Vaz • Typifications of names of taxa of Bauhinia
TAXON 60 (5) • October 2011: 1464–1474
N O M E N C L AT U R E
Edited by John McNeill & Anthony E. Orchard
Typification of names of taxa of Bauhinia L. (Leguminosae : Cercideae)
from Brazil
Angela Maria Studart da Fonseca Vaz
Instituto Brasileiro de Geografia e Estatística (IBGE); current address: Instituto de Pesquisas Jardim Botânico do Rio de Janeiro, Rua
Pacheco Leão, 915, CEP 22460-050, Rio de Janeiro, Brazil; avaz@jbrj.gov.br
Abstract An alphabetical list of all the validly published names of Brazilian taxa of Bauhinia L. s.str. (= Bauhinia subg.
Bauhinia) including those first published in Pauletia Cav. is presented with details of all types including 30 new typifications.
All names are assigned to their accepted taxa following the classification for Leguminosae tribe Cercideae published by Lewis
& Forest in 2005.
Keywords Bauhinia ; Brazil; infrageneric taxa; taxonomy; typification
INTRODUCTION
Wunderlin & al. (1987) proposed a reorganization of the
tribe Cercideae of Leguminosae subfamily Caesalpinioideae
into five genera (Cercis L., Adenolobus (Harv. ex Benth.
& Hook. f.) Torre & Hill., Griffonia Baill., Brenierea Humbert,
Bauhinia L.). They classified the genus Bauhinia into 4 subgenera, 22 sections, 12 new subsections, and 30 new series, with an
estimate of about 300 species that were not listed. Wunderlin
(1976) had previously listed and typified 33 validly published
generic names referable to the tribe Cercideae. Lewis & Forest (2005) presented a new classification for tribe Cercideae.
Their treatment differs in that Barklya F. Muell., Gigasiphon
Drake, Lasiobema (Korth.) Miq., Lysiphyllum (Benth.) de Wit,
Phanera Lour., Piliostigma Hochst. and Tylosema (Schweinf.)
Torre & Hillc., treated as subdivisions of Bauhinia by Wunderlin & al. (1987), are considered distinct genera.
A Bauhinia species monograph or checklist for the Neotropics is still needed.
Three of the four subgenera of Bauhinia proposed by Wunderlin & al. (1987) occur in Brazil: (1) Bauhinia subg. Bauhinia
(57 spp.) with B. sect. Bauhinia, B. sect. Amaria (S. Mutis)
Endl., and B. sect. Pauletia (Cav.) DC.; (2) Bauhinia subg.
Elayuna (Raf.) Wunderlin & al. (3 spp.) with only B. sect.
Benthamia Fortunato & Wunderlin; and (3) Bauhinia subg.
Phanera (Lour.) Wunderlin & al. (ca. 40 spp.) with B. sect.
Caulotretus DC. and B. sect. Schnella (Raddi) Benth.
Bauhinia sects. Bauhinia, Amaria, and Pauletia were included by Lewis & Forest (2005) in their newly circumscribed
Bauhinia L. s.str., as they recognized B. subg. Phanera at generic rank and included the Neotropical taxa Schnella Raddi
and Caulotretus Rich. ex Schott within it. The Neotropical
B. sect. Benthamia was not mentioned.
Bentham (1870) presented a treatment with 64 species of
Bauhinia s.l. occurring at Brazil. In more recent times, Vaz
(1979) published a taxonomic treatment covering 23 native
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Brazilian species of Bauhinia sect. Caulotretus and Vaz
& Tozzi (2003a, b, 2005) published one that recognized 53 species of Bauhinia sect. Pauletia and Vaz & al. (2010) presented a
key for the identification of Brazilian species of Bauhinia s.str.
This paper presents a list of validly published names referring to taxa of Bauhinia sensu Lewis & Forest (2005) in alphabetical order of basionym and includes 24 new typifications.
All validly published combinations under Bauhinia are also
included in the alphabetical list with cross-reference to their
basionyms, as are two under Pauletia which are not names of
species with the same epithet as under Bauhinia. The accepted
names, are presented in bold-italics and the synonyms in italics.
TYPIFICATIONS
Bauhinia acreana Harms in Notizbl. Königl. Bot. Gart. Berlin
6: 307. 1915 – Type: Bolivia, “Gebiet des Rio Acre, Cobija”,
Ule 9441 (B, destroyed [photo in RB! ex F negative 1561];
lectotype, designated here, L!; isolectotype G [photo in
RB!]).
Bauhinia aculeata Vell., Fl. Flumin.: 171. 1829 (‘1825’) nom.
illeg., non L., 1753, nec Cav., 1799 – Type: Brazil, Rio de
Janeiro, “tum maritimis, tum mediterraneis”, in protologue
(lectotype, designated here, [original icon in Biblioteca
Nacional, Rio de Janeiro, of] Fl. Flumin. 4: tab. 83. 1831
(‘1827’)). [= Bauhinia forficata Link]
The Flora Fluminensis manuscript was finished by Vellozo and presented by him to the Lisbon Court in 1790 (Borgmeier, 1937: 79) but the first part of the text was printed only in
1825 and distributed (and hence effectively published) in 1829,
long after the author’s death in 1811 (Carauta, 1973; Stafleu
& Cowan 1986; Lima, 1995). The original illustrations are
kept at the National Library, Rio de Janeiro. Bauhinia aculeata
was given the diagnosis “B. caule aculeato. (tab. 83ª T. 4)” by
TAXON 60 (5) • October 2011: 1464–1474
Vellozo (1829) and this phrase is a reference to the original
material preceding the description of the species, together with
the type locality (stated above). The type material of Vellozo no
longer exists. Brother Arrabida, the editor of the Icones (Vellozo, 1831), wrote in the preface that the illustrations were done
by Vellozo, himself, so the original illustration of Bauhinia
aculeata is original material for the name. There is no need to
indicate an epitype as Vellozo’s figure of B. aculeata is elucidative of the flowers, fruit, and vegetative characters.
This name is a later homonym of B. aculeata L., the name
of a recognised Neotropical species that does not occur in Brazil.
Bauhinia acuruana Moric., Pl. Nouv. Amer. 6: 77. 1840 ≡ Pauletia acuruana (Moric.) Schmitz in Bull. Jard. Bot. Natl.
Belg. 43: 387. 1973 – Type: Brazil, Bahia, Serra Açuruá,
Blanchet 2825 (holotype G; isotypes NY!, P!).
Moricand’s herbarium is now part of the general herbarium
at Geneva (G) (Stafleu & Cowan, 1981).
Bauhinia acuruana var. nitida Benth. in Martius, Fl. Bras.
15(2): 187. 1870 – Type: Brazil, Minas Gerais, S. Romão,
Gardner 4536 (lectotype, designated here, K, Herb.
Hook. 1867 [photo in RB!]; isolectotype OXF!). Brazil,
Minas Gerais, Claussen s.n. (remaining syntype, not located). [= Bauhinia acuruana Moric.]
Wunderlin annotated on the sheet, the collection Gardner
4536 (K) as lectotype of Bauhinia acuruana var. nitida and his
unpublished designation is effectively published here.
Bauhinia affinis Vogel in Linnaea 10: 594. 1836 ≡ Pauletia
affinis (Vogel) A. Schmitz in Bull. Jard. Bot. Natl. Belg.
43: 388. 1973 – Type: Brazil, Santa Catarina, Chamisso
s.n. (B, destroyed [photo in RB! ex F negative 1564]; lectotype LE; designated by Fortunato, 1996: 405; isolectotype
fragm. BAB).
Bauhinia albicans Vogel in Linnaea 13: 304. 1839 – Type:
Brazil, Rio de Janeiro, “Sellow inter Rio et Cabo Frio” (B,
destroyed [photo in RB! ex F negativo 1565]; neotype, designated here: Brazil, Rio de Janeiro, Saquarema, Morro
dos Pregos, estrada do Jundiá, próximo da Fazenda Santa
Barbara, 22°51′53″ S, 42°33′10″ W, 12. I. 2005, H.C. de
Lima & al., 6232 (RB).
The type labelled “Sellow 201, inter Rio et Cabo Frio”, deposited in Berlin, was destroyed during the Second World War.
There is no evidence of isotypes of this collection. The neotype,
designated above, was collected at Saquarema, which is located
between the Rio de Janeiro and Cabo Frio municipalities. This
locality is in Regiao dos Lagos, in Rio de Janeiro state. See
also Rodrigues & Vaz (2008) for a detailed description and
illustration of B. albicans.
Bauhinia amplifolia Ducke in Bol. Técn. Inst. Agron. N. 2:
16. 1944 – Type: Brazil, Amazonas, Tabatinga, Solimões
River, Ducke 1594 (lectotype MG 17988!, designated
by Vaz & Tozzi, 2005: 484; isolectotypes RB!, R!, SI).
Peru: Iquitos, Ducke Herb. Amaz. Musaei Paraensis 7552
Vaz • Typifications of names of taxa of Bauhinia
(remaining syntype MG!; isosyntype INPA!). [= Bauhinia
tarapotensis Benth.]
Bauhinia angulicaulis Harms in Bot. Jahrb. Syst. 33, Beibl.
72: 20. 1903 ≡ Pauletia angulicaulis (Harms) A. Schmitz
in Bull. Jard. Bot. Natl. Belg. 43: 388. 1973 –Type: Brazil,
Goiás, “Fazenda do Palmital”, Glaziou 21018 (B, destroyed
[photo in RB! ex F negative 1563]; lectotype, designated
here, C!; isolectotypes K! [photo in RB!], G [photo in
RB!]).
Bauhinia armata Thunb., Pl. Bras. 3: 33. Mai 1821 – Type: Brazil, leg. Westin & Freyreiss s.n. (holotype UPS-THUNB
9902 [photo in RB! ex UPS]) [= Bauhinia forficata Link].
The text on top of the sheet reads “Mus. Westiniam”. The
text on the reverse reads “Brasilia. Westin. Jan. Febr. Freyreiss”.
According to Mats Hjertson (pers. comm.) it is unclear if the
collection was made jointly, or if it was made by only one of
the two collectors mentioned, both of whom collected in Brazil.
Bauhinia aromatica Ducke in Anais Acad. Brasil. Ci. 31: 295.
1959 – Type: Brazil, Ceará, Serra de Maranguape, riacho
Pirapora, Ducke 2337, 1ª collection, 15-04-1954 (lectotype
EAC!, designated by Vaz & Tozzi, 2003b: 87; isolectotype MG 24481!). Brazil, Ceará, Ducke 2337, 2ª collection: 20-07-1955 (remaining syntype MG!). [= Bauhinia
cheilantha]
Bauhinia aureopunctata Ducke in Arch. Jard. Bot. Rio de
Janeiro 4: 53. 1925 – Type: Brazil, Pará, Tapajós river, Vila
Braga, Ducke Herb. Jard. Bot. Rio 16959 (lectotype RB!,
designated by Vaz & Tozzi, 2003b: 80). Brazil, Pará, lugar
Francez, Tapajós river, Ducke Herb. Jard. Bot. Rio 11119
(remaining syntype RB!); Bella Vista, Tapajós river, Ducke
Herb. Amaz. Musaei Paraensis 15823 (remaining syntype
MG; isosyntype RB!). According to Egler (1963), the locality “lugar Francez”, corresponds to the lower Tapajós river.
Bauhinia bauhinioides (Mart.) J.F. Macbr. ≡ Perlebia bauhinioides Mart.
Bauhinia benthamiana Taub. in Engler & Prantl, Nat. Pflanzenfam. 3 (3): 149. 1892 ≡ Bauhinia macrostachya Benth.
[= Bauhinia ungulata L.]
Bauhinia bicolor (Bong.) Steud. ≡ Pauletia bicolor Bong.
Bauhinia bicuspidata Benth. in Martius, Fl. Bras. 15(2): 193.
1870 ≡ Pauletia bicuspidata (Benth.) A. Schmitz in Bull.
Jard. Bot. Natl. Belg. 43: 388. 1973 ≡ Bauhinia longicuspis
var. bicuspidata (Benth.) Wunderlin ex Lewis, Legumes
Bahia: 93. 1987 – Type: Brazil, Amazonas “in sylvis Yapurensibus prov. Alto Amazonas”, Martius 1202 (holotype
M [photo in RB! ex M]).
Bauhinia bombaciflora Ducke in Arch. Jard. Bot. Rio de Janeiro 3: 104. 1922 – Type: Brazil, Pará, Itaboca, Tocantins
1465
Vaz • Typifications of names of taxa of Bauhinia
river, near the Itaboca waterfall, Ducke 16236 (holotype
MG!; isotypes R!, RB!).
Bauhinia bongardii Steud. ≡ Pauletia ferruginea Bong. [=
Bauhinia brevipes Vogel]
Bauhinia brachycalyx Ducke in Trop. Woods 90: 12. 1947 –
Type: Brazil, Amazonas, region of the middle Purus river,
boca do Pauini, “in Musaei Paraensis horto culta florebat”,
Ducke 2045 (holotype RB!; isotype MG!).
Bauhinia brasiliensis Vogel in Linnaea 13: 302. 1839 – Type:
Sellow (syntype, not located); Brazil, Rio de Janeiro,
Luschnath s.n. (syntype LE, pers. comm. A.A. Oskolski,
14. Jun. 2001; fragment and photo BAB, apud Fortunato,
1996: 408). [= Bauhinia forficata Link subsp. forficata]
Bauhinia brasiliensis var. longiflora Wawra, Bot. Ergebn.: 3–4,
pl. 35. 1866 – Type: not located. [= Bauhinia forficata
Link, fide Bentham in Martius, 1870]
Bauhinia breviloba Benth. in Martius, Fl. Bras. 15(2): 197.
1870 ≡ Pauletia breviloba (Benth.) A. Schmitz in Bull.
Jard. Bot. Natl. Belg. 43: 388. 1973 – Type: Brazil, Piauí,
Barra do Jardim, Gardner 1937 (syntype K; isosyntype
OXF! [photos in RB!]); frontier between the States of Rio
de Janeiro and Espírito Santo, Rio Itabapoana, Principe
Maximilianus Neovidensis (not located); Bahia, Serra Jacobina, Blanchet 3293 (syntype B, destroyed [photo in RB!
ex F negative 1570], OXF! [photo in RB!], BM, G: photos
in RB!; [see also Fortunato, 1996]). [= Bauhinia forficata
Link subsp. forficata]
Bauhinia brevipes Vogel in Linnaea 13: 307. 1839 – Type:
Brazil, Minas Gerais, Pitangui (sphalm. Pitargui), 18. Dec.
1818 (fr), Sellow s.n. (B, destroyed [photo in RB! ex F
negative 1568]).
Bauhinia burchellii Benth. in Martius, Fl. Bras. 15(2): 186.
1870 ≡ Pauletia burchellii (Benth.) A. Schmitz in Bull.
Jard. Bot. Natl. Belg. 43: 394. 1973 – Type: Brazil, Goiás,
between Funil and São João, Burchell Cat. Geogr. Pl. Bras.
Tropicae 8953 (holotype K H395/83-13 [photo in RB! ex
NY negative 1582; photo A!]).
Bauhinia caatingae Harms—see under Bauhinia catingae
Harms.
Bauhinia caloneura Malme in Bih. Kongl. Svenska Vetensk.Akad. Handl. 25, Afd. 3, no. 11: 29, pl. 2, fig. 4. 1900
– Type: Brazil, Mato Grosso, Cuiabá, Malme I: 1138
(lectotype S!, designated by Vaz, 1995: 2). Brazil, Cuiabá,
Malme I: 1138* (remaining syntypes S!; isosyntype R!).
Note: original illustration at S!.
Bauhinia campestris Malme in Ark. Bot. 5(5): 10. 1905 – Type:
Brazil, Mato Grosso, “Santa Anna da Chapada” (currently
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TAXON 60 (5) • October 2011: 1464–1474
Chapada dos Guimarães), Malme II: 2364b (lectotype S!,
designated by Vaz, 1995: 2). Brazil, Mato Grosso, Malme
II: 2364 and 2364a (remaining syntypes S! [photos in RB]).
Bauhinia candelabriformis R.S. Cowan in Los Angeles
County Mus. Contr. Sci. 13: 4. 1957 – Type: Brazil, Goiás,
7 km south of Veadeiros (currently town of Alto Paraíso),
Chapada dos Veadeiros, 47°30′ W, 14°30′ S, Dawson 14581
(holotype R!; isotype RB!).
Bauhinia candicans Benth. in Martius, Fl. Bras. 15(2): 201.
1870 ≡ Pauletia candicans (Benth.) A. Schmitz in Bull.
Jard. Natl. Belg. 43: 389. 1973 – Type: “ad Missiones uruguayensis”, Tweedie s.n. (lectotype K, selected by Fortunato, 1987: 550, photo in A! ex NY negative 1636). [=
Bauhinia forficata subsp. pruinosa (Vogel) Fortunato
& Wunderlin]
Bauhinia cataholo Hoehne in Relat. Commiss. Linhas Telegr.
Estratég. Matto Grosso Annexo 5, Bot. 8: 34, t. 140. 1919 –
Type: Brazil, Mato Grosso, Juruena river, Hoehne Comm.
Rondon 1989 (holotype R!, photo in RB!). [= Bauhinia
ungulata var. cuyabensis (Bong.) Vaz]
Bauhinia catingae Harms in Bot. Jahrb. Syst. 42: 209. 1908
– Type: Brazil, Bahia, Tamburi, Ule 7277 (lectotype, designated here, G [photo in RB!]; isolectopype K [photo in
RB!]; holotype B, destroyed [photo in F negative 1571!]).
The epithet “catingae” is a variant spelling of “caatingae”,
a type of vegetation found in northeastern Brazil. [≡ Bauhinia
caatingae orth. mut. (Vaz & Tozzi, 2005)].
Bauhinia chapadensis Malme in Ark. Bot. 5(5): 13. 1905 –
Type: Brazil, Mato Grosso, “Santa Anna da Chapada (currently Chapada dos Guimarães), Malme II: 3409a (lectotype S!, designated by Vaz, 1995: 3). Brazil, Mato Grosso,
ibidem, Malme II: 3409 and 3409b (remaining syntypes S!
[photos in RB!]). [= Bauhinia ungulata var. cuyabensis
(Bong.) Vaz]
Bauhinia cheilantha (Bong.) Steud. ≡ Pauletia cheilantha
Bong.
Bauhinia choriophylla Vogel in Linnaea 13: 307. 1839 – Type:
Brazil, Rio de Janeiro, Sellow (holotype B, destroyed
[photo in RB! ex NY negative 1584]; lectotype, designated here, K Herb. Hook. 1867; isolectotype E-GL!). [=
Bauhinia rufa (Bong.) Steud.]
Bauhinia cinnamomea DC., Prodr. 2: 517. 1825 – Type: Guayana Francesa, Cayenna: Martin s.n. (holotype P; isotype
M [photo in RB! ex F negative 1572]).
Bauhinia conwayi Rusby in Bull. New York Bot. Gard. 8: 92.
1912 – Type: Bolivia, Tumupasa, R.S. Williams 485 (holotype NY! [photo in RB! ex NY negative 4983]; isotype
BM!).
TAXON 60 (5) • October 2011: 1464–1474
Bauhinia cordata Vogel in Linnaea 11: 308. 1839 ≡ Bauhinia
rufa var. cordata (Vogel) Bentham in Martius, Fl. Bras.
15(2): 187. 1870 ≡ Pauletia rufa var. cordata (Vogel)
A. Schmitz in Bull. Jard. Bot. Natl. Belg. 43(3–4): 392.
1973 – Type: Brazil, Minas Gerais, “pr. Pompeo fusa aliisq.
locis”, Sellow s.n. (B, destroyed; lectotype, designated
here, K Herb. Benth. 1854 [photo in RB! ex K negative
16423 and NY negative 1586]; isolectotype E-GL!). [=
Bauhinia holophylla (Bong.) Steud.]
Bauhinia corniculata Benth. in Martius, Fl. Bras. 15(2): 198.
1870 ≡ Pauletia corniculata (Benth.) A. Schmitz in Bull.
Jard. Bot. Natl. Belg. 43: 394. 1973 – Type: Brazil, Pará,
Óbidos, Spruce 497 (holotype K [photo in RB! ex F negative 1573]).
Bauhinia corumbensis S. Moore in Trans. Linn. Soc. London, Bot. 4(2): 348. 1895 – Type: Brazil, Mato Grosso,
Corumbá, S. Moore 1039 (holotype, not located; isotype
NY! [photo in RB!]). [= Bauhinia mollis (Bong.) D. Dietr.]
Bauhinia cupulata Benth. in Martius, Fl. Bras. 15(2): 188. 1870
≡ Pauletia cupulata (Benth.) A. Schmitz in Bull. Jard. Bot.
Natl. Belg. 43: 394. 1973 – Type: Brazil, Piauí, banks of the
Gurgea river, Gardner 2529 (lectotype, designated here,
K!; isolectotypes K, MANCH!, OXF!, W! [photos in RB!
ex negatives IPA 1197 and F 1577]). Brazil, Goiás, “prope
Goyaz [currently Goiás Velho], Cavalcante, Conceição, aliosque locos”, Burchell s.n.; “in eadem provinciae?”, Pohl
s.n. (remaining syntypes, not located).
Wunderlin annotated Gardner 2529 (K) as a lectotype of
B. cupulata here accepted.
Bauhinia cupulata var. longifolia Benth. in Martius, Fl. Bras.
15(2): 189. 1870 – Type: Brazil, Goiás, “in sylvis humidis
Sertão d’Amaro-Leite”, Weddel 2875 (holotype P! [photo
in RB!]). [= Bauhinia cupulata Benth.]
Bauhinia curvula Benth. in Martius, Fl. Bras. 15(2): 194. 1870
≡ Pauletia curvula (Benth.) A. Schmitz in Bull. Jard. Bot.
Natl. Belg. 43: 389. 1973 – Type: Brazil, Goiás, Goiás
Velho, “in campis prope Goyaz”, Weddell 2590 (lectotype
K [photo in RB! ex NY negative 1595], designated by Vaz
& Tozzi, 2003b: 95). Brazil, Pohl 1223 (remaining syntype
M [photo in RB! ex F negative 6218]).
Bauhinia cuyabensis (Bong.) Steud. ≡ Pauletia cuyabensis
Bong.
Bauhinia cuyabensis var. ferruginea Chodat & Hassl. in Bull.
Herb. Boissier, ser. 2, 4: 689. 1904 (‘cujabensis’) – Type:
Paraguay, “ad marginem silvae, Arroyo Mocoy, Sierra de
Maracayu“, E. Hassler 5380 (syntype G; isosyntype NY,
[photo in RB!]); Paraguay, “in dumetis pr. Conception“,
E. Hassler 7192 (syntype G; isosyntype NY [photo in RB!]);
ibdem, E. Hassler 7192a (syntype G; isosyntype NY [photo
in RB!]). [= Bauhinia ungulata var. cuyabensis (Bong.) Vaz]
Vaz • Typifications of names of taxa of Bauhinia
Bauhinia cuyabensis f. vestita Chodat & Hassl. in Bull. Herb.
Boissier, ser. 2, 4: 689. 1904 (‘cujabensis’) – Type: Paraguai, “in campo Apepu (Tapiraguay)“, E. Hassler 5953
(syntype G; isosyntype NY, photo in RB!); idem “ad marginem silvarum in valle fluminis Y-aca“, E. Hassler 6654
(syntype G; isosyntype NY, photo in RB!). [= Bauhinia
ungulata var. cuyabensis (Bong.) Vaz]
Bauhinia dodecandra (Bong.) Steud. ≡ Pauletia dodecandra
Bong. [= Bauhinia holophylla (Bong.) Steud.]
Bauhinia dubia G. Don, Gen. Hist. 2: 463. 1832, non Bauhinia
dubia Vogel in Linnaea 13: 314. 1839 ≡ Pauletia dubia
(G. Don) A. Schmitz in Bull. Jard. Bot. Natl. Belg. 43:
394. 1973 – Type locality: Brazil, “Native of Maranham”,
“Gardner 6008” on a label (lectotype designated here,
BM No. 535635! [photo in RB!]).
Wunderlin annotated as lectotype of B. dubia the specimen at BM formally designated as such above. In publishing
the name, Don wrote “B.? dubia”, being unsure as to its generic
assignment. No collection data were provided beyond the statement “Native of Maranham” along with the field characters
“Flowers white” and “Shrub 1 to 2 feet”. Don included in his
description “leaves alternate, cordate, acuminated, glabrous
and obsoletely 3-nerved” and these leaf characters are represented on the specimen designated here as lectotype. There are
at least two sheets annotated as “6008, Bauhinia sp. a shrub
6 feet high—dry woods Maranham, june/41 [i.e., 1841]” and
attributed to Gardner: one sheet is at Kew with fruits, and the
other at BM [BM No. 535635] with flowers. The BM sheet
corresponds closely with the description in the protologue that
refers to flower characters but does not mention fruit. Bentham
(1870) cited under Bauhinia dubia two collections from the same
locality (“habitat”): “in sylvis siccis circa Maranhão: Gardner
6008, G. Don [s.n.]”. The specimens at BM and K do not belong
to the same collection. According to Urban (1906), Gardner
only went to Maraham in 1841, long after Don had published
Bauhinia dubia (1832). Examining the sheet from BM that corresponds exactly to the original description, I prefer to choose
this material as lectotype of Bauhinia dubia, rather than treating
it as a neotype, because I believe it has been wrongly annotated
as to collector. I am conviced that this collection is original
material, and probably was made by George Don himself, who
had been in Maranhão state, in 1822–1823 (Urban, 1906).
Bauhinia dumosa Benth. in Martius, Fl. Bras. 15(2): 194. 1870
≡ Pauletia dumosa (Benth.) A. Schmitz in Bull. Jard. Bot.
Natl. Belg. 43: 394. 1973 – Type: Brazil, Goiás, between
Goiás and Cavalcante, Burchell Cat. Geogr. Pl. Bras. Tropicae 7790 (lectotype K 76 [photo in RB! ex NY negative
1594], designated by Vaz & Tozzi, 2003b: 101). Brazil,
Bahia, Juazeiro, São Francisco River, “in sylvis inundatis”, Martius s.n. (remaining syntype M [photo in RB! ex
F negative 6219; this sheet was wrongly determinated by
Bentham, it is a sheet of Bauhinia bauhinioides]); Brazil, Minas Gerais, “inter Diamantina et Rio Paranayba”,
Burchell s.n. (not located).
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Vaz • Typifications of names of taxa of Bauhinia
Bauhinia dumosa var. viscidula (Harms) Vaz & R. Marquete
≡ Bauhinia viscidula Harms
Bauhinia elongipes R.S. Cowan in Los Angeles County Mus.
Contr. Sci. 13: 6. 1957 – Type: Brazil, Goiás, Formosa,
“forest and forest margin along road 22–23 km, east of
Formoso, region of southern Serra Dourada”, 48°50′ W,
13°45′ S, Dawson 15051 (holotype R!; isotype NY! [photo
in RB!]).
Bauhinia ferruginea (Bong.) D. Dietr. ≡ Pauletia ferruginea
Bong.
Bauhinia forficata Link, Enum. Hort. Berol. Alt. 1: 404. 16
Mar–30 Jun 1821 subsp. forficata ≡ Pauletia forficata
(Link) A. Schmitz in Bull. Jard. Bot. Natl. Belg. 43: 389.
1973 – Type: “Hab. in Brasilia” (not located).
The original description refers to a shrub cultivated at the
Berlin Botanical Garden from seeds collected in Brazil and
donated by Prince Maximiliano de Neuwied. The holotype
was illustrated (Link & Otto, 1825: tab. 36).
Bauhinia forficata subsp. pruinosa (Vogel) Fortunato & Wunderlin ≡ Bauhinia pruinosa Vogel
Bauhinia forficata var. latifolia Benth. in Martius, Fl. Bras:
15(2): 201 ≡ Pauletia forficata var. latifolia (Benth.)
A. Schmitz in Bull. Jard. Bot. Natl. Belg. 43: 390. 1973
– Type: Brazil, São Paulo, Amparo, G.A. Lindberg 403
(holotype S! [photo in RB!]). [= Bauhinia forficata Link
var. forficata]
Bauhinia forficata var. longiflora (Bong.) Benth. ≡ Pauletia
longiflora Bong.
Bauhinia forficata var. platypetala (Burch. ex Benth.) Wunderlin ≡ Bauhinia platypetala Burch. ex Benth.
Bauhinia fusconervis (Bong.) Steud. ≡ Pauletia fusco-nervis
Bong.
Bauhinia gardneri Benth. in Martius, Fl. Bras. 15(2): 186. 1870
≡ Pauletia gardneri (Benth.) A. Schmitz in Bull. Jard. Bot.
Natl. Belg. 43: 395. 1973 – Type: Brazil, Goiás: “Mission
of Douro”, Gardner 3123 (lectotype, designated here,
K [photo in RB!]; isolectotypes BM!, K! [photo in RB!],
OXF!, W!). Brazil, Arrayas, Gardner 3696 (remaining
syntype K [photo in RB!]).
Wunderlin annotated Gardner 3123 (K) as a lectotype of
B. gardneri here accepted.
Bauhinia geminata Vogel in Linnaea 13: 305. 1839 ≡ Pauletia geminata (Benth.) Schmitz in Bull. Jard. Bot. Natl.
Belg. 43: 395. 1973 – Type: Brazil, “Brasilia meridionalis”, Sellow s.n. (B, destroyed [photo in RB! ex F negative 1580], no duplicate located). [= Bauhinia longifolia
(Bong.) Steud.]
1468
TAXON 60 (5) • October 2011: 1464–1474
Bauhinia goyazensis Harms in Bot. Jahrb. Syst. 33, Beibl.
72: 21. 1903 – Type: Brazil, Goiás, Fazenda do Paranana,
Glaziou 21012 (B, destroyed [photo in RB! ex F negative
1582, NY negative 1619]; lectotype, designated here, K!
[photo in RB!]).
Bauhinia grandifolia (Bong.) Steud. ≡ Pauletia grandifolia
Bong.
Bauhinia hagenbeckii Harms in Bot. Jahrb. Syst. 33, Beibl.
72: 21. 1903 – Type: Brazil, “Gran Chaco”, Hagenbeck s.n.
(B, destroyed [photo in RB! ex F negative 1585]; lectotype
F!, designated by Wunderlin, 1968: 245).
Bauhinia hassleriana Chodat in Chodat & Hassl. in Bull. Herb.
Boissier, ser. 2, 4: 690. 1904, type not cited.
Chodat (l.c.) segregated Bauhinia hassleriana into four
forms (f. angustifolia, f. intermedia, f. latifolia, f. acuminata)
and a variety (var. marginata) in the same paper, citing their
types, all of them from Paraguay. See Wunderlin (1968) for
discussion. [ = Bauhinia hagenbeckii]
Bauhinia heterandra Benth. in Martius, Fl. Bras. 15(2): 196.
1870 – Type: Brazil, Ceará, near Icó, Gardner 1565 (lectotype, designated here, K [photo in RB! ex NY negative 1597]; isolectotypes G, K [photo in RB!]). Brazil,
Bahia, Joazeiro, Martius 2333 (remaining syntypes M,
two sheets; H. Esser, pers. comm.); Piauí, Boa Esperança,
Gardner 2156 (syntype K [photo in RB!]; isosyntypes
G [photo in RB!], OXF!); Goiás, Goiás Velho, Burchell
Cat. Geogr. Pl. Bras. Tropicae 73AA (syntype K [photo
in RB!]); between Funil and S. João, Burchell Cat. Geogr.
Pl. Bras. Tropicae 8896 et 8868-2 (syntypes, K [photos in
RB!]). [= Bauhinia pentandra (Bong.) D. Dietr.]
Wunderlin annotated Gardner 1565 (K) as a lectotype of
B. heterandra here accepted.
Bauhinia hiemalis Malme in Ark. Bot. 5(5): 13. 1905 – Type:
Brazil. Mato Grosso, Cuiabá, Malme II: 1628 (lectotype S!,
designated by Vaz, 1995: 2). Brazil, idem, Malme II: 1643a
(remaining syntype S!); ibidem, 1643b (remaining syntype
S!; isosyntypes R!); ibidem, 1643d (remaining syntype S),
ibidem 1643 f (remaining syntype S!). [= Bauhinia ungulata var. cuyabensis (Bong.) Vaz]
Bauhinia hirsuta (Bong.) Vogel ≡ Pauletia hirsuta Bong.
Bauhinia hirsutiflora Vaz ≡ Pauletia hirsuta Bong.
Bauhinia holophylla (Bong.) Steud. ≡ Pauletia holophylla
Bong.
Bauhinia holophylla var. paraensis Ducke in Arch. Jard. Bot. Rio
de Janeiro 4: 52. 1925 ≡ Bauhinia longicuspis var. paraensis
(Ducke) Wunderlin in Ann. Missouri Bot. Gard. 60 (2):
571. 1973 – Type: Brazil, Pará, Vila Braga, Tapajós river,
Ducke Herb. Amaz. Musaei Paraensis 16909 (lectotype
TAXON 60 (5) • October 2011: 1464–1474
RB!, designated by Vaz & Tozzi, 2003b: 110; isolectotype
MG) [= Bauhinia longicuspis Spruce ex Benth.]. Brazil,
Pará, Santarém, Serra Piroca, Ducke Herb. Amaz. Musaei Paraensis 16347 (remaining syntype MG; isosyntype
RB!) [= Bauhinia longicuspis Spruce ex Benth.]. Brazil,
Mato Grosso, heads of the Novo river, affluent of Arinos,
Kuhlman 406 (remaining syntype R 2626!) [= Bauhinia
longicuspis Spruce ex Benth.]; Brazil, Pará, “in vicinibus
Santarem”, Spruce named on label as Bauhinia longistyla
(remaining syntypes, RB 5103!, FI!) [= Bauhinia dubia
G. Don].
Bauhinia integerrima Benth. in Martius, Fl. Bras. 15(2): 197.
1870 ≡ Pauletia integerrima (Benth.) A. Schmitz in Bull.
Jard. Bot. Natl. Belg. 43: 394. 1973 – Type: Brazil, Bahia,
“ad Villa da Caxoeira”, Martius Iter. Brasil. 2182 (holotype
M [photo in RB! ex F negative 6221]).
Bauhinia intermedia Vogel in Linnaea 11: 309. 1839 ≡ Bauhinia
rufa var. intermedia (Vogel) Benth. in Martius, Fl. Bras.
15(2): 187. 1870 ≡ Pauletia rufa var. intermedia (Vogel)
A. Schmitz in Bull. Jard. Bot. Natl. Belg. 43(3–4): 392.
1973 – Type: “Brasilia”, Sellow s.n. (B, destroyed; lectotype, designated here, K Herb. Hook. 1867 [photo in RB!,
negative NY 1584]; isolectotype E-GL!). [= Bauhinia rufa
(Bong.) Steud.]
Bauhinia lamprophylla Harms in Bot. Jahrb. Syst. 33, Beibl.
72: 22. 1903 – Type: Brazil, Goiás, “Serra dos Cristais
dans le campo”, Glaziou 21008 (B Herb. Urban, destroyed
[photo in RB! ex F negative 1592]; lectotype, designated
here, K [photo in RB!, NY negative 1620]; isolectotypes
C!, G [photo in RB!];).
Bauhinia leptantha Malme in Ark. Bot. 5(5): 11. 1905 – Type:
Brazil, Mato Grosso, Corumbá, Dec. 1902 (bt/fl), Malme
II: 2730 (lectotype S!, designated by Vaz, 1995: 2; isolectotypes R!, S!). Brazil, idem, Malme II: 2730a (remaining syntype S!); Malme II: 2730b (remaining syntype S!);
Malme II: 2730c (remaining syntype S!).
Bauhinia longicuspis Spruce ex Benth. in Martius, Fl. Bras.
15(2): 185. 1870 ≡ Pauletia longicuspis (Benth.) A. Schmitz
in Bull. Jard. Bot. Natl. Belg. 43: 395. 1973 – Type: Brazil, Amazonas, Rio Negro capoeira near mouth of river
Caburés, Spruce 1978 (holotype K [photo in RB! ex NY
negative 1580]).
Bauhinia longicuspis var. bicuspidata (Benth.) Wunderlin ex
Lewis ≡ Bauhinia bicuspidata Benth.
Bauhinia longicuspis var. paraensis (Ducke) Wunderlin ≡
Bauhinia holophylla var. paraensis Ducke
Bauhinia longiflora (Bong.) Steud. ≡ Pauletia longiflora Bong.
Bauhinia longifolia (Bong.) Steud. ≡ Pauletia longifolia Bong.
Vaz • Typifications of names of taxa of Bauhinia
Bauhinia longifolia var. acuminata Micheli in Vidensk. Meddel. Naturhist. Foren. Kjøbenhavn 1875: 118. 1875. Unlocated opus. [Uncertain identity]
Bauhinia longipedicellata Ducke in Arch. Jard. Bot. Rio de
Janeiro 3: 105. 1922 – Type: Brazil, Pará, Pimental, Tapajoz river, Ducke Herb. Amaz. Musaei Paraensis. 17064
(lectotype MG!, designated by Vaz & Tozzi, 2003b: 114;
isolectotype RB 11140!). Brazil, Pará, region of the lower
waterfalls of the Tapajóz rive, Ducke Herb. Amaz. Musaei
Paraensis 16864 (remaining syntype MG!; isosyntype RB
11142!); Pará, Santarém, colonia Poço Branco [Santarém,
according to Egler, 1963], Ducke Herb. Amaz. Musaei
Paraensis 17101 (remaining syntype MG!; isosyntype
RB11141!).
Bauhinia macrostachya Benth. in J. Bot. (Hooker) 2: 96. 1840
≡ Bauhinia benthamiana Taubert in Engler & Prantl, Nat.
Pflanzenfam. 3 (3): 149. 1892 ≡ Pauletia macrostachya
(Benth.) A. Schmitz in Bull. Jard. Bot. Natl. Belg. 43: 391.
1973 – Type: Guyana, Essequibo, Schomburgk 71 (holotype K! [photo in RB! ex NY negative 1593]). [= Bauhinia
ungulata L. var. ungulata]
Bauhinia macrostachya var. obtusifolia Ducke in Arch. Jard.
Bot. Rio de Janeiro 3: 106. 1922 ≡ Bauhinia ungulata
var. obtusifolia (Ducke) Vaz in Rodriguésia 54(83): 127.
2003 – Type: Brazil, Pará, Alcobaça (currently Tucuruí
apud Egler, 1963), Tocantins river, Ducke Herb. Amaz.
Musaei Paraensis 16192 (lectotype RB 11146!, designed
by Vaz & Tozzi, 2003b: 127; isolectotype MG). Brazil,
Pará, Forte Ambé, Xingu river, Ducke 10411 (remaining
syntype MG); Pará, between Belém and Bragança, railway,
Peixe Boi Station, Siqueira s.n. (remaining syntype MG
8763!); idem, Santo Antonio do Prata, Goeldi (remaining
syntype MG 7295!).
Bauhinia macrostachya var. parvifolia Ducke in Arch. Jard.
Bot. Rio de Janeiro 3: 106. 1922 ≡ Bauhinia ungulata var.
parvifolia (Ducke) Vaz in Rodriguésia 54(83): 127. 2003.
– Type: Brazil, Pará, “in rupibus siccis humo vix obtectis
prope cataractum Itaboca”, Tocantins river, Ducke Herb.
Amaz. Musaei Paraensis 16232 (lectotype RB 11137!, designed by Vaz & Tozzi, 2003b: 127). Brazil, Maranhão,
Codó, Ducke Herb. Gener. Mus. Pará 583 (remaining syntype RB!); idem, Pedreiras, Lisboa s/n Herb. Mus. Goeldi
2307 (remaining syntypes, MG; isosyntype RB!).
Bauhinia macrostachya var. tenuifolia Ducke in Arch. Jard.
Bot. Rio de Janeiro 3: 106. 1922 – Type: Brazil, Pará,
Belém, Huber Herb. Amaz. Musaei Paraensis 2103 (lectotype RB 11132!, designated by Vaz & Tozzi, 2003b: 124;
isolectotype MG). Type: Brazil, Pará, Peixeboi, Bragança
railway, Siqueira (remaining syntype MG 8272!; isosyntype RB!); Pará, Almeirim, Ducke Herb. Amaz. Museu
Goeldi 17236 (remaining syntype MG!; isosyntype RB!).
[= Bauhinia ungulata L.]
1469
Vaz • Typifications of names of taxa of Bauhinia
Bauhinia malacotricha Harms in Bot. Jahrb. Syst. 33,
Beibl. 72: 22. 1903 – Type: Brazil, Goiás, Chapadão dos
Veadeiros, Glaziou 21016 (B, destroyed [photo in RB! ex
F negative 1595]; lectotype designated here, P!; isolectotype K).
Bauhinia malacotrichoides R.S. Cowan in Los Angeles County
Mus. Contr. Sci. 13: 8. 1957 – Type: Brazil, Goiás, 21 km
north of S. João da Aliança, region of the Chapada dos
Veadeiros, Dawson 14293 (holotype R! [photo in RB!]).
Bauhinia marginata (Bong.) Steud. ≡ Pauletia marginata
Bong.
Bauhinia membranacea Benth. in Martius, Fl. Bras. 15(2):
187. 1870 ≡ Pauletia membranacea (Benth.) A. Schmitz
in Bull. Jard. Bot. Natl. Belg. 43: 395. 1973 – Type: Brazil, Goiás: “in sylvis humidis ad radices Serra de Santa
Brida” Gardner 3695 (holotype K [photo in RB!]; isotypes
K [photo in RB!], OXF!).
Bauhinia microphylla Vogel in Linnaea 15(2): 301. 1839 – Type:
Brazil, “ad Faz. da Piedade”, Sellow 950 (B, destroyed
[photo in RB! ex F negative 1596]); lectotype, designated
here, K Herb. Hook. 1867 [photo in RB!]). [= Bauhinia
bauhinioides (Mart.) J.F. Macbr.]
Bauhinia mollis (Bong.) D. Dietr. ≡ Pauletia mollis Bong.
Bauhinia nitida Benth. in Martius, Fl. Bras. 15 (2): 184. 1870
≡ Pauletia nitida (Benth.) A. Schmitz in Bull. Jard. Bot.
Natl. Belg. 43: 395. 1973 – Type: Brazil, Tocantins, Natividade, “Dry wooded campos”, Gardner 3122 (B, destroyed
[photo in RB!]); lectotype, designated here, K; isolectotypes BM!, OXF!). Brazil, Tocantins, “ad Porto Imperial”
[Porto Real in sched., currently Porto Nacional], Burchell
Cat. Geogr. Pl. Bras. Tropicae 8398 (remaining syntype
GRAH [photo in RB! ex K negative 16415]). [= Bauhinia
dubia G. Don]
Wunderlin annotated Gardner 3122 (K) as a lectotype of
B. nitida here accepted.
Bauhinia obtusata Vogel in Linnaea 13: 305. 1839 ≡ Pauletia
obtusata (Vogel) A. Schmitz in Bull. Jard. Bot. Natl. Belg.
43: 391. 1973 – Type: Brazil, Minas Gerais, “pr. Marianna”,
Sellow 5428 (B, destroyed [photo in RB!], fragment F!
[photo in RB! ex F negative 1600]). [= Bauhinia longifolia
(Bong.) Steud.]
Bauhinia odoratissima Moric., Pl. Nouv. Amér.: 78. 1840, non
Newm. ex G. Don, 1832 ≡ Pauletia odoratissima (Moric.)
A. Schmitz in Bull. Jard. Bot. Natl. Belg. 43: 395. 1973
– Type: Brazil, Bahia, Ilhéus, Blanchet 2387 (lectotype,
designated here, K [photo in RB! ex NY negative 1599];
isolectotypes, A!, BM [photo in RB!], OXF!, W!). Brazil,
Bahia, Nagé, Blanchet 2010 (remaining syntype, not located). [= Bauhinia integerrima Benth.]
1470
TAXON 60 (5) • October 2011: 1464–1474
Bauhinia olfersiana Vogel in Linnaea 13: 306. 1839 ≡ Bauhinia
cuyabensis var. olfersiana (Vogel) Kuntze in Revis. Gen. Pl.
3(2): 53. 1898 – Type not mentioned. According to Bentham
(1870), a synonym of Bauhinia cuyabensis (Bong.) Steud.
Bauhinia ovata (Bong.) Vogel ≡ Pauletia ovata Bong.
Bauhinia pentandra (Bong.) D. Dietr. ≡ Pauletia pentandra
Bong.
Bauhinia pinheroi Wunderlin in Phytologia 62: 89. 1987 –
Type: Brazil, Bahia, km 25 on the Guaratinga–São Paulinho road, Pinheiro 2081 (holotype CEPEC!; isotypes
K, US); Brazil, Bahia, Fazenda Pau-Brazil, ca. 5 km NW
of Itamaraju, Mattos Silva & al. 529 (paratype CEPEC!;
duplicate RB! [photo in RB!]).
Bauhinia platypetala Burch. ex Benth. in Martius, Fl. Bras.
15(2): 198. 1870 ≡ Bauhinia forficata var. platypetala
(Burch. ex Benth.) Wunderlin in Ann. Missouri Bot.
Gard. 60(2): 571. 1973 ≡ Pauletia platypetala (Burch. ex
Benth.) A. Schmitz in Bull. Jard. Bot. Natl. Belg. 43: 396.
1973 – Type: Brazil, Tocantins, Porto Real (currently Porto
Nacional), Burchell Cat. Geogr. Pl. Bras. Tropicae 8563
(lectotype, designated here, K Herb. Hook. 1867 [photo
in RB!, ex NY negative 1602]). Brazil, Goiás, “in sylvis
prope Natividade et Conceição”, Gardner 3118 (remaining
syntype OXF! [isosyntype photo in RB!]); idem, idem,
Gardner 3700 (remaining syntype OXF! [isosyntype photo
in RB!]).
Bauhinia platyphylla Benth. in Martius, Fl. Bras. 15(2): 185.
1870, non Zipp. ex Span. in Linnaea 15: 201. 1841 – Type:
Brazil, Goiás, “inter Cavalcante et Conceição”, Burchell
Cat. Geogr. Pl. Bras. Tropicae 8086 (holotype K [photo
in RB! ex NY negative 1581]; isotype P [photo in RB! ex
F negative 39861]).
Bauhinia pruinosa Vogel in Linnaea 13: 301. 1839 ≡ Bauhinia
forficata subsp. pruinosa (Vogel) Fortunato & Wunderlin in Darwiniana 27: 550. 1987 (‘1986’) – Type: Brazil,
“Bras. merid.”, Sellow 2468 (B, destroyed [photo in RB!
ex F negative 1607!], F [fragment, fide Fortunato, 1987).
Bauhinia pulchella Benth. in Martius, Fl. Bras. 15(2): 190.
1870 ≡ Pauletia pulchella (Benth.) A. Schmitz in Bull.
Jard. Bot. Natl. Belg. 43: 391. 1973 – Type: Brazil, Piaui,
Oeiras, Gardner 2150 (lectotype, designated here, K!
[photo in RB!]; isolectotypes BM!, GRAH!, L!, MANCH!,
OXF!, W!). Brazil, Minas Gerais, “in campis Taboleira
ad Barreiras, Martius 1406 (remaining syntypes, M, two
sheets; H. Esser, pers. comm.); Minas Gerais, “in campis
siccis ejusdem prov.: Riedel, Claussen, Pohl ” (remaining
syntype, not located); Bahia, “in inundatis sylvis, prope
Joazeiro ad Rio S. Francisco”, Martius s.n. (remaining syntypes, M, three specimens; H. Esser, pers. comm.); Bahia,
Serra Açurua, Blanchet 2897 (remaining syntype BM!).
TAXON 60 (5) • October 2011: 1464–1474
Bauhinia pulchella var. parvifolia Benth. in Martius, Fl. Bras
15(2): 190. 1870 – Type: Brazil. Minas Gerais, Minas
Novas, Martius s.n. (holotype M). [=Bauhinia pulchella]
Bauhinia recurva R.S. Cowan in Los Angeles County Mus.
Contr. Sci. 13: 10. 1957 – Type: Brazil, Goiás, São João
da Aliança, region of the Chapada dos Veadeiros, highway north of São João da Aliança km 19–19,5; 47°30′ W,
14°30′ S, Dawson 14387 (holotype R!; isotype NY!). [=
Bauhinia longifolia (Bong.) Steud.]
Bauhinia rhodacantha Desv. in Ann. Sci. Nat. (Paris), ser. 1, 9:
429. 1826 – Type: “Hab. in Brasilia” (not located, P?). The
original diagnosis is too unclear to determine the species.
Bauhinia rufa (Bong.) Steud. ≡ Pauletia rufa Bong.
Bauhinia rufa var. cordata (Vogel) Benth. ≡ Bauhinia cordata
Vogel
Bauhinia rufa var. dodecandra (Bong.) Benth. ≡ Bauhinia dodecandra Bong.
Bauhinia rufa var. intermedia (Vogel) Benth. ≡ Bauhinia intermedia Vogel
Bauhinia selloviana Vogel in Linnaea 13: 306. 1839 – Type:
Brazil, Minas Gerais, “pr. Marianna”, Sellow 1902 (B, destroyed [photo in RB! ex F negative 1575]). [= Bauhinia
fusconervis]
Bauhinia smilacifolia Burch. ex Benth. in Martius, Fl. Bras.
15(2): 183. 1870 – Type: Brazil, Goiás, between Cavalcante
and Conceição, “in campis elevatis petrosis”, Burchell Cat.
Geogr. Pl. Bras. Tropicae 8076 (lectotype, designated
here, K [photo in RB! ex NY negative 1579]). Brazil, Tocantins, “prope Arrayas”, Gardner 3698 (remaining syntype K Herb. Hook. [photo in RB! ex Kew negative 16425];
isosyntypes BM!, E!, MANCH!, OXF!).
Wunderlin annotated Burchell 8076 (K) as a lectotype of
B. smilacifolia, a choice that is effectively published here.
Bauhinia stenocardia Standl. in Trop. Woods 33: 12. 1933 –
Type: Brazil, Pará, Aramanahy, lower Tapajóz, Monteiro
da Costa 238 (holotype F; isotype RB!). [= Bauhinia longicuspis Spruce ex Benth.]
Bauhinia straussiana Harms in Notizbl. Königl. Bot. Gart.
Berlin 6(59): 308 1915 – Type: “Bolivia: Gebiet des Alto
Acre, Cobija, Jan. 1912, Ule 9442” (holotype B, destroyed
[photos in RB!, SI! ex F negative 1617]). [ = Bauhinia
conwayi Rusby]
Bauhinia subclavata Benth. in Martius, Fl. Bras. 15(2): 188.
1870 ≡ Pauletia subclavata (Benth.) A. Schmitz in Bull.
Jard. Bot. Natl. Belg. 43: 396. 1973 – Type: Brazil, Piauí,
between the Camindé and Oeiras rivers, Gardner 2154
Vaz • Typifications of names of taxa of Bauhinia
(holotype K, barcode K000056822 [photo in RB!]; isotypes B, destroyed [photo in RB! ex F negative 1623], K
[photo in RB! ex K negative 16427], NY! [photo in RB!
ex NY negative 12714], OXF!, P!).
Bauhinia tarapotensis Benth. in Martius, Fl. Bras. 15(2):
198. 1870 ≡ Pauletia tarapotensis (Benth.) A. Schmitz in
Bull. Jard. Bot. Natl. Belg. 43: 396. 1973 – Type: Peru,
San Martin, Tarapoto, Spruce 4417 (holotype K, 2 sheets,
flower and fruit [photos in RB!]; isotypes E!, GRAH!,
NY!, OXF!, W!).
Bauhinia tenella Benth. in Martius, Fl. Bras. 15(2): 195. 1870
≡ Pauletia tenella (Benth.) A. Schmitz in Bull. Jard. Bot.
Natl. Belg. 43: 396. 1973 – Type: Brazil, Tocantins, Arraias, Gardner 3701 (lectotype designated here, K; isolectotypes, E!, K [2 sheets, photos in RB! ex K], MANCH!,
OXF!, P!, W!). Brazil, Goiás, Cavalcante-Conceição,
Burchell Cat. Geogr. Pl. Bras. Tropicae 8000 (remaining
syntype: K, photo in RB!); Brazil, idem, Pohl s.n. (not
located).
Wunderlin annotated Burchell 8000 (K) as lectotype of
B. tenella, a choice that is effectively published here.
Bauhinia tumupasensis Rusby in Bull. New York Bot. Gard.
28(8): 93. 1912 – Holotype: Bolivia, Tumupasa, Willams
495 (NY!; isotype BM!). [= Bauhinia conwayi Rusby]
Bauhinia ungulata L., Sp. Pl.: 374. 1753 ≡ Pauletia ungulata
(L.) A. Schmitz in Bull. Jard. Bot. Natl. Belg. 43: 393. 1973
– Type: “Habitat in America.” Herb. Clifford 157, Bauhinia
no. 3 (lectotype BM, designated by Wunderlin in Turland
& Jarvis, 1997: 466).
Bauhinia ungulata var. cuyabensis (Bong.) Vaz (‘cuiabensis’)
≡ Pauletia cuyabensis Bong.
Bauhinia ungulata var. obtusifolia (Ducke) Vaz ≡ Bauhinia
macrostachya var. obtusifolia Ducke
Bauhinia ungulata var. parvifolia (Ducke) Vaz ≡ Bauhinia
macrostachya var. parvifolia Ducke
Bauhinia urocalyx Harms in Notizbl. Königl. Bot. Gart. Berlin
6(59): 308. 1915 – Type: Brazil, Acre, “Lago, Jurua-Miry
[sic], Rio Juruá Sup.”, Ule 5542 (B, destroyed [photo in RB!
ex F negative 1625]; lectotype, designated here, HBG!).
The type locality Juruá-Mirim river is at Porto Walter municipality (D. Daly, pers. comm.).
Bauhinia vespertilio S. Moore in Trans. Linn. Soc. London,
Bot. ser 2, 4: 347. 1895 – Type: Brazil, Mato Grosso, Santa
Cruz, S. Moore iter matogrossense 587 (holotype BM;
isotype NY! [photo in RB! ex NY negative 12713]).
Bauhinia viridiflora Ducke in Arch. Jard. Bot. Rio de Janeiro
3: 105. 1922, non Backer in Bull. Jard. Bot. Buitenzorg,
1471
Vaz • Typifications of names of taxa of Bauhinia
ser. 3, 2: 323. 1920 ≡ Bauhinia viridiflorens Ducke in Arch.
Jard. Bot. Rio de Janeiro 5: 129. 1930 – Type: Brazil, Pará,
Alcobaça Railway, Breu Branco Station, Tocantins river,
Ducke Herb. Amaz. Musaei Paraensis 15597 (lectotype,
designated here, MG!; isolectotypes RB 11121!, R!). Brazil, “prope Santo Antonio do Prata, inter Belém et Bragança”, Snethlage s.n. (remaining syntype, not located).
[= Bauhinia dubia G. Don]
Bauhinia viscidula Harms in Bot. Jahrb. Syst. 33 Beibl. 72: 22.
1903 ≡ Pauletia viscidula (Harms) A. Schmitz in Bull. Jard.
Bot. Natl. Belg. 43: 393. 1973 ≡ Bauhinia dumosa var. viscidula (Harms) Vaz & R. Marquete in Revista Brasil. Bot.
16(1): 97. 1993 – Type: Brazil, Goiás, between “Ciganos and
As Brancas”, Glaziou 21010 (lectotype RB!, designated by
Vaz & Marquete, 1993: 105; isolectotype K!). Remaining
syntype: Brazil, Goiás, Glaziou 21012ª (K! [photo in RB!]).
Pauletia bicolor Bong. in Mém. Acad. Imp. Sci. Saint-Pétersbourg, Sér. 6, Sci. Math. 4, Seconde Pt. Sci. Nat. 2: 117.
tab. 6. 1838 ≡ Bauhinia bicolor (Bong.) Steud., Nomencl.
Bot., ed. 2, 1: 191 (‘291’). early Aug. 1840 [“B. bicolor
(Bong.) Walpers”, Repert. 1: 848. 1842, later isonym] –
Type locality: “In campis siccis Brasiliae. Flor. Octobri”
(not located, LE?).
Pauletia cheilantha Bong. in Mém. Acad. Imp. Sci. SaintPétersbourg, Sér. 6, Sci. Math. 4, Seconde Pt. Sci. Nat. 2:
120. 1838 ≡ Bauhinia cheilantha (Bong.) Steud., Nomencl.
Bot., ed. 2, 1: 191 (‘291’). early Aug. 1840 [“B. cheilantha
(Bong.) D. Dietr.”, Syn. Pl. 2: 1476. Dec.1840; “B. cheilantha (Bong.) Walpers”, Repert. 1: 848. 1842, later isonyms]
– Type: Brazil, Mato Grosso, “in fruticetis prope Cuyabá”
[Cuiabá], collector not cited, Riedel 773 (lectotype, designated here, LE [photo in BAB!, RB!]).
Pauletia cuyabensis Bong. in Mém. Acad. Imp. Sci. SaintPétersbourg, Sér. 6, Sci. Math. 4, Seconde Pt. Sci. Nat. 2:
125. 1838 ≡ Bauhinia cuyabensis (Bong.) Steud., Nomencl.
Bot., ed. 2, 1: 191 (‘291’). early Aug. 1840 [“B. cuyabensis
(Bong.) D. Dietr.”, Syn. Pl. 2: 1476. Dec. 1840; “B. cuyabensis (Bong.) Walpers”, Repert. 1: 848. 1842, later isonyms] ≡ Bauhinia ungulata var. cuyabensis (Bong.) Vaz
in Rodriguésia 54: 125. 2003 (‘cuiabensis’) – Type: Brazil,
Mato Grosso, “in fruticetis humidis prope Cuyabá et in
montibus Chapada Cuiabá dictis, floret februario”, Riedel
829 (lectotype, designated here, LE [photos in BAB! and
RB!]; isolectotype P!).
Pauletia dodecandra Bong. in Mém. Acad. Imp. Sci. SaintPétersbourg, Sér. 6, Sci. Math. 4, Seconde Pt. Sci. Nat.
2: 129. 1838 ≡ Bauhinia dodecandra (Bong.) Steud., Nomencl. Bot., ed. 2, 1: 191 (‘291’). early Aug. 1840 – Type:
Brazil, Mato Grosso “in campis siccis Camapuensibus”
[Camapuã], “floret octobri”, Riedel s.n. (lectotype, designated here, LE [photo in RB! ex BAB]). [= Bauhinia
holophylla (Bong.) Steud.]
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TAXON 60 (5) • October 2011: 1464–1474
Pauletia ferruginea Bong., in Mém. Acad. Imp. Sci. SaintPétersbourg, Sér. 6, Sci. Math. 4, Seconde Pt. Sci. Nat. 2:
119. 1838 ≡ Bauhinia ferruginea (Bong.) D. Dietr., Syn. Pl.
2: 1476. Dec. 1840 (non Roxb. 1832) ≡ Bauhinia bongardii
Steud., Nomencl. Bot., ed. 2, 1: 191 (‘291’). early Aug. 1840
≡ Pauletia bongardii (Steud.) A. Schmitz in Bull. Jard. Bot.
Natl. Belg. 43(3–4): 392. 1973, nom. superfl. – Type locality: Brazil, “in sterilibus prope Cuyaba. Flor. februario”
(LE?). [= Bauhinia brevipes Vogel]
Pauletia fusconervis Bong. in Mém. Acad. Imp. Sci. SaintPétersbourg, Sér. 6, Sci. Math. 4, Seconde Pt. Sci. Nat. 2:
116. 1838 (‘fusco-nervis’) ≡ Bauhinia fusconervis (Bong.)
Steud., Nomencl. Bot., ed. 2, 1(1/2): 191 (‘291’) early Aug.
1840 [“B. fusconervis (Bong.) D. Dietr., Syn. Pl. 2: 1476.
Dec. 1840; “B. fusconervis (Bong.) Walpers”, Repert.
1: 848. 1842, later isonyms] – Type: Brazil, “in sylvis
Morro Grande, inter Parahyba et Parahybuna”, May 1829,
Langsdorff s.n. (lectotype, designated here, LE [photos
in BAB!, RB!]).
Pauletia grandifolia Bong in Mém. Acad. Imp. Sci. Saint-Pétersbourg, Sér. 6, Sci. Math. 4, Seconde Pt. Sci. Nat. 2: 130.
1838 ≡ Bauhinia grandifolia (Bong.) Steud., Nomencl.
Bot., ed. 2, 1: 191 (‘291’). early Aug. 1840.– Type: Brazil,
Amazonas, “in fruticetis siccis prope Borba”, Jul. 1828,
Riedel 1293 (lectotype, designated here, LE (2 sheets),
[photos in BAB!, RB!]).
Pauletia hirsuta Bong. in Mém. Acad. Imp. Sci. Saint-Pétersbourg, Sér. 6, Sci. Math. 4, Seconde Pt. Sci. Nat. 2: 134. 1838
≡ Bauhinia hirsuta (Bong.) Vogel, non Weinm.≡ Bauhinia
hirsutiflora Vaz in Atas Soc. Bot. Brasil, Secc. Rio de Janeiro 3(20): 3. 1995, nom. illeg. – Type: Brazil, Mato Grosso,
“prope Cuyabá”, Riedel 825 (B, destroyed [photo in RB! ex
F negative 1588]). [= Bauhinia vespertilio]
Pauletia holophylla Bong. in Mém. Acad. Imp. Sci. SaintPétersbourg, Sér. 6, Sci. Math. 4, Seconde Pt. Sci. Nat. 2:
121. 1838 ≡ Bauhinia holophylla (Bong.) Steud., Nomencl.
Bot., ed. 2, 1: 191 (‘291’). early Aug. 1840 – Type: Brazil,
Mato Grosso, “Hab. in sylvis siccis Serrado dictis prope
Camapua”, Nov. 1826, Riedel s.n. (LE, G!, OXF!).
Pauletia longiflora Bong. in Mém. Acad. Imp. Sci. Saint-Pétersbourg, Sér. 6, Sci. Math. 4, Seconde Pt. Sci. Nat. 2: 128.
1838 ≡ Bauhinia longiflora (Bong.) Steud., Nomencl. Bot.,
ed. 2, 1: 191 (‘291’). early Aug. 1840 ≡ Bauhinia forficata
var. longiflora (Bong.) Benth. in Martius, Fl. Bras. 15(2):
201. 1870 ≡ Pauletia forficata var. longiflora (D. Dietr)
A. Schmitz in Bull. Jard. Bot. Natl. Belg. 43: 390. 1973 –
Type: Brazil. Rio de Janeiro, Bongard s.n. (holotype LE,
fide Fortunato, 1996: 408). [= Bauhinia forficata Link
subsp. forficata].
Pauletia longifolia Bong. in Mém. Acad. Imp. Sci. Saint-Pétersbourg, Sér. 6, Sci. Math. 4, Seconde Pt. Sci. Nat. 2: 122.
TAXON 60 (5) • October 2011: 1464–1474
1838 ≡ Bauhinia longifolia (Bong.) Steud., Nomencl. Bot.,
ed. 2, 1: 191 (‘291’). early Aug. 1840 [“B. longifolia (Bong.)
D. Dietr., Syn. Pl. 2: 1476. Dec. 1840; “B. longifolia Walpers”, Repert. 1: 848. 1842, later isonyms] – Type: Brazil,
Mato Grosso, “habitat in humidiusculis Cuyabá”, collector
not mentioned (LE).
Vaz • Typifications of names of taxa of Bauhinia
bauhinioides (Mart.) A. Schmitz in Bull. Jard. Bot. Natl.
Belg. 43: 388. 1973 – Type: Brazil, Minas Gerais, “in Alagadisso, San Francisco, Martius s.n. (syntypes, M [2 sheets
no. 1284 and no. 1285], fide Fortunato, 1987: 543, 544).
ACKNOWLEDGEMENTS
Pauletia marginata Bong. in Mém. Acad. Imp. Sci. SaintPétersbourg, Sér. 6, Sci. Math. 4, Seconde Pt. Sci. Nat. 2:
121. 1838 ≡ Bauhinia marginata (Bong.) Steud., Nomencl.
Bot., ed. 2, 1: 191 (‘291’). early Aug. 1840 [“B. marginata
(Bong.) D. Dietr.”, Syn. Pl. 2 1475. Dec. 1840; “B. marginata Walpers”, Repert. 1: 848. 1842, later isonyms] – Type
locality: Brazil, São Paulo, “in campis siccis Rio Pardo”,
collector not mentioned (LE).
Pauletia mollis Bong. in Mém. Acad. Imp. Sci. Saint-Pétersbourg, Sér. 6, Sci. Math. 4, Seconde Pt. Sci. Nat. 2: 133.
1838 ≡ Bauhinia mollis (Bong.) D. Dietr., Syn. Pl. 2: 1475.
1840 – Type locality: “In petrosis siccis prope Cuyabá et
Camapua”. Brazil, Mato Grosso, Cuiabá, Bongard 786,
I-1827 (syntype LE, fide Fortunato, 1984); Camapuã, Bongard 620, X-1826 (syntype LE, fide Fortunato, 1984).
The author would like to thank G.P. Lewis (Royal Botanic Gardens, Kew) for reviewing an early version of this manuscript. Also I
am in debt to Dan H. Nicolson (Smithsonian Institution, Washington,
D.C.), John McNeill (Royal Botanic Garden, Edinburgh), Jefferson
Prado (Instituto de Botânica, São Paulo, SP), Hans-Joachim Esser
(Botanische Staatssammlung München, Munich), Mats Hjertson
(Museum of Evolution, Uppsala University), and an anonymous reviewer for their corrections and criticism. I acknowledge Ana Tozzi
(University of Campinas, São Paulo state); Renée Fortunato (INTA,
Castelar, Buenos Aires) and Vidal F. Mansano (Rio de Janeiro Botanical Garden) for suggestions. The author thanks the curators of
the consulted herbaria and specially BM, F, K, and UPS for sending
photographs of specimens.
LITERATURE CITED
Pauletia ovata Bong. in Mém. Acad. Imp. Sci. Saint-Pétersbourg, Sér. 6, Sci. Math. 4, Seconde Pt. Sci. Nat. 2: 132.
1838 ≡ Bauhinia ovata (Bong.) Vogel in Linnaea 13: 300.
1839 [“B. ovata (Bong.) D. Dietr.”, Syn. Pl. 2: 1475. Dec.
1840; “B. ovata Walpers”, Repert. 1: 849. 1842, later isonyms] – Type locality: “Hab. in Brasília”.
Bentham (1870: 197) examined a collection of Langsdorff
that probably is the original material of P. ovata (not located).
The material cited by Vogel (1839): “Sellow, leg. inter Campos
et Victoria” (photo in RB!) is not part of the original material,
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Pauletia pentandra Bong. in Mém. Acad. Imp. Sci. Saint-Pétersbourg, Sér. 6, Sci. Math. 4, Seconde Pt. Sci. Nat. 2: 126.
1838 ≡ Bauhinia pentandra (Bong.) D. Dietr., Syn. Pl. 2:
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1: 848. 1842, later isonym] – Type locality: Brazil, Mato
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Angela Maria Studart Da Fonseca Vaz