kenyan coastal forests .......................................................... 8
New Records .................................................................................................... 11
Exotic species ................................................................................................... 11
Conclusion....................................................................................................... 13
Checklist ................................................................................................................ 15
Part 1 Lycophytes ............................................................................................. 16
Part 2 Monilophytes......................................................................................... 16
Part 3 Gymnosperm ......................................................................................... 20
Part 4 Angiosperms .......................................................................................... 21
Acknowledgments ................................................................................................ 188
References ............................................................................................................ 189
Introduction
Africa is among the world’s major centers of endemism and species-rich biodiversity regions (Burgess and Clarke 2000; Linder 2001). It hosts several centers of diversity with
the eastern moist forests, which include coastal forests of East Africa and Eastern Arc
forests as one of the significant centers of diversity (Lovett 1998). Another important
center is the Congo Basin rainforest, which forms the second largest extent of continuous rainforest in the world (Linder 2014). Hamilton (1974, 1982) presented four centers of endemism in Africa, which include Kivu, the East African coast, Cameroon Ga-
Checklist of coastal forests of Kenya
3
bon area, and possibly small refugia in West Africa. Brenan (1978), working with data
arranged according to country, identified the same regions as being rich in endemism.
Coastal forests of Eastern Africa are ranked among the 35 identified world’s biodiversity hotspots due to the concentration of many endemic species and habitat loss
(Mittermeier et al. 1998, 2004; Olson and Dinerstein 1998). Together with the Eastern Arc mountains, they contain approximately 2000 endemic plant species (WWF–
US 2003b ). These forests have lived for millions of years, of which they existed as a
continuous belt of forest between the East and West coast referred to as the ancient
Pan-African forest. However, fluctuation of climate, during the last 2 million years,
caused their fragmentation, leading to loss of some lowland dry forests in Africa (Hawthorne et al. 1981). Today, these forests have lost considerable amounts of their wilderness due to anthropogenic pressures leading to small fragments up to less than 5 km2,
with the most extensive patches recorded in the Kenya coastal forests (Wass 1995;
Habel et al. 2017).
Loss of habitat due to human activities is the most severe threat to biodiversity
and has become a major global environmental problem (Collinge 1996), hence, plant
diversity protection has attracted more attention. Coastal forests of East Africa are
ranked among the world’s ten most threatened forest hotspots because they have lost
more than 90% of their original habitat. Similarly, Brooks et al. (2002) and Azeria
et al. (2007) pointed out that Northern Kenya and Southern Tanzania coastal forests
should be highly prioritized because they are in danger of losing most of their biodiversity in coming times.
Floristic studies, however, provide a basic outline for plant conservation. Based on
these studies, it is possible to determine the condition of an ecosystem, the primary relationships of species with each other or with the environment, and the identity of rare
species or widespread species (Ivanauskas et al. 2007). Therefore, they must be carried
out not only in a particular area but also over time (Mota et al. 2017). In spite the coastal forests of Kenya acting as significant reservoirs of carbon and biodiversity, and supporting the livelihoods of rural people, plant species diversity remains poorly studied.
Despite a substantial amount of floristic research having been performed over the last
decades on some of the Kenyan coastal relicts, (Robertson and Luke 1993; Lehmann
and Kioko 2005; Luke 2005), a comprehensive study of the whole coastal region of
Kenya is still extremely urgent due to its vast area and large number of threatened and
endemic taxa. In addition, these forests are vital in providing ecological services at
local, national, and global levels; it is therefore, crucial to understand their composition. Here we aim at presenting a detailed checklist of vascular plants recorded in the
whole coastal region of Kenya. We achieved this through numerous field investigations
carried out from 2015 to 2018, checking of plant species data from published monographs or literature (Beentje 1994; Luke 2005; Flora of Tropical East Africa (FTEA)
1952–2012), and obtaining data documented in the BRAHMS (Botanical Research
and Herbarium Management System) database at the East African Herbarium (EA).
We also document endemic and threatened species found in this region.
4
Veronicah M. Ngumbau et al. / PhytoKeys 147: 1–191 (2020)
Material and methods
Study area
The study was carried out in the Kenyan coastal region located in the southeast part
of Kenya (Fig. 1). Coastal forests are mainly distributed in four counties in the coastal
province of Kenya, lying within the latitude of 1°40' to about 4°40' south and between
0–ca. 840 m a.s.l. especially at around Kilibasi (Moomaw 1960; Robertson and Luke
1993). They are part of the larger coastal forests of East Africa which cover an area of
approximately 3170 km2 (Azeria et al. 2007; Wegner et al. 2009) of which, Kenyan
coastal forests cover about 787 km2 (Younge et al. 2002; Burgess et al. 2003). They
contain various forest patches with Arabuko Sokoke, and Shimba hills National Reserves forests being the largest remaining forests patches (Burgess et al. 1998, 2003;
Younge et al. 2002; Matiku et al. 2013). Younge et al. (2002), estimated a total area
covered by the coastal woodland/bushland to be 120 000 ha. Of this, 114 460 ha are
in national reserves, 50 790 ha is in forest reserves, 16 000 ha remains ungazetted,
and 10–200 ha are in the sacred Kaya Forests. The coastal region of Kenya supports
a mosaic of different vegetation types. Much of the area supports bushland or thicket
habitats, transition woodland, swampy forests, edaphic grasslands habitats, moist forests, and dry forests. The littoral vegetation includes mangrove vegetation along some
parts of the coast and shoreline vegetation (Githitho 2004).
Plant collection and nomenclature
Plant species data for the checklist of Kenya coastal forests were obtained from three
primary sources. Firstly, collection records collected during floristic surveys carried out
between 2015 and 2018. Different forest patches were visited and surveyed at different
seasons to mainly cover the area where there had been few or no previous collections.
The investigations included random walks while collecting fertile plant specimens.
Details of collected specimens were recorded in a notebook. Information such as habit,
habitat, location, and collector details were recorded as well. All collected materials
were then processed (pressed and dried) and later deposited at the East Africa herbarium (EA). Duplicates were stored at Wuhan Botanical Garden (HIB). Secondly, plant
species data were obtained from the BRAHMS (Botanical Research and Herbarium
Management System) database in the East Africa herbarium (EA). Lastly, plant species data for coastal forests of Kenya were assembled from monographs and published
literature (Beentje 1994; Luke 2005; FTEA 1952–2012).
Further information about the species recorded at the coastal forests was checked
on specimens found in the Global Biodiversity Information Facility (GBIF) (www.
gbif.org). Identification of collected specimens was made using taxonomic monographs/floras (FTEA) also through checking and comparing with identified specimens
Checklist of coastal forests of Kenya
5
Figure 1. A map showing the study area and distribution of forest patches along the Kenyan coastline.
6
Veronicah M. Ngumbau et al. / PhytoKeys 147: 1–191 (2020)
at the East African Herbarium (EA). Family and species circumscription, as well as
spelling, authorities, and synonyms of scientific names, were updated based on online databases such as African plant database (www.villege.ch/musinfo/bd/cjb/africa),
Catalogue of life (Roskov et al. 2019), and Tropicos (www.tropicos.org). Finally, we
identified threatened and near threatened species based on the International Union for
Conservation of Nature (IUCN) Red List of Threatened Species (IUCN 2019).
Results and conclusion
Species diversity
A total of 2489 vascular plant species and 100 infraspecific taxa (subspecies and
varieties) belonging to 981 genera and 176 families are comprehensively listed and
presented here. This represents 39.55%, 63.78%, and 78.67%, respectively, of the
total number of taxa found in Kenya flora (FTEA 1952–2012). Also, it represents
more than half the total number of plant species found at the coastal forests of East
Africa (WWF–US 2003b). 120, 89, and 72 entries represent threatened and near
threatened, endemic, and exotic species of the coastal forests of Kenya, respectively
(Tables 5, 7, 9). Three new species were discovered and described from these relicts;
Adenia angulosa Passifloraceae (Ngumbau et al. 2017) recorded from the coastal
forests of Kenya and Tanzania, Zehneria monocarpa, Curcubitaceae (Ngumbau et al.
unpublished) from Kenyan coastal relicts, and Croton kinondoensis, Euphorbiaceae,
(Ngumbau et al. in press). Morphology and ITS sequence of Croton kinondoensis
placed it under the Adenophorus group, which is known to be endemic to Madagascar
and Comoros Islands. This new entity represented an independent dispersal of Croton
from Madagascar to Africa mainland (Ngumbau et al. in press).
The top 5 most species-rich families for coastal forests of Kenya are Fabaceae
(226 species), Poaceae (207 species), Rubiaceae (137 species), Malvaceae (117 species), and Cyperaceae (111 species). The top 5 most species-rich genera are Cyperus
(48 species, Cyperaceae), Euphorbia (28 species, Euphorbiaceae), Justicia (27 species,
Acanthaceae), Ipomoea (25 species, Convolvulaceae), and Crotalaria (23 species, Fabaceae) (Table 1).
Growth forms
The growth forms (trees, shrubs, lianas, and herbs) have widely different mechanical
architectures, which can also vary phenotypically with the environment (Rowe and
Speck 2005). In the coastal forests of Kenya, herbs represent the highest percentage of
life form with 48.09%, followed by shrubs 19.96%, trees 15.79%, climbers 8.28%,
lianas, 6.07%, and epiphytes 1.81% (Table 2).
Checklist of coastal forests of Kenya
7
Table 1. The 10 largest families and genera of the vascular plants of Kenya coastal region.
Family
Fabaceae
Poaceae
Rubiaceae
Malvaceae
Cyperaceae
Acanthaceae
Apocynaceae
Euphorbiaceae
Asteraceae
Lamiaceae
Orchidaceae
Genus
80
72
63
30
18
30
53
26
49
18
23
Species
226
207
137
117
111
108
101
87
82
67
60
Genus
Cyperus
Euphorbia
Justicia
Ipomoea
Indigofera
Crotalaria
Berlaria
Ficus
Combretum
Hibiscus
Grewia
Species
48
28
27
25
24
23
19
19
19
18
17
Table 2. Growth habit of the plants of Kenya coastal region.
Habit
Trees
Shrubs
Lianas
Climbers
Herbs
Epiphytes
Total
No. of species
393
497
151
206
1197
45
2489
Percentage
15.79%
19.96%
6.07%
8.28%
48.09%
1.81%
100%
Table 3. IUCN category and number of species representing in each category.
IUCN category
Extinct in wild (EW)
Critically endangered (CR)
Endangered (EN)
Vulnerable (VU)
Near threatened (NT)
Total
No. of Species
1
11
30
52
26
120
Percentage
0.83%
9.17%
25%
43.33%
21.67%
100%
Species in need of conservation attention
The Red List and Red Data species system is an approach developed by the IUCN
for evaluating the conservation status of species, and in particular for identifying and
documenting species in need of conservation attention (IUCN 2019). According to
this system, 120 species from the coastal forests of Kenya (Table 5), belonging to 39
families and 98 genera were categorized as Extinct in the wild (EW), Critically Endangered (CR), Endangered (EN), Vulnerable (VU), and Near Threatened (NT). Of
120 species recorded, one taxon was recorded as extinct in the wild, 11 were critically
endangered, 20 endangered, 52 vulnerable, and 26 near threatened (Table 3). The top
three families which had most of its members threatened and near threatened were
8
Veronicah M. Ngumbau et al. / PhytoKeys 147: 1–191 (2020)
Table 4. Families with the largest to the smallest number of threatened and near threatened taxa and their
percentage representatives.
Family
Acanthaceae
Fabaceae
Annonaceae
Rubiaceae
Moraceae
Asphodelaceae
Loranthaceae
Araceae
Asteraceae
Malvaceae
Orchidaceae
Burseraceae
Combretaceae
Euphorbiaceae
Melastomataceae
Meliaceae
Putranjivaceae
Sapotaceae
Anacardiaceae
Apocynaceae
Burmanniaceae
Buxaceae
Canellaceae
Connaraceae
Crassulaceae
Ebenaceae
Gesneriaceae
Lamiaceae
Lythraceae
Menispermaceae
Montiniaceae
Myristicaceae
Picrodendraceae
Rhamnaceae
Rutaceae
Salicaceae
Sapindaceae
Zosteraceae
Totals
No. of species
21
17
14
9
5
4
4
3
3
3
3
2
2
2
2
2
2
2
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
120
Percentage (%)
17.5
14.17
11.67
7.5
4.17
3.33
3.33
2.5
2.5
2.5
2.5
1.67
1.67
1.67
1.67
1.67
1.67
1.67
0.83
0.83
0.83
0.83
0.83
0.83
0.83
0.83
0.83
0.83
0.83
0.83
0.83
0.83
0.83
0.83
0.83
0.83
0.83
0.83
100
Acanthaceae 21 species (17.5%), followed by Fabaceae 18 species (14.17%), and Annonaceae 14 species (11.67%) (Table 4).
Endemic species of Kenyan coastal forests
From our results, 551 species were endemic to coastal forests of East Africa. 185 plant
species are endemic to Kenya and Tanzania coastal forests and 89 species are only
Checklist of coastal forests of Kenya
9
Table 5. List of threatened and near threatened plant taxa found in Kenya coastal region.
Family
Crassulaceae
Acanthaceae
Annonaceae
Burmanniaceae
Combretaceae
Euphorbiaceae
Gesneriaceae
Lamiaceae
Loranthaceae
Malvaceae
Rubiaceae
Acanthaceae
Asteraceae
Asphodelaceae
Araceae
Ebenaceae
Fabaceae
Loranthaceae
Malvaceae
Montiniaceae
Orchidaceae
Rhamnaceae
Rubiaceae
Acanthaceae
Annonaceae
Species
Kalanchoe fadeniorum Raadts
Justicia drummondii Vollesen
Megalochlamys tanaensis Vollesen
Uvaria puguensis D.M. Johnson
Afrothismia baerae Cheek
Combretum tenuipetiolatum Wickens
Euphorbia tanaensis P.R.O. Bally
Streptocarpus ionanthus subsp. rupicola (B. L.
Burtt) Christenh.
Karomia gigas (Faden) Verdc.
Taxillus wiensii Polhill
Cola porphyrantha Brenan
Vangueriopsis shimbaensis A.P. Davis & Q. Luke
Asystasia linearis S. Moore
Barleria lukei I. Darbysh.
Barleria maculata S. Moore
Blepharis kenyensis Vollesen
Cephalophis lukei Vollesen
Barleria whytei S. Moore
Justicia faulknerae Vollesen
Justicia breviracemosa Vollesen
Monanthotaxis faulknerae Verdc.
Toussaintia orientalis Verdc.
Uvariodendron gorgonis Verdc.
Uvaria faulknerae Verdc.
Aspilia macrorrhiza Chiov.
Aloe kilifiensis Christian
Stylochaeton bogneri Mayo
Diospyros shimbaensis F. White
Newtonia erlangeri (Harms) Brenan
Crotalaria laburnoides var. nudicarpa Polhill
Cynometra lukei Beentje
Gigasiphon macrosiphon (Harms) Brenan
Bauhinia mombassae Vatke
Dalbergia gloveri Q. Luke, ined.
Englerina ramulosa (Sprague) Polhill & Wiens
Agelanthus microphyllus Polhill & Wiens
Cola octoloboides Brenan
Grevea eggelingii var. keniensis (Verdc.) Verdc.
Habenaria stylites Rchb.f. & S. Moore
Polystachya teitensis P.J. Cribb
Ziziphus robertsoniana Beentje
Multidentia sclerocarpa (K. Schum.) Bridson
Barleria usambarica Lindau
Justicia galeata Hedrén
Asystasia minutiflora Ensermu & Vollesen
Barleria maritima I. Darbysh.
Blepharis pratensis S. Moore
Dicliptera inconspicua I. Darbysh.
Anisotes galanae (Baden) Vollesen
Justicia brevipila Hedrén
Justicia heterotricha Mildbr.
Isolona cauliflora Verdc.
Uvariodendron kirkii Verdc.
Life form
Herb
Herb
Herb
Liana
Herb
Tree
Tree
Herb
Tree
Shrub
Tree
Tree
Herb
Herb
Herb
Herb
Herb
Herb
Shrub
Shrub
Liana
Tree
Tree
Liana
Herb
Herb
Herb
Tree
Tree
Herb
Tree
Tree
Shrub
Liana
Shrub
Shrub
Tree
Shrub
Herb
Epiphytic herb
Tree
Shrub
Herb
shrub
Herb
Herb
Herb
Herb
Shrub
Herb
Herb
Tree
Shrub
IUCN
Version
Extinct in the wild
3.1
Critically Endangered 3.1
Critically Endangered 3.1
Critically Endangered 3.1
Critically Endangered 3.1
Critically Endangered 2.3
Critically Endangered 3.1
Critically Endangered 3.1
Critically Endangered
Critically Endangered
Critically Endangered
Critically Endangered
Endangered
Endangered
Endangered
Endangered
Endangered
Endangered
Endangered
Endangered
Endangered
Endangered
Endangered
Endangered
Endangered
Endangered
Endangered
Endangered
Endangered
Endangered
Endangered
Endangered
Endangered
Endangered
Endangered
Endangered
Endangered
Endangered
Endangered
Endangered
Endangered
Endangered
Vulnerable
Vulnerable
Vulnerable
Vulnerable
Vulnerable
Vulnerable
Vulnerable
Vulnerable
Vulnerable
Vulnerable
Vulnerable
2.3
3.1
3.1
3.1
3.1
3.1
3.1
3.1
3.1
3.1
3.1
3.1
3.1
3.1
3.1
3.1
3.1
3.1
3.1
2.3
3.1
3.1
3.1
2.3
3.1
3.1
3.1
3.1
3.1
3.1
3.1
3.1
3.1
3.1
3.1
3.1
3.1
3.1
3.1
3.1
3.1
3.1
3.1
3.1
3.1
10
Family
Annonaceae
Veronicah M. Ngumbau et al. / PhytoKeys 147: 1–191 (2020)
Species
Xylopia arenaria Engl.
Mkilua fragrans Verdc.
Apocynaceae
Pleioceras orientale Vollesen
Asteraceae
Emilia bellioides (Chiov.) C. Jeffrey
Araceae
Gonatopus petiolulatus (Peter) Bogner
Asphodelaceae Aloe massawana Reynolds
Aloe ukambensis Reynolds
Buxaceae
Buxus obtusifolia (Mildbr.) Hutch.
Canellaceae
Warburgia stuhlmannii Engl.
Euphorbiaceae Pycnocoma littoralis Pax
Fabaceae
Newtonia paucijuga (Harms) Brenan
Sesbania speciosa Taub.
Angylocalyx braunii Harms
Cynometra webberi Baker f.
Dalbergia vacciniifolia Vatke
Diospyros greenwayi F. White
Prioria msoo (Harms) Breteler
Loranthaceae
Erianthemum alveatum (Sprague) Danser
Lythraceae
Ammannia parkeri var. longifolia (Verdc.) S.A.
Graham & Gandhi
Malvaceae
Hibiscus holstii Mwachala
Melastomataceae Memecylon verruculosum Brenan
Warneckea mouririifolia (Brenan) Borhidi
Menispermaceae Cissampelos nigrescens Diels var. nigrescens
Moraceae
Dorstenia tayloriana Rendle var. tayloriana
Ficus faulkneriana C.C. Berg
Myristicaceae
Cephalosphaera usambarensis (Warb.) Warb.
Orchidaceae
Ansellia africana Lindl.
Picrodendraceae Aristogeitonia monophylla Airy shaw
Putranjivaceae
Drypetes natalensis var. leiogyna Brenan
Drypetes usambarica var. trichogyna Radcl.-Sm.
Rubiaceae
Pavetta linearifolia Bremek.
Tarenna drummondii Bridson
Vangueria pallidiflora (Bullock) Lantz
Afrocanthium kilifiense (Bridson) Lantz
Pavetta tarennoides S. Moore
Rothmannia macrosiphon (K. Schum. ex Engl.)
Bridson
Rutaceae
Vepris sansibarensis (Engl.) Mziray
Sapindaceae
Chytranthus obliquinervis Radlk. ex Engl.
Sapotaceae
Mimusops riparia Engl.
Pouteria pseudoracemosa (J.H. Hemsl.) L.Gaut.
Zosteraceae
Zostera capensis Setch.
Acanthaceae
Chlamydacanthus lindavianus H. Winkl.
Justicia anisophylla (Mildbr.) Brummitt
Anacardiaceae
Lannea schweinfurthii var. acutifoliolata (Engl.)
Kokwaro
Annonaceae
Uvaria kirkii Oliv. ex Hook. f.
Uvaria denhardtiana Engl. & Diels
Asteranthe asterias (S. Moore) Engl. & Diels
Huberantha stuhlmannii (Engl.) Chaowasku
Ophrypetalum odoratum Diels
Araceae
Callopsis volkensii Engl.
Asteraceae
Brachylaena huillensis O. Hoffm.
Asphodelaceae Aloe deserti A. Berger
Burseraceae
Commiphora obovata Chiov.
Life form
Shrub
Shrub
Tree
Herb
Herb
Herb
Herb
Shrub
Tree
Shrub
Tree
Herb
Tree
Tree
Liana
Tree
Tree
Shrub
Herb
IUCN
Vulnerable
Vulnerable
Vulnerable
Vulnerable
Vulnerable
Vulnerable
Vulnerable
Vulnerable
Vulnerable
Vulnerable
Vulnerable
Vulnerable
Vulnerable
Vulnerable
Vulnerable
Vulnerable
Vulnerable
Vulnerable
Vulnerable
Version
3.1
3.1
3.1
3.1
3.1
3.1
3.1
3.1
3.1
2.3
2.3
3.1
3.1
2.3
2.3
2.3
2.3
3.1
3.1
Shrub
Shrub
Shrub
Climber
Herb
Tree
Tree
Epiphytic herb
Shrub
Shrub
Tree
Shrub
Shrub
Shrub
Shrub
Shrub
Tree
Vulnerable
Vulnerable
Vulnerable
Vulnerable
Vulnerable
Vulnerable
Vulnerable
Vulnerable
Vulnerable
Vulnerable
Vulnerable
Vulnerable
Vulnerable
Vulnerable
Vulnerable
Vulnerable
Vulnerable
3.1
3.1
3.1
3.1
3.1
3.1
2.3
3.1
2.3
2.3
2.3
3.1
2.3
2.3
2.3
2.3
2.3
Shrub
Tree
Tree
Tree
Herb
Shrub
Herb
Tree
Vulnerable
Vulnerable
Vulnerable
Vulnerable
Vulnerable
Near Threatened
Near Threatened
Near Threatened
2.3
2.3
3.1
2.3
3.1
3.1
3.1
2.3
Shrub
Shrub
Shrub
Shrub
Shrub
Herb
Tree
Shrub
Shrub
Near Threatened
Near Threatened
Near Threatened
Near Threatened
Near Threatened
Near Threatened
Near Threatened
Near Threatened
Near Threatened
3.1
3.1
3.1
3.1
3.1
3.1
2.3
3.1
2.3
Checklist of coastal forests of Kenya
Family
Burseraceae
Combretaceae
Connaraceae
Fabaceae
Meliaceae
Moraceae
Rubiaceae
Salicaceae
Species
Commiphora pseudopaolii J.B. Gillett
Conocarpus lancifolius Engl. & Diels
Ellipanthus madagascariensis (G. Schellenb.)
Capuron ex Keraudren
Dialium orientale Baker f.
Dalbergia eremicola Polhill
Dalbergia melanoxylon Guill. & Perr.
Erythrina sacleuxii Hua
Lovoa swynnertonii Baker f.
Pseudobersama mossambicensis (Sim) Verdc.
Dorstenia hildebrandtii Engl. var. hildebrandtii
Dorstenia goetzei Engl.
Dorstenia warneckei Engl.
Coffea pseudozanguebariae Bridson
Bivinia jalbertii Tul.
11
Life form
Tree
Tree
Tree
IUCN
Near Threatened
Near Threatened
Near Threatened
Version
2.3
2.3
2.3
Shrub
Shrub
Shrub
Tree
Tree
Tree
Herb
Herb
Herb
Shrub
Tree
Near Threatened
Near Threatened
Near Threatened
Near Threatened
Near Threatened
Near Threatened
Near Threatened
Near Threatened
Near Threatened
Near Threatened
Near Threatened
2.3
3.1
2.3
3.1
3.1
3.1
3.1
3.1
3.1
3.1
2.3
Table 6. Number of endemic species in coastal of East Africa forests found in Kenya coastal forests.
Region
Kenyan coastal forests
Kenyan and Tanzanian coastal forests
East Africa coastal endemic
No. of endemic species
90
185
551
found in Kenya coastal forests (Tables 6, 7). In terms of life forms herbs (44 species,
49.44%) had the highest number of endemic species recorded at the coastal region followed by shrubs (24 species, 26.97%), trees (13 species, 14.6%), climbers and lianas
recorded the least numbers of endemic species. (Table 8).
New records
Several new species have been discovered recently from the coastal forests of Kenya,
Adenia angulosa (Ngumbau et al. 2017), Dovyalis keniensis (Williams 2017),
Zehneria monocarpa (Ngumbau et al. unpublished) and Croton kinondoensis (Ngumbau
et al. in press) belonging to family Passifloraceae, Salicaceae, Cucurbitaceae and
Euphorbiaceae respectively. In addition, one species from family Poaceae, Panicum
peteri, was collected and recorded for the first time in Kenya.
Exotic species
A total of 72 introduced, cultivated or naturalized species are recorded in this study
belonging to 67 genera and 32 families, for Kenya coastal region (Table 9). The toprich exotic plant families include: Fabaceae (13 species), Asteraceae (7 species) and
Solanaceae (6 species). Out of the 72 plant species, herbs had the highest number of
exotic species (35) 48.61%, followed by trees (21) 29.17%, shrubs (12) 16.67% and
climbers (4) 5.55%.
12
Veronicah M. Ngumbau et al. / PhytoKeys 147: 1–191 (2020)
Table 7. The endemic plants of Kenya coastal forest.
Family
Acanthaceae
Amaranthaceae
Ancistrocladaceae
Annonaceae
Apocynaceae
Asparagaceae
Asteraceae
Burmanniaceae
Capparaceae
Caryophyllaceae
Commelinaceae
Convolvulaceae
Crassulaceae
Cucurbitaceae
Cyperaceae
Dichapetalaceae
Euphorbiaceae
Fabaceae
Lamiaceae
Species
Blepharis kenyensis Vollesen
Dicliptera maculata subsp. A
Justicia drummondii Vollesen
Justicia sp. aff. fittonioides S. Moore
Justicia sp. F
Megalochlamys tanaensis Vollesen
Hermbstaedtia gregoryi C.B. Clarke
Ancistrocladus robertsoniorum J.Léonard
Polyceratocarpus sp. ?nov.
Uvariodendron schmidtii Q. Luke ined.
Xylopia keniensis D. M. Johnson
Echidnopsis ericiflora Lavranos
Huernia andreaeana (Rauh) L.C. Leach
Huernia archeri L.C.Leach
Raphionacme sp. cf. jurensis N.E. Br.
Ceropegia racemosa var. voiensis Masinde
Sansevieria ballyi L.E. Newton
Sansevieria francisii Chahinian
Sansevieria nitida Chahinian
Athroisma pusillum T. Erikss.
Afrothismia baerae Cheek
Maerua mungaii Beentje
Thilachium roseomaculatum Y.B. Harv. & Vollesen
Polycarpaea grahamii Turrill
Polycarpaea tenuistyla Turrill
Aneilema succulentum Faden
Aneilema tanaense Faden
Merremia sp. C of FTEA
Kalanchoe ballyi Raym.-Hamet ex Cufod.
Kalanchoe fadeniorum Raadts
Cucumis sp. (Oreosyce sp. A of FTEA)
Zehneria monocarpa G.W. Hu, V.M. Ngumbau & Q.F. Wang, sp. nov.
Bulbostylis densecaespitosa (Lye) R. W. Haines
Bulbostylis hispidula subsp. intermedia (Lye) R.W. Haines
Cyperus boreobellus Lye
Cyperus kwaleensis Lye
Cyperus microumbellatus Lye
Dichapetalum fadenii Breteler
Dichapetalum sp. ?nov.
Croton kinondoensis G.W. Hu, V.M. Ngumbau & Q.F. Wang, sp. nov.
Euphorbia fluminis S. Carter
Euphorbia tanaensis P.R.O. Bally
Euphorbia taruensis S. Carter
Erythrococca pentagyna Radcl.-Sm.
Dichrostachys cinerea subsp. keniensis Brenan & Brummitt
Aeschynomene sp. B
Bauhinia mombassae Vatke
Cynometra greenwayi Brenan
Dalbergia gloveri Q. Luke, ined.
Vigna membranacea subsp. hapalantha (Harms) Verdc.
Crotalaria grata Polhill
Abrus sp. A of FTEA
Endostemon wakefieldii (Baker) M. Ashby
Life form
Herb
Herb
Herb
Herb
Herb
Herb
Herb
Liana
Tree
Tree
Tree
Herb
Herb
Herb
Climber
Herb
Herb
Herb
Herb
Herb
Herb
Shrub
Herb
Herb
Herb
Herb
Herb
Herb
Herb
Herb
Climber
Climber
Herb
Herb
Herb
Herb
Herb
Liana
Shrub
Shrub
Shrub
Tree
Herb
Shrub
Shrub
Shrub
Shrub
Tree
Liana
Herb
Shrub
Herb
Herb
Checklist of coastal forests of Kenya
Family
Lamiaceae
Lamiaceae
Loranthaceae
Lythraceae
Malvaceae
Montiniaceae
Nyctaginaceae
Ochnaceae
Olacaceae
Orchidaceae
Passifloraceae
Plumbaginaceae
Poaceae
Portulacaceae
Rubiaceae
Rutaceae
Salicaceae
Sapotaceae
Solanaceae
Urticaceae
Vitaceae
13
Species
Plectranthus auriglandulosus A.J.Paton
Leucas tsavoensis var. kilifiensis Sebald
Taxillus wiensii Polhill
Ammannia parkeri var. longifolia (Verdc.) S.A. Graham & Gandhi
Cola octoloboides Brenan
Grevea eggelingii var. keniensis (Verdc.) Verdc.
Boerhavia sp A of FTEA
Ochna sp. 17 of KTSL
Strombosiopsis pentamera Q. Luke, ined.
Eulophia serrata P.J. Cribb
Turnera thomasii (Urb.) Story
Plumbago stenophylla Wilmot-Dear
Eleusine semisterilis S.M.Phillips
Eragrostis sp. A
Portulaca coralloides S.M.Phillips
Tricalysia bridsoniana Robbr. var. bridsoniana
Keetia lukei Bridson
Mitriostigma greenwayi Bridson
Pavetta tarennoides S.Moore
Psydrax robertsoniae Bridson
Psydrax sp. A of FTEA
Canthium mrimaense (Verdc.) Lantz
Rytigynia sp. I of FTEA
Rytigynia sp. L of FTEA
Spermacoce sp. B of FTEA
Vangueriopsis shimbaensis A.P. Davis & Q. Luke
Vepris sp. (Diphasia sp. A of FTEA)
Vepris robertsoniae Q. Luke, ined.
Vepris sp. nr. stolzii I. Verd.
Dovyalis keniensis E. V. Williams
Synsepalum subverticillatum (E.A. Bruce) T.D.Penn.
Synsepalum sp. cf. subcordatum De Wild.
Solanum malindiense Voronts.
Pouzolzia fadenii Friis & Jellis
Cyphostemma sp. I of FTEA
Cyphostemma sp. G
Cyphostemma sp. L of FTEA
Life form
Herb
Herb
Shrub
Herb
Tree
Shrub
Herb
Shrub
Tree
Herb
Shrub
Herb
Herb
Herb
Herb
Shrub
Shrub
Shrub
Shrub
Shrub
Shrub
Shrub
Shrub
Tree
Herb
Tree
Tree
Tree
Tree
Shrub
Shrub
Tree
Shrub
Herb
Climber
Climber
Climber
Table 8. Life form representative for endemic species.
Life form
Climbers
Herbs
Lianas
Shrubs
Trees
Total
No. of species
6
44
3
24
13
90
Percentage presentation
5.62%
49.44%
3.37%
26.97%
14.6%
100
Conclusion
Kenya coastal forests host a large number of species and provide a unique habitat for
many species of special concern and endemic plant species. It harbors hundreds of
14
Veronicah M. Ngumbau et al. / PhytoKeys 147: 1–191 (2020)
Table 9. Exotic plants recorded at the coastal region of Kenya.
Family
Acanthaceae
Amaranthaceae
Anacardiaceae
Annonaceae
Apiaceae
Apocynaceae
Araucariaceae
Arecaceae
Asteraceae
Bixaceae
Cactaceae
Cassuarinaceae
Chrysobalanaceae
Combretaceae
Cucurbitaceae
Euphorbiaceae
Fabaceae
Lamiaceae
Lamiaceae
Malvaceae
Melastomataceae
Meliaceae
Species
Indoneesiella echioides (L.) Sreem.
Ruellia tuberosa L.
Ruspolia hypocrateriformis (Vahl) Milne-Redh.
Justicia gendarussa Burm. f.
Gomphrena globosa L.
Amaranthus dubius Mart. ex Thell.
Atriplex semibaccata R.Br.
Mangifera indica L.
Anacardium occidentale L.
Cananga odorata (Lam.) Hook. f. & Thomson
Foeniculum vulgare Mill.
Asclepias curassavica L.
Araucaria cunninghamii Aiton ex D. Don
Areca catechu L
Cocos nucifera L.
Synedrella nodiflora (L.) Gaertn.
Tridax procumbens L.
Ageratum conyzoides L.
Acanthospermum hispidum DC.
Flaveria trinervia (Spreng.) C. Mohr
Bidens pilosa L.
Galinsoga parviflora Cav.
Bixa orellana L.
Opuntia ficus-indica (L.) Mill.
Casuarina equisetifolia L.
Dactyladenia barteri (Hook. f. ex Oliv.) Prance & F. White
Terminalia catappa L.
Conocarpus lancifolius Engl. ex Engl. & Diels
Luffa cylindrica M. Roem.
Citrullus lanatus (Thunb.) Matsum. & Nakai
Euphorbia heterophylla L.
Croton blanchetianus Baill.
Centrosema pubescens Benth.
Desmodium adscendens (Sw.) DC. var. adscendens
Senna timoriensis (DC.) H.S.Irwin & Barneby
Caesalpinia pulcherrima (L.) Sw.
Leucaena leucocephala (Lam.) de Wit
Prosopis juliflora (Sw.) DC.
Lysiloma latisiliquum (L.) Benth.
Tamarindus indica L.
Samanea saman (Jacq.) Merr.
Senna siamea (Lam.) H. S. Irwin & Barneby
Parkinsonia aculeata L.
Pithecellobium dulce (Roxb.) Benth.
Mimosa pudica var. unijuga (Duchass. & Walp.) Griseb.
Ocimum basilicum L.
Mesosphaerum suaveolens (L.) Kuntze
Corchorus parviflorus (Benth.) Domin
Gossypium barbadense L.
Gossypium hirsutum L.
Clidemia hirta (L.) D. Don
Melia azedarach L.
Toona ciliata M. Roem.
Growth form
Herb
Herb
Herb
Herb
Herb
Herb
Herb
Tree
Tree
Tree
Herb
Herb
Tree
Herb
Tree
Herb
Herb
Herb
Herb
Herb
Herb
Herb
Shrub
Shrub
Tree
Tree
Tree
Tree
Climber
Climber
Herb
Shrub
Climber
Herb
Shrub
Shrub
Shrub
Shrub
Tree
Tree
Tree
Tree
Tree
Tree
Herb
Herb
Herb
Herb
Shrub
Shrub
Shrub
Tree
Tree
Checklist of coastal forests of Kenya
Family
Meliaceae
Moringaceae
Myrtaceae
Oxalidaceae
Papaveraceae
Passifloraceae
Piperaceae
Sapindaceae
Solanaceae
Talinaceae
Verbenaceae
Species
Azadirachta indica A.Juss.
Moringa oleifera Lam.
Syzygium cumini (L.) Skeels
Oxalis latifolia Kunth
Oxalis barrelieri L.
Oxalis corniculata L.
Argemone mexicana L.
Passiflora foetida L.
Piper betle L.
Sapindus trifoliatus L.
Lycopersicon esculentum Mill.
Datura metel L.
Lycopersicon esculentum Mill.
Physalis angulata L.
Physalis minima L.
Solanum americanum Mill.
Talinum paniculatum (Jacq.) Gaertn.
Lantana camara L.
Stachytarpheta urticifolia Sims
15
Growth form
Tree
Tree
Tree
Herb
Herb
Herb
Herb
Climber
Herb
Tree
Herb
Herb
Herb
Herb
Herb
Herb
Herb
Shrub
Shrub
plant species that are only found in coastal forests of East Africa. In addition, they are
home to almost half of plant species found in Kenya flora (39.55%), and more than
half of species found in the coastal forests of East Africa (65%). Despite this high plant
diversity relevance, they have undergone a history of anthropogenic disturbance since
the colonial era (Moomaw 1960). Today the rapidly increasing human population
along the coast of Kenya demands more agricultural land, more wood products for
fuel, house construction, and industry, consequentially threatening the small areas of
intact forest and woodland. This therefore, has resulted in large number of threatened
and near threatened plants in this region. High conservation measures should be highly
taken into action to ensure the protection of large numbers of endemic and threatened
species that are at the brink of extinction. Besides, conservation of these relicts will not
only provide the protection of species rich diversity of the flora but also of the fauna,
including humans which depend on it. Importantly, it will prevent soil erosion and the
drying up of moist microhabitats as a result protecting catchment areas and wetlands.
This study hence provides not only a comprehensive list of the plant species that have
been recorded in this region but also endemic species and species of special concern to
guide conservationists and policymakers towards conserving the remaining patches. It
also provides a prerequisite to ecology and other thorough work towards understanding plant diversity and conservation of the region.
Checklist
This checklist is designed for use by anybody interested in the plants of the Kenya coastal
region. However, it will be most useful to professionals and students who have some botanical knowledge. Information such as voucher number, habit, habitat, altitude is record-
16
Veronicah M. Ngumbau et al. / PhytoKeys 147: 1–191 (2020)
ed for each species. Species without voucher numbers were not included in the checklist.
Species preceded by “?” means that there is doubt over the correct identification, those
preceded with “letter” means that they have not been described and finally vouchers documented as “sr” means that they were sight recorded. EA means the East African Herbarium, Nairobi, Kenya; HIB means the Herbarium of Wuhan Botanical Garden, Wuhan,
China, and SAJIT means Sino Africa Joint Investigation Team. The families are divided
into four groups, namely lycophytes, monilophytes, gymnosperms, and angiosperms.
Families of lycophytes and monilophytes are organized based on the PPG I system (PPG
I 2016), while those of gymnosperms are organized based on Christenhusz et al. (2011)
and families of angiosperms are organized based on the APG IV system (APG IV 2016).
Part 1 Lycophytes
F1. Selaginellaceae
1 Genus, 3 Species
Selaginella eublepharis A. Braun Habit: Herb. Habitat: Coastal forest and bushland,
0–300 m. Vouchers: Robertson & Luke 5840, Hildebrandt 1961, Drummond &
Hemsley 4258, Wamukoya G K7–x936 (EA).
Selaginella mittenii Baker Habit: Herb. Habitat: Wet rocks of forest streams, waterfalls in evergreen forest, ca. 20 m. Voucher: Luke et al. 4521 (EA).
Selaginella perpusilla Baker Habit: Herb. Habitat: In Hyphaene grassland in shade
of grasses, 7–400 m. Vouchers: Drummond & Hemsley 4034, Faden & Evans
70/441, Rawlins 38, Faden RB 70/441 (EA).
Part 2 Monilophytes
F2. Aspleniaceae
1 Genus, 5 Species
Asplenium affine var. mettenii (Kuhn) Tardieu Habit: Herb. Habitat: Forest, ca.
50 m. Voucher: Kresten 29 (EA).
Asplenium buettneri Hieron. Habit: Epiphytic herb. Habitat: Forest, ca. 280 m.
Vouchers: Ngumbau V & Mwadime N V016 (EA, HIB), Luke WRQ & Robertson SA 2743B (EA).
Asplenium erectum Bory ex Willd. Habit: Epiphytic herb. Habitat: Forest, ca. 400 m.
Voucher: HD van Someren 116 (EA).
Asplenium nidus L. Habit: Epiphytic herb. Habitat: Moist forest, 0–275 m. Vouchers: Ngumbau V & Mwadime N V067 (EA, HIB), Luke et al. 4529 & 4512, Luke
& Robertson 2693 (EA).
Checklist of coastal forests of Kenya
17
Asplenium emarginatum P. Beauv. Habit: Herb. Habitat: Moist forest, ca. 40 m.
Vouchers: Luke WRQ 1645, Faden & Faden 72/224 (EA).
F3. Azollaceae
1 Genus, 1 Species
Azolla nilotica Decne. ex Mett. Habit: Herb. Habitat: River, lake edges, and temporary pools, ca. 10 m. Voucher: Festo L, Luke Q & P 2751 (EA).
F4. Blechnaceae
1 Genus, 1 Species
Stenochlaena tenuifolia (Desv.) Moore Habit: Epiphytic herb. Habitat: Swamp forest, ca. 0–140 m. Vouchers: Ngumbau V & Mwadime N V0229 (EA, HIB), Luke
WRQ & Mbinda J 6278 (EA).
F5. Dryopteridaceae
1 Genus, 1 Species
Bolbitis auriculata (Lam.) Alston Habit: Herb. Habitat: Moist forest, ca. 300 m.
Vouchers: Luke WRQ & Robertson SA 2750, Luke 1648 (EA).
F6. Davalliaceae
1 Genus, 1 Species
Davallia denticulata (Burm. f.) Mett. ex Kuhn Habit: Epiphyte. Habitat: Bushland
on rocks, ca. 2–410 m. Vouchers: Ngumbau V & Mwadime N V0305 (EA, HIB),
Robertson SA & Luke WRQ 6046 (EA).
F7. Dennstaedtiaceae
2 Genera, 2 Species
Microlepia speluncae (L.) T. Moore Habit: Herb. Habitat: Forests edge, 350–700 m.
Voucher: Luke WRQ & PA 8199 (EA).
Pteridium aquilinum (L.) Kuhn Habit: Herb. Habitat: Seasonally wet grassland,
0–150 m. Vouchers: Magogo FC & Glover PE 25, Bally & Smith 14340 (EA).
F8. Hymenophyllaceae
2 Genera, 2 Species
Crepidomanes chevalieri (Christ) Ebihara & Dubuisson Habit: Epiphytic herb.
Habitat: Moist forest, 150–500 m. Voucher: Faden et al. 70/812 (EA).
18
Veronicah M. Ngumbau et al. / PhytoKeys 147: 1–191 (2020)
Didymoglossum erosum (Willd.) J.P. Roux Habit: Low epiphytic or lithophytic herb.
Habitat: Shaded rocks or trunk of trees, ca. 230 m. Voucher: Luke WRQ & Robertson SA 2760 (EA).
F9. Nephrolepidaceae
1 Genus, 2 Species
Nephrolepis acutifolia (Desv.) Christ Habit: Herb. Habitat: Rocky cliff or AcaciaCommiphora bushland, ca. 230 m. Voucher: Luke & Robertson 2530 (EA).
Nephrolepis biserrata (Sw.) Schott Habit: Herb. Habitat: Evergreen forest or streamside forest, ca. 417 m. Vouchers: Ngumbau V & Mwadime N V0322 (EA, HIB),
Luke WRQ & Robertson SA 2750, Magogo & Glover 518, Drummond & Hemsley 1221 (EA).
F10. Ophioglossaceae
1 Genus, 5 Species
Ophioglossum costatum R. Br. Habit: Herb. Habitat: Dry deciduous woodland, ca.
10 m. Vouchers: Faden & Faden 74/1119, Festo 2731 (EA).
Ophioglossum latifolium (Prantl) J. E. Burrows Habit: Herb. Habitat: In seasonally
wet shallow sandy soils and deciduous woodland, ca. 10 m. Vouchers: Faden &
Faden 74/1121, Faden RB 70/439B (EA).
Ophioglossum lusoafricanum Welw. ex Prantl Habit: Herb. Habitat: In marshy areas
among tall grasses, grassland, and woodland, ca. 150 m. Voucher: Faden 70/440 (EA).
Ophioglossum nudicaule var. robustum J. S. Licht. Habit: Herb. Habitat: In shallow
sandy soils, dry tropical woodland, and savannah, ca. 10 m. Voucher: Faden &
Faden 74/1120 (EA).
Ophioglossum reticulatum L. Habit: Herb. Habitat: Grassland, moist woodland,
montane grassland among rocks, and forest margins, ca. 0–167 m. Vouchers: Magogo & Glover 896, Schippers RR 196 (EA).
F11. Pteridaceae
8 Genera, 13 Species
Acrostichum aureum L. Habit: Herb. Habitat: Woodland, ca. 253 m. Voucher: SAJIT–005915 (EA, HIB).
Adiantum capillus-veneris L. Habit: Herb. Habitat: Road banks and riverbanks, ca.
200 m. Voucher: Luke WRQ 3969 (EA).
Adiantum comorense (Tardieu) Verdc. Habit: Herb. Habitat: Rainforest, rivulet,
road banks, and deep shade, 100–700 m. Vouchers: Gilbert et al. 4964, Magogo
& Clover 726, Luke 3600 (EA).
Adiantum incisum Forssk. Habit: Herb. Habitat: Riverside, path sides, and riverine
forest, ca. 86 m. Vouchers: Ngumbau V & Mwadime N V0477 (EA, HIB), Kassner 137 (EA).
Checklist of coastal forests of Kenya
19
Adiantum lunulatum Burm. f. Habit: Terrestrial or lithophytic herb. Habitat: Lowland forest or woodland, 160 m. Voucher: Luke WRQ 3147 (EA).
Ceratopteris thalictroides (L.) Brongn. Habit: Herb. Habitat: Swampy areas, swamp
forests, marshes, ca. 15 m. Vouchers: SAJIT–006213 & 005485 (EA, HIB), Luke
WRQ & PA 4577 (EA).
Cheilanthes involuta (Sw.) Schelpe & N. C. Anthony Habit: Herb. Habitat: Woodland, 90–450 m. Vouchers: SAJIT–006131 (EA, HIB), Napper 2180, Magogo FC
& Glover PE 754 (EA).
Pellaea dura (Willd.) Hook. Habit: Herb. Habitat: Forest or woodland, ca. 90 m.
Vouchers: Verdcourt 5290, Luke WRQ & Robertson SA 2786A (EA).
Pityrogramma calomelanos (L.) Link Habit: Herb. Habitat: Riverbanks, forest, ca.
150 m. Voucher: Luke 5273 (EA).
Pteris atrovirens Willd. Habit: Herb. Habitat: Shady places, ca. 60 m. Vouchers:
Faden RB 226, Luke WRQ 2936 (EA).
Pteris hamulosa (Christ) Christ Habit: Terrestrial herb. Habitat: In shade on wet forest floors, ca. 100 m. Vouchers: Luke WRQ 2937, Luke & Robertson 2729 (EA).
Pteris tripartita Sw. Habit: Terrestrial or epiphytic herb. Habitat: Moist area, ca.
60 m. Voucher: Faden RB 218 (EA).
Vittaria elongata Sw. Habit: Epiphytic or lithophytic herb. Habitat: Low altitude
forest, ca. 10–200 m. Vouchers: Robertson SA & Luke WRQ 5168, Magogo &
Glover 222, Faden & Evans 70/414, Faden et al. 69/486 (EA).
F12. Psilotaceae
1 Genus, 1 Species
Psilotum nudum (L.) P. Beauv. Habit: Herb. Habitat: Dry riverbeds, under cliff overhangs, on rocks, and in rock crevices, 0–100 m. Voucher: Dale in FD 3584 (EA).
F13. Polypodiaceae
4 Genera, 5 Species
Microgramma mauritiana (Willd.) Tardieu Habit: Epiphytic herb. Habitat: Rainforest, evergreen thicket, and riverine forest, ca. 30–300 m. Vouchers: SAJIT–004664
& 005433, Ngumbau V & Mwadime N V0178 (EA, HIB), Gillett 18700, Polhill
& Paulo 832, Luke WRQ & Robertson SA 2799 (EA).
Microsorum punctatum (L.) Copel. Habit: Epiphytic, lithophytic or terrestrial herb.
Habitat: Forest, evergreen thicket or beneath hedges in wet areas, 20–350 m.
Vouchers: Verdcourt 3919, Luca, Jeffery & Kirika 230, Faden RB 69/484 (EA).
Microsorum scolopendria (Burm. f.) Copel. Habit: Epiphytic, lithophytic or terrestrial herb. Habitat: Evergreen forest and thicket, 0–297 m. Vouchers: Ngumbau V
& Mwadime N V0152 (EA, HIB), Verdcourt 3914, Polhill & Paulo 858, Faden
RB 69/488 (EA).
Platycerium alcicorne Desv. Habit: Epiphytic herb. Habitat: Evergreen thickets, 30–200
m. Vouchers: Robertson & Luke 6362, Verdcourt 2408, RB & AJ Faden 72/66 (EA).
20
Veronicah M. Ngumbau et al. / PhytoKeys 147: 1–191 (2020)
Platycerium angolense Welw. ex Hook. Habit: Epiphytic herb. Habitat: Mixed evergreen forest, woodland, ca. 5–150 m. Vouchers: RB & AJ Faden 74/1144, Luke
WRQ & Robertson SA 2811 (EA).
F14. Tectariaceae
1 Genus, 1 Species
Tectaria puberula (Desv.) C. Chr. Habit: Herb. Habitat: Moist forest, 1–150 m.
Vouchers: Faden RB 70/808, Faden RB, Evans A & Msafiri F 952 (EA).
F15. Thelypteridaceae
4 Genera, 5 Species
Chrismatopteris holttumii N. Quansah & D.S. Edwards Habit: Herb. Habitat: Forest, 150 m. Voucher: Faden RB 70/422B (EA).
Christella dentata (Forssk.) Brownsey & Jermy Habit: Herb. Habitat: Along stream
banks in forest or forest floor, 0–100 m. Voucher: Faden RB 70/419 (EA).
Christella hispidula (Decne.) Holttum Habit: Herb. Habitat: Terrestrial on forest
floor away from water or along stream banks in forests, ca. 150 m. Voucher: Luke
WRQ & Robertson SA 2710 (EA).
Cyclosorus interruptus (Willd.) H. Itô Habit: Herb. Habitat: Grassland swamps,
0–50 m. Voucher: Luke WRQ & PA 4580 (EA).
Sphaerostephanos arbuscula subsp. africanus Holttum Habit: Herb. Habitat: Shady
valley, near streams or forest fringing, ca. 300 m. Vouchers: Faden RB 70/413,
Drummond RB & Hemsley JH 1203 (EA).
F16. Lygodiaceae
1 Genus, 2 Species
Lygodium kerstenii Kuhn Habit: Herb. Habitat: Lowland forests, near sea level.
Voucher: Kersten 76 (EA).
Lygodium microphyllum (Cav.) R. Br. Habit: Climber. Habitat: Swampy forest and
thicket, ca. 10–200 m. Vouchers: Gardner in FD 1456, Drummond & Hemsley
1208, Robertson SA & Luke WRQ 5146 (EA).
Part 3 Gymnosperm
F17. Zamiaceae
1 Genus, 1 species
Encephalartos hildebrandtii A. Braun & Bouché Habit: Tree. Habitat: Coastal evergreen bushland and lowland forest, ca. 31 m. Vouchers: Mwangangi OM, Waiganjo T & Kamau P 3800 (EA).
Checklist of coastal forests of Kenya
21
Part 4 Angiosperms
F18. Acanthaceae
30 Genera, 108 Species
Anisotes galanae (Baden) Vollesen Habit: Shrub. Habitat: Acacia and Acacia-Commiphora bushland, 25–550 m. Voucher: Robertson 5284 (EA): Vulnerable.
Anisotes parvifolius Oliv. Habit: Shrub. Habitat: Acacia-Terminalia and AcaciaCommiphora bushland, 25–650 m. Vouchers: Bally, PRO 16793, Drummond &
Hemsley 4265 (EA).
Anisotes tanensis Baden Habit: Shrub. Habitat: Bushland & limestone escarpments,
250–600 m. Voucher: Taylor WE 181 (EA).
Asystasia ansellioides C.B. Clarke Habit: Herb. Habitat: Open grassland or Hyphaene grassland, 15–200 m. Vouchers: Luke WRQ & PA 4509, Jeffrey GM 5, Bally
6105, Rawlins 699 (EA).
Asystasia gangetica subsp. micrantha (Nees) Ensermu Habit: Herb. Habitat: Forest
margin, woodland, wooded grassland, bushland, and sandy seashores, ca. 300 m.
Vouchers: Ngumbau V & Mwadime N V0259 (EA, HIB), Jeffrey GW 500, Harvey et al. 54, Robertson & Faulkner 2303 (EA).
Asystasia guttata (Forssk.) Brummitt Habit: Herb. Habitat: Isolated inselberg in
ACB, ca. 600 m. Voucher: Luke WRQ 1396 (EA).
Asystasia leptostachya Lindau Habit: Herb. Habitat: Swampy areas of rainforest, ca.
50 m. Voucher: Luke WRQ 1650 (EA).
Asystasia linearis S. Moore Habit: Herb. Habitat: Tall grassland on cotton soil, 25–
200 m. Voucher: Gregory s.n. (EA): Endangered.
Asystasia minutiflora Ensermu & Vollesen Habit: Herb. Habitat: Lowland
wet evergreen forest, lowland semi-deciduous forest, and thicket, ca. 400 m.
Vouchers: Bidgood, Abdallah & Vollesen s.n., Luke 1650, Magogo & Glover
1101 (EA).
Asystasia moorei Ensermu Habit: Herb. Habitat: Lowland evergreen forest, ca.
300 m. Voucher: Luke WRQ & Robertson SA 2703 (EA).
Asystasia mysorensis (Roth) T. Anderson Habit: Herb. Habitat: Lowland forest, ca.
100 m. Vouchers: J Makin 14616, M Hucks 649 (EA).
Asystasia sp. B Habit: Herb. Habitat: Riverine forest, 450 m. Voucher: Battiscombe
250 (EA).
Avicennia marina (Forssk.) Vierh. Habit: Shrub. Habitat: Inland fringes of mangrove associations, sandy dunes, mud of tidal rivers, and salty creeks, sea level.
Vouchers: Graham RM 251, Greenway 8933 (EA).
Barleria acanthoides Vahl Habit: Herb. Habitat: Bushland, ca. 690 m. Voucher: Ossent J 18 (EA).
Barleria argentea Balf. f. Habit: Herb. Habitat: Woodland, semi-desert bushland,
grassland, disturbed, and eroded areas, ca. 100 m. Voucher: Gillett 19514 (EA).
Barleria gracilispina (Fiori) I. Darbysh. Habit: Herb. Habitat: Woodland, grassland, and degraded areas, ca. 30 m. Voucher: Gillett & Gachathi 20522 (EA).
22
Veronicah M. Ngumbau et al. / PhytoKeys 147: 1–191 (2020)
Barleria inclusa I. Darbysh. Habit: Herb. Habitat: Open woodland, grassland or dry
rocky, ca. 400 m. Voucher: Drummond RB & Hemsley 4138 (EA).
Barleria linearifolia Rendle Habit: Herb. Habitat: Bushland and wooded grassland,
ca. 30 m. Voucher: Robertson 1816 (EA).
Barleria lukei I. Darbysh. Habit: Climber. Habitat: Forest, ca. 264 m. Vouchers:
Ngumbau V & Mwadime N V099 (EA, HIB), Drummond & Hemsley 3976,
Magogo & Glover, Robertson & Luke 1955 (EA): Endangered.
Barleria maculata S. Moore Habit: Herb. Habitat: Dry rocky, wooded grassland, ca.
80 m. Vouchers: Drummond & Hemsley 1953, Luke 3492 (EA): Endangered.
Barleria marginata Oliv. Habit: Herb. Habitat: Open bushland and lightly wooded grassland, 100–650 m. Vouchers: Luke, Mbinda & Madudu 6346, Tweedie
2197 (EA).
Barleria maritima I. Darbysh. Habit: Herb. Habitat: Lowland bushland, scrub on
coral rag, and along the shoreline, ca. 15 m. Vouchers: Symes 189, Mainwaring
3045, Napier 2301, Tweedie 3165, Rayner 259 (EA).
Barleria paolioides I. Darbysh. Habit: Shrub. Habitat: Open woodland and grassland, 30–850 m. Voucher: Williams Sangai 832 (EA).
Barleria polhillii I. Darbysh. subsp. polhillii Habit: Shrub. Habitat: Dry woodland,
bushland, grassland, and grazed areas, 0–200 m. Voucher: Luke Q 5472 (EA).
Barleria ramulosa C.B. Clarke Habit: Shrub. Habitat: Dry open bushland and
wooded grassland, 6–152 m. Vouchers: SAJIT–005909 (EA, HIB), Curtis 6,
Drummond & Hemsley 4082, Luke & Robertson 2567 (EA).
Barleria repens Nees Habit: Shrub. Habitat: Coastal dry forest and lowland woodland, 0–100 m. Vouchers: Graham 2102, Luke & Robertson 2598 (EA).
Barleria sp. E Habit: Herb. Habitat: Bushland, 0–100 m. Vouchers: Rawlins 771,
Bally 16859 & 16864 (EA).
Barleria submollis Lindau Habit: Herb. Habitat: Bushland or wooded grassland, ca.
182 m. Vouchers: Ngumbau V & Mwadime N V0349 (EA, HIB), Robertson SA
& Luke WRQ 6022, Kassner T 283 (EA).
Barleria taitensis S. Moore Habit: Herb. Habitat: Bushland, grassland, and margin
of riverine grassland, ca. 620 m. Vouchers: Bally PRO 16890, Mwadime N 6 (EA).
Barleria usambarica Lindau Habit: Herb. Habitat: Lowland forest margins, bushland, and grassland, 0–600 m. Vouchers: Robertson 6558, RB & AJ Faden
74/1157 (EA): Vulnerable.
Barleria ventricosa Hochst. ex Nees Habit: Herb. Habitat: Woodland and along forest margins, 700 m. Voucher: Luke WRQ & PA 6023 (EA).
Barleria whytei S. Moore Habit: Herb. Habitat: Coastal forest on coral rag, 0–10
m. Vouchers: Drummond & Hemsley 3923, Luke 2907, Luke & Saidi 5829
(EA): Endangered.
Blepharis kenyensis Vollesen Habit: Herb. Habitat: Coastal bushland, 0–150 m.
Vouchers: Kimeu JM 532, Graham RM 1630 (EA): Endangered, Endemic.
Blepharis maderaspatensis (L.) B. Heyne ex Roth Habit: Herb. Habitat: Grassland,
bushland, woodland, and forest habitats, ca. 95 m. Vouchers: SAJIT–006127 &
004654 (EA, HIB), Robertson SA & Luke WRQ 6028, Jeffrey GM 15 (EA).
Checklist of coastal forests of Kenya
23
Blepharis pratensis S. Moore Habit: Herb. Habitat: Bushland, 0–50 m. Vouchers: Taylor WE s.n., Polhill & Paulo 690, Makin 416, Luke 3064, 3398 & 3562
(EA): Vulnerable.
Cephalophis lukei Vollesen Habit: Herb. Habitat: Wet evergreen lowland forest, 25–
400 m. Vouchers: Harvey & Vollens 41, Luke 3377 (EA).
Chlamydacanthus lindavianus H. Winkl. Habit: Herb. Habitat: Coastal forest, 10–
250 (–600) m. Vouchers: Ngumbau V & Mwadime N V0421 (EA, HIB), Luke
WRQ & Robertson SA 2787, Luke 2941, Luke et al. 6273 (EA): Near Threatened.
Crabbea velutina S. Moore Habit: Herb. Habitat: Woodland, bushland, riverine forests, and thicket, ca. 35 m. Voucher: Luke Q 1485 (EA).
Crossandra mucronata Lindau Habit: Herb. Habitat: Woodland, bushland, upland
forests, and dry riverine forest, 50–300 m. Voucher: Whyte A s.n. (EA).
Crossandra pungens Lindau Habit: Herb. Habitat: Evergreen or semi-evergreen
coastal forest and thicket, 0–500 m. Vouchers: Festo L, Luke Q & P 2693, Drummond & Hemsley 3755A, Gillespie 312, Rawlins 71 (EA).
Crossandra stenostachya (Lindau) C.B. Clarke Habit: Herb. Habitat: Bushland and
riverine forest, ca. 60 m. Vouchers: Faden 74/986, Festo 2283 (EA).
Dicliptera albicaulis (S. Moore) S. Moore Habit: Herb. Habitat: Open Acacia woodland, short grassland, roadsides, ca. 800 m. Voucher: Mumiukha 29 (EA).
Dicliptera heterostegia Nees Habit: Herb. Habitat: Coastal riverine forest, forest margins, secondary woodland, and thicket, ca. 300 m. Voucher: Magogo FC & Glover
PE 137 (EA).
Dicliptera inconspicua I. Darbysh. Habit: Herb. Habitat: Lowland moist semi-deciduous forest and riverine forest, 0–450 m. Voucher: Festo L, Luke Q & P 2692
(EA): Vulnerable.
Dicliptera maculata subsp. A Habit: Herb. Habitat: Lowland moist forest or roadsides, ca. 300 m. Vouchers: Luke 2922, Luke stone & Baer 8243 (EA): Endemic.
Duosperma crenatum (Lindau) P.G. Mey. Habit: Shrub. Habitat: Deciduous woodland and scrub, ca. 800 m. Voucher: Bally PRO 16824 (EA).
Duosperma grandiflorum Vollesen Habit: Shrub. Habitat: Bushland, woodland, and
wooded grassland, ca. 300 m. Voucher: Mungai et al. 239/83 (EA).
Dyschoriste hildebrandtii (S. Moore) S. Moore Habit: Shrub. Habitat: Woodland,
wooded grassland, and bushland, ca. 100 m. Vouchers: Drummond & Hemsley
4124, Robertson SA & Luke WRQ 5981 (EA).
Ecbolium amplexicaule S. Moore Habit: Herb. Habitat: Forest, thicket, riverine forest, and sisal plantations, 0–400 m. Vouchers: Luke Q 1465, Bally 13065, Robertson 5902, Drummond & Hemsley 3921 (EA).
Ecbolium subcordatum C.B. Clarke var. subcordatum Habit: Herb. Habitat: Lowland Acacia and Acacia-Commiphora bushland, riverbanks, and river-beds, ca. 182
m. Vouchers: Bally PRO 16757, Luke et al. 644, Greenway 9818 (EA).
Ecbolium subcordatum var. glabratum Vollesen Habit: Herb. Habitat: Low bushland
and riverbanks, 60–152 m. Vouchers: Bally PRO 16819, Luke et al. TPR 387 (EA).
Ecbolium viride (Forssk.) Alston Habit: Herb. Habitat: Bushland, riverbeds, and
coastal bushland on sand dunes, 0–50 m. Voucher: Faden 74/1114 (EA).
24
Veronicah M. Ngumbau et al. / PhytoKeys 147: 1–191 (2020)
Elytraria acaulis (L.f.) Lindau Habit: Herb. Habitat: Alluvial grassland with AcaciaCombretum-Dobera thicket, 25–800 m. Vouchers: Greenway & Rawlins 9460,
Faden 74/1059 (EA).
Elytraria minor Dokosi Habit: Herb. Habitat: Evergreen and semi-deciduous forest, secondary thickets, and cultivated areas, 15–450 (–650) m. Vouchers: SAJIT–006026 (EA, HIB), Greenway PJ & Rawlins SP 9366, Robertson SA & Luke
WRQ 5986, Rawlins 34, Faden 71/798 (EA).
Hygrophila auriculata (Schumach.) Heine Habit: Herb. Habitat: Seasonal pools or
other stagnant water bodies, ca. 156 m. Vouchers: SAJIT–005982, Ngumbau V
& Mwadime N V0435 (EA, HIB), Magogo FC & Glover PE 528, Drummond &
Hemsley 3927 (EA).
Hypoestes forskaolii subsp. hildebrandtii (Lindau) I. Darbysh. Habit: Herb. Habitat:
Dry grassland and open bushland, ca. 382 m. Vouchers: SAJIT–006066, Ngumbau V & Mwadime N V0365 (EA, HIB), Magogo FC & Glover PE 138 (EA).
Indoneesiella echioides (L.) Sreem. Habit: Herb. Habitat: Bushland and grassland,
ca. 2 m. Voucher: Ngumbau V & Mwadime N V0518 (EA, HIB): Exotic.
Justicia anisophylla (Mildbr.) Brummitt Habit: Shrubby herb. Habitat: Lowland
and semi-deciduous forest, ca. 200 m. Vouchers: Faden 77/421, Luke 29142 (EA):
Near Threatened.
Justicia baravensis C.B. Clarke Habit: Herb. Habitat: Sandy foreshores, coastal
bushland, 0–15 m. Vouchers: Robertson & Brummit 6763, Kokwaro 3900 (EA).
Justicia betonica L. Habit: Herb. Habitat: Grassland, bushland, and woodland, ca.
253 m. Vouchers: Ngumbau V & Mwadime N V0121 (EA, HIB), Robertson SA
& Luke WRQ 5979 (EA).
Justicia brevipila Hedrén Habit: Herb. Habitat: Bushland and grassland, 25–350 m.
Vouchers: Luke et al. TRP785, Robertson & Luke 5728 (EA): Vulnerable.
Justicia breviracemosa Vollesen Habit: Shrub. Habitat: Lowland forest and riverine
forest, 10–250 m. Voucher: Luke 4271 (EA): Endangered.
Justicia caerulea Forssk. Habit: Herb. Habitat: Grassland, bushland, riverine, and
scrub, ca. 140 m. Vouchers: Thomas 38, RB & AJ Faden 74/985 (EA).
Justicia calyculata Deflers Habit: Herb. Habitat: Woodland, bushland, and grassland, ca. 25 m. Vouchers: Robertson SA & Luke WRQ 6039, Luke 3355 (EA).
Justicia capensis Thunb. Habit: Herb. Habitat: Forest or forest grassland, 5–715 m.
Vouchers: Luke & Robertson 1810 & 2029 (EA).
Justicia debilis (Forssk.) Vahl Habit: Herb. Habitat: Forest, ca. 186 m. Vouchers:
Ngumbau V & Mwadime N V0336 (EA, HIB), Festo L & Luke Q 2436 (EA).
Justicia drummondii Vollesen Habit: Herb. Habitat: Lowland forest, 75–200 m.
Vouchers: Drummond & Hemsley 3792, Robertson & Luke 5985, Luke 4508
(EA): Critically Endangered, Endemic.
Justicia engleriana Lindau Habit: Shrub. Habitat: Lowland to low montane evergreen forest, ca. 372 m. Vouchers: SAJIT–006150 (EA, HIB), Mbinda J & Luke
WRQ 259, Magogo & Glover 139 (EA).
Justicia faulknerae Vollesen Habit: Shrub. Habitat: Semi-deciduous lowland forest,
150–450 m. Vouchers: Luke 1590, Luke & Robertson 1819 (EA): Endangered.
Checklist of coastal forests of Kenya
25
Justicia fittonioides S. Moore Habit: Herb. Habitat: Dry or semi-deciduous lowland
forest, ca. 240 m. Vouchers: Ngumbau V & Mwadime N V107 (EA, HIB), Luke
WRQ 1651, Robertson 6560 (EA).
Justicia flava (Forssk.) Vahl Habit: Herb. Habitat: Forest margin and clearings,
woodland, grassland, and bushland, (25–) 350 m. Vouchers: SAJIT–006110 (EA,
HIB), Festo L & Luke WRQ 2443 (EA).
Justicia galeata Hedrén Habit: Herb. Habitat: Forest and shrubland, ca. 274 m.
Voucher: Dale IR 1077 (EA).
Justicia gendarussa Burm. f. Habit: Shrub. Habitat: Abandoned gardens, ca.
0–300 m. Vouchers: Dale 2015, Reitsma 448 (EA): Naturalized.
Justicia heterotricha Mildbr. Habit: Herb. Habitat: Forest, ca. 240 m. Vouchers:
Luke & Robertson 2770, Luke WRQ 2931 (EA): Vulnerable.
Justicia inaequifolia Brummitt Habit: Shrub. Habitat: Evergreen or semi deciduous lowland forest or coral rag thicket, ca. 420 m. Vouchers: SAJIT–006088 (EA,
HIB), Luke WRQ & Robertson SA 2699, BR Adams 87, Luke 1828, Magogo &
Glover 136, Robertson SA & Polhill 4833 (EA).
Justicia nyassana Lindau Habit: Herb. Habitat: Lowland forest, riverine forest, and
woodland, ca. 100 m. Voucher: Drummond & Hemsley 3872 (EA).
Justicia odora (Forssk.) Lam. Habit: Herb. Habitat: Dry wooded grassland, open
woodland, and scrub, 0–300 m. Voucher: Field CR 59 (EA).
Justicia pseudorungia Lindau Habit: Shrub. Habitat: Lowland to montane evergreen
forest, and riverine forest, 15 m. Vouchers: SAJIT–006220 (EA, HIB), Taylor WE
s.n., Robertson 5085, Faden & Evans 71/736 (EA).
Justicia scandens Vahl Habit: Herb. Habitat: Forest, bushland, and lake shores, ca.
50 m. Voucher: Luke & Robertson 2690 (EA).
Justicia schimperiana subsp. campestris Vollesen Habit: Shrub. Habitat: Bushland
and dry riverine forest, ca. 35 m. Vouchers: Luke Q 1521, Robertson & Luke
5328, Drummond & Hemsley 4119 (EA).
Justicia sp. aff. fittonioides S. Moore. Habit: Herb. Habitat: Forest, ca. 150 m.
Voucher: Luke WRQ & Robertson SA 5985 (EA): Endemic.
Justicia sp. F of FTEA Habit: Herb. Habitat: Lowland forest, 200–300 m. Voucher:
Robertson & Luke 5989 (EA): Endemic.
Justicia stachytarphetoides (Lindau) C.B. Clarke Habit: Shrub. Habitat: Lowland
forest, riverine forest, thicket, and swamp forest, 5–400 m. Vouchers: MG Gilbert
et al. 5909, Robertson & Luke 5337, BR Adams 78 (EA).
Justicia striata (Klotzsch) Bullock Habit: Herb. Habitat: Lowland to montane forest, ca. 239 m. Vouchers: Ngumbau V & Mwadime N V0114 & 0354 (EA, HIB),
Magogo FC & Glover PE 878, Drummond & Hemsley 3829 (EA).
Lankesteria alba Lindau Habit: Herb. Habitat: Forest, 25–500 m. Vouchers: SAJIT–006186 & 005933 (EA, HIB), Luke WRQ 870, Greenway & Rawlins 9351,
Luke & Robertson 2623, Drummond & Hemsley 1126 (EA).
Megalochlamys revoluta (Lindau) Vollesen subsp. revoluta Habit: Shrubby
herb. Habitat: Bushland or grassland thicket, ca. 250 m. Voucher: Bally PRO
16778 (EA).
26
Veronicah M. Ngumbau et al. / PhytoKeys 147: 1–191 (2020)
Megalochlamys tanaensis Vollesen Habit: Shrubby herb. Habitat: Lowland forest,
ca. 25 m. Vouchers: Gillett et al. 19922, Luke 10727 (EA): Critically Endangered, Endemic.
Megalochlamys trinervia (C.B. Clarke) Vollesen Habit: Shrubby herb. Habitat:
Woodland or bushland, 25–200(–400) m. Vouchers: Brenan et al. 14774, Greenway & Rawlins 9459, Bally PRO16750, Luke 10726, Luke TPR247 (EA).
Monothecium aristatum (Nees) T. Anderson Habit: Herb. Habitat: Lowland and medium altitude wet evergreen forest or swampy forest, ca. 200 m. Vouchers: Greenway & Rawlins 9358, K & L Holm 8, Robertson & Luke 5970, BR Adams 86 (EA).
Phaulopsis gediensis Mankt. Habit: Herb. Habitat: Lowland forest, thicket, and
shrubland, 0–500 m. Vouchers: Robertson SA 6575, Luke WRQ 3968, Drummond & Hemsley 1091, Luke et al. 3372 (EA).
Pseuderanthemum hildebrandtii Lindau Habit: Shrub. Habitat: Coastal lowland,
medium altitude forest, thicket, and bushland, ca. 199 m. Vouchers: Ngumbau V
& Mwadime M V0402 (EA, HIB), Meester V de 336, Magogo FC & Glover PE
721 (EA).
Pseuderanthemum subviscosum (C.B. Clarke) Stapf Habit: Herb. Habitat: Forest,
riverine forest, and bushland, ca. 365 m. Vouchers: Luke 3947, Glover et al. 1154,
Luke WRQ 886, Magogo & Glover 75 (EA).
Rhinacanthus dichotomus (Lindau) I. Darbysh. var. dichotomus Habit: Herb. Habitat:
Bushland, coastal thicket, margins of cultivation, and roadsides, ca. 199 m. Vouchers:
Ngumbau V & Mwadime N V0481 (EA, HIB), Festo L, Luke Q & P 2694 (EA).
Rhinacanthus pulcher Milne-Redh. Habit: Herb. Habitat: Bushland, 225–552 m.
Voucher: Gillett 21100 (EA).
Rhinacanthus rotundifolius C.B. Clarke Habit: Herb. Habitat: Bushland, 0–350 m.
Vouchers: Gillett 19527, Luke et al. in TPR 640 (EA).
Ruellia amabilis S. Moore Habit: Herb. Habitat: Lowland and riverine forest, 25–
450 m. Voucher: Gillett & Kibuwa 19949 (EA).
Ruellia patula Jacq. Habit: Herb. Habitat: Bushland, woodland, and riverine forest, 60–
400 m. Vouchers: Harvey & Vollens 52, Curtis A 13, Magogo & Glover 274 (EA).
Ruellia prostrata Poir. Habit: Herb. Habitat: Grassland, bushland, and woodland, ca.
10 m. Vouchers: Luke et al. TRP775, M Robertson 6363 (EA).
Ruellia tuberosa L. Habit: Herb. Habitat: Roadside or secondary grassland, ca.
150 m. Vouchers: Luke PA & WRQ 3076, Napier 6195, Luke 3076 (EA): Exotic.
Ruspolia hypocrateriformis (Vahl) Milne-Redh. Habit: Shrub. Habitat: Bushland,
0–100 m. Voucher: Bally 13702 (EA).
Sclerochiton boivinii (Baill.) C.B. Clarke Habit: Shrub. Habitat: Moist forest, 0–400
m. Vouchers: SAJIT–006181 & 005546, Ngumbau V & Mwadime N V012 (EA,
HIB), Luke WRQ 891, Dale 3664, Drummond & Hemsley 1143, Polhill & Robertson 4795 (EA).
Sclerochiton vogelii (Nees) T. Anderson subsp. holstii (Lindau) Napper Habit:
Shrub. Habitat: Sub-tropical and tropical moist lowland, 0–500 m. Vouchers: SAJIT–004648, Ngumbau V & Mwadime N V022 (EA, HIB).
Checklist of coastal forests of Kenya
27
Thunbergia alata Bojer ex Sims Habit: Climber. Habitat: Dry bushland to disturbed
and secondary vegetation, ca. 15 m. Vouchers: SAJIT–006206 (EA, HIB), Magogo FC & Glover PE 310, Fabus 23 (EA).
Thunbergia kirkii Hook. f. Habit: Shrub. Habitat: Evergreen, semi-evergreen lowland,
and lower montane forest, ca. 152 m. Vouchers: SAJIT–005541 (EA, HIB), Luke
WRQ 1628, Gardner 1407, Magogo & Glove 619, Robertson & Luke 6454 (EA).
Thunbergia schimbensis S. Moore Habit: Herb. Habitat: Woodland or wooded grassland, ca. 297 m. Vouchers: Ngumbau V & Mwadime N V0157 (EA, HIB), Luke
Q 1487, Drummond & Hemsley 4085, Verdcourt 1934 (EA).
Thunbergia stelligera Lindau Habit: Climber. Habitat: Lowland forest, 100–650 m.
Vouchers: Luke WRQ 3115, Luke & Robertson 2645 (EA).
Trichaulax mwasumbii Vollesen Habit: Herb. Habitat: Woodland or coastal thicket,
50–200 m. Vouchers: Ngumbau V & Mwadime N V017 (EA, HIB), Verdcourt
3607, Faden 71/800, Luke 1947 (EA).
Whitfieldia elongata (P. Beauv.) De Wild. & T. Durand Habit: Shrub. Habitat: Evergreen lowland and lower montane forest, ca. 150 m. Vouchers: Mwadime N 16, Luke
WRQ 888, Drummond & Hemsley 1146, Bally 13711, Faden RB 70/809 (EA).
Whitfieldia orientalis Vollesen Habit: Shrub. Habitat: Evergreen lowland forest,
220–370 m. Vouchers: Faden 71/767, Luke 893B, Luke 4729 (EA).
F19. Achariaceae
3 Genera, 3 Species
Lindackeria bukobensis Gilg Habit: Shrub. Habitat: Forest or forest edge, often in
open mixed forest, ca. 50 m. Voucher: J Adanson 271 (EA).
Rawsonia lucida Harv. & Sond. Habit: Tree. Habitat: Lowland and upland rainforest, dry evergreen forest, semi-swamp, and riverine forest, ca. 237 m. Vouchers:
SAJIT–006027, Ngumbau V & Mwadime N V0168 (EA, HIB), Luke WRQ &
Robertson SA 217 & 4821(EA).
Xylotheca tettensis (Klotzsch) Gilg var. kirkii (Oliv.) Wild Habit: Shrub. Habitat: Lowland woodland, bushland, and wooded grassland, 0–700 m. Vouchers: Ngumbau V
& Mwadime N V0218 & 0542 (EA, HIB), Dale in FD 3879, Luke 3083 (EA).
F20. Aizoaceae
3 Genera, 5 Species
Sesuvium portulacastrum (L.) L. Habit: Herb. Habitat: Maritime shores, coastal
dunes, and beaches, 0–200 m. Vouchers: Luke Q 6147, RM Graham 2221, Luke
PA & WRQ 5648 (EA).
Trianthema ceratosepala Volkens & Irmsch Habit: Herb. Habitat: Bushland, 135 m.
Voucher: Bally 16715 (EA).
Trianthema portulacastrum L. Habit: Herb. Habitat: Cultivated ground, 0–250 m.
Vouchers: Linder 2634, Verdcourt B 3617 (EA).
28
Veronicah M. Ngumbau et al. / PhytoKeys 147: 1–191 (2020)
Trianthema triquetra Willd. Habit: Herb. Habitat: Grassland, bushland, woodland,
and saline sands, ca. 5 m. Vouchers: Rawlins SP 828, Drummond & Hemsley
4090, Luke Q 6147 (EA).
Zaleya pentandra (L.) C. Jeffrey Habit: Herb. Habitat: Woodland, bushland or grassland, ca. 30 m. Vouchers: Reitsma J 413, Drummond & Hemsley 1046, Muchiri
J 490 (EA).
F21. Alismataceae
2 Genera, 2 Species
Burnatia enneandra Micheli Habit: Herb. Habitat: Edges of slow flowing rivers and
streams, shallow lakes, waterholes, and swamps, ca. 5 m. Voucher: Mwadime N
2573 (EA).
Limnophyton obtusifolium (L.) Miq. Habit: Herb. Habitat: Swamps & pools, 15 m.
Vouchers: Luke Q 5635, RM Graham 2164, Hildebrandt 1912 (EA).
F22. Amaranthaceae
20 Genera, 38 Species
Achyranthes aspera L. var. aspera Habit: Herb. Habitat: Disturbed areas, ca. 0–240 m.
Vouchers: Brown ES 1412, Robertson SA 3799, Hooper & Townsend 1163 (EA).
Achyranthes aspera var. sicula L. Habit: Herb. Habitat: Disturbed areas, ca. 0–300 m.
Voucher: Magogo FC & Glover PE 153 (EA).
Aerva lanata (L.) Juss. ex Schult. Habit: Herb. Habitat: Cultivated area, disturbed
area, bushland, and grassland, ca. 186 m. Vouchers: SAJIT–006137, Ngumbau V
& Mwadime N V0347 (EA, HIB), Luke WRQ 5468, Magogo FC & Glover PE
924, MacNaughton 96 (EA).
Allmaniopsis fruticulosa Suess. Habit: Herb. Habitat: Bushland, 110–200 m. Voucher: Gillett 16528 (EA).
Alternanthera pungens Kunth Habit: Herb. Habitat: Roadsides, grassland, ca.
0–200 m. Vouchers: Jeffery GW 522, Rawlins 839 (EA).
Alternanthera sessilis (L.) R.Br. ex DC. Habit: Herb. Habitat: Damp forests and
abandoned cultivation, ca. 0–300 m. Vouchers: Magogo FC & Glover PE 110,
Drummond & Hemsley 4174 (EA).
Amaranthus cruentus L. Habit: Herb. Habitat: Cultivated lands, forest margins, ca.
20 m. Vouchers: Wakefield, Jeffery GW 431 (EA).
Amaranthus dubius Mart. ex Thell. Habit: Herb. Habitat: Cultivated area, waste places,
roadsides, and grassland, ca. 0–50 m. Voucher: Jeffery GW 441 (EA): Naturalized.
Amaranthus graecizans subsp. silvestris (Vill.) Brenan Habit: Herb. Habitat: Cultivated ground, grassland, ca. 0–350 m. Voucher: Luke WRQ & PA 5694 (EA).
Amaranthus spinosus L. Habit: Herb. Habitat: Cultivated ground ca. 2 m. Voucher:
Mwadime N & Chesire C 256 (EA).
Amaranthus viridis L. Habit: Herb. Habitat: Cultivated and disturbed areas, ca. 2 m.
Voucher: Sturrock 2008 B (EA).
Checklist of coastal forests of Kenya
29
Atriplex farinosa subsp. keniensis (Brenan) Friis & M.G. Gilbert Habit: Shrub.
Habitat: In sand at or near high tide mark. Vouchers: Festo L, Luke Q & P 2794,
Graham 2166 (EA).
Atriplex semibaccata R.Br. Habit: Herb. Habitat: Cultivated area, ca. 10 m. Voucher:
Kennox M 121 (EA): Introduced.
Celosia argentea L. Habit: Herb. Habitat: Cultivated lands, grassland, rivers, ca.
610 m. Vouchers: Weruer 993, Greenway & Rawlins 9461 (EA).
Celosia hastata Lopr. Habit: Herb. Habitat: Bushland and woodland, 3–106 m.
Vouchers: Ngumbau V & Mwadime N V0482 (EA, HIB), Festo L & Luke Q
2675, Magogo FC & Glover PE 914 (EA).
Celosia schweinfurthiana Schinz Habit: Herb. Habitat: Lowland forest, clearings,
especially near water, ca. 0–50 m. Vouchers: Robertson SA & Luke 5859, Magogo
FC & Glover PE 915 (EA).
Celosia stuhlmanniana Schinz Habit: Herb. Habitat: Shaded places, forest, and
scrub, ca. 60 m. Voucher: Nyange M 508 (EA).
Celosia trigyna L. Habit: Herb. Habitat: Cultivated lands or forest clearings, 0–100 m.
Vouchers: Jeffery GW 445, McCrag in CM 9170 (EA).
Centemopsis gracilenta (Hiern) Schinz Habit: Herb. Habitat: Woodland or cultivated lands, ca. 50 m. Voucher: Rawlins 776 (EA).
Centrostachys aquatica (R.Br.) Moq. Habit: Herb. Habitat: Pools, waterholes, rivers,
and lake shores, ca. 20 m. Voucher: Hooper & Townsend 1227 (EA).
Cyathula braunii Gilg ex Schinz Habit: Herb. Habitat: Forest undergrowth, ca.
360 m. Vouchers: SAJIT–005931, Ngumbau V & Mwadime N V0368 (EA,
HIB), Gillett 18714, Luke WRQ et al. 3328 (EA).
Cyathula coriacea Schinz Habit: Herb. Habitat: Thicket, woodland, and roadside,
30–300 m. Voucher: Greenway 8864 (EA).
Cyathula prostrata (L.) Blume var. prostrata Habit: Herb. Habitat: Deciduous to
evergreen vine thickets, ca. 304 m. Vouchers: SAJIT–006155, Ngumbau V &
Mwadime N V0294 (EA, HIB), Luke WRQ 4714 (EA).
Digera muricata (L.) Mart. subsp. muricata Habit: Herb. Habitat: Waste ground,
grassland, and bushland, ca. 150 m. Vouchers: RM Graham in FD 648 & 1993 (EA).
Gomphrena celosioides Mart. Habit: Herb. Habitat: Roadsides, coastal grassland, and
cultivated area, ca. 50–300 m. Vouchers: Festo L, Luke Q & P 2703, Hooper &
Townsend 1168, Robertson SA 3480 (EA).
Gomphrena globosa L. Habit: Herb. Habitat: On grassland, ca. 100 m. Vouchers:
Jeffery GW 468, WE Taylor (EA): Naturalized.
Hermbstaedtia gregoryi C.B. Clarke Habit: Herb. Habitat: Sand dune, 20 m. Vouchers: Frazier J 1101, Moomaw 1562 (EA): Endemic.
Neocentema alternifolia (Schinz) Schinz Habit: Herb. Habitat: Roadside or cultivation area, 0–50 m. Voucher: Kabuye CHS, Gilbet VC & Robertson 43 (EA).
Nothosaerva brachiata (L.) Wight Habit: Herb. Habitat: Waste ground, grassland,
and bushland, ca. 300 m. Vouchers: Thuiru 78, Luke WRQ 3501 (EA).
Psilotrichum cyathuloides Suess. & Launert Habit: Herb. Habitat: Forests, 0–150 m.
Voucher: Rawlins 394 (EA).
30
Veronicah M. Ngumbau et al. / PhytoKeys 147: 1–191 (2020)
Psilotrichum fallax C.C. Towns. Habit: Herb. Habitat: Forests on limestone,
0–380 m. Vouchers: Ngumbau V and Mwadime N V109 (EA, HIB), Magogo &
Glover 4141, Adams 71, Herbarium Techniques Course III 078 (EA).
Psilotrichum gnaphalobryum (Hochst.) Schinz Habit: Herb. Habitat: Open Acacia
and Acacia-Commiphora bushland, 140–790 m. Voucher: Luke PA, WRQ et al.
3833 (EA).
Psilotrichum majus Peter Habit: Herb. Habitat: Forest undergrowth and margins,
ca. 285 m. Vouchers: Ngumbau V & Mwadime N V0122 (EA, HIB), Luke PA,
WRQ 4306, Magogo FC & Glover PE 79 & 261 (EA).
Psilotrichum scleranthum Thwaites Habit: Herb. Habitat: Forest margin, grassland,
and coconut plantation, ca. 417 m. Vouchers: Ngumbau V & Mwadime N V0318
(EA, HIB), Luke Q & Robertson SA 5495, Ross KS 142 (EA).
Psilotrichum sericeum (J. Koenig ex Roxb.) Dalzell Habit: Herb. Habitat: Abandoned land, grassland, and bushland, 15–420 m. Vouchers: Robertson SA 3801,
Pakia M 113, GM Mungai & SM Rucina 307 (EA).
Pupalia lappacea var. argyrophylla C.C. Towns. Habit: Herb. Habitat: Coastal
bushland, grassland, and cultivated area, 0–530 m. Vouchers: SAJIT–006184,
Ngumbau V & Mwadime N V0319 (EA, HIB), Festo L & Luke Q 2525, Magogo
& Glover 125, Moggridge 157, Gillespie 383 (EA).
Pupalia lappacea var. velutina (Moq.) Hook.f. Habit: Herb. Habitat: Woodland,
grassland, thickets or roadsides, ca. 200 m. Vouchers: Luke WRQ & Robertson SA
1528, Luke WRQ 3327 (EA).
Salsola africana (Brenan) Botsch. Habit: Herb. Habitat: Semi-desert scrub, ca.
90 m. Voucher: Battiscombe 223 (EA).
Tecticornia indica (Willd.) K. Sheph. & Paul G. Wilson Habit: Herb. Habitat: Salt
marshes by the sea and mangrove swamps, ca. 5 m. Vouchers: Gillett JB 20439,
Luke Q 6149 (EA).
Volkensinia prostrata (Volkens ex Gilg) Schinz Habit: Herb. Habitat: Bushland or
grassland, ca. 180 m. Voucher: Gillett 19505 (EA).
F23. Amaryllidaceae
4 Genera, 5 Species
Crinum politifolium R. Wahlstr. Habit: Herb. Habitat: Riverine vegetation or damp
places, ca. 86 m. Vouchers: Ngumbau V & Mwadime N V0480 (EA, HIB), Luke
WRQ & PA 4581 (EA).
Crinum stuhlmannii Baker subsp. stuhlmannii Habit: Herb. Habitat: Grassland
or wooded grassland, ca. 0–300 m. Vouchers: Magogo & Glover 468, Jeffery &
Kirika 251, Graham K. 1844 (EA).
Cryptostephanus haemanthoides Pax Habit: Herb. Habitat: Deciduous bushland, ca.
20 m. Vouchers: Luke TPR 574, Robertson SA 5371D (EA).
Cyrtanthus sanguineus subsp. wakefieldii (Sealy) Nordal Habit: Herb. Habitat:
Woodland, ca. 0–250 m. Vouchers: Walker in Bally 7406, Wakefield 42 (EA).
Checklist of coastal forests of Kenya
31
Scadoxus multiflorus (Martyn) Raf. subsp. multiflorus Habit: Herb. Habitat:
Grassland, riverine vegetation, coastal rocks or montane vegetation, ca. 0–350 m.
Voucher: Magogo FC & Glover PE 367 (EA).
F24. Anacardiaceae
7 Genera, 13 Species
Anacardium occidentale L. Habit: Tree. Habitat: Cultivated area, abandoned cultivated areas, ca. 250 m. Vouchers: RE Perdue & SP Kibuwa 10116, Napier 3274,
R Karisa 24 (EA): Naturalized.
Lannea alata (Engl.) Engl. Habit: Tree. Habitat: Bushland and coastal forests edges,
ca. 88–450 m. Vouchers: SAJIT–006442 (EA, HIB), Dale IR 3885, Polhill &
Paulo 598, Luke WRQ 950 (EA).
Lannea cotoneaster (Chiov.) Sacleux Habit: Shrub. Habitat: Bushland, 75–600 m.
Voucher: Njoroge Thairu 86 (EA).
Lannea schweinfurthii var. acutifoliolata (Engl.) Kokwaro Habit: Tree. Habitat: Coastal forests, 1–460 m. Vouchers: SAJIT–004658 & 004663, Ngumbau V & Mwadime
N V0361 & 0333 (EA, HIB) Greenway 9799, RM Graham in FD 1784, Gilbert
MG & Kuchar P 5866, Luke WRQ & Robertson SA 2775 (EA): Near Threatened.
Lannea schweinfurthii var. stuhlmannii (Engl.) Kokwaro Habit: Tree. Habitat:
Woodland, bushland, dry forests, and river valleys, ca. 20–450 m. Vouchers:
Drummond & Hemsley 4166, Luke WRQ 920, Robertson SA 3498 (EA).
Lannea triphylla (Hochst. ex A. Rich.) Engl. Habit: Tree. Habitat: Grassland, and
bushland, ca. 70 m. Vouchers: Drummond & Hemsley 4102, JB Gillet 16486 (EA).
Lannea welwitschii var. ciliolata Engl. Habit: Tree. Habitat: Lowland rainforest,
riverine forest, and mountain slopes, 110–450 m. Vouchers: Verdcourt in EAH
11322, Magogo & Glover 177, Faden & Faden 74/318 (EA).
Mangifera indica L. Habit: Tree. Habitat: Cultivated area, abandoned cultivated areas, ca. 100 m. Voucher: Bally 13945 (EA): Cultivated.
Ozoroa insignis subsp. reticulata (Baker f.) J.B. Gillett Habit: Tree. Habitat: Wooded grassland, ca. 20–400 m. Vouchers: Magogo FC & Glover PE 133, RM Graham in CM 14563 (EA).
Ozoroa obovata (Oliv.) R. Fern. & A. Fern. Habit: Tree. Habitat: Bushland, woodland, and forest edges, ca. 88 m. Vouchers: SAJIT–006448, 005484 (EA, HIB),
Robertson SA 4240, Robertson SA 7761, Jeffery 150, Rawlins 654 (EA).
Sclerocarya birrea subsp. caffra (Sond.) Kokwaro Habit: Tree. Habitat: Wooded
grassland, riverine woodland, and bushland, ca. 5 m. Vouchers: SAJIT–006111
(EA, HIB), Sangai GW 15779, Gray M & Luke WRQ 271 (EA).
Sclerocarya gillettii Kokwaro Habit: Tree. Habitat: Bushland, 150–300 m. Voucher:
Robertson 1846 (EA).
Searsia crenulata (A. Rich.) Moffett Habit: Tree. Habitat: Evergreen bushland,
woodland, riverine associations, and forests edges, ca. 1–300 m. Vouchers: Ngumbau V & Mwadime N V0359, Robertson SA 4217, Jeffrey 154 (EA).
32
Veronicah M. Ngumbau et al. / PhytoKeys 147: 1–191 (2020)
Sorindeia madagascariensis DC. Habit: Tree. Habitat: Riverine, coastal, and upland forests, ca. 0–297 m. Vouchers: SAJIT–006085 & 005918, Ngumbau V &
Mwadime N V073 (EA, HIB), Mwadime N 11, Magogo FC & Estes R 1195, RM
Graham in FD 1575, Gillet & Kibuwa 19945, Greenway & Rawlins 9356 (EA).
F25. Ancistrocladaceae
1 Genus, 1 Species
Ancistrocladus robertsoniorum J. Léonard Habit: Liana. Habitat: Forest in disturbed
areas, 60–80 m. Vouchers: Luke WRQ 1641, Drummond & Hemsley 3936, Polhill & Robertson 4786, Robertson 3649 (EA): Endemic.
F26. Annonaceae
17 Genera, 37 Species
Annona senegalensis Pers. subsp. senegalensis Habit: Shrub. Habitat: Grassland,
woodland and forest, ca. 20–300 m. Vouchers: SAJIT–005602 & 005547, Ngumbau V & Mwadime N V0283 (EA, HIB), Katz SS 21, Rawlins S 645, Magogo FC
& Glover PE 50 (EA).
Artabotrys brachypetalus Benth. Habit: Liana. Habitat: Riverine vegetation, sometimes in dry woodland, 0–150 m. Voucher: Festo L, Luke Q & P 2724 (EA).
Artabotrys modestus subsp. macranthus Verdc. Habit: Liana. Habitat: Forest, woodland to grassland, 60–490 m. Vouchers: Ngumbau V & Mwadime N V0431 &
V0320 (EA, HIB), Festo L, Luke Q & P 2753, Luke WRQ 3104 (EA).
Artabotrys monteiroae Oliv. Habit: Liana. Habitat: Moist forest edge or moist grassland, ca. 270 m. Vouchers: Ngumbau V & Mwadime N V0504, V0536 & V040
(EA, HIB), Luke, Kibet & Amini 6303, Robertson SA, Beentje HJ & Khayota 21
& 94 (EA).
Asteranthe asterias (S. Moore) Engl. & Diels subsp. asterias Habit: Shrub. Habitat:
Forest, 0–60 (–500) m. Vouchers: SAJIT–005449 & 006182, Ngumbau V &
Mwadime N V0236 & V0417 (EA, HIB), Verdcourt B 11355, Luke WRQ &
Robertson SA 524 (EA).
Cananga odorata (Lam.) Hook. f. & Thomson Habit: Tree. Habitat: Plantations,
0–750 m. Vouchers: Jeffery GW 365, RM Graham in FD 2342 (EA): Cultivated.
Huberantha stuhlmannii (Engl.) Chaowasku Habit: Shrub. Habitat: Shrubland,
forest, and savannah, under 100 m. Vouchers: Festo L & Luke Q 2521, Rawlins
262 & 303B (EA): Near Threatened.
Isolona cauliflora Verdc. Habit: Tree. Habitat: Moist forests to evergreen bushland,
20–500 m. Vouchers: SAJIT–005564 (EA, HIB), Luke WRQ & Robertson SA
1803, Luke WRQ 2930 (EA): Vulnerable.
Lettowianthus stellatus Diels Habit: Tree. Habitat: Forest and riverine forest, 10–
280 m. Vouchers: SAJIT–005578 (EA, HIB), Luke WRQ & Robertson SA 2768,
Luke WRQ 3304 (EA).
Checklist of coastal forests of Kenya
33
Mkilua fragrans Verdc. Habit: Shrub. Habitat: Moist forests, 0–400 m. Vouchers: SAJIT–006082, 005532, 005535 & 005919, Ngumbau V & Mwadime N
V059 (EA, HIB), Faden RB & Faden AJ 77/572, Robertson SA, Luke WRQ 235
(EA): Vulnerable.
Monanthotaxis buchananii (Engl.) Verdc. Habit: Liana. Habitat: Forest, bushland,
and grassland, 100–400 m. Vouchers: SAJIT–004645 & 006002, Ngumbau V
& Mwadime N V0301 (EA, HIB), Robertson SA & Luke WRQ 4779, 2743A,
Drummond & Hemsley 4485, Verdcourt 1953 (EA).
Monanthotaxis faulknerae Verdc. Habit: Liana. Habitat: Forest and savannah, 30–
370 m. Vouchers: Festo L, Laizer G & P 2635, Luke WRQ 3351, Luke WRQ, PA
3822 (EA): Endangered.
Monanthotaxis fornicata (Baill.) Verdc. Habit: Liana. Habitat: Forest, woodland,
and bushland, 0–450 m. Vouchers: SAJIT–005447, 005448 & 005599 (EA,
HIB), Festo L, Luke Q & P 2635, Battiscombe E 1044, Robertson SA 4252 (EA).
Monanthotaxis trichocarpa (Engl. & Diels) Verdc. Habit: Liana. Habitat: In evergreen
forest, 100–300 m. Vouchers: SAJIT–005920, Ngumbau V & Mwadime N V0302
(EA, HIB), Robertson SA 3697, Verdcourt 1895, Magogo & Glover 288 (EA).
Monodora grandidieri Baill. Habit: Shrub. Habitat: Forest, bushland, and grassland,
10–290 m. Vouchers: Ngumbau V & Mwadime N V0434 (EA, HIB), Festo L,
Luke Q & P 2804, Dale 2030, Kokwaro JO 3974, Robertson SA 4263 (EA).
Monodora junodii Engl. & Diels Habit: Shrub. Habitat: On rocky outcrops, woodland, and dry areas, 10–15 m. Vouchers: Robertson SA 5708 & 5889, Faden RB
71/273, Luke WRQ 3560 (EA).
Ophrypetalum odoratum Diels Habit: Shrub. Habitat: Woodland, forest, 0–600 m.
Vouchers: Verdcourt B 1075 & 3952, Drummond & Hemsley 3981, Greenway
PJ 9634 (EA): Near Threatened.
Polyceratocarpus sp. ?nov. Habit: Tree. Habitat: Forest, 300–370 m. Vouchers: Luke
1621 & 881, Luke & Stone 8245 (EA): Endemic.
Sphaerocoryne gracilis (Engl. & Diels) Verdc. subsp. gracilis Habit: Shrub. Habitat:
Moist to dry forest, and shrubland, 30–350 m. Vouchers: SAJIT–005922, Ngumbau V & Mwadime N V0315 (EA, HIB), Luke WRQ & Robertson SA 2597,
Robert SA & Luke WRQ 5190 (EA).
Toussaintia orientalis Verdc. Habit: Tree. Habitat: Shrubland and forest, 140–610 m.
Voucher: Luke WRQ & Robertson SA 1038 (EA): Endangered.
Uvaria acuminata Oliv. Habit: Liana. Habitat: Bushland, forest, and woodland, 0–810
m. Vouchers: SAJIT–005496, Ngumbau V & Mwadime N V0161 (EA, HIB),
Kuchar P 13454, Robertson SA 6456, Luke WRQ & Robertson SA 516 (EA).
Uvaria denhardtiana Engl. & Diels Habit: Shrub. Habitat: Open scrub or coastal
bushland, 0–300 m. Vouchers: Kirika P, Mbale M & Mbatha M 780, J Adamson
260 (EA): Near Threatened.
Uvaria faulknerae Verdc. Habit: Liana. Habitat: Bushland or woodland, 45–180 m.
Vouchers: Robertson SA & Luke WRQ 5688, Verdcourt 3605, Luke WRQ 118
& 3468 (EA): Endangered.
34
Veronicah M. Ngumbau et al. / PhytoKeys 147: 1–191 (2020)
Uvaria kirkii Oliv. ex Hook. f. Habit: Shrub. Habitat: Bushland, grassland, and
woodland, 0–450 m. Vouchers: Ngumbau V & Mwadime N V0467 (EA, HIB),
Festo L, Luke Q 2557, Rawlins 361(EA): Near Threatened.
Uvaria leptoclados Oliv. subsp. leptoclados Habit: Shrub. Habitat: Bushland, 30–
457 m. Vouchers: Hawthorne W 510, Hildebrandt 1971, RM Graham in FD
1763 (EA).
Uvaria lucida Benth. subsp. lucida Habit: Shrub. Habitat: Forest, grassland, and
bushland, ca. 0–400 m. Vouchers: SAJIT–005460, Ngumbau V & Mwadime N
V0296 (EA, HIB), Robertson SA 4775, Muchiri J 514, Robertson SA 4232 (EA).
Uvaria puguensis D.M. Johnson Habit: Liana. Habitat: Evergreen coastal forest,
200–390 m. Vouchers: Ngumbau V & Mwadime N V060 & 0364 (EA, HIB),
Nyange M 525 (EA): Critically Endangered.
Uvaria welwitschii (Hiern) Engl. & Diels Habit: Liana. Habitat: Evergreen coastal
forests, 2–300 m. Vouchers: SAJIT–005582, Ngumbau V & Mwadime M V0472
(EA, HIB), Festo L & Luke Q 2526, Verdcourt 2124, Robertson SA 2786, Luke
WRQ 536 (EA).
Uvariodendron gorgonis Verdc. Habit: Tree. Habitat: Lowland rainforest, 180–800
m. Vouchers: Ngumbau V & Mwadime M V0528 (EA, HIB), Brenan JPM, Brenan JH, Gillett JB, Kanuri K & Chomba W 14601, Verdcourt B 1890 & 1911
(EA): Endangered.
Uvariodendron kirkii Verdc. Habit: Shrub. Habitat: Evergreen coastal forest, often on
coral, 5–450 m. Vouchers: SAJIT–005580 & 005445, Ngumbau V & Mwadime
N V0516 (EA, HIB), Luke WRQ 1044 & 3408 (EA): Vulnerable.
Uvariodendron schmidtii Q. Luke, ined. Habit: Tree. Habitat: Lowland rainforest,
ca. 390 m. Voucher: Luke WRQ 2919 (EA): Endemic.
Uvariodendron sp. nov. 1 of CFS Habit: Tree. Habitat: Forest. Voucher: Luke 1654 (EA).
Uvariodendron sp. nov. 2 of CFS Habit: Tree. Habitat: Forest. Voucher: Luke
2929 (EA).
Xylopia aethiopica (Dunal) A. Rich. Habit: Tree. Habitat: Forest, 40–280 m. Vouchers: Luke WRQ 3378, Luke WRQ & Robertson SA 2709 (EA).
Xylopia arenaria Engl. Habit: Shrub. Habitat: Woodland and open coastal bushland,
30–600 m. Vouchers: Luke WRQ & Robertson SA 2134, 6000 & 1686, Moggridge 389, Gisau 10, Trump 96 (EA): Vulnerable.
Xylopia holtzii Engl. Habit: Tree. Habitat: Forest and woodland, 50–400 m. Vouchers: SAJIT–005445, Ngumbau V & Mwadime N V0257 (EA, HIB), Robertson
SA & Abio Gafo L 6812 (EA).
Xylopia keniensis D. M. Johnson Habit: Tree. Habitat: Coastal forest, ca. 450 m. Vouchers: Luke 13949, Tim P et al. NMK 722, Luke & Robertson 2723 (EA): Endemic.
F27. Apiaceae
2 Genera, 2 Species
Centella asiatica (L.) Urb. Habit: Herb. Habitat: Damp grassland along river and forest clearings, ca. 150 m. Voucher: Luke WRQ & Robertson SA 2753 (EA).
Checklist of coastal forests of Kenya
35
Foeniculum vulgare Mill. Habit: Herb. Habitat: Naturalized in disturbed places, ca.
30 m. Voucher: Rexton K 32.F (EA): Exotic.
F28. Apocynaceae
53 Genera, 101 Species
Adenium obesum (Forssk.) Roem. & Schult. Habit: Shrub. Habitat: Woodland and
dry forest, 20–700 m. Vouchers: SAJIT–004679 & 005913 (EA, HIB), Greenway
PJ & Rawlins SP 8924 (EA).
Alafia caudata Stapf Habit: Liana. Habitat: Forest and bushland, 0–800 m. Vouchers: Ngumbau V & Mwadime N V0378 (EA, HIB), Magogo & Glover 791, Luke
& Robertson 1680 & 985 (EA).
Alafia microstylis K. Schum. Habit: Liana. Habitat: Forest, riverine forest, and bushland, 30–250 m. Vouchers: Mwadime N 22, Luke WRQ & Robertson SA 2818,
Robertson & Luke 5505 (EA).
Ancylobothrys petersiana (Klotzsch) Pierre Habit: Liana. Habitat: Forest, bushland,
and thicket, ca. 50 m. Vouchers: SAJIT–004643, 005430 & 005590 (EA, HIB),
Robertson SA 4227, Beentje 2286, Kuchar 13531 (EA).
Ancylobothrys tayloris (Stapf ) Pichon Habit: Liana. Habitat: Forest, riverine forest,
and bushland, ca. 110 m. Vouchers: Ngumbau V & Mwadime N V0488 & 0358
(EA, HIB), Luke WRQ et al. 6196, Drummond & Hemsley 3935 (EA).
Asclepias aurea (Schltr.) Schltr. Habit: Herb. Habitat: Grassland or waterlogged
grassland, ca. 0–200 m. Vouchers: SAJIT–005518 (EA, HIB), Magogo FC &
Glover PE 120 (EA).
Asclepias curassavica L. Habit: Herb. Habitat: Forest and grassland, 0–50 m. Voucher: Jeffery K332 (EA): Naturalized.
Aspidoglossum masaicum (N.E.Br.) Kupicha Habit: Herb. Habitat: Forest, grassland, ca. 206 m. Vouchers: Ngumbau V & Mwadime N V0206 (EA, HIB), Luke
WRQ 3110 (EA).
Baissea myrtifolia (Benth.) Pichon Habit: Liana. Habitat: Forest and bushland,
0–450 m. Vouchers: Ngumbau V & Mwadime N V0325 & 0104 (EA, HIB),
Dale 3580, Luke 1602, Musyoki & Hansen 1004 (EA).
Baseonema gregoryi Schltr. & Rendle Habit: Liana. Habitat: Dry bushland or evergreen forest, ca. 620 m. Voucher: Mwadime N 4 (EA).
Calotropis gigantea (L.) W. T. Aiton Habit: Shrub. Habitat: Bare and degraded lands,
1–100 m. Vouchers: SAJIT–005440 (EA, HIB), Gloves 15300, Tweedie 4032 (EA).
Calotropis procera (Aiton.) W. T. Aiton Habit: Shrub. Habitat: Degraded roadsides,
ca. 65 m. Voucher: SAJIT–006439 (EA, HIB).
Caralluma arachnoidea (P.R.O. Bally) M.G. Gilbert var. arachnoidea Habit: Herb.
Habitat: Grassland, bushland, and riverine scrub, 200–300 m. Voucher: Robertson SA & Luke WRQ 6029 (EA).
Caralluma arachnoidea var. breviloba (P.R.O. Bally) M. G. Gilbert Habit: Herb.
Habitat: Grassland, bushland, and riverine scrub, ca. 400 m. Voucher: Luke WRQ
& Robertson SA2109 (EA).
36
Veronicah M. Ngumbau et al. / PhytoKeys 147: 1–191 (2020)
Carissa bispinosa (L.) Desf. ex Brenan Habit: Shrub. Habitat: Forest and woodland,
0–400 m. Vouchers: Luke Q 1469, Robertson & Luke 5367, Luke & Robertson
2144 (EA).
Carissa spinarum L. Habit: Shrub. Habitat: Bushland, riverine forest, and thicket.
Voucher: Luke WRQ & PA sr.
Carissa tetramera (Sacleux) Stapf Habit: Shrub. Habitat: Forest, thicket, and bushland, 0–450 m. Vouchers: Luke Q 1519, Drummond & Hemsley 4179 (EA).
Carvalhoa campanulata K. Schum. Habit: Shrub. Habitat: Moist forest, ca. 350 m.
Vouchers: Ngumbau V & Mwadime N V0311 (EA, HIB), Luke WRQ & Robertson SA 2727, Robertson 4673 (EA).
Catharanthus roseus (L.) G. Don Habit: Herb. Habitat: Roadsides in dry savannah,
urban open spaces, and in cultivated areas, ca. 100 m. Voucher: Spjut RW & Ensor
PD 2631 (EA).
Ceropegia abyssinica Decne. var. abyssinica Habit: Climber. Habitat: Bushland, 80
m. Voucher: Nyange M 371 (EA).
Ceropegia ampliata E. Mey. var. oxyloba H. Huber Habit: Climber. Habitat: Coastal
forest edges, bushland, and rocky ground, ca. 115 m. Voucher: Archer PG 401 (EA).
Ceropegia aristolochioides Decne. Habit: Climber. Habitat: Bushland, thicket, and
dry forest, ca. 15 m. Vouchers: Luke 4052, Kabuye CHS, Gilbert VC & Robertson
SA 84/66 (EA).
Ceropegia distincta N.E.Br. Habit: Climber. Habitat: Bushland, coral reef, woodland, and riverine, 0–100 m. Vouchers: Hawthorne 217, Masinde 860, Bally
12191 (EA).
Ceropegia galeata H. Huber Habit: Climber. Habitat: Bushland, 50–250 m. Vouchers: Bally 12184 & S220, Bayliss 3 (EA).
Ceropegia inornata P.R.O. Bally ex Masinde Habit: Climber. Habitat: Coastal forest and bushland, ca. 50 m. Vouchers: Robertson et al. 6742, Masinde 814 (EA).
Ceropegia konasita Masinde Habit: Climber. Habitat: Bushland and thickets, ca.
60 m. Voucher: Mbinda & Pakia 218 (EA).
Ceropegia nilotica Kotschy Habit: Climber. Habitat: Mixed deciduous woodland,
ca. 300 m. Vouchers: Robertson SA & Luke WRQ 6060, Moomaw JC 1616,
Masinde 816 (EA).
Ceropegia racemosa var. voiensis Masinde Habit: Herb. Habitat: Bushland and
thicket, ca. 50–650 m. Voucher: Archer 402 (EA).
Ceropegia seticorona var. dilatiloba P.R.O. Bally Habit: Climber. Habitat: Bushland, 50 m. Vouchers: Kabuye CHS, Gilbert VC & Robertson SA 84/66, Drummond & Hemsley 3889 (EA).
Ceropegia somalensis Chiov. Habit: Climber. Habitat: Bushland, ca. 300 m. Voucher: Adams 145 (EA).
Chlorocyathus monteiroae Oliv. Habit: Climber. Habitat: Bushland, 300–800 m. Vouchers: Drummond & Hemsley 4227, Robertson & Luke 6167, Bock H 16047 (EA).
Cryptolepis africana (Bullock) Venter & R.L. Verh. Habit: Climber. Habitat: Coastal lowland forest, 0–100 m. Vouchers: Drummond & Hemsley 3836, Luke &
Mbinda 5978, Festo L, Luke Q & P 2784 (EA).
Checklist of coastal forests of Kenya
37
Cryptolepis apiculata K. Schum. Habit: Climber. Habitat: Lowland forest and
shrubby thickets in grassland, 30–350 m. Vouchers: Ngumbau V & Mwadime N
V0373 & V0211 (EA, HIB), Saufferer 788, Kimeu JM 688, Graham 1526 (EA).
Cryptolepis hypoglauca K. Schum. Habit: Climber. Habitat: Moist evergreen lowland and coastal forest, ca. 220 m. Vouchers: Ngumbau V & Mwadime N V0326
(EA, HIB), Drummond & Hemsley 3917, Rawlins 892 (EA).
Cryptolepis nigrescens (Wennberg) L. Joubert & Bruy Habit: Liana. Habitat: Secondary forest and savannah, ca. 106 m. Voucher: Bally PRO 1995 (EA).
Cynanchum crassiantherae Liede Habit: Climber. Habitat: Bushland and coastal
dunes, near sea level. Voucher: Festo et al 2777 (EA).
Cynanchum gerrardii (Harv.) Liede subsp. gerrardii Habit: Climber. Habitat: Dry
bushland, 2 m. Vouchers: SAJIT–006246 (EA, HIB), Luke Q 1471 (EA).
Cynanchum hastifolium subsp. longirostrum Goyder Habit: Climber. Habitat:
Bushland, ca. 200 m. Voucher: Tweedie 3199 (EA).
Cynanchum insipidum (E. Mey.) Liede & Khanum Habit: Herb. Habitat: Open
grassland, bushland, woodland, and forest, ca. 20 m. Vouchers: SA Robertson
6721, Magogo FC & Glover PE 967 (EA).
Cynanchum resiliens (B.R. Adams & R.W.K. Holland) Goyder Habit: Climber. Habitat:
Bushland, 40–500 m. Vouchers: Drummond & Hemsley 4060, Gillett 16473 (EA).
Cynanchum viminale (L.) L. subsp. viminale Habit: Climber. Habitat: Dry scrub,
rocky area, woodland, and riverine, ca. 590 m. Vouchers: Luke WRQ & Robertson SA 1471, Adams 146, Makin s.n in EA 13047 (EA).
Cynanchum viminale subsp. odontolepis (Balf.f.) Goyder (Balf.f.) Goyder Habit:
Climber. Habitat: Dry coastal forest or woodland, 0–200 (–800) m. Vouchers:
SAJIT–006446, Ngumbau V & Mwadime N V0494 (EA, HIB), Adams 144,
Robertson & Luke 5611 (EA).
Desmidorchis retrospiciens Ehrenb. Habit: Herb. Habitat: Semi-desert, ca. 0–100 m.
Voucher: Gillett 19504 (EA).
Dictyophleba lucida (K. Schum.) Pierre Habit: Liana. Habitat: Forest, riverine, ca.
410 m. Vouchers: Ngumbau V & Mwadime N V0308 & 058, SAJIT–005510
(EA, HIB), Luke WRQ & Robertson SA 234 (EA).
Echidnopsis ericiflora Lavranos Habit: Herb. Habitat: Dry bushland, 250–450 m.
Voucher: Powys & Heath 793 (EA): Endemic.
Echidnopsis sharpei A.C. White & B. Sloane Habit: Herb. Habitat: Rocky slopes and
woodland, ca. 450 m. Vouchers: Faden & Faden 74/1045, Luke WRQ 919 (EA).
Edithcolea grandis N.E.Br. Habit: Herb. Habitat: Dry woodland, mostly on rocky
ground, 200 m. Vouchers: SAJIT–005910 (EA, HIB), Karisa R 40 (EA).
Fockea angustifolia K. Schum. Habit: Climber. Habitat: Bushland and coastal thicket,
ca. 350 m. Vouchers: Luke WRQ et al. 6188, Drummond & Hemsley 4045 (EA).
Funtumia africana (Benth.) Stapf Habit: Tree. Habitat: Forest and riverine, 30–
150 m. Voucher: Luke WRQ 890 (EA).
Gymnema sylvestre (Retz.) R.Br. ex Sm. Habit: Liana. Habitat: Forest margins and
dry bushland, ca. 410 m. Vouchers: Ngumbau V & Mwadime N V0307 & V039
(EA, HIB), Cunningham-van Someren GR Sh 14, MD Graham 1970 (EA).
38
Veronicah M. Ngumbau et al. / PhytoKeys 147: 1–191 (2020)
Holarrhena pubescens Wall. ex G. Don Habit: Tree. Habitat: Forest, riverine, woodland, and bushland, ca. 407 m. Vouchers: SAJIT–006064 (EA, HIB), Luke Q
1409, Gachathi 50/88 (EA).
Huernia andreaeana (Rauh) L.C. Leach Habit: Herb. Habitat: Dry deciduous bushland, ca. 400 m. Voucher: Rauh Ke867 (EA).
Huernia archeri L.C. Leach Habit: Herb. Habitat: Rocky bushland, 200–300 m.
Vouchers: Luke & Robertson 2546 & 6041, Robertson 7061 (EA): Endemic.
Hunteria zeylanica (Retz.) Gardner ex Thwaites Habit: Tree. Habitat: Coral rag
forest, riverine forest, and coastal bushland, 0–350 m. Vouchers: Mwadime N 33,
Omino EA 89 (EA).
Kanahia laniflora (Forssk.) R.Br. Habit: Shrub. Habitat: Seasonal watercourses.
Voucher: Kibuwa 2465 (EA).
Landolphia eminiana K. Schum. ex Hallier f. Habit: Liana. Habitat: Riverine,
woodland, and thicket, ca. 300 m. Voucher: Lap LJ 307 (EA).
Landolphia kirkii Dyer Habit: Liana. Habitat: Forest, Woodland, bushland, thicket,
and wooded grassland, ca. 60–270 m. Vouchers: Luke Q 1508, Luke WRQ &
Robertson SA 2821, Kokwaro 3960 (EA).
Landolphia watsoniana Romburgh Habit: Liana. Habitat: Moist forest, 0–250 m.
Vouchers: Ngumbau V & Mwadime N V006 & V0226 (EA, HIB), Luke WRQ
& Robertson SA 2820, Luke 3124 (EA).
Marsdenia crinita Oliv. Habit: Liana. Habitat: Forest, ca. 142 m. Vouchers: SAJIT–006007 (EA, HIB), Nyange M 574 (EA).
Marsdenia faulknerae (Bullock) Omlor Habit: Liana. Habitat: Forest margins,
0–400 m. Voucher: Luke & Luke 4749 (EA).
Marsdenia macrantha (Klotzsch) Schltr. Habit: Liana. Habitat: Woodland and forest margins, ca. 60 m. Vouchers: Festo L, Luke Q & P 2791, Robertson & Luke
5536 (EA).
Marsdenia rubicunda N.E.Br. Habit: Liana. Habitat: Bushland and riverine, ca.
20 m. Vouchers: Magogo FC & Glover PE 877, RM Graham 2335, Kimeu JM
698, Robertson SA 3436 (EA).
Marsdenia stelostigma K. Schum. Habit: Liana. Habitat: Bushland, 0–300 m.
Voucher: Mungai et al. 93 (EA).
Marsdenia taylorii Schltr. & Rendle Habit: Liana. Habitat: Dry forest, ca. 200 m.
Vouchers: Luke 4695, Taylor s.n. (EA).
Mascarenhasia arborescens A. DC. Habit: Tree. Habitat: River banks, forest,
0–750 m. Vouchers: Ngumbau V & Mwadime N V0374 (EA, HIB), Magogo FC
& Estes R 1206 (EA).
Mondia ecornuta (N.E.Br.) Bullock Habit: Liana. Habitat: Coastal bushland, woodland, and mangrove forest edge, 0–700 m. Vouchers: Ngumbau V & Mwadime N
V0381 (EA, HIB), Luke WRQ 1637, Hawthorne W 434 (EA).
Mondia whitei (Hook.f.) Skeels Habit: Liana. Habitat: Bushland, 5–250 m. Vouchers: Hawthorne W 529, Robertson SA & Luke WRQ 2352, Robertson SA, Beentje
HJ, Luke WRQ & Khayota B 41 (EA).
Checklist of coastal forests of Kenya
39
Oncinotis tenuiloba Stapf Habit: Liana. Habitat: Forest, riverine, swamp forest, and secondary forest, ca. 20 m. Vouchers: Luke & Robertson 2325, Luke
10720 (EA).
Orbea denboefii (Lavranos) Bruyns Habit: Herb. Habitat: Woodland, ca. 700 m.
Voucher: Heath & Powys 790 (EA).
Orbea distincta (E.A. Bruce) Bruyns Habit: Herb. Habitat: Bushland, ca. 230 m.
Vouchers: Luke & Robertson 2337A, Ritchie s.n. in Bally S52, Robertson SA
6902 (EA).
Oxystelma bornouense R.Br. Habit: Climber. Habitat: Riverine, ca. 60 m. Vouchers:
GRC van Someren 1072, Greenway 9245 (EA).
Pachycarpus bisacculatus (Oliv.) Goyder Habit: Herb. Habitat: Waterlogged grassland or woodland, ca. 100 m. Vouchers: SAJIT–005521 (EA, HIB), Luke WRQ
3162, Drummond & Hemsley 1102 (EA).
Pentatropis nivalis (J.F. Gmel.) D.V. Field & J.R.I. Wood Habit: Climber. Habitat:
Bushland and riverine forests, ca. 20 m. Vouchers: SAJIT–005903, Ngumbau V
& Mwadime N V0454 (EA, HIB), Luke Q 6153 (EA).
Pergularia daemia (Forssk.) Chiov. Habit: Climber. Habitat: Dry bushland, savannah or forest margins, 0–150 m. Vouchers: SAJIT–006210 & 005936 (EA, HIB),
Magogo FC & Glover PE 868 (EA).
Pleiocarpa pycnantha (K. Schum.) Stapf Habit: Tree. Habitat: Evergreen forest, riverine, and swamp forest, ca. 300 m. Vouchers: Luke WRQ 893A & 1405, RB
Faden & AJ Faden 77/751 (EA).
Pleioceras orientale Vollesen Habit: Tree. Habitat: Coastal thicket, 300–750 m.
Voucher: Luke et al. 9445 (EA): Vulnerable.
Raphionacme sp. cf. jurensis N.E. Br. Habit: Climber. Habitat: Woodland. Voucher:
Archer 520 (EA): Endemic.
Raphionacme splendens Schltr. subsp. splendens Habit: Herb. Habitat: Forest,
woodland, and grassland, ca. 80 m. Vouchers: Luke et al. 4494, Luke & Robertson
2331, Adamson 220, Bally 5916, Robertson 6647 (EA).
Rauvolfia mombasiana Stapf Habit: Shrub. Habitat: Riverine forest and coastal forest, 0–550m. Vouchers: SAJIT–005462, Ngumbau V & Mwadime N V116 &
0225 (EA, HIB), Mussa Mandia 64 (EA).
Saba comorensis (Bojer ex A.DC.) Pichon Habit: Liana. Habitat: Closed-forest,
fringing forest, and savannah woodland, ca. 5–80 m. Vouchers: SAJIT–006093
(EA, HIB), Elliot CW 1348, Magogo FC & Glover PE 145 (EA).
Schizostephanus alatus Hochst. ex K. Schum. Habit: Climber. Habitat: Bushland
or degraded forest patches, ca. 10–300 m. Vouchers: Mwadime N 5, Luke 2454,
Robertson 4939, RB Faden, AJ & Faden JB, Gillett & N Gachathi 77/427 (EA).
Schizozygia coffaeoides Baill. Habit: Shrub. Habitat: Moist forest, ca. 204 m. Vouchers: SAJIT–005943 & 005579, Ngumbau V & Mwadime N V063 & 0212 (EA,
HIB), Robertson SA & Luke WRQ 5184 (EA).
Secamone gracilis N.E. Br. Habit: Climber. Habitat: Coastal forest, 0–500 m. Voucher: Hawthorne 191 (EA).
40
Veronicah M. Ngumbau et al. / PhytoKeys 147: 1–191 (2020)
Secamone parvifolia (Oliv.) Bullock Habit: Liana. Habitat: Thickets and coastal forest, ca. 245 m. Vouchers: Ngumbau V & Mwadime N V0269 (EA, HIB), Kokwaro JO 3562, Polhill & Paulo 613 (EA).
Secamone punctulata Decne. Habit: Liana. Habitat: Thickets, riverine, and coastal forest, ca. 5–200 m. Vouchers: SAJIT–006084, Ngumbau V & Mwadime N V0410
& 0513 (EA, HIB), Graham RM FD 2004, Gerhardt K & Steiner M 50 (EA).
Secamone retusa N.E.Br. Habit: Liana. Habitat: Lowland and coastal forest, 0–350 m.
Vouchers: SAJIT–004659, Ngumbau V & Mwadime N V0364, 0458 & 0513
(EA, HIB), Magogo FC & Glover PE 840, Luke 3344, Greenway 1082, Harvey
et al. 53 (EA).
Secamone stuhlmannii K. Schum. Habit: Climber. Habitat: Thickets, ca. 0–800 m.
Voucher: Reitsma J 407 (EA).
Stathmostelma pauciflorum K. Schum Habit: Herb. Habitat: Waterlogged grassland,
ca. 0–400 m. Vouchers: Dale IR K3544, Kassner J 159 (EA).
Strophanthus courmontii Sacleux ex Franch. Habit: Liana. Habitat: Riverine forest
and thickets, ca. 40–280 m. Vouchers: Ngumbau V & Mwadime N V100 & 0386
(EA, HIB), Perdue & Kibuwa 10221, Luke et al. 136, Faden 74/301 (EA).
Strophanthus kombe Oliv. Habit: Liana. Habitat: Evergreen forest and dense bushland,
ca. 30 m. Vouchers: Kimeu JM 664, Luke WRQ et al. 4604G, JB Gillet 21065 (EA).
Strophanthus petersianus Klotzsch Habit: Liana. Habitat: Coastal forest and woodland, 0–650 m. Vouchers: Graham 1712, Robertson 4122 (EA).
Strophanthus zimmermannianus Monach. Habit: Liana. Habitat: Moist forest, 0–800
m. Vouchers: Ngumbau V & Mwadime N V0123 (EA, HIB), Luke PA, WRQ
4528, Luke WRQ 877 & 5276, Magogo & Glover 4911, Luke et al. 4528 (EA).
Tabernaemontana elegans Stapf Habit: Tree. Habitat: Forest, coastal bushland, and
thicket, 0–400 m. Vouchers: SAJIT–005492 (EA, HIB), Kimeu JM 526, Magogo
FC & Glover PE 746 (EA).
Tabernaemontana pachysiphon Stapf Habit: Tree. Habitat: Moist forest and riverine
forest, 0–400 m. Vouchers: SAJIT–005514 (EA, HIB), Magogo FC & Glover PE
201 (EA).
Tacazzea apiculata Oliv. Habit: Liana. Habitat: Swamp and lake bank forests, ca.
30 m. Voucher: Luke et al. 778 (EA).
Vincetoxicum apiculatum (K. Schum.) Meve & Liede Habit: Climber. Habitat:
Coastal, riverine forest, 0–300 m. Vouchers: Robertson SA 6894, Roberson &
Luke 6313, Drummond & Hemsley 3941 (EA).
Vincetoxicum conspicuum (N.E. Br.) Meve & Liede Habit: Climber. Habitat: Moist
forest, ca. 65 m. Vouchers: Luke & Pakia 7458, Luke & Robertson 2309, Nyange
M 519 (EA).
Vincetoxicum cernuum (Decne.) Meve & Liede Habit: Climber. Habitat: Bare
ground, coral thickets, and sand dunes, 0–100 m. Vouchers: SAJIT–005586 (EA,
HIB), Vorontsova MS 110, Napper 1273 (EA).
Vincetoxicum stenolobum (K. Schum.) Meve & Liede. Habit: Climber. Habitat:
Coastal, riverine forest, 0–300 m. Vouchers: Luke et al. TPR 782, Luke & Luke
5975, Kasnner 395 (EA).
Checklist of coastal forests of Kenya
41
Vincetoxicum tenuipedunculatum (K. Schum.) Meve & Liede Habit: Climber.
Habitat: Upland and coastal forest, ca. 80–240 m. Vouchers: SAJIT–005988 (EA,
HIB), Archer PG 511 (EA).
Voacanga africana Stapf Habit: Tree. Habitat: Forest or riverine forest, ca. 60 m.
Vouchers: Luke WRQ & PA 5403, Drummond & Hemsley 3902, Luke 3829,
Nyange M 504 (EA).
Wrightia demartiniana Chiov. Habit: Shrub. Habitat: Dry bushland, ca. 620 m.
Vouchers: Mwadime N 2429, Robertson 1771 (EA).
Xysmalobium stocksii N.E.Br. Habit: Herb. Habitat: Grassland and woodland, ca.
350 m. Vouchers: Luke WRQ 3098B, Magogo & Clover 481 (EA).
F29. Aponogetonaceae
1 Genus, 1 Species
Aponogeton abyssinicus var. albiflorus Lye Habit: Herb. Habitat: Temporary pools,
ca. 35 m. Voucher: Luke Q 1489 (EA).
F30. Araceae
11 Genera, 16 Species
Amorphophallus maximus (Engl.) N.E.Br. subsp. maximus Habit: Herb. Habitat:
Woodland and bushland, ca. 50 m. Voucher: Hildebrandt 2018 (EA).
Anchomanes abbreviatus Engl. Habit: Herb. Habitat: Evergreen forest, 0–800 m.
Vouchers: SAJIT–006158, 005511 & 005979 (EA, HIB), Luke WRQ & Robertson SA 4711, Magogo FC & Glover PE 566 (EA).
Callopsis volkensii Engl. Habit: Herb. Habitat: Forest, 45–800 m. Vouchers: SAJIT–005543, Ngumbau V & Mwadime N V030 (EA, HIB), Magogo FC & Glover PE 1041 (EA): Near Threatened.
Culcasia orientalis Mayo Habit: Climber. Habitat: Riverine and secondary forest,
0–750 m. Vouchers: Ngumbau V & Mwadime N V0372 (EA, HIB), Greenway
PJ, Rawlins SP 9360, Magogo FC & Glover PE 622 (EA).
Gonatopus boivinii (Decne.) Engl. Habit: Herb. Habitat: Forest, woodland, and
wooded grassland, ca. 70 m. Vouchers: Kimeu JM, Meso M Ot 601, Magogo FC
& Glover PE 580, Robertson 5679 (EA).
Gonatopus marattioides (Peter) Bogner Habit: Herb. Habitat: Dry lowland forest
and woodland, 3–350 m. Voucher: Luke WRQ et al. 3331 (EA): Vulnerable.
Gonatopus petiolulatus (Peter) Bogner Habit: Herb. Habitat: Evergreen forest,
0–400 m. Voucher: Luke WRQ & PA 4499 (EA).
Lemna aequinoctialis Welw. Habit: Herb. Habitat: Rock pools, small pools, and
rice fields, ca. 350 m. Vouchers: Drummond & Hemsley 4141, Polhill & Paulo
845(EA).
Pistia stratiotes L. Habit: Herb. Habitat: Open still freshwater, 15–182 m. Vouchers: SAJIT–006224 & 006022 (EA, HIB), Luke PA, WRQ 5623, Magogo FC &
Glover PE 908 (EA).
42
Veronicah M. Ngumbau et al. / PhytoKeys 147: 1–191 (2020)
Spirodela polyrrhiza (L.) Schleid. Habit: Herb. Habitat: Still water, ca. 450 m.
Voucher: Gillett & Kibuwa 19956 (EA).
Stylochaeton bogneri Mayo Habit: Herb. Habitat: Coastal evergreen forest, ca. 207 m.
Vouchers: SAJIT–006022 (EA, HIB), Luke WRQ & PA 4502 (EA): Endangered.
Stylochaeton borumensis N.E. Br. Habit: Herb. Habitat: Bushland, woodland, and
grassland, 0–200 m. Vouchers: SAJIT–005986 & 005987 (EA, HIB).
Stylochaeton salaamicus N.E.Br. Habit: Herb. Habitat: Forest or grassland, 30–
250 m. Vouchers: Rawlins SP 149, Beentje HJ 2364, Luke WRQ & Robertson SA
2694 & 6186 (EA).
Stylochaeton sp. cf. milneanus Mayo Habit: Herb. Habitat: Forest. Voucher: R & L
6104 (EA).
Wolffia arrhiza (L.) Horkel ex Wimm. Habit: Herb. Habitat: Ditches and pools, ca.
0–350 m. Voucher: Drummond & Hemsley 4164 (EA).
Zamioculcas zamiifolia (Lodd.) Engl. Habit: Herb. Habitat: Forest, woodland,
bushland, and grassland, 0–610 m. Vouchers: Luke Q 1547, Magogo FC & Glover PE 1129 (EA).
F31. Araliaceae
1 Genus, 1 Species
Cussonia zimmermannii Harms Habit: Tree. Habitat: Forest and woodland, 20–
400 m. Vouchers: Hawthorne W 553, Magogo FC & Glover PE 680, Drummond
& Hemsley 1198, RM Graham in FD 1806, Robertson SA 3631 (EA).
F32. Araucariaceae
1 Genus, 1 Species
Araucaria cunninghamii Aiton ex D. Don Habit: Tree. Habitat: Dry forest and
thicket, ca. 15 m. Voucher: Perdue RE 10034 (EA): Cultivated.
F33. Arecaceae
7 Genera, 8 Species
Areca catechu L. Habit: Tree. Habitat: Open secondary forest, ca. 0–100 m. Voucher:
DP Kariuki EA 16085 (EA): Cultivated.
Borassus aethiopum Mart. Habit: Tree. Habitat: Riverine flats, coastal plains, and
open secondary forest, ca. 0–200 m. Voucher: Luke WRQ & Robertson SA sr.
Cocos nucifera L. Habit: Tree. Habitat: Tropical seashores, ca. 0–100 m. Voucher:
Verdcourt B 2120 (EA): Cultivated.
Elaeis guineensis Jacq. Habit: Tree. Habitat: Open forest, ca. 0–200 m. Vouchers:
Luke PA, WRQ 4517, Shimba Hills Survey Unit 56 (EA).
Hyphaene compressa H. Wendl. Habit: Tree. Habitat: Open grassland, 150 m.
Voucher: Luke WRQ & Robertson SA sr.
Checklist of coastal forests of Kenya
43
Hyphaene coriacea Gaertn. Habit: Shrub. Habitat: Savannah and grassland, 0–300m.
Vouchers: Kirika P 576, Shimba Hills Survey Unit 42 (EA).
Phoenix reclinata Jacq. Habit: Tree. Habitat: Open forests, savannah woodland, or
low scrub thickets, ca. 30 m. Voucher: Moomaw JC 1289 (EA).
Raphia farinifera (Gaertn.) Hyl. Habit: Tree. Habitat: Moist swampy ground, ca.
0–250 m. Voucher: Magogo FC & Glover PE sr.
F34. Aristolochiaceae
2 Genera, 3 Species
Aristolochia albida Duch. Habit: Climber. Habitat: Forest grassland, clearings, and
edges, ca. 260 m. Vouchers: SAJIT–005595, Ngumbau V & Mwadime N V004
(EA, HIB), Festo L, Luke Q & P 2673 (EA).
Aristolochia bracteolata Lam. Habit: Climber. Habitat: Riverine forest, 30–100 m.
Vouchers: AB Katende 1796, Q Luke et al. TPR 479 (EA).
Hydnora sinandevu Beentje & Q. Luke Habit: Herb. Habitat: Scattered tree grassland, ca. 2 m. Vouchers: Luke 3033, Bally 2051 & 10434 (EA).
F35. Asparagaceae
7 Genera, 40 Species
Albuca abyssinica Jacq. Habit: Herb. Habitat: In grassland, ca. 382 m. Vouchers:
Ngumbau V & Mwadime N V0357 (EA, HIB), Luke WRQ 3153 (EA).
Asparagus africanus Lam. Habit: Climber. Habitat: Forest edges and bushy wooded
areas, ca. 2 m. Vouchers: SAJIT–006236 (EA, HIB), Rawlins SP 84 (EA).
Asparagus buchananii Baker Habit: Climber. Habitat: Grassland and wooded grassland, 0–150 m. Vouchers: Williams 299, Graham RM 1931 (EA).
Asparagus falcatus L. Habit: Climber. Habitat: Bushland and dry forest, 0–850 m.
Vouchers: SAJIT–006428 (EA, HIB), Greenway 9623, Luke WRQ et al. 6193,
Bally 5837, Greenway 9623 (EA).
Asparagus faulknerae Sebsebe Habit: Climber. Habitat: Bushland, dry forest,
and woodland, 0–250 (–450) m. Vouchers: Bally & Smith B14364, Bjornstad
211 (EA).
Asparagus flagellaris (Kunth) Baker Habit: Shrub. Habitat: Grassland and wooded
bushland, 0–100 m. Vouchers: Mwadime N 612, Waterman 1093 (EA).
Asparagus humilis Engl. Habit: Herb. Habitat: Coral outcrops, salt flats near mangrove, 0–15 m. Vouchers: Luke WRQ 3029, De Meester 19/79 (EA).
Asparagus leptocladodius Chiov. Habit: Shrub. Habitat: Grassland or bushed grassland, 0–750 m. Voucher: Ngweno 7C (EA).
Asparagus scaberulus A. Rich. Habit: Climber. Habitat: Woodland, grassland, and
bushed grassland, (60)–350 m. Voucher: Makin in EA 13055 (EA).
Asparagus setaceus (Kunth) Jessop Habit: Climber. Habitat: Woodland and grassland, ca. 50 m. Voucher: Luke WRQ & Robertson SA 989 (EA).
44
Veronicah M. Ngumbau et al. / PhytoKeys 147: 1–191 (2020)
Chlorophytum angustissimum (Poelln.) Nordal Habit: Herb. Habitat: Grassland, ca.
50 m. Voucher: Luke 3025 (EA).
Chlorophytum cameronii var. pterocaulon (Welw. ex Baker) Nordal Habit: Herb.
Habitat: Woodland and grassland, ca. 350 m. Vouchers: Drummond & Hemsley
4156, Polhill & Paulo 834 (EA).
Chlorophytum filipendulum subsp. amaniense (Engl.) Nordal & A.D. Poulsen
Habit: Herb. Habitat: Forest, 30–650 m. Vouchers: Ngumbau V & Mwadime N
V021 & V0407 (EA, HIB), Magogo & Glover 784, Polhill & 795 (EA).
Chlorophytum holstii Engl. Habit: Herb. Habitat: Forest or woodland, ca. 80 m.
Vouchers: Robertson & Luke 6240, Gray M 435 (EA).
Chlorophytum suffruticosum Baker Habit: Herb. Habitat: Rocks and termite
mounds, ca. 10 m Vouchers: Luke Q 1553, Drummond & Hemsley 1012 (EA).
Chlorophytum tenerrimum Peter ex Poelln. Habit: Herb. Habitat: Lowland forest
and thickets, 70–150 m. Vouchers: SAJIT–006130 (EA, HIB), Drummond &
Hemsley 3794, Polhill & Paulo 851, Musyoki & Hansen 994 (EA).
Dipcadi longifolium (Lindl.) Baker Habit: Herb. Habitat: On sand along the coast,
0–30 m. Vouchers: SAJIT–006242 & 004675 (EA, HIB), Luke WRQ 3118, Bally
8877, Gillett 20836, Greenway 10458 (EA).
Dipcadi viride (L.) Moench Habit: Herb. Habitat: Grassland, bushland, and woodland, ca. 500 m. Vouchers: Magogo FC & Glover PE 600, Meester-Manger Cats
V de 42 (EA).
Dracaena aletriformis (Haw.) Bos Habit: Shrub. Habitat: Evergreen or semi-deciduous forest, 50–300 m. Vouchers: Verdcourt 3606, Robertson & Luke 4781,
Schmidt 1712 (EA).
Dracaena fragrans (L.) Ker Gawl. Habit: Shrub. Habitat: Moist forest, ca. 210 m.
Vouchers: Ngumbau V & Mwadime N V0391 (EA, HIB), Robertson SA
4207 (EA).
Dracaena laxissima Engl. Habit: Climber. Habitat: Moist forest or riverine forest, ca.
1–400 m. Voucher: Luke WRQ & Robertson SA 242A (EA).
Dracaena mannii Baker Habit: Shrub. Habitat: Moist forest, riverine forest, forest margins, and secondary bushland, ca. 300 m. Vouchers: SAJIT–005505 (EA,
HIB), Robertson SA & Luke WRQ 5185 (EA).
Drimia altissima (L.f.) Ker Gawl. Habit: Herb. Habitat: Grassland, bushed grassland,
ca. 30 m. Vouchers: Magogo FC & Glover PE 213, Irwin 453, Jeffrey 293 (EA).
Drimia macrocarpa Stedje Habit: Herb. Habitat: Grassland and bushland, ca. 35 m.
Vouchers: Magogo FC & Glover PE 30, Archer 531, Drummond & Hemsley
1030 (EA).
Ledebouria kirkii (Baker) Stedje & Thulin Habit: Herb. Habitat: Grassland or
bushland, ca. 0–300 m. Voucher: Gilbert 6048 (EA).
Sansevieria itumea (Mbugua) Jankalski Habit: Herb. Habitat: Thickets lowlands of
the coastal region. Voucher: Bally & Smith B14404A (EA).
Sansevieria arborescens Cornu ex Gérôme & Labroy Habit: Herb. Habitat: Evergreen coastal forest or evergreen bushland, 0–600 m. Vouchers: Reitsma J 543,
Luke WRQ et al. 5703 (EA).
Checklist of coastal forests of Kenya
45
Sansevieria bagamoyensis N.E.Br. Habit: Herb. Habitat: Open forest, coastal bushland, and thicket, 0–400 m. Vouchers: Graham 1808, Faden et al. 77/460 (EA).
Sansevieria ballyi L.E. Newton Habit: Herb. Habitat: Rock faces, ca. 180 m. Vouchers: Mrs Robertson SA 7602, Newton 5856, Newton 5594 (EA): Endemic.
Sansevieria conspicua N.E.Br. Habit: Herb. Habitat: Littoral forest and evergreen
bushland, 2–150 m. Vouchers: Luke 3092, Drummond & Hemsley 3845, Rawlins SP 382, Robertson & Luke 2284 (EA).
Sansevieria ehrenbergii Schweinf. ex Baker Habit: Herb. Habitat: Dry wooded grassland, dry bushland, and thickets, ca. 20 m. Voucher: Mrs Robertson SA 7723 (EA).
Sansevieria fischeri (Baker) Marais Habit: Herb. Habitat: Dry bushland, 250–
550 m. Voucher: Bally PRO 16893 (EA).
Sansevieria forskaoliana (Schult. f.) Hepper & J.R.I. Wood Habit: Herb. Habitat:
Dry or evergreen bushland and grassland, ca. 100 m. Voucher: Robertson 1778 (EA).
Sansevieria francisii Chahinian Habit: Herb. Habitat: Coastal forest, low altitude.
Voucher: Horwood 432 (EA): Endemic.
Sansevieria gracilis N.E.Br. Habit: Herb. Habitat: Dry bushland and thicket,
0–100 m. Voucher: Luke WRQ & Robertson SA 1478 (EA).
Sansevieria hyacinthoides (L.) Druce Habit: Herb. Habitat: Dry bushland or riverine forest, 50–600 m. Voucher: Verdcourt 3220 (EA).
Sansevieria kirkii Baker Habit: Herb. Habitat: Forest, thicket, and riverine fringe,
5–180 m. Vouchers: Luke 4012 & 4013 (EA).
Sansevieria nitida Chahinian Habit: Herb. Habitat: No data. Voucher: Chahinian
301 (EA): Endemic.
Sansevieria perrotii Warb. Habit: Herb. Habitat: Bushland and bushed grassland,
550 m. Voucher: Sangai GW 15602 (EA).
Sansevieria powelii N.E.Br. Habit: Herb. Habitat: Dry bushland and scattered tree
grassland, ca. 650 m. Vouchers: Faden RB & AJ 71/810, Perdue RE & Kibuwa SP
10126, Gillett JB 19970 (EA).
Sansevieria raffillii N.E.Br. Habit: Herb. Habitat: Dry bushland, woodland or dry
forest, 0–500 m. Voucher: Polhill R & Paulo S 682 (EA).
F36. Asphodelaceae
1 Genus, 12 Species
Aloe deserti A. Berger Habit: Shrub. Habitat: Deciduous thickets, ca. 259 m. Voucher: Bally PRO 8866 (EA): Near Threatened.
Aloe kilifiensis Christian Habit: Herb. Habitat: Bushland, 3–380 m. Vouchers: Magogo FC & Glover PE 430, Jeffery GW 710, Frazier 881 (EA): Endangered.
Aloe massawana Reynolds Habit: Herb. Habitat: Open thickets, 0–20 m. Voucher:
SA Robertson 6365 (EA): Vulnerable.
Aloe microdonta Chiov. Habit: Herb. Habitat: Bushland, ca. 60 m. Voucher: Gillett
16544 (EA).
Aloe parvidens M.G. Gilbert & Sebsebe Habit: Herb. Habitat: Acacia woodland, ca.
560 m. Voucher: Tweedie 1199 (EA).
46
Veronicah M. Ngumbau et al. / PhytoKeys 147: 1–191 (2020)
Aloe penduliflora Baker Habit: Shrub. Habitat: Bushland, ca. 645 m. Voucher: Robertson & Luke 6159 (EA).
Aloe rabaiensis Rendle Habit: Shrub. Habitat: Bushland, 18–500 m. Vouchers: Robertson SA & Luke WRQ 2211, Verdcourt 3193, Bally 8537, Hooper & Transcend
1156 (EA).
Aloe ruspoliana Baker Habit: Herb. Habitat: Bushland, open rocky hillsides, ca.
400 m. Voucher: Bally & Smith 14990 (EA).
Aloe secundiflora Engl. var. secundiflora Habit: Herb. Habitat: Grassland, woodland, ca. 200 m. Voucher: Mwendia CW 16247 (EA).
Aloe ukambensis Reynolds Habit: Herb. Habitat: Woodland, ca. 600 m. Voucher:
Gilbert MG 1739 (EA): Vulnerable.
Aloe vituensis Baker Habit: Shrub. Habitat: Bushland, ca. 290 m. Voucher: Newton
& Powys 3569 (EA).
Aloe volkensii Engl. subsp. volkensii Habit: Shrub. Habitat: Riverine woodland,
ca. 10–410 m. Vouchers: Luke WRQ & Robertson SA 1026, Robertson SA
6379 (EA).
F37. Asteraceae
49 Genera, 82 Species
Acanthospermum hispidum DC. Habit: Herb. Habitat: Disturbed places, ca. 260 m.
Voucher: Waaijenberg H 8 (EA): Naturalized.
Acmella uliginosa (Sw.) Cass. Habit: Herb. Habitat: Forest, swampy area, and grassland, ca. 40 m. Vouchers: Luke WRQ & PA 6032, Nyange M 457 (EA).
Adenostemma viscosum J.R. Forst. & G. Forst. Habit: Herb. Habitat: Wet ground
and riverine, ca. 300 m. Vouchers: Luke WRQ & Robertson SA 2755, Magogo
FC & Glover PE 102 & 373 (EA).
Ageratum conyzoides L. Habit: Herb. Habitat: Woodland, grassland, and cultivated
land, ca. 300 m. Voucher: Magogo FC & Glover PE 371 (EA): Naturalized.
Aspilia kotschyi (Sch. Bip. ex Hochst.) Oliv. Habit: Herb. Habitat: Grassland, waste
land, bushland, and woodland, ca. 350 m. Vouchers: Ngumbau V & Mwadime N
V0241 (EA, HIB), Rawlins SP 817 (EA).
Aspilia macrorrhiza Chiov. Habit: Herb. Habitat: Sand dunes or coral, 0–10 m.
Voucher: Luke, PA & WRQ 6148 (EA): Endangered.
Aspilia mossambicensis (Oliv.) Wild Habit: Shrub. Habitat: Swamps, forest, woodland, bushland, and grassland, ca. 45–200 m. Vouchers: SAJIT–006147, Ngumbau V & Mwadime N V0196 (EA, HIB), Festo L, Luke WRQ & PA 2802, Magogo FC & Glover PE 325 (EA).
Athroisma pusillum T. Erikss. Habit: Herb. Habitat: Woodland, 350–500 m. Vouchers: Drummond & Hemsley 4073, Faden & Faden 72/120 (EA): Endemic.
Bidens holstii (O. Hoffm.) Sherff Habit: Herb. Habitat: Forest, bushland, and grassland, ca. 700 m. Voucher: Luke & Robertson 2050 (EA).
Bidens pilosa L. Habit: Herb. Habitat: Neglected places, ca. 90 m. Vouchers: Magogo
FC & Glover PE 825, Craig M 9179 (EA): Naturalized.
Checklist of coastal forests of Kenya
47
Bidens schimperi Sch.Bip. ex Walp. Habit: Herb. Habitat: Disturbed areas, woodland, bushland, and grassland, 250–372 m. Vouchers: SAJIT–006146, Ngumbau
V & Mwadime N V0149 (EA, HIB), Gillett 18677, Someren, HD van 43 (EA).
Bidens taylorii (S. Moore) Sherff Habit: Herb. Habitat: Grassland, bushland, and forest,
ca. 60 m. Vouchers: Robertson SA & Luke WRQ 6051, Magogo & Glover 651 (EA).
Blepharispermum ellenbeckii Cufod. Habit: Shrub. Habitat: Bushland, ca. 150 m.
Voucher: Mungai & Rucina 404/84 (EA).
Blepharispermum minus S. Moore Habit: Shrub. Habitat: Bushland and grassland,
30–450 m. Vouchers: Luke et al. TPR 650, Bally 16886 (EA).
Blepharispermum zanguebaricum Oliv. & Hiern Habit: Shrub. Habitat: Dry forest
and bushland, ca. 150 m. Vouchers: Robertson SA & Beentje HJ 4061, Robertson
SA, Beentje HJ, Luke WRQ & Khayota B 152 (EA).
Blumea axillaris (Lam.) DC. Habit: Herb. Habitat: Swampy, riverine, and grassland,
ca. 1–300 m. Voucher: Magogo FC & Glover PE 526 (EA).
Bothriocline longipes (Oliv. & Hiern) N.E.Br. Habit: Herb. Habitat: Wooded grassland or riverine vegetation, ca. 200 m. Voucher: Gilbert VC 3505 (EA).
Brachylaena huillensis O. Hoffm. Habit: Tree. Habitat: Forest and bushland, ca.
500 m. Vouchers: Katz SS 75/33, Luke WRQ 915 (EA): Near Threatened.
Crassocephalum crepidioides (Benth.) S. Moore Habit: Herb. Habitat: Riverine, forest, and cultivation area, ca. 45 m. Vouchers: Luke WRQ et al. 4728, Makokha D,
Malombe I & Saidi C 1567 (EA).
Cyanthillium cinereum (L.) H. Rob. Habit: Herb. Habitat: Cultivated area and grassland, ca. 243 m. Vouchers: Ngumbau V & Mwadime N V024 (EA, HIB), Luke
WRQ & PA 5691, Kirika P & Muthoka P 738, Rawlins 430, Perdue & Kibuwa
10037, Robertson 3307 (EA).
Dicoma tomentosa Cass. Habit: Herb. Habitat: Grassland, bushland, woodland, and
rocky area, ca. 400 m. Voucher: van Someren 946 (EA).
Eclipta prostrata (L.) L. Habit: Herb. Habitat: Swampy and riverine area, ca.
23 m. Vouchers: Kirika P, Muthoka P & Mbale M 755, Bally PRO & Smith AR
14373 (EA).
Emilia bellioides (Chiov.) C.Jeffrey Habit: Herb. Habitat: Sand dunes and coral,
0–10 m. Vouchers: Luke WRQ 5464, Rawlins 105, Greenway & Rawlins 9421
(EA): Vulnerable.
Emilia coccinea (Sims) G. Don Habit: Herb. Habitat: Grassland and bushland, ca.
186 m. Vouchers: Ngumbau V & Mwadime N V0348 (EA, HIB), Fukuoka N
284, Cunningham-van Someren GR Sh41 (EA).
Emilia sonchifolia (L.) DC. ex Wight Habit: Herb. Habitat: Swampy, grassland, and
cultivation area, 1–600 m. Vouchers: Jeffery 780, Greenway & Rawlins 9375 (EA).
Emilia tricholepis C. Jeffrey Habit: Herb. Habitat: Woodland, ca. 5–200 m. Voucher: Brathany C 58 (EA).
Erigeron aegyptiacus L. Habit: Herb. Habitat: Grassland, cultivation, riverbeds,
floodplains, ca. 100 m. Voucher: Njoroge Thairu 38 (EA).
Erigeron bonariensis L. Habit: Herb. Habitat: Grassland, 0–500 m. Voucher: Magogo FC & Glover PE 344 (EA).
48
Veronicah M. Ngumbau et al. / PhytoKeys 147: 1–191 (2020)
Erythrocephalum marginatum (O. Hoffm.) S. Ortiz & A.P. Cout. Habit: Herb.
Habitat: Forest, ca. 53 m. Vouchers: SAJIT–004651, Ngumbau V & Mwadime N
V0130 (EA, HIB), Magogo FC & Glover PE 937, Simpson BL 388 (EA).
Erythrocephalum minus Oliv. Habit: Herb. Habitat: Grassland, forest, woodland,
and roadsides, ca. 45 m. Vouchers: Graham RM 1928, Drummond & Hemsley
4146, Luke & Luke 3980 (EA).
Ethulia angustifolia Bojer ex DC. Habit: Herb. Habitat: Cultivation, riverine, roadside, and grassland, ca. 210 m. Vouchers: Ngumbau V & Mwadime N V0376
(EA, HIB), Festo L, Luke Q & P 2803 (EA).
Ethulia faulknerae C. Jeffrey Habit: Herb. Habitat: Bushland, coral, 0–15 m. Vouchers: Luke WRQ 15341, Gregory, Festo 2803 (EA).
Ethulia gracilis Delile Habit: Herb. Habitat: A common weed, ca. 256 m. Vouchers:
Fukuoka N 517, Kirika P 934, Kuchar P 7267 (EA).
Flaveria trinervia (Spreng.) C. Mohr Habit: Herb. Habitat: Waste ground, swampy,
and cultivation. Voucher: Drummond & Hemsley 1047 (EA): Naturalized.
Galinsoga parviflora Cav. Habit: Herb. Habitat: Disturbed areas and roadside, ca.
0–100 m. Voucher: Jeffery GW 776 (EA): Naturalized.
Geigeria acaulis Benth. & Hook.f. ex Oliv. & Hiern Habit: Herb. Habitat: Grassland and bushland, ca. 250 m. Voucher: Gillett & Gachathi 20527 (EA).
Grangea maderaspatana (L.) Poir. Habit: Herb. Habitat: Riverine, ca. 5–100 m.
Voucher: Luke et al. TPR 144 (EA).
Grauanthus linearifolius (O. Hoffm.) Fayed Habit: Herb. Habitat: Grassland and forest,
1–150 m. Vouchers: Greenway & Rawlins 9455, Hooper & Townsend 1181 (EA).
Gutenbergia cordifolia Benth. ex Oliv. var. cordifolia Habit: Herb. Habitat: Grassland,
riverine, forest, roadside, and cultivated area, ca. 300 m. Voucher: Boyle B 132 (EA).
Gutenbergia pembensis S. Moore Habit: Herb. Habitat: Woodland, grassland, and
forest, 1–400 m. Vouchers: Ngumbau V & Mwadime N V0461 (EA, HIB), Simpson, BL 76, Magogo FC & Glover PE 1084, Boyle B 132, Luke & Robertson
1780, Kassner 366 (EA).
Gymnanthemum coloratum (Willd.) H. Rob. & B. Kahn Habit: Shrub. Habitat:
Woodland or riverine, 160–250 m. Vouchers: Magogo FC & Glover PE 1142,
Faden 77/705 (EA).
Gynura colorata Peter ex F. G. Davies Habit: Herb. Habitat: Forest and rocky area,
0–850 m. Voucher: Musyoki BM & Hansen OJ 974 (EA).
Gynura pseudochina (L.) DC. Habit: Herb. Habitat: Grassland and woodland, ca.
0–100m. Vouchers: Festo L & Luke WRQ 2691, Kassner 475 (EA).
Gynura scandens O. Hoffm. Habit: Climber. Habitat: Forest, riverine and woodland, ca.
1–200 m. Vouchers: Magogo FC & Glover PE 975, Luke & Robertson 1789 (EA).
Helichrysum glumaceum DC. Habit: Herb. Habitat: Grassland or dry bushland, ca.
350 m. Vouchers: Luke WRQ & Robertson SA 2541, Drummond RB & Hemsley
JH 4047 (EA).
Jeffreycia hildebrandtii (Vatke) H. Rob., S. C. Keeley & Skvarla Habit: Shrub. Habitat: Forest margins, swampy area, riverine, bushland, and roadsides, 1–700 m. Vouchers: Kirika P, Muthoka P & Mbale M 749, Magogo FC & Glover PE 171 (EA).
Checklist of coastal forests of Kenya
49
Jeffreycia zanzibarensis (Less.) H. Rob., S. C. Keeley & Skvarla Habit: Herb. Habitat: Forest margins, bushland, and roadsides, ca. 96 m. Vouchers: SAJIT–006126
(EA, HIB), Adamson 11, Drummond & Hemsley 1155 (EA).
Kleinia abyssinica (A. Rich.) A. Berger var. abyssinica Habit: Herb. Habitat: Bushland, ca. 60 m. Vouchers: SAJIT–004666 (EA, HIB), Simpson BL 390 (EA).
Kleinia abyssinica var. hildebrandtii (Vatke) C. Jeffrey Habit: Herb. Habitat: Rocky,
grassland, bushland, woodland, and forest, 20–250 m. Vouchers: Luke WRQ &
Robertson SA 964, Musyoki & Hansen 971 (EA).
Kleinia implexa (P.R.O. Bally) C. Jeffrey Habit: Herb. Habitat: Bushland and rocky
area, ca. 340 m. Vouchers: Luke WRQ & Robertson SA 2566, Drummond &
Hemsley 4092 (EA).
Kleinia odora (Forssk.) DC. Habit: Shrub. Habitat: Bushland, ca. 75 m. Voucher:
Bally PRO 16891 (EA).
Kleinia schweinfurthii (Oliv. & Hiern) A. Berger Habit: Herb. Habitat: Bushland,
ca. 200 m. Voucher: Luke & Luke 4427 (EA).
Laggera brevipes Oliv. & Hiern Habit: Herb. Habitat: Grassland, woodland, and
waste ground, 1–200 m. Voucher: Magogo FC & Glover PE 1137 (EA).
Launaea cornuta (Hochst. ex Oliv. & Hiern) C. Jeffrey Habit: Herb. Habitat:
Grassland and cultivated area, ca. 227 m. Vouchers: Ngumbau V & Mwadime N
V0275 (EA, HIB), Magogo FC & Glover PE 813 (EA).
Launaea intybacea (Jacq.) Beauverd Habit: Herb. Habitat: Riverbed, coral, and
cultivated area, ca. 6 m. Vouchers: Greenway 9267, Mwadime N & Chesire C
195 (EA).
Launaea nana (Baker) Chiov. Habit: Herb. Habitat: Grassland, ca. 450 m. Voucher:
Luke WRQ et al. 4604E (EA).
Launaea sarmentosa (Willd.) Sch.Bip. ex Kuntze Habit: Herb. Habitat: High water
mark, beach, and sandy shores, 0–15 m. Vouchers: SAJIT–006260 (EA, HIB),
Rawlins 184, Robertson 6136, Drummond & Hemsley 4001 (EA).
Microglossa hildebrandtii O. Hoffm. Habit: Liana. Habitat: Forest and bushland,
1–600 m. Vouchers: Robertson SA & Luke WRQ 5591, Magogo FC & Glover
PE 737, Luke WRQ 1811 (EA).
Mikania chenopodiifolia Willd. Habit: Climber. Habitat: Forest, swampy, and riverine, 0–300 m. Voucher: Magogo FC & Glover PE 1126 (EA).
Nicolasia nitens (O. Hoffm.) Eyles Habit: Herb. Habitat: Bushland, riverine, and
temporary pools, 300–600 m. Voucher: Mungai et al. 284/83 (EA).
Orbivestus homilanthus (S. Moore) H. Rob. Habit: Shrub. Habitat: Forest margin,
bushland, and beach, 0–15 m. Voucher: Gillespie JB 204 (EA).
Pluchea dioscoridis (L.) DC. Habit: Shrub. Habitat: Riverine, woodland, and
swampy, ca. 61 m. Vouchers: Sangai GW EA 15795, Kibuwa 2466, Robertson &
Luke 5816, Polhill & Paulo 830 (EA).
Pluchea sordida (Vatke) Oliv. & Hiern Habit: Shrub. Habitat: Riverine, ca. 256 m.
Vouchers: Luke Q 5651, Kuchar P 11974, Kirika P 932 (EA).
Porphyrostemma monocephala (E.A. Bruce) Leins Habit: Herb. Habitat: Grassland,
ca. 45 m. Voucher: Makin 422 (EA).
50
Veronicah M. Ngumbau et al. / PhytoKeys 147: 1–191 (2020)
Pseudoconyza viscosa (Mill.) D’Arcy Habit: Herb. Habitat: Riverine forest, ca.
1–300 m. Vouchers: Luke 3354, Jeffery GW 703, Magogo FC & Glover PE
388 (EA).
Sigesbeckia orientalis L. Habit: Herb. Habitat: Cultivated and ruderal area, ca.
240 m. Vouchers: Greenway PJ 9178, Pierce T, David & James (EA).
Solanecio angulatus (Vahl) C. Jeffrey Habit: Climber. Habitat: In rivers and streams
or woodland, 250–600 m. Voucher: Luke WRQ & PA sr.
Solanecio cydoniifolius (O. Hoffm.) C.Jeffrey Habit: Climber. Habitat: Forest margin, cultivated area, and coral, ca. 3 m. Vouchers: van Someren VGL Sh 3, Wakefield s.n., Mwadime & Chesire 278 (EA).
Sphaeranthus africanus L. Habit: Herb. Habitat: Riverine, 1–450 m. Vouchers: Luke
WRQ 2938, Nyange M 489 (EA).
Sphaeranthus bullatus Mattf. Habit: Herb. Habitat: Riverine and rocky area, ca.
60 m. Voucher: Luke & Luke 3827a (EA).
Sphaeranthus kirkii Oliv. & Hiern var. kirkii Habit: Herb. Habitat: Riverine, ca.
137 m. Vouchers: Fukuoka N 270, Magogo & Glover 760 (EA).
Sphaeranthus kirkii var. cyathuloides (O. Hoffm.) Beentje Habit: Herb. Habitat:
Bushland, 1–1750 m. Voucher: Thomas 125 (EA).
Sphaeranthus spathulatus A. Peter Habit: Herb. Habitat: Ponds and streams,
1–250 m. Voucher: Festo L & Luke Q 2577 (EA).
Sphaeranthus ukambensis Vatke & O. Hoffm. Habit: Herb. Habitat: Riverine, bushland, grassland, and woodland, 1–450 m. Voucher: Tana R. Primate Res. Expedition 298 (EA).
Sphaeranthus zavattarii Cufod. Habit: Herb. Habitat: Grassland and swampy area,
100–450 m. Voucher: Makin EA 14692 (EA).
Synedrella nodiflora (L.) Gaertn. Habit: Herb. Habitat: Waste land or cultivated
land, ca. 213 m. Vouchers: SAJIT–006071, Ngumbau V & Mwadime N V0335
(EA, HIB), Magogo FC & Glover PE 722 (EA): Naturalized.
Tridax procumbens L. Habit: Herb. Habitat: Weed of cultivated areas, 213–
381 m. Vouchers: Waaijenberg H 10, Magogo FC & Glover PE 723 & 351
(EA): Naturalized.
Vernonia colorata (Willd.) Drake subsp. colorata Habit: Tree. Habitat: Woodland
or riverine, ca. 60 m. Vouchers: Faden & Faden 77/705, Nyange M 506 (EA).
Vernonia popeana C. Jeffrey Habit: Herb. Habitat: Bushland, wooded grassland, and
woodland, 0–150 m. Voucher: Luke & Robertson 2514 (EA).
Vernonia wakefieldii Oliv. Habit: Shrub. Habitat: Bushland, ca. 350 m. Vouchers:
SAJIT–005429 (EA, HIB), Robertson SA & Beentje HJ 4062, Drummond &
Hemsley 4058 (EA).
Vernoniastrum aemulans (Vatke) H. Rob. Habit: Herb. Habitat: Roadside, cultivations, grassland, woodland, and bushland, ca. 80–375 m. Vouchers: Ngumbau V
& Mwadime N V0147 (EA, HIB), Waaijenbrg H 3, Magogo FC & Glover PE
716, 1087, 398 & 500 (EA).
Wollastonia biflora (L.) DC. Habit: Shrub. Habitat: Bushland, coral, and sandy
shores, 0–15 m. Voucher: Festo L & Luke WRQ 2454 (EA).
Checklist of coastal forests of Kenya
51
F38. Balsaminaceae
1 Genus, 1 Species
Impatiens walleriana Hook. f. Habit: Herb. Habitat: Upland and coastal rainforest,
ca. 200 m. Vouchers: SAJIT–005953, Ngumbau V & Mwadime N V0397 (EA,
HIB), Magogo FC & Glover PE 129, Luke 1336 (EA).
F39. Basellaceae
1 Genus, 2 Species
Basella alba L. Habit: Climber. Habitat: Moist places, ca. 136 m. Voucher: SAJIT–005968 (EA, HIB).
Basella paniculata Volkens Habit: Climber. Habitat: Bushland and forest, ca. 20 m.
Vouchers: Robertson SA 5720, Adams BR 59 (EA).
F40. Begoniaceae
1 Genus, 1 Species
Begonia wakefieldii Gilg ex Engl. Habit: Herb. Habitat: Forest and bushland, ca.
136–280 m. Vouchers: SAJIT–005952 (EA, HIB), Luke WRQ & Robertson SA
478, Luke WRQ 1634 (EA).
F41. Bignoniaceae
5 Genera, 5 Species
Fernandoa magnifica Seem. Habit: Tree. Habitat: Coastal thicket and woodland,
5–450 m. Vouchers: SAJIT–006157, Ngumbau V & Mwadime N V0385 (EA,
HIB), Magogo FC & Glover PE 168 (EA).
Kigelia africana (Lam.) Benth. Habit: Tree. Habitat: Grassland and woodland, ca.
300 m. Voucher: Jeffery 111 (EA).
Markhamia zanzibarica (Bojer ex DC.) K. Schum. Habit: Shrub. Habitat: Coastal
thicket and woodland, ca. 230 m. Vouchers: Ngumbau V & Mwadime N V0466
(EA, HIB), Shimba Hills Survey Unit 90, Kimeu JM 536 (EA).
Spathodea campanulata P. Beauv. subsp. nilotica (Seem.) Bidgood Habit: Shrub.
Habitat: Scrubland and secondary forests, ca. 100 m. Voucher: Luke WRQ
3089 (EA).
Stereospermum kunthianum Cham. Habit: Tree. Habitat: Dry areas of deciduous
forest and woodland, ca. 20 m. Voucher: Magogo FC & Glover PE 642 (EA).
F42. Bixaceae
2 Genera, 2 Species
Bixa orellana L. Habit: Shrub. Habitat: Cultivated, naturalized, ca. 30–150 m.
Vouchers: SAJIT–005493 (EA, HIB), Spjut RW 2637 (EA): Naturalized.
52
Veronicah M. Ngumbau et al. / PhytoKeys 147: 1–191 (2020)
Cochlospermum religiosum (L.) Alston Habit: Tree. Habitat: Dry forests. Voucher:
Graham RM 2334 (EA).
F43. Boraginaceae
7 Genera, 28 Species
Argusia argentea (L.f.) Heine Habit: Shrub. Habitat: Sea shore, ± sea level. Voucher:
Greenway 10442 (EA).
Cordia crenata subsp. meridionalis Warfa Habit: Shrub. Habitat: Bushland, ca.
60 m. Voucher: Thairu 110 (EA).
Cordia faulknerae Verdc. Habit: Liana. Habitat: Riverine bushland, 0–250 m.
Vouchers: SA Robertson & WRQ Luke 5732, Luke 1573 (EA).
Cordia goetzei Gürke Habit: Tree. Habitat: Coastal forest, rainforest, riverine forest,
and ground water forest, 5–60 m. Vouchers: Mwadime N & Ndungu K 27, RB
Faden & AJ Faden 77/657 (EA).
Cordia guineensis Thonn. subsp. mutica Verdc. Habit: Shrub. Habitat: Limestone
outcrops, 0–10 (–50) m. Vouchers: SAJIT–006185 (EA, HIB), Mwadime N 46,
Luke Q 1413 (EA).
Cordia monoica Roxb. Habit: Tree. Habitat: Dry woodland, ca. 15 m. Voucher: Mariette Steiner 290 (EA).
Cordia sinensis Lam. Habit: Tree. Habitat: Grassland and bushland, ca. 60 m. Voucher: Drummond & Hemsley 4185 (EA).
Cordia somaliensis Baker Habit: Shrub. Habitat: Coastal bushland, high-tidemark,
and coral rag, 0–15 (–150) m. Vouchers: Kirika P, Muthoka P & Mbale M 758,
Greenway & Rawlins 8904 (EA).
Cordia sp. B of FTEA Habit: Shrub. Habitat: Acacia-Combretum woodland. Vouchers: Muchiri 567, Kuchar 13691 (EA).
Cordia subcordata Lam. Habit: Tree. Habitat: Shore, high-tidemark, and mangrove
swamps, 0–15 (–75) m. Vouchers: Kimeu JM 667, Gillett 20837, Verdcourt
1070 (EA).
Cordia torrei E.S. Martins Habit: Tree. Habitat: Forest and rock outcrop, 200–420 m.
Vouchers: Verdcourt 2414, Luke & Robertson 1876 (EA).
Ehretia amoena Klotzsch Habit: Shrub. Habitat: Woodland, 0–600 m. Vouchers:
Ngumbau V & Mwadime N V0507 (EA, HIB), Luke WRQ et al. 6192 (EA).
Ehretia bakeri Baker Habit: Shrub. Habitat: Forest, bushland, 199–700 m. Vouchers: SAJIT–006456, Ngumbau V & Mwadime N V0424 (EA, HIB), Shimba Hills
Survey Unit 84, R & L 2047, Drummond & Hemsley 4201 (EA).
Ehretia cymosa var. silvatica (Gürke) Brenan Habit: Tree. Habitat: Forest, grassland,
and bushland, ca. 10 m. Voucher: Luke WRQ 3073 (EA).
Euploca ovalifolia (Forsk.) Diane & Hilger Habit: Herb. Habitat: Bushland, woodland, and grassland, ca. 15 m. Voucher: Luke Q 5657 (EA).
Euploca strigosa (Willd.) Diane & Hilger Habit: Herb. Habitat: Woodland, grassland, and along road sides, ca. 186 m. Vouchers: SAJIT–006134, Ngumbau V &
Mwadime N V0343 (EA, HIB), Luke Q 1550, Graham RM 1901 (EA).
Checklist of coastal forests of Kenya
53
Heliotropium benadirense Chiov. Habit: Herb. Habitat: Sand dunes and sandy places, 0–120 m. Voucher: Luke Q 5488 (EA).
Heliotropium gorinii Chiov. Habit: Herb. Habitat: Above high tide and roadsides,
0–15 m. Vouchers: Verdcourt 3909, Rayner 296a, Greenway 13138 (EA).
Heliotropium pectinatum Vaupel subsp. pectinatum Habit: Herb. Habitat: Grassland,
and Commiphora-Acacia bushland, 15–620 m. Voucher: Polhill & Paulo 593 (EA).
Heliotropium simile Vatke Habit: Herb. Habitat: Bushland and scrub, ca. 150 m.
Voucher: Mungai et al. 112 (EA).
Heliotropium sp. A of FTEA Habit: Herb. Habitat: Grassland with scattered Acacia,
ca. 96 m. Voucher: Gillett 16522 (EA).
Heliotropium steudneri Vatke subsp. steudneri Habit: Herb. Habitat: Bushland and
grassland, 0–200 m. Vouchers: Ngumbau V & Mwadime N V0455 (EA, HIB),
Luke Q 5658 (EA).
Heliotropium zeylanicum (Burm.f.) Lam. Habit: Herb. Habitat: Bushland or grassland, ca. 20 m. Voucher: Luke Q 5659 (EA).
Hilsenbergia nemoralis (Gürke) J.S. Mill. Habit: Tree. Habitat: Forest, woodland,
and coral, 0–600 m. Vouchers: Drummond & Hemsley 1106, Magogo & Glover
727, J Adamson 289 (EA).
Hilsenbergia petiolaris (Lam.) J. S. Mill. Habit: Shrub. Habitat: Forest, woodland,
and coral, 0–30 (–660) m. Vouchers: SAJIT–005604, Ngumbau V & Mwadime
N V0298 (EA, HIB), Bridson 129, Jeffery K75, Verdcourt 1081, Festo L & Luke
Q 2561 (EA).
Hilsenbergia teitensis (Gürke) J. S. Mill. Habit: Shrub. Habitat: Bushland and forest,
60–850 m. Vouchers: SAJIT–005422 (EA, HIB), Drummond & Hemsley 4267 (EA).
Trichodesma indicum (L.) Lehm. Habit: Herb. Habitat: Grassland and coconut
plantations, 0–45 m. Voucher: Linder 2650 (EA).
Trichodesma zeylanicum (Burm. f.) R. Br. Habit: Herb. Habitat: Forest, 15 m.
Voucher: GW Sangai 15597 (EA).
F44. Burmanniaceae
1 Genus, 1 Species
Afrothismia baerae Cheek Habit: Herb. Habitat: Evergreen coastal, 360–400 m.
Voucher: Baer S s.n. (EA): Critically Endangered, Endemic.
F45. Burseraceae
2 Genera, 13 Species
Boswellia neglecta S. Moore Habit: Tree. Habitat: Bushland, ca. 400–620 m. Vouchers: Mwadime N2, Luke WRQ 196, Robertson SA & Khayota B 6416 (EA).
Commiphora africana (A. Rich.) Engl. var. africana Habit: Tree. Habitat: Grassland
and bushland, ca. 110 m. Vouchers: Kuchar P 12902, Kibuwa SP 1224 (EA).
Commiphora africana var. glaucidula (Engl.) J. B. Gillett Habit: Tree. Habitat:
Bushed grassland, ca. 450 m. Vouchers: Rawlins SP 130, Luke WRQ 915 (EA).
54
Veronicah M. Ngumbau et al. / PhytoKeys 147: 1–191 (2020)
Commiphora africana var. oblongifoliolata (Engl.) J. B. Gillett Habit: Tree. Habitat: Bushed grassland, ca. 20 m. Vouchers: Gillett JB 20354, Cunningham van
Someren GR Sh 82, Kibuwa SP 19845 (EA).
Commiphora campestris subsp. glabrata (Engl.) J. B. Gillett Habit: Tree. Habitat:
Bushland, 10–200 m. Voucher: Faden 77/476 (EA).
Commiphora edulis subsp. boiviniana (Engl.) J. B. Gillett Habit: Tree. Habitat:
Bushland and thickets dominated by Sapotaceae, ca. 70 m. Vouchers: van Someren
VGL Sh 61, Robertson SA 4083 (EA).
Commiphora eminii subsp. zimmermannii (Engl.) J. B. Gillett Habit: Tree. Habitat: Forest, 5–110 m. Vouchers: Luke WRQ & Robertson SA 212, RB & AJ
Faden77/650 (EA).
Commiphora erosa Vollesen Habit: Tree. Habitat: Woodland, 90–400 m. Voucher:
Robertson SA 1849 (EA).
Commiphora kua (R. Br. ex Royle) Vollesen Habit: Tree. Habitat: Coastal thicket,
2–360 m. Vouchers: SAJIT–006237 (EA, HIB), Robertson SA 4082, Polhill R &
Paulo S 495 (EA).
Commiphora obovata Chiov. Habit: Shrub. Habitat: Coastal bushland, 50–100 m.
Vouchers: Gillett JB 20353, Mahasi AS in EAH 14890 (EA): Near Threatened.
Commiphora pseudopaolii J. B. Gillett Habit: Tree. Habitat: Bushland, 60–600 m.
Vouchers: Gillett 16374, Verdcourt & Polhill 2709 (EA): Near Threatened.
Commiphora pteleifolia Engl. Habit: Shrub. Habitat: Coastal thicket, ca. 10 m. Vouchers: Luke Q 1477, RB Faden, AJ Faden, JB Gillett & N Gachathi 77/475 (EA).
Commiphora sennii Chiov. Habit: Shrub. Habitat: Bushland, ca. 200 m. Voucher:
Gillet & Gachathi 20535 (EA).
Commiphora unilobata J. B. Gillett & Vollesen Habit: Shrub. Habitat: Bushland,
ca. 70 m. Vouchers: Gillett 21108 & 21109 (EA).
Commiphora zanzibarica (Baill.) Engl. Habit: Tree. Habitat: Coastal thickets and
coral, 2–510 m. Vouchers: Willan 341, RB & SA Faden 77/650, SA Robertson
3663 (EA).
F46. Buxaceae
1 Genus, 1 Species
Buxus obtusifolia (Mildbr.) Hutch. Habit: Shrub. Habitat: Forest, 280–450 m.
Vouchers: SAJIT–005537, Ngumbau V & Mwadime N V0496 (EA, HIB), Mutanga JG & Kamau 6, Hawthorne W 139 (EA): Vulnerable.
F47. Cactaceae
2 Genera, 2 Species
Opuntia ficus-indica (L.) Mill. Habit: Shrub. Habitat: Dry arid areas, 0–300 m.
Voucher: Luke WRQ & PA sr: Cultivated.
Checklist of coastal forests of Kenya
55
Rhipsalis baccifera (Sol.) Stearn Habit: Epiphytic herb. Habitat: Wet forests and
riverine forest, ca. 145 m. Vouchers: Faden RB & Beentje HJ 85/47, Magogo FC
& Glover PE 633 (EA).
F48. Calophyllaceae
1 Genus, 1 Species
Calophyllum inophyllum L. Habit: Tree. Habitat: Rocky and sandy seashores,
0–20 m. Voucher: Robertson SA 4926 (EA).
F49. Campanulaceae
2 Genera, 4 Species
Lobelia fervens Thunb. subsp. fervens Habit: Herb. Habitat: Grassland, forest margins, roadsides, streamside or on coastal sand, ca. 136 m. Vouchers: SAJIT–005960
& 005945 (EA, HIB), Magogo FC & Glover PE 734, Polhill & Paulo 488 (EA).
Lobelia fervens subsp. recurvata (E. Wimm.) Thulin Habit: Herb. Habitat: On
moist ground in grassland or woodland, ca. 15 m. Vouchers: Mwadime N & Luke
WRQ 784 & 419 (EA).
Wahlenbergia abyssinica (Hochst. ex A. Rich.) Thulin subsp. abyssinica Habit:
Herb. Habitat: Cultivated or waste ground, ca. 300 m. Vouchers: Magogo FC &
Glover PE 121, Thulin M 300 (EA).
Wahlenbergia napiformis (A. DC.) Thulin Habit: Herb. Habitat: Woodland, grassland, and roadsides, ca. 300 m. Vouchers: Luke WRQ & Robertson SA 2823,
Thulin M 298 (EA).
F50. Canellaceae
1 Genus, 1 Species
Warburgia stuhlmannii Engl. Habit: Tree. Habitat: Forest and woodland, 20–225 m.
Vouchers: Luke WRQ 15340, Graham 2208 (EA): Vulnerable.
F51. Cannabaceae
2 Genera, 3 Species
Celtis mildbraedii Engl. Habit: Tree. Habitat: Lowland rainforest, ca. 160–300 m.
Vouchers: Mwadime N 2, Luke WRQ & Robertson SA 2810, RB Faden & AJ
Faden 77/678, RB Faden, AJ Faden & N Gachathi 77/526 (EA).
Celtis philippensis Blanco Habit: Tree. Habitat: Lowland, groundwater, and riverine forest, ca. 30–207 m. Vouchers: SAJIT–006021 (EA, HIB), Mwadime N 30,
Luke WRQ & Robertson SA 504, Thomas Mwadime in Mrs Robertson SA 7807,
Drummond & Hemsley 4014, Verdcourt 2405 (EA).
56
Veronicah M. Ngumbau et al. / PhytoKeys 147: 1–191 (2020)
Trema orientalis (L.) Blume Habit: Tree. Habitat: Margins of lowland and upland
rainforest, ca. 280 m. Vouchers: SAJIT–005457 (EA, HIB), Jeffery 193 (EA).
F52. Capparaceae
7 Genera, 37 Species
Boscia angustifolia A. Rich. var. angustifolia Habit: Tree. Habitat: Woodland and
bushland, ca. 200 m. Voucher: Robertson SA & Luke Q 4721 (EA).
Boscia coriacea Pax Habit: Tree. Habitat: Bushland and grassland, ca. 150 m. Vouchers: Kuchar P 13709, Drummond RB 4093, Graham RM 1586 (EA).
Boscia keniensis Beentje Habit: Shrub. Habitat: Bushland, ca. 100–350 m. Vouchers:
Bally 2024, Robertson SA 319 (EA).
Boscia mossambicensis Klotzsch Habit: Tree. Habitat: Woodland, bushland, and
thicket, ca. 630 m. Voucher: Luke PA WRQ 4400 (EA).
Boscia salicifolia Oliv. Habit: Tree. Habitat: Woodland, bushland, bamboo thicket,
and grassland, ca. 300 m. Vouchers: Burstyne P 70, Bally J 13944 (EA).
Cadaba carneoviridis Gilg & Gilg-Ben. Habit: Shrub. Habitat: Bushland and grassland, 30–600 m. Vouchers: Gilbert MG 5862, Robertson SA 2086 (EA).
Cadaba farinosa Forssk. subsp. farinosa Habit: Shrub. Habitat: Bushland and grassland, 0–300 m. Vouchers: Greenway PJ 9440, Moomaw JC 1578 (EA).
Cadaba farinosa subsp. adenotricha (Gilg & Gilg-Ben.) R. A. Graham Habit:
Shrub. Habitat: Bushland and grassland, 0–400 m. Vouchers: Luke WRQ et al.
5411, Rayner RW 163 (EA).
Cadaba gillettii R.A. Graham Habit: Shrub. Habitat: Bushland, 100–750 m. Voucher: Mrs J Adamson 103 (EA).
Cadaba glandulosa Forssk. Habit: Shrub. Habitat: Bushland and grassland, ca.
600 m. Voucher: Greenway 8866 (EA).
Cadaba rotundifolia Forssk. Habit: Shrub. Habitat: Bushland, 0–400 m. Voucher:
SAJIT–005425 (EA, HIB).
Cadaba ruspolii Gilg Habit: Shrub. Habitat: Bushland and woodland, ca. 180 m.
Voucher: Bally 2010 (EA).
Capparis erythrocarpos var. rosea (Klotzsch) De Wolf Habit: Shrub. Habitat:
Woodland and bushland, 0–400 m. Vouchers: SAJIT–006432, Ngumbau V &
Mwadime N V0497 (EA, HIB), Greenway PJ & Rawlins SP 9353 (EA).
Capparis fascicularis DC. var. fascicularis Habit: Shrub. Habitat: Bushland and
grassland, 600 m. Voucher: Battiscombe 800 (EA).
Capparis fascicularis var. scheffleri (Gilg & Benedict) DeWolf Habit: Shrub. Habitat: Forest and bushland, ca. 0–400 m. Vouchers: Ngumbau V & Mwadime N
V0445 (EA, HIB), Sangai GW 15690, Luke WRQ 3475 (EA).
Capparis sepiaria var. fischeri (Pax) DeWolf Habit: Shrub. Habitat: Bushland and
grassland, 180 m. Voucher: Battiscombe 269 (EA).
Capparis sepiaria var. stuhlmannii (Gilg) DeWolf Habit: Shrub. Habitat: Woodland, bushland, and riverine forest, 0–300 m. Vouchers: Rawlins SP 168, Magogo
FC & Glover PE 684 (EA).
Checklist of coastal forests of Kenya
57
Capparis sepiaria var. subglabra (Oliv.) DeWolf Habit: Shrub. Habitat: Bushland,
0–200 m. Vouchers: Luke Q 1441, Mrs. J Adamson 299 RM in FD 1612, Bally
1991 (EA).
Capparis spinosa var. aegyptia (Lam.) Boiss. Habit: Liana. Habitat: Bushland and
coral outcrops, 0–200 m. Vouchers: SAJIT–005575 (EA, HIB), Taiti S 19 (EA).
Capparis tomentosa Lam. Habit: Liana. Habitat: Bushland, grassland, and riverine,
0–200 m. Vouchers: Ngumbau V & Mwadime N V0190 (EA, HIB), Kuchar P
12903, Sangai GW EA 15762 (EA).
Capparis viminea Hook. f. & Thomson ex Oliv. var. viminea Habit: Shrub. Habitat: Forest, ca. 50 m. Voucher: Luke & Robertson 1562 (EA).
Cladostemon kirkii (Oliv.) Pax & Gilg Habit: Tree. Habitat: Forest or thicket,
0–800 m. Vouchers: Festo L & Luke Q 2562, Dale IR 3795 (EA).
Maerua angolensis DC. Habit: Tree. Habitat: Woodland, bushland, and grassland,
0–300 m. Vouchers: Robertson SA 7763 & 5075, Drummond & Hemsley
4218 (EA).
Maerua calantha Gilg Habit: Shrub. Habitat: Bushland, riverine, and, woodland,
45–460 m. Vouchers: Gillett JB 20401, F Thomas 65 (EA).
Maerua crassifolia Forssk. Habit: Tree. Habitat: Bushland or semi-desert scrub, ca.
180 m. Voucher: Robertson SA & Luke WRQ 5585 (EA).
Maerua decumbens (Brongn.) DeWolf Habit: Shrub. Habitat: Bushland, 50 m.
Voucher: Robertson SA 7635 (EA).
Maerua denhardtiorum Gilg Habit: Shrub. Habitat: Bushland, 90–780 m. Voucher:
Sampson 1(EA).
Maerua endlichii Gilg & Gilg-Ben. Habit: Shrub. Habitat: Bushland, ca. 23 m.
Voucher: Ngumbau V & Mwadime N V0456 (EA, HIB).
Maerua glauca Chiov. Habit: Shrub. Habitat: Bushland and thicket, below 100 m.
Vouchers: Luke Q & Robertson 2148, Greenway 8964 (EA).
Maerua grantii Oliv. Habit: Shrub. Habitat: Woodland, bushland, and grassland, ca.
50 m. Voucher: Rawlins SP 452 (EA).
Maerua holstii Pax Habit: Liana. Habitat: Lowland rainforest and woodland, 30–
450 m. Vouchers: Ngumbau V & Mwadime N V0486 (EA, HIB), Gillett JB
20387 (EA).
Maerua kaessneri Gilg & Gilg-Ben. Habit: Shrub. Habitat: Bushland, 150–350 m.
Voucher: Robertson SA & Luke WRQ 5685 (EA).
Maerua kirkii (Oliv.) F. White Habit: Shrub. Habitat: Woodland, bushland, and riverine forest, ca. 100 m. Vouchers: Luke WRQ & Robertson SA 183, Luke WRQ
3330 (EA).
Maerua macrantha Gilg Habit: Shrub. Habitat: Woodland and bushland, 30–50 m.
Vouchers: Festo L, Luke Q & P 2709, Rawlins 395, Bally 2155, Drummond &
Hemsley 4038 (EA).
Maerua mungaii Beentje Habit: Shrub. Habitat: Woodland, 100–150 m. Voucher:
Parker I 513/H (EA): Endemic.
Maerua sessiliflora Gilg Habit: Shrub. Habitat: Deciduous bushland, semi-desert
scrub, 300–750 m. Voucher: Bally 2187 (EA).
58
Veronicah M. Ngumbau et al. / PhytoKeys 147: 1–191 (2020)
Maerua triphylla A. Rich. var. triphylla Habit: Shrub. Habitat: Forest, ca. 181 m.
Vouchers: Ngumbau V & Mwadime N V0223 (EA, HIB).
Maerua triphylla var. calophylla (Gilg) DeWolf Habit: Shrub. Habitat: Bushland,
grassland, and riverine, ca. 60–200 m. Vouchers: Ngumbau V & Mwadime N
V0412 (EA, HIB), Robertson SA & Luke WRQ 5575 (EA).
Maerua triphylla var. johannis (Volkens & Gilg) DeWolf Habit: Shrub. Habitat: Bushland and grassland, ca. 200 m. Vouchers: Bally 14389, Drummond
3868 (EA).
Maerua triphylla var. pubescens (Klotzsch) DeWolf Habit: Shrub. Habitat: Rainforest, bushland, and grassland, ca. 20 m. Vouchers: Adamson J 6138, Roberson SA
4256 (EA).
Ritchiea capparoides (Andrews) Britten Habit: Shrub. Habitat: Forest and bushland,
0–300 m. Vouchers: Ngumbau V & Mwadime N V0379 (EA, HIB), Hooper SS
& Townsend CC 1214 (EA).
Thilachium africanum Lour. Habit: Shrub. Habitat: Woodland, bushland, and grassland, 25–300 m. Vouchers: Luke WRQ & PA 5696, Graham MD 1536 (EA).
Thilachium roseomaculatum Y.B. Harv. & Vollesen Habit: Herb. Habitat: Denser
forest parts, 50–150 m. Vouchers: Harvey, Mwachala & Vollesen 49, Robertson &
Luke 6251, Luke 3454 & 3472 (EA): Endemic.
Thilachium thomasii Gilg Habit: Shrub. Habitat: Woodland and bushland, ca. 50 m.
Vouchers: SAJIT–004646 (EA, HIB), Rawlins SP 133 & 9426, Drummond &
Hemsley 4052 (EA).
F53. Cardiopteridaceae
1 Genus, 1 Species
Leptaulus holstii (Engl.) Engl. Habit: Shrub. Habitat: Lowland rainforests, ca. 289 m.
Vouchers: Ngumbau V & Mwadime N V032 (EA, HIB), Luke WRQ 1852 (EA).
F54. Caryophyllaceae
1 Genus, 5 Species
Polycarpaea corymbosa (L.) Lam. Habit: Herb. Habitat: Roadside and grassland, ca.
30 m. Voucher: Festo L, Luke Q & P 2653 (EA).
Polycarpaea eriantha Hochst. ex A. Rich. Habit: Herb. Habitat: Grassland, ca.
480 m. Voucher: Magogo FC & Estes R 1246 (EA).
Polycarpaea grahamii Turrill Habit: Herb. Habitat: Near the shore. Voucher: Graham MD 1617 (EA): Endemic.
Polycarpaea spicata Wight ex Arn. Habit: Herb. Habitat: Woodland, 0–500 m.
Voucher: Luke WRQ 5460 (EA).
Polycarpaea tenuistyla Turrill Habit: Herb. Habitat: Bushland, ca. 25 m. Voucher:
Musyoki BM & Hansen OJ 982 (EA): Endemic.
Checklist of coastal forests of Kenya
59
F55. Casuarinaceae
1 Genus, 1 Species
Casuarina equisetifolia L. Habit: Tree. Habitat: Coastal bushland and seashore, ca.
2 m. Vouchers: SAJIT–006252 (EA, HIB), Greenway 9619, MacNaughton159,
Whellan 1851 (EA): Naturalized.
F56. Celastraceae
12 Genera, 29 Species
Apodostigma pallens (Planch. ex Oliv.) R. Wilczek var. pallens Habit: Liana.
Habitat: Forest, woodland, and riverine, ca. 0–239 m. Vouchers: Ngumbau V &
Mwadime N V078 & V0306 (EA, HIB), Magogo FC & Glover PE 969, Luke
WRQ 3601 (EA).
Apodostigma pallens var. dummeri N. Hallé Habit: Liana. Habitat: Forest, ca. 20 m.
Vouchers: Mwadime N 18, Luke WRQ 16135 (EA).
Elachyptera holtzii (Loes. ex Harms) R. Wilczek Habit: Liana. Habitat: Evergreen
forest to semi-evergreen forest, 120–460 m. Voucher: Luke WRQ 3601 (EA).
Elachyptera parvifolia (Oliv.) N. Hallé Habit: Liana. Habitat: Forest, woodland,
ca. 30 m. Vouchers: SAJIT–005527 (EA, HIB), Festo L, Luke Q & P 2696 (EA).
Elaeodendron schlechterianum (Loes.) Loes. Habit: Shrub. Habitat: Woodland and
forest, 0–410 m. Vouchers: Ngumbau V & Mwadime N V0303 (EA, HIB), Luke
Q 1265, van Someren VGL Sh103 (EA).
Elaeodendron schweinfurthianum (Loes.) Loes. Habit: Shrub. Habitat: Coastal
bushland and forest, 0–30 m. Voucher: Magogo FC & Glover PE 1133 (EA).
Gymnosporia buchananii Loes. Habit: Shrub. Habitat: Forest, 60–400 m. Vouchers:
Luke WRQ & Robertson SA 500, Magogo FC & Glover PE 267 (EA).
Gymnosporia gracilis Loes. subsp. gracilis Habit: Shrub. Habitat: Evergreen forest
margins and thicket, 0–500 m. Vouchers: SAJIT–005464 (EA, HIB), Magogo FC &
Glover PE 289, Adamson J 5834, Spjut 4550, Brenan, Gillett & Kanuri 14675 (EA).
Gymnosporia heterophylla (Eckl. & Zeyh.) Loes. Habit: Shrub. Habitat: Forest margins and woodland thicket, 0–100 m. Vouchers: Luke Q 1546, Graham RM FD
Ox352 (EA).
Gymnosporia masindei (Gereau) Jordaan Habit: Shrub. Habitat: Bushland, 30–
245 m. Voucher: Luke WRQ & Robertson SA 2498 (EA).
Gymnosporia putterlickioides Loes. Habit: Tree. Habitat: Woodland and riverine, ca.
30 m. Vouchers: Luke Q 1430, Festo, L & Luke Q 2600, Polhill & Paulo 873 (EA).
Gymnosporia senegalensis (Lam.) Loes. Habit: Shrub. Habitat: Woodland, grassland, and riverine, ca. 45 m. Voucher: Nash LT 40 (EA).
Loeseneriella africana var. richardiana (Cambess.) R. Wilczek ex N. Hallé Habit:
Liana. Habitat: Riverine forest and thickets, ca. 372 m. Vouchers: SAJIT–006159
(EA, HIB), Mwadime N et al. 615, Luke WRQ 1392 (EA).
60
Veronicah M. Ngumbau et al. / PhytoKeys 147: 1–191 (2020)
Loeseneriella crenata (Klotzsch) R. Wilczek ex N. Hallé var. crenata Habit: Liana.
Habitat: Forest, bushland, 0–80 m. Voucher: Luke Q 1419 (EA).
Maytenus undata (Thunb.) Blakelock Habit: Tree. Habitat: Forest, riverine, and
woodland, 0–407 m. Vouchers: SAJIT–006054 & 005549, Ngumbau V &
Mwadime N V0525 (EA, HIB), Luke WRQ & PA 6255 (EA).
Mystroxylon aethiopicum (Thunb.) Loes. subsp. aethiopicum Habit: Tree. Habitat:
Coastal woodland, 0–250 m. Vouchers: SAJIT–005901, Ngumbau V & Mwadime
N V0128 (EA, HIB), Magogo FC & Glover PE 507 (EA).
Pleurostylia africana Loes. Habit: Tree. Habitat: Woodland and forest, 0–500 m.
Vouchers: Luke Q 1429, RB Faden & AJ Faden 77/707 (EA).
Pristimera andongensis var. volkensii (Loes.) N. Hallé & B. Mathew Habit: Liana.
Habitat: Bushland and woodland, ± 100 m. Vouchers: Mwadime N 19, Gray M
& Luke WRQ 302 (EA).
Pristimera longipetiolata (Oliv.) N. Hallé Habit: Liana. Habitat: Forest margins,
0–15 m. Voucher: RB & AJ Faden 74/1148 (EA).
Salacia elegans Welw. ex Oliv. Habit: Liana. Habitat: Forest and woodland, 0–250 m.
Vouchers: SAJIT–005990 (EA, HIB), Luke Q 1448, Luke WRQ & Robertson SA
2739 (EA).
Salacia erecta (G. Don) Walp. Habit: Liana. Habitat: Evergreen forest, ca. 50 m.
Voucher: Luke WRQ & Robertson SA 521 (EA).
Salacia lehmbachii Loes. Habit: Shrub. Habitat: Forest, ca. 45 m. Vouchers: SAJIT–005554 (EA, HIB), Luke WRQ et al. 4725 (EA).
Salacia leptoclada Tul. Habit: Liana. Habitat: Forest, ca. 70 m. Vouchers: Luke Q
1502, Robertson SA & Luke WRQ 6024 (EA).
Salacia madagascariensis (Lam.) DC. Habit: Liana. Habitat: Forest and woodland,
0–600 m. Vouchers: Ngumbau V & Mwadime N V0140, SAJIT–006176 (EA,
HIB), Luke Q 43 (EA).
Salacia sp. cf. erecta (G. Don) Walp. Habit: Shrub. Habitat: Forest. Voucher: Birch
62/222 (EA).
Salacia stuhlmanniana Loes. Habit: Habitat: Forest, riverine, bushland, and woodland, 0–36 m. Vouchers: SAJIT–006175 (EA, HIB), Robertson SA 5470 (EA).
Simicratea welwitschii (Oliv.) N. Hallé Habit: Liana. Habitat: Forest, ca. 210 m.
Vouchers: Ngumbau V & Mwadime N V0377 (EA, HIB), Mwadime N 20, Luke
WRQ & Robertson SA 2819 (EA).
Simirestis goetzei (Loes.) N. Hallé ex R. Wilczek Habit: Liana. Habitat: Forest, ca.
90 m. Voucher: Kimeu JM 689 (EA).
Simirestis scheffleri (Loes.) N. Hallé Habit: Liana. Habitat: Forest, ca. 100 m. Voucher: Moggridge 519 (EA).
F57. Ceratophyllaceae
1 Genus, 1 Species
Ceratophyllum demersum L. Habit: Herb. Habitat: Water bodies, lakes, rivers, and
swamps, ca. 0–15 m. Vouchers: SAJIT–006227 (EA, HIB), Rawlins SP 428 (EA).
Checklist of coastal forests of Kenya
61
F58. Chrysobalanaceae
2 Genera, 2 Species
Dactyladenia barteri (Hook. f. ex Oliv.) Prance & F. White Habit: Tree. Habitat: Forest, ca. 410 m. Voucher: Ngumbau V & Mwadime N V0304 (EA,
HIB): Cultivated.
Hirtella zanzibarica Oliv. subsp. zanzibarica Habit: Tree. Habitat: Forest, ca.
407 m. Vouchers: SAJIT–006030, 005517 & 006063, Ngumbau V & Mwadime
M V0250 (EA, HIB), Luke WRQ & Robertson SA 239 (EA).
F59. Cleomaceae
4 Genera, 9 Species
Cleome gynandra L. Habit: Herb. Habitat: Waste, disturbed or cultivated ground, ca.
20 m. Voucher: Jeffery K206 (EA).
Coalisina tenella (L. f.) Roalson & J.C. Hall Habit: Herb. Habitat: Hyphaene grassland, ca. 5–360 m. Voucher: Jeffery K206 (EA).
Sieruela allamani (Chiov.) Roalson & J.C. Hall Habit: Herb. Habitat: Bushland,
semi-desert scrub, 150–175 m. Vouchers: Mrs J Adamson 616, Gillett 19508 (EA).
Sieruela briquetii (Polhill) Roalson & J.C. Hall Habit: Herb. Habitat: Bushland
and roadside, ca. 0–210 m. Vouchers: Simpson BL 361, Magogo FC & Glover PE
512 & 1205 (EA).
Sieruela hirta (Klotzsch) Roalson & J. C. Hall Habit: Herb. Habitat: Bushland and
grassland, ca. 175 m. Voucher: Festo L & Luke Q 2705 (EA).
Sieruela parvipetala (R.A. Graham) Roalson & J.C. Hall Habit: Herb. Habitat:
Acacia-Commiphora-Lannea-Boswellia bushland 120–420 m. Vouchers: Mungai
GM & Rucina SM 416/84, JB Gillett 16400, J Makin 14472 (EA).
Sieruela strigosa (Bojer) Roalson & J.C. Hall Habit: Herb. Habitat: Sandy seashores, ± 0 m. Vouchers: SAJIT–006262 (EA, HIB), Greenway PJ & Rawlins SP
9422 (EA).
Sieruela usambarica (Pax ex Engl.) Roalson & J.C. Hall Habit: Herb. Habitat:
Rainforest and bushland, ca. 30 m. Vouchers: Simpson BL 156, Robertson SA &
Luke WRQ 6020 (EA).
Stylidocleome brachycarpa (Vahl ex DC.) Roalson & J.C. Hall Habit: Herb. Habitat: Acacia-Commiphora bushland. Vouchers: Lucas 34, JB Gillett 21126/B (EA).
F60. Clusiaceae
1 Genus, 3 Species
Garcinia buchananii Baker Habit: Tree. Habitat: Evergreen forest, often riverine,
and thickets, ca. 85 m. Vouchers: Ngumbau V & Mwadime N V0485 (EA, HIB),
Luke WRQ & Robertson SA 1642 & 1899 (EA).
Garcinia livingstonei T. Anderson Habit: Tree. Habitat: Woodland, thicket, and
grassland, ca. 5 m. Vouchers: SAJIT–006092 (EA, HIB), Robertson SA 4314 (EA).
62
Veronicah M. Ngumbau et al. / PhytoKeys 147: 1–191 (2020)
Garcinia volkensii Engl. Habit: Shrub. Habitat: Evergreen forest, ca. 210 m. Vouchers: Ngumbau V & Mwadime N V0390 (EA, HIB), Luke Q & Robertson 2655,
Luke WRQ 3973 (EA).
F61. Colchicaceae
2 Genera, 2 Species
Gloriosa superba L. var. superba Habit: Herb. Habitat: Forest margin and coastal bushland, ca. 290 m Vouchers: SAJIT–006261 & 005544, Ngumbau V &
Mwadime N V0287 (EA, HIB), Magogo FC & Glover PE 973 (EA).
Iphigenia oliveri Engl. Habit: Herb. Habitat: Grassland and bushland, ca. 0–30 m.
Vouchers: Adams 12, Luke et al. 5925, Luke & Luke 4379 (EA).
F62. Combretaceae
5 Genera, 31 Species
Combretum aculeatum Vent. Habit: Shrub. Habitat: Bushland, woodland, and riverine, ca. 15 m. Vouchers: Mwadime N 460, 489 & 656, Luke WRQ 121 (EA).
Combretum apiculatum Sond. subsp. apiculatum Habit: Tree. Habitat: Woodland,
ca. 150 m. Vouchers: Msafiri F & Mwendwa H 441, Luke WRQ 3471 (EA).
Combretum butyrosum (G. Bertol.) Tul. Habit: Liana. Habitat: Riverine forest, 50–
450 m. Vouchers: Ngumbau V & Mwadime N V0453 (EA, HIB), Festo L, Luke
Q & P 2755 (EA).
Combretum chionanthoides Engl. & Diels Habit: Liana. Habitat: Forest margin,
50–170 m. Vouchers: Ngumbau V & Mwadime N V0413 (EA, HIB), Brenan
JPM & JH Gillet JB 14657 (EA).
Combretum constrictum (Benth.) M. A. Lawson Habit: Climber. Habitat: Swamps
and riverine forest, ca. 15 m. Vouchers: RM Graham in FD 2243, Festo 2419 (EA).
Combretum contractum Engl. & Diels Habit: Shrub. Habitat: Bushland, 150–396
m. Vouchers: Graham RM 2047, Bally 8162, Dale 2049 (EA).
Combretum exalatum Engl. Habit: Shrub. Habitat: Bushland, ca. 300 m. Vouchers:
Festo L, Luke Q & P 2789, Robertson SA 6396 (EA).
Combretum falcatum (Welw. ex Hiern) Jongkind Habit: Liana. Habitat: Bushland
and forest margins, 0–70 m. Vouchers: Ngumbau V & Mwadime N V0530 &
V0436 (EA, HIB), Luke WRQ 894 (EA).
Combretum hereroense Schinz subsp. hereroense Habit: Shrub. Habitat: Grassland
and bushland, ca. 70 m. Vouchers: Ngumbau V & Mwadime N V0448 (EA,
HIB), Dale in FD 3666, Robertson SA & Luke WRQ 5715 (EA).
Combretum hereroense subsp. volkensii (Engl.) Wickens Habit: Shrub. Habitat:
Grassland or bushland, 50–500 m. Vouchers: Power 10994, Hildebrandt 2337,
Luke 916 (EA).
Combretum holstii Engl. Habit: Liana. Habitat: Woodland and riverine forest, ca.
0–300 m. Voucher: Robertson SA 5084 (EA).
Checklist of coastal forests of Kenya
63
Combretum illairii Engl. Habit: Liana. Habitat: Forest, woodland, and bushland,
30–800 m. Vouchers: SAJIT–004665 & 005569, Ngumbau V & Mwadime N
V0228 (EA, HIB), Kuchar P 13482 (EA).
Combretum molle R.Br. ex G. Don Habit: Tree. Habitat: Wooded grassland, 30–340
m. Vouchers: Spjut RW 3809, Shimba Hills Survey Unit 30, Drummond RB &
Hemsley JH 4101 (EA).
Combretum padoides Engl. & Diels Habit: Liana. Habitat: Riverine forest, swamp forest, 0–280 m. Vouchers: SAJIT–006070, Ngumbau V & Mwadime N V0324 (EA,
HIB), Robertson SA & Luke WRQ 5647, Luke WRQ & Robertson SA 2718 (EA).
Combretum paniculatum Vent. Habit: Liana. Habitat: Forest, woodland, and riverine forest, 10–156 m. Vouchers: Ngumbau V & Mwadime N V0429 (EA, HIB),
Graham RM 1694, Rawlins SP 253 (EA).
Combretum pentagonum M.A. Lawson Habit: Liana. Habitat: Forest and thicket,
20–153 m. Vouchers: Ngumbau V & Mwadime N V0505 (EA, HIB), Robertson
SA & Luke WRQ 4516 (EA).
Combretum schumannii Engl. Habit: Tree. Habitat: Forest, riverine forest, and
woodland, 0–140 m. Vouchers: Ngumbau V & Mwadime N V0538 (EA, HIB),
Kuchar P 13548 (EA).
Combretum tenuipetiolatum Wickens Habit: Tree. Habitat: Coastal forest, ca.
100 m. Voucher: Robertson SA & Luke WRQ 2289 (EA): Critically Endangered.
Combretum umbricola Engl. Habit: Liana. Habitat: Riverine forest and coastal
woodland, ca. 280 m. Voucher: Luke WRQ & Robertson SA 2717 (EA).
Conocarpus lancifolius Engl. & Diels Habit: Tree. Habitat: Semi-desert coastal zone,
10–570 m. Voucher: Gillett JB 20314 (EA): Near Threatened, Cultivated.
Lumnitzera racemosa Willd. Habit: Shrub. Habitat: Mangrove swamps, 0–30 m.
Vouchers: Mrs Robertson SA 7747, Luke Q 5649, Brathay 1982 Expedition
55 (EA).
Pteleopsis myrtifolia (M.A. Lawson) Engl. & Diels Habit: Tree. Habitat: Riverine, forest, woodland, bushland, and grassland, 0–150 m. Vouchers: Ngumbau
V & Mwadime N V0530 (EA, HIB), Luke WRQ & Robertson SA 2822, Faden
70/256 (EA).
Pteleopsis tetraptera Wickens Habit: Tree. Habitat: Bushland, wooded grassland, and
forest, 0–300 m. Vouchers: Luke WRQ & Robertson SA 5531, Luke WRQ &
Robertson SA 2796 (EA).
Terminalia boivinii Tul. Habit: Shrub. Habitat: Bushland and forest, 0–30 m.
Vouchers: SAJIT–005585 (EA, HIB), Festo L, Luke Q & P 2699, Moomaw 1602,
Donald 96 in FD 2496 (EA).
Terminalia brevipes Pamp. Habit: Tree. Habitat: Riverine forest and bushland, 20–
280 m. Vouchers: Brathay Expedition 46, Tweedie 1866 (EA).
Terminalia catappa L. Habit: Tree. Habitat: Sandy or rocky beaches, ca. 5 m. Vouchers: Ndakala J 407, Faden RB & Faden AJ 77/390 (EA): Naturalized.
Terminalia parvula Pamp. Habit: Tree. Habitat: Bushland, ca. 60 m. Voucher: Bally
16708 (EA).
64
Veronicah M. Ngumbau et al. / PhytoKeys 147: 1–191 (2020)
Terminalia prunioides M.A. Lawson Habit: Tree. Habitat: Bushland and riverine
thicket, 30–360 m. Vouchers: Ngumbau V & Mwadime N V0453 (EA, HIB),
Rawlins SP 335, Luke WRQ & Robertson SA 2113 (EA).
Terminalia sambesiaca Engl. & Diels Habit: Tree. Habitat: Forest, riverine forest,
and woodland, 70–830 m. Vouchers: Mohamed Abdulla 1159, Verdcourt 3934,
Dale in FD 3660 (EA).
Terminalia sp. aff. spinosa Engl. Habit: Tree. Habitat: Forest. Voucher: Luke et al.
TPR766 (EA).
Terminalia spinosa Engl. Habit: Tree. Habitat: Bushland, woodland, and grassland,
0–330 m. Vouchers: SAJIT–006440 (EA, HIB), Rawlins SP 246, Beentje HJ
2275 (EA).
F63. Commelinaceae
5 Genera, 31 Species
Aneilema aequinoctiale (P. Beauv.) Loudon Habit: Herb. Habitat: Moist or dry evergreen forests, 0–150 m. Vouchers: Robertson SA & Luke WRQ 5156, RB Faden
& AJ Faden 290 & 1270, Polhill & Paulo 825 (EA).
Aneilema calceolus Brenan Habit: Herb. Habitat: Evergreen forest, 10–450 m.
Vouchers: Faden & Faden 77/378, 74/1069 & 77/565 (EA).
Aneilema clarkei Rendle Habit: Herb. Habitat: Evergreen forest, woodland, and
thicket, 0–250 m. Vouchers: Luke Q 1457, Faden & Evans 71/714, Faden &
Faden 77/714 (EA).
Aneilema hockii De Wild. subsp. hockii Habit: Herb. Habitat: Bushland, woodland,
and grassland, 15–160 m. Vouchers: Luke & Robertson 2565, Faden & Faden
74/1065 (EA).
Aneilema indehiscens Faden subsp. indehiscens Habit: Herb. Habitat: Bushland and
thicket, ca. 15 m. Vouchers: Faden & Faden 74/380 & 74/1184 (EA).
Aneilema lamuense Faden Habit: Herb. Habitat: Sand dunes, 0–50 m. Voucher:
Faden & Faden 74/1083 (EA).
Aneilema nyasense C.B. Clarke Habit: Herb. Habitat: Forest, swamp, riverine, and
bushland, 50–300 m. Vouchers: Luke WRQ et al. 3379 & 4735 (EA).
Aneilema petersii (Hassk.) C.B. Clarke subsp. petersii Habit: Herb. Habitat: Grassland, bushland, wooded grassland, and woodland, 0–500 m. Vouchers: Luke
WRQ 3470, Q Luke & SA Robertson 2495, SA Robertson & Q Luke 6320 (EA).
Aneilema succulentum Faden Habit: Herb. Habitat: Bushland, forest, and woodland,
10–600 m. Vouchers: Faden et al. 72/39, Faden & Faden 74/1152 (EA): Endemic.
Aneilema tanaense Faden Habit: Herb. Habitat: Bushland, 10–250 m. Vouchers:
Faden & Faden 77/582, 77/ 738 & 74/1185 (EA): Endemic.
Aneilema taylorii C.B. Clarke Habit: Herb. Habitat: Riverine forest, 30–400 m.
Vouchers: Verdcourt 2410 & 1921, Faden & Faden 77/611 (EA).
Coleotrype bruechneriana Mildbr. Habit: Herb. Habitat: Forest and riverine, ca.
150–320 m. Vouchers: Luke WRQ & PA 8193, Brenan et al. 14587, Luke &
Luke 8193 (EA).
Checklist of coastal forests of Kenya
65
Commelina africana subsp. zanzibarica Faden Habit: Herb. Habitat: Forest, riverine, and grassland, 0–250 m. Vouchers: Faden & Faden 74/1123, Greenway &
Rawlins 9343 (EA).
Commelina benghalensis L. Habit: Herb. Habitat: Palm forest, 0–400 m. Vouchers:
Graham RM 2005, Meester-Manger Cats V de 38, Robertson SA 5221 (EA).
Commelina bracteosa Hassk. Habit: Herb. Habitat: Forest, woodland, grassland,
bushland, and cultivated lands, 45–305 m. Vouchers: Ngumbau V & Mwadime N
V0290 & 084 (EA, HIB), Luke Q 1456, Robertson SA & Luke WRQ 5984 (EA).
Commelina erecta L. Habit: Herb. Habitat: Forest and riverine forest, 0–550 m.
Vouchers: Ngumbau V & Mwadime N V055 &0338 (EA, HIB), Robertson
7386, Faden & Faden 74/1068 (EA).
Commelina forskaolii Vahl Habit: Herb. Habitat: Grassland, bushland, woodland, forest, cultivated lands, and riverine, ca. 30 m. Voucher: Faden & Faden
74/1249 (EA).
Commelina livingstonii C.B. Clarke Habit: Herb. Habitat: Grassland, woodland,
and bushland, ca. 100 m. Voucher: Faden & Faden 74/1189 (EA).
Commelina lukei Faden Habit: Herb. Habitat: Forest, bushland, grassland, and roadside, ca. 5 m. Vouchers: Verdcourt 1056 & 1851, SA Robertson 7377, Q Luke &
et al. 7080 (EA).
Commelina mascarenica C.B. Clarke Habit: Herb. Habitat: Roadside, grassland,
and open bushland, ca. 230 m. Vouchers: Ngumbau V & Mwadime N V0230
(EA, HIB), Rayner 294 (EA).
Commelina petersii Hassk. Habit: Herb. Habitat: Moist forest, ca. 297 m. Voucher:
Ngumbau V & Mwadime N V075 (EA, HIB).
Commelina sp. aff. bracteosa Hassk. Habit: Herb. Habitat: Forest. Voucher: R & L
6018 (EA).
Commelina sp. aff. erecta L. Habit: Herb. Habitat: Forest. Voucher: K et al.
TPR638 (EA).
Commelina zambesica C.B. Clarke Habit: Herb. Habitat: Grassland, woodland, and
forest, ca. 186 m. Vouchers: Ngumbau V & Mwadime N V0339 (EA, HIB), Magogo & Glover 70/562 (EA).
Cyanotis axillaris (L.) D. Don ex Sweet Habit: Herb. Habitat: Wooded grassland
and water bodies, 5–300 m. Voucher: Faden & Faden 74/1211 (EA).
Cyanotis foecunda DC. ex Hassk. Habit: Herb. Habitat: Woodland, bushland, thickets, and grassland, ca. 230 m. Vouchers: Robertson SA & Luke WRQ 6151, Luke
Q & SA Robertson 2523 (EA).
Cyanotis lanata Benth. Habit: Herb. Habitat: Grassland, bushland, and woodland,
ca. 50 m. Voucher: Polhill R & Paulo S 787 (EA).
Cyanotis repens Faden & D. M. Cameron subsp. repens Habit: Herb. Habitat: Dry
or moist forest, ca. 250 m. Vouchers: Robertson SA & Luke WRQ 6017, Luke Q
1455, Faden & Faden 74/1174, Adams 98 (EA).
Cyanotis sp. 1 Habit: Herb. Habitat: Forest. Voucher: Rawlins 455 (EA).
Murdannia blumei (Hassk.) Brenan Habit: Herb. Habitat: Swamps, ± 10 m. Vouchers: Greenway PJ & Rawlins 9374, Luke Q 5684 (EA).
66
Veronicah M. Ngumbau et al. / PhytoKeys 147: 1–191 (2020)
Murdannia simplex (Vahl) Brenan Habit: Herb. Habitat: Grassland, bushland, and
woodland, forest, ca. 239 m. Vouchers: SAJIT–005993 (EA, HIB), Magogo FC
& Glover PE 366 (EA).
F64. Connaraceae
4 Genera, 6 Species
Agelaea pentagyna (Lam.) Baill. Habit: Liana. Habitat: Riverine forest, coastal forest,
and woodland, 0–150 m. Voucher: Luke WRQ 902 (EA).
Connarus longistipitatus Gilg Habit: Liana. Habitat: Woodland, ca. 300 m. Vouchers: Luke WRQ & Robertson SA 2728, Graham Q 290 in FD 1695 (EA).
Ellipanthus madagascariensis (G. Schellenb.) Capuron ex Keraudren Habit: Tree.
Habitat: Forest, 0–100 m. Vouchers: SAJIT–005929 & 005513 (EA, HIB), Faden
RB & Faden AJ 77/570, Luke WRQ & Robertson SA 216 (EA): Near Threatened.
Rourea coccinea subsp. boiviniana (Baill.) Jongkind Habit: Shrub. Habitat: Coastal
forest, bushland, riverine, and woodland, 0–750 m. Vouchers: Ngumbau V &
Mwadime N V0492 (EA, HIB), Kuchar P 13484, Luke WRQ 3128 (EA).
Rourea minor (Gaertn.) Alston Habit: Liana. Habitat: Woodland, riverine, and
forest, ca. 250 m. Vouchers: Luke WRQ & Robertson SA 2722, Drummond &
Hemsley 4034A (EA).
Rourea orientalis Baill. Habit: Shrub. Habitat: Forest, woodland, and bushland,
ca. 156 m. Vouchers: SAJIT–005478, Ngumbau V & Mwadime N V0432 (EA,
HIB), Luke Q 1506, Luke WRQ & Robertson SA 514 (EA).
F65. Convolvulaceae
14 Genera, 44 Species
Astripomoea hyoscyamoides (Vatke) Verdc. var. hyoscyamoides Habit: Herb. Habitat: Grassland, bushland, roadside, and cultivated area, ca. 50–100 m. Vouchers:
Robertson SA & Luke WRQ 6197, Jeffrey 8, Fukuoka N 276 (EA).
Astripomoea malvacea (Klotzsch) A. Meeuse var. malvacea Habit: Herb. Habitat:
Woodland and grassland, 60 m. Voucher: Kassner 437 (EA).
Convolvulus jefferyi Verdc. Habit: Climber. Habitat: Grassland, forest, and sand
dunes, 0–150 m. Vouchers: Luke Q 5447, Jeffery 321, Irwin 271 (EA).
Cressa cretica L. Habit: Herb. Habitat: Coastal dunes or sandy places, ± 0 m. Vouchers: Luke Q 5474, Rawlins 182 & 391 (EA).
Evolvulus alsinoides (L.) L. Habit: Herb. Habitat: Grassland, woodland, roadside,
and cultivated area, ca. 100 m. Vouchers: Magogo FC & Glover PE 817, Battiscombe 774 (EA).
Hewittia malabarica (L.) Suresh Habit: Climber. Habitat: Grassland, bushland, forest edges, and waste places, ca. 0–200 m. Vouchers: SAJIT–006205, Ngumbau V
& Mwadime N V0286 & 0396 (EA, HIB), Magogo FC & Glover PE 73 (EA).
Ipomoea albivenia (Lindl.) Sweet Habit: Climber. Habitat: Coastal bushland, 50–
200 m. Voucher: Wakefield (EA).
Checklist of coastal forests of Kenya
67
Ipomoea aquatica Forssk. Habit: Herb. Habitat: Swampy, ca. 50 m. Voucher: Greenway & Rawlins 9450 (EA).
Ipomoea barteri A. Chev. Habit: Herb. Habitat: Grassland and woodland, 0–200 m.
Voucher: Whyte (EA).
Ipomoea bullata Oliv. Habit: Climber. Habitat: Bushland, ca. 280 m. Voucher: Luke
WRQ & Robertson SA 2212 (EA).
Ipomoea cairica (L.) Sweet Habit: Climber. Habitat: Seashores, forest clearings, and
grassland, ca. 15 m. Voucher: Simpson BL 118 (EA).
Ipomoea coptica (L.) Roth ex Roem. & Schult. var. coptica Habit: Herb. Habitat:
Grassland, woodland, roadside, and cultivated area, ca. 190 m. Vouchers: Drummond & Hemsley 4239, Rawlins 826 (EA).
Ipomoea coptica var. acuta Choisy. Habit: Climber. Habitat: Cultivated area,
0–10 m. Voucher: Festo L & Luke Q 2583 (EA).
Ipomoea eriocarpa R.Br. Habit: Climber. Habitat: Grassland and cultivated ground,
ca. 0–150 m. Vouchers: Festo L & Luke Q 2468, Magogo FC & Glover PE
1059 (EA).
Ipomoea ficifolia Lindl. Habit: Climber. Habitat: Bushland and cultivated ground,
20–390 m. Vouchers: Herbarium Techn. Course II 054, Irwin 277, Festo L &
Luke Q 2527 (EA).
Ipomoea garckeana Vatke Habit: Climber. Habitat: Bushland and cultivated ground,
30–350 m. Vouchers: Drummond & Hemsley 4083, Jeffery 544 (EA, HIB).
Ipomoea hildebrandtii Vatke subsp. orientalis Verdc. Habit: Climber. Habitat:
Grassland or bushland, ca. 304 m. Vouchers: Bally PRO 16798, Irwin 156 (EA).
Ipomoea irwiniae Verdc. Habit: Climber. Habitat: Bushland and grassland, 0–500 m.
Vouchers: Irwin PH 276, Luke WRQ & PA 5688, Jeffery 49 (EA).
Ipomoea kotschyana Hochst. ex Choisy Habit: Herb. Habitat: Grassland, ± 0 m.
Vouchers: Tweedie 1057 & 1874, Irwin 273 (EA).
Ipomoea marginata (Desr.) Verdc. Habit: Climber. Habitat: Swampy, wooded grassland, and woodland, ca. 100 m. Vouchers: Magogo FC & Glover PE 999, Festo L
& Luke Q 2360 (EA).
Ipomoea mauritiana Jacq. Habit: Liana. Habitat: Rainforest, riverine, and bushland,
ca. 65 m. Vouchers: Irwin 440, Mwadime N & Chesire 236 (EA).
Ipomoea mombassana Vatke subsp. mombassana Habit: Climber. Habitat: Bushland and woodland, ca. 0–304 m. Vouchers: Kuchar P 11819, Irwin PH 164 &
280 (EA).
Ipomoea obscura (L.) Ker-Gawl. Habit: Climber. Habitat: Bushland, grassland, and
woodland, ca. 20 m. Vouchers: Gray M & Luke WRQ 292, Festo 2647 9 (EA).
Ipomoea ochracea (Lindl.) G. Don Habit: Herb. Habitat: Forest, ca. 60–365 m.
Vouchers: Drummond & Hemsley 4079, Polhill R & Paulo S 917 (EA).
Ipomoea pes-caprae subsp. brasiliensis (L.) Ooststr. Habit: Herb. Habitat: Sandy
shores, and roadside, 0–750 m. Vouchers: SAJIT–006251 (EA, HIB), Verdcourt
1175, Kirika P, Muthoka P & Mbale M 761 (EA).
Ipomoea pes-tigridis L. Habit: Climber. Habitat: Grassland and thickets, 0–183 m.
Vouchers: Ngumbau V & Mwadime N V0193 (EA, HIB), Luke Q 5632 (EA).
68
Veronicah M. Ngumbau et al. / PhytoKeys 147: 1–191 (2020)
Ipomoea pileata Roxb. Habit: Climber. Habitat: Rainforest, bushland, and grassland,
ca. 75 m. Vouchers: Graham 1963, Jeffery 270 (EA).
Ipomoea shupangensis Baker Habit: Climber. Habitat: Forest, bushland, and coral,
ca. 20 m. Vouchers: Mwadime N 43, Pakia M et al. 986, Luke WRQ 10704 (EA).
Ipomoea ticcopa Verdc. Habit: Herb. Habitat: Grassland, 0–25 m. Vouchers: SAJIT–006245 (EA, HIB), Tweedie 982 (EA).
Ipomoea urbaniana (Dammer) Hallier f. Habit: Climber. Habitat: Forest, grassland,
ca. 10 m. Vouchers: Robertson SA 4208, Irwin 394, Luke 3078 (EA).
Ipomoea wightii var. kilimandschari (Dammer) Verdc. Habit: Climber. Habitat:
Open forest and scrub, ca. 50 m. Voucher: Graham RM 509 (EA).
Ipomoea violacea L. Habit: Herb. Habitat: Bushland or coral, ± 0 m. Vouchers: Jeffery 381, Bally 1894, Irwin 260 (EA).
Jacquemontia ovalifolia (Choisy) Hallier f. Habit: Climber. Habitat: Sandy beaches,
10–750 m. Vouchers: Kirika P, Muthoka P & Mbale M 756, Festo 2447, Hooper
& Townsend 1194 (EA).
Jacquemontia paniculata (Burm. f.) Hallier f. Habit: Climber. Habitat: Grassland,
0–360 m. Vouchers: Ngumbau V & Mwadime N V0231 (EA, HIB), Magogo FC
& Glover PE 1048 (EA).
Jacquemontia tamnifolia (L.) Griseb. Habit: Climber. Habitat: Grassland and cultivated place, ca. 272 m. Vouchers: Ngumbau V & Mwadime N V0272 (EA, HIB),
Magogo FC & Glover PE 861 (EA).
Merremia ampelophylla Hallier f. var. ampelophylla Habit: Herb. Habitat: Bushland
or roadside, ca. 20 m. Vouchers: C van Someren in EAH 10560, Festo 2645 (EA).
Merremia semisagitta (Peter) Dandy Habit: Herb. Habitat: Not known, ca. 20m.
Voucher: Gregory (EA).
Merremia sp. C of FTEA Habit: Herb. Habitat: Woodland, ca. 90 m. Voucher: Rawlins 864 (EA): Endemic.
Metaporana densiflora (Hallier f.) N. E. Br. Habit: Liana. Habitat: Forest edge,
50–250 m. Vouchers: Ngumbau V & Mwadime N V0562 (EA, HIB), Mwadime
N 24, Drummond & Hemsley 4245 (EA).
Operculina turpethum (L.) Silva Manso Habit: Herb. Habitat: Swampy, ca. 100 m.
Voucher: Rawlins 188 (EA).
Seddera hirsuta Hallier f. var. hirsuta Habit: Herb. Habitat: Bushland, 90–460 m.
Voucher: Drummond & Hemsley 4069 (EA).
Seddera suffruticosa (Schinz) Hallier f. Habit: Herb. Habitat: Grassland, 0–250 m.
Voucher: Drummond & Hemsley 3768 (EA).
Stictocardia incompta (Hallier f.) Hallier f. Habit: Climber. Habitat: Mangrove forest, bushland, 0–800 m. Voucher: Luke WRQ 1322 (EA).
Stictocardia macalusoi (Mattei) Verdc. Habit: Climber. Habitat: Sand dunes and sea
shore, ± 0 m. Voucher: Luke Q 5482 (EA).
Xenostegia tridentata (L.) D. F. Austin & Staples Habit: Climber. Habitat: Grassland, wooded and grassland, 30–213 m. Vouchers: Ngumbau V & Mwadime N
V0330 (EA, HIB), Magogo FC & Glover PE 151, Robertson SA 4225 (EA).
Checklist of coastal forests of Kenya
69
F66. Costaceae
1 Genus, 1 Species
Costus afer Ker Gawl. Habit: Herb. Habitat: Moist forests at forest edges, 160–210 m.
Vouchers: Ngumbau V & Mwadime N V0375 (EA, HIB), Magogo FC & Glover
PE 563, Drummond RB & Hemsley JH 1200 (EA).
F67. Crassulaceae
1 Genus, 7 Species
Kalanchoe ballyi Raym.-Hamet ex Cufod. Habit: Herb. Habitat: Coastal woodland,
ca. 85 m. Vouchers: Ngumbau V & Mwadime N V0414 (EA, HIB), Robertson
SA & Luke WRQ 5139 (EA): Endemic.
Kalanchoe fadeniorum Raadts Habit: Herb. Habitat: Semi-evergreen thicket on
sandy soils, ca. 360 m. Voucher: RB & AJ Faden 77/777 (EA): Extinct in the
wild, Endemic.
Kalanchoe glaucescens Britten Habit: Herb. Habitat: Bushland, grassland, and forest,
ca. 550 m. Voucher: Reitsma J 508 (EA).
Kalanchoe nyikae Engl. subsp. nyikae Habit: Herb. Habitat: Bushland, ca. 304 m.
Vouchers: Jeffery GW 248, Luke WRQ & Robertson SA 2223 (EA).
Kalanchoe lateritia Engl. var. lateritia Habit: Herb. Habitat: Woodland and lowland forest, 1–200 m. Vouchers: Robertson SA & Luke WRQ 6488, 6002 &
4925, Drummond & Hemsley 3790 (EA).
Kalanchoe obtusa Engl. Habit: Herb. Habitat: Evergreen to semi-deciduous forest,
1–450 m. Voucher: Robertson SA 5442 (EA).
Kalanchoe rotundifolia (Haw.) Haw. Habit: Herb. Habitat: Bushland, 500–700 m.
Voucher: Rauh W 860 (EA).
F68. Cucurbitaceae
18 Genera, 48 Species
Cephalopentandra ecirrhosa (Cogn.) C. Jeffrey Habit: Climber. Habitat: Deciduous
woodland and bushland, ca. 45 m. Voucher: Bally PRO 2057 (EA).
Citrullus lanatus (Thunb.) Matsum. & Nakai Habit: Climber. Habitat: Grassland
and bushland ca. 50 m. Vouchers: Jeffery GW 135, Drummond & Hemsley 44
(EA): Cultivated.
Coccinia grandiflora Cogn. ex Engl. Habit: Climber. Habitat: Lowland and upland
rainforest, 100–269 m. Vouchers: Ngumbau V & Mwadime N V003 (EA, HIB),
Kimeu JM 522, Drummond & Hemsley 3874 (EA).
Coccinia grandis (L.) Voigt Habit: Climber. Habitat: Deciduous bushland and woodland, 0–199 m. Vouchers: SAJIT–006116 & 006222, Ngumbau V & Mwadime
N V0427 (EA, HIB), Festo L, Luke Q & P 2800, Drummond & Hemsley
3901 (EA).
70
Veronicah M. Ngumbau et al. / PhytoKeys 147: 1–191 (2020)
Coccinia microphylla Gilg Habit: Climber. Habitat: Lowland rainforest, ca. 180 m.
Vouchers: Milne-Redhead E & Taylor P 7104, Drummond & Hemsley 4087 &
4040 (EA).
Coccinia pwaniensis Holstein Habit: Climber. Habitat: Lowland rainforest, 80–
300 m. Vouchers: Drummond & Hemsley 1078 & 3953, Saunders, FC Magogo
& P Glover 493 (EA).
Coccinia trilobata (Cogn.) C. Jeffrey Habit: Climber. Habitat: Wooded grassland
and deciduous bushland, ca. 30 m. Voucher: Simpson BL 383 (EA).
Corallocarpus ellipticus Chiov. Habit: Climber. Habitat: Bushland on coral sand
near sea level. Vouchers: Robertson SA 3809, Bogdan 4723, Lucas, Jeffrey & Kirrika 266 (EA).
Corallocarpus epigaeus (Rottler) C. B. Clarke Habit: Climber. Habitat: Wooded
grassland and deciduous bushland, ca. 550 m. Voucher: Williams JG 12588 (EA).
Ctenolepis cerasiformis (Stocks) Hook. f. Habit: Climber. Habitat: Deciduous bushland and thicket, ca. 5 m. Vouchers: Kimeu JM 657, Rawlins 843, Mwadime N
438 (EA).
Cucumis aculeatus Cogn. Habit: Climber. Habitat: Forest edge, ca. 142 m. Voucher:
SAJIT–005999 (EA, HIB).
Cucumis dipsaceus Ehrenb. ex Spach Habit: Climber. Habitat: Bushland and woodland, ca. 425 m. Voucher: Fukuoka N 250 (EA).
Cucumis engleri (Gilg) Ghebret. & Thulin Habit: Climber. Habitat: Bushland, wooded
grassland, and grassland, ca. 330 m. Voucher: Drummond & Hemsley 4220 (EA).
Cucumis ficifolius A. Rich. Habit: Climber. Habitat: Deciduous bushland and grassland, 0–300 m. Voucher: Jeffery GW 696 (EA).
Cucumis maderaspatanus L. Habit: Climber. Habitat: Seasonal swamps, flood plain,
and valley grassland, ca. 30 m. Vouchers: Cunningham-van Someren GR 280,
Nyange M 406 (EA).
Cucumis prophetarum subsp. dissectus (Naudin) C. Jeffrey Habit: Climber. Habitat: Grassland, ca. 60 m. Vouchers: Luke WRQ & PA 4574B, Drummond &
Hemsley 3994 & 4206 (EA).
Cucumis sacleuxii Paill. & Bois Habit: Climber. Habitat: Rain and swamp or other
ground water forest, ca. 210 m. Vouchers: Ngumbau V & Mwadime N V0371
(EA, HIB), Rawlins SP 56, Luke WRQ & PA 5687 (EA).
Cucumis sp. (Oreosyce sp. A of FTEA) Habit: Climber. Habitat: Bushland. Vouchers:
Rawlins, Gillespie 242 (EA): Endemic.
Cyclantheropsis parviflora (Cogn.) Harms Habit: Climber. Habitat: Lowland evergreen forest, deciduous woodland, and bushland, ca. 70 m. Voucher: Ngumbau V
& Mwadime N V0443 (EA, HIB).
Diplocyclos palmatus (L.) C. Jeffrey Habit: Climber. Habitat: Swamps, ground water
forest, and seasonal swamp grasslands, ca. 0– 50 m. Vouchers: Sangai GW 15723,
Polhill & Paulo 720 (EA).
Diplocyclos tenuis (Klotzsch) C. Jeffrey Habit: Climber. Habitat: Coastal bushland,
0–90 m. Voucher: Robertson SA 3423 (EA).
Checklist of coastal forests of Kenya
71
Eureiandra orientalis Q. Luke, ined. Habit: Climber. Habitat: Coastal forest and
bushland, 0–100 m. Vouchers: Ngumbau V & Mwadime N V0409 & V0332
(EA, HIB), Gray M & Luke WRQ 282 (EA).
Gerrardanthus grandiflorus Gilg ex Cogn. Habit: Climber. Habitat: Lowland rainforest, ca. 80 m. Vouchers: Ngumbau V & Mwadime N V072 (EA, HIB), Drummond & Hemsley 3958 (EA).
Gerrardanthus lobatus (Cogn.) C. Jeffrey Habit: Climber. Habitat: Deciduous
bushland, woodland, and wooded grassland, 0–400 m. Vouchers: Drummond &
Hemsley 41158, Robertson SA 6398 (EA).
Kedrostis abdallae Zimm. Habit: Climber. Habitat: Forest, ca. 35 m. Voucher: Luke
Q 1529 (EA).
Kedrostis foetidissima (Jacq.) Cogn. Habit: Climber. Habitat: Deciduous and semievergreen woodland, ca. 400 m. Vouchers: Faden RB & AJ 74/1196, Robertson
SA 3491 (EA).
Kedrostis heterophylla Zimm. Habit: Climber. Habitat: Lowland rainforest, 40–
300 m. Vouchers: SAJIT–006138 (EA, HIB), Luke WRQ 1833, Luke WRQ &
Robertson SA 2813 (EA).
Kedrostis leloja (Forssk. ex J.F Gmel.) C. Jeffrey Habit: Climber. Habitat: Deciduous
bushland and wooded grassland, 10–550 m. Vouchers: SAJIT–006129, Ngumbau
V & Mwadime N V0245 (EA, HIB), Luke WRQ & Luke Q 2458 (EA).
Lagenaria siceraria (Molina) Standl. Habit: Climber. Habitat: Cultivated area.
Voucher: Waaijenberg 29 (EA).
Lagenaria sphaerica (Sond.) Naudin Habit: Climber. Habitat: Evergreen and deciduous forests, 0–50 m. Vouchers: Katz SS 197/75, Drummond & Hemsley 3903,
Nyange M 473 (EA).
Luffa cylindrica M. Roem. Habit: Climber. Habitat: Hyphaene woodland, 5–15 m.
Vouchers: Festo L & Luke Q 2476, Leauthaud C 120 (EA): Naturalized.
Momordica anigosantha Hook. f. Habit: Climber. Habitat: Lowland, groundwater
forest and forest remnants, ca. 300 m. Vouchers: Ngumbau V & Mwadime N
V0221 (EA, HIB), Faden RB 70/822 (EA).
Momordica boivinii Baill. Habit: Climber. Habitat: Deciduous bushland, thicket,
woodland, and wooded grassland, 0–200 m. Vouchers: SAJIT–005951, Ngumbau
V & Mwadime N V0345 (EA, HIB), Luke & Robertson 2291, Magogo FC &
Glover PE 215 (EA).
Momordica calantha Gilg Habit: Climber. Habitat: Forests edge and valley grasslands, ca. 100 m. Vouchers: Ngumbau V & Mwadime N V119 (EA, HIB), Kabuye CHS, Gilbert VC & Robertson A 115, Magogo FC & Glover PE 306 (EA).
Momordica charantia L. Habit: Climber. Habitat: Lowland rainforest and riverine
forest, 100 m. Voucher: Verdcourt 3942 (EA).
Momordica leiocarpa Gilg Habit: Climber. Habitat: Lowland rainforest, 60–300 m.
Vouchers: Drummond & Hemsley 1196, Verdcourt 1875, Luke WRQ 2890 (EA).
Momordica littorea Thulin Habit: Climber. Habitat: Bushland, 10–150 m. Voucher:
Luke Q 5440 (EA).
72
Veronicah M. Ngumbau et al. / PhytoKeys 147: 1–191 (2020)
Momordica peteri Zimm. Habit: Climber. Habitat: Lowland rainforest, ca. 280 m.
Vouchers: Reitsma J 344, Luke WRQ 1638 (EA).
Momordica rostrata Zimm. Habit: Climber. Habitat: Deciduous bushland, thicket,
woodland, and wooded grassland, 15–350 m. Vouchers: Ngumbau V & Mwadime
N V0447 (EA, HIB), Kuchar P 12907, Drummond & Hemsley 4110 (EA).
Momordica spinosa (Gilg) Chiov. Habit: Climber. Habitat: Deciduous bushland and
semi-desert scrub, ca. 330 m. Voucher: Bally (EA).
Momordica trifoliolata Hook. f. Habit: Climber. Habitat: Deciduous bushland and
woodland, ca. 0–86 m. Vouchers: Ngumbau V & Mwadime N V0478 (EA, HIB),
Stuhlmann 6970 (EA).
Peponium vogelii (Hook. f.) Engl. Habit: Climber. Habitat: Lowland and upland
rainforest, 80–207 m. Vouchers: Ngumbau V & Mwadime N V065 (EA, HIB),
Verdcourt B 2121, Magogo FC & Estes R 1211 (EA).
Pilogyne scabra (L. f.) W. J. de Wilde & Duyfjes Habit: Climber. Habitat: Damp
places, 80–272 m. Vouchers: Ngumbau V & Mwadime N V106 (EA, HIB), Brenan JPM & Brenan JH 14642 (EA).
Trochomeria macrocarpa (Sond.) Harv. Habit: Climber. Habitat: Deciduous woodland on sandy soils, ca. 120 m. Voucher: SAJIT–005489 (EA, HIB).
Zehneria monocarpa G.W. Hu, V.M. Ngumbau & Q.F. Wang, sp. nov Habit:
Climber. Habitat: Bushland, ca. 199 m. Vouchers: SAJIT 007172 & 007173,
Ngumbau V & Mwadime N V0399 (EA, HIB), Mwadime 2404 (EA): Endemic.
Zehneria pallidinervia (Harms) C. Jeffrey Habit: Climber. Habitat: Deciduous
bushland, ca. 0–88 m. Vouchers: SAJIT–006443 & 006241 (EA, HIB), Beentje
HJ 2345 (EA).
Zehneria peneyana (Naudin) Schweinf. & Asch. Habit: Climber. Habitat: Swampy
and marshy areas, 20–425 m. Vouchers: Ngumbau V & Mwadime N V0555 (EA,
HIB), Festo L & Luke Q 2453 (EA).
Zehneria thwaitesii (Schweinf.) C. Jeffrey Habit: Climber. Habitat: Lowland grassland, margins of rivers, 30–223 m. Vouchers: Ngumbau V & Mwadime N V0540
& 0124 (EA, HIB), Robertson SA 5854 (EA).
F69. Cymodoceaceae
2 Genera, 2 Species
Thalassodendron ciliatum (Forssk.) Hartog Habit: Herb. Habitat: Shallow waters,
0–33 m. Voucher: SAJIT–006253 (EA, HIB).
Cymodocea rotundata Ehrenb. & Hempr. ex Asch. Habit: Herb. Habitat: Clear water, 0–10 m. Vouchers: SAJIT–006255 & 006266 (EA, HIB).
F70. Cyperaceae
18 Genera, 111 Species
Alinula paradoxa (Cherm.) Goetgh. & Vorster Habit: Herb. Habitat: In and near
rice fields, 0–30 m. Voucher: Coppejans 5690 (EA).
Checklist of coastal forests of Kenya
73
Bolboschoenus maritimus (L.) Palla Habit: Herb. Habitat: In and near swamps,
0–30 m. Vouchers: SAJIT–005912 (EA, HIB), Luke Q 5630 (EA).
Bulbostylis abortiva (Steud.) C.B. Clarke Habit: Herb. Habitat: Shallow soil over
rock, grassland, ca. 50 m. Voucher: Kaesnner 346 (EA).
Bulbostylis afroorientalis (Lye) R.W. Haines Habit: Herb. Habitat: Grassland with
scattered trees, 80–300 m. Vouchers: Luke & Luke 4387, Lye & Katende 6288,
Faden & Evans 71/631 (EA).
Bulbostylis argenteobrunnea C.B. Clarke Habit: Herb. Habitat: Mixed grasslands
and cleared bushland, ca. 150 m. Vouchers: Bogdan 5339, Magogo FC & Esters
R 1177 (EA).
Bulbostylis barbata (Rottb.) C.B. Clarke Habit: Herb. Habitat: Mixed woodland
and bushland, ca. 0–120 m. Vouchers: Polhill & Paulo 847, Kirika P, Muthoka P
& Mbale M 753 (EA).
Bulbostylis boeckeleriana (Schweinf.) Beetle var. boeckeleriana Habit: Herb. Habitat: Dry grassland, ca. 0–500 m. Vouchers: Luke WRQ & Kabuye CHS 5120,
Luke WRQ & Robertson SA 5120 (EA).
Bulbostylis boeckeleriana var. transiens (K. Schum.) R.W. Haines Habit: Herb.
Habitat: Grassland and open bushland, 100–400 m. Vouchers: Moomaw 931,
Robertson 5120 (EA).
Bulbostylis burchellii (Ficalho & Hiern) C.B. Clarke Habit: Herb. Habitat: Open
coastal forests and marginal mangrove vegetation, 10–75 m. Vouchers: Rexton
Karisa s.n., Karisa L 52 (EA).
Bulbostylis contexta (Nees) Bodard Habit: Herb. Habitat: Rocky area, ca. 0–400 m.
Vouchers: Gilbert MG & Kuchar P 5899, Magogo FC & Glover PE 356, Drummond & Hemsley1116 (EA).
Bulbostylis densecaespitosa (Lye) R. W. Haines Habit: Herb. Habitat: Coastal forest,
50–300 m. Vouchers: Moormaid JC 931, Luke & Robertson 2199, Robertson et
al. 7010 (EA): Endemic.
Bulbostylis filamentosa (Vahl) C. B. Clarke Habit: Herb. Habitat: Seasonally wet
habitats, grassland, and crevices in rock faces, 0–326 m. Vouchers: Magogo FC &
Glover PE 23, Lye & Katende 6283, Magogo & Glover 231 (EA).
Bulbostylis hispidula (Vahl) R. W. Haines subsp. hispidula Habit: Herb. Habitat:
Coastal wooded grassland, 0–100 m. Voucher: Mwachala G et al. 856 (EA).
Bulbostylis hispidula subsp. filiformis (C. B. Clarke) R. W. Haines Habit: Herb. Habitat: Grassland, bushland on sandy dunes, dry banks, roadsides, and wet rock crevices, ca. 0–50 m. Vouchers: Rawlins s.n., Robertson SA 3222, Kuchar P 7205 (EA).
Bulbostylis hispidula subsp. halophila (Lye) R. W. Haines Habit: Herb. Habitat:
Short grassland, sandy beaches, and abandoned cultivation, 0–15 m. Vouchers:
Moomaw JC 816, Cunningham-van Someren GR Sh107 (EA).
Bulbostylis hispidula subsp. intermedia (Lye) R. W. Haines Habit: Herb. Habitat: Dry grassland and shallow soil on rocks, near sea level. Voucher: Lye 6280
(EA): Endemic.
Bulbostylis microelegans (Lye) R. W. Haines Habit: Herb. Habitat: Grassland and
scrub, ca. 10 m. Voucher: Festo L & Luke WRQ 2729 (EA).
74
Veronicah M. Ngumbau et al. / PhytoKeys 147: 1–191 (2020)
Bulbostylis pilosa (Willd.) Cherm. Habit: Herb. Habitat: Seasonally flooded wooded
grassland, mangrove swamps, and bushland, 1–400 m. Vouchers: Ngumbau V &
Mwadime N V0150 (EA, HIB), Drummond RB & Hemsley JH 1174 (EA).
Courtoisina assimilis (Steud.) Maquet Habit: Herb. Habitat: Streamside, ditches,
and seasonal pools, ca. 500 m. Voucher: Faden RB & Faden AJ 116 (EA).
Cyperus afrodunensis Lye Habit: Herb. Habitat: On sand dunes and sandy soil near
seashore. Voucher: Bogdan A 2537 (EA).
Cyperus alopecuroides Rottb. Habit: Herb. Habitat: In swamps and seasonally wet
grasslands, ca. 20 m. Voucher: Rawlins SP 633 (EA).
Cyperus amabilis Vahl Habit: Herb. Habitat: In grassland and wooded grassland,
ca. 0–15 m. Vouchers: Rawlins SP 88, Magogo FC & Glover PE 1024, RB & AJ
Faden 71 (EA).
Cyperus articulatus L. Habit: Herb. Habitat: In swamps, lakeshores, wet grasslands, and
pools, ca. 0–190 m. Vouchers: Sangai GW 15839, Luke PA & WRQ 4087 (EA).
Cyperus blysmoides Hochst. ex C. B. Clarke Habit: Herb. Habitat: Seasonally wet
habitats, flooded grasslands, and, swampy areas, ca. 30 m. Voucher: Kabuye et al.
TPR 449 (EA).
Cyperus boreobellus Lye Habit: Herb. Habitat: On damp shallow sandy soil and
rocky pools, 350–400 m. Vouchers: Drummond & Hemsley 4153, Luke 5582A
(EA): Endemic.
Cyperus chordorrhizus Chiov. Habit: Herb. Habitat: On sand dunes, sea level.
Vouchers: Luke Q 5646, Frazier J 896 (EA).
Cyperus colymbetes Kotschy & Peyr. Habit: Herb. Habitat: Muddy areas, on
swampy ground, 10–400 m. Vouchers: Kabuye et al. TPR 382, Luke PA & WRQ
4388 (EA).
Cyperus compressus L. Habit: Herb. Habitat: In roadside ditches, drainage trenches,
permanent, and seasonal pools, ca. 0–300 m. Vouchers: Gilbert MG & Kuchar
5898, Magogo FC & Glover PE 1064 (EA).
Cyperus crassipes Vahl Habit: Herb. Habitat: On shores and sand dunes, 0–60 m.
Vouchers: SAJIT–006263 (EA, HIB), Greenway PJ & Rawlins SP 9293, Greenway PG 11336 (EA).
Cyperus cuspidatus Kunth Habit: Herb. Habitat: Grassland, ca. 0–30 m. Vouchers:
Langridge WP 139, Greenway PJ 10840 (EA).
Cyperus denudatus L. f. Habit: Herb. Habitat: Riversides, floodplains, swamps, damp
grassland, and moist rock crevices, ca. 0–150 m. Voucher: Magogo FC & Glover
PE 339 (EA).
Cyperus difformis L. Habit: Herb. Habitat: In swamps, alongside water edge,
0–198 m. Voucher: Magogo FC & Glover PE 556 (EA).
Cyperus diurensis Boeckeler Habit: Herb. Habitat: In grassland, woodland, and
rocky outcrops, 15–200 m. Vouchers: Greenway PJ & Rawlins SP 8956, Adams
64, L Festo & WRQ Luke 2492 (EA).
Cyperus dubius Rottb. subsp. dubius Habit: Herb. Habitat: Near the sea on sandy
beaches, 0–300 m. Vouchers: Robertson SA 7299, Robertson SA et al. 270 (EA).
Checklist of coastal forests of Kenya
75
Cyperus dubius var. macrocephalus (C.B. Clarke) Kük. Habit: Herb. Habitat: Riverine, dry bushland, and grassland, 50–380 m. Vouchers: Kabuye CHS, Gilbert
VC & Robertson A 84/84A, Magogo FC & Glover PE 307A, Luke 4004 (EA).
Cyperus exaltatus Retz. Habit: Herb. Habitat: Grasslands and pond margins, ca.
0–30 m. Vouchers: Kuchar P 13556, Luke WRQ et al. 5718, Luke WRQ 3464 (EA).
Cyperus foliaceus C.B. Clarke Habit: Herb. Habitat: Woodlands, seasonally wet
habitats, swamps, along streams, and pools, 0–198 m. Vouchers: Magogo FC &
Glover PE 103 & 574 (EA).
Cyperus frerei C.B. Clarke Habit: Herb. Habitat: Sand dunes, sandy mangrove
shores, 0–10 m. Voucher: Mwadime N & Luke WRQ 2612 (EA).
Cyperus grandis C.B. Clarke Habit: Herb. Habitat: Swamps, in stagnant or moving
water, 0–400 m. Vouchers: Harper GB 113/55, Drummond RB & Hemsley JH
1092 (EA).
Cyperus haspan L. Habit: Herb. Habitat: Swampy or marshy sites, seasonally wet
grassland, 0–426 m. Vouchers: Mwachala G et al. 857, Gilbert MG & Kuchar P.
5894, Sangai GW 15794 (EA).
Cyperus hemisphaericus Boeckeler Habit: Herb. Habitat: Open grassland, Brachystegia woodland, and wooded grassland, 0–140 m. Vouchers: Mamur JC 981, Luke
WRQ 3199 (EA).
Cyperus holstii Kük. Habit: Herb. Habitat: Seasonally wet grassland, alongside pools, and
swamps, 0–250 m. Vouchers: Verdcourt 3959, Luke & Grey 4027, Luke 4605 (EA).
Cyperus involucratus Rottb. Habit: Herb. Habitat: Swampy grassland and lake
shores, 30 m. Vouchers: Brathany 1982 Expedition 49 (EA).
Cyperus kaessneri C.B. Clarke Habit: Herb. Habitat: Seasonally damp habitats and
bushland, 0–400 m. Vouchers: Drummond & Hemsley 3745, Adams BR 7 (EA).
Cyperus kwaleensis Lye Habit: Herb. Habitat: On shallow sandy soils over outcropping rocks, ca. 360 m. Voucher: Drummond & Hemsley 4204 (EA): Endemic.
Cyperus laevigatus L. Habit: Herb. Habitat: Damp sandy places, ca. 45 m. Vouchers:
Festo L & Luke WRQ 2439, Makin J 417 (EA).
Cyperus latifolius Poir. Habit: Herb. Habitat: In swamps, marshes, boggy grasslands,
and roadside, 0–195 m. Voucher: Magogo FC & Glover PE 576 (EA).
Cyperus laxus subsp. buchholzii (Boeckeler) Lye Habit: Herb. Habitat: Forest, secondary vegetation, on stream banks, 100 m. Vouchers: Greenway & Rawlins 9364,
Robertson & Luke 5951 (EA).
Cyperus ligularis L. Habit: Herb. Habitat: Swamps, marshes, and creek margins.
Voucher: Vera de Measter 209 (EA).
Cyperus longus var. pallidus Boeckeler Habit: Herb. Habitat: Lake edges, swamps,
and temporary pools, ca. 65 m. Voucher: Nattrass MS 1521 (EA).
Cyperus luteus Boeckeler Habit: Herb. Habitat: Wet grassland and secondary forest,
0–228 m. Vouchers: Ngumbau V & Mwadime N V048 (EA, HIB), Musyoki &
Hansen 996 (EA).
Cyperus meeboldii Kük. Habit: Herb. Habitat: Wet habitats or in mud, 30–353 m.
Vouchers: Luke & Gray 4050, Mwachala G et al 808 (EA).
76
Veronicah M. Ngumbau et al. / PhytoKeys 147: 1–191 (2020)
Cyperus microumbellatus Lye Habit: Herb. Habitat: In swampy area, ca. 380 m.
Voucher: Magogo FC & Glover PE 323 (EA): Endemic.
Cyperus michelianus subsp. pygmaeus (Rottb.) Asch. & Graebn. Habit: Herb.
Habitat: Seasonally wet habitats, damp sandy, ca. 30 m. Voucher: Faden & Faden
72/139 (EA).
Cyperus mollipes (C. B. Clarke) K. Schum. Habit: Herb. Habitat: Grassland, ca.
0–20 m. Vouchers: Napper D 2179, Magogo FC & Glover PE 316, Waayenberg
H 17, Kabuye et al. TPR 249, Luke WRQ & Robertson SA 2195 (EA).
Cyperus niveus var. leucocephalus (Kunth) Fosberg Habit: Herb. Habitat: Dry grassland, 0–200 m. Vouchers: Mwadime N 2652, Ross K 164, Drummond & Hemsley 4171 (EA).
Cyperus oblongoincrassatus Kük. Habit: Herb. Habitat: Rocky sites, dry AcaciaCommiphora bushland, ca. 350 m. Voucher: Drummond & Hemsley 4097 (EA).
Cyperus pectinatus Vahl Habit: Herb. Habitat: Swamps, lake margins, and shallow
water, ca. 15 m. Vouchers: SAJIT–006223 (EA, HIB), Mwadime N 2662 (EA).
Cyperus prolifer Lam. Habit: Herb. Habitat: Swamp edges, streamside, and seasonally flooded grasslands, 0–450 m. Vouchers: Ngumbau V & Mwadime N V0255
(EA, HIB), Magogo FC & Glover PE 119 (EA).
Cyperus renschii Boeckeler var. renschii Habit: Herb. Habitat: Forests, forest swamps,
along forest streams, or roadside, (0–) 150 m. Vouchers: Ngumbau V & Mwadime
N V110 (EA, HIB) Magogo FC & Glover PE 135, Moomaw 1063 (EA).
Cyperus rotundus L. subsp. rotundus Habit: Herb. Habitat: Roadsides, sandy fields,
and cultivated ground, ca. 260 m. Voucher: Waaijenberg H 6 (EA).
Cyperus rotundus subsp. tuberosus (Rottb.) Kük. Habit: Herb. Habitat: Roadsides,
sandy fields, and cultivated ground, 140–300 m. Vouchers: Robertson SA et al.
342, Faden RB & Evans A 627 (EA).
Cyperus rubicundus Vahl Habit: Herb. Habitat: Seasonally wet habitats, 0–353 m.
Vouchers: Bryan R Adames 54, Mwachala G et al. 814 (EA).
Cyperus squarrosus L. Habit: Herb. Habitat: Grassland and roadside, ca. 200 m.
Vouchers: Festo L & Luke WRQ 2474, Luke et al. 6325 (EA).
Cyperus tenax Boeckeler Habit: Herb. Habitat: Seasonally flooded grassland, swampy
grassland, mangrove edge, and woodland, 0–297 m. Vouchers: Ngumbau V &
Mwadime N V0151 (EA, HIB), Luke PA & WRQ 5676B (EA).
Cyperus zanzibarensis C. B. Clarke Habit: Herb. Habitat: No data. Voucher: Taylor
s.n. (EA).
Cyperus zollingeri Steud. Habit: Herb. Habitat: In seasonally wet habitats, 0–650 m.
Vouchers: Ngumbau V & Mwadime N V0537 (EA, HIB), Gillespie 155, Faden et
al. 71/645, Luke et al. 5912 (EA).
Diplacrum africanum C. B. Clarke Habit: Herb. Habitat: On bare sand or mud in
marshy grassland, ca. 60–400 m. Voucher: Luke & Luke 5963 (EA).
Eleocharis atropurpurea (Retz.) J. Presl & C. Presl Habit: Herb. Habitat: Seasonal
pools, seepage areas, and rice paddies ca. 5 m. Voucher: Mwadime N 2568 (EA).
Eleocharis dulcis (Burm.f.) Trin. ex Hensch. Habit: Herb. Habitat: Swamps and shallow parts of lakes, 0–100 m. Voucher: Kirika P, Mbale M & Mbatha M 778 (EA).
Checklist of coastal forests of Kenya
77
Eleocharis geniculata (L.) Roem. & Schult. Habit: Herb. Habitat: Lake shores and
swamps, 0–150 m. Voucher: Luke PA & WRQ 5640 (EA).
Fimbristylis bisumbellata (Forssk.) Bubani Habit: Herb. Habitat: Mud flats and
sandy riverbanks, 0–150 m. Voucher: Sangai GW 15848 (EA).
Fimbristylis bivalvis (Lam.) Lye Habit: Herb. Habitat: Streams, roadsides, and bushland, 1–100 m. Voucher: Taylor s.n. (EA).
Fimbristylis cymosa R.Br. Habit: Herb. Habitat: Sandy foreshores, edges of mangrove swamps, coral rock, and saline marshes, 0–45 m. Vouchers: Robertson SA &
Abuodha J 6916, Polhill & Paulo 897 (EA).
Fimbristylis dichotoma (L.) Vahl Habit: Herb. Habitat: Margins of rice fields and
swamps, ca. 0–150 m. Vouchers: Luke WRQ et al. 6186, Drummond & Hemsley
1164 (EA).
Fimbristylis ferruginea (L.) Vahl Habit: Herb. Habitat: Sandy soils by the coast,
0–50 m. Vouchers: Kuchar P 10052, Smith, Beentje & Muasya 244 (EA).
Fimbristylis littoralis Gaudich. Habit: Herb. Habitat: Dry river beds and drying
swamp, ca. 0–300 m. Vouchers: Luke WRQ et al. 5712, Polhill & Paulo 897,
Makin 418 (EA).
Fimbristylis pilosa Vahl Habit: Herb. Habitat: Seasonally flooded grassland and
bushed grasslands, ca. 30 m. Vouchers: Luke et al. 6317, Gillespie 277 & 280 (EA).
Fimbristylis polytrichoides (Retz.) Vahl Habit: Herb. Habitat: Seasonally inundated
mangrove swamp, 0–15 m. Vouchers: Rawlins SP 12080, Bogdan 4708, Polhill &
Paulo 896 (EA).
Fimbristylis quinquangularis subsp. macroglumis (Lye) Verdc. Habit: Herb. Habitat: Marshy grassland, 65 m. Voucher: Luke WRQ 17419 (EA).
Fimbristylis ovata (Burm.f.) J. Kern Habit: Herb. Habitat: Plain grassland, seasonally wooded grassland, and grazed fallow areas, 0–381 m. Vouchers: Magogo &
Glover 384, Adams BR 13 (EA).
Fimbristylis triflora (L.) K. Schum. ex Engl. Habit: Herb. Habitat: Tidal mud,
wooded grassland, and mangrove swamps, 0–45 m. Vouchers: Kirika P & Muthoka P 734, Luke & Mbinda 5880, Polhill & Paulo 900 (EA).
Fuirena ciliaris (L.) Roxb. Habit: Herb. Habitat: Coastal streams, pools, seasonal
swamp edges, wet grassland, and weed in rice fields, 0–300 (–500) m. Voucher:
Gilbert MG & Kuchar P 5897 (EA).
Fuirena claviseta Peter Habit: Herb. Habitat: Swamp, river, stream banks, and drainage ditches, 0–300 (–500) m. Voucher: Luke WRQ & PA 4385 (EA).
Fuirena ochreata Nees ex Kunth Habit: Herb. Habitat: Seasonally wet grassland,
edge of permanent swamp and stream, 0–450 m. Vouchers: SAJIT–005487 (EA,
HIB), Magogo FC & Glover PE 163 (EA).
Fuirena umbellata Rottb. Habit: Herb. Habitat: Seasonally wet grassland, swamp
forest, stream, and lake banks, ca. 0–213 m. Vouchers: Luke WRQ & PA 6027,
Magogo & Glover 736 (EA).
Kyllinga cartilaginea K. Schum. Habit: Herb. Habitat: Lowland forest, coconut grooves,
beach crest, dunes, and Brachystegia woodland, 0–50 (–200) m. Vouchers: Ngumbau
V & Mwadime N V097 (EA, HIB), Townsend CC & Hooper SS 1212 (EA).
78
Veronicah M. Ngumbau et al. / PhytoKeys 147: 1–191 (2020)
Kyllinga crassipes Boeckeler Habit: Herb. Habitat: Seasonally wet grassland, old cultivations, and woodlands, 0–200 m. Voucher: Bogdan 4703 (EA).
Kyllinga erecta Schumach. Habit: Herb. Habitat: Wet depressions, swamps, lake,
pool, and dam fringes, 0–353 m. Vouchers: Mwachala G et al. 809, Luke WRQ et
al. 5716, Gilbert & Kuchar 5896 (EA).
Kyllinga microstyla C.B. Clarke Habit: Herb. Habitat: Bushland or scattered tree
grassland, ca. 300 m. Voucher: Faden & Evans 71/628 (EA).
Kyllinga nervosa var. flava (C.B. Clarke) Lye Habit: Herb. Habitat: Bushland, 50–
250 m. Voucher: Wamukoya 80 (EA).
Kyllinga polyphylla Willd. ex Kunth var. polyphylla Habit: Herb. Habitat: Moist
grassland, ca. 426 m. Vouchers: Kuchar P 12875, Magogo FC & Glover PE
108 (EA).
Kyllinga polyphylla var. elatior (Kunth) Kük. Habit: Herb. Habitat: Grassland,
60 m. Voucher: Adams BR 19 (EA).
Mariscus longibracteatus Cherm. Habit: Herb. Habitat: Streamside, permanently or
seasonally swampy, and moist sites in shade, ca. 10 m. Vouchers: Luke WRQ et al.
5628, Mwachala G et al. 852, Festo 2424 (EA).
Mariscus maderaspatanus (Willd.) Napper Habit: Herb. Habitat: Grassland or
roadsides, 0–200 m. Vouchers: Luke & Mbinda 5973A, Jeffrey 14, Luke, Mbinda
& Madudu 6375 (EA).
Mariscus psilostachys C.B. Clarke Habit: Herb. Habitat: Dry grassland and rocky
outcrops, 0–100 m. Voucher: Luke WRQ & PA 4505 (EA).
Mariscus sumatrensis (Retz.) J. Raynal Habit: Herb. Habitat: Forest clearings, path
sides, grassland, and woodland, 0–381 m. Vouchers: Magogo FC & Glover PE
351, Magogo FC & Estes R 1218 (EA).
Oxycaryum cubense (Poepp. & Kunth) Palla Habit: Herb. Habitat: Floating in lakes,
swamps, and pools, ca. 0–300 m. Voucher: Kabuye et al. TPRs 531 (EA).
Pycreus hildebrandtii C.B. Clarke Habit: Herb. Habitat: Floating in lakes, swamps,
and pools, 0–800 m. Voucher: Festo L & Luke WRQ 2608 (EA).
Pycreus macrostachyos (Lam.) J. Raynal Habit: Herb. Habitat: Lakes, ponds, and at
river edges. Vouchers: Luke Q 5631, Luke WRQ & PA 5728 (EA).
Pycreus pelophilus (Ridl.) C.B. Clarke Habit: Herb. Habitat: Near water, ca. 15 m.
Voucher: Festo L & Luke WRQ 2606 (EA).
Pycreus polystachyos (Rottb.) P. Beauv. subsp. polystachyos Habit: Herb. Habitat:
Margins of rivers, riverbanks, lake margins, and sandy beaches, 0–200 m. Vouchers: Luke WRQ et al. 5719, Drummond RB & Hemsley JH 1163 (EA).
Pycreus polystachyos var. laxiflorus (Benth.) C.B. Clarke Habit: Herb. Habitat:
Margins of rivers, and sandy beaches, ca. 55 m. Vouchers: Luke WRQ 17417,
Mwadime N & Chesire C 168 (EA).
Pycreus pumilus (L.) Nees Habit: Herb. Habitat: Along swampy and streamside, ca.
50 m. Vouchers: Kirika P & Muthoka P 736, Luke et al. 6324, Gillet & Kuchar
5900 (EA).
Queenslandiella hyalina (Vahl) Ballard Habit: Herb. Habitat: Grassland and bushland on coral rag, 0–30 m. Vouchers: Schlieben 12140, Gillespie 14 (EA).
Checklist of coastal forests of Kenya
79
Remirea maritima Aubl. Habit: Herb. Habitat: Sandy seashores, just above the high
tide level. Voucher: Robertson 3675 (EA).
Rhynchospora perrieri Cherm. Habit: Herb. Habitat: Swamp areas, roadside ditches,
alongside streams, and in damp places, ca. 300 m. Voucher: Bogdan A 5347 (EA).
Schoenoplectiella articulata (L.) Lye Habit: Herb. Habitat: Grassland, wooded grassland, and Acacia bushland zone, 0–300 m. Vouchers: Verdcourt 3958, Kuchar P
12876, Kirika P, Muthoka P & Mbale M 743 (EA).
Schoenoplectiella juncea (Willd.) Lye Habit: Herb. Habitat: Seasonal pools, swamps,
along drainage lines, and stagnant water, 0–600 m. Voucher: Gillespie 282 (EA).
Schoenoplectiella lateriflora (J.F. Gmel.) Lye Habit: Herb. Habitat: Swamps, pond,
lake shores, and seasonal pools, ca. 0–300 m. Vouchers: Gillespie 283, Sangai in
EA 15793 (EA).
Schoenoplectus scirpoides (Schrad.) Browning Habit: Herb. Habitat: Lakes, riverine
fringes, and coastal saltmarsh, 0–200 m. Voucher: Luke 5642 (EA).
Scleria achtenii De Wild. Habit: Herb. Habitat: Occasionally in permanent water,
0–60 m. Voucher: Luke & Luke 5994 (EA).
Scleria bambariensis Cherm. Habit: Herb. Habitat: Swamps and seasonally wet
grassland, ca. 60 m. Vouchers: Bogdan A 3344, Luke & Luke 5993 (EA).
Scleria boivinii Steud. Habit: Herb. Habitat: Swampy forest and lowland forest, ca.
60 m. Voucher: Luke et al. 5920 (EA).
Scleria foliosa Hochst. ex A. Rich. Habit: Herb. Habitat: Swamp edges, seasonally
damp areas, and rice fields, ca. 30 m. Voucher: Luke & Gray 4058 (EA).
Scleria hildebrandtii Boeckeler Habit: Herb. Habitat: Grassland and lowland forest,
30–215 m. Vouchers: Luke & Luke 5964, Hildebrandt 2044 (EA).
Scleria lagoensis Boeckeler Habit: Herb. Habitat: Grassland, Terminalia-Combretum
and Brachystegia-Uapaca woodland, and forest edge grassland, ca. 750 m. Voucher:
Luke & Luke 5962 (EA).
Scleria lithosperma (L.) Sw. Habit: Herb. Habitat: Evergreen forest, forest, and plantation edges, 20–200 m. Vouchers: Ngumbau V & Mwadime N V087 (EA, HIB),
Magogo FC & Glover PE 98 (EA).
Scleria parvula Steud. Habit: Herb. Habitat: Seasonally flooded grassland and swampy
stream banks, ca. 60 m. Vouchers: Bogdan 3344, Luke WRQ & PA 4078 & 5993 (EA).
Scleria racemosa Poir. Habit: Herb. Habitat: Swampy ground in forest, lake shores,
and swamps, muddy valley bottoms, ca. 0–28 m. Vouchers: SAJIT–006217 (EA,
HIB), Luke WRQ 3607, Magogo & Glover 539, Kirika P 949 (EA).
F71. Dichapetalaceae
2 Genera, 9 Species
Dichapetalum arenarium Breteler Habit: Liana. Habitat: Coastal forest and bushland, 0–450 m. Vouchers: Ngumbau V & Mwadime N V0361 (EA, HIB), Luke
WRQ & Robertson SA 501 (EA).
Dichapetalum fadenii Breteler Habit: Shrub. Habitat: Forest on limestone outcrops,
200–220 m. Voucher: Faden et al. 77/416 (EA): Endemic.
80
Veronicah M. Ngumbau et al. / PhytoKeys 147: 1–191 (2020)
Dichapetalum fructuosum Hiern Habit: Liana. Habitat: Coastal forest, ca. 200 m.
Vouchers: Luke WRQ & Robertson SA 2707, Magogo & Glover 179 (EA).
Dichapetalum madagascariense Poir. var. madagascariense Habit: Liana. Habitat: Evergreen forest, 200 m. Vouchers: Magogo FC & Glover PE 250, Greenway
9652, Verdcourt 1899 (EA).
Dichapetalum mossambicense (Klotzsch) Engl. Habit: Liana. Habitat: Coastal
forest, forest edges, coastal, and secondary bushland, 0–400 m. Vouchers: SAJIT–006028, Ngumbau V & Mwadime N V0334 (EA, HIB), Luke WRQ &
Robertson SA 522 (EA).
Dichapetalum ruhlandii Engl. Habit: Liana. Habitat: Forest and associated bushland, 50–382 m. Vouchers: SAJIT–006079 (EA, HIB), Luke WRQ 871, Verdcourt 1880 (EA).
Dichapetalum sp. ?nov. Habit: Shrub. Habitat: Moist forest, ca. 231 m. Voucher:
Ngumbau V & Mwadime N V113 (EA, HIB): Endemic.
Dichapetalum zenkeri Engl. Habit: Shrub. Habitat: Lowland mostly evergreen forest, 30–700 m. Vouchers: Ngumbau V & Mwadime N V0428 & 0529 (EA, HIB),
Graham MD 2132, Kimeu JM 691 (EA).
Tapura fischeri Engl. Habit: Tree. Habitat: Forest or sometimes riverine, ca. 220 m.
Vouchers: Luke WRQ 873, Gillett & Kibuwa, Faden RB & Evans A 71/730 (EA).
F72. Didiereaceae
1 Genus, 1 Species
Calyptrotheca taitensis (Pax & Vatke) Brenan Habit: Shrub. Habitat: Deciduous
bushland, open woodland, and grassland, ca. 30 m. Voucher: Robertson 6600 (EA).
F73. Dilleniaceae
1 Genus, 2 Species
Tetracera boiviniana Baill. Habit: Shrub. Habitat: Coastal woodland and thicket,
50–350 m. Vouchers: SAJIT–005464 & 005490 (EA, HIB), Robertson SA &
Luke Q 5712 (EA).
Tetracera litoralis Gilg Habit: Shrub. Habitat: Lowland rainforest and dry evergreen
forest, 0–50 (–500) m. Vouchers: SAJIT–005515 & 005977, Ngumbau V &
Mwadime N V0367 & V045 (EA, HIB), Simpson BL 375 (EA).
F74. Dioscoreaceae
2 Genera, 7 Species
Dioscorea asteriscus Burkill Habit: Climber. Habitat: Rainforest, lowland dry evergreen forest, and coastal evergreen bushland, 0–160 m. Vouchers: SAJIT–006003
& 005947 (EA, HIB), Kabuye CHS 84, Kirika P, Nyamongo D & Sanyanyi S
39664, Luke WRQ 2878 (EA).
Checklist of coastal forests of Kenya
81
Dioscorea buchananii Benth. Habit: Climber. Habitat: Riverine forest and near
mangrove swamps, ca. 300 m. Vouchers: Mutanga JG & Kamau P 22, Kuchar P
12003 (EA).
Dioscorea dumetorum (Kunth) Pax Habit: Climber. Habitat: Edges of lowland rainforest, dry evergreen forest, and evergreen bushlands, 0–650 m. Vouchers: SAJIT–006015 & 005524, Ngumbau V & Mwadime N V079 & 0252 (EA, HIB),
Katz SS 32, Magogo FC & Glover PE 1062 (EA).
Dioscorea hirtiflora subsp. orientalis Milne-Redh. Habit: Climber. Habitat: Dry
evergreen forest, thickets, and rocky places, 0–570 m. Vouchers: Jeffery GW 276,
Robertson SA & Luke WRQ 5158 (EA).
Dioscorea quartiniana A. Rich. var. quartiniana Habit: Climber. Habitat: Lowland
rainforest, riverine forest, and forest edges, ca. (50–) 300 m. Vouchers: Mwadime
N 26, Robertson SA 4934 (EA).
Dioscorea sansibarensis Pax Habit: Climber. Habitat: Lowland rainforest, riverine
forest, and coastal evergreen bushland, ca. 0–300 m. Vouchers: SAJIT–005927
(EA, HIB), Brenan JPM & JB Gillet 14644, Gillett JB 18713 (EA).
Tacca leontopetaloides (L.) Kuntze Habit: Herb. Habitat: Grassland, bushland or
woodland, ca. 0–227 m. Vouchers: SAJIT–005375, Ngumbau V & Mwadime
N V0281 (EA, HIB), Magogo FC & Glover PE 484, Napier 6294, RM Graham
2120 (EA).
F75. Droseraceae
1 Genus, 1 Species
Drosera indica L. Habit: Herb. Habitat: Wet mud in swamps, pools, and streams, ca.
30 m. Voucher: Festo L, Luke Q & P 2728 (EA).
F76. Ebenaceae
2 Genera, 15 Species
Diospyros abyssinica (Hiern) F. White subsp. abyssinica Habit: Tree. Habitat: Rainforest and evergreen forest, 0–385 m. Vouchers: Luke WRQ 1855, Muasya JM
708 (EA).
Diospyros amaniensis Gürke Habit: Tree. Habitat: Rainforest or ground water forest,
250–500 m. Vouchers: Luke WRQ & Kahumbu P 4546G, Magogo & Glover 82,
White 11191 (EA).
Diospyros bussei Gürke Habit: Tree. Habitat: Wet bushland or wooded grassland,
0–600 m. Vouchers: Dale IR K3539, Drummond & Hemsley 4183 (EA).
Diospyros consolatae Chiov. Habit: Tree. Habitat: Semi-evergreen or evergreen dry
forest, thicket, and woodland, 0–300 m. Vouchers: Dale IR K3768, Greenway
9815 (EA).
Diospyros greenwayi F. White Habit: Tree. Habitat: Semi-deciduous and evergreen
forest, 0–400 m. Vouchers: SAJIT–005571 & 006062, Ngumbau V & Mwadime
82
Veronicah M. Ngumbau et al. / PhytoKeys 147: 1–191 (2020)
N V0233 (EA, HIB), Mohamed HB 3860, Luke WRQ & Robertson SA 219
(EA): Vulnerable.
Diospyros kabuyeana F. White Habit: Tree. Habitat: Evergreen, semi-deciduous lowland forest, and sometimes riverine, 20–700 m. Vouchers: Dale IR 3426, Luke
WRQ & Robertson SA 525 (EA).
Diospyros loureiroana subsp. rufescens (Caveney) Verdc. Habit: Tree. Habitat: Evergreen bushland, 0–750 m. Vouchers: SAJIT–005479 (EA, HIB), Luke WRQ &
Robertson SA 2779, RM Graham 323 in FD 1821 (EA).
Diospyros mespiliformis Hochst. ex A.DC Habit: Tree. Habitat: Moist semi-deciduous forest and riverine forest, ca. 0–250 m. Voucher: Luke WRQ & Robertson
SA 535 (EA).
Diospyros natalensis (Harv.) Brenan subsp. natalensis Habit: Tree. Habitat: Moist
semi-deciduous forest, dry evergreen forest, and thicket, ca. 0–250 m. Vouchers:
Luke Q 1568, Drummond RB & Hemsley JH 4018, Faden 77/690 (EA).
Diospyros occulta F. White Habit: Tree. Habitat: Evergreen forest, 0–300 m. Vouchers: Luke WRQ 1849, Luke & Robertson 1700 (EA).
Diospyros shimbaensis F. White Habit: Tree. Habitat: Coastal evergreen, 0–450 m.
Vouchers: SAJIT–005545 (EA, HIB), Luke WRQ 1850 (EA): Endangered.
Diospyros squarrosa Klotzsch Habit: Tree. Habitat: Woodland and bushland, and
hillsides, 0–100 m. Vouchers: SAJIT–005477 (EA, HIB), Luke Q 1423, Drummond RB & Hemsley JH 4018 (EA).
Euclea divinorum Hiern Habit: Shrub. Habitat: Grassland, open bushland, and
thicket, 0–200 m. Voucher: Robertson SA & Luke WRQ 74 (EA).
Euclea natalensis subsp. obovata F. White Habit: Shrub. Habitat: Grassland, evergreen, and deciduous thicket, 0–301 m. Vouchers: SAJIT–005971 (EA, HIB),
Gardner HM 1417, Cunningham-van Someren (EA).
Euclea racemosa subsp. schimperi (A. DC.) F. White Habit: Shrub. Habitat: Grassland bushland, woodland, and riverine forest or scrub, ca. 2 m. Voucher: SAJIT–006112 (EA, HB).
F77. Eriocaulaceae
1 Genus, 1 Species
Eriocaulon elegantulum Engl. Habit: Herb. Habitat: Seasonal ponds and ditches, ca.
0–50 m. Vouchers: Ngumbau V & Mwadime N V0549 (EA, HIB), Magogo FC
& Glover PE 355 (EA).
F78. Erythroxylaceae
2 Genera, 3 Species
Erythroxylum emarginatum Thonn. Habit: Shrub. Habitat: Evergreen forest, coastal forest, and transitional bushland, ca. 2 m. Voucher: Mwadime N & Chesire C 184 (EA).
Erythroxylum platyclados Bojer Habit: Tree. Habitat: Evergreen bushland on coral,
sand dunes, and wooded grassland, 12–180 m. Voucher: Luke Q 1418 (EA).
Checklist of coastal forests of Kenya
83
Nectaropetalum kaessneri Engl. Habit: Tree. Habitat: Dry evergreen forest, thickets,
and waste ground, 0–390 m. Voucher: Dale in FD 3874 (EA).
F79. Euphorbiaceae
26 Genera, 87 Species
Acalypha bussei Hutch. Habit: Herb. Habitat: Coastal forest, 0–160 m. Vouchers:
Drummond & Hemsley 3993, Gillett 18640, Verdcourt 1079 (EA).
Acalypha ciliata Forssk. Habit: Herb. Habitat: Wooded grassland, deciduous, and
coastal bushland, 0–150 m. Voucher: Gillespie 219 (EA).
Acalypha echinus Pax & K. Hoffm. Habit: Shrub. Habitat: Riverine forest, 60–
300 m. Vouchers: Festo L, Luke Q & P 2757, Pakia M et al. 988, Drummond &
Hemsley 3340 (EA).
Acalypha engleri Pax Habit: Shrub. Habitat: Forest undergrowth, edges, and associated
bushland, (100–) 400–500 m. Vouchers: Ngumbau V & Mwadime N V018 (EA,
HIB), Luke WRQ & Robertson SA 1556, Magogo FC & Glover PE 140 (EA).
Acalypha fruticosa Forssk. var. fruticosa Habit: Shrub. Habitat: Coastal deciduous
bushland and thicket, ca. 0–272 m. Vouchers: Ngumbau V & Mwadime N V043
(EA, HIB), Kuchar P 10145 (EA).
Acalypha indica L. Habit: Herb. Habitat: Rivers, base of outcrops, and ruderal, ca.
0–100 m. Vouchers: Gillespie 40, Faden RB & Faden AJ 74/1170 & 74/1019 (EA).
Acalypha lanceolata var. glandulosa (Müll.Arg.) Radcl.-Sm. Habit: Herb. Habitat:
Grassland, 0–100 m. Vouchers: SAJIT–006179 (EA, HIB), Luke WRQ & PA
4510, Polhill & Paulo 537 (EA).
Acalypha neptunica Müll.Arg. var. neptunica Habit: Shrub. Habitat: Forest undergrowth, edges, and associated bushland, ca. 36 m. Vouchers: SAJIT–006178 &
005551 (EA, HIB), Luke WRQ & Robertson SA 2698 (EA).
Acalypha neptunica var. pubescens (Pax) Hutch. Habit: Shrub. Habitat: Thickets
and dry forest understory, 0–150 m. Voucher: Rawlins 203 (EA).
Acalypha ornata Hochst. ex A. Rich. Habit: Shrub. Habitat: Forest undergrowth, edges,
wooded grassland, deciduous woodland, and thicket, 0–276 m. Vouchers: Ngumbau
V & Mwadime N V044 & 0220 (EA, HIB), Magogo FC & Glover PE 485A (EA).
Acalypha paniculata Miq. Habit: Shrub. Habitat: Forest, open places, edges, and
riverine, ca. 100–280 m. Vouchers: SAJIT–006152, Ngumbau V & Mwadime N
V115 (EA, HIB), Robertson SA & Luke WRQ 5159 (EA).
Alchornea laxiflora (Benth.) Pax & K. Hoffm. Habit: Shrub. Habitat: Evergreen
forest and riverine thickets near coast, ca. 10 m. Vouchers: SAJIT–005512 (EA,
HIB), Luke Q 1566, Magogo FC & Glover PE 72 (EA).
Argomuellera mjikendae Q. Luke, ined. Habit: Shrub. Habitat: Bushland, ca. 86 m.
Vouchers: Ngumbau V & Mwadime N V0471 (EA, HIB), Luke WRQ et al.
5702 (EA).
Caperonia fistulosa Beille Habit: Herb. Habitat: Swamps, flood plains, and other
seasonally flooded areas, 0–360 m. Vouchers: SAJIT–006195 (EA, HIB), Luke
WRQ & Robertson SA 2815 (EA).
84
Veronicah M. Ngumbau et al. / PhytoKeys 147: 1–191 (2020)
Cavacoa aurea (Cavaco) J. Léonard Habit: Tree. Habitat: Forest, ca. 100 m. Vouchers: Ngumbau V & Mwadime N V0382 (EA, HIB), Thomas Mwadime in Mrs
Robertson SA 7790, Luke WRQ & Robertson SA 2697 (EA).
Croton blanchetianus Baill. Habit: Shrub. Habitat: Forest. Voucher: Luke WRQ et
al. 4333 (EA).
Croton dichogamus Pax Habit: Shrub. Habitat: Dry forest, bushland, and thicket, ca.
253 m. Vouchers: SAJIT–005902, FOKP–10002 (EA, HIB).
Croton kinondoensis G.W. Hu, V.M. Ngumbau & Q.F. Wang, sp. nov. Habit:
Shrub. Habitat: Forest edge, ca. 5 m. Vouchers: SAJIT 007225, Ngumbau V
& Mwadime N V0520 (EA, HIB), Malombe I, Mwadime N & Saidi KI 1647
(EA): Endemic.
Croton megalocarpoides Friis & M.G. Gilbert Habit: Tree. Habitat: In semi-evergreen coastal forest, and bushland, 5–275 m. Vouchers: Robertson SA & Luke
WRQ 5551, Muchiri 458 (EA, HIB).
Croton menyharthii Pax Habit: Shrub. Habitat: Deciduous bushland and thicket, ca.
0–50 m. Vouchers: Drummond & Hemsley 4116, Gillespie 378 (EA).
Croton pseudopulchellus Pax Habit: Shrub. Habitat: Dry evergreen forest, deciduous woodland, bushland, and thicket, ca. 0–59 m. Vouchers: Ngumbau V &
Mwadime N V0440 (EA, HIB), Festo L & Luke Q 2596, Ross KS 133 (EA).
Croton sylvaticus Hochst. Habit: Tree. Habitat: Secondary forest, forest edges, along
rivers, and around lakes, ca. 60–410 m. Vouchers: Ngumbau V & Mwadime N
V0300 & 0111 (EA, HIB), Luke WRQ & Robertson SA 515 (EA).
Croton talaeporos Radcl.–Sm. Habit: Tree. Habitat: Coastal bushland and wooded
grassland, 0–100 m. Vouchers: Muchiri J 565, J Adamson in Bally 5935 (EA).
Dalechampia parvifolia Lam. var. parvifolia Habit: Climber. Habitat: Coastal
bushland, ca. 0–600 m. Vouchers: Magogo FC & Glover PE 864, Luke Q 5679,
Luke Q 1462, Graham RM 1904, Drummond & Hemsley 4089, Gray M & Luke
WRQ 270, Drummond & Hemsley 1054 (EA).
Dalechampia trifoliata Peter ex Verdc. & Greenway Habit: Climber. Habitat: Deciduous and coastal bushland or woodland, 60–600 m. Vouchers: Drummond &
Hemsley 4089, Field 3 (EA).
Erythrococca kirkii (Müll.Arg.) Prain Habit: Shrub. Habitat: Forest edges, coastal bushland, and thicket, ca. 0–181 m. Vouchers: Ngumbau V & Mwadime N
V0224 (EA, HIB), Magogo FC & Glover PE 219 (EA).
Erythrococca pentagyna Radcl.-Sm. Habit: Shrub. Habitat: Moist coastal forest, 60–
160 m. Vouchers: Ngumbau V & Mwadime N V0464 (EA, HIB), Luke WRQ
3136, Luke WRQ & Robertson SA 1911, Luke WRQ et al. 6191, Drummond &
Hemsley 3944, Kokwaro 3931, Faden 77/524 (EA): Endemic.
Erythrococca pubescens Radcl.-Sm. Habit: Shrub. Habitat: Forest, ca. 0–380 m.
Vouchers: Ngumbau V & Mwadime N V0464, Drummond & Hemsley
4168 (EA).
Erythrococca usambarica Prain Habit: Shrub. Habitat: Primary and secondary evergreen forest, ca. 300–400 m. Vouchers: Luke WRQ 872, Gillett JB & Robertson
SA 23996 (EA).
Checklist of coastal forests of Kenya
85
Euphorbia acalyphoides Hochst. ex Boiss. Habit: Herb. Habitat: Bushland, ca.
10 m. Voucher: Festo L, Luke Q & P 2625 (EA).
Euphorbia breviarticulata Pax var. breviarticulata Habit: Shrub. Habitat: AcaciaCommiphora bushland, ca. 60–365 m. Vouchers: Bally & Smith 14404, Kuchar P
7323 (EA).
Euphorbia buruana Pax Habit: Herb. Habitat: Bushland, ca. 100 m. Voucher: M van
Leeuwen 19 (EA).
Euphorbia bussei var. kibwezensis (N.E.Br.) S. Carter Habit: Tree. Habitat: Deciduous woodland. Voucher: Polhill R & Paulo S 816 (EA).
Euphorbia candelabrum Trémaux ex Kotschy var. candelabrum Habit: Tree. Habitat: Wooded grassland, ca. (15–) 450 m. Vouchers: Robertson SA, Luke WRQ &
Awimbo J 5248, Luke WRQ 954 (EA).
Euphorbia candelabrum var. bilocularis (N.E.Br.) S. Carter Habit: Tree. Habitat:
Open wooded grassland, ca. 550 m. Voucher: Bally E 212 (EA).
Euphorbia crotonoides Boiss. Habit: Herb. Habitat: Open woodland or scattered
bushland, ca. 186 m. Voucher: Ngumbau V & Mwadime N V0337 (EA, HIB).
Euphorbia cryptospinosa P.R.O. Bally Habit: Herb. Habitat: Open dry deciduous
bushland, ca. 230 m. Voucher: Luke WRQ & Robertson SA 2527 (EA).
Euphorbia cuneata Vahl subsp. cuneata Habit: Shrub. Habitat: Coastal forest and often cultivated as a hedge plant, 0–160 m. Vouchers: Tweedie 1024, Gillespie 62 (EA).
Euphorbia fluminis S. Carter Habit: Shrub. Habitat: Dry deciduous woodland, ca.
15 m. Voucher: Gillett JB 19534 (EA): Endemic.
Euphorbia furcata N.E.Br. Habit: Shrub. Habitat: Acacia bushland, ca. 300 m.
Voucher: Bally 1326 (EA).
Euphorbia gossypina Pax subsp. gossypina Habit: Shrub. Habitat: In Acacia-Commiphora bushland, ca. 30 m. Vouchers: Drummond & Hemsley 4044, Faden RB &
Faden A74/1054 (EA).
Euphorbia heterochroma subsp. tsavoensis S. Carter Habit: Shrub. Habitat: Deciduous bushland, ca. 550 m. Voucher: Luke WRQ 940 (EA).
Euphorbia heterophylla L. Habit: Herb. Habitat: Disturbed localities, cultivated, and
wasted land, ca. 500 m. Voucher: Magogo FC & Glover PE 984 (EA): Naturalized.
Euphorbia hirta L. Habit: Herb. Habitat: Roadsides and waste places, ca. 20 m.
Vouchers: Magogo FC & Glover PE 724, Polhill & Paulo 535 (EA).
Euphorbia indica Lam. Habit: Herb. Habitat: Grassland and seasonally water-logged
places, ca. 5 m. Vouchers: SAJIT–006099 (EA, HIB), Tweedie 3138 (EA).
Euphorbia invenusta (N.E.Br.) Bruyns Habit: Herb. Habitat: Deciduous woodland,
thicket, ca. 50 m. Vouchers: Robertson SA & Luke WRQ 4464, Greenway PJ &
Rawlins SP 9457 (EA).
Euphorbia kalisana S. Carter Habit: Herb. Habitat: Acacia-Commiphora bushland,
ca. 100 m. Voucher: Faden 74/998 (EA).
Euphorbia kaessneri Pax Habit: Shrub. Habitat: Grassland and bushland near the
coast, 20–100 m. Vouchers: SAJIT–006441 (EA, HIB), Mungai GM 166/80 (EA).
Euphorbia neovirgata Bruyns Habit: Herb. Habitat: Bushland, 50–200 m. Voucher:
Bally 12196 (EA).
86
Veronicah M. Ngumbau et al. / PhytoKeys 147: 1–191 (2020)
Euphorbia nyikae Pax ex Engl. var. nyikae Habit: Tree. Habitat: Deciduous woodland, 250–700 m. Vouchers: Bally 5781, Balslev 47 (EA).
Euphorbia nyikae var. neovolkensii (Pax) S. Carter Habit: Tree. Habitat: Sand
dunes, seashores, and deciduous woodland, 2–250 m. Vouchers: Robertson SA &
Luke WRQ 5377, Luke WRQ et al. 4331 (EA).
Euphorbia pereskiifolia Houllet ex Baill. Habit: Shrub. Habitat: Bushland, ca.
136 m. Vouchers: SAJIT–005950 (EA, HIB), Robertson SA & Luke WRQ
5538 (EA).
Euphorbia systyloides Pax Habit: Herb. Habitat: Bushland, ca. 95 m. Voucher: SAJIT–006132 (EA, HIB).
Euphorbia tanaensis P.R.O. Bally & S. Carter Habit: Tree. Habitat: Semi-deciduous
swampy forest, ca. 15 m. Voucher: Robertson SA 4327 (EA): Critically Endangered, Endemic.
Euphorbia taruensis S. Carter Habit: Herb. Habitat: Leaf-litter amongst rocks beneath trees, 150–480 m. Voucher: Lauranos & Newton 12315 (EA): Endemic.
Euphorbia tenuispinosa Gilli var. tenuispinosa Habit: Shrub. Habitat: Acacia-Commiphora bushland and evergreen forest, 0–350 m. Vouchers: Sangai GW 15629,
Drummond & Hemsley 4182 (EA).
Euphorbia tirucalli L. Habit: Tree. Habitat: Woodland, ca. 50 m. Vouchers: Jeffery
GW 701, Festo L, Luke Q & P 2643 (EA).
Euphorbia pervittata S. Carter Habit: Shrub. Habitat: Open deciduous bushland,
ca. 500 m. Voucher: Mungai et al. 367 (EA).
Euphorbia wakefieldii N.E.Br. Habit: Tree. Habitat: Bushland and forest remnants,
50–275 m. Vouchers: Ngumbau V & Mwadime N V0416 (EA, HIB), Bally PRO
s.n., Robertson SA & Luke WRQ 4536B (EA).
Excoecaria bussei (Pax) Pax Habit: Tree. Habitat: Deciduous woodland, bushland,
and wooded grassland, ca. 5 m. Voucher: Festo L & Luke Q 2553 (EA).
Excoecaria madagascariensis (Baill.) Müll.Arg. Habit: Shrub. Habitat: Forest and
thicket, commonly along rivers, ca. 50 m. Voucher: Dale in FD 3651 (EA).
Givotia gosai Radcl.–Sm. Habit: Shrub. Habitat: Acacia-Commiphora open bushland, 50–600 m. Voucher: Gillett 16487 (EA).
Jatropha curcas L. Habit: Tree. Habitat: Grassland, ca. 0–200 m. Voucher: Jeffery
GW 418 (EA).
Jatropha hildebrandtii Pax var. hildebrandtii Habit: Shrub. Habitat: Dunes in open
coastal bushland, 0–230 m. Voucher: Robertson SA 3793 (EA).
Jatropha mollis Pax Habit: Shrub. Habitat: Acacia-Commiphora bushland, ca. 90 m.
Voucher: Gillett 16430 (EA).
Jatropha multifida L. Habit: Shrub. Habitat: Cultivated as a hedge plant and garden
ornamental. Voucher: Bally PRO 2156 (EA).
Jatropha prunifolia Pax Habit: Herb. Habitat: Dry coastal forest, thicket, and bushland, (20–) 60–350 m. Vouchers: Ngumbau V & Mwadime N V0156 (EA, HIB),
Luke WRQ & Robertson SA 1472, Greenway PJ & Kanuri K 12756 (EA).
Jatropha spicata Pax Habit: Herb. Habitat: Deciduous bushland and thicket, ca.
45 m. Voucher: Sangai GW 961 (EA).
Checklist of coastal forests of Kenya
87
Jatropha stuhlmannii Pax subsp. stuhlmannii Habit: Herb. Habitat: Grassland and
coastal bushland, 0–100 m. Voucher: Gillett JB 20383 (EA).
Jatropha velutina Pax & K. Hoffm. Habit: Herb. Habitat: Thicket and in Acacia-Commiphora bushland, 300–350 m. Voucher: Robertson SA & Luke WRQ 2213 (EA).
Macaranga capensis (Baill.) Sim var. capensis Habit: Tree. Habitat: Evergreen forest,
(10–) 75 m. Voucher: Luke WRQ & Robertson SA 2759 (EA).
Mallotus oppositifolius (Geiseler) Müll.Arg. Habit: Shrub. Habitat: Moist forests,
5–36 m. Vouchers: SAJIT–006183 & 005917, Ngumbau V & Mwadime N V0517
(EA, HIB), Luke Q 1401, Thomas Mwadime & Mrs Robertson SA 7798 (EA).
Micrococca mercurialis (L.) Benth. Habit: Herb. Habitat: Woodland and bushland,
ca. 36 m. Vouchers: SAJIT–006180 (EA, HIB), Festo L & Luke Q 2467, Luke
WRQ 3338 (EA).
Micrococca scariosa Prain Habit: Shrub. Habitat: Forest and associated thickets,
10–170 m. Vouchers: Ngumbau V & Mwadime N V029 (EA, HIB), Faden RB
216 (EA).
Plesiatropha sp. (Mildbraedia sp. A of FTEA) Habit: Tree. Habitat: Bushland or
forest edges, 70 m. Vouchers: Moggridge 398, Padwa B715 (EA).
Necepsia castaneifolia subsp. kimbozensis (Radcl.-Sm.) Bouchat & J.Léonard
Habit: Tree. Habitat: Moist forests, ca. 150 m. Voucher: Malombe I, Mwadime N
& Saidi 1654 (EA).
Plesiatropha carpinifolia (Pax) Breteler var. carpinifolia Habit: Shrub. Habitat:
Forest, forest edges, and coastal woodland, 0–272 m. Vouchers: SAJIT–005923,
Ngumbau V & Mwadime N V042 (EA, HIB), Luke WRQ & Robertson SA 1492,
Ross KS 196, Festo L & Luke Q 2533 (EA).
Pycnocoma littoralis Pax Habit: Shrub. Habitat: Dry coastal forest, ca. 90 m. Vouchers: Vorontsova MS 117, Luke WRQ & Robertson SA 2832 (EA): Vulnerable.
Ricinodendron heudelotii var. tomentellum (Hutch. & E.A.) Radcl.–Sm. Habit:
Tree. Habitat: Semi-deciduous and dry evergreen forest, ca. 220–600 m. Vouchers:
RB & AJ Faden 74/312 & 71/771 (EA).
Shirakiopsis trilocularis (Pax & K. Hoffm.) Esser Habit: Tree. Habitat: Evergreen
forest, ca. 210 m. Voucher: Luke WRQ & PA 4067 (EA).
Spirostachys africana Sond. Habit: Tree. Habitat: Deciduous woodland, bushland, and
wooded grassland, ca. 15 m. Vouchers: Robertson SA 4236, Dale in FD 3769 (EA).
Spirostachys venenifera (Pax) Pax Habit: Tree. Habitat: Along rivers in deciduous
and coastal bushland, ca. 0–30 m. Voucher: Verdcourt 5295A (EA).
Suregada zanzibariensis Baill. Habit: Shrub. Habitat: Coastal forest, woodland,
and bushland, 0–213 m. Vouchers: SAJIT–006204, 005948, 005466 & 004650,
Ngumbau V & Mwadime N V0328 (EA, HIB), Kimeu JM, Meso M & Ot 624,
Luke WRQ 3154 (EA).
Tannodia tenuifolia var. glabrata Prain Habit: Shrub. Habitat: Lowland evergreen
forest and riverine, 160–660 m. Vouchers: Hawthorne 178, Faden et al. 77/514,
Luke WRQ 3146 (EA).
Tragia furialis Bojer Habit: Climber. Habitat: Forest and disturbed places, 0–250 m.
Vouchers: SAJIT–006117 (EA, HIB), Luke Q 1403 (EA).
88
Veronicah M. Ngumbau et al. / PhytoKeys 147: 1–191 (2020)
Tragia glabrescens Pax Habit: Herb. Habitat: Coastal bushland and thickets, 170–
365 m. Vouchers: Kuchar 7342, Polhil & Paulo 491, Kassner 466 (EA).
Tragia hildebrandtii Müll.Arg. Habit: Herb. Habitat: Riverine, flood plains or disturbed places, 15 m. Voucher: Magogo FC & Glover PE 1000 (EA).
Tragia kirkiana Müll.Arg. Habit: Climber. Habitat: Wooded and open grassland, ca.
183 m. Vouchers: Ngumbau V & Mwadime N V0195 (EA, HIB), Magogo FC &
Glover PE 1136 (EA).
Tragia plukenetii Radcl.–Sm. Habit: Climber. Habitat: Riverine, floodplains, and
valley grassland, 0–100 m. Vouchers: Rawlins 83, JM Reitsma 410 (EA).
Tragiella natalensis (Sond.) Pax & K. Hoffm. Habit: Climber. Habitat: Forest edges,
bushland, riverine, lakesides and disturbed places, ca. 80–230 m. Vouchers: Ngumbau V & Mwadime N V061 (EA, HIB), Magogo FC & Glover PE 408 (EA).
F80. Fabaceae
80 Genera, 226 Species
Abrus precatorius subsp. africanus Verdc. Habit: Liana. Habitat: Grassland, thicket,
and bushland, ca. 15 m. Vouchers: Tweedie 1249, Festo L, Luke Q & P 2708,
Magogo FC & Glover PE 644 (EA).
Abrus sp. A of FTEA Habit: Herb. Habitat: Forest edge and burnt grassland, 300–
420 m. Vouchers: Magogo & Glover 548 & 122, Luke WRQ 3163 (EA): Endemic.
Aeschynomene cristata var. pubescens J. Léonard Habit: Shrub. Habitat: Swamps,
edges of dams, lakes, and rivers, ca. 350 m. Vouchers: Drummond & Hemsley
3930 & 4177, Greenway & Rawlins 9448 (EA).
Aeschynomene cristata Vatke var. cristata Habit: Shrub. Habitat: Swamps in grassland, ca. 5–30 m. Voucher: Festo L & Luke Q 2455 (EA).
Aeschynomene indica L. Habit: Shrub. Habitat: Mostly in wet places, floodplain
grassland, wooded grassland, and desert grassland, ca. 96 m. Vouchers: Polhill &
Paulo 667, Gillespie 157, JB Gillett 16508 (EA).
Aeschynomene uniflora E. Mey. var. uniflora Habit: Shrub. Habitat: Swamps,
marshy grassland, riverside, and bushland, ca. 15 m. Vouchers: Drummond &
Hemsley 4016, Bogdan 2617, Rawlins 36, Magogo FC & Glover PE 757, Luke
Q 5671 (EA).
Aeschynomene sp. B of FTEA Habit: Shrub. Habitat: Roadsides, ca. 25 m. Vouchers:
Rawlins 410, Patterson 32 (EA): Endemic.
Afzelia quanzensis Welw. Habit: Tree. Habitat: Thickets, woodland, and lowland dry
evergreen forest, ca. 0–300 m. Vouchers: SAJIT–005470 (EA, HIB), Battiscombe
35, Hildebrandt 1967, Jeffery 115, Gilbert MG & Kanuri K 5863, Magogo FC
& Glover PE 49 (EA).
Albizia adianthifolia (Schum.) W. Wight var. adianthifolia Habit: Tree. Habitat:
Lowland rainforest, deciduous woodland, and wooded grassland, ca. 30 m. Vouchers: Hildebrandt 1936, Gardner 1443, RM Graham 312, Cunningham-van Someren GR Sh49 (EA).
Checklist of coastal forests of Kenya
89
Albizia anthelmintica Brongn. Habit: Tree. Habitat: In dry open woodland, ca. 50–
110 m. Vouchers: Ngumbau V & Mwadime N V0492 (EA, HIB), Rukunga G,
Mungai G & Muthaura C 019/04 (EA).
Albizia glaberrima var. glabrescens (Oliv.) Brenan Habit: Tree. Habitat: Lowland
rainforest, riverine forest, and coastal evergreen bushlands, 0–760 m. Vouchers:
Dale in FH 3540, Mulwa PCW 58 (EA).
Albizia gummifera (J.F. Gmel.) C.A.Sm. var. gummifera Habit: Tree. Habitat: Rainforest and riverine forest, ca. 30 m. Vouchers: Shimba Hills Survey Unit 64, Luke
WRQ 192 (EA).
Albizia versicolor Oliv. Habit: Tree. Habitat: Deciduous woodland, bushland, and
wooded grassland, ca. 0–153 m. Vouchers: Ngumbau V & Mwadime N V0508
(EA, HIB), St. Barbe Baker 43, CW Elliot Q 87 in FH 1677 & in CM 14009, RM
Graham N331 in FH 1724 (EA).
Alysicarpus glumaceus subsp. macalusoi (Mattei) Verdc. Habit: Herb. Habitat: Sand
dunes close to the sea, 0–10 m. Vouchers: Rawlins 47 & 99, Greenway & Rawlins
9285, Luke Q 5465 (EA).
Alysicarpus glumaceus var. intermedius Verdc. Habit: Herb. Habitat: Lowland and
coastal bushland, 0–300 m. Vouchers: Bogdan A 2601, Drummond & Hemsley
3779 (EA).
Alysicarpus ovalifolius (Schumach.) J. Léonard Habit: Herb. Habitat: Grassland, ca.
60 m. Vouchers: Tweedie 963, Rawlins 621, Polhill & Paulo 662 (EA).
Alysicarpus vaginalis (L.) DC. var. vaginalis Habit: Herb. Habitat: Old cultivations,
roadsides, bushland, and open woodland, ca. 36 m. Vouchers: SAJIT–006168
(EA, HIB), Drummond & Hemsley 1063 & 3840, Luke Q 5682 (EA).
Alysicarpus vaginalis var. villosus Verdc. Habit: Herb. Habitat: Not known, ca. 200–
420 m. Voucher: Magogo & Glover 58 (EA).
Angylocalyx braunii Harms Habit: Tree. Habitat: Understorey of lowland rainforest,
dry evergreen, and riverine forest, 0–800 m. Vouchers: SAJIT–005555, Ngumbau
V & Mwadime N V031 (EA, HIB), Drummond & Hemsley 1145, Greenway
& Rawlins 9368, CW Elliot in FD 1550, Robertson SA & Luke WRQ 5173
(EA): Vulnerable.
Bauhinia mombassae Vatke Habit: Shrub. Habitat: Forest in riverine areas, ca. 210 m.
Vouchers: Ngumbau V & Mwadime N V086 (EA, HIB), Wakefield, Hildebrandt
2006, Kelly 1230 & in CM 13911, Faden RB & AJ, Gillett JB & Gachathi N
77/442, Luke WRQ & Robertson SA 2793 (EA): Endangered, Endemic.
Bauhinia taitensis Taub. Habit: Shrub. Habitat: Deciduous bushland, 330–610 m.
Voucher: Parker I 307 (EA).
Brachystegia spiciformis Benth. Habit: Tree. Habitat: Deciduous woodland, ca.
15 m. Vouchers: Greenway 9657, Jeffery 102, Moomaw 940, Kimeu JM 527,
Magogo FC & Glover PE 456 (EA).
Caesalpinia angolensis (Oliv.) Herend. & Zarucchi Habit: Liana. Habitat: Lowland rainforest, swamp forest, and coastal evergreen bushland, ca. 20 m. Voucher:
Mwadime N 1 (EA).
90
Veronicah M. Ngumbau et al. / PhytoKeys 147: 1–191 (2020)
Caesalpinia bonduc (L.) Roxb. Habit: Shrub. Habitat: Near seashores, 0–15 m.
Vouchers: Ngumbau V & Mwadime N V0519 (EA, HIB), RM Graham R 144 in
FH 1562, F Thomas 199, Bally 2192 in CM 11617, Kirika P 203 (EA).
Caesalpinia insolita (Harms) Brenan & J. B. Gillett Habit: Tree. Habitat: In a
wooded gulley, ca. 150 m. Vouchers: Dale in FH 3572 & in CM 9273, Brenan,
Gillett, Kanuri & Chomba 14599, Robertson SA & Luke WRQ 5982 (EA).
Caesalpinia pulcherrima (L.) Sw. Habit: Shrub. Habitat: Near beach, ca. 5 m.
Voucher: SAJIT–005456 (EA, HIB): Cultivated.
Caesalpinia trothae Harms subsp. trothae Habit: Shrub. Habitat: Deciduous bushland, ca. 150 m. Vouchers: RM Graham U 760 in FH 2239 & in CM 13918,
Bamps P 6369 (EA).
Caesalpinia volkensii Harms Habit: Shrub. Habitat: Margins of lowland rainforest, 90–417 m. Vouchers: Ngumbau V & Mwadime N V0321 (EA, HIB), Birch
62/146, Kabuye CHS, Gilbert VC & Robertson SA 110 (EA).
Canavalia africana Dunn Habit: Liana. Habitat: Low altitude evergreen forest,
5–150 m. Voucher: Kassner 203 (EA).
Canavalia cathartica Thouars Habit: Liana. Habitat: Coastal bushland on coral,
0–30 (–420) m. Vouchers: SAJIT–006016, Ngumbau V & Mwadime N V0187
(EA, HIB), Moomaw 860, Verdcourt 1071, Thorold 1554, Magogo FC & Glover
PE 64 (EA).
Canavalia rosea (Sw.) DC. Habit: Liana. Habitat: Beaches and edges of coastal bushland, 0–30 m. Vouchers: SAJIT–006257 (EA, HIB), Birch 62/75, Tweedie 1010,
Elliot in FD 3364 (EA).
Cassia abbreviata subsp. beareana (Holmes) Brenan Habit: Tree. Habitat: Margin
of evergreen forest, 460–1200 m. Vouchers: SAJIT–005998 & 006001, Ngumbau
V & Mwadime N V0180 (EA, HIB), Jeffery 96, J Adamson 280 in Bally 5971,
Kuchar P 13699, Magogo FC & Glover PE 875, Festo L & Luke Q 2421 (EA).
Cassia abbreviata subsp. kaessneri (Baker f.) Brenan Habit: Tree. Habitat: Wooded
grassland and deciduous bushland, ca. 350 m. Vouchers: Drummond & Hemsley
4059, Luke WRQ & Robertson SA 2526 (EA).
Cassia afrofistula Brenan var. afrofistula Habit: Tree. Habitat: Coastal evergreen
bushland, 0–90 m. Vouchers: Ngumbau V & Mwadime N V0521 (EA, HIB), J
Adamson 192 in Bally 6092, Tweedie 996, RM Graham Z 303 in FH 1791 (EA).
Centrosema pubescens Benth. Habit: Climber. Habitat: Forest, ca. 250 m. Voucher:
Ngumbau V & Mwadime N V0243 (EA, HIB): Naturalized.
Chamaecrista absus (L.) H. S. Irwin & Barneby Habit: Herb. Habitat: Grassland,
deciduous bushland, and waste ground, ca. 140 m. Vouchers: Ngumbau V &
Mwadime N V0232 (EA, HIB), Greenway & Rawlins 8959, Kirika P, Muthoka P
& Mbale M 745 (EA).
Chamaecrista fallacina (Chiov.) Lock Habit: Herb. Habitat: Grassland, ca. 100 m.
Voucher: Drummond & Hemsley 3766 (EA).
Chamaecrista mimosoides (L.) Greene Habit: Herb. Habitat: Forest margins, wooded grassland, grassland, cultivated, and waste ground, ca. 0–227 m. Vouchers:
Ngumbau V & Mwadime N V0279 (EA, HIB), Tweedie 992, Moggridge 481,
Checklist of coastal forests of Kenya
91
RM Graham DD453 in FH 1992, Jex Blake in Napier 2267 in CM 4980, Festo L
& Luke Q 2452, Luke WRQ et al. 5709 (EA).
Chamaecrista telfairiana (Hook.f.) Lock Habit: Herb. Habitat: Forest margin, ca.
10 m. Vouchers: Luke Q 6134, Magogo FC & Glover PE 341 (EA).
Chamaecrista zambesica (Oliv.) Lock Habit: Herb. Habitat: Grassland and cultivated land, 80–460 m. Vouchers: Jeffery 232, Jeffery 288, Whyte, Festo L Luke Q
& P 2627 (EA).
Clitoria falcata Lam. Habit: Climber. Habitat: Forest, ca. 15–280 m. Vouchers:
SAJIT–006230, Ngumbau V & Mwadime N V0249 (EA, HIB), Nyange M
456 (EA).
Clitoria ternatea L. Habit: Climber. Habitat: Grassland, scrub, bushland, and old cultivations, 0–80 m. Vouchers: Battiscombe 267, Magogo FC & Glover PE 991 (EA).
Cordyla africana Lour. Habit: Tree. Habitat: Riverine and ground water forest, ca.
150 m. Vouchers: Dale 3852, Luke WRQ 882 (EA).
Craibia brevicaudata (Vatke) Dunn subsp. brevicaudata Habit: Tree. Habitat:
Evergreen coastal forest and fringing forest, 0–382 m. Vouchers: SAJIT–006077
(EA, HIB), Battiscombe 466, Greenway 9493, Muchiri J 426 (EA).
Crotalaria axillaris Aiton Habit: Shrub. Habitat: Forest margins, deciduous woodland, bushland, and on coral crags, ca. 0–226 m. Vouchers: Ngumbau V &
Mwadime N V046 (EA, HIB), Polhill & Paulo 843, Festo L & Luke Q 2539,
Magogo FC & Glover PE 142 (EA).
Crotalaria barkae subsp. teitensis (Sacleux) Polhill Habit: Herb. Habitat: Deciduous woodland and bushland, ca. 243 m. Vouchers: Greenway 10436, Gillett JB
19548 (EA).
Crotalaria barkae subsp. zimmermannii (Baker f.) Polhill Habit: Herb. Habitat:
Grassland and forest edges, 0–700 m. Vouchers: Hacker 122, Rawlins in EAH
11206, Greenway & Rawlins in EAH 11335, Soest LV 16013 (EA).
Crotalaria bernieri Baill. Habit: Herb. Habitat: Grassland, coral rags, and foreshores, 0–100 m. Vouchers: Thorold 1562, Tweedie 1002, Greenway & Rawlins
9464 (EA).
Crotalaria deserticola subsp. orientalis Polhill Habit: Herb. Habitat: Coastal grassland and open woodland, ca. 0–243 m. Vouchers: RM Graham Q537 in FD 1938,
Thorold 1557, Greenway & Rawlins 8960 (EA).
Crotalaria emarginata Benth. Habit: Herb. Habitat: Grassland, swamp margins,
cultivated ground, and roadsides, 0–300 m. Vouchers: Ngumbau V & Mwadime
N V0288 (EA, HIB), Verdcourt 1105, Drummond & Hemsley 1033, Greenway
& Rawlins 8958, Magogo FC & Glover PE 99 (EA).
Crotalaria emarginella Vatke Habit: Herb. Habitat: Deciduous bushland, semi-desert grassland, upland evergreen bushland, and grassland, ca. 750 m. Voucher:
Verdcourt 2095 (EA).
Crotalaria goodiiformis Vatke Habit: Shrub. Habitat: Upland rainforest, dry evergreen forest, deciduous woodland, bushland, and wooded grassland, ca. 75 m.
Vouchers: Drummond & Hemsley 3869, Luke Q 1526, Robertson SA & Luke
WRQ 5998 (EA).
92
Veronicah M. Ngumbau et al. / PhytoKeys 147: 1–191 (2020)
Crotalaria grata Polhill Habit: Shrub. Habitat: Deciduous coastal bushland and
grassland, below 50 m. Vouchers: MacNaughton 138 in FD 2717, Rawlins 781,
Polhill & Paulo 583 (EA): Endemic.
Crotalaria hyssopifolia Klotzsch Habit: Herb. Habitat: Deciduous woodland and grassland, ca. 10 m. Vouchers: RM Graham in FD 2026, Festo L & Luke Q 2607 (EA).
Crotalaria juncea L. Habit: Herb. Habitat: Deciduous bushland and grassland, ca.
100 m. Voucher: Jeffery 530 (EA).
Crotalaria kirkii Baker Habit: Herb. Habitat: Grassland and Brachystegia woodland,
ca. 100 m. Vouchers: RM Graham 273, Hildebrandt 1932, Thorold 1558 (EA).
Crotalaria laburnifolia L. subsp. laburnifolia Habit: Shrub. Habitat: Deciduous bushland, grassland, and secondary scrub, 0–360 m. Voucher: Polhill & Paulo 707 (EA).
Crotalaria laburnoides Klotzsch var. laburnoides Habit: Herb. Habitat: Grassland,
sand dunes, coral outcrops, and bushland near coast, ca. 10 m. Vouchers: SAJIT–006097 & 006259 (EA, HIB), Bogdan 5354, Irwin 173, Bogdan 2536 (EA).
Crotalaria laburnoides var. nudicarpa Polhill Habit: Herb. Habitat: Dry coastal
scrub, beach dunes, and woodland, 0–10 m. Vouchers: Courtald 21, Tweedie 2201
(EA): Endangered.
Crotalaria malindiensis Polhill Habit: Herb. Habitat: Woodland and Acacia bushland, grassland, and dunes near seashore, 70–230 m. Vouchers: Bogdan 2523,
Tweedie 939, Gillespie 220, Festo L & Luke Q 2554 (EA).
Crotalaria microcarpa Benth. Habit: Herb. Habitat: Brachstegia woodland, deciduous
bushland, and grassland, ca. 100 m. Vouchers: Verdcourt 1928, Bork 15408 (EA).
Crotalaria patula Polhill Habit: Herb. Habitat: Deciduous bushland, grassland, cultivated ground, and roadsides, 125–420 m. Vouchers: Drummond & Hemsley
4055, Polhill & Paulo 912 (EA).
Crotalaria polysperma Kotschy Habit: Herb. Habitat: Grassland, woodland, often in
disturbed places. Voucher: Luke WRQ & PA sr.
Crotalaria retusa L. var. retusa Habit: Herb. Habitat: Coastal grassland, waste places,
and old cultivations, 0–240 m. Vouchers: SAJIT–005454, Ngumbau V & Mwadime
N V0274 (EA, HIB), Drummond & Hemsley 3898, Festo L & Luke Q 2356 (EA).
Crotalaria retusa var. tunguensis (Lima) Polhill Habit: Herb. Habitat: Coastal
grassland, bushland, coral crags, and salt marsh edges, 0–150 m. Vouchers: Napier
3283, Verdcourt 1097, Rawlins 875, Ndakala J 120 (EA).
Crotalaria rhynchocarpa Polhill Habit: Shrub. Habitat: Dunes and secondary coastal
bushland, less than 50 m. Vouchers: Polhill & Paulo 706, CW Elliot in FD 3375,
Verdcourt 2120, Greenway PJ & Rawlins SP 9437 (EA).
Crotalaria vallicola Baker f. Habit: Herb. Habitat: Forest, ca. 136 m. Voucher: SAJIT–005954 (EA, HIB).
Crotalaria vasculosa Benth. Habit: Herb. Habitat: Grassland and woodland, ca.
20 m. Vouchers: Jeffery 847, Tweedie 1049, Luke 3411 (EA).
Cynometra greenwayi Brenan Habit: Tree. Habitat: Sand dunes with scattered coral
rock outcrops, ca. 8 m. Voucher: Greenway PJ 10440 (EA): Endemic.
Cynometra lukei Beentje Habit: Tree. Habitat: Forest, 1–50 m. Voucher: Q Luke &
SA Robertson 1267 (EA): Endangered.
Checklist of coastal forests of Kenya
93
Cynometra suaheliensis (Taub.) Baker f. Habit: Tree. Habitat: Lowland dry evergreen forest, riverine forest, and coastal evergreen bushland, 0–150 m. Vouchers:
RM Graham U 767 in FH 2238, Polhill & Paulo 737 & 824, Robertson SA 7257,
Moomaw JC 1250 (EA).
Cynometra webberi Baker f. Habit: Tree. Habitat: Evergreen woodland and deciduous woodland, 0–150 m. Vouchers: Drummond & Hemsley 4257, RM Graham
B 699 in FH 2168, Polhill & Paulo 838, Graham RM 2168, Luke WRQ & Robertson SA 2814 (EA): Vulnerable.
Dalbergia boehmii Taub. subsp. boehmii Habit: Tree. Habitat: Dry evergreen, deciduous types of forest, woodland, bushland, and thicket, ca. 0–110 m. Vouchers:
Ngumbau V & Mwadime N V0489 (EA, HIB), Wakefield (EA).
Dalbergia bracteolata Baker Habit: Liana. Habitat: Coastal and deciduous thicket,
ca. 100 m. Vouchers: Kassner 220, Luke WRQ & Robertson SA 2780 (EA).
Dalbergia eremicola Polhill Habit: Shrub. Habitat: Bushlands and thickets, 75–
810 m. Voucher: Bally PRO & Melville R 15252 (EA): Near Threatened.
Dalbergia gloveri Q. Luke, ined. Habit: Liana. Habitat: Coastal forest, 30–320 m.
Voucher: Luke WRQ & Baer S 8247 (EA): Endangered, Endemic.
Dalbergia melanoxylon Guill. & Perr. Habit: Tree. Habitat: Deciduous woodland
deciduous, coastal bushland, and wooded grassland, ca. 0–300 m. Vouchers:
Drummond & Hemsley 4139, Greenway 8968, Robertson SA & Luke WRQ
5518, Magogo FC & Glover PE 841 (EA): Near Threatened.
Dalbergia arbutifolia Baker subsp. arbutifolia Habit: Shrub. Habitat: Woodland,
ca. 0–300 m. Voucher: Chidzinga S & Mwadime N 752 (EA).
Dalbergia vacciniifolia Vatke Habit: Liana. Habitat: Coastal bushland and thicket,
0–100 m. Vouchers: Bally 8896, Dale in FD 3774, Polhill & Paulo 697, Bally
PRO 13260 & 13070, Robertson SA 4220 (EA): Vulnerable.
Dendrolobium umbellatum (L.) Benth. Habit: Shrub. Habitat: Sandy beaches, near
the sea. Vouchers: SAJIT–006104, Ngumbau V & Mwadime N V0511 (EA,
HIB), Napier 3294 in CM 6307 (EA).
Derris trifoliata Lour. Habit: Climber. Habitat: Mangrove swamps, coastal bushland, and
thicket, 0–30 m. Vouchers: Rawlins 886, Polhill & Paulo 686, Dale in FD 3844 (EA).
Desmodium adscendens (Sw.) DC. Habit: Herb. Habitat: Old cultivated fields, and
grassland, ca. 15 m. Vouchers: SAJIT–006215 (EA, HIB), Magogo & Glover 534,
Luke WRQ & Robertson SA 2692 (EA).
Desmodium barbatum var. dimorphum (Baker) B. G. Schub. Habit: Herb. Habitat:
Grassland, woodland, ca. 372 m. Vouchers: SAJIT–006163 (EA, HIB), Magogo
FC & Glover PE 1266 (EA).
Desmodium barbatum var. procumbens B. G. Schub. Habit: Herb. Habitat: Open
and wooded grassland, ca. 35–1800 m. Vouchers: Kassner 121, Magogo FC &
Glover PE 1 (EA).
Desmodium gangeticum (L.) DC. Habit: Herb. Habitat: Deciduous woodland, riverine forest, and swampy places, ca. 183 m. Vouchers: SAJIT–006174, Ngumbau
V & Mwadime N V0209 (EA, HIB), Thorold 1533, Kassner 391, Magogo FC &
Glover PE 385 (EA).
94
Veronicah M. Ngumbau et al. / PhytoKeys 147: 1–191 (2020)
Desmodium ramosissimum G. Don Habit: Herb. Habitat: Grassland or dry sandy
open areas, ca. 0–150 m. Vouchers: Magogo & Glover 27, Luke WRQ 3114 (EA).
Desmodium salicifolium var. densiflorum B. G. Schub. Habit: Herb. Habitat:
Grassland, ca. 80 m. Vouchers: SAJIT–006197 (EA, HIB), Nyange M 372 (EA).
Desmodium triflorum (L.) DC Habit: Herb. Habitat: Roadsides and lawns, ca.
0–100 m. Voucher: Luke WRQ 3120 (EA).
Desmodium velutinum (Willd.) DC. Habit: Herb. Habitat: Forest edges, deciduous woodland, wooded, and open grassland, ca. 156 m. Vouchers: Ngumbau V
& Mwadime N V0430 (EA, HIB), Drummond & Hemsley 3800, Rawlins 37,
Magogo FC & Glover PE 1074 (EA).
Dialium orientale Baker f. Habit: Tree. Habitat: Lowland dry evergreen forest and
coastal evergreen bushland, 0–60 m. Vouchers: SAJIT–006458 (EA, HIB), Elliot
in FH 1489 in CM 16435, Dale in FH 3898, J Adamson 342 in Bally 3836, Rawlins SP 339 (EA): Near Threatened.
Dichrostachys cinerea (L.) Wight & Arn. subsp. cinerea Habit: Shrub. Habitat: Deciduous bushland, scrub, wooded grassland, and deciduous woodland, ca. 80 m.
Vouchers: SAJIT–005588 (EA, HIB), Drummond & Hemsley 1178 (EA).
Dichrostachys cinerea subsp. keniensis Brenan & Brummitt Habit: Shrub. Habitat:
Bushland. Voucher: Magogo FC & Estes R 1237 (EA): Endemic.
Dolichos sericeus subsp. glabrescens Verdc. Habit: Climber. Habitat: Bushland.
Voucher: Nash LT 32 (EA).
Dolichos trilobus L. Habit: Climber. Habitat: Grassland, bushland, and forest, ca.
0–450 m. Vouchers: Ngumbau V & Mwadime N V0352 (EA, HIB), RM Graham
in FD 2225 & 1952, Jeffery 327, Magogo FC & Glover PE 1265 (EA).
Entada leptostachya Harms Habit: Liana. Habitat: Forest, ca. 200 m. Voucher: Robertson SA, Davis S & Morley J 7098 (EA)
Entada rheedii Spreng. Habit: Liana. Habitat: Lowland rainforest and evergreen
bushland, 0–600 m. Vouchers: Ngumbau V & Mwadime N V0490 (EA, HIB),
RM Graham NN757 in FH 2227, Magogo FC & Glover PE 626 (EA).
Eriosema glomeratum (Guill. & Perr.) Hook. f. Habit: Herb. Habitat: Grassland,
scrub on coral, and sandy soils, ca. 227 m. Vouchers: SAJIT–005938 & 006162,
Ngumbau V & Mwadime N V0285 (EA, HIB), Verdcourt 1885, Drummond &
Hemsley 1032, Rawlins, Magogo FC & Glover PE 390 (EA).
Eriosema parviflorum E. Mey. Habit: Herb. Habitat: Grassland, thicket, and swampy
grassland, ca. 222 m. Vouchers: SAJIT–005991, Ngumbau V & Mwadime N
V082 (EA, HIB), RM Graham in FD 1922, Kassner 444, Jeffery 252, Magogo
FC & Glover PE 338 (EA).
Eriosema pauciflorum Klotzsch var. pauciflorum Habit: Herb. Habitat: Seasonally burnt grasslands, Brachystegia woodland, coastal dry forest, and roadsides, ca.
40 m. Vouchers: Magogo & Glover 404, Nyange M 342 (EA).
Erythrina abyssinica DC. Habit: Tree. Habitat: Wooded grassland and deciduous
woodland, ca. 200 m. Voucher: Magogo FC & Glover PE 68 (EA).
Erythrina sacleuxii Hua Habit: Tree. Habitat: Bushland, open woodland, and evergreen coastal forest, 0–450 m. Vouchers: Ngumbau V & Mwadime N V0465
Checklist of coastal forests of Kenya
95
(EA, HIB), Greenway 9646, CW Elliot in FD 1376, Greenway & Rawlins 9361,
Cunningham-van Someren GR in FD 9816 (EA).
Erythrophleum suaveolens (Guill. & Perr.) Brenan Habit: Tree. Habitat: Riverine
and lowland rainforest, ca. 0–245 m. Vouchers: SAJIT–006053, Ngumbau V &
Mwadime N V0266 (EA, HIB), Verdcourt 2129, CW Elliot 1493, Luke WRQ &
Robertson SA 220 (EA).
Galactia argentifolia S. Moore Habit: Herb. Habitat: Grassland, 0–450 m. Vouchers: SAJIT–005958 (EA, HIB), Drummond & Hemsley 1137, Bogdan 3908,
Graham in FD 1977, Magogo FC & Glover PE 403 (EA).
Galactia striata var. villosa (Wight & Arn.) Verdc. Habit: Herb. Habitat: Grassland
and coastal bushland, 0–120 m. Voucher: Rawlins in EAH 11270 (EA).
Gigasiphon macrosiphon (Harms) Brenan Habit: Tree. Habitat: Lowland rainforest,
30–600 m. Vouchers: Ngumbau V & Mwadime N V0526 (EA, HIB), TAM Gardner 1, Verdcourt 2404, Verdcourt 3935, Robertson SA 7579 (EA): Endangered.
Hymenaea verrucosa Gaertn. Habit: Tree. Habitat: Lowland dry evergreen forest,
woodland, and coastal evergreen bushland, 15–240 m. Vouchers: SAJIT–007184
(EA, HIB), RM Graham B263 in FH Ox. 289 & in CM 13951, Trump 100,
Kimeu JM 522, Saufferer S 861, Luke WRQ & Robertson SA 220, RM Graham
B263 in FH Ox. 289 & in CM 1395 (EA).
Indigofera brachynema J. B. Gillett Habit: Herb. Habitat: Dry grassland and AcaciaCommiphora bushland, ca. 186 m. Vouchers: Ngumbau V & Mwadime N V0342
(EA, HIB), Greenway & Rawlins 9295, Luke Q 6142 (EA).
Indigofera charlierana Schinz Habit: Herb. Habitat: Sandy coastal grassland,
0–230 m. Vouchers: Whyte, Tweedie 950, Bogdan 2524, Luke Q 6141 (EA).
Indigofera colutea (Burm. f.) Merr. Habit: Herb. Habitat: Grassland, cultivated area,
and bushland, ca. 0–20 m. Vouchers: Tweedie 1003, Robertson SA 3328 (EA).
Indigofera congesta Baker Habit: Herb. Habitat: Grassland, seasonal swamps, and
old cultivation, ca. 136 m. Vouchers: SAJIT–005957 (EA, HIB), Whyte, Graham
RM 577 (EA).
Indigofera cuneata Oliv. Habit: Herb. Habitat: Scattered tree grassland and cultivation especially on sand, 0–365 m. Vouchers: Makin 429, Drummond & Hemsley
3861, Moomaw 1132, Gillett JB 20348 (EA).
Indigofera dendroides Jacq. Habit: Herb. Habitat: Grassland, cultivated ground, seasonal swamps, and flood plains, ca. 10 m. Vouchers: Rawlins 68 & 343, Magogo
FC & Glover PE 1138 (EA).
Indigofera dyeri Britten Habit: Herb. Habitat: Weed of cleared land and bushland,
ca. 20 m. Vouchers: Mainwaring in Napier 3040, Robertson SA 3298 (EA).
Indigofera hirsuta L. Habit: Herb. Habitat: Cultivated and waste areas, ca. 36 m.
Vouchers: SAJIT–006172 (EA, HIB), MacNaughton 97, Jeffery 35, Luke WRQ
& PA 5730 (EA).
Indigofera kirkii Oliv. Habit: Herb. Habitat: Between Mangroves and forest, ca. 2 m.
Voucher: Luke 3032 (EA).
Indigofera longimucronata Baker f. Habit: Herb. Habitat: Rocky outcrops, lowland
forests, and coconut groves, 0–350 m. Vouchers: Drummond & Hemsley 4178,
96
Veronicah M. Ngumbau et al. / PhytoKeys 147: 1–191 (2020)
Whyte A, Jeffery 233, Schrire BD, Robertson SA & Stirton CH 2573, Magogo
FC & Estes R 1208 (EA).
Indigofera longiracemosa Baill. Habit: Herb. Habitat: Coastal areas, 0–100 m.
Vouchers: Mearns 2169, Swynnerton 281, Gisau in EAH 11207, Luke Q 1491,
Luke WRQ & PA 5722A (EA).
Indigofera malindiensis J.B. Gillett Habit: Herb. Habitat: Wooded sand dunes
and on riverbanks, 0–450 m. Vouchers: Rawlins 941, Hacker 87, Rawlins SP
610 (EA).
Indigofera microcarpa Desv. Habit: Herb. Habitat: Cultivated area, on the shores of
receding lakes and rivers, 0–200 m. Vouchers: Jeffery 457, Rawlins 365, Luke Q
6120 (EA).
Indigofera paniculata Vahl ex Pers. subsp. paniculata Habit: Herb. Habitat: Grassland, dried up swamps, and old cultivation, ca. 0–372 m. Vouchers: SAJIT–006165
& 006171 (EA, HIB), Drummond & Hemsley 3965, Whyte, Mearns 2164, Magogo FC & Glover PE 290 (EA).
Indigofera schimperi var. baukeana (Vatke) J. B. Gillett Habit: Herb. Habitat:
Grassland, ca. 100 m. Voucher: Drummond & Hemsley 3744 (EA).
Indigofera spicata Forssk. Habit: Herb. Habitat: Common in disturbed grassland,
cultivated areas, and waste place, 0–200 m. Voucher: Thorold in FD 1577 (EA).
Indigofera strobilifera (Hochst.) Baker subsp. strobilifera Habit: Herb. Habitat: Deciduous woodland, bushland, and waste places, ca. 36 m. Vouchers: SAJIT–006170 (EA, HIB), Rawlins 374 (EA).
Indigofera strobilifera subsp. lanuginosa (Baker f.) J. B. Gillett Habit: Herb. Habitat: Sandy grassland near the coast, 0–15 m. Vouchers: Hildebrandt 927/b, Bogdan 2572, Festo L & Luke Q 2464 (EA).
Indigofera tanganyikensis var. paucijuga J. B. Gillett Habit: Herb. Habitat: Grassland, ca. 10–350 m. Vouchers: Greenway PJ & Rawlins SP 8921, Drummond RB
& Hemsley JH 4056 (EA).
Indigofera tinctoria L. var. tinctoria Habit: Herb. Habitat: Bush margins, cultivation, and secondary growth, ca. 100 m. Voucher: Peter 45187 (EA).
Indigofera trita var. subulata (Poir.) Ali Habit: Herb. Habitat: Secondary growth,
ca. 0–237 m. Vouchers: Ngumbau V & Mwadime N V0189 (EA, HIB), Tweedie
988, Magogo FC & Glover PE 685 (EA).
Indigofera vohemarensis Baill. Habit: Herb. Habitat: Grassy and stony slopes, ca.
20 m. Vouchers: Napier 3310, Robertson SA 3482 & 3296 (EA).
Indigofera wituensis Baker f. var. wituensis Habit: Herb. Habitat: Grassland,
0–300 m. Vouchers: A Whyte, Polhill & Paulo 889, Rawlins 73 (EA).
Indigofera zanzibarica J. B. Gillett Habit: Herb. Habitat: Damp open places, often on sand, 0–100 m. Vouchers: SAJIT–006189 (EA, HIB), Rawlins 59 & 193,
Faden AJ & RB 1128 (EA).
Indigofera zenkeri Baker f. Habit: Herb. Habitat: Woodland, grassland and, cultivated area, ca. 0–300 m. Vouchers: Verdcourt 1879, OM Mwangangi 1315 (EA).
Julbernardia magnistipulata (Harms) Troupin Habit: Tree. Habitat: Lowland rainforest, riverine forest, and coastal evergreen bushland, ca. 0–206 m. Vouchers:
Checklist of coastal forests of Kenya
97
Ngumbau V & Mwadime N V0501 (EA, HIB), Drummond & Hemsley 4039,
Sulemani in FH 3236, Polhill & Paulo 803, Robertson SA 6992 (EA).
Lablab purpureus subsp. uncinatus Verdc. Habit: Climber. Habitat: Variety of habitats. Voucher: Polhill RM & Paulo S 699 (EA).
Leucaena leucocephala (Lam.) de Wit Habit: Tree. Habitat: A cultivated plant, ca.
50 m. Voucher: Bally 6363 (EA): Cultivated.
Lysiloma latisiliquum (L.) Benth. Habit: Tree. Habitat: Along the lake shores, ca.
300 m. Voucher: Magogo FC & Glover PE 132 (EA): Exotic.
Macrotyloma axillare var. glabrum (E. Mey.) Verdc. Habit: Climber. Habitat: Mixed
woodland, grassland, often near rivers, ca. 266 m. Vouchers: SAJIT–005940, Ngumbau V & Mwadime N V0292 & 050 (EA, HIB), Magogo FC & Glover PE 364 (EA).
Macrotyloma uniflorum var. benadirianum (Chiov.) Verdc. Habit: Climber. Habitat: Sand dunes, thin soil on coral rag, sea level. Vouchers: Greenway 9268, Rawlins 4 & 41 (EA).
Macrotyloma uniflorum var. verrucosum Verdc. Habit: Climber. Habitat: Grassland
with scattered Hyphaene, Sterculia, dry Acacia, Commiphora thickets, 15–540 m.
Vouchers: Drummond & Hemsley 1191, Tweedie 1200, Polhill & Paulo 886,
Magogo FC & Glover PE 200 (EA).
Microcharis garissaensis (J.B. Gillett) Schrire Habit: Herb. Habitat: Acacia-Commiphora open bushland, 100–700 m. Vouchers: Gillett 16530 & 16445 (EA).
Microcharis latifolia Benth. Habit: Herb. Habitat: Seasonally damp grassy places,
0–500 m. Vouchers: Rawlins 74, Festo L & Luke Q 2574 (EA).
Microcharis microcharoides var. latistipulata (J.B. Gillett) Schrire Habit: Herb.
Habitat: Forest, grassland, and savannah, ca. 100 m. Vouchers: Adamson T 53,
Drummond RB & Hemsley JH 3784 (EA).
Millettia lasiantha Dunn Habit: Liana. Habitat: Coastal and riverine evergreen forest, 10–300 m. Vouchers: Ngumbau V & Mwadime N V0392 & V062 (EA,
HIB), Drummond & Hemsley 1122, RM Graham in FD 1801, Festo L, Luke Q
& P 2775, Luke WRQ 3201 (EA).
Millettia usaramensis Taub. Habit: Tree. Habitat: Wooded grassland and margins of
lowland forest, 10–700 m. Vouchers: SAJIT–005481, Ngumbau V & Mwadime
N V0531 (EA, HIB), Wakefield, RM Graham in FD 1779, Drummond & Hemsley 3885, Robertson SA & Luke WRQ 5188 (EA).
Mimosa pigra L. Habit: Herb. Habitat: Swamps, especially along rivers and lake shores,
ca. 15 m. Vouchers: SAJIT–006229 (EA, HIB), Jeffery K165, Luke WRQ 3148 (EA).
Mimosa pudica var. unijuga (Duchass. & Walp.) Griseb. Habit: Herb. Habitat:
Probably an introduced weed, ca. 200 m. Vouchers: Jeffery K228, Magogo FC &
Glover PE 662 (EA): Naturalized.
Mucuna gigantea subsp. quadrialata (Baker) Verdc. Habit: Liana. Habitat: Bushland and forest edges, ca. 40 m. Vouchers: RM Graham in FD 1937, Bally 1883,
Nyange 580 (EA).
Mucuna pruriens (L.) DC. var. pruriens Habit: Liana. Habitat: Course grassland, bushland, riverine forest, forest edges, and abandoned cultivations, 0–700 m. Vouchers:
Verdcourt 1873, RM Graham in FD 1960, Tweedie 1038, Bogdan 255A (EA).
98
Veronicah M. Ngumbau et al. / PhytoKeys 147: 1–191 (2020)
Mundulea sericea (Willd.) A. Chev. Habit: Shrub. Habitat: Lowland evergreen forest and deciduous bushland, ca. 10 m. Vouchers: Drummond & Hemsley 4208,
Simpson BL 160, Shimba Hills Survey Unit 19 (EA).
Neptunia oleracea Lour. Habit: Herb. Habitat: Freshwater of pools, lakes, and
swamps, ca. 30 m. Vouchers: Mrs J Adamson 434 in Bally 6133, Leauthaud C 11,
TPR781 (EA).
Newtonia erlangeri (Harms) Brenan Habit: Tree. Habitat: Deciduous woodland,
Euphorbia bushland, and thickets, ca. 35 m. Vouchers: Power in EAH 10971,
Greenway & Rawlins 9435, Luke Q 1473 (EA): Endangered.
Newtonia paucijuga (Harms) Brenan Habit: Tree. Habitat: Lowland rainforest and riverine forest, 75–300 m. Vouchers: Battiscombe 93, Webber 607 in CM 16977, Moggridge 384, Hawthorne W 483, Luke WRQ & Robertson SA 215 (EA): Vulnerable.
Ophrestia hedysaroides (Willd.) Verdc. Habit: Liana. Habitat: Bushland, forest edges, grassland, and old cultivations, ca. 100 m. Vouchers: Verdcourt 1903, Kirk,
RM Graham in FD 1914, Luke WRQ et al. 4334 (EA).
Ormocarpum kirkii S. Moore Habit: Shrub. Habitat: Acacia-Commiphora or coastal
bushland, grassland, ca. 20–153 m. Vouchers: Ngumbau V & Mwadime N V0509
(EA, HIB), Rawlins 273, Robertson SA 4226 (EA).
Ormocarpum sennoides subsp. zanzibaricum Brenan & J.B. Gillett Habit: Shrub.
Habitat: Scrub on coral rock near shore, woodland, and shady places by streams,
0–600 m. Vouchers: SAJIT–006024 & 005566, Ngumbau V & Mwadime N
V071 (EA, HIB), CG van Someren Sh127, RM Graham B 488 in FD 1980, Rawlins 710, Beentje 3738 (EA).
Paramacrolobium coeruleum (Taub.) J. Léonard Habit: Tree. Habitat: Lowland
rainforest, 8–360 m. Vouchers: Ngumbau V & Mwadime N V0362 (EA, HIB),
Dale 3628, Greenway 9804, Pakia M 974 (EA).
Parkia filicoidea Oliv. Habit: Tree. Habitat: Lowland rainforest and riverine forest, ca. 250 m. Vouchers: Hildebrandt 1975, Battiscombe 1 & Cunningham-van
Someren GR Sh 56 (EA).
Parkinsonia aculeata L. Habit: Tree. Habitat: Semi-dry vegetation, ca. 100 m.
Voucher: SAJIT–005423 (EA, HIB): Cultivated.
Parkinsonia anacantha Brenan Habit: Shrub. Habitat: Semi-desert scrub with Acacia mellifera, ca. 300 m. Voucher: LC Edwards 168 (EA).
Philenoptera bussei (Harms) Schrire Habit: Tree. Habitat: In riverine vegetation and
mixed deciduous woodland, ca. 0–200 m. Vouchers: Ngumbau V & Mwadime N
V0506 (EA, HIB), Magogo FC & Glover PE 656, Luke Q 1510 (EA).
Piliostigma thonningii (Schumach.) Milne-Redh. Habit: Tree. Habitat: Woodland,
wooded grassland, and bushland, 5–227 m. Vouchers: SAJIT–006105 & 005451,
Ngumbau V & Mwadime N V0280 (EA, HIB), Magogo FC & Glover PE 652 (EA).
Pithecellobium dulce (Roxb.) Benth. Habit: Tree. Habitat: Coastal thickets, 0–500 m.
Voucher: SAJIT–006121 (EA, HIB): Cultivated.
Prioria msoo (Harms) Breteler Habit: Tree. Habitat: Ground water forest, ca. 95 m.
Vouchers: Ngumbau V & Mwadime N V0484 (EA, HIB), Katende AB 1759
(EA): Vulnerable.
Checklist of coastal forests of Kenya
99
Prosopis juliflora (Sw.) DC. Habit: Shrub. Habitat: Acacia-Commiphora dry bushland, ca. 100 m. Vouchers: Johnsson S et al. 154, E Mwangi 38 (EA): Cultivated.
Platycelyphium voense (Engl.) Wild Habit: Tree. Habitat: Scattered in Acacia-Commiphora deciduous bushland, ca. 80 m. Voucher: Drummond & Hemsley 4132 (EA).
Pseudarthria hookeri Wight & Arn. Habit: Herb. Habitat: Woodland, ca. 227 m.
Vouchers: Ngumbau V & Mwadime N V0284 (EA, HIB), Magogo FC & Glover
PE 1115 (EA).
Pseudoeriosema borianii subsp. longipedunculatum Verdc. Habit: Herb. Habitat:
Bushland, grassland, and seasonal swampy grasslands, 45–300 m. Voucher: Rawlins 838 (EA).
Pseudoprosopis euryphylla subsp. puguensis Brenan Habit: Liana. Habitat: Forest,
ca. 30–120 m. Vouchers: Luke WQR 3466, Robertson 6232 & 5808, Nyange M
& Luke Q 0591 (EA).
Pseudovigna argentea (Willd.) Verdc. Habit: Climber. Habitat: Grassland, 0–45 m.
Vouchers: Ngumbau V & Mwadime N V0242 (EA, HIB), Irwin 257, RM Graham in FD 1907, Jeffery 241, Magogo FC & Glover PE 883 (EA).
Rhynchosia albissima Gand. Habit: Herb. Habitat: Grassland, 0–200 m. Voucher:
Drummond & Hemsley 3763 (EA).
Rhynchosia congensis subsp. orientalis Verdc. Habit: Climber. Habitat: Coastal
grassland and bushland, 0–45(–780) m. Vouchers: SAJIT–006108 (EA, HIB),
RM Graham in FD 1595, Tweedie 1051, Graham MD 2111 (EA).
Rhynchosia densiflora (Roth) DC. Habit: Climber. Habitat: Forest, 20–75 m.
Voucher: Ngumbau V & Mwadime N V0125 (EA, HIB).
Rhynchosia hirta (Andrews) Meikle & Verdc. Habit: Climber. Habitat: Grassland,
scrub, woodland, and evergreen forest margins, ca. 280 m. Vouchers: Ngumbau V
& Mwadime N V0254 (EA, HIB), Magogo FC & Glover PE 446 (EA).
Rhynchosia micrantha Harms Habit: Herb. Habitat: Grassland, bushland, and abandoned cultivations, ca. 5 m. Vouchers: SAJIT–006101 (EA, HIB), Bogdan 4731,
Bogdan 2600, Thorold 1578 (EA).
Rhynchosia minima (L.) DC. var. minima Habit: Climber. Habitat: Bushland, Acacia scrub, and grassland particularly in coastal areas, ca. 0–200 m. Vouchers: Symes
165, Rawlins 887 & 76, Luke WRQ & PA 6018 (EA).
Rhynchosia minima var. nuda (DC.) Kuntze Habit: Climber. Habitat: Grassland,
bushland, and dry Acacia scrub, ca. 350 m. Vouchers: Drummond & Hemsley
4136, Magogo FC & Glover PE 992 (EA).
Rhynchosia sublobata (Schumach.) Meikle Habit: Climber. Habitat: Forest, 10–
600 m. Vouchers: Ngumbau V & Mwadime N V0208 (EA, HIB), Nyange M
326 (EA).
Rhynchosia velutina var. discolor (Baker) Verdc. Habit: Climber. Habitat: Bushland
and Brachystegia woodland, 0–30 m. Vouchers: Rawlins 860, Jeffery 843 (EA).
Rhynchosia velutina Wight & Arn. var. velutina Habit: Climber. Habitat: Bushland near coast, sandy beaches and dunes, coral rocks, 0–30 (–360) m. Vouchers:
SAJIT–006109 (EA, HIB), Drummond & Hemsley 1021, Rawlins 876, Bogdan
4695, Luke Q 1443, Magogo FC & Glover PE 867 (EA).
100
Veronicah M. Ngumbau et al. / PhytoKeys 147: 1–191 (2020)
Rhynchosia viscosa (Roth) DC. var. viscosa Habit: Climber. Habitat: Coarse grassland and marshy places, near sea level. Vouchers: Ngumbau V & Mwadime N
V0244 (EA, HIB), Thorold 1553, Rawlins 884, Whyte (EA).
Rhynchosia viscosa var. breviracemosa (Hauman) Verdc. Habit: Climber. Habitat:
Forest edges, bushland, grassland, and cultivated land, 0–200 m. Vouchers: RM
Graham in FD 2027, Tweedie 2205, Jeffery 236, Kirika P, Mbale M & Mbatha
M 771 (EA).
Rothia hirsuta (Guill. & Perr.) Baker Habit: Herb. Habitat: Cultivated ground and
disturbed places, ca. 30 m. Vouchers: Bogdan 2624, Festo L, Luke Q & P 2684,
Robertson SA 3312 (EA).
Samanea saman (Jacq.) Merr. Habit: Tree. Habitat: Forest. Voucher: Robertson SA
& Luke Q 5425 (EA): Naturalized.
Scorodophloeus fischeri (Taub.) J. Léonard Habit: Tree. Habitat: Lowland dry evergreen forest and riverine forest, 30–670 m. Vouchers: Wakefield, Luke WRQ &
Robertson SA 214 (EA).
Senegalia adenocalyx (Brenan & Exell) Kyal. & Boatwr. Habit: Shrub. Habitat:
Bushland and secondary thickets, 45–450 m. Vouchers: Jeffery K311 & Battiscombe 781, Jeffery K495, Kuchar P 13518 (EA).
Senegalia brevispica (Harms) Seigler & Ebinger Habit: Shrub. Habitat: Grassland,
ca. 50–400 m. Voucher: Gerhardt K & Steiner M 78 (EA).
Senegalia goetzei (Harms) Kyal. & Boatwr. Habit: Tree. Habitat: Wooded grassland
and woodland, ca. 18 m. Voucher: Luke WRQ & Robertson SA 5546 (EA).
Senegalia mellifera (Vahl) Seigler & Ebinger subsp. mellifera Habit: Tree. Habitat:
Dry scrub and deciduous bushland, ca. 300 m. Vouchers: Jeffery K490, Robertson
SA 4235, Gillett JB 20351 (EA).
Senegalia mellifera subsp. detinens (Burch.) Kyal. & Boatwr. Habit: Tree. Habitat:
Deciduous bushland and woodland, ca. 300 m. Voucher: Geffery GM 362 (EA).
Senegalia pentagona (Schumach.) Kyal. & Boatwr. Habit: Liana. Habitat: Evergreen forest, ca. 400 m. Vouchers: Verdcourt B 2122 (EA), Luke WRQ & Robertson SA sr.
Senegalia polyacantha subsp. campylacantha (Hochst. ex. A. Rich.) Kyal. & Boatwr. Habit: Tree. Habitat: In wooded grassland and woodland, ca. 213 m. Voucher:
Unknown 69 (EA).
Senegalia rovumae (Oliv.) Kyal. & Boatwr. Habit: Tree. Habitat: Riverine forest and
saline water swamp forest, 6–700 m. Vouchers: Greenway & Rawlins 9432, Luke
WRQ & Robertson SA 1453 (EA).
Senegalia senegal (L.) Britton var. senegal Habit: Tree. Habitat: Wooded grassland,
ca. 120 m. Vouchers: Elliott CW 1373, Verdcourt B 1916 (EA).
Senegalia senegal var. kerensis (Schweinf.) Kyal. & Boatwr. Habit: Tree. Habitat:
Wooded grassland, ca. 243 m. Voucher: King JM 15 (EA).
Senna alexandrina var. obtusata (Brenan) Lock Habit: Shrub. Habitat: Bush on
sand, ca. 10 m. Voucher: Jeffery 329 (EA).
Senna longiracemosa (Vatke) Lock Habit: Shrub. Habitat: Dry scrub, deciduous bushland, and semi desert scrub, ca. 300 m. Voucher: Drummond & Hemsley 4071 (EA).
Checklist of coastal forests of Kenya
101
Senna obtusifolia (L.) H.S. Irwin & Barneby Habit: Herb. Habitat: Grassland, roadsides, and on waste ground, ca. 100 m. Vouchers: RM Graham DD 774 in FH
2285 & in CM 1362 (EA).
Senna occidentalis (L.) Link Habit: Herb. Habitat: Waste places and disturbed area,
ca. 100 m. Voucher: Luke WRQ & PA 5729 (EA).
Senna siamea (Lam.) H. S. Irwin & Barneby Habit: Shrubby tree. Habitat: forest,
ca. 50 m. Vouchers: Perdue & Kibuwa 10168, P Kuchar 13401 (EA): Cultivated.
Senna singueana (Delile) Lock Habit: Shrub. Habitat: Woodland, wooded grassland, and bushland, ca. 150 m. Vouchers: Drummond & Hemsley 1149, Klotzli F
422, Magogo FC & Glover PE 268 (EA).
Senna timoriensis (DC.) H.S. Irwin & Barneby Habit: Shrub. Habitat: Open forest
and lowland, ca. 50 m. Voucher: Mrs Robertson SA 7768 (EA).
Sesbania bispinosa (Jacq.) W. Wight Habit: Herb. Habitat: Marshes, ditches, a weed
in cultivation area and in rice fields, 0–300 m. Vouchers: Hildebrandt 1990, RM
Graham in FD 2107, Rawlins 305, Kirika P 204 (EA).
Sesbania sericea (Willd.) Link Habit: Herb. Habitat: Marshy ground and old rice
fields, 0–850 m. Vouchers: Jeffery 486, Sampson 60 (EA).
Sesbania speciosa Taub. Habit: Herb. Habitat: Swamps, ditches, and riverbanks, ca.
100 m. Vouchers: Sampson 56, Gardner in FD 1368 (EA): Vulnerable.
Sophora inhambanensis Klotzsch Habit: Shrub. Habitat: Sandy foreshores, ca. 10 m.
Vouchers: RM Graham 274, CW Elliot in FD 1375, Greenway & Rawlins 8917 (EA).
Sophora tomentosa L. Habit: Shrub. Habitat: Sandy foreshores. Vouchers: Thorold
1550, Napier in CM 6305 (EA).
Spathionema kilimandscharicum Taub. Habit: Climber. Habitat: Bushland, ca. 270
m. Vouchers: Ngumbau V & Mwadime N V0442 (EA, HIB), Luke WRQ &
Robertson SA 2192 (EA).
Sphenostylis stenocarpa (A. Rich.) Harms Habit: Climber. Habitat: Grassland,
wooded grassland, and woodland, ca. 183 m. Vouchers: Ngumbau V & Mwadime
N V0205 (EA, HIB), R RM Graham in FD 1936 (EA).
Stylosanthes erecta P. Beauv. Habit: Herb. Habitat: Sandy places near the coast usually just above loose dunes, sea level. Vouchers: D Davies 78, Bogdan 2525, Greenway 9492 (EA).
Stylosanthes fruticosa (Retz.) Alston Habit: Herb. Habitat: Grassland and bushland,
ca. 137 m. Vouchers: Drummond & Hemsley 3942, Magogo FC & Glover PE
95, Kirika P 941 (EA).
Tamarindus indica L. Habit: Tree. Habitat: Woodland, wooded grassland and deciduous bushland, ca. 297 m. Vouchers: Ngumbau V & Mwadime N V0155 (EA,
HIB), Bogdan 2556, Magogo FC & Glover PE 884 (EA): Naturalized.
Tephrosia linearis (Willd.) Pers. Habit: Herb. Habitat: Grassland, rocky slopes, cultivation, and fallows, ca. 186 m. Vouchers: Ngumbau V & Mwadime N V0344
(EA, HIB), Patterson 40, Luke WRQ 3311 (EA).
Tephrosia noctiflora Baker Habit: Herb. Habitat: Grassland, cultivation, and thickets, ca. 213 m. Vouchers: Ngumbau V & Mwadime N V0329 (EA, HIB), Jeffery
271, Thorold 1556, Luke WRQ & PA 5722B (EA).
102
Veronicah M. Ngumbau et al. / PhytoKeys 147: 1–191 (2020)
Tephrosia pentaphylla (Roxb.) G. Don Habit: Herb. Habitat: Black cotton soil and
gravelly hillsides, 0–300 m. Vouchers: Wakefield, Rawlins 733, Magogo FC &
Glover PE 988 (EA).
Tephrosia pumila (Lam.) Pers. var. pumila Habit: Herb. Habitat: Short grassland,
Acacia-Commiphora bushland, and in cultivated land, ca. 30 m. Vouchers: Tweedie
1030, Magogo FC & Glover PE 931 (EA).
Tephrosia pumila var. aldabrensis (J.R. Drumm. & Hemsl.) Bosman & A.J.P. de
Haas Habit: Herb. Habitat: Bushland and grassland, near the sea. Vouchers: Verdcourt 2138, Greenway 10457, Rawlins 17 (EA).
Tephrosia purpurea subsp. dunensis Brummitt Habit: Herb. Habitat: Sandy sea and
lake shores. Vouchers: Hacker 77, Greenway & Rawlins 8923 & 8914 (EA).
Tephrosia purpurea subsp. leptostachya (DC.) Brummitt Habit: Herb. Habitat:
Grassland and plantations, ca. 297 m. Vouchers: SAJIT–004653, Ngumbau V &
Mwadime N V0148 (EA, HIB), Thorold 1534, RM Graham in FD 1789, Jeffery
459, Magogo FC & Glover PE 889 (EA).
Tephrosia stormsii De Wild. var. stormsii Habit: Herb. Habitat: Grassland, cultivated land, and deciduous woodland, ca. 0–100 m. Vouchers: Butler 25, MacNaughton 53, Thorold 1579, Luke WRQ et al. 6194 (EA).
Tephrosia villosa subsp. ehrenbergiana (Schweinf.) Brummitt Habit: Herb. Habitat: Acacia-Commiphora bushland, ca. 181 m. Vouchers: Ngumbau V & Mwadime
N V0227 (EA, HIB), Napier 3263 (EA).
Tephrosia vogelii Hook.f. Habit: Herb. Habitat: Grassland, forest margins, waste
ground, and old cultivation, ca. 50 m. Voucher: Donald in FD 2369 (EA).
Teramnus labialis subsp. arabicus Verdc. Habit: Climber. Habitat: Grassland and
bushland, ca. 30 m. Vouchers: Whyte, Agnew ADQ 9763 (EA).
Teramnus repens subsp. gracilis (Chiov.) Verdc. Habit: Herb. Habitat: Bushland,
0–50 m. Vouchers: Thorold 1574, Tweedie 1015, Greenway & Rawlins 9355 (EA).
Tetrapleura tetraptera (Schum. & Thonn.) Taub. Habit: Tree. Habitat: Lowland
rainforest, ca. 80 m. Voucher: Drummond & Hemsley 3960 (EA).
Vachellia bussei (Harms ex Y. Sjöstedt) Kyal. & Boatwr. Habit: Tree. Habitat: Scrubland, bushland, ca. 50–400 m. Vouchers: Bally PRO 8553, Robertson SA 6402 (EA).
Vachellia etbaica subsp. platycarpa (Brenan) Kyal. & Boatwr. Habit: Tree. Habitat:
Deciduous bushland, dry scrub, and grassland, ca. 350 m. Vouchers: Robertson SA
4120, Drummond & Hemsley 4114 (EA).
Vachellia etbaica subsp. uncinata (Brenan) Kyal. & Boatwr. Habit: Tree. Habitat: Wooded grassland, ca. 60 m. Vouchers: Hornby 3108, Graham RM 226,
Moomaw JC 1412 (EA).
Vachellia nilotica subsp. indica (Benth.) Kyal. & Boatwr. Habit: Tree. Habitat:
Cultivated area, ca. 5 m. Voucher: Mrs Robertson SA 7744 (EA).
Vachellia nilotica subsp. leiocarpa (Brenan) Kyal. & Boatwr. Habit: Tree. Habitat:
Grassland near the coast and woodland, ca. 100 m. Vouchers: Bogdan 2612, Dale
3832, Mrs J Adamson 437 in Bally 6136 (EA).
Checklist of coastal forests of Kenya
103
Vachellia nilotica subsp. subalata (Vatke) Kyal. & Boatwr. Habit: Tree. Habitat:
Wooded grassland and deciduous bushland, ca. 15 m. Vouchers: Magogo FC &
Glover PE 863, Robertson SA 4968 (EA).
Vachellia robusta subsp. usambarensis (Taub.) Kyal. & Boatwr. Habit: Tree. Habitat: Woodland, 10–250 m. Voucher: Robertson SA 4379 (EA).
Vachellia sieberiana (DC.) Kyal. & Boatwr. var. sieberiana Habit: Tree. Habitat:
Deciduous woodland, wooded grassland, and riverine forest, ca. 153 m. Vouchers: Ngumbau V & Mwadime N V0510 (EA, HIB), RM Graham 2281, Luke
WRQ 3158 (EA).
Vachellia stuhlmannii (Taub.) Kyal. & Boatwr. Habit: Shrub. Habitat: Woodland
and grassland, 100–750 m. Voucher: Robertson SA & Luke Q 5624 (EA).
Vachellia tortilis subsp. raddiana (Savi) Kyal. & Boatw Habit: Tree. Habitat: Bushland, 50–100 m. Vouchers: RM Graham W309 in FH 1800, Greenway 9444 (EA).
Vachellia tortilis subsp. spirocarpa (Hochst. ex. A. Rich.) Kyal. & Boatwr. Habit:
Tree. Habitat: Deciduous woodland, ca. 600 m. Voucher: Kuchar P 13716 (EA).
Vachellia zanzibarica (S. Moore) Kyal. & Boatwr. Habit: Tree. Habitat: Woodland
and wooded grassland, 0–460 m. Vouchers: Bogdan 3303, RM Graham L119 in
FH 1620 in CM 13991, Sangai GRW 15609 (EA).
Vigna friesiorum var. angustifolia Verdc. Habit: Herb. Habitat: Grassland, deciduous bushland, and Combretum woodland, ca. 480 m. Vouchers: Tweedie 1226,
Kassner 424 (EA).
Vigna hosei (Craib) Backer Habit: Climber. Habitat: Grass at roadsides and wastelands, 0–200 m. Vouchers: SAJIT–006203 (EA, HIB), Whyte, Magogo FC &
Estes R 1159 (EA).
Vigna macrorhyncha (Harms) Milne-Redh. Habit: Climber. Habitat: Grassland,
bushland, woodland, and forest, ca. 30 m. Vouchers: Napier 3363 in CM 6308,
Meester V 92, Robertson SA 3356 (EA).
Vigna membranacea subsp. caesia (Chiov.) Verdc. Habit: Climber. Habitat: AcaciaCommiphora bushland, coastal mixed bushland, ca. 0–400 m. Vouchers: Polhill &
Paulo 718, Jeffery GW 407, Gillett JB 16870 (EA).
Vigna membranacea subsp. hapalantha (Harms) Verdc. Habit: Climber. Habitat:
Coastal grassland in thicket, ca. 15 m. Vouchers: Jex-Blake in Napier 2276, Polhill
& Paulo 877, Tweedie 940, Gillet JB 20324 (EA): Endemic.
Vigna reticulata Hook.f. Habit: Climber. Habitat: Grassland, bushland, usually on
damp or swampy ground, ca. 0–700 m. Vouchers: RM Graham in FD 1905,
Tweedie 1061, Jeffery GW 297 (EA).
Vigna unguiculata subsp. dekindtiana (Harms) Verdc. Habit: Climber. Habitat:
Grassland, swamp, riverine forest, and dry scrub, ca. 0–300 m. Vouchers: Napier
3366 in CM 6309, RM Graham in FD 2101, Power 10969 (EA).
Vigna unguiculata subsp. pubescens (R. Wilczek) Pasquet Habit: Climber. Habitat: Bushland, grassland, and grassland with scattered trees, ca. 50 m. Vouchers:
Tweedie 2223, Jeffery 57, Sampson 52 (EA).
104
Veronicah M. Ngumbau et al. / PhytoKeys 147: 1–191 (2020)
Vigna unguiculata var. spontanea (Schweinf.) Pasquet Habit: Climber. Habitat:
Bushland, 100 m. Voucher: Magogo FC & Glover PE 471 (EA).
Vigna vexillata (L.) A. Rich. var. vexillata Habit: Climber. Habitat: Grassland, grassland with scattered trees, and thicket, ca. 70 m. Vouchers: Brathay 1982 Expedn
35, Soest LV 193 (EA).
Vigna vexillata var. angustifolia (Schumach. & Thonn.) Baker Habit: Climber.
Habitat: Grassland, ca. 500 m. Voucher: Magogo FC & Glover PE 994 (EA).
Wajira praecox (Verdc.) Thulin & Lavin Habit: Climber. Habitat: Dry Acacia, Terminalia-Commiphora bushland and semi-desert associations, ca. 375 m. Vouchers:
Greenway 9821, Luke WRQ & Robertson SA 2536 (EA).
Xeroderris stuhlmannii (Taub.) Mendonça & E. P. Sousa Habit: Tree. Habitat: Deciduous woodland and bushland, ca. 100 m. Vouchers: Kassner 158, Dale in FD 3843 (EA).
Zornia apiculata Milne-Redh. Habit: Herb. Habitat: Forest in dry scrub, grasslands,
and rocky places, ca. 20 m. Vouchers: Jeffery 231, Festo 2547 (EA).
Zornia capensis subsp. tropica Milne-Redh. Habit: Herb. Habitat: Dry scrub and
cultivated ground, ca. 0–350 m. Vouchers: Drummond & Hemsley 4145, Rawlins
in EAH 12068 (EA).
Zornia glochidiata Rchb. ex DC. Habit: Herb. Habitat: Grassland, rocky places,
often cultivated ground or roadsides, ca. 50 m. Voucher: Jeffery 845 (EA).
Zornia pratensis Milne-Redh. subsp. pratensis Habit: Herb. Habitat: Grassland on
hillsides, ca. 400 m. Voucher: Drummond & Hemsley 1113 (EA).
F81. Flagellariaceae
1 Genus, 1 Species
Flagellaria guineensis Schumach. Habit: Climber. Habitat: Forest fringes and thicket, 0–400 m. Vouchers: SAJIT–005932, Ngumbau V & Mwadime N V0247 (EA,
HIB), Magogo FC & Glover PE 128 (EA).
F82. Francoaceae
1 Genus, 1 Species
Bersama abyssinica Fresen. subsp. abyssinica Habit: Tree. Habitat: Lowland bushland, savannah, and gallery forests, ca. 246 m. Voucher: Luke WRQ & Robertson
SA 2724 (EA).
F83. Gelsemiaceae
1 Genus, 2 Species
Mostuea brunonis Didr. Habit: Shrub. Habitat: Lowland rainforest and riverine forest, ca. 86 m. Vouchers: SAJIT–006075, Ngumbau V & Mwadime N V0468
Checklist of coastal forests of Kenya
105
(EA, HIB), Luke WRQ 889, Luke Q 1554, Drummond & Hemsley 3805, RM
Graham & Dale 3809 (EA).
Mostuea microphylla Gilg Habit: Shrub. Habitat: Coastal evergreen bushland and
lowland rainforest, 0–350 m. Vouchers: Ngumbau V & Mwadime N V0493 (EA,
HIB), WE Taylor & Bally 5824, Luke Q 1496 (EA).
F84. Gentianaceae
4 Genera, 5 Species
Canscora alata (Roth) Wall. Habit: Herb. Habitat: Streamside on wet mud, 0–600 m.
Voucher: Luke WRQ 3316 (EA).
Enicostema axillare (Poir. ex Lam.) A. Raynal subsp. axillare Habit: Herb. Habitat: Moist or swampy sites in grassland, lake shores, and sparse woodland, ca. 200
m. Vouchers: Magogo FC & Glover PE 1049, Greenway & Rawlins 9453, Luke
6358 (EA).
Enicostema axillare subsp. latilobum (N. E. Br.) A. Raynal Habit: Herb. Habitat:
Grassland, 0–550 m. Vouchers: Greenway 4968, Magogo & Glover 1049 (EA).
Exacum oldenlandioides (S. Moore) Klack. Habit: Herb. Habitat: Moist sites grassland on river banks, ca. 54 m. Vouchers: Ngumbau V & Mwadime N V0554 (EA,
HIB), Magogo FC & Glover PE 357 (EA).
Exochaenium platypterum (Baker) Schinz Habit: Herb. Habitat: Damp sites woodland or grassland, ca. 300 m. Voucher: Luke WRQ et al. 7455 (EA).
Exochaenium pumilum (Baker) A. W. Hill Habit: Herb. Habitat: Damp and often
sandy sites in grassland or woodland, ca. 450 m. Voucher: Luke & Luke 5996 (EA).
F85. Gesneriaceae
1 Genus, 2 Species
Streptocarpus ionantha subsp. rupicola (B. L. Burtt) Christenh. Habit: Herb. Habitat: Rocky moist areas, ca. 200 m. Vouchers: Ngumbau V & Mwadime N V0408
(EA, HIB), Mabberley 718, Faden & Beentje 85/30 (EA): Critically Endangered.
Streptocarpus pallidiflorus C.B. Clarke Habit: Herb. Habitat: Rock crevices or on
banks in forest or unshaded areas, rarely epiphytic, 750 m. Vouchers: Luke &
Robertson 20152, Luke 4705 (EA).
F86. Gisekiaceae
1 Genus, 2 Species
Gisekia africana (Lour.) Kuntze Habit: Herb. Habitat: Open bushland and sand
dunes. Vouchers: Rawlins SP 11578, Greenway & Rawlins 9291, Rawlins
155 (EA).
106
Veronicah M. Ngumbau et al. / PhytoKeys 147: 1–191 (2020)
Gisekia pharnaceoides L. var. pharnaceoides Habit: Herb. Habitat: Grassland, bushland. Vouchers: Muchiri J 490, Napier 3395, Jeffery K190, GRW Sangai 951 (EA).
F87. Goodeniaceae
1 Genus, 2 Species
Scaevola plumieri (L.) Vahl Habit: Shrub. Habitat: Coastal sand dunes. Vouchers:
Kirika P, Muthoka P & Mbale M 762, Verdcourt 3910, Greenway 13134 (EA).
Scaevola taccada (Gaertn.) Roxb. Habit: Shrub. Habitat: Strand vegetation on
sand and coral rock. Vouchers: Napper 3288 & 1298, Drummond & Hemsley
1059 (EA).
F88. Hernandiaceae
2 Genera, 2 Species
Gyrocarpus americanus Jacq. Habit: Tree. Habitat: Evergreen forest and derived secondary forest, 15–510 m. Voucher: Ngumbau V & Mwadime N V0405 (EA, HIB).
Hernandia nymphaeifolia (J. Presl) Kubitzki Habit: Tree. Habitat: Sandy foreshores
near high-water mark in littoral bushland, sea level. Vouchers: Drummond &
Hemsley 3986, Bock in EAH 16317 (EA).
F89. Hydrocharitaceae
6 Genera, 8 Species
Enhalus acoroides (L. f.) Royle Habit: Herb. Habitat: On sand or mud in sheltered
water, up to 4 m deep. Vouchers: Frazier 912, Greenway & Rawlins 9377 (EA).
Halophila gaudichaudii J. Kuo Habit: Herb. Habitat: On sand and mud in sheltered
sea water, up to 2 m depth. Vouchers: Isaac 116, 100 & 21 (EA).
Halophila ovalis (R. Br.) Hook. f. Habit: Herb. Habitat: On fine mud to coarse coral
rubble, mid-tidal level to 12 m. Vouchers: SAJIT–006265 (EA, HIB), Isaac 66,
Gillespie 317 (EA).
Halophila stipulacea (Forssk.) Asch. Habit: Herb. Habitat: On sand and mud in
sheltered sea water, up to about 7 m deep. Vouchers: Isaac 61, Rayner 288, Greenway & Rawlins 9325 (EA).
Lagarosiphon cordofanus Casp. Habit: Herb. Habitat: Still or slow flowing freshwater, ca. 30 m. Vouchers: Polphill & Paulo 846, Festo 2714 (EA).
Najas horrida A. Braun ex Magnus Habit: Herb. Habitat: Lakes, river-banks, and
swamps, ca. 5 m. Voucher: Starzenski 50 (EA).
Ottelia exserta (Ridl.) Dandy Habit: Herb. Habitat: Still freshwater or rarely wet
mud, ca. 50 m. Vouchers: Hooper & Towsend 1205, Gillespie 339 & 340 (EA).
Thalassia hemprichii (Ehrenb. ex Solms) Asch. Habit: Herb. Habitat: Fine mud to
clean coral sand on reef platforms, and sublittoral flats, low water mark to 5 m
depth. Vouchers: Frazier 885 & 871, Gillespie 319 (EA).
Checklist of coastal forests of Kenya
107
F90. Hydroleaceae
1 Genus, 1 Species
Hydrolea zeylanica (L.) Vahl Habit: Herb. Habitat: Pond margins, streamside, open
forest, and swampy area, 0–100 m. Vouchers: Ngumbau V & Mwadime N V0546
(EA, HIB), Mwadime N & Luke WRQ 2572 (EA).
F91. Hypericaceae
3 Genera, 3 Species
Harungana madagascariensis Lam. ex Poir. Habit: Tree. Habitat: Lowland and upland rainforest, ca. 0–305 m. Vouchers: SAJIT–006167 & 005944, Ngumbau V
& Mwadime N V0293 (EA, HIB), Luke WRQ & Robertson SA 222 (EA).
Psorospermum febrifugum Spach Habit: Shrub. Habitat: Deciduous woodland and
wooded grassland, ca. 50 m. Vouchers: Luke WRQ & Robertson SA 225, Donald
36 (EA).
Vismia orientalis Engl. Habit: Tree. Habitat: Forest, ca. 229 m. Vouchers: SAJIT–006046, Ngumbau V & Mwadime N V0162 (EA, HIB), Luke WRQ &
Robertson SA 519, Graham 2119 (EA).
F92. Hypoxidaceae
2 Genera, 2 Species
Curculigo pilosa subsp. major (Baker) Wiland Habit: Herb. Habitat: Grassland and
woodland, ca. 350 m. Voucher: Magogo FC & Glover PE 469 (EA).
Hypoxis angustifolia Lam. var. angustifolia Habit: Herb. Habitat: Grassland and
open woodland, ca. 136 m. Vouchers: SAJIT–005946 (EA, HIB), Magogo FC &
Glover PE 211 (EA).
Hypoxis angustifolia var. luzuloides (Robyns & Tournay) Wiland Habit: Herb.
Habitat: Grassland, swampy grassland, scattered tree grassland, and secondary
grassland, ca. 445 m. Voucher: Magogo & Glover 498 (EA).
F93. Icacinaceae
2 Genera, 4 Species
Iodes usambarensis Sleumer Habit: Liana. Habitat: Edges of lowland rainforest, sometimes persisting on cultivated ground, below 100 m. Vouchers: SAJIT–006221
(EA, HIB), Luke Q 1406, Rawlins 205 (EA).
Pyrenacantha kaurabassana Baill. Habit: Climber. Habitat: Dry scrub and bushland to deciduous woodland, ca. 65 m. Vouchers: SAJIT–006438 (EA, HIB),
Mwangangi OM 1299, Drummond & Hemsley 1023 (EA).
Pyrenacantha malvifolia Engl. var. malvifolia Habit: Liana. Habitat: Bushland.
Voucher: Moomaw JC 1001 (EA).
108
Veronicah M. Ngumbau et al. / PhytoKeys 147: 1–191 (2020)
Pyrenacantha vogeliana Baill. Habit: Climber. Habitat: Moist forest, ca. 249 m.
Vouchers: Ngumbau V & Mwadime N V001 (EA, HIB), Robertson SA & Luke
WRQ 5162 (EA).
F94. Iridaceae
2 Genera, 2 Species
Gladiolus dalenii subsp. andongensis (Baker) Goldblatt Habit: Herb. Habitat: Grassland and light woodland, ca. 300 m. Voucher: Magogo FC & Glover PE 581 (EA).
Afrosolen schimperi (Aschers. & Klatt) Goldblatt & J. C. Manning Habit: Herb.
Habitat: Moist habitats, along streams, and seasonal marshes, ca. 20 m. Vouchers:
Battiscombe 231, JB Gillespie 176 (EA).
F95. Kewaceae
1 Genus, 1 Species
Kewa bowkeriana (Sond.) Christenhusz Habit: Herb. Habitat: Saline sandy maritime
inland soils. Vouchers: Festo L & Luke Q 2585, Greenway & Rawlins 8912 (EA).
F96. Lamiaceae
18 Genera, 67 Species
Aeollanthus zanzibaricus S. Moore Habit: Herb. Habitat: Rocks or shallow soil overlying rocks, 200–500 m. Vouchers: Festo L, Luke Q & P 2631, Magogo FC &
Glover PE 903 (EA).
Basilicum polystachyon (L.) Moench Habit: Herb. Habitat: Waterside and seasonally
flooded areas, ca. 300 m. Vouchers: Mutangah 38, RB & AJ Faden 74/1279 (EA).
Clerodendrum cephalanthum Oliv. subsp. cephalanthum Habit: Shrub. Habitat:
Coastal bushland, 0–330 m. Vouchers: Ngumbau V & Mwadime N V0418 (EA,
HIB), Mwadime N 14, Magogo & Estes 1223, Drummond & Hemsley 3879 (EA).
Clerodendrum hildebrandtii Vatke var. fischeri (Gürke) Verdc. Habit: Shrub. Habitat: Wooded grassland, secondary bush, 5–230 m. Vouchers: Magogo FC & Glover PE 1005, Festo L & Luke Q 2456 (EA).
Clerodendrum robustum Klotzsch var. robustum Habit: Shrub. Habitat: Wet grasslands, woodland, and forest clearings, 0–450 m. Vouchers: Jeffrey K140, Maggridge 138, Battscombe 50 & Gardener in FD 1454, Luke WRQ et al. 6189 (EA).
Clerodendrum tricholobum Gürke Habit: Shrub. Habitat: Riverine bushland, woodland, and grassland mosaic, 0–300 m. Vouchers: J Adamson 312, Robertson &
Luke 5617, Whyte (EA).
Endostemon albus A. J. Paton, Harley & M. M. Harley Habit: Herb. Habitat: Grassy
places, streamside in coastal, and deciduous bushland, ca. 15–350 m. Vouchers: Luke
WRQ & PA 6021, PA & WRQ Luke 6021, Drummond & Hemsley 4172 (EA).
Endostemon ctenoneurus Harley Habit: Herb. Habitat: Bushland, 0–400 m. Voucher: Festos L & Luke Q 2556 (EA).
Checklist of coastal forests of Kenya
109
Endostemon gracilis (Benth.) M. Ashby Habit: Herb. Habitat: Acacia-Commiphora bushland, Brachystegia woodland, and cultivations, 0–350 m. Vouchers: SAJIT–006135 (EA, HIB), Kirika P, Muthoka P & Mbale M 740 (EA).
Endostemon tereticaulis (Poir.) M. Ashby Habit: Herb. Habitat: Short grassland,
coastal, and Acacia-Commiphora bushland, ca. 30 m. Vouchers: Magogo FC &
Glover PE 687, Greenway 9486 (EA).
Endostemon wakefieldii (Baker) M. Ashby Habit: Herb. Habitat: Brachystegia woodland, ca. 100 m. Voucher: Graham 1739 (EA): Endemic.
Hoslundia opposita Vahl Habit: Shrub. Habitat: Open areas in grassland, woodland or
forest, ca. 100 m. Vouchers: Gillett JB 20412, Magogo FC & Glover PE 176 (EA).
Karomia gigas (Faden) Verdc. Habit: Tree. Habitat: Relict forest on limestone rocks,
ca. 140 m. Vouchers: Faden et al. 77/439, Gillett 22775 (EA).
Leonotis nepetifolia (L.) R.Br. var. nepetifolia Habit: Shrub. Habitat: Grassland,
bushland, and disturbed places. Voucher: Robertson SA 4974 (EA).
Leucas glabrata (Vahl) J. C. Manning & Goldblatt Habit: Herb. Habitat: Damp
ground in riverine, ca. 30 m. Voucher: Luke et al. TPR 501 (EA).
Leucas martinicensis (Jacq.) R.Br. Habit: Herb. Habitat: Grassland and bushland,
ca. 382 m. Vouchers: Ngumbau V & Mwadime N V0370 (EA, HIB), Graham
RM 2044 (EA).
Leucas nubica Benth. Habit: Herb. Habitat: Grassland, coastal or deciduous bushland, ca. 30 m. Voucher: Luke et al. TPR 550 (EA).
Leucas tomentosa Gürke Habit: Shrub. Habitat: Bushland and rough grassland, ca.
200 m. Voucher: Mungai, Mutangah & Rucina 335/83 (EA).
Leucas tsavoensis var. kilifiensis Sebald Habit: Herb. Habitat: Brachystegia woodland or
degraded woodland, 30–150 m. Voucher: Polhill RM & Paulo S 815 (EA): Endemic.
Leucas urticifolia var. angustifolia Sebald Habit: Herb. Habitat: Wooded grassland
and abandoned cultivated areas, ca. 30 m. Vouchers: Luke et al. TPR 549, Brand
s.n. (EA).
Mesosphaerum pectinatum (L.) Kuntze Habit: Shrub. Habitat: Edges of rivers, lakes,
swamps, thickets or disturbed habitats. Voucher: Luke WRQ & Robertson SA
2764 (EA).
Mesosphaerum suaveolens (L.) Kuntze Habit: Herb. Habitat: Disturbed places, roadsides, and cultivated ground, ca. 30 m. Voucher: Magogo FC & Glover PE 876
(EA): Naturalized.
Ocimum americanum L. Habit: Herb. Habitat: Roadsides, 0–500 m. Vouchers: Patterson GD 31, Magogo FC & Glover PE 880 (EA).
Ocimum basilicum L. Habit: Herb. Habitat: Cultivated and disturbed ground, ca.
420 m. Vouchers: Bally 13279, Jeffery 208 (EA): Cultivated.
Ocimum filamentosum Forssk. Habit: Herb. Habitat: Brachystegia woodland, Combretum or Acacia-Commiphora or coastal bushland, and grassland, ca. 195 m.
Vouchers: Ngumbau V & Mwadime N V0457 (EA, HIB), Magogo FC & Glover
PE 922 (EA).
Ocimum fischeri Gürke Habit: Herb. Habitat: Acacia-Commiphora bushland, scrub,
and rocky ground, 100–500 m. Voucher: Drummond & Hemsley 4123 (EA).
110
Veronicah M. Ngumbau et al. / PhytoKeys 147: 1–191 (2020)
Ocimum gratissimum L. var. gratissimum Habit: Herb. Habitat: Dry riverine forest,
semi-deciduous forest, ca. 30–160 m. Vouchers: Robertson SA et al. 175, Kimberly Medley 271 (EA).
Ocimum obovatum E. Mey. ex Benth. Habit: Herb. Habitat: Grassland, open Acacia
or Brachystegia woodland, ca. 100 m. Vouchers: Moomaw JC 1135, Ossent 73 (EA).
Orthosiphon hanningtonii (Baker) A.J. Paton Habit: Herb. Habitat: SetariaSporobolus grassland, and Terminalia wooded grassland, 75–110 m. Voucher: Luke
& Luke 6307 (EA).
Orthosiphon parvifolius Vatke Habit: Herb. Habitat: Seasonally waterlogged grassland, ca. 35 m. Voucher: Luke Q 1488 (EA).
Orthosiphon rubicundus (D. Don) Benth. Habit: Herb. Habitat: Woodland, ca.
150 m. Voucher: Luke WRQ 3138 (EA).
Orthosiphon thymiflorus (Roth) Sleesen Habit: Herb. Habitat: Wooded grassland
and deciduous woodland, ca. 59 m. Voucher: Ngumbau V & Mwadime N V0439
(EA, HIB).
Platostoma africanum P. Beauv. Habit: Herb. Habitat: Cleared or swampy areas in
open woodland, 100–400 m. Vouchers: Luke WRQ et al. 4722, Drummond &
Hemsley 3877, Magogo & Glover 377, Spjut 2736 (EA).
Plectranthus aegyptiacus (Forssk.) C.Chr. Habit: Herb. Habitat: Acacia-Commiphora bushland, cultivated, and disturbed ground, 0–1300 m. Voucher: Robertson SA
& Luke WRQ 6052 (EA).
Plectranthus amboinicus (Lour.) Spreng. Habit: Herb. Habitat: Acacia-Commiphora
bushland, and wooded grassland, ca. 30 m. Voucher: Luke & Robertson 2568 (EA).
Plectranthus auriglandulosus A.J. Paton Habit: Herb. Habitat: Dry coastal forest
and bushland, 40–550 m. Vouchers: Luke 1585, Newton 3772, Maikweki, Mturi
& Abio 605 (EA): Endemic.
Plectranthus bojeri (Benth.) Hedge Habit: Herb. Habitat: Grassland, woodland,
open rocky areas, and forest clearings, ca. 120 m. Vouchers: Robertson SA & Luke
WRQ 6003, Robertson 6482 (EA).
Plectranthus dupuisii (Briq.) A. J. Paton Habit: Herb. Habitat: Grassland or woodland, and seasonally flooded areas, 100–500 m. Voucher: Robertson & Luke
5871 (EA).
Plectranthus flaccidus (Vatke) Gürke Habit: Herb. Habitat: Dunes in coastal scrub
or evergreen forest, 0–150 (–450) m. Vouchers: Luke Q 5668, Verdcourt 1860,
Faden 74 (EA).
Plectranthus hadiensis (Forssk.) Schweinf. ex Sprenger Habit: Herb. Habitat: Acacia-Commiphora bushland, deciduous woodland, and dry evergreen bushland, ca.
800 m. Voucher: Luke WRQ & Robertson SA 2828 (EA).
Plectranthus lasianthus (Gürke) Vollesen Habit: Herb. Habitat: Acacia-Commiphora
bushland, 100 m. Voucher: Festo L, Luke Q & P 2637 (EA).
Plectranthus leptophyllus (Baker) A. J. Paton Habit: Herb. Habitat: Openings and
margins of closed evergreen forest, ca. 450 m. Voucher: Cokefield (EA).
Plectranthus longipes Baker Habit: Herb. Habitat: Dry bushland or woodland, 70 m.
Voucher: Robertson SA & Luke WRQ 6004 (EA).
Checklist of coastal forests of Kenya
111
Plectranthus tetragonus Gürke Habit: Herb. Habitat: Acacia-Commiphora bushland,
ca. 80 m. Voucher: Luke WRQ & PA sr.
Plectranthus prostratus Gürke Habit: Herb. Habitat: Acacia scrub, ca. 60 m. Vouchers: Rawlins SP 766, Faden & Faden 74/1051 (EA).
Premna chrysoclada (Bojer) Gürke Habit: Shrub. Habitat: Coastal Brachystegia
woodland, scrub, and thicket, 0–450 m. Vouchers: SAJIT–005603, Robertson SA
4224, Mutanga JG & Kamau P 24 (EA).
Premna discolor Verdc. var. discolor Habit: Shrub. Habitat: Jurassic limestone outcrop with Pandanus, Euphorbia, Gyrocarpus, Ficus and Cola, ca. 260 m. Vouchers:
Luke & Robertson 1886, Luke 1324 (EA).
Premna discolor var. dianiensis Verdc. Habit: Shrub. Habitat: Very mixed dry evergreen forests, 0–12 m. Vouchers: Friis 1211, Gillett & Kibuwa 19854, Robertson
& Luke 5893 (EA).
Premna gracillima Verdc. Habit: Liana. Habitat: Deciduous coastal thicket, 400–
660 m. Vouchers: Ngumbau V & Mwadime N V017 (EA, HIB), Robertson SA &
Luke WRQ 1820, Luke WRQ 2935 (EA).
Premna hildebrandtii Gürke Habit: Shrub. Habitat: Dry coastal evergreen forest,
0–200 m. Vouchers: SAJIT–005589 (EA, HIB), Moggridge JY 97 (EA).
Premna senensis Klotzsch Habit: Climber. Habitat: Acacia woodland, thicket, and
riverine woodland, (100–) 300 m. Vouchers: Luke WRQ 3134, Magogo & Glover
530 (EA).
Premna serratifolia L. Habit: Shrub. Habitat: Sandy shores, thicket, and riverine
forest, 0–375 m. Vouchers: Magogo & Estes 1242, Magogo & Glover 1090 (EA).
Premna velutina Gürke Habit: Shrub. Habitat: Grassland, forest, riverine woodland,
18–500 m. Vouchers: Bally 16827, Kibuwa 2458, Greenway & Rawlins 9370 (EA).
Premna resinosa subsp. holstii (Gürke) Verdc. Habit: Shrub. Habitat: Coastal evergreen
forest and thicket on coral, 0–20 (–580) m. Vouchers: Ngumbau V & Mwadime N
V0350 (EA, HIB), Gillet 18642, GW Jeffery 462, RB & AJ Faden 74/1116 (EA).
Rotheca microphylla (Blume) Callm. & Phillipson Habit: Shrub. Habitat: Coastal
thicket, forest scrub, and Brachystegia-Hymenaea woodland, 5–450 m. Vouchers: SAJIT–005567 (EA, HIB), Magogo FC & Glover PE 926, Gillett JB 20340,
Drummond & Hemsley 3786, Musyoki & Hansen 1016, Jarman in Gillespie 70,
Mwadime N 671 (EA).
Rotheca rupicola (Verdc.) Verdc. Habit: Shrub. Habitat: Acacia-Cordia open bushland, ca. 160 m. Voucher: Hemming 83/111 (EA).
Rotheca sansibarensis (Gürke) Steane & Mabb. Habit: Shrub. Habitat: Evergreen
forest and edges, ca. 229 m. Vouchers: Ngumbau V & Mwadime N V0159 (EA,
HIB), Magogo & Glover 855, Musyoki & Hansek 1037, Luke WRQ 3598 (EA).
Tinnea aethiopica subsp. litoralis Vollesen Habit: Shrub. Habitat: Grassland and
deciduous woodland, 0–750 m. Vouchers: Ngumbau V & Mwadime N V0289
(EA, HIB), Festo L, Luke Q & P 2632, Greenway 9655 (EA).
Tinnea aethiopica subsp. stolzii (Robyns & Lebrun) Vollesen Habit: Shrub. Habitat: Grassland and deciduous woodland, ca. 0–100 m. Voucher: Luke & Robertson 1930 (EA).
112
Veronicah M. Ngumbau et al. / PhytoKeys 147: 1–191 (2020)
Vitex doniana Sweet Habit: Tree. Habitat: Combretum woodland and elephant grassland, ca. 0–36 m. Vouchers: SAJIT–006194 & 005558 (EA, HIB), RM Graham
756 in FD 2226 (EA).
Vitex ferruginea Schumach. & Thonn. Habit: Tree. Habitat: Bushland and dry lowland
forest, 0–570 m. Vouchers: SAJIT–005498, Ngumbau V & Mwadime N V0543
(EA, HIB), Luke WRQ 3133, Napier 3304, Polhill & Paulo 723, Rawlins (EA).
Vitex mombassae Vatke Habit: Tree. Habitat: Scrub, thicket, and Brachystegia woodland, ca. 20–305 m. Vouchers: SAJIT–005475, Ngumbau V & Mwadime N
V0297 (EA, HIB), Drummond & Hemsley 1156, Magogo & Glover 505, RB &
AJ Faden 77/633 (EA).
Vitex payos (Lour.) Merr. Habit: Tree. Habitat: Wooded grassland, Acacia woodland,
and rocky outcrops, ca. 30 m. Voucher: Magogo & Glover 504 (EA).
Vitex schliebenii Moldenke Habit: Shrub. Habitat: Bushland, Acacia scrub, thicket,
and riverine, 5–500 m. Vouchers: Polhill & Paulo 649, RB & AJ Faden 74/1191,
Gillespie 91 (EA).
Vitex strickeri Vatke & Hildebr. Habit: Shrub. Habitat: Bushland, thickets, woodland,
and forest edges, ca. 0–400 m. Vouchers: Kuchar P 13655, RM Graham 814 (EA).
Vitex trifolia L. Habit: Shrub. Habitat: Open vegetation near shore, 0–20 m. Vouchers: Brenan et al. 14522, Templar 23, Jeffery 348 (EA).
Vitex zanzibarensis Vatke Habit: Tree. Habitat: Deciduous coastal thicket and dry
forest, 0–600 m. Vouchers: SAJIT–006091 (EA, HIB), SA Robertson & Luke
4910, Conservator of forests H65/43 (EA).
Volkameria acerbiana Vis. Habit: Shrub. Habitat: Acacia woodland, scrub, and sand
dunes, ca. 0–180 m. Vouchers: RM Graham in FD 2237, Battiscombe 229, Werner in Battiscombe 997, Festo L, Luke Q & P 2671 (EA).
Volkameria glabra (E. Mey.) Mabb. & Y.W. Yuan Habit: Shrub. Habitat: Littoral
sand dunes associations, coastal bushland, and evergreen thicket on coral cliffs, ca.
0–50 (–480) m. Vouchers: SAJIT–006177 (EA, HIB), Napier 3303 in CM 6237,
Tweedie 2373, Gillespie 174 (EA).
F97. Lauraceae
1 Genus, 1 Species
Cassytha filiformis L. Habit: Climber. Habitat: Bushland, scrub, mixed woodland,
thickets, and forest patches, ca. 0–445 m. Vouchers: Allen JD 15123, Verdcourt
3960 (EA).
F98. Lecythidaceae
1 Genus, 1 Species
Barringtonia racemosa (L.) Spreng. Habit: Tree. Habitat: In forest by rivers, mangroves, and bordering grassland, 0–450 (–750) m. Vouchers: Ngumbau V &
Mwadime N V0394 (EA, HIB), Magogo FC & Glover PE 891 (EA).
Checklist of coastal forests of Kenya
113
F99. Lentibulariaceae
1 Genus, 5 Species
Utricularia arenaria A. DC. Habit: Herb. Habitat: Boggy grassland, damp open
sandy ground, and rock pavements. Vouchers: Magogo FC & Glover PE 320,
Polhill & Paulo 902 (EA).
Utricularia firmula Welw. ex Oliv. Habit: Herb. Habitat: Boggy grassland. Voucher:
Magogo FC & Glover PE 321 (EA).
Utricularia gibba subsp. exoleta (R.Br.) P. Taylor Habit: Herb. Habitat: Shallow water or mud in swamps, pools, and ditches. Voucher: Rawlins in EAH 2125 (EA).
Utricularia inflexa Forssk. Habit: Herb. Habitat: Still or slow flowing water in pools,
lakes, and rivers, ca. 15 m. Vouchers: SAJIT–006228 (EA, HIB), Leauthaud C &
Geopar 27, Drummond & Hemsley 4017 (EA).
Utricularia stellaris L.f. Habit: Herb. Habitat: Still or slowly flowing water in lakes,
marshes, and rivers. Voucher: Greenway 9242 (EA).
F100. Linaceae
1 Genus, 1 Species
Hugonia castaneifolia Engl. Habit: Liana. Habitat: Dry evergreen or riverine forest,
7–150 m. Vouchers: SAJIT–006009, Ngumbau V & Mwadime N V0145 (EA,
HIB), Greenway 9638 & 10839, Jeffery K300, Magogo FC & Glover PE 37 (EA).
F101. Linderniaceae
8 Genera, 11 Species
Artanema longifolium var. amplexicaule Vatke Habit: Herb. Habitat: Wet grassland,
forest swamps, and plantations, 0–800 m. Vouchers: Hildebrandt 2001b, Graham
2217, Luke & Robertson 2334 (EA).
Craterostigma hirsutum S. Moore Habit: Herb. Habitat: Shallow soil over rock where
water runs during rains, ca. 350 m. Voucher: Drummond & Hemsley 4149 (EA).
Craterostigma newtonii (Engl.) Eb. Fisch., Schäferh. & Kai Müll. Habit: Herb.
Habitat: Wet grassland, marshes, banks of streams, and coasts, ca. 20 m. Voucher:
Polhill & Paulo 893 (EA).
Crepidorhopalon goetzei (Engl.) Eb. Fisch. Habit: Herb. Habitat: Boggy places besides streams, ca. 140 m. Voucher: Luke & Gray 4061 (EA).
Crepidorhopalon hepperi Eb. Fisch. Habit: Herb. Habitat: Secondary grassland and
wet forests, ca. 150 m. Vouchers: Luke 2899 & 3796, Luke & Gray 4061(EA).
Crepidorhopalon whytei (Skan) Eb. Fisch Habit: Herb. Habitat: Riversides and
marshy area, ca. 200 m. Voucher: Drummond & Hemsley 4007 (EA).
Linderniella bolusii (Hiern) Eb. Fisch., Schäferh. & Kai Müll. Habit: Herb. Habitat: Wet, shallow, sandy soil surrounding outcrops, ca. 250 m. Vouchers: Polhill &
Paulo 486, Drummond & Hemsley 4150 (EA).
114
Veronicah M. Ngumbau et al. / PhytoKeys 147: 1–191 (2020)
Lindernia parviflora (Roxb.) Haines Habit: Herb. Habitat: Sandy, muddy river, permanent pools and rice fields, ca. 27 m. Vouchers: SAJIT–005491, Ngumbau V
& Mwadime N V0556 (EA, HIB), Tweedie 1039, Luke WRQ & Robertson SA
2751 (EA).
Stemodiopsis rivae Engl. Habit: Herb. Habitat: In rock crevices, ca. 600 m. Voucher:
Mwadime N & Luke WRQ 2428 (EA).
Torenia thouarsii (Cham. & Schltdl.) Kuntze Habit: Herb. Habitat: Wet sandy
soil, often shady, and by streams or pools, ca. 95 m. Vouchers: SAJIT–006140 &
005486 (EA, HIB), Graham 1704, Drummond & Hemsley 4012, Luke WRQ
1652 (EA).
Vandellia humilis (Bonati) Eb. Fisch., Schäferh. & Kai Müll. Habit: Herb. Habitat:
Margins of pools and water holes, rice fields, and streamside in lowland forests,
0–400 m. Vouchers: Ngumbau V & Mwadime N V0553 (EA, HIB), Drummond
& Hemsley 4008, Luke & Robertson 2714, Luke 3044, Luke WRQ 2925 (EA).
F102. Loganiaceae
1 Genus, 12 Species
Strychnos angolensis Gilg Habit: Liana. Habitat: Moist forest and riverine forest, ca.
407 m. Vouchers: SAJIT–006034 (EA, HIB), Luke WRQ 2926 (EA).
Strychnos decussata (Pappe) Gilg Habit: Tree. Habitat: Dry forest and woodlands,
ca. 500 m. Voucher: Luke WRQ & Robertson SA 57 (EA).
Strychnos henningsii Gilg Habit: Tree. Habitat: Upland and lowland rainforest,
semi-evergreen bushland, lowland dry evergreen forest, and riverine forest, 340 m.
Vouchers: SAJIT–006444 (EA, HIB), Kuchar P 13523, Drummond & Hemsley
4226 (EA).
Strychnos kasengaensis De Wild. Habit: Liana. Habitat: Riverine forest, ca. 200 m.
Vouchers: Swynnerton R JM 44, Moreau JC 1072 (EA).
Strychnos madagascariensis Poir. Habit: Tree. Habitat: Bushland and open woodland, 0–450 m. Vouchers: SAJIT–005525 (EA, HIB), MacNaughton 90 in FD
2622, Dale 3803, Drummond & Hemsley 4077, Muchiri J 441(EA).
Strychnos madagascariensis Poir. F. “b” Habit: Tree. Habitat: Forest. Voucher: Robertson SA 5657 (EA).
Strychnos madagascariensis Poir. F. “e” Habit: Tree. Habitat: Forest. Voucher: Ross
KS 119 (EA).
Strychnos mellodora S. Moore Habit: Tree. Habitat: Found in riverine and moist
montane forest, ca. 400 m. Voucher: Luke WRQ & Robertson SA 2740 (EA).
Strychnos mitis S. Moore Habit: Tree. Habitat: Forests and coastal bushland, ca.
20 m. Vouchers: Robertson SA & Luke WRQ 5511, Luke WRQ & Robertson
SA 2100 (EA).
Strychnos panganensis Gilg Habit: Liana. Habitat: Lowland rainforest and coastal
evergreen bushland, 50–500 m. Vouchers: SAJIT–005488 & 004644 (EA, HIB),
Luke Q 1480, Magogo FC & Glover PE 459, RM Graham 1581, Swynnerton 9,
Moggridge 503 (EA).
Checklist of coastal forests of Kenya
115
Strychnos scheffleri Gilg Habit: Liana. Habitat: Lowland rainforest, ca. 200 m.
Vouchers: Ngumbau V & Mwadime N V066 (EA, HIB), Luke WRQ & Robertson SA 1415, RM Graham 2045, Drummond & Hemsley 1094, CF Elliott 327,
Luke WRQ 884 (EA).
Strychnos spinosa Lam. Habit: Tree. Habitat: Dry woodland and thickets, ca. 400 m.
Vouchers: SAJIT–006000, Ngumbau V & Mwadime N V0216 (EA, HIB),
Kuchar P 12901, Luke WRQ & Robertson SA 2791 (EA).
Strychnos spinosa subsp. volkensii (Gilg) E.A. Bruce Habit: Shrub. Habitat: Coastal
evergreen bushland, ca. 10 m. Vouchers: Adamson J 5976, RM Graham 1770,
Mrs J Adamson 285 in Bally 5976, F Thomas 198 (EA).
Strychnos usambarensis Gilg ex Engl. Habit: Tree. Habitat: Lowland and upland
rainforest, ca. 0–300 m. Vouchers: Sangai GW 986, Luke PA & WRQ 3565 (EA).
F103. Lophiocarpaceae
1 Genus, 1 Species
Corbichonia decumbens (Forssk.) Exell Habit: Herb. Habitat: Woodland, cultivated,
and disturbed ground, ca. 375 m. Voucher: Luke WRQ & PA 6019 (EA).
F104. Loranthaceae
11 Genera, 27 Species
Agelanthus heteromorphus (A. Rich.) Polhill & Wiens Habit: Shrub. Habitat:
Coastal and deciduous forest, ca. 150 m. Vouchers: Graham RM 1598, Luke &
SA Robertson 2231 (EA).
Agelanthus kayseri (Engl.) Polhill & Wiens Habit: Shrub. Habitat: Coastal bushland
and mangrove stands, 0–600 m. Vouchers: Robertson, SA & Luke WRQ 5713,
Wiens 452 (EA).
Agelanthus longipes (Baker & Sprague) Polhill & Wiens Habit: Shrub. Habitat:
Coastal forest and associated bushland, 30–650 m. Vouchers: Ngumbau V &
Mwadime N V0101 (EA, HIB), Luke 4023, Magogo & Glover 968, Musyoki &
Hansen 999 (EA).
Agelanthus microphyllus Polhill & Wiens Habit: Shrub. Habitat: Deciduous bushland, ca. 30 m. Voucher: Moomaw 1033A (EA): Endangered.
Agelanthus oehleri (Engl.) Polhill & Wiens Habit: Shrub. Habitat: Drier habitats,
ca. 20 m. Voucher: Gilbert MG & Kuchar P 5928 (EA).
Agelanthus sansibarensis (Engl.) Polhill & Wiens subsp. sansibarensis Habit:
Shrub. Habitat: Coastal bushland, mangrove swamps, and forest, ca. 181 m.
Vouchers: Ngumbau V & Mwadime N V0222 (EA, HIB), Magogo & Glover
194, Verdcourt 2127, Kuchar P 12869, Magogo FC & Glover PE 61, Jeffery GW
122 (EA).
Agelanthus subulatus (Engl.) Polhill & Wiens Habit: Shrub. Habitat: Forest to
woodland, bushland, and wooded grassland, 10–380 m. Vouchers: Festo L &
Luke Q 2560, Luke WRQ 3106, Polhill 4823, MG Gilbert & Kuchar 5853 (EA).
116
Veronicah M. Ngumbau et al. / PhytoKeys 147: 1–191 (2020)
Emelianthe panganensis (Engl.) Danser subsp. panganensis Habit: Shrub. Habitat:
Deciduous and coastal bushland, ca. 0–200 m. Vouchers: Robertson SA 4037,
RM Graham in FD 1818 (EA).
Emelianthe panganensis subsp. commiphorae Wiens & Polhill Habit: Shrub. Habitat: Coastal and deciduous bushland or wooded grassland, ca. 15 m. Vouchers: SA
Robertson 4037, Festo L, Luke Q & P 2799 (EA).
Englerina heckmanniana (Engl.) Polhill & Wiens Habit: Shrub. Habitat: Woodland, ca. 700 m. Voucher: Rawlins SP 934 (EA).
Englerina ramulosa (Sprague) Polhill & Wiens Habit: Shrub. Habitat: Coastal
bushland, 0–60 m. Vouchers: Tweedie 2386, SA Robertson 3600, Gillespie 184
(EA): Endangered.
Erianthemum alveatum (Sprague) Danser Habit: Shrub. Habitat: Coastal bushland,
woodland, and wooded grassland, 0–300 m. Vouchers: Wiens 4532, Greenway
9791, SA Robertson & Luke 4925A (EA): Vulnerable.
Erianthemum dregei (Eckl. & Zeyh.) Tiegh. Habit: Shrub. Habitat: Woodland,
bushland, wooded grassland, and disturbed places, ca. 0–396 m. Vouchers: SAJIT–005598, Ngumbau V & Mwadime N V033 (EA, HIB), Robertson SA &
Luke WRQ 5576, Magogo FC & Glover PE 966, Wiens 4527 (EA).
Helixanthera kirkii (Oliv.) Danser Habit: Shrub. Habitat: Coastal bushland
and mixed dry bushland, ca. 10 m. Vouchers: Festo L & Luke Q 2563, Polhill
4811 (EA).
Oliverella hildebrandtii (Engl.) Tiegh. Habit: Shrub. Habitat: Deciduous and coastal bushland, ca. 0–30 m. Vouchers: Faden, Gillett & Gachachi 77/471, Festo L,
Luke Q & P 2792, Magogo FC & Glover PE 686 (EA).
Oncella ambigua (Engl.) Tiegh. Habit: Shrub. Habitat: Coastal bushland and riverine forest, 0–300 m. Vouchers: Festo L, Luke Q & P 2681, Gillett & Kibuwa
19913, Luke et al. in TPR 68, RM Graham in FD 1799 (EA).
Oncella curviramea (Engl.) Danser Habit: Shrub. Habitat: Coastal forest and bushland, ca. 0–80 m. Vouchers: Robertson SA & Luke WRQ 5555, Polhill & SA
Robertson 4846, Bally & Smith 14379, SA Robertson 6880 (EA).
Oncocalyx cordifolius Wiens & Polhill Habit: Shrub. Habitat: Coastal bushland, on
various hosts, near sea level. Vouchers: Gilbert MG 6834, SA Robertson & Luke
5559, MG Gilbert & Kuchar 5871, Buck in EA 16032 (EA).
Oncocalyx fischeri (Engl.) M.G. Gilbert Habit: Shrub. Habitat: Woodland, ca. 10 m.
Voucher: Festo L, Laizer G, Luke Q & P 2742 (EA).
Oncocalyx kelleri (Engl.) M.G. Gilbert Habit: Shrub. Habitat: Deciduous bushland
or commonly riverine, ca. 15 m. Voucher: Polhill & Paulo 500 (EA).
Oncocalyx ugogensis (Engl.) Wiens & Polhill Habit: Shrub. Habitat: Deciduous
bushland, ca. 15 m. Voucher: Gillett 16375 (EA).
Plicosepalus curviflorus (Benth. ex Oliv.) Tiegh. Habit: Shrub. Habitat: Deciduous
bushland, ca. 30 m. Voucher: Luke WRQ 5681 (EA).
Plicosepalus kalachariensis (Schinz) Danser Habit: Shrub. Habitat: Mixed woodland, ca. 30 m. Voucher: Luke & SA Robertson 2368 (EA).
Checklist of coastal forests of Kenya
117
Plicosepalus meridianus (Danser) Wiens & Polhill Habit: Shrub. Habitat: Deciduous
bushland, ca. 100 m. Vouchers: RB & AJ Faden 74/1022, Luke WRQ 3513 (EA).
Plicosepalus sagittifolius (Engl.) Danser Habit: Shrub. Habitat: Coastal bushland,
ca. 450 m. Voucher: Karisa R9 (EA).
Spragueanella rhamnifolia (Engl.) Balle Habit: Shrub. Habitat: Forest along coast
or inland along rivers, ca. 30 m. Vouchers: SAJIT–004649 (EA, HIB), Verdcourt
1864, Wiens 4525.
Taxillus wiensii R. M. Polhill Habit: Shrub. Habitat: Dry evergreen coastal forest,
100–150 m. Vouchers: Wiens D 4526, Polhill 4825, Luke & SA Robertson 2173,
SA Robertson & Polhill 6861 (EA): Critically Endangered, Endemic.
F105. Lythraceae
4 Genera, 12 Species
Ammannia auriculata Willd. Habit: Herb. Habitat: Mud by dams and ponds, wooded
riverine flats, and damp grassland, ca. 5 m. Vouchers: F Thomas 70, Festo 2351 (EA).
Ammannia capitellata (C. Presl) S.A. Graham & Gandhi subsp. capitellata Habit:
Herb. Habitat: Edges of permanent water or rice fields, ca. 50 m. Vouchers: Gilbert & Kuchar 588, Luke Q 5661 (EA).
Ammannia parkeri (Verdc.) S. A. Graham & Gandhi var. parkeri Habit: Herb.
Habitat: Edges of water pans in open bushland, grassland, and flooded grassland,
30–460 m. Vouchers: Luke WRQ & Robertson SA 1550, Luke Q 1550, Luke et
al. TPR 761 (EA).
Ammannia parkeri var. longifolia (Verdc.) S. A. Graham & Gandhi Habit: Herb.
Habitat: Found in flooded grassland habitat, ca. 10 m. Vouchers: Festo L, Luke Q
& P 2740, Whyte (EA): Vulnerable, Endemic.
Ammannia pedicellata (Hiern) S. A. Graham & Gandhi Habit: Herb. Habitat:
Seasonally flooded freshwater swamps or rice fields, 0–150 m. Vouchers: Luke Q
5634, Luke 3034, Greenway & Rawlins 9451 (EA).
Ammannia prieuriana Guill. & Perr. Habit: Herb. Habitat: Sandy, bank of the river,
and edge of swamp. Voucher: Luke WRQ 3606 (EA).
Ammannia radicans (Guill. & Perr.) S. A. Graham & Gandhi var. radicans Habit:
Herb. Habitat: Swampy ground, grassy places on lava, pool edges, and rice-fields,
ca. 95 m. Vouchers: SAJIT–006139 (EA, HIB), Luke WRQ 3315, SA Robertson
3512, Whyte (EA).
Ammannia radicans var. floribunda (Sond.) S. A. Graham & Gandhi Habit: Herb.
Habitat: Marshes, swampy grassland, muddy ditches, riverbanks, and rice-fields,
ca. 60 m. Vouchers: Polhill & Paulo 891, Festo L & Luke Q 2581 (EA).
Ammannia senegalensis Lam. Habit: Herb. Habitat: Damp ground, seasonal freshwater short grass, and sedge swamp, 0–12 (–240) m. Vouchers: Hooper & Townsend
1127, Luke Q 5625 (EA).
Ammannia stuhlmannii (Koehne) S. A. Graham & Gandhi Habit: Herb. Habitat:
Open grassland, Brachystegia woodland, and irrigated rice fields, 15–75 m. Vouch-
118
Veronicah M. Ngumbau et al. / PhytoKeys 147: 1–191 (2020)
ers: Rawlins 865, Hooper & Townsend 1244, Polhill & Paulo 533, Festo L, Luke
Q & P 2624 (EA).
Ammannia urceolata Hiern Habit: Herb. Habitat: Floodplain grassland, ca. 75 m.
Vouchers: Luke et al. in TPR 124 & 780 (EA).
Lawsonia inermis L. Habit: Herb. Habitat: Temporarily flooded rocky river, riverine
thicket, also near coast, ca. 30 m. Voucher: Polhill & Paulo 673 (EA).
Pemphis acidula J. R. Forst. & G. Forst. Habit: Shrub. Habitat: Littoral at or even
below high-water mark, coral rag thicket, and drier sides of swamp, 0–5 m. Vouchers: Kimeu JM 650, Kibuwa 1204, Williams Sangai 865, Greenway & Rawlins
8873 (EA).
Sonneratia alba Sm. Habit: Tree. Habitat: Frequent in salty muds of mangrove
swamps, ca. 5 m. Vouchers: SAJIT–006264 (EA, HIB), Kuchar P 13607 (EA).
F106. Malpighiaceae
3 Genera, 5 Species
Acridocarpus alopecurus Sprague Habit: Liana. Habitat: Dry evergreen and riverine forest, 20–500 m. Vouchers: Ngumbau V & Mwadime N V090 (EA, HIB), Verdcourt
3920A, RM Graham in FD 1786, Luke Q 1408, Luke WQR et al. 4604F (EA).
Acridocarpus zanzibaricus A. Juss. Habit: Shrub. Habitat: Forest, 0–180 m. Vouchers: SAJIT–005568 (EA, HIB), VG van Someren 1793, Battiscombe 801 (EA).
Triaspis mozambica A. Juss. Habit: Liana. Habitat: Coastal and riverine bushland or
thicket, dry evergreen forest, and lowland rainforest, 0–800 m. Vouchers: Ngumbau V & Mwadime N V0411 (EA, HIB), Drummond & Hemsley 3910, Jeffery
451, Festo L, Luke Q & P 2623 (EA).
Triaspis niedenzuiana Engl. Habit: Liana. Habitat: Deciduous bushland, sometimes
on stony hillsides or rock outcrops, ca. 500 m. Voucher: Kassner 533 (EA).
Tristellateia africana S. Moore Habit: Liana. Habitat: Coastal bushland, thicket,
and mangrove swamp, 0–50 m. Vouchers: Drummond & Hemsley 3980, VG van
Someren 1791, Polhill & Paulo 688, Festo L, Luke Q & P 2725 (EA).
F107. Malvaceae
30 Genera, 117 Species
Abelmoschus ficulneus (L.) Wight & Arn. Habit: Herb. Habitat: Grassland on clay, flood
plains, riverbanks, and roadsides, ca. 20 m. Voucher: Robertson & Luke 5372 (EA).
Abelmoschus moschatus Medik. Habit: Herb. Habitat: Riverine vegetation, ca. 28
m. Vouchers: SAJIT–006207 & 006208, Ngumbau V & Mwadime N V0541
(EA, HIB).
Abutilon guineense (Schumach.) Baker f. & Exell Habit: Herb. Habitat: Grassland,
bushland, and wooded bushland, ca. 30 m. Voucher: Kabuye et al. TPR 322 (EA).
Abutilon mauritianum subsp. zanzibaricum (Mast.) Verdc. Habit: Herb. Habitat:
Coastal thicket, bushland, sandy grassland, in banana, and coconut plantations,
Checklist of coastal forests of Kenya
119
0–400 m. Vouchers: SAJIT–006114 (EA, HIB), Magogo & Glover 798, Wright
13, Jeffery 267, Magogo FC & Glover PE 302 (EA).
Abutilon pannosum (G. Forst.) Schltdl. var. pannosum Habit: Herb. Habitat: Bushland, 30–100 m. Voucher: Festo L, Luke Q & P 2626 (EA).
Abutilon pannosum var. scabrum Verdc. Habit: Herb. Habitat: Bushland and thicket, 15–105 m. Vouchers: BL Simpson 283, Thairu 18, F Thomas 155 (EA).
Abutilon rotundifolium Mattei Habit: Herb. Habitat: On coral with Commiphora,
dunes, and calcareous sandstone, 0–5 m. Vouchers: Rawlins 117, Greenway &
Rawlins 9417, Gillespie 115, Luke Q 5473 (EA).
Abutilon sp. nr. wituense Baker f. Habit: Herb. Habitat: Hyphaene thicket and grassland, 30 m. Vouchers: Kabuye et al. TPR 44, 03A (EA).
Abutilon wituense Baker f. Habit: Herb. Habitat: Riverine gallery forest, thicket, and
dry sand rivers, 30–115 m. Vouchers: Luke & Robertson 2303, Kabuye et al. TPR
66, Thomas 10 (EA).
Adansonia digitata L. Habit: Tree. Habitat: Dry forest or bushland, ca. 250 m.
Vouchers: SAJIT–004671 & 005597 (EA, HIB), Kuchar P 11856 (EA).
Rhodognaphalon schumannianum A. Robyns Habit: Tree. Habitat: Woodland, forest, bushland, 20–700 m. Voucher: Magogo FC & Glover PE 675 (EA).
Byttneria fruticosa K. Schum. ex Engl. Habit: Tree. Habitat: Moist forest, especially
riverine habitats, 10–400 m. Vouchers: Luke WRQ & Robertson SA 2706, Faden
et al. 77/525, Luke 3303 (EA).
Carpodiptera africana Mast. Habit: Tree. Habitat: Evergreen thicket and dry evergreen mixed forest, 0–550 m. Vouchers: Salim bin Said 135, J Adamson 305,
Polhill & Paulo 757, Awanda 5 (EA).
Christiana africana DC. Habit: Tree. Habitat: Forest along river banks and mangrove forest, 0–300 m. Vouchers: Luke WRQ 900, Luke 2916, Luke & Robertson
542 (EA).
Cienfuegosia hildebrandtii Garcke Habit: Herb. Habitat: Commiphora-Acacia-Thespesia bushland, 100–450 m. Vouchers: Luke Q 1266, Q Jones 69/164, Kassner
439, Luke & Robertson 2138 (EA).
Cola minor Brenan Habit: Tree. Habitat: Coastal evergreen forest or thicket, 50–
250 m. Vouchers: Ngumbau V & Mwadime N V0533 (EA, HIB), Luke WRQ &
PA 4500, Langridge 1/57, Verdcourt 1863, Polhill & Robertson 4830, Simpson
BL 5 (EA).
Cola octoloboides Brenan Habit: Tree. Habitat: Evergreen forest on limestone, 200–
400 m. Vouchers: Faden et al. 70/943, Adams 125, Robertson et al. MDE 301,
Robertson SA & Luke WRQ 4798 (EA): Endangered, Endemic.
Cola porphyrantha Brenan Habit: Tree. Habitat: Evergreen forest, 45–950 m. Vouchers: Luke 1835, Magogo & Glover 1080, Brenan et al. 14557, Brenan JPM, Gillett
JB, Kanuri K & Chomba W 14562, Luke WRQ et al. 4604C (EA): Critically
Endangered.
Cola pseudoclavata Cheek Habit: Tree. Habitat: Riverine or coastal evergreen or semideciduous forest, 30–600 m. Vouchers: Ngumbau V & Mwadime N V0524 (EA,
120
Veronicah M. Ngumbau et al. / PhytoKeys 147: 1–191 (2020)
HIB), Medley 344, Mohamed Abdullah in FD 3353, Verdcourt 1855, Greenway
PJ & Rawlins SP 895 (EA).
Cola uloloma Brenan Habit: Tree. Habitat: Coastal forest, 30–500 m. Vouchers: Robertson & Luke 4787, Drummond & Hemsley 3972, Luke & Robertson 494, Luke
WRQ 2934 (EA).
Corchorus aestuans L. Habit: Herb. Habitat: Grassland on heavy alluvial soils, ca.
10–300 m. Vouchers: Ngumbau V & Mwadime N V0278 (EA, HIB), Drummond & Hemsley 1089, Gillett 18637, Gardner 2280, Sangai GW 976 (EA).
Corchorus baldaccii Mattei Habit: Herb. Habitat: Commiphora woodland, ca. 30 m.
Voucher: Polhill & Paulo 602 (EA).
Corchorus fascicularis Lam. Habit: Herb. Habitat: Seasonally flooded grassland or
wooded grassland, ca. 10 m. Voucher: Luke et al. in TPR 457 (EA).
Corchorus gillettii Bari Habit: Herb. Habitat: Open bushland on sand, 50–650 m.
Voucher: Gillet 16335 (EA).
Corchorus olitorius L. Habit: Herb. Habitat: Weed of cultivated area, 30–300 m.
Voucher: Magogo FC & Glover PE 1054 (EA).
Corchorus parviflorus (Benth.) Domin Habit: Herb. Habitat: Woodland, ca. 30 m.
Voucher: Dale IR 1154 (EA): Exotic.
Corchorus pseudoolitorius Islam & Zaid Habit: Herb. Habitat: Riverine vegetation,
10–600 m. Vouchers: Polhill & Paulo 829, Festo L & Luke Q 2449 (EA).
Corchorus schimperi Cufod. Habit: Herb. Habitat: Grassland on sand or wet clay.
Voucher: Mungai et al. 440/84 (EA).
Corchorus trilocularis L. Habit: Herb. Habitat: Grassland, roadsides, and disturbed
places, ca. 10 m. Vouchers: Festo L & Luke Q 2451, Magogo FC & Glover PE
869 (EA).
Dombeya kirkii Mast. Habit: Shrub. Habitat: Forest edge, Acacia bushland, bushland
on rocky slopes, ca. 600 m. Vouchers: Gillett 16843, Mwachala G 281 (EA).
Dombeya taylorii Baker f. Habit: Tree. Habitat: Thicket or wooded grassland, lowland evergreen forest, 0–500 m. Vouchers: Ngumbau V & Mwadime N V0532
(EA, HIB), Roberston & Luke 5826, Whyte s.n., Verdcourt 1901, Graham RM
350, Robertson SA & Luke Q 5826, Gray M & Luke WRQ 273 (EA).
Gossypioides kirkii (Mast.) J. B. Hutch. Habit: Liana. Habitat: Bushland, mixed
coastal forest, and Brachystegia woodland, 0–450 m. Vouchers: SAJIT–006014,
Ngumbau V & Mwadime N V0251 (EA, HIB), Verdcourt 1871, Polhill & Paulo
842, Obunyali & Omondi W 304 (EA).
Gossypium barbadense L. Habit: Shrub. Habitat: Gardens and experimental plots,
ca. 5 m. Voucher: Magogo & Glover 985 (EA): Cultivated.
Gossypium herbaceum L. Habit: Shrub. Habitat: Woodland, ca. 10 m. Voucher: Luke
Q 5486 (EA).
Gossypium hirsutum L. Habit: Shrub. Habitat: Cultivated in gardens and coconut
plantations, 0–150 m. Voucher: Greenway 10455 (EA): Cultivated.
Gossypium somalense (Gürke) J. B. Hutch., Silow & S. G. Stephens Habit: Shrub.
Habitat: Acacia-Commiphora thicket and Acacia-Commiphora-Delonix bushland,
(300–) 400–700 m. Voucher: Gillett 21101 (EA).
Checklist of coastal forests of Kenya
121
Grewia balensis Kirkup & Sebsebe Habit: Shrub. Habitat: Acacia bushland and coastal forest, 0–10 m. Vouchers: J Adamson in Bally 5973, Gillespie 211 & 218 (EA).
Grewia calymmatosepala K. Schum. Habit: Shrub. Habitat: Forest and coastal forest, ca. 50 m. Vouchers: SAJIT–005934, Ngumbau V & Mwadime N V068 (EA,
HIB), Drummond & Hemsley 3933 & 1123, Luke & Robertson 1925, Luke &
Robertson 1925 (EA).
Grewia capitellata Bojer Habit: Shrub. Habitat: Forest, forest edges, on coral rock,
and sand near the shoreline, 0–500 m. Vouchers: SAJIT–005526, Ngumbau V
& Mwadime N V0316 (EA, HIB), Luke & Robertson 1717 & 2716, Greenway
10842, Brenan, JPM, JH, Gillet JB et al. 14670 (EA).
Grewia damine Gaertn. Habit: Shrub. Habitat: Woodland and thicket, ca. 650 m.
Voucher: Kuchar P 11832 (EA).
Grewia densa K. Schum. Habit: Shrub. Habitat: Forest, including riverine, and Acacia-Commiphora bushland, 0–450 m. Vouchers: Dale 2036, Luke et al. in TPR
662, Gillett & Kihuwa 19991 (EA).
Grewia forbesii Harv. ex Mast. Habit: Liana. Habitat: Thicket, forest, woodland, and
secondary vegetation, ca. 237 m. Vouchers: Ngumbau V & Mwadime N V0175
(EA, HIB), Robertson SA 4237, Rawlins 782 (EA).
Grewia glandulosa Vahl Habit: Shrub. Habitat: Dry forest and bush on coral rag, 0–
20 m. Vouchers: SAJIT–005570 & 006119 (EA, HIB), Bridson 131, Bally & AR
Smith 14382, Greenway 10453 (EA).
Grewia holstii Burret Habit: Liana. Habitat: Forest, forest edges, clearings, thicket,
and bushland, 0–650 m. Vouchers: SAJIT–005497, Ngumbau V & Mwadime
N V052 (EA, HIB), Gillespie 202, Sjut & Ensor 2601, Drummond & Hemsley
3962, Mrs Robertson SA & Luke Q 5586 (EA).
Grewia microcarpa K. Schum. Habit: Shrub. Habitat: Woodland, thicket, AcaciaCommiphora bushland, and riverine vegetation, 0–700 m. Vouchers: RM Graham
1760, RB Faden et al. 77/462, Gisau 38 (EA).
Grewia pedunculata K. Schum. Habit: Shrub. Habitat: Dry bushland, wooded grassland, and dense thicket, 0–400 m. Vouchers: Drummond & Hemsley 4041, Verdcourt 5304, Luke & Robertson 1497, Robertson SA & Luke WRQ 6030, Luke
Q 1497 (EA).
Grewia plagiophylla K. Schum. Habit: Tree. Habitat: Forest, riverine thicket, coral
rag thicket, and Acacia-Commiphora bushland, 0–500 m. Vouchers: Ngumbau V
& Mwadime N V0192 (EA, HIB), Robertson 3543, Goyder et al. 4045, Robertson 4043, Kuchar P 13657 (EA).
Grewia praecox subsp. latiovata C. Whitehouse Habit: Shrub. Habitat: Coastal
bush forest, thorny bushland, and scattered tree grassland, 0–550 m. Vouchers:
RB & AJ Faden 72/89, Luke et al. in TPR 264, Luke et al. in TPR 422 (EA).
Grewia stuhlmannii K. Schum. Habit: Shrub. Habitat: Forest, forest edges, and
thicket, 0–500 m. Vouchers: Hooper & Townsend 1099, Drummond & Hemsley
3851, Medley 257, Greenway PJ in EAH 12202, Rawlins SP 352 (EA).
Grewia triflora (Bojer) Walp. Habit: Shrub. Habitat: Coastal forest, thicket, often
on coral rag, 0–30 (–100) m. Vouchers: SAJIT–006118 (EA, HIB), Drummond
122
Veronicah M. Ngumbau et al. / PhytoKeys 147: 1–191 (2020)
& Hemsley 3984, RB & AJ Faden 74/1256, Gillespie 71, Mrs Robertson SA
7756 (EA).
Grewia tristis K. Schum. Habit: Shrub. Habitat: Dry Acacia-Commiphora bushland,
0–650 m. Vouchers: RB & AJ Faden 74/1282, Drummond & Hemsley 4057 (EA).
Grewia truncata Mast. Habit: Shrub. Habitat: Thicket, bushland, and along river
courses, ca. 0–186 m. Vouchers: Ngumbau V & Mwadime N V0356 (EA, HIB),
Verdcourt 1872, Gillespie 88 (EA).
Grewia villosa Willd Habit: Shrub. Habitat: Woodland and bushland, ca. 0–59 m.
Vouchers: Ngumbau V & Mwadime N V0438 (EA, HIB), Gillespie 216 (EA).
Heritiera littoralis Aiton Habit: Tree. Habitat: Mangrove swamps and forest on coral rag,
0–5 m. Vouchers: Abdulla 1139, MacNaughton 9246, Faden & Faden 77/646 (EA).
Hermannia boranensis K. Schum. Habit: Herb. Habitat: Acacia-Commiphora bushland, 90–650 m. Voucher: Mungai 202 (EA).
Hermannia exappendiculata (Mast.) K. Schum. Habit: Herb. Habitat: Acacia-Commiphora bushland, roadsides, old cultivation, coastal forest, and grassland, ca. 5 m.
Vouchers: Gardner 1425, Gillett JB 203 (EA).
Hermannia fischeri K. Schum. Habit: Herb. Habitat: Bushland, 300–550 m.
Vouchers: Bally 16659, Drummond & Hemsley 4098, Tweedie 2570, Parker I
GM/523/H (EA).
Hermannia glanduligera K. Schum. ex Schinz Habit: Herb. Habitat: Road edges,
old cultivation, and edges of seasonally wet areas, ca. 30 m. Voucher: Greenway PJ
& Kanuri 12905 (EA).
Hermannia kirkii Mast. Habit: Herb. Habitat: Forest, ca. 225–552 m. Voucher: Gillett JB 21097 (EA).
Hermannia pseudofischeri Cheek Habit: Herb. Habitat: Dry bushland, ca. 150 m.
Vouchers: Njoroge Thairu 98, Linder 3658 (EA).
Hermannia uhligii Engl. Habit: Herb. Habitat: Acacia-Commiphora bushland or
grassland, ca. 30 m. Voucher: Luke et al. in TPR 218 (EA).
Hibiscus articulatus Hochst. ex A. Rich. Habit: Herb. Habitat: Woodland, grassland,
and along roadsides, ca. 152 m. Voucher: Magogo FC & Glover PE 986 (EA).
Hibiscus calyphyllus Cav. Habit: Shrub. Habitat: Thickets, roadsides in forests, riverine forests, and forest edges, ca. 30 m. Voucher: Festo L, Luke Q & P 2688 (EA).
Hibiscus cannabinus L. Habit: Herb. Habitat: Grassland. Voucher: Luke WRQ &
Robertson SA sr.
Hibiscus ceratophorus Thulin Habit: Herb. Habitat: Bushland, 0–200 m. Vouchers:
Mwadime N & Luke WRQ 2591 & 571 (EA).
Hibiscus faulknerae Vollesen Habit: Shrub. Habitat: Bushland, 0–300 m. Vouchers:
Ngumbau V & Mwadime N V0235 (EA, HIB), Drummond & Hemsley 3855, Magogo & Glover 797, Hooper & Townsend 1209, Festo L, Luke Q & P 2543 (EA).
Hibiscus greenwayi Baker f. Habit: Shrub. Habitat: Acacia-Commiphora bushland,
ca. 0–200 m. Vouchers: Polhill & Paulo 835, Verdcourt 3904 (EA).
Hibiscus hildebrandtii Sprague & Hutch. Habit: Herb. Habitat: Grassland and
bushland, 0–500 m. Vouchers: Drummond & Hemsley 4244, Adamson 52, Kabuye et al. in TPR 442, Festo L, Luke Q & P 2707, Graham RM 611 (EA).
Checklist of coastal forests of Kenya
123
Hibiscus holstii Mwachala Habit: Liana. Habitat: Bushland and forest on limestone,
70–500 m. Vouchers: Ngumbau V & Mwadime N V027 (EA, HIB), Luke &
Robertson 1666, 1858 & 2266 (EA): Vulnerable.
Hibiscus micranthus L. f. Habit: Herb. Habitat: Grassland and bushland, ca. 92 m.
Vouchers: SAJIT–006136 (EA, HIB), Greenway PJ 10849 (EA).
Hibiscus panduriformis Burm. f. Habit: Herb. Habitat: Seasonally flooded areas in
grassland and woodland, ca. 5 m. Vouchers: SAJIT–006102 (EA, HIB), Hooper
& Townsend 1235 (EA).
Hibiscus physaloides Guill. & Perr. Habit: Herb. Habitat: Woodland and abandoned
cultivations ca. 227 m. Vouchers: Ngumbau V & Mwadime N V0271 (EA, HIB),
Magogo & Estes 1202, Napier 6354, Luke WRQ et al. 4730 (EA).
Hibiscus rhabdotospermus Garcke Habit: Herb. Habitat: Sandy or stony ground and
among rocks on inselbergs, 300–700 m. Vouchers: van Someren 887, Mungai &
Rucina 490/84 (EA).
Hibiscus schizopetalus (Mast.) Hook. f. Habit: Shrub. Habitat: Coastal bushland
and forest, 0–200 m. Vouchers: Ngumbau V & Mwadime N V0384 (EA, HIB),
Jex-Blake s.n., Drummond & Hemsley 4256, Robertson & Luke 5811, Festo L,
Luke Q & P 2661, Luke WRQ & PA 4578 (EA).
Hibiscus sidiformis Baill. Habit: Herb. Habitat: Rocky areas in dry woodland, ca.
350 m. Voucher: Gilbert & Thulin 1738 (EA).
Hibiscus surattensis L. Habit: Herb. Habitat: Coastal habitats such as sand dunes, ca.
15 m. Voucher: Luke Q 5666 (EA).
Hibiscus sp. aff. lunariifolius Willd. Habit: Herb. Habitat: Forest, ca. 0–10 m.
Vouchers: SA Robertson 4247, Mwadime & Chesire 302 (EA).
Hibiscus tiliaceus L. Habit: Shrub. Habitat: Sandy seashores and mangrove swamps,
0–20 m. Vouchers: Graham 738, Jeffery 142, Abdulla 2197 (EA).
Hibiscus vitifolius L. Habit: Herb. Habitat: Acacia bushland, grassland, woodland,
and forest edges, ca. 620 m. Vouchers: Drummond & Hemsley 3989, Festo L,
Luke Q & P 2522 (EA).
Leptonychia usambarensis K. Schum. Habit: Tree. Habitat: Moist forest, 200–350 m.
Vouchers: Luke & Robertson 2721, Robertson SA & Luke WRQ 4605 (EA).
Melhania annua Thulin Habit: Herb. Habitat: Sand dunes and eroded sand dunes,
0–50 m. Vouchers: Faden & Faden 74/1107, Gillepie 75 (EA).
Melhania denhardtii K. Schum. Habit: Herb. Habitat: Open Acacia-Commiphora bushland, 5–250 m. Vouchers: Donald 2499, Graham 1641, Luke & Robertson 2524 (EA).
Melhania ovata (Cav.) Spreng. Habit: Herb. Habitat: Bushland, ca. 30 m. Voucher:
Reitsma J 293 (EA).
Melhania parviflora Chiov. Habit: Herb. Habitat: Acacia-Commiphora woodland
and coastal bushland, 0–850 m. Vouchers: Rawlins 800, Verdcourt 3950 (EA).
Melhania rotundata Hochst. ex Mast. Habit: Herb. Habitat: Open to dense AcaciaCommiphora woodland or bushland, ca. 100 m. Vouchers: Gillet 16437, Luke
WRQ & Robertson SA 2524 (EA).
Melhania velutina Forssk. Habit: Herb. Habitat: Wooded grassland, bushland, and
riverine forest, 0–700 m. Vouchers: Polhill & Paulo 874, Rawlins SP 82 (EA).
124
Veronicah M. Ngumbau et al. / PhytoKeys 147: 1–191 (2020)
Melochia corchorifolia L. Habit: Herb. Habitat: Roadsides, and grassland, 0–227 m.
Vouchers: Ngumbau V & Mwadime N V0273 (EA, HIB), Jeffery K518, Drummond & Hemsley 3928, Luke et al. TPR 522, Kirika P, Mbale M & Mbatha M
772, Magogo FC & Glover PE 160 (EA).
Melochia melissifolia Benth. Habit: Herb. Habitat: Swamps and swamp edges or less
frequently sandy soils, ca. 30 m. Vouchers: SAJIT–005994 & 006199 (EA, HIB),
Robertson & Luke 6345, Graham 1900, Luke & Luke 5995 (EA).
Nesogordonia holtzii (Engl.) Capuron ex L. C. Barnett & Dorr Habit: Tree. Habitat: Evergreen coastal forest, 1–500 m. Vouchers: Dale 1127, Luke & Robertson
2627, Barnett, Dorr & Check 518, Robertson SA & Luke WRQ 5526 (EA).
Pavonia arenaria var. microphylla (Ulbr.) Verdc. Habit: Herb. Habitat: Woodland,
bushland, flood plains, and rocky slopes with dense grass cover, ca. 0–150 m.
Voucher: Kabuye et al. TPR 219 (EA).
Pavonia blepharicarpa N.A. Brummitt & Vollesen Habit: Herb. Habitat: Wooded
grassland, 100–550 m. Voucher: Robertson 1815 (EA).
Pavonia elegans Garcke Habit: Herb. Habitat: Mixed dry evergreen bushland, ca.
90 m. Vouchers: Verdcourt 3903, JB Gillett 16420 (EA).
Pavonia ellenbeckii Gürke Habit: Herb. Habitat: Wooded grassland, ca. 100 m.
Voucher: Drummond & Hemsley 3761 (EA).
Pavonia leptocalyx (Sond.) Ulbr. Habit: Herb. Habitat: Marshy grassland, mixed
coastal bushland, and thicket, 5–300 m. Vouchers: SAJIT–005980, Ngumbau V
& Mwadime N V0181 (EA, HIB), Robertson & Luke 5995, RM Graham 805 in
FD 2353, Greenway 9261, Robertson SA & Luke WRQ 5186 (EA).
Pavonia sennii Chiov. Habit: Herb. Habitat: Woodland, 35–120 m. Voucher: Festo
L, Luke Q & P 2597 (EA).
Roifia dictyocarpa (Webb) Verdc. Habit: Herb. Habitat: Bushland with grassland
and floodplain bushland, 200–750 m. Voucher: Gillett & Gachathi 20543 (EA).
Senra incana Cav. Habit: Herb. Habitat: Bushland, scattered tree grassland, 35–
120 m. Vouchers: Thairu 92, Kabuye et al. TPR 744 (EA).
Sida acuta Burm. f. Habit: Herb. Habitat: Grassland, bushland, and forest edges, 0–150 m.
Vouchers: Drummond & Hemsley 4237, Magogo FC & Glover PE 660 (EA).
Sida chrysantha Ulbr. Habit: Herb. Habitat: Coastal grassland, wooded grassland, and
bushland, 0–100 m. Vouchers: Rawlins 767, Drummond & Hemsley 4240 (EA).
Sida cordifolia L. Habit: Herb. Habitat: Very mixed bushland and thicket, evergreen
forest, and cultivated area, 0–450 m. Vouchers: Drummond & Hemsley 4236 &
4240, MacNaughton 93, Rawlins 175 & 767 (EA).
Sida ovata Forssk. Habit: Herb. Habitat: Dry areas ca. 5–15 m. Vouchers: Luke
WRQ et al. 5706, Robertson SA 6848 (EA).
Sida rhombifolia var. serratifolia (R. Wilczek & Steyaert) Verdc. Habit: Herb.
Habitat: Forest margin, and scrub, 0–200 m. Voucher: Robertson SA 3439 (EA).
Sida spinosa L. Habit: Herb. Habitat: Wastelands, ca. 10 m. Voucher: Festo L & Luke
Q 2448 (EA).
Sida tanaensis Vollesen Habit: Herb. Habitat: Open bushland and scattered tree
grassland, 30–250 m. Voucher: Kabuye et al. TPR 346 (EA).
Checklist of coastal forests of Kenya
125
Sterculia appendiculata K. Schum Habit: Tree. Habitat: Coastal and lowland riverine, 5–750 m. Vouchers: Ngumbau V & Mwadime N V0476 (EA, HIB), Gillet
& Kibuwa 19948, Bally 2036, Dale 3772, Birch WH s.n., Mulwa PCW 60 (EA).
Sterculia foetida L. Habit: Tree. Habitat: Lowland dry woodlands, 0–100 m. Voucher: Robertson SA 3662 (EA): Cultivated.
Sterculia rhynchocarpa K. Schum Habit: Tree. Habitat: Woodland and bushland,
ca. 10–70 m. Vouchers: Ngumbau V & Mwadime N V0446 (EA, HIB), Gathii S
142, Wamukoya K7–x62a (EA).
Sterculia schliebenii Mildbr. Habit: Tree. Habitat: Coastal evergreen forest, 20–
400 m. Vouchers: Luke 896, Faden, Evans & Mahasi 70/826, Luke & Robertson
1739, Luke WRQ & Robertson SA 534, Hawthorne W 276 (EA).
Thespesia danis Oliv. Habit: Shrub. Habitat: Riverine grassland, coastal bushland, and
thicket, 0–450 m. Vouchers: SAJIT–006449, Ngumbau V & Mwadime N V0129
(EA, HIB), Verdcourt 3217, Tweedie 1007, Magogo FC & Glover PE 700 (EA).
Thespesia populnea (L.) Corrêa Habit: Shrub. Habitat: Mangrove swamps, 0–6 m.
Vouchers: SAJIT–006249 (EA, HIB), Thomas s.n., Greenway & Rawlins 9484 (EA).
Triumfetta flavescens Hochst. ex A. Rich. var. flavescens Corrêa Habit: Shrub. Habitat: Open bushland, 225–552 m. Vouchers: P Kuchar, F Msafiri & R Seet 5840,
J Mutanga & J Muasya 261, JB Gillett 21114 (EA).
Triumfetta heterocarpa Sprague & Hutch. Habit: Herb. Habitat: Acacia-Commiphora bushland on sandy, silt or laterite soils, ca. 30 m. Voucher: GC van Someren
880 (EA).
Triumfetta rhomboidea Jacq. Habit: Herb. Habitat: Grassland, ca. 95 m. Vouchers:
SAJIT–006141 (EA, HIB), Magogo FC & Glover PE 455 (EA).
Triumfetta tomentosa Bojer Habit: Herb. Habitat: Forests, swamp margins, and
bushland, ca. 60 m. Vouchers: Magogo FC & Estes R 1191, Nyange M 510 (EA).
Urena procumbens L. Habit: Herb. Habitat: Roadsides and waste places, 0–200 m.
Voucher: SAJIT–006196 (EA, HIB).
Urena lobata var. sinuata (L.) Borss. Waalk. Habit: Herb. Habitat: Damp grassland
and areas of cultivations, 0–15 m. Voucher: Luke 2915 (EA).
Waltheria indica L. Habit: Herb. Habitat: Dry grassland, cultivated, and disturbed
places, 0–300 m. Vouchers: SAJIT–006128 (EA, HIB), Katz SS KNMUS/
H197/75, Magogo FC & Glover PE 31 (EA).
F108. Melastomataceae
7 Genera, 12 Species
Antherotoma debilis (Sond.) Jacques-Fél. Habit: Herb. Habitat: Valley grassland, ca.
10 m. Voucher: RM Graham LL 731 in FD 2183 (EA).
Antherotoma senegambiensis (Guill. & Perr.) Jacques-Fél. Habit: Herb. Habitat:
Along river banks and forest margins. Voucher: Reitsma J 239 (EA).
Clidemia hirta (L.) D. Don Habit: Shrub. Habitat: Moist or wet mixed forest and
thickets, ca. 290 m. Vouchers: SAJIT–005504 (EA, HIB), Luke WRQ & Robertson SA 2712 (EA).
126
Veronicah M. Ngumbau et al. / PhytoKeys 147: 1–191 (2020)
Heterotis rotundifolia (Sm.) Jacq.-Fél. Habit: Herb. Habitat: Margins of rain forest,
riverine forest, flood plains, and valley grassland, ca. 0–426 m. Vouchers: Ngumbau V & Mwadime N V083 & 0264 (EA, HIB), RM Graham in FD 2189, Bally
8898, Purse-glove 1727, Rawlins SP 342, Magogo FC & Glover PE 112 (EA).
Memecylon fragrans A. Fern. & R. Fern. Habit: Shrub. Habitat: Coastal dry evergreen forest, 60–100 m. Vouchers: SAJIT–005976 & 006161, Ngumbau V &
Mwadime N V0239 (EA, HIB), Drummond & Hemsley 3859, Verdcourt 1180,
Perdue & Kibuwa 10010, Jeffery GW 616 (EA).
Memecylon verruculosum Brenan Habit: Shrub. Habitat: Riverine forest and lowland
evergreen forest, 300–750 m. Vouchers: Luke WRQ 2939, Gardner in FD 1447,
Drummond & Hemsley 3971, Magogo & Glover 237 (EA): Vulnerable.
Tristemma mauritianum J. F. Gmel. Habit: Shrub. Habitat: Marshy clearings in
rainforest and swampy riverine forest, ca. 290 m. Vouchers: Glover & Etes 1153,
Luke WRQ & Robertson SA 2715 (EA).
Warneckea amaniensis Gilg Habit: Shrub. Habitat: Lowland rainforest and riverine
forest, 40–600 m. Vouchers: SAJIT–006068 (EA, HIB), Gardner in FD 1409,
Drummond & Hemsley 3931, Moomaw 1276, Brenan JPM & JH GilletJB
14593, Stone RD & Luke WRQ 2415 (EA).
Warneckea hedbergiorum Borhidi Habit: Shrub. Habitat: Moist forest, ca. 250 m.
Vouchers: Ngumbau V & Mwadime N V0238 (EA, HIB), Stone RD & Luke
WRQ 2416 (EA).
Warneckea mouririifolia (Brenan) Borhidi Habit: Shrub. Habitat: Lowland dry evergreen forest and rainforest, 45–100 m. Vouchers: SAJIT–005924 (EA, HIB),
Drummond & Hemsley 3857, RM Graham in FD 1527, Polhill & Paulo 852,
Luke Q 1501, Luke WRQ & Robertson SA 2805, Festo L, Luke PA & WRQ
2723 (EA): Vulnerable.
Warneckea melindensis (A. Fern. & R. Fern.) R. D. Stone & Q. Luke Habit: Shrub.
Habitat: Coastal forest and thicket, 10–100 m. Vouchers: Dale IR 3835, Luke
WRQ 4574A, Graham 1951, Jeffery 570, Faden 71/751 (EA).
Warneckea sansibarica (Taub.) Jacq.-Fél. subsp. sansibarica Habit: Shrub. Habitat: Lowland dry evergreen forest and deciduous woodland, 10–100 m. Vouchers:
Donald 1 in FD 2327, Trump 1361, Gisau in EAH 10985, Robertson SA & Abio,
LG 6815, Luke WRQ & Robertson SA 2783, Gillett JB 20425, Robertson SA &
Luke WRQ 5479 (EA).
Warneckea sansibarica subsp. buchananii (Gilg) Borhidi Habit: Tree. Habitat:
Forest, ca. 30 m. Vouchers: Mohamed Abdullah 3351, Perdue RE & Kibuwa SP
10019 (EA).
F109. Meliaceae
9 Genera, 16 Species
Azadirachta indica A. Juss. Habit: Tree. Habitat: Dry Habitat, ca. 0–20 m. Voucher:
Hume L 10 (EA): Cultivated.
Checklist of coastal forests of Kenya
127
Ekebergia capensis Sparrm. Habit: Tree. Habitat: Variable habitats, ca. 10 m. Voucher: Luke Q 1514 (EA).
Lovoa swynnertonii Baker f. Habit: Tree. Habitat: Lowland and mid altitude rainforest, ca. 180 m. Vouchers: Verdcourt 3936a, White 11334, Luke WRQ & Robertson SA 2731 (EA): Near Threatened.
Melia azedarach L. Habit: Tree. Habitat: Secondary grassland, thicket, and waste
ground, ca. 100 m. Voucher: Perdue & Kibuwa 10118 (EA): Cultivated.
Pseudobersama mossambicensis (Sim) Verdc. Habit: Tree. Habitat: Understorey and
edges of moist lowland forest, 60–300 m. Vouchers: SAJIT–005519, 005553 &
005972, Ngumbau V & Mwadime N V085 (EA, HIB), White 11319, Greenway
9814, Trump 141, Robertson SA 4529 & 3460 (EA): Near Threatened.
Toona ciliata M. Roem. Habit: Tree. Habitat: Thickets, forests, ca. 400 m. Voucher:
Lennox K 10566 (EA): Cultivated.
Trichilia emetica Vahl Habit: Tree. Habitat: Coastal forest, drier types of riparian forest, and wooded grassland, ca. 10–407 m. Vouchers: SAJIT–006042 (EA, HIB),
White 11399, Sangai GW 971, Robertson SA 4238A (EA).
Turraea floribunda Hochst. Habit: Shrub. Habitat: Evergreen forest and riverine
forest, ca. 100–237 m. Vouchers: SAJIT–005509 & 005583, Ngumbau V &
Mwadime N V0169 (EA, HIB), Brenan et al. 14622, Battiscombe 111, Robertson
SA & Luke WRQ 5493, Luke WRQ & Robertson SA 2772 (EA).
Turraea holstii Gürke Habit: Tree. Habitat: Moist to dry forest, ca. 99 m. Voucher:
Kimeu JM 645 (EA).
Turraea mombassana Hiern ex C. DC. subsp. mombassana Habit: Shrub. Habitat: Evergreen forest, coastal scrub, and secondary forest, ca. 0–400 m. Vouchers: SAJIT–005925 & 005548, Ngumbau V & Mwadime N V0188 (EA, HIB),
Drummond & Hemsley 1108, RM Graham 1947, Rawlins 230, Luke WRQ &
Robertson SA 502, Rawlins SP 11261 (EA).
Turraea nilotica Kotschy & Peyr. Habit: Tree. Habitat: Woodland, bushland, and
wooded grassland, ca. 100 m. Vouchers: Luke Q 5641, Magogo FC & Glover PE
96, Greenway & Rawlins 8905 (EA).
Turraea parvifolia Deflers Habit: Shrub. Habitat: Acacia-Commiphora bushland and
banks of seasonal watercourses, ca. 260 m. Voucher: Hemming 83/42 (EA).
Turraea robusta Gürke Habit: Tree. Habitat: Thickets and Brachystagia woodland, ca.
30 m. Voucher: Jeffery GW 446 (EA).
Turraea wakefieldii Oliv. Habit: Tree. Habitat: Coastal forest and secondary forest,
near sea level. Vouchers: Ngumbau V & Mwadime N V0201 (EA, HIB), Drummond & Hemsley 4187, Luke Q 1402, Robertson SA 5069 (EA).
Xylocarpus granatum J. Koenig Habit: Tree. Habitat: In tidal mud of mangrove
swamps. Vouchers: RM Graham in FD 2236, Sangai 849, Greenway & Rawlins
8869 (EA).
Xylocarpus moluccensis (Lam.) M. Roem. Habit: Tree. Habitat: In coastal scrub,
above high-water mark. Vouchers: Kibuwa 1203, RB & AJ Faden 77/553, Rawlins
144, Kuchar P 13584 (EA).
128
Veronicah M. Ngumbau et al. / PhytoKeys 147: 1–191 (2020)
F110. Menispermaceae
9 Genera, 11 Species
Albertisia undulata (Hiern) Forman Habit: Liana. Habitat: Dry thicket, 50–450 m.
Vouchers: Luke WRQ & Robertson SA 2744 & 1745, NMK Mrima/Dzombo
Exp. 218 & 311 (EA).
Anisocycla blepharosepala subsp. tanzaniensis Vollesen Habit: Liana. Habitat: Bushland, 1–150 m. Vouchers: Robertson SA 5485, Festo L, Luke Q & P 2660 (EA).
Chasmanthera dependens Hochst. Habit: Liana. Habitat: Rainforest, regrowth, and
fringing forest, ca. 600 m. Voucher: Festo L & Luke Q 2559 (EA).
Cissampelos nigrescens Diels var. nigrescens Habit: Climber. Habitat: Moist forest,
ca. 15 m. Vouchers: SAJIT–006218 (EA, HIB), Luke WRQ 3125 (EA): Vulnerable.
Cissampelos pareira L. Habit: Climber. Habitat: Forest, deciduous woodland,
and secondary vegetation, ca. 15 m. Vouchers: SAJIT–006142, Ngumbau V &
Mwadime N V0263 (EA, HIB), Luke Q 1452, Robertson SA & Luke WRQ
6035, Jeffery K360 (EA).
Dioscoreophyllum volkensii Engl. Habit: Climber. Habitat: Lowland rainforest,
0–15 m. Vouchers: SAJIT–005538 (EA, HIB), Faden RB 69/489, Drummond &
Hemsley 4037 (EA).
Jateorhiza palmata (Lam.) Miers Habit: Climber. Habitat: Lowland rainforest
and riverine forest, 0–231 m. Vouchers: Ngumbau V & Mwadime N V0331
(EA, HIB), Kirika P, Mbale M & Mbatha M 770, Faden RB 70/430, Graham
1797 (EA).
Tiliacora funifera (Miers) Oliv. Habit: Liana. Habitat: Riverine, evergreen forest,
and moist areas in woodland, ca. 20 m. Vouchers: Mwadime N 17, Robertson SA
& Luke WRQ 5169 (EA).
Tinospora caffra (Miers) Troupin Habit: Climber. Habitat: Lowland, upland rainforest, deciduous bushland, and often near rock outcrops, ca. 350 m. Vouchers:
Robertson SA & Luke WRQ 5978, Drummond & Hemsley 4155 (EA).
Tinospora oblongifolia (Engl.) Troupin Habit: Liana. Habitat: Lowland rainforest
and coastal evergreen bushland, 0–304 m. Vouchers: Ngumbau V & Mwadime N
V0523 (EA, HIB), Drummond & Hemsley 3905 & 3244, van Someren Sh 115,
Robertson SA & Luke WRQ 1410, (EA).
Triclisia sacleuxii (Pierre) Diels Habit: Liana. Habitat: Evergreen forest, 50–500 m.
Vouchers: Ngumbau V & Mwadime N V0474 (EA, HIB), Mahasi AS EA 14893,
Mwadime N 51 (EA).
F111. Menyanthaceae
1 Genus, 1 Species
Nymphoides forbesiana (Griseb.) Kuntze Habit: Herb. Habitat: Shallow water at
the edge of lakes, pools, and rivers, ca. 0–30 m. Vouchers: Gillett 18725, Sacleux
2306, Whyte 40 (EA).
Checklist of coastal forests of Kenya
129
F112. Metteniusaceae
2 Genera, 2 Species
Apodytes dimidiata E. Mey. ex Arn. Habit: Tree. Habitat: Widespread from coastal
evergreen bushland to upland rainforest, ca. 0–200 m. Vouchers: SAJIT–006145
(EA, HIB), Luke Q 1427, Robertson SA 4218, Polhill & Paulo 820 (EA).
Rhaphiostylis beninensis (Hook. f. ex Planch.) Planch. ex Benth. Habit: Liana.
Habitat: Lowland rainforests. Voucher: Luke WRQ 8327B (EA).
F113. Molluginaceae
2 Genera, 4 Species
Glinus oppositifolius (L.) Aug. DC. Habit: Herb. Habitat: Dry regions, seasonally
swampy, and dry river beds, ca. 40 m. Vouchers: SAJIT–006212 (EA, HIB), Luke
Q 1425, Rawlins SP 289, Jeffery K466 (EA).
Glinus lotoides L. Habit: Herb. Habitat: Woodlands, mixed savannah woodland,
grassland, riverine forest, and river banks ca. 15 m. Voucher: Reitsma J 413 (EA).
Glinus setiflorus Forssk. Habit: Herb. Habitat: Dry river bed, bushland, and woodland, ca. 90 m. Vouchers: Greenway 9236, JB Gillett 16414 (EA).
Hypertelis umbellata (Forsk.) Thulin Habit: Herb. Habitat: Roadsides, cultivated
areas. Vouchers: Townsend CC & Hooper SS 1224, Jeffery K607 (EA).
F114. Montiniaceae
1 Genus, 1 Species
Grevea eggelingii var. keniensis (Verdc.) Verdc. Habit: Shrub. Habitat: Evergreen
forest, near sea-level. Vouchers: SAJIT–006089 & 005444 (EA, HIB), Napier
3348 (EA): Endangered, Endemic.
F115. Moraceae
8 Genera, 34 Species
Antiaris toxicaria var. subsp. welwitschii (Engl.) C.C. Berg usambarensis (Engl.)
C. C. Berg Habit: Tree. Habitat: Rainforest, evergreen forests, and riverine, ca.
300 m. Voucher: Verdcourt 3937 (EA).
Dorstenia alta Engl. Habit: Shrub. Habitat: Undershrub in evergreen forest and near
streams, up to 800 m. Vouchers: Ngumbau V & Mwadime N V0400 (EA, HIB),
Luke WRQ & Robertson SA 2695, Drummond & Hemsley 1199, Faden et al.
70/944, B Adams 103 (EA).
Dorstenia barnimiana Schweinf. Habit: Herb. Habitat: Wooded and open grassland, ca. 300 m. Voucher: Luke WRQ & Robertson SA 2560 (EA).
Dorstenia cuspidata var. brinkmaniana Hijman Habit: Herb. Habitat: Deciduous,
coastal bushland, thickets, wooded grassland, and along rivers, ca. 250 m. Vouchers: Mason in EAH 15404, Luke 1577 (EA).
130
Veronicah M. Ngumbau et al. / PhytoKeys 147: 1–191 (2020)
Dorstenia foetida (Forssk.) Schweinf. Habit: Herb. Habitat: Rock outcrops and
open places in deciduous, and succulent bushland, ca. 100 m. Voucher: Faden
74/1017 (EA).
Dorstenia goetzei Engl. Habit: Herb. Habitat: Forest in wet shaded places and often
among rocks, ca. 200 m. Vouchers: Luke WRQ & Robertson SA 1893, Faden
71/790, B Adams 82 (EA): Near Threatened.
Dorstenia hildebrandtii Engl. Habit: Herb. Habitat: Open forest to woodland, bushland, and succulent thickets, ca. 0–330 m. Vouchers: Drummond & Hemsley
4254, Luke WRQ & Robertson SA 1464 (EA).
Dorstenia kameruniana Engl. Habit: Shrub. Habitat: Undergrowth of evergreen forests, often near streams, ca. 231 m. Vouchers: SAJIT–005528, Ngumbau V &
Mwadime N V112 (EA, HIB), Dale in FD 3545, Drummond & Hemsley 1154,
Faden et al. 70/947, Robertson SA 5083 (EA).
Dorstenia tayloriana Rendle var. tayloriana Habit: Herb. Habitat: Lowland evergreen forest, up to 600 m. Vouchers: Ngumbau V & Mwadime N V0312 &
V0475 (EA, HIB), WE Taylori, Dale IR 3827 (EA): Vulnerable.
Dorstenia tayloriana var. laikipiensis (Rendle) Hijman Habit: Herb. Habitat: Evergreen forest and coastal forests, ca. 240 m. Vouchers: SAJIT–005530, Ngumbau V
& Mwadime N V0108 (EA, HIB), Gardner in FD 1440, Drummond & Hemsley
3959, Faden 74/1271, Faden RB 70/427, Malombe I, Mwandime N & Saidi
1663, Luke PA & WRQ 5980 (EA).
Dorstenia warneckei Engl. Habit: Herb. Habitat: Forest in shady and sometimes wet
places, 200–750 m. Vouchers: Glover et al. 1155, Luke WRQ & Robertson SA
471, Luke WRQ 1644 (EA): Near Threatened.
Ficus bubu Warb. Habit: Tree. Habitat: Forest, riverine, lakesides, and other ground water forest, ca. 100 m. Vouchers: Dale 2014, Luke WRQ & Robertson SA 2788 (EA).
Ficus craterostoma Warb. ex Mildbr. & Burret Habit: Tree. Habitat: Along the margins of evergreen forest, ca. 36 m. Voucher: SAJIT–006190 (EA, HIB).
Ficus exasperata Vahl Habit: Tree. Habitat: Forest, often at edges, in rocky places, and
along rivers, ca. 200 m. Vouchers: Ngumbau V & Mwadime N V0403 (EA, HIB),
Gillett 18636, BR Adams 102, Robertson SA 4239 (EA).
Ficus faulkneriana C.C. Berg Habit: Tree. Habitat: Coastal bushland and wooded
grassland, 0–450 m. Voucher: Magogo & Glover 51 (EA): Vulnerable.
Ficus ingens (Miq.) Miq. Habit: Tree. Habitat: Wooded savannah, sometimes in
rocky places. Voucher: Robertson SA 5216 (EA).
Ficus lingua subsp. depauperata (Sim) C.C. Berg Habit: Tree. Habitat: Forest,
coastal bushland, and coral outcrops, 0–50 m. Vouchers: Gillet & Kibuwa 19904,
Perdue & Kibuwa 10103, Greenway & Rawlins 9346 (EA).
Ficus lutea Vahl Habit: Tree. Habitat: Deciduous woodland, wooded grassland, and
forest. Voucher: Robertson SA & Luke WRQ 5145 (EA).
Ficus natalensis Hochst. Habit: Tree. Habitat: Riverine and coastal forests or swamp forest, ca. 200 m. Vouchers: Luke WRQ & Robertson SA 190, Robertson SA 5068 (EA).
Checklist of coastal forests of Kenya
131
Ficus ottoniifolia subsp. ulugurensis (Mildbr. & Burret) C.C. Berg Habit: Tree.
Habitat: Rainforest, riverine, and coastal bushland, ca. 100 m. Vouchers: Verdcourt 3949, BR Adams 116, Luke WRQ 880 (EA).
Ficus polita Vahl Habit: Tree. Habitat: Evergreen forests, bushland, semi-deciduous
forest, and coastal rainforest. Voucher: Luke WRQ & PA sr.
Ficus recurvata De Wild. Habit: Tree. Habitat: Lowland forest, riverine, swampy forest,
and flood plains, 0–550 m. Vouchers: Jeffery 497, Magogo FC & Glover PE 69 (EA).
Ficus sansibarica Warb. subsp. sansibarica Habit: Tree. Habitat: Evergreen forest,
including drier types, and coastal bushland, ca. 0–250 m. Vouchers: Bally & Smith
14397, Dale 2008, Verdcourt 2409, Luke Q 1428, Gray M & Luke WRQ 290,
Mulwa PCW 57 (EA).
Ficus scassellatii Pamp. subsp. scassellatii Habit: Tree. Habitat: Rainforest, lakesides, riverine, and ground water forest, ca. 300 m. Vouchers: Polhill & Paulo 807,
Saufferer S 896 (EA).
Ficus stuhlmannii Warb. Habit: Tree. Habitat: Wooded grassland, bushland, and often
along lakesides, ca. 0–400 m. Vouchers: Ngumbau V & Mwadime N V0539 (EA,
HIB), Robertson SA & Luke WRQ 5863, Pakia M 979, Dale in FD 3634 (EA).
Ficus sur Forssk. Habit: Tree. Habitat: Woodland, riverine, and semi-deciduous forest, ca. 150 m. Vouchers: Brenan JPM, Gillett JB 14573, Luke WRQ & Robertson
SA 249 (EA).
Ficus sycomorus L. Habit: Tree. Habitat: Forest edges, lakesides, riverine, and rock outcrops, ca. 0–250 m. Vouchers: Greenway & Rawlins 9362, Mulwa PCW 13 (EA).
Ficus tremula Warb. subsp. tremula Habit: Tree. Habitat: Lowland dry evergreen
forest, woodland, and coastal bushland, 0–600 m. Vouchers: Greenway 9800, Verdcourt 3288, Gillett & Kibuwa 19893 (EA).
Ficus usambarensis Warb. Habit: Tree. Habitat: Woodland, ca. 500 m. Vouchers:
Ngumbau V & Mwadime N V0240 (EA, HIB), Mulwa PCW 57 (EA).
Ficus wakefieldii Hutch. Habit: Tree. Habitat: Riverine, lakesides, and wooded grassland, ca. 230 m. Vouchers: Jembe SM & Morimoto Y 47, Robertson SA & Luke
WRQ 4469 (EA).
Maclura africana (Bureau) Corner Habit: Tree. Habitat: Bushland and dry evergreen forest, 0–720 m. Vouchers: Bally 8908, Drummond & Hemsley 3922, RM
Graham in FD 1953, Luke Q 1444, Wamukoya K7x57a (EA).
Milicia excelsa (Welw.) C.C. Berg Habit: Tree. Habitat: Rainforest, lowland evergreen forest, riverine, and ground water forest, ca. 0–300 m. Vouchers: Dale in FD
1072, CW Elliot 212, Robertson SA 4221 (EA).
Morus mesozygia Stapf Habit: Tree. Habitat: Semi deciduous forests, ca. 20 m.
Voucher: Thomas Mwadime in Mrs Robertson SA 7785 (EA).
Streblus usambarensis (Engl.) C.C.Berg Habit: Shrub. Habitat: Forest and often riverine, 30–700 m. Vouchers: SAJIT–006187 & 005534 (EA, HIB), Brenan, Gillett
et al. in EAH 14534, Faden, Evans & Msafiri 70/950, Gilbert & May 5341, Luke
WRQ & Robertson SA 528 (EA).
132
Veronicah M. Ngumbau et al. / PhytoKeys 147: 1–191 (2020)
Trilepisium madagascariense DC. Habit: Tree. Habitat: Rainforest, wetter evergreen
forests, riverine, and ground water forest, ca. 100 m. Voucher: RM Graham in FD
1643 (EA).
F116. Moringaceae
1 Genus, 1 Species
Moringa oleifera Lam. Habit: Tree. Habitat: Cultivations, grassland on black clay,
and bushland on coral, ca. 15 m. Voucher: Verdcourt B 1102 (EA): Naturalized.
F117. Myristicaceae
1 Genus, 1 Species
Cephalosphaera usambarensis (Warb.) Warb. Habit: Tree. Habitat: Rainforest on
steep slopes, also lowland streamside forest, ca. 200 m. Vouchers: Luke & Robertson 2704A & 2704B (EA): Vulnerable.
F118. Myrtaceae
2 Genera, 7 Species
Eugenia capensis subsp. multiflora Verdc. Habit: Shrub. Habitat: Dry evergreen forest, bushland, coastal thicket, and mangrove swamp edges, 0–350 m. Vouchers:
Robertson & Luke 5165 & 5640, Sampson 30 (EA).
Eugenia nigerina A. Chev. Habit: Shrub. Habitat: Thicket and river beds, 18–360 m.
Vouchers: Drummond & Hemsley 4143, Robertson & Luke 6083 & 5325, Festo
L & Luke WRQ 2537 (EA).
Eugenia tanaensis Verdc. Habit: Shrub. Habitat: On rocks at high water mark of
river, (5–) 800 m. Vouchers: Drummond & Hemsley 4260, Robertson & Luke
5813 (EA).
Eugenia verdcourtii Byng Habit: Shrub. Habitat: Dry evergreen forest and bushland, coastal thicket, mangrove swamp edges, flood plains, 0–350 m. Vouchers: Ngumbau V & Mwadime N V0317 (EA, HIB), Luke Q 1439, Verdcourt
5283 (EA).
Syzygium cordatum subsp. shimbaense Verdc. Habit: Tree. Habitat: Forest edges,
grassland, and scattered trees, (100–) 300–450 m. Vouchers: SAJIT–005557 (EA,
HIB), RM Graham Q251 in FD 1775, Brenan et al. 14559, Magogo & Glover
104, Luke WRQ & PA 5735 (EA).
Syzygium cumini (L.) Skeels Habit: Tree. Habitat: Coastal bushland and savannahs
on coral rock, ca. 200 m. Vouchers: Perdue & Kibuwa 10115, Magogo FC &
Glover PE 859 (EA): Cultivated.
Syzygium guineense (Willd.) DC. ?subsp. nov, Habit: Tree. Habitat: Forest. Voucher: Luke WRQ & PA 4511 (EA).
Syzygium guineense subsp. afromontanum F. White Habit: Tree. Habitat: Evergreen
forest, ca. 100 m. Voucher: Luke WRQ & Robertson SA 2683 (EA).
Checklist of coastal forests of Kenya
133
F119. Nyctaginaceae
2 Genera, 6 Species
Boerhavia coccinea Mill. Habit: Herb. Habitat: Disturbed, rocky, and swampy
ground, ca. 0–10 m. Vouchers: Tweedie 933, Frazier J 2221 (EA).
Boerhavia diffusa L. Habit: Herb. Habitat: Disturbed ground in cultivation, waste
places, and grassland, ca. 65 m. Vouchers: SAJIT–005592 (EA, HIB), Moomaw
1417, Polhill & Paulo 534, Bally 13268 (EA).
Boerhavia erecta L. Habit: Herb. Habitat: Disturbed sandy, bushland, waste ground,
cultivation, and on roadsides, ca. 213 m. Vouchers: SA Robertson 3335, Gerhardt
K & Steiner M 44, Magogo FC & Glover PE 725 (EA).
Boerhavia sp A of FTEA Habit: Herb. Habitat: Bushland and forest, ca. 10 m.
Voucher: Luke Q 5463 (EA): Endemic.
Pisonia aculeata L. Habit: Liana. Habitat: Found along coasts, hedges, rain forests,
and open forests, ca. 20 m. Voucher: Mwadime N 9 (EA).
Pisonia grandis R. Br. Habit: Shrub. Habitat: Coral outcrops along the coast, shoreline. Vouchers: Frazier 2251 & 2252 (EA).
F120. Nymphaeaceae
1 Genus, 2 Species
Nymphaea lotus L. Habit: Herb. Habitat: Sheltered water, ca. 200 m. Vouchers: Verdcourt 3195, Festo L, Luke Q & P 2678 (EA).
Nymphaea nouchali var. caerulea (Savigny) Verdc. Habit: Herb. Habitat: Pools or
lake edges, also swamps, seasonal ponds, and waterholes, ca. 0–700 m. Vouchers:
Bally 8558, Gillett JB 20426 (EA).
Nymphaea nouchali var. zanzibariensis (Casp.) Verdc. Habit: Herb. Habitat: Lowland streams, also ponds on granitic rocks, ca. 0–650 m. Vouchers: SAJIT–005474
& 006231 (EA, HIB), Magogo & Glover 578, Magogo & Glover 155, Hildebrandt 1921 (EA).
F121. Ochnaceae
4 Genera, 12 Species
Brackenridgea zanguebarica Oliv. Habit: Shrub. Habitat: Coastal bushland, wooded
grassland, Brachystegia woodland, and lowland Julbernadia evergreen forest, ca.
0–200 m. Vouchers: Ngumbau V & Mwadime N V0499 & V095 (EA, HIB),
Greenway 9802, Drummond & Hemsley 1107, Hildebrandt 1966, Reitsma J
225, Robertson SA 5071 (EA).
Campylospermum reticulatum (P. Beauv.) Farron Habit: Shrub. Habitat: Evergreen
forest, rainforest, and riverine forest, ca. 300 m. Vouchers: SAJIT–006065 (EA,
HIB), Magogo & Glover 898, Luke & Robertson 2736, Robertson 7392 (EA).
Campylospermum sacleuxii (Tiegh.) Farron Habit: Shrub. Habitat: Lowland evergreen forest, riverine forest, ground water forest, and semi-dry forest, 40–750 m.
134
Veronicah M. Ngumbau et al. / PhytoKeys 147: 1–191 (2020)
Vouchers: Spjut & Ensor 2728, Magogo & Glover 166, Luke 4542, Luke WRQ
et al. 4716 (EA).
Ochna apetala Verdc. Habit: Shrub. Habitat: Lowland rainforest, deciduous forest,
and sometimes riverine, 5–200 m. Vouchers: Ngumbau V & Mwadime N V091
(EA, HIB), Robertson & Luke 6348, Luke & Robertson 1674 & 2639 (EA).
Ochna atropurpurea DC. Habit: Shrub. Habitat: Various mixed thicket and bushland, 0–450 m. Vouchers: SAJIT–006041, Ngumbau V & Mwadime N V0276
(EA, HIB), Verdcourt 2347, Drummond & Hemsley 1013, Polhill & Paulo 592,
Roberts SA & Luke Q 5584, Luke WRQ & Robertson SA 531 (EA).
Ochna holstii Engl. Habit: Shrub. Habitat: Understorey of evergreen forest and
among rocks, ca. 480 m. Vouchers: SAJIT–005985 (EA, HIB), Luke WRQ &
Robertson SA 1010 (EA).
Ochna holtzii Gilg Habit: Shrub. Habitat: Brachystegia woodland, thicket, and intermediate evergreen rainforest, 15–700 m. Vouchers: Luke & Robertson 2128,
Swynnerton 16 & 10 (EA).
Ochna kirkii subsp. multisetosa Verdc. Habit: Shrub. Habitat: Semi deciduous forest and groundwater forest, 100–600 m. Vouchers: Drummond & Hemsley 4013,
Greenway & Rawlins 9367, Rawlins 222, Luke Q 1411 (EA).
Ochna macrocalyx Oliv. Habit: Shrub. Habitat: Coral rag forest, ca. 10 m. Vouchers:
Luke & Mbinda 5847, Robertson SA & Luke Q 1557 (EA).
Ochna sp. 17 of KTSL Habit: Shrub. Habitat: Thickets on dune above high water
mark, 1–50 m. Vouchers: Gilbert & Kuchar 5874, Kuchar 13719, MacNaughton
2567, Oxtoby EA 15356 (EA): Endemic.
Ochna thomasiana Engl. & Gilg Habit: Shrub. Habitat: Evergreen thicket, forest on
coral rock, and coastal bushland, 0–400 m. Vouchers: SAJIT–005942 & 006244,
Ngumbau V & Mwadime N V0323 (EA, HIB), Drummond & Hemsley 4154,
Gardner in FD 1428, Robertson & Luke 5317, Magogo FC & Glover PE 12,
Robertson SA & Beentje HJ 5631 (EA).
Sauvagesia erecta L. Habit: Herb. Habitat: Grassland, 20–60 m. Vouchers: Nyange
M & Luke Q 0548, Luke & Mbinda 5749, L & L 3795 (EA).
F122. Olacaceae
2 Genera, 2 Species
Olax obtusifolia De Wild. Habit: Tree. Habit: Lowland forest, ca. 200 m. Vouchers:
Luke WRQ & Robertson SA 2771, Luke, Stone & Baer 8244 (EA).
Strombosiopsis pentamera Q. Luke, ined. Habit: Tree, 390 m. Habitat: Forest.
Voucher: Luke WRQ & Robertson SA 2725 (EA): Endemic.
F123. Oleaceae
2 Genera, 7 Species
Jasminum fluminense Vell. subsp. fluminense Habit: Liana. Habitat: Forests, woodlands, bushlands, and grassland, ca. 210 m. Voucher: Wakefield (EA).
Checklist of coastal forests of Kenya
135
Jasminum meyeri-johannis Engl. Habit: Liana. Habitat: Forest and bushland, ca.
690 m. Voucher: Battiscombe 261 (EA).
Jasminum schimperi Vatke Habit: Liana. Habitat: Rainforests, wooded grassland, and
bushland, ca. 400 m. Voucher: Luke WRQ 3157 (EA).
Jasminum stenolobum Rolfe Habit: Liana. Habitat: Brachystegia woodland often on termite mounds, ca. 183 m. Vouchers: Ngumbau V & Mwadime N V0197 (EA, HIB),
Malombe I, Mwadime N & Saidi 1653, Luke WRQ & Robertson SA 2782 (EA).
Jasminum streptopus E. Mey. Habit: Liana. Habitat: Scrub and woodland ca. 100 m.
Vouchers: Luke WRQ & Robertson SA 2798, Graham 2147 (EA).
Olea europaea subsp. cuspidata (Wall. ex G. Don) Cif. Habit: Tree. Habitat: Bushland, woodland, ca. 80 m. Voucher: Luke WRQ & Robertson SA 2497 (EA).
Olea woodiana subsp. disjuncta P. S. Green Habit: Tree. Habitat: Forest, ca. 238 m.
Vouchers: Ngumbau V & Mwadime N V0185 (EA, HIB), Luke WRQ & Robertson SA 505 (EA).
F124. Onagraceae
1 Genus, 7 Species
Ludwigia abyssinica A. Rich. Habit: Herb. Habitat: In swampy places, ca. 40 m.
Vouchers: Luke WRQ & Robertson SA 2756, Nyange M 576 (EA).
Ludwigia adscendens subsp. diffusa (Forssk.) P. H. Raven Habit: Herb. Habitat: In
wet places, ca. 36 m. Vouchers: SAJIT–006201 (EA, HIB), Sangai GW 15607 (EA).
Ludwigia erecta (L.) H. Hara Habit: Herb. Habitat: Wet areas, ca. 30 m. Voucher:
Gray M, PA & Luke Q 431 (EA).
Ludwigia jussiaeoides Desr. Habit: Herb. Habitat: Swampy areas, ca. 25 m. Vouchers: Graham 2099, Joanna in CM 5977, Luke WRQ 15336 (EA).
Ludwigia perennis L. Habit: Herb. Habitat: Swamps, flood plains of lakes, and probably
in other wet habitats, ca. 0–30 m. Vouchers: Whyte, Festo L, Luke Q & P 2701 (EA).
Ludwigia sp. aff. stolonifera (Guill. & Perr.) P. H. Raven Habit: Herb. Habitat: Forest. Voucher: Luke WRQ & Robertson SA 2777 (EA).
Ludwigia stenorraphe subsp. macrosepala (Brenan) P. H. Raven Habit: Herb. Habitat: Forest, ca. 240 m. Voucher: Luke WRQ & Robertson SA 2766 (EA).
F125. Opiliaceae
2 Genera, 2 Species
Opilia amentacea Roxb. Habit: Liana. Habitat: Coastal bushland and riverine forest
to upland rainforest, ca. 0–300 m. Vouchers: Ngumbau V & Mwadime N V0270,
V0426 & V041 (EA, HIB), Kimeu JM, Meso M, Ot 613, Magogo FC & Glover
PE 1053, Verdcourt 3609 (EA).
Pentarhopalopilia umbellulata (Baill.) Hiepko Habit: Liana. Habitat: Coastal forest, Brachystegia woodland, and coastal bushland, 0–100 m. Vouchers: Ngumbau
V & Mwadime N V0498 (EA, HIB), RM Graham in FD1830 & 2301, Gray M
& Luke WRQ 341 (EA).
136
Veronicah M. Ngumbau et al. / PhytoKeys 147: 1–191 (2020)
F126. Orchidaceae
24 Genera, 61 Species
Acampe pachyglossa Rchb.f. Habit: Herb. Habitat: Epiphytic on trees in deciduous
and coastal bushland, 0–800 m. Vouchers: Ngumbau V & Mwadime N V0202
(EA, HIB), Copley 25, Dale in FD 3773, Tweedie 634, Tweedie EM 87 (EA).
Aerangis hologlottis (Schltr.) Schltr. Habit: Herb. Habitat: Epiphytic on twigs and
small branches of trees in coastal forests, 0–500 m. Vouchers: Castelino in EAH
18/63, Archer 522, Luke WRQ 1640 (EA).
Aerangis kirkii (Rchb.f.) Schltr. Habit: Herb. Habitat: Coastal bush and riverine forest at low altitudes inland, on small trees, and bushes, 0–450 m. Vouchers: Ngumbau V & Mwadime N V0139 (EA, HIB), Magogo & Glover 955, RM Graham B
442 & FD 2146, Rawlins SP 11259 (EA).
Aerangis kotschyana (Rchb.f.) Schltr. Habit: Herb. Habitat: Wooded grassland,
woodland, and sometimes in forest, 0–200 m. Vouchers: WE Taylori, Musyoki
BM & Hansen OJ 963, Robertson SA & Luke WRQ 4850 (EA).
Angraecum cultriforme Summerh. Habit: Herb. Habitat: Brachystegia woodland, dry
evergreen forest, and evergreen bushland, ca. 0–400 m. Vouchers: Boscawen in
Moreau 724, Robertson SA & Luke WRQ 2450 (EA).
Angraecum dives Rolfe Habit: Herb. Habitat: Coastal woodland, evergreen forest, scrub,
and on exposed coral rocks, 0–50 m. Vouchers: SAJIT–006460 & 005591 (EA,
HIB), Irwin 92, Polhill & Paulo 691, Rathbun in EAH 14989, Muchiri J 506 (EA).
Angraecum eburneum subsp. giryamae (Rendle) Senghas & P.J. Cribb Habit: Epiphytic or lithophytic herb. Habitat: Thickets on outcropping rocks, on rocks by
sea, and on cliff faces, ca. 0–350 m. Vouchers: Drummond & Hemsley 4181,
Greenway 4969, Dale in FD 3566, Jeffery GW 699 (EA).
Angraecum teres Summerh. Habit: Epiphytic herb. Habitat: Epiphytic on bole of
Cedrela odorata, and on small trees by the sea, 0–600 m. Voucher: Luke WRQ &
Robertson SA 2661 (EA).
Angraecum viride Kraenzl. Habit: Herb. Habitat: In rainforest, ca. 150 m. Voucher:
Luke WRQ 1848 (EA).
Ansellia africana Lindl. Habit: Epiphytic herb. Habitat: Epiphytic or rarely on rocks
in open woodland, wooded grassland, and on Hyphaene palm trees, ca. 0–381 m.
Vouchers: Ngumbau V & Mwadime N V0545 (EA, HIB), Magogo FC & Glover
PE 1078, Reitsma J 495, Copley 123 (EA): Vulnerable.
Bolusiella sp. Habit: Epiphyte. Habitat: Forest. Voucher: Luke WRQ 5270 (EA).
Bonatea rabaiensis (Rendle) Rolfe Habit: Herb. Habitat: Lowland dry evergreen forest on sand or evergreen bushland, ca. 0–304 m. Vouchers: Jeffery GW 238, Luke
WRQ & PA 4507 (EA).
Bulbophyllum intertextum Lindl. Habit: Herb. Habitat: Epiphytic in rainforest, riverine, and moist forest, ca. 300 m. Voucher: Boscawen in Moreau 99 (EA).
Bulbophyllum maximum (Lindl.) Rchb. f. Habit: Herb. Habitat: Epiphyte in open
woodland and riverine forests, ca. 0–365 m. Vouchers: Luke WRQ 1580, Magogo
FC & Glover PE 582, Rawlings 763 (EA).
Checklist of coastal forests of Kenya
137
Bulbophyllum scaberulum (Rolfe) Bolus Habit: Herb. Habitat: Epiphyte in riverine
and montane forests, ca. 100–250 m. Vouchers: Cunningham van Someren in
Moreau 414, Greenway 9807, Luke WRQ & Robertson SA 2621, Robertson SA
& Luke WRQ 6013 (EA).
Calyptrochilum christyanum (Rchb. f.) Summerh. Habit: Herb. Habitat: Forest,
woodland, and grassland, ca. 300 m. Vouchers: Kirk 12, Malombe I, Mwadime N
& Saidi 1669 (EA).
Cynorkis kirkii Rolfe Habit: Herb. Habitat: On rocky seepage slopes, usually in
shade, ca. 326 m. Voucher: Luke WRQ & Pakia M 5278 (EA).
Cyrtorchis arcuata (Lindl.) Schltr. Habit: Herb. Habitat: Woodland and forest, ca.
0–800 m. Vouchers: Verdcourt & Bayliss 2413, Jeffery GW 806, Robertson SA &
Luke WRQ 2064 (EA).
Disperis aphylla Kraenzl. subsp. aphylla Habit: Herb. Habitat: Evergreen forest, ca.
380 m. Vouchers: CG van Someren Sh 8, Magogo FC & Glover PE 981 (EA).
Eulophia angolensis (Rchb. f.) Summerh. var. angolensis Habit: Herb. Habitat:
Swampy area, ca. 20 m. Vouchers: Dale IR 3548, Luke WRQ & PA 3798 (EA).
Eulophia cucullata (Afzel. ex Sw.) Steud. Habit: Herb. Habitat: Grassland, wooded
grassland, bushland, and woodland, ca. 100–200 m. Vouchers: SAJIT–005520 (EA,
HIB), Lucas, Jeffery & Kirrika 246, Hawthorne W 109, Luke WRQ 3155 (EA).
Eulophia horsfallii (Bateman) Summerh. Habit: Herb. Habitat: Riverine, evergreen
forest, and along streams, ca. 30 m. Vouchers: Bally PRO & Smith 14360, Luke
WRQ & PA 3954 (EA).
Eulophia livingstoneana (Rchb. f.) Summerh. Habit: Herb. Habitat: Grassland or
in deciduous woodland of Brachystegia, ca. 0–442 m. Vouchers: Magogo & Glover
851, Magogo FC & Glover PE 203 (EA).
Eulophia petersii (Rchb. f.) Rchb. f. Habit: Herb. Habitat: Thickets, and bushland,
ca. 0–330 m. Vouchers: SAJIT–006239 (EA, HIB), Drummond & Hemsley 4083
& 4232, Luke WRQ 1081 (EA).
Eulophia schweinfurthii Kraenzl. Habit: Herb. Habitat: In grassland, thicket, bushland, and woodland, ca. 40 m. Vouchers: Kassner 359, Luke WRQ 3097 (EA).
Eulophia serrata P. J. Cribb Habit: Herb. Habitat: Commiphora-Acacia bushland, ca.
150–200 m. Vouchers: Robertson SA 5198 & 4037B (EA): Endemic.
Eulophia speciosa (R. Br.) Bolus Habit: Herb. Habitat: In grassland, deciduous bushland, and woodland, ca. 0–335 m. Vouchers: Ngumbau V & Mwadime N V0199
(EA, HIB), Verdcourt 1191, Luke WRQ & Robertson SA 1816, Magogo FC &
Glover PE 480 (EA).
Eulophia taitensis Pfennig & P. J. Cribb Habit: Herb. Habitat: Thickets, coastal
bushland, and woodland. Voucher: RM Graham 388 (EA).
Habenaria armatissima Rchb. f. Habit: Herb. Habitat: Forest, open marshy ground,
deciduous thicket, and mixed dry woodland, ca. 20 m. Vouchers: Jeffery 247,
Rawlins SP 806 (EA).
Habenaria boiviniana Kraenzl. & Schltr. Habit: Herb. Habitat: At margins of lowland rainforest and coastal evergreen bushland, up to 400 m. Voucher: Drummond & Hemsley 3967 (EA).
138
Veronicah M. Ngumbau et al. / PhytoKeys 147: 1–191 (2020)
Habenaria kilimanjari Rchb. f. Habit: Herb. Habitat: Periodically flooded grassland, often with scattered bushes, ca. 90–152 m. Vouchers: RM Graham in FD
1596, Dale in FD 3547, Jex-Blake in CM 5996, Dale IR 3547 (EA).
Habenaria plectromaniaca Rchb.f. & S. Moore Habit: Herb. Habitat: Marshy grassland or at edge of forests, 30–420 m. Vouchers: RM Graham in FD 1593 & 1934,
Adams BR 33, Luke WRQ 3317 (EA).
Habenaria stylites Rchb. f. & S. Moore subsp. stylites Habit: Herb. Habitat: Grassland, lowland rainforest, and wooded grassland, ca. 100–335 m. Vouchers: GM
11200, Cunningham-van Someren GR Sh 17 (EA): Endangered.
Habenaria subarmata Rchb. f. Habit: Herb. Habitat: Grassland or open woodland,
ca. 5 m. Vouchers: Rawlins in EAH 11258, Festo 2550 (EA).
Habenaria trilobulata Schltr. Habit: Herb. Habitat: Open woodland and lowland
dry evergreen forest, ca. 304 m. Vouchers: Ngumbau V & Mwadime N V092 (EA,
HIB), Drummond & Hemsley 3809, Tweedie 635 (167), Luke WRQ & Robertson SA 1794, Luke WRQ 3116 (EA).
Jumellea filicornoides (De Wild.) Schltr. Habit: Herb. Habitat: Riverine forest, savannah trees, and on rocks, ca. 600 m. Vouchers: Luke & Robertson 1826, Luke
WRQ & Robertson SA 1826 (EA).
Microcoelia aphylla (Thouars) Summerh. Habit: Epiphyte. Habitat: In thickets of
Acacia-Commiphora, in scrub, and riverine forest on Barringtonia racemosa, ca.
65 m. Vouchers: SAJIT–006436 (EA, HIB), RM Graham 1637 (EA).
Microcoelia exilis Lindl. Habit: Herb. Habitat: In riverine forest and woodland, ca.
0–800 m. Vouchers: Drummond & Hemsley 3890, Polhill & Paulo 636, Rawlins
SP 11268 (EA).
Microcoelia megalorrhiza (Rchb. f.) Summerh. Habit: Herb. Habitat: Woodland
and riverine thicket, ca. 100–500 m. Vouchers: Luke WRQ & Robertson SA 1762
& 2022, Robertson SA, Beentje HJ, Luke WRQ & Khayota B268 (EA).
Microcoelia obovata Summerh. Habit: Herb. Habitat: Woodland or dry riverine forest, ca. 0–500 m. Vouchers: Saunders PJR in EAH 11247, Luke WRQ & Robertson SA 2680 & 2018 (EA).
Microcoelia physophora (Rchb. f.) Summerh. Habit: Herb. Habitat: Coastal bushland and woodland, 0–500 m. Vouchers: JB Smart s.n., Luke WRQ 3480 (EA).
Microcoelia smithii (Rolfe) Summerh. Habit: Herb. Habitat: Evergreen forest, on
Euphorbia sp, 0–500 m. Vouchers: WE Taylor s.n., Beentje HJ 2309 (EA).
Nervilia bicarinata (Blume) Schltr. Habit: Herb. Habitat: Evergreen forest, ca. 7 m.
Vouchers: Luke WRQ 2404, Bytebier B 630 (EA).
Nervilia crociformis (Zoll. & Moritzi) Seidenf. Habit: Herb. Habitat: Evergreen forest, grassland, and riverine forest, ca. 150 m. Voucher: Luke WRQ 3129 (EA).
Nervilia kotschyi (Rchb. f.) Schltr. var. kotschyi Habit: Herb. Habitat: Scrub and in shady
places in deciduous woodland, ca. 0–442 m. Vouchers: Ngumbau V & Mwadime N
V037 (EA, HIB), Magogo & Glover 265, Adams BR 28, Luke WRQ 3596 (EA).
Nervilia petraea (Afzel. ex Sw.) Summerh. Habit: Herb. Habitat: Brachystegia woodland, grassland, and pine plantations, ca. 381 m. Vouchers: Luke WRQ 3119,
Magogo FC & Glover PE 1112 & 560 (EA).
Checklist of coastal forests of Kenya
139
Oeceoclades maculata (Lindl.) Lindl. Habit: Herb. Habitat: Dry and moist habitats
including evergreen deciduous forests, ca. 0–250 m. Vouchers: Luke WRQ & PA
4506, Luke WRQ & Robertson SA 6007 (EA).
Oeceoclades saundersiana (Rchb. f.) Garay & P. Taylor Habit: Herb. Habitat: Forest and thickets, ca. 0–280 m. Vouchers: Ngumbau V & Mwadime N V015 (EA,
HIB), Forestry Dept., Archr 411, Castelino H347/62, Robertson SA & Luke
WRQ 6007, Evans M 15122 (EA).
Oeceoclades zanzibarica (Summerh.) Garay & P. Taylor Habit: Terrestrial or lithophtic herb. Habitat: Thickets, ca. 5 m. Vouchers: Archer PG 728, Mwadime N
& Cherise C 229 (EA).
Platycoryne crocea Rolfe Habit: Herb. Habitat: Grassland, 0–360 m. Vouchers: CG
van Someren Sh 4, Jeffery 629, Luke WRQ & Pakia M 7460A (EA).
Polystachya concreta (Jacq.) Garay & H. R. Sweet Habit: Herb. Habitat: Rainforest,
ca. 0–280 m. Vouchers: SAJIT–005431, Ngumbau V & Mwadime N V013 (EA,
HIB), Piers 62/29, Luke WRQ 1597 (EA).
Polystachya cultriformis (Thouars) Lindl. ex Spreng. Habit: Herb. Habitat: Evergreen forest, ca. 335 m. Voucher: Bayliss RDA H93/57/2 (EA).
Polystachya fischeri Rchb. f. ex Kraenzl. Habit: Herb. Habitat: Epiphytic in dry evergreen forest, ca. 310 m. Vouchers: WE Taylor, Pearce TR 616 (EA).
Polystachya lindblomii Schltr. Habit: Herb. Habitat: Sub montane, riverine evergreen
forest, and high rainfall woodland, ca. 69 m. Voucher: Miyawa DO 1155 (EA).
Polystachya modesta Rchb. f. Habit: Herb. Habitat: In riverine forest and woodland,
ca. 470 m. Vouchers: Bayliss RDA 3, Robertson SA 5806 (EA).
Polystachya teitensis P. J. Cribb Habit: Herb. Habitat: Forest, ca. 700 m. Vouchers:
SAJIT–005434 (EA, HIB), Luke WRQ & Robertson SA 2049 (EA): Endangered.
Rhipidoglossum rutilum (Rchb. f.) Schltr. Habit: Herb. Habitat: In forests, ca.
100 m. Voucher: Luke WRQ & Robertson SA sr.
Solenangis wakefieldii (Rolfe) P. J. Cribb & J. Stewart Habit: Herb. Habitat: In
coastal and lowland bushland, 0–300 m. Vouchers: SAJIT–005426 (EA, HIB),
Wakefield, Archer PG 447 (EA).
Tridactyle bicaudata (Lindl.) Schltr. Habit: Herb. Habitat: In forest and woodland
areas, ca. 600 m. Vouchers: Irwin PH 497, Luke WRQ et al. 7083, Luke WRQ &
Robertson SA 2035 (EA).
Vanilla ramosa Rolfe Habit: Climber. Habitat: Moist forest, ca. 280 m. Vouchers:
Ngumbau V & Mwadime N V010 (EA, HIB), Luke WRQ 3047 (EA).
Vanilla roscheri Rchb. f. Habit: Climber. Habitat: Coastal bushland, coral rock,
mangrove swamps, and open evergreen inland scrub, 0–750 m. Vouchers: SAJIT–006445 (EA, HIB), Bally 10462, Verdcourt 3223, Polhill & Paulo 876 (EA).
F127. Orobanchaceae
11 Genera, 21 Species
Alectra orobanchoides Benth. Habit: Herb. Habitat: In open woodland, scrub, rocky areas, and cultivated land, ca. 220 m. Voucher: Luke WRQ & Robertson SA 2765 (EA).
140
Veronicah M. Ngumbau et al. / PhytoKeys 147: 1–191 (2020)
Alectra picta (Hiern) Hemsl. Habit: Herb. Habitat: Riverine Brachystegia woodland,
amongst shrubs, and sedges at edges of river, ca. 30 m. Vouchers: Robertson &
Luke 6213, Luke 1332 (EA).
Alectra sessiliflora (Vahl) Kuntze Habit: Herb. Habitat: Grassland and cultivation
areas, ca. 400 m. Vouchers: Luke 1332, Drummond & Hemsley 3970 (EA).
Buchnera hispida Buch.-Ham. ex D. Don Habit: Herb. Habitat: Grassland and woodland, ca. 0–100 m. Vouchers: Drummond & Hemsley 3741, Rawlins SP 65 (EA).
Buchnera leptostachya Benth. Habit: Herb. Habitat: In wet grassland, swamps, and
coastal bush, 0–500 m. Vouchers: Swynnerton 403, Graham 2115, Festo L, Luke
Q & P 2810 (EA).
Buttonia natalensis McKen ex Benth. Habit: Liana. Habitat: Open woodlands,
among rocks, and on riverbanks, ca. 30–518 m. Voucher: Robertson SA & Luke
WRQ 6061 (EA).
Cistanche tinctoria (Forssk.) Deflers Habit: Herb. Habitat: Dry stony ground or
semi desert scrub, ca. 20 m. Vouchers: Verdcourt 1100, Taiti S 542 (EA).
Cycnium adonense E. Mey. ex Benth. subsp. adonense Habit: Herb. Habitat: Grassland and grassland woodland, ca. 0–360 m. Vouchers: Drummond & Hemsley
1192, Magogo FC & Glover PE 26, Festo L, Luke Q & P 2651 (EA).
Cycnium tubulosum (L. f.) Engl. Habit: Herb. Habitat: Damp grassland, along
marshes, and river banks, ca. 5 m. Voucher: Festo L & Luke Q 2551 (EA).
Cycnium veronicifolium (Vatke) Engl. Habit: Herb. Habitat: Grassland, wooded
grassland, and bushland, ca. 0–300 m. Vouchers: CW Elliot in FD 3373, Robertson SA & Luke WRQ 6054 (EA).
Ghikaea speciosa (Rendle) Diels Habit: Herb. Habitat: Dry grassland and AcaciaCommiphora scrub, ca. 350 m. Voucher: van Someren 878 (EA).
Harveya kenyensis Hepper Habit: Herb. Habitat: Dry forests, 30–450 m. Voucher:
Robertson SA & Luke WRQ 6065 (EA).
Harveya obtusifolia (Benth.) Vatke Habit: Herb. Habitat: Thickets, bushland, fields, and
orchards, ca. 350 m. Vouchers: Drummond & Hemsley 3865, Jeffery GW 268 (EA).
Micrargeria filiformis (Schumach. & Thonn.) Hutch. & Dalziel Habit: Herb.
Habitat: Swamps, damp and wet areas in grassland, and seepages in woodland, ca.
0–20 m. Vouchers: Greenway 4963, Joy Adamson in Bally B 6141, Luke WRQ
15342 (EA).
Nesogenes africanus G. Taylor Habit: Herb. Habitat: In shade of bushes on limestone
rock, ca. 10 m. Voucher: Robertson SA 5805 (EA).
Pseudosopubia hildebrandtii Engl. Habit: Herb. Habitat: Grassland, grassland with
trees, rocky, river, and swamp margins, ca. 0–100 m. Vouchers: Drummond &
Hemsley 3748, Rawlins SP 801 (EA).
Striga asiatica (L.) Kuntze Habit: Herb. Habitat: Semi-parasitic on roots of wild
grasses and cultivated maize, ca. 0–365 m. Vouchers: SAJIT–006051, Ngumbau
V & Mwadime N V0341 (EA, HIB), Drummond & Hemsley 3839, Burstyn P
75/77, Magogo FC & Glover PE 1089 (EA).
Striga forbesii Benth. Habit: Herb. Habitat: Low lying grassy places, ca. 10–70 m. Vouchers: Gilbert & Kibuwa 19932, Festo L, Luke Q & P 2711, Luke WRQ 3489 (EA).
Checklist of coastal forests of Kenya
141
Striga gesnerioides (Willd.) Vatke Habit: Herb. Habitat: In permanent grassland
parasitizing grass roots, ca. 20 m. Voucher: Polhill & Paulo 579 (EA).
Striga latericea Vatke Habit: Herb. Habitat: Seasonally wet places among grass, ca.
100 m. Vouchers: Drummond & Hemsley 4247, Archer PG 445, Magogo FC &
Glover PE 1050 (EA).
Striga pubiflora Klotzsch Habit: Herb. Habitat: Woodland among grass, in salt
marshes, and on roots of grasses, ca. 100–304 m. Vouchers: Hildebrandt 1907,
Moomaw 953, Magogo & Glover 849, Thomas M 23 (EA).
F128. Oxalidaceae
2 Genera, 4 Species
Biophytum umbraculum Welw. Habit: Herb. Habitat: Brachystegia woodland, open
grassland, and marsh edges, ca. 0–381 m. Vouchers: Ngumbau V & Mwadime N
V0346 (EA, HIB), Jeffery 342, Magogo FC & Glover PE 951, Rawlins SP 66 (EA).
Oxalis barrelieri L. Habit: Herb. Habitat: Weed of roadsides and cultivated areas, ca.
100 m. Voucher: Robertson SA 3389 (EA): Naturalized.
Oxalis corniculata L. Habit: Herb. Habitat: Arable land and waste places, 0–396 m.
Vouchers: Magogo FC & Glover PE 667 & 496 (EA): Naturalized.
Oxalis latifolia Kunth Habit: Herb. Habitat: Forests and forest edges, arable fields,
and waste places. Voucher: Jeffery GW 527 (EA).
F129. Pandanaceae
1 Genus, 2 Species
Pandanus kirkii Rendle Habit: Tree. Habitat: Sandy beaches and on coral near the
sea, 0–10 m. Vouchers: Greenway 9642, St. John 26594, Beentje 3737 (EA).
Pandanus rabaiensis Rendle. Habit: Tree. Habitat: Riverine forest or along streams
in evergreen forest, 1–100 m. Vouchers: Agnew 9742, St. John 26596, Dominic U
16189, Luke WRQ & Robertson SA 2827 (EA).
F130. Papaveraceae
1 Genus, 1 Species
Argemone mexicana L. Habit: Herb. Habitat: Open ground or along the roads, ca.
0–200 m. Voucher: Robertson SA 3774 (EA): Naturalized.
F131. Passifloraceae
7 Genera, 19 Species
Adenia aculeata (Oliv.) Engl. subsp. aculeata Habit: Liana. Habitat: Dry forests, ca.
20 m. Voucher: Gilbert MG & Kuchar P 5870 (EA).
Adenia aculeata subsp. manganiana (Chiov.) W.J.de Wilde Habit: Liana. Habitat:
Coastal scrub, 0–100 m. Voucher: Bally 5951 (EA).
142
Veronicah M. Ngumbau et al. / PhytoKeys 147: 1–191 (2020)
Adenia angulosa G.W. Hu & Q.F. Wang Habit: Liana. Habitat: Coastal moist forest,
forest margin, ca. 350 m. Vouchers: SAJIT–005956, 006061, 006074, 006078,
005975 & 006156, Ngumbau V & Mwadime N V0171 (EA, HIB).
Adenia globosa Engl. subsp. globosa Habit: Liana. Habitat: Deciduous and dry evergreen bushland, often in rocky places, ca. 100 m. Vouchers: Drummond & Hemsley 4053, Ndakala J 363, Graham RM 863 (EA).
Adenia cissampeloides (Planch. ex Hook.) Harms Habit: Liana. Habitat: Forest and
bushland of various types, ca. 0–442 m. Vouchers: SAJIT–005959, 006083 &
006209, Ngumbau V & Mwadime N V0268 (EA, HIB), RM Graham A 494 in
FD1873, Robertson SA & Luke WRQ 5626 (EA).
Adenia keramanthus Harms Habit: Herb. Habitat: Deciduous woodland, bushland, dry evergreen, and coastal bushland, ca. 0–330 m. Vouchers: Drummond &
Hemsley 4233, Luke WRQ & Robertson SA 1551 (EA).
Adenia kirkii (Mast.) Engl. Habit: Liana. Habitat: Lowland evergreen forest and
grassland, 0–700 m. Vouchers: SAJIT–006429, Ngumbau V & Mwadime N
V0441 & V005 (EA, HIB), Rawlins 891, Jeffery K 569, Rawlins 292, Robertson
SA & Luke WRQ 5375 (EA).
Adenia lindiensis Harms Habit: Liana. Habitat: Edges of evergreen forest and bushland,
0–304 m. Vouchers: VG van Someren 87, Robertson SA & Luke WRQ 5170 (EA).
Adenia lobata subsp. rumicifolia (Engl. & Harms) Lye Habit: Liana. Habitat: In
moister places, in forest edges, semi swamp forest, riverine forest, and thicket, 0–320
m. Vouchers: SAJIT–006008, Ngumbau V & Mwadime N V0522 (EA, HIB), VG
van Someren 97, Mwadime N 29, Luke WRQ 3107, Luke WRQ 1599 (EA).
Adenia venenata Forssk. Habit: Liana. Habitat: Wooded grassland, deciduous woodland, and bushland, ca. 20 m. Vouchers: Rawlins 109, Rawlins SP 372 (EA).
Basananthe hanningtoniana (Mast.) W.J.de Wilde Habit: Climber. Habitat: Wide
variety of open habitats and deciduous bushland, ca. (0–) 400 m. Vouchers: Rawlins SP 340, Magogo FC & Glover PE 852 (EA).
Basananthe lanceolata (Engl.) W. J.de Wilde Habit: Climber. Habitat: Grassland,
bushland, and disturbed places, 0–100 m. Vouchers: Graham RM 2022, Kuchar P
11753, Drummond & Hemsley 3856 (EA).
Basananthe zanzibarica (Mast.) W.J.de Wilde Habit: Climber. Habitat: Lowland
dry evergreen forest and coastal bushland, 0–450 m. Vouchers: SAJIT–006039,
Ngumbau V & Mwadime N V070 (EA, HIB), Drummond & Hemsley 3956,
Napper 1373, Verdcourt 3924, Festo L, Luke Q & P 2716, Magogo FC & Glover
PE 438 (EA).
Loewia tanaensis Urb. Habit: Shrub. Habitat: Woodland, ca. 280 m. Voucher:
Mwadime N & Luke WRQ 2436 (EA).
Passiflora foetida L. Habit: Climber. Habitat: Thickets, ca. 10–15 m. Vouchers: Luke
WRQ 3059 & 3069 (EA): Naturalized.
Schlechterina mitostemmatoides Harms Habit: Liana. Habitat: Lowland dry evergreen, riverine forest, and coastal bushland, 0–700 m. Vouchers: SAJIT–005998
& 006430, Ngumbau V & Mwadime N V0213 (EA, HIB), Magogo & Glover
287, Dale in FD 3643, Rawlins in EAH 11269, Muchiri J 451 (EA).
Checklist of coastal forests of Kenya
143
Turnera thomasii (Urb.) Story Habit: Shrub. Habitat: Dry sandy places, ca. 25 m.
Vouchers: Mwadime N & Luke WRQ 2435, F Thomas 47 (EA): Endemic.
Tricliceras brevicaule (Urb.) R. Fern. var. brevicaule Habit: Herb. Habitat: Wooded
grassland, ca. 200 m. Vouchers: Luke WRQ et al. 3623 & 3839 (EA).
Tricliceras xylorhizum Verdc. Habit: Herb. Habitat: Acacia-Commiphora bushland,
ca. 230 m. Voucher: Luke & Robertson 2516 (EA).
F132. Pedaliaceae
4 Genera, 6 Species
Dicerocaryum zanguebarium (Lour.) Merr. Habit: Herb. Habitat: Coastal dune,
0–500 m. Vouchers: SAJIT–006250 (EA, HIB), Luke Q 6143 (EA).
Pedalium murex L. Habit: Herb. Habitat: On limestone in short grass near the coast,
old sisal plantations, 0–440 m. Vouchers: SAJIT–004678 (EA, HIB), Napier
3265 (EA).
Sesamothamnus busseanus Engl. Habit: Shrub. Habitat: Acacia-Commiphora scrub,
ca. 300 m. Voucher: Bally 4375 (EA).
Sesamum angolense Welw. Habit: Herb. Habitat: Disturbed grassland and roadsides,
ca. 215 m. Voucher: SAJIT–005939 (EA, HIB).
Sesamum angustifolium (Oliv.) Engl. Habit: Herb. Habitat: In cultivated, waste areas, native gardens, roadsides, and short grassland, ca. (15–) 600 m. Voucher: Battiscombe 258 (EA).
Sesamum indicum L. Habit: Herb. Habitat: Grassland and roadsides, ca. 10 m.
Voucher: Jeffery K 24 (EA).
F133. Peraceae
1 Genus, 1 Species
Clutia abyssinica var. usambarica Pax & K. Hoffm. Habit: Shrub. Habitat: Dry
evergreen forest, ca. 300 m. Voucher: Cunningham-van Someren GR Sh 84 (EA).
F134. Phyllanthaceae
10 Genera, 30 Species
Antidesma membranaceum Müll. Arg. Habit: Shrub. Habitat: Rainforest, drier evergreen forest, thickets, and wooded grassland, ca. 10–410 m. Vouchers: Gillett
18669, Wakefield (EA).
Antidesma venosum E. Mey. ex Tul. Habit: Shrub. Habitat: In riverine vegetation,
woodland, and wooded grassland, ca. 0–237 m. Vouchers: SAJIT–005949,
005499 & 005560, Ngumbau V & Mwadime N V0173 (EA, HIB), Luke WRQ
& PA 238 (EA).
Antidesma vogelianum Müll. Arg. Habit: Shrub. Habitat: Rainforest and drier evergreen forest, ca. 0–187 m. Vouchers: Ngumbau V & Mwadime N V089 (EA,
HIB), Robertson SA & Luke WRQ 5160 (EA).
144
Veronicah M. Ngumbau et al. / PhytoKeys 147: 1–191 (2020)
Bridelia atroviridis Müll. Arg. Habit: Tree. Habitat: Forest edges, associated bushland, thicket, and rivers, ca. 70–300 m. Vouchers: SAJIT–006011, Ngumbau V
& Mwadime N V076 & 0172 (EA, HIB), Drummond & Hemsley 4025, Faden
70/255 (EA).
Bridelia cathartica Bertol. Habit: Shrub. Habitat: Woodland, bushland, and thicket,
ca. 0–300 m. Vouchers: SAJIT–006106 & 004660, Ngumbau V & Mwadime N
V0191 (EA, HIB), Magogo & Glover 1067, Polhill & Paulo 746, Gillespie 146,
Gray M & Luke WRQ 342 (EA).
Bridelia micrantha (Hochst.) Baill. Habit: Tree. Habitat: Evergreen forest, associated bushland, thickets, along rivers, lakes, and swamps, ca. 50–400 m. Vouchers:
Magogo & Glover 165, Luke WRQ & Robertson SA 228 (EA).
Cleistanthus beentjei Q. Luke, ined. Habit: Tree. Habitat: Moist forest, 30–500 m.
Vouchers: Luke WRQ & Robertson SA 2684 & 1934, Robertson SA & Luke
WRQ 6343A (EA).
Cleistanthus schlechteri (Pax) Hutch. Habit: Tree. Habitat: Lowland dry evergreen
forest and riverine, 0–760 m. Vouchers: Faden & Faden 77/737, Graham RM
1652 (EA).
Cyathogyne usambarensis (Verdc.) J. Léonard Habit: Shrub. Habitat: Lowland dry
evergreen forest and bushland, 200–244 m. Vouchers: SAJIT–004656, Ngumbau
V & Mwadime N V0398 (EA, HIB), Faden & Evans 70/797, Lavranos & Newton
12292, Festo L, Luke Q & P 2765, Thomas Mwadime in Mrs SA Robertson 7792,
Luke WRQ 3144 (EA).
Flueggea virosa (Roxb. ex Willd.) Royle subsp. virosa Habit: Shrub. Habitat: Locally common in wide associations, mainly forest edges, bushland, and thickets,
0–230 m. Vouchers: Musyoki & Hansen 961, Magogo FC & Glover PE 295 (EA).
Margaritaria discoidea var. nitida (Pax) Radcl.-Sm. Habit: Tree. Habitat: Dry evergreen forest, deciduous woodland, thickets, and disturbed places, ca. 20–237 m.
Vouchers: Ngumbau V & Mwadime N V0186 (EA, HIB), Magogo & Glover 678,
Luke Q 1461(EA).
Margaritaria discoidea var. triplosphaera Radcl.-Sm. Habit: Tree. Habitat: Dry evergreen forest and disturbed places, 9–300 m. Vouchers: Kassner 146, Drummond
& Hemsley 1140 (EA).
Meineckia fruticans (Pax) G. L. Webster var. fruticans Habit: Shrub. Habitat: Forest
and coastal bushland, 6–200 m. Vouchers: SAJIT–006081, Ngumbau V & Mwadime
N V0420 (EA, HIB), Gillett et al. 14645, Faden RB 70/433, Luke Q 1500 (EA).
Meineckia fruticans var. engleri (Pax) G. L. Webster Habit: Shrub. Habitat: On
lowland wet evergreen and riparian forest, 30–220 m. Vouchers: Faden & Evans
70/735, Verdcourt 1078 (EA).
Phyllanthus amarus Schumach. & Thonn. Habit: Herb. Habitat: Cultivated ground
and disturbed places, ca. 2 m. Vouchers: Verdcourt 3908, Tweedie 1245, Sangai
GW in EA 15752 (EA).
Phyllanthus chevalieri Beille Habit: Herb. Habitat: On black clay soil, in grassland,
flood plain grassland, and Acacia woodland, ca. 15–200 m. Vouchers: Polhill &
Paulo 509, Luke 4620 (EA).
Checklist of coastal forests of Kenya
145
Phyllanthus harrisii Radcl.-Sm. Habit: Herb. Habitat: Waste ground, swamps, rice
fields, dry stream beds, riverine forest, and old cultivations, 0–150 m. Vouchers:
SAJIT–005992, Ngumbau V & Mwadime N V0551 (EA, HIB), Tweedie 1041,
Rawlins 674, Hooper & Townsend 1198, Magogo FC & Glover PE 363 (EA).
Phyllanthus kaessneri Hutch. Habit: Shrub. Habitat: Coastal bushland, Brachystegia
woodland, shady evergreen forest, and disturbed places, 0–90 m. Voucher: Polhill
& Paulo 774 (EA).
Phyllanthus leucocalyx Hutch. Habit: Herb. Habitat: Open or rocky generally damp
places in a variety of associations, disturbed places, ca. 0–407 m. Vouchers: SAJIT–006047 (EA, HIB), Magogo & Glover 245, Gillespie 287, RM Graham
1726 (EA).
Phyllanthus maderaspatensis L. var. maderaspatensis Habit: Herb. Habitat: Deciduous woodland, bushland, wooded grassland places, cultivated, and disturbed places, ca. 0–300 m. Vouchers: Magogo & Glover 1057, Festo L & Luke Q 2470 (EA).
Phyllanthus mittenianus Hutch. Habit: Shrub. Habitat: Moist forest, ca. 300 m.
Vouchers: SAJIT–005984 (EA, HIB), Luke WRQ & PA 5404 (EA).
Phyllanthus nummulariifolius Poir. Habit: Herb. Habitat: Woodland, wooded
grassland, forest edges, extending to upland grassland, and bushland, 0–382 m.
Vouchers: SAJIT–006073 (EA, HIB), Mwangangi 1282, Luke WRQ & Robertson SA 2749 (EA).
Phyllanthus ovalifolius Forssk. Habit: Shrub. Habitat: Deciduous woodlands, thickets, and evergreen rainforest, ca. 86 m. Vouchers: Ngumbau V & Mwadime N
V0470 (EA, HIB), Luke Q 1564 (EA).
Phyllanthus pinnatus (Wight) G.L. Webster Habit: Shrub. Habitat: Coastal, lowland
mixed bushland, wooded grassland, and sometimes riverine, 0–125 m. Voucher:
Polhill & Paulo 685 (EA).
Phyllanthus physocarpus Müll.Arg. Habit: Shrub. Habitat: Evergreen forest, ca.
450 m. Vouchers: Magogo & Glover 260, Spjut 4553, Cain in EAH. 13663, Luke
WRQ & Robertson SA 503 (EA).
Phyllanthus reticulatus Poir. var. reticulatus Habit: Shrub. Habitat: Forest edges,
riverine, maritime, edges of swamps, and lakes, 0–213 m. Vouchers: Ngumbau V
& Mwadime N V0327 (EA, HIB), Sangai 962 (EA).
Phyllanthus reticulatus var. glaber (Thwaites) Müll.Arg. Habit: Shrub. Habitat:
Coastal forests, 0–600 m. Vouchers: Bally 5790, Drummond & Hemsley 1022,
Verdcourt 2111, Luke WRQ et al. 5412 (EA).
Phyllanthus somalensis Hutch. Habit: Shrub. Habitat: Acacia-Commiphora bushland, in thickets, temporary pools, and swampy areas, 120–300 (–570) m. Vouchers: Bally 16865, Lucas 60 (EA).
Phyllanthus suffrutescens Pax Habit: Herb. Habitat: Grassland and open bushland
often in rocky places, ca. 100 m. Voucher: Gillett 16866 (EA).
Phyllanthus welwitschianus Müll. Arg. Habit: Shrub. Habitat: Coastal bushland,
dry forest undergrowth, Brachystegia and mixed woodland, ca. 0–224 m. Vouchers: Ngumbau V & Mwadime N V096 (EA, HIB), Maggridge 144, Rawlins 696,
Tweedie 1859, Bock K 715 (EA).
146
Veronicah M. Ngumbau et al. / PhytoKeys 147: 1–191 (2020)
Thecacoris spathulifolia (Pax) Leandri Habit: Shrub. Habitat: Dry evergreen forest,
wooded grassland, and thickets, 15–700 m. Vouchers: Dale in FD 3560, Polhill
& Paulo 772 (EA).
Wielandia fadenii (Radcl.-Sm.) Petra Hoffm. & McPherson Habit: Shrub. Habitat: Lowland evergreen forest, ca. 220 m. Vouchers: Dale in FD 3665, Bally &
Smith 14396, Hawthorne 200, Thomas Mwandime in Mrs. SA Robertson 7793,
Festo L, Luke Q & P 2761, Faden, R.B, Githui, M. & Evans, A.N. 71/263 (EA).
F135. Phytolaccaceae
1 Genus, 1 Species
Rivina humilis L. Habit: Herb. Habitat: A weed of closed forests or forest margins, ca.
3 m. Voucher: Mrs Robertson SA 7692 (EA).
F136. Picrodendraceae
2 Genera, 2 Species
Aristogeitonia monophylla Airy Shaw Habit: Tree. Habitat: Forest, ca. 199 m. Vouchers: Ngumbau V & Mwadime N V0406 (EA, HIB), Faden RB, Evans A & Msafiri
F 70/949 (EA): Vulnerable.
Oldfieldia somalensis (Chiov.) Milne-Redh. Habit: Tree. Habitat: Lowland dry evergreen and semi deciduous forest, 30–500 m. Vouchers: Perdue & Kibuwa 10005,
Greenway & Rawlins 9352, Ross KS 193, Kirika P, Nyamongo D & Sanyanyi S
04/16/2008 (EA).
F137. Piperaceae
2 Genera, 2 Species
Peperomia pellucida (L.) Kunth Habit: Herb. Habitat: Seasonally swampy areas in
grassland, scattered tree grassland, and evergreen thicket, ca. 0–30 m. Vouchers:
Schlieben 12142, RB & AJ Faden 74/1213 (EA).
Piper betle L. Habit: Climber. Habitat: Humid forest, ca. 240 m. Vouchers: Robertson SA 7421, Luke WRQ 1647 (EA): Cultivated.
F138. Plantaginaceae
4 Genera, 5 Species
Bacopa crenata (Benth.) Hepper Habit: Herb. Habitat: Grassy swamps and salt
marshes, ca. 0–36 m. Vouchers: SAJIT–006198, Ngumbau V & Mwadime N
V0552 (EA, HIB), Drummond & Hemsley 4009 & 1186, Magogo & Glover
130, Luke WRQ & Robertson SA 2754 (EA).
Bacopa floribunda (R.Br.) Wettst. Habit: Herb. Habitat: Wet places and rice fields,
ca. 2–350 m. Vouchers: Drummond & Hemsley 4010 & 4144, Polhill & Paulo
888, Festo L & Luke Q 2579 (EA).
Checklist of coastal forests of Kenya
147
Limnophila indica (L.) Druce Habit: Herb. Habitat: Marshes, pools, along riversides
or forest paths, 0–200 m. Voucher: Festo L & Luke Q 2566 (EA).
Scoparia dulcis L. Habit: Herb. Habitat: In cultivated and waste ground, 0–50 m.
Vouchers: Magogo & Glover 1034, Mwachala & Vollesen 57, Kirika P, Mbale M
& Mbatha M 779 (EA).
Stemodia serrata Benth. Habit: Herb. Habitat: Moist, grassy places, flood plains,
drying out during dry season, ca. 50 m. Voucher: Luke et al. 562 (EA).
F139. Plumbaginaceae
1 Genus, 1 Species
Plumbago stenophylla Wilmot-Dear Habit: Herb. Habitat: Coastal forest, ca. 0–30
m. Vouchers: Graham RM 615 & 2103 (EA): Endemic.
F140. Poaceae
72 Genera, 207 Species
Acroceras attenuatum Renvoize Habit: Herb. Habitat: In shade, ca. 0–300 m. Vouchers: Bogdan A 4714, Magogo & Glover 772, Rawlins 300, Bogdan 4714 (EA).
Alloteropsis papillosa Clayton Habit: Herb. Habitat: Seasonally wet grassland, particularly on black clay, ca. 0–400 m. Vouchers: Robertson SA 3396, Moomaw JC
1165 (EA).
Andropogon africanus Franch. Habit: Herb. Habitat: Seasonally flooded grassland.
Voucher: Allan 189 (EA).
Andropogon canaliculatus Schumach. Habit: Herb. Habitat: Mainly in moist or
swampy places, ca. 0–300 m. Voucher: Magogo FC & Glover PE 612 (EA).
Andropogon chinensis (Nees) Merr. Habit: Herb. Habitat: Deciduous bushland
and wooded grassland, 0–200 m. Vouchers: Thomas Mwambiri 10, Bogdan A
3316 (EA).
Andropogon heterantherus Stapf Habit: Herb. Habitat: Coastal scrub, 0–100 m.
Vouchers: Wilson J 32, Bogdan 3310 (EA).
Andropogon schirensis Hochst. ex A. Rich. Habit: Herb. Habitat: Deciduous bushland and wooded grassland, ca. 100 m. Voucher: Moomaw JC 1217 (EA).
Aristida adscensionis L. Habit: Herb. Habitat: Waste places, ca. 0–150 m. Vouchers:
Magogo FC & Glover PE 704, Kirika P 85 (EA).
Aristida barbicollis Trin. & Rupr. Habit: Herb. Habitat: Deciduous bushland and
open places, ca. 0–351 m. Vouchers: JM 15, Magogo FC & Glover PE 329 (EA).
Aristida kelleri Hack. Habit: Herb. Habitat: Deciduous bushland, 250–450 m.
Voucher: Verdcourt 2101 (EA).
Aristida mutabilis Trin. & Rupr. Habit: Herb. Habitat: Acacia-Commiphora deciduous bushland and open semi-desert bushland, ca. 100–304 m. Voucher: Magogo
FC & Glover PE 521 (EA).
Aristida sieberiana Trin. ex Spreng. Habit: Herb. Habitat: Deciduous bushland, ca.
300 m. Voucher: Pratt in EAH 13812 (EA).
148
Veronicah M. Ngumbau et al. / PhytoKeys 147: 1–191 (2020)
Aristida stenostachya Clayton Habit: Herb. Habitat: Deciduous thicket and bushland, 0–426 m. Vouchers: Magogo FC & Glover PE 692, Muchiri J 475, Bogdan
5322 (EA).
Axonopus flexuosus (Peter) Troupin Habit: Herb. Habitat: Damp or swampy soils,
0–152 m. Vouchers: Luke PA & WRQ 3611, Bogdan 5348 & 3343 (EA).
Bothriochloa bladhii (Retz.) S. T. Blake Habit: Herb. Habitat: Streamside, swamp margins, and cracking clays, ca. 0–75 m. Voucher: Magogo FC & Glover PE 964 (EA).
Bothriochloa insculpta (Hochst. ex A. Rich.) A. Camus Habit: Herb. Habitat: Overgrazed and disturbed places, ca. 50 m. Voucher: Tateoka 3023 (EA).
Brachiaria brizantha (A. Rich.) Stapf Habit: Herb. Habitat: Deciduous woodland,
wooded grassland, and upland grassland, ca. 300 m. Vouchers: Taylor 1212, Magogo FC & Glover PE 206 (EA).
Brachiaria chusqueoides (Hack.) Clayton Habit: Herb. Habitat: Coastal bushland
and forest in shade, rarely inland, 0–274 m. Vouchers: Luke WRQ & Robertson
SA 2193, Magogo & Glover 770, Gillespie 303 (EA).
Brachiaria deflexa (Schumach.) C. E. Hubb. ex Robyns Habit: Herb. Habitat: Deciduous bushland, margins or riverine forest, 0–300 m. Vouchers: Luke PA &
WRQ 6138, Magogo FC & Glover PE 814 (EA).
Brachiaria dictyoneura (Fig. & De Not.) Stapf Habit: Herb. Habitat: Wooded grassland and deciduous bushland, ca. 300 m. Vouchers: Moomaw JC 970, Magogo
FC & Glover PE 901 (EA).
Brachiaria distachya (L.) Stapf Habit: Herb. Habitat: Weedy places, 0–300 m. Vouchers: Allan 117, Bogdan 3333, Graham MD 13, Luke WRQ & PA 6261 (EA).
Brachiaria leersioides (Hochst.) Stapf Habit: Herb. Habitat: Waysides, old farmland, and weedy places, ca. 0–200 m. Vouchers: Patternson GD 29, Polhill &
Paulo 745 (EA).
Brachiaria leucacrantha (K. Schum.) Stapf Habit: Herb. Habitat: Deciduous coastal bushland and disturbed places, ca. 0–250 m. Vouchers: Greenway PJ & Rawlins
SP 11337, Magogo FC & Glover PE 714, Drummond & Hemsley 3812 (EA).
Brachiaria lindiensis (Pilg.) Clayton Habit: Herb. Habitat: In shade of open forest,
0–250 m. Voucher: Faden RB, Evans A & Rathbun G 71/690 (EA).
Brachiaria longiflora Clayton Habit: Herb. Habitat: Coastal bushland, 0–30 m.
Vouchers: Hacker JB 81, Faden 71/807, Gillespie 124 (EA).
Brachiaria reptans (L.) C.A. Gardner & C.E. Hubb. Habit: Herb. Habitat: Roadsides and weedy places, 0–400 m. Vouchers: Leauthaud C 123, Church 85, Bogdan 5329, Faden 74/116 (EA).
Brachiaria rugulosa Stapf Habit: Herb. Habitat: Streamside, ca. 15 m. Voucher:
Bogdan 4698 (EA).
Brachiaria serrifolia (Hochst.) Stapf Habit: Herb. Habitat: Deciduous bushland, ca.
400 m. Voucher: Luke WRQ & Robertson SA 2176 (EA).
Cenchrus biflorus Roxb. Habit: Herb. Habitat: A noxious weed of old farm land and
waste places, 0–650 m. Voucher: GR Williams 818 (EA).
Cenchrus ciliaris L. Habit: Herb. Habitat: Deciduous bushland and wooded grassland, ca. 0–350 m. Voucher: Luke PA & WRQ 6133 (EA).
Checklist of coastal forests of Kenya
149
Cenchrus echinatus L. Habit: Herb. Habitat: Roadside and waste places, 0–350 m.
Vouchers: Spjut RW 2630, Luke WRQ & PA 6266, Magogo & Glover 1060 &
816, Bally 4704 (EA).
Cenchrus mitis Andersson Habit: Herb. Habitat: Coastal bushland on sandy soils,
0–100 m. Vouchers: Kimeu JM 663, Magogo FC & Glover PE 816, Drummond
& Hemsley 3816, Greenaway 10832, Polhill & Paulo 574 (EA).
Cenchrus pedicellatus subsp. unispiculus (Brunken) Morrone Habit: Herb. Habitat: Coastal bushland and disturbed sites, ca. 100 m. Voucher: Jeffrey 538 (EA).
Cenchrus polystachios (L.) Morrone. subsp. polystachios Habit: Herb. Habitat: Old
farmland and disturbed places, ca. 0–303 m. Vouchers: Ngumbau V & Mwadime
N V036 (EA, HIB), Pettersson GD 6, Magogo FC & Glover PE 824, Polhill &
Paulo 744 (EA).
Cenchrus polystachios subsp. atrichus (Stapf & C.E. Hubb.) Morrone Habit: Herb.
Habitat: Seasonally flooded grassland on sand or clay, 0–420 m. Vouchers: DKS
Grant 878, Rawlins 401 (EA).
Cenchrus purpureus (Schumach.) Morrone Habit: Herb. Habitat: Riverine sites, valley bottoms, and forest margins, ca. 0–300 m. Voucher: Allan 506 (EA).
Chloris barbata Sw. Habit: Herb. Habitat: Waste places or disturbed ground usually
near the coast, 0–400 m. Vouchers: Davis RM 1256, Luke WRQ & PA 6263,
Napier 6379, Leah & Bayliss 10294 (EA).
Chloris gayana Kunth Habit: Herb. Habitat: Open grassland, ca. 120 m. Voucher:
Magogo FC & Glover PE sr.
Chloris mossambicensis K. Schum. Habit: Herb. Habitat: Grassland with scattered
trees or bushland, 200–381 m. Vouchers: Patterson GD 24, Magogo FC & Glover
PE 823, Drummond & Hemsley 4189, Bogdan 5429 (EA).
Chloris pycnothrix Trin. Habit: Herb. Habitat: Grassland, deciduous woodland, and
disturbed ground, ca. 200 m. Voucher: Magogo FC & Glover PE 240 (EA).
Chloris virgata Sw. Habit: Herb. Habitat: Scattered tree grassland, bushland, and
disturbed habitats, ca. 10 m. Voucher: Rawlins 682 (EA).
Chloris woodii Renvoize Habit: Herb. Habitat: Deciduous bushland, ca. 270 m.
Voucher: Wood D 1363 (EA).
Chrysopogon plumulosus Hochst. Habit: Herb. Habitat: Deciduous bushland and
sub-desert grassland, 0–350 m. Voucher: Graham A4 (EA).
Cleistachne sorghoides Benth. Habit: Herb. Habitat: On fallow land following cultivation, ca. 0–50 m. Vouchers: Allan 96, Moggridge 455, Bogdan 2549 (EA).
Coelorachis lepidura Stapf Habit: Herb. Habitat: Swampy grassland, 0–300 m.
Voucher: Allan 188 (EA).
Coix lacryma-jobi L. Habit: Herb. Habitat: Wet places in grassland, ca. 90 m. Voucher: Luke WRQ 3305 (EA).
Ctenium concinnum Nees Habit: Herb. Habitat: Open grassy places in deciduous
bushland, 250 m. Vouchers: Moomaw JC 1156, Bogdan A 3905 (EA).
Cymbopogon caesius (Hook. & Arn.) Stapf Habit: Herb. Habitat: Wooded grassland, deciduous bushland, and open grassland, 0–322 m. Vouchers: Magogo FC
& Glover PE 557, Drummond & Hemsley 1073 (EA).
150
Veronicah M. Ngumbau et al. / PhytoKeys 147: 1–191 (2020)
Cymbopogon commutatus (Steud.) Stapf Habit: Herb. Habitat: Deciduous bushland
and sub-desert grassland, ca. 100 m. Voucher: Praet van (EA).
Cynodon dactylon (L.) Pers. Habit: Herb. Habitat: Roadsides, old farmland, and
weedy, 0–410 m. Voucher: Bogdan 2531 (EA).
Cynodon nlemfuensis Vanderyst var. nlemfuensis Habit: Herb. Habitat: Open places
in deciduous bushland or forest, ca. 300 m. Voucher: Luke WRQ & PA 6258 (EA).
Cynodon nlemfuensis var. robustus Clayton & Harlan Habit: Herb. Habitat: Clearings in forest, deciduous bushland, cattle paddocks, and old cultivation, ca. 300 m.
Voucher: Verdcourt 3944 (EA).
Cyrtococcum trigonum (Retz.) A. Camus Habit: Herb. Habitat: In forest shade,
0–400 m. Vouchers: Luke WRQ & PA 8196, Drummond & Hemsley 1193, Bogdan 4713, Greenway & Rawlins 9368 (EA).
Dactyloctenium aegyptium (L.) Willd. Habit: Herb. Habitat: Grassland, open woodland, roadsides, and waste ground, ca. 10 m. Voucher: Magogo FC & Glover PE
1081 (EA).
Dactyloctenium aristatum Link Habit: Herb. Habitat: A seashore grass on sand and
exposed coral outcrops, sea level. Vouchers: Leauthaud C 39, Greenway 9263,
Rawlins 121, Gillespie 238 (EA).
Dactyloctenium australe Steud. Habit: Herb. Habitat: Bushland and grassland, ca.
0–800 m. Voucher: Bogdan A 5430 (EA).
Dactyloctenium ctenoides (Steud.) Bosser Habit: Herb. Habitat: Sand dunes, coral
outcrops on or near the seashore, sea level. Vouchers: Allan 30, Echlin 7, Bogdan
2532 (EA).
Dactyloctenium geminatum Hack. Habit: Herb. Habitat: Coastal bushland, 0–170 m.
Vouchers: Luke Q 6139, Bogdan 2597, Kassner 451, Gillespie 160 (EA).
Dactyloctenium giganteum B.S. Fisher & Schweick. Habit: Herb. Habitat: Roadsides, old cultivations, and other disturbed sites, ca. 200 m. Voucher: Sampson
21 (EA).
Dactyloctenium pilosum Stapf Habit: Herb. Habitat: On rocks by the sea, sea level.
Vouchers: Bogdan 4704, Harker 41a (EA).
Dactyloctenium scindicum Boiss. Habit: Herb. Habitat: Dry grassland and open
bushland, ca. 20 m. Voucher: Luke WRQ & PA 6119 (EA).
Daknopholis boivinii (A. Camus) Clayton Habit: Herb. Habitat: Coastal bushland,
near the seashore. Voucher: Hacker 127 (EA).
Dichanthium annulatum (Forssk.) Stapf Habit: Herb. Habitat: Dry open places
subject to grazing or disturbances, 0–800 m. Voucher: Hitchcock 25163 (EA).
Dichanthium foveolatum (Delile) Roberty Habit: Herb. Habitat: Deciduous bushland or sub-desert grassland, 0–800 m. Voucher: Heady HF 1358 (EA).
Digitaria abyssinica (Hochst. ex A. Rich.) Stapf Habit: Herb. Habitat: Cultivated
areas, ca. 20 m. Vouchers: Polhill & Paulo 532, Luke WRQ & PA 5937 (EA).
Digitaria argyrotricha (Andersson) Chiov. Habit: Herb. Habitat: Seashore and
coastal bushland, 0–365 m. Vouchers: Kuchar P 13588, Luke WRQ & PA 6262,
Drummond & Hemsley 3821, Greenway 10819, Bogdan 5698 (EA).
Checklist of coastal forests of Kenya
151
Digitaria aridicola Napper Habit: Herb. Habitat: Dry open deciduous bushland,
200–600 m. Voucher: Luke WRQ & Robertson SA 2187 (EA).
Digitaria ciliaris (Retz.) Koeler Habit: Herb. Habitat: Roadsides and weedy places,
0–200 m. Voucher: Hacker JB 142 (EA).
Digitaria diagonalis (Nees) Stapf Habit: Herb. Habitat: Open grassy places in a wide
range of habitats from waterlogged, 0–200 m. Voucher: Magogo FC & Glover PE
233 (EA).
Digitaria gymnotheca Clayton Habit: Herb. Habitat: In shade of trees, bushes or
rocks on sandy soils, ca. 0–30 m. Voucher: Bogdan A 2589 (EA).
Digitaria longiflora (Retz.) Pers. Habit: Herb. Habitat: Open places in deciduous
bushland, disturbed soils, old farmlands, and path sides, ca. 200 m. Voucher:
Drummond & Hemsley 4198 (EA).
Digitaria milanjiana (Rendle) Stapf Habit: Herb. Habitat: Recorded from a wide
range of habitats, ca. 10 m. Vouchers: Luke Q 6135, Magogo FC & Glover PE
296, Drummond & Hemsley 3825, Makin 298 (EA).
Digitaria nuda Schumach. Habit: Herb. Habitat: Open weedy places, ca. 300 m.
Vouchers: Church 87, Drummond & Hemsley 1079, Allan 182 (EA).
Digitaria pennata (Hochst.) T. Cooke Habit: Herb. Habitat: Dry open deciduous
bushland, ca. 100 m. Voucher: Praet van U.N.D.P/F.A.O (EA).
Digitaria perrottetii (Kunth) Stapf Habit: Herb. Habitat: Deciduous bushland,
particularly on old farmland, and disturbed places, 0–100 m. Voucher: Bogdan
4718 (EA).
Digitaria velutina (Forssk.) P. Beauv. Habit: Herb. Habitat: A weed of path sides,
farmland, and open places, 0–228 m. Voucher: Ngumbau V & Mwadime N V054
(EA, HIB).
Dignathia gracilis Stapf Habit: Herb. Habitat: Coastal bushland, 0–15 m. Vouchers:
Luke Q 6155, Greenway 10841, Polhill & Paulo 650, Greenway 9497 (EA).
Dignathia hirtella Stapf Habit: Herb. Habitat: Deciduous bushland, 300–800 m.
Voucher: Ndakala J Gang P 279 (EA).
Diheteropogon amplectens (Nees) Clayton Habit: Herb. Habitat: Deciduous bushland, coastal bushland, and wooded grassland, ca. 0–320 m. Voucher: Magogo FC
& Glover PE 57 (EA).
Dinebra retroflexa var. condensata S.M. Phillips Habit: Herb. Habitat: Grassland
and open woodland, ca. 300 m. Voucher: Moomaw 1380 (EA).
Echinochloa colona (L.) Link Habit: Herb. Habitat: A weedy species of muddy
or swampy places, ca. 0–85 m. Vouchers: Luke WRQ & PA 6269, Nyange M
507 (EA).
Echinochloa haploclada (Stapf ) Stapf Habit: Herb. Habitat: Stream banks and dry
river beds, ca. 5–85 m. Vouchers: Leauthaud C 5, Luke WRQ & PA 6259 (EA).
Echinochloa stagnina (Retz.) P. Beauv. Habit: Herb. Habitat: Swamps and standing
water, ca. 5 m. Voucher: Leauthaud C 8 (EA).
Eleusine indica (L.) Gaertn. Habit: Herb. Habitat: Roadsides, waste ground, and cultivated land, 0–213 m. Vouchers: Luke WRQ & PA 6256, Leauthaud C 18 (EA).
152
Veronicah M. Ngumbau et al. / PhytoKeys 147: 1–191 (2020)
Eleusine semisterilis S.M. Phillips Habit: Herb. Habitat: Dry open ground, ca.
300 m. Voucher: Allan 507 (EA): Endemic.
Enteropogon barbatus C.E. Hubb. Habit: Herb. Habitat: Not precisely recorded, ca.
150 m. Vouchers: VG van Someren in Herb, Amani 9863 & Bogdan 4506 (EA).
Enteropogon macrostachyus (Hochst. ex A. Rich.) Munro ex Benth. Habit: Herb. Habitat: Disturbed places, particularly in Acacia-Commiphora bushland, ca. 150–400 m.
Vouchers: Luke Q & Kabuye CHS 5121, Magogo FC & Glover PE 888 (EA).
Enteropogon rupestris (J.A. Schmidt) A. Chev. Habit: Herb. Habitat: Scattered tree
grassland, ca. 300 m. Voucher: Moomaw JC 1378 (EA).
Enteropogon sechellensis (Benth.) T. Durand & Schinz Habit: Herb. Habitat:
Coastal bushland, sea level. Vouchers: Drummond & Hemsley 3915, Greenway
10835, 9441 (EA).
Eragrostiella bifaria (Vahl) Bor Habit: Herb. Habitat: Semi-desert grassland or dry
bushland, and often among rocks, ca. 300 m. Voucher: Sheldrick D 103 (EA).
Eragrostis aethiopica Chiov. Habit: Herb. Habitat: Disturbed ground and weedy
places, ca. 100 m. Voucher: Mwadime N & Luke WRQ 2635 (EA).
Eragrostis unioloides (Retz.) Nees ex Steud. Habit: Herb. Habitat: Path sides and
cultivated land, ca. 300 m. Voucher: Bally 8936 (EA).
Eragrostis ambleia Clayton Habit: Herb. Habitat: Deciduous bushland, 200–500 m.
Voucher: Hacker 161b (EA).
Eragrostis cilianensis (All.) Vignolo ex Janch. Habit: Herb. Habitat: Path sides,
farmland, and overgrazed places, ca. 100 m. Voucher: Polhill & Paulo 906 (EA).
Eragrostis ciliaris (L.) R.Br. Habit: Herb. Habitat: Farmland, clearings, and overgrazed, ca. 0–70 m. Vouchers: Leauthaud C 37, Magogo FC & Glover PE 820,
Bogdan 2618 (EA).
Eragrostis cylindriflora Hochst. Habit: Herb. Habitat: Overgrazed places and dry
sandy riverbeds, ca. 90 m. Voucher: Greenway & Kanuri 12872 (EA).
Eragrostis exasperata Peter Habit: Herb. Habitat: Moist grassland, streamside, and
seasonally flooded places, ca. 300 m. Voucher: Heady 1375 (EA).
Eragrostis homomalla Nees Habit: Herb. Habitat: Moist soils bordering depressions,
ca. 200 m. Voucher: Estes 29 (EA).
Eragrostis inamoena K. Schum. Habit: Herb. Habitat: Floodplain grassland and
swampy grassland, ca. 300 m. Voucher: Moomaw JC 1158 (EA).
Eragrostis lappula Nees Habit: Herb. Habitat: Brachystegia woodland, ca. 350 m.
Voucher: JA Allan 108A (EA).
Eragrostis perbella K. Schum. Habit: Herb. Habitat: Coastal bushland, 0–450 m.
Vouchers: Bogdan A 3320, Moomaw 1159, Magogo & Glover 847, Linton 194 (EA).
Eragrostis pilosa (L.) P. Beauv. Habit: Herb. Habitat: Roadsides, old farmland, and
weedy places, ca. 300 m. Voucher: Hitchcock 25153 (EA).
Eragrostis racemosa (Thunb.) Steud. Habit: Herb. Habitat: Moist vegetation types,
ca. 300 m. Voucher: Magogo FC & Glover PE 229 (EA).
Eragrostis sennii Chiov. Habit: Herb. Habitat: Roadside, sea level. Voucher: Pearsson
in EAH 15302 (EA).
Checklist of coastal forests of Kenya
153
Eragrostis superba Peyr. Habit: Herb. Habitat: Deciduous bushland or wooded grassland, often in disturbed places, 0–410 m. Voucher: Luke WRQ & PA 6270 (EA).
Eragrostis sp. A of FTEA Habit: Herb. Habitat: Bushland. Voucher: Greenway 9259
(EA): Endemic.
Eragrostis tenuifolia (A. Rich.) Steud. Habit: Herb. Habitat: Forest, ca. 50 m.
Voucher: A Bogdan 4717 (EA).
Eriochloa fatmensis (Hochst. & Steud.) Clayton Habit: Herb. Habitat: Swampy
places and damp depressions, 0–300 m. Vouchers: RB & AJ Faden 74/1163, Magogo FC & Glover PE 696 (EA).
Eriochloa meyeriana (Nees) Pilg. Habit: Herb. Habitat: Swampy places and streamside, ca. 0–300 m. Voucher: Luke WRQ & PA 6265 (EA).
Eriochloa parvispiculata C.E. Hubb. Habit: Herb. Habitat: Damp places in coastal
bushland and near cultivation, 0–300 m. Vouchers: Leauthaud C 34, Luke WRQ
& PA 6264, Allan 37, Bogdan 3308, Polhill & Paulo 502 (EA).
Eriochloa stapfiana Clayton Habit: Herb. Habitat: Swampy places and streamside,
0–800 m. Voucher: Polhill & Paulo 831 (EA).
Eustachys paspaloides (Vahl) Lanza & Mattei Habit: Herb. Habitat: Deciduous
bushland and dry grassland, 300 m. Voucher: Magogo FC & Glover PE 372 (EA).
Hackelochloa granularis (L.) Kuntze Habit: Herb. Habitat: Disturbed soils around
habitations and old farmland, 0–123 m. Vouchers: Luke WRQ 3301, Drummond
& Hemsley 1090, Bogdan 3906 (EA).
Halopyrum mucronatum (L.) Stapf Habit: Herb. Habitat: Coastal sand dunes,
sea level. Vouchers: Luke Q 6113, Polhill & Paulo 766, Greenway & Rawlins
8913 (EA).
Heteropogon contortus (L.) P. Beauv. ex Roem. & Schult. Habit: Herb. Habitat:
Deciduous bushland and wooded grassland, 0–20 m. Vouchers: Magogo FC &
Glover PE 399, Drummond & Hemsley 4196 (EA).
Heteropogon melanocarpus (Elliott) Benth. Habit: Herb. Habitat: Abandoned cultivation, bushland and Brachystegia wooded grassland, ca. 80 m. Vouchers: Drummond & Hemsley 3949, Bogdan 2552 & 2614 (EA).
Holcolemma inaequale Clayton Habit: Herb. Habitat: Coastal bushland, 10–200 m.
Vouchers: Polhill & Paulo 564 & 619 (EA).
Hylebates chlorochloe (K. Schum.) Napper Habit: Herb. Habitat: Shady places in
forest or bushland, 50–300 m. Vouchers: Ngumbau V & Mwadime N V023 (EA,
HIB), Magogo FC & Glover PE 944 & 641, Bogdan 3909 (EA).
Hyparrhenia filipendula (Hochst.) Stapf Habit: Herb. Habitat: Wide range of soil
and vegetation types, particularly those subject to disturbances, ca. 200 m. Vouchers: Hacker JB 140, Magogo FC & Ester R 1212A (EA).
Hyparrhenia poecilotricha (Hack.) Stapf Habit: Herb. Habitat: Deciduous bushland, 0–304 m. Vouchers: Drummond RB & Hemsley JH 1081, Ross K 102 (EA).
Hyparrhenia rufa (Nees) Stapf Habit: Herb. Habitat: Deciduous bushland and
wooded grassland, ca. 100 m. Vouchers: Magogo FC & Glover PE 658, Verdcourt
1868 (EA).
154
Veronicah M. Ngumbau et al. / PhytoKeys 147: 1–191 (2020)
Hyperthelia dissoluta (Nees ex Steud.) Clayton Habit: Herb. Habitat: Deciduous
bushland and wooded grassland, ca. 0–326 m. Vouchers: Magogo FC & Glover
PE 63, Mwadime N & Gaya H 71 (EA).
Imperata cylindrica (L.) Raeusch. Habit: Herb. Habitat: Cultivated areas, ca.
0–381 m. Vouchers: Kimeu JM Meso M & Ot 606, Magogo FC & Glover PE
381, Drummond & Hemsley 1031 (EA).
Ischaemum rugosum Salisb. Habit: Herb. Habitat: Damp soils, pond margins, and
rice fields, 0–50 m. Voucher: Greenway & Rawlins 9482 (EA).
Leersia hexandra Sw. Habit: Herb. Habitat: In shallow water, ca. 40 m. Vouchers:
Leauthaud C 7, Nyange M 339 (EA).
Leptaspis zeylanica Nees ex Steud. Habit: Herb. Habitat: Ground layer in forest, ca.
400 m. Voucher: Drummond RB & Hemsley JH 1127 (EA).
Leptochloa chinensis (L.) Nees Habit: Herb. Habitat: Aquatic or semi-aquatic, in damp
hollows and in shallow water at pond margins, 0–400 m. Voucher: Sampson 63 (EA).
Leptochloa fusca (L.) Kunth Habit: Herb. Habitat: Shallow margins of lakes and rivers, 0–100 m. Voucher: Bogdan 2605 (EA).
Leptochloa obtusiflora Hochst. Habit: Herb. Habitat: Deciduous bushland and grassland, ca. 0–300 m. Vouchers: Magogo FC & Glover PE 871, Moomaw 1160 (EA).
Leptochloa panicea (Retz.) Ohwi Habit: Herb. Habitat: Deciduous bushland, wooded grassland, plantations, roadsides, and waste places, 0–300 m. Vouchers: Magogo FC & Glover PE 1051, Bogdan 5431 & 3305, Rawlins 446 (EA).
Leptochloa uniflora Hochst. ex A. Rich. Habit: Herb. Habitat: Wooded grassland
and deciduous bushland, 0–100 m. Vouchers: Magogo FC & Glover PE 368,
Verdcourt 1913, Bogdan 3910, Gillespie 195 (EA).
Leptothrium senegalense (Kunth) Clayton Habit: Herb. Habitat: Deciduous bushland and sub-desert grassland, ca. 5 m. Voucher: Luke Q 6117(EA).
Lepturus radicans (Steud.) A. Camus Habit: Herb. Habitat: Sandy in grassland,
0–300 m. Vouchers: Luke Q 6118, JA Allan 94, Napper 1665, Bogdan 4711 (EA).
Lepturus repens (J.R. Forst.) R.Br. Habit: Herb. Habitat: Coastal sands above highwater mark. Vouchers: JA Allan 97, Bogdan 3642, Longridge 24 (EA).
Megastachya mucronata (Poir.) P. Beauv. Habit: Herb. Habitat: Forest shade, ca.
0–400 m. Vouchers: SAJIT–005516, Ngumbau V & Mwadime N V0103 (EA,
HIB), Kirika P, Nyamongo D & Sanyanyi S 04/13/2008, Robertson SA 3394,
Drummond & Hemsley 3835, Bogdan 3912, Greenway & Rawlins 9347 (EA).
Melinis repens (Willd.) Zizka subsp. repens Habit: Herb. Habitat: Disturbed areas,
ca. 43 m. Vouchers: Magogo FC & Glover PE 717, Luke WRQ 16133 (EA).
Olyra latifolia L. Habit: Herb. Habitat: Edges of clearings and path sides in forests,
ca. 300 m. Vouchers: Ngumbau V & Mwadime N V0164 (EA, HIB), Verdcourt
B 3930, Lucas, Jeffery & Kirika 229, JA Allan 501 (EA).
Oplismenus burmannii (Retz.) P. Beauv. Habit: Herb. Habitat: In shade of forest or
bushland, 0–300 m. Vouchers: Brathys Expenditure 67, Drummond & Hemsley
3998, Hacker 5, Bogdan 4716 (EA).
Oplismenus compositus (L.) P. Beauv. Habit: Herb. Habitat: Forest shade, ca. 200 m.
Voucher: Magogo FC & Glover PE 940 (EA).
Checklist of coastal forests of Kenya
155
Oplismenus hirtellus (L.) P. Beauv. Habit: Herb. Habitat: Forest shade, ca. 0–246 m.
Vouchers: Luke WRQ et al. 6181, Magogo & Glover 963 (EA).
Oropetium minimum (Hochst.) Pilg. Habit: Herb. Habitat: Dry grassland and open
deciduous bushland, ca. 780 m. Vouchers: Legesse A 119, Pratt DJ 699 & 402 (EA).
Oryza eichingeri Peter Habit: Herb. Habitat: forest, ca. 300 m. Vouchers: Luke WRQ
& Robertson SA 2295, Magogo FC & Glover PE 769 (EA).
Oryza longistaminata A. Chev. & Roehr. Habit: Herb. Habitat: Swamp and flood plain
grassland, ca. 0–100 m. Vouchers: Leauthaud C 23, JA Allan 111, Sampson 58 (EA).
Oryza punctata Kotschy ex Steud. Habit: Herb. Habitat: Swampy soils by stream
banks and pond margins, 0–280 m. Vouchers: SAJIT–006225, Ngumbau V &
Mwadime N V0258 (EA, HIB), Leauthaud C 33, Luke WRQ 3602, Taylor 1211,
Polhill & Paulo 484, Bogdan 5428 (EA).
Panicum atrosanguineum Hochst. ex A. Rich. Habit: Herb. Habitat: Disturbed
places and old farmland in deciduous bushland, ca. 100 m. Voucher: Magogo &
Glover 187 (EA).
Panicum brevifolium L. Habit: Herb. Habitat: Forest shade, ca. 15 m. Vouchers:
SAJIT–006216 (EA, HIB), Magogo FC & Glover PE 114, Bogdan 3341, Drummond & Hemsley 1128 (EA).
Panicum deustum Thunb. Habit: Herb. Habitat: Forest, deciduous bushland, and
grassland, ca. 0–70 m. Vouchers: Magogo FC & Glover PE 281, Allan 165 (EA).
Panicum genuflexum Stapf Habit: Herb. Habitat: Wooded grassland or deciduous
bushland, ca. 0–100 m. Vouchers: Drummond & Hemsley 3814, Bogdan 3300,
Gillespie 193, Muchiri J 550 (EA).
Panicum hippothrix K. Schum. ex Engl. Habit: Herb. Habitat: Deciduous bushland
and margins of cultivation, ca. 0–80 m. Vouchers: Rawlins SP 399, Drummond &
Hemsley 3823, Bogdan 3307, Moomaw 994, Heady 1378 (EA).
Panicum hirtum Lam. Habit: Herb. Habitat: Deciduous bushland or wooded grassland, ca. 30 m. Voucher: Gilllespie 295 (EA).
Panicum infestum Andersson Habit: Herb. Habitat: Grassland, bushland or deciduous woodland, ca. 0–266 m. Vouchers: Ngumbau V & Mwadime N V047 (EA,
HIB), Luke Q 6121, Magogo FC & Glover PE 828 (EA).
Panicum laticomum Nees Habit: Herb. Habitat: Forest shade, 0–800 m. Vouchers:
Ngumbau V & Mwadime N V053 (EA, HIB), Robertson SA & Luke WRQ 5496,
Magogo FC & Glover PE 457, Verdcourt 1874 & 1080 (EA).
Urochloa maxima (Jacq.) R. D. Webster subsp. maxima Habit: Herb. Habitat:
Woodland, deciduous bushland, roadsides, and river banks, ca. 0–381 m. Vouchers: Ngumbau V & Mwadime N V102 (EA, HIB), Luke PA & WRQ 6114, Magogo FC & Glover PE 395 (EA).
Panicum parvifolium Lam. Habit: Herb. Habitat: Swamps, ca. 0–100 m. Vouchers:
Luke WRQ et al. 4332, Bogdan 5346 (EA).
Panicum peteri Pilg. Habit: Herb. Habitat: Forests and Brachystegia woodland, 100–
800 m. Voucher: Ngumbau V & Mwadime N V007 (EA, HIB).
Panicum pinifolium Chiov. Habit: Herb. Habitat: Coastal bushland and sand dunes,
sea level. Vouchers: Greenway & Rawlins 9286, Rawlins & Oseni (EA).
156
Veronicah M. Ngumbau et al. / PhytoKeys 147: 1–191 (2020)
Panicum pleianthum Peter Habit: Herb. Habitat: Forest shade, 50–550 m. Vouchers:
Ngumbau V & Mwadime N V011 (EA, HIB), Robertson SA 5124, Robertson SA
3395A, Magogo & Glover 236, Bogdan 3913, Verdcourt 3916 (EA).
Panicum stoloniferum Poir. Habit: Herb. Habitat: Bush or forest shade or sandy soils,
ca. 0–300 m. Vouchers: Magogo FC & Glover PE 332, Drummond & Hemsley
3826 (EA).
Panicum trichoides Sw. Habit: Herb. Habitat: Forest shade, 0–900 m. Vouchers:
Moggridge 448, Graham 45 (EA).
Paspalum glumaceum Clayton Habit: Herb. Habitat: Damp places and forest margins, ca. 0–100 m. Vouchers: Magogo FC & Glover PE 664, Allan 110 & 5 (EA).
Paspalum scrobiculatum L. Habit: Herb. Habitat: Damp places and forest margins,
ca. 50 m. Vouchers: Magogo FC & Glover PE 324 & 106 (EA).
Paspalum vaginatum Sw. Habit: Herb. Habitat: Coastal salt marshes, but sometimes
also in inland marshes, 0–100 m. Vouchers: Allan 114, Bogdan 3314 (EA).
Perotis hildebrandtii Mez Habit: Herb. Habitat: Open places, roadsides, and wasteland, ca. 0–400 m. Vouchers: Magogo FC & Glover PE 369, Greenway 10834 (EA).
Perotis patens Gand. Habit: Herb. Habitat: Path sides and weedy places, ca. 300 m.
Vouchers: Drummond & Hemsley 1099, Magogo FC & Glover PE 109 (EA).
Phragmites australis (Cav.) Trin. ex Steud. subsp. australis Habit: Herb. Habitat:
Shallow water of streams, rivers lakes and in swampy places, 0–300 m. Vouchers:
Moomaw 961, Battiscombe 266, Kimeu JM, Meso M & Otieno V 616 (EA).
Rhytachne rottboellioides Desv. ex Ham. Habit: Herb. Habitat: In swamps and seasonally wet grassland, 0–300 m. Voucher: Moomaw JC 1155 (EA).
Rottboellia cochinchinensis (Lour.) Clayton Habit: Herb. Habitat: Disturbed places,
0–110 m. Voucher: Magogo FC & Glover PE 1135 (EA).
Saccharum spontaneum subsp. aegyptiacum (Willd.) Hack. Habit: Herb. Habitat:
Damp soils, often fringing rivers and lakes, ca. 100 m. Vouchers: Hacker JB 93,
DC Edwards 53 (EA).
Sacciolepis curvata (L.) Chase Habit: Herb. Habitat: Damp shady places subject
to disturbances, ca. 0–100 m. Vouchers: Kirika P, Muthoka P & Mbale M 750,
Magogo FC & Glover PE 672, Bogdan 3309, Greenway 10836, Polhill & Paulo
620 (EA).
Schizachyrium brevifolium (Sw.) Buse Habit: Herb. Habitat: Damp or shady places,
ca. 200 m. Vouchers: Bogdan 5351, 2611 & 4700 (EA).
Schizachyrium exile (Hochst.) Pilg. Habit: Herb. Habitat: Bushland and woodland,
ca. 600 m. Voucher: Allan 75 (EA).
Schizachyrium rupestre (K. Schum.) Stapf Habit: Herb. Habitat: Moist places in
coastal bushland, ca. 100 m. Voucher: Allan 75 (EA).
Schizachyrium sanguineum (Retz.) Alston Habit: Herb. Habitat: Bushland and
woodland, ca. 0–365 m. Vouchers: Festo L & Luke Q 2614, Magogo FC & Glover PE 511 (EA).
Schoenefeldia transiens (Pilg.) Chiov. Habit: Herb. Habitat: Wooded grassland, deciduous bushland, and dry grassland, 5–15 m. Vouchers: Polhill & Paulo 823, RB
& AJ Faden74/1167 (EA).
Checklist of coastal forests of Kenya
157
Setaria incrassata (Hochst.) Hack. Habit: Herb. Habitat: Swamps to stony hillsides
or margins of upland evergreen forest, ca. 0–457 m. Vouchers: Patterson G 27,
Ross K 17 (EA).
Setaria sulcata Raddi Habit: Herb. Habitat: Shady places and around forest, ca.
200 m. Vouchers: Ngumbau V & Mwadime N V034 (EA, HIB), Magogo FC &
Glover PE 393 & 787, Allan 500 (EA).
Setaria obtusifolia (Delile) Morrone Habit: Herb. Habitat: Marshy soils or shallow
water, ca. 0–25 m. Vouchers: Leauthaud C 24, Allan 166 (EA).
Setaria pumila (Poir.) Roem. & Schult. Habit: Herb. Habitat: Path sides, bare
patches, and grazing area, ca. 0–300 m. Voucher: Magogo FC & Glover PE
1085 (EA).
Setaria punctata (Burm.f.) Veldkamp Habit: Herb. Habitat: On wet soils or in water, 0–400 m. Voucher: Magogo FC & Glover PE 115 (EA).
Setaria sphacelata (Schumach.) Stapf & C.E. Hubb. ex Moss Habit: Herb. Habitat: Deciduous bushland or wooded grassland, swamps, and riversides, ca. 20 m.
Vouchers: Magogo FC & Glover PE 23, Moomaw JC 1254 (EA).
Sorghum bicolor subsp. verticilliflorum (Steud.) de Wet ex Wiersema & J. Dahlb.
Habit: Herb. Habitat: Swampy soils, streamside, disturbed places, and old farmland, 0–60 m. Vouchers: Greenway 8962, Moomaw JC 962 (EA).
Sorghum versicolor Andersson Habit: Herb. Habitat: Bushland, deciduous bushland,
and wooded grassland, 0–400 m. Vouchers: Petterson GD 28, Kirika P, Mbale M
& Mbatha M 767, Drummond & Hemsley 3775, Polhill & Paulo 711 (EA).
Sporobolus congoensis Franch. Habit: Herb. Habitat: Wooded grassland, ca. 304 m.
Vouchers: Magogo FC & Glover PE 550, Moomaw 1269 (EA).
Sporobolus coromandelianus (Retz.) Kunth Habit: Herb. Habitat: Rare open places,
0–100 m. Voucher: Hitchcock 25158 (EA).
Sporobolus ioclados (Trin.) Nees Habit: Herb. Habitat: Saline grasslands, seashore
sands, and sandy patches in mangrove swamps, ca. 0–100 m. Voucher: Polhill &
Paulo 687 (EA).
Sporobolus microprotus Stapf Habit: Herb. Habitat: Deciduous bushland, 0–100 m.
Voucher: Bogdan 3331A (EA).
Sporobolus pyramidalis P. Beauv. Habit: Herb. Habitat: Disturbed areas, ca.
0–266 m. Vouchers: Ngumbau V & Mwadime N V051 (EA, HIB), Luke WRQ
& PA 6268 (EA).
Sporobolus spicatus (Vahl) Kunth Habit: Herb. Habitat: Grassland and open bushland, ca. 0–100 m. Voucher: Bogdan A 4707 (EA).
Sporobolus stolzii Mez Habit: Herb. Habitat: Roadsides and waste places in deciduous bushland, ca. 0–80 m. Vouchers: JA Allan 81, Drummond & Hemsley 3799,
Boyle B 127 (EA).
Sporobolus subglobosus A. Chev. Habit: Herb. Habitat: Coastal bushland, up to
300 m. Voucher: Moomaw JC 1096 (EA).
Sporobolus tenuissimus (Mart. ex Schrank) Kuntze Habit: Herb. Habitat: Roadsides
and waste places, ca. 0–200 m. Vouchers: JA Allan 8, Burtt Davy 2695, Bogdan
2542, Moomaw JC 1278 (EA).
158
Veronicah M. Ngumbau et al. / PhytoKeys 147: 1–191 (2020)
Sporobolus virginicus (L.) Kunth Habit: Herb. Habitat: Sandy seashores, 0–300 m.
Vouchers: SAJIT–006248 (EA, HIB), Bogdan 2632, Gillespie 110, Greenway &
Rawlins 8911, Luke Q 6112 (EA).
Stenotaphrum dimidiatum (L.) Brongn. Habit: Herb. Habitat: Seashore, usually in
shade, 0–10 m. Vouchers: Ngumbau V & Mwadime N V0512 (EA, HIB), Drummond & Hemsley 3996, Bogdan 3645, Greenway & Rawlins 9465 (EA).
Tetrapogon roxburghiana (Schult.) P. M. Peterson Habit: Herb. Habitat: Wooded
grassland, bushland, and disturbed habitats, ca. 30 m. Vouchers: Hacker JB 111,
Luke WRQ & PA 6260, Drummond & Hemsley 3849 (EA).
Tetrapogon bidentatus Pilg. Habit: Herb. Habitat: Deciduous bushland or grassland,
ca. 30–65 m. Vouchers: TPR 282, RB & AJ Faden 74/1049 (EA).
Tetrapogon cenchriformis (A. Rich.) Clayton Habit: Herb. Habitat: Deciduous
bushland or grassland, ca. 50 m. Voucher: J Makin 14667 (EA).
Tetrapogon tenellus (J. Koenig ex Roxb.) Chiov. Habit: Herb. Habitat: Deciduous
bushland or grassland, ca. 0–65 m. Vouchers: Leauthaud C 6, Magogo FC &
Glover PE 881, Drummond & Hemsley 4127 (EA).
Themeda triandra Forssk. Habit: Herb. Habitat: Deciduous bushland, ca. 0–381 m.
Vouchers: Robertson SA & Luke WRQ 5113, Magogo FC & Glover PE 827 (EA).
Trachypogon spicatus (L.f.) Kuntze Habit: Herb. Habitat: Deciduous bushland or
wooded grassland, ca. 0–370 m. Voucher: Magogo FC & Glover PE 330 (EA).
Tragus berteronianus Schult. Habit: Herb. Habitat: Overgrazed places, ca. 0–100 m.
Voucher: Magogo FC & Glover PE 932 (EA).
Tragus heptaneuron Clayton Habit: Herb. Habitat: Deciduous bushland, 0–800 m.
Vouchers: Bogdan A 5333, Polhill & Paulo 515 (EA).
Tragus racemosus (L.) All. Habit: Herb. Habitat: Deciduous bushland, ca. 200 m.
Vouchers: Amani 9868A, VGL van Someren (EA).
Trichoneura mollis (Kunth) Ekman Habit: Herb. Habitat: Dry bushland, 100–
230 m. Vouchers: Koss 114, Makin (EA).
Urochloa mosambicensis (Hack.) Dandy Habit: Herb. Habitat: Wooded grassland,
deciduous bushland, and disturbed sites, ca. 0–100 m. Voucher: Allan 79 (EA).
Urochloa panicoides P. Beauv. Habit: Herb. Habitat: Old farmland, overgrazed, and
deciduous bushland, ca. 0–170 m. Voucher: Leauthaud C 36 (EA).
Urochloa rudis Stapf Habit: Herb. Habitat: Coastal bushland, 0–30 m. Vouchers:
Allan 151 & 173, Polhill & Paulo 589 (EA).
Urochloa sclerochlaena Chiov. Habit: Herb. Habitat: Coastal and semi-desert bushland, ca. 200 m. Voucher: Thairu 125 (EA).
Urochloa setigera (Retz.) Stapf Habit: Herb. Habitat: Lowland and riverine forest,
0–900 m. Voucher: Greenway & Rawlins 9365 (EA).
Urochloa trichopus (Hochst.) Stapf Habit: Herb. Habitat: Wooded grassland and
coastal bushland, 0–100 m. Vouchers: Drummond & Hemsley 3841, Bogdan
2529, Gillespie 10, Luke Q 6140 (EA).
Vossia cuspidata (Roxb.) Griff. Habit: Herb. Habitat: In or close to water, often floating, 0–200 m. Voucher: Leauthaud C 12 (EA).
Checklist of coastal forests of Kenya
159
F141. Polygalaceae
3 Genera, 17 Species
Carpolobia goetzei Gürke Habit: Shrub. Habitat: Moist or dry forest, riverine forest, ca. 0–210 m. Vouchers: SAJIT–006453, Ngumbau V & Mwadime N V0389
(EA, HIB), Drummond & Hemsley 1135, Adams 93, Festo L, Luke Q & P 2759,
Magogo FC & Glover PE 269 (EA).
Polygala amboniensis Gürke Habit: Herb. Habitat: Bushland, secondary grassland,
and coastal dunes, 0–100 m. Vouchers: SAJIT–006098 (EA, HIB), Polhill & Paulo 594, Luke & Saidi 6250, Luke WRQ 3337 & 2796 (EA).
Polygala arenaria Willd. Habit: Herb. Habitat: Grassland, scattered tree grassland, ruderal sites, ca. 0–200 m. Vouchers: Makin 425, Luke 3572, Luke WRQ
2453A(EA).
Polygala conosperma Bojer Habit: Herb. Habitat: Wooded grassland, 0–10 m.
Vouchers: Blake 2278, Luke 3323 (EA).
Polygala erioptera DC. Habit: Herb. Habitat: Grassland, sandy, and muddy roadside,
30 m. Voucher: Klotzli F et al. 601 (EA).
Polygala fischeri Gürke Habit: Herb. Habitat: Various habitats, ca. 0–300 m. Vouchers: Loveridge JP 13, Drummond & Hemsley 1074 (EA).
Polygala irregularis Boiss. Habit: Herb. Habitat: Acacia-Commiphora bushland on
dunes or white sand, 0–500 m. Vouchers: Luke 5441, Luke PA & WRQ 5441 (EA).
Polygala kilimandjarica Chodat Habit: Herb. Habitat: Acacia-Commiphora bushland, secondary bushland, ca. 0–300 m. Vouchers: Luke 3494, Festo L & Luke Q
2658, Luke WRQ 3156 (EA).
Polygala meonantha Chodat Habit: Herb. Habitat: Scattered tree grassland, Acacia
bushland, woodland, 0–700 m. Voucher: Harvey, Mwachala & Vollesen 59 (EA).
Polygala paniculata L. Habit: Herb. Habitat: Cultivated land, grassland, path and
roadsides, ca. 350 m. Voucher: Magogo & Estes 1231 (EA).
Polygala petitiana A. Rich. Habit: Herb. Habitat: Brachystegia-Julbernadia woodland, grassland, and cultivated ground, ca. 15 m. Vouchers: Robertson 3313, Luke
PA & WRQ 5974, Luke 10344K (EA).
Polygala sadebeckiana Gürke Habit: Herb. Habitat: Riverine forest, bushland, and
forest margin, ca. 10–200 m. Voucher: Magogo & Glover 996 (EA).
Polygala sansisbarensis Gürke Habit: Herb. Habitat: Wet places and seasonally wet
grassland, 0–10 m. Vouchers: Tweedie 3181, Luke & Gray 4054, 2nd Herbarium
Technique Course 075, Gillett JB 20433 (EA).
Polygala senensis Klotzsch Habit: Herb. Habitat: Scattered tree grassland, 0–500 m.
Voucher: Gillet 16535 (EA).
Polygala sphenoptera Fresen. Habit: Herb. Habitat: Evergreen bushland, wooded
grassland, ca. 0–227 m. Vouchers: SAJIT–005941, Ngumbau V & Mwadime N
V0277 (EA, HIB), Drummond & Hemsley 1074, Festo L, Luke Q & P 2636 (EA).
Polygala stenopetala Klotzsch Habit: Herb. Habitat: Brachystegia-Julbernadia woodland, grassland, and cultivated ground, 0–280 m. Vouchers: SAJIT–005937,
160
Veronicah M. Ngumbau et al. / PhytoKeys 147: 1–191 (2020)
Ngumbau V & Mwadime N V0248 (EA, HIB), Verdcourt 3911, Magogo &
Glover 394 (EA).
Securidaca longipedunculata Fresen. Habit: Shrub. Habitat: Wooded and bushed
grassland, ca. 0–300 m. Vouchers: SAJIT–005480 (EA, HIB), Greenway 9006,
Magogo FC & Glover PE 169 (EA).
F142. Polygonaceae
3 Genera, 7 Species
Antigonon leptopus Hook. & Arn. Habit: Climber. Habitat: Thickets, rainforest
margins, and coastal sand dunes, 0–600 m. Vouchers: Kimeu JM 538, Starzenska
6 (EA).
Oxygonum atriplicifolium (Meisn.) Martelli Habit: Herb. Habitat: Hedgerows, cultivated, and waste ground. Vouchers: Bogdan 3298, Verdcourt 1087, Tweedie 942,
Verdcourt B 2112 (EA).
Oxygonum sagittatum R.A. Graham Habit: Herb. Habitat: Sand ground, disturbed
area, grassland, ca. 150 m. Voucher: Kokwaro JO 3571 (EA).
Oxygonum salicifolium Dammer Habit: Herb. Habitat: Grasslands and disturbed
ground, 0–156 m. Vouchers: SAJIT–004676, Ngumbau V & Mwadime N V0433
(EA, HIB), Kassner 262, RM Graham 2138, Simpson BL 259 (EA).
Oxygonum stuhlmannii Dammer Habit: Herb. Habitat: Roadsides, ca. 88 m. Vouchers: SAJIT–006452 (EA, HIB), Reitsma J 419 (EA).
Persicaria decipiens (R.Br.) K.L. Wilson Habit: Herb. Habitat: Waste land and roadsides, ca. 0–213 m. Vouchers: Luke WRQ 3599, Simpson BL 259, Magogo FC &
Glover PE 921 (EA).
Persicaria senegalensis f. albotomentosa (R.A. Graham) K.L. Wilson Habit: Herb.
Habitat: In damp places, lakes and rivers, ca. 60 m. Vouchers: Perdue RE & Kibuwa SP 10 & 159, Luke PA & WRQ 3826 (EA).
F143. Pontederiaceae
1 Genus, 1 Species
Monochoria africana (Solms) N.E.Br. Habit: Herb. Habitat: Margins of rice fields
and cleared mangrove forest, 0–100 m. Voucher: Greenway & Rawlins 9483 (EA).
F144. Portulacaceae
1 Genus, 11 Species
Portulaca ciferrii Chiov. Habit: Herb. Habitat: Sand patches on coral rag pavement,
ca. 5 m. Voucher: Greenway & Rawlins 9420 (EA).
Portulaca coralloides S.M. Phillips Habit: Herb. Habitat: Deciduous bushland on
coral and between coastal dunes, near sea level. Vouchers: Greenway & Rawlins
9423, Rawlins 94 (EA): Endemic.
Checklist of coastal forests of Kenya
161
Portulaca fascicularis Peter Habit: Herb. Habitat: Coastal deciduous bushland,
0–450 m. Voucher: Faden et al. 70/939 (EA).
Portulaca foliosa Ker Gawl. Habit: Herb. Habitat: Seasonally dry river beds or sandbanks in rivers, ca. 20 m. Voucher: Katz SS 75/2/11 (EA).
Portulaca greenwayi M.G. Gilbert Habit: Herb. Habitat: Deciduous bushland, 50–
150 m. Voucher: Ochung & Koech 86 (EA).
Portulaca kermesina N.E.Br. var. kermesina Habit: Herb. Habitat: Bushland, ca.
0–25 m. Vouchers: Rawlins SP 94 & 932, Graham RM 1623 (EA).
Portulaca kermesina var. lutea (Poelln.) S.M. Phillips Habit: Herb. Habitat: Grassland, wooded grassland, ca. 30 m. Voucher: Luke et al. TPR 253 (EA).
Portulaca oblonga Peter Habit: Herb. Habitat: Open bushland and disturbed weedy
places, ca. 0–65 m. Vouchers: Polhill & Paulo 640, RB & AJ Faden 74/1037 (EA).
Portulaca oleracea L. Habit: Herb. Habitat: Seasonally flooded grassland or sandy
areas, ca. 0–20 m. Vouchers: Kassner 463, Tweedie 935, Robertson SA 3755 (EA).
Portulaca peteri Poelln. Habit: Herb. Habitat: Deciduous bushland and thicket, ca.
30 m. Voucher: Luke et al. TPR401 (EA).
Portulaca quadrifida L. Habit: Herb. Habitat: Open disturbed grounds, ca. 50 m.
Vouchers: Robertson SA 3756 & 3455 (EA).
Portulaca wightiana Wall. ex Wight & Arn. Habit: Herb. Habitat: Rock outcrops,
0–100 m. Voucher: RB & AJ Faden 72/93 (EA).
F145. Primulaceae
1 Genus, 1 Species
Myrsine melanophloeos (L.) R. Br. Habit: Tree. Habitat: Upland forest, riverine, swamp
forest, open woodland, and thickets, ca. 100 m. Voucher: Sangai GW 1125 (EA).
F146. Putranjivaceae
1 Genus, 4 Species
Drypetes natalensis var. leiogyna Brenan Habit: Tree. Habitat: Lowland dry evergreen,
semi-deciduous forest, and riverine forest, 15–380 m. Vouchers: SAJIT–005442 &
006095 (EA, HIB), Magogo FC & Glover PE 286 (EA): Vulnerable.
Drypetes parvifolia (Müll.Arg.) Pax & K. Hoffm. Habit: Tree. Habitat: Evergreen semideciduous coastal and riverine forest, 0–450 m. Vouchers: SAJIT–005539, 005550,
005935 & 006192 (EA, HIB), Verdcourt 1893, RB & AJ Faden 71/773 (EA).
Drypetes reticulata Pax Habit: Tree. Habitat: Evergreen forest and thicket, 0–500 m.
Vouchers: Greenway 9621, Simpson 28 (EA).
Drypetes usambarica (Pax) Hutch. var. usambarica Habit: Tree. Habitat: Rainforest, 400 m. Voucher: Polhill R & Robertson 4836 (EA).
Drypetes usambarica var. mrimae Radcl.-Sm. Habit: Tree. Habitat: Lowland evergreen forest, 100–250 m. Vouchers: Luke WRQ & Robertson SA 523, Robertson
SA & Luke Q 5842, RB & AJ Faden 77/677 (EA).
162
Veronicah M. Ngumbau et al. / PhytoKeys 147: 1–191 (2020)
Drypetes usambarica var. trichogyna Radcl.-Sm. Habit: Tree. Habitat: Lowland to submontane moist forest. Voucher: Luke WRQ & Robertson SA 250 (EA): Vulnerable.
F147. Ranunculaceae
1 Genus, 1 Species
Clematis sigensis Engl. Habit: Climber. Habitat: Forest edges and bushland, ca. 350–
500 m. Vouchers: Luke WRQ 1625, Gillett JB 18715 (EA).
F148. Rhamnaceae
5 Genera, 9 Species
Berchemia discolor (Klotzsch) Hemsl. Habit: Tree. Habitat: Grassland and open
woodland, ca. 5 m. Voucher: Mrs Robertson SA 7743 (EA).
Colubrina asiatica (L.) Brongn. Habit: Shrub. Habitat: Just above high-tide level.
Vouchers: SAJIT–005596 (EA, HIB), Verdcourt 3963 & 1073, Ossent 239, Luke
Q 5686 (EA).
Lasiodiscus mildbraedii Engl. Habit: Tree. Habitat: Dry evergreen rainforest, dry
forest, and woodland, ca. 20 m. Voucher: Thomas Mwadime in Mrs Robertson
SA 7782 (EA).
Lasiodiscus pervillei subsp. ferrugineus (Verdc.) Figueiredo Habit: Shrub. Habitat:
Lowland evergreen forest, ca. 100 m. Vouchers: Verdcourt 1181, Greenway &
Rawlins 8954, Dale in FD 8834, Rawlins SP 241, Luke Q 1555 (EA).
Scutia myrtina (Burm.f.) Kurz Habit: Shrub. Habitat: Forest margins, bushland,
thicket, and wooded grassland, ca. 0–370 m. Vouchers: SAJIT–005573 (EA,
HIB), Polhill & Paulo 497, Robertson SA 4377 (EA).
Ziziphus mauritiana Lam. Habit: Tree. Habitat: Disturbed areas, ca. 30 m. Vouchers:
SAJIT–005438 & 005439 (EA, HIB), RM Graham 245, Bally PRO 215 (EA).
Ziziphus mucronata Willd. Habit: Tree. Habitat: Open scrubland, woodland, forest margins, and riverine vegetation, ca. 5 m. Vouchers: Mrs Robertson SA 7717,
Magogo FC & Glover PE 887 (EA).
Ziziphus pubescens Oliv. Habit: Tree. Habitat: Coastal forest, sand forest, and riverine fringes, ca. 0–300 m. Voucher: Verdcourt B 2410 (EA).
Ziziphus robertsoniana Beentje Habit: Tree: Habitat: Moist semi-deciduous forest
on coral rag, 10–160 m. Vouchers: Robertson SA 3645, 4274, 4310, 4889, 4675,
5097 & 6601, Mrima-Dzombo Expedition 170 (EA): Endangered.
F149. Rhizophoraceae
4 Genera, 5 Species
Bruguiera gymnorhiza (L.) Lam. Habit: Tree. Habitat: Intertidal mud-flats and estuaries, up to 10 m. Vouchers: SAJIT–005584 (EA, HIB), Greenway PJ & Rawlins
SP 8871 (EA).
Checklist of coastal forests of Kenya
163
Cassipourea celastroides Alston Habit: Shrub. Habitat: On rocky hillsides, ca.
290 m. Vouchers: Kuchar P 13438, Robertson SA 5197, Luke WRQ & Robertson
SA 283 (EA).
Cassipourea euryoides Alston Habit: Tree. Habitat: Coastal forest, scrub or rarely in
secondary bushland, 80–400 m. Vouchers: SAJIT–006012 & 006045, Ngumbau
V & Mwadime N V0144 (EA, HIB), RM Graham 1627, Drummond & Hemsley
3964, Dale 3549, Gilbert MG & Kuchar P 5832, Greenway PJ 8934 (EA).
Ceriops tagal (Perr.) C.B. Rob. Habit: Tree. Habitat: Intertidal mud flats and estuaries, sea level. Vouchers: MacNaughton 129 in Forestry Dept. 2715, Dale IR
1072 (EA).
Rhizophora mucronata Lam. Habit: Tree. Habitat: Intertidal mud-flats of shores and
estuaries, sea level. Vouchers: SAJIT–006247 (EA, HIB), Jeffery K136, Ross KS
203 (EA).
F150. Rubiaceae
63 Genera, 137 Species
Afrocanthium kilifiense (Bridson) Lantz Habit: Shrub. Habitat: Dry lowland forest or Brachystegia woodland, 30–130 m. Vouchers: Ngumbau V & Mwadime
N V0154 (EA, HIB), Luke Q 1516, Luke WRQ & Robertson SA 2789, RM
Graham in FD 1711, Musyoki & Hansen 1012, Rawlins 856 (EA): Vulnerable.
Afrocanthium peteri (Bridson) Lantz Habit: Shrub. Habitat: Forest or riverine thicket, 0–450 m. Voucher: Luke et al. TPR 659 (EA).
Afrocanthium pseudoverticillatum (S. Moore) Lantz Habit: Shrub. Habitat: Bushland, ca. 0–297 m. Vouchers: SAJIT–006037, Ngumbau V & Mwadime N V0141
(EA, HIB), Festo L, Luke Q & P 2786 & 2601, Luke WRQ & Robertson SA
2790, Kassner 383, Faden 71/713, Robertson & Luke 5616 (EA).
Agathisanthemum bojeri Klotzsch Habit: Herb. Habitat: Woodland, grassland, and
riverine forest, 25–50 m. Vouchers: SAJIT–006050 & 005473 (EA, HIB), Luke
WRQ & Robertson SA 2757 (EA).
Aidia abeidii S.E. Dawson & Gereau Habit: Tree. Habitat: Forest, 8–360 m. Vouchers: Luke Q 2948, 8317, Festo L, Luke Q & P 2762 (EA).
Breonadia salicina (Vahl) Hepper & J.R.I. Wood Habit: Tree. Habitat: Gallery forest by rivers, 0–100 m. Vouchers: Sampson 8, Luke WRQ 3597 (EA).
Bullockia mombazensis (Baill.) Razafim., Lantz & B. Bremer Habit: Shrub. Habitat: Coastal bushland, wooded grassland, and evergreen forest, ca. 256 m. Vouchers: SAJIT–004642, Ngumbau V & Mwadime N V0137 (EA, HIB), Spjut RW
4579, Gillet & Kibuwa 20023, Gillespie 374, Luke Q 1437 (EA).
Bullockia setiflora (Hiern) Razafim., Lantz & B. Bremer Habit: Shrub. Habitat:
Woodland, riverine forests, 60–750 m. Vouchers: SAJIT–006125 (EA, HIB), Luke
WRQ & Robertson SA 2808, Faden 77/675, Polhill & Robertson 4842 (EA).
Calycosiphonia spathicalyx (K. Schum.) Robbr. Habit: Shrub. Habitat: Forest, ca.
250–400 m. Vouchers: Magogo & Glover 1100, Luke WRQ 8330 (EA).
164
Veronicah M. Ngumbau et al. / PhytoKeys 147: 1–191 (2020)
Canthium glaucum Hiern subsp. glaucum Habit: Shrub. Habitat: Coastal forest,
10–225 m. Vouchers: Ngumbau V & Mwadime N V0463 (EA, HIB), Gisau SOK
22, RM Graham in FD 2352, Langridge 30, Luke WRQ 3159, Graham RM
1327 (EA).
Canthium mrimaense (Verdc.) Lantz Habit: Shrub. Habitat: Evergreen forest, moist
semi-deciduous forest, 10–200 m. Vouchers: Ngumbau V & Mwadime N V0500
(EA, HIB), Robertson 7, Luke 5897, Robertson et al. in MDE 14, Robertson &
Luke 5868, Festo L & Luke Q 2599, Luke WRQ & PA 4386 (EA): Endemic.
Catunaregam nilotica (Stapf ) Tirveng. Habit: Shrub. Habitat: Riverine, thicket edges, woodland, coastal bushland, and scattered tree grassland, ca. 300 m. Vouchers:
Greenway 9660, Spjut 3953, J Adamson 287, Robertson SA 3530 (EA).
Catunaregam spinosa (Thunb.) Tirveng. Habit: Shrub. Habitat: Lowland evergreen
forest, 70–290 m. Vouchers: SAJIT–006018, Ngumbau V & Mwadime N V0176
(EA, HIB), Verdcourt 1859, Faden 70/245, Gillett & Kibuwa 19930, Festo L,
Luke Q & P 2756 (EA).
Chassalia umbraticola Vatke subsp. umbraticola Habit: Shrub. Habitat: Moist
forest, 0–450 (–800) m. Vouchers: SAJIT–004655 & 005563, Ngumbau V &
Mwadime N V077 & V0210 (EA, HIB), Luke Q 1518, Luke WRQ 8321, Napper 1377, Verdcourt 3921 (EA).
Chassalia umbraticola subsp. geophila Verdc. Habit: Shrub. Habitat: Moist forest,
ca. 250 m. Voucher: Mrs SA Robertson & Luke Q 4511 (EA).
Cladoceras subcapitatum (K. Schum. & K. Krause) Bremek. Habit: Shrub. Habitat: Evergreen forest and bushland, ca. 0–200 m. Vouchers: Joanna in CM 5952,
Luke WRQ & Robertson SA 2648 (EA).
Coffea pseudozanguebariae Bridson Habit: Shrub. Habitat: Forest and coastal bushland, 0–800 m. Vouchers: Ngumbau V & Mwadime N V0495 (EA, HIB), Kibuwa 1225, Magogo & Glover 421, Adams 105, Goodrich J 17167, Luke WRQ
3479 (EA): Near Threatened.
Coffea rhamnifolia (Chiov.) Bridson Habit: Shrub. Habitat: Acacia-Commiphora
bushland, ca. 95 m. Vouchers: Makin in EAH 13041, Gillett 16505 (EA).
Coffea sessiliflora Bridson subsp. sessiliflora Habit: Shrub. Habitat: Forest, 0–450 m.
Vouchers: Magogo & Glover 209, Dale in FD 3817, Rawlins 216, Festo L, Luke
Q & P 2669, Luke WRQ 3478, Katende AB 1769 (EA).
Coptosperma graveolens (S. Moore) Degreef var. graveolens Habit: Shrub. Habitat:
Forest, ca. 40 m. Vouchers: Luke Q 1417, Ross KS 126 (EA).
Coptosperma littorale (Hiern) Degreef Habit: Shrub. Habitat: Coastal bushland or
foreshore, 0–105 m. Voucher: RM Graham in FD 2209 (EA).
Coptosperma supra-axillare (Hemsl.) Degreef Habit: Shrub. Habitat: Forest and
also inner borders of mangrove swamps, 0–400 m. Vouchers: Ngumbau V &
Mwadime N V094 & V0153 (EA, HIB), Magogo & Glover 590/B, RM Graham
in FD 2129, Rawlins 855, Festo L, Luke Q & P 2743, Robertson SA & Luke
WRQ 6008 (EA).
Checklist of coastal forests of Kenya
165
Cordylostigma longifolium (Klotzsch) Groeninckx & Dessein Habit: Herb. Habitat: Deciduous woodland, ca. 15–400 m. Vouchers: Polhill & Paulo 603, Magogo
& Glover 826, Whyte (EA).
Cordylostigma obtusilobum (Hiern) Groeninckx & Dessein Habit: Herb. Habitat:
Coarse grassland, 0–270 m. Vouchers: Napier in CM 6291, Hildebrandt 1968,
Jeffery 250, Magogo FC & Glover PE 995 (EA).
Cordylostigma prolixipes (S. Moore) Groeninckx & Dessein Habit: Herb. Habitat:
Rocky places, bushland, and grassland, ca. 120 m. Voucher: RM Graham in FD
1720 (EA).
Cordylostigma virgatum (Willd.) Groeninckx & Dessein Habit: Herb. Habitat:
Coastal bushland, deciduous woodland, and rice fields, 0–600 m. Vouchers: Verdcourt 3286, Rawlins 614 & 650, Magogo FC & Glover PE 473 (EA).
Cremaspora triflora subsp. confluens (K. Schum.) Verdc. Habit: Shrub. Habitat: Evergreen forest, fringing forest, and bush thicket, 0–700 m. Vouchers: SAJIT–006087, Ngumbau V & Mwadime N V074 (EA, HIB), Dale in FD 3559,
RM Graham in FD 2072, Rawlins 280, Mutanga JG, Kamau 7, Magogo FC &
Glover PE 76 (EA).
Crossopteryx febrifuga (Afzel. ex G. Don) Benth. Habit: Tree. Habitat: Deciduous woodland and wooded grassland, ca. 0–400 m. Vouchers: SAJIT–005482
& 005973, Ngumbau V & Mwadime N V0138 (EA, HIB), Drummond &
Hemsley 1209, RM Graham in FD 1696, Donald in FD 2372, Luke WRQ
8323 (EA).
Didymosalpinx norae (Swynn.) Keay Habit: Shrub. Habitat: Evergreen forest, secondary forest, forest edges, 190–810 m. Vouchers: SAJIT–006031, Ngumbau V
& Mwadime N V0310 (EA, HIB), Verdcourt 1854, Magogo & Glover 225 &
857, Luke WRQ 878 (EA).
Diodella sarmentosa (Sw.) Bacigalupo & Cabral ex Borhidi Habit: Herb. Habitat:
Evergreen forest, 0–280 m. Vouchers: SAJIT–005989, Ngumbau V & Mwadime
N V0260 (EA, HIB), Drummond RB & Hemsley JH 1205 (EA).
Diodia aulacosperma K. Schum. var. aulacosperma Habit: Herb. Habitat: Grassland and bushland, 0–30 m. Vouchers: Davis 91, Tweedie 3163, Verdcourt 2115,
Luke Q 5647 (EA).
Diodia aulacosperma var. angustata Verdc. Habit: Herb. Habitat: Grassland and
bushland, 0–5 m. Vouchers: Napier 3321 in CM 6276, Rawlins 779, Gillespie 28,
Festo L, Luke Q & P 2779 (EA).
Empogona ovalifolia (Hiern) Tosh & Robbr. var. ovalifolia Habit: Shrub. Habitat:
Coastal evergreen or mixed formations, 0–80(–150) m. Vouchers: SAJIT–005572,
Ngumbau V & Mwadime N V0136 (EA, HIB), Gillett 21047, Ng’weno 10, Moggridge 333, Luke Q 1432 (EA).
Empogona ovalifolia var. glabrata (Oliv.) Tosh & Robbr. Habit: Shrub. Habitat:
Dry thickets, wooded grassland, and evergreen forest, ca. 400 m. Vouchers: Brenan, Gillett, Kanuri & Chomba 14620, Tweedie 3192 (EA).
166
Veronicah M. Ngumbau et al. / PhytoKeys 147: 1–191 (2020)
Empogona ovalifolia var. taylorii (S. Moore) Tosh & Robbr. Habit: Shrub. Habitat:
Thicket, dry forest, and bushland, ca. 0–300 m. Vouchers: RM Graham in FD
2172, WE Taylor, Robertson SA, Luke WRQ & Awimbo J 5237 (EA).
Eumachia abrupta (Hiern) Delprete & J.H. Kirkbr. var. abrupta Habit: Shrub.
Habitat: Evergreen forest, Combretum, Acacia and Brachystegia woodland, ca.
30–200 m. Vouchers: SAJIT–005565, Ngumbau V & Mwadime N V0351 (EA,
HIB), Magogo & Glover 375, Kassner 393, Bally 8850, Graham MD 2339 (EA).
Eumachia abrupta var. parvifolia (Verdc.) C.M. Taylor Habit: Shrub. Habitat: Presumably evergreen forest, ca. 50 m. Vouchers: Wakefield, Swynnerton 257 (K29),
RM Graham in FD 2339 (EA).
Feretia apodanthera subsp. keniensis Bridson Habit: Shrub. Habitat: Coastal forest
and bushland, 0–30 m. Vouchers: Gillett 18651, Bally 12195, MacNaughtan 9 (EA).
Galiniera saxifraga (Hochst.) Bridson Habit: Tree. Habitat: Forest, ca. 2 m. Voucher: SAJIT–006239 (EA, HIB).
Gardenia fiorii Chiov. Habit: Shrub. Habitat: Commiphora-Acacia open scrub, 60–
750 m. Vouchers: Gillett 16501, Makin in EAH 13049 (EA).
Gardenia posoquerioides S. Moore Habit: Shrub. Habitat: Evergreen forest, Brachystegia woodland, ca. 250–400 m. Vouchers: SAJIT–006124 (EA, HIB), Drummond & Hemsley 3875, Magogo & Glover 939, van Someren 313, Luke WRQ
1649 (EA).
Gardenia ternifolia subsp. jovis-tonantis (Welw.) Verdc. Habit: Tree. Habitat:
Grassland, bushland, Brachystegia and Acacia woodland, 0–2100 m. Voucher: Bally & Smith 14346 (EA).
Gardenia transvenulosa Verdc. Habit: Shrub. Habitat: Dry lowland evergreen forest,
woodland, and bushland, 10–450 (–700) m. Vouchers: Greenway 9809, RB & AJ
Faden 71/748, Moggridge 135, Musyoki BM & Hansen OJ 997 (EA).
Gardenia volkensii K. Schum. var. volkensii Habit: Tree. Habitat: Thicket and dry
woodland, ca. 0–160 m. Vouchers: Polhill & Paulo 642, Graham RM 1584 (EA).
Geophila obvallata subsp. ioides (K. Schum.) Verdc. Habit: Herb. Habitat: Coastal
evergreen forest and lowland rainforest, ca. 80–350 m. Vouchers: Drummond &
Hemsley 3892, Verdcourt 3920, Magogo & Glover 1018, Luke WRQ 1631 (EA).
Geophila repens (L.) I.M. Johnst. Habit: Herb. Habitat: Evergreen forest floors, 80–
300 m. Vouchers: SAJIT–006148 (EA, HIB), Verdcourt 3939A, Drummond &
Hemsley 3801, Napper 1379, Magogo FC & Glover PE 806 (EA).
Guettarda speciosa L. Habit: Tree. Habitat: On sand and coral, sea level. Vouchers:
Bally 8906, Napier in CM 3268, Verdcourt 1060 (EA).
Heinsenia diervilleoides K. Schum. subsp. diervilleoides Habit: Tree. Habitat: Rainforest and moist evergreen forest, ca. 375 m. Vouchers: SAJIT–006069, Ngumbau
V & Mwadime N V056 (EA, HIB), Magogo & Glover 309, Luke WRQ & Robertson SA 2734 (EA).
Heinsia crinita subsp. parviflora (K. Schum. & K. Krause) Verdc. Habit: Shrub.
Habitat: Bushland and degraded coastal Brachystegia, 15–660 m. Vouchers: SAJIT–006454, Ngumbau V & Mwadime N V0165 (EA, HIB), Magogo & Glover
865, Hildebrandt 1983, Polhill & Paulo 840 (EA).
Checklist of coastal forests of Kenya
167
Heinsia zanzibarica (Bojer) Verdc. Habit: Shrub. Habitat: Forest edges, 90–480 (–700)
m. Vouchers: Ngumbau V & Mwadime N V064 & 0177 (EA, HIB), Drummond
& Hemsley 1110, Battiscombe 72, Magogo & Glover 1065, Luke WRQ 8322 (EA).
Hymenodictyon parvifolium Oliv. Habit: Tree. Habitat: Mixed bushland and thicket,
ca. 0–200 m. Vouchers: Drummond & Hemsley 4230, Robertson SA 4234 (EA).
Ixora narcissodora K. Schum. Habit: Shrub. Habitat: Riverine forest, thickets, and
seashore, ca. 240–400 m. Vouchers: SAJIT–006040, Ngumbau V & Mwadime N
V0313 & 014 (EA, HIB), Drummond & Hemsley 1153, Magogo & Glover 78,
Faden 72/63, Luke WRQ 8324 (EA).
Keetia gueinzii (Sond.) Bridson Habit: Liana. Habitat: Forest, woodland, often on
swampy ground, ca. 90–300 m. Vouchers: Ngumbau V & Mwadime N V026
(EA, HIB), Drummond & Hemsley 1138, Magogo FC & Glover PE 571 (EA).
Keetia lukei Bridson Habit: Shrub. Habitat: Mixed forest, 40–400 m. Vouchers:
Luke 3080, 3397 & 3302, Luke & Robertson 2634 (EA).
Keetia venosa (Oliv.) Bridson Habit: Liana. Habitat: Forest edges and scrub, ca.
275 m. Vouchers: SAJIT–005500, 005506 & 006035, Ngumbau V & Mwadime
N V0174 (EA, HIB), Spjut 4582, Gillet 18704, Gardner 1435, Magogo FC &
Glover PE 66 (EA).
Keetia zanzibarica (Klotzsch) Bridson subsp. zanzibarica Habit: Shrub. Habitat:
Coastal bushland, thickets, and forest edges, 0–500 m. Vouchers: SAJIT–005465,
Ngumbau V & Mwadime N V081 (EA, HIB), Bally & Smith 14351, Jeffery
K104, Hooper & Townsend 1210, Luke WRQ & Robertson SA 223 (EA).
Kohautia caespitosa subsp. amaniensis (K. Krause) Govaerts Habit: Herb. Habitat:
Bushland, ca. 420 m. Vouchers: Magogo FC & Glover PE 666, GM Mungai &
SM Rucina 223/84 (EA).
Kraussia kirkii (Hook.f.) Bullock Habit: Shrub. Habitat: Forest and coastal bushland, 0–480 m. Vouchers: Dale in FD 3668, J Adamson 309 in Bally 5815, Power,
Festo L, Luke Q & P 2680 (EA).
Kraussia speciosa Bullock Habit: Shrub. Habitat: Forest, ca. 10–600 m. Vouchers:
Magogo & Glover 435, Greenway & Rawlins 9348, Rawlins 326, Robertson SA
& Luke WRQ 5532 (EA).
Lamprothamnus zanguebaricus Hiern Habit: Shrub. Habitat: Wooded grassland,
coastal bushland, thicket, and woodland, 0–300 m. Vouchers: Ngumbau V &
Mwadime N V0451 (EA, HIB), Polhill & Paulo 908, Jex-Blake in Bally 5704,
Polhill & Paulo 695, Kokwaro JO 3984 (EA).
Leptactina platyphylla (Hiern) Wernham Habit: Shrub. Habitat: Evergreen forest, woodland, and secondary bushland, 45–400 m. Vouchers: Ngumbau V &
Mwadime N V0146 (EA, HIB), Robertson SA 4671 (EA).
Meyna tetraphylla subsp. comorensis (Robyns) Verdc. Habit: Shrub. Habitat:
Coastal bush on coral, dry or moist evergreen forest, 0–200 m. Voucher: Gillett &
Kibuwa 19843 (EA).
Mitracarpus hirtus (L.) DC. Habit: Herb. Habitat: Waste places, roadsides, ca.
418 m. Vouchers: Ngumbau V & Mwadime N V0126 (EA, HIB), Luke WRQ &
PA 5692 (EA).
168
Veronicah M. Ngumbau et al. / PhytoKeys 147: 1–191 (2020)
Mitriostigma greenwayi Bridson Habit: Shrub. Habitat: Coastal forest, on coral rock,
0–150 m. Vouchers: SAJIT–005581, Ngumbau V & Mwadime N V0473 (EA, HIB),
Brenan, Gillett et al. 14669, Hawthorne 247, Faden et al. 77/531 (EA): Endemic.
Multidentia sclerocarpa (K. Schum.) Bridson Habit: Tree. Habitat: Forest, 100–
550 m. Vouchers: SAJIT–006058, Ngumbau V & Mwadime N V0367 (EA,
HIB), Luke & Robertson 530, Luke PA & WRQ 4327 (EA): Endangered.
Mussaenda monticola K. Krause var. monticola Habit: Shrub. Habitat: Evergreen
forest, 45–820 m. Vouchers: SAJIT–006080 (EA, HIB), Magogo & Glover 1040,
Spjut & Ensor 2739, Faden 74/1264, Luke WRQ & Robertson SA 1915, Luke
WRQ et al. 4723 (EA).
Nichallea soyauxii (Hiern) Bridson Habit: Tree. Habitat: Forest edges or coastal
bushland, 0–350 m. Vouchers: Magogo & Glover 799, Gillett & Kibuwa 19857,
Dale in FD 3670, Omondi W & Obunyali C 308 (EA).
Oldenlandia affinis subsp. fugax (Vatke) Verdc. Habit: Herb. Habitat: Grassland,
swampy ground, bushland, and evergreen forest margins, ca. 0–240 m. Vouchers:
Ngumbau V & Mwadime N V049 (EA, HIB), Drummond & Hemsley 1076,
GM Jeffery 30, Gillespie 364, Magogo FC & Glover PE 1030 (EA).
Oldenlandia corymbosa L. var. corymbosa Habit: Herb. Habitat: Grassland and
bushland, ca. 228 m. Voucher: Ngumbau V & Mwadime N V0120 (EA, HIB).
Oldenlandia cryptocarpa Chiov. Habit: Herb. Habitat: Coastal bushland, ca. 15 m.
Voucher: Polhill & Paulo 601 (EA).
Oldenlandia fastigiata Bremek. var. fastigiata Habit: Herb. Habitat: Grassland,
thickets, and open Acacia woodland, ca. 0–200 m. Vouchers: Verdcourt 1870,
Luke WRQ & PA 5723 (EA).
Oldenlandia fastigiata var. pseudopentodon Verdc. Habit: Herb. Habitat: Open
damp grassland, ca. 15 m. Voucher: Gillett & Kibuwa 19900 (EA).
Oldenlandia fastigiata var. somala (Chiov. ex Bremek.) Verdc. Habit: Herb. Habitat:
Grassland areas in Acacia bushland, ca. 60 m. Vouchers: Thairu 77, Thomas 23 (EA).
Oldenlandia goreensis var. trichocaula Bremek. Habit: Herb. Habitat: Edges of
swamps, 0–375 m. Vouchers: Ngumbau V & Mwadime N V0550 (EA, HIB),
Drummond & Hemsley 1162, Magogo & Glover 378, Luke Q 5624 (EA).
Oldenlandia herbacea (L.) Roxb. var. herbacea Habit: Herb. Habitat: Grassland,
thicket, and bushland, ca. 0–300 m. Vouchers: Polhill & Paulo 485, Thulin M
301(EA).
Oldenlandia ichthyoderma Cufod. Habit: Herb. Habitat: Acacia-Commiphora open
bushland, 100 m. Voucher: Gillett 16518 (EA).
Oldenlandia johnstonii (Oliv.) K. Schum. ex Engl. Habit: Herb. Habitat: Dry evergreen forest and wooded grassland, ca. 350 m. Voucher: Luke WRQ & Robertson
SA 2702 (EA).
Oldenlandia lancifolia var. scabridula Bremek. Habit: Herb. Habitat: In moist or
aquatic habitats, ca. 24 m. Vouchers: Ngumbau V & Mwadime N V0547 (EA,
HIB), Gerhardt K & Steiner M 162, Luke WRQ & Pakia M 7459 (EA).
Checklist of coastal forests of Kenya
169
Oldenlandia richardsonioides (K. Schum.) Verdc. var. richardsonioides Habit:
Herb. Habitat: On consolidated dunes, ca. 6 m. Vouchers: Gillespie 254, Luke Q
6127, Greenway & Rawlins 9289, Rawlins 107 (EA).
Oldenlandia rosulata var. littoralis Verdc. Habit: Herb. Habitat: Forest, ca. 50 m.
Voucher: Symes 133 (EA).
Oxyanthus goetzei subsp. keniensis Bridson Habit: Shrub. Habitat: Bushland, forest,
ca. 15 m. Vouchers: SAJIT–005542 (EA, HIB), JM Reitsma 200, RB & AJ Faden
77/662, Robertson 4882, Gray 106, Luke 5272 (EA).
Oxyanthus pyriformis subsp. longitubus Bridson Habit: Tree. Habitat: Forest, 240–
450 m. Vouchers: SAJIT–006059 & 006151 (EA, HIB), Drummond & Hemsley
3961, Magogo & Glover 811, Magogo & Glover 812, Herbarium Techniques
Course II–099 (EA).
Oxyanthus zanguebaricus (Hiern) Bridson Habit: Shrub. Habitat: Forest, 60–
240 m. Vouchers: Ngumbau V & Mwadime N V0214 (EA, HIB), Gisau SOK 25,
Rawlins 225 & 259, Luke WRQ et al. 3329 (EA).
Paracephaelis trichantha (Baker) De Block Habit: Shrub. Habitat: In coastal bushland, besides rivers, ca. 20 m. Vouchers: Luke Q 1436, Faden 74/1104, Marquis P,
Festo L, Luke Q & P 2787 (EA).
Pavetta crebrifolia Hiern var. crebrifolia Habit: Shrub. Habitat: Coastal scrub or
forest, ca. 0–304 m. Vouchers: Verdcourt 1904, Polhill & Paulo 801, Festo L, Luke
Q & P 2655, Spjut RW 4584 (EA).
Pavetta crebrifolia var. pubescens Bridson Habit: Shrub. Habitat: Forest edge, ca.
30 m. Vouchers: Gillespie 327, Robertson SA & Luke WRQ 5606 (EA).
Pavetta linearifolia Bremek. Habit: Shrub. Habitat: Coastal bushland or besides rivers, 50–100 m. Vouchers: Polhill & Paulo 821, Leroy 1039, Adamson 308 in Bally
5999, Luke, WRQ & Robertson SA 2508 (EA): Vulnerable.
Pavetta sansibarica subsp. trichosphaera (Bremek.) Bridson Habit: Shrub. Habitat: Bushland and forest, ca. 380 m. Vouchers: Luke WRQ et al. 4720, Luke WRQ
8326 & 4315 (EA).
Pavetta sepium var. merkeri (K. Krause) Bridson Habit: Shrub. Habitat: Forest, ca.
183 m. Vouchers: Ngumbau V & Mwadime N V0203 (EA, HIB), Hawthorne
186, Luke 5280 (EA).
Pavetta sphaerobotrys subsp. tanaica (Bremek.) Bridson Habit: Shrub. Habitat: Riverine forest, 30–350 m. Vouchers: Hamewood 8, Kibuwa 2500, Thairu 45 (EA).
Pavetta stenosepala K. Schum. subsp. stenosepala Habit: Shrub. Habitat: Evergreen
forest, thickets, and bushland, 0–340 m. Vouchers: SAJIT–005978 & 005502,
Ngumbau V & Mwadime N V105 (EA, HIB), Drummond & Hemsley 1207,
RM Graham in FD 2149, Rawlins 425, Luke WRQ & Robertson SA 517, Luke
WRQ & Robertson SA 225 (EA).
Pavetta tarennoides S. Moore Habit: Shrub. Habitat: Undergrowth of coastal forest,
300–440 m. Vouchers: Brown 736, Magogo & Glover 40, Christensen 188, Luke
WRQ & Robertson SA 2719 (EA): Vulnerable, Endemic.
170
Veronicah M. Ngumbau et al. / PhytoKeys 147: 1–191 (2020)
Pavetta uniflora Bremek. Habit: Shrub. Habitat: Coastal woodland or coastal bushland, 0–150 m. Vouchers: Bally 16739, RM Graham in FD 2136, Polhill & Paulo
859, Graham RM 1856 (EA).
Pentas zanzibarica (Klotzsch) Vatke Habit: Herb. Habitat: Open grassland and forest edges, 0–600 m. Vouchers: Ngumbau V & Mwadime N V0198 (EA, HIB),
Verdcourt 1867, Luke WRQ 8319 (EA).
Pentodon pentandrus var. minor Bremek. Habit: Herb. Habitat: Swamp lakes and
river margins, ca. 0–50 m. Vouchers: SAJIT–005522 & 006226 (EA, HIB), Verdcourt 1887, Polhill & Paulo 890, Polhill & Paulo 538, Kirika P, Muthoka P &
Mbale M 746, Luke WRQ & PA 5732 (EA).
Polysphaeria lanceolata Hiern subsp. lanceolata Habit: Shrub. Habitat: Riverine
forest, ca. 15 m. Voucher: Faden et al. 77/485 (EA).
Polysphaeria multiflora Hiern subsp. multiflora Habit: Shrub. Habitat: Margins of
shrubby thickets, mangrove swamps, forest, and woodland, ca. 0–200 m. Vouchers: Drummond & Hemsley 3913, Kassner 362, Bally 13051(EA).
Polysphaeria multiflora subsp. pubescens Verdc. Habit: Shrub. Habitat: Riverine
forest and bushland, ca. 60 m. Vouchers: Homewood 32, Makin 14 (EA).
Polysphaeria parvifolia Hiern Habit: Shrub. Habitat: Dry evergreen forest, woodland, coastal bushland, and scrub, 0–500 m. Vouchers: SAJIT–005540, 005495
& 005446 (EA, HIB), Drummond & Hemsley 1109, Wakefield, RM Graham in
FD 1522, Robertson SA 4229 (EA).
Psychotria amboniana K. Schum. var. amboniana Habit: Shrub. Habitat: Coastal
forest, thickets, scrub, and grassland with scattered trees, 0–300 m. Vouchers: SAJIT–006043, Ngumbau V & Mwadime N V0132 (EA, HIB), Lucas, Jeffery & Kirrika
243, Wakefield, Linder 2660, Rawlins 426, Robertson SA 4231, Luke Q 1451 (EA).
Psychotria amboniana var. velutina (E.M.A. Petit) Verdc. Habit: Shrub. Habitat:
Coastal bushland, 0–160 m. Vouchers: RM Graham in FD 2337, Gillett & Kibuwa 19928, Rawlins 427 (EA).
Psychotria capensis subsp. riparia (K. Schum. & K. Krause) Verdc. Habit: Shrub.
Habitat: Evergreen bushland, coastal bushland, and wooded grassland, ca. 20 m.
Vouchers: Ngumbau V & Mwadime N V0309 (EA, HIB), Mwadime N 39, RM
Graham in FD 2185 (EA).
Psychotria capensis var. puberula (E.M.A. Petit) Verdc. Habit: Shrub. Habitat:
Coastal forest and thicket, ca. 80 m. Vouchers: Drummond & Hemsley 3951,
Magogo & Glover 324 (EA).
Psychotria crassipetala E.M.A. Petit Habit: Shrub. Habitat: Evergreen forest, ca.
30 m. Voucher: Dale in FD 3831 (EA).
Psychotria faucicola K. Schum. Habit: Shrub. Habitat: Rainforest and forest edges,
15–750 m. Vouchers: SAJIT–006188, Ngumbau V & Mwadime N V0353 (EA,
HIB), Napper 1375, Faden 70/242, Magogo & Glover 972, Faden RB 677A,
Luke WRQ 868 (EA).
Psychotria holtzii (K. Schum.) E.M.A. Petit var. holtzii Habit: Shrub. Habitat: Evergreen forest, 0–360 (450) m. Vouchers: Verdcourt 1884, Drummond & Hemsley
3895 & 1190, Verdcourt 3915 (EA).
Checklist of coastal forests of Kenya
171
Psychotria lauracea (K. Schum.) E.M.A. Petit Habit: Shrub. Habitat: Various types
of evergreen forest, 0–268 m. Vouchers: SAJIT–005508 & 005983, Ngumbau V
& Mwadime N V038 (EA, HIB), Drummond & Hemsley 1180, Luke WRQ &
Robertson SA 245 (EA).
Psychotria leucopoda E.M.A. Petit Habit: Shrub. Habitat: Rainforest, lowland fringing forest, and coastal evergreen forest, 45–300 m. Vouchers: SAJIT–006010,
Ngumbau V & Mwadime N V0166 (EA, HIB), VG van Someren 7180, CF Elliot
202, Wakefield, Luke PA & WRQ 6295, Luke WRQ 897 (EA).
Psychotria punctata Vatke Habit: Shrub. Habitat: Coastal bushland, savannah, and
forest, mostly on sand or coral rag, 0–45 (–280) m. Vouchers: SAJIT–005576,
006233, 005600 & 006238 (EA, HIB), Napier 3252 in CM 6282, Tweedie 1045,
Greenway & Rawlins 8907 (EA).
Psychotria schliebenii var. parvipaniculata E.M.A. Petit Habit: Shrub. Habitat: Evergreen forest, ca. 60 m. Voucher: Dale in FD 3570 (EA).
Psychotria schliebenii var. sessilipaniculata E.M.A. Petit Habit: Shrub. Habitat:
Evergreen forest, ca. 0–442 m Vouchers: RM Graham in FD 2031, Magogo &
Glover 14, Magogo & Glover 259 (EA).
Psychotria tanganyicensis Verdc. subsp. tanganyicensis Habit: Shrub. Habitat: Rainforest, riverine forest, 300–1530 m. Vouchers: SAJIT–006160 (EA, HIB), Luke
WRQ 1847, Drummond & Hemsley 1194, Magogo & Glover 440, Moomaw
1140 (EA).
Psydrax faulknerae Bridson Habit: Tree. Habitat: Coastal bush, thicket or Brachystegia woodland, 0–725 m. Vouchers: Ngumbau V & Mwadime N V0267 & V093
(EA, HIB), Magogo & Glover 726, RM Graham in FD 1823, Moggridge 332,
Luke WRQ & Robertson SA 1543 (EA).
Psydrax kaessneri (S. Moore) Bridson Habit: Shrub. Habitat: Riverine thickets and
forest edges, (20–) 90–300 m. Vouchers: Gilllett & Kibuwa 19927, Medley 414,
Luke & Robertson 1532, Festo L, Luke Q & P 2672 (EA).
Psydrax polhillii Bridson Habit: Shrub. Habitat: Thickets and Brachystegia woodland,
15–350 m. Vouchers: Drummond & Hemsley 4163, Faden & Evans 71/708,
Luke WRQ 9466 (EA).
Psydrax recurvifolia (Bullock) Bridson Habit: Shrub. Habitat: Forest or at swamp
edge, 0–10 m. Vouchers: RM Graham in FD 1751, Moggridge 534, SA Robertson
4652, Robertson SA & Luke WRQ 5993 (EA).
Psydrax robertsoniae Bridson Habit: Shrub. Habitat: Thicket and forest, ca. 5 m.
Vouchers: Moggridge 334, Hawthorne 273, SA Robertson 6152, Luke WRQ
3142 (EA): Endemic.
Psydrax schimperiana (A. Rich.) Bridson Habit: Tree. Habitat: Forest, thicket, and
bushland, ca. 15 m. Vouchers: RM Graham in FD 1995, Moomaw JC 1077 (EA).
Psydrax sp. A of FTEA Habit: Shrub. Habitat: Sand dune thicket. Voucher: Luke &
Robertson 1368 (EA): Endemic.
Pyrostria bibracteata (Baker) Cavaco Habit: Tree. Habitat: Bushland or forest edges,
0–400 m. Vouchers: Ngumbau V & Mwadime N V0133 (EA, HIB), Magogo &
Glover 294, Perdue & Kibuwa 10022, Gillett 20346, Festo L Luke Q 2523 (EA).
172
Veronicah M. Ngumbau et al. / PhytoKeys 147: 1–191 (2020)
Pyrostria phyllanthoidea (Baill.) Bridson Habit: Shrub. Habitat: Acacia-Commiphora
bushland, ca. 20 m. Vouchers: Drummond & Hemsley 4170, Luke Q 1435 (EA).
Rhodopentas bussei (K. Krause) Kårehed & B. Bremer Habit: Shrub. Habitat:
Grassland, bushland, and woodland dry evergreen forest, ca. 0–400 m. Vouchers:
Ngumbau V & Mwadime N V0217 & V088 (EA, HIB), Drummond & Hemsley
3813, Graham in FD 2065, Napier 3361 in CM 6286, Luke Q 1483 (EA).
Rhodopentas parvifolia (Hiern) Kårehed & B. Bremer Habit: Shrub. Habitat: Dry
bushland, 0–300 m. Vouchers: Robertson & Luke 1778, V de Meester-Manger
Cats 243, Luke WRQ et al. 5418 (EA).
Richardia scabra L. Habit: Herb. Habitat: Grassland, ca. 36 m. Voucher: SAJIT–006169 (EA, HIB).
Rothmannia macrosiphon (K. Schum. ex Engl.) Bridson Habit: Tree. Habitat: Forest, 60–450 m. Vouchers: SAJIT–005928, Ngumbau V & Mwadime N V069
(EA, HIB), Drummond & Hemsley 3938 & 3875, RM Graham in FD1984,
Luke WRQ & Robertson SA 312 & 235 (EA): Vulnerable.
Rothmannia ravae (Chiov.) Bridson Habit: Tree. Habitat: In thicket or sometimes
forest, 45–600 m. Vouchers: Dale K 206A, Gillespie 205, Festo L, Luke Q & P
2774, Luke WRQ 2932 (EA).
Rutidea fuscescens Hiern Habit: Liana. Habitat: Evergreen forest, ca. 370 m. Voucher: Luke WRQ 895 (EA).
Rytigynia celastroides (Baill.) Verdc. var. celastroides Habit: Shrub. Habitat: Woodland, open forest, relict thickets, and lowland rainforest, 0–750 m. Vouchers: SAJIT–005467, 005468 & 005469, Ngumbau V & Mwadime N V0134 (EA, HIB),
Verdcourt 1902, RM Graham BB383 in FD 1750, RM Graham N824 in FD
2336, Luke WRQ & Robertson SA 2795 (EA).
Rytigynia decussata (K. Schum.) Robyns Habit: Shrub. Habitat: Acacia-Combretum
and Brachystegia woodlands, thickets, and open grassland, ca. 75–300 m. Vouchers:
Ngumbau V & Mwadime N V0131 (EA, HIB), Robertson & Luke 5183 (EA).
Rytigynia parvifolia Verdc. Habit: Shrub. Habitat: Bushland and scattered tree grassland, 30–180 m. Vouchers: Rawlins 726, Greenway 9489, Luke Q 1450 (EA).
Rytigynia sp. I of FTEA Habit: Shrub. Habitat: Coastal riverine forest, ca. 6 m.
Vouchers: Homewood 62 & 64 (EA): Endemic.
Rytigynia sp. L of FTEA Habit: Shrub. Habitat: Moist lowland forest, ca. 30 m.
Voucher: Robertson & Luke 6309 (EA): Endemic.
Spermacoce filituba (K. Schum.) Verdc. Habit: Herb. Habitat: Grassland with scattered trees, sea shore bushland on sand, and coconut groves, 0–600 m. Vouchers: SAJIT–006052 (EA, HIB), RM Graham in AD 1629, Polhill & Paulo 550,
Whyte, Luke Q 5670, Magogo FC & Glover PE 715, Tiede 9 & 11 (EA).
Spermacoce laevis Lam. Habit: Herb. Habitat: Grassland, along roadsides, and in
rice field, ca. 0–418 m. Vouchers: Ngumbau V & Mwadime N V0127 (EA, HIB),
Magogo FC & Glover PE 661 (EA).
Spermacoce pusilla Wall. Habit: Herb. Habitat: Seasonally damp grassland, bushland, ca. 120 m. Voucher: McKeag in EAH 9180 (EA).
Checklist of coastal forests of Kenya
173
Spermacoce sp. B of FTEA Habit: Herb. Habitat: Evergreen forest, ca. 225 m. Voucher: Bally 8546 (EA): Endemic.
Tarenna drummondii Bridson Habit: Tree. Habitat: Forest or open woodland, 100–
460 m. Vouchers: SAJIT–006072 & 006149, Ngumbau V & Mwadime N V0163
(EA, HIB), Drummond & Hemsley 3876, Magogo & Glover 492, Faden et al.
77/411, Luke WRQ & Kahumbu P 4546F (EA): Vulnerable.
Tarenna pavettoides subsp. friesiorum (K. Krause) Bridson Habit: Tree. Habitat:
Forest. Voucher: Cunningham-van Someren GR Sh 245 (EA).
Triainolepis africana subsp. hildebrandtii (Vatke) Verdc. Habit: Shrub. Habitat: Mostly in coastal bushland near high-tide mark, 0 (–150) m. Vouchers:
SAJIT–006107 & 005450 (EA, HIB), Verdcourt 3961, Ossent 242, Tweedie
2377 (EA).
Tricalysia bridsoniana Robbr. var. bridsoniana Habit: Shrub. Habitat: Cynometra
forest, below 300 m. Vouchers: Spjut & Ensor 2789, Brenan, Gillett, Kanuri &
Chomba 14685, Pdwa 26, Brenan JPM 14685, Luke WRQ 9463 (EA): Endemic.
Tricalysia congesta (Oliv.) Hiern Habit: Shrub. Habitat: Sclerophyllous thicket, fireprotected woodland, and riverine forest, ca. 400 m. Voucher: Verdcourt 5289 (EA).
Tricalysia microphylla Hiern Habit: Shrub. Habitat: Forest, ca. 300 m. Vouchers:
Ngumbau V & Mwadime N V0314 (EA, HIB), Faden 77/716, Magogo & Glover 941, Gillett & Gachathi 77/413, Robertson SA & Luke, WRQ 5480, Luke
WRQ 9457 (EA).
Tricalysia pallens Hiern Habit: Shrub. Habitat: Evergreen forests, forest margins,
and thickets, ca. 229 m. Vouchers: SAJIT–005443, Ngumbau V & Mwadime N
V0160 (EA, HIB), Drummond & Hemsley 1197, Luke WRQ 867 (EA).
Uncaria africana G. Don subsp. africana Habit: Liana. Habitat: Secondary forest,
ca. 18 m. Voucher: Luke WRQ & Robertson SA 2752 (EA).
Vangueria infausta subsp. rotundata (Robyns) Verdc. Habit: Tree. Habitat: Grassland with forest patches, ca. 10–200 m. Vouchers: Ngumbau V & Mwadime N
V0200 (EA, HIB), Magogo FC & Glover PE 181, Luke Q 1511 (EA).
Vangueria loranthifolia K. Schum. Habit: Shrub. Habitat: Cynometra forest, thicket,
woodland, and bushland, 0–600 m. Vouchers: SAJIT–005974, Ngumbau V &
Mwadime N V0234 & 0460 (EA, HIB), Napier 3305 in C.M.6281, Spjut & Ensor 2671, Polhill & Paulo 652, Gillett JB 20431, Luke WRQ 9464 (EA).
Vangueria pallidiflora (Bullock) Lantz Habit: Shrub. Habitat: Evergreen forest,
thickets, 0–600 m. Vouchers: SAJIT–006057 (EA, HIB), Drummond & Hemsley
3940, Magogo & Hemsley 441, Spjut 2731, Festo L, Luke Q & P 2657, Luke
WRQ & Kahumbu P 4546D (EA): Vulnerable.
Vangueria randii subsp. acuminata Verdc. Habit: Shrub. Habitat: Evergreen forest,
(0–) 30–300 m. Vouchers: SAJIT–005536 (EA, HIB), Magogo & Glover 437,
Gillett 21053, Gillett & Kibuwa 19895 (EA).
Vangueriopsis shimbaensis A. P. Davis & Q. Luke Habit: Tree. Habitat: Lowland
forest, 360 m. Vouchers: Luke & Lehman 10894, Luke WQR 8316 (EA): Critically Endangered, Endemic.
174
Veronicah M. Ngumbau et al. / PhytoKeys 147: 1–191 (2020)
F151. Rutaceae
5Genera, 17 Species
Clausena anisata (Willd.) Hook.f. ex Benth. Habit: Tree. Habitat: Evergreen and
fringing forest, 0–400 m. Vouchers: SAJIT–006044, Ngumbau V & Mwadime N
V0170 (EA, HIB), Luke WRQ & Robertson SA 533 (EA).
Vepris (Diphasia sp. A of FTEA) Habit: Tree. Habitat: Moist forest, ca. 5 m. Voucher:
SAJIT–006094 (EA, HIB), Luke WRQ 3035 (EA): Endemic.
Harrisonia abyssinica Oliv. Habit: Shrub. Habitat: Dry evergreen forest, forest edges,
and clearings, 35–120 m. Vouchers: SAJIT–005459 & 005458 (EA, HIB), Magogo FC & Glover PE 21, Burstyn P 89 (EA).
Toddalia asiatica (L.) Lam. Habit: Liana. Habitat: Forest edges, bushland, and wooded grassland, ca. 1–400 m. Vouchers: SAJIT–005452, Ngumbau V & Mwadime
N V0237 (EA, HIB), Kokwaro 1983, Magogo FC & Glover PE 257 (EA).
Vepris amaniensis (Engl.) Mziray Habit: Shrub. Habitat: Rainforest or forest-edge
thickets, ca. 360 m. Vouchers: Magogo & Glover 1102, Lavranos J & Newton
12290, Malombe I, Mwadime N & Saidi 1652 (EA).
Vepris eugeniifolia (Engl.) I. Verd. Habit: Shrub. Habitat: Coastal forest and bushland on coral rock, 0–750 m. Vouchers: Ngumbau V & Mwadime N V0299 (EA,
HIB), Drummond & Hemsley 4063, Polhill & Paulo 610, Magogo FC & Glover
PE 693 (EA).
Vepris glomerata var. glabra Kokwaro Habit: Shrub. Habitat: Wooded grassland and
deciduous bushland, ca. 30 m. Voucher: Dale in FD 3853 (EA).
Vepris lanceolata (Lam.) G. Don Habit: Tree. Habitat: Coastal evergreen thicket or
on loose sandy soil along the beaches, 0–30 m. Voucher: Schultka K 149 (EA).
Vepris robertsoniae Q. Luke, ined. Habit: Tree. Habitat: Lowland coastal forest, ca.
35 m. Vouchers: Luke Q 1539, Luke WRQ & Robertson SA 2806 (EA).
Vepris sansibarensis (Engl.) Mziray Habit: Shrub. Habitat: Coastal thicket and forest, 1–760 m. Vouchers: Dale in FD 3661, Moomaw 1674, Rawlins 762, Muchiri
J 440, Luke Q 1560, Luke WRQ 3135 (EA): Vulnerable.
Vepris simplicifolia (Engl.) Mziray Habit: Tree. Habitat: Dry forest, riverine thicket
or woodland, ca. 15 m. Vouchers: Ngumbau V & Mwadime N V0560 (EA, HIB),
Karin Gerhardt & Mariette Steiner 263 (EA).
Vepris sp. nr. stolzii I. Verd. Habit: Tree. Habitat: Lowland and upland forest, ca.
100 m. Voucher: Luke WRQ & PA 4501 (EA).
Vepris trichocarpa (Engl.) Mziray Habit: Tree. Habitat: Coastal and upland forests,
1–2300 m. Vouchers: SAJIT–004657 (EA, HIB), Greenway 10826, Luke WRQ
& Mbinda J 5969 (EA).
Zanthoxylum chalybeum Engl. Habit: Tree. Habitat: Dry bushland and wooded
grassland, ca. 3 m. Vouchers: SAJIT–006447, Ngumbau V & Mwadime N V0158
(EA, HIB), Gilbert MG, Kuchar P 5873 (EA).
Zanthoxylum holtzianum (Engl.) P.G. Waterman Habit: Tree. Habitat: Coastal forest, bushland, sometimes on coral, 1–230 m. Vouchers: Ngumbau V & Mwadime
Checklist of coastal forests of Kenya
175
N V0535 (EA, HIB), Drummond & Hemsley 4006, Dale in FD 3567, Rawlins in
EAH 11279, Robertson SA 3775, Festo L, Luke Q & P 2785 (EA).
Zanthoxylum paracanthum (Mildbr.) J. O. Kokwaro Habit: Tree. Habitat: Evergreen
forest, bushland, and riverine, ca. 140 m. Vouchers: Ngumbau V & Mwadime N
V0534 (EA, HIB), Faden 234, Luke WRQ & Robertson SA 2801 (EA).
F152. Salicaceae
10 Genera, 15 Species
Bivinia jalbertii Tul. Habit: Tree. Habitat: Open forest and coastal bushland, up to
200 m. Vouchers: Ngumbau V & Mwadime N V0246 (EA, HIB), Rawlins SP
321, Luke WRQ & Robertson SA 510 (EA): Near Threatened.
Casearia gladiiformis Mast. Habit: Tree. Habitat: Dry evergreen, riverine, secondary forest, coastal woodland, and bushland, ca. 600 m. Vouchers: Luke WRQ &
Robertson SA 1926, Luke WRQ & Robertson SA 511 (EA).
Dovyalis abyssinica (A. Rich.) Warb. Habit: Shrub. Habitat: Dry evergreen forest,
and open wooded grassland, ca. 15 m. Voucher: Gerhardt K & Steiner M 42 (EA).
Dovyalis hispidula Wild Habit: Shrub. Habitat: Riverine thickets and Brachystegia
woodland, ca. 150 m. Voucher: Rawlins SP 240 (EA).
Dovyalis keniensis E. V. Williams Habit: Shrub. Habitat: In Afzelia-Mimusops forest,
ca. 142 m. Vouchers: SAJIT–006004 (EA, HIB), Trump EC 109, Luke WRQ
3150 (EA).
Dovyalis macrocalyx (Oliv.) Warb. Habit: Tree. Habitat: Rainforest, dry evergreen,
riverine forest, bushland, and wooded grassland, ca. 0–400 m. Vouchers: SAJIT–006004 (EA, HIB), Sangai GW 15650, Luke WRQ 3160 (EA).
Flacourtia indica (Burm.f.) Merr. Habit: Tree. Habitat: Woodland, wooded grassland and bushland, ca. 0–350 m. Vouchers: SAJIT–005606 & 006433 (EA, HIB),
Festo L & Luke Q 2291, Greenway & Rawlins 8877 (EA).
Homalium abdessammadii Asch. & Schweinf. Habit: Tree. Habitat: Forest or forest
edge, ca. 200 m. Vouchers: Luke WRQ & PA 4582, Greenway PJ & Rawlins SP
9427 (EA).
Homalium longistylum Mast. Habit: Tree. Habitat: Forest, forest edge, and riverine
forest, 600–800 m. Voucher: Luke WRQ & Robertson SA 2281 (EA).
Ludia mauritiana J.F. Gmel. Habit: Tree. Habitat: Dry evergreen forest and coastal
bushland, ca. 0–300 m. Vouchers: SAJIT–005587 & 005441 (EA, HIB), Dale IR
3864, Greenway 9627 (EA).
Oncoba spinosa Forssk. Habit: Tree. Habitat: Forest edges, riverine forest, bushland,
and Brachystegia woodland, ca. 300 m. Voucher: Festo L & Luke Q 2535 (EA).
Populus ilicifolia (Engl.) Rouleau Habit: Tree. Habitat: Riverine forest, mudflats,
and sandbanks, ca. 30 m. Voucher: Gillett & Kibuwa 19912/A (EA).
Scolopia rhamniphylla Gilg Habit: Shrub. Habitat: Rainforest or dry evergreen forest, ca. 300 m. Vouchers: Robertson SA 5457, Festo L & Luke Q 2529, Dale in
FD 3866, Verdcourt 1896 (EA).
176
Veronicah M. Ngumbau et al. / PhytoKeys 147: 1–191 (2020)
Scolopia zeyheri (Nees) Harv. Habit: Tree. Habitat: Dry evergreen forest, riverine
forest, and bushland, ca. 0–20 m. Vouchers: Luke WRQ & Robertson SA 1035,
Luke 10769 (EA).
Trimeria grandifolia (Hochst.) Warb. subsp. tropica (Burkill) Sleumer Habit: Tree.
Habitat: Dry evergreen or riverine forest, ca. 297 m. Vouchers: SAJIT–005494 &
006006, Ngumbau V & Mwadime N V0142 (EA, HIB), Luke WRQ & Robertson SA 507 (EA).
F153. Salvadoraceae
3 Genera, 4 Species
Azima tetracantha Lam. Habit: Shrub. Habitat: Scrub, seasonal rivers, and coastal
bushland, ca. 30 m. Voucher: Verdcourt 1179 (EA).
Dobera glabra (Forssk.) Poir. Habit: Tree. Habitat: Dry forest, ca. 30–40 m. Vouchers: Kimberly Medley 386, Robertson SA 7734 (EA).
Dobera loranthifolia (Warb.) Harms Habit: Tree. Habitat: In scrub and wooded
grassland, 0–810 m. Voucher: Luke WRQ & PA sr.
Salvadora persica L. var. persica Habit: Shrub. Habitat: On saline, sandy or loamy
soils, along large seasonal rivers, ca. 0–150 m. Vouchers: SAJIT–005906 & 006451
(EA, HIB), Jeffery 476 (EA).
Salvadora persica var. cyclophylla (Chiov.) Cuf. Habit: Shrub. Habitat: In scrub,
wooded grassland, just above high tide level, ca. 0–50 m. Vouchers: Unknown collector in CM 242, Verdcourt 3599, Birch 62/170B, Gillespie 65, Boll K 16031(EA).
F154. Santalaceae
2 Genera, 2 Species
Thesium subaphyllum Engl. Habit: Herb. Habitat: Grassland and on rocks, ca. 60–
380 m. Vouchers: Faden, Evans & Mahasi 70/821, Luke WRQ 3312 (EA).
Viscum triflorum DC. Habit: Herb. Habitat: Evergreen forest, 0–150 m. Voucher:
Wiens 4538 (EA).
F155. Sapindaceae
20 Genera, 25 Species
Allophylus chirindensis Baker f. Habit: Tree. Habitat: Evergreen forest, 350–850 m.
Vouchers: Magogo & Glover 1098, Luke WRQ et al. 4715 (EA).
Allophylus pervillei Blume f. pervillei Habit: Shrub. Habitat: Fringing forest, forest clumps, ground water forest, and coastal Brachystegia, 0–550 m. Vouchers:
SAJIT–006193 & 006144 (EA, HIB), SA Robertson 3502, RM Graham in FD
2126, Rawlins 275, Mutangah JG & Kamau P 14, Robertson SA 3502 (EA).
Allophylus pervillei f. trifoliolatus Radlk. Habit: Shrub. Habitat: Coastal forest
bushland and thicket, 0–400 m. Vouchers: Drummond & Hemsley 3952, Bally
4685, Rawlins 349 (EA).
Checklist of coastal forests of Kenya
177
Allophylus rubifolius var. alnifolius (Baker) Friis & Vollesen Habit: Shrub. Habitat:
Evergreen thicket and dry forest edges, 0–700 m. Vouchers: Polhill & Paulo 576,
Ross KS 118 (EA).
Allophylus rubifolius var. dasystachys (Gilg) Verdc. Habit: Tree. Habitat: Dry evergreen forest, open woodland, bushland, and cultivations, 50–150 m. Vouchers:
Luke Q 5669, Saufferer S 864 (EA).
Aporrhiza paniculata Radlk. Habit: Tree. Habitat: Riverine forest, and ground water
forest, ca. 20 m. Vouchers: Luke WRQ & Robertson SA 2762, Magogo & Glover
615, 753, F Hughes (EA).
Blighia unijugata Baker Habit: Tree. Habitat: Evergreen forest, riverine or forest
margins, ca. 0–20 m. Vouchers: Brenan et al. 14629, Mwadime N 55, Shimba
Hills Survey Unit 75 (EA).
Camptolepis ramiflora (Taub.) Radlk. Habit: Tree. Habitat: Trichilia, Garcinia, Hyphaene forest, 10–100 m. Vouchers: SA Robertson & Luke 5476, SA Robertson &
Baer 7229, Mwadime N 49 (EA).
Cardiospermum microcarpum Kunth Habit: Climber. Habitat: Coastal grassland, ca.
86 m. Vouchers: Ngumbau V & Mwadime N V0479 (EA, HIB), Kassner 339 (EA).
Chytranthus obliquinervis Radlk. ex Engl. Habit: Tree. Habitat: Lowland evergreen
forest, 5–400 m. Vouchers: SAJIT–006086 (EA, HIB), Verdcourt 3955, Drummond & Hemsley 1120, Luke Q 1515 (EA): Vulnerable.
Chytranthus prieurianus Baill. Habit: Shrub. Habitat: Evergreen forest or forest on
coral rag, 0–600 m. Vouchers: Drummond & Hemsley 3926, SA Robertson &
Luke 4911, SA Robertson & Baer 7191 (EA).
Deinbollia borbonica Scheff. Habit: Shrub. Habitat: Riverine Acacia thorn bush,
evergreen thicket, and low evergreen forest, 0–210 m. Vouchers: Ngumbau V &
Mwadime N V0395 (EA, HIB), Polhill & Paulo 861, Omondi W & Obunyali C
305 (EA).
Dodonaea viscosa (L.) Jacq. subsp. viscosa Habit: Shrub. Habitat: Coastal bushland,
on sand dunes, and coral rocks, 0–30 (–75) m. Vouchers: SAJIT–005593 & 006258
(EA, HIB), Strange 74, RM Graham, Harry van der Hagen & Simpson 88 (EA).
Dodonaea viscosa subsp. angustifolia (L.f.) J.G. West Habit: Shrub. Habitat: Margins
of evergreen forest, ca. 100 m. Vouchers: Gillett JB 20317, Robertson SA 7660 (EA).
Glenniea africana (Radlk.) Leenh. Habit: Tree. Habitat: Wet evergreen forest, riparian
forest, and bushland, ca. 30 m. Vouchers: Battiscombe 57, Luke WRQ 2929A (EA).
Haplocoelopsis africana F.G. Davies Habit: Tree. Habitat: Lowland wet, dry forest,
and thickets, 40–700 m. Vouchers: Dale IR 3820, Mrima-Dzombo Exped. 142,
Luke & SA Robertson 1433 & 247 (EA).
Haplocoelum foliolosum subsp. mombasense (Bullock) Verdc. Habit: Tree. Habitat:
Hillside grassland, Acacia-Combretum woodland, seashore bushland, 0–300 m. Vouchers: RB & AJ Faden 77/722, SA Robertson & Luke 5719, SA Robertson 6435 (EA).
Haplocoelum inoploeum Radlk. Habit: Tree. Habitat: Coastal forest edges, semi-evergreen bushland, and thicket on coral rag, 0–600 m. Vouchers: Greenway 9622,
SA Robertson 3866, Polhill & Paulo 671, Luke Q 1438, Sangai GW in EAH
15763 (EA).
178
Veronicah M. Ngumbau et al. / PhytoKeys 147: 1–191 (2020)
Lecaniodiscus fraxinifolius subsp. scassellatii (Chiov.) Friis Habit: Tree. Habitat:
Coastal forest, bushland, and riverine forest, 0–150 m. Vouchers: Verdcourt 1177,
Greenway & Rawlins 9479 (EA).
Lecaniodiscus fraxinifolius subsp. vaughanii (Dunkley) Friis Habit: Tree. Habitat:
Forest, ca. 199 m. Vouchers: Ngumbau V & Mwadime N V0423 (EA, HIB).
Lepisanthes senegalensis (Poir.) Leenh. Habit: Tree. Habitat: Evergreen lowland and
sub montane forest, 0–100 m. Voucher: Rawlins 353 (EA).
Macphersonia gracilis var. hildebrandtii (O. Hoffm.) Capuron Habit: Tree. Habitat: Coastal bushland, thicket, and forest edges, 0–100 m. Vouchers: Wamukoya
x41, Luke & SA Robertson 265 (EA).
Majidea zanguebarica J. Kirk ex Oliv. Habit: Tree. Habitat: Dry evergreen forest
fringes, riverine, and coastal bushland, 0–300 m. Vouchers: SAJIT–005577 (EA,
HIB), Gillett & Kibuwa 19874, Polhill & Paulo 803, Verdcourt 2128 (EA).
Pancovia golungensis (Hiern) Exell & Mendonça Habit: Tree. Habitat: Moist evergreen lowland forest and riverine forest, 0–500 m. Vouchers: Ngumbau V &
Mwadime N V0388 (EA, HIB), Drummond & Hemsley 1130, Dale in FD
3875, SA Robertson & Luke 5478, Mwadime N 53, Luke WRQ & Robertson
SA 2824 (EA).
Pancovia sp. Habit: Tree. Habitat: Forest. Voucher: Luke WRQ & Robertson SA
2803 (EA).
Paullinia pinnata L. Habit: Liana. Habitat: Forest margins, gallery forest, moist thicket, and scrub, 0–1600 m. Vouchers: SAJIT–006019, Ngumbau V & Mwadime N
V0262 & 080 (EA, HIB), Magogo & Glover 284 (EA).
Sapindus trifoliatus L. Habit: Tree. Habitat: Probably not naturalized anywhere,
about 0–200 m. Voucher: van Someren 316 (EA): Cultivated.
Stadmania oppositifolia Lam. subsp. oppositifolia Habit: Tree. Habitat: Dry evergreen forest and coastal bushland on coral rag, 0–5 m. Vouchers: RM Graham in
FH 2232, Gillett & SA Robertson 24025 & 24028, AJ Faden 77/387, Mwadime
N 48 (EA).
Zanha golungensis Hiern Habit: Tree. Habitat: Deciduous woodland and rainforest, ca.
300 m. Vouchers: RB & AJ Faden 72/71, Luke WRQ & Robertson SA 2732 (EA).
F156. Sapotaceae
8 Genera, 20 Species
Chrysophyllum viridifolium J.M. Wood & Franks Habit: Tree. Habitat: Evergreen
lowland forests, low to moderate elevation. Voucher: Robertson SA & Luke WRQ
5208 (EA).
Inhambanella henriquezii (Engl. & Warb.) Dubard Habit: Tree. Habitat: Lowland
rainforest and groundwater forest, 10–300 m. Vouchers: Verdcourt 3936B, Rawlins in EAH 25/58, Greenway & Rawlins 8957, Mwadime N 60, Luke Q 1513,
Luke WRQ & Baer S 8242 (EA).
Manilkara discolor (Sond.) J.H. Hemsl. Habit: Tree. Habitat: Lowland and upland
dry evergreen forest, ca. 20 m. Voucher: Mwadime N 50 (EA).
Checklist of coastal forests of Kenya
179
Manilkara mochisia (Baker) Dubard Habit: Tree. Habitat: Deciduous bushland and
thickets, 0–300 m. Vouchers: Drummond & Hemsley 4217, Dale in FD 3663,
Polhill & Paulo 677 (EA).
Manilkara sansibarensis (Engl.) Dubard Habit: Tree. Habitat: Lowland rainforest
and lowland dry evergreen forest, 0–300 m. Vouchers: Ngumbau V & Mwadime
N V0487 (EA, HIB), Suleman in FD 877, RM Graham 809 in FD 2313, RM
Graham 695 in FD 2161, Luke WRQ 3126 (EA).
Manilkara sulcata (Engl.) Dubard Habit: Tree. Habitat: Lowland dry evergreen
forest, coastal woodlands, evergreen bushlands, and thickets, 0–600 m. Vouchers: Drummond & Hemsley 4231, RM Graham in FD 2145, Kuchar P 13430,
Shimba Hills Survey Unit 107 (EA).
Manilkara sp. cf. discolor (Sond.) J.H. Hemsl. Habit: Tree. Habitat: Forest. Voucher: Luke WRQ 3139A (EA).
Mimusops aedificatoria Mildbr. Habit: Tree. Habitat: Lowland rainforest, riverine
forest, and groundwater forest, ca. 200 m. Vouchers: Templer 3, Dale in FD 3869,
Luke WRQ & Robertson SA 538 (EA).
Mimusops riparia Engl. Habit: Tree. Habitat: Riverine, ca. 120 m. Voucher: PJ
Greenway 4473 (EA): Vulnerable
Mimusops obtusifolia Lam. Habit: Tree. Habitat: Coastal evergreen bushland or dry
evergreen forest, 1–400 m. Vouchers: SAJIT–006235, 005574 & 005374 (EA,
HIB), CW Elliot in FD 1499, Battiscombe 232, Greenway & Rawlins 9481,
Kimeu, J.M. 524 (EA).
Mimusops somalensis Chiov. Habit: Tree. Habitat: Coastal woodlands and evergreen
bushlands, 150–300 m. Vouchers: Bally 8557, Luke WRQ & Robertson SA 4631,
Luke WRQ & Robertson SA 211 (EA).
Pouteria alnifolia (Baker) Roberty var. alnifolia Habit: Tree. Habitat: Lowland
rainforest, groundwater forest, and riverine forest, 0–600 m. Vouchers: Thomas
Mwadime in Mrs SA Robertson 7791, Luke WRQ & Robertson SA 242B, Drummond & Hemsley 4635, Verdcourt 1881 (EA).
Pouteria pseudoracemosa (J.H. Hemsl.) L.Gaut. Habit: Tree. Habitat: Old coral rag,
forest fragment, ca. 20 m. Voucher: Thomas Mwadime in Mrs SA Robertson 7791
(EA): Vulnerable.
Sideroxylon inerme subsp. diospyroides (Baker) J.H. Hemsl. Habit: Tree. Habitat:
Thickets on the sea shore, 0–900 m. Vouchers: SAJIT–006240 & 006241(EA,
HIB), Napier in CM 6411 & in CM 6272, J Adamson 263 in Bally 5956, Greenway & Rawlins 8874 (EA).
Synsepalum brevipes (Baker) T.D.Penn. Habit: Tree. Habitat: Lowland rainforest and
riverine forest, ca. 0–300 m. Vouchers: SAJIT–005930 (EA, HIB), Drummond &
Hemsley 4032, RM Graham in FD 2050, Luke Q 1404, Luke WRQ 1623 (EA).
Synsepalum kaessneri (Engl.) T.D.Penn. Habit: Tree. Habitat: Lowland rainforest
and riverine forest, less than 300 m. Vouchers: Kassner 398, Robertson SA 6563,
Pakia M 915 (EA).
Synsepalum msolo (Engl.) T.D.Penn. Habit: Tree. Habitat: Lowland rainforest, ca.
30 m. Voucher: Sampson 51 (EA).
180
Veronicah M. Ngumbau et al. / PhytoKeys 147: 1–191 (2020)
Synsepalum sp. cf. subcordatum De Wild. Habit: Tree. Habitat: Moist forest, 240–
400 m. Vouchers: Luke WRQ 1846, Moomaw 1056, Faden et al. 69/487 (EA).
Synsepalum subverticillatum (E.A. Bruce) T.D.Penn. Habit: Shrub. Habitat: Lowland
rainforest, 0–100 m. Vouchers: Ngumbau V & Mwadime N V0483 (EA, HIB),
Dale in FD 3583, Dale 1138 in CM 11481, Greenway & Rawlins 9431, Festo L,
Luke Q & P 2760, Luke WRQ & Robertson SA 2804, Abdallah G 3344 (EA).
Vitellariopsis kirkii (Baker) Dubard Habit: Shrub. Habitat: Lowland dry evergreen
forest and coastal evergreen bushlands, 0–300 (–360) m. Vouchers: MacNaughton
85 in FD 2620, Davies RM 1257, Robertson SA & Luke WRQ 6034 (EA).
F157. Scrophulariaceae
1 Genus, 1 Species
Anticharis senegalensis (Walp.) Bhandari Habit: Herb. Habitat: In semi desert, an
ephemeral, ca. 200 m. Voucher: D Wood 1350 (EA).
F158. Simaroubaceae
1 Genus, 1 Species
Quassia undulata (Guill. & Perr.) D.Dietr. Habit: Tree. Habitat: Thickets, open
grassland, wooded grassland, montane forest, riverine, and semi-swamp forests,
(0–) 150 m. Vouchers: Drummond & Hemsley 1173A, Faden & Faden 72/70,
Faden RB 70/835 (EA).
F159. Smilacaceae
1 Genus, 1 Species
Smilax anceps Willd. Habit: Liana. Habitat: Forest edges, clearings, secondary associations of bushland, thicket, commonly along rivers, lakesides or marshes, ca.
0–400 m. Vouchers: SAJIT–005970, Ngumbau V & Mwadime N V0295 (EA,
HIB), Magogo & Glover 1063, Robertson SA & Luke WRQ 5181 (EA).
F160. Solanaceae
8 Genera, 18 Species
Capsicum annuum L. Habit: Herb. Habitat: Secondary vegetation, forests edge, riverine thicket, and roadsides, ca. 150 m. Vouchers: Magogo FC & Glover PE 564,
MacNaughton 127 (EA).
Datura metel L. Habit: Herb. Habitat: Roadside weed or ruderal side, ca.
0–100 m. Vouchers: Graham RM 1935, Napier 6299, Thomas 154, Kassner
280 (EA): Naturalized.
Lycium shawii Roem. & Schult. Habit: Shrub. Habitat: Rocky outcrops, montane
scrub, riverine thickets, woodland, and bushland, ca. 15 m. Voucher: Luke et al.
TPR 405 (EA).
Checklist of coastal forests of Kenya
181
Lycopersicon esculentum Mill. Habit: Herb. Habitat: Cultivated. Voucher: Waaijenberg H 25 (EA): Cultivated.
Nicandra physalodes (L.) Gaertn. Habit: Herb. Habitat: Waste places and near dwellings, ca. 130 m. Voucher: Fukuoka N 267 (EA): Cultivated.
Physalis angulata L. Habit: Herb. Habitat: Damp areas, weed of cultivated crops, and
ruderal of waste places, ca. 90 m. Vouchers: SAJIT–006115, Ngumbau V & Mwadime
N V0291 (EA, HIB), Luke WRQ & PA 5724, Luke 1338 (EA): Naturalized.
Physalis minima L. Habit: Herb. Habitat: Field edges, waste ground near houses, and
roadsides. Vouchers: Brathay 1982, Expedition 40 (EA): Naturalized.
Solanum americanum Mill. Habit: Herb. Habitat: Common weed of cultivation,
grassland, open bush, forest clearings, shrubland, and upland rainforest, 0–200 m.
Vouchers: Robertson 3454, Polhill R & Paulo S 828 (EA).
Solanum campylacanthum Hochst. ex A. Rich. Habit: Shrub. Habitat: Ubiquitous
weed of roadsides, abandoned cultivation, wooded grassland, bushland, dunes,
and forest edges, ca. 5 m. Vouchers: Gillespie 24, Vorontsova MS 107 (EA).
Solanum dasyphyllum Schumach. & Thonn. Habit: Herb. Habitat: Forest, savannah, grassland, or wasteland, ca. 30 m. Vouchers: Ngumbau V & Mwadime N
V0265 (EA, HIB), Robertson SA & Luke WRQ 5177, Luke Q 1399 (EA).
Solanum goetzei Dammer Habit: Herb. Habitat: Dry to moist forest, riverine forest,
coastal thicket, woodland, wooded grassland, and bushland, ca. 0–750 m. Vouchers: Polhill 4818, Polhill & Paulo 750, BR Adams 76 (EA).
Solanum malindiense Voronts. Habit: Shrub. Habitat: Coastal bush, dunes and sand,
often on coral, 0–50 m. Vouchers: Faden & Faden 74/1072, Luke & Luke 10326,
Polhill & Paulo 709, Vorontsova MS 115 (EA).
Solanum pampaninii Chiov. Habit: Shrub. Habitat: Coastal sand dunes and shores,
0–9 m. Vouchers: Gillespie 120, Rawlins 25 & 156 (EA).
Solanum sp. cf. monotanthum Damer Habit: Herb. Habitat: Shore. Voucher: Robertson 3776 (EA).
Solanum stipitatostellatum Dammer Habit: Shrub. Habitat: Forest understorey,
open forest, forest edges or disturbed ground, ca. 300 m. Voucher: Verdcourt
3931 (EA).
Solanum usaramense Dammer Habit: Shrub. Habitat: Coastal bushland, thickets,
disturbed places, 0–500 m. Voucher: Gillett & Kibuwa 19892 (EA).
Solanum zanzibarense Vatke Habit: Shrub. Habitat: Moist or dry forest, forest edges,
and rocky outcrops, 0–700 m. Vouchers: Ngumbau V & Mwadime N V0527
(EA, HIB), Polhill & Paulo 850, Magogo FC & Glover PE 185 (EA).
Withania somnifera (L.) Dunal Habit: Herb. Habitat: Disturbed soil, along roadsides, in cultivated land, ca. 5 m. Voucher: Luke PA & WRQ 6145 (EA).
F161. Sphenocleaceae
1 Genus, 1 Species
Sphenoclea zeylanica Gaertn. Habit: Herb. Habitat: Forest, up to 350 m. Vouchers:
SAJIT–006200 (EA, HIB), Luke WRQ & PA 4575 (EA).
182
Veronicah M. Ngumbau et al. / PhytoKeys 147: 1–191 (2020)
F162. Surianaceae
1 Genus, 1 Species
Suriana maritima L. Habit: Shrub. Habitat: Seashore just above the high-water
mark, the beach crest, sea-level. Vouchers: Drummond & Hemsley 1044, Luke
3019, Kimeu JM 656 (EA).
F163. Talinaceae
1 Genus, 3 Species
Talinum paniculatum (Jacq.) Gaertn. Habit: Herb. Habitat: Moist or wet fields,
thickets. Voucher: Someren HD van 7172 (EA): Naturalized.
Talinum portulacifolium (Forssk.) Asch. ex Schweinf. Habit: Herb. Habitat: Grassland and thicket, ca. 0–300 m. Vouchers: RB & AJ Faden 74/1195, Parker I
517 (EA).
Talinum tenuissimum Dinter Habit: Herb. Habitat: Disturbed places in bushland,
50–600 m. Vouchers: Ngumbau V & Mwadime N V0340 (EA, HIB), Robertson
3484, Luke 3066, Luke WRQ et al. 6195 (EA).
F164. Tamaricaceae
1 Genus, 1 Species
Tamarix senegalensis DC. Habit: Shrub. Habitat: Sandy desert and sea shore, ca.
100 m. Voucher: Johansson S, Gachathi N & Mulatya J 99 (EA).
F165. Thymelaeaceae
2 Genera, 3 Species
Gnidia latifolia (Oliv.) Gilg Habit: Shrub. Habitat: Evergreen bushland and wooded
grassland, ca. 50–304 m. Vouchers: Drummond & Hemsley 4040, Festo L &
Luke Q 2555 (EA).
Synaptolepis alternifolia Oliv. Habit: Liana. Habitat: Woodland and wooded grassland, 30–420 m. Vouchers: Luke WRQ 3099, Luke TPR794 (EA).
Synaptolepis kirkii Oliv. Habit: Liana. Habitat: Lowland dry evergreen forest, Brachystegia woodland, coastal and secondary bushland or thicket, 0–450 m. Vouchers:
SAJIT–006143 (EA, HIB), RM Graham 1805, Polhill & Paulo 776, Verdcourt
1931, Robertson SA 5070, Simpson BL 326 (EA).
F166. Typhaceae
1 Genus, 1 Species
Typha domingensis Pers. Habit: Herb. Habitat: Swamps, dams, lakes, and rivers,
ca. 0–250 m. Vouchers: SAJIT–005914 (EA, HIB), Hildebrandt 1229, Luke Q
5643 (EA).
Checklist of coastal forests of Kenya
183
F167. Urticaceae
4 Genera, 6 Species
Laportea interrupta (L.) Chew Habit: Herb. Habitat: Lowland rainforest, often along
roads, riverine forest, and moist places in wooded grassland, 0–550 m. Voucher:
Gillespie 406 (EA).
Laportea lanceolata (Engl.) Chew Habit: Herb. Habitat: Lowland rainforest and
moist woodland, 50–400 m. Vouchers: SAJIT–006025 & 006153 (EA, HIB),
Verdcourt 1940, BR Adams 27, Luke Q 1467 (EA).
Pilea holstii Engl. Habit: Herb. Habitat: In lowland rainforest, ca. 200 m. Vouchers:
Gilbert et al. 4963, BR Adams 85, Luke WRQ 2928 (EA).
Pouzolzia fadenii Friis & Jellis Habit: Herb. Habitat: Lowland rainforest, ca.
0–220 m. Vouchers: Verdcourt 1858, Faden 71/792 & 74/1092, Luke WRQ &
Robertson SA 1915 (EA).
Urera sansibarica Engl. Habit: Liana. Habitat: Lowland rainforest, often on limestone, and climbing over rocks, 5–600 m. Vouchers: Ngumbau V & Mwadime N
V057 (EA, HIB), Magogo & Glover 613, Verdcourt 3939C, Musyoki & Hansen
954, Mwadime N 41, Saufferer S 1590 (EA).
Urera trinervis (Hochst.) Friis & Immelman Habit: Liana. Habitat: Along margins
and in clearings of lowland evergreen forest, ca. 300 m. Vouchers: SAJIT–006191
(EA, HIB), Mwadime N 32 (EA).
F168. Vahliaceae
1 Genus, 2 Species
Vahlia dichotoma (Murray) Kuntze Habit: Herb. Habitat: Grassland or woodland,
ca. 10–1370 m. Vouchers: Thulin 303, Polhill & Paulo 590 (EA).
Vahlia digyna (Retz.) Kuntze Habit: Herb. Habitat: Wet and heavy soils or on saline
flats, ca. 140 m. Vouchers: Makin, RB & AJ Faden 74/ 990 (EA).
F169. Verbenaceae
4 Genera, 5 Species
Lantana camara L. Habit: Shrub. Habitat: Roadsides or degraded lands, ca. 100 m.
Voucher: Magogo FC & Glover PE 434 (EA): Naturalized.
Lantana viburnoides (Forssk.) Vahl subsp. viburnoides Habit: Shrub. Habitat:
Woodland and bushland, ca. 40 m. Voucher: Magogo FC & Glover PE 283 (EA).
Lippia carviodora Meikle Habit: Shrub. Habitat: Acacia and Commiphora bushland,
0–700 m. Voucher: Jeffrey H21/49 (EA).
Phyla nodiflora (L.) Greene Habit: Herb. Habitat: Grassy area, ca. 23 m. Voucher:
Kirika P, Muthoka P & Mbale M 754 (EA).
Stachytarpheta urticifolia (Salisb.) Sims Habit: Shrub. Habitat: Roadside weed,
grassland, bushland, and forest, ca. 0–100 m. Vouchers: Bock in EAH 15433,
Church 73, Magogo & Glover 892 (EA): Naturalized.
184
Veronicah M. Ngumbau et al. / PhytoKeys 147: 1–191 (2020)
F170. Violaceae
2 Genera, 10 Species
Afrohybanthus enneaspermus (L.) Flicker Habit: Herb. Habitat: Ploughed lands,
riverbanks, forest margins, and disturbed area, ca. 0–335 m. Vouchers: Adams BR
53, Magogo FC & Glover PE 730, Drummond & Hemsley 4042, Luke Q 1459,
RM Graham in FD 1944, Magogo & Estes 1185, Jeffrey 298, Graham RM 1983
& 205 (EA).
Rinorea angustifolia subsp. ardisiiflora (Oliv.) Grey-Wilson Habit: Tree. Habitat:
Evergreen forest, ca. 100–300 m. Vouchers: Ngumbau V & Mwadime N V0383
(EA, HIB), Hawthorne 327, Luke PA & WRQ Luke WRQ 4321 (EA).
Rinorea angustifolia subsp. albersii (Engl.) Grey-Wilson Habit: Tree. Habitat: Evergreen forest, ca. 0–420 m. Voucher: Robertson SA, Luke Q & Kh 297 (EA).
Rinorea arborea (Thouars) Baill. Habit: Tree. Habitat: Lowland evergreen forest,
0–850 m. Vouchers: SAJIT–006076, Ngumbau V & Mwadime N V0404 (EA,
HIB), Polhill & Paulo 798, Verdcourt 1077, Mohamed Abdulla in FD 3846,
Gathii S 127, Luke WRQ 875 (EA).
Rinorea elliptica (Oliv.) Kuntze Habit: Tree. Habitat: Forest, 50–600 m. Vouchers:
Bally 2043, Mohamed Abdullah in FD 3349, Rawlins 70, Festo L, Luke Q & P
2666 (EA).
Rinorea ferruginea Engl. Habit: Shrub. Habitat: Evergreen Forest, ca. 150–400 m.
Vouchers: Magogo & Glover 405, Drummond & Hemsley 1104 & 3963, Luke
WRQ & Robertson SA 2747 (EA).
Rinorea ilicifolia (Welw. ex Oliv.) Kuntze var. ilicifolia Habit: Shrub. Habitat: Lowland and sub montane evergreen forest, 0–500 m. Vouchers: SAJIT–005533, Ngumbau V & Mwadime N V0401 (EA, HIB), Mwadime N 6, Festo L & Luke Q 2538,
Gillet 21069, Drummond & Hemsley 4262, Greenway & Rawlins 8950 (EA).
Rinorea sp. nov. Habit: Tree. Habitat: Forest, ca. 300 m. Voucher: Luke WRQ et al.
6184 (EA).
Rinorea sp. nr. beniensis Engl. Habit: Tree. Habitat: Forest, ca. 420 m. Vouchers:
Luke WRQ & Robertson SA 2758, Mrima/Dzombo Expedition 297 (EA).
Rinorea squamosa subsp. kaessneri (Engl.) Grey-Wilson Habit: Tree. Habitat: Evergreen lowland forest, 30–450 m. Vouchers: SAJIT–006033 (EA, HIB), Kassner
310, Drummond &Hemsley 3858, Mohamed Abdullah in FD 3346, Festo L,
Luke Q & P 2686 (EA).
Rinorea subintegrifolia (P. Beauv.) Kuntze Habit: Shrub. Habitat: Evergreen forest,
ca. 50 m. Voucher: Luke WRQ, PA, Parkia M & Kahumbu P 4736 (EA).
F171. Vitaceae
5 Genera, 31 Species
Ampelocissus africana (Lour.) Merr. Habit: Liana. Habitat: Seasonally swampy
grassland, coastal and deciduous thicket, ca. 0–183 m. Vouchers: SAJIT–005453,
Checklist of coastal forests of Kenya
185
Ngumbau V & Mwadime N V0204 (EA, HIB), Drummond & Hemsley 1024,
Kirika P, Mbale M & Mbatha M 768 (EA).
Ampelocissus obtusata subsp. kirkiana (Planch.) Wild & R.B. Drumm. Habit: Liana. Habitat: Woodland, bushland, thicket, and riverine vegetation, ca. 60–243 m.
Vouchers: Ngumbau V & Mwadime N V025, Luke & Robertson 1685 & 1760,
Luke WRQ & Robertson SA 2746 (EA).
Cayratia gracilis (Guill. & Perr.) Suess. Habit: Climber. Habitat: Forest, 15–120 m.
Vouchers: L& L 6009, Kimeu JM KEFRI 457 (EA).
Cayratia ibuensis (Hook.f.) Suess. Habit: Climber. Habitat: Edge of small forest
patches, derived thickets, cleared land, and papyrus swamps, ca. 100 m. Voucher:
Mutangah & Muasya 296 (EA).
Cissus adeyana Masinde & L.E. Newton Habit: Climber. Habitat: Scattered scrub
on rocky ground, ca. 300 m. Voucher: Mungai et al. 247 (EA).
Cissus albiporcata Masinde & L.E. Newton Habit: Climber. Habitat: Bushland in
rocky areas, ca. 50 m. Vouchers: Kassner 504, RM Graham in FD 1559 (EA).
Cissus aphylla Chiov. Habit: Climber. Habitat: Open bushland to dense mixed
thicket of Acacia, Commiphora, Terminalia, Grewia, etc., 100–750 m. Voucher:
Mutang’ah 63 (EA).
Cissus aphyllantha Gilg Habit: Climber. Habitat: Bushland, 240–360 m. Voucher:
Luke WRQ & Robertson SA 52 (EA).
Cissus aralioides subsp. orientalis Verdc. Habit: Climber. Habitat: Coastal and riverine evergreen forest, 60–210 m. Vouchers: Ngumbau V & Mwadime N V0380
(EA, HIB), Verdcourt 3935B, Moggridge 170, SA Robertson 6727 (EA).
Cissus integrifolia (Baker) Planch. Habit: Climber. Habitat: Evergreen forest,
woodland, bushland, and grassland, (5–) 10–300 m. Vouchers: Ngumbau V &
Mwadime N V0215 (EA, HIB), Verdcourt 1906, RM Graham D434, Rawlins in
EAH 11276, Kimeu JM 533, Magogo FC & Glover PE 197 (EA).
Cissus phymatocarpa Masinde & L.E. Newton Habit: Climber. Habitat: Coastal
bush, thickets on coral rock, and forest edges, ca. 0–225 m. Vouchers: Magogo &
Glover 997, Masinde 326, Smith SS 12 (EA).
Cissus quinquangularis Chiov. Habit: Climber. Habitat: Forest, woodland, grassland
with scattered trees, 0–300 m. Vouchers: Ngumbau V & Mwadime N V002 (EA,
HIB), Bally 5789, Brenan et al. 14660, Verdcourt 3602, Jeffery GW 137, Masinde
PS 331 (EA).
Cissus rotundifolia Vahl var. rotundifolia Habit: Climber. Habitat: CommiphoraAcacia scrub, dry forest and forest edges, 0–304 m. Vouchers: Bally & Smith
14366, Magogo FC & Glover PE 676, Frazier J 903 (EA).
Cissus rotundifolia var. ferrugineo-pubescens Verdc. Habit: Climber. Habitat: Bushland, 300–365 m. Vouchers: Verdcourt 2351, Kuchar 7345 (EA, HIB).
Cissus sciaphila Gilg Habit: Climber. Habitat: Lowland forest, riverine forest fringes,
and woodland, 0–450 m. Vouchers: Adams 101, Robertson & Luke 5394 (EA).
Cissus sylvicola Masinde & L.E. Newton Habit: Climber. Habitat: Evergreen forest, also forest on rocky hills, coral limestone, and thicket, 5–500 m. Vouchers:
186
Veronicah M. Ngumbau et al. / PhytoKeys 147: 1–191 (2020)
SAJIT–006013 (EA, HIB), Verdcourt 2406, Sangai in EAH 15735, Masinde 355,
Luke Q 1522, Masinde PS 357 (EA).
Cyphostemma adenocaule (Steud. ex A. Rich.) Desc. ex Wild & R.B. Drumm. subsp. adenocaule Habit: Climber. Habitat: Fringing forest, clearings in evergreen
forest, bushland, thicket, grassland, ca. 0–300 m. Vouchers: SAJIT–005476 (EA,
HIB), Polhill & Paulo 733, Magogo FC & Glover PE 74 (EA).
Cyphostemma buchananii (Planch.) Desc. ex Wild & R.B. Drumm. Habit: Climber. Habitat: Coastal thicket on old coral reefs, bushland, wooded grassland, and
woodland, ca. 10 m. Vouchers: SA Robertson 3548, Magogo & Glover 945, Haller
25, fide Wild & Drummond (EA).
Cyphostemma cyphopetalum var. nodiglandulosum (T.C.E.Fr.) Verdc. Habit:
Climber. Habitat: Forest, grassland margins, grassland with scattered trees and
derived waste places. Voucher: Nash LT 47 (EA).
Cyphostemma duparquettii (Planch.) Desc. Habit: Climber. Habitat: Lowland forest, 10–600 m. Vouchers: Ngumbau V & Mwadime N V0194 (EA, HIB), Verdcourt 1907, Robertson SA 6842 (EA).
Cyphostemma dysocarpum subsp. pwani Verdc. Habit: Climber. Habitat: Bushland near shore and on open rocky sites, 0–several m. Vouchers: Tweedie 944, de
Meester 48, SA Robertson 6710, Rawlins SP 1 (EA).
Cyphostemma engleri (Gilg) Desc. Habit: Climber. Habitat: Low Commiphora-Acacia-Combretum-Terminalia-Boscia bushland, (20–) 350–600 m. Vouchers: Drummond & Hemsley 4140, Kassner 502, Vercourt 5303 (EA).
Cyphostemma hildebrandtii (Gilg) Desc. ex Wild & R.B. Drumm. Habit: Climber.
Habitat: Grassland, scrub, and thicket, ca. 15–200 m. Vouchers: Ngumbau V &
Mwadime N V0544 (EA, HIB), Drummond & Hemsley 1086, RM Graham A
549 in FD 1933, Luke Q 1540 (EA).
Cyphostemma kirkianum (Planch.) Desc. ex Wild & R.B. Drumm. subsp. kirkianum
Habit: Climber. Habitat: Rainforest or lowland evergreen forest on coral, ca. 0–600 m.
Vouchers: Brenan et al. 14507, Robertson & Luke 5887, SA Robertson 5915 (EA).
Cyphostemma pachyanthum (Gilg & M. Brandt) Desc. Habit: Climber. Habitat:
Rainforest, ca. 200 m. Vouchers: Luke & Robertson 1896, Luke WRQ & Robertson SA 2254 (EA).
Cyphostemma sp. G Habit: Climber. Habitat: Coastal bushland. Voucher: Luke &
Robertson 1284 (EA).
Cyphostemma sp. I of FTEA Habit: Climber. Habitat: Moist forest, ca. 304 m.
Vouchers: Spjut RW & Ensor PD 2682, Robertson SA & Luke WRQ 5148 (EA).
Cyphostemma sp. L Habit: Climber. Habitat: Forest, ca. 70 m. Voucher: Luke &
Robertson 1691 (EA).
Cyphostemma ternatum (Forssk.) Desc. Habit: Climber. Habitat: Open bushland
and grassland, 30–230 m. Voucher: Luke & Robertson 1186 (EA).
Cyphostemma zimmermannii Verdc. Habit: Climber. Habitat: Streamside in rainforest, also Brachystegia patches, ca. 330 m. Vouchers: SAJIT–006060, Ngumbau V
& Mwadime N V028 & 0253 (EA, HIB), GG van Someren Sh 71, Magogo &
Checklist of coastal forests of Kenya
187
Glover 74, Hawthorne 208, Luke WRQ & PA, Kahumbu & Lehman 4688, Luke
WRQ & Robertson SA 2711 (EA).
Rhoicissus revoilii Planch. Habit: Liana. Habitat: Grassland, bushland, and woodland, ca. 0–410 m. Vouchers: SAJIT–005424 (EA, HIB), RM Graham 496 in
FD1871, Katz SS 75/63, Luke WRQ & Robertson SA 2786D (EA).
Rhoicissus tridentata (L.f.) Wild & R.B. Drumm. Habit: Liana. Habitat: Often associated with rocky places, ca. 240 m. Vouchers: Luke WRQ & Robertson SA
2786C & 543 (EA).
F172. Ximeniaceae
1 Genus, 2 species
Ximenia americana L. Habit: Shrub. Habitat: Wooded grassland, deciduous, and
coastal bushland, ca. 50 m. Vouchers: SAJIT–004652 (EA, HIB), Moggridge 215,
Simpson BL 296 (EA).
Ximenia caffra var. natalensis Sond. Habit: Shrub. Habitat: Dry woodland, bushland, and wooded grassland, ca. 10 m. Vouchers: Drummond & Hemsley 4167,
Mwadime N & Luke WRQ 546 & 743, Robertson SA 4317 (EA).
F173. Xyridaceae
1 Genus, 3 Species
Xyris anceps Lam. var. anceps Habit: Herb. Habitat: Perennial swamps and pools
near the coast, 0–100 m. Vouchers: Ngumbau V & Mwadime N V0548 (EA,
HIB), SA Robertson 3509 (EA).
Xyris anceps var. minima (Steud.) Lock Habit: Herb. Habitat: Seasonal pools,
marshes, and rice fields, ca. 0–200 m. Vouchers: Luke & SA Robertson 2713,
Magogo & Glover 354 (EA).
Xyris parvula Malme Habit: Herb. Habitat: Sandy seasonally wet places near the sea,
0–10 m. Voucher: RM Graham 2113 (EA).
F174. Zingiberaceae
2 Genera, 6 Species
Aframomum amaniense Loes. Habit: Herb. Habitat: Moist forest undergrowth, ca.
304 m. Voucher: Gardner FD 1461 (EA).
Aframomum angustifolium (Sonn.) K. Schum. Habit: Herb. Habitat: Forest undergrowth, forest margins, and gallery forest, ca. 300 m. Voucher: Drummond RB &
Hemsley JH 1177 (EA).
Aframomum orientale Lock Habit: Herb. Habitat: Coastal forest and abandoned
cultivation, 100–400 m. Voucher: Magogo FC & Glover PE 263 (EA).
Aframomum alboviolaceum (Ridl.) K. Schum. Habit: Herb. Habitat: Moist savannas, often in Brachystegia woodland, ca. 375 m. Voucher: Padwa 446 (EA).
188
Veronicah M. Ngumbau et al. / PhytoKeys 147: 1–191 (2020)
Siphonochilus brachystemon (K. Schum.) B.L. Burtt Habit: Herb. Habitat: Drier evergreen forests, ca. 50–300 m. Vouchers: SAJIT–005531 (EA, HIB), Luke
WRQ 3137 (EA).
Siphonochilus kirkii (Hook.f.) B.L. Burtt Habit: Herb. Habitat: Woodland, ca.
450 m. Vouchers: Ngumbau V & Mwadime N V0207 (EA, HIB), Magogo FC &
Glover PE 919 (EA).
F175. Zosteraceae
1 Genus, 1 Species
Zostera capensis Setch. Habit: Herb. Habitat: In shore sandbanks or mudflats in
fairly shallow water, 0–6 m. Vouchers: Greenway & Rawlins 8902, Isaac 105 &
98, fide Isaac 1968, Isaac A109 (EA): Vulnerable.
F176. Zygophyllaceae
3 Genera, 8 Species
Balanites aegyptiaca (L.) Delile Habit: Tree. Habitat: Wooded and scattered tree
grassland or deciduous bushland. Voucher: Perdue RE & Kibuwa SP 10096 (EA).
Balanites maughamii subsp. acuta Sands Habit: Tree. Habitat: Coastal forest, riverine thicket, and groundwater forest, ca. 0–400 m. Vouchers: Ngumbau V &
Mwadime N V0459 & 0355 (EA, HIB), Gilbert 6038 (EA).
Balanites pedicellaris Mildbr. & Schltr. Habit: Tree. Habitat: Bushland, woodland
or grassland, ca. 0–450 m. Vouchers: Ngumbau V & Mwadime N V0449 (EA,
HIB), Robertson SA & Luke WRQ 6058 (EA).
Balanites rotundifolia (Tiegh.) Blatt. Habit: Tree. Habitat: Bushland and grassland,
0–800 m. Vouchers: Luke Q 1536, Luke & Roberson 1536 (EA).
Tetraena simplex (L.) Beier & Thulin Habit: Herb. Habitat: Open places in deciduous bushland, 0–600 m. Vouchers: RB & AJ Faden 74/783, Luke WRQ
5462 (EA).
Tribulus cistoides L. Habit: Herb. Habitat: Sand and coral along coast, also open
places inland, 0–800 m. Vouchers: Verdcourt 3616, Lucas GLl, Jeffrey C & Kirika
265 (EA).
Tribulus parvispinus C. Presl var. parvispinus Habit: Herb. Habitat: Open disturbed area, ca. 2 m. Voucher: Mwadime N & Chesire C 259 (EA).
Tribulus terrestris L. Habit: Herb. Habitat: Bushland, tree grassland, ca. 40–420 m.
Vouchers: TPR 739, Mugai GM & Rucina SM 491/84 (EA).
Acknowledgments
We sincerely thank all the institutes that made it possible for us to carry out the field investigations by processing the necessary permits; National Museums of Kenya (NMK),
Kenya Forest Service (KFS), and Kenya Wildlife Service (KWS). We are grateful to the
Checklist of coastal forests of Kenya
189
following NMK staffs, the late Daniel Odhimbo, Kennedy Matheka, Paul Kirika, Peris
Kamau, Paul Musili, Brenda Nyaboke, Mathias Mbale, and John Kimeu, for their help
with the fieldwork and plant identification. Our gratitude is expressed to Roy Gereau and anonymous reviewer for their incredible suggestions and comments towards
improving the manuscript. We also express our gratitude to Jane Chilande for her
assistance in extracting data from BRAHMS database. This work was financially supported by grants from the Backbone Talents Project of Wuhan Botanical Garden, CAS
(Y655301M01), and from Sino-Africa Joint Research Center, CAS (SAJC201614).
References
African Plant Database (2020) (version 3.4.0) Conservatoire et Jardin botaniques de la Ville de
Genève and South African National Biodiversity Institute, Pretoria. [Retrieved 20.01.2020]
http://www.ville-ge.ch/musinfo/bd/cjb/africa/
APG IV (2016) An update of the Angiosperm Phylogeny Group classification for the orders
and families of flowering plants: APG IV. Botanical Journal of the Linnean Society 181(1):
1–20. https://doi.org/10.1111/boj.12385
Azeria ET, Sanmartín I, Ås S, Carlson A, Burgess N (2007) Biogeographic patterns of the East
African coastal forest vertebrate fauna. Biodiversity and Conservation 16(4): 883–912.
https://doi.org/10.1007/s10531-006-9022-0
Beentje HJ (1994) Kenya trees, shrubs and lianas. National Museums of Kenya, Nairobi.
Brenan JPM (1978) Some aspects of the phytogeography of tropical Africa. Annals of the Missouri Botanical Garden 65(2): 437–478. https://doi.org/10.2307/2398859
Brooks TM, Mittermeier RA, Mittermeier CG, Da Fonseca GA, Rylands AB, Konstant WR,
Flick P, Pilgrim J, Oldfield S, Magin G, Hilton-Taylor C (2002) Habitat loss and extinction in the hotspots of biodiversity. Conservation Biology 16(4): 909–923. https://doi.
org/10.1046/j.1523-1739.2002.00530.x
Burgess ND, Clarke GP (2000) Coastal forests of eastern Africa, IUCN-The World Conservation Union, Publications Services Unit.
Burgess N, Clarke GP, Rodgers WA (1998) Coastal forests of eastern Africa: Status, endemism
patterns and their potential causes. Biological Journal of the Linnean Society. Linnean Society of London 64(3): 337–367. https://doi.org/10.1111/j.1095-8312.1998.tb00337.x
Burgess ND, Doggart N, Doody K, Negussie G, Sumbi P, Perkin A (2003) New information
on the lowland coastal forests of eastern Africa. Oryx 37: 280–281.
Christenhusz MJM, Zhang XC, Schneider H (2011) A linear sequence of extant families and
genera of lycophytes and ferns. Phytotaxa 19(1): 7–54. https://doi.org/10.11646/phytotaxa.19.1.2
Collinge SK (1996) Ecological consequences of habitat fragmentation: Implications for landscape architecture and planning. Landscape and Urban Planning 36(1): 59–77. https://
doi.org/10.1016/S0169-2046(96)00341-6
FTEA [Eds] (1952−2012) Flora of Tropical East Africa. Royal Botanic Gardens, Kew.
Githitho A (2004) The coastal terrestrial forests of Kenya. A report on resources threats and
investments.
190
Veronicah M. Ngumbau et al. / PhytoKeys 147: 1–191 (2020)
Habel JC, Casanova ICC, Zamora C, Teucher M, Hornetz B, Shauri H, Ronald KM, Lens L
(2017) East African coastal forest under pressure. Biodiversity and Conservation 26(11):
2751–2758. https://doi.org/10.1007/s10531-017-1375-z
Hamilton AC (1974) Distribution patterns of forest trees in Uganda and their historical significance. Vegetation 29(1): 21–35. https://doi.org/10.1007/BF02390893
Hamilton AC (1982) Environmental history of East Africa. Academic press, London.
Hawthorne WD, Hunt K, Russell A (1981) Kaya; An Ethnobotanical Perspective: Report of
the Oxford ethnobotanical expedition to Kenya.
IUCN (2019) The IUCN Red List of Threatened Species. Version 2019−1. http://www.iucnredlist.org. [Downloaded on 21 March 2019]
Ivanauskas NM, Rodrigues RR, Souza VC (2007) The importance of the regional floristic diversity for the forest restoration successfulness. In: Rodrigues RR, Martins SV, Gandolfi S
(Eds) High Diversity Forest Restoration in Degraded Areas. New York, 63−76.
Lehmann I, Kioko E (2005) Lepidoptera diversity, floristic composition and structure of three
Kaya forests on the south coast of Kenya. Journal of East African Natural History 94(1):
121–164. https://doi.org/10.2982/0012-8317(2005)94[121:LDFCAS]2.0.CO;2
Linder HP (2001) Plant diversity and endemism in sub-Saharan tropical Africa. Journal of
Biogeography 28(2): 169–182. https://doi.org/10.1046/j.1365-2699.2001.00527.x
Linder HP (2014) The evolution of African plant diversity. Frontiers in Ecology and Evolution
2: 1–38. https://doi.org/10.3389/fevo.2014.00038
Lovett JC (1998) Eastern tropical African centre of endemism: A candidate for World Heritage Status. Journal of East African Natural History 87(1): 359–367. https://doi.
org/10.2982/0012-8317(1998)87[359:ETACOE]2.0.CO;2
Luke Q (2005) Annotated checklist of the plants of the Shimba Hills, Kwale District, Kenya.
Journal of East African Natural History 94(1): 5–121. https://doi.org/10.2982/00128317(2005)94[5:ACOTPO]2.0.CO;2
Matiku P, Caleb M, Callistus O (2013) The impact of participatory forest management on local community livelihoods in the Arabuko-Sokoke forest, Kenya. Conservation & Society
11(2): 112–129. https://doi.org/10.4103/0972-4923.115724
Mittermeier RA, Myers N, Thomsen JB, Da Fonseca GA, Olivieri S (1998) Biodiversity hotspots
and major tropical wilderness areas: Approaches to setting conservation priorities. Conservation Biology 12(3): 516–520. https://doi.org/10.1046/j.1523-1739.1998.012003516.x
Mittermeier RA, Robles GP, Hoffmann M, Pilgrim J, Brooks T, Mittermeier CG, Lamoreux
J, da Fonseca GAB (2004) Hotspots revisited: Earth’s biologically richest and most endangered terrestrial ecoregions. Cemex, Mexico City.
Moomaw JC (1960) A study of the plant ecology of the Coast Region of Kenya Colony, British
East Africa.
Mota TJ, Carvalho FA, Ivanauskas NM, Eisenlohr PV (2017) On the relevance of floristic and
quantitative studies to the restoration of degraded areas: the case of the Atlantic Forest
hotspot Running title: Insights for restoration in Atlantic Forest. Environmental Sciences
(Ruse) 4(1): 42–53. https://doi.org/10.3934/environsci.2017.1.42
Ngumbau VM, Nyange M, Dai C, Zhong ZX, Wei N, Malombe I, Hu GW, Wang QF (2017)
Adenia angulosa (Passifloraceae), a new species from coastal forests of Kenya and Tanzania.
Phytotaxa 313(1): 137–142. https://doi.org/10.11646/phytotaxa.313.1.10
Checklist of coastal forests of Kenya
191
Ngumbau VM, Nyange M, Wei N, Benjamin W, Paul EB, Malombe I, Hu GW, Wang QF (in
press) A New Species of Croton (Euphorbiaceae) from a Madagascan Lineage Discovered
in Coastal Kenya.
Ngumbau VM, Nyange M, Wei N, Malombe I, Hu GW, Wang QF (unpublished) Zehneria
monocarpa (Cucurbitaceae), a new species from the relicts of Kenya’s coastal forests.
Olson DM, Dinerstein E (1998) The Global 200: A representation approach to conserving
the Earth’s most biologically valuable ecoregions. Conservation Biology 12(3): 502–515.
https://doi.org/10.1046/j.1523-1739.1998.012003502.x
PPG I (2016) A community-derived classification for extant lycophytes and ferns. Journal of
Systematics and Evolution 54(6): 563–603. https://doi.org/10.1111/jse.12229
Robertson SA, Luke WRQ (1993) Kenya Coastal Forests. Report to Director NMK. World
Wide Fund for Nature, Nairobi.
Roskov Y, Ower G, Orrell T, Nicolson D, Bailly N, Kirk PM, Bourgoin T, DeWalt RE, Decock
W, Nieukerken E van, Zarucchi J, Penev L [Eds] (2019) Species 2000 & ITIS Catalogue
of Life, 2019 Annual Checklist. Species 2000: Naturalis, Leiden. Digital resource at www.
catalogueoflife.org/annual-checklist/2019
Rowe N, Speck T (2005) Plant growth forms: An ecological and evolutionary perspective. The
New Phytologist 166(1): 61–72. https://doi.org/10.1111/j.1469-8137.2004.01309.x
Wass P (1995) Kenya’s indigenous forests, IUCN, Gland, and Cambridge in collaboration
with ODA.
Wegner G, Howell KM, Davenport TRB, Burgess ND (2009) The forgotten ‘Coastal Forests’
of Mtwara, Tanzania: A biologically impoverished and yet important ecosystem. Journal
of East African Natural History 98(2): 167–210. https://doi.org/10.2982/028.098.0202
Williams EV (2017) Dovyalis keniensis (Salicaceae), a new species from the coastal forests of
Kenya. Kew Bulletin 72(4): 1–51. https://doi.org/10.1007/s12225-017-9723-4
WWF−US (2003b) Ecoregional reports: Northern Zanzibar-Inhambane coastal Forest Mosaic.
Eastern and Southern Africa Bioregions.
Younge A, Negussie G, Burgess N (2002) Eastern Africa Coastal Forest Programme. Regional
Workshop Report February, 4−7.