LAKE NYASA
CLIMATIC REGION
FLORISTIC CHECKLIST
Roy E. Gereau,1 Asukile R. Kajuni,2
Tim R. B. Davenport,3 and
Henry J. Ndangalasi4
ABSTRACT
The Lake Nyasa Climatic Region described in this study is composed of the Livingstone, Kipengere, Mbeya, and
Umalila ranges, Mporoto Ridge, Mt. Rungwe, and the Kitulo Plateau. The region is located on the rift escarpment at the
northeast end of Lake Nyasa. Floristic studies in the area reveal many unique plants, some of which are endemic to this
region. The climate of the region is governed by altitude and its proximity to Lake Nyasa, which results in generation
of high convectional rainfall. The high rainfall and diversity of habitats interacting with soils and other environmental
factors in the area are responsible for the richness of the flora, and current identifications of vascular plant taxa have
yielded a total of 170 families, 733 genera, and 1735 species. The Lake Nyasa Climatic Region, therefore, has nearly
14% of the estimated total of 12,500 vascular plant species in all of Tropical East Africa (H. Beentje, pers. comm.). The
largest families in the flora are Fabaceae (s. str. excluding Caesalpiniaceae and Mimosaceae; 156 species in 42 genera),
Asteraceae (139/55), Poaceae (124/50), Rubiaceae (119/49), and Orchidaceae (97/28); the largest genera are Crotalaria L. (Fabaceae, 30 species), Indigofera L. (Fabaceae, 22 species), Vernonia Schreb. (Asteraceae, 21 species), Cyperus
L. (Cyperaceae, 21 species), Habenaria Willd. (Orchidaceae, 18 species), and Plectranthus L’Hér. (Lamiaceae, 18 species). Taxa here reported as new to East Africa are Cephalanthus natalensis Oliv. (Rubiaceae), Cerastium dregeanum
Fenzl (Caryophyllaceae), Cynorkis symoensii Geerinck & Tournay (Orchidaceae), Dianthus zeyheri Sond. subsp. zeyheri
(Caryophyllaceae), Panicum inaequilatum Stapf & C. E. Hubb. (Poaceae), Rhynchosia caribaea (Jacq.) DC. var. caribaea (Fabaceae), Tricalysia longituba De Wild. var. longituba (Rubiaceae), Tricalysia ngalaensis Robbr. (Rubiaceae), and
Triumfetta angolensis Sprague & Hutch. (Tiliaceae); new to Tanzania from elsewhere in East Africa are Erica kingaensis
Engl. subsp. bequaertii (De Wild.) R. Ross (Ericaceae), Peperomia fernandopoiana C. DC. var. butaguensis (De Wild.)
Düll (Piperaceae), and Raphidiocystis jeffreyana A. Fern. & R. Fern. (Cucurbitaceae).
The Lake Nyasa Climatic Region is located in
the Iringa and Mbeya regions of Tanzania (frontispiece), and covers approximately 1200 km2
(120,000 hectares) between latitude 8°50' and
9°32'S and longitude 33°15' and 34°09'E (Fig. 1).
It extends to the southwest from the Eastern Arc
Mountains of East Africa (Fig. 2), which are characterized by small and fragmented mountain blocs
each having a patchwork of dense tropical forests
and montane vegetation. It encompasses Kitulo
National Park, Mpanga/Kipengere Game Reserve,
and the following gazetted forest reserves: Mt.
Rungwe, Chimala Scarp, Umalila, Sawago, Mporoto Ridge, Irenga, Irungu, Chuvwi, Masukulu,
Kabulo, Kyosa, Mswima, Iyondo, Mpara, Mbeya
Range, North Usafwa, Ileje Range, and Kyejo. The
Lake Nyasa Climatic Region lies on the junction
of the eastern and western arms of the Great Rift
Valley of Africa, which runs from Mozambique
through Malawi, Tanzania, Kenya, Ethiopia, and
Eritrea into Saudi Arabia.
A closer scrutiny of the study area reveals
many similarities to the Eastern Arc Mountains,
which include the Udzungwa, Uluguru, Mahenge,
Malundwe, Rubeho, Ukaguru, Nguru, Nguu, East
Usambara, West Usambara, South Pare, and North
Pare mountains in Tanzania, and the Taita Hills in
Kenya (Platts et al., 2011). These discontinuous
blocs of mountains described as “Galapagos Islands
of Africa” have recently been a taxonomist’s paradise, and new discoveries are being made regularly.
Although our study area forms a southern extension of this ecologically important zone (Burgess
et al., 2004), it has only recently started receiving
attention. It is possible that the climatic influences associated with altitude and Lake Nyasa are
responsible for the botanical differences between
this area and the Eastern Arc Mountains (Fig. 2).
However, recent and ongoing work by the Wildlife
Conservation Society (Davenport, 2004, 2005,
2006; Jones et al., 2005; Davenport et al., 2006)
has shown that this separation is not as marked
1
Missouri Botanical Garden, P.O. Box 299, St. Louis, Missouri 63166–0299, U.S.A. roy.gereau@mobot.org.
U.S. Agency for International Development, P.O. Box 9130, Dar es Salaam, Tanzania. akajuni@usaid.gov.
3
Wildlife Conservation Society, P.O. Box 922, Zanzibar, Tanzania. tdavenport@wcs.org.
4
Botany Department, University of Dar es Salaam, P.O. Box 35060, Dar es Salaam, Tanzania. hjndangalasi@udsm.
ac.tz; hjndangalasi@yahoo.com.
2
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Gereau et al.
Figure 1. Map of the study area: The Lake Nyasa Climatic Region of southwestern Tanzania, with selected localities
indicated. Lower contour line at 1500 m above sea level; upper contour line at 2000 m; lightly stippled area, land at
1500–2000 m; darkly stippled area, land above 2000 m; triple-dotted line, border between Tanzania (on north and east,
above and to right) and Malawi (on south and west, below and to left).
zoologically as botanically. From the Makambako
Gap the study area runs southwest to the northern
tip of Lake Nyasa and forms a continuous range
of high-altitude mountain ranges that divide the
water catchment area into two drainage patterns:
the northbound flowing into the Indian Ocean and
the southbound flowing into Lake Nyasa. Several
rivers and streams fed by numerous tributaries
drain the flanks of the ridge, including Ruaha,
Rumakali, Kiwira, Lufirio, Mbaka, Chimala,
Kimani, Songwe, and Mbarali.
Elton and Cotteril (1879) wrote the first account
of the floristic richness of the Kitulo Plateau, originally known as the Elton Plateau. Thereafter in the
20th century, several workers made attempts to
describe some major resources including vascular
plants from part of the areas surrounding the plateau. However, interest in the Kitulo Plateau was
particularly intense due to the uniqueness of its
LakeNyasa_5-18.indd 2
resources and to the area’s great potential for intensive agricultural development, sheep and dairy
farming. The following studies have been particularly useful in providing insights into the natural
resources of the Kitulo Plateau and surrounding
areas: Gillman (1927), Greenway (1955), Hedberg
(1957), Kerfoot (1964), Napper (1965), Prins and
Prins-Lampert (1974), Cribb and Leedal (1982),
TISCO (1983), Lovett et al. (1994), Lovett and
Prins (1994), Davenport (2002, 2006), Davenport
and Ndangalasi (2003), Davenport and Bytebier
(2004), and Davenport et al. (2006, 2008).
The Lake Nyasa Climatic Region is of great
ecological and phytogeographic importance for a
number of reasons. The Indian Ocean drainage
from this area is the source of the Ruaha, Mbarali,
and Chimala rivers and numerous other streams
that feed and provide constant recharge to the
Usangu wetlands and floodplains that are impor-
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Lake Nyasa Climatic Region
Floristic Checklist
3
Figure 2. The Neogene Volcanics, Eastern Arc Mountains, and Lake Nyasa Climatic Region. Shaded areas between
dashed lines indicate the Eastern Arc Mountains, with the Neogene Volcanics to the north and the Lake Nyasa Climatic
Region to the southwest.
tant rice growing areas, producing over 60% of rice
sold in urban market stalls in the country (Kamukala & Crafter, 1993). Much of the Usangu Plains
have recently been gazetted as an extension of the
Ruaha National Park and provide important refuge
and grazing grounds for wildlife; the Usangu wetlands feed into the Great Ruaha River, whose waters
are an important source of hydroelectric power,
supplying over 60% of all electricity consumed in
the country through the national grid from dams
LakeNyasa_5-18.indd 3
impounded at Mtera and Kidatu (Power et al.,
1997; NORPLAN, 2002). The irrigation schemes
for the Kilombero sugar plantations and rice farms
are dependent on waters from the Ruaha River,
and the whole Rufiji Delta with its rich marine and
other aquatic resources depends on water resources
from the Southern Highlands. The southern drainage from the region is the source of the Rumakali,
Songwe, Kiwira, Mbaka, and Lufirio rivers, which
water Rungwe District with its important cash
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Figure 3. Average monthly precipitation for Kitulo Dairy Farm (1965–1994). Source: TISCO (1983), with additional
data obtained from Kitulo Dairy Farm Meteorological Station in 1999.
crops of coffee, tea, avocados, and bananas; Kyela
District with its high-quality and fragrant rice,
abundant fruits, and cocoa; and Lake Nyasa with
its high diversity of cichlid fish. The Kitulo Plateau has been described as harboring “one of the
great floral spectacles of the world,” where rolling
montane grassland and patches of forest provide
a unique environment and unparalleled botanical
display with a plethora of globally significant flora
and fauna (Davenport, 2002; Davenport & Bytebier, 2004). Management of natural and physical
resources requires knowledge about the biology,
ecology, and their interrelationship in space and
time. Information about these resources is essential
in order to guide decision-making processes about
appropriate land uses and how best to manage and
preserve them (Mwakilema & Davenport, 2005).
Therefore, the Wildlife Conservation Society is
working to understand better the flora, fauna,
invertebrates, and other associated resources found
in the region (Davenport, 2004, 2006), which is so
critical at this point as a result of increased human
populations and globalization of the economy. The
open market economy has provided incentives
for people to increase their agricultural produce
through cultivating more land and encroaching on
areas that originally were reserved for conservation
purposes. Uncontrolled cultivation for agricultural
LakeNyasa_5-18.indd 4
crops results in loss of an important genetic pool
from the original, indigenous flora due to change in
the natural vegetation structure and composition.
As more of the natural environment is converted
to agricultural land, it means a loss in indigenous
natural flora and fauna. The loss in indigenous
natural resources through extinction is tantamount
to a lost opportunity for science and human use, as
we cannot recreate the natural world.
CLIMATE OF STUDY AREA
Pratt and Gwynne (1977) described the climatic
conditions of the study area as having one single
season rainfall from November to May and a dry
season from June to October. The mean annual
rainfall is variable depending on altitude and
proximity to Lake Nyasa but lies between 1200
and 2950 mm. The number of rainy days ranges
between 117 in Kyela along Lake Nyasa and 159
days on the Kitulo Plateau (Atlas of Tanzania,
1967; TISCO, 1983). Long-term climatic data
have been collected and kept at Kitulo Dairy Farm,
which over a 30-year period show that December
to April receives the heaviest rainfall, averaging
231.9 mm per month, and June to September
receives the least amount, with average monthly
precipitation below 10 mm (Fig. 3).
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Lake Nyasa Climatic Region
Floristic Checklist
5
Figure 4.
Mean monthly maximum and minimum and soil temperatures for Kitulo Dairy Farm (1965–1982).
Source: TISCO (1983).
The plateau has a relatively uniform cool, moist
climate, cool temperatures, low evapotranspiration
rates, few sunshine hours per day, and brisk winds;
the mean minimum and maximum temperatures
are 3.1°C and 16.5°C, respectively. The soil temperatures show similar trends, with both air and
soil temperatures being coolest in July and warmest
in October (Fig. 4). During the cool season (June,
July, and August), the air temperatures frequently
dip below freezing and ground frosts occur almost
every night. The lowest daily mean minimum temperature recorded at Kitulo was –9.5°C in July of
1973. Mean grass minimum temperatures are normally below 0°C for the months of June through
September.
The maximum hours of sunshine and radiation
occur during the dry, cool season, at which time the
sky is relatively clear (Figs. 5 and 6). Solar radiation peaks in August, which is also the month of
maximum daily sunshine hours (7.5 hours). Both
hours of sunshine and radiation start to decrease in
September and reach their minimum in January.
The weather records from Kitulo generally
represent the climatic conditions obtained in
the highlands and high-altitude areas within the
Lake Nyasa Climatic Region. Kitulo Dairy Farm
has maintained uninterrupted long-term meteorological data collection that is quite reliable and
therefore used to illustrate the weather conditions
LakeNyasa_5-18.indd 5
found in the entire study area. However, localized
differences resulting from physiographic conditions
provide some exceptions to this general pattern. To
the southwest of Mt. Rungwe, for example, convectional rain from Lake Nyasa continues throughout
most of the year (Table 1).
GEOLOGY AND SOILS
Geologically, the rock formations in the study
area are derived from the Ubendian system of
stratigraphical units that have been aged in the
Late Archaean era about 2000 to 1600 million
years ago. Superimposed on the Ubendian Belt are
zones of intense recent Proterozoic deformations
(Early Proterozoic dates between 1600 million
years and 750 million years), which have obscured
and in some cases completely obliterated Late
Archaean structures. These structures were further
complicated by extensive continued faulting activity during the Karoo Rift episodes, where certain
areas are still seismically active (Harpum, 1970).
The Ubendian system is characterized by greater
amounts of metamorphosed igneous rocks and by
the presence of charnockitic rocks. For example,
the rock formation of the Livingstone Mountains
is of anorthositic composition with intrusive basic
and ultra-basic complexes. As an example of the
superimposition of more recent Proterozoic defor-
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Figure 5.
Mean daily sunshine (hours/day) for Kitulo Dairy Farm (1965–1982). Source: TISCO (1983).
Figure 6.
Mean solar radiation (gm cal/sq cm/day) for Kitulo Dairy Farm (1965–1982). Source: TISCO (1983).
LakeNyasa_5-18.indd 6
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Lake Nyasa Climatic Region
Floristic Checklist
7
Table 1. Maximum (Ma) and minimum (Mi) mean
monthly temperature (°C) and mean monthly rainfall
(R) in millimeters from southwest of Mt. Rungwe near
Syukula.
Ma
Mi
R
Jan
23
15
312
Feb
24
16
173
Mar
25
16
262
Apr
24
14
361
May
22
13
152
June
22
11
78
July
22
9
97
Aug
23
12
12
Sep
25
14
31
Oct
26
16
9
Nov
27
16
160
Dec
26
16
326
mations, the soils of Kitulo Plateau and much of
the adjacent areas in the region are derived from
volcanic pumice as a result of volcanic eruptions
from Rungwe and Ngosi volcanos and are well
drained. The plateau is composed of Pliocene
basaltic and phonolitic lavas to an average depth of
150 m, which overlay pre-Karoo formations (Harkin, 1960). The soils range from humic ferrallitic
on various parent materials to humic ferrisols of
high-altitude regions (Atlas of Tanzania, 1967). In
all, six principal soil types have been identified for
Kitulo and are related to physiographic factors as
shown in Table 2.
ECOLOGY
Much of the area of the Lake Nyasa Climatic
Region lies in eco-climatic zone II and a small area
in zone I (Pratt & Gwynne, 1977). Zone I is characterized by an Afroalpine climate governed mostly
by altitude and is dominated by Afroalpine moorland and montane grassland above the forest line,
while eco-climatic zone II comprises highlands with
a tropical climate. The Lake Nyasa Climatic Region
has the largest and one of the most important
Afromontane and Afroalpine grassland communities in the country (White, 1983). The land in this
eco-climatic zone is the center of forest industry,
although it also has large tracts under intensive
subsistence agriculture or under cultivation for
LakeNyasa_5-18.indd 7
cash crops like tea and coffee. The dominant vegetation includes moist forests and derived grassland
and bushland with or without glades. The indigenous montane and upper montane forests in these
areas are the sources of economically important
large forest trees including Ficalhoa laurifolia
Hiern (Theaceae), Juniperus procera Hochst. ex
Engl. (Cupressaceae), Ocotea usambarensis Engl.
(Lauraceae), Podocarpus latifolius (Thunb.) R.
Br. ex Mirb. (Podocarpaceae), and Pouteria adolfifriedericii (Engl.) A. Meeuse subsp. australis (J.
H. Hemsl.) L. Gaut. (Sapotaceae). There are also
significant areas of bamboo forest dominated by
Sinarundinaria alpina (K. Schum.) C. S. Chao &
Renvoize (Poaceae). Bushland or evergreen shrubland are the commonest derivatives where shifting
cultivation has been practiced or where forest has
been disturbed but not burned with sufficient regularity to maintain grassland.
COLLECTING HISTORY
James Elton, the British consul to Mozambique,
was probably the first botanical explorer of the
Lake Nyasa Climatic Region (Moffett, 1958); he
crossed the Kitulo Plateau in late October 1877 and
published the first account of its floristic composition (Elton & Cotteril, 1879). German botanists,
most notably W. Goetze and A. Stolz, carried
out most subsequent plant collection in the area
through the end of World War I; Engler and Goetze
(1902) included Mt. Rungwe in their work on the
montane vegetation of Tanzania. After World War
I, the area received relatively sporadic attention
from plant collectors (with the notable exception
of H. J. Schlieben in Njombe District) until Mary
Richards began her extensive collecting in the
Mbeya and Njombe highlands in 1956, continuing through much of the 1960s (Cribb & Leedal,
1982). Many of the plant collectors working in the
area from the 1970s until the present have been
associated with the Flora of Tropical East Africa
(see Beentje & Smith, 2001), the National Herbarium of Tanzania, the University of Dar es Salaam,
the Missouri Botanical Garden, or the Wildlife Conservation Society. Despite a relatively large number
of collections, floristic checklists and/or vegetation
descriptions have been published only for Mbeya
Peak (Kerfoot, 1964) and the Kitulo Plateau (Prins
& Prins-Lampert, 1974; Lovett et al., 1994).
CONSERVATION STATUS
Although much of the land area in the Lake Nyasa
Climatic Region is gazetted as Forest Reserves and
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Gereau et al.
Table 2.
Kitulo Plateau Soil Types in relation to physiographic regions. Source: TISCO (1983).
Soil type
Physiographic region found
Black organic loam underlain by pumice
Undulating to hilly plateau
Black organic loam over very dark grey loam
over pumice
Undulating to hilly plateau
Shallow, black organic loam over loamy pumice
over clay loam
Rolling to steeply dissected highlands
Black peaty loam over pumice
Flat to almost flat floodplains
Shallow black loam over clay over weathered
schistose bedrock
Hilly to steeply dissected highlands
Shallow, very dark brown stony loam over bedrock;
very dark grey or black loam over bedrock
Steeply dissected to mountainous
theoretically accorded a legal protection status,
there are serious problems that lead to continued
loss of indigenous montane forests. Continued
conversion to farmland and pine plantations and
uncontrolled human encroachment and cultivation
have completely altered the natural vegetation patterns. Some information is available on the extent
of loss of montane forest habitat from illegal harvesting, fuelwood collection, and logging (Machaga
et al., 2004, 2005), and this loss is also the subject
of ongoing research; it is obvious that thinning
and opening up of the forest has occurred in all
areas. Some forests that were once almost impenetrable have been opened up in the last 30 years
for potato and pyrethrum farming. The reduction
in the size of these once intact indigenous montane
forests poses serious threats to their capacity to
ensure a stable, high-quality supply and recharge
of water in the hydrological cycle. In addition, the
indigenous montane forests play other important
functions in the prevention of soil erosion, loss of
topsoil, and damage from landslides, as well as
providing valuable non-timber forest products and
medicines for thousands of people (Machaga et al.,
2004, 2005). Only the intact natural forest can
sustain the intricate processes of water movements
through vegetation and soil. The bryophyte and
lichen epiphytes on woody plants are particularly
important to ensure the interception and storage of
large amounts of rainwater that are released slowly
and recharge the water table. The Usangu Plain
and Kilombero agricultural irrigation schemes, the
Mtera and Kidatu hydroelectric power stations,
and the Rufiji Basin depend entirely on the water
LakeNyasa_5-18.indd 8
from the Lake Nyasa Climatic Region, as do the
Kyela rice schemes. Many long-term residents of
the surrounding districts of Rungwe, Mbeya Rural,
Mbeya Urban, Makete, Njombe, Ileje, and Kyela
have witnessed changing hydrologic patterns and
noted a reduction in the volume of water in most
perennial rivers, and many once perennial streams
have become intermittent.
Continued degradation of the area’s high watershed values and the presence of many endemic and
unique plants in the area have stimulated reconsideration of the protection status of many key
areas. Following the gazetting of Kitulo National
Park in 2004, it is now extremely important that
other areas of this region be accorded better protection due to the current lack of management
strategies and the increase in inappropriate land
use practices.
Resource use associated with the increase in the
human and livestock populations is often unsustainable, resulting in environmental degradation.
There is a growing awareness of the importance of
conservation, but progress takes time. The environmental management schemes in Makete and
Njombe districts have yielded positive results, but
are not practiced throughout the study area. However, in October 2002 the government, as a result
of increased pressure arising from an increase of
human activities resulting in unsustainable use
of resources in the Usangu Plains and the surrounding areas, announced the gazettement of
the Mpanga/Kipengere area as a protected area
(Game Reserve). The Game Reserve covers an area
of 1574.25 km2 in part of the study area abutting
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Lake Nyasa Climatic Region
Floristic Checklist
the three districts of Njombe, Makete, and Mbarali
(Staddon et al., 2003). The Mpanga/Kipengere
Game Reserve comprises very important watercatchment areas.
The Kitulo Plateau, lying between the rugged
peaks of the Kipengere, Uporoto, and Livingstone
mountains, was gazetted as a national park in 2004
(Davenport, 2002; Davenport & Bytebier, 2004) in
order to ensure the protection and preservation of
an area that supports the largest and most important montane grassland community in Tanzania
(Mwakilema & Davenport, 2005). Much of the
park is located at ca. 2600 m on well-watered
volcanic soils. However, the park also covers significant amounts of montane forest (Davenport et
al., 2006). Kitulo National Park is located in the
headwaters and forms the most important watersheds for the Great Ruaha River and is also well
known for its floral significance.
ORGANIZATION, FORMAT, AND NOTATION OF CHECKLIST
Families are listed alphabetically within the divisions Pteridophyta, Pinophyta (Gymnospermae),
and Magnoliophyta (Angiospermae). Within Pteridophyta and Pinophyta, family circumscriptions
follow Kubitzki (1990). Within Magnoliophyta,
family circumscriptions for Liliopsida (Monocotyledonae) follow Dahlgren (1985); those for
Magnoliopsida (Dicotyledonae) follow Cronquist
(1981), with the exception of Hebenstretia L.,
which is here included in the Scrophulariaceae.
Genera, species, and infraspecific taxa are listed
alphabetically within families. One or more species
appearing without specific epithets at the end of
genera (e.g., “Xysmalobium sp.” appearing twice
under Asclepiadaceae) are known to be different
from other species in the same genus cited in the
checklist but have not been positively identified,
and thus may be either new to science or disjunct
from another geographic area and not yet reported
from the Lake Nyasa Climatic Region.
A single reference is given to support the use of
each species (or infraspecific taxon) name in the
Lake Nyasa Climatic Region; this is not the original place of publication of the name unless that is
the principal literature used for specimen identification or the only literature available. Publications
other than major African floras are listed in full
in the Literature Cited. Selected major African
floras are cited using the following standard abbreviations: F.T.A. = Flora of Tropical Africa (Oliver,
1868–1877; Dyer, 1896–1913; Prain, 1916–1934;
Hill, 1937; see also Stafleu & Cowan, 1981);
F.T.E.A. = Flora of Tropical East Africa (Turrill &
Milne-Redhead, 1952–1958; Hubbard & Milne-
LakeNyasa_5-18.indd 9
9
Redhead, 1958–1966; Milne-Redhead & Polhill,
1966–1972; Polhill, 1973–1978, 1982–1997;
Beentje, 1998–2007, 2010; Beentje & Ghazanfar,
2005–2009; see also Beentje & Smith, 2001);
F.W.T.A. = Flora of West Tropical Africa, second
edition (Keay, 1954, 1958; Hepper, 1963–1972);
Fl. Zamb. = Flora Zambesiaca (Exell & Wild,
1960–1961; Exell et al., 1963, 1966; Brenan,
1970; Exell & Launert, 1970; Fernandes et al.,
1971; Launert, 1978–1989; Launert & Pope,
1989–1991; Pope, 1992–1995, 1996–2005; Pope
& Martins, 2005; Pope et al., 2006; Timberlake
et al., 2007; Timberlake & Martins, 2008–2010).
Elevational ranges are taken directly from herbarium labels; a single number indicates that only
one collection exists, or that only one label indicated
elevation. Abbreviations cited for mountain ranges
under “Distribution” in the checklist are as follows
(Kipengere and Umalila ranges not included due
to insufficient collections): KP = Kitulo Plateau;
MP = Mbeya Range (including Mbeya Peak and
Mt. Loleza); MR = Mporoto Ridge (including Ngozi
Crater); NL = Northern Livingstone Mountains
(i.e., north of Rumakali River); RM = Rungwe
Mountain; SL = Southern Livingstone Mountains
(i.e., south of Rumakali River).
A single representative specimen is cited for
each species or infraspecific taxon unless the type
specimen was collected within the Lake Nyasa Climatic Region, in which case the type and one other
specimen (if any) are cited. Herbaria of deposit
are indicated for type specimens. Most other cited
specimens are deposited in DSM, EA, K, MO, or
NHT.
LAKE NYASA CLIMATIC REGION
FLORISTIC CHECKLIST
PTERIDOPHYTA
ASPLENIACEAE
1 genus and 9 species
Asplenium abyssinicum Fée; Beentje, F.T.E.A.
(Aspleniaceae): 62. 2008.
Terrestrial herb. Montane forest. 1950–2400
m.
Distribution: MR—Gereau 4620
Asplenium aethiopicum (Burm. f.) Bech.; Beentje,
F.T.E.A. (Aspleniaceae): 58. 2008.
Epiphytic or epilithic herb. Montane forest.
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10
1920–2400 m.
Distribution: KP, MR, NL, RM, SL—Gereau
3727
Asplenium anisophyllum Kunze; Beentje, F.T.E.A.
(Aspleniaceae): 36. 2008.
Epiphytic or epilithic herb. Montane forest.
2100–2150 m.
Distribution: SL—Gereau 5130
Asplenium dregeanum Kunze; Beentje, F.T.E.A.
(Aspleniaceae): 43. 2008.
Epiphyte. Montane forest. 1500–1790 m.
Distribution: NL, RM—Gereau 4170
Asplenium friesiorum C. Chr.; Beentje, F.T.E.A.
(Aspleniaceae): 27. 2008.
Terrestrial or epiphytic herb. Montane forest.
1950–2450 m.
Distribution: MR, RM—Gereau 4588
Asplenium hypomelas Kuhn; Beentje, F.T.E.A.
(Aspleniaceae): 48. 2008.
Terrestrial, epiphytic, or epilithic herb. Montane forest. 2040–2200 m.
Distribution: MR—Mwasumbi 16315
Asplenium monanthes L.; Beentje, F.T.E.A. (Aspleniaceae): 16. 2008.
Terrestrial or epiphytic herb. Montane forest.
2200–2460 m.
Distribution: KP, NL, RM—Gereau 4116
Asplenium protensum Schrad.; Beentje, F.T.E.A.
(Aspleniaceae): 41. 2008.
Epiphyte and on wet riverbanks. Montane forest. 1800–2380 m.
Distribution: KP, MR, RM, SL—Gereau 4589
Asplenium theciferum (Kunth) Mett.; Beentje,
F.T.E.A. (Aspleniaceae): 55. 2008.
Epiphytic or epilithic herb. Montane forest, tree
clumps in grassland. 1800–2380 m.
Distribution: KP, MR, RM, SL—Gereau 3728
CYATHEACEAE
Gereau et al.
Cyathea manniana Hook.; Edwards, F.T.E.A.
(Cyatheaceae): 9. 2005.
Tree fern. Montane forest. 1880–2100 m.
Distribution: MR, NL, RM—Gereau 4227
Cyathea mossambicensis Baker; Edwards, F.T.E.A.
(Cyatheaceae): 7. 2005.
Tree fern. Montane forest. 1560–1710 m.
Distribution: RM—Mwasumbi 16293
Cyathea thomsonii Baker; Edwards, F.T.E.A.
(Cyatheaceae): 12. 2005.
Tree fern. Montane forest. 1940 m.
Distribution: RM—WCS SHCP 6
DRYOPTERIDACEAE
5 genera and 8 species
Athyrium schimperi Moug. ex Fée; Verdcourt,
F.T.E.A. (Woodsiaceae): 8. 2003.
Terrestrial herb. Stream banks in grassland.
2560–2570 m.
Distribution: KP, MP—Gereau 3166
Dryoathyrium boryanum (Willd.) Ching; Verdcourt, F.T.E.A. (Woodsiaceae): 2. 2003.
Terrestrial herb. Montane forest, riverine forest.
1250–2460 m.
Distribution: MP, NL—Gereau 4121
Dryopteris athamantica (Kunze) Kuntze; Roux,
F.T.E.A. (Dryopteridaceae): 38. 2007.
Terrestrial herb. Montane grassland. 2080–
2140 m.
Distribution: SL—Suleiman 144
Dryopteris pentheri (Krasser) C. Chr.; Roux,
F.T.E.A. (Dryopteridaceae): 4. 2007.
Terrestrial herb. Montane forest and woodland,
riverine forest. 1250–2350 m.
Distribution: MP, RM, SL—Suleiman 100
Dryopteris sp.
Terrestrial herb. Thicket edges in montane
grassland. 2600–2650 m.
Distribution: KP—Gereau 3198
1 genus and 4 species
Cyathea dregei Kunze; Edwards, F.T.E.A.
(Cyatheaceae): 8. 2005.
Tree fern. Riverine forest in montane grassland.
1810 m.
Distribution: SL—Harder 1506
LakeNyasa_5-18.indd 10
Polystichum transvaalense N. C. Anthony; Roux,
F.T.E.A. (Dryopteridaceae): 32. 2007.
Terrestrial herb. Montane forest. 2150 m.
Distribution: MR—Gereau 4619
Polystichum wilsonii H. Christ; Roux, F.T.E.A.
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Floristic Checklist
(Dryopteridaceae): 30. 2007.
Terrestrial herb. Montane forest. 2460 m.
Distribution: NL—Gereau 4119
Tectaria gemmifera (Fée) Alston; Shaffer-Fehre,
F.T.E.A. (Dryopteridaceae): 4. 2007.
Terrestrial herb. Riverine forest. 1250 m.
Distribution: MP—Kayombo 2349
EQUISETACEAE
1 genus and 1 species
Equisetum ramosissimum Desf.; Verdcourt,
F.T.E.A. (Equisetaceae): 1. 1999.
Terrestrial herb. Roadsides in miombo woodland, riverine forest. 1150–1500 m.
Distribution: MP, SL—Kayombo 2459
11
LYCOPODIACEAE
2 genera and 3 species
Huperzia dacrydioides (Baker) Pic. Serm.; Verdcourt, F.T.E.A. (Lycopodiaceae): 5. 2005.
Epiphyte. Montane forest. 1960–1980 m.
Distribution: MR—Mwasumbi 16368
Huperzia ophioglossoides (Lam.) Rothm.; Verdcourt, F.T.E.A. (Lycopodiaceae): 7. 2005.
Epiphyte. Montane forest. 2640 m.
Distribution: NL—Gereau 4635
Lycopodium clavatum L. subsp. clavatum; Verdcourt, F.T.E.A. (Lycopodiaceae): 11. 2005.
Epiphyte. Montane forest.
Distribution: RM—Ndangalasi 1013
NEPHROLEPIDACEAE
GLEICHENIACEAE
1 genus and 1 species
1 genus and 1 species
Sticherus flagellaris (Bory ex Willd.) Ching subsp.
tomentosus (Reimers) Verdc.; Verdcourt,
F.T.E.A. (Gleicheniaceae): 5. 2000.
Climbing terrestrial herb. Cliff face in montane
forest. 2070 m.
Distribution: NL—Bracebridge 76
Nephrolepis undulata (Afzel. ex Sw.) J. Sm.; Verdcourt, F.T.E.A. (Oleandraceae): 5. 2001.
Terrestrial herb. Montane grassland, riverine
forest, miombo woodland, cultivated fields. 1500–
2440 m.
Distribution: MP, MR, RM, SL—Suleiman 186
OLEANDRACEAE
HYMENOPHYLLACEAE
1 genus and 1 species
Hymenophyllum capillare Desv.; Beentje, F.T.E.A.
(Hymenophyllaceae): 3. 2008.
Epiphyte. Riverine forest. 1710–1740 m.
Distribution: RM—Mwasumbi 16335
LOMARIOPSIDACEAE
1 genus and 2 species
Elaphoglossum aubertii (Desv.) T. Moore; Mickel,
F.T.E.A. (Lomariopsidaceae): 12. 2002.
Terrestrial herb. Forest patches in upland moor.
2620–2770 m.
Distribution: RM—Mwasumbi 16265
Elaphoglossum macropodium (Fée) T. Moore;
Mickel, F.T.E.A. (Lomariopsidaceae): 18.
2002.
Epiphyte. Montane forest. 1750 m.
Distributon: RM—Kindeketa 2983
LakeNyasa_5-18.indd 11
2 genera and 2 species
Arthropteris monocarpa (Cordem.) C. Chr.; Verdcourt, F.T.E.A. (Oleandraceae): 10. 2001.
Epiphyte. Montane forest. 1790–2400 m.
Distribution: NL, RM, SL—Gereau 4342
Oleandra distenta Kunze; Verdcourt, F.T.E.A.
(Oleandraceae): 13. 2001.
Epiphyte. Montane forest. 1790 m.
Distribution: NL—Gereau 4172
OPHIOGLOSSACEAE
1 genus and 2 species
Ophioglossum reticulatum L.; Burrows, F.T.E.A.
(Ophioglossaceae): 15. 2001.
Terrestrial herb. Riverine forest. 1850 m.
Distribution: SL—Gereau 4050
Ophioglossum vulgatum L.; Burrows, F.T.E.A.
(Ophioglossaceae): 15. 2001.
Terrestrial herb. Montane grassland. 2570–
2590 m.
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Gereau et al.
Distribution: KP—Gereau 3083
OSMUNDACEAE
1 genus and 1 species
Osmunda regalis L.; Verdcourt, F.T.E.A. (Osmundaceae): 1. 1999.
Terrestrial herb. Stream banks in montane
grassland. 2560 m.
Distribution: KP—Gereau 3179
POLYPODIACEAE
4 genera and 4 species
Drynaria volkensii Hieron.; Verdcourt, F.T.E.A.
(Polypodiaceae): 12. 2001.
Epiphyte. Montane forest, riverine forest.
1850–2140 m.
Distribution: SL—Suleiman 131
Lepisorus excavatus (Bory ex Willd.) Ching; Verdcourt, F.T.E.A. (Polypodiaceae): 17. 2001.
Epiphytic or epilithic herb. Tree clumps in
grassland, montane forest, miombo woodland.
1800–2380 m.
Distribution: KP, MP, MR, SL—Suleiman 125A
Loxogramme abyssinica (Baker) M. G. Price;
Verdcourt, F.T.E.A. (Polypodiaceae): 33. 2001.
Epiphyte. Montane forest, riverine forest.
980–2150 m.
Distribution: NL, RM, SL—Gereau 5156
Pleopeltis macrocarpa (Bory ex Willd.) Kaulf.;
Verdcourt, F.T.E.A. (Polypodiaceae): 26. 2001.
Epiphyte. Montane forest. 2080–2140 m.
Distribution: SL—Suleiman 125
PTERIDACEAE
4 genera and 5 species
Adiantum capillus-veneris L.; Verdcourt, F.T.E.A.
(Adiantaceae): 61. 2002.
Terrestrial herb. Riverine forest. 1150 m.
Distribution: MP—Kayombo 2440
Adiantum poiretii Wikstr.; Verdcourt, F.T.E.A.
(Adiantaceae): 62. 2002.
Terrestrial herb. Wet riverbanks in montane
forest, riverine forest in miombo. 1275–2380 m.
Distribution: KP, MP—Kayombo 2360
LakeNyasa_5-18.indd 12
Cheilanthes farinosa (Forssk.) Kaulf.; Verdcourt,
F.T.E.A. (Adiantaceae): 28. 2002.
Terrestrial herb. Riverine forest. 1275–1300 m.
Distribution: MP, MR—Kayombo 2359
Pellaea pectiniformis Baker; Verdcourt, F.T.E.A.
(Adiantaceae): 9. 2002.
Epilithic herb. Miombo woodland. 1780 m.
Distribution: SL—Gereau 5101
Pteris catoptera Kunze var. catoptera; Verdcourt,
F.T.E.A. (Pteridaceae): 25. 2002.
Terrestrial herb. Montane forest, riverine forest.
1250–2350 m.
Distribution: MP, RM—Kayombo 2347
SCHIZAEACEAE
1 genus and 2 species
Mohria lepigera (Baker) Baker; Verdcourt, F.T.E.A.
(Schizaeaceae): 8. 2000.
Terrestrial herb. Stream banks in montane
grassland. 2560–2570 m.
Distribution: KP—Gereau 3167
Mohria vestita Baker; Verdcourt, F.T.E.A. (Schizaeaceae): 7. 2000.
Terrestrial herb. Rocky montane grassland.
2488–2770 m.
Distribution: RM, SL—Suleiman 179
SELAGINELLACEAE
1 genus and 3 species
Selaginella eublepharis A. Braun; Verdcourt,
F.T.E.A. (Selaginellaceae): 10. 2005.
Terrestrial herb or epiphyte. Miombo woodland. 620 m.
Distribution: SL—Harder 1518
Selaginella goudotiana Spring var. abyssinica
(Spring) Bizzarri; Verdcourt, F.T.E.A. (Selaginellaceae): 19. 2005.
Terrestrial herb. Montane forest. 1560–2350
m.
Distribution: RM, SL—Mwasumbi 16297
Selaginella kraussiana (Kunze) A. Braun; Verdcourt, F.T.E.A. (Selaginellaceae): 18. 2005.
Terrestrial herb. Montane forest, bamboo forest, montane grassland. 1750–2760 m.
5/31/12 3:59 PM
Lake Nyasa Climatic Region
Floristic Checklist
Distribution: KP, RM—Abeid 2897
13
MAGNOLIOPHYTA
THELYPTERIDACEAE
3 genera and 4 species
Amauropelta bergiana (Schltdl.) Holttum var. bergiana; Verdcourt, F.T.E.A. (Thelypteridaceae):
6. 2006.
Terrestrial herb. Montane forest. 2460 m.
Distribution: NL—Gereau 4120
Christella chaseana (Schelpe) Holttum; Verdcourt,
F.T.E.A. (Thelypteridaceae): 17. 2006.
Epilithic herb. Riverine forest. 980 m.
Distribution: NL—Gereau 5163
Christella dentata (Forssk.) Brownsey & Jermy;
Verdcourt, F.T.E.A. (Thelypteridaceae): 13.
2006.
Epilithic herb. Riverine forest. 980–1400 m.
Distribution: MP, NL—Gereau 5164
Pneumatopteris unita (Kunze) Holttum; Verdcourt, F.T.E.A. (Thelypteridaceae): 32. 2006.
Terrestrial herb. Montane forest. 1880 m.
Distribution: NL—Gereau 4226
ACANTHACEAE
20 genera and 53 species
Acanthopale confertiflora (Lindau) C. B. Clarke;
Vollesen, F.T.E.A. (Acanthaceae 1): 217. 2008.
Shrub. Montane forest and clearings. 1800–
2230 m.
Distribution: MR, NL, RM, SL—Gereau 4423
Acanthus ueleensis De Wild.; Vollesen, F.T.E.A.
(Acanthaceae 1): 94. 2008.
Terrestrial herb. Montane and riverine forest.
1250–2040 m.
Distribution: MP, MR, RM, SL—Kayombo
2345
Asystasia albiflora Ensermu; Ensermu & Vollesen,
F.T.E.A. (Acanthaceae 2): 455. 2010.
Terrestrial herb. Riverine forest and thickets,
woodland. 1200–1250 m.
Distribution: MP—Kayombo 2346
1 genus and 1 species
Asystasia gangetica (L.) T. Anderson subsp.
micrantha (Nees) Ensermu; Ensermu & Vollesen, F.T.E.A. (Acanthaceae 2): 459. 2010.
Scrambling terrestrial herb. Riverine forest and
bushland. 474–1700 m.
Distribution: MP, NL, RM, SL—Mwasumbi
16554
Juniperus procera Hochst. ex Endl.; Melville,
F.T.E.A. (Gymnospermae): 16. 1958.
Tree. Montane forest, stream banks in montane
grassland. 2300–2700 m.
Distribution: KP—Congdon 20
Barleria lactiflora Brummitt & Seyani; Darbyshire,
F.T.E.A. (Acanthaceae 2): 392. 2010.
Terrestrial herb. Miombo woodland. 1200–
1210 m.
Distribution: MP, SL—Gereau 4297
PINOPHYTA
CUPRESSACEAE
PODOCARPACEAE
2 genera and 2 species
Afrocarpus falcatus (Thunb.) C. N. Page; Lovett
et al., 2006: 9.
Tree. Montane forest with bamboo. 2630 m.
Distribution: RM—Abeid 2994
Podocarpus latifolius (Thunb.) R. Br. ex Mirb.;
Lovett et al., 2006: 10.
Tree. Montane forest. 1450–2380 m.
Distribution: KP, RM—Abeid 2821
LakeNyasa_5-18.indd 13
Blepharis grandis C. B. Clarke; Vollesen, F.T.E.A.
(Acanthaceae 1): 120. 2008.
Terrestrial herb. Grassland, woodland, riverine
forest. 1250–1400 m.
Distribution: MP—Kayombo 1061
Blepharis katangensis De Wild.; Vollesen, F.T.E.A.
(Acanthaceae 1): 125. 2008.
Terrestrial herb. Grassland. 1300–1400 m.
Distribution: MP—Kayombo 1047
Blepharis longifolia Lindau; Vollesen, F.T.E.A.
(Acanthaceae 1): 116. 2008.
Terrestrial herb. Shrubland. 1250 m.
Distribution: MP—J. Lovett 4581
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14
Brachystephanus coeruleus S. Moore subsp.
apiculatus Champl.; Darbyshire, F.T.E.A.
(Acanthaceae 2): 472. 2010.
Herbaceous climber. Montane forest. 2000 m.
Distribution: MP—P. Lovett 445
Brillantaisia cicatricosa Lindau; Vollesen, F.T.E.A.
(Acanthaceae 1): 159. 2008.
Terrestrial herb. Riverine forest. 1250–1400 m.
Distribution: MP—Kayombo 2351
Dicliptera brevispicata I. Darbysh.; Darbyshire,
F.T.E.A. (Acanthaceae 2): 714. 2010.
Terrestrial herb. Woodland, coffee plantations.
1300–1500 m.
Distribution: MP—J. Lovett 4803
Dicliptera capitata Milne-Redh.; Darbyshire,
F.T.E.A. (Acanthaceae 2): 707. 2010.
Terrestrial herb. Miombo woodland. 1400 m.
Distribution: MP—P. Lovett 97
Dicliptera carvalhoi Lindau subsp. nemorum
(Milne-Redh.) I. Darbysh.; Darbyshire, F.T.E.A.
(Acanthaceae 2): 710. 2010.
Herbaceous climber. Riverine forest. 1150 m.
Distribution: MP—Kayombo 2453
Dicliptera maculata Nees subsp. maculata; Darbyshire, F.T.E.A. (Acanthaceae 2): 701. 2010.
Shrub. Montane forest. 2110 m.
Distribution: MR—Gereau 4546
Duosperma livingstoniense Vollesen; Vollesen,
F.T.E.A. (Acanthaceae 1): 254. 2008.
Terrestrial herb. Miombo woodland. 1530 m.
Distribution: SL—Gereau 4350 (type: DSM,
EA, K, MO, NHT, PRE)
Duosperma tanzaniense Vollesen; Vollesen,
F.T.E.A. (Acanthaceae 1): 253. 2008.
Terrestrial herb. Woodland, riverine forest.
1150–1400 m.
Distribution: MP, MR—Bidgood 2105 (type:
BR, C, CAS, DSM, K, NHT, UPS), Kayombo 2448
Dyschoriste albiflora Lindau; Vollesen, F.T.E.A.
(Acanthaceae 1): 188. 2008.
Shrub. Riverine forest. 1200 m.
Distribution: MP, MR—J. Lovett 4409
Dyschoriste nyassica Gilli; Vollesen, F.T.E.A.
(Acanthaceae 1): 191. 2008.
LakeNyasa_5-18.indd 14
Gereau et al.
Terrestrial herb or shrub. Riverine forest. 525
m.
Distribution: SL—Gilli 522 (type: K, W)
Dyschoriste trichocalyx (Oliv.) Lindau subsp.
verticillaris (C. B. Clarke) Vollesen; Vollesen,
F.T.E.A. (Acanthaceae 1): 192. 2008.
Terrestrial herb. Shrubland. 1300 m.
Distribution: MP—J. Lovett 4806
Hygrophila schulli (Buch.-Ham.) M. R. Almeida &
S. M. Almeida; Vollesen, F.T.E.A. (Acanthaceae
1): 176. 2008.
Terrestrial herb. Woodland, grassland, seasonal
swamps. 1250–1400 m.
Distribution: MP—Kayombo 984
Hypoestes aristata (Vahl) Sol. ex Roem. & Schult.;
Darbyshire, F.T.E.A. (Acanthaceae 2): 724.
2010.
Terrestrial herb or shrub. Montane forest, forest
edges, riverine forest. 1350–2400 m.
Distribution: MP, MR, RM—Mwasumbi 16208
Hypoestes forskaolii (Vahl) R. Br. subsp. forskaolii;
Darbyshire, F.T.E.A. (Acanthaceae 2): 726.
2010.
Terrestrial herb or shrub. Montane forest,
woodland. 1100–2110 m.
Distribution: MP, MR, NL—Mwasumbi 16312
Hypoestes triflora (Forssk.) Roem. & Schult.; Darbyshire, F.T.E.A. (Acanthaceae 2): 720. 2010.
Terrestrial herb. Montane forest, riverine forest.
1250–1980 m.
Distribution: MP, MR, RM—Kayombo 2467
Isoglossa bracteosa Mildbr.; Darbyshire, F.T.E.A.
(Acanthaceae 2): 638. 2010.
Terrestrial herb. Montane forest. 2280–2460
m.
Distribution: NL, RM—Mwasumbi 16236
Isoglossa grandiflora C. B. Clarke; Darbyshire,
F.T.E.A. (Acanthaceae 2): 645. 2010.
Herb or shrub. Riverine forest. 1150–1400 m.
Distribution: MP—Kayombo 2442
Isoglossa gregoryi (S. Moore) Lindau; Darbyshire,
F.T.E.A. (Acanthaceae 2): 621. 2010.
Terrestrial herb. Disturbed montane forest.
1980 m.
Distribution: MR—Gereau 4624
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Floristic Checklist
Isoglossa multinervis I. Darbysh.; Darbyshire,
F.T.E.A. (Acanthaceae 2): 635. 2010.
Shrub. Montane forest. 2500 m.
Distribution: MR—Kayombo 1105 (type: K,
MO), Aleljung 536
Isoglossa strigulosa C. B. Clarke; Darbyshire,
F.T.E.A. (Acanthaceae 2): 628. 2010.
Terrestrial herb, often climber or straggler.
Montane forest, bamboo forest. 2000–2400 m.
Distribution: KP, MP, MR, RM—Mwasumbi
16513
Justicia acutifolia Hedrén; Vollesen, F.T.E.A.
(Acanthaceae 2): 591. 2010.
Terrestrial herb. Rocky grassland. 2100–2320
m.
Distribution: KP, SL—Richards 14014 (type:
BR, K), Gereau 3990
Justicia betonica L.; Vollesen, F.T.E.A. (Acanthaceae 2): 549. 2010.
Terrestrial herb. Riverine forest. 1210–1500 m.
Distribution: MP, SL—Gereau 4390
Justicia flava (Forssk.) Vahl; Vollesen, F.T.E.A.
(Acanthaceae 2): 532. 2010.
Terrestrial herb. Riverine forest. 1210–1250 m.
Distribution: SL—Gereau 4256
Justicia linearispica C. B. Clarke; Vollesen, F.T.E.A.
(Acanthaceae 2): 539. 2010.
Terrestrial herb. Woodland. 1720 m.
Distribution: MP—J. Lovett 3505
Justicia mollugo C. B. Clarke; Vollesen, F.T.E.A.
(Acanthaceae 2): 589. 2010.
Terrestrial herb. Riverine forest, grassland.
1850–1940 m.
Distribution: NL, SL—Gereau 4677
Justicia nuttii C. B. Clarke; Vollesen, F.T.E.A.
(Acanthaceae 2): 561. 2010.
Terrestrial herb. Grassland. 2360–2640 m.
Distribution: KP, MR, NL—Suleiman 38
Justicia nyassana Lindau; Vollesen, F.T.E.A.
(Acanthaceae 2): 539. 2010.
Terrestrial herb. Riverine forest. 1400–1600 m.
Distribution: MP—Kayombo 2477
Justicia phyllostachys C. B. Clarke; Vollesen,
F.T.E.A. (Acanthaceae 2): 586. 2010.
LakeNyasa_5-18.indd 15
15
Terrestrial herb. Miombo woodland. 1700 m.
Distribution: NL—Bidgood 152
Justicia striata (Klotzsch) Bullock; Vollesen,
F.T.E.A. (Acanthaceae 2): 584. 2010.
Terrestrial herb. Riverine forest, miombo
woodland. 850–1800 m.
Distribution: MP, SL—Gereau 4280
Justicia striolata Mildbr.; Vollesen, F.T.E.A. (Acanthaceae 2): 576. 2010.
Terrestrial herb. Riverine forest, montane forest. 1350–1710 m.
Distribution: MP, RM—Mwasumbi 16273
Justicia sp.
Terrestrial herb. Montane forest edges. 2300 m.
Distribution: SL—Iwarsson 1212
Note: In Justicia sect. Harnieria, fide Hedrén
(1984 annotation, NHT).
Lepidagathis eriocephala Lindau; Darbyshire,
F.T.E.A. (Acanthaceae 2): 320. 2010.
Terrestrial herb. Woodland, abandoned cultivation. 1300–1400 m.
Distribution: MP—Goetze 1083 (type: BM,
BR), Kayombo 992
Lepidagathis lanatoglabra C. B. Clarke; Darbyshire, F.T.E.A. (Acanthaceae 2): 321. 2010.
Terrestrial herb. Grassland, woodland, abandoned cultivation. 1200–1400 m.
Distribution: MP—Kayombo 2417
Mimulopsis solmsii Schweinf.; Vollesen, F.T.E.A.
(Acanthaceae 1): 222. 2008.
Terrestrial herb. Montane forest. 2500 m.
Distribution: MR—Kayombo 1103
Nelsonia canescens (Lam.) Spreng.; Vollesen,
F.T.E.A. (Acanthaceae 1): 18. 2008.
Straggling terrestrial herb. Riverine forest,
woodland, lawns near houses. 1300–1620 m.
Distribution: MP—J. Lovett 4753
Ruspolia seticalyx (C. B. Clarke) Milne-Redh.;
Vollesen, F.T.E.A. (Acanthaceae 2): 486. 2010.
Suffrutex. Riverine forest. 1275–1300 m.
Distribution: MP—Kayombo 2361
Sclerochiton obtusisepalus C. B. Clarke; Vollesen,
F.T.E.A. (Acanthaceae 1): 83. 2008.
Shrub. Montane forest. 1800 m.
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Gereau et al.
Distribution: RM—Kibure 1394
Thunbergia alata Bojer ex Sims; Vollesen, F.T.E.A.
(Acanthaceae 1): 56. 2008.
Trailing terrestrial herb or vine. Woodland,
grassland, riverine forest, abandoned cultivation.
1250–2600 m.
Distribution: KP, MP, MR, SL—Suleiman 114
Thunbergia gregoryi S. Moore; Vollesen, F.T.E.A.
(Acanthaceae 1): 52. 2008.
Trailing terrestrial herb. Roadsides. 1600–1900
m.
Distribution: MP, NL—Iwarsson 1143
Thunbergia kirkiana T. Anderson; Vollesen,
F.T.E.A. (Acanthaceae 1): 43. 2008.
Terrestrial herb. Woodland. 1300–1600 m.
Distribution: MP—J. Lovett 3614
Thunbergia lancifolia T. Anderson; Vollesen,
F.T.E.A. (Acanthaceae 1): 70. 2008.
Terrestrial herb. Woodland, roadsides, cultivated fields. 1250–2000 m.
Distribution: MP—J. Lovett 5005
Thunbergia lathyroides Burkill; Vollesen, F.T.E.A.
(Acanthaceae 1): 68. 2008.
Terrestrial herb. Woodland. 1400 m.
Distribution: MP—J. Lovett 3292A
Thunbergia oblongifolia Oliv.; Vollesen, F.T.E.A.
(Acanthaceae 1): 73. 2008.
Terrestrial herb. Bamboo forest. 2340 m.
Distribution: KP—WCS SHCP 354
Thunbergia petersiana Lindau; Vollesen, F.T.E.A.
(Acanthaceae 1): 63. 2008.
Terrestrial herb. Riverine forest. 1160 m.
Distribution: SL—Gereau 3696
AIZOACEAE
1 genus and 1 species
Psammotropha myriantha Sond.; Jeffrey, F.T.E.A.
(Aizoaceae): 11. 1961.
Terrestrial herb. Rocky outcrops in montane
grassland. 2600–2700 m.
Distribution: KP—Gereau 2827
ALANGIACEAE
1 genus and 1 species
LakeNyasa_5-18.indd 16
Alangium chinense (Lour.) Harms; Verdcourt,
F.T.E.A. (Alangiaceae): 3. 1958.
Tree. Montane forest edges and clearings.
1840–1860 m.
Distribution: RM—Abeid 2767
AMARANTHACEAE
6 genera and 7 species
Achyranthes aspera L. var. pubescens (Moq.) C. C.
Towns.; Townsend, F.T.E.A. (Amaranthaceae):
102. 1985.
Terrestrial herb. Montane forest, riverine forest.
1400–2000 m.
Distribution: MP—J. Lovett 4678
Achyranthes aspera L. var. sicula L.; Townsend,
F.T.E.A. (Amaranthaceae): 104. 1985.
Terrestrial herb. Abandoned cultivation, roadsides, trailsides in montane forest. 1250–2460 m.
Distribution: MP, MR, NL—Gereau 4521
Achyropsis laniceps C. B. Clarke; Townsend,
F.T.E.A. (Amaranthaceae): 108. 1985.
Terrestrial herb. Miombo woodland. 1400–
1800 m.
Distribution: MP—P. Lovett 111
Aerva lanata (L.) Juss. ex Schult.; Townsend,
F.T.E.A. (Amaranthaceae): 85. 1985.
Straggling herb. Riverine forest. 1210 m.
Distribution: SL—Gereau 4267
Celosia trigyna L.; Townsend, F.T.E.A. (Amaranthaceae): 12. 1985.
Terrestrial herb. Abandoned cultivation, woodland. 1250 m.
Distribution: MP—J. Lovett 4456
Cyathula cylindrica Moq.; Townsend, F.T.E.A.
(Amaranthaceae): 64. 1985.
Herbaceous climber. Montane forest. 2000–
2200 m.
Distribution: MP, MR—Gereau 4522
Cyathula polycephala Baker; Townsend, F.T.E.A.
(Amaranthaceae): 61. 1985.
Scrambling herb. Montane forest edges. 2360
m.
Distribution: MR—Gereau 3760
Pandiaka rubrolutea (Lopr.) C. C. Towns.;
Townsend, F.T.E.A. (Amaranthaceae): 115.
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Lake Nyasa Climatic Region
Floristic Checklist
1985.
Terrestrial herb. Riverine forest. 1350 m.
Distribution: MP—J. Lovett 4653
AMARYLLIDACEAE
3 genera and 4 species
Boophone disticha (L. f.) Herb.; Nordal, F.T.E.A.
(Amaryllidaceae): 21. 1982.
Terrestrial herb. Woodland. 1300 m.
Distribution: MP—J. Lovett 4913
Cyrtanthus breviflorus Harv.; Nordal, F.T.E.A.
(Amaryllidaceae): 26. 1982.
Terrestrial herb. Montane grassland and heathland. 2600–2730 m.
Distribution: KP, MP, RM—Abeid 2998
Scadoxus multiflorus (Martyn) Raf. subsp. multiflorus; Nordal, F.T.E.A. (Amaryllidaceae): 5.
1982.
Terrestrial herb. Woodland, riverine forest,
montane forest, edges of cultivation. 1250–2260
m.
Distribution: MP, SL—Gereau 3533
Scadoxus puniceus (L.) Friis & Nordal; Nordal,
F.T.E.A. (Amaryllidaceae): 5. 1982.
Terrestrial herb. Open woodland, cultivated
fields. 1400–1700 m.
Distribution: MP—J. Lovett 3355
Note: J. Lovett 3351, with narrow involucral
bracts, is a possible hybrid with S. multiflorus
subsp. multiflorus.
17
Distribution: MP, SL—Kayombo 2412
Rhus anchietae Hiern vel sp. aff.; Kokwaro,
F.T.E.A. (Anacardiaceae): 35. 1986.
Shrub. Montane forest. 1740 m.
Distribution: RM—Abeid 2930
Rhus brenanii Kokwaro; Kokwaro, F.T.E.A. (Anacardiaceae): 38. 1986.
Shrub, scandent shrub, or liana. Montane forest
and forest edges, miombo woodland. 1400–2600
m.
Distribution: MP, MR, RM, SL—Mwasumbi
16242
Rhus glaucescens A. Rich.; Moffett, 1993: 76.
Shrub. Riverine forest. 1500 m.
Distribution: SL—Gereau 4787
Rhus longipes Engl. var. elgonensis Kokwaro; Kokwaro, F.T.E.A. (Anacardiaceae): 38. 1986.
Shrub. Riverine forest, grassland, montane forest edges. 1810–2340 m.
Distribution: RM, SL—Harder 1474
Rhus longipes Engl. var. longipes; Kokwaro,
F.T.E.A. (Anacardiaceae): 37. 1986.
Shrub. Miombo woodland, lakeshores, wet
meadows, disturbed montane forest. 474–1790 m.
Distribution: MP, NL, RM—Harder 1517
Rhus longipes Engl. var. schinoides R. Fern.; Kokwaro, F.T.E.A. (Anacardiaceae): 38. 1986.
Shrub or small tree. Riverine forest, thickets.
1810–2140 m.
Distribution: SL—Suleiman 95
ANACARDIACEAE
5 genera and 12 species
Lannea katangensis Van der Veken; Kokwaro,
F.T.E.A. (Anacardiaceae): 16. 1986.
Shrublet. Woodland. 1200–1400 m.
Distribution: MP—Kayombo 1020
Rhus ochracea Meikle; Kokwaro, F.T.E.A. (Anacardiaceae): 35. 1986.
Shrub. Lower montane forest. 2015 m.
Distribution: MR—Gereau 7022
Lannea schimperi (Hochst. ex A. Rich.) Engl.;
Kokwaro, F.T.E.A. (Anacardiaceae): 19. 1986.
Tree. Miombo woodland. 1350 m.
Distribution: MP—Kayombo 1092
Rhus pyroides Burch. var. gracilis (Engl.) Burtt
Davy; Kokwaro, F.T.E.A. (Anacardiaceae): 32.
1986.
Shrub or small tree. Montane forest, bamboo
forest, heathland. 1800–2890 m.
Distribution: KP, MP, NL, RM—Abeid 2997
Ozoroa insignis Delile subsp. reticulata (Baker f.)
J. B. Gillett; Kokwaro, F.T.E.A. (Anacardiaceae): 5. 1986.
Tree. Rocky gorges, lakeshore, miombo woodland. 474–1390 m.
Rhus pyroides Burch. var. pyroides; Moffett, 1993:
43.
Shrub or small tree. Thickets, wooded grassland, miombo woodland, disturbed montane
forest, riverbanks, lakeshores. 474–2250 m.
LakeNyasa_5-18.indd 17
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18
Gereau et al.
Distribution: MP, RM, SL—Kayombo 2369
Rhus quartiniana A. Rich.; Kokwaro, F.T.E.A.
(Anacardiaceae): 33. 1986.
Tree. Disturbed montane forest. 1790 m.
Distribution: RM—Abeid 2906
Sorindeia madagascariensis Thouars ex DC.; Kokwaro, F.T.E.A. (Anacardiaceae): 46. 1986.
Tree. Riverine forest, lakeshore. 474–480 m.
Distribution: NL, SL—Gereau 3656
Trichoscypha ulugurensis Mildbr. subsp. submontana (Van der Veken) Kokwaro; Kokwaro
F.T.E.A. (Anacardiaceae): 51. 1986.
Tree. Montane forest. 1750 m.
Distribution: NL—Bracebridge 107
Trichoscypha ulugurensis Mildbr. subsp. ulugurensis; Kokwaro, F.T.E.A. (Anacardiaceae): 51.
1986.
Tree. Montane forest. 1830 m.
Distribution: RM—Kibure 1408
ANNONACEAE
(Annonaceae): 48. 1971.
Tree. Miombo woodland. 1200 m.
Distribution: MP—J. Lovett 4040
Uvaria lucida Bojer ex Benth. subsp. lucida; Verdcourt, F.T.E.A. (Annonaceae): 18. 1971.
Liana. Lakeshore shrubland, riverine forest.
474–840 m.
Distribution: SL—Mwasumbi 16592
ANTHERICACEAE
1 genus and 16 species
Chlorophytum affine Baker var. affine; Nordal et
al., F.T.E.A. (Anthericaceae): 24. 1997.
Terrestrial herb. Rocky mountain slopes and
grassland. 2740–2800 m.
Distribution: KP—J. Lovett 3670
Chlorophytum blepharophyllum Schweinf. ex
Baker; Nordal et al., F.T.E.A. (Anthericaceae):
51. 1997.
Terrestrial herb. Grassland, woodland, abandoned cultivation. 1250–1600 m.
Distribution: MP—J. Lovett 3531
5 genera and 6 species
Annona senegalensis Pers. subsp. senegalensis;
Verdcourt, F.T.E.A. (Annonaceae): 115. 1971.
Shrub or small tree. Rocky lakeshore, woodland, abandoned cultivation. 474–1320 m.
Distribution: MP, SL—Mwasumbi 16560
Artabotrys monteiroae Oliv.; Verdcourt, F.T.E.A.
(Annonaceae): 62. 1971.
Liana or shrub. Woodland, disturbed riverine
forest. 1100–1200 m.
Distribution: MP—Kayombo 2455
Artabotrys stolzii Diels; Verdcourt, F.T.E.A.
(Annonaceae): 64. 1971.
Liana. Riverine forest, montane forest. 1210–
1880 m.
Distribution: MP, NL, SL—Gereau 4252
Friesodielsia obovata (Benth.) Verdc.; Verdcourt,
F.T.E.A. (Annonaceae): 86. 1971.
Tree or shrub, sometimes lianescent. Riverine
forest, woodland. 580–1200 m.
Distribution: MP, SL—Kayombo 2460
Hexalobus monopetalus (A. Rich.) Engl. & Diels
var. obovatus Brenan; Verdcourt, F.T.E.A.
LakeNyasa_5-18.indd 18
Chlorophytum cameronii (Baker) Kativu var. cameronii; Nordal et al., F.T.E.A. (Anthericaceae):
31. 1997.
Terrestrial herb. Rocky woodland, riverine
thickets. 1200–2000 m.
Distribution: MP—J. Lovett 3477
Chlorophytum cameronii (Baker) Kativu var.
pterocaulon (Welw. ex Baker) Nordal; Nordal
et al., F.T.E.A. (Anthericaceae): 32. 1997.
Terrestrial herb. Woodland, montane grassland, rocky mountain slopes. 1200–2800 m.
Distribution: KP, MP—Goldblatt 8192A
Chlorophytum collinum (Poelln.) Nordal; Nordal
et al., F.T.E.A. (Anthericaceae): 23. 1997.
Terrestrial herb. Rocky mountain slopes. 2800
m.
Distribution: KP—Goldblatt 8192
Chlorophytum colubrinum (Baker) Engl.; Nordal
et al., F.T.E.A. (Anthericaceae): 19. 1997.
Terrestrial herb. Woodland. 1200–1250 m.
Distribution: MP—J. Lovett 4253
Chlorophytum comosum (Thunb.) Jacques; Nordal
et al., F.T.E.A. (Anthericaceae): 55. 1997.
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Lake Nyasa Climatic Region
Floristic Checklist
Terrestrial herb. Riverine forest, montane
grassland. 1210–2320 m.
Distribution: MP, SL—Gereau 4421
Chlorophytum floribundum Baker; Nordal et al.,
F.T.E.A. (Anthericaceae): 47. 1997.
Terrestrial herb. Riverine forest, abandonded
cultivation. 580–1250 m.
Distribution: MP, SL—Gereau 3823
Chlorophytum gallabatense Schweinf. ex Baker;
Nordal et al., F.T.E.A. (Anthericaceae): 45.
1997.
Terrestrial herb. Riverine forest and thickets,
woodland. 580–1310 m.
Distribution: MP, SL—Gereau 3824
Chlorophytum macrophyllum (A. Rich.) Asch.;
Nordal et al., F.T.E.A. (Anthericaceae): 49.
1997.
Terrestrial herb. Stream banks in montane
grassland. 2560 m.
Distribution: KP—Gereau 3183
Chlorophytum nubicum (Baker) Kativu; Nordal et
al., F.T.E.A. (Anthericaceae): 15. 1997.
Terrestrial herb. Woodland. 1200 m.
Distribution: MP—J. Lovett 3959
Chlorophytum nyasae (Rendle) Kativu; Nordal et
al., F.T.E.A. (Anthericaceae): 13. 1997.
Terrestrial herb. Montane grassland. 2120 m.
Distribution: SL—Gereau 4319
Chlorophytum rubribracteatum (De Wild.) Kativu
vel sp. aff.; Nordal et al., F.T.E.A. (Anthericaceae): 36. 1997.
Terrestrial herb. Woodland. 1250–1400 m.
Distribution: MP—J. Lovett 3650
Chlorophytum sphacelatum (Baker) Kativu var.
milanjianum (Rendle) Nordal; Nordal et al.,
F.T.E.A. (Anthericaceae): 35. 1997.
Terrestrial herb. Miombo woodland. 1200 m.
Distribution: MP—J. Lovett 4060
Chlorophytum stolzii (K. Krause) Kativu; Nordal
et al., F.T.E.A. (Anthericaceae): 15. 1997.
Terrestrial herb. Cultivated fields, grassland.
1100–1400 m.
Distribution: MP—Kayombo 994
Chlorophytum subpetiolatum (Baker) Kativu; Nordal et al., F.T.E.A. (Anthericaceae): 36. 1997.
LakeNyasa_5-18.indd 19
19
Terrestrial herb. Woodland. 1300–1950 m.
Distribution: MP—J. Lovett 3787
Chlorophytum zingiberastrum Nordal & A. D.
Poulsen; Nordal et al., F.T.E.A. (Anthericaceae): 48. 1997.
Terrestrial herb. Woodland, riverine forest.
1200 m.
Distribution: MP—J. Lovett 4301
APIACEAE
13 genera and 28 species
Agrocharis incognita (C. Norman) Heywood &
Jury; Townsend, F.T.E.A. (Umbelliferae): 30.
1989.
Terrestrial herb. Agricultural fields. 2200 m.
Distribution: MR—Gereau 4511
Agrocharis melanantha Hochst.; Townsend,
F.T.E.A. (Umbelliferae): 33. 1989.
Terrestrial herb. Wet montane grassland. 2660
m.
Distribution: KP—Abeid 2788
Agrocharis pedunculata (Baker f.) Heywood &
Jury; Townsend, F.T.E.A. (Umbelliferae): 32.
1989.
Terrestrial herb. Montane grassland. 1860–
2590 m.
Distribution: KP, MP, SL—Gereau 3870
Alepidea peduncularis Steud. ex A. Rich.;
Townsend, F.T.E.A. (Umbelliferae): 20. 1989.
Terrestrial herb. Wooded grassland, montane
grassland. 1700–2700 m.
Distribution: KP, MP, MR, NL—Suleiman 18
Anthriscus sylvestris (L.) Hoffm. var. sylvestris;
Townsend, F.T.E.A. (Umbelliferae): 26. 1989.
Terrestrial herb. Roadsides. 2300 m.
Distribution: MR—Macha 78
Centella asiatica (L.) Urb.; Townsend, F.T.E.A.
(Umbelliferae): 15. 1989.
Terrestrial herb. Gardens, ditch banks. 1740
m.
Distribution: MP—Batty 756
Cryptotaenia africana (Hook. f.) Drude;
Townsend, F.T.E.A. (Umbelliferae): 54. 1989.
Terrestrial herb. Montane forest. 1790–2110
m.
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20
Gereau et al.
Distribution: MR, NL—Mwasumbi 16198
Distribution: SL—Iwarsson 1210
Heracleum abyssinicum (Boiss.) C. Norman;
Townsend, F.T.E.A. (Umbelliferae): 121. 1989.
Terrestrial herb. Thickets and stream banks in
montane grassland. 2550–2650 m.
Distribution: KP—Gereau 3197
Pimpinella aff. alismatifolia C. C. Towns.;
Townsend, F.T.E.A. (Umbelliferae): 61. 1989.
Terrestrial herb. Rocky thickets, rocky grassland. 1800–2570 m.
Distribution: KP, NL—Gereau 3199
Heteromorpha arborescens Cham. & Schltdl. var.
abyssinica (Hochst. ex A. Rich.) H. Wolff; Winter & Wyk, 1996: 227.
Tree or shrub. Montane forest, riverine forest,
woodland. 1250–2420 m.
Distribution: KP, MP, NL, RM, SL—Gereau
4796
Pimpinella buchananii H. Wolff; Townsend,
F.T.E.A. (Umbelliferae): 63. 1989.
Terrestrial herb. Woodland, montane grassland. 1200–2500 m.
Distribution: MP, NL, SL—Gereau 4382
Heteromorpha involucrata Conrath; Townsend,
F.T.E.A. (Umbelliferae): 40. 1989.
Terrestrial herb. Miombo woodland, wooded
grassland, riverine forest, montane grassland.
1200–2600 m.
Distribution: KP, MP, MR—Gereau 7038
Lefebvrea grantii S. Droop; Townsend, F.T.E.A.
(Umbelliferae): 113. 1989.
Terrestrial herb. Riverine forest, woodland. 1250
m.
Distribution: MP—J. Lovett 4512
Peucedanum claessensii C. Norman; Townsend,
F.T.E.A. (Umbelliferae): 103. 1989.
Terrestrial herb. Montane forest, thickets in
montane grassland, bushland. 1430–2560 m.
Distribution: KP, MR, RM, SL—Gereau 4748
Peucedanum dispersum C. C. Towns.; Townsend,
F.T.E.A. (Umbelliferae): 101. 1989.
Terrestrial herb. Montane grassland. 2360 m.
Distribution: KP, MR—Hooper 1758 (type: K),
Gereau 3776
Peucedanum cf. kerstenii Engl.; Townsend,
F.T.E.A. (Umbelliferae): 102. 1989.
Terrestrial herb. Montane grassland. 1810 m.
Distribution: SL—Harder 1495
Peucedanum lynesii C. Norman; Townsend,
F.T.E.A. (Umbelliferae): 96. 1989.
Terrestrial herb. Montane grassland and shrubland. 2350–2500 m.
Distribution: KP—Brummitt 18129
Peucedanum sp.
Terrestrial herb. Montane grassland. 2300 m.
LakeNyasa_5-18.indd 20
Pimpinella caffra (Eckl. & Zeyh.) D. Dietr. subsp.
conopodioides C. C. Towns.; Townsend,
F.T.E.A. (Umbelliferae): 65. 1989.
Terrestrial herb. Rocky montane grassland.
2590 m.
Distribution: KP, MP, RM—Hooper 1755 (type:
K), Abeid 3052
Pimpinella duridentata C. C. Towns.; Townsend,
F.T.E.A. (Umbelliferae): 61. 1989.
Terrestrial herb. Montane grassland. 2700 m.
Distribution: NL—Gereau 4085
Pimpinella kyimbilaensis H. Wolff; Townsend,
F.T.E.A. (Umbelliferae): 65. 1989.
Terrestrial herb. Thickets in montane grassland. 2550 m.
Distribution: KP—Gereau 3150
Pimpinella ledermannii H. Wolff subsp. engleriana
(H. Wolff) C. C. Towns.; Townsend, F.T.E.A.
(Umbelliferae): 58. 1989.
Terrestrial herb. Miombo woodland. 1780 m.
Distribution: SL—Gereau 5104
Pimpinella rigidistyla C. C. Towns.; Townsend,
F.T.E.A. (Umbelliferae): 64. 1989.
Terrestrial herb. 2610 m.
Distribution: MP—Kerfoot 1743 (type: EA)
Pimpinella cf. rigidistyla C. C. Towns.; Townsend,
F.T.E.A. (Umbelliferae): 64. 1989.
Terrestrial herb. Riverine forest, rocky grassland. 1850–2320 m.
Distribution: SL—Gereau 4041
Sanicula elata Buch.-Ham. ex D. Don; Townsend,
F.T.E.A. (Umbelliferae): 17. 1989.
Terrestrial herb. Montane forest. 2360–2460
m.
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Lake Nyasa Climatic Region
Floristic Checklist
Distribution: MP, MR, NL—Gereau 4113
Sium repandum Welw. ex Hiern; Townsend,
F.T.E.A. (Umbelliferae): 73. 1989.
Terrestrial herb. Stream banks.
Distribution: RM—Richards 14271
Steganotaenia araliacea Hochst.; Townsend,
F.T.E.A. (Umbelliferae): 115. 1989.
Tree. Riverine forest, montane forest edges.
1740–2070 m.
Distribution: MR, RM—St. Clair-Thompson
738
Steganotaenia hockii (C. Norman) C. Norman;
Townsend, F.T.E.A. (Umbelliferae): 117. 1989.
Terrestrial herb. Miombo woodland, grassland.
1400–2250 m.
Distribution: MP—J. Lovett 4868
APOCYNACEAE
8 genera and 10 species
Carissa spinarum L.; Omino, F.T.E.A. (Apocynaceae 1): 12. 2002.
Lianescent shrub. Riverine forest, miombo
woodland. 980–1530 m.
Distribution: MP, NL, SL—Gereau 4291
Carvalhoa campanulata K. Schum.; Omino,
F.T.E.A. (Apocynaceae 1): 54. 2002.
Shrub. Montane forest. 1450–1980 m.
Distribution: MR, RM—Mwasumbi 16349
Diplorhynchus condylocarpon (Müll. Arg.) Pichon;
Omino, F.T.E.A. (Apocynaceae 1): 56. 2002.
Tree. Woodland. 1200–1400 m.
Distribution: MP—Kayombo 2408
Holarrhena pubescens Wall. ex G. Don; Omino,
F.T.E.A. (Apocynaceae 1): 92. 2002.
Shrub. Lakeshore forest and bushland. 474–
480 m.
Distribution: SL—Gereau 3650
Landolphia buchananii (Hallier f.) Stapf; Omino,
F.T.E.A. (Apocynaceae 1): 19. 2002.
Liana or shrub. Riverine forest, montane forest.
840–1950 m.
Distribution: MP, RM, SL—Gereau 3687
Rauvolfia caffra Sond.; Omino, F.T.E.A. (Apocynaceae 1): 61. 2002.
LakeNyasa_5-18.indd 21
21
Tree. Riverine forest, pond margins in montane
grassland. 1290–1840 m.
Distribution: MP, SL—Gereau 3922
Rauvolfia mannii Stapf; Omino, F.T.E.A. (Apocynaceae 1): 63. 2002.
Shrub. Montane forest. 1750 m.
Distribution: RM—Kindeketa 2976
Saba comorensis (Bojer ex A. DC.) Pichon; Omino,
F.T.E.A. (Apocynaceae 1): 34. 2002.
Liana. Lakeshore shrubland. 474–480 m.
Distribution: SL—Mwasumbi 16603
Tabernaemontana stapfiana Britten;
F.T.E.A. (Apocynaceae 1): 49. 2002.
Tree. Riverine forest. 2000 m.
Distribution: MP—J. Lovett 4990
Omino,
Tabernaemontana ventricosa Hochst. ex A. DC.;
Omino, F.T.E.A. (Apocynaceae 1): 49. 2002.
Tree. Riverine forest. 890–1160 m.
Distribution: SL—Gereau 3698
AQUIFOLIACEAE
1 genus and 1 species
Ilex mitis (L.) Radlk. var. mitis; Verdcourt, F.T.E.A.
(Aquifoliaceae): 3. 1968.
Tree or shrub. Montane forest. 1450–2340 m.
Distribution: KP, MR, NL, RM, SL—Mwasumbi 16303
ARACEAE
2 genera and 2 species
Amorphophallus abyssinicus (A. Rich.) N. E. Br.
subsp. unyikae (Engl. & Gehrm.) Ittenb. ex
Govaerts & Frodin; Ittenbach, 2003: 81.
Terrestrial herb. Edges of cultivation, woodland. 1250–1300 m.
Distribution: MP—J. Lovett 3692
Culcasia falcifolia Engl.; Mayo, F.T.E.A. (Araceae): 18. 1985.
Epiphyte. Montane forest. 1750 m.
Distribution: RM—Abeid 2898
ARALIACEAE
3 genera and 6 species
Cussonia arborea Hochst. ex A. Rich.; Bamps,
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22
1974: 104.
Tree. Montane woodland, miombo woodland.
910–2000 m.
Distribution: MP, RM, SL—Gereau 4069
Cussonia spicata Thunb.; Bamps, 1974: 115.
Tree. Montane forest, forest/woodland transition. 1620–2100 m.
Distribution: RM, SL—Gereau 4060
Polyscias fulva (Hiern) Harms; Bamps, 1974: 122.
Tree. Riverine forest, montane forest. 1350–
1900 m.
Distribution: MP, RM—Kayombo 1076
Schefflera lukwangulensis (Tennant) Bernardi;
Bamps, 1974: 132.
Tree. Montane forest. 1610 m.
Distribution: RM—Kibure 1399
Schefflera myriantha (Baker) Drake; Bamps,
1974: 133.
Liana or tree. Montane forest. 1730–2015 m.
Distribution: MR, NL—Gereau 4145
Schefflera stolzii Harms; Bamps, 1974: 135.
Liana. Montane forest. 1740–2360 m.
Distribution: MR, RM, SL—Stolz 2044 (type:
BM, K, MO, P, WAG), Abeid 2929
ARECACEAE
1 genus and 1 species
Phoenix reclinata Jacq.; Dransfield, F.T.E.A. (Palmae): 15. 1986.
Tree. Montane grassland and forest edges, riverine forest. 890–2120 m.
Distribution: SL—Gereau 4344
ARISTOLOCHIACEAE
1 genus and 1 species
Aristolochia hockii De Wild.; Verdcourt, F.T.E.A.
(Aristolochiaceae): 7. 1986.
Terrestrial herb. Miombo woodland, abandoned cultivation. 1200–1250 m.
Distribution: MP—J. Lovett 4212
ASCLEPIADACEAE
18 genera and 38 species
Asclepias randii S. Moore; Brown, F.T.A. 4(1): 351.
LakeNyasa_5-18.indd 22
Gereau et al.
1902.
Terrestrial herb. Montane grassland. 2150 m.
Distribution: SL—Gereau 5139
Asclepias sp.
Terrestrial herb. Montane grassland. 2800 m.
Distribution: KP—Goldblatt 8215
Aspidoglossum angustissimum (K. Schum.) Bullock; Bullock, 1952: 418.
Terrestrial herb. Wooded grassland. 1250 m.
Distribution: MP—J. Lovett 4221
Brachystelma kituloense Goyder; Goyder, 1990:
729.
Terrestrial herb. Montane grassland. 2250–
2800 m.
Distribution: KP—Brummitt 18092 (type:
DSM, EA, K, MO, PRE), Richards 18501
Ceropegia claviloba Werderm.; Huber, 1957: 153.
Twining vine. Montane forest. 1700–1880 m.
Distribution: NL, RM—Gereau 4224
Ceropegia meyeri-johannis Engl.; Huber, 1957:
154.
Twining vine. Montane forest. 2380 m.
Distribution: KP—Gereau 3129
Ceropegia papillata N. E. Br. var. cordiloba (Werderm.) H. Huber; Huber, 1957: 152.
Twining vine. Montane forest/woodland transition. 2140 m.
Distribution: SL—Gereau 4002
Ceropegia racemosa N. E. Br.; Huber, 1957: 97.
Terrestrial herb. Miombo woodland. 1200 m.
Distribution: MP—J. Lovett 3969
Ceropegia stenantha K. Schum.; Huber, 1957:
123.
Terrestrial herb. Rocky montane grassland.
2700 m.
Distribution: KP—Gereau 2837
Cryptolepis oblongifolia (Meisn.) Schltr.; Brown,
F.T.A. 4(1): 249. 1902.
Shrub. Woodland, riverine forest. 1200–1400
m.
Distribution: MP—Lovett 4054
Cynanchum rungweense Bullock; Liede, 1996a:
335.
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Lake Nyasa Climatic Region
Floristic Checklist
23
Herbaceous climber. Montane forest, tree
clumps in grassland. 2200–2380 m.
Distribution: KP, MR—Gereau 3724
Liana. Montane forest, forest/woodland transition. 1800–2140 m.
Dregea abyssinica K. Schum.; Bullock, F.W.T.A. 2:
97. 1963.
Liana. Riverine forest. 1300–1400 m.
Distribution: MP—Kayombo 1025
Raphionacme aff. lanceolata Schinz; Brown, F.T.A.
4(1): 274. 1902.
Glossostelma carsonii (N. E. Br.) Bullock; Bullock,
1952: 415.
Terrestrial herb. Miombo woodland. 1780–
1790 m.
Distribution: SL—Gereau 3876
Gomphocarpus physocarpus E. Mey.; Bullock,
F.W.T.A. 2: 92. 1963.
Terrestrial herb. 2870 m.
Distribution: KP—WCS SHCP 373
Gongronema angolense (N. E. Br.) Bullock; Bullock, F.W.T.A. 2: 98. 1963.
Herbaceous climber.
Distribution: MR—Stolz 548
Pachycarpus concolor E. Mey.; Smith, 1988: 411.
Terrestrial herb. Montane grassland. 2550 m.
Distribution: KP—Goldblatt 8219
Pachycarpus distinctus (N. E. Br.) Bullock; Bullock, 1953b: 335.
Terrestrial herb. Disturbed woodland. 1910 m.
Distribution: SL—Gereau 4073
Pachycarpus goetzei (K. Schum.) Bullock; Bullock, 1953b: 338.
Terrestrial herb. Miombo woodland. 1700 m.
Distribution: NL—Bidgood 154
Pachycarpus lineolatus (Decne.) Bullock; Bullock,
1953b: 333.
Terrestrial herb. Miombo woodland. 1200–
1600 m.
Distribution: MP—J. Lovett 5038
Pachycarpus spurius (N. E. Br.) Bullock; Bullock,
1953b: 338.
Terrestrial herb. Woodland, abandoned cultivation. 1200–1250 m.
Distribution: MP—J. Lovett 4028
Periploca linearifolia Quart.-Dill. & A. Rich.; Venter, 2003: 103.
LakeNyasa_5-18.indd 23
Distribution: RM, SL—Suleiman 104
Terrestrial herb. Miombo woodland. 1200–
1250 m.
Distribution: MP—J. Lovett 3972
Raphionacme longituba E. A. Bruce; Bruce, 1937:
419.
Terrestrial herb. Miombo woodland. 1200 m.
Distribution: MP—J. Lovett 3881
Raphionacme welwitschii Schltr. & Rendle; Brown,
F.T.A. 4(1): 273. 1902.
Terrestrial herb. Grassland, woodland. 1200–
1600 m.
Distribution: MP—Gereau 3493
Schizoglossum alpestre K. Schum.; Brown, F.T.A.
4(1): 364. 1902.
Terrestrial herb. Montane grassland. 2800 m.
Distribution: KP—Goldblatt 8197
Schizoglossum sp.
Terrestrial herb. Montane grassland. 2800 m.
Distribution: KP—Goldblatt 8195
Stathmostelma gigantiflorum K. Schum.; Bullock,
1953a: 52.
Terrestrial herb. Woodland. 1250 m.
Distribution: MP—J. Lovett 3879
Stathmostelma spectabile (N. E. Br.) Schltr.; Bullock, 1953a: 56.
Terrestrial herb. Riverine forest edge. 1160 m.
Distribution: SL—Gereau 3697
Tacazzea apiculata Oliv.; Venter, 2003: 106.
Liana. Montane forest edges. 2300 m.
Distribution: MR, NL—Magogo 2400
Tacazzea conferta N. E. Br.; Venter, 2003: 106.
Liana. Montane forest edges. 2280–2650 m.
Distribution: MP, MR, NL—Iwarsson 1159
Tylophora aff. apiculata K. Schum.; Brown, F.T.A.
4(1): 407. 1903.
Herbaceous climber. Montane forest. 1850 m.
Distribution: RM—Bidgood 82
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24
Tylophora conspicua N. E. Br.; Brown, F.T.A. 4(1):
405. 1903.
Herbaceous climber. Riverine forest. 1320 m.
Distribution: MP—Gereau 6122
Tylophora erubescens (Liede & Meve) Liede;
Liede, 1996b: 206.
Herbaceous climber. Montane forest. 1920 m.
Distribution: NL—Gereau 4212
Xysmalobium kaessneri S. Moore; Moore, 1908:
295.
Terrestrial herb. Miombo woodland. 1200 m.
Distribution: MP—J. Lovett 3944
Xysmalobium reticulatum N. E. Br.; Brown, F.T.A.
4(2): 303. 1902.
Terrestrial herb. Miombo woodland. 1790 m.
Distribution: SL—Gereau 3909
Xysmalobium undulatum (L.) W. T. Aiton; Goyder,
2003: 131.
Terrestrial herb. Montane grassland. 2080–
2140 m.
Distribution: SL—Suleiman 115
Xysmalobium sp.
Terrestrial herb. Riverbanks in montane grassland. 2560 m.
Distribution: KP—Gereau 3178
Xysmalobium sp.
Terrestrial herb. Thin soil among boulders.
2600–2800 m.
Distribution: KP, MP—Gereau 3502
Gereau et al.
Distribution: MP, RM, SL—J. Lovett 4809
Asparagus flagellaris (Kunth) Baker; Sebsebe,
F.T.E.A. (Asparagaceae): 10. 2006.
Terrestrial herb. Abandoned cultivation,
miombo woodland. 1250–1400 m.
Distribution: MP—J. Lovett 3750
Asparagus laricinus Burch.; Sebsebe, F.T.E.A.
(Asparagaceae): 5. 2006.
Shrub. Montane bushland, scree slopes. 1980–
2300 m.
Distribution: MR, SL—Mwasumbi 16545
Asparagus setaceus (Kunth) Jessop; Sebsebe,
F.T.E.A. (Asparagaceae): 9. 2006.
Liana. Montane forest and thickets, riverine
forest. 1500–2230 m.
Distribution: NL, RM, SL—Harder 1504
Asparagus virgatus Baker; Sebsebe, F.T.E.A.
(Asparagaceae): 4. 2006.
Shrub. Scree slopes. 2060–2120 m.
Distribution: SL—Gereau 4445
ASPHODELACEAE
4 genera and 8 species
Aloe leedalii S. Carter; Carter, F.T.E.A. (Aloaceae):
9. 1994.
Terrestrial herb. Rocky slopes. 2950 m.
Distribution: KP—Leedal 5503
1 genus and 7 species
Aloe nuttii Baker; Carter, F.T.E.A. (Aloaceae): 9.
1994.
Terrestrial herb. Rocky slopes, montane grassland. 2500–2800 m.
Distribution: KP, MP—P. Lovett 93
Asparagus africanus Lam. var. puberulus (Baker)
Sebsebe; Sebsebe, F.T.E.A. (Asparagaceae): 8.
2006.
Shrub. Edges of cultivation. 1300 m.
Distribution: MP—J. Lovett 3548
Bulbine abyssinica A. Rich.; Whitehouse, F.T.E.A.
(Asphodelaceae): 11. 2002.
Terrestrial herb. Woodland, montane grassland, rocky ridges. 1200–2850 m.
Distribution: KP, MP, SL—Abeid 2792
Asparagus asparagoides (L.) Druce; Sebsebe,
F.T.E.A. (Asparagaceae): 19. 2006.
Liana. Montane forest. 2300 m.
Distribution: MR—Gereau 4581
Kniphofia goetzei Engl.; Whitehouse, F.T.E.A.
(Asphodelaceae): 4. 2002.
Terrestrial herb. Montane forest. 1980–2500
m.
Distribution: MR, NL, SL—Gereau 4629
ASPARAGACEAE
Asparagus buchananii Baker; Sebsebe, F.T.E.A.
(Asparagaceae): 14. 2006.
Shrub. Miombo woodland. 1300–2140 m.
LakeNyasa_5-18.indd 24
Kniphofia grantii Baker; Whitehouse, F.T.E.A.
(Asphodelaceae): 7. 2002.
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Lake Nyasa Climatic Region
Floristic Checklist
Terrestrial herb. Woodland, montane grassland. 1400–2500 m.
Distribution: KP, MP—J. Lovett 5040
Kniphofia paludosa Engl.; Whitehouse, F.T.E.A.
(Asphodelaceae): 9. 2002.
Terrestrial or aquatic herb. Wet places in
montane grassland, in running water of streams,
swamps. 2300–2770 m.
Distribution: KP—Goetze 1278 (type: B
[destroyed]), Gereau 3182
Kniphofia princeae (A. Berger) Marais; Whitehouse, F.T.E.A. (Asphodelaceae): 6. 2002.
Terrestrial herb. Montane grassland. 2500 m.
Distribution: MP—P. Lovett 89
Trachyandra saltii (Baker) Oberm.; Whitehouse,
F.T.E.A. (Asphodelaceae): 16. 2002.
Terrestrial herb. Montane grassland. 2150–
2800 m.
Distribution: KP, SL—Gereau 5140
ASTERACEAE
55 genera and 139 species
Acanthospermum hispidum DC.; Beentje & Hind,
F.T.E.A. (Compositae 3): 766. 2005.
Terrestrial herb. Disturbed montane forest.
Distribution: RM—WCS SHCP 233
Acmella caulirhiza Delile; Beentje & Hind, F.T.E.A.
(Compositae 3): 730. 2005.
Terrestrial herb. Shallow ponds in montane
grassland, riverine forest, edges of cultivation.
1200–2080 m.
Distribution: MP, RM, SL—Gereau 3917
Adenostemma caffrum DC.; Smith, F.T.E.A. (Compositae 3): 821. 2005.
Terrestrial herb. 2080–2140 m.
Distribution: SL—Suleiman 124
Adenostemma mauritianum DC.; Smith, F.T.E.A.
(Compositae 3): 824. 2005.
Terrestrial herb. Montane forest, Eucalyptus
plantations, riverine brushland. 1660–2300 m.
Distribution: MR, RM, SL—Abeid 2886
Ageratinastrum polyphyllum (Baker) Mattf.; Jeffrey & Beentje, F.T.E.A. (Compositae 1): 276.
2000.
Terrestrial herb. Montane grassland. 2488 m.
LakeNyasa_5-18.indd 25
25
Distribution: SL—Suleiman 167
Ageratum conyzoides L.; Smith, F.T.E.A. (Compositae 3): 827. 2005.
Terrestrial herb. Open woodland, edges of cultivation. 1250–1400 m.
Distribution: MP, RM—J. Lovett 3687
Ageratum houstonianum Mill. var. houstonianum;
Smith, F.T.E.A. (Compositae 3): 830. 2005.
Terrestrial herb.
Distribution: RM—Semsei 1553
Anisopappus chinensis (L.) Hook. & Arn. subsp.
buchwaldii (O. Hoffm.) S. Ortiz, Paiva & Rodr.
Oubiña; Beentje, F.T.E.A. (Compositae 2): 344.
2002.
Terrestrial herb. Montane forest edges, grassland. 1980–2500 m.
Distribution: KP, MP, MR, NL, RM—Gereau
4593
Artemisia afra Jacq. ex Willd.; Beentje, F.T.E.A.
(Compositae 2): 529. 2002.
Shrub or woody-based herb. Montane grassland. 1960–2840 m.
Distribution: KP, RM—Suleiman 29
Aspilia mossambicensis (Oliv.) Wild; Beentje &
Hind, F.T.E.A. (Compositae 3): 751. 2005.
Shrub. Riverine forest edges, bamboo forest,
grassland, woodland. 1200–2000 m.
Distribution: MP, NL, SL—Kayombo 2354
Aspilia pluriseta Schweinf.; Beentje & Hind,
F.T.E.A. (Compositae 3): 749. 2005.
Terrestrial herb. Edges of cultivation, woodland. 1200–1400 m.
Distribution: MP—J. Lovett 3847
Aster tansaniensis W. Lippert; Beentje, F.T.E.A.
(Compositae 2): 470. 2002.
Terrestrial herb. Montane grassland. 2230–
2500 m.
Distribution: KP, RM, SL—Gereau 3591
Athrixia rosmarinifolia (Sch. Bip. ex Walp.) Oliv.
& Hiern var. rosmarinifolia; Beentje, F.T.E.A.
(Compositae 2): 456. 2002.
Terrestrial herb. Bushland. 2360 m.
Distribution: KP, MP—Myembe 56
Athrixia subsimplex Brenan; Beentje, F.T.E.A.
(Compositae 2): 456. 2002.
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26
Terrestrial herb. Montane grassland. 1800–
2700 m.
Distribution: KP, MP, MR—Machaga 25
Berkheya echinacea (Harv.) O. Hoffm. ex Burtt
Davy subsp. polyacantha (Baker) Roessler;
Halliday & Wilmot-Dear, F.T.E.A. (Compositae
1): 291. 2000.
Terrestrial herb. Montane grassland. 1940–
2840 m.
Distribution: KP, MP, RM, SL—Suleiman 24
Gereau et al.
Bidens steppia (Steetz) Sherff; Mesfin, F.T.E.A.
(Compositae 3): 801. 2005.
Terrestrial herb. Grassland, miombo woodland,
abandoned cultivation. 1250–1800 m.
Distribution: MP—J. Lovett 4619
Blumea axillaris (Lam.) DC.; Beentje, F.T.E.A.
(Compositae 2): 319. 2002.
Terrestrial herb. Riverine forest, woodland.
560–1250 m.
Distribution: MP, SL—J. Lovett 3761
Berkheya zeyheri Oliv. & Hiern; Halliday &
Wilmot-Dear, F.T.E.A. (Compositae 1): 293.
2000.
Terrestrial herb. Rocky mountain slopes.
2250–2800 m.
Distribution: KP, MP—Goldblatt 8202
Bothriocline inyangana N. E. Br. var. amplexicaulis (Muschl.) C. Jeffrey; Jeffrey & Beentje,
F.T.E.A. (Compositae 1): 147. 2000.
Shrub. Montane forest edges, grassland, roadsides. 1950–2240 m.
Distribution: MR, RM—Gereau 4520
Bidens flagellata (Sherff) Mesfin; Mesfin, F.T.E.A.
(Compositae 3): 796. 2005.
Terrestrial herb. Montane grassland. 2140–
2150 m.
Distribution: SL—Suleiman 155
Bothriocline inyangana N. E. Br. var. inyangana;
Jeffrey & Beentje, F.T.E.A. (Compositae 1):
147. 2000.
Shrub. Abandoned cultivation. 2000 m.
Distribution: MP—P. Lovett 252
Bidens kilimandscharica (O. Hoffm.) Sherff; Mesfin, F.T.E.A. (Compositae 3): 783. 2005.
Terrestrial herb. Montane forest. 2110 m.
Distribution: MR—Gereau 4566
Bothriocline mbalensis (Wild & G. V. Pope) C. Jeffrey; Jeffrey & Beentje, F.T.E.A. (Compositae
1): 145. 2000.
Terrestrial herb. Miombo woodland. 1550–
1800 m.
Distribution: MP—Gereau 7002
Bidens ochracea (O. Hoffm.) Sherff; Mesfin,
F.T.E.A. (Compositae 3): 789. 2005.
Terrestrial herb. Rocky montane grassland.
2740 m.
Distribution: KP—Abeid 3075
Bidens pilosa L.; Mesfin, F.T.E.A. (Compositae 3):
806. 2005.
Terrestrial herb. Riverine forest, montane forest, woodland. 1160–1900 m.
Distribution: MP, NL, RM, SL—Gereau 3678
Bidens pinnatipartita (O. Hoffm.) Wild; Mesfin,
F.T.E.A. (Compositae 3): 783. 2005.
Soft-wooded shrub. Montane forest edges,
montane grassland. 1940–2300 m.
Distribution: MP, MR, SL—Gereau 4657
Bidens schimperi Sch. Bip. ex Walp.; Mesfin,
F.T.E.A. (Compositae 3): 803. 2005.
Terrestrial herb. Grassland, montane forest.
1300–2400 m.
Distribution: MP, RM—Mwasumbi 16268
LakeNyasa_5-18.indd 26
Bothriocline pectinata (O. Hoffm.) Wild & G. V.
Pope; Jeffrey & Beentje, F.T.E.A. (Compositae
1): 155. 2000.
Shrub or woody-based herb. Montane forest.
1750–2354 m.
Distribution: RM, SL—Abeid 2925
Brachythrix glomerata (Mattf.) C. Jeffrey; Jeffrey
& Beentje, F.T.E.A. (Compositae 1): 278. 2000.
Terrestrial herb. Miombo woodland. 1200–
2140 m.
Distribution: MP, SL—Gereau 4409
Brachythrix stolzii (S. Moore) Wild & G. V. Pope;
Jeffrey & Beentje, F.T.E.A. (Compositae 1):
277. 2000.
Terrestrial herb. Montane forest edges, grassland. 1980–2525 m.
Distribution: MR, NL, RM—Stolz 2595 (type:
BM, K, MO), Abeid 2830
Carduus nyassanus (S. Moore) R. E. Fr. subsp.
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Floristic Checklist
27
kikuyorum (R. E. Fr.) C. Jeffrey; Jeffrey &
Beentje, F.T.E.A. (Compositae 1): 49. 2000.
Terrestrial herb. Montane forest, grassland.
2400 m.
Distribution: KP, MR, RM—Richards 6461
Jeffrey & Beentje, F.T.E.A. (Compositae 3):
610. 2005.
Terrestrial herb. Riverine forest, montane forest. 1210–1670 m.
Distribution: RM, SL—Gereau 4298
Chrysanthemoides monilifera (L.) Norl. subsp.
septentrionalis Norl.; Beentje, F.T.E.A. (Compositae 2): 535. 2002.
Shrub. Scree slopes. 2060–2120 m.
Distribution: SL—Gereau 4438
Crassocephalum montuosum (S. Moore) MilneRedh.; Jeffrey & Beentje, F.T.E.A. (Compositae
3): 602. 2005.
Terrestrial herb. Grassland, montane forest.
1700–2100 m.
Distribution: MR, RM—Richards 6551
Cineraria deltoidea Sond.; Jeffrey & Beentje,
F.T.E.A. (Compositae 3): 563. 2005.
Terrestrial herb. Montane grassland, edges of
cultivation. 2200–2550 m.
Distribution: KP, MP, MR—Gereau 3154
Cirsium buchwaldii O. Hoffm.; Jeffrey & Beentje,
F.T.E.A. (Compositae 1): 54. 2000.
Terrestrial herb. Stream banks in montane
grassland. 2400–2590 m.
Distribution: KP—Suleiman 13
Conyza gouanii (L.) Willd.; Beentje, F.T.E.A.
(Compositae 2): 493. 2002.
Terrestrial herb. Open woodland, riverine forest. 1650–1850 m.
Distribution: SL—Gereau 4047
Conyza limosa O. Hoffm.; Beentje, F.T.E.A. (Compositae 2): 508. 2002.
Terrestrial herb. Pond margins, stream banks,
seasonally waterlogged grassland. 1250–2550 m.
Distribution: KP, MP, SL—Gereau 3913
Conyza pyrrhopappa Sch. Bip. ex A. Rich.;
Beentje, F.T.E.A. (Compositae 2): 499. 2002.
Shrub. Woodland, riverine forest. 1300–1780
m.
Distribution: MP—J. Lovett 4874
Conyza stricta Willd.; Beentje, F.T.E.A. (Compositae 2): 500. 2002.
Terrestrial herb. Riverine forest. 1500 m.
Distribution: SL—Gereau 3950
Conyza tigrensis Oliv. & Hiern; Beentje, F.T.E.A.
(Compositae 2): 497. 2002.
Terrestrial herb. Montane grassland. 2230–
2650 m.
Distribution: KP, MR, SL—Gereau 3770
Crassocephalum crepidioides (Benth.) S. Moore;
LakeNyasa_5-18.indd 27
Crassocephalum rubens (Juss. ex Jacq.) S. Moore
var. rubens; Jeffrey & Beentje, F.T.E.A. (Compositae 3): 609. 2005.
Terrestrial herb. Disturbed forest, woodland,
grassland, cultivation. 1100–2100 m.
Distribution: MP, MR, RM—J. Lovett 3782
Crassocephalum rubens (Juss. ex Jacq.) S. Moore
var. sarcobasis (DC.) C. Jeffrey & Beentje; Jeffrey & Beentje, F.T.E.A. (Compositae 3): 609.
2005.
Terrestrial herb. Miombo woodland. 1400 m.
Distribution: MP—P. Lovett 110
Crassocephalum vitellinum (Benth.) S. Moore; Jeffrey & Beentje, F.T.E.A. (Compositae 3): 606.
2005.
Terrestrial herb. Montane forest, woodland,
roadsides. 1400–2300 m.
Distribution: MP, RM—J. Lovett 4882
Crepis newii Oliv. & Hiern subsp. oliveriana
(Kuntze) C. Jeffrey & Beentje; Jeffrey &
Beentje, F.T.E.A. (Compositae 1): 70. 2000.
Terrestrial herb. Montane grassland, miombo
woodland, cultivation, roadsides. 1740–2600 m.
Distribution: KP, MP, MR, RM, SL—Suleiman
26
Dichrocephala chrysanthemifolia (Blume) DC. var.
chrysanthemifolia; Beentje, F.T.E.A. (Compositae 2): 461. 2002.
Terrestrial herb. Montane grassland, montane
forest edges. 2300–2640 m.
Distribution: KP, MP, NL—Gereau 4642
Dichrocephala integrifolia (L. f.) Kuntze subsp.
integrifolia; Beentje, F.T.E.A. (Compositae 2):
459. 2002.
Terrestrial herb. Coffee plantation, old cultivation, riverine forest, montane forest edges.
5/31/12 3:59 PM
28
1250–2420 m.
Distribution: MP, RM—J. Lovett 5008
Dicoma anomala Sond.; Jeffrey & Beentje, F.T.E.A.
(Compositae 1): 14. 2000.
Terrestrial herb. Grassland, woodland. 1200–
2500 m.
Distribution: KP, MP—Kayombo 2431
Dicoma kirkii Harv. subsp. vaginata (O. Hoffm.)
G. V. Pope; Jeffrey & Beentje, F.T.E.A. (Compositae 1): 17. 2000.
Terrestrial herb. Wooded grassland. 980 m.
Distribution: SL—Kibure 1484
Echinops giganteus A. Rich.; Jeffrey & Beentje,
F.T.E.A. (Compositae 1): 39. 2000.
Terrestrial herb. Montane grassland, cultivation. 1650–2500 m.
Distribution: MP, MR—P. Lovett 494
Echinops lanatus C. Jeffrey & Mesfin; Jeffrey &
Beentje, F.T.E.A. (Compositae 1): 39. 2000.
Terrestrial herb. Montane grassland. 2400–
2850 m.
Distribution: KP, MR, NL—Bidgood 121 (type:
K, NHT), Abeid 2860
Elephantopus scaber L. subsp. plurisetus (O.
Hoffm.) Philipson; Jeffrey & Beentje, F.T.E.A.
(Compositae 1): 280. 2000.
Terrestrial herb. Woodland, grassland, riverine
forest. 1200–2040 m.
Distribution: MP, MR, SL—Gereau 3964
Emilia abyssinica (Sch. Bip. ex A. Rich.) C. Jeffrey var. abyssinica; Jeffrey & Beentje, F.T.E.A.
(Compositae 3): 575. 2005.
Terrestrial herb. Miombo woodland. 1200 m.
Distribution: MP—J. Lovett 4064
Emilia caespitosa Oliv.; Jeffrey & Beentje, F.T.E.A.
(Compositae 3): 581. 2005.
Terrestrial herb. Miombo woodland. 1475 m.
Distribution: MP—Kayombo 2398
Emilia coccinea (Sims) G. Don; Jeffrey & Beentje,
F.T.E.A. (Compositae 3): 583. 2005.
Terrestrial herb. Abandoned cultivation, woodland. 1200–1800 m.
Distribution: MP, RM—Kayombo 1001
Emilia integrifolia Baker; Jeffrey & Beentje,
F.T.E.A. (Compositae 3): 592. 2005.
LakeNyasa_5-18.indd 28
Gereau et al.
Terrestrial herb. Grassland, montane forest.
1250–2660 m.
Distribution: KP, MP, MR—Gereau 3107
Emilia tenera (O. Hoffm.) C. Jeffrey; Jeffrey &
Beentje, F.T.E.A. (Compositae 3): 592. 2005.
Terrestrial herb. Montane grassland. 1800–
2740 m.
Distribution: KP, RM, SL—J. Lovett 774
Erythrocephalum longifolium Benth. ex Oliv.; Jeffrey & Beentje, F.T.E.A. (Compositae 1): 27.
2000.
Terrestrial herb. Wooded grassland, miombo
woodland, abandoned cultivation. 1250–1900 m.
Distribution: MP, MR—Gereau 7050
Felicia boehmii O. Hoffm.; Beentje, F.T.E.A. (Compositae 2): 475. 2002.
Terrestrial herb. Miombo woodland. 1400–
1475 m.
Distribution: MP—Kayombo 2397
Galinsoga parviflora Cav.; Beentje & Hind,
F.T.E.A. (Compositae 3): 763. 2005.
Terrestrial herb. Riverine forest, edges of cultivation, weed of bean fields. 1250–1500 m.
Distribution: MP, SL—Gereau 3961
Gerbera piloselloides (L.) Cass.; Jeffrey & Beentje,
F.T.E.A. (Compositae 1): 12. 2000.
Terrestrial herb. Montane forest remnants,
woodland. 1320–1840 m.
Distribution: MP, SL—Gereau 3549
Gerbera viridifolia (DC.) Sch. Bip. subsp. viridifolia; Jeffrey & Beentje, F.T.E.A. (Compositae
1): 10. 2000.
Terrestrial herb. Montane grassland. 1810–
2700 m.
Distribution: KP, NL, SL—Taylor 8527
Guizotia scabra (Vis.) Chiov. subsp. scabra;
Beentje & Hind, F.T.E.A. (Compositae 3): 771.
2005.
Soft-wooded shrub. Montane bushland. 2280
m.
Distribution: MR—Mwasumbi 16168
Gutenbergia cordifolia Benth. ex Oliv. var. cordifolia; Jeffrey & Beentje, F.T.E.A. (Compositae
1): 124. 2000.
Terrestrial herb. Roadsides. 474–480 m.
Distribution: SL—Mwasumbi 16588
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Lake Nyasa Climatic Region
Floristic Checklist
Haplocarpha thunbergii Less.; Halliday & WilmotDear, F.T.E.A. (Compositae 1): 287. 2000.
Terrestrial herb. Montane grassland. 2320–
2570 m.
Distribution: KP, SL—Gereau 3997
Helichrysum abietinum O. Hoffm.; Beentje,
F.T.E.A. (Compositae 2): 417. 2002.
Subshrub. Montane grassland, heathland.
2550–2960 m.
Distribution: KP, RM—Goetze 1151 (type: BR,
K), Abeid 3008
Helichrysum aureum (Houtt.) Merr. var. monocephalum (DC.) Hilliard; Beentje, F.T.E.A.
(Compositae 2): 437. 2002.
Terrestrial herb. Montane grassland. 2640 m.
Distribution: KP, NL—Gereau 4648
Helichrysum foetidum (L.) Moench; Beentje,
F.T.E.A. (Compositae 2): 434. 2002.
Terrestrial herb. Montane grassland, montane
forest/woodland transition, riverine forest. 2170–
2900 m.
Distribution: KP, MR, SL—Gereau 3999
Helichrysum forskahlii (J. F. Gmel.) Hilliard & B.
L. Burtt var. forskahlii; Beentje, F.T.E.A. (Compositae 2): 415. 2002.
Terrestrial herb or shrub. Heathland, montane
grassland, woodland. 1400–2770 m.
Distribution: KP, MP, MR, RM, SL—Mwasumbi
16259
Helichrysum globosum Sch. Bip.; Beentje, F.T.E.A.
(Compositae 2): 417. 2002.
Terrestrial herb. Montane grassland. 2550 m.
Distribution: KP—Abeid 3108
Helichrysum herbaceum (Andrews) Sweet; Beentje,
F.T.E.A. (Compositae 2): 434. 2002.
Terrestrial herb. Montane grassland. 2120–
2700 m.
Distribution: KP, NL, RM, SL—Gereau 4322
Helichrysum kalandanum Lisowski; Beentje,
F.T.E.A. (Compositae 2): 427. 2002.
Terrestrial herb. Montane forest, montane
grassland, woodland. 1890–2360 m.
Distribution: KP, MP, MR, NL, SL—Gereau
4221
Helichrysum kirkii Oliv. & Hiern; Beentje, F.T.E.A.
(Compositae 2): 429. 2002.
LakeNyasa_5-18.indd 29
29
Terrestrial herb. Montane grassland, miombo
woodland. 1280–2840 m.
Distribution: KP, MP, SL—Suleiman 19
Helichrysum milne-redheadii Brenan vel sp. aff.;
Lisowski, 1989: 166.
Subshrub. Montane grassland. 1940 m.
Distribution: SL—Gereau 4658
Helichrysum nitens Oliv. & Hiern; Beentje, F.T.E.A.
(Compositae 2): 433. 2002.
Terrestrial herb. Montane grassland. 2500–
2740 m.
Distribution: KP, MP—Gereau 3203
Helichrysum nudifolium (L.) Less.; Beentje,
F.T.E.A. (Compositae 2): 420. 2002.
Terrestrial herb. Montane grassland. 2230–
2700 m.
Distribution: NL, SL—Gereau 3588
Helichrysum odoratissimum (L.) Sweet; Beentje,
F.T.E.A. (Compositae 2): 410. 2002.
Terrestrial herb. Montane grassland, montane
shrubland. 1760–2360 m.
Distribution: MR, NL—Gereau 3769
Helichrysum quartinianum A. Rich.; Beentje,
F.T.E.A. (Compositae 2): 438. 2002.
Terrestrial herb. Miombo woodland. 1820 m.
Distribution: SL—Gereau 3907
Helichrysum schimperi (Sch. Bip. ex A. Rich.)
Moeser; Beentje, F.T.E.A. (Compositae 2): 418.
2002.
Straggling terrestrial herb. Montane bushland,
montane forest. 1780–2240 m.
Distribution: MR, RM—Mwasumbi 16507
Helichrysum setosum Harv.; Beentje, F.T.E.A.
(Compositae 2): 436. 2002.
Terrestrial herb. Pyrethrum fields. 2200 m.
Distribuition: MR, RM—Gereau 4524
Helichrysum tillandsiifolium O. Hoffm.; Beentje,
F.T.E.A. (Compositae 2): 412. 2002.
Subshrub. Rocky montane grassland, woodland. 1920–2740 m.
Distribution: KP, NL—Wiland 149A
Hypericophyllum compositarum Steetz; Beentje &
Hind, F.T.E.A. (Compositae 3): 720. 2005.
Terrestrial herb. Miombo woodland. 1600 m.
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Gereau et al.
Distribution: MP—P. Lovett 229
Hypericophyllum elatum (O. Hoffm.) N. E. Br.;
Beentje & Hind, F.T.E.A. (Compositae 3): 720.
2005.
Terrestrial herb. Riverine forest. 1200 m.
Distribution: MP—J. Lovett 4401
Inula mannii (Hook. f.) Oliv. & Hiern; Beentje,
F.T.E.A. (Compositae 2): 326. 2002.
Terrestrial herb or suffrutex. Montane forest.
2354–2500 m.
Distribution: MR, RM—Kayombo 1107
Inula rungwensis Beentje; Beentje, F.T.E.A. (Compositae 2): 328. 2002.
Terrestrial herb. Montane grassland. 2600 m.
Distribution: KP—J. Lovett 3655 (type: K, MO)
Inula shirensis Oliv.; Beentje, F.T.E.A. (Compositae
2): 323. 2002.
Terrestrial herb. Woodland, grassland. 1250–
1400 m.
Distribution: MP—Kayombo 983
Inula stolzii Mattf.; Beentje, F.T.E.A. (Compositae
2): 326. 2002.
Terrestrial herb. Montane grassland. 2590 m.
Distribution: KP—Abeid 3053
Inula sp.
Terrestrial herb. Miombo woodland. 2000 m.
Distribution: MP—P. Lovett 257
Lactuca inermis Forssk.; Jeffrey & Beentje,
F.T.E.A. (Compositae 1): 83. 2000.
Terrestrial herb. Fallow fields, woodland, montane grassland. 1250–2340 m.
Distribution: MP, MR, RM—Mwasumbi 16528
Laggera brevipes Oliv. & Hiern; Beentje, F.T.E.A.
(Compositae 2:) 355. 2002.
Terrestrial herb. Disturbed woodland. 1200 m.
Distribution: MP—Kayombo 2422
Launaea cornuta (Hochst. ex Oliv. & Hiern) C.
Jeffrey; Jeffrey & Beentje, F.T.E.A. (Compositae
1): 104. 2000.
Terrestrial herb. Montane forest. 1920 m.
Distribution: RM—Ndangalasi SHCP 111
Launaea rarifolia (Oliv. & Hiern) Boulos; Jeffrey
& Beentje, F.T.E.A. (Compositae 1): 101. 2000.
Terrestrial herb. Woodland. 1400 m.
LakeNyasa_5-18.indd 30
Distribution: MP—J. Lovett 3361
Lopholaena dolichopappa (O. Hoffm.) S. Moore;
Jeffrey & Beentje, F.T.E.A. (Compositae 3):
570. 2005.
Terrestrial herb. Montane bushland. 2200–
2300 m.
Distribution: MR—Mwasumbi 16544
Lopholaena trianthema (O. Hoffm.) B. L. Burtt;
Jeffrey & Beentje, F.T.E.A. (Compositae 3):
572. 2005.
Shrub. Woodland. 1400 m.
Distribution: MP—J. Lovett 3331
Melanthera pungens Oliv. & Hiern var. albinervia
(O. Hoffm.) Beentje; Beentje & Hind, F.T.E.A.
(Compositae 3): 742. 2005.
Terrestrial herb. Woodland, riverine forest.
1200–1790 m.
Distribution: MP, SL—Gereau 3902
Microglossa pyrifolia (Lam.) Kuntze; Beentje,
F.T.E.A. (Compositae 2): 482. 2002.
Shrub. Edges of cultivation. 1250 m.
Distribution: MP—J. Lovett 3703
Nidorella microcephala Steetz; Beentje, F.T.E.A.
(Compositae 2): 490. 2002.
Terrestrial herb. Edges of cultivation. 1290 m.
Distribution: MP—Gereau 6114
Osteospermum monocephalum (Oliv. & Hiern)
Norl.; Beentje, F.T.E.A. (Compositae 2): 532.
2002.
Terrestrial herb. Montane grassland and shrubland. 1810–2340 m.
Distribution: MP, SL—Harder 1494
Pleiotaxis pulcherrima Steetz; Jeffrey & Beentje,
F.T.E.A. (Compositae 1): 22. 2000.
Terrestrial herb. Miombo woodland. 1400 m.
Distribution: MP—P. Lovett 103
Schistostephium artemisiifolium Baker; Beentje,
F.T.E.A. (Compositae 2): 526. 2002.
Terrestrial herb. Montane grassland. 2500 m.
Distribution: MP—P. Lovett 5
Senecio doryphorus Mattf.; Jeffrey & Beentje,
F.T.E.A. (Compositae 3): 634. 2005.
Terrestrial herb. Montane grassland. 2700 m.
Distribution: NL—Gereau 4077
5/31/12 3:59 PM
Lake Nyasa Climatic Region
Floristic Checklist
Senecio erubescens Aiton; Jeffrey & Beentje,
F.T.E.A. (Compositae 3): 667. 2005.
Terrestrial herb. Montane grassland, abandoned cultivation. 2600–2700 m.
Distribution: KP—Taylor 8531
Senecio immixtus C. Jeffrey; Jeffrey & Beentje,
F.T.E.A. (Compositae 3): 640. 2005.
Terrestrial herb. Montane grassland. 2570 m.
Distribution: KP—Gereau 3098
Senecio inornatus DC.; Jeffrey & Beentje, F.T.E.A.
(Compositae 3): 633. 2005.
Terrestrial herb. Montane grassland. 2360–
2570 m.
Distribution: KP, MR—Gereau 3782
Senecio cf. mesogrammoides O. Hoffm.; Jeffrey &
Beentje, F.T.E.A. (Compositae 3): 649. 2005.
Terrestrial herb. Montane grassland. 1940 m.
Distribution: SL—Gereau 4692
Senecio microalatus C. Jeffrey; Jeffrey & Beentje,
F.T.E.A. (Compositae 3): 634. 2005.
Terrestrial herb. Stream bank in montane
grassland. 2310 m.
Distribution: MP—Milne-Redhead 10214
(type: K).
Senecio pergamentaceus Baker; Jeffrey & Beentje,
F.T.E.A. (Compositae 3): 643. 2005.
Terrestrial herb. Miombo woodland. 1790 m.
Distribution: SL—Gereau 3880
Senicio purpureus L.; Jeffrey & Beentje, F.T.E.A.
(Compositae 3): 668. 2005.
Terrestrial herb. Montane grassland. 2590 m.
Distribution: KP, RM—Abeid 3048
Senecio ruwenzoriensis S. Moore; Jeffrey & Beentje,
F.T.E.A. (Compositae 3): 643. 2005.
Terrestrial herb. Riverine forest. 1850 m.
Distribution: SL—Gereau 4048
Senecio cf. semiamplexifolius De Wild.; Jeffrey &
Beentje, F.T.E.A. (Compositae 3): 642. 2005.
Terrestrial herb. Montane grassland. 2400–
2840 m.
Distribution: KP—Abeid 2787
Senecio aff. subsessilis Oliv. & Hiern; Jeffrey &
Beentje, F.T.E.A. (Compositae 3): 627. 2005.
Terrestrial herb. Montane forest and grassland.
LakeNyasa_5-18.indd 31
31
2200–2830 m.
Distribution: KP, MR—Kayombo 1102
Senecio syringifolius O. Hoffm.; Jeffrey & Beentje,
F.T.E.A. (Compositae 3): 621. 2005.
Herbaceous climber. Montane forest. 1960–
1980 m.
Distribution: MR—Mwasumbi 16363
Senecio urundensis S. Moore; Jeffrey & Beentje,
F.T.E.A. (Compositae 3): 642. 2005.
Terrestrial herb. Woodland. 1720 m.
Distribution: MP—J. Lovett 3506
Solanecio mannii (Hook. f.) C. Jeffrey; Jeffrey &
Beentje, F.T.E.A. (Compositae 3): 678. 2005.
Tree or shrub. Riverine forest, montane forest.
1600–2240 m.
Distribution: MP, MR, RM—J. Lovett 4831
Sonchus bipontini Asch.; Jeffrey & Beentje,
F.T.E.A. (Compositae 1): 95. 2000.
Terrestrial herb. Montane grassland. 2100 m.
Distribution: MR—Richards 6540
Sonchus luxurians (R. E. Fr.) C. Jeffrey; Jeffrey &
Beentje, F.T.E.A. (Compositae 1): 96. 2000.
Terrestrial herb. Woodland, wet montane
grassland. 1720–2660 m.
Distribution: KP, MP—Abeid 2790
Sphaeranthus randii S. Moore var. randii; Beentje,
F.T.E.A. (Compositae 2): 386. 2002.
Emergent aquatic herb. Seasonal ponds in
montane grassland. 1810–1840 m.
Distribution: SL—Gereau 3916
Stoebe kilimandscharica O. Hoffm.; Beentje,
F.T.E.A. (Compositae 2): 452. 2002.
Shrub. Disturbed montane grassland, heathland. 2500–2890 m.
Distribution: KP, RM—Gereau 3139
Tolpis capensis (L.) Sch. Bip.; Jeffrey & Beentje,
F.T.E.A. (Compositae 1): 66. 2000.
Terrestrial herb. Miombo woodland, clearings in montane forest, roadsides in cultivation.
1790–2350 m.
Distribution: MR, RM, SL—Gereau 3881
Tridax procumbens L.; Beentje & Hind, F.T.E.A.
(Compositae 3): 764. 2005.
Terrestrial herb. Roadsides, abandoned cultivation. 474–1250 m.
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Gereau et al.
Distribution: MP, SL—Gereau 3788
Vernonia adoensis Sch. Bip. ex Walp.; Jeffrey &
Beentje, F.T.E.A. (Compositae 1): 245. 2000.
Shrub. Woodland, thickets near cultivation,
disturbed lower montane forest. 1300–1350 m.
Distribution: MP, RM—Gereau 6096
Vernonia alticola G. V. Pope; Jeffrey & Beentje,
F.T.E.A. (Compositae 1): 205. 2000.
Terrestrial herb. Montane grassland. 2700 m.
Distribution: KP—Taylor 8526
Distribution: MP, MR, SL—Gereau 4691
Vernonia meiostephana C. Jeffrey; Jeffrey &
Beentje, F.T.E.A. (Compositae 1): 211. 2000.
Terrestrial herb. Old cultivation. 1250 m.
Distribution: MP—J. Lovett 4548
Vernonia melleri Oliv. & Hiern var. superba (O.
Hoffm.) C. Jeffrey; Jeffrey & Beentje, F.T.E.A.
(Compositae 1): 233. 2000.
Terrestrial herb. Woodland, edges of cultivation. 1250 m.
Distribution: MP—J. Lovett 4114
Vernonia calvoana (Hook. f.) Hook. f. subsp.
leucocalyx (O. Hoffm.) C. Jeffrey; Jeffrey &
Beentje, F.T.E.A. (Compositae 1): 238. 2000.
Shrub. Disturbed montane forest, forest edges.
1980–2100 m.
Distribution: MR, RM—Gereau 4627
Vernonia musofensis S. Moore var. miamensis (S.
Moore) G. V. Pope; Jeffrey & Beentje, F.T.E.A.
(Compositae 1): 258. 2000.
Terrestrial herb. Miombo woodland. 1600 m.
Distribution: MP—P. Lovett 238
Vernonia calyculata S. Moore; Jeffrey & Beentje,
F.T.E.A. (Compositae 1): 207. 2000.
Terrestrial herb. Miombo woodland. 1790 m.
Distribution: SL—Gereau 3900
Vernonia myriantha Hook. f.; Jeffrey & Beentje,
F.T.E.A. (Compositae 1): 195. 2000.
Shrub. Riverine forest. 1500–2250 m.
Distribution: MP, MR, RM—J. Lovett 4692
Vernonia exsertiflora Baker var. exsertiflora; Jeffrey & Beentje, F.T.E.A. (Compositae 1): 181.
2000.
Shrub. Bushland, woodland, grassland. 1250–
1400 m.
Distribution: MP—J. Lovett 4580
Vernonia perrottetii Sch. Bip. ex Walp.; Jeffrey &
Beentje, F.T.E.A. (Compositae 1): 256. 2000.
Terrestrial herb. Grassland. 1100 m.
Distribution: MP—Carter 2474
Vernonia glabra (Steetz) Vatke var. glabra; Jeffrey
& Beentje, F.T.E.A. (Compositae 1): 230. 2000.
Terrestrial herb. Woodland. 1250 m.
Distribution: MP—J. Lovett 4566
Vernonia glabra (Steetz) Vatke var. laxa (Steetz)
Brenan; Jeffrey & Beentje, F.T.E.A. (Compositae 1): 230. 2000.
Terrestrial herb. Grassland. 1100 m.
Distribution: MP—Carter 2466
Vernonia jelfiae S. Moore var. albida G. V. Pope;
Jeffrey & Beentje, F.T.E.A. (Compositae 1):
253. 2000.
Terrestrial herb. Woodland, grassland. 1250–
1300 m.
Distribution: MP—J. Lovett 4617
Vernonia karaguensis Oliv. & Hiern; Jeffrey &
Beentje, F.T.E.A. (Compositae 1): 200. 2000.
Shrub or terrestrial herb. Montane forest edges,
montane grassland, woodland. 1250–2110 m.
LakeNyasa_5-18.indd 32
Vernonia praemorsa Muschl.; Jeffrey & Beentje,
F.T.E.A. (Compositae 1): 224. 2000.
Terrestrial herb. Montane grassland. 2250 m.
Distribution: MP—J. Lovett 4848
Vernonia pteropoda Oliv. & Hiern; Jeffrey &
Beentje, F.T.E.A. (Compositae 1): 184. 2000.
Terrestrial herb. Montane forest. 1760 m.
Distribution: MR, RM—Kibure 1380
Vernonia suprafastigiata Klatt vel sp. aff.; Jeffrey
& Beentje, F.T.E.A. (Compositae 1): 198. 2000.
Shrub. Miombo woodland. 1300 m.
Distribution: MP—J. Lovett 4808
Vernonia syringifolia O. Hoffm. vel sp. aff.; Jeffrey
& Beentje, F.T.E.A. (Compositae 1): 190. 2000.
Straggling shrub. Montane forest edges. 2110
m.
Distribution: MR—Gereau 4552
Vernonia aff. thomsoniana Oliv. & Hiern; Jeffrey &
Beentje, F.T.E.A. (Compositae 1): 192. 2000.
Shrub. Grassland. 1300 m.
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Lake Nyasa Climatic Region
Floristic Checklist
Distribution: MP—P. Lovett 381
Vernonia tolypophora Mattf.; Jeffrey & Beentje,
F.T.E.A. (Compositae 1): 241. 2000.
Shrub. Roadsides, edges of cultivation, woodland, montane grassland. 1300–2200 m.
Distribution: MP, MR, RM, SL—Gereau 4772
Vernonia ugandensis S. Moore; Jeffrey & Beentje,
F.T.E.A. (Compositae 1): 261. 2000.
Terrestrial herb. Miombo woodland. 1530 m.
Distribution: SL—Gereau 4375
Vernonia usafuensis O. Hoffm.; Jeffrey & Beentje,
F.T.E.A. (Compositae 1): 232. 2000.
Terrestrial herb. Montane grassland, montane
forest edges. 1800–2600 m.
Distribution: MP, MR, RM—Goetze 1039 (type:
BR, K, P), P. Lovett 492
BALANOPHORACEAE
1 genus and 1 species
Sarcophyte sanguinea Sparrm. subsp. piriei
(Hutch.) B. Hansen; Hansen, F.T.E.A. (Balanophoraceae): 3. 1993.
Parasitic terrestrial herb. Woodland, Eucalyptus plantation, roadsides in coffee plantation.
1310–1500 m.
Distribution: MP—Schmidt 1128
BALSAMINACEAE
1 genus and 15 species
Impatiens assurgens Baker f.; Grey-Wilson,
F.T.E.A. (Balsaminaceae): 57. 1982.
Terrestrial herb. Woodland, montane grassland. 1300–2500 m.
Distribution: MP, MR, SL—Gereau 4756
Impatiens austrotanzanica Grey-Wilson; GreyWilson, F.T.E.A. (Balsaminaceae): 66. 1982.
Terrestrial herb. Montane forest, forest/woodland transition, montane grassland. 1550–2360 m.
Distribution: MR, RM, SL—Kibure 1414
Impatiens confusa Grey-Wilson subsp. confusa;
Grey-Wilson, F.T.E.A. (Balsaminaceae): 45.
1982.
Terrestrial herb. Montane forest. 1750–1980
m.
Distribution: MR, NL, RM—Jones & Pócs
6331/P (type: K), Mwasumbi 16359
LakeNyasa_5-18.indd 33
33
Impatiens cribbii (Grey-Wilson) Grey-Wilson;
Grey-Wilson, F.T.E.A. (Balsaminaceae): 65.
1982.
Terrestrial herb. Near streams in montane forest, montane grassland. 2380–2590 m.
Distribution: KP—Cribb 10764 (type: DSM,
EA, K), Goldblatt 8238
Impatiens eryaleia Launert subsp. eryaleia; GreyWilson, F.T.E.A. (Balsaminaceae): 30. 1982.
Terrestrial herb. Montane forest. 1900–2350
m.
Distribution: NL, RM, SL—Gereau 4206
Impatiens eryaleia Launert subsp. mbeyaensis
Grey-Wilson; Grey-Wilson, F.T.E.A. (Balsaminaceae): 30. 1982.
Terrestrial herb. Montane forest. 1560–1710
m.
Distribution: MR, RM—Cribb 11312 (type:
DSM, EA, K), Mwasumbi 16285
Impatiens flammea Gilg; Grey-Wilson, F.T.E.A.
(Balsaminaceae): 71. 1982.
Terrestrial herb. Montane forest. 2460 m.
Distribution: NL, RM—Goetze 1147 (type: B),
Gereau 4115
Impatiens gomphophylla Baker f.; Grey-Wilson,
F.T.E.A. (Balsaminaceae): 64. 1982.
Terrestrial herb. Montane grassland, tree
clumps, montane forest edges, woodland. 1300–
2760 m.
Distribution: KP, MP, MR, RM—Abeid 2826
Impatiens hochstetteri Warb. subsp. hochstetteri;
Grey-Wilson, F.T.E.A. (Balsaminaceae): 46.
1982.
Terrestrial herb. Montane forest, heathland.
1560–2810 m.
Distribution: KP, NL, RM—Mwasumbi 16294
Impatiens hydrogetonoides Launert subsp.
hydrogetonoides; Grey-Wilson, F.T.E.A. (Balsaminaceae): 39. 1982.
Terrestrial herb. Montane forest. 2000 m.
Distribution: MP—J. Lovett 3851
Impatiens hydrogetonoides Launert subsp. kituloensis Grey-Wilson; Grey-Wilson, F.T.E.A.
(Balsaminaceae): 39. 1982.
Terrestrial herb. Stream banks, montane grassland, montane forest edges. 2380–2960 m.
Distribution: KP—Cribb 10771 (type: DSM,
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34
EA, K), Goldblatt 8237
Impatiens leedalii Grey-Wilson; Grey-Wilson,
F.T.E.A. (Balsaminaceae): 54. 1982.
Terrestrial herb. Montane forest. 2380–2775
m.
Distribution: KP—Cribb 10770 (type: DSM,
EA, K), Goldblatt 8263
Impatiens polyantha Gilg; Grey-Wilson, F.T.E.A.
(Balsaminaceae): 54. 1982.
Terrestrial herb. Montane grassland, roadsides.
1900–2600 m.
Distribution: MP, MR, RM, SL—Gereau 3499
Impatiens polyantha Gilg × austrotanzanica
Grey-Wilson; Grey-Wilson, F.T.E.A. (Balsaminaceae): 57. 1982.
Terrestrial herb. Montane forest. 1960–2640
m.
Distribution: MR, NL—Gereau 4634
Impatiens polyantha Gilg × rosulata Grey-Wilson;
Grey-Wilson, F.T.E.A. (Balsaminacaeae): 55.
1982.
Terrestrial herb. Montane grassland with tree
clumps. 2360 m.
Distribution: MR—Gereau 3743
Impatiens pseudozombensis Grey-Wilson; GreyWilson, F.T.E.A. (Balsaminaceae): 36. 1982.
Terrestrial herb. Montane forest. 1920–2240
m.
Distribution: MR, SL—Gereau 5129
Impatiens rosulata Grey-Wilson; Grey-Wilson,
F.T.E.A. (Balsaminaceae): 57. 1982.
Terrestrial herb. Montane grassland. 2400–
2740 m.
Distribution: KP, NL—Cribb 10761 (type:
DSM, EA, K), Suleiman 23
Impatiens rubromaculata Warb. subsp. grandiflora Grey-Wilson; Grey-Wilson, F.T.E.A.
(Balsaminaceae): 35. 1982.
Terrestrial herb. Montane forest. 2000–2354
m.
Distribution: MP, MR, NL, RM—Richards 9774
(type: B, K), Mwasumbi 16214
Impatiens volkensii Warb. subsp. volkensii; Frimodt-Møller & Grey-Wilson, 1999: 180.
Terrestrial herb or soft-wooded shrub. Stream
LakeNyasa_5-18.indd 34
Gereau et al.
banks in montane forest, riverine brushland.
1660–1790 m.
Distribution: NL, RM—Gereau 4168
BASELLACEAE
1 genus and 1 species
Basella alba L.; Verdcourt, F.T.E.A. (Basellaceae):
2. 1968.
Twining herb. Montane forest. 1870–1980 m.
Distribution: MR, RM—Mwasumbi 16362
BEGONIACEAE
1 genus and 5 species
Begonia johnstonii Oliv. ex Hook. f.; Sands &
Beentje, F.T.E.A. (Begoniaceae): 12. 2006.
Terrestrial herb. Montane grassland. 2140–
2150 m.
Distribution: SL—Suleiman 151
Begonia meyeri-johannis Engl.; Plana, F.T.E.A.
(Begoniaceae): 4. 2006.
Epilithic or epiphytic herb. Montane forest.
1500–2500 m.
Distribution: MR, RM, SL—Mwasumbi 16223
Begonia princeae Gilg; Sands & Beentje, F.T.E.A.
(Begoniaceae): 14. 2006.
Terrestrial herb. Woodland, abandoned cultivation. 1200–1350 m.
Distribution: MP—J. Lovett 5003
Begonia aff. rumpiensis Kupicha; Kupicha, Fl.
Zamb. 4: 502. 1978.
Terrestrial herb. Rocks in riverine forest. 1160–
1320 m.
Distribution: MP, SL—Gereau 3689
Begonia sutherlandii Hook. f.; Sands & Beentje,
F.T.E.A. (Begoniaceae): 17. 2006.
Epiphytic and epilithic herb. Montane forest
and clearings, riverine forest. 1320–2380 m.
Distribution: KP, MP, RM, SL—Gereau 3603
BERBERIDACEAE
1 genus and 1 species
Berberis holstii Engl.; Polhill, F.T.E.A. (Berberidaceae): 1. 1966.
Shrub. Heathland. 2460 m.
5/31/12 3:59 PM
Lake Nyasa Climatic Region
Floristic Checklist
35
Distribution: KP, MP, MR, RM, SL—Suleiman
Distribution: RM—Geilinger s.n.
14
BIGNONIACEAE
3 genera and 3 species
Kigelia africana (Lam.) Benth.; Bidgood et al.,
F.T.E.A. (Bignoniaceae): 43. 2006.
Tree. Open woodland. 1350 m.
Distribution: MP—Lovett 4769
Markhamia obtusifolia (Baker) Sprague; Bidgood
et al., F.T.E.A. (Bignoniaceae): 35. 2006.
Tree or shrub. Woodland, edges of cultivation,
riverine forest. 830–1400 m.
Distribution: MP, SL—Kayombo 2449
Tecomaria capensis (Thunb.) Spach subsp. nyassae (Oliv.) Brummitt; Bidgood et al., F.T.E.A.
(Bignoniaceae): 25. 2006.
Tree or shrub. Wooded grassland, riverine forest, montane forest. 1500–2750 m.
Distribution: MP, MR, NL, RM, SL—Suleiman
132
BOMBACACEAE
1 genus and 1 species
Adansonia digitata L.; Beentje, F.T.E.A. (Bombacaceae): 4. 1989.
Tree. Secondary forest on lakeshore. 474 m.
Distribution: SL—Gereau 3666
BORAGINACEAE
7 genera and 10 species
Cordia africana Lam.; Verdcourt, F.T.E.A. (Boraginaceae): 31. 1991.
Tree. Woodland, old cultivation. 1250–1400 m.
Distribution: MP—Kayombo 955
Cynoglossum amplifolium Hochst ex A. DC.; Verdcourt, F.T.E.A. (Boraginaceae): 104. 1991.
Terrestrial herb. Montane grassland. 2400–
2700 m.
Distribution: KP, NL, RM—Abeid 2796
Cynoglossum coeruleum A. DC. subsp. johnstonii
(Baker) Verdc.; Verdcourt, F.T.E.A. (Boraginaceae): 109. 1991.
Terrestrial herb. Montane grassland, woodland.
1200–2400 m.
LakeNyasa_5-18.indd 35
Cynoglossum lanceolatum Forssk.; Verdcourt,
F.T.E.A. (Boraginaceae): 106. 1991.
Terrestrial herb. Miombo woodland. 1530 m.
Distribution: SL—Gereau 4377
Ehretia cymosa Thonn.; Verdcourt, F.T.E.A. (Boraginaceae): 37. 1991.
Tree or shrub. Riverine forest, montane forest.
1800–1825 m.
Distribution: MP, RM—Kayombo 1112
Heliotropium ovalifolium Forssk.; Verdcourt,
F.T.E.A. (Boraginaceae): 75. 1991.
Terrestrial herb. Edges of cultivation. 1290 m.
Distribution: MP—Gereau 6115
Lithospermum afromontanum Weim.; Verdcourt,
F.T.E.A. (Boraginaceae): 77. 1991.
Terrestrial herb. Montane forest edges, montane grassland and heathland. 2080–2830 m.
Distribution: KP, MP, NL, SL—Suleiman 136
Myosotis vestergrenii Stroh; Verdcourt, F.T.E.A.
(Boraginaceae): 87. 1991.
Terrestrial herb. Montane grassland. 2750 m.
Distribution: KP, NL—Sturtz 232
Trichodesma physaloides (Fenzl) A. DC.; Verdcourt, F.T.E.A. (Boraginaceae): 100. 1991.
Terrestrial herb. Abandoned cultivation, grassland. 1250–2250 m.
Distribution: MP—J. Lovett 4608
Trichodesma zeylanicum (Burm. f.) R. Br.; Verdcourt, F.T.E.A. (Boraginaceae): 92. 1991.
Terrestrial herb. Woodland. 1200–1250 m.
Distribution: MP—Kayombo 2418
BRASSICACEAE
2 genera and 2 species
Capsella bursa-pastoris (L.) Medik.; Jonsell,
F.T.E.A. (Cruciferae): 29. 1982.
Terrestrial herb. Cultivated fields. 2280 m.
Distribution: MR—Mwasumbi 16162
Cardamine africana L.; Jonsell, F.T.E.A. (Cruciferae): 38. 1982.
Terrestrial herb. Montane forest. 2320–2460
5/31/12 3:59 PM
36
Gereau et al.
Distribution: SL—Gereau 3808
m.
Distribution: NL, SL—Gereau 4117
BUDDLEJACEAE
2 genera and 5 species
Buddleja axillaris Willd. ex Roem. & Schult.;
Leeuwenberg, 1979: 27.
Liana. Riverine forest. 1210 m.
Distribution: SL—Gereau 4261
Buddleja pulchella N. E. Br. vel sp. aff.; Leeuwenberg, 1979: 68.
Tree. Montane forest. 2120 m.
Distribution: NL—Bracebridge 73
Buddleja salviifolia (L.) Lam.; Leeuwenberg,
1979: 77.
Shrub or tree. Montane forest. 1780–2500 m.
Distribution: KP, MP, RM—P. Lovett 478
Nuxia congesta R. Br. ex Fresen.; Bruce & Lewis,
F.T.E.A. (Loganiaceae): 44. 1960.
Tree. Riverine forest, tree clumps in montane
grassland. 2200–2360 m.
Distribution: MP, MR, RM—Gereau 3764
Nuxia floribunda Benth.; Bruce & Lewis, F.T.E.A.
(Loganiaceae): 43. 1960.
Tree. Riverine forest, montane forest edges.
474–2350 m.
Distribution: MR, NL, RM, SL—Mwasumbi
16609
BURSERACEAE
1 genus and 4 species
Commiphora africana (A. Rich.) Engl. var.
rubriflora (Engl.) Wild; Gillett, F.T.E.A. (Burseraceae): 47. 1991.
Tree. Riverine forest. 1250–1290 m.
Distribution: MP—Gereau 6111
Commiphora eminii Engl. subsp. zimmermannii
(Engl.) J. B. Gillett; Gillett, F.T.E.A. (Burseraceae): 57. 1991.
Tree. Riverine forest. 1210 m.
Distribution: SL—Gereau 4275
Commiphora mollis (Oliv.) Engl.; Gillett, F.T.E.A.
(Burseraceae): 54. 1991.
Tree. Lakeshore forest, riverine forest. 480–580
m.
LakeNyasa_5-18.indd 36
Commiphora mossambicensis (Oliv.) Engl.; Gillett,
F.T.E.A. (Burseraceae): 58. 1991.
Tree. Woodland. 1200–1400 m.
Distribution: MP—Gereau 6116
CAESALPINIACEAE
10 genera and 25 species
Afzelia quanzensis Welw.; Brenan, F.T.E.A. (Leguminosae–Caesalpinioideae): 125. 1967.
Tree. Lakeshore forest. 480 m.
Distribution: SL—Gereau 3653
Bauhinia petersiana Bolle; Brenan, F.T.E.A.
(Leguminosae–Caesalpinioideae): 211. 1967.
Tree. Woodland, riverine forest edges. 1250–
1400 m.
Distribution: MP—Kayombo 968
Bauhinia thonningii Schumach.; Wilczek, 1952:
275.
Tree. Woodland. 1200–1530 m.
Distribution: MP, SL—Gereau 3826
Brachystegia bussei Harms; Hoyle, F.T.E.A.
(Leguminosae–Caesalpinioideae): 170. 1967.
Tree. Miombo woodland. 1300–1400 m.
Distribution: MP—Kayombo 1030
Brachystegia floribunda Benth.; Hoyle, F.T.E.A.
(Leguminosae–Caesalpinioideae): 179. 1967.
Tree. Miombo woodland. 1300–1800 m.
Distribution: MP—P. Lovett 59
Brachystegia longifolia Benth.; Hoyle, F.T.E.A.
(Leguminosae–Caesalpinioideae): 188. 1967.
Shrub or tree. Lakeshore thickets, woodland.
474–1400 m.
Distribution: MP, SL—Mwasumbi 16583
Brachystegia microphylla Harms; Hoyle, F.T.E.A.
(Leguminosae–Caesalpinioideae): 174. 1967.
Tree. Miombo woodland. 1780–1790 m.
Distribution: SL—Gereau 3871
Brachystegia spiciformis Benth.; Hoyle, F.T.E.A.
(Leguminosae–Caesalpinioideae): 167. 1967.
Tree. Miombo woodland, old cultivation.
1250–1790 m.
Distribution: MP, SL—Kayombo 2370
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Lake Nyasa Climatic Region
Floristic Checklist
Brachystegia utilis Hutch. & Burtt Davy; Hoyle,
F.T.E.A.
(Leguminosae–Caesalpinioideae):
172. 1967.
Tree. Miombo woodland. 1300–1400 m.
Distribution: MP—Kayombo 1042
Brachystegia wangermeeana De Wild.; Hoyle,
F.T.E.A.
(Leguminosae–Caesalpinioideae):
192. 1967.
Tree. Miombo woodland. 1780 m.
Distribution: SL—Gereau 5102
Brachystegia sp.
Tree. Miombo woodland. 1780 m.
Distribution: SL—Gereau 5099
Caesalpinia decapetala (Roth) Alston; Brenan,
F.T.E.A. (Leguminosae–Caesalpinioideae): 36.
1967.
Liana. Riverine forest, montane forest. 1400–
1875 m.
Distribution: MP, RM—J. Lovett 4690
Cassia abbreviata Oliv. subsp. abbreviata; Brenan,
F.T.E.A. (Leguminosae–Caesalpinioideae): 59.
1967.
Shrub. Miombo woodland. 980 m.
Distribution: SL—Gereau 5159
Cassia thyrsoidea Brenan; Brenan, F.T.E.A. (Leguminosae–Caesalpinioideae): 62. 1967.
Tree. Riverine forest edges. 1500 m.
Distribution: SL—Gereau 3962
Chamaecrista comosa E. Mey. var. capricornia
(Steyaert) Lock; Lock, Fl. Zamb. 3(2): 143.
2007.
Terrestrial herb. Wooded grassland. 1860 m.
Distribution: SL—Gereau 3930
37
3(2): 138. 2007.
Terrestrial herb. Riverine forest, miombo
woodland, montane grassland. 1250–1940 m.
Distribution: MP, SL—Gereau 4775
Chamaecrista wittei (Ghesq.) Lock; Lock, Fl.
Zamb. 3(2): 141. 2007.
Terrestrial herb. Montane brushland. 1450–
2040 m.
Distribution: RM, SL—Gereau 4750
Julbernardia globiflora (Benth.) Troupin; Brenan,
F.T.E.A.
(Leguminosae–Caesalpinioideae):
147. 1967.
Tree. Miombo woodland. 1180–1400 m.
Distribution: MP, NL—Kayombo 2411
Pterolobium stellatum (Forssk.) Brenan; Brenan,
F.T.E.A. (Leguminosae–Caesalpinioideae): 42.
1967.
Liana. Thicket woodland, riverine forest.
1275–1600 m.
Distribution: MP—Kayombo 2479
Senna occidentalis (L.) Link; Lock, Fl. Zamb.
3(2): 152. 2007.
Shrub. Riverine forest. 1200 m.
Distribution: MP—J. Lovett 4405
Senna septemtrionalis (Viv.) H. S. Irwin & Barneby; Lock, Fl. Zamb. 3(2): 158. 2007.
Shrub. Eroded hillsides, montane forest. 1300–
1810 m.
Distribution: MP, RM, SL—Gereau 4075
Senna singueana (Delile) Lock; Lock, Fl. Zamb.
3(2): 160. 2007.
Tree. Woodland. 1200–1400 m.
Distribution: MP—Kayombo 2405
Chamaecrista mimosoides (L.) Greene; Lock, Fl.
Zamb. 3(2): 136. 2007.
Terrestrial herb. Woodland, grassland,
abandoned cultivation, secondary shrubland.
1250–1760 m.
Distribution: MP, SL—Gereau 4379
Tamarindus indica L.; Brenan, F.T.E.A. (Leguminosae–Caesalpinioideae): 153. 1967.
Tree. Lakeshore forest. 480 m.
Distribution: SL—Gereau 3655
Chamaecrista parva (Steyaert) Lock; Lock, Fl.
Zamb. 3(2): 147. 2007.
Terrestrial herb. Montane grassland. 2400–
2640 m.
Distribution: KP—Suleiman 53
5 genera and 16 species
Chamaecrista stricta E. Mey.; Lock, Fl. Zamb.
LakeNyasa_5-18.indd 37
CAMPANULACEAE
Canarina eminii Asch. ex Schweinf.; Thulin,
F.T.E.A. (Campanulaceae): 4. 1976.
Epiphyte or vine. Montane forest. 1700–2360
m.
Distribution: MR, NL, RM—Abeid 2774
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38
Cyphia brachyandra Thulin; Thulin, F.T.E.A.
(Lobeliaceae): 52. 1984.
Herbaceous climber. Riverine forest. 1500 m.
Distribution: SL—Gereau 4419
Cyphia lasiandra Diels; Thulin, F.T.E.A. (Lobeliaceae): 55. 1984.
Herbaceous climber. Montane grassland.
2080–2400 m.
Distribution: KP, SL—Suleiman 113
Cyphia rupestris E. Wimm.; Thulin, F.T.E.A.
(Lobeliaceae): 53. 1984.
Terrestrial herb. Montane forest edges, montane grassland. 2320–2740 m.
Distribution: KP, MP, SL—Gereau 3972
Cyphia ubenensis Engl.; Thulin, F.T.E.A. (Lobeliaceae): 52. 1984.
Terrestrial herb. Montane grassland.
Distribution: KP—Leedal 5397
Lobelia giberroa Hemsl.; Thulin, F.T.E.A. (Lobeliaceae): 9. 1984.
Giant rosette herb. Riverine forest, woodland,
montane forest edges. 1810–2640 m.
Distribution: KP, MP, MR, NL, RM, SL—
Gereau 4651
Lobelia mildbraedii Engl.; Thulin, F.T.E.A. (Lobeliaceae): 14. 1984.
Giant rosette herb. Swamps and stream banks
in montane grassland. 2500–2560 m.
Distribution: KP—Taylor 8524
Lobelia trullifolia Hemsl. subsp. trullifolia; Thulin,
F.T.E.A. (Lobeliaceae): 21. 1984.
Terrestrial herb. Marshes in montane grassland. 2550 m.
Distribution: KP—Abeid 3116
Monopsis stellarioides (C. Presl) Urb. subsp.
schimperiana (Urb.) Thulin; Thulin, F.T.E.A.
(Lobeliaceae): 47. 1984.
Terrestrial herb. Riverine forest. 1850 m.
Distribution: SL—Gereau 4043
Gereau et al.
Wahlenberbia capillacea (L. f.) A. DC. subsp.
tenuior (Engl.) Thulin; Thulin, F.T.E.A. (Campanulaceae): 15. 1976.
Terrestrial herb. Montane grassland, lava flows.
2400–2740 m.
Distribution: KP, RM—Suleiman 34
Wahlenbergia capitata (Baker) Thulin; Thulin,
F.T.E.A. (Campanulaceae): 19. 1976.
Terrestrial herb. Woodland. 1475 m.
Distribution: MP—Kayombo 2394
Wahlenbergia denticulata (Burch.) A. DC.; Thulin, F.T.E.A. (Campanulaceae): 15. 1976.
Terrestrial herb. Montane grassland. 2500 m.
Distribution: KP—J. Lovett 779
Wahlenbergia huttonii (Sond.) Thulin; Thulin,
F.T.E.A. (Campanulaceae): 17. 1976.
Terrestrial herb. Montane grassland. 2488–
2740 m.
Distribution: KP, SL—Suleiman 169
Wahlenbergia krebsii Cham. subsp. arguta (Hook.
f.) Thulin; Thulin, F.T.E.A. (Campanulaceae):
13. 1976.
Terrestrial herb. Rocky montane grassland.
2560 m.
Distribution: KP, RM—Gereau 3184
Wahlenbergia virgata Engl.; Thulin, F.T.E.A.
(Campanulaceae): 9. 1976.
Terrestrial herb. Woodland, wet grassland.
1300–2000 m.
Distribution: KP, MP, NL—J. Lovett 4701
CANNABACEAE
1 genus and 1 species
Cannabis sativa L.; Verdcourt, F.T.E.A. (Cannabaceae): 1. 1975.
Terrestrial herb. Edges of cultivated fields, disturbed montane grassland. 2200–2780 m.
Distribution: KP, MR—Abeid 2808
CANNACEAE
Wahlenbergia abyssinica (Hochst. ex A. Rich.)
Thulin subsp. abyssinica; Thulin, F.T.E.A.
(Campanulaceae): 25. 1976.
Terrestrial herb. Montane grassland. 2400–
2610 m.
Distribution: KP—Suleiman 33
LakeNyasa_5-18.indd 38
1 genus and 1 species
Canna indica L.; Lock, F.T.E.A. (Musaceae [Cannaceae]): 7. 1993.
Terrestrial herb. Montane forest. 1790 m.
Distribution: RM—WCS SHCP 34
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Lake Nyasa Climatic Region
Floristic Checklist
CAPPARACEAE
5 genera and 5 species
Boscia salicifolia Oliv.; Elffers et al., F.T.E.A.
(Capparidaceae): 52. 1964.
39
Cerastium octandrum Hochst. ex A. Rich.; Turrill,
F.T.E.A. (Caryophyllaceae): 21. 1956.
Terrestrial herb. Montane grassland. 2570–
2600 m.
Distribution: KP—Gereau 3112
Tree. Miombo woodland. 1200 m.
Distribution: MP—J. Lovett 3807
Capparis tomentosa Lam.; Elffers et al., F.T.E.A.
(Capparidaceae): 62. 1964.
Liana. Riverine forest. 1290 m.
Distribution: MP—Gereau 6113
Cleome monophylla L.; Elffers et al., F.T.E.A.
(Capparidaceae): 5. 1964.
Terrestrial herb. Woodland, roadsides. 1300–
1600 m.
Distribution: MP, SL—Gereau 3951
Maerua triphylla A. Rich. var. pubescens
(Klotzsch) DeWolf; Elffers et al., F.T.E.A.
(Capparidaceae): 45. 1964.
Shrub. Lakeshore forest. 474 m.
Distribution: SL—Gereau 3662
Ritchiea albersii Gilg; Elffers et al., F.T.E.A. (Capparidaceae): 21. 1964.
Shrub or tree. Montane forest. 1790 m.
Distribution: RM—WCS SHCP 72
Dianthus zeyheri Sond. subsp. zeyheri; Hooper,
1959: 47.
Terrestrial herb. Montane grassland. 2120–
2488 m.
Distribution: SL—Gereau 4310
Note: New to East Africa, from South Africa
(Transvaal Drakensberg).
Silene burchellii Otth ex DC.; Turrill, F.T.E.A.
(Caryophyllaceae): 33. 1956.
Terrestrial herb. Montane grassland. 2250–
2640 m.
Distribution: KP, MP—Suleiman 46
CECROPIACEAE
1 genus and 1 species
Myrianthus holstii Engl.; Berg, F.T.E.A. (Moraceae
[Cecropiaceae]): 88. 1989.
Tree. Riverine forest, montane forest. 1350–
2040 m.
Distribution: MP, RM, SL—Gereau 4721
CELASTRACEAE
CARICACEAE
1 genus and 1 species
Cylicomorpha parviflora Urb.; Hemsley, F.T.E.A.
(Caricaceae): 2. 1958.
Tree. Montane forest. 1670 m.
Distribution: NL—Bracebridge 57
CARYOPHYLLACEAE
3 genera and 5 species
Cerastium dregeanum Fenzl; Sonder, 1860: 130.
Terrestrial herb. Rocky mountain slopes. 2800
m.
Distribution: KP—Goldblatt 8216
Note: New to East Africa, from South Africa.
Cerastium indicum Wight & Arn.; Turrill, F.T.E.A.
(Caryophyllaceae): 19. 1956.
Terrestrial herb. Montane forest edges. 2320 m.
Distribution: SL—Gereau 3970
LakeNyasa_5-18.indd 39
4 genera and 9 species
Catha edulis Forssk.; Robson & Blakelock, F.T.E.A.
(Celastraceae): 25. 1994.
Tree. Riverine forest, scree slopes, montane forest. 1500–2230 m.
Distribution: MR, RM, SL—Mwasumbi 16176
Gymnosporia buchananii Loes.; Jordaan & van
Wyk, 2006: 521.
Tree. Riverine forest. 840 m.
Distribution: SL—Kibure 1459
Gymnosporia harveyana Loes. subsp. stolzii (N.
Robson) Jordaan; Jordaan & van Wyk, 2004:
844.
Shrub, sometimes scandent. Montane forest.
2040–2100 m.
Distribution: RM, SL—Gereau 4715
Gymnosporia heterophylla (Eckl. & Zeyh.) Loes.;
Jordaan & van Wyk, 2006: 520.
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40
Shrub or tree. Montane forest, bamboo thicket,
woodland. 1810–2380 m.
Distribution: KP, MP, NL, RM, SL—Gereau
4128
Gymnosporia livingstonei Jordaan; Jordaan & van
Wyk, 2004: 851.
Subshrub. Wooded grassland. 1860 m.
Distribution: SL—Gereau 3925 (type: K, MO)
Gymnosporia senegalensis (Lam.) Loes.; Jordaan
& van Wyk, 2006: 519.
Tree or shrub. Woodland, montane forest edges.
1200–1780 m.
Distribution: MP, RM, SL—Kayombo 2436
Maytenus acuminata (L. f.) Loes.; Robson &
Blakelock, F.T.E.A. (Celastraceae): 19. 1994.
Tree, shrub. Montane forest, forest edges.
1960–2260 m.
Distribution: MR, RM, SL—Mwasumbi 16348
Gereau et al.
CHRYSOBALANACEAE
2 genera and 3 species
Hirtella zanzibarica Oliv. subsp. zanzibarica;
Prance, 2002: 994.
Tree. Riverine forest. 500–980 m.
Distribution: SL—Kibure 1488
Parinari curatellifolia Planch. ex Benth.; Prance &
White, 1988: 112.
Tree. Protea woodland, miombo woodland,
coffee plantations. 1300–2000 m.
Distribution: MP, SL—Gereau 3853
Parinari excelsa Sabine; Prance & White, 1988:
112.
Tree. Riverine forest, montane forest, bamboo
thicket. 1620–2100 m.
Distribution: MP, NL, RM, SL—Gereau 3928
CLUSIACEAE
Maytenus undata (Thunb.) Blakelock; Robson &
Blakelock, F.T.E.A. (Celastraceae): 18. 1994.
Tree or shrub, sometimes scandent. Montane
forest, bamboo forest, forest edges, riverine forest.
1300–2700 m.
Distribution: MP, NL, RM—Kayombo 2476
4 genera and 9 species
Mystroxylon aethiopicum (Thunb.) Loes.; Robson
& Blakelock, F.T.E.A. (Celastraceae): 21. 1994.
Tree, shrub. Riverine forest, woodland, montane forest. 980–2320 m.
Distribution: MP, NL, RM, SL—Suleiman 137
Garcinia kingaensis Engl.; Bamps et al., F.T.E.A.
(Guttiferae): 22. 1978.
Shrub or tree. Riverine forest, montane forest.
980–2000 m.
Distribution: MP, RM, SL—Gereau 5152
CHENOPODIACEAE
1 genus and 3 species
Chenopodium ambrosioides L.; Brenan, F.T.E.A.
(Chenopodiaceae): 10. 1954.
Terrestrial herb. Riverine forest. 1350 m.
Distribution: MP—J. Lovett 4755
Chenopodium procerum Hochst. ex Moq.; Brenan,
F.T.E.A. (Chenopodiaceae): 11. 1954.
Terrestrial herb. Cultivated fields. 2280 m.
Distribution: MR—Mwasumbi 16163
Chenopodium schraderianum Schult.; Brenan,
F.T.E.A. (Chenopodiaceae): 12. 1954.
Terrestrial herb. Cultivated fields. 1700 m.
Distribution: MP—J. Lovett 3494
LakeNyasa_5-18.indd 40
Garcinia buchananii Baker; Bamps et al., F.T.E.A.
(Guttiferae): 24. 1978.
Tree. Riverine forest, woodland. 474–1400 m.
Distribution: MP, SL—Mwasumbi 16591
Garcinia volkensii Engl.; Bamps et al., F.T.E.A.
(Guttiferae): 23. 1978.
Tree. Montane forest. 1920 m.
Distribution : NL—Bracebridge 49
Harungana madagascariensis Lam. ex Poir.;
Bamps et al., F.T.E.A. (Guttiferae): 32. 1978.
Shrub or tree. Montane forest. 2040 m.
Distribution: RM—WCS SHCP 157
Hypericum conjungens N. Robson; Bamps et al.,
F.T.E.A. (Guttiferae): 30. 1978.
Shrub. Montane forest. 2130 m.
Distribution: MR—Mwasumbi 16191A
Hypericum peplidifolium A. Rich.; Bamps et al.,
F.T.E.A. (Guttiferae): 31. 1978.
Terrestrial herb. Montane grassland, riverine
forest. 1300–2590 m.
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Lake Nyasa Climatic Region
Floristic Checklist
Distribution: KP, MP, MR, RM, SL—Gereau
4430
Hypericum quartinianum A. Rich.; Bamps et al.,
F.T.E.A. (Guttiferae): 29. 1978.
Shrub. Riverine forest. 1650 m.
Distribution: MP—J. Lovett 4922
Hypericum revolutum Vahl subsp. revolutum;
Bamps et al., F.T.E.A. (Guttiferae): 28. 1978.
Shrub. Montane forest, rocky mountain slopes,
stream banks in montane grassland. 2340–2800
m.
Distribution: KP, RM—Abeid 3003
Psorospermum febrifugum Spach var. febrifugum;
Milne-Redhead, F.T.E.A. (Hypericaceae): 18.
1953.
Shrub. Miombo woodland, lakeside forest and
thickets. 474–1450 m.
Distribution: MP, SL—Gereau 5153
Psorospermum febrifugum Spach var. ferrugineum
(Hook. f.) Keay & Milne-Redh.; Milne-Redhead, F.T.E.A. (Hypericaceae): 18. 1953.
Shrub. Miombo woodland, grassland. 1470–
1950 m.
Distribution: MP, MR, SL—Gereau 3855
41
Combretum pentagonum M. A. Lawson; Wickens,
F.T.E.A. (Combretaceae): 57. 1973.
Liana. Riverine forest, lakeshore shrubland.
474–1160 m.
Distribution: SL—Gereau 3705
Combretum psidioides Welw. subsp. psidioides;
Wickens, F.T.E.A. (Combretaceae): 37. 1973
Tree. Woodland. 1300–1400 m.
Distribution: MP—Kayombo 977
Combretum zeyheri Sond.; Wickens, F.T.E.A.
(Combretaceae): 31. 1973.
Tree or shrub. Grassland. 1300–1400 m.
Distribution: MP—Kayombo 986
Terminalia kaiseriana F. Hoffm.; Wickens, F.T.E.A.
(Combretaceae): 84. 1973.
Tree. Open woodland. 1300–1400 m.
Distribution: MP—Kayombo 1002
Terminalia sambesiaca Engl. & Diels; Wickens,
F.T.E.A. (Combretaceae): 91. 1973.
Tree. Lakeshore forest. 480 m.
Distribution: SL—Gereau 3652
COMMELINACEAE
5 genera and 16 species
COLCHICACEAE
2 genera and 2 species
Gloriosa superba L. var. superba; Hoenselaar,
F.T.E.A. (Colchicaceae): 6. 2005.
Terrestrial herb or herbaceous climber. Riverine forest, montane forest, woodland, edges of
cultivation. 1200–2140 m.
Distribution: MP, MR, NL, SL—Suleiman 120
Wurmbea tenuis (Hook. f.) Baker subsp. goetzei
(Engl.) B. Nord.; Hoenselaar, F.T.E.A. (Colchicaceae): 14. 2005.
Terrestrial herb. Montane grassland, open
woodland. 1720–2800 m.
Distribution: KP, MP—Gereau 3497
COMBRETACEAE
2 genera and 6 species
Combretum molle R. Br. ex G. Don; Wickens,
F.T.E.A. (Combretaceae): 33. 1973.
Tree. Woodland, riverine forest. 1200–1680 m.
Distribution: MP, SL—Kayombo 2406
LakeNyasa_5-18.indd 41
Aneilema aequinoctiale (P. Beauv.) G. Don;
Ensermu & Faden, 1997: 351.
Terrestrial herb. Riverine forest. 1650 m.
Distribution: MP—J. Lovett 4921
Aneilema dispermum Brenan; Brenan, F.W.T.A. 3:
31. 1968.
Terrestrial herb. Montane forest. 2000–2150
m.
Distribution: MR, RM—Mwasumbi 16231
Aneilema hirtum A. Rich.; Ensermu & Faden,
1997: 360.
Terrestrial herb. Woodland, riverine forest.
1250–1800 m.
Distribution: MP—P. Lovett 222
Aneilema pedunculosum C. B. Clarke; Faden,
1991: 147.
Terrestrial herb. Riverine forest. 1500 m.
Distribution: SL—Gereau 4403
Commelina africana L. subsp. africana; Faden,
F.T.E.A. (Commelinaceae) [in press].
5/31/12 3:59 PM
42
Terrestrial herb. Montane grassland, cultivated
fields. 1825–2800 m.
Distribution: KP, MP, MR, NL, RM, SL—
Gereau 3586
Commelina benghalensis L.; Ensermu & Faden,
1997: 373.
Terrestrial herb. Riverine forest, miombo
woodland. 1210–1950 m.
Distribution: MP, SL—Gereau 4290
Commelina diffusa Burm. f. subsp. diffusa; Brenan,
F.W.T.A. 3: 47. 1968.
Terrestrial herb. Montane forest. 1790 m.
Distribution: NL—Gereau 4165
Gereau et al.
Distribution: MP—J. Lovett 3621
Floscopa glomerata (Willd. ex Schult. & Schult. f.)
Hassk. subsp. glomerata; Brenan, F.W.T.A. 3:
28. 1968.
Terrestrial herb. Seasonally waterlogged grassland, woodland. 1250–1400 m.
Distribution: MP—Kayombo 970
Murdannia simplex (Vahl) Brenan; Ensermu &
Faden, 1997: 346.
Terrestrial herb. Edges of agricultural fields.
1300 m.
Distribution: MP—J. Lovett 3554
CONNARACEAE
Commelina hockii De Wild.; Faden, 2001: 403.
Terrestrial herb. Woodland, abandoned cultivation. 1250–1450 m.
Distribution: MP—J. Lovett 3724
Commelina kituloensis Faden; Faden, 2001: 400.
Terrestrial herb. Montane grassland, rocky
ridges, woodland. 1600–2800 m.
Distribution: KP, MP, MR, NL, RM—Faden
96/438 (type: BR, EA, K, MO, NHT, PRE, US),
Gereau 3181
Commelina neurophylla C. B. Clarke; Clarke,
F.T.A. 8: 53. 1901.
Terrestrial herb. Miombo woodland, montane
grassland. 2000–2500 m.
Distribution: KP, MP—P. Lovett 273
Commelina subulata Roth; Ensermu & Faden,
1997: 364.
Terrestrial herb. Stream banks. 1825 m.
Distribution: MR—Cribb 11389
Commelina triangulispatha Mildbr.; Faden, 1994:
305.
Terrestrial herb. Roadside banks.
Distribution: KP—Hepper 5318
Cyanotis barbata D. Don; Ensermu & Faden,
1997: 341.
Terrestrial herb. Montane grassland. 2280–
2600 m.
Distribution: KP, MP, MR—Mwasumbi 16167
Cyanotis longifolia Benth.; Ensermu & Faden,
1997: 340.
Terrestrial herb. Woodland. 1200–1600 m.
LakeNyasa_5-18.indd 42
1 genus and 3 species
Rourea coccinea (Schumach. & Thonn.) Benth.
subsp. boiviniana (Baill.) Jongkind; Jongkind,
1989: 332.
Scandent shrub. Disturbed montane forest.
Distribution: RM—Ndangalasi 1026
Rourea orientalis Baill.; Jongkind, 1989: 348.
Shrub. Miombo woodland, lakeshore shrubland. 474–980 m.
Distribution: SL—Gereau 5154
Rourea thomsonii (Baker) Jongkind; Jongkind,
1989: 359.
Liana. Montane forest. 1740–1780 m.
Distribution: RM—Kibure 1372
CONVOLVULACEAE
4 genera and 13 species
Astripomoea malvacea (Klotzsch) A. Meeuse var.
volkensii (Dammer) Verdc.; Verdcourt, F.T.E.A.
(Convolvulaceae): 76. 1963.
Trailing terrestrial herb. Edges of cultivation.
1350 m.
Distribution: MP—J. Lovett 3533
Dichondra repens J. R. Forst. & G. Forst.; Verdcourt, F.T.E.A. (Convolvulaceae): 12. 1963.
Creeping terrestrial herb. Montane forest.
Distribution: RM—WCS SHCP 249
Evolvulus alsinoides (L.) L.; Verdcourt, F.T.E.A.
(Convolvulaceae): 18. 1963.
Terrestrial herb. Grassland. 530 m.
Distribution: SL—Gilli 433
5/31/12 3:59 PM
Lake Nyasa Climatic Region
Floristic Checklist
Ipomoea alpina Rendle; Verdcourt, F.T.E.A. (Convolvulaceae): 104. 1963.
Terrestrial herb. Miombo woodland. 1530 m.
Distribution: SL—Gereau 4365
Ipomoea barteri Baker; Verdcourt, F.T.E.A. (Convolvulaceae): 119. 1963.
Herbaceous climber. Abandoned cultivation.
1250 m.
Distribution: MP—J. Lovett 4072
Ipomoea crassipes Hook. var. crassipes; Verdcourt,
F.T.E.A. (Convolvulaceae): 98. 1963.
Trailing terrestrial herb. Woodland. 1200 m.
Distribution: MP—J. Lovett 3878
Ipomoea involucrata P. Beauv. var. involucrata;
Verdcourt, F.T.E.A. (Convolvulaceae): 104.
1963.
Trailing terrestrial herb. Montane grassland,
montane forest. 1940–2530 m.
Distribution: KP, SL—Gereau 4774
43
Herbaceous climber. Woodland. 1400–1500 m.
Distribution: MP—J. Lovett 3414
CORNACEAE
1 genus and 1 species
Cornus volkensii Harms; Lovett et al., 2006: 62.
Tree. Montane forest. 1760–2230 m.
Distribution: MR, RM, SL—Gereau 4608
COSTACEAE
1 genus and 1 species
Costus macranthus K. Schum.; Lock, F.T.E.A.
(Zingiberaceae): 6. 1985.
Terrestrial herb. Woodland. 1250–1400 m.
Distribution: MP—J. Lovett 3572
CRASSULACEAE
2 genera and 5 species
Ipomoea marginata (Desr.) Verdc.; Verdcourt,
1987: 658.
Herbaceous climber. Wet grassland. 1250 m.
Distribution: MP—J. Lovett 4575
Crassula cf. alticola R. Fern.; Fernandes, Fl.
Zamb. 7(1): 15. 1983.
Terrestrial herb. Montane grassland. 2488 m.
Distribution: SL—Suleiman 171
Ipomoea pes-caprae (L.) R. Br. subsp. brasiliensis
(L.) Ooststr.; Verdcourt, F.T.E.A. (Convolvulaceae): 121. 1963.
Creeping terrestrial herb. Sandy lakeshore. 474
m.
Distribution: SL—Mwasumbi 16561
Crassula schimperi Fisch. & C. A. Mey. subsp.
phyturus (Mildbr.) R. Fern.; Wickens, F.T.E.A.
(Crassulaceae): 8. 1987.
Terrestrial herb. Thin soil among boulders.
2600 m.
Distribution: MP—Gereau 3504
Ipomoea prismatosyphon Welw. var. prismatosyphon; Verdcourt, F.T.E.A. (Convolvulaceae):
142. 1963.
Terrestrial herb. Woodland. 1250 m.
Distribution: MP—J. Lovett 4615
Crassula similis Baker f. vel sp. aff.; Fernandes, Fl.
Zamb. 7(1): 29. 1983.
Terrestrial herb. Woodland, montane grassland. 1200–2488 m.
Distribution: MP, SL—Suleiman 170
Ipomoea tenuirostris Choisy subsp. tenuirostris;
Verdcourt, F.T.E.A. (Convolvulaceae): 101.
1963.
Straggling terrestrial herb. Woodland. 1250 m.
Distribution: MP, RM—J. Lovett 4410
Crassula vaginata Eckl. & Zeyh.; Wickens,
F.T.E.A. (Crassulaceae): 13. 1987.
Terrestrial herb. Montane grassland, secondary
woodland. 2000–2500 m.
Distribution: MP—P. Lovett 502
Ipomoea welwitschii Vatke ex Hallier f.; Verdcourt,
F.T.E.A. (Convolvulaceae): 119. 1963.
Trailing terrestrial herb. Woodland. 1250 m.
Distribution: MP—J. Lovett 3651
Crassula vaginata Eckl. & Zeyh. × alba Forssk.;
Wickens, F.T.E.A. (Crassulaceae): 15. 1987.
Terrestrial herb. Montane grassland, roadsides.
2400–2730 m.
Distribution: KP, NL, RM—Gereau 4099
Ipomoea wightii (Wall.) Choisy var. wightii; Verdcourt, F.T.E.A. (Convolvulaceae): 110. 1963.
LakeNyasa_5-18.indd 43
Kalanchoe
crenata
(Andrews)
Haw.
subsp.
5/31/12 3:59 PM
44
Gereau et al.
nyassensis R. Fern.; Wickens, F.T.E.A. (Crassulaceae): 45. 1987.
Terrestrial herb. Montane forest. 2500 m.
Distribution: MP—P. Lovett 485
CUCURBITACEAE
11 genera and 15 species
Coccinia adoensis (Hochst. ex A. Rich.) Cogn.; Jeffrey, F.T.E.A. (Cucurbitaceae): 65. 1967.
Herbaceous climber. Montane forest remnants,
riverine forest, lakeside forest, woodland, abandoned cultivation. 474–2160 m.
Distribution: MP, SL—Gereau 3535
Coccinia grandiflora Cogn.; Jeffrey, F.T.E.A.
(Cucurbitaceae): 64. 1967.
Herbaceous climber. Abandoned cultivation.
1250 m.
Distribution: MP—J. Lovett 3701
Coccinia senensis (Klotzsch) Cogn.; Jeffrey,
F.T.E.A. (Cucurbitaceae): 66. 1967.
Herbaceous climber. Riverine forest. 580 m.
Distribution: SL—Gereau 3817
Cucumella aetheocarpa C. Jeffrey vel sp. aff.; Jeffrey, F.T.E.A. (Cucurbitaceae): 114. 1967.
Herbaceous climber. Woodland. 1200 m.
Distribution: MP—J. Lovett 3977
Lagenaria abyssinica (Hook. f.) C. Jeffrey; Jeffrey,
F.T.E.A. (Cucurbitaceae): 50. 1967.
Herbaceous climber. Woodland. 1300–1400 m.
Distribution: MP—Kayombo 1003
Momordica foetida Schumach.; Jeffrey, F.T.E.A.
(Cucurbitaceae): 29. 1967.
Herbaceous climber. Woodland, montane forest
edges. 1300–2250 m.
Distribution: MP, RM—J. Lovett 3449
Mukia maderaspatana (L.) M. Roem.; Jeffrey,
F.T.E.A. (Cucurbitaceae): 115. 1967.
Herbaceous climber. Riverine forest, woodland,
grassland. 1250–1400 m.
Distribution: MP—Kayombo 996
Oreosyce africana Hook. f.; Jeffrey, F.T.E.A.
(Cucurbitaceae): 110. 1967.
Herbaceous climber. Abandoned cultivation, riverine forest, montane bushland, bamboo
thicket. 1250–2300 m.
LakeNyasa_5-18.indd 44
Distribution: MP, MR, NL, RM, SL—Mwasumbi 16550
Peponium vogelii (Hook. f.) Engl.; Jeffrey, F.T.E.A.
(Cucurbitaceae): 81. 1967.
Herbaceous climber. Montane forest edges, riverine forest. 1500–2260 m.
Distribution: MP, SL—Gereau 3531
Raphidiocystis jeffreyana R. Fern. & A. Fern.; Jeffrey, F.T.E.A. (Cucurbitaceae): 55. 1967.
Herbaceous climber. Montane forest. 1710–
1750 m.
Distribution: RM—Abeid 2777
Note: New to Tanzania, from Uganda, Democratic Republic of Congo, Cameroon, Gabon, and
Angola.
Telfairia pedata (Sm. ex Sims) Hook.; Jeffrey,
F.T.E.A. (Cucurbitaceae): 15. 1967.
Herbaceous climber. 1750 m.
Distribution: RM—WCS SHCP 3
Trochomeria macrocarpa (Sond.) Hook. f.; Jeffrey,
F.T.E.A. (Cucurbitaceae): 87. 1967.
Straggling herb. Woodland, riverine forest.
1400–1500 m.
Distribution: MP—J. Lovett 3294A
Zehneria minutiflora (Cogn.) C. Jeffrey; Jeffrey,
F.T.E.A. (Cucurbitaceae): 126. 1967.
Herbaceous climber. Montane forest. 1790 m.
Distribution: NL, RM—Gereau 4159
Zehneria cf. oligosperma C. Jeffrey; Jeffrey,
F.T.E.A. (Cucurbitaceae): 124. 1967.
Herbaceous climber. Montane forest. 2000 m.
Distribution: MP—P. Lovett 442
Zehneria scabra (L. f.) Sond.; Jeffrey, F.T.E.A.
(Cucurbitaceae): 122. 1967.
Herbaceous climber. Montane forest, grassland,
coffee plantations. 1340–2400 m.
Distribution: MP, MR, RM—Mwasumbi 16244
CUSCUTACEAE
1 genus and 1 species
Cuscuta kilimanjari Oliv. var. kilimanjari;
Verdcourt, F.T.E.A. (Convolvulaceae): 6. 1963.
Twining parasitic herb. Riverine forest. 1275–
1400 m.
Distribution: MP—Kayombo 2364
5/31/12 3:59 PM
Lake Nyasa Climatic Region
Floristic Checklist
45
Terrestrial herb. Abandoned cultivation. 1250
CYPERACEAE
15 genera and 59 species
Ascolepis capensis (Kunth) Ridl.; Beentje, F.T.E.A.
(Cyperaceae): 273. 2010.
Terrestrial herb. Montane grassland. 2560–
2590 m.
Distribution: KP—Gereau 2851
Ascolepis hemisphaerica Peter ex Goetgh.; Beentje,
F.T.E.A. (Cyperaceae): 273. 2010.
Terrestrial herb. Wooded grassland. 1270–
1860 m.
Distribution: MP, SL—Gereau 3488
Bulbostylis atrosanguinea (Boeckeler) C. B.
Clarke; Verdcourt, F.T.E.A. (Cyperaceae): 75.
2010.
Terrestrial herb. Rocky mountain slopes. 2700
m.
Distribution: KP—Richards 18446
m.
Distribution: MP—J. Lovett 4082
Bulbostylis macra (Ridl.) C. B. Clarke; Verdcourt,
F.T.E.A. (Cyperaceae): 72. 2010.
Terrestrial herb. Woodland. 1300–1600 m.
Distribution: MP—J. Lovett 3615
Bulbostylis pusilla (Hochst. ex A. Rich.) C. B.
Clarke subsp. congolensis (De Wild.) R. W.
Haines; Verdcourt, F.T.E.A. (Cyperaceae): 95.
2010.
Terrestrial herb. Woodland. 1200 m.
Distribution: MP—J. Lovett 3999A
Carex chlorosaccus C. B. Clarke; Verdcourt,
F.T.E.A. (Cyperaceae): 432. 2010.
Terrestrial herb. Montane forest. 1790 m.
Distribution: NL—Gereau 4155
Bulbostylis buchananii C. B. Clarke; Verdcourt,
F.T.E.A. (Cyperaceae): 92. 2010.
Terrestrial herb. Woodland. 1200 m.
Distribution: MP—J. Lovett 3999
Carex cyrtosaccus C. B. Clarke; Verdcourt,
F.T.E.A. (Cyperaceae): 444. 2010.
Terrestrial herb. Riverbanks in montane grassland. 2560 m.
Distribution: KP—Gereau 3176
Bulbostylis densa (Wall.) Hand.–Mazz. subsp.
afromontana (Lye) R. W. Haines; Verdcourt,
F.T.E.A. (Cyperaceae): 100. 2010.
Terrestrial herb. Road banks in cultivation,
montane grassland. 2200–2500 m.
Distribution: KP, MR—Gereau 4533
Carex echinochloe Kunze subsp. nyasensis (C. B.
Clarke) Lye; Verdcourt, F.T.E.A. (Cyperaceae):
431. 2010.
Terrestrial herb. Riverine forest. 1210 m.
Distribution: SL—Gereau 4250
Bulbostylis filamentosa (Vahl) C. B. Clarke; Verdcourt, F.T.E.A. (Cyperaceae): 101. 2010.
Terrestrial herb. Miombo woodland. 1530 m.
Distribution: SL—Gereau 4360
Carex lycurus K. Schum. subsp. lycurus; Verdcourt, F.T.E.A. (Cyperaceae): 429. 2010.
Terrestrial herb. Riverine forest, riverbanks in
montane grassland. 1500–2590 m.
Distribution: KP, MP—Gereau 3155
Bulbostylis hispidula (Vahl) R. W. Haines subsp.
brachyphylla (Cherm.) R. W. Haines; Verdcourt, F.T.E.A. (Cyperaceae): 82. 2010.
Terrestrial herb. Miombo woodland. 1790 m.
Distribution: SL—Gereau 3896
Carex petitiana A. Rich.; Verdcourt, F.T.E.A.
(Cyperaceae): 439. 2010.
Terrestrial herb. Riverbanks and thickets in
montane grassland. 2560–2650 m.
Distribution: KP—Gereau 3193
Bulbostylis hispidula (Vahl) R. W. Haines subsp.
filiformis (C. B. Clarke) R. W. Haines; Verdcourt, F.T.E.A. (Cyperaceae): 83. 2010.
Terrestrial herb. Woodland. 1200 m.
Distribution: MP—J. Lovett 3999B
Carex spicatopaniculata C. B. Clarke; Verdcourt,
F.T.E.A. (Cyperaceae): 431. 2010.
Terrestrial herb. Montane forest edges. 2230 m.
Distribution: SL—Gereau 3573
Bulbostylis hispidula (Vahl) R. W. Haines subsp.
hispidula; Verdcourt, F.T.E.A. (Cyperaceae):
82. 2010.
Carex steudneri Boeckeler; Verdcourt, F.T.E.A.
(Cyperaceae): 435. 2010.
Terrestrial herb. Montane forest edges. 2320 m.
LakeNyasa_5-18.indd 45
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46
Gereau et al.
Distribution: KP, SL—Gereau 3986
Coleochloa setifera (Ridl.) Gilly var. setifera;
Hoenselaar, F.T.E.A. (Cyperaceae): 377. 2010.
Terrestrial herb. Miombo woodland, montane
forest/woodland transition. 1780–2140 m.
Distribution: SL—Gereau 4006
Cyperus ajax C. B. Clarke; Hoenselaar, F.T.E.A.
(Cyperaceae): 202. 2010.
Terrestrial herb. Montane forest. 2200–2500
m.
Distribution: MP, RM—Mwasumbi 16227
Cyperus aff. albopilosus (C. B. Clarke) Kük.;
Hoenselaar, F.T.E.A. (Cyperaceae): 155. 2010.
Terrestrial herb. Wooded grassland. 1860 m.
Distribution: SL—Gereau 3926
Cyperus amabilis Vahl; Hoenselaar, F.T.E.A.
(Cyperaceae): 158. 2010.
Terrestrial herb. Abandoned cultivation. 1250
m.
Distribution: MP—J. Lovett 4173
Cyperus articulatus L.; Hoenselaar, F.T.E.A.
(Cyperaceae): 208. 2010.
Terrestrial herb. 1970 m.
Distribution: RM—WCS SHCP 52
Cyperus cyperoides (L.) Kuntze; Beentje, F.T.E.A.
(Cyperaceae): 223. 2010.
Terrestrial herb. Montane forest, grassland,
riverine forest, woodland, roadsides. 474–2150 m.
Distribution: MP, NL, SL—Suleiman 165
Cyperus dichrostachyus Hochst. ex A. Rich.;
Hoenselaar, F.T.E.A. (Cyperaceae): 198. 2010.
Terrestrial herb. Swampy depressions in montane grassland, riverine forest. 1500–2590 m.
Distribution: KP, NL—Gereau 2847
Cyperus distans L. f.; Beentje, F.T.E.A. (Cyperaceae): 249. 2010.
Terrestrial herb. Riverine forest, montane
grassland, woodland/grassland transition, seasonal
ponds, forest clearings, abandoned cultivation.
1160–2230 m.
Distribution: MP, NL, SL—Gereau 4270
Cyperus esculentus L.; Hoenselaar, F.T.E.A.
(Cyperaceae): 227. 2010.
Terrestrial herb. Miombo woodland, riverine
forest, edges of cultivation, grassland, roadsides.
LakeNyasa_5-18.indd 46
474–1500 m.
Distribution: MP, SL—Gereau 3957
Cyperus exaltatus Retz. var. dives (Delile) C. B.
Clarke; Beentje, F.T.E.A. (Cyperaceae): 245.
2010.
Terrestrial herb. Woodland, inundated grassland. 1270–1400 m.
Distribution: MP—Gereau 3492
Cyperus glaucophyllus Boeckeler; Beentje, F.T.E.A.
(Cyperaceae): 198. 2010.
Terrestrial herb. Forest clearings, forest/
woodland transition, scree slopes, riverine forest.
1850–2140 m.
Distribution: NL, SL—Gereau 4001
Cyperus involucratus Rottb.; Hoenselaar, F.T.E.A.
(Cyperaceae): 187. 2010.
Terrestrial herb. Riverine forest. 980–1400 m.
Distribution: MP, NL—Mwangoka 663
Cyperus laxus Lam. subsp. sylvestris (Ridl.) Lye;
Hoenselaar, F.T.E.A. (Cyperaceae): 201. 2010.
Terrestrial herb. Montane forests, riverbanks.
1200–2230 m.
Distribution: MP, SL—Gereau 4428
Cyperus luteus Boeckeler; Hoenselaar, F.T.E.A.
(Cyperaceae): 171. 2010.
Terrestrial herb. Woodland, cultivated fields.
1350–2200 m.
Distribution: MP, MR—Gereau 4525
Cyperus mollipes (C. B. Clarke) K. Schum.;
Hoenselaar & Beentje, F.T.E.A. (Cyperaceae):
180. 2010.
Terrestrial herb. Riverine forest, miombo
woodland, montane grassland. 580–2230 m.
Distribution: SL—Gereau 3596
Cyperus niveus Retz. var. leucocephalus (Kunth)
Fosberg; Hoenselaar, F.T.E.A. (Cyperaceae):
170. 2010.
Terrestrial herb. Woodland. 1200 m.
Distribution: MP—J. Lovett 3986
Cyperus nyererei Lye; Hoenselaar, F.T.E.A.
(Cyperaceae): 176. 2010.
Terrestrial herb. Woodland, montane grassland, scree slopes. 1600–2740 m.
Distribution: KP, MP, SL—Wingfield 529 (type:
DSM, K), Gereau 3507
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Floristic Checklist
47
Cyperus pluribracteatus (Kük.) Govaerts; Hoenselaar, F.T.E.A. (Cyperaceae): 223. 2010.
Terrestrial herb. Woodland. 1200 m.
Distribution: MP—J. Lovett 4048
F.T.E.A. (Cyperaceae): 123. 2010.
Terrestrial herb. Riverbanks in montane grassland. 2550 m.
Distribution: KP—Gereau 3149
Cyperus rigidifolius Steud.; Hoenselaar, F.T.E.A.
(Cyperaceae): 165. 2010.
Terrestrial herb. Montane grassland. 1810–
2600 m.
Distribution: KP, MP, MR, SL—Gereau 2852
Kyllinga brevifolia Rottb. var. lurida (Kük.)
Beentje; Beentje, F.T.E.A. (Cyperaceae): 315.
2010.
Terrestrial herb. Montane grassland. 2230–
2360 m.
Distribution: KP, MR, SL—Gereau 3740
Cyperus rotundus L.; Beentje, F.T.E.A. (Cyperaceae): 211. 2010.
Terrestrial herb. Grassland, riverine forest.
1500–2230 m.
Distribution: MP, SL—Gereau 3592
Cyperus tenuiculmis Boeckeler var. guineensis
(Nelmes) S. S. Hooper; Hoenselaar, F.T.E.A.
(Cyperaceae): 243. 2010.
Terrestrial herb. Seasonal ponds in montane
grassland. 1840 m.
Distribution: SL—Gereau 3915
Kyllinga comosipes (Mattf. & Kük.) Napper;
Beentje, F.T.E.A. (Cyperaceae): 338. 2010.
Terrestrial herb. Thin soil among boulders.
2600 m.
Distribution: MP—Gereau 3506
Kyllinga crassipes Boeckeler; Beentje, F.T.E.A.
(Cyperaceae): 342. 2010.
Terrestrial herb. Riverine forest. 1160–1500 m.
Distribution: SL—Gereau 3941
Cyperus tenuiculmis Boeck. var. tenuiculmis;
Hoenselaar, F.T.E.A. (Cyperaceae): 243. 2010.
Terrestrial herb. Riverine forest, montane
grassland. 1160–2120 m.
Distribution: SL—Gereau 4316
Kyllinga odorata Vahl var. major (C. B. Clarke)
Chiov.; Beentje, F.T.E.A. (Cyperaceae): 340.
2010.
Terrestrial herb. Montane forest, abandoned
cultivation. 1250–2110 m.
Distribution: MP, MR, NL—Gereau 4563
Cyperus tomaiophyllus K. Schum.; Hoenselaar,
F.T.E.A. (Cyperaceae): 239. 2010.
Terrestrial herb. Montane forest edges. 2360 m.
Distribution: MR—Gereau 3783
Kyllinga odorata Vahl var. odorata; Beentje,
F.T.E.A. (Cyperaceae): 340. 2010.
Terrestrial herb. Montane forest. 1900 m.
Distribution: NL—Gereau 4218
Ficinia filiformis (Lam.) Schrad.; Muasya, F.T.E.A.
(Cyperaceae): 118. 2010.
Terrestrial herb. Montane grassland, rocky outcrops. 2610–2800 m.
Distribution: KP, RM—Gereau 2822
Kyllinga tanzaniae Lye; Beentje, F.T.E.A. (Cyperaceae): 343. 2010.
Terrestrial herb. Grassland, miombo woodland.
1270–1400 m.
Distribution: MP—Gereau 3485
Fimbristylis complanata (Retz.) Link; Verdcourt,
F.T.E.A. (Cyperaceae): 50. 2010
Terrestrial herb. Grassland, woodland. 1250–
1270 m.
Distribution: MP—Gereau 3483
Kyllinga tenuifolia Steud.; Beentje, F.T.E.A.
(Cyperaceae): 29. 2010.
Terrestrial herb. Riverine forest, miombo
woodland. 1200 m.
Distribution: MP—J. Lovett 3990
Fimbristylis dichotoma (L.) Vahl; Verdcourt,
F.T.E.A. (Cyperaceae): 57. 2010.
Terrestrial herb. Miombo woodland, riverine
forest. 474–1800 m.
Distribution: MP, NL, SL—Gereau 3682
Kyllingiella polyphylla (A. Rich.) Lye; Beentje,
F.T.E.A. (Cyperaceae): 128. 2010.
Terrestrial herb. Grassland. 1270 m.
Distribution: MP—Gereau 3489
Isolepis costata Hochst. ex A. Rich.; Muasya,
LakeNyasa_5-18.indd 47
Lipocarpha comosa J. Raynal vel sp. aff.; Beentje,
F.T.E.A. (Cyperaceae): 347. 2010.
5/31/12 3:59 PM
48
Gereau et al.
Terrestrial herb. Woodland. 1250 m.
Distribution: MP—J. Lovett 4332
Pycreus capillifolius (A. Rich.) C. B. Clarke;
Hoenselaar, F.T.E.A. (Cyperaceae): 284. 2010.
Terrestrial herb. Road banks in cultivation.
2200 m.
Distribution: MR—Gereau 4534
Pycreus elegantulus (Steud.) C. B. Clarke; Hoenselaar, F.T.E.A. (Cyperaceae): 303. 2010.
Terrestrial herb. Swampy depressions in
montane grassland, riverine forest, abandoned
cultivation. 1250–2590 m.
Distribution: KP, MP, SL—Gereau 2848
(Cyperaceae): 407. 2010.
Terrestrial herb. Miombo woodland. 1260 m.
Distribution: SL—Gereau 4306
Scleria racemosa Poir.; Verdcourt, F.T.E.A.
(Cyperaceae): 413. 2010.
Terrestrial herb. Riverine forest. 474–1500 m.
Distribution: NL—Gereau 5169
Scleria woodii C. B. Clarke var. woodii; Verdcourt,
F.T.E.A. (Cyperaceae): 391. 2010.
Terrestrial herb. Grassland. 1270 m.
Distribution: MP—Gereau 3484
DIOSCOREACEAE
Pycreus flavescens (L.) P. Beauv. ex Rchb.; Hoenselaar, F.T.E.A. (Cyperaceae): 292. 2010.
Terrestrial herb. Riverine forest, wooded grassland. 1200–1500 m.
Distribution: MP, SL—Gereau 3943
Pycreus macrostachyos (Lam.) J. Raynal; Hoenselaar, F.T.E.A. (Cyperaceae): 288. 2010.
Terrestrial herb. Riverine forest, wooded grassland. 1250–1350 m.
Distribution: MP—J. Lovett 4658
Schoenoplectus corymbosus (Roth ex Roem. &
Schult.) J. Raynal; Beentje, F.T.E.A. (Cyperaceae): 25. 2010.
Terrestrial herb. Seasonal ponds in montane
grassland. 1840 m.
Distribution: SL—Gereau 3918
Schoenoxiphium ludwigii Hochst.; Verdcourt,
F.T.E.A. (Cyperaceae): 420. 2010.
Terrestrial herb. Riverbanks in montane grassland. 2560 m.
Distribution: KP—Gereau 3172
Schoenoxiphium sparteum (Wahlenb.) C. B.
Clarke; Verdcourt, F.T.E.A. (Cyperaceae): 418.
2010.
Terrestrial herb.
Distribution: MP—Napper 1696
Scleria bulbifera Hochst. ex A. Rich.; Verdcourt,
F.T.E.A. (Cyperaceae): 386. 2010.
Terrestrial herb. Miombo woodland. 1200–
1600 m.
Distribution: MP, SL—Gereau 4307
Scleria lagoensis Boeckeler; Verdcourt, F.T.E.A.
LakeNyasa_5-18.indd 48
1 genus and 5 species
Dioscorea buchananii Benth.; Milne-Redhead,
F.T.E.A. (Dioscoreaceae): 19. 1975.
Herbaceous climber. Woodland. 1300–1600 m.
Distribution: MP—J. Lovett 4286
Dioscorea cochleariapiculata De Wild.; MilneRedhead, F.T.E.A. (Dioscoreaceae): 22. 1975.
Herbaceous climber. Woodland. 1250–1400 m.
Distribution: MP—Kayombo 962
Dioscorea dumetorum (Kunth) Pax; Milne-Redhead, F.T.E.A. (Dioscoreaceae): 21. 1975.
Herbaceous climber. Riverine forest, woodland.
1160–1200 m.
Distribution: MP, SL—Gereau 3692
Dioscorea quartiniana A. Rich.; Milne-Redhead,
F.T.E.A. (Dioscoreaceae): 23. 1975.
Herbaceous climber. Riverine forest, montane
forest. 1210–1850 m.
Distribution: NL, SL—Gereau 4285
Dioscorea schimperiana Hochst. ex Kunth; MilneRedhead, F.T.E.A. (Dioscoreaceae): 14. 1975.
Herbaceous climber. Riverine forest, woodland.
1200–1500 m.
Distribution: MP, NL, SL—Gereau 3948
DIPSACACEAE
2 genera and 2 species
Cephalaria pungens Szabó; Napper, F.T.E.A. (Dipsacaceae): 4. 1968.
Terrestrial herb. Montane grassland. 2500–
2700 m.
5/31/12 3:59 PM
Lake Nyasa Climatic Region
Floristic Checklist
Distribution: KP, MP, NL—Gereau 4098
Scabiosa austroafricana Heine; Napper, F.T.E.A.
(Dipsacaceae): 9. 1968.
Terrestrial herb. Montane grassland, woodland.
1720–2600 m.
Distribution: KP, MP—Gereau 3088
DIPTEROCARPACEAE
1 genus and 2 species
Monotes adenophyllus Gilg subsp. adenophyllus;
Verdcourt, F.T.E.A. (Dipterocarpaceae): 7.
1989.
Tree. Woodland. 1200–1310 m.
Distribution: MP—J. Lovett 4312
Monotes africanus A. DC.; Verdcourt, F.T.E.A.
(Dipterocarpaceae): 8. 1989.
Tree. Miombo woodland. 1530–1790 m.
Distribution: SL—Gereau 3873
DRACAENACEAE
2 genera and 5 species
Dracaena afromontana Mildbr.; Mwachala,
F.T.E.A. (Dracaenaceae): 6. 2007.
Tree. Montane forest. 1985–2380 m.
Distribution: KP, RM, SL—Abeid 2816
Dracaena laxissima Engl.; Mwachala, F.T.E.A.
(Dracaenaceae): 3. 2007.
Shrub, liana. Riverine forest, montane forest.
1810–2110 m.
Distribution: MP, MR, SL—Gereau 4555
Dracaena mannii Baker; Mwachala, F.T.E.A.
(Dracaenaceae): 7. 2007.
Shrub. Lakeshore bushland, montane forest.
485–2000 m.
Distribution: RM, SL—Mwasumbi 16613
Dracaena steudneri Engl.; Mwachala, F.T.E.A.
(Dracaenaceae): 8. 2007.
Tree or shrub. Woodland, riverine forest.
1300–2000 m.
Distribution: MP, NL, RM—Gereau 6108
Sansevieria kirkii Baker; Mbugua, F.T.E.A. (Dracaenaceae): 30. 2007.
Terrestrial herb or suffrutex. Riverine forest,
woodland. 580–1250 m.
Distribution: MP, SL—Gereau 3825
LakeNyasa_5-18.indd 49
49
DROSERACEAE
1 genus and 2 species
Drosera madagascariensis DC.; Laundon, F.T.E.A.
(Droseraceae): 5. 1959.
Terrestrial herb. Stream bank in montane
grassland. 2740 m.
Distribution: KP—Abeid 3067
Drosera pilosa Exell & J. R. Laundon; Laundon,
F.T.E.A. (Droseraceae): 4. 1959.
Terrestrial herb. Montane grassland. 2488–
2700 m.
Distribution: NL, RM, SL—Gereau 4086
EBENACEAE
2 genera and 6 species
Diospyros abyssinica (Hiern) F. White subsp.
abyssinica; White & Verdcourt, F.T.E.A.
(Ebenaceae): 22. 1996.
Tree. Montane forest. 1900 m.
Distribution: RM—Bracebridge 45
Diospyros mespiliformis Hochst. ex A. DC.; White
& Verdcourt, F.T.E.A. (Ebenaceae): 40. 1996.
Tree. Valley forest. 1200 m.
Distribution: MP—J. Lovett 4248
Diospyros natalensis (Harv.) Brenan; White &
Verdcourt, F.T.E.A. (Ebenaceae): 13. 1996.
Shrub. Riverine forest. 980 m.
Distribution: SL—Gereau 5155
Diospyros whyteana (Hiern) F. White; White &
Verdcourt, F.T.E.A. (Ebenaceae): 28. 1996.
Tree or shrub. Montane forest. 1750–2380 m.
Distribution: KP, MR, NL, RM, SL—Mwasumbi 16194
Diospyros zombensis (B. L. Burtt) F. White; White
& Verdcourt, F.T.E.A. (Ebenaceae): 27. 1996.
Tree or shrub. Riverine forest. 860–1210 m.
Distribution: SL—Kibure 1450
Euclea racemosa L. subsp. schimperi (A. DC.) F.
White; White & Verdcourt, F.T.E.A. (Ebenaceae): 46. 1996.
Shrub. Miombo woodland, riverine forest.
474–1530 m.
Distribution: MP, NL, SL—Gereau 4799
5/31/12 3:59 PM
50
ERICACEAE
2 genera and 6 species
Agarista salicifolia (Comm. ex Lam.) G. Don;
Beentje, F.T.E.A. (Ericaceae): 2. 2006.
Tree. Miombo woodland, tree clumps in montane grassland. 1730–2800 m.
Distribution: KP, MR, RM, SL—Gereau 5096
Erica benguelensis (Welw. ex Engl.) E. G. H. Oliv.
var. albescens (R. Ross) E. G. H. Oliv.; Beentje,
F.T.E.A. (Ericaceae): 10. 2006.
Tree. Montane shrubland. 2100–2540 m.
Distribution: KP, MR, NL—Abeid 2873
Erica benguelensis (Welw. ex Engl.) E. G. H. Oliv.
var. benguelensis; Beentje, F.T.E.A. (Ericaceae): 9. 2006.
Tree. Montane heathland and forest edges.
2340–2350 m.
Distribution: RM—Abeid 2990
Erica kingaensis Engl. subsp. bequaertii (De
Wild.) R. Ross; Beentje, F.T.E.A. (Ericaceae):
19. 2006.
Shrub. Stream banks in montane grassland.
2550 m.
Distribution: KP—Gereau 3157
Note: New to Tanzania, from Rwenzori Mountains in Uganda and Democratic Republic of
Congo.
Erica kingaensis Engl. subsp. kingaensis; Beentje,
F.T.E.A. (Ericaceae): 18. 2006.
Shrub or subshrub. Grassland, heathland, secondary shrubland. 1760–2770 m.
Distribution: KP, MR, NL, RM—Mwasumbi
16254
Gereau et al.
1940–2770 m.
Distribution: KP, MR, NL, RM, SL—Mwasumbi 16261
Erica whyteana Britten; Beentje, F.T.E.A. (Ericaceae): 11. 2006.
Subshrub. Stream banks and swamps in montane grassland. 2500–2830 m.
Distribution: KP—Abeid 2856
ERIOCAULACEAE
1 genus and 1 species
Eriocaulon schimperi Körn. ex Ruhland; Phillips,
F.T.E.A. (Eriocaulaceae): 15. 1997.
Terrestrial herb. Ravines and stream banks in
montane grassland. 2560–2600 m.
Distribution: KP, MP—Gereau 3116
ERIOSPERMACEAE
1 genus and 1 species
Eriospermum abyssinicum Baker; Whitehouse,
F.T.E.A. (Eriospermaceae): 4. 1996.
Terrestrial herb. Miombo woodland, rocky
mountain slopes. 1200–2800 m.
Distribution: KP, MP—J. Lovett 3811
ERYTHROXYLACEAE
1 genus and 1 species
Erythroxylum emarginatum Thonn.; Verdcourt,
F.T.E.A. (Erythroxylaceae): 3. 1984.
Shrub. Riverine forest. 1320 m.
Distribution: MP—Gereau 6125
EUPHORBIACEAE
Erica mannii (Hook. f.) Beentje subsp. pallidiflora
(Engl.) E. G. H. Oliv.; Beentje, F.T.E.A. (Ericaceae): 17. 2006.
Shrub. Woodland. 1300–1790 m.
Distribution: MP, SL—Kayombo 2481
Erica mannii (Hook. f.) Beentje subsp. usambarensis (Alm. & T. C. E. Fr.) Beentje; Beentje,
F.T.E.A. (Ericaceae): 17. 2006.
Shrub. Rocky mountaintop grassland. 2960 m.
Distribution: RM—Abeid 3005
Erica silvatica (Engl.) Beentje; Beentje, F.T.E.A.
(Ericaceae): 19. 2006.
Subshrub. Heathland, montane grassland.
LakeNyasa_5-18.indd 50
19 genera and 51 species
Acalypha fimbriata Schumach. & Thonn.; Radcliffe-Smith, F.T.E.A. (Euphorbiaceae 1): 198.
1987.
Terrestial herb. Roadsides. 474–490 m.
Distribution: SL—Gereau 3792
Acalypha ornata Hochst. ex A. Rich.; RadcliffeSmith, F.T.E.A. (Euphorbiaceae 1): 190. 1987.
Shrub. Riverine forest, woodland, abandoned
cultivation. 1160–1500 m.
Distribution: MP, SL—Gereau 3700
Acalypha petiolaris Hochst.; Prain & Hutchinson,
5/31/12 3:59 PM
Lake Nyasa Climatic Region
Floristic Checklist
1913: 18.
Terrestrial herb. Woodland, cultivated fields,
lakeshore bushland. 474–1600 m.
Distribution: MP, SL—Mwasumbi 16577
Acalypha polymorpha Müll. Arg.; Hutchinson,
F.T.A. 6(1): 894. 1912.
Terrestrial herb. Grassland, woodland, abandoned cultivation. 1250–1940 m.
Distribution: MP, SL—Gereau 4759
Acalypha psilostachya Hochst. var. glandulosa
Hutch.; Radcliffe-Smith, F.T.E.A. (Euphorbiaceae 1): 204. 1987.
Shrub, often scandent, or perennial herb. Montane forest, riverine forest, scree slopes, woodland.
1500–2400 m.
Distribution: MP, MR, SL—Gereau 3637
Acalypha psilostachya Hochst. var. psilostachya;
Radcliffe-Smith, F.T.E.A. (Euphorbiaceae 1):
204. 1987.
Shrub, often scandent, or perennial herb.
Montane forest, tree clumps in grassland, riverine
forest, roadsides in cultivation, tree plantations.
1160–2460 m.
Distribution: KP, MP, MR, NL, RM, SL—Mwasumbi 16226
Antidesma membranaceum Müll. Arg.; RadcliffeSmith, F.T.E.A. (Euphorbiaceae 2): 574. 1988.
Tree or shrub. Abandoned cultivation. 1250 m.
Distribution: MP—J. Lovett 3748
Antidesma venosum E. Mey. ex Tul.; RadcliffeSmith, F.T.E.A. (Euphorbiaceae 2): 573. 1988.
Tree or shrub. Woodland, riverine forest, lakeshore forest. 474–1350 m.
Distribution: MP, SL—Gereau 3665
Bridelia brideliifolia (Pax) Fedde; Radcliffe-Smith,
F.T.E.A. (Euphorbiaceae 1): 126. 1987.
Tree. Montane forest, woodland/forest transition. 1900–2110 m.
Distribution: MR, NL, SL—Gereau 4743
Bridelia cathartica G. Bertol.; Radcliffe-Smith,
F.T.E.A. (Euphorbiaceae 1): 123. 1987.
Shrub. Miombo woodland, wooded grassland,
riverine forest. 1200–1400 m.
Distribution: MP—J. Lovett 4359
Bridelia micrantha (Hochst.) Baill.; RadcliffeSmith, F.T.E.A. (Euphorbiaceae 1): 127. 1987.
LakeNyasa_5-18.indd 51
51
Tree. Montane forest, woodland, riverine forest,
edges of cultivation. 1160–1920 m.
Distribution: MP, NL, RM, SL—Kibure 1402
Bridelia scleroneura Müll. Arg.; Radcliffe-Smith,
F.T.E.A. (Euphorbiaceae 1): 122. 1987.
Shrub. Miombo woodland. 1200 m.
Distribution: MP—Kayombo 2409
Clutia abyssinica Jaub. & Spach var. abyssinica;
Radcliffe-Smith, F.T.E.A. (Euphorbiaceae 1):
334. 1987.
Shrub. Montane forest, woodland, abandoned
cultivation. 1250–2470 m.
Distribution: KP, MP, MR, NL, SL—Gereau
4712
Clutia abyssinica Jaub. & Spach var. pedicellaris
(Pax) Pax; Radcliffe-Smith, F.T.E.A. (Euphorbiaceae 1): 336. 1987.
Shrub. Montane forest, forest edges, miombo
woodland. 1400–2400 m.
Distribution: MP, MR, RM—Mwasumbi 16181
Clutia angustifolia Knauf; Radcliffe-Smith,
F.T.E.A. (Euphorbiaceae 1): 339. 1987.
Shrub. Miombo woodland, grassland. 980–
1940 m.
Distribution: NL, SL—Gereau 4653
Clutia paxii Knauf; Radcliffe-Smith, F.T.E.A.
(Euphorbiaceae 1): 338. 1987.
Shrub. Montane forest, stream banks in montane grassland. 2400–2665 m.
Distribution: KP, MP—Gereau 3126
Clutia whytei Hutch.; Radcliffe-Smith, F.T.E.A.
(Euphorbiaceae 1): 333. 1987.
Shrub. Wooded grassland. 1860 m.
Distribution: SL—Gereau 3924
Clutia sp.
Shrub. Montane forest edges, scree slopes.
1980–2120 m.
Distribution: SL—Gereau 4330
Clutia sp.
Shrub. Riverbanks in montane grassland.
2500–2660 m.
Distribution: KP—Taylor 8520
Croton macrostachyus Hochst. ex Delile; RadcliffeSmith, F.T.E.A. (Euphorbiaceae 1): 149. 1987.
Tree. Woodland, riverine forest. 1300–1850 m.
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52
Gereau et al.
Distribution: MP, RM, SL—Gereau 4025
Croton scheffleri Pax; Radcliffe-Smith, F.T.E.A.
(Euphorbiaceae 1): 147. 1987.
Shrub. Riverine forest. 1200 m.
Distribution: MP—J. Lovett 3836
Drypetes gerrardii Hutch. var. gerrardii; RadcliffeSmith, F.T.E.A. (Euphorbiaceae 1): 98. 1987.
Tree. Montane forest. 1900–2260 m.
Distribution: NL, RM—Abeid 2778
Drypetes gerrardii Hutch. var. tomentosa Radcl.Sm.; Radcliffe-Smith, F.T.E.A. (Euphorbiaceae
1): 98. 1987.
Tree. Riverine forest. 1450 m.
Distribution: MP—J. Lovett 4898
Drypetes natalensis (Harv.) Hutch. var. natalensis;
Radcliffe-Smith, F.T.E.A. (Euphorbiaceae 1):
92. 1987.
Tree. Montane forest. 1780–1955 m.
Distribution: NL, RM—Kibure 1428
Erythrococca trichogyne (Müll. Arg.) Prain; Radcliffe-Smith, F.T.E.A. (Euphorbiaceae 1): 274.
1987.
Shrub. Montane forest. 1780–2300 m.
Distribution: MR, RM, SL—Mwasumbi 16539
Euphorbia daviesii E. A. Bruce; Carter, F.T.E.A.
(Euphorbiaceae 2): 437. 1988.
Terrestrial herb. Montane grassland. 1810–
2800 m.
Distribution: KP, SL—R. M. Davies E.17 (type:
K), Harder 1493
Euphorbia grantii Oliv.; Carter, F.T.E.A. (Euphorbiaceae 2): 457. 1988.
Shrub. 1400 m.
Distribution: MP—Michelmore 973
Euphorbia heterophylla L.; Carter, F.T.E.A.
(Euphorbiaceae 2): 431. 1988.
Terrestrial herb. Abandoned cultivation, lakeshore, riverine forest. 474–1500 m.
Distribution: MP, SL—Gereau 4397
Euphorbia hirta L.; Carter, F.T.E.A. (Euphorbiaceae 2): 415. 1988.
Terrestrial herb. Riverine forest, roadsides.
474–1350 m.
Distribution: MP, SL—Gereau 3787
LakeNyasa_5-18.indd 52
Euphorbia ugandensis Pax; Carter, F.T.E.A.
(Euphorbiaceae 2): 441. 1988.
Shrub. Montane forest. 2040–2640 m.
Distribution: MR, NL, RM, SL—Gereau 4708
Flueggea virosa (Roxb. ex Willd.) Royle; RadcliffeSmith, F.T.E.A. (Euphorbiaceae 1): 68. 1987.
Shrub. Riverine forest, lakeshore forest, abandoned cultivation. 480–1500 m.
Distribution: MP, SL—Gereau 3937
Macaranga capensis (Baill.) Benth. ex Sim var.
capensis; Lovett et al., 2006: 85.
Tree. Montane forest, riverine forest. 840–1760
m.
Distribution: RM, SL—Kibure 1436
Macaranga capensis (Baill.) Benth. ex Sim var.
kilimandscharica (Pax) Friis & M. G. Gilbert;
Lovett et al., 2006: 85.
Tree. Montane forest. 1900–2230 m.
Distribution: MR, NL, SL—Gereau 4215
Macaranga conglomerata Brenan; RadcliffeSmith, F.T.E.A. (Euphorbiaceae 1): 246. 1987.
Tree. Miombo woodland. 620 m.
Distribution: SL—Harder 1523
Margaritaria discoidea (Baill.) G. L. Webster var.
nitida (Pax) Radcl.-Sm.; Radcliffe-Smith,
F.T.E.A. (Euphorbiaceae 1): 67. 1987.
Tree or shrub. Woodland, riverine forest. 580–
1200 m.
Distribution: MP, SL—Harder 1510
Margaritaria discoidea (Baill.) G. L. Webster var.
triplosphaera Radcl.-Sm.; Radcliffe-Smith,
F.T.E.A. (Euphorbiaceae 1): 67. 1987.
Shrub. Miombo woodland. 620 m.
Distribution: SL—Harder 1509
Monadenium orobanchoides Bally vel sp. aff.;
Carter, F.T.E.A. (Euphorbiaceae 2): 544. 1988.
Terrestrial herb. Montane grassland. 1860–
2200 m.
Distribution: SL—Gereau 3931
Monadenium pedunculatum S. Carter; Carter,
F.T.E.A. (Euphorbiaceae 2): 542. 1988.
Terrestrial herb. Woodland, montane grassland. 1250–1810 m.
Distribution: MP, SL—Harder 1491
Neoboutonia macrocalyx Pax; Radcliffe-Smith,
5/31/12 3:59 PM
Lake Nyasa Climatic Region
Floristic Checklist
F.T.E.A. (Euphorbiaceae 1): 232. 1987.
Tree. Montane forest. 1960–2300 m.
Distribution: MP, MR, SL—Mwasumbi 16360
Phyllanthus amarus Schumach & Thonn.; Radcliffe-Smith, F.T.E.A. (Euphorbiaceae 1): 58.
1987.
Terrestrial herb. Montane forest. 1900 m.
Distribution: NL—Gereau 4222
Phyllanthus arvensis Müll. Arg.; Radcliffe-Smith,
F.T.E.A. (Euphorbiaceae 1): 57. 1987.
Terrestrial herb. Riverbanks in montane grassland. 2550–2890 m.
Distribution: KP—Gereau 3115
Phyllanthus capillaris Schumach. & Thonn.;
Hutchinson, F.T.A. 6(1): 709. 1912.
Terrestrial herb. Riverine forest, montane forest. 1210–1790 m.
Distribution: RM, SL—Gereau 4249
Phyllanthus micromeris Radcl.-Sm.; RadcliffeSmith, F.T.E.A. (Euphorbiaceae 1): 59. 1987.
Terrestrial herb. Montane forest. 2300 m.
Distribution: MR—Gereau 4585
Phyllanthus muellerianus (Kuntze) Exell; Radcliffe-Smith, F.T.E.A. (Euphorbiaceae 1): 24.
1987.
Shrub. Montane forest. 2075 m.
Distribution: RM—Semsei 1563
53
Distribution: SL—Gereau 4705
Pseudolachnostylis maprouneifolia Pax var.
dekindtii (Pax) Radcl.-Sm.; Radcliffe-Smith,
F.T.E.A. (Euphorbiaceae 1): 81. 1987.
Tree. Riverine forest in miombo woodland.
1200–1250 m.
Distribution: MP—J. Lovett 4360
Pseudolachnostylis maprouneifolia Pax var.
maprouneifolia; Radcliffe-Smith, F.T.E.A.
(Euphorbiaceae 1): 81. 1987.
Tree. Coffee plantation, miombo woodland.
1300–1410 m.
Distribution: MP—Kayombo 1058
Pseudolachnostylis maprouneifolia Pax var.
polygyna (Pax & K. Hoffm.) Radcl.-Sm.;
Radcliffe-Smith, F.T.E.A. (Euphorbiaceae 1):
83. 1987.
Tree. Miombo woodland. 1300 m.
Distribution: MP—J. Lovett 4909
Ricinus communis L.; Radcliffe-Smith, F.T.E.A.
(Euphorbiaceae 1): 322. 1987.
Shrub. Riverine forest, woodland, montane forest. 1310–1980 m.
Distribution: MP, RM—Kayombo 1109
Tragia furialis Bojer ex Prain; Radcliffe-Smith,
F.T.E.A. (Euphorbiaceae 1): 316. 1987.
Herbaceous climber. Woodland. 1250 m.
Distribution: MP—J. Lovett 4424
Phyllanthus niruroides Müll. Arg.; RadcliffeSmith, F.T.E.A. (Euphorbiaceae 1): 60. 1987.
Terrestrial herb. Miombo woodland. 1200 m.
Distribution: MP—J. Lovett 4139
Tragia plukenetii Radcl.-Sm.; Radcliffe-Smith,
F.T.E.A. (Euphorbiaceae 1): 296. 1987.
Terrestrial herb. Riverine forest. 1275 m.
Distribution: MP—Kayombo 2376
Phyllanthus paxii Hutch.; Radcliffe-Smith,
F.T.E.A. (Euphorbiaceae 1): 56. 1987.
Terrestrial herb. Miombo woodland, grassland,
riverine forest. 1500–1940 m.
Distribution: SL—Gereau 4666
Tragiella anomala (Prain) Pax & K. Hoffm.;
Radcliffe-Smith, F.T.E.A. (Euphorbiaceae 1):
321. 1987.
Herbaceous climber. Montane forest. 2200 m.
Distribution: MR—Mwasumbi 16202
Phyllanthus pseudoniruri Müll. Arg.; RadcliffeSmith, F.T.E.A. (Euphorbiaceae 1): 48. 1987.
Terrestrial herb. Woodland. 1250 m.
Distribution: MP—J. Lovett 4074
Uapaca kirkiana Müll. Arg.; Radcliffe-Smith,
F.T.E.A. (Euphorbiaceae 2): 567. 1988.
Tree. Woodland. 1260–1800 m.
Distribution: MP, NL, SL—Kayombo 2390
Phyllanthus ukagurensis Radcl.-Sm.; RadcliffeSmith, F.T.E.A. (Euphorbiaceae 1): 44. 1987.
Terrestrial herb. Trailsides in montane forest.
2040 m.
Uapaca nitida Müll. Arg.; Radcliffe-Smith,
F.T.E.A. (Euphorbiaceae 2): 568. 1988.
Tree. Woodland. 620–1400 m.
Distribution: MP, SL—Harder 1524
LakeNyasa_5-18.indd 53
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54
FABACEAE
42 genera and 156 species
Abrus melanospermus Hassk. subsp. tenuiflorus
(Spruce ex Benth.) D. Harder; Harder, Fl.
Zamb. 3(3): 216. 2007.
Lianescent shrub. Riverine forest. 580 m.
Distribution: SL—Gereau 3802
Adenocarpus mannii (Hook. f.) Hook. f.;
Milne-Redhead, F.T.E.A. (Leguminosae–Papilionoideae 2): 1009. 1971.
Shrub. Stream banks, montane grassland and
heathland. 1950–2960 m.
Distribution: KP, MP, MR, RM—Abeid 2877
Aeschynomene abyssinica (A. Rich.) Vatke; Verdcourt, F.T.E.A. (Leguminosae–Papilionoideae
1): 396. 1971.
Shrublet. Montane grassland. 2300–2610 m.
Distribution: KP, MP—Pócs 87063
Aeschynomene mimosifolia Vatke; Verdcourt,
F.T.E.A. (Leguminosae–Papilionoidede 1):
392. 1971.
Shrub. Shrubby montane grassland. 2300–
2400 m.
Distribution: MP—P. Lovett 501
Aeschynomene nyassana Taub.; Verdcourt,
F.T.E.A. (Leguminosae–Papilionoideae 1):
403. 1971.
Terrestrial herb. Miombo woodland, rocky
scrubland. 1780–2000 m.
Distribution: MP, SL—Gereau 3908
Aeschynomene nyikensis Baker; Verdcourt, F.T.E.A.
(Leguminosae–Papilionoideae 1): 394. 1971.
Shrub. Riverine forest. 1500 m.
Distribution: RM, SL—Gereau 4792
Aeschynomene schimperi Hochst. ex A. Rich.; Verdcourt, F.T.E.A. (Leguminosae–Papilionoideae
1): 376. 1971.
Shrub. Wet grassland. 1300–1400 m.
Distribution: MP—Kayombo 1043
Aeschynomene schliebenii Harms var. mossambicensis (Baker f.) Verdc.; Verdcourt, F.T.E.A.
(Leguminosae–Papilionoideae 1): 395. 1971.
Shrub. Miombo woodland on steep rocky slope.
1850 m.
Distribution: MR—Carter 2585
LakeNyasa_5-18.indd 54
Gereau et al.
Aeschynomene stolzii Harms; Verdcourt, F.T.E.A.
(Leguminosae–Papilionoideae 1): 388. 1971.
Subshrub or perennial herb. Montane grassland. 2500 m.
Distribution: KP, NL, RM—J. Lovett 776
Aeschynomene cf. trigonocarpa Taub. ex Baker f.;
Verdcourt, F.T.E.A. (Leguminosae–Papilionoideae 1): 399. 1971.
Subshrub. Montane grassland. 2570 m.
Distribution: KP—Gereau 3094
Alysicarpus glumaceus (Vahl) DC. subsp.
glumaceus; Verdcourt, F.T.E.A. (Leguminosae–
Papilionoideae 1): 498. 1971.
Terrestrial herb. Abandoned cultivation. 1250
m.
Distribution: MP—J. Lovett 4077
Alysicarpus rugosus (Willd.) DC. subsp. perennirufus J. Léonard; Verdcourt, F.T.E.A.
(Leguminosae–Papilionoideae 1): 496. 1971.
Terrestrial herb. Grassland. 1250 m.
Distribution: MP—J. Lovett 4443
Amphicarpaea africana (Hook. f.) Harms; Verdcourt, F.T.E.A. (Leguminosae–Papilionoideae
2): 511. 1971.
Herbaceous climber. Montane forest. 2080 m.
Distribution: MR—Mwasumbi 16192
Argyrolobium macrophyllum Harms; Polhill,
F.T.E.A. (Leguminosae–Papilionoideae 2):
1002. 1971.
Terrestrial herb. Roadsides in montane grassland. 1940–2140 m.
Distribution: SL—Suleiman 145
Argyrolobium rupestre (E. Mey.) Walp. subsp.
aberdaricum (Harms) Polhill; Polhill, F.T.E.A.
(Leguminosae–Papilionoideae 2): 1006. 1971.
Terrestrial herb. Montane grassland. 2320–
2740 m.
Distribution: KP, NL, SL—Gereau 4095
Argyrolobium stolzii Harms; Polhill, F.T.E.A.
(Leguminosae–Papilionoideae 2): 1006. 1971.
Terrestrial herb. Montane heathland. 2620–
2770 m.
Distribution: RM—Mwasumbi 16263
Argyrolobium tomentosum (Andrews) Druce; Polhill, F.T.E.A. (Leguminosae–Papilionoideae 2):
1007. 1971.
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Lake Nyasa Climatic Region
Floristic Checklist
Terrestrial herb or shrub. Montane grassland
and bushland. 1560–2300 m.
Distribution: MP, RM, SL—Mwasumbi 16277
Argyrolobium vaginiferum Harms; Polhill, F.T.E.A.
(Leguminosae–Papilionoideae 2): 1008. 1971.
Terrestrial herb or subshrub. Montane grassland and marsh. 2500–2840 m.
Distribution: KP, NL, RM—Goetze 1134 (type:
K), Abeid 2783
Astragalus atropilosulus (Hochst.) Bunge subsp.
burkeanus (Benth. ex Harv.) J. B. Gillett; Gillett, F.T.E.A. (Leguminosae–Papilionoideae 2):
1056. 1971.
Shrub or perennial herb. Montane grassland,
edges of cultivated fields. 2200–2840 m.
Distribution: KP, MP, MR—Suleiman 40
Centrosema pubescens Benth.; Verdcourt, F.T.E.A.
(Leguminosae–Papilionoideae 2): 520. 1971.
Herbaceous climber. Grassland. 1300–1400 m.
Distribution: MP—Kayombo 974
Craibia brevicaudata (Vatke) Dunn subsp. baptistarum (Büttner) J. B. Gillett; Gillett, F.T.E.A.
(Leguminosae–Papilionoideae 1): 151. 1971.
Tree. Riverine forest, montane forest. 1500–
1950 m.
Distribution: MP, RM, SL—J. Lovett 4814
Crotalaria aculeata De Wild. subsp. aculeata; Polhill, Fl. Zamb. 3(7): 186. 2003.
Shrublet. Grassland. 1250–1300 m.
Distribution: MP—J. Lovett 4236
Crotalaria adolfi Harms; Polhill, F.T.E.A. (Leguminosae–Papilionoideae 2): 861. 1971.
Shrub. Riverine forest, montane forest. 900–
2000 m.
Distribution: NL, SL—Stolz 2238 (type: BM,
MO, P, Z), Richards 9856
Crotalaria agatiflora Schweinf. subsp. vaginifera
Polhill; Polhill, F.T.E.A. (Leguminosae–Papilionoideae 2): 853. 1971.
Shrub. Riverine forest, montane forest edges.
1210–2590 m.
Distribution: MP, SL—Procter 1513 (type: EA,
K), Gereau 4247
Crotalaria alexandri Baker f.; Polhill, F.T.E.A.
(Leguminosae–Papilionoideae 2): 992. 1971.
Terrestrial herb. Woodland. 1250 m.
LakeNyasa_5-18.indd 55
55
Distribution: MP—J. Lovett 4504
Crotalaria anthyllopsis Welw. ex Baker; Polhill,
F.T.E.A. (Leguminosae–Papilionoideae 2):
927. 1971.
Terrestrial herb. Abandoned cultivation. 1300–
1400 m.
Distribution: MP—Kayombo 1018
Crotalaria argyrolobioides Baker; Polhill, F.T.E.A.
(Leguminosae–Papilionoideae 2): 975. 1971.
Terrestrial herb. Miombo woodland. 1600 m.
Distribution: MP—P. Lovett 234
Crotalaria becquetii Baker f. ex R. Wilczek subsp.
becquetii; Polhill, F.T.E.A. (Leguminosae–
Papilionoideae 2): 856. 1971.
Shrub. Montane forest and shrubland, riverine
forest. 1500–2240 m.
Distribution: MP, MR, NL, RM—Gereau 4198
Crotalaria brevidens Benth. var. intermedia
(Kotschy) Polhill; Polhill, F.T.E.A. (Leguminosae–Papilionoideae 2): 909. 1971.
Shrub. Montane grassland. 2280 m.
Distribution: MR—Mwasumbi 16169
Crotalaria cephalotes Steud. ex A. Rich.; Polhill,
F.T.E.A. (Leguminosae–Papilionoideae 2):
983. 1971.
Terrestrial herb. Woodland, grassland. 1100–
1400 m.
Distribution: MP—Kayombo 1013
Crotalaria chirindae Baker f.; Polhill, F.T.E.A.
(Leguminosae–Papilionoideae 2): 923. 1971.
Shrub. Edges of cultivated fields. 2200 m.
Distribution: MR—Gereau 4502
Crotalaria cleomifolia Welw. ex Baker; Polhill,
F.T.E.A. (Leguminosae–Papilionoideae 2):
922. 1971.
Shrub or perennial herb. Montane forest edges
and clearings. 1900–2140 m.
Distribution: MR, NL, RM, SL—Suleiman 121
Crotalaria hyssopifolia Klotzsch; Polhill, F.T.E.A.
(Leguminosae–Papilionoideae 2): 991. 1971.
Terrestrial herb. Miombo woodland. 1600–
1800 m.
Distribution: MP—P. Lovett 236
Crotalaria incana L. subsp. purpurascens (Lam.)
Milne-Redh.; Polhill, F.T.E.A. (Leguminosae–
5/31/12 3:59 PM
56
Papilionoidede 2): 870. 1971.
Terrestrial herb. Cultivated fields, waste
ground. 1250–2100 m.
Distribution: MP, MR—Kayombo 2464
Crotalaria lachnocarpoides Engl.; Polhill, F.T.E.A.
(Leguminosae–Papilionoideae 2): 879. 1971.
Shrub or perennial herb. Montane forest edges,
grassland, edges of cultivation. 1250–2250 m.
Distribution: MP, MR, RM, SL—Suleiman 147
Crotalaria natalitia Meisn. var. natalitia; Polhill,
F.T.E.A. (Leguminosae–Papilionoideae 2):
872. 1971.
Shrub. Woodland, riverine forest. 1250–1710
m.
Distribution: MP, RM, SL—Gereau 4798
Crotalaria natalitia Meisn. var. rutshuruensis De
Wild.; Polhill, F.T.E.A. (Leguminosae–Papilionoideae 2): 872. 1971.
Shrub. Montane forest edges and bushland.
2040–2300 m.
Distribution: MP, MR, SL—Gereau 4749
Crotalaria cf. nudiflora Polhill; Polhill, Fl. Zamb.
3(7): 151. 2003.
Terrestrial herb. Miombo woodland. 1780 m.
Distribution: SL—Gereau 5105
Crotalaria ochroleuca G. Don; Polhill, F.T.E.A.
(Leguminosae–Papilionoideae 2): 908. 1971.
Terrestrial herb. Woodland, abandoned cultivation. 1250 m.
Distribution: MP—J. Lovett 4648
Crotalaria oocarpa Baker subsp. oocarpa; Polhill,
F.T.E.A. (Leguminosae–Papilionoideae 2):
968. 1971.
Terrestrial herb. Roadsides in miombo woodland, disturbed riverine forest. 1150–1500 m.
Distribution: MP, SL—Kayombo 2454
Crotalaria recta Steud. ex A. Rich.; Polhill, F.T.E.A.
(Leguminosae–Papilionoideae 2): 957. 1971.
Terrestrial herb. Riverine forest, montane forest margins, wet grassland, woodland, cultivated
fields. 1250–2000 m.
Distribution: MP, RM—Kayombo 2377
Crotalaria ringoetii Baker f.; Polhill, F.T.E.A.
(Leguminosae–Papilionoideae 2): 987. 1971.
Terrestrial herb. Miombo woodland. 1800 m.
LakeNyasa_5-18.indd 56
Gereau et al.
Distribution: MP—P. Lovett 51
Crotalaria rogersii Baker f.; Polhill, F.T.E.A.
(Leguminosae–Papilionoideae 2): 906. 1971.
Terrestrial herb. Lakeshore sand. 480 m.
Distribution: SL—Gereau 3659
Crotalaria seemeniana Harms; Polhill, F.T.E.A.
(Leguminosae–Papilionoideae 2): 975. 1971.
Terrestrial herb. Montane grassland, roadsides,
tree plantations, cultivated ground. 1500–2290 m.
Distribution: MR, RM—Greenway 3581
Crotalaria senegalensis (Pers.) Bacle ex DC.; Polhill, F.T.E.A. (Leguminosae–Papilionoideae 2):
967. 1971.
Terrestrial herb. Cultivated fields. 1250 m.
Distribution: MP—J. Lovett 4623
Crotalaria shirensis (Baker f.) Milne-Redh.; Polhill, F.T.E.A. (Leguminosae–Papilionoideae 2):
946. 1971.
Terrestrial herb. Miombo woodland. 1800 m.
Distribution: MP—P. Lovett 243
Crotalaria sparsifolia Baker; Polhill, F.T.E.A.
(Leguminosae–Papilionoideae 2): 988. 1971.
Terrestrial herb. Miombo woodland. 1475 m.
Distribution: MP—Kayombo 2393
Crotalaria spartea R. Br. ex Baker; Polhill, F.T.E.A.
(Leguminosae–Papilionoideae 2): 937. 1971.
Terrestrial herb. Grassland, abandoned cultivation. 1250–1500 m.
Distribution: MP, SL—Gereau 4405
Crotalaria sphaerocarpa Perr. ex DC.; Polhill,
F.T.E.A. (Leguminosae–Papilionideae 2): 942.
1971.
Terrestrial herb. Open woodland. 1650–1680
m.
Distribution: SL—Gereau 5111
Crotalaria subcapitata De Wild. subsp. subcapitata; Polhill, 1982: 197.
Terrestrial herb. Montane grassland. 1940 m.
Distribution: SL—Gereau 4667
Crotalaria teretifolia Milne-Redh.; Polhill, F.T.E.A.
(Leguminosae–Papilionoideae 2): 946. 1971.
Terrestrial herb. Grassland. 1250 m.
Distribution: MP—J. Lovett 4390
Crotalaria valida Baker; Polhill, F.T.E.A. (Legu-
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Lake Nyasa Climatic Region
Floristic Checklist
57
minosae–Papilionoideae 2): 879. 1971.
Terrestrial herb. Montane grassland. 1810–
1860 m.
Distribution: KP, SL—Gereau 3923
469. 1971.
Shrub. Riverine forest, edges of rice fields.
600–1210 m.
Distribution: MR, SL—Gereau 4251
Dalbergia arbutifolia Baker subsp. arbutifolia;
Polhill, F.T.E.A. (Leguminosae–Papilionoideae
1): 108. 1971.
Liana. Riverine forest, lakeshore thickets.
474–1400 m.
Distribution: MP, SL—Gereau 3674
Desmodium tortuosum (Sw.) DC.; Schubert,
F.T.E.A. (Leguminosae–Papilionoideae 1):
474. 1971.
Terrestrial herb. Lakeshore forest. 474 m.
Distribution: SL—Gereau 3672
Dalbergia fischeri Taub.; Polhill, F.T.E.A. (Leguminosae–Papilionoideae 1): 110. 1971.
Liana or scandent shrub. Riverine forest, lakeshore woodland. 474–1210 m.
Distribution: MP, SL—Mwasumbi 16589
Dalbergia lactea Vatke; Polhill, F.T.E.A. (Leguminosae–Papilionoideae 1): 111. 1971.
Liana or shrub. Montane forest, riverine forest.
1290–2100 m.
Distribution: MP, RM, SL—Gereau 3613
Desmodium adscendens (Sw.) DC.; Schubert,
F.T.E.A. (Leguminosae–Papilionoideae 1):
461. 1971.
Terrestrial herb. Grassland. 1360 m.
Distribution: RM—Hooper 845
Desmodium appressipilum B. G. Schub.; Schubert,
F.T.E.A. (Leguminosae–Papilionoideae 1):
458. 1971.
Terrestrial herb. Woodland, montane grassland. 1250–2100 m.
Distribution: KP, MP—Richards 14184 (type:
A, K), J. Lovett 4505
Desmodium barbatum (L.) Benth. var. argyreum
(Welw. ex Baker) B. G. Schub.; Schubert,
F.T.E.A. (Leguminosae–Papilionoideae 1):
479. 1971.
Woody-based herb. Woodland. 1400–1500 m.
Distribution: MP—J. Lovett 3400
Desmodium repandum (Vahl) DC.; Schubert,
F.T.E.A. (Leguminosae–Papilionoideae 1):
465. 1971.
Terrestrial herb, often scrambling. Montane
grassland, montane forest. 1780–2360 m.
Distribution: MR, NL, RM, SL—Suleiman 102
Desmodium salicifolium (Poir.) DC.; Schubert,
F.T.E.A. (Leguminosae–Papilionoideae 1):
LakeNyasa_5-18.indd 57
Dolichos
kilimandscharicus
Taub.
subsp.
kilimandscharicus; Verdcourt, F.T.E.A. (Leguminosae–Papilionoideae 2): 685. 1971.
Terrestrial herb. Montane grassland, montane
forest edges, woodland. 1300–2250 m.
Distribution: MP, RM, SL—Gereau 3583
Dolichos sericeus E. Mey. subsp. sericeus; Verdcourt, F.T.E.A. (Leguminosae–Papilionoideae
2): 680. 1971.
Herbaceous climber. Montane forest edges,
shrubland. 2180–2400 m.
Distribution: MR—Mwasumbi 16175
Dolichos simplicifolius Hook. f.; Verdcourt, F.T.E.A.
(Leguminosae–Papilionoideae 2): 692. 1971.
Terrestrial herb. Seasonally inundated grassland, woodland. 1270–1310 m.
Distribution: MP—Gereau 3478
Dolichos trinervatus Baker; Verdcourt, F.T.E.A.
(Leguminosae–Papilionoideae 2): 688. 1971.
Terrestrial herb. Miombo woodland. 1200–
1500 m.
Distribution: MP—Gereau 7001
Dolichos xiphophyllus Baker; Verdcourt, F.T.E.A.
(Leguminosae–Papilionoideae 2): 691. 1971.
Terrestrial herb. Grassland. 1250 m.
Distribution: MP—Lovett 3645
Dumasia villosa DC.; Verdcourt, F.T.E.A. (Leguminosae–Papilionoideae 2): 513. 1971.
Herbaceous climber. Montane forest. 1790–
2100 m.
Distribution: MR, RM—Mwasumbi 16310
Eriosema buchananii Baker f. var. buchananii;
Verdcourt, F.T.E.A. (Leguminosae–Papilionoideae 2): 776. 1971.
Terrestrial herb. Montane grassland. 2120 m.
Distribution: SL—Gereau 4329
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58
Eriosema buchananii Baker f. var. subprostratum
Verdc.; Verdcourt, F.T.E.A. (Leguminosae–
Papilionoideae 2): 777. 1971.
Terrestrial herb. Montane grassland. 2250–
2840 m.
Distribution: KP, MR, RM—Abeid 2780
Eriosema ellipticum Welw. ex Baker; Verdcourt,
F.T.E.A. (Leguminosae–Papilionoideae 2):
768. 1971.
Shrub or subshrub. Woodland, wooded grassland. 900–1400 m.
Distribution: MP, RM, SL—Kibure 1483
Gereau et al.
Terrestrial herb. Grassland.
Distribution: RM—Eggeling 6518
Eriosema shirense Baker f.; Verdcourt, F.T.E.A.
(Leguminosae–Papilionoideae 2): 798. 1971.
Terrestrial herb. Woodland. 1200–1790 m.
Distribution: MP, SL—Gereau 4381
Eriosema ukingense Harms; Verdcourt, F.T.E.A.
(Leguminosae–Papilionoideae 2): 777. 1971.
Terrestrial herb. Rocky mountain slopes. 2800
m.
Distribution: KP, MP—Goldblatt 8207
Eriosema flexuosum Staner; Verdcourt, F.T.E.A.
(Leguminosae–Papilionoideae 2): 804. 1971.
Terrestrial herb. Montane grassland. 2500 m.
Distribution: KP—Abeid 3110
Eriosema verdickii De Wild.; Verdcourt, F.T.E.A.
(Leguminosae–Papilionoideae 2): 803. 1971.
Terrestrial herb. Woodland. 1250–1790 m.
Distribution: MP, SL—Gereau 3893
Eriosema lebrunii Staner & De Craene; Verdcourt,
F.T.E.A. (Leguminosae–Papilionoideae 2):
802. 1971.
Terrestrial herb. Montane grassland. 2120–
2488 m.
Distribution: SL—Suleiman 181
Erythrina abyssinica Lam.; Verdcourt, F.T.E.A.
(Leguminosae–Papilionoideae 2): 555. 1971.
Tree. Montane forest remnants, forest edges,
riverine forest, woodland, wooded grassland.
1300–1840 m.
Distribution: MP, SL—Gereau 3547
Eriosema
macrostipulum
Baker
f.
var.
macrostipulum; Verdcourt, F.T.E.A. (Leguminosae–Papilionoideae 2): 789. 1971.
Terrestrial herb. Woodland. 1200 m.
Distribution: MP—J. Lovett 3867
Flemingia grahamiana Wight & Arn.; Verdcourt,
F.T.E.A. (Leguminosae–Papilionoideae 2):
806. 1971.
Terrestrial herb. Wooded grassland. 980 m.
Distribution: SL—Kibure 1485
Eriosema montanum Baker f. var. brevipedunculatum;
Verdcourt,
F.T.E.A.
(Leguminosae–Papilionoideae 2): 78. 1971.
Shrub. Edges of cultivated fields. 2200 m.
Distribution: MR—Gereau 4509
Humularia drepanocephala (Baker) P. A. Duvign.
var. drepanocephala; Verdcourt, F.T.E.A.
(Leguminosae–Papilionoideae 1): 431. 1971.
Terrestrial herb. Woodland. 1400–2250 m.
Distribution: MP—J. Lovett 4853
Eriosema montanum Baker f. var. montanum;
Verdcourt, F.T.E.A. (Leguminosae–Papilionoideae 2): 780. 1971.
Shrub. Montane forest, forest edges. 2040–
2100 m.
Distribution: MR, SL—Gereau 4751
Indigofera arrecta Hochst. ex. A. Rich.; Gillett,
F.T.E.A. (Leguminosae–Papilionoideae 1):
307. 1971.
Woody-based herb. Woodland, montane grassland. 1350–2520 m.
Distribution: KP, MP—J. Lovett 4691
Eriosema nutans Schinz; Verdcourt, F.T.E.A.
(Leguminosae–Papilionoideae 2): 775. 1971.
Terrestrial herb. Montane grassland. 2230–
2360 m.
Distribution: MR, SL—Gereau 3773
Indigofera asterocalycina Gilli; Gillett, F.T.E.A.
(Leguminosae–Papilionoideae 1): 276. 1971.
Shrub. Cultivated cassava fields. 474–480 m.
Distribution: SL—Mwasumbi 16575
Eriosema parviflorum E. Mey. subsp. parviflorum;
Verdcourt, F.T.E.A. (Leguminosae–Papilionoideae 2): 779. 1971.
LakeNyasa_5-18.indd 58
Indigofera astragalina DC.; Gillett, F.T.E.A.
(Leguminosae–Papilionoideae 1): 310. 1971.
Terrestrial herb. Lakeshore shrubland, abandoned cultivation. 474–1250 m.
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Lake Nyasa Climatic Region
Floristic Checklist
Distribution: MP, SL—Mwasumbi 16576
Indigofera atricephala J. B. Gillett; Gillett, F.T.E.A.
(Leguminosae–Papilionoideae 1): 284. 1971.
Terrestrial herb. Disturbed montane forest.
2280–2400 m.
Distribution: RM—Mwasumbi 16243
Indigofera atriceps Hook. f. subsp. atriceps; Gillett, F.T.E.A. (Leguminosae–Papilionoideae 1):
282. 1971.
Shrub. Miombo woodland, montane forest
edges, riverine forest. 1500–2400 m.
Distribution: MR, RM, SL—Gereau 4501
Indigofera atriceps Hook. f. subsp. glandulosissima (R. E. Fr.) J. B. Gillett; Gillett, F.T.E.A.
(Leguminosae–Papilionoideae 1): 282. 1971.
Woody-based herb. Miombo woodland. 1300–
1800 m.
Distribution: MP—P. Lovett 60
Indigofera atriceps Hook. f. vel sp. aff., Taxon “B”;
Gillett, F.T.E.A. (Leguminosae–Papilionoideae
1): 284. 1971.
Terrestrial herb. Montane grassland. 2100–
2600 m.
Distribution: KP—J. Lovett 1854
Indigofera demissa Taub.; Gillett, F.T.E.A. (Leguminosae–Papilionoideae 1): 225. 1971.
Terrestrial herb. Abandoned cultivation. 480–
900 m.
Distribution: NL—Pócs 87649
Indigofera dendroides Jacq.; Gillett, F.T.E.A.
(Leguminosae–Papilionoideae 1): 242. 1971.
Woody-based terrestrial herb. Wooded grassland, montane thickets. 1400–2400 m.
Distribution: MP, MR—Kayombo 2356
Indigofera emarginella Steud. ex A. Rich.; Gillett,
F.T.E.A. (Leguminosae–Papilionoideae 1):
300. 1971.
Shrublet. Miombo woodland. 1200–1790 m.
Distribution: MP, SL—Kayombo 2424
Indigofera fulvopilosa Brenan; Gillett, F.T.E.A.
(Leguminosae–Papilionideae 1): 257. 1971.
Terrestrial herb. Premontane shrubland. 1610
m.
Distribution: RM—Mwasumbi 16344
Indigofera fuscosetosa Baker; Gillett, F.T.E.A.
LakeNyasa_5-18.indd 59
59
(Leguminosae–Papilionoideae 1): 279. 1971.
Terrestrial herb. Montane grassland, premontane shrubland. 1610–1940 m.
Distribution: RM, SL—Mwasumbi 16339
Indigofera homblei Baker f. & Martin subsp.
homblei; Gillett, F.T.E.A. (Leguminosae–Papilionoideae 1): 289. 1971.
Shrub. Miombo woodland, riverine forest
edges. 1475–1800 m.
Distribution: MP, SL—Kayombo 2391
Indigofera masaiensis J. B. Gillett; Gillett, F.T.E.A.
(Leguminosae–Papilionoideae 1): 264. 1971.
Scrambling terrestrial herb. Bushy montane
grassland. 2100 m.
Distribution: MP—P. Lovett 406
Indigofera mimosoides Baker; Gillett, F.T.E.A.
(Leguminosae–Papilionoideae 1): 262. 1971.
Terrestrial herb. Montane shrubland. 1760–
2540 m.
Distribution: KP, MP, NL—Abeid 2871
Indigofera paniculata Vahl ex Pers. subsp.
paniculata; Gillett, F.T.E.A. (Leguminosae–
Papilionoideae 1): 232. 1971.
Shrub. Riverine forest edges in miombo woodland. 1500 m.
Distribution: SL—Gereau 4411
Indigofera rhynchocarpa Welw. ex Baker var.
rhynchocarpa; Gillett, F.T.E.A. (Leguminosae–
Papilionoideae 1): 297. 1971.
Shrub. Miombo woodland, field edges, and
abandoned cultivation. 1250–1800 m.
Distribution: MP—P. Lovett 250
Indigofera roseocaerulea Baker f.; Gillett, F.T.E.A.
(Leguminosae–Papilionoideae 1): 299. 1971.
Shrub. Montane forest edges, bushy montane
grassland. 1780–2230 m.
Distribution: MP, MR—Mwasumbi 16174
Indigofera schliebenii Harms; Gillett, F.T.E.A.
(Leguminosae–Papilionoideae 1): 284. 1971.
Terrestrial herb. Montane grassland. 1940 m.
Distribution: SL—Gereau 4687
Indigofera subulifera Welw. ex Baker; Gillett,
F.T.E.A. (Leguminosae–Papilionoideae 1):
274. 1971.
Terrestrial herb. Miombo woodland. 1790 m.
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Gereau et al.
Distribution: SL—Gereau 3904
Indigofera trachyphylla Benth. ex Oliv.; Gillett,
F.T.E.A. (Leguminosae–Papilionoideae 1):
278. 1971.
Terrestrial herb. Miombo woodland. 1530 m.
Distribution: SL—Gereau 4373
Indigofera ugandensis Baker f.; Gillett, F.T.E.A.
(Leguminosae–Papilionoideae 1): 273. 1971.
Terrestrial herb. Montane grassland. 1920 m.
Distribution: SL—Gereau 4671
Indigofera zenkeri Harms ex Baker f.; Gillett,
F.T.E.A. (Leguminosae–Papilionoideae 1):
265. 1971.
Terrestrial herb. Lakeshore shrubland. 474–
480 m.
Distribution: SL—Mwasumbi 16580
Kotschya africana Endl.; Verdcourt, F.T.E.A.
(Leguminosae–Papilionoideae 1): 412. 1971.
Shrub. Riverine forest. 1500 m.
Distribution: SL—Gereau 4780
Kotschya coalescens Dewit & P. A. Duvign.; Verdcourt, F.T.E.A. (Leguminosae–Papilionoideae
1): 425. 1971.
Shrub. Shrubby grassland. 2600 m.
Distribution: MP, MR—Iwarsson 1187
Kotschya recurvifolia (Taub.) F. White subsp.
recurvifolia; Verdcourt, F.T.E.A. (Leguminosae–Papilionoideae 1): 417. 1971.
Shrub or tree. Montane heathland, forest edges,
wooded grassland. 2100–2770 m.
Distribution: MP, RM, SL—Mwasumbi 16260
Kotschya strigosa (Benth.) Dewit & P. A. Duvign.
var. strigosa; Verdcourt, F.T.E.A. (Leguminosae–Papilionoideae 1): 423. 1971.
Shrub or tree. Woodland, montane grassland,
bamboo forest. 1400–2200 m.
Distribution: MP, RM, SL—Gereau 4689
Kotschya thymodora (Baker f.) Wild subsp.
septentrionalis Verdc.; Verdcourt, F.T.E.A.
(Leguminosae–Papilionoideae 1): 416. 1971.
Shrub. Montane grassland and heathland,
bushed grassland. 1940–2960 m.
Distribution: KP, RM, SL—Abeid 2867
Lonchocarpus capassa Rolfe; Polhill, F.T.E.A.
(Leguminosae–Papilionoideae 1): 66. 1971.
LakeNyasa_5-18.indd 60
Tree. Miombo woodland, lakeshore forest.
480–1300 m.
Distribution: MP, SL—Gereau 3651
Lotononis angolensis Welw. ex Baker; Milne-Redhead, F.T.E.A. (Leguminosae–Papilionoideae
2): 814. 1971.
Terrestrial herb. Montane grassland, roadsides
in miombo woodland. 1500–1940 m.
Distribution: RM, SL—Mwasumbi 16342
Lotononis stolzii Harms; Milne-Redhead, F.T.E.A.
(Leguminosae–Papilionoideae 2): 816. 1971.
Terrestrial herb. Montane grassland. 2300–
2600 m.
Distribution: KP, NL, SL—Stolz 2602 (type:
BM, BOL, C, EA, K, MO, PRE), J. Lovett 1855
Lotus discolor E. Mey.; Gillett, F.T.E.A. (Leguminosae–Papilionoideae 2): 1047. 1971.
Terrestrial herb. Woodland, montane grassland. 1400–2740 m.
Distribution: KP, MP—J. Lovett 3398
Lotus subdigitatus Boutique; Gillett, F.T.E.A.
(Leguminosae–Papilionoideae 2): 1047. 1971.
Terrestrial herb. Montane grassland. 2500 m.
Distribution: RM—Mwasumbi 16252
Macrotyloma axillare (E. Mey.) Verdc.; Verdcourt,
F.T.E.A. (Leguminosae–Papilionoideae 2):
586. 1971.
Herbaceous climber. Miombo woodland. 1780
m.
Distribution: SL—Gereau 5094
Macrotyloma ellipticum (R. E. Fr.) Verdc.; Verdcourt, F.T.E.A. (Leguminosae–Papilionoideae
2): 589. 1971.
Terrestrial herb. Woodland. 1600 m.
Distribution: RM—Stolz 1335
Macrotyloma fimbriatum (Harms) Verdc.; Verdcourt, F.T.E.A. (Leguminosae–Papilionoideae
2): 591. 1971.
Terrestrial herb. Miombo woodland. 1530 m.
Distribution: SL—Gereau 4369
Microcharis ephemera (J. B. Gillet) Schrire;
Schrire, 1992: 166.
Terrestrial herb. Woodland. 1300–1600 m.
Distribution: MP—P. Lovett 99
Millettia oblata Dunn subsp. intermedia J. B.
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Floristic Checklist
Gillett; Gillett, F.T.E.A. (Leguminosae–Papilionoideae 1): 143. 1971.
Tree. Montane forest. 1780 m.
Distribution: RM—Bracebridge 29
Millettia oblata Dunn subsp. stolzii J. B. Gillett;
Gillett, F.T.E.A. (Leguminosae–Papilionoideae
1): 143. 1971.
Tree. Montane forest. 1400–1500 m.
Distribution: RM—Stolz 183 (type: K, MO),
Davies 301
Mucuna pruriens (L.) DC. var. pruriens; Verdcourt, F.T.E.A. (Leguminosae–Papilionoideae
2): 567. 1971.
Herbaceous climber. Riverine forest. 1250 m.
Distribution: MP—J. Lovett 4481
Neonotonia wightii (Graham ex Wight & Arn.)
J. A. Lackey subsp. wightii; Mackinder, Fl.
Zamb. 3(5): 56. 2001.
Herbaceous climber. Roadsides in cultivation.
2200 m.
Distribution: MR—Gereau 4505
Neorautanenia mitis (A. Rich.) Verdc.; Verdcourt,
F.T.E.A. (Leguminosae–Papilionoideae 2):
700. 1971.
Terrestrial herb, sometimes climbing. Woodland, riverine forest edges. 1250–1600 m.
Distribution: MP, SL—J. Lovett 4288
Ormocarpum kirkii S. Moore; Gillett, F.T.E.A.
(Leguminosae–Papilionoideae 1): 362. 1971.
Shrub or small tree. Wooded grassland, roadsides. 490–880 m.
Distribution: SL—Kibure 1487
Otholobium foliosum (Oliv.) C. H. Stirt.; Verdcourt,
Fl. Zamb. 3(6): 46. 2000.
Shrub. Bamboo forest edges, montane grassland. 2270–2360 m.
Distribution: KP, NL—Bracebridge 111
Pericopsis angolensis (Baker) Meeuwen; Polhill,
F.T.E.A. (Leguminosae–Papilionoideae 1): 41.
1971.
Tree. Woodland. 1200–1400 m.
Distribution: MP—Kayombo 2414
Pseudarthria hookeri Wight & Arn. var. hookeri;
Verdcourt, F.T.E.A. (Leguminosae–Papilionoideae 1): 484. 1971.
Shrub or perennial herb. Woodland, montane
LakeNyasa_5-18.indd 61
61
grassland, riverine forest. 1200–2440 m.
Distribution: MP, MR, SL—Suleiman 188
Pterocarpus angolensis DC.; Polhill, F.T.E.A.
(Leguminosae–Papilionoideae 1): 89. 1971.
Tree. Miombo woodland. 1200–1300 m.
Distribution: MP—J. Lovett 4908
Pterocarpus tinctorius Welw.; Polhill, F.T.E.A.
(Leguminosae–Papilionoideae 1): 86. 1971.
Tree. Miombo woodland. 1200 m.
Distribution: MP—Kayombo 2438
Rhynchosia caribaea (Jacq.) DC. var. caribaea;
Verdcourt, Fl. Zamb. 3(5): 195. 2001.
Herbaceous climber. Riverine forest. 1100 m.
Distribution: MP—Mwangoka 664
Note: New to East Africa, from Malawi,
Mozambique, Zimbabwe, and South Africa.
Rhynchosia clivorum S. Moore subsp. pycnantha
(Harms) Verdc.; Verdcourt, Fl. Zamb. 3(5):
179. 2001.
Shrub. Montane grassland and bushland.
2100–2550 m.
Distribution: MP, RM—Kerfoot 4117
Rhynchosia goetzei Harms var. goetzei; Verdcourt,
F.T.E.A. (Leguminosae–Papilionoideae 2):
744. 1971.
Shrub. Woodland. 1400–1500 m.
Distribution: MP—J. Lovett 3384
Rhynchosia luteola (Hiern) K. Schum. var. luteola;
Verdourt, F.T.E.A. (Leguminosae–Papilionoideae 2): 732. 1971.
Herbaceous climber. Riverine forest. 1300 m.
Distribution: MP—J. Lovett 4779
Rhynchosia nyikensis Baker; Verdcourt, F.T.E.A.
(Leguminosae–Papilionoideae 2): 729. 1971.
Herbaceous climber. Montane forest. 2040 m.
Distribution: SL—Gereau 4738
Sesbania macrantha Welw. ex E. Phillips & Hutch.
var. acrantha; Gillett, F.T.E.A. (Leguminosae–
Papilionoideae 1): 341. 1971.
Shrub. Wooded grassland, woodland, cultivated fields. 1250–1980 m.
Distribution: MP—Kayombo 2463
Sesbania sesban (L.) Merr. var. nubica Chiov.; Gillett, F.T.E.A. (Leguminosae–Papilionoideae 1):
339. 1971.
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Gereau et al.
Shrub. Sandy lakeshore. 474–490 m.
Distribution: SL—Mwasumbi 16562
Sphenostylis stenocarpa (Hochst. ex A. Rich.)
Harms; Verdcourt, F.T.E.A. (Leguminosae–
Papilionoideae 2): 672. 1971.
Herbaceous climber. Miombo woodland. 1530
m.
Distribution: SL—Gereau 4355
Tephrosia aequilata Baker; Gillett, F.T.E.A. (Leguminosae–Papilionoideae 1): 207. 1971.
Shrub. Montane forest. 1760 m.
Distribution: RM—WCS SHCP 260
Terrestrial herb. Grassland. 1250 m.
Distribution: MP—J. Lovett 4231
Tephrosia richardsiae J. B. Gillett; Gillett, F.T.E.A.
(Leguminosae–Papilionoideae 1): 178. 1971.
Terrestrial herb. Woodland. 1250 m.
Distribution: MP—J. Lovett 4340
Tephrosia stormsii De Wild. var. pilosa Brummitt;
Gillett, F.T.E.A. (Leguminosae–Papilionoideae
1): 198. 1971.
Terrestrial herb. Roadsides. 474–480 m.
Distribution: SL—Mwasumbi 16572
Tephrosia decora Welw. ex Baker; Gillett, F.T.E.A.
(Leguminosae–Papilionoideae 1): 180. 1971.
Terrestrial herb. Woodland. 1200 m.
Distribution: MP—Kayombo 2415
Tephrosia vogelii Hook. f.; Gillett, F.T.E.A. (Leguminosae–Papilionoideae 1): 210. 1971.
Terrestrial herb or soft-wooded shrub. Montane
forest edges. 2100 m.
Distribution: RM—WCS SHCP 178
Tephrosia interrupta Hochst. & Steud. ex Engl.
subsp. elongatiflora Harms ex J. B. Gillett; Gillett, F.T.E.A. (Leguminosae–Papilionoideae 1):
206. 1971.
Shrub. Montane forest edges. 2110 m.
Distribution: MR—Gereau 4550
Teramnus uncinatus (L.) Sw. subsp. axilliflorus
(Kotschy) Verdc.; Verdcourt, F.T.E.A. (Leguminosae–Papilionoideae 2): 540. 1971.
Herbaceous climber. Woodland. 1300–1400 m.
Distribution: MP—Kayombo 997
Tephrosia interrupta Hochst. & Steud. ex Engl.
subsp. mildbraedii (Harms) J. B. Gillett; Gillett, F.T.E.A. (Leguminosae–Papilionoideae 1):
206. 1971.
Shrub. Edges of cultivated fields. 2200 m.
Distribution: MR—Gereau 4538
Tephrosia nyikensis Baker; Gillett, F.T.E.A. (Leguminosae–Papilionoideae 1): 203. 1971.
Shrub. Miombo woodland. 1700 m.
Distribution: MP—Kayombo 1113
Tephrosia paniculata Welw. ex Baker; Gillett,
F.T.E.A. (Leguminosae–Papilionoideae 1):
176. 1971.
Shrub. Stream banks, edges of cultivated fields.
1500–2200 m.
Distribution: MP, MR, RM—Mwasumbi 16276
Tephrosia paucijuga Harms; Gillett, F.T.E.A.
(Leguminosae–Papilionoideae 1): 195. 1971.
Terrestrial herb. Montane grassland. 2440 m.
Distribution: SL—Suleiman 187
Tephrosia purpurea (L.) Pers. subsp. leptostachya
(DC.) Brummitt; Gillett, F.T.E.A. (Leguminosae–Papilionoideae 1): 186. 1971.
LakeNyasa_5-18.indd 62
Trifolium repens L.; Gillett, F.T.E.A. (Leguminosae–Papilionoideae 2): 1016. 1971.
Terrestrial herb. Montane grassland. 2400 m.
Distribution: KP—Suleiman 42
Trifolium semipilosum Fresen. var. semipilosum;
Gillett, F.T.E.A. (Leguminosae–Papilionoideae
2): 1027. 1971.
Terrestrial herb. Montane grassland. 2360 m.
Distribution: MP, MR—Gereau 3748
Trifolium simense Fresen.; Gillett, F.T.E.A. (Leguminosae–Papilionoideae 2): 1022. 1971.
Terrestrial herb. Montane grassland. 2250–
2640 m.
Distribution: KP, MP, RM—Suleiman 56
Trifolium tembense Fresen.; Gillett, F.T.E.A.
(Leguminosae–Papilionoideae 2): 1033. 1971.
Terrestrial herb. Montane grassland. 2400–
2570 m.
Distribution: KP—Gereau 3105
Trifolium wentzelianum Harms; Gillett, F.T.E.A.
(Leguminosae–Papilionoideae 2): 1026. 1971.
Terrestrial herb. Montane grassland. 2250–
2590 m.
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Floristic Checklist
Distribution: KP, MP, MR, RM—Mwasumbi
16165
Vicia paucifolia Baker subsp. malosana (Baker)
Verdc.; Verdcourt, F.T.E.A. (Leguminosae–
Papilionoideae 2): 1070. 1971.
Terrestrial herb. Montane grassland. 2700 m.
Distribution: NL—Gereau 4090
Vigna frutescens A. Rich. subsp. frutescens; Verdcourt, F.T.E.A. (Leguminosae–Papilionoideae
2): 648. 1971.
Herbaceous climber. Woodland. 1400–1500 m.
Distribution: MP—J. Lovett 3379
Vigna triphylla (R. Wilczek) Verdc.; Verdcourt,
F.T.E.A. (Leguminosae–Papilionoideae 2):
663. 1971.
Twining herb. Miombo woodland. 1530 m.
Distribution: SL—Gereau 4378
Vigna unguiculata (L.) Walp. subsp. dekindtiana
(Harms) Verdc.; Verdcourt, F.T.E.A. (Leguminosae–Papilionoideae 2): 644. 1971.
Herbaceous climber. Open woodland. 1650–
1680 m.
Distribution: SL—Gereau 5118
Vigna verdcourtii Pasquet; Pasquet, 2007: 53
Herbaceous climber. Edges of pyrethrum fields.
2200 m.
Distribution: MR—Gereau 4499
Vigna vexillata (L.) A. Rich. var. vexillata;
Verdcourt, F.T.E.A. (Leguminosae–Papilionoideae 2): 653. 1971.
Herbaceous climber. Woodland, cultivated
fields, riverine forest. 1250–1400 m.
Distribution: MP—Kayombo 967
63
Dovyalis abyssinica (A. Rich.) Warb.; Sleumer,
F.T.E.A. (Flacourtiaceae): 61. 1975.
Shrub or tree. Riverine forest, montane forest.
1250–1990 m.
Distribution: MP, RM—Kayombo 4078
Dovyalis hebecarpa (Gardner) Warb.; Sleumer,
1972: 69.
Shrub. Montane forest. 2040 m.
Distribution: SL—Gereau 4730
Flacourtia indica (Burm. f.) Merr.; Sleumer,
F.T.E.A. (Flacourtiaceae): 57. 1975.
Tree or shrub. Montane forest, riverine forest,
miombo woodland. 1150–2110 m.
Distribution: MP, MR, RM, SL—Kayombo
2446
Kiggelaria africana L.; Sleumer, F.T.E.A. (Flacourtiaceae): 31. 1975.
Tree or shrub. Montane forest, forest edges and
clearings. 2200–2500 m.
Distribution: KP, MP, MR, RM, SL—Suleiman
10
Lindackeria fragrans (Gilg) Gilg; Sleumer, F.T.E.A.
(Flacourtiaceae): 25. 1975.
Tree or shrub. Woodland. 1200–1310 m.
Distribution: MP—J. Lovett 5018
Oncoba spinosa Forssk.; Sleumer, F.T.E.A. (Flacourtiaceae): 16. 1975.
Shrub. Woodland, riverine forest. 1100–1400
m.
Distribution: MP—Kayombo 979
8 genera and 10 species
Rawsonia lucida Harv. & Sond.; Sleumer, F.T.E.A.
(Flacourtiaceae): 4. 1975.
Tree or shrub. Riverine forest, montane forest.
474–1900 m.
Distribution: MP, NL, RM, SL—Mwasumbi
16599
Aphloia theiformis (Vahl) Benn.; Lovett et al.,
2006: 95.
Tree. Montane forest. 1850–1970 m.
Distribution: KP, RM—Kibure 1395
Rawsonia reticulata Gilg; Sleumer, F.T.E.A. (Flacourtiaceae): 6. 1975.
Tree or shrub. Montane forest. 1750–2070 m.
Distribution: NL, RM—Gereau 4205
FLACOURTIACEAE
Casearia battiscombei R. E. Fr.; Sleumer, F.T.E.A.
(Flacourtiaceae): 49. 1975.
Tree. Montane forest, riverine forest. 1780–
2880 m.
Distribution: MP, MR, RM—Mwasumbi 16302
LakeNyasa_5-18.indd 63
FUMARIACEAE
1 genus and 1 species
Fumaria abyssinica Hammar; Lucas, F.T.E.A.
(Fumariaceae): 3. 1962.
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64
Terrestrial herb. Cultivated pyrethrum fields.
2200 m.
Distribution: MR, RM—Gereau 4512
GENTIANACEAE
4 genera and 12 species
Chironia laxiflora Baker; Sileshi, F.T.E.A. (Gentianaceae): 44. 2002.
Terrestrial herb. Montane grassland, miombo
woodland. 1800–2350 m.
Distribution: MP, MR, RM, SL—Mwasumbi
16204
Faroa pusilla Baker; Sileshi, F.T.E.A. (Gentianaceae): 34. 2002.
Terrestrial herb. Woodland. 1250–1350 m.
Distribution: MP—J. Lovett 4491
Sebaea bojeri Griseb. vel sp. aff.; Sileshi, F.T.E.A.
(Gentianaceae): 12. 2002.
Terrestrial herb. Wooded grassland. 1250 m.
Distribution: MP—J. Lovett 4508
Sebaea brachyphylla Griseb.; Sileshi, F.T.E.A.
(Gentianaceae): 7. 2002.
Terrestrial herb. Montane grassland and stream
banks. 2500–2720 m.
Distribution: KP, NL—Gereau 4630
Gereau et al.
(Gentianaceae): 59. 2002.
Terrestrial herb. Montane grassland and heathland. 2400–2850 m.
Distribution: KP, RM—Suleiman 35
Swertia squamigera Sileshi; Sileshi, F.T.E.A. (Gentianaceae): 63. 2002.
Terrestrial herb. Montane grassland and forest
margins. 1980–2600 m.
Distribution: MP, MR, NL, RM—Mgaza 478
(type: EA, K), Mhoro 1394
Swertia usambarensis Engl. var. curtioides (Gilg)
Sileshi; Sileshi, F.T.E.A. (Gentianaceae): 55.
2002.
Terrestrial herb. Montane grassland. 2400 m.
Distribution: KP, MP—Suleiman 57
Swertia usambarensis Engl. var. usambarensis;
Sileshi, F.T.E.A. (Gentianaceae): 53. 2002.
Terrestrial herb. Montane grassland. 2000 m.
Distribution: KP, MR—J. Lovett 1770
Swertia welwitschii Engl.; Sileshi, F.T.E.A. (Gentianaceae): 55. 2002.
Terrestrial herb. Disturbed montane grassland.
2280 m.
Distribution: MR—Mwasumbi 16164
GERANIACEAE
Sebaea grandis (E. Mey.) Steud.; Sileshi, F.T.E.A.
(Gentianaceae): 13. 2002.
Terrestrial herb. Montane grassland, abandoned cultivation. 1200–1940 m.
Distribution: MP, SL—Gereau 4674
Sebaea leiostyla Gilg; Sileshi, F.T.E.A. (Gentianaceae): 9. 2002.
Terrestrial herb. Montane grassland. 2500 m.
Distribution: KP, MR—J. Lovett 762
Sebaea longicaulis Schinz; Sileshi, F.T.E.A. (Gentianaceae): 9. 2002.
Terrestrial herb. Stream banks in montane
grassland. 2830 m.
Distribution: KP—Abeid 2855
Sebaea pumila (Baker) Schinz; Sileshi, F.T.E.A.
(Gentianaceae): 16. 2002.
Terrestrial herb. Miombo woodland. 1800 m.
Distribution: MP—P. Lovett 248
Swertia kilimandscharica Engl.; Sileshi, F.T.E.A.
LakeNyasa_5-18.indd 64
2 genera and 5 species
Geranium aculeolatum Oliv.; Kokwaro, F.T.E.A.
(Geraniaceae): 2. 1971.
Terrestrial herb. Montane forest. 2100 m.
Distribution: MR—Mwasumbi 16304
Geranium incanum Burm. f.; Kokwaro, F.T.E.A.
(Geraniaceae): 3. 1971.
Terrestrial herb. Montane grassland and forest
edges. 2250–2700 m.
Distribution: KP, MP, MR, NL, RM—Suleiman
39
Geranium vagans Baker subsp. vagans; Kokwaro,
F.T.E.A. (Geraniaceae): 5. 1971.
Terrestrial herb. Montane grassland. 2250–
2900 m.
Distribution: KP, MR, RM—Gereau 3739
Pelargonium luridum (Andrews) Sweet; F.T.E.A.
(Geraniaceae): 15. 1971.
Terrestrial herb. Woodland, montane grass-
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Lake Nyasa Climatic Region
Floristic Checklist
land. 1950–2550 m.
Distribution: KP, MP—J. Lovett 3426
Pelargonium whytei Baker; Kokwaro, F.T.E.A.
(Geraniaceae): 16. 1971.
Terrestrial herb. Montane forest. 2500 m.
Distribution: MP, MR—P. Lovett 484A
GESNERIACEAE
1 genus and 4 species
Streptocarpus buchananii C. B. Clarke; Darbyshire, F.T.E.A. (Gesneriaceae): 23. 2006.
Epiphyte. Montane forest. 1500–2600 m.
Distribution: MR, NL, RM—Gereau 4161
Streptocarpus eylesii S. Moore subsp. brevistylus
Hilliard & B. L. Burtt; Darbyshire, F.T.E.A.
(Gesneriaceae): 44. 2006.
Terrestrial herb.
Distribution: MP, SL—Richards 14085
Streptocarpus goetzei Engl.; Darbyshire, F.T.E.A.
(Gesneriaceae): 40. 2006.
Epiphytic or epilithic herb. Montane forest.
1500–1900 m.
Distribution: NL, RM—Goetze 854 (type: BR),
Gereau 4209
Streptocarpus mbeyensis I. Darbysh.; Darbyshire,
F.T.E.A. (Gesneriaceae): 39. 2006.
Terrestrial herb. Riverine forest, woodland.
1200–1350 m.
Distribution: MP—J. Lovett 4016 (type: BR, K,
MO), J. Lovett 3886
GROSSULARIACEAE
1 genus and 1 species
Choristylis rhamnoides Harv.; Verdcourt, F.T.E.A.
(Escalloniaceae): 3. 1973.
Lianescent shrub. Montane forest and remnants. 1840–2500 m.
Distribution: MP, SL—Gereau 3551
65
Distribution: KP, RM—Gereau 3158
HAMAMELIDACEAE
1 genus and 2 species
Trichocladus ellipticus Eckl. & Zeyh. subsp.
malosanus (Baker) Verdc.; Verdcourt, F.T.E.A.
(Hamamelidaceae): 2. 1971.
Shrub or tree. Riverine forest, montane forest.
980–2010 m.
Distribution: NL, RM, SL—Gereau 4197
Trichocladus goetzei Engl.; Lovett et al., 2006:
111.
Tree. Montane forest, bamboo forest. 1500–
2160 m.
Distribution: NL, RM—Kibure 1367
HIPPOCRATEACEAE
2 genera and 2 species
Elachyptera parvifolia (Oliv.) N. Hallé; Hallé &
Mathew, F.T.E.A. (Celastraceae): 49. 1994.
Liana. Riverine forest edge. 580 m.
Distribution: SL—Gereau 3819
Reissantia indica (Willd.) N. Hallé var. orientalis N. Hallé & B. Mathew; Hallé & Mathew,
F.T.E.A. (Celastraceae): 52. 1994.
Liana. Lakeshore forest. 474 m.
Distribution: SL—Gereau 3664
HYACINTHACEAE
5 genera and 5 species
Albuca abyssinica Jacq.; Stedje, F.T.E.A. (Hyacinthaceae): 22. 1996.
Terrestrial herb. Woodland, grassland, montane forest. 1250–2400 m.
Distribution: KP, MP, SL—Suleiman 60
Drimia elata Jacq.; Stedje, F.T.E.A. (Hyacinthaceae): 17. 1996.
Terrestrial herb. Miombo woodland. 1200 m.
Distribution: MP—J. Lovett 4061
GUNNERACEAE
1 genus and 1 species
Gunnera perpensa L.; Boutique & Verdcourt,
F.T.E.A. (Haloragaceae): 3. 1973.
Terrestrial herb. Stream banks in montane
grassland, bushland. 1490–2660 m.
LakeNyasa_5-18.indd 65
Drimiopsis botryoides Baker subsp. botryoides;
Stedje, F.T.E.A. (Hyacinthaceae): 9. 1996.
Terrestrial herb. Edges of cultivation. 1300 m.
Distribution: MP—J. Lovett 3556
Ledebouria revoluta (L. f.) Jessop; Stedje, F.T.E.A.
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66
(Hyacinthaceae): 15. 1996.
Terrestrial herb. Miombo woodland, edges of
cultivated fields, montane grassland. 1200–2150
m.
Distribution: MP, SL—Gereau 5141
Gereau et al.
Distribution: MP—Wiland 154
ICACINACEAE
1 genus and 1 species
Ornithogalum tenuifolium Redouté; Stedje,
F.T.E.A. (Hyacinthaceae): 27. 1996.
Terrestrial herb. Riverine forest. 1350 m.
Distribution: MP—J. Lovett 4526
Apodytes dimidiata E. Mey. ex Arn. var. dimidiata;
Lucas, F.T.E.A. (Icacinaceae): 6. 1968.
Tree or shrub. Montane forest, riverine forest,
miombo woodland. 980–2320 m.
Distribution: MR, SL—Gereau 3984
HYDROSTACHYACEAE
IRIDACEAE
1 genus and 1 species
10 genera and 23 species
Hydrostachys polymorpha Klotzsch vel sp. aff.;
Verdcourt, F.T.E.A. (Hydrostachyaceae): 3.
1986.
Aquatic herb. Rocks in fast-flowing stream.
1160 m.
Distribution: SL—Gereau 3706
Aristea abyssinica Pax; Goldblatt, F.T.E.A. (Iridaceae): 10. 1996.
Terrestrial herb. Montane grassland. 1940–
2740 m.
Distribution: NL, SL—Gereau 4760
HYPOXIDACEAE
2 genera and 5 species
Curculigo pilosa (Schumach. & Thonn.) Engl.
subsp. pilosa; Wiland-Szymanska & Nordal,
F.T.E.A. (Hypoxidaceae): 22. 2006.
Terrestrial herb. Weed in cultivated fields. 520
m.
Distribution: SL—Wiland 164
Hypoxis filiformis Baker; Wiland-Szymanska &
Nordal, F.T.E.A. (Hypoxidaceae): 8. 2006.
Terrestrial herb. Woodland, montane grassland, rocky slopes. 1720–2800 m.
Distribution: KP, MP, MR—Gereau 3745
Hypoxis fischeri Pax var. zernyi (Schulze) Wiland
& Nordal; Wiland-Szymanska & Nordal,
F.T.E.A. (Hypoxidaceae): 14. 2006.
Terrestrial herb. Woodland. 1950–2250 m.
Distribution: MP—J. Lovett 3473
Aristea nyikensis Baker; Goldblatt, F.T.E.A. (Iridaceae): 6. 1996.
Terrestrial herb. Miombo woodland, wooded
grassland. 1790–1900 m.
Distribution: MR, SL—Gereau 3892
Aristea torulosa Klatt; Weimarck, 1940: 27.
Terrestrial herb. Montane grassland. 2120–
2830 m.
Distribution: KP, NL, SL—Gereau 4311
Note: Goldblatt (pers. comm.) regards this
as the earliest name for the species treated as A.
woodii N. E. Br. in F.T.E.A. (Iridaceae): 8. 1996.
Crocosmia aurea (Pappe ex Hook.) Planch. subsp.
aurea; Goldblatt, F.T.E.A. (Iridaceae): 44.
1996.
Terrestrial herb. Montane forest. 1500–2200
m.
Distribution: MR, NL, RM, SL—Abeid 2775
Hypoxis nyasica Baker; Wiland-Szymanska &
Nordal, F.T.E.A. (Hypoxidaceae): 16. 2006.
Terrestrial herb. Woodland. 1950 m.
Distribution: MP—J. Lovett 3421
Dierama cupuliflorum Klatt subsp. caudatum
Marais; Goldblatt, F.T.E.A. (Iridaceae): 30.
1996.
Terrestrial herb. Montane grassland. 2000–
2600 m.
Distribution: KP, MR, RM—Thulin 1244 (type:
K, UPS), Gillett 17716
Hypoxis polystachya Welw. ex Baker; Wiland-Szymanska & Nordal, F.T.E.A. (Hypoxidaceae):
10. 2006.
Terrestrial herb. Miombo woodland. 1180 m.
Dierama densiflorum Marais; Goldblatt, F.T.E.A.
(Iridaceae): 29. 1996.
Terrestrial herb. Open grassland. 1800–2200
m.
LakeNyasa_5-18.indd 66
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67
Distribution: MP, MR—Bullock 3978 (type: K),
Eggeling 6123
2320 m.
Distribution: SL—Gereau 3980
Dietes iridioides (L.) Sweet ex Klatt; Goldblatt,
F.T.E.A. (Iridaceae): 11. 1996.
Terrestrial herb. Riverine forest. 980–2340 m.
Distribution: MP, RM, SL—Gereau 5170
Lapeirousia erythrantha (Klotzsch ex Klatt) Baker;
Goldblatt, F.T.E.A. (Iridaceae): 23. 1996.
Terrestrial herb. Riverine forest edge. 1850 m.
Distribution: SL—Gereau 4038
Gladiolus atropurpureus Baker; Goldblatt,
F.T.E.A. (Iridaceae): 58. 1996.
Terrestrial herb. Woodland, montane grassland. 1400–2600 m.
Distribution: KP, MP, MR—Gereau 3757
Moraea callista Goldblatt; Goldblatt, F.T.E.A. (Iridaceae): 15. 1996.
Terrestrial herb. Montane grassland. 2320–
2740 m.
Distribution: KP, SL—Richards 7562 (type:
K), Gereau 3981
Gladiolus curtifolius Marais; Goldblatt, F.T.E.A.
(Iridaceae): 60. 1996.
Terrestrial herb. Woodland, montane grassland.
Distribution: MP, SL—Goetze 1448 (type: B),
Stolz 2330
Gladiolus dalenii Van Geel subsp. andongensis
(Baker) Goldblatt; Goldblatt, F.T.E.A. (Iridaceae): 71. 1996.
Terrestrial herb. Woodland, riverine forest,
heathland, montane grassland. 1200–2770 m.
Distribution: MP, RM, SL—Gereau 7054
Gladiolus dalenii Van Geel subsp. dalenii; Goldblatt, F.T.E.A. (Iridaceae): 70. 1996.
Terrestrial herb. Montane grassland. 2400–
2580 m.
Distribution: KP, MP—P. Lovett 15
Gladiolus gregarius Welw. ex Baker; Goldblatt,
F.T.E.A. (Iridaceae): 62. 1996.
Terrestrial herb. Miombo woodland. 1200–
1530 m.
Distribution: MP, SL—Gereau 4380
Gladiolus laxiflorus Baker; Goldblatt, F.T.E.A.
(Iridaceae): 55. 1996.
Terrestrial herb. Montane grassland. 2600 m.
Distribution: KP—Leedal 4849
Gladiolus sulculatus Goldblatt; Manning & Goldblatt, 2009: 45.
Terrestrial herb. Montane grassland. 2500–
2600 m.
Distribution: KP, MR—J. Lovett 806 (type:
DSM, K, MO), J. Lovett 1870
Hesperantha cf. petitiana (A. Rich.) Baker; Goldblatt, F.T.E.A. (Iridaceae): 33. 1996.
Terrestrial herb. Rocky montane grassland.
LakeNyasa_5-18.indd 67
Moraea schimperi (Hochst.) Pic. Serm.; Goldblatt,
F.T.E.A. (Iridaceae): 18. 1996.
Terrestrial herb. Woodland, montane grassland. 1800–2700 m.
Distribution: KP, MP, NL, SL—Gereau 4080
Moraea tanzanica Goldblatt; Goldblatt, F.T.E.A.
(Iridaceae): 22. 1996.
Terrestrial herb. Montane grassland. 2320–
2700 m.
Distribution: KP, NL, RM, SL—Stolz 2412
(type: K), Suleiman 45
Moraea thomsonii Baker; Goldblatt, F.T.E.A.
(Iridaceae): 17. 1996.
Terrestrial herb. Montane grassland. 2440 m.
Distribution: KP—Leedal 5521
Moraea ventricosa Baker; Goldblatt, F.T.E.A. (Iridaceae): 21. 1996.
Terrestrial herb. Woodland. 1250–1300 m.
Distribution: MP—J. Lovett 4621
Moraea verdickii De Wild.; Goldblatt, F.T.E.A. (Iridaceae): 21. 1996.
Terrestrial herb. Montane grassland. 2600 m.
Distribution: KP—Bidgood 137
Romulea camerooniana Baker; Goldblatt, F.T.E.A.
(Iridaceae): 36. 1996.
Terrestrial herb. Montane grassland. 2320–
2740 m.
Distribution: KP, SL—Gereau 3978
Savannosiphon euryphyllus (Harms) Goldblatt
& Marais; Goldblatt, F.T.E.A. (Iridaceae): 27.
1996.
Terrestrial herb. Roadsides in coffee plantation.
1310–1400 m.
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Gereau et al.
Distribution: MP—Gereau 7006
JUNCACEAE
2 genera and 2 species
Juncus dregeanus Kunth subsp. bachitii (Hochst.
ex Steud.) Hedb.; Haines & Lye, 1983: 32.
Terrestrial herb. Stream banks in montane
grassland. 2130 m.
Distribution: MR, RM—Wingfield 282
Luzula abyssinica Parl.; Haines & Lye, 1983: 35.
Terrestrial herb. Montane grassland. 2700–
2900 m.
Distribution: KP, MR, RM—Pócs 6510/A
Clinopodium simense (Benth.) Kuntze; Ryding,
F.T.E.A. (Lamiaceae): 123. 2009.
Terrestrial herb. Thicket near waterfall in
montane grassland. 2600–2650 m.
Distribution: KP—Gereau 3190
Clinopodium uhligii (Gürke) Ryding var. obtusifolium (Avetta) Ryding; Ryding, F.T.E.A.
(Lamiaceae): 123. 2009.
Terrestrial herb. Montane grassland and forest
edges. 2000–2700 m.
Distribution: KP, MP, NL, RM—Gereau 4647
Endostemon villosus (Briq.) M. Ashby; Paton,
F.T.E.A. (Lamiaceae): 201. 2009.
Shrub. Miombo woodland. 1450 m.
Distribution: MP—J. Lovett 4919
LAMIACEAE
21 genera and 57 species
Achyrospermum carvalhoi Gürke var. carvalhoi;
Polhill, F.T.E.A. (Lamiaceae): 34. 2009.
Shrub. Montane forest, riverine forest.
890–1710 m.
Distribution: RM, SL—Mwasumbi 16287
Achyrospermum cryptanthum Baker; Polhill,
F.T.E.A. (Lamiaceae): 35. 2009.
Shrub. Montane forest. 2040–2400 m.
Distribution: MR, RM, SL—Gereau 4703
Aeollanthus subacaulis (Baker) Hua & Briq. var.
linearis (Burkill) Ryding; Ryding, F.T.E.A.
(Lamiaceae): 371. 2009.
Terrestrial herb. Montane grassland, riverine
forest, miombo woodland. 1200–2488 m.
Distribution: MP, SL—Suleiman 177
Aeollanthus subacaulis (Baker) Hua & Briq. var.
subacaulis; Ryding, F.T.E.A. (Lamiaceae):
371. 2009.
Terrestrial herb. Woodland. 1250 m.
Distribution: MP—J. Lovett 4326
Ajuga integrifolia Buch.-Ham. ex D. Don; Lukhoba, F.T.E.A. (Lamiaceae): 15. 2009.
Terrestrial herb. Montane grassland. 2570 m.
Distribution: KP—Gereau 3106
Clinopodium myrianthum (Baker) Ryding; Ryding, F.T.E.A. (Lamiaceae): 120. 2009.
Shrub. Montane forest edges. 2300 m.
Distribution: MR—Gereau 4591
LakeNyasa_5-18.indd 68
Haumaniastrum caeruleum (Oliv.) P. A. Duvign.
& Plancke; Paton, F.T.E.A. (Lamiaceae): 213.
2009.
Terrestrial herb. Woodland. 1200 m.
Distribution: MP—Kayombo 2427
Haumaniastrum uniflorum A. J. Paton; Paton,
F.T.E.A. (Lamiaceae): 213. 2009.
Terrestrial herb. Grassland.
Distribution: MR, NL—Davies 818
Haumaniastrum villosum (Benth.) A. J. Paton;
Paton, F.T.E.A. (Lamiaceae): 220. 2009.
Terrestrial herb. Miombo woodland. 1400 m.
Distribution: MP—P. Lovett 101
Hoslundia opposita Vahl; Paton, F.T.E.A. (Lamiaceae): 193. 2009.
Shrub. Riverine forest, miombo woodland.
1160–1500 m.
Distribution: MP, RM, SL—Gereau 3938
Leonotis decadonta Gürke var. decadonta; Iwarsson & Harvey, F.T.E.A. (Lamiaceae): 100.
2009.
Shrub.
Distribution: SL—Iwarsson 1216
Leonotis decadonta Gürke var. porotoensis Iwarsson & Y. B. Harv.; Iwarsson & Harvey, F.T.E.A.
(Lamiaceae): 100. 2009.
Shrub. Montane grassland and forest margins.
2300–2650 m.
Distribution: MR, NL—Iwarsson 1160 (type:
BR, DSM, EA, K, NHT, S, UPS), Richards 9727
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Floristic Checklist
Leonotis goetzei Gürke; Iwarsson & Harvey,
F.T.E.A. (Lamiaceae): 102. 2009.
Shrub. Montane grassland and shrubland,
roadsides. 1500–2800 m.
Distribution: MR, NL—Iwarsson 1193
Leonotis grandis Iwarsson & Y. B. Harv.; Iwarsson
& Harvey, F.T.E.A. (Lamiaceae): 97. 2009.
Shrub. Riverine forest. 1500 m.
Distribution: MP, NL—J. Lovett 4695
Leonotis myricifolia Iwarsson & Y. B. Harv.; Iwarsson & Harvey, F.T.E.A. (Lamiaceae): 101.
2009.
Shrub.
Distribution: KP, MP—Myembe 40
Leonotis ocymifolia (Burm. f.) Iwarsson var.
raineriana (Vis.) Iwarsson; Iwarsson & Harvey,
F.T.E.A. (Lamiaceae): 103. 2009.
Shrub or perennial herb. Montane forest edges,
secondary bushland, woodland. 1300–2640 m.
Distribution: MP, MR, NL—Gereau 4643
Leucas deflexa Hook f. vel sp. aff.; Bramley,
F.T.E.A. (Lamiaceae): 73. 2009.
Terrestrial herb. Abandoned cultivation. 1250
m.
Distribution: MP—J. Lovett 4191
Leucas tettensis Vatke; Bramley, F.T.E.A. (Lamiaceae): 91. 2009.
Terrestrial herb. Riverine forest. 1250–1350 m.
Distribution: MP—J. Lovett 4662
Micromeria imbricata (Forssk.) C. Chr. var.
imbricata; Ryding, F.T.E.A. (Lamiaceae): 117.
2009.
Terrestrial herb or subshrub. Montane grassland. 1940–2400 m.
Distribution: MP, MR, SL—Gereau 4680
Ocimum decumbens Gürke; Paton, F.T.E.A.
(Lamiaceae): 170. 2009.
Terrestrial herb. Montane grassland. 2230 m.
Distribution: SL—Gereau 3589
Ocimum fimbriatum Briq.; Paton, F.T.E.A. (Lamiaceae): 156. 2009.
Terrestrial herb. Woodland, rocky grassland.
1250–1810 m.
Distribution: MP, SL—Harder 1497
Ocimum gratissimum L. subsp. gratissimum;
LakeNyasa_5-18.indd 69
69
Paton, F.T.E.A. (Lamiaceae): 152. 2009.
Terrestrial herb. Woodland, riverine forest,
disturbed montane forest. 1100–1790 m.
Distribution: MP, RM—Abeid 2915
Ocimum obovatum E. Mey. ex Benth. subsp. cordatum (A. J. Paton) A. J. Paton; Paton, F.T.E.A.
(Lamiaceae): 169. 2009.
Terrestrial herb. Montane grassland, forest
edges. 2650 m.
Distribution: MP—Goyder 3847
Orthosiphon schimperi Benth.; Paton, F.T.E.A.
(Lamiaceae): 182. 2009.
Terrestrial herb. Woodland, riverine thickets.
1200–2000 m.
Distribution: MP, SL—Gereau 4065
Platostoma rotundifolium (Briq.) A. J. Paton;
Paton, F.T.E.A. (Lamiaceae): 231. 2009.
Shrub or perennial herb. Montane grassland,
woodland, forest edges, roadsides in cultivation.
1475–2240 m.
Distribution: MP, MR, NL, RM, SL—Kayombo
2403
Plectranthus adenophorus Gürke; Paton, F.T.E.A.
(Lamiaceae): 280. 2009.
Terrestrial herb. Montane grassland, mountain
ridges. 2100–2850 m.
Distribution: KP, MP, NL—P. Lovett 12
Plectranthus alboviolaceus Gürke; Paton, F.T.E.A.
(Lamiaceae): 246. 2009.
Shrub. Montane forest. 2150 m.
Distribution: MR—Gereau 4618
Plectranthus autranii (Briq.) A. J. Paton; Paton,
F.T.E.A. (Lamiaceae): 326. 2009.
Shrub. Montane forest.
Distribution: MR—Goyder 3861
Plectranthus daviesii (E. A. Bruce) B. Mathew;
Paton, F.T.E.A. (Lamiaceae): 312. 2009.
Terrestrial herb.
Distribution: RM—Davies R4 (type: K, SRGH),
Mgaza 468
Plectranthus defoliatus Hochst ex Benth.; Paton,
F.T.E.A. (Lamiaceae): 297. 2009.
Terrestrial herb. Montane grassland. 2300 m.
Distribution: MP, MR—Iwarsson 1083
Plectranthus densus N. E. Br.; Paton, F.T.E.A.
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Gereau et al.
(Lamiaceae): 292. 2009.
Terrestrial herb. Montane grassland. 2150 m.
Distribution: SL—Gereau 5138
Plectranthus djalonensis (A. Chev.) A. J. Paton;
Paton, F.T.E.A. (Lamiaceae): 286. 2009.
Terrestrial herb. Montane forest, miombo
woodland. 1300–1660 m.
Distribution: MP, RM—Kayombo 1008
Plectranthus gracilis Suess.; Paton, F.T.E.A.
(Lamiaceae): 269. 2009.
Terrestrial herb.
Distribution: RM—Abdullah 1349
Plectranthus ignotus A. J. Paton; Paton, F.T.E.A.
(Lamiaceae): 278. 2009.
Terrestrial herb. Mountain slopes. 2080–2140
m.
Distribution: MP, SL—Suleiman 87
Plectranthus laxiflorus Benth.; Paton, F.T.E.A.
(Lamiaceae): 250. 2009.
Terrestrial herb. Montane forest. 2280–2400
m.
Distribution: RM—Mwasumbi 16217
Plectranthus megacalyx A. J. Paton; Paton,
F.T.E.A. (Lamiaceae): 316. 2009.
Terrestrial herb. Montane grassland.
Distribution: MR—Geilinger 2884 (type: K),
Leedal 3835
Plectranthus parvicalyx A. J. Paton; Paton,
F.T.E.A. (Lamiaceae): 316. 2009.
Terrestrial herb. Montane grassland. 2400–
2600 m.
Distribution: KP, MP—J. Lovett 802
Plectranthus pubescens Baker; Paton, F.T.E.A.
(Lamiaceae): 272. 2009.
Terrestrial herb. Woodland. 1300–1600 m.
Distribution: MP, SL—J. Lovett 4285
Plectranthus punctatus (L. f.) L’Hér. subsp. edulis
(Vatke) J. Paton; Paton, F.T.E.A. (Lamiaceae):
350. 2009.
Terrestrial herb. Montane grassland. 2360 m.
Distribution: MR, RM—Gereau 3755
Plectranthus punctatus (L. f.) L’Hér. subsp. punctatus; Paton, F.T.E.A. (Lamiaceae): 349. 2009.
Terrestrial herb.
LakeNyasa_5-18.indd 70
Distribution: MR—Goyder 3860
Plectranthus rungwensis A. J. Paton; Paton,
F.T.E.A. (Lamiaceae): 252. 2009.
Shrub. Montane forest. 2100–2350 m.
Distribution: MR, RM—Mwasumbi 16222
(type: DSM, K, MO), Richards 6514
Plectranthus schizophyllus Baker; Paton, F.T.E.A.
(Lamiaceae): 333. 2009.
Terrestrial herb. Montane grassland, miombo
woodland. 1500–2700 m.
Distribution: KP, NL, SL—Gereau 4418
Plectranthus stolzii Gilli; Paton, F.T.E.A. (Lamiaceae): 254. 2009.
Terrestrial herb. Montane forest. 2040 m.
Distribution: MP, SL—Gereau 4731
Plectranthus viphyensis Brummitt & Seyani subsp.
viphyensis; Paton, F.T.E.A. (Lamiaceae): 276.
2009.
Terrestrial herb. Woodland, montane grassland. 1400–1810 m.
Distribution: MP, SL—Harder 1496
Pycnostachys meyeri Gürke; Bramley, F.T.E.A.
(Lamiaceae): 412. 2009.
Shrub or woody-based herb. Montane forest
and brushland. 1610–1700 m.
Distribution: RM—Mwasumbi 16338
Pycnostachys ruandensis De Wild.; Bramley,
F.T.E.A. (Lamiaceae): 389. 2009.
Terrestrial herb. Cultivated fields. 2280 m.
Distribution: MR—Mwasumbi 16161
Pycnostachys schliebenii Mildbr.; Bramley,
F.T.E.A. (Lamiaceae): 410. 2009.
Shrub. Montane forest. 2000 m.
Distribution: RM—Mwasumbi 16232
Pycnostachys sphaerocephala Baker; Bramley,
F.T.E.A. (Lamiaceae): 402. 2009.
Shrub. Woodland. 1400–1500 m.
Distribution: MP—J. Lovett 3413
Pycnostachys aff. urticifolia Hook.; Bramley,
F.T.E.A. (Lamiaceae): 411. 2009.
Shrub. Montane forest edges. 2200 m.
Distribution: MR—Gereau 4537
Salvia nilotica Juss. ex Jacq.; Polhill, F.T.E.A.
(Lamiaceae): 110. 2009.
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Lake Nyasa Climatic Region
Floristic Checklist
Terrestrial herb. Montane grassland. 2640 m.
Distribution: KP, MP, NL—Gereau 4638
Scutellaria schweinfurthii Briq. subsp. paucifolia
(Baker) A. J. Paton; Paton, F.T.E.A. (Lamiaceae): 28. 2009.
Terrestrial herb. Woodland. 1400–2250 m.
Distribution: MP—J. Lovett 4862
Stachys aculeolata Hook. f.; Harvey, F.T.E.A.
(Lamiaceae): 42. 2009.
Herbaceous climber. Montane forest edges.
2200–2300 m.
Distribution: MR, SL—Gereau 3532
Stachys glandulibracteata Y. B. Harv.; Harvey,
F.T.E.A. (Lamiaceae): 45. 2009.
Terrestrial herb. Streamsides in montane grassland. 1950–2100 m.
Distribution: MP, MR, RM—Richards 17016
(type: EA, K), Napper 1171
71
Distribution: SL—Gereau 3639
Cassytha pondoensis Engl.; Verdcourt, F.T.E.A.
(Lauraceae): 17. 1996.
Parasitic herb. Secondary shrubland. 1760 m.
Distribution: NL—Gereau 4245
Ocotea usambarensis Engl.; Verdcourt, F.T.E.A.
(Lauraceae): 10. 1996.
Tree. Montane forest. 1620–2050 m.
Distribution: NL, RM—Abeid 2822
LEEACEAE
1 genus and 1 species
Leea guineensis G. Don; Verdcourt, F.T.E.A. (Vitaceae [Leeaceae]): 141. 1993.
Terrestrial herb. Riverine forest. 890 m.
Distribution: SL—Kibure 1469
LENTIBULARIACEAE
Stachys pseudonigricans Gürke; Harvey, F.T.E.A.
(Lamiaceae): 48. 2009.
Terrestrial herb. Woodland. 1300–2000 m.
Distribution: MP, SL—Gereau 4064
Tetradenia tanganyikae Phillipson; Phillipson,
F.T.E.A. (Lamiaceae): 359. 2009.
Shrub. Riverine forest, woodland. 1150–1500
m.
Distribution: MP—Kayombo 2441
1 genus and 1 species
Utricularia welwitschii Oliv.; Taylor, F.T.E.A.
(Lentibulariaceae): 10. 1973.
Terrestrial herb. Marshy montane grassland.
2550 m.
Distribution: KP—Abeid 3118
LILIACEAE
1 genus and 1 species
Teucrium goetzei Gürke; Paton, F.T.E.A. (Lamiaceae): 14. 2009.
Terrestrial herb. Montane grassland. 1700–
1940 m.
Distribution: SL—Goetze 1188a (type: BM),
Gereau 4652
Lilium formosanum Wallace; Grimshaw, F.T.E.A.
(Liliaceae): 3. 2005.
Herb. Montane forest. 2100 m.
Distribution: MR—Mwasumbi 16308
Tinnea aethiopica Kotschy ex Hook. f. subsp.
stolzii (Robyns & Lebrun) Vollesen; Willis,
F.T.E.A. (Lamiaceae): 20. 2009.
Shrub. Woodland, riverine forest. 1200–1910
m.
Distribution: MP, SL—Gereau 4071
1 genus and 3 species
LAURACEAE
2 genera and 3 species
Cassytha filiformis L.; Verdcourt, F.T.E.A. (Lauraceae): 15. 1996.
Parasitic herb. Lakeshore forest. 474 m.
LakeNyasa_5-18.indd 71
LINACEAE
Linum keniense T. C. E. Fr.; Smith, F.T.E.A. (Linaceae): 10. 1966.
Terrestrial herb. Montane grassland.
Distribution: RM—Procter 1449
Linum thunbergii Eckl. & Zeyh.; Smith, F.T.E.A.
(Linaceae): 10. 1966.
Terrestrial herb. Montane grassland. 2570–
2700 m.
Distribution: KP, MP, NL—Gereau 4087
Linum volkensii Engl.; Smith, F.T.E.A. (Linaceae):
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72
Gereau et al.
8. 1966.
Terrestrial herb. Montane grassland. 1940 m.
Distribution: SL—Gereau 4662
LOGANIACEAE
2 genera and 4 species
Anthocleista grandiflora Gilg; Leeuwenberg, Fl.
Zamb. 7(1): 329. 1983.
Tree. Montane forest. 1610–1680 m.
Distribution: RM—Kibure 1398
Strychnos innocua Delile; Leeuwenberg, Fl. Zamb.
7(1): 361. 1983.
Tree. Miombo woodland. 1200 m.
Distribution: MP—J. Lovett 3883
Strychnos mellodora S. Moore vel sp. aff.; Leeuwenberg, Fl. Zamb. 7(1): 366. 1983.
Tree. Montane forest. 1920 m.
Distribution: NL—Bracebridge 59
Strychnos spinosa Lam.; Leeuwenberg, Fl. Zamb.
7(1): 371. 1983.
Tree. Riverine and lakeshore thickets, woodland, abandoned cultivation. 474–1250 m.
Distribution: MP, SL—Mwasumbi 16602
LORANTHACEAE
7 genera and 14 species
Agelanthus bipartitus Balle ex Polhill & Wiens;
Polhill & Wiens, F.T.E.A. (Loranthaceae): 59.
1999.
Parasitic shrub. Montane forest.
Distribution: RM, SL—Stolz 2475 (type: BM,
BR, FHO, K, MO, P), Richards 17622
Agelanthus fuellebornii (Engl.) Polhill & Wiens;
Polhill & Wiens, F.T.E.A. (Loranthaceae): 61.
1999.
Parasitic shrub. Woodland, sandy lakeshores.
478–1700 m.
Distribution: MP, SL—Gereau 6084
Agelanthus nyasicus (Baker & Sprague) Polhill &
Wiens; Polhill & Wiens, F.T.E.A. (Loranthaceae): 62. 1999.
Parasitic shrub. Riverine forest, coffee plantations. 1300–1390 m.
Distribution: MP—Gereau 7039
Agelanthus subulatus (Engl.) Polhill & Wiens;
LakeNyasa_5-18.indd 72
Polhill & Wiens, F.T.E.A. (Loranthaceae): 46.
1999.
Parasitic shrub. Montane forest. 1900 m.
Distribution: NL—Gereau 4183
Agelanthus zizyphifolius (Engl.) Polhill & Wiens
subsp. vittatus (Engl.) Polhill & Wiens; Polhill
& Wiens, F.T.E.A. (Loranthaceae): 51. 1999.
Parasitic shrub. Montane forest edges. 1980 m.
Distribution: SL—Gereau 3623
Englerina cordata Polhill & Wiens; Polhill &
Wiens, F.T.E.A. (Loranthaceae): 32. 1999.
Parasitic shrub. Woodland. 2110 m.
Distribution: SL—Gereau 4018
Englerina inaequilatera (Engl.) Gilli; Polhill &
Wiens, F.T.E.A. (Loranthaceae): 31. 1999.
Parasitic shrub. Montane forest, forest edges.
1850–2640 m.
Distribution: KP, MP, MR, NL, RM, SL—
Gereau 4644
Erianthemum taborense (Engl.) Tiegh.; Polhill &
Wiens, F.T.E.A. (Loranthaceae): 98. 1999.
Parasitic shrub. Woodland. 1200–1475 m.
Distribution: MP—Kayombo 2437
Globimetula anguliflora (Engl.) Danser; Polhill &
Wiens, F.T.E.A. (Loranthaceae): 79. 1999.
Parasitic shrub. Miombo woodland. 1500 m.
Distribution: MP—Gereau 6985
Oedina pendens (Engl. & K. Krause) Polhill &
Wiens; Polhill & Wiens, F.T.E.A. (Loranthaceae): 86. 1999.
Parasitic shrub. Montane forest. 1910 m.
Distribution: NL, RM—Stolz 1103 (type: B, K,
WAG), Gereau 4213
Phragmanthera cornetii (Dewèvre) Polhill &
Wiens; Polhill & Wiens, F.T.E.A. (Loranthaceae): 108. 1999.
Parasitic shrub. Woodland. 1400 m.
Distribution: MP—J. Lovett 3340
Phragmanthera dschallensis (Engl.) M. G. Gilbert;
Polhill & Wiens, F.T.E.A. (Loranthaceae): 110.
1999.
Parasitic shrub. Woodland. 1200–1600 m.
Distribution: MP—Kayombo 2396
Phragmanthera usuiensis (Oliv.) M. G. Gilbert;
Polhill & Wiens, F.T.E.A. (Loranthaceae): 105.
5/31/12 3:59 PM
Lake Nyasa Climatic Region
Floristic Checklist
73
1999.
Parasitic shrub. Miombo woodland, montane
forest. 1780–1890 m.
Distribution: RM, SL—Gereau 5107
Steyaert) Verdc.; Verdcourt, F.T.E.A. (Malvaceae): 158. 2009.
Shrub. Montane forest. 2080 m.
Distribution: MR—Mwasumbi 16188
Vanwykia rubella Polhill & Wiens; Polhill & Wiens,
F.T.E.A. (Loranthaceae): 85. 1999.
Parasitic shrub. Woodland. 1300–1400 m.
Distribution: MP—Kayombo 1041
Thespesia garckeana F. Hoffm.; Verdcourt,
F.T.E.A. (Malvaceae): 104. 2009.
Tree or shrub. Riverine forest, woodland.
1100–1410 m.
Distribution: MP, SL—Kayombo 960
MALVACEAE
5 genera and 9 species
MELASTOMATACEAE
2 genera and 7 species
Hibiscus debeerstii De Wild. & T. Durand; Mwachala, F.T.E.A. (Malvaceae): 53. 2009.
Shrub. Riverine forest and edges of miombo
woodland. 1200–1500 m.
Distribution: MP, SL—Kayombo 2366
Hibiscus fuscus Garcke; Mwachala, F.T.E.A. (Malvaceae): 49. 2009.
Shrub. Montane forest. 1870 m.
Distribution: RM—WCS SHCP 231
Hibiscus ludwigii Eckl. & Zeyh.; Mwachala,
F.T.E.A. (Malvaceae): 65. 2009.
Scrambling shrub. Montane forest, woodland.
1400–2100 m.
Distribution: MP, MR—Mwasumbi 16318
Hibiscus aff. micranthus L. f.; Mwachala, F.T.E.A.
(Malvaceae): 47.
Shrub or perennial herb. Montane forest, Eucalyptus plantations, riverine forest. 1200–2100 m.
Distribution: MP, MR, RM—Mwasumbi 16337
Malvaviscus penduliflorus DC.; Turner & Mendenhall, 1993: 449.
Cultivated shrub. Garden in village. 1250 m.
Distribution: MP—Kayombo 2337
Pavonia kilimandscharica Gürke; Verdcourt,
F.T.E.A. (Malvaceae): 14. 2009.
Shrub. Montane forest. 2080 m.
Distribution: MR, RM—Mwasumbi 16187
Pavonia urens Cav.; Verdcourt, F.T.E.A. (Malvaceae): 11. 2009.
Shrub. Montane forest, forest edges, riverine
forest. 1400–2360 m.
Distribution: MP, MR, RM, SL—Gereau 4707
Sida rhombifolia L. var. serratifolia (R. Wilczek &
LakeNyasa_5-18.indd 73
Antherotoma naudinii Hook. f.; Wickens, F.T.E.A.
(Melastomataceae): 9. 1975.
Terrestrial herb. Woodland, grassland, riverine
forest, abandoned cultivation. 1250–1940 m.
Distribution: MP, SL—Gereau 4668
Dissotis cryptantha Baker; Wickens, F.T.E.A.
(Melastomataceae): 56. 1975.
Shrub.
Distribution: RM—Goetze 1138
Dissotis melleri Hook. f. var. greenwayi (A. Fern. &
R. Fern.) & R. Fern.; Wickens, F.T.E.A. (Melastomataceae): 46. 1975.
Shrub. Montane forest edges, bamboo thicket.
1810–2350 m.
Distribution: MR, NL, RM, SL—Gereau 3971
Dissotis pachytricha Gilg ex R. E. Fr. var. orientalis A. Fern. & R. Fern.; Wickens, F.T.E.A.
(Melastomataceae): 54. 1975.
Shrub. Thickets in montane forest. 1840 m.
Distribution: SL—Gereau 3541
Dissotis princeps (Kunth) Triana var. princeps;
Wickens, F.T.E.A. (Melastomataceae): 51.
1975.
Shrub. Open woodland. 1650–1680 m.
Distribution: SL—Gereau 5117
Dissotis senegambiensis (Guill. & Perr.) Triana var.
alpestris (Taub.) A. Fern. & R. Fern.; Wickens,
F.T.E.A. (Melastomataceae): 50. 1975.
Terrestrial herb. Montane grassland. 1940 m.
Distribution: SL—Gereau 4684
Dissotis trothae Gilg; Wickens, F.T.E.A. (Melastomataceae): 55. 1975.
Shrub. Stream banks in montane grassland.
2080–2140 m.
5/31/12 3:59 PM
74
Gereau et al.
Distribution: SL—Suleiman 119
edges. 1500–1660 m.
Distribution: RM—Mwasumbi 16326
MELIACEAE
4 genera and 9 species
Lepidotrichilia volkensii (Gürke) J.-F. Leroy ex
Styles & F. White; Styles & White, F.T.E.A.
(Meliaceae): 37. 1991.
Tree or shrub. Riverine forest, montane forest.
1730–2400 m.
Distribution: MP, MR, NL, RM—Abeid 2900
Toona ciliata M. Roem.; Styles & White, F.T.E.A.
(Meliaceae): 45. 1991.
Tree. Lakeshore forest and thickets; naturalized. 474–490 m.
Distribution: SL—Mwasumbi 16559
Trichilia dregeana Sond.; Styles & White, F.T.E.A.
(Meliaceae): 34. 1991.
Tree. Riverine forest. 1350 m.
Distribution: MP—J. Lovett 4757
Trichilia emetica Vahl; Styles & White, F.T.E.A.
(Meliaceae): 34. 1991.
Tree. Lakeshore forest, riverine forest, coffee
plantations. 474–1300 m.
Distribution: MP, SL—Gereau 3671
Trichilia aff. prieuriana A. Juss.; Styles & White,
F.T.E.A. (Meliaceae): 30. 1991.
Tree. Riverine forest. 1275–1400 m.
Distribution: MP—Kayombo 2378
Turraea floribunda Hochst.; Styles & White,
F.T.E.A. (Meliaceae): 21. 1991.
Tree. Montane forest.
Distribution: RM—Ndangalasi 977
Turraea holstii Gürke; Styles & White, F.T.E.A.
(Meliaceae): 13. 1991.
Shrub. Woodland, riverine forest. 1250–1600
m.
Distribution: MP—Kayombo 2352
MELIANTHACEAE
1 genus and 1 species
Bersama abyssinica Fresen. subsp. abyssinica;
Verdcourt, F.T.E.A. (Melianthaceae): 4. 1958.
Tree. Riverine forest, montane forest, scree
slopes. 1650–2260 m.
Distribution: MP, MR, SL—Suleiman 138
Bersama abyssinica Fresen. subsp. paulliniodes
(Planch.) Verdc.; Verdcourt, F.T.E.A. (Melianthaceae): 5. 1958.
Tree. Riverine forest, montane forest. 1770–
2040 m.
Distribution: NL, SL—Gereau 4033
MENISPERMACEAE
3 genera and 3 species
Stephania abyssinica (Quart.-Dill. & A. Rich.)
Walp. var. abyssinica; Troupin, F.T.E.A. (Menispermaceae): 22. 1956.
Herbaceous climber. Tree clumps in montane
grassland. 2360 m.
Distribution: MR, RM—Gereau 3718
Tiliacora funifera (Miers) Oliv.; Troupin, F.T.E.A.
(Menispermaceae): 9. 1956.
Liana. Disturbed montane forest. 1790–1910
m
Distribution: RM—Abeid 2910
Tinospora caffra (Miers) Troupin; Troupin,
F.T.E.A. (Menispermaceae): 19. 1956.
Herbaceous climber. Lakeshore forest. 474 m.
Distribution: SL—Gereau 3670
MIMOSACEAE
7 genera and 18 species
Turraea robusta Gürke; Styles & White, F.T.E.A.
(Meliaceae): 8. 1991.
Tree. Riverine forest. 1500–1780 m.
Distribution: RM, SL—Kibure 1427
Acacia amythethophylla Steud. ex A. Rich.; Asfaw
& Thulin, 1989: 86.
Tree. Woodland, abandoned cultivation. 1100–
1530 m.
Distribution: MP, SL—Kayombo 2385
Turraea stolzii Harms; Styles & White, F.T.E.A.
(Meliaceae): 13. 1991.
Shrub. Riverine brushland, montane forest
Acacia erubescens Welw. ex Oliv.; Brenan, F.T.E.A.
(Leguminosae–Mimosoideae): 88. 1959.
Shrub or tree. Woodland. 1200 m.
LakeNyasa_5-18.indd 74
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Lake Nyasa Climatic Region
Floristic Checklist
Distribution: MP—J. Lovett 3857
Acacia polyacantha Willd. subsp. campylacantha
(Hochst. ex A. Rich.) Brenan; Brenan, F.T.E.A.
(Leguminosae–Mimosoideae): 88. 1959.
Tree. Woodland, riverine forest, lakeshore
forest, cultivated fields. 480–1400 m.
Distribution: MP, SL—Gereau 3660
Acacia pseudofistula Harms vel sp. aff.; Brenan,
F.T.E.A. (Leguminosae–Mimosoideae): 122.
1959.
Shrub. Woodland. 1200–1300 m.
Distribution: MP—P. Lovett 379
Acacia schweinfurthii Brenan & Exell var.
schweinfurthii; Brenan, F.T.E.A. (Leguminosae–Mimosoideae): 99. 1959.
Liana. Riverine forest. 1350 m.
Distribution: MP—J. Lovett 3604
Acacia sieberiana DC. var. woodii (Burtt Davy)
Keay & Brenan; Asfaw & Thulin, 1989: 90.
Tree. Woodland, riverine forest. 1250–1600 m.
Distribution: MP—Gereau 6092
Albizia amara (Roxb.) Boivin; Brenan, F.T.E.A.
(Leguminosae–Mimosoideae): 151. 1959.
Tree. Montane forest. 1510 m.
Distribution: RM—WCS SHCP 320
Albizia antunesiana Harms; Brenan, F.T.E.A.
(Leguminosae–Mimosoideae): 148. 1959.
Tree. Woodland. 1280–1400 m.
Distribution: MP—Kayombo 2367
Albizia gummifera (J. F. Gmel.) C. A. Sm. var.
gummifera; Brenan, F.T.E.A. (Leguminosae–
Mimosoideae): 157. 1959.
Tree. Riverine forest. 1300–1500 m.
Distribution: MP, RM—J. Lovett 4788
Albizia harveyi E. Fourn.; Brenan, F.T.E.A. (Leguminosae–Mimosoideae): 149. 1959.
Tree. Woodland. 1250 m.
Distribution: MP—J. Lovett 4368
Albizia lebbeck (L.) Benth.; Brenen, F.T.E.A.
(Leguminosae–Mimosoideae): 147. 1959.
Tree.
Distribution: NL—Davies 645
Albizia schimperiana Oliv. var. schimperiana; Brenan, F.T.E.A. (Leguminosae–Mimosoideae):
LakeNyasa_5-18.indd 75
75
154. 1959.
Tree. Riverine forest. 1400 m.
Distribution: MP, NL—J. Lovett 4686
Albizia versicolor Welw. ex Oliv.; Brenan, F.T.E.A.
(Leguminosae–Mimosoideae): 146. 1959.
Tree. Woodland. 1300–1400 m.
Distribution: MP—Gereau 6100
Dichrostachys cinerea (L.) Wight & Arn. subsp.
nyassana (Taub.) Brenan; Brenan & Brummitt,
Fl. Zamb. 3(1): 40. 1970.
Tree or shrub. Woodland. 1200–1310 m.
Distribution: MP—Gereau 6103
Entada abyssinica Steud. ex A. Rich.; Brenan,
F.T.E.A. (Leguminosae–Mimosoideae): 13.
1959.
Tree. Roadsides, thickets, miombo woodland,
cultivation. 1250–1475 m.
Distribution: MP—Kayombo 2384
Faidherbia albida (Delile) A. Chev.; Barnes &
Fagg, 2003: 3.
Tree. Coffee plantations. 1250–1400 m.
Distribution: MP—Kayombo 957
Mimosa pigra L.; Brenan, F.T.E.A. (Leguminosae–
Mimosoideae): 43. 1959.
Lianescent shrub. Lakeshore forest. 480 m.
Distribution: SL—Gereau 3658
Parkia filicoidea Welw. ex Oliv.; Brenan, F.T.E.A.
(Leguminosae–Mimosoideae): 7. 1959.
Tree. Riverine forest. 1160 m.
Distribution: SL—Gereau 3703
MOLLUGINACEAE
1 genus and 1 species
Glinus lotoides L. var. virens Fenzl; Gonçalves, Fl.
Zamb. 4: 538. 1978.
Terrestrial herb. Woodland. 1250 m.
Distribution: MP—J. Lovett 3775
MONIMIACEAE
1 genus and 1 species
Xymalos monospora (Harv.) Baill. ex Warb.; Verdcourt, F.T.E.A. (Monimiaceae): 1. 1968.
Shrub or tree. Montane forest, riverine forest.
1780–2300 m.
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76
Distribution: MP, MR, NL, RM, SL—Gereau
4745
MORACEAE
3 genera and 13 species
Dorstenia benguellensis Welw.; Hijman, F.T.E.A.
(Moraceae): 38. 1989.
Terrestrial herb. Miombo woodland, riverine
forest edges. 1300–1850 m.
Distribution: MP, SL—Gereau 3901
Dorstenia cuspidata Hochst. ex A. Rich.; Hijman,
F.T.E.A. (Moraceae): 40. 1989.
Terrestrial herb. Miombo woodland, riverine
forest. 1160–1530 m.
Distribution: SL—Gereau 4372
Dorstenia psilurus Welw.; Hijman, F.T.E.A. (Moraceae): 31. 1989.
Terrestrial herb. Riverine forest. 1320 m.
Distribution: MP—Gereau 6121
Dorstenia schliebenii Mildbr.; Hijman, F.T.E.A.
(Moraceae): 28. 1989.
Terrestrial herb. Montane forest. 1730 m.
Distribution: NL—Gereau 4147
Ficus exasperata Vahl; Berg, F.T.E.A. (Moraceae):
52. 1989.
Tree. Riverine forest, woodland, coffee plantations, abandoned cultivation. 1250–1400 m.
Distribution: MP—Gereau 6098
Gereau et al.
Tree. Riverine forest. 1160–1850 m.
Distribution: MP, SL—Gereau 3694
Ficus sycomorus L.; Berg, F.T.E.A. (Moraceae): 54.
1989.
Tree. Woodland, riverine forest, coffee plantations. 1250–1370 m.
Distribution: MP—Gereau 6106
Ficus thonningii Blume; Berg, F.T.E.A. (Moraceae):
73. 1989.
Tree. Riverine forest. 1500 m.
Distribution: SL—Gereau 4795
Ficus vallis-choudae Delile; Berg, F.T.E.A. (Moraceae): 58. 1989.
Tree. Riverine forest. 580–1150 m.
Distribution: MP, SL—Kibure 1448
Trilepisium madagascariense DC.; Berg, F.T.E.A.
(Moraceae): 17. 1989.
Tree. Riverine forest. 890–1680 m.
Distribution: MP, RM, SL—Kayombo 1097
MUSACEAE
1 genus and 1 species
Ensete ventricosum (Welw.) Cheesman; Lock,
F.T.E.A. (Musaceae): 3. 1993.
Terrestrial herb. Woodland. 1750–1950 m.
Distribution: MP, RM—Lovett 3431
MYRICACEAE
Ficus ingens (Miq.) Miq.; Berg, F.T.E.A. (Moraceae): 60. 1989.
Tree. Lakeshore bushland. 474–480 m.
Distribution: SL—Kibure 1490
Ficus natalensis Hochst.; Berg, F.T.E.A. (Moraceae): 71. 1989.
Tree. Riverine forest. 1290 m.
Distribution: MP—Gereau 6109
Ficus ottoniifolia (Miq.) Miq. subsp. ulugurensis
(Warb. ex Mildbr. & Burret) C. C. Berg; Berg,
F.T.E.A. (Moraceae): 77. 1989.
Tree. Riverine forest, montane forest. 1250–
1840 m.
Distribution: MP, RM—Kayombo 2480
Ficus sur Forssk.; Berg, F.T.E.A. (Moraceae): 56.
1989.
LakeNyasa_5-18.indd 76
1 genus and 2 species
Morella salicifolia (Hochst. ex A. Rich.) Verdc. &
Polhill subsp. kilimandscharica (Engl.) Verdc.
& Polhill; Polhill & Verdcourt, F.T.E.A. (Myricaceae): 9. 2000.
Tree. Montane forest, tree clumps in grassland.
1670–2380 m.
Distribution: KP, MP, MR, RM, SL—Gereau
3736
Morella serrata (Lam.) Killick; Polhill & Verdcourt,
F.T.E.A. (Myricaceae): 4. 2000.
Tree. Riverine forest. 1500 m.
Distribution: SL—Gereau 4782
MYROTHAMNACEAE
1 genus and 1 species
5/31/12 3:59 PM
Lake Nyasa Climatic Region
Floristic Checklist
Myrothamnus flabellifolius Welw.; Bywater,
F.T.E.A. (Myrothamnaceae): 1. 1984.
Shrub. Rocky mountain slopes. 2320 m.
Distribution: SL—Gereau 3993
MYRSINACEAE
3 genera and 4 species
Embelia schimperi Vatke; Halliday, F.T.E.A.
(Myrsinaceae): 13. 1984.
Liana. Montane forest, riverine forest. 1500–
2430 m.
Distribution: MR, RM, SL—Abeid 2814
Maesa lanceolata Forssk.; Halliday, F.T.E.A.
(Myrsinaceae): 3. 1984.
Shrub or tree. Montane forest, riverine forest,
miombo woodland, bamboo thicket. 1500–2430
m.
Distribution: KP, MP, MR, NL, SL—Suleiman
133
Myrsine africana L.; Halliday, F.T.E.A. (Myrsinaceae): 6. 1984.
Shrub. Montane heathland and shrubland.
2620–2820 m.
Distribution: KP, RM—Mwasumbi 16253
Myrsine melanophloeos (L.) R. Br.; Lovett et al.,
2006: 188.
Shrub or tree. Montane forest edges, miombo
woodland. 620–2700 m.
Distribution: MR, NL, RM, SL—Gereau 5122
MYRTACEAE
1 genus and 3 species
Syzygium cordatum Hochst. subsp. cordatum;
Verdcourt, F.T.E.A. (Myrtaceae): 74. 2001.
Tree or shrub. Riverine forest, secondary shrubland. 1530–1810 m.
Distribution: MP, NL, SL—Gereau 4243
Syzygium guineense (Willd.) DC. subsp.
afromontanum F. White; Verdcourt, F.T.E.A.
(Myrtaceae): 79. 2001.
Tree. Riverine forest, lower montane forest,
woodland. 1400–1810 m.
Distribution: MP, NL, SL—Harder 1502
Syzygium guineense (Willd.) DC. subsp. guineense;
Verdcourt, F.T.E.A. (Myrtaceae): 78. 2001.
Tree. Riverine forest, woodland, edges of
LakeNyasa_5-18.indd 77
77
cultivation. 474–1400 m.
Distribution: MP, SL—Mwasumbi 16590
Syzygium guineense (Willd.) DC. subsp. macrocarpum (Engl.) F. White; Verdcourt, F.T.E.A.
(Myrtaceae): 79. 2001.
Tree. Riverine forest. 1300 m.
Distribution: MP—J. Lovett 3594
Syzygium masukuense (Baker) R. E. Fr. subsp.
masukuense; Verdcourt, F.T.E.A. (Myrtaceae):
82. 2001.
Tree. Montane forest. 2010 m.
Distribution: RM—WCS SHCP 14
OCHNACEAE
1 genus and 7 species
Ochna afzelii R. Br. ex Oliv. subsp. mechowiana
(O. Hoffm.) N. Robson; Verdcourt, F.T.E.A.
(Ochnaceae): 34. 2005.
Shrub. Miombo woodland. 1780 m.
Distribution: SL—Gereau 5108
Ochna holstii Engl.; Verdcourt, F.T.E.A. (Ochnaceae): 23. 2005.
Tree. Riverine forest, montane forest. 1450–
1990 m.
Distribution: MP, RM—Kibure 1413
Ochna oxyphylla N. Robson; Verdcourt, F.T.E.A.
(Ochnaceae): 24. 2005.
Shrub or tree. Forest patches above treeline,
montane forest edges. 2150–2320 m.
Distribution: MR, SL—Gereau 3982
Ochna polyneura Gilg vel sp. aff.; Verdcourt,
F.T.E.A. (Ochnaceae): 25. 2005.
Suffrutex. Riverine forest. 1200–1300 m.
Distribution: MP—J. Lovett 4906
Ochna puberula N. Robson; Verdcourt, F.T.E.A.
(Ochnaceae): 26. 2005.
Tree or shrub. Woodland. 1400–2000 m.
Distribution: MP, SL—Gereau 4021
Ochna richardsiae N. Robson; Verdcourt, F.T.E.A.
(Ochnaceae): 31. 2005.
Suffrutex. Woodland. 1300–1600 m.
Distribution: MP—J. Lovett 3612
Ochna schweinfurthiana F. Hoffm.; Verdcourt,
F.T.E.A. (Ochnaceae): 33. 2005.
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78
Gereau et al.
Shrub or tree. Woodland. 1300–1400 m.
Distribution: MP—J. Lovett 4810
OLACACEAE
3 genera and 3 species
Olax dissitiflora Oliv.; Lucas, F.T.E.A. (Olacaceae):
9. 1968.
Shrub. Lakeshore bushland. 474–490 m.
Distribution: SL—Mwasumbi 16571
Strombosia scheffleri Engl.; Lucas, F.T.E.A.
(Olacaceae): 11. 1968.
Tree. Riverine forest, montane forest. 910–1950
m.
Distribution: MP, RM, NL, SL—Kayombo 1071
Olea capensis L. subsp. macrocarpa (C. H. Wright)
I. Verd.; Lovett et al., 2006: 197.
Tree. Montane forest. 1840–2340 m.
Distribution: NL, RM—Abeid 2993
Schrebera alata (Hochst.) Welw.; Turrill, F.T.E.A.
(Oleaceae): 4. 1952.
Tree. Riverine forest, montane thickets, woodland, abandoned cultivation. 1250–1850 m.
Distribution: RM, SL—Gereau 4026
Schrebera trichoclada Welw.; Turrill, F.T.E.A.
(Oleaceae): 2. 1952.
Tree or shrub. Woodland. 1200–1400 m.
Distribution: MP—J. Lovett 4337
OLINIACEAE
Ximenia americana L.; Lucas, F.T.E.A. (Olacaceae): 3. 1968.
Tree. Miombo woodland. 1350 m.
Distribution: MP—Kayombo 1091
OLEACEAE
3 genera and 8 species
Jasminum abyssinicum Hochst. ex DC.; Turrill,
F.T.E.A. (Oleaceae): 18. 1952.
Liana. Montane forest. 1580–1900 m.
Distribution: KP, RM—Mwasumbi 16331
Jasminum fluminense Vell.; Turrill, F.T.E.A. (Oleaceae): 19. 1952.
Liana. Disturbed montane forest.
Distribution: RM—Ndangalasi 961
Jasminum odoratissimum L. subsp. goetzeanum
(Gilg) P. S. Green; Kupicha, Fl. Zamb. 7(1):
310. 1983.
Lianescent shrub. Montane forest and thickets,
tree clumps in grassland. 1650–2360 m.
Distribution: MR, RM, SL—Gereau 4716
Jasminum schimperi Vatke; Kupicha, Fl. Zamb.
7(1): 318. 1983.
Liana. Wooded grassland. 1270 m.
Distribution: MP—Gereau 3476
Jasminum streptopus E. Mey.; Kupicha, Fl. Zamb.
7(1): 315. 1983.
Twining shrub. Woodland, abandoned cultivation, road banks. 1200–1400 m.
Distribution: MP—Gereau 7007
LakeNyasa_5-18.indd 78
1 genus and 1 species
Olinia rochetiana A. Juss.; Verdcourt, F.T.E.A.
(Oliniaceae): 2. 1975.
Tree. Montane forest, riverine forest, montane
heathland. 1700–2770 m.
Distribution: MP, NL, RM, SL—Abeid 2811
ONAGRACEAE
3 genera and 4 species
Epilobium capense Buchinger ex Hochst.; Raven,
Fl. Zamb. 4: 343. 1978.
Terrestrial herb. Montane forest edges. 2360 m.
Distribution: MR—Gereau 3729
Epilobium salignum Hausskn.; Raven, Fl. Zamb.
4: 345. 1978.
Terrestrial herb. Montane grassland, roadsides.
2210–2550 m.
Distribution: KP, MR, RM—Suleiman 43
Ludwigia abyssinica A. Rich.; Raven, Fl. Zamb.
4: 340. 1978.
Terrestrial herb. Wet meadows, riverine forest.
580–1250 m.
Distribution: MP, SL—Gereau 3804
Oenothera glazioviana Micheli; Dietrich et al.,
1997: 107.
Shrub. Secondary brushland. 1610 m.
Distribution: RM—Mwasumbi 16336
OPILIACEAE
1 genus and 1 species
5/31/12 3:59 PM
Lake Nyasa Climatic Region
Floristic Checklist
Opilia amentacea Roxb.; Hiepko, 1989: 358.
Liana. Montane forest and thickets, lakeshore
forest. 474–2230 m.
Distribution: MR, NL, SL—Gereau 4214
ORCHIDACEAE
28 genera and 97 species
Aerangis carnea J. L. Stewart; Stewart, F.T.E.A.
(Orchidaceae 3): 558. 1989.
Epiphyte. Riverine forest. 1400–1830 m.
Distribution: MP, MR—Cunningham s.n. (type:
K), J. Lovett 4819
Aerangis oligantha Schltr.; Stewart, F.T.E.A.
(Orchidaceae 3): 565. 1989.
Epiphyte. Montane forest. 1800 m.
Distribution: MR, RM—Stolz 2399 (type: K),
Pettersson 485
Angraecum chamaeanthus Schltr.; Cribb, F.T.E.A.
(Orchidaceae 3): 494. 1989.
Epiphyte. Montane forest.
Distribution: RM—Richards 14306
Angraecum stolzii Schltr.; Cribb, F.T.E.A. (Orchidaceae 3): 498. 1989.
Epiphyte. Montane forest. 2000–2500 m.
Distribution: MR, RM—Stolz 2508 (type: B,
K), Cribb 11318
Ansellia africana Lindl.; Cribb, F.T.E.A. (Orchidaceae 2): 402. 1984.
Epiphyte. Montane forest. 1675 m.
Distribution: RM—WCS SHCP 254
Bonatea stereophylla (Kraenzl.) Summerh.; Summerhayes, F.T.E.A. (Orchidaceae 1): 140.
1968.
Terrestrial herb. Montane grassland. 1500 m.
Distribution: RM—Goetze 1339 (type: B
[destroyed])
Brachycorythis pleistophylla Rchb. f.; Summerhayes, F.T.E.A. (Orchidaceae 1): 22. 1968.
Terrestrial herb. Miombo woodland, abandoned cultivation. 1780–1790 m.
Distribution: MP, SL—Gereau 3879A
Brownleea parviflora Harv. ex Lindl.; Summerhayes, F.T.E.A. (Orchidaceae 1): 179. 1968.
Terrestrial herb. Montane grassland, cypress
plantation. 1800–2740 m.
LakeNyasa_5-18.indd 79
79
Distribution: KP, MP, MR, RM—J. Lovett 1875
Bulbophyllum flectens P. J. Cribb & P. Taylor;
Cribb, F.T.E.A. (Orchidaceae 2): 307. 1984.
Epiphyte. Montane forest.
Distribution: RM—Leedal 1341
Bulbophyllum longiflorum Thouars;
F.T.E.A. (Orchidaceae 2): 304. 1984.
Epiphyte. Forest.
Distribution: RM—Cribb 10666
Cribb,
Bulbophyllum mahonii Rolfe; Cribb, F.T.E.A.
(Orchidaceae 2): 310. 1984.
Epiphyte. Montane forest.
Distribution: NL—Cribb 10626
Bulbophyllum cf. mahonii Rolfe; Cribb, F.T.E.A.
(Orchidaceae 2): 310. 1984.
Epiphyte. Disturbed montane forest.
Distribution: RM—Ndangalasi 1020
Calanthe sylvatica (Thouars) Lindl.; Cribb,
F.T.E.A. (Orchidaceae 2): 282. 1984.
Terrestrial herb. Montane forest. 1620–2600
m.
Distribution: MR, NL, RM—Suleiman 76
Corycium nigrescens Sond.; Cribb & Kurzweil,
1994: 555.
Terrestrial herb. Montane grassland.
Distribution: KP—Leedal 4964
Cynorkis anacamptoides Kraenzl.; Summerhayes,
F.T.E.A. (Orchidaceae 1): 32. 1968.
Terrestrial herb. Marshes and stream banks in
montane grassland. 2400–2600 m.
Distribution: KP, RM, SL—Suleiman 49
Cynorkis kaessneriana Kraenzl.; Summerhayes,
F.T.E.A. (Orchidaceae 1): 33. 1968.
Terrestrial herb. Open volcanic ridges, Protea
woodland. 1800–2550 m.
Distribution: RM—Abeid 2838
Cynorkis rungweensis Schltr.; Summerhayes,
F.T.E.A. (Orchidaceae 1): 35. 1968.
Terrestrial herb. Montane grassland. 2000 m.
Distribution: RM—Stolz 2532 (type: K)
Cynorkis symoensii Geerinck & Tournay; Geerinck,
1984: 165.
Terrestrial herb. Open volcanic ridges. 1800–
2240 m.
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80
Distribution: RM—Bidgood 96
Note: New to East Africa, from Malawi and
Rwanda.
Diaphananthe meliantha P. J. Cribb; Cribb,
F.T.E.A. (Orchidaceae 3): 540. 1989.
Epiphyte. Montane forest. 2070 m.
Distribution: NL—Gereau 4141
Disa aconitoides Sond. subsp. concinna (N. E. Br.)
H. P. Linder; Linder, 1981: 91.
Terrestrial herb. Woodland. 1250–1400 m.
Distribution: MP—J. Lovett 5037
Disa aconitoides Sond. subsp. goetzeana (Kraenzl.)
H. P. Linder; Linder, 1981: 92.
Terrestrial herb. Miombo woodland. 1790 m.
Distribution: SL—Gereau 3883
Gereau et al.
(Orchidaceae 1): 168. 1968.
Terrestrial herb. Montane grassland. 2150–
2700 m.
Distribution: KP, MR, RM—Gereau 3781
Disa rungweensis Schltr.; Summerhayes, F.T.E.A.
(Orchidaceae 1): 175. 1968.
Terrestrial herb. Rocky mountain slopes.
2400–2960 m.
Distribution: RM—Stolz 2437 (type: K), Richards 14302
Disa stolzii Schltr.; Summerhayes, F.T.E.A. (Orchidaceae 1): 156. 1968.
Terrestrial herb. Montane grassland, rocky
slopes. 2600–2800 m.
Distribution: KP, RM—Stolz 2506 (type: K),
Abeid 3092
Disa equestris Rchb. f. vel sp. aff.; Linder, 1981:
99.
Terrestrial herb. Rocky mountain summit.
2800 m.
Distribution: KP—Goldblatt 8206
Disa walleri Rchb. f.; Summerhayes, F.T.E.A.
(Orchidaceae 1): 169. 1968.
Terrestrial herb. Miombo woodland. 1450 m.
Distribution: MP—Gereau 6982
Disa erubescens Rendle; Summerhayes, F.T.E.A.
(Orchidaceae 1): 157. 1968.
Terrestrial herb. Montane grassland. 2400–
2700 m.
Distribution: KP, NL, SL—Suleiman 172
Disa walteri Schltr.; Summerhayes, F.T.E.A.
(Orchidaceae 1): 176. 1968.
Terrestrial herb. Montane grassland. 2200–
2800 m.
Distribution: MP, SL—Goetze 925 (type: B
[destroyed]), Kerfoot 1632
Disa fragrans Schltr. subsp. fragrans; Linder,
1999: 197.
Terrestrial herb. Montane grassland. 2740 m.
Distribution: KP, MP—Wingfield 952
Disa hamatopetala Rendle; Summerhayes,
F.T.E.A. (Orchidaceae 1): 177. 1968.
Terrestrial herb. Montane grassland.
Distribution: KP—Willan 176
Disa longilabris Schltr.; Summerhayes, F.T.E.A.
(Orchidaceae 1): 177. 1968.
Terrestrial herb. Montane grassland. 2550–
2760 m.
Distribution: KP, RM, SL—Goetze 1222, 1226
(syntypes: B [destroyed]), Abeid 2837
Disa ochrostachya Rchb. f.; Summerhayes,
F.T.E.A. (Orchidaceae 1): 164. 1968.
Terrestrial herb. Montane grassland. 2440 m.
Distribution: SL—Suleiman 183
Disa robusta N. E. Br.; Summerhayes, F.T.E.A.
LakeNyasa_5-18.indd 80
Disa welwitschii Rchb. f. vel sp. aff.; Summerhayes, F.T.E.A. (Orchidaceae 1): 160. 1968.
Terrestrial herb. Rocky montane grassland.
2720 m.
Distribution: KP—Abeid 3055
Disperis anthoceros Rchb. f.; Verdcourt, F.T.E.A.
(Orchidaceae 1): 229. 1968.
Terrestrial herb. Montane forest, woodland.
1250–2140 m.
Distribution: MP, SL—Suleiman 103
Eulophia clavicornis Lindl. var. nutans (Sond.) A.
V. Hall; Cribb, F.T.E.A. (Orchidaceae 3): 450.
1989.
Terrestrial herb. Montane grassland. 2640 m.
Distribution: KP—Abeid 3081
Eulophia elegans Schltr.; Cribb, F.T.E.A. (Orchidaceae 3): 446. 1989.
Terrestrial herb. Wet grassland at riverine
forest edge. 1600 m.
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Lake Nyasa Climatic Region
Floristic Checklist
81
Distribution: MP, RM—Stolz 1980 (type: B, K,
Z), Kerfoot 1725
Distribution: MR, RM—Stolz 1148 (type: K,
S), Pettersson 472
Euolphia fridericii (Rchb. f.) A. V. Hall; Cribb,
F.T.E.A. (Orchidaceae 3): 471. 1989.
Terrestrial herb. Montane grassland. 1940 m.
Distribution: SL—Gereau 4672
Habenaria kyimbilae Schltr.; Summerhayes,
F.T.E.A. (Orchidaceae 1): 87. 1968.
Terrestrial herb. Montane grassland. 2230–
2600 m.
Distribution: KP, SL—Gereau 4436
Eulophia guineensis Lindl.; Cribb, F.T.E.A. (Orchidaceae 3): 428. 1989.
Terrestrial herb. Abandoned cultivation. 1250
m.
Distribution: MP—J. Lovett 3934
Eulophia horsfallii (Bateman) Summerh.; Cribb,
F.T.E.A. (Orchidaceae 3): 428. 1989.
Terrestrial herb. Riverine forest. 1850 m.
Distribution: RM, SL—Gereau 4045
Eulophia kyimbilae Schltr.; Cribb, F.T.E.A. (Orchidaceae 3): 446. 1989.
Terrestrial herb. Grassland, forest edges.
Distribution: MP, MR—Cribb 11305
Eulophia odontoglossa Rchb. f.; Cribb, F.T.E.A.
(Orchidaceae 3): 458. 1989.
Terrestrial herb. Miombo woodland, Cupressus
plantations, maize fields. 1540–1790 m.
Distribution: RM, SL—Gereau 3878
Eulophia streptopetala Lindl.; Cribb, F.T.E.A.
(Orchidaceae 3): 469. 1989.
Epilithic herb. Riverine forest. 1500 m.
Distribution: SL—Gereau 3946
Habenaria clavata (Lindl.) Rchb. f.; Summerhayes, F.T.E.A. (Orchidaceae 1): 109. 1968.
Terrestrial herb. Woodland. 1250–1790 m.
Distribution: MP, SL—Gereau 3890
Habenaria cornuta Lindl.; Summerhayes, F.T.E.A.
(Orchidaceae 1): 106. 1968.
Terrestrial herb. Montane grassland, cypress
plantation. 1750–2400 m.
Distribution: KP, MP, RM—Suleiman 51
Habenaria goetzeana Kraenzl.; Summerhayes,
F.T.E.A. (Orchidaceae 1): 114. 1968.
Terrestrial herb. Miombo woodland. 1790 m.
Distribution: SL—Gereau 3888
Habenaria isoantha Schltr.; Summerhayes,
F.T.E.A. (Orchidaceae 1): 100. 1968.
Terrestrial herb. Grassland. 1600–1800 m.
LakeNyasa_5-18.indd 81
Habenaria leucoceras Schltr.; Summerhayes,
F.T.E.A. (Orchidaceae 1): 90. 1968.
Terrestrial herb. Montane grassland. 2000 m.
Distribution: MP, NL—Stolz 2490 (type: K),
Procter 1911
Habenaria leucotricha Schltr.; Summerhayes,
F.T.E.A. (Orchidaceae 1): 123. 1968.
Terrestrial herb. Woodland. 1200 m.
Distribution: MP—J. Lovett 3875
Habenaria lithophila Schltr.; Summerhayes,
F.T.E.A. (Orchidaceae 1): 132. 1968.
Terrestrial herb. Rocky mountain slopes. 2800
m.
Distribution: RM—Stolz 2505 (type: K)
Habenaria macrura Kraenzl.; Summerhayes,
F.T.E.A. (Orchidaceae 1): 126. 1968.
Terrestrial herb. Montane grassland. 2400–
2600 m.
Distribution: KP—Suleiman 52
Habenaria malacophylla Rchb. f.; Summerhayes,
F.T.E.A. (Orchidaceae 1): 73. 1968.
Terrestrial herb. Montane forest. 2120–2600
m.
Distribution: MR, SL—Gereau 4337
Habenaria occlusa Summerh.; Summerhayes,
F.T.E.A. (Orchidaceae 1): 67. 1968.
Terrestrial herb. Montane grassland. 2100–
2590 m.
Distribution: KP, MP—Richards 14178 (type:
K), Suleiman 20
Habenaria odorata Schltr.; Summerhayes,
F.T.E.A. (Orchidaceae 1): 121. 1968.
Terrestrial herb. Montane grassland. 2100 m.
Distribution: NL—Stolz 2488 (type: K)
Habenaria petitiana (A. Rich.) T. Durand &
Schinz; Summerhayes, F.T.E.A. (Orchidaceae
1): 56. 1968.
Terrestrial herb. Scree slopes, montane
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82
grassland. 2060–2120 m.
Distribution: RM, SL—Gereau 4447
Habenaria praestans Rendle; Summerhayes,
F.T.E.A. (Orchidaceae 1): 70. 1968.
Terrestrial herb. Riverine forest, miombo
woodland, roadsides in cultivation. 1500–1900 m.
Distribution: MP, MR, NL, SL—Gereau 4407
Habenaria schimperiana Hochst. ex A. Rich.;
Summerhayes, F.T.E.A. (Orchidaceae 1): 89.
1968.
Terrestrial herb. Woodland. 1200 m.
Distribution: MP—J. Lovett 4243
Habenaria uhehensis Schltr.; Summerhayes,
F.T.E.A. (Orchidaceae 1): 77. 1968.
Terrestrial herb. Grassland. 2440 m.
Distribution: MP, RM—Nicholson DSM2546
Habenaria cf. welwitschii Rchb. f.; Summerhayes,
F.T.E.A. (Orchidaceae 1): 83. 1968.
Terrestrial herb. Montane grassland. 2400 m.
Distribution: KP—Suleiman 54
Habenaria xanthochlora Schltr.; Summerhayes,
F.T.E.A. (Orchidaceae): 56. 1968.
Terrestrial herb. Montane grassland. 2590–
2600 m.
Distribution: KP—Lovett 1859
Holothrix nyasae Rolfe; Summerhayes, F.T.E.A.
(Orchidaceae 1): 7. 1968.
Terrestrial herb. Montane grassland. 2400–
2740 m.
Distribution: KP, MP, NL—Suleiman 50
Holothrix triloba (Rolfe) Kraenzl.; Summerhayes,
F.T.E.A. (Orchidaceae 1): 11. 1968.
Terrestrial herb. Montane grassland. 2100–
2250 m.
Distribution: RM—Procter 1341
Liparis bowkeri Harv.; Cribb & Bowden, F.T.E.A.
(Orchidaceae 2): 299. 1984.
Terrestrial or epilithic herb. Miombo woodland,
riverine forest, scree slopes. 1210–2120 m.
Distribution: SL—Gereau 4406
Liparis deistelii Schltr.; Cribb & Bowden, F.T.E.A.
(Orchidaceae 2): 299. 1984.
Terrestrial or epiphytic herb. Forest.
Distribution: RM—Richards 14352
LakeNyasa_5-18.indd 82
Gereau et al.
Liparis nervosa (Thunb.) Lindl.; Cribb & Bowden,
F.T.E.A. (Orchidaceae 2): 298. 1984.
Terrestrial herb. Miombo woodland. 1530 m.
Distribution: SL—Gereau 4370
Liparis rungweensis Schltr.; Cribb & Bowden,
F.T.E.A. (Orchidaceae 2): 297. 1984.
Terrestrial herb. Montane grassland. 1600–
2960 m.
Distribution: MP, NL, RM—Stolz 2438 (type:
K), Gereau 4089
Liparis stolzii Schltr.; Cribb & Bowden, F.T.E.A.
(Orchidaceae 2): 297. 1984.
Epiphyte. Forest. 1600 m.
Distribution: RM—Stolz 1152 (type: K)
Malaxis schliebenii (Mansf.) Summerh.; Cribb,
F.T.E.A. (Orchidaceae 2): 286. 1984.
Terrestrial herb. Miombo woodland. 1530–
1790 m.
Distribution: SL—Gereau 4366
Margelliantha leedalii P. J. Cribb; Cribb, F.T.E.A.
(Orchidaceae 3): 541. 1989.
Epiphyte. Montane forest. 2070 m.
Distribution: NL, RM—Gereau 4139
Mystacidium tanganyikense Summerh.; Cribb,
F.T.E.A. (Orchidaceae 3): 594. 1989.
Epiphyte. Montane forest. 2070 m.
Distribution: MP, NL—Gereau 4140
Neobolusia stolzii Schltr.; Summerhayes, F.T.E.A.
(Orchidaceae 1): 13. 1968.
Terrestrial herb. Grassland. 1850–2600 m.
Distribution: MR, RM—Stolz 1076 (type: K),
Wingfield 173
Platycoryne crocea (Schweinf. ex Rchb. f.) Rolfe
subsp. ochrantha (Schltr.) Summerh.; Summerhayes, F.T.E.A. (Orchidaceae 1): 148.
1968.
Terrestrial herb. Miombo woodland. 1200 m.
Distribution: MP, NL—J. Lovett 3991
Polystachya albescens Ridl. subsp. imbricata
(Rolfe) Summerh.; Cribb, F.T.E.A. (Orchidaceae 2): 355. 1984.
Terrestrial or epiphytic herb. Montane forest.
1750 m.
Distribution: RM—Semsei 1594
Polystachya cultriformis (Thouars) Lindl. ex
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Lake Nyasa Climatic Region
Floristic Checklist
83
Spreng.; Cribb, F.T.E.A. (Orchidaceae 2): 341.
1984.
Epiphyte. Montane forest, riverine forest. 2000
2650–2800 m.
Distribution: MP, RM: Stolz 2504 (type: K),
Wingfield 936
Distribution: MP, RM—J. Lovett 4987
Satyrium crassicaule Rendle; Summerhayes,
F.T.E.A. (Orchidaceae 1): 193. 1968.
Terrestrial herb. Riverbanks and marshes in
montane grassland. 1860–2600 m.
Distribution: KP, MP, SL—Gereau 3863
m.
Polystachya fusiformis (Thouars) Lindl.; Cribb,
F.T.E.A. (Orchidaceae 2): 371. 1984.
Epiphyte. Montane forest. 2070 m.
Distribution: NL—Gereau 4138
Polystachya malilaensis Schltr.; Cribb, F.T.E.A.
(Orchidaceae 2): 398. 1984.
Epiphytic or epilithic herb. Montane forest,
riverine forest. 1850–1980 m.
Distribution: MP, SL—Gereau 4037
Polystachya modesta Rchb. f.; Cribb, F.T.E.A.
(Orchidaceae 2): 363. 1984.
Epiphyte. Riverine forest. 1210 m.
Distribution: SL—Gereau 4266
Pterygodium ukingense Schltr.; Hunt, F.T.E.A.
(Orchidaceae 1): 216. 1968.
Terrestrial herb. Montane grassland. 2000 m.
Distribution: NL—Stolz 2609 (type: K)
Satyrium acutirostrum Summerh.; Summerhayes,
F.T.E.A. (Orchidaceae 1): 196. 1968.
Terrestrial herb. Montane grassland. 1950–
2610 m.
Distribution: KP, MP, RM—Suleiman 21
Satyrium anomalum Schltr.; Summerhayes,
F.T.E.A. (Orchidaceae 1): 205. 1968.
Terrestrial herb. Miombo woodland. 1790 m.
Distribution: SL—Gereau 3887
Satyrium buchananii Schltr.; Summerhayes,
F.T.E.A. (Orchidaceae 1): 199. 1968.
Terrestrial herb. Montane grassland, riverine
forest, miombo woodland. 1790–2700 m.
Distribution: KP, NL, RM, SL—Suleiman 65
Satyrium chlorocorys Rchb. f. ex Rolfe; Summerhayes, F.T.E.A. (Orchidaceae 1): 206. 1968.
Terrestrial herb. Montane grassland, miombo
woodland. 1610–2400 m.
Distribution: KP, MP, RM, SL—Gereau 3866
Satyrium comptum Summerh.; Summerhayes,
F.T.E.A. (Orchidaceae 1): 208. 1968.
Terrestrial herb. Cracks in rock face.
LakeNyasa_5-18.indd 83
Satyrium longicauda Lindl.; Summerhayes,
F.T.E.A. (Orchidaceae 1): 198. 1968.
Terrestrial herb. Montane grassland. 2610 m.
Distribution: KP, MP—Kerfoot 1607
Satyrium microcorys Schltr.; Summerhayes,
F.T.E.A. (Orchidaceae 1): 207. 1968.
Terrestrial herb. Rocky montane grassland.
2000–2570 m.
Distribution: KP, MP, RM—Stolz 2436 (type:
K), Gereau 3165
Satyrium miserum Kraenzl. vel sp. aff.; Summerhayes, F.T.E.A. (Orchidaceae 1): 207. 1968.
Terrestrial herb. Rocky montane grassland.
2730 m.
Distribution: KP—Abeid 3079
Satyrium monadenum Schltr.; Summerhayes,
F.T.E.A. (Orchidaceae 1): 195. 1968.
Terrestrial herb. Montane grassland. 2400–
2840 m.
Distribution: KP, RM—Stolz 1031 (type: K),
Abeid 2785
Satyrium neglectum Schltr.; Summerhayes,
F.T.E.A. (Orchidaceae 1): 197. 1968.
Terrestrial herb. Montane grassland, disturbed
shrubland. 1850–2740 m.
Distribution: KP, MP, NL, RM, SL—Gereau
4231
Satyrium princeae Kraenzl.; Summerhayes,
F.T.E.A. (Orchidaceae 1): 189. 1968.
Terrestrial herb. Montane grassland. 2400–
2550 m.
Distribution: KP, RM, SL—Suleiman 30
Satyrium rhynchantoides Schltr.; Summerhayes,
F.T.E.A. (Orchidaceae 1): 195. 1968.
Terrestrial herb. Montane grassland. 2400 m.
Distribution: RM—Stolz 1074, 2435 (syntypes:
K)
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84
Satyrium sceptrum Schltr.; Summerhayes, F.T.E.A.
(Orchidaceae 1): 197. 1968.
Terrestrial herb. Montane grassland. 2000–
2570 m.
Distribution: KP, RM—Stolz 2434 (type: EA,
K), Gereau 3091
Gereau et al.
OXALIDACEAE
2 genera and 4 species
Biophytum abyssinicum Steud. ex A. Rich.;
Kabuye, F.T.E.A. (Oxalidaceae): 12. 1971.
Terrestrial herb. Woodland. 1250 m.
Distribution: MP—J. Lovett 4073
Satyrium sphaeranthum Schltr.; Summerhayes,
F.T.E.A. (Orchidaceae 1): 208. 1968.
Terrestrial herb. Montane grassland, montane
forest. 2080–2600 m.
Distribution: KP, MP, RM, SL—Suleiman 111
Biophytum umbraculum Welw.; Lourteig, 1981:
451.
Terrestrial herb. Fallow fields. 520–1620 m.
Distribution: RM, SL—Mboya 241
Satyrium trinerve Lindl.; Hall, 1999: 155.
Terrestrial herb. Montane grassland, miombo
woodland. 1790–1860 m.
Distribution: SL—Gereau 3882
Oxalis corniculata L.; Kabuye, F.T.E.A. (Oxalidaceae): 3. 1971.
Terrestrial herb. Montane forest.
Distribution: MP, RM—Bullock 3987
Satyrium volkensii Schltr.; Summerhayes, F.T.E.A.
(Orchidaceae 1): 202. 1968.
Terrestrial herb. Woodland, montane grassland. 1300–1860 m.
Distribution: MP, SL—Gereau 3879
Oxalis obliquifolia Steud. ex A. Rich.; Kabuye,
F.T.E.A. (Oxalidaceae): 6. 1971.
Terrestrial herb. Rocky mountain summit,
montane grassland. 2590–2600 m.
Distribution: KP, MP—Gereau 3500
Schizochilus sulphureus Schltr.; Summerhayes,
F.T.E.A. (Orchidaceae 1): 27. 1968.
Terrestrial herb. Montane grassland. 2400–
2750 m.
Distribution: KP, MP, RM—Stolz 1075 (type:
K), Kerfoot 1631
Stenoglottis fimbriata Lindl.; Summerhayes,
F.T.E.A. (Orchidaceae 1): 29. 1968.
Epiphyte. Montane forest and openings. 1800
m.
Distribution: NL, RM—Bidgood 98
Ypsilopus erectus (P. J. Cribb) P. J. Cribb & J. L.
Stewart; Cribb, F.T.E.A. (Orchidaceae 3): 628.
1989.
Epiphyte. Woodland.
Distribution: RM—Moreau 871
PASSIFLORACEAE
2 genera and 3 species
Adenia lindiensis Harms; Wilde, F.T.E.A. (Passifloraceae): 39. 1975.
Herbaceous climber. Riverine forest. 580 m.
Distribution: SL—Gereau 3814
Adenia stenodactyla Harms; Wilde, F.T.E.A. (Passifloraceae): 37. 1975.
Herbaceous climber. Woodland, montane
thickets. 1300–1920 m.
Distribution: MP, SL—Gereau 3609
Passiflora edulis Sims; Wilde, F.T.E.A. (Passifloraceae): 15. 1975.
Herbaceous climber. Montane forest edges.
1720–1850 m.
Distribution: RM—Bracebridge 91
OROBANCHACEAE
1 genus and 1 species
Orobanche minor Sm.; Graham, F.T.E.A. (Orobanchaceae): 5. 1957.
Parasitic terrestrial herb. Woodland, riverine
forest, montane forest and shrubland, abandoned
cultivation. 1250–2500 m.
Distribution: MP, SL—Suleiman 182
PEDALIACEAE
1 genus and 2 species
Sesamum angolense Welw.; Bruce, F.T.E.A. (Pedaliaceae): 20. 1953.
Herb or subshrub. Woodland. 1200–1460 m.
Distribution: MP—Kayombo 2428
Sesamum angustifolium (Oliv.) Engl.; Bruce,
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F.T.E.A. (Pedaliaceae): 19. 1953.
Terrestrial herb. Woodland, forest edges. 1200–
1520 m.
Distribution: MP, RM—J. Lovett 4242
PIPERACEAE
2 genera and 5 species
Peperomia fernandopoiana C. DC. var. butaguensis (De Wild.) Düll.; Verdcourt, F.T.E.A.
(Piperaceae): 21. 1996.
Epiphyte. Montane forest. 1750 m.
Distribution: RM—Kindeketa 2972
Note: New to Tanzania, from Kenya, Uganda,
Democratic Republic of Congo, and Rwanda.
85
Distribution: KP, MR, SL—Gereau 4271
PLANTAGINACEAE
1 genus and 2 species
Plantago lanceolata L.; Verdcourt, F.T.E.A. (Plantaginaceae): 6. 1971.
Terrestrial herb. Montane grassland. 2520 m.
Distribution: KP—WCS SHCP 407
Plantago palmata Hook. f.; Verdcourt, F.T.E.A.
(Plantaginaceae): 2. 1971.
Terrestrial herb. Montane forest. 2015 m.
Distribution: MR, RM—Gereau 7010
PLUMBAGINACEAE
Peperomia retusa (L. f.) A. Dietr.; Verdcourt,
F.T.E.A. (Piperaceae): 13. 1996.
Epiphyte. Montane forest. 1780 m.
Distribution: RM—Bracebridge 99A
Peperomia tetraphylla (G. Forst.) Hook. & Arn.;
Verdcourt, F.T.E.A. (Piperaceae): 12. 1996.
Epiphyte. Montane forest. 1780 m.
Distribution: RM—Bracebridge 99
1 genus and 1 species
Plumbago zeylanica L.; Wilmot-Dear, F.T.E.A.
(Plumbaginaceae): 6. 1976.
Shrub. Montane forest edges. 1760 m.
Distribution: RM—WCS SHCP 171
POACEAE
50 genera and 124 species
Piper capense L. f.; Verdcourt, F.T.E.A. (Piperaceae): 5. 1996.
Shrub. Montane forest, tree clumps in grassland. 1720–2460 m.
Distribution: MP, MR, NL, RM, SL—Gereau
4713
Agrostis continuata Stapf; Clayton, F.T.E.A. (Gramineae 1): 111. 1970.
Terrestrial herb. Stream banks in montane
grassland. 2560 m.
Distribution: KP—Gereau 3113
Piper umbellatum L.; Verdcourt, F.T.E.A. (Piperaceae): 8. 1996.
Shrub. Riverine forest. 1160–1210 m.
Distribution: SL—Gereau 4260
Agrostis kilimandscharica Mez; Clayton, F.T.E.A.
(Gramineae 1): 110. 1970.
Terrestrial herb. Montane grassland, seasonal
swamps, rock face near waterfall. 2570–2920 m.
Distribution: KP, MP, RM—Gereau 3082
PITTOSPORACEAE
1 genus and 2 species
Pittosporum abyssinicum Delile; Lovett et al.,
2006: 201.
Tree. Montane forest edges, tree clumps in
montane grassland. 2410–2700 m.
Distribution: KP, NL, RM—Kibure 1412
Pittosporum viridiflorum Sims; Lovett et al., 2006:
201.
Tree or shrub. Montane forest, riverine
forest, miombo woodland, lakeshore shrubland.
474–2470 m.
LakeNyasa_5-18.indd 85
Agrostis lachnantha Nees; Clayton, F.T.E.A. (Gramineae 1): 106. 1970.
Terrestrial herb. Open woodland. 1650–1680
m.
Distribution: SL—Gereau 5113
Andropogon amethystinus Steud.; Clayton & Renvoize, F.T.E.A. (Gramineae 3): 772. 1982.
Terrestrial herb. Road beds in cultivation,
montane grassland. 2200–2840 m.
Distribution: KP, MR—Abeid 2853
Andropogon chinensis (Nees) Merr.; Clayton &
Renvoize, F.T.E.A. (Gramineae 3): 779. 1982.
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Gereau et al.
Terrestrial herb. Riverine forest. 1500 m.
Distribution: SL—Gereau 4806
Andropogon chrysostachyus Steud.; Clayton &
Renvoize, F.T.E.A. (Gramineae 3): 773. 1982.
Terrestrial herb. Montane grassland. 2525–
2770 m.
Distribution: KP, RM—Abeid 2831
Andropogon distachyos L.; Clayton & Renvoize,
F.T.E.A. (Gramineae 3): 770. 1982.
Terrestrial herb. Open and disturbed places in
montane forest, montane grassland. 2200–2300
m.
Distribution: MR—Gereau 4616
Andropogon mannii Hook. f.; Clayton & Renvoize,
F.T.E.A. (Gramineae 3): 774. 1982.
Terrestrial herb. Montane grassland, stream
banks. 2440–2740 m.
Distribution: KP, MR, RM—Gereau 3170
Andropogon schirensis Hochst. ex A. Rich.; Clayton & Renvoize, F.T.E.A. (Gramineae 3): 779.
1982.
Terrestrial herb. Montane grassland, woodland.
1260–2320 m.
Distribution: MP, SL—Suleiman 162
Andropterum stolzii (Pilg.) C. E. Hubb.; Clayton &
Renvoize, F.T.E.A. (Gramineae 3): 752. 1982.
Terrestrial herb. Open woodland, montane
grassland. 1650–2300 m.
Distribution: MR, SL—Gereau 4762
Arthraxon micans (Nees) Hochst.; Clayton & Renvoize, F.T.E.A. (Gramineae 3): 742. 1982.
Terrestrial herb. Montane grassland, road
banks. 1940–2200 m.
Distribution: MR, SL—Gereau 4776
Arundinella nepalensis Trin.; Hubbard, F.T.E.A.
(Gramineae 2): 409. 1974.
Terrestrial herb. Eucalyptus woodland. 1250
m.
Distribution: MP—J. Lovett 4425
Bewsia biflora (Hack.) Gooss.; Clayton, F.T.E.A.
(Gramineae 2): 286. 1974.
Terrestrial herb. Miombo woodland. 1260–
1530 m.
Distribution: SL—Gereau 4361
Bothriochloa insculpta (Hochst. ex A. Rich.) A.
LakeNyasa_5-18.indd 86
Camus; Clayton & Renvoize, F.T.E.A. (Gramineae 3): 720. 1982.
Terrestrial herb. Roadsides in miombo woodland. 1500 m.
Distribution: SL—Gereau 3955
Brachiaria decumbens Stapf; Clayton & Renvoize,
F.T.E.A. (Gramineae 3): 586. 1982.
Terrestrial herb. Abandoned cultivation. 1250
m.
Distribution: MP—J. Lovett 4543
Brachiaria deflexa (Schumach.) C. E. Hubb. ex
Robyns; Clayton & Renvoize, F.T.E.A. (Gramineae 3): 598. 1982.
Terrestrial herb. Roadsides. 474–490 m.
Distribution: SL—Gereau 3789
Brachiaria jubata (Fig. & De Not.) Stapf; Clayton & Renvoize, F.T.E.A. (Gramineae 3): 580.
1982.
Terrestrial herb. Woodland. 1250–1950 m.
Distribution: MP—J. Lovett 3519
Brachiaria reptans (L.) C. A. Gardner & C. E.
Hubb.; Clayton & Renvoize, F.T.E.A. (Gramineae 3): 591. 1982.
Terrestrial herb. Miombo woodland. 1200 m.
Distribution: MP—J. Lovett 4320
Brachiaria scalaris Pilg.; Clayton & Renvoize,
F.T.E.A. (Gramineae 3): 594. 1982.
Terrestrial herb. Woodland. 2000 m.
Distribution: MP—P. Lovett 268
Brachiaria serrata (Thunb.) Stapf; Clayton &
Renvoize, F.T.E.A. (Gramineae 3): 588. 1982.
Terrestrial herb. Miombo woodland. 1200–
1790 m.
Distribution: MP, SL—Gereau 3898
Brachypodium flexum Nees; Clayton, F.T.E.A.
(Gramineae 1): 71. 1970.
Scrambling terrestrial herb. Montane forest,
stream banks in montane grassland. 2040–2570
m.
Distribution: KP, SL—Gereau 4728
Briza minor L.; Hubbard, F.T.E.A. (Gramineae 1):
53. 1970.
Terrestrial herb. Cultivated and waste ground.
2000 m.
Distribution: RM—Semsei 1570
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Bromus leptoclados Nees; Clayton, F.T.E.A. (Gramineae 1): 68. 1970.
Terrestrial herb. Montane grassland. 2360–
2570 m.
Distribution: KP, MR—Gereau 3777
Cymbopogon nardus (L.) Rendle; Clayton & Renvoize, F.T.E.A. (Gramineae 3): 764. 1982.
Terrestrial herb. Miombo woodland. 1260–
1500 m.
Distribution: SL—Gereau 4413
Cynodon nlemfuensis Vanderyst; Clayton, F.T.E.A.
(Gramineae 2): 319. 1974.
Terrestrial herb. Seasonally inundated grassland. 1270 m.
Distribution: MP—Gereau 3490
87
Terrestrial herb. Montane grassland, clearings in montane forest, scree slopes, woodland.
1900–2230 m.
Distribution: NL, SL—Gereau 4442
Digitaria perrottetii (Kunth) Stapf; Clayton &
Renvoize, F.T.E.A. (Gramineae 3): 649. 1982.
Terrestrial herb. Cultivated areas, woodland.
1250–1500 m.
Distribution: MP—Kayombo 2343
Digitaria rukwae Clayton; Clayton & Renvoize,
F.T.E.A. (Gramineae 3): 646. 1982.
Terrestrial herb. Trails and edges of cultivation
in montane grassland. 2470–2590 m.
Distribution: KP—Gereau 3127
Digitaria abyssinica (Hochst. ex A. Rich.) Stapf;
Clayton & Renvoize, F.T.E.A. (Gramineae 3):
641. 1982.
Terrestrial herb. Montane grassland, valley
grassland. 1270–2360 m.
Distribution: MR, SL—Suleiman 157
Digitaria scalarum (Schweinf.) Chiov.; Goetghebeur & Van der Veken, Fl. Zamb. 10(3): 159.
1989.
Terrestrial herb. Seasonal pond margins in
montane grassland, agricultural fields. 1840–2300
m.
Distribution: MR, SL—Gereau 3911
Digitaria diagonalis (Nees) Stapf var. hirsuta (De
Wild. & T. Durand) Stapf ex Troupin; Clayton
& Renvoize, F.T.E.A. (Gramineae 3): 626.
1982.
Terrestrial herb. Miombo woodland. 1790 m.
Distribution: SL—Gereau 3897
Digitaria ternata (Hochst. ex A. Rich.) Stapf;
Clayton & Renvoize, F.T.E.A. (Gramineae 3):
630. 1982.
Terrestrial herb. Montane grassland, road beds,
woodland, abandoned cultivation. 1250–2200 m.
Distribution: MP, MR, SL—Gereau 4777
Digitaria diagonalis (Nees) Stapf var. uniglumis
(Hochst. ex A. Rich.) Pilg.; Clayton & Renvoize, F.T.E.A. (Gramineae 3): 626. 1982.
Terrestrial herb. Miombo woodland. 1200–
1530 m.
Distribution: MP, SL—Gereau 4384
Digitaria thouarsiana (Flüggé) A. Camus; Clayton & Renvoize, F.T.E.A. (Gramineae 3): 631.
1982.
Terrestrial herb. Clearings in montane forest.
1900 m.
Distribution: NL—Gereau 4192
Digitaria gazensis Rendle; Clayton & Renvoize,
F.T.E.A. (Gramineae 3): 643. 1982.
Terrestrial herb. Roadsides in miombo woodland. 1500 m.
Distribution: SL—Gereau 3956
Eleusine coracana (L.) Gaertn. subsp. africana
(Kenn.-O’Byrne) Hilu & de Wet; Hilu, 1994:
411.
Terrestrial herb. Roadsides. 474–490 m.
Distribution: SL—Gereau 3791
Digitaria myurus Stapf; Clayton & Renvoize,
F.T.E.A. (Gramineae 3): 627. 1982.
Terrestrial herb. Montane grassland. 2100–
2700 m.
Distribution: KP, MP, SL—Stolz 2619 (type: K,
US), Kerfoot 1609
Elionurus muticus (Spreng.) Kuntze; Clayton &
Renvoize, F.T.E.A. (Gramineae 3): 837. 1982.
Terrestrial herb. Montane grassland. 2250 m.
Distribution: MP—J. Lovett 4859
Digitaria pearsonii Stapf; Clayton & Renvoize,
F.T.E.A. (Gramineae 3): 643. 1982.
LakeNyasa_5-18.indd 87
Eragrostis capensis (Thunb.) Trin.; Clayton,
F.T.E.A. (Gramineae 2): 221. 1974.
Terrestrial herb. Montane grassland. 1940 m.
Distribution: SL—Gereau 4768
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88
Eragrostis congesta Oliv.; Clayton, F.T.E.A. (Gramineae 2): 224. 1974.
Terrestrial herb. Montane grassland. 1940 m.
Distribution: SL—Gereau 4766
Eragrostis exasperata Peter; Clayton, F.T.E.A.
(Gramineae 2): 220. 1974.
Terrestrial herb. Roadsides. 474–490 m.
Distribution: SL—Gereau 3798
Eragrostis hispida K. Schum.; Clayton, F.T.E.A.
(Gramineae 2): 201. 1974.
Terrestrial herb. Clearings in riverine forest,
woodland. 1250–1850 m.
Distribution: MP, SL—Gereau 4052
Eragrostis macilenta (A. Rich.) Steud.; Clayton,
F.T.E.A. (Gramineae 2): 235. 1974.
Terrestrial herb. Riverine forest, edges of cultivation. 1500–1700 m.
Distribution: MP, SL—Gereau 3959
Eragrostis mollior Pilg. ex R. E. Fr.; Clayton,
F.T.E.A. (Gramineae 2): 202. 1974.
Terrestrial herb. Miombo woodland. 1530 m.
Distribution: SL—Gereau 4383
Eragrostis racemosa (Thunb.) Steud.; Clayton,
F.T.E.A. (Gramineae 2): 230. 1974.
Terrestrial herb. Montane grassland, roadsides
in villages. 1360–2700 m.
Distribution: KP, SL—Suleiman 164
Eragrostis schweinfurthii Chiov. var. schweinfurthii; Phillips, 1995: 117.
Terrestrial herb. Montane grassland, edges of
cultivation, road beds. 1940–2590 m.
Distribution: KP, MR, SL—Gereau 4769
Eragrostis setulifera Pilg.; Clayton, F.T.E.A. (Gramineae 2): 205. 1974.
Terrestrial herb. Disturbed cultivated areas.
1250 m.
Distribution: MP—Kayombo 2465
Eragrostis tenuifolia (A. Rich.) Hochst. ex Steud.;
Clayton, F.T.E.A. (Gramineae 2): 238. 1974.
Terrestrial herb. Montane grassland, abandoned cultivation. 1250–2150 m.
Distribution: MP, SL—Suleiman 159
Eulalia villosa Nees; Clayton & Renvoize, F.T.E.A.
(Gramineae 3): 713. 1982.
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Gereau et al.
Terrestrial herb. Montane grassland. 2140–
2150 m.
Distribution: SL—Suleiman 152
Exotheca abyssinica (Hochst. ex A. Rich.)
Andersson; Clayton & Renvoize, F.T.E.A. (Gramineae 3): 821. 1982.
Terrestrial herb. Montane grassland, openings in montane forest, roadsides in cultivation.
2200–2860 m.
Distribution: KP, MR, RM—Abeid 2859
Festuca abyssinica Hochst. ex A. Rich. var. abyssinica; Clayton, F.T.E.A. (Gramineae 1): 60.
1970.
Terrestrial herb. Montane grassland. 2560–
2840 m.
Distribution: KP—Abeid 2852
Festuca africana (Hack.) Clayton; Clayton, 1985:
727.
Terrestrial herb. Montane forest. 1700–2130
m.
Distribution: MR, RM—Abeid 2931
Festuca caprina Nees; Clayton, F.T.E.A. (Gramineae 1): 62. 1970.
Terrestrial herb. Montane grassland. 2560–
2660 m.
Distribution: KP, MP, NL—Abeid 2803
Festuca engleri Pilg.; Phillips, 1995: 27.
Terrestrial herb. Cypress plantation. 2100 m.
Distribution: MP, RM—Geilinger 2337
Helictotrichon elongatum (Hochst. ex A. Rich.) C.
E. Hubb.; Clayton, F.T.E.A. (Gramineae 1): 89.
1970.
Terrestrial herb. Montane grassland. 1940 m.
Distribution: SL—Gereau 4764
Helictotrichon milanjianum (Rendle) C. E. Hubb.;
Clayton, F.T.E.A. (Gramineae 1): 88. 1970.
Terrestrial herb. Clearings in montane forest.
2360 m.
Distribution: NL—Gereau 4126
Helictotrichon umbrosum (Hochst. ex Steud.) C.
E. Hubb.; Clayton, F.T.E.A. (Gramineae 1):
88. 1970.
Terrestrial herb. Montane grassland. 2570 m.
Distribution: KP—Gereau 3108
Hyparrhenia anamesa Clayton; Clayton & Ren-
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Floristic Checklist
voize, F.T.E.A. (Gramineae 3): 800. 1982.
Terrestrial herb. Montane grassland. 1940 m.
Distribution: SL—Gereau 4753
Hyparrhenia cymbaria (L.) Stapf; Clayton & Renvoize, F.T.E.A. (Gramineae 3): 804. 1982.
Terrestrial herb. Montane grassland, openings
in montane forest, cultivated areas. 1250–2200 m.
Distribution: MP, MR, SL—Kayombo 2341
Hyparrhenia dregeana (Nees) Stapf ex Stent;
Clayton & Renvoize, F.T.E.A. (Gramineae 3):
809. 1982.
Terrestrial herb. Montane grassland near forest
edge. 2560 m.
Distribution: KP—Abeid 2872
Hyparrhenia filipendula (Hochst.) Stapf; Clayton
& Renvoize, F.T.E.A. (Gramineae 3): 803.
1982.
Terrestrial herb. Riverine forest edges. 1250–
1500 m.
Distribution: MP, SL—Gereau 4412
Hyparrhenia nyassae (Rendle) Stapf; Clayton &
Renvoize, F.T.E.A. (Gramineae 3): 793. 1982.
Terrestrial herb. Riverine forest. 1500 m.
Distribution: SL—Gereau 4807
Hyparrhenia pilgeriana C. E. Hubb.; Clayton &
Renvoize, F.T.E.A. (Gramineae 3): 807. 1982.
Terrestrial herb. Montane shrubland. 1560–
2300 m.
Distribution: MR, RM—Mwasumbi 16301
Hyparrhenia quarrei Robyns; Clayton & Renvoize,
F.T.E.A. (Gramineae 3): 799. 1982.
Terrestrial herb. Montane grassland near forest
edge. 2540 m.
Distribution: KP—Abeid 2865
Hyparrhenia rufa (Nees) Stapf; Clayton & Renvoize, F.T.E.A. (Gramineae 3): 794. 1982.
Terrestrial herb. Riverine forest, miombo
woodland, abandoned cultivation. 1250–1800 m.
Distribution: MP, SL—Gereau 4808
Isachne mauritiana Kunth; Clayton, F.T.E.A.
(Gramineae 2): 434. 1974.
Terrestrial herb. Clearings in montane forest.
1790 m.
Distribution: NL—Gereau 4152
Koeleria capensis Nees; Clayton, F.T.E.A. (Gra-
LakeNyasa_5-18.indd 89
89
mineae 1): 79. 1970.
Terrestrial herb. Montane grassland. 2040–
2800 m.
Distribution: KP, MP, MR, RM—Stolz 2517A
Loudetia simplex (Nees) C. E. Hubb.; Clayton,
F.T.E.A. (Gramineae 2): 418. 1974.
Terrestrial herb. Montane grassland. 2140–
2150 m.
Distribution: SL—Suleiman 163
Melinis minutiflora P. Beauv.; Zizka, 1988: 92.
Terrestrial herb. Montane grassland. 1940–
2530 m.
Distribution: KP, SL—Abeid 2866
Melinis repens (Willd.) Zizka; Zizka, 1988: 55.
Terrestrial herb. Woodland. 1250 m.
Distribution: MP—J. Lovett 4542
Melinis scabrida (K. Schum.) Hack.; Zizka, 1988:
80.
Terrestrial herb. Montane grassland. 2140–
2150 m.
Distribution: SL—Suleiman 165A
Microchloa caffra Nees; Renvoize, F.T.E.A. (Gramineae 2): 316. 1974.
Terrestrial herb. Montane grassland. 2320 m.
Distribution: SL—Gereau 3989
Microchloa kunthii Desv.; Renvoize, F.T.E.A. (Gramineae 2): 314. 1974.
Terrestrial herb. Montane grassland, valley
grassland. 1270–2570 m.
Distribution: KP, MP—Gereau 3479
Oplismenus burmannii (Retz.) P. Beauv.; Clayton &
Renvoize, F.T.E.A. (Gramineae 3): 542. 1982.
Terrestrial herb. Woodland, riverine forest.
1250 m.
Distribution: MP—J. Lovett 4480
Oplismenus compositus (L.) P. Beauv.; Clayton &
Renvoize, F.T.E.A. (Gramineae 3): 542. 1982.
Terrestrial herb. Riverine forest. 1275–1600 m.
Distribution: MP—Kayombo 2363
Oplismenus hirtellus (L.) P. Beauv.; Clayton &
Renvoize, F.T.E.A. (Gramineae 3): 542. 1982.
Terrestrial herb. Montane forest. 1710–2110
m.
Distribution: MR, RM, SL—Gereau 4736
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Oplismenus undulatifolius (Ard.) P. Beauv.; Clayton & Renvoize, F.T.E.A. (Gramineae 3): 544.
1982.
Terrestrial herb. Montane forest.
Distribution: RM—St. Clair-Thompson 1009
Terrestrial herb. Montane grassland, edges of
cultivation. 2230–2470 m.
Distribution: KP, SL—Gereau 3125
Note: New to East Africa, from Mozambique,
Malawi, and Zimbabwe.
Oxytenanthera abyssinica (A. Rich.) Munro; Clayton, F.T.E.A. (Gramineae 1): 11. 1970.
Bamboo. Villages. 2000 m.
Distribution: NL—Wiland 152
Panicum issongense Pilg.; Clayton & Renvoize,
F.T.E.A. (Gramineae 3): 495. 1982.
Terrestrial herb. Montane forest. 2180 m.
Distribution: MR, RM—Mwasumbi 16195
Panicum adenophorum K. Schum.; Clayton &
Renvoize, F.T.E.A. (Gramineae 3): 469. 1982.
Scrambling terrestrial herb. Montane thickets.
1840 m.
Distribution: SL—Gereau 3544
Panicum lukwangulense Pilg.; Clayton & Renvoize, F.T.E.A. (Gramineae 3): 467. 1982.
Terrestrial herb. Montane grassland and thicket
edges. 2290–2740 m.
Distribution: KP, MR, RM—Gereau 3124
Panicum calvum Stapf; Clayton & Renvoize,
F.T.E.A. (Gramineae 3): 493. 1982.
Scrambling terrestrial herb. Montane forest.
2040 m.
Distribution: SL—Gereau 4722
Panicum maximum Jacq.; Clayton & Renvoize,
F.T.E.A. (Gramineae 3): 471. 1982.
Terrestrial herb. Riverine forest, abandoned
cultivation. 1200–1250 m.
Distribution: MP, SL—Gereau 4295
Panicum claytonii Renvoize; Clayton & Renvoize,
F.T.E.A. (Gramineae 3): 467. 1982.
Scrambling terrestrial herb. Montane forest,
forest edges, scree slopes. 1980–2230 m.
Distribution: NL, SL—Gereau 4134
Panicum delicatulum Fig. & De Not.; Clayton &
Renvoize, F.T.E.A. (Gramineae 3): 491. 1982.
Terrestrial herb. Riverine forest, road banks.
1500–2200 m.
Distribution: MR, SL—Gereau 4529
Panicum ecklonii Nees; Clayton & Renvoize,
F.T.E.A. (Gramineae 3): 466. 1982.
Terrestrial herb. Rocky mountain slopes, edges
of cultivation. 2470–2800 m.
Distribution: KP—Gereau 3123
Panicum gracilicaule Rendle; Clayton & Renvoize,
F.T.E.A. (Gramineae 3): 474. 1982.
Terrestrial herb. Woodland. 1350 m.
Distribution: MP—J. Lovett 4554
Panicum haplocaulos Pilg.; Clayton & Renvoize,
F.T.E.A. (Gramineae 3): 487. 1982.
Terrestrial herb. Seasonally waterlogged grassland. 1250 m.
Distribution: MP—J. Lovett 4224
Panicum inaequilatum Stapf & C. E. Hubb.; Renvoize, Fl. Zamb. 10(3): 38. 1989.
LakeNyasa_5-18.indd 90
Panicum mitopus K. Schum.; Clayton & Renvoize,
F.T.E.A. (Gramineae 3): 498. 1982.
Terrestrial herb. Riverbanks in montane forest.
1880 m.
Distribution: NL—Gereau 4225
Panicum trichocladum Hack. ex K. Schum.; Clayton & Renvoize, F.T.E.A. (Gramineae 3): 473.
1982.
Terrestrial herb. Riverine forest. 1600 m.
Distribution: MP—P. Lovett 225
Paspalum scrobiculatum L.; Clayton & Renvoize,
F.T.E.A. (Gramineae 3): 610. 1982.
Terrestrial herb. Montane grassland, roadsides
in miombo woodland, abandoned cultivation.
1250–2150 m.
Distribution: MP, SL—Suleiman 160
Pennisetum clandestinum Hochst. ex Chiov.; Clayton & Renvoize, F.T.E.A. (Gramineae 3): 675.
1982.
Terrestrial herb. Roadsides in montane grassland. 2600 m.
Distribution: KP—Gereau 3205
Pennisetum macrourum Trin.; Clayton & Renvoize,
F.T.E.A. (Gramineae 3): 689. 1982.
Terrestrial herb. Riverine forest, montane forest
edges. 1500–2360 m.
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Distribution: MR, SL—Gereau 4805
91
Distribution: KP, MP—Gereau 2845
Pennisetum purpureum Schumach.; Clayton &
Renvoize, F.T.E.A. (Gramineae 3): 677. 1982.
Terrestrial herb. Grassland, cultivated and
abandoned fields, riverine forest. 1150–1400 m.
Distribution: MP—Kayombo 2342
Rottboellia cochinchinensis (Lour.) Clayton; Clayton & Renvoize, F.T.E.A. (Gramineae 3): 853.
1982.
Terrestrial herb. Woodland. 1300–1400 m.
Distribution: MP—Kayombo 1009
Pennisetum sphacelatum (Nees) T. Durand &
Schinz; Clayton & Renvoize, F.T.E.A. (Gramineae 3): 689. 1982.
Terrestrial herb. Montane grassland near forest
edge. 2530 m.
Distribution: KP—Abeid 2869
Sacciolepis transbarbata Stapf; Clayton & Renvoize, F.T.E.A. (Gramineae 3): 456. 1982.
Terrestrial herb. Miombo woodland. 1400–
1790 m.
Distribution: MP, SL—Gereau 3874
Pennisetum thunbergii Kunth; Clayton & Renvoize, F.T.E.A. (Gramineae 3): 687. 1982.
Terrestrial herb. Montane grassland. 2080–
2140 m.
Distribution: SL—Suleiman 109
Pennisetum unisetum (Nees) Benth.; Clayton &
Renvoize, F.T.E.A. (Gramineae 3): 681. 1982.
Terrestrial herb. Woodland, riverine forest.
1250 m.
Distribution: MP—J. Lovett 4541
Pentaschistis natalensis Stapf; Clayton, F.T.E.A.
(Gramineae 1): 125. 1970.
Terrestrial herb. Montane grassland and heathland. 2100–2880 m.
Distribution: KP, MP, MR, RM—Abeid 3001
Phragmites mauritianus Kunth; Clayton, F.T.E.A.
(Gramineae 1): 120. 1970.
Terrestrial herb. Wet meadows in miombo
woodland, riverine forest. 840–1980 m.
Distribution: MP, SL—P. Lovett 276
Poa schimperiana Hochst. ex A. Rich.; Clayton,
F.T.E.A. (Gramineae 1): 48. 1970.
Terrestrial herb. Montane grassland. 2570 m.
Distribution: KP, MR—Gereau 3085
Poecilostachys oplismenoides (Hack.) Clayton;
Clayton, Fl. Zamb. 10(3): 3. 1989.
Terrestrial herb. Riverine forest, montane
forest. 1560–1780 m.
Distribution: RM—Mwasumbi 16299
Polypogon schimperianus (Hochst. ex Steud.)
Cope; Phillips, 1995: 45.
Terrestrial herb. Riverine forest, stream banks
in montane grassland. 1500–2590 m.
LakeNyasa_5-18.indd 91
Sacciolepis typhura (Stapf) Stapf; Clayton & Renvoize, F.T.E.A. (Gramineae 3): 460. 1982.
Terrestrial herb. Stream banks in montane
grassland. 2560–2570 m.
Distribution: KP—Gereau 3169
Setaria homonyma (Steud.) Chiov.; Clayton &
Renvoize, F.T.E.A. (Gramineae 3): 536. 1982.
Terrestrial herb. Riverine forest, grassland.
1200–1350 m.
Distribution: MP—J. Lovett 4645
Setaria plicatilis (Hochst.) Hack. ex Engl.; Clayton & Renvoize, F.T.E.A. (Gramineae 3): 538.
1982.
Terrestrial herb. Montane forest. 1750 m.
Distribution: RM—Abeid 2899
Setaria pumila (Poir.) Roem. & Schult.; Clayton &
Renvoize, F.T.E.A. (Gramineae 3): 530. 1982.
Terrestrial herb. Paths in montane forest, roadsides in miombo woodland, abandoned cultivation.
1200–2110 m.
Distribution: MP, MR, SL—Gereau 4564
Setaria sphacelata (Schumach.) M. B. Moss ex
Stapf & C. E. Hubb.; Clayton & Renvoize,
F.T.E.A. (Gramineae 3): 527. 1982.
Terrestrial herb. Woodland, seasonally inundated grassland, fallow fields. 1270–2300 m.
Distribution: MP, MR—Gereau 3482
Sinarundinaria alpina (K. Schum. ex Engl.) C. S.
Chao & Renvoize; Lovett et al., 2006: 289.
Bamboo. Montane forest edges, heathland.
2000–2730 m.
Distribution: MR, NL, RM—Gereau 4535
Sorghastrum bipennatum (Hack.) Pilg.; Clayton &
Renvoize, F.T.E.A. (Gramineae 3): 731. 1982.
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92
Gereau et al.
Terrestrial herb. Grassland, miombo woodland.
1250–1800 m.
Distribution: MP—P. Lovett 116
Sporobolus africanus (Poir.) Robyns & Tournay;
Clayton, F.T.E.A. (Gramineae 2): 375. 1974.
Terrestrial herb. Montane grassland, clearings
in montane forest, edges of cultivation. 1900–2500
m.
Distribution: KP, NL, SL—Gereau 4770
Sporobolus myrianthus Benth.; Clayton, F.T.E.A.
(Gramineae 2): 383. 1974.
Terrestrial herb. Riverine forest, woodland,
abandoned cultivation. 1200–1600 m.
Distribution: MP—Gereau 3491
Sporobolus natalensis (Steud.) T. Durand &
Schinz; Clayton, F.T.E.A. (Gramineae 2): 374.
1974.
Terrestrial herb. Montane grassland. 2080–
2140 m.
Distribution: SL—Suleiman 110
Sporobolus pyramidalis P. Beauv.; Clayton,
F.T.E.A. (Gramineae 2): 373. 1974.
Terrestrial herb. Seasonal pond margins in
montane grassland, abandoned cultivation. 1250–
1840 m.
Distribution: MP, SL—Gereau 3912
Sporobolus subtilis Kunth; Clayton, F.T.E.A. (Gramineae 2): 386. 1974.
Terrestrial herb. Montane grassland. 2100 m.
Distribution: RM—Stolz 1073
Themeda triandra Forssk.; Clayton & Renvoize,
F.T.E.A. (Gramineae 3): 829. 1982.
Terrestrial herb. Montane grassland, miombo
woodland, edges of cultivation. 1260–2800 m.
Distribution: KP, SL—Suleiman 153
Trachypogon spicatus (L. f.) Kuntze; Clayton &
Renvoize, F.T.E.A. (Gramineae 3): 709. 1982.
Terrestrial herb. Montane grassland. 2320 m.
Distribution: SL—Gereau 3994
Trichopteryx elegantula (Hook. f.) Stapf; Clayton,
F.T.E.A. (Gramineae 2): 410. 1974.
Terrestrial herb. Rocky woodland. 1300–1600
m.
Distribution: MP—J. Lovett 4380
Trichopteryx
LakeNyasa_5-18.indd 92
marungensis
Chiov.;
Clayton,
F.T.E.A. (Gramineae 2): 411. 1974.
Terrestrial herb. Montane grassland. 1940 m.
Distribution: SL—Gereau 4754
Tripogon major Hook. f.; Phillips, F.T.E.A. (Gramineae 2): 291. 1974.
Terrestrial herb. Rocky woodland. 2100 m.
Distribution: SL—Gereau 4019
Tristachya bequaertii De Wild.; Clayton, F.T.E.A.
(Gramineae 2): 428. 1974.
Terrestrial herb. Woodland. 1200 m.
Distribution: MP—J. Lovett 3940
Urochloa trichopus (Hochst.) Stapf; Clayton &
Renvoize, F.T.E.A. (Gramineae 3): 604. 1982.
Terrestrial herb. Roadsides. 474–490 m.
Distribution: SL—Gereau 3799
Zonotriche inamoena (K. Schum.) Clayton; Clayton, F.T.E.A. (Gramineae 2): 429. 1974.
Terrestrial herb. Miombo woodland. 1260–
1530 m.
Distribution: SL—Gereau 4385
POLYGALACEAE
3 genera and 5 species
Muraltia flanaganii Bolus; Paiva, F.T.E.A. (Polygalaceae): 8. 2007.
Suffrutex. Rocky mountain ridges. 2800 m.
Distribution: KP, MR—Goldblatt 8191
Polygala exelliana Troupin; Paiva, F.T.E.A. (Polygalaceae): 41. 2007.
Shrub. Montane forest edges, shrubland.
2110–2350 m.
Distribution: MR, RM—Gereau 4549
Polygala macrostigma Chodat; Paiva, F.T.E.A.
(Polygalaceae): 40. 2007.
Terrestrial herb. Woodland, montane grassland. 1400–1940 m.
Distribution: MP, SL—Gereau 4685
Polygala virgata Thunb. var. decora (Sond.) Harv.;
Paiva, F.T.E.A. (Polygalaceae): 56. 2007.
Shrub. Bamboo forest.
Distribution: KP, MP—Brenan 8205
Securidaca longipedunculata Fresen.; Paiva,
F.T.E.A. (Polygalaceae): 6. 2007.
Shrub or tree. Miombo woodland. 474–1475
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Lake Nyasa Climatic Region
Floristic Checklist
m.
Distribution: MP, NL—Kayombo 2430
93
1300–2580 m.
Distribution: KP, MP, RM—Abeid 2840
POLYGONACEAE
PRIMULACEAE
4 genera and 8 species
2 genera and 6 species
Antigonon leptopus Hook. & Arn.; Graham,
F.T.E.A. (Polygonaceae): 38. 1958.
Lianescent shrub. Lakeshore forest edges.
474–490 m.
Distribution: SL—Gereau 3795
Anagallis brevipes P. Taylor; Taylor, F.T.E.A.
(Primulaceae): 13. 1958.
Terrestrial herb. Wet soil in montane grassland.
2250 m.
Distribution: RM—Greenway 8278 (type: EA,
K)
Fagopyrum snowdenii (Hutch. & Dandy) S. P.
Hong; Hong, 1988: 295.
Terrestrial herb. Riverbanks in montane grassland. 2560–2570 m.
Distribution: KP—Gereau 3164
Persicaria decipiens (R. Br.) K. L. Wilson; Nogueira
et al., Fl. Zamb. 9(3): 9. 2006.
Terrestrial or emergent aquatic herb. Riverine
forest. 1210–1500 m.
Distribution: SL—Gereau 4269
Persicaria madagascariensis (Meisn.) S. Ortiz
& Paiva; Nogueira et al., Fl. Zamb. 9(3): 12.
2006.
Terrestrial herb. Damp soil in woodland. 1300
m.
Distribution: MP—J. Lovett 3446
Anagallis djalonis A. Chev.; Taylor, F.T.E.A.
(Primulaceae): 17. 1958.
Terrestrial herb. Roadsides in montane grassland, riverine forest. 1500–1940 m.
Distribution: RM, SL—Gereau 4669
Anagallis hexamera P. Taylor; Taylor, F.T.E.A.
(Primulaceae): 15. 1958.
Terrestrial or aquatic herb. Swamp. 2100 m.
Distribution: MP—Richards 17024
Anagallis kingaensis Engl.; Taylor, F.T.E.A. (Primulaceae): 11. 1958.
Terrestrial herb. Stream banks in montane
grassland. 2100–2700 m.
Distribution: KP, RM—Goldblatt 8186
Persicaria nepalensis (Meisn.) H. Gross; Nogueira
et al., Fl. Zamb. 9(3): 5. 2006.
Terrestrial herb. Edges of pyrethrum fields.
2200 m.
Distribution: MR—Gereau 4513
Ardisiandra primuloides Knuth; Tayor, F.T.E.A.
(Primulaceae): 2. 1958.
Terrestrial herb. Shaded rocky ground. 1000–
2250 m.
Distribution: RM—Stolz 924 (type: K, UPS),
Geilinger 2365
Rumex abyssinicus Jacq.; Graham, F.T.E.A.
(Polygonaceae): 7. 1958.
Terrestrial herb. Montane forest and grassland,
seasonal marshland. 1770–2840 m.
Distribution: KP, RM, SL—Abeid 2876
Ardisiandra wettsteinii R. Wagner; Taylor, F.T.E.A.
(Primulaceae): 4. 1958.
Terrestrial herb. Montane forest, montane
grassland and shrubland. 2120–2740 m.
Distribution: KP, MR, RM, SL—Gereau 4343
Rumex bequaertii De Wild.; Graham, F.T.E.A.
(Polygonaceae): 8. 1958.
Terrestrial herb. Rocky montane grassland.
2960 m.
Distribution: KP, MR—St. Clair-Thompson
820
Rumex usambarensis (Dammer) Dammer; Graham, F.T.E.A. (Polygonaceae): 7. 1958.
Terrestrial herb. Woodland, montane grassland.
LakeNyasa_5-18.indd 93
PROTEACEAE
2 genera and 9 species
Faurea rochetiana (A. Rich.) Chiov. ex Pic. Serm.;
Marner, F.T.E.A. (Proteaceae): 3. 1993.
Tree or shrub. Woodland, riverine forest.
1200–2500 m.
Distribution: KP, MP, SL—Kayombo 2399
Faurea wentzeliana Engl.; Marner, F.T.E.A. (Pro-
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94
teaceae): 6. 1993.
Tree. Woodland, montane grassland. 1520–
1590 m.
Distribution: NL, RM—Bracebridge 23
Protea gaguedi J. F. Gmel.; Brummitt, F.T.E.A.
(Proteaceae): 17. 1993.
Tree or shrub. Woodland, montane grassland.
1300–2140 m.
Distribution: MP, SL—Kayombo 2402
Protea heckmanniana Engl. subsp. heckmanniana; Brummitt, F.T.E.A. (Proteaceae): 25.
1993.
Suffrutex. Montane grassland. 2120–2700 m.
Distribution: MP, SL—Gereau 4324
Protea humifusa Beard; Brummitt, F.T.E.A. (Proteaceae): 26. 1993.
Suffrutex. Montane grassland, rocky ridges.
2010–2800 m.
Distribution: KP, MP, RM—Napper 1173 (type:
EA, K), Taylor 8523
Protea linearifolia Engl.; Brummitt, F.T.E.A. (Proteaceae): 26. 1993.
Suffrutex. Rocky mountain slopes. 2600 m.
Distribution: KP—Brummitt 18136
Protea rupestris R. E. Fr.; Brummitt, F.T.E.A.
(Proteaceae): 17. 1993.
Tree. Miombo woodland. 1580 m.
Distribution: SL—Gereau 4809
Protea welwitschii Engl.; Brummitt, F.T.E.A. (Proteaceae): 19. 1993.
Shrub. Montane grassland, miombo woodland,
secondary shrubland. 1760–2230 m.
Distribution: NL, SL—Gereau 3906
Protea wentzeliana Engl.; Brummitt, F.T.E.A.
(Proteaceae): 22. 1993.
Shrub. Tree clumps in montane grassland.
2360 m.
Distribution: MR—Gereau 3738
Gereau et al.
Distribution: MP, SL—Gereau 4363
Clematis hirsuta Guill. & Perr.; Milne-Redhead &
Turrill, F.T.E.A. (Ranunculaceae): 6. 1952.
Liana, lianescent shrub, or herbaceous climber.
Woodland, riverine forest, montane forest, bamboo
thicket. 840–2090 m.
Distribution: MP, MR, NL, RM, SL—Gereau
4600
Clematis simensis Fresen.; Milne-Redhead & Turrill, F.T.E.A. (Ranunculaceae): 2. 1952.
Herbaceous climber. Montane forest and grassland. 2100–2400 m.
Distribution: MP, MR—Mwasumbi 16316
Clematis uhehensis Engl.; Brummitt, 2000: 105.
Terrestrial herb. Montane grassland, rocky
hills, woodland. 1200–2800 m.
Distribution: KP, MP, MR, RM, SL—Suleiman
16
Clematis villosa DC. subsp. oliveri (Hutch.) Brummitt; Brummitt, 2000: 104.
Woody-based terrestrial herb. Montane grassland, riverine forest. 1100–1940 m.
Distribution: MP, RM, SL—Gereau 4773
Clematis welwitschii Hiern ex Kuntze; MilneRedhead & Turrill, F.T.E.A. (Ranunculaceae):
3. 1952.
Herbaceous climber. Riverine forest. 1210 m.
Distribution: SL—Gereau 4288
Delphinium leroyi Franch. ex Huth; Milne-Redhead & Turrill, F.T.E.A. (Ranunculaceae): 20.
1952.
Terrestrial herb. Montane grassland. 1940–
2590 m.
Distribution: KP, MP, RM, SL—Suleiman 17
4 genera and 13 species
Ranunculus multifidus Forssk.; Milne-Redhead &
Turrill, F.T.E.A. (Ranunculaceae): 19. 1952.
Terrestrial and emergent aquatic herb.
Riverbanks and ponds in montane grassland,
streambeds in montane forest, woodland, riverine
forest. 1400–2550 m.
Distribution: KP, MP, MR, RM, SL—Gereau
4625
Clematis chrysocarpa Welw. ex Oliv. subsp. bijuga
Brummitt; Brummitt, 2000: 102.
Terrestrial herb. Miombo woodland. 1400–
1530 m.
Ranunculus oreophytus Delile; Milne-Redhead &
Turrill, F.T.E.A. (Ranunculaceae): 14. 1952.
Terrestrial herb. Stream banks in montane
grassland. 2500 m.
RANUNCULACEAE
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Distribution: KP—Goldblatt 8187
Ranunculus raeae Exell; Milne-Redhead & Turrill,
F.T.E.A. (Ranunculaceae): 16. 1952.
Terrestrial herb. Montane grassland. 2400–
2600 m.
Distribution: KP—Suleiman 47
Ranunculus volkensii Engl.; Milne-Redhead &
Turrill, F.T.E.A. (Ranunculaceae): 14. 1952.
Terrestrial or aquatic herb. Stream banks and
swamps in montane grassland. 2500–2550 m.
Distribution: KP—Goldblatt 8248
Thalictrum rhynchocarpum Quart.-Dill. & A.
Rich.; Milne-Redhead & Turrill, F.T.E.A.
(Ranunculaceae): 11. 1952.
Terrestrial herb. Montane grassland, riverbank
thickets, montane forest, scree slopes, lava flows.
1980–2560 m.
Distribution: KP, MR, RM, SL—Gereau 3720
Thalictrum zernyi Ulbr.; Milne-Redhead & Turrill,
F.T.E.A. (Ranunculaceae): 9. 1952.
Terrestrial herb. Riverbanks in montane forest
and grassland, forest edges in montane grassland.
2070–2590 m.
Distribution: KP, NL, SL—Gereau 4136
RHAMNACEAE
4 genera and 6 species
Phylica emirnensis (Tul.) Pillans; Johnston,
F.T.E.A. (Rhamnaceae): 9. 1972.
Suffrutex. Rocky mountain ridges. 2600–2800
m.
Distribution: KP—Goldblatt 8194
Rhamnus prinoides L’Hér.; Johnston, F.T.E.A.
(Rhamnaceae): 18. 1972.
Shrub or tree. Montane forest, bamboo forest.
1590–2400 m.
Distribution: NL, RM—Abeid 2812
Rhamnus staddo A. Rich.; Johnston, F.T.E.A.
(Rhamnaceae): 18. 1972.
Shrub. Tree clumps in montane grassland,
bamboo forest. 2360–2630 m.
Distribution: KP, MR—Gereau 3719
Scutia myrtina (Burm. f.) Kurz; Johnston, F.T.E.A.
(Rhamnaceae): 21. 1972.
Liana. Disturbed montane forest.
LakeNyasa_5-18.indd 95
95
Distribution: RM—Ndangalasi 959
Ziziphus abyssinica Hochst. ex A. Rich.; Johnston,
F.T.E.A. (Rhamnaceae): 27. 1972.
Tree. Woodland. 1200–1400 m.
Distribution: MP—Kayombo 2433
Ziziphus mucronata Willd. subsp. mucronata;
Johnston, F.T.E.A. (Rhamnaceae): 26. 1972.
Tree. Riverine forest. 840 m.
Distribution: SL—Kibure 1457
RHIZOPHORACEAE
1 genus and 2 species
Cassipourea gummiflua Tul.; Lewis, F.T.E.A. (Rhizophoraceae): 15. 1956.
Tree or shrub. Montane forest. 1750–1965 m.
Distribution: RM—Abeid 2819
Cassipourea malosana (Baker) Alston; Lewis,
F.T.E.A. (Rhizophoraceae): 11. 1956.
Tree. Montane forest. 1885–2320 m.
Distribution: NL, RM—Kibure 1363
ROSACEAE
6 genera and 11 species
Alchemilla kiwuensis Engl.; Graham, F.T.E.A.
(Rosaceae): 14. 1960.
Terrestrial herb. Trails and stream banks in
montane forest, roadsides. 1790–2130 m.
Distribution: MR, NL, RM—Gereau 4163
Cliffortia nitidula R. E. Fr. & T. C. E. Fr.; Graham,
F.T.E.A. (Rosaceae): 42. 1960.
Shrub. Montane grassland, riversides. 2550–
2740 m.
Distribution: KP, RM—Prins 200
Fragaria vesca L.; Graham, F.T.E.A. (Rosaceae):
20. 1960.
Terrestrial herb. Heathland. 2950 m.
Distribution: KP—Richards 6589
Hagenia abyssinica J. F. Gmel.; Graham, F.T.E.A.
(Rosaceae): 43. 1960.
Tree. Montane forest, mostly at edges. 1920–
2230 m.
Distribution: MR, RM, SL—Gereau 3570
Prunus africana (Hook. f.) Kalkman; Lovett et al.
2006: 207.
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96
Tree. Riverine forest, montane forest. 1600–
2700 m.
Distribution: MP, NL, RM, SL—Harder 1507
Rubus apetalus Poir.; Graham, F.T.E.A. (Rosaceae): 39. 1960.
Scrambling shrub. Montane forest. 2080–2300
m.
Distribution: MP, RM, SL—Suleiman 146
Rubus ellipticus Sm.; Graham, F.T.E.A. (Rosaceae): 25. 1960.
Scrambling shrub. Montane forest. 1780–2250
m.
Distribution: RM—Abeid 2919
Rubus kirungensis Engl.; Graham, F.T.E.A. (Rosaceae): 36. 1960.
Scrambling shrub.
Distribution: RM—MacInnes 408
Rubus niveus Thunb.; Graham, F.T.E.A. (Rosaceae): 40. 1960.
Scrambling shrub. Edges of cultivation. 1250
m.
Distribution; MP—J. Lovett 3843
Rubus pinnatus Willd.; Graham, F.T.E.A. (Rosaceae): 37. 1960.
Scrambling shrub. Montane grassland, riverine
forest, disturbed montane forest. 1780–1940 m.
Distribution: MP, RM, SL—J. Lovett 4876
Rubus porotoensis R. A. Graham; Graham, F.T.E.A.
(Rosaceae): 35. 1960.
Scrambling shrub. Montane forest edges.
2100–2340 m.
Distribution: MR—Richards 9700 (type: K),
Michelmore 965
Gereau et al.
Anthospermum herbaceum L. f.; Verdcourt,
F.T.E.A. (Rubiaceae 1): 325. 1976.
Terrestrial herb. Montane grassland.
Distribution: RM—Ndangalasi 994
Anthospermum rosmarinus K. Schum.; Verdcourt,
F.T.E.A. (Rubiaceae 1): 329. 1976.
Shrub. Montane grassland. 1940–2150 m.
Distribution: NL, SL—Gereau 4660A
Anthospermum usambarense K. Schum.; Verdcourt, F.T.E.A. (Rubiaceae 1): 331. 1976.
Shrub. Montane grassland, roadsides. 1940–
2770 m.
Distribution: KP, MP, NL, RM, SL—Abeid
2870
Anthospermum whyteanum Britten; Verdcourt,
F.T.E.A. (Rubiaceae 1): 331. 1976.
Subshrub. Montane grassland. 2440 m.
Distribution: SL—Suleiman 185
Catunaregam spinosa (Thunb.) Tirveng. subsp.
spinosa; Verdcourt, F.T.E.A. (Rubiaceae 2):
499. 1988.
Shrub. Lakeshore forest. 474 m.
Distribution: SL—Gereau 3663
Catunaregam spinosa (Thunb.) Tirveng. subsp.
taylorii (S. Moore) Verdc.; Verdcourt, F.T.E.A.
(Rubiaceae 2): 500. 1988.
Shrub or tree. Woodland. 1250–1400 m.
Distribution: MP—Kayombo 966
Cephalanthus natalensis Oliv.; Ridsdale, 1976:
183.
Liana. Montane forest. 1900–2040 m.
Distribution: NL, SL—Gereau 4733
Note: New to East Africa, from South Africa
and Malawi.
RUBIACEAE
49 genera and 119 species
Afrocanthium parasiebenlistii (Bridson) Lantz vel
sp. aff.; Lantz & Bremer, 2004: 278.
Tree. Montane forest edges, riverine forest.
1320–2230 m.
Distribution: MP, SL—Gereau 3580
Agathisanthemum quadricostatum Bremek.; Verdcourt, F.T.E.A. (Rubiaceae 1): 256. 1976.
Terrestrial herb. Miombo woodland. 1530 m.
Distribution: SL—Gereau 4376
LakeNyasa_5-18.indd 96
Chamaepentas nobilis (S. Moore) Kårehed & B.
Bremer; Kårehed & Bremer, 2007: 1075.
Terrestrial herb. Woodland. 1200–1250 m.
Distribution: MP—J. Lovett 4227
Chassalia discolor K. Schum. subsp. discolor;
Verdcourt, F.T.E.A. (Rubiaceae 1): 127. 1976.
Shrub. Submontane forest. 1630 m.
Distribution: RM—WCS SHCP–142
Chassalia parvifolia K. Schum.; Verdcourt,
F.T.E.A. (Rubiaceae 1): 130. 1976.
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Lake Nyasa Climatic Region
Floristic Checklist
Shrub. Montane forest, riverine forest. 1560–
2260 m.
Distribution: MR, RM, SL—Gereau 4710
Chassalia subochreata (De Wild.) Robyns; Verdcourt, F.T.E.A. (Rubiaceae 1): 131. 1976.
Tree. Montane forest. 1950 m.
Distribution: RM—Bracebridge 88
Coffea canephora Pierre ex A. Froehner; Bridson,
F.T.E.A. (Rubiaceae 2): 710. 1988.
Shrub. Riverine forest. 1300 m.
Distribution: MP—Kayombo 2353
Coffea mufindiensis Hutch. ex Bridson subsp.
mufindiensis; Bridson, F.T.E.A. (Rubiaceae 2):
720. 1988.
Shrub. Montane forest, riverine forest. 2000–
2080 m.
Distribution: MP, MR, SL—Gereau 4735
Cremaspora triflora (Thonn.) K. Schum. subsp.
confluens (K. Schum.) Verdc.; Verdcourt,
F.T.E.A. (Rubiaceae 2): 735. 1988.
Shrub. Riverine forest. 474–910 m.
Distribution: SL—Mwasumbi 16600
Cremaspora triflora (Thonn.) K. Schum. subsp.
triflora; Verdcourt, F.T.E.A. (Rubiaceae 2):
733. 1988.
Shrub. Stream banks in miombo woodland.
980 m.
Distribution: SL—Harder 1529
Crossopteryx febrifuga (Afzel. ex G. Don) Benth.;
Verdcourt, F.T.E.A. (Rubiaceae 2): 457. 1988.
Tree or shrub. Woodland, lakeshore forest.
480–1400 m.
Distribution: MP, SL—Gereau 3646
Diodia sarmentosa Sw.; Verdcourt, F.T.E.A. (Rubiaceae 1): 336. 1976.
Terrestrial herb. Woodland/montane forest
transition. 2140 m.
Distribution: SL—Gereau 4012
Dolichopentas decora (S. Moore) Kårehed & B.
Bremer var. decora Moore; Kårehed & Bremer,
2007: 1076.
Terrestrial herb. Woodland, montane grassland. 1700–2520 m.
Distribution: KP, MP, MR, SL—Gereau 4434
Dolichopentas lindenioides (S. Moore) Kårehed &
LakeNyasa_5-18.indd 97
97
B. Bremer; Kårehed & Bremer, 2007: 1076.
Terrestrial herb. Rocky mountain slopes,
montane grassland. 2400–2820 m.
Distribution: KP, MP, MR, NL, RM—Suleiman
64
Fadogia ancylantha Hiern; Verdcourt, F.T.E.A.
(Rubiaceae 3): 799. 1991.
Suffrutex or shrub. Woodland, abandoned
cultivation. 1200–1720 m.
Distribution: MP, SL—Kayombo 2482
Fadogia elskensii De Wild.; Verdcourt, F.T.E.A.
(Rubiaceae 3): 790. 1991.
Suffrutex. Woodland. 2000 m.
Distribution: SL—Gereau 4066
Fadogia stenophylla Hiern subsp. odorata (K.
Krause) Verdc.; Verdcourt, F.T.E.A. (Rubiaceae
3): 794. 1991.
Terrestrial herb. Montane grassland, woodland.
1200–2250 m.
Distribution: MP, SL—Gereau 5135
Fadogia triphylla Baker var. giorgii (De Wild.)
Verdc.; Verdcourt, F.T.E.A. (Rubiaceae 3): 797.
1991.
Shrub. Miombo woodland. 1500 m.
Distribution: MP—Gereau 6996
Fadogia triphylla Baker var. triphylla; Verdcourt,
F.T.E.A. (Rubiaceae 3): 796. 1991.
Terrestrial herb or suffrutex. Woodland, roadsides in village. 1200–1400 m.
Distribution: MP, SL—Gereau 3567
Galiniera saxifraga (Hochst.) Bridson; Bridson,
F.T.E.A. (Rubiaceae 2): 696. 1988.
Shrub or tree. Montane forest, riverine forest.
1560–2300 m.
Distribution: MR, NL, SL, RM—Suleiman 75
Galium bussei K. Schum. & K. Krause; Verdcourt,
F.T.E.A. (Rubiaceae 1): 399. 1976.
Terrestrial herb. Montane grassland, rocky
mountain ridges, woodland, edges of cultivation.
1250–2700 m.
Distribution: KP, MP, SL—Suleiman 184
Galium chloroionanthum K. Schum.; Verdcourt,
F.T.E.A. (Rubiaceae 1): 388. 1976.
Terrestrial herb, often climbing. Montane forest
edges. 2700 m.
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98
Distribution: MR, RM—Goetze 1162 (type:
BM, K), Aleljung 516
Galium ossirwaense K. Krause; Verdcourt, F.T.E.A.
(Rubiaceae 1): 400. 1976.
Terrestrial herb. Montane grassland. 2260–
2700 m.
Distribution: KP, MR, NL, RM—Suleiman 62
Gereau et al.
Tree or lianescent shrub. Woodland, montane
forest. 1200–1920 m.
Distribution: MP, SL—Kayombo 2478
Ixora narcissodora K. Schum.; Bridson, F.T.E.A.
(Rubiaceae 2): 616. 1988.
Shrub. Riverine forest. 980–1210 m.
Distribution: SL—Gereau 4276
Galium scabrellum K. Schum.; Verdcourt, F.T.E.A.
(Rubiaceae 1): 401. 1976.
Terrestrial herb, often climbing. Montane
grassland, woodland, scree slopes. 1280–2135 m.
Distribution: MP, RM, SL—Kayombo 2371
Ixora scheffleri K. Schum. & K. Krause subsp.
scheffleri; Bridson, F.T.E.A. (Rubiaceae 2):
613. 1988.
Tree or shrub. Montane forest. 1750–2040 m.
Distribution: MR, NL, RM, SL—Abeid 2776
Galium stenophyllum Baker; Verdcourt, F.T.E.A.
(Rubiaceae 1): 397. 1976.
Terrestrial herb. Riverine forest, miombo
woodland. 1300–1800 m.
Distribution: MP—J. Lovett 3591
Keetia gueinzii (Sond.) Bridson; Bridson, F.T.E.A.
(Rubiaceae 3): 911. 1991.
Shrub or liana. Riverine forest, montane forest,
woodland. 1100–2300 m.
Distribution: KP, MP, MR, NL, SL—Mwangoka
659
Galium tanganyikense Ehrend. & Verdc.; Verdcourt, F.T.E.A. (Rubiaceae 1): 397. 1976.
Terrestrial herb. Montane grassland. 2490–
2590 m.
Distribution: KP, NL—Stolz 2282 (type: BM,
K, MO), Taylor 8521
Gardenia subacaulis Stapf & Hutch.; Verdcourt,
F.T.E.A. (Rubiaceae 2): 510. 1988.
Suffrutex. Woodland. 1250 m.
Distribution: MP—J. Lovett 3561
Gardenia ternifolia Schumach. & Thonn. subsp.
jovis-tonantis (Welw.) Verdc.; Verdcourt,
F.T.E.A. (Rubiaceae 2): 509. 1988.
Tree or shrub. Woodland. 1200–1400 m.
Distribution: MP—Kayombo 2434
Hedythyrsus thamnoideus (K. Schum.) Bremek.;
Verdcourt, F.T.E.A. (Rubiaceae 1): 250. 1976.
Shrub or perennial herb. Montane grassland,
rocky mountain ridges. 2150–2800 m.
Distribution: KP, RM, SL—Gereau 5134
Heinsenia diervilleoides K. Schum. subsp. diervilleoides; Verdcourt, F.T.E.A. (Rubiaceae 2):
732. 1988.
Tree. Montane forest. 1780–1980 m.
Distribution: RM—Kibure 1371
Hymenodictyon floribundum (Hochst. & Steud.)
B. L. Rob.; Verdcourt, F.T.E.A. (Rubiaceae 2):
452. 1988.
LakeNyasa_5-18.indd 98
Keetia venosa (Oliv.) Bridson; Bridson, F.T.E.A.
(Rubiaceae 3): 914. 1991.
Lianescent shrub. Riverine forest. 1210 m.
Distribution: SL—Gereau 4286
Kohautia coccinea Royle; Verdcourt, F.T.E.A.
(Rubiaceae 1): 235. 1976.
Terrestrial herb. Woodland, abandoned cultivation. 1250–2000 m.
Distribution: MP—J. Lovett 4605
Kohautia longifolia Klotzsch var. longifolia; Verdcourt, F.T.E.A. (Rubiaceae 1): 233. 1976.
Terrestrial herb. Grassland. 1550 m.
Distribution: RM—Aleljung 352
Mitracarpus hirtus (L.) DC.; Verdcourt, Fl. Zamb.
5(1): 188. 1989.
Terrestrial herb. Disturbed riverine forest.
1150 m.
Distribution: MP—Kayombo 2451
Mitragyna rubrostipulata (K. Schum.) Havil.;
Lovett et al., 2006: 217.
Tree. Montane forest. 1900 m.
Distribution: RM—Kibure 1362
Multidentia crassa (Hiern) Bridson & Verdc. var.
crassa; Bridson, F.T.E.A. (Rubiaceae 3): 847.
1991.
Tree or shrub. Miombo woodland, riverine
forest. 1200–1920 m.
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Lake Nyasa Climatic Region
Floristic Checklist
Distribution: MP, RM—Kayombo 1093
Multidentia fanshawei (Tennant) Bridson vel sp.
aff.; Bridson, F.T.E.A. (Rubiaceae 3): 843.
1991.
Tree. Montane forest. 1860 m.
Distribution: RM—Bracebridge 81
Mussaenda arcuata Lam ex Poir.; Bridson,
F.T.E.A. (Rubiaceae 2): 461. 1988.
Liana. Miombo woodland, secondary shrubland, wooded grassland, riverine forest, montane
forest. 950–1760 m.
Distribution: NL, RM, SL—Gereau 3849
Oldenlandia corymbosa L. var. caespitosa (Benth.)
Verdc.; Verdcourt, F.T.E.A. (Rubiaceae 1): 310.
1976.
Terrestrial herb. Sandy woodland edges. 1920
m.
Distribution: MR—W. H. Lewis 6088
Note: This collection is somewhat intermediate
with O. capensis L. f. var. capensis.
Oldenlandia echinulosa K. Schum. var. echinulosa; Verdcourt, Fl. Zamb. 5(1): 130. 1989.
Terrestrial herb. Cultivated fields. 1700–2000
m.
Distribution: MP, RM—Mwasumbi 16234
Oldenlandia echinulosa K. Schum. var. pellucida
(Hiern) Verdc.; Verdcourt, Fl. Zamb. 5(1):
130. 1989.
Terrestrial herb. Cultivated fields, riverine
forest. 1600–1800 m.
Distribution: MP, MR—P. Lovett 217
Oldenlandia friesiorum Bremek.; Verdcourt,
F.T.E.A. (Rubiaceae 1): 282. 1976.
Terrestrial herb. Montane grassland. 1950–
2360 m.
Distribution: MR, NL, RM—Gereau 3715
Oldenlandia herbacea (L.) Roxb. var. herbacea;
Verdcourt, F.T.E.A. (Rubiaceae 1): 305. 1976.
Terrestrial herb. Riverine bushland. 1600 m.
Distribution: RM—Aleljung 321
Oldenlandia herbacea (L.) Roxb. var. holstii (K.
Schum.) Bremek.; Verdcourt, F.T.E.A. (Rubiaceae 1): 306. 1976.
Terrestrial herb. 1980 m.
Distribution: MP—Leedal 4259
LakeNyasa_5-18.indd 99
99
Oldenlandia rosulata K. Schum.; Verdcourt,
F.T.E.A. (Rubiaceae 1): 290. 1976.
Terrestrial herb. Montane grassland, miombo
woodland. 1800–1940 m.
Distribution: MP, SL—Gereau 4679
Oldenlandia rupicola (Sond.) Kuntze var. rupicola;
Verdcourt, F.T.E.A. (Rubiaceae 1): 284. 1976.
Prostrate herb or subshrub. Montane heathland. 2730 m.
Distribution: RM—Abeid 2989
Otiophora caerulea (Hiern) Bullock; Verdcourt,
F.T.E.A. (Rubiaceae 1): 321. 1976.
Terrestrial herb. Woodland, montane grassland. 1700–2500 m.
Distribution: MP—J. Lovett 4170
Otiophora stolzii (Verdc.) Verdc.; Verdcourt,
F.T.E.A. (Rubiaceae 1): 320. 1976.
Terrestrial herb. Montane grassland. 2080–
2520 m.
Distribution: KP, SL—Stolz 2467 (type: BM,
BOL, BR, EA, K, MO), Suleiman 85
Otiophora villicaulis Mildbr. var. villicaulis;
Verdcourt, F.T.E.A. (Rubiaceae 1): 322. 1976.
Terrestrial herb. Miombo woodland, montane
grassland. 1530–2230 m.
Distribution: SL—Gereau 4431
Oxyanthus speciosus DC. subsp. stenocarpus (K.
Schum.) Bridson; Bridson, F.T.E.A. (Rubiaceae
2): 529. 1988.
Tree or shrub. Montane forest, riverine forest.
1760–2200 m.
Distribution: MP, MR, NL, RM, SL—Gereau
4727
Pauridiantha paucinervis (Hiern) Bremek.; Verdcourt, F.T.E.A. (Rubiaceae 1): 153. 1976.
Shrub. Montane forest. 1560–2320 m.
Distribution: KP, RM, SL—Gereau 4200
Pauridiantha udzungwaensis Ntore & Dessein;
Ntore et al., 2003: 67.
Lianescent shrub or small tree. Montane forest.
1910–2700 m.
Distribution: RM, SL—Gereau 4702
Pavetta cataractarum S. Moore; Bridson, F.T.E.A.
(Rubiaceae 2): 647. 1988.
Shrub. Riverine forest. 1160 m.
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100
Distribution: SL—Gereau 3702
Pavetta comostyla S. Moore subsp. nyassica (Bremek.) Bridson; Bridson, F.T.E.A. (Rubiaceae
2): 656. 1988.
Shrub or tree. Montane forest. 1560–2110 m.
Distribution: MR, RM—Suleiman 73
Pavetta crassipes K. Schum.; Bridson, F.T.E.A.
(Rubiaceae 2): 670. 1988.
Tree or shrub. Woodland. 1200–1410 m.
Distribution: MP, SL—Kayombo 1012
Pavetta gardeniifolia Hochst. ex A. Rich. var.
gardeniifolia; Bridson, F.T.E.A. (Rubiaceae 2):
677. 1988.
Shrub. Riverine forest. 1200–1500 m.
Distribution: MP, SL—Gereau 4399
Pavetta gardeniifolia A. Rich. var. subtomentosa
K. Schum.; Bridson, F.T.E.A. (Rubiaceae 2):
678. 1988.
Shrub. Riverine forest. 1210 m.
Distribution: SL—Gereau 4259
Gereau et al.
Bridson, F.T.E.A. (Rubiaceae 2): 663. 1988.
Shrub. Montane forest. 2000–2320 m.
Distribution: MP, MR, SL—Gereau 4575
Pentanisia monticola (K. Krause) Verdc.; Verdcourt, F.T.E.A. (Rubiaceae 1): 170. 1976.
Terrestrial herb. Miombo woodland, riverine
forest. 1300–1800 m.
Distribution: MP—Kayombo 2400
Pentanisia schweinfurthii Hiern; Verdcourt,
F.T.E.A. (Rubiaceae 1): 168. 1976.
Terrestrial herb. Montane grassland, woodland.
1300–2000 m.
Distribution: MP, SL—Gereau 3867
Pentas purpurea Oliv. subsp. purpurea; Verdcourt,
F.T.E.A. (Rubiaceae 1): 207. 1976.
Terrestrial herb. Woodland, grassland, rocky
ridges. 1700–2660 m.
Distribution: KP, MP, MR, RM, SL—Abeid
2843
Pavetta hymenophylla Bremek.; Bridson, F.T.E.A.
(Rubiaceae 2): 632. 1988.
Tree. Montane forest. 1750–1780 m.
Distribution: RM—Abeid 2770
Phyllopentas schimperiana (Vatke) Kårehed & B.
Bremer; Kårehed & Bremer, 2007: 1076.
Shrub or terrestrial herb. Montane grassland,
montane forest patches. 2080–2525 m.
Distribution: RM, SL—Gereau 3985
Pavetta kyimbilensis Bremek. var. iringensis (Bremek.) Bridson; Bridson, F.T.E.A. (Rubiaceae
2): 667. 1988.
Shrub or tree. Montane forest. 1780–2230 m.
Distribution: MR, RM, SL—Gereau 4586
Phyllopentas schumanniana (K. Krause) Kårehed
& B. Bremer; Kårehed & Bremer, 2007: 1076.
Shrub, sometimes lianescent. Montane forest,
woodland/forest transition. 1900–2260 m.
Distribution: NL, SL—Gereau 4711
Pavetta kyimbilensis Bremek. var. kyimbilensis;
Bridson, F.T.E.A. (Rubiaceae 2): 667. 1988.
Shrub. Montane forest. 2100–2620 m.
Distribution: MR, RM—Stolz 2332 (type: BM,
BR, EA, K, MO, P); Mwasumbi 16215
Phyllopentas ulugurica (Verdc.) Kårehed & B.
Bremer; Kårehed & Bremer, 2007: 1076.
Terrestrial herb. Grassland, montane forest.
2000–2100 m.
Distribution: MP—J. Lovett 3499
Pavetta lasiobractea K. Schum.; Bridson, F.T.E.A.
(Rubiaceae 2): 667. 1988.
Tree or shrub. Montane forest. 1800–2040 m.
Distribution: MR, NL, RM, SL—Harder 1503
Polysphaeria dischistocalyx Brenan; Verdcourt,
F.T.E.A. (Rubiaceae 2): 578. 1988.
Shrub. Riverine forest. 580–1210 m.
Distribution: SL—Gereau 4257
Pavetta schumanniana F. Hoffm. ex K. Schum.;
Bridson, F.T.E.A. (Rubiaceae 2): 672. 1988.
Tree or shrub. Woodland, edges of cultivation.
1200–1530 m.
Distribution: MP, SL—Kayombo 1051
Psychotria capensis (Eckl.) Vatke; Verdcourt, Fl.
Zamb. 5(1): 13. 1989.
Shrub or tree. Riverine forest. 474–1350 m.
Distribution: MP, SL—Gereau 4265
Pavetta subumbellata Bremek. var. subumbellata;
LakeNyasa_5-18.indd 100
Psychotria ealaensis De Wild.; Verdcourt, F.T.E.A.
(Rubiaceae 1): 57. 1976.
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Lake Nyasa Climatic Region
Floristic Checklist
Liana. Montane forest. 1910–2040 m.
Distribution: NL, SL—Gereau 4208
Psychotria eminiana (Kuntze) E. M. A. Petit
var. eminiana; Verdcourt & Bridson, F.T.E.A.
(Rubiaceae 3): 924. 1991.
Shrub or subshrub. Woodland, edges of cultivation. 1160–1600 m.
Distribution: MP, SL—Kayombo 1007
Psychotria goetzei (K. Schum.) E. M. A. Petit;
Verdcourt, F.T.E.A. (Rubiaceae 1): 44. 1976.
Shrub. Montane forest. 2200 m.
Distribution: MR—Gereau 4606
Psychotria heterosticta E. M. A. Petit var. plurinervata E. M. A. Petit; Verdcourt, F.T.E.A.
(Rubiaceae 1): 102. 1976.
Shrub. Montane forest. 1960–1980 m.
Distribution: MR—Mwasumbi 16346
Psychotria kirkii Hiern var. diversinodula Verdc.;
Verdcourt, F.T.E.A. (Rubiaceae 1): 98. 1976.
Subshrub. Montane forest. 1960–1980 m.
Distribution: MR—Mwasumbi 16350
Psychotria kirkii Hiern var. nairobiensis (Bremek.)
Verdc.; Verdcourt, F.T.E.A. (Rubiaceae 1): 94.
1976.
Shrub. Lakeshore forest. 480 m.
Distribution: SL—Gereau 3661
Psychotria kirkii Hiern var. swynnertonii (Bremek.)
Verdc.; Verdcourt, F.T.E.A. (Rubiaceae 1): 96.
1976.
Suffrutex. Riverine forest. 1350 m.
Distribution: MP—J. Lovett 4664
Psychotria linearisepala E. M. A. Petit var. linearisepala; Verdcourt, F.T.E.A. (Rubiaceae 1): 85.
1976.
Shrub. Riverine forest, miombo woodland.
1200–1530 m.
Distribution: MP, SL—Gereau 4263
101
Subshrub. Riverine forest. 980 m.
Distribution: SL—Gereau 5165
Psychotria schliebenii E. M. A. Petit; Verdcourt,
F.T.E.A. (Rubiaceae 1): 80. 1976.
Lianescent shrub. Riverine forest. 1560–1710
m.
Distribution: RM—Mwasumbi 16282
Psychotria zombamontana (Kuntze) E. M. A.
Petit; Verdcourt, Fl. Zamb. 5(1): 16. 1989.
Shrub or tree. Montane forest, bamboo forest,
riverine forest, scree slopes, shrubland. 1580–2600
m.
Distribution: MP, MR, NL, RM, SL—Abeid
2923
Psydrax livida (Hiern) Bridson; Bridson, F.T.E.A.
(Rubiaceae 3): 898. 1991.
Shrub. Riverine forest, woodland. 1200–1210
m.
Distribution: MP, SL—Gereau 4287
Psydrax parviflora (Afzel.) Bridson subsp. rubrocostata (Robyns) Bridson; Bridson, F.T.E.A.
(Rubiaceae 3): 898. 1991.
Tree. Montane forest. 1725–2040 m.
Distribution: RM, SL—Gereau 4706
Psydrax whitei Bridson; Bridson, F.T.E.A. (Rubiaceae 3): 899. 1991.
Tree. Montane forest edges. 2230–2380 m.
Distribution: KP, SL—Gereau 3575
Pygmaeothamnus zeyheri (Sond.) Robyns var. rogersii Robyns; Verdcourt, F.T.E.A. (Rubiaceae
3): 778. 1991.
Suffrutex. Montane grassland. 2230 m.
Distribution: SL—Gereau 3594
Pyrostria lobulata Bridson; Bridson, F.T.E.A.
(Rubiaceae 3): 887. 1991.
Shrub. Riverine forest. 980 m.
Distribution: SL—Harder 1525
Psychotria mahonii C. H. Wright var. puberula (E.
M. A. Petit) Verdc.; Verdcourt, F.T.E.A. (Rubiaceae 1): 60. 1976.
Shrub or tree. Montane forest. 1690–2400 m.
Distribution: MR, RM, SL—Suleiman 77
Rhodopentas bussei (K. Krause) Kårehed & B.
Bremer; Kårehed & Bremer, 2007: 1076.
Shrub or subshrub. Woodland, edges of cultivation, riverine forest. 1160–1250 m.
Distribution: MP, SL—Gereau 3701
Psychotria peduncularis (Salisb.) Steyerm. var.
nyassana (K. Krause) Verdc.; Verdcourt,
F.T.E.A. (Rubiaceae 1): 74. 1976.
Rhodopentas parvifolia (Hiern) Kårehed & B.
Bremer; Kårehed & Bremer, 2007: 1076.
Lianescent shrub. Riverine forest. 1210 m.
LakeNyasa_5-18.indd 101
5/31/12 3:59 PM
102
Distribution: SL—Gereau 4274
Gereau et al.
Distribution: SL—Gereau 4696
Rothmannia fischeri (K. Schum.) Bullock subsp.
fischeri; Bridson, F.T.E.A. (Rubiaceae 2): 513.
1988.
Tree. Riverine forest. 910–1320 m.
Distribution: MP, SL—Gereau 6123
Rytigynia monantha (K. Schum.) Robyns var.
lusakati Verdc.; Verdcourt, F.T.E.A. (Rubiaceae 3): 826. 1991.
Shrub. Riverine forest. 1210 m.
Distribution: SL—Gereau 4246
Rubia cordifolia L. subsp. conotricha (Gand.)
Verdc.; Verdcourt, F.T.E.A. (Rubiaceae 1): 381.
1976.
Herbaceous climber. Montane forest, tree
clumps in montane grassland, riverine forest,
woodland. 1150–2360 m.
Distribution: KP, MP, MR, NL, SL—Kayombo
2443
Rytigynia monantha (K. Schum.) Robyns var.
monantha; Verdcourt, F.T.E.A. (Rubiaceae 3):
826. 1991.
Shrub. Woodland. 1530–1970 m.
Distribution: SL—Gereau 4023
Rutidea fuscescens Hiern subsp. fuscescens; Bridson, F.T.E.A. (Rubiaceae 2): 610. 1988.
Liana. Riverine forest. 980 m.
Distribution: SL—Gereau 5167
Rutidea orientalis Bridson; Bridson, F.T.E.A.
(Rubiaceae 2): 606. 1988.
Lianescent shrub or liana. Montane forest, riverine forest. 1560–1900 m.
Distribution: NL, RM—Gereau 4182
Rytigynia adenodonta (K. Schum.) Robyns var.
adenodonta; Verdcourt, F.T.E.A. (Rubiaceae
3): 819. 1991.
Shrub. Montane forest. 2200–2300 m.
Distribution: MP, MR—Mwasumbi 16541
Rytigynia adenodonta (K. Schum.) Robyns var.
reticulata (Robyns) Verdc.; Verdcourt, F.T.E.A.
(Rubiaceae 3): 819. 1991.
Shrub or tree. Riverine forest. 1780 m.
Distribution: MP, RM—J. Lovett 4873
Rytigynia bugoyensis (K. Krause) Verdc. subsp.
glabriflora Verdc.; Verdcourt, F.T.E.A. (Rubiaceae 3): 835. 1991.
Shrub. Montane forest. 1780 m.
Distribution: RM—Abeid 2766
Rytigynia obscura Robyns; Verdcourt, F.T.E.A.
(Rubiaceae 3): 811. 1991.
Tree or shrub. Montane forest, woodland/forest
transition. 1900–2360 m.
Distribution: MR, NL, SL—Gereau 4336
Rytigynia stolzii Robyns; Verdcourt, F.T.E.A.
(Rubiaceae 3): 812. 1991.
Shrub or tree. Riverine forest, montane forest.
1160–1980 m.
Distribution: MP, RM, SL—Gereau 3685
Rytigynia uhligii (K. Schum. & K. Krause) Verdc.;
Verdcourt, F.T.E.A. (Rubiaceae 3): 818. 1991.
Tree or shrub. Disturbed montane forest,
miombo woodland. 1550–2260 m.
Distribution: MP, NL, RM—Gereau 7004
Spermacoce dibrachiata Oliv.; Verdcourt, F.T.E.A.
(Rubiaceae 1): 370. 1976.
Terrestrial herb. Montane grassland, secondary
shrubland, woodland, abandoned cultivation.
1300–2140 m.
Distribution: MP, NL, SL—Suleiman 122
Spermacoce natalensis Hochst.; Verdcourt,
F.T.E.A. (Rubiaceae 1): 359. 1976.
Terrestrial herb. Montane grassland and shrubland, scree slopes. 1760–2140 m.
Distribution: NL, SL—Gereau 4238
Rytigynia flavida Robyns; Verdcourt, F.T.E.A.
(Rubiaceae 3): 826. 1991.
Tree or shrub. Montane forest. 1860–2260 m.
Distribution: MR, NL—Suleiman 72
Spermacoce princeae (K. Schum.) Verdc.; Verdcourt, F.T.E.A. (Rubiaceae 1): 362. 1976.
Terrestrial herb. Montane forest and grassland.
1790–2140 m.
Distribution: NL, SL—Gereau 4173
Rytigynia cf. flavida Robyns; Verdcourt, F.T.E.A.
(Rubiaceae 3): 826. 1991.
Tree. Montane forest. 2040 m.
Spermacoce pusilla Wall.; Verdcourt, F.T.E.A.
(Rubiaceae 1): 356. 1976.
Terrestrial herb. Woodland. 1250–1800 m.
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Distribution: MP—P. Lovett 242
Spermacoce sphaerostigma (A. Rich.) Oliv.; Verdcourt, F.T.E.A. (Rubiaceae 1): 367. 1976.
Terrestrial herb. Woodland, riverine forest.
1200–1600 m.
Distribution: MP, SL—Gereau 4282
Tarenna pavettoides (Harv.) Sim subsp. affinis (K.
Schum.) Bridson; Bridson, F.T.E.A. (Rubiaceae
2): 589. 1988.
Shrub or tree. Montane forest. 1780 m.
Distribution: RM—Kibure 1422
Temnocalyx nodulosus Robyns; Verdcourt,
F.T.E.A. (Rubiaceae 3): 761. 1991.
Shrub or tree. Bamboo thicket, montane forest
edges. 2590–2850 m.
Distribution: MR, NL, RM—Stolz 2296 (type:
BM, EA, K, MO), Abeid 2996
Tricalysia acocantheroides K. Schum.; Robbrecht,
F.T.E.A. (Rubiaceae 2): 556. 1988.
Shrub. Montane forest. 1900–2110 m.
Distribution: MP, MR, NL—Gereau 4553
Tricalysia longituba De Wild. var. longituba; Robbrecht, 1982: 320.
Shrub. Riverine forest. 980 m.
Distribution: SL—Gereau 5145
Note: New to East Africa, from Zambia and
Democratic Republic of Congo.
Tricalysia ngalaensis Robbr.; Robbrecht, 1979:
277.
Shrub. Riverine forest. 580 m.
Distribution: SL—Gereau 3815
Note: New to East Africa, from Malawi.
103
abandoned cultivation. 830–2000 m.
Distribution: MP, SL—Gereau 4785
Vangueria madagascariensis J. F. Gmel.; Verdcourt, F.T.E.A. (Rubiaceae 3): 849. 1991.
Shrub or tree. Montane forest, miombo woodland. 1450–2110 m.
Distribution: MP, MR—Gereau 4570
Vangueria volkensii K. Schum.; Verdcourt, F.T.E.A.
(Rubiaceae 3): 854. 1991.
Tree. Riverine forest. 2000 m.
Distribution: MP—J. Lovett 4985
RUTACEAE
4 genera and 7 species
Clausena anisata (Willd.) Hook. f. ex Benth.; Kokwaro, F.T.E.A. (Rutaceae): 49. 1982.
Shrub or small tree. Montane forest, riverine
forest, forest clearings, woodland. 1400–2500 m.
Distribution: KP, MP, RM, SL—Abeid 2883
Toddalia asiatica (L.) Lam.; Kokwaro, F.T.E.A.
(Rutaceae): 3. 1982.
Liana or lianescent shrub. Tree clumps in
grassland, montane forest. 1700–2360 m.
Distribution: MP, MR, RM—Abeid 2927
Vepris hanangensis (Kokwaro) Mziray; Lovett et
al., 2006: 236.
Shrub. Riverine forest. 1300–1400 m.
Distribution: MP—J. Lovett 5009
Vepris nobilis (Delile) Mziray; Lovett et al., 2006:
236.
Tree. Riverine forest. 1320 m.
Distribution: MP—Gereau 6118
Tricalysia pallens Hiern; Robbrecht, F.T.E.A.
(Rubiaceae 2): 548. 1988.
Tree. Lower montane forest. 1470 m.
Distribution: NL—Bracebridge 110
Vepris simplicifolia (Engl.) Mziray; Lovett et al.,
2006: 237.
Tree. Montane forest. 1920–2150 m.
Distribution: NL—Bracebridge 58
Tricalysia verdcourtiana Robbr.; Robbrecht,
F.T.E.A. (Rubiaceae 2): 549. 1988.
Shrub or treelet. Montane forest. 1780–1900
m.
Distribution: NL, RM—Abeid 2772
Vepris stolzii I. Verd.; Lovett et al., 2006: 237.
Tree. Montane forest. 1750–1900 m.
Distribution: NL, RM—Gereau 4187
Vangueria infausta Burch.; Verdcourt, F.T.E.A.
(Rubiaceae 3): 851. 1991.
Tree or shrub. Woodland, riverine forest,
LakeNyasa_5-18.indd 103
Zanthoxylum deremense (Engl.) Kokwaro;
Kokwaro, F.T.E.A. (Rutaceae): 44. 1982.
Shrub or tree. Montane forest. 1720–2030 m.
Distribution: NL, RM—Abeid 2882
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104
SANTALACEAE
3 genera and 7 species
Gereau et al.
(Sapindaceae): 82. 1998.
Tree or shrub. Riverine forest. 580–1500 m.
Distribution: SL—Gereau 4393
Osyridicarpos schimperianus (Hochst. ex A. Rich.)
A. DC.; Polhill, F.T.E.A. (Santalaceae): 21.
2005.
Scrambling shrub or liana. Riverine forest,
woodland. 1150–1250 m.
Distribution: MP, SL—Kayombo 2456
Allophylus chaunostachys Gilg; Davies & Verdcourt, F.T.E.A. (Sapindaceae): 84. 1998.
Tree or shrub, sometimes lianescent. Montane
forest, woodland. 1530–2450 m.
Distribution: MR, NL, RM, SL—Suleiman 107
Osyris lanceolata Hochst & Steud.; Polhill,
F.T.E.A. (Santalaceae): 23. 2005.
Shrub or tree. Riverine forest, woodland,
montane thickets. 980–2140 m.
Distribution: KP, MP, SL—Gereau 7005
Allophylus congolanus Gilg; Davies & Verdcourt,
F.T.E.A. (Sapindaceae): 92. 1998.
Shrub, sometimes lianescent. Riverine forest,
miombo woodland. 1100–1530 m.
Distribution: MP, SL—Mwangoka 660
Thesium fimbriatum A. W. Hill; Polhill, F.T.E.A.
(Santalaceae): 14. 2005.
Terrestrial herb. Miombo woodland, montane
grassland. 1400–1940 m.
Distribution: MP, SL—Gereau 4664
Allophylus fulvotomentosus Gilg; Davies & Verdcourt, F.T.E.A. (Sapindaceae): 82. 1998.
Tree or shrub, sometimes lianescent. Woodland. 1100–1260 m.
Distribution: MP—Kayombo 2358
Thesium kilimandscharicum Engl.; Polhill,
F.T.E.A. (Santalaceae): 7. 2005.
Suffrutex. Montane grassland, rocky ridges.
2500–2960 m.
Distribution: KP, RM—Taylor 8522
Allophylus richardsiae Exell; Davies & Verdcourt,
F.T.E.A. (Sapindaceae): 92. 1998.
Tree or shrub. Woodland, riverine forest, abandoned cultivation. 1250–1600 m.
Distribution: MP—J. Lovett 3393
Thesium subaphyllum Engl.; Polhill, F.T.E.A.
(Santalaceae): 12. 2005.
Terrestrial herb. Miombo woodland. 1790 m.
Distribution: SL—Gereau 3899
Allophylus tanzaniensis F. G. Davies; Davies &
Verdcourt, F.T.E.A. (Sapindaceae): 87. 1998.
Shrub. Riverine forest. 1200 m.
Distribution: MP—J. Lovett 4307
Thesium triflorum Thunb. ex L. f.; Polhill, F.T.E.A.
(Santalaceae): 6. 2005.
Straggling shrub. Montane forest. 2040 m.
Distribution: SL—Gereau 4739
Blighia unijugata Baker; Davies & Verdcourt,
F.T.E.A. (Sapindaceae): 25. 1998.
Tree. Lakeshore bushland, riverine forest.
474–840 m.
Distribution: SL—Mwasumbi 16587
Thesium unyikense Engl.; Polhill, F.T.E.A. (Santalaceae): 16. 2005.
Terrestrial herb. Woodland. 1400–1500 m.
Distribution: MP—J. Lovett 3401
Deinbollia borbonica Scheff. f. glabrata Radlk.;
Davies & Verdcourt, F.T.E.A. (Sapindaceae):
71. 1998.
Shrub. Riverine forest. 880 m.
Distribution: SL—Kibure 1452
SAPINDACEAE
5 genera and 12 species
Allophylus abyssinicus (Hochst.) Radlk.; Davies &
Verdcourt, F.T.E.A. (Sapindaceae): 78. 1998.
Tree. Montane forest. 1750–2110 m.
Distribution: MR, RM—Abeid 2909
Allophylus africanus P. Beauv. var. griseotomentosus (Gilg) Verdc.; Davies & Verdcourt, F.T.E.A.
LakeNyasa_5-18.indd 104
Deinbollia kilimandscharica Taub. var. adusta
(Radlk.) Verdc.; Davies & Verdcourt, F.T.E.A.
(Sapindaceae): 71. 1998.
Tree. Montane forest. 1730 m.
Distribution: NL—Bracebridge 55
Dodonaea viscosa Jacq. var. angustifolia (L. f.)
Benth.; Davies & Verdcourt, F.T.E.A. (Sapindaceae): 9. 1998.
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Floristic Checklist
Shrub or tree. Montane thickets, secondary
forest, riverine forest. 1500–2510 m.
Distribution: NL, RM, SL—Suleiman 139
Paullinia pinnata L.; Davies & Verdcourt, F.T.E.A.
(Sapindaceae): 102. 1998.
Liana. Riverine forest, lakeshore bushland,
abandoned cultivation. 474–1400 m.
Distribution: MP, SL—Kayombo 1024
105
Pouteria alnifolia (Baker) Roberty var. alnifolia;
Lovett et al., 2006: 263.
Tree. Disturbed montane forest.
Distribution: RM—Ndangalasi 960
Synsepalum brevipes (Baker) T. D. Penn.; Lovett
et al., 2006: 265.
Tree. Riverine forest. 830–920 m.
Distribution: SL—Kibure 1476
SAPOTACEAE
SCROPHULARIACEAE
5 genera and 9 species
13 genera and 22 species
Chrysophyllum gorungosanum Engl.; Hemsley,
F.T.E.A. (Sapotaceae): 10. 1968.
Alectra sessiliflora (Vahl) Kuntze; Ghazanfar,
F.T.E.A. (Scrophulariaceae): 112. 2008.
Terrestrial herb. Roadsides in cultivation,
riverine forest, woodland. 1250–2200 m.
Distribution: KP, MP, MR—Gereau 4516
Tree. Montane forest. 1840 m.
Distribution: RM—Kibure 1389
Englerophytum magalismontanum (Sond.) T. D.
Penn.; Lovett et al., 2006: 254.
Shrub or small tree. Miombo woodland,
montane forest. 980–1800 m.
Distribution: NL, SL—Harder 1528
Englerophytum natalense (Sond.) T. D. Penn.;
Lovett et al., 2006: 255.
Tree. Riverine forest. 880–1400 m.
Distribution: MP, SL—Kibure 1453
Mimusops kummel Bruce ex A. DC.; Hemsley,
F.T.E.A. (Sapotaceae): 54. 1968.
Tree. Riverine forest. 474–480 m.
Distribution: RM, SL—Mwasumbi 16596
Mimusops obtusifolia Lam.; Lovett et al., 2006:
259.
Shrub. Riverine forest, miombo woodland.
620–980 m.
Distribution: SL—Gereau 5151
Mimusops zeyheri Sond.; Hemsley, F.T.E.A. (Sapotaceae): 56. 1968.
Tree. Riverine forest. 1350 m.
Distribution: MP—J. Lovett 4667
Pouteria adolfi-friedericii (Engl.) A. Meeuse subsp.
australis (J. H. Hemsl.) L. Gaut.; Lovett et al.,
2006: 261.
Tree. Riverine forest, montane forest. 1840–
2200 m.
Distribution: MP, RM—Semsei 1532 (type: EA,
K), Kayombo 1078
LakeNyasa_5-18.indd 105
Buchnera cryptocephala (Baker) Philcox; Philcox,
F.T.E.A. (Scrophulariaceae): 126. 2008.
Terrestrial herb. Woodland, montane grassland. 1400–2300 m.
Distribution: MP, SL—J. Lovett 4159
Buchnera decandollei Govaerts; Govaerts, 1996:
14.
Terrestrial herb. Wet grassland. 1250–1400 m.
Distribution: MP—Kayombo 971
Buchnera lastii Engl. subsp. lastii; Philcox,
F.T.E.A. (Scrophulariaceae): 133. 2008.
Terrestrial herb. 2250 m.
Distribution: MP—J. Lovett 4841
Buchnera quadrifaria Baker; Philcox, F.T.E.A.
(Scrophulariaceae): 129. 2008.
Terrestrial herb. Montane grassland. 1940 m.
Distribution: KP, SL—Gereau 4779
Buchnera rungwensis Engl.; Philcox, F.T.E.A.
(Scrophulariaceae): 129. 2008.
Terrestrial herb. Montane grassland, rocky
ridges. 1940–2960 m.
Distribution: KP, RM, SL—Stolz 2439 (type:
K), Suleiman 36
Buchnera speciosa Skan; Philcox, F.T.E.A.
(Scrophulariaceae): 136. 2008.
Terrestrial herb. Riverine forest. 1500 m.
Distribution: MP, SL—Gereau 4800
Buchnera trilobata Skan vel sp. aff.; Philcox, Fl.
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106
Zamb. 8(2): 102. 1990.
Terrestrial herb. Woodland. 1400–1500 m.
Distribution: MP—J. Lovett 3408
Cycnium adonense E. Mey. ex Benth.; Hepper,
F.T.E.A. (Scrophulariaceae): 168. 2008.
Terrestrial herb. Woodland, scree slopes. 1200–
2000 m.
Distribution: MP, SL—Gereau 3638
Diclis ovata Benth.; Hepper & Ghazanfar, F.T.E.A.
(Scrophulariaceae): 20. 2008.
Terrestrial herb. Abandoned cultivation, open
mountain slopes, riverine forest. 1250–1980 m.
Distribution: MP, SL—Gereau 3642
Halleria lucida L.; Hepper, F.T.E.A. (Scrophulariaceae): 26. 2008.
Shrub or tree. Riverine forest, montane forest.
1780–2500 m.
Distribution: MP, MR, NL, RM, SL—Harder
1477
Hebenstretia angolensis Rolfe; Ghazanfar, F.T.E.A.
(Scrophulariaceae): 197. 2008.
Terrestrial herb or shrub. Woodland, grassland.
1300–2600 m.
Distribution: KP, MP—Suleiman 41
Hedbergia abyssinica (Hochst. ex Benth.) Molau;
Philcox, F.T.E.A. (Scrophulariaceae): 195.
2008.
Terrestrial herb. Montane forest, montane
grassland. 2200–2700 m.
Distribution: KP, MR, NL—Gereau 4604
Jamesbrittenia micrantha (Klotzsch) Hilliard;
Ghazanfar, F.T.E.A. (Scrophulariaceae): 32.
2008.
Terrestrial herb. Abandoned cultivation,
riverine forest. 1250–1350 m.
Distribution: MP—J. Lovett 3779
Lindernia nummulariifolia (D. Don) Wettst.; Philcox, F.T.E.A. (Scrophulariaceae): 80. 2008.
Terrestrial herb. Abandoned cultivation. 1250–
1600 m.
Distribution: MP—J. Lovett 4023
Lindernia rupestris Engl.; Philcox, F.T.E.A.
(Scrophulariaceae): 72. 2008.
Terrestrial herb. Miombo woodland. 1200–
1800 m.
LakeNyasa_5-18.indd 106
Gereau et al.
Distribution: MP—P. Lovett 247
Micrargeria filiformis (Schumach. & Thonn.)
Hutch. & Dalziel; Ghazanfar, F.T.E.A. (Scrophulariaceae): 187. 2008.
Terrestrial herb. Wet grassland. 1250 m.
Distribution: MP—J. Lovett 4570
Sopubia mannii Skan var. linearifolia O. J. Hansen; Philcox, F.T.E.A. (Scrophulariaceae): 178.
2008.
Terrestrial herb. Montane grassland. 1940–
2840 m.
Distribution: KP, MP, SL—Richards 14183
(type: EA, K), Suleiman 168
Sopubia ramosa (Hochst.) Hochst.; Philcox,
F.T.E.A. (Scrophulariaceae): 180. 2008.
Terrestrial herb or suffrutex. Riverine forest,
woodland. 1300–2000 m.
Distribution: MP, SL—Gereau 4801
Striga asiatica (L.) Kuntze; Hepper, F.T.E.A.
(Scrophulariaceae): 148. 2008.
Terrestrial herb. Woodland. 1250 m.
Distribution: MP—J. Lovett 4331
Veronica abyssinica Fresen.; Hepper, F.T.E.A.
(Scrophulariaceae): 102. 2008.
Terrestrial herb. Montane grassland, montane
forest edges. 1900–2640 m.
Distribution: KP, MR, NL, SL—Suleiman 116
Veronica glandulosa Hochst. ex Benth.; Hepper,
F.T.E.A. (Scrophulariaceae): 104. 2008.
Terrestrial herb. Montane grassland. 2570 m.
Distribution: KP—Gereau 3092
SIMAROUBACEAE
1 genus and 1 species
Harrisonia abyssinica Oliv.; Stannard, F.T.E.A.
(Simaroubaceae): 2. 2000.
Shrub. Lakeshore shrubland. 474–490 m.
Distribution: SL—Mwasumbi 16553
SMILACACEAE
1 genus and 2 species
Smilax anceps Willd.; Cowley, F.T.E.A. (Smilacaceae): 2. 1989.
Liana. Montane forest and thickets, riverine
forest, edges of cultivation. 1250–2140 m.
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Lake Nyasa Climatic Region
Floristic Checklist
Distribution: MP, NL, SL—Gereau 4202
Smilax aspera L.; Cowley, F.T.E.A. (Smilacaceae):
2. 1989.
Liana. Montane thickets, forest edges. 1840–
2120 m.
Distribution: SL—Gereau 4315
SOLANACEAE
5 genera and 11 species
Cestrum aurantiacum Lindl.; Gonçalves, Fl.
Zamb. 8(4): 11. 2005.
Shrub. Riverine forest, abandoned cultivation.
1250–1400 m.
Distribution: MP—Kayombo 1028
Discopodium penninervium Hochst.; Gonçalves,
Fl. Zamb. 8(4): 63. 2005.
Shrub. Montane forest. 1720–2350 m
Distribution: MR, RM—Mwasumbi 16225
107
Distribution: MP, MR, NL, RM, SL—Gereau
4515
Solanum incanum L.; Gonçalves, Fl. Zamb. 8(4):
112. 2005.
Shrub or suffrutex. Roadsides, edges of cultivation, woodland, heathland. 474–2700 m.
Distribution: KP, MP, SL—Gereau 3786
Solanum schumannianum Dammer; Gonçalves,
Fl. Zamb. 8(4): 90. 2005.
Shrub. Montane forest. 1900–2370 m.
Distribution: MR, NL, RM, SL—Mwasumbi
16356
Solanum terminale Forssk.; Gonçalves, Fl. Zamb.
8(4): 77. 2005.
Liana or shrub. Montane forest, forest edges.
2260–2500 m.
Distribution: MP, SL—Gereau 3522
STERCULIACEAE
Nicandra physalodes (L.) Gaertn.; Gonçalves, Fl.
Zamb. 8(4): 26. 2005.
Terrestrial herb. Weed of cultivation. 1250 m.
Distribution: MP—Kayombo 2466
Physalis peruviana L.; Gonçalves, Fl. Zamb. 8(4):
47. 2005.
Terrestrial herb. Riverine forest. 1500 m.
Distribution: MP—J. Lovett 4694
Solanum aculeastrum Dunal; Gonçalves, Fl.
Zamb. 8(4): 107. 2005.
Shrub. Montane forest. 1870 m.
Distribution: RM—WCS SHCP 117
Solanum aculeatissimum Jacq.; Gonçalves, Fl.
Zamb. 8(4): 118. 2005.
Shrub or perennial herb. Rocky ridges, riverine
forest. 1850–2600 m.
Distribution: MP, SL—Gereau 3494
5 genera and 7 species
Cola greenwayi Brenan vel sp. aff.; Cheek, F.T.E.A.
(Sterculiaceae): 42. 2007.
Tree. Montane forest, riverine forest. 890–1630
m.
Distribution: RM, SL—Bracebridge 41
Dombeya acutangula Cav.; Cheek, F.T.E.A. (Sterculiaceae): 62. 2007.
Shrub or tree. Riverine forest, roadsides. 1150–
1350 m.
Distribution: MP, NL—J. Lovett 4660
Dombeya burgessiae Gerrard ex Harv.; Cheek,
F.T.E.A. (Sterculiaceae): 64. 2007.
Shrub or tree. Woodland, riverine forest,
montane forest, secondary forest. 1250–2300 m.
Distribution: MP, MR, RM—Abeid 2911
Solanum americanum Mill.; Gonçalves &
Edmonds, Fl. Zamb. 8(4): 81. 2005.
Terrestrial herb. Riverine forest, montane
forest, abandoned cultivation. 1250–2230 m.
Distribution: MP, SL—Gereau 4426
Dombeya torrida (J. F. Gmel.) Bamps subsp. erythroleuca (K. Schum.) Seyani; Cheek, F.T.E.A.
(Sterculiaceae): 67. 2007.
Shrub or tree. Montane forest, montane grassland. 1900–2350 m.
Distribution: MR, RM, SL—Gereau 4765
Solanum anguivi Lam.; Gonçalves, Fl. Zamb.
8(4): 93. 2005.
Shrub. Montane forest, roadsides in cultivation,
grassland, riverine forest. 1210–2300 m.
Leptonychia usambarensis K. Schum.; Dorr,
F.T.E.A. (Sterculiaceae): 102. 2007.
Tree. Montane forest. 1700 m.
Distribution: NL—Bracebridge 64
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Gereau et al.
Sterculia quinqueloba (Garcke) K. Schum.; Cheek,
F.T.E.A. (Sterculiaceae): 12. 2007.
Tree. Woodland. 1250–1300 m.
Distribution: MP—Kayombo 1096
2100–2685 m.
Waltheria indica L.; Dorr, F.T.E.A. (Sterculiaceae):
127. 2007.
Herb or suffrutex. Miombo woodland. 1200–
1250 m.
Distribution: MP—J. Lovett 4067
Terrestrial herb or suffrutex. Woodland. 1400–
1800 m.
TACCACEAE
1 genus and 1 species
Tacca leontopetaloides (L.) Kuntze; Carter,
F.T.E.A. (Taccaceae): 1. 1962.
Terrestrial herb. Woodland, lakeshore forest.
474–1250 m.
Distribution: MP, SL—Gereau 3801
THEACEAE
1 genus and 1 species
Ficalhoa laurifolia Hiern; Verdcourt, F.T.E.A.
(Theaceae): 8. 1962.
Tree. Montane forest, riverine forest. 1640–
2600 m.
Distribution: MR, NL, RM, SL—Gereau 4609
THYMELAEACEAE
4 genera and 9 species
Dais cotinifolia L.; Peterson, F.T.E.A. (Thymelaeaceae): 16. 1978.
Tree. Woodland. 2080 m.
Distribution: SL—Gereau 4016
Gnidia chrysantha (Solms) Gilg; Petersen, F.T.E.A.
(Thymelaeaceae): 26. 1978.
Terrestrial herb or suffrutex. Miombo woodland, grassland. 1450–1700 m.
Distribution: MP, RM—Gereau 6980
Gnidia fastigiata Rendle; Petersen, F.T.E.A.
(Thymelaeaceae): 21. 1978.
Terrestrial herb or suffrutex. Montane grassland, rocky ridges. 2000–2800 m.
Distribution: KP, MP, RM, SL—Gereau 5137
Gnidia glauca (Fresen.) Gilg; Peterson, F.T.E.A.
(Thymelaeaceae): 32. 1978.
Shrub. Montane forest edges, wooded grassland.
LakeNyasa_5-18.indd 108
Distribution: KP, MP, NL—Gereau 4641
Gnidia involucrata Steud. ex A. Rich.; Peterson,
F.T.E.A. (Thymelaeaceae): 25. 1978.
Distribution: MP—J. Lovett 3524
Gnidia kraussiana Meisn.; Peterson, F.T.E.A.
(Thymelaeaceae): 29. 1978.
Terrestrial herb. Montane grassland. 2230 m.
Distribution: SL—Gereau 3602
Gnidia usafuae Gilg; Peterson, F.T.E.A. (Thymelaeaceae): 26. 1978.
Shrub. Montane grassland and forest edges.
Distribution: MP, RM—Semsei 1541
Peddiea polyantha Gilg; Peterson,
(Thymelaeaceae): 11. 1978.
F.T.E.A.
Shrub or tree. Montane forest. 1900–2260 m.
Distribution: MR, NL, RM, SL—Abeid 2846
Synaptolepis alternifolia Oliv.; Peterson, F.T.E.A.
(Thymelaeaceae): 6. 1978.
Liana. 1740 m.
Distribution: RM—WCS SHCP 28
TILIACEAE
4 genera and 16 species
Corchorus tridens L.; Andrews, F.T.E.A. (Tiliaceae): 113. 2001.
Terrestrial herb. Riverine forest. 1350 m.
Distribution: MP—J. Lovett 4657
Grewia bicolor Juss.; Whitehouse, F.T.E.A. (Tiliaceae): 42. 2001.
Tree. Riverine forest. 580 m.
Distribution: SL—Gereau 3822
Grewia flavescens Juss.; Whitehouse, F.T.E.A.
(Tiliaceae): 33. 2001.
Shrub or liana. Woodland, grassland, riverine
forest, abandoned cultivation. 1200–1600 m.
Distribution: MP—Kayombo 2413
Grewia similis K. Schum.; Whitehouse, F.T.E.A.
(Tiliaceae): 13. 2001.
Shrub or small tree. Riverine forest. 880 m.
Distribution: SL—Kibure 1454
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Lake Nyasa Climatic Region
Floristic Checklist
Grewia stolzii Ulbr.; Whitehouse, F.T.E.A. (Tiliaceae): 22. 2001.
Treelet. Opening in montane forest. 1920 m.
Distribution: MR, SL—Gereau 6126
Sparrmannia ricinocarpa (Eckl. & Zeyh.) Kuntze;
Verdcourt, F.T.E.A. (Tiliaceae): 93. 2001.
Straggling terrestrial herb or shrub. Road
banks, montane grassland, montane forest. 2040–
2700 m.
Distribution: KP, MP, MR, NL, SL—Suleiman
66
Triumfetta amuletum Sprague; Cheek, F.T.E.A.
(Tiliaceae): 70. 2001.
Trailing shrub. Grassland, miombo woodland.
1200–1400 m.
Distribution: MP—Kayombo 1015
Triumfetta angolensis Sprague & Hutch.; Wild, Fl.
Zamb. 2: 71. 1963.
Terrestrial herb. Miombo woodland. 1800 m.
Distribution: MP—P. Lovett 20
Note: New to East Africa, from Zambia,
Malawi, Mozambique, Zimbabwe, South Africa,
and Angola.
Triumfetta annua L.; Cheek, F.T.E.A. (Tiliaceae):
79. 2001.
Terrestrial herb. Woodland. 1250–2080 m.
Distribution: MP, MR—Mwasumbi 16191
Triumfetta b rachyceras K. Schum.; Cheek,
F.T.E.A. (Tiliaceae): 89. 2001.
Shrub. Montane forest edges. 1770 m.
Distribution: RM—WCS SHCP 48
Triumfetta pentandra A. Rich.; Cheek, F.T.E.A.
(Tiliaceae): 85. 2001.
Terrestrial herb. Riverine forest. 1500 m.
Distribution: SL—Gereau 4791
Triumfetta pilosa Roth; Cheek, F.T.E.A. (Tiliaceae): 91. 2001.
Shrub, subshrub, herb. Miombo woodland,
riverine forest, disturbed montane forest, montane
grassland. 1530–1940 m.
Distribution: MP, RM, SL—Gereau 4386
Triumfetta rhomboidea Jacq.; Cheek, F.T.E.A.
(Tiliaceae): 83. 2001.
Terrestrial herb, suffrutex, shrub. Montane
forest, riverine forest, woodland, grassland, edges
LakeNyasa_5-18.indd 109
109
of cultivation. 1200–2200 m.
Distribution: MP, MR, SL—Suleiman 90
Triumfetta setulosa Mast.; Cheek, F.T.E.A. (Tiliaceae): 76. 2001.
1963. Terrestrial herb. Riverine forest. 1210 m.
Distribution: MR, SL—Gereau 4255
Triumfetta tomentosa Bojer; Cheek, F.T.E.A. (Tiliaceae): 90. 2001.
Shrub. Woodland, riverine forest. 1200–1600
m.
Distribution: MP—J. Lovett 4419
Triumfetta welwitschii Mast. var. welwitschii;
Cheek, F.T.E.A. (Tiliaceae): 74. 2001.
Terrestrial herb. Montane grassland, woodland.
1250–2250 m.
Distribution: MP—J. Lovett 4870
TURNERACEAE
1 genus and 2 species
Tricliceras lobatum (Urb.) R. Fern.; Fernandes, Fl.
Zamb. 4: 363. 1978.
Terrestrial herb. Roadsides. 474–490 m.
Distribution: SL—Gereau 3797
Tricliceras longipedunculatum (Mast.) R. Fern.
var. longepidunculatum; Fernandes, Fl. Zamb.
4: 359. 1978.
Terrestrial herb. Woodland, riverine forest.
1200–1400 m.
Distribution: MP—J. Lovett 5021
ULMACEAE
2 genera and 3 species
Celtis africana Burm. f.; Polhill, F.T.E.A. (Ulmaceae): 4. 1966.
Tree. Montane forest. 1800 m.
Distribution: NL, RM—Bracebridge 54
Celtis gomphophylla Baker; Lovett et al., 2006:
279.
Tree. Riverine forest. 1320–1480 m.
Distribution: MP—Lovett 4894
Trema orientalis (L.) Blume; Polhill, F.T.E.A.
(Ulmaceae): 10. 1966.
Tree. Miombo woodland, riverine forest,
montane forest, abandoned cultivation. 620–1830
m.
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110
Gereau et al.
9 genera and 13 species
(Urticaceae): 27. 1989.
Terrestrial herb. Montane forest, riverine forest.
1790–1980 m.
Distribution: MR, NL—Gereau 4153
Boehmeria macrophylla Hornem.; Friis, F.T.E.A.
(Urticaceae): 44. 1989.
Shrub. Riverine forest, montane forest. 1210–
2040 m.
Distribution: MP, NL, RM, SL—Gereau 4277
Pilea usambarensis Engl.; Friis, F.T.E.A. (Urticaceae): 36. 1989.
Terrestrial herb. Montane forest, riverine forest.
1790–1850 m.
Distribution: NL, SL—Gereau 4164
Droguetia iners (Forssk.) Schweinf. subsp. iners;
Friis, F.T.E.A. (Urticaceae): 57. 1989.
Straggling terrestrial herb. Tree clumps in
montane grassland, hedge beside cultivated field.
2080–2360 m.
Distribution: MR, NL—Gereau 3749
Pouzolzia mixta Solms; Friis, F.T.E.A. (Urticaceae): 48. 1989.
Shrub. Lakeshore forest and shrubland.
480–495 m.
Distribution: RM, SL—Gereau 3647
Distribution: MP, NL, RM, SL—Harder 1522
URTICACEAE
Elatostema monticola Hook. f.; Friis, F.T.E.A.
(Urticaceae): 41. 1989.
Terrestrial herb. Montane forest. 2460 m.
Distribution: NL, RM—Gereau 4112
Elatostema welwitschii Engl.; Friis, F.T.E.A. (Urticaceae): 39. 1989.
Terrestrial herb. Riverine forest. 1790 m.
Distribution: NL—Gereau 4160
Girardinia diversifolia (Link) Friis; Friis, F.T.E.A.
(Urticaceae): 13. 1989.
Terrestrial herb. Woodland, riverine forest.
1275–1400 m.
Distribution: MP—Kayombo 1004
Laportea alatipes Hook. f.; Friis, F.T.E.A. (Urticaceae): 16. 1989.
Terrestrial herb. Montane forest. 2400–2460
m.
Distribution: MP, NL—Gereau 4122
Laportea interrupta (L.) Chew; Friis, F.T.E.A.
(Urticaceae): 17. 1989.
Terrestrial herb. Abandoned cultivation. 1250
m.
Distribution: MP—J. Lovett 4037
Pilea rivularis Wedd.; Friis, F.T.E.A. (Urticaceae):
29. 1989.
Terrestrial herb. Montane forest. 2120–2400
m.
Distribution: MP, SL—Gereau 4331
Pilea tetraphylla (Steud.) Blume; Friis, F.T.E.A.
LakeNyasa_5-18.indd 110
Procris crenata C. B. Rob.; Friis, F.T.E.A. (Urticaceae): 42. 1989.
Epiphyte. Riverine forest. 1710–1750 m.
Distribution: RM—Mwasumbi 16334
Urera hypselodendron (Hochst. ex A. Rich.)
Wedd.; Friis, F.T.E.A. (Urticaceae): 9. 1989.
Liana. Montane forest and grassland. 1780–
2300 m.
Distribution: KP, MR, RM—Mwasumbi 16307
VALERIANACEAE
1 genus and 1 species
Valeriana capensis Thunb.; Kokwaro, F.T.E.A.
(Valerianaceae): 6. 1968.
Terrestrial herb. Montane grassland. 2360–
2550 m.
Distribution: KP, MR—Gereau 3775
VERBENACEAE
6 genera and 17 species
Clerodendrum eriophyllum Gürke; Verdcourt,
F.T.E.A. (Verbenaceae): 120. 1992.
Shrub. Woodland. 1200–1250 m.
Distribution: MP—J. Lovett 4232
Clerodendrum frutectorum S. Moore; Verdcourt,
F.T.E.A. (Verbenaceae): 103. 1992.
Shrub. Riverine forest, woodland. 1200–1350
m.
Distribution: MP—J. Lovett 4408
Clerodendrum johnstonii Oliv. subsp. johnstonii;
Verdcourt, F.T.E.A. (Verbenaceae): 118. 1992.
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Lake Nyasa Climatic Region
Floristic Checklist
Scrambling shrub. Tree clumps in grassland,
montane forest. 1960–2360 m.
Distribution: MR, SL—Gereau 3751
Lantana camara L.; Verdcourt, F.T.E.A. (Verbenaceae): 39. 1992.
Shrub. Montane forest. 2010 m.
Distribution: RM—WCS SHCP 182
Lantana trifolia L.; Verdcourt, F.T.E.A. (Verbenaceae): 46. 1992.
Shrub. Miombo woodland. 1800 m.
Distribution: MP—P. Lovett 42
Lantana ukambensis (Vatke) Verdc.; Verdcourt,
F.T.E.A. (Verbenaceae): 43. 1992.
Terrestrial herb or suffrutex. Woodland. 1200
m.
Distribution: MP—J. Lovett 4255
Lippia abyssinica (Otto & A. Dietr.) Cufod.; Verdcourt, F.T.E.A. (Verbenaceae): 29. 1992.
Shrub or perennial herb. Riverine forest edges,
montane grassland. 1500–2140 m.
Distribution: SL—Gereau 4401
Lippia javanica (Burm. f.) Spreng.; Verdcourt,
F.T.E.A. (Verbenaceae): 30. 1992.
Shrub. Roadsides in cultivation, montane
forest. 1300–2200 m.
Distribution: MP, MR, RM—Gereau 4517
Lippia plicata Baker; Verdcourt, F.T.E.A. (Verbenaceae): 34. 1992.
Shrub. Woodland, riverine forest, montane
grassland, montane forest, abandoned cultivation.
1200–2010 m.
Distribution: MP, RM, SL—Gereau 4655
Premna senensis Klotzsch; Verdcourt, F.T.E.A.
(Verbenaceae): 78. 1992.
Shrub, sometimes lianescent. Woodland, lakeshore forest, rocky beaches. 474–1200 m.
Distribution: MP, SL—Gereau 3654
Rotheca alata (Gürke) Verdc.; Fernandes & Verdcourt, 2000: 148.
Shrub. Abandoned cultivation. 2000 m.
Distribution: MP—P. Lovett 253
Rotheca myricoides (Hochst.) Steane & Mabb.
subsp. austromonticola (Verdc.) Verdc.; Fernandes & Verdcourt, 2000: 151.
Shrub or subshrubby terrestrial herb. Montane
LakeNyasa_5-18.indd 111
111
grassland, woodland, Cupressus plantations.
1250–2550 m.
Distribution: MP, MR, RM—Richards 14342
(type: K), Abeid 2834
Rotheca myricoides (Hochst.) Steane & Mabb.
subsp. myricoides; Fernandes & Verdcourt,
2000: 151.
Shrub or treelet. Montane forest, woodland,
lakeshore forest, grassland, abandoned cultivation.
474–2350 m.
Distribution: MP, MR, RM, SL—Suleiman 150
Vitex buchananii Baker ex Gürke; Verdcourt,
F.T.E.A. (Verbenaceae): 54. 1992.
Shrub. Montane forest. 1760 m.
Distribution: RM—WCS SHCP 338
Vitex doniana Sweet; Verdcourt, F.T.E.A. (Verbenaceae): 62. 1992.
Tree. Riverine forest, woodland, abandoned
cultivation. 580–1600 m.
Distribution: MP, SL—Kayombo 973
Vitex fischeri Gürke; Verdcourt, F.T.E.A. (Verbenaceae): 59. 1992.
Tree. Abandoned cultivation. 1250 m.
Distribution: MP—J. Lovett 3778
Vitex mombassae Vatke; Verdcourt, F.T.E.A. (Verbenaceae): 64. 1992.
Tree. Woodland. 1250–1400 m.
Distribution: MP—Kayombo 1095
Vitex payos (Lour.) Merr. var. glabrescens (W.
Piep.) Moldenke; Verdcourt, F.T.E.A. (Verbenaceae): 63. 1992.
Tree. Riverine forest. 580 m.
Distribution: SL—Gereau 3810
VIOLACEAE
2 genera and 2 species
Hybanthus enneaspermus (L.) F. Muell. var. latifolius (De Wild.) Engl.; Grey-Wilson, F.T.E.A.
(Violaceae): 34. 1986.
Shrub or subshrub. Montane forest. 1790–
2400 m.
Distribution: NL, RM—Gereau 4151
Viola abyssinica Steud. ex Oliv.; Grey-Wilson,
F.T.E.A. (Violaceae): 25. 1986.
Terrestrial herb. Montane forest, montane
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112
grassland. 2040–2750 m.
Distribution: KP, MR, NL, RM, SL—Gereau
4723
Note: Most collections are somewhat intermediate with V. eminii (Engl.) R. E. Fr.
VISCACEAE
1 genus and 4 species
Viscum congdonii Polhill & Wiens; Polhill & Wiens,
F.T.E.A. (Viscaceae): 19. 1999.
Parasitic shrub. Montane forest and woodland.
2230–2350 m.
Distribution: KP, MR—Wiens 6985 (type: BR,
DSM, EA, K, MO, SRGH), Mwasumbi 16173
Viscum goetzei Engl.; Polhill & Wiens, F.T.E.A.
(Viscaceae): 18. 1999.
Parasitic shrub. Montane forest. 2350 m.
Distribution: KP—Goetze 1273 (type: B, BR,
K), Wiens 6986
Viscum loranthicola Polhill & Wiens; Polhill &
Wiens, F.T.E.A. (Viscaceae): 16. 1999.
Parasitic shrub. Woodland. 1310 m.
Distribution: MP—Gereau 6101
Viscum shirense Sprague; Polhill & Wiens, F.T.E.A.
(Viscaceae): 23. 1999.
Parasitic shrub. Montane forest. 1920 m.
Distribution: SL—Gereau 3619
VITACEAE
5 genera and 25 species
Ampelocissus africana (Lour.) Merr.; Verdcourt,
F.T.E.A. (Vitaceae): 5. 1993.
Herbaceous climber or liana. Riverine forest,
lakeshore forest, woodland. 474–1400 m.
Distribution: MP, SL—Gereau 4283
Ampelocissus obtusata (Welw. ex Baker) Planch.
subsp. kirkiana (Planch.) Wild & R. B. Drumm.;
Verdcourt, F.T.E.A. (Vitaceae): 8. 1993.
Herbaceous climber. Woodland. 1200 m.
Distribution: MP—J. Lovett 4051
Ampelocissus obtusata (Welw. ex Baker) Planch.
subsp. obtusata; Verdcourt, F.T.E.A. (Vitaceae): 7. 1993.
Terrestrial herb. Miombo woodland. 1200 m.
Distribution: MP—J. Lovett 4044
LakeNyasa_5-18.indd 112
Gereau et al.
Cayratia gracilis (Guill. & Perr.) Suess.; Verdcourt,
F.T.E.A. (Vitaceae): 138. 1993.
Herbaceous climber. Riverine forest, abandoned cultivation. 1250–1500 m.
Distribution: MP, SL—Gereau 4395
Cissus cornifolia (Baker) Planch.; Verdcourt,
F.T.E.A. (Vitaceae): 36. 1993.
Shrub. Acacia woodland. 1250–1300 m.
Distribution: MP—J. Lovett 3571
Cissus faucicola Wild & R. B. Drumm.; Verdcourt,
F.T.E.A. (Vitaceae): 47. 1993.
Herbaceous climber. Montane forest, riverine
forest. 1250–2300 m.
Distribution: MP, SL—Kayombo 2344
Cissus integrifolia (Baker) Planch.; Verdcourt,
F.T.E.A. (Vitaceae): 15. 1993.
Liana. Lakeshore shrubland and forest.
474–490 m.
Distribution: SL—Gereau 3667
Cissus oliveri (Engl.) Gilg; Verdcourt, F.T.E.A.
(Vitaceae): 31. 1993.
Shrub. Montane forest. 1860 m.
Distribution: RM—WCS SHCP 34
Cissus rubiginosa (Welw. ex Baker) Planch.; Verdcourt, F.T.E.A. (Vitaceae): 26. 1993.
Herbaceous climber. Riverine forest. 580 m.
Distribution: SL—Gereau 3811
Cissus sciaphila Gilg vel sp. aff.; Verdcourt,
F.T.E.A. (Vitaceae): 30. 1993.
Liana. Riverine forest. 1210 m.
Distribution: SL—Gereau 4284
Cyphostemma aff. bambuseti (Gilg & M. Brandt)
Desc. ex Wild & R. B. Drumm.; Verdcourt,
F.T.E.A. (Vitaceae): 119. 1993.
Herbaceous climber. Riverine forest. 1210–
1600 m.
Distribution: MP, SL—Gereau 4289
Cyphostemma buchananii (Planch.) Desc. ex Wild
& R. B. Drumm.; Verdcourt, F.T.E.A. (Vitaceae): 99. 1993.
Herbaceous climber or prostrate terrestrial
herb. Riverine forest, montane forest. 1780–2140
m.
Distribution: RM, SL—Suleiman 94
Cyphostemma crotalarioides (Planch.) Desc. ex
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Lake Nyasa Climatic Region
Floristic Checklist
Wild & R. B. Drumm.; Verdcourt, F.T.E.A.
(Vitaceae): 81. 1993.
Terrestrial herb. Miombo woodland. 1200–
1530 m.
Distribution: MP, SL—Gereau 3834
Cyphostemma cyphopetalum (Fresen.) Desc. ex
Wild & R. B. Drumm.; Verdcourt, F.T.E.A.
(Vitaceae): 121. 1993.
Herbaceous climber. Montane forest, woodland, abandoned cultivation. 1250–2350 m.
Distribution: MP, MR, RM—Gereau 4601
Cyphostemma gigantophyllum (Gilg & M. Brandt)
Desc. ex Wild & R. B. Drumm.; Verdcourt,
F.T.E.A. (Vitaceae): 68. 1993.
Herbaceous climber. Riverine forest, miombo
woodland. 1500–1620 m.
Distribution: SL—Gereau 3939
Cyphostemma heterotrichum (Gilg & R. E. Fr.)
Desc. ex Wild & R. B. Drumm.; Verdcourt,
F.T.E.A. (Vitaceae): 77. 1993.
Terrestrial herb. Woodland, abandoned cultivation. 1200–1500 m.
Distribution: MP—J. Lovett 4356
Cyphostemma hildebrandtii (Gilg) Desc. ex Wild &
R. B. Drumm.; Verdcourt, F.T.E.A. (Vitaceae):
66. 1993.
Herbaceous climber. Riverine forest. 580 m.
Distribution: SL—Gereau 3816
Cyphostemma hypoleucum (Harv.) Desc. ex Wild
& R. B. Drumm. vel sp. aff.; Wild & Drummond, Fl. Zamb. 2: 484. 1966.
Herbaceous climber. Scree slopes. 2060–2120
m.
Distribution: SL—Gereau 4439
Cyphostemma junceum (Webb) Desc. ex Wild &
R. B. Drumm. subsp. jatrophoides (Welw. ex
Baker) Verdc.; Verdcourt, F.T.E.A. (Vitaceae):
62. 1993.
Terrestrial herb. Woodland, edges of cultivation. 1300 m.
Distribution: MP—J. Lovett 3552
Cyphostemma kilimandscharicum (Gilg) Desc. ex
Wild & R. B. Drumm. var. jaegeri (Gilg & M.
Brandt) Verdc.; Verdcourt, F.T.E.A. (Vitaceae):
110. 1993.
Herbaceous climber. Montane forest. 1790 m.
Distribution: RM—Abeid 2917
LakeNyasa_5-18.indd 113
113
Cyphostemma kilimandscharicum (Gilg) Desc. ex
Wild & R. B. Drumm. var. kilimandscharicum;
Verdcourt, F.T.E.A. (Vitaceae): 108. 1993.
Herbaceous climber or liana. Montane forest,
tree clumps in grassland. 1730–2400 m.
Distribution: MP, MR, NL—Gereau 4144
Cyphostemma masukuense (Baker) Desc. ex Wild
& R. B. Drumm. subsp. masukuense; Verdcourt, F.T.E.A. (Vitaceae): 105. 1993.
Herbaceous climber or weak liana. Montane
forest, woodland. 1950–2110 m.
Distribution; MP, MR, RM—Gereau 4554
Cyphostemma princeae (Gilg & M. Brandt) Desc.
ex Wild & R. B. Drumm.; Verdcourt, F.T.E.A.
(Vitaceae): 79. 1993.
Terrestrial herb. Woodland, riverine forest,
abandoned cultivation. 1200–1720 m.
Distribution: MP—J. Lovett 4078
Cyphostemma pseudosesquipedale Verdc.; Verdcourt, F.T.E.A. (Vitaceae): 83. 1993.
Terrestrial herb. Woodland. 1200 m.
Distribution: MP—J. Lovett 3953
Cyphostemma vandenbrandeanum (Dewit) Desc.
ex Wild & R. B. Drumm.; Verdcourt, F.T.E.A.
(Vitaceae): 102. 1993.
Herbaceous climber. Scree slopes, rocky mountain grassland. 1980–2140 m.
Distribution: SL—Gereau 3643
Cyphostemma sp.
Herbaceous climber. Riverine forest. 1210 m.
Distribution: SL—Gereau 4254
Rhoicissus tridentata (L. f.) Wild & R. B. Drumm.;
Verdcourt, F.T.E.A. (Vitaceae): 9. 1993.
Liana. Montane grassland, montane forest
edges, thickets, riverine forest, woodland, abandoned cultivation. 1100–2140 m.
Distribution: MP, NL, RM, SL—Suleiman 96
XYRIDACEAE
1 genus and 1 species
Xyris obscura N. E. Br.; Lock, F.T.E.A. (Xyridaceae): 10. 1999.
Terrestrial herb. Wet montane grassland.
2550–2740 m.
Distribution: KP, RM—Gereau 3187
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114
ZINGIBERACEAE
2 genera and 2 species
Aframomum zambesiacum (Baker) K. Schum.;
Lock, F.T.E.A. (Zingiberaceae): 25. 1985.
Terrestrial herb. Stream banks in woodland.
1650–1680 m.
Distribution: RM, SL—Gereau 5120
Siphonochilus rhodesicus (T. C. E. Fr.) Lock;
Lock, F.T.E.A. (Zingiberaceae): 20. 1985.
Terrestrial herb. Miombo woodland. 1200 m.
Distribution: MP—J. Lovett 4677
ZYGOPHYLLACEAE
1 genus and 1 species
Balanites aegyptiacus (L.) Delile; Sands, F.T.E.A.
(Balanitaceae): 6. 2003.
Tree. Woodland, termite mounds in grassland.
1250–1400 m.
Distribution: MP—J. Lovett 4916
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