Catalogue of

Palaearctic Coleoptera
Volume 7

Curculionoidea I

Edited by
I. LÖBL & A. SMETANA

Apollo Books
 Catalogue of
Palaearctic Coleoptera

Volume 7

Curculionoidea I

DEDICATION

The present volume is dedicated to Lothar Dieckmann (25.08.1920 in Leipzig - 9.02.1990 in

Eberswalde), who as few others, shared his outstanding knowledge of weevils with all who showed
interest. Thus, he influenced modern study of Central European curculionoids more than any of his
contemporaries. His work, most of the time realized under repressive totalitarian regime, demon-
strates primacy of honest interest in beetles over the actually generalized competition pressure in
professional biology.
Lepidapion argentatum (Gerstaecker, 1854)
 Catalogue of
Palaearctic Coleoptcra
Volume 7

Curculionoidea I

Edited by
I. LÖBL & A. SMETANA

Apollo Books
Stenstrup, 2011
© Copyright 2011 by Apollo Books, Stenstrup, Denmark.

All rights reserved. No part of this book may be reproduced or translated in any form,
by print, photoprint, microfilm, microfiche, disc, CD-Rom or any other means without
written permission from the publisher.

Cover illustration: Olga Mansilla.

Edited by: I. Löbl & A. Smetana.

Printed by: Vinderup Bogtrykkeri Aps.

Published by:
Apollo Books
Kirkeby Sand 19
DK-5771 Stenstrup
Denmark
Telephone + 45 62 26 37 37
Telefax + 45 62 26 37 80
E-mail: apollobooks@vip.cybercity.dk
Web site: www.apollobooks.com

Chapters from this publication should be cited as follows:

Alonso-Zarazaga, M. A., 2011: Nemonychidae, pp. 90. - In I. Löbl & A. Smetana (ed.):
Catalogue of Palaearctic Coleoptera, Vol. 7. Stenstrup, Apollo Books 373pp.

ISBN 978-87-88757-93-4 (Vol. 7)

ISBN 978-87-88757-71-2 (Vols 1-8)

Date of publication: 01.02.2011.

 CONTENTS

Introduction .......... ; 7
Taxonomic information ; 10
Distributional information 11
Biographical information -, 16
Acknowledgements 17
Authors ,.....,.,...., iw «> ÄJt
Mailing a n d e-mail addresses of the authors ;: 19
Comments of the Editors ,„,.„ „.. t t..*i*..^ 21
Errata to Volumes 1 to 6 ,. .....'.'...' 23
New nomenciatural and taxonomic acts, and comments * .< 64
Catalogue ...^..w.*- *u iuuM.* 90
Nemonychidae 90
Anthribidae ^..^„^..^^ .^ 90
Belidae ;* KWu.wwvtoà 109
Rhynchitidae i'. 109
Attelabidae „J, .^ 129
142
Brentidae «•»-•*• w
Apionidae ,.,. r fr, 148
Nanophyidae 177
Brachyceridae 182
Dryophthoridae 185
Erirhinidae 192
Raymondionymidae r.« 198
Curculionidae IHA s.*i 201
Platypodinae i......v. 201
Scolytinae - 204
Baridinae 251
References -...». 262
Index to family-group names - 358
Index to genus-group names 360
 7

INTRODUCTION
Structured knowledge requires categorisation of observations and phenomena, and one of these phenomena is
life. Structured knowledge of life, as it evolved and is present on our planet, requires cataloguing.
Taxonomy has a profound impact in many fields, including genetics, physiology, ecology, plant protection,
nature conservation, and legislation. At the core of taxonomic information are taxonomic catalogues that serve as
guides to the diversity of life. They summarize the work in systematics and taxonomy that enable assessment of
faunal diversity and classification, and provide information on historical taxonomic actions. Catalogues are also
essential in achieving a universal language in taxonomy that assures unambiguous transfer of biological
information.
The ever increasing number of taxa described since the tenth edition of Linné's Systema Naturae was probably
the main motivation for compiling catalogues in the past. Since the middle of the 19 century numerous
catalogues were published, usually covering restricted parts of the world and served as checklists rather than
works that we would qualify today as full catalogues. While knowledge is dynamic, catalogues are static and
reflect a time restricted information. Therefore, there is always a need of newer, up-to-date, and complete
catalogues. The first world catalogue of Coleoptera was Gemminger & Harold's Catalogus Coleopterorum
issued from 1868 to 1874. It was followed by Junk & Schenkling's (editors) Coleopterorum Catalogus,
published in separate parts from 1910 to 1940. At present, even though it is desperately obsolete, the
Coleopterorum Catalogus remains the only source of comprehensive taxonomic information on a world basis for
many families of beetles. The only catalogue covering all groups of Coleoptera occurring in the Palaearctic
region is Winkler's 1924 - 1932 Catalogus Coleopterorum regionis palaearcticae. However, the Junk &
Schenkung and Winkler catalogues differ. The former one is approaching the definition of a "full" catalogue,
while the latter is rather a checklist. They both share two serious omissions. The primary sources of taxonomic
acts were not consistently checked, and the information on distribution was not detailed enough to satisfy the
needs of biologists. In addition, many rules subsequently adopted by the International Code of Zoological
Nomenclature were not respected. Due to the lack of modern catalogues, the language in taxonomic literature
remains inadequate in many respects. In particular, unavailable names are regularly used as available and
sometimes even regarded as valid, various spellings are used for the same taxon, and references to taxonomic
actions are inexact to a high degree for papers published during the 20th century.
Undoubtedly, the Coleoptera of the Palaearctic Region were over the years studied by more professional and
amateur biologists than those in other parts of the World, and the results may exceed those of the study of many
other arthropods. It is therefore remarkable that an overview of the present knowledge of the Palaearctic
Coleoptera is badly lagging behind other groups, such as the Nearctic Coleoptera, or Palaearctic Diptera.
Systematic studies of many groups of Palaearctic Coleoptera, even in view of their popularity for biologists,
have to start with searches for data scattered throughout the literature of the past 250 years.
A full catalogue provides information concerning all published names within a group, their classification,
taxonomic history, and so on. Ideally, the present Catalogue should include such data. However, considering the
number of taxa and the low number of present day taxonomists, along with difficulties in accessing primary
literature, a full catalogue of all Palaearctic beetles would not be completed within a reasonable period of time.
Having this in mind, the aim of this Catalogue is to provide a tool that meets the most urgent needs: 1) a list of
available names, both valid and invalid, of taxa occurring in the Palaearctic Region, in their verified orthography
and with correct publication dates, 2) a complete list of verified references to primary descriptions, and 3)
informative distributions of the species and subspecies. Thus, the Catalogue is expected to respond to questions
related to biodiversity and to the stability of the taxonomic nomenclature.
This Catalogue is a collective work involving many authors. For its realisation, it was necessary to achieve a
reasonable consensus in very diverse fields. The Editors appreciate the understanding of the need of consistency
by the contributing authors. The most controversial points concern the spelling of names, and the delimitation of
the Palaearctic Region and its main subdivisions.
8 INTRODUCTION
The first point is purely formal. The alternatives adopted by the Editors and presented below result from the
consensus achieved during the discussions with colleagues. The Editors are aware of the difficulty in satisfying
all opinions and appeal to the understanding of the user.
The second point pertains to the presentation of distributional data. The Paiaearctic Region, as it is defined for
the purpose of this Catalogue, includes some areas that are usually considered to belong to the Afrotropical,
Oriental, and Pacific Regions respectively, i.e. die south of the Arabian Peninsula and Suqutra, all of Pakistan,
die Himalayan part of India including Arunachal Pradesh, Bhutan, and Nepal, the south of China, and the Pacific
Islands of Japan. The main reasons for inclusion of diese areas are as follows:
Old records of "Arabia" and "China" may pertain to any states on the Arabian Peninsula and to Jordan, or to any
part of the People's Republic of China, respectively.
1. Recent field work in the Himalaya and in die mountains of mainland China and Taiwan provides evidence of
altitudinal faunal transition in Coleoptera, as well as in otiier insect groups. While subtropical climate witii
dominant Oriental taxa prevails at low elevations, significant faunal changes are found already at elevations
of 1500 m, and almost "pure Paiaearctic" taxa are present at elevations of 2500 m and above. Thus, it
appears illusory to draw simple biogeographical frontiers in any of diese parts of die world.
2. The Catalogue includes more information. This is an obvious advantage in the absence of modern catalogues
covering the Afrotropical and Oriental regions.

The Catalogue of Paiaearctic Coleoptera is published in a number of independent volumes, each having its own
Reference section and Index. The taxonomic, geographical and bibliographical information in dûs volume is
presented as in the previous Volumes.
The present volume treats the smaller families of die polyphagous superfamily Curculionoidea, and four
subfamilies of die family Curculionidae, die Plastypodinae, Scolytinae, and Baridinae. Unlike the previous
volumes, die adopted arrangement of families and subfamilies witiiin the superfamily is not consistent with J. F.
Lawrence & A. F. Newton's 1995 classification (in J. Pakaluk & S. A. Slipinski (eds): Biology, phylogeny, and
classification of Coleoptera. Papers celebrating the 8(f birthday of Roy A. Crowson. Volume 2. Warszawa:
Muzeum i Instytut Zoologii PAN). The adopted classification is commented in die chapter "New nomenclatural
and taxonomic acts and comments.
The volume includes about 8320 names of taxa and 2530 primary references to the genus- and species-groups
names.
The Editors adopted the policy that the gap between the publication date of each volume of the Catalogue and
the date of die corresponding entry should be as short as possible, and not exceed three years. The first Volume
contains the available genus-group and species-group names in Archostemata, Myxophaga and Adephaga
published before January 1, 2000; die second Volume contains the available names in Staphyliniformia
published before January 1,2002; the third Volume contains the available names in Scarabaeoidea, Dascilloidea,
Buprestoidea and Byrrhoidea published before January 1, 2004; the fourth Volume contains the available names
in Elateroidea, Derodontoidea, Bostrichoidea, Lymexyloidea, Cleroidea, and Cucujoidea published before
January 1, 2005; the fifth Volume contains the available names in Tenebrionoidea published before January 1,
2007; the sixth Volume contains the available names in Chrysomeloidea published before January 1, 2009. The
present Volume is extended to contain die available names of die included groups of Curculionoidea published
before January 1, 2010.
An index of the species-group names was not provided for practical reasons associated witii die printing and
bookbinding costs. Therefore, as for the previous volumes, an electronic version of species-group names indexes
is available on the web sites of the Apollo Books (www.apollobooks.com) and of the Muséum d'histoire
naturelle in Geneva (www.ville-ge.ch/musinfo/mhng).
The Editors were repeatedly asked to make die Catalogue available electronically, either as an online interactive
database, or as CD-Rom. After careful consideration they decided to publish the work only in book form for the
following reasons: Information in taxonomy, unlike that in most other biological fields, remains useful for
 INTRODUCTION 9
extremely long periods of time, extending over centuries. Taxonomic work deals commonly with sources
published throughout the 19th and 20th centuries and often even earlier. Consequently, catalogues that summarize
taxonomic work are consulted over a very long period of time. The simple, although often ignored, reason for the
durability of taxonomic data is in the fact that taxonomy provides the language indispensable for unambiguous
conveyance of biological information. The bulk of correctly recorded data in Catalogues remains informative
even under a continuous inflow of additional new taxa, and even if the assignments and ranks of taxa and the
validity or invalidity of names are changing, and the known distributions of species become gradually more
precise. It is therefore necessary to insure access to taxonomic work, including Catalogues, not only in spatial
but also in temporal dimensions. Experience shows that printed texts may remain available for centuries, while
life expectancy of electronic information is unknown. Ideally, works like this Catalogue, should be available in
both print- and electronic formats; however, the real world is not ideal and the production of printed Catalogues
is associated with considerable costs. The consequence of providing low-cost online or CD-Rom editions would
have a serious negative economic impact on the production of printed version. We believe that under the
globally inadequate institutional interest for the needs of taxonomy, the livelihood of those who are willing to
accept the commercially hazardous production of printed taxonomic publications should be protected. We
advocate the growth and continuation of taxonomy, not its demise.
10

TAXONOMIC INFORMATION
The present Catalogue includes all available names, both valid and invalid, of extant beetle taxa described before
January 1, 2010 and known to occur in the Palaearctic Region, as it is defined below. The higher classification,
from suborder down to subfamily, is based on the work of Lawrence & Newton (1995): Families and
subfamilies ofColeoptera (with selected genera, notes, references and data on family-group names). However,
the classification proposed in this work is not taken as dogma, and changes are accepted when considered well
founded. All taxa below subfamily rank are arranged alphabetically within the higher taxon and the synonyms
follow the respective valid name alphabetically.
Extinct taxa, names rejected by the ICZN {International Code of Zoological Nomenclature, Fourth Edition.
London: International Trust for Zoological Nomenclature, 1999), misspellings, misidentifications and other
nomina nuda are not included in the Catalogue. However, concepts that are important for nomenclatural
purposes, may be included. Similarly, all infrasubspecific names, such as those established as "morpha", "natio",
"race", "subvariety" and "aberration", or proposed as variety and form of a subspecies or another variety, and
names published, e.g., as A-us b-us c-us, but specified in the text that they are actually proposed for a "natio" or
"race", etc., are not considered subspecific and are therefore excluded from the Catalogue. Names proposed as
varieties and forms before 1961 are included, if deemed subspecific under the provisions of the ICZN, Article
45.6. Unjustified emendations are available and may be included.
The currently valid names of the family-group taxa include die name of the autiior and tiie year of me
publication. Their synonyms are not listed.
The names of die genus-group taxa are given with the name of die aumor, and the year and page of publication.
The page given is the page where die name and die actual description of the taxon is printed. The type species of
all genus-group names are given in tiieir original combination. If die type species is currently regarded as a
junior synonym, die valid senior synonym is given in brackets in its original combination.
The names of the species-group taxa are given witii the name of die aumor, and me year and page of publication.
The page given is die page where the name and die actual description of die taxon is printed. In bi-languaged
Chinese/English and Japanese/English publications both respective pages of the actual description may be given.
For species-group taxa subsequently transferred to another genus, die name of die original genus is given in
parentiieses, following the page of publication.
Some authors (e.g., V. Apfelbeck, H. John) published the same description twice, or even more times, in separate
papers. Such publications produce, de facto, primary homonyms and objective synonyms. The first publication
in such cases is referred to as indicated above, followed by the mark = , die year and first page of die subsequent
description/s in square brackets. This is particularly important for taxa mat are erroneously associated witii tiieir
junior description.
The following symbols, all given in square brackets following die page of publication, or the original
combination when applicable, are used for taxonomic information: HN for homonyms, RN for replacement
names, NO for nomina oblita, NP for nomina protecta, DA for doubtful assignment, and EA for erroneous
assignment.
Taxa considered incertae sedis and nomina dubia are listed separately at me end of the nearest applicable taxon.
Taxonomic and nomenclatural acts published after December 31, 2009 are considered only when tiiey concern
taxa described on or before that date.
 11
DISTRIBUTIONAL INFORMATION
The limits of the Palaearctic region, as those of other biogeographical regions, are arbitrarily defined (Map 1 ).
For practical reasons, the boundaries of the Palaearctic Region, as they were established for the Catalogue (see
above), usually follow national boundaries. The region includes Europe, Africa north of the Sahara, and Asia
except for the part that is arbitrarily defined as belonging to the Oriental Region.
For each species and subspecies an outline of its present distribution is given. Fossil records are not considered.
The information is given by means of symbols, presented at three levels.
The first level is the subdivision of the Palaearctic Region into three main parts, Europe (letter E, bold), North
Africa (letter N, bold) and Asia (letter A, bold).
Europe includes the Azores, Iceland and Turkey west of the Bosporus. The eastern boundaries are a matter of
controversy. In the Catalogue, Europe includes Russia west of the main ridge of the Ural mountains, the Permsk
Oblast, Bashkortostan Republic and Orenburskaya Oblast, and the small part of Kazakhstan west of the Ural
River. It includes the Caucasian republics of Georgia, Armenia and Azerbaijan. The south-eastern boundaries are
the political boundaries of the Asian part of Turkey, Iran, Kazakhstan, and the Caspian and Black seas.
Yugoslavia has been recently split into two independent countries: Serbia and Montenegro. At the time of the
split almost all distributional records for the respective species were finished, using the symbol YU. It would
have been an unreasonably time consuming effort to retroactively differentiate these records, therefore, as a
compromise, the term Yugoslavia is no longer used in the list of geographical symbols on page 14, but records
for Serbia and Montenegro are still listed under the symbol YU in the body of the Catalogue.
North Africa includes Morocco (incl. Western Sahara), Algeria, Tunisia, Libya and Egypt west of the Suez
Canal, and the Canary and Madeira islands.
Asia includes Sinai and the Arabian Peninsula (including Suqutra), Turkey east of the Bosporus, the Middle East
and Central Asian countries, Russia east of the main ridge of the Ural mountains, Korea, Japan (including
Ryukyu [= Nansei] Islands and the Japanese Pacific Islands), the entire People's Republic of China, Taiwan,
Bhutan, Nepal, North India along the base of the Himalaya (Arunachal Pradesh, Uttarakhand (= Uttaranchal),
northwestern area of former Uttar Pradesh), Himachal Pradesh), Jammu & Kashmir and all of Pakistan. Thus,
India is the only state for which the strict political boundaries are not respected. Large parts of Uttar Pradesh
south of Nepal are overpopulated plains. Information on Coleoptera from this North Indian state is based almost
exclusively on its Himalayan districts lying west of Nepal, which are in the present state Uttarakhand.
The second level of the geographic information is provided by two-letter symbols for countries, major areas of
Russia and North Indian states, and by three-letter symbols for provinces of mainland China and for Taiwan (see
Table 1, Map 3).
The symbols are arranged in alphabetical order within the first-level subdivision. Some, usually older,
distributional records cannot be accommodated within the structure of the Catalogue (e.g., Arabia, Caucasus,
North India, Siberia). Such information is given in quotation marks (e.g., "Caucasus") behind the last symbol of
the respective first-level symbol. Russia is subdivided into six major sub-regions (Map 2), each of which has its
own two-letter symbol. These, as well as the symbol RU for Russia, are used only when more detailed
information is not available. Similarly, the symbol CH for the People's Republic of China, as well as the seven
two-letter symbols for China's major regions, are used only in the absence of more detailed geographical
information.
In general, the published distributional information is based on both identified material in collections and on
published records, scattered in an enormous number of taxonomic and faunistic papers that are virtually
impossible to review in their entirety. Revisions of collections reveal a high proportion of misidentifications,
attaining 40% of specimens in some large museums. The degree of identification reliability and of the records
derived from identifications, is a function of faunal diversity and quality of systematic revisions. Thus, the
reliability in general increases from south to north and from poorly studied groups to "popular" groups. At
present, a number of modern catalogues or check-lists, covering the beetle faunas of many European and some
extra-European countries or archipelagos, are available. The use of data contained in these and other faunistic
 DISTRIBUTIONAL INFORMATION
works is left to the discretion of the authors, who may also add unpublished information available to them. The
second-level geographic information is not necessarily exhaustive, it should rather be considered as a base for
future faunal research.
The third-level geographic information concerns species and subspecies with restricted distribution. Taxa of this
category may be strict endemites, or taxa comparatively widely distributed in one area but restricted in another
area. For example, the distributional record of a species widely distributed in North Africa with isolated
occurrence on Pantelleria would appear as follows: E: IT (Pantelleria) N: AG MO TU. The third-level
information is facultative. It is given in parentheses after the respective second-level symbol. The official
language of the respective state is used for records in languages using the Latin alphabet, or it is transliterated
from the Cyrillic alphabet. Records in languages using non-Latin or Cyrillic characters (e.g., Chinese or Japanese
pictographs) are translated into English, and the translated geographical terms are spelled as closely as possible
to those used in the Times Atlas®, or in other well-known sources. Detailed geographical information may refer
to natural geographical features such as islands, mountains, lakes, valleys, caves, or to administrative entities,
such as districts.
The extralimital distribution of some Palaearctic species is indicated by three letter symbols in bold,
cosmopolitan species by the symbol COS, all located at the end of the respective geographical information (see
Table 1). Introductions are indicated by the letter "i" (e.g., Ei: GB). The extralimital regions for the needs of the
Catalogue are defined as follows (see Map 1): Nearctic (NAR): north of Mexico; Neotropical (NTR): south of
the United States; Afrotropical (AFR): south of the North African states included in the Palaearctic Region;
Oriental (ORR): areas south of the People's Republic of China and Taiwan, areas south of the Himalaya in
India, the Philippines, Malaysia and Indonesia south to the Lydekker line; Australian (AUS): south of the
Lydekker line, Pacific.

Map 1. The limits of the geographic Regions as defined for the purpose of this Cataloj
 DISTRIBUTIONAL INFORMATION 13

Map 2. Subdivisions of Russia

Map 3. Subdivisions and provinces of the People's Republic of China

14 DISTRIBUTIONAL INFORMATION

Table 1: GEOGRAPHICAL SYMBOLS

E Europe KZ Kazakhstan
AB Azerbaijan LA Latvia
AL Albania LS Liechtenstein
AN Andorra LT Lithuania
AR Armenia LU Luxembourg
AU Austria MA Malta
AZ Azores MC Macedonia
BE Belgium MD Moldavia
BH Bosnia Herzegovina ME Montenegro
BU Bulgaria NL The Netherlands
BY Belarus NR Norway
CR Croatia NT Russia: North European Territory
CT Russia: Central European Territory PL Poland
CZ Czech Republic PT Portugal
DE Denmark RO Romania
EN Estonia RU Russia
FA Faeroe Islands SB Serbia
FI Finland SK Slovakia
FR France (incl. Corsica, Monaco) SL Slovenia
GB Great Britain (incl. Channel Islands) SP Spain (incl. Gibraltar)
GE Germany SR Svalbard (Spitzbergen)
GG Georgia ST Russia: South European Territory
GR Greece (incl. Crete) SV Sweden
HU Hungary sz Switzerland
IC Iceland TR Turkey
IR Ireland UK Ukraine
IT Italy (incl. Sardinia, Sicily, San Marino) YU Serbia and Montenegro

N North Africa LB Libya

AG Algeria MO Morocco (incl. Western Sahara)
CI Canary Islands MR Madeira Archipelago
EG Egypt TU Tunisia

A Asia IQ Iraq
AE Arab Emirates IS Israel
AF Afghanistan JA Japan
AP India: Arunachal Pradesh JO Jordan
BA Bahrain KA India: Kashmir
BT Bhutan KI Kyrgysstan
CE China: Central Territory KU Kuwait
CH China KZ Kazakhstan
CY Cyprus LE Lebanon
ES Russia: East Siberia MG Mongolia
FE Russia: Far East NE China: Northeast Territory
HP India: Himachal Pradesh NC North Korea
IN Iran NO China: Northern Territory
 DISTRIBUTIONAL INFORMATION 15

NP Nepal SW China: Southwestern Territory

NW China: Northwest Territory SY Syria
OM Oman TD Tajikistan
PA Pakistan TM Turkmenistan
QA Qatar (incl. United Arab Emirates) TR Turkey
RU Russia UP India: Uttarakhand (= Uttaranchal), Uttar
SA Saudi Arabia Pradesh
SC South Korea UZ Uzbekistan
SD India: Sikkim, Darjeeling District WP China: Western Plateau
SE China: Southeastern Territory (incl. WS Russia: west Siberia
Macao, Hongkong) YE Yemen (incl. Suqutra)
SI Egypt: Sinai

CHINA: PROVINCES, AUTONOMOUS

REGIONS OR MUNICIPALITIES, AND TAIWAN
ANH Anhui (Anhwei) LIA Liaoning
BEI Beijing (Peking or Peiping) MAC Macao
CHQ Chongqing NIN Ningxia (Ningsia)
FUJ Fujian (Fukien) NMO Nei Mongol (Inner Mongolia)
GAN Gansu (Kansu) QIN Qinghai (Tsinghai)
GUA Guandong (Kwantung) SCH Sichuan (Szechwan)
GUI Guizhou (Kweichow) SHA Shaanxi (Shensi)
GUX Guangxi (Kwangsi) SHG Shanghai
HAI Hainan SHN Shandong (Shantung)
HEB Hebei (Hopeh) SHX Shanxi (Shansi)
HEI Heilongjiang (Heilungkiang) TAI Taiwan (Formosa)
HEN Henan (Honana) TIA Tianjin (Tsiensin)
HKG Hongkong XIN Xinjiang (Sinkiang)
HUB Hubei (Hupeh) XIZ Xizang (Tibet)
HUN Hunan YUN Yunnan
JIA Jiangsu (Kiangsu) ZHE Zhejiang (Chekiang)
JIL Jilin (Kirin)
JIX Jiangxi (Kiangsi)

WORLD ZOOGEOGRAPHICAL REGIONS

AFR Afrotropical Region
AUR Australian Region
NAR Nearctic Region
NTR Neotropical Region
ORR Oriental Region
16
BIBLIOGRAPHIC INFORMATION
References are given to, and only to, primary sources of the genus-group and species-group names included in
the Catalogue. No bibliographical reference is given to family-group names, to designation of type-species, to
secondary taxonomic acts, or to sources of any other information. Names of all authors, if applicable, are given
for each reference. An effort was put forth to unify the spellings of the names. For names that appeared in
publications in two or more different orthographies, the one used in the Catalogue is that which was considered
to be correct after thorough verification (e.g., the spelling Josef Müller for the author who also published as
Giuseppe Müller). In such cases the alternative spelling follows the original one in brackets. The original name is
used for authors that subsequently adopted additional names, or honorary epithets (e.g., A. Semenov, and not A.
Semenov-Tian-Shanskyi). Initials are given for all authors, even if they are missing in the original publication.
Names published in two or more alternative spellings, that seem to be equally correct, are spelled following
Schmitt, Hübner & Gaedike (1998): Nomina Auctorum. Auflösung von Abkürzungen taxonomischer Autoren-
Namen. Nova Supplementa Entomologica 11: 3-189. An exception is made for F. L. Castelnau de Laporte, who
is referred to as Laporte in recent major works. Therefore, the name Laporte is arbitrarily used in the Catalogue.
Names in Cyrillic are transliterated as for any other Cyrillic text, unless an alternative spelling was preferred by
the author, or the alternative name is generally used in the literature (e.g., Jakobson, Tschitschérine). It should be
noted that lowercase prepositions such as French "de" and German "von" are not given with the author's name in
the list of the taxa.

Titles of references are given in full, i.e. not abbreviated, with subtitles, as originally published. Text written in
Cyrillic is transliterated. Titles in languages using non-Latin or non-Cyrillic characters are translated into English
and are presented in square brackets, followed by the indication of the original language in parentheses.
Titles of periodicals are given in full. Series numbers are given in parentheses, and always precede the boldface
volume number. The numbers of separate issues are only provided when they are paginated separately, in which
case they are in parentheses following the volume number. Unnumbered pages are indicated in square brackets.
The titles of some periodicals vary over the years. For these periodicals the titles are given as they appeared for
the respective items referred to (e.g., Coleopterologische Rundschau I Koleopterologische Rundschau).
Contrarily, the journal Entomologische Blätter had several subtitles during its existence, the last of them being
Entomologische Blätter für Biologie und Systematik der Käfer. Since there is no danger of confusion with any
other periodical, the journal is cited simply as Entomologische Blätter. If the year of the volume differs from die
actual year of publication, the former follows die volume number and is given is square brackets. If periodicals
have no volume number, me year of me volume is given.
The Bulletin de la Société Entomologique de France is one of the more important periodicals in tiie field of
systematics. The title changed almost continuously for more than 60 years, appearing usually as Bulletin des
séances, Bulletin trimestriel or Bulletin entomologique. The first numbered volume, 37, was published 100 years
after the creation of the Annales and the Bulletin of die French Entomological Society in 1832. To avoid
confusion, this periodical is referred to as Bulletin de la Société Entomologique de France, and, from 1832 to
1931, is identified by the year in which it was published. From me year 1932, the volume number is given.
The Bulletin de la Société Entomologique de France was published from 1873 to 1894, in addition to me well
known quarterly version, in a bi-mensuel version "Bulletin Bimensuel". The pagination of die two versions are
not identical, the publication of the bi-mensuel version preceded the quarterly one. The Bulletin Bimensuel was
ignored by most coleopterists.
For ease of access, references of the same author and year were placed in order based on the first page number of
the paper, with Roman numerals being placed last, with respect to the number of the respective issue. Books
were placed first for a given year, unless a page range was cited, in which case the first-page rule was followed.
Papers of the same year and with the same first page numbers were placed in order of their last page number.
Thus the references are not arranged chronologically witiiin each year.
 17

ACKNOWLEDGEMENTS
As for the previous volumes of the Catalogue, the assistance of libraries was one of the prerequisites for
realisation of the present work. The most important libraries to be mentioned are: Muséum d'histoire naturelle,
Genève, Switzerland (Mrs. Christele Mougin); Agriculture and Agri-Food Canada (Entomological Library)
Ottawa, Canada (Mrs Patricia Madaire and Mr. Steve Gamman): Nârodni muzeum, Dept. of Entomology,
Kunratice, Czech Republic (Mrs Irena Chalupovâ), and Naturhistorisches Museum, Wien Austria (Mr. Wolfgang
Brunnbauer).

Special thanks are due to M. A. ALONSO-ZARAZAGA, Madrid, Spain who, in addition of his work as coordinator,
and author of a major part of the present volume, assisted in various ways other coauthors. He also provided a
number of old overlooked names of taxa listed in Anthribidae and Scolytinae under nomina dubia. Patrice
BOUCHARD and Yves BOUSQUET, both from Ottawa, Canada, were helpful in solving some difficult bibliographic
issues.

In addition, the Editors and authors of the present volume are indepted to many individuals who commented on
different parts of the Catalogue, added distributional information, completed references, and helped in various
ways. These include also coauthors and are as follows: Maxwell V. L. BARCLAY, London, UK; Cosimo
BAVIERA, Messina, Italy; Roger A. BEAVER, Chiangmai, Thailand; Aleä BEZDÈK, Ceské Budëjovice, Czech
Republic; Donald E. BRIGHT, Fort Collins, Colorado, USA; Anthony I. COGNATO, East Lansing, Michigan,
USA; Enzo COLONNELLI, Rome, Italy; Anthony DAVIES, Ottawa, Canada; Massimo FACCOLI, Legnaro, Italy;
Beat FORSTER, Zürich, Switzerland; Heiko GEBHARDT, Tübingen, Germany; Theodoor HEUERMAN, Wageningen,
The Netherlands; Dariusz IWAN, Warszawa, Poland; Josef JELINEK, Praha, Czech Republic; Bjarte H. JORDAL,
Bergen, Norway; Mark Ju. KALASHIAN, Erevan, Armenia; late Izya KERZNER, St. Petersburg, Russia; Hannes
KREHAN, Wien, Austria; Lawrence R. KIRKENDALL, Bergen, Norway; Boris A. KOROTYAEV, St. Petersburg,
Russia; Ren Li, Beijing, China; Anastâsios LEGÄKIS, Athens, Greece; Âke LINDELÖW, Uppsala, Sweden; late
Luigi MAGNANO, Poggibonsi, Italy; Michail Y. MANDELSHTAM, St. Petersburg, Russia; Massimo MEREGALLI,
Rivalta, Italy; Otto MERKL, Budapest, Hungary; David MIFSUD, Msida, Malta; Michael MORRIS, London, UK;
Vitaliy NAZARENKO, Kiev, Ukraine; Giuseppe B. OSELLA, Aquila, Italy; Sang-Wook PARK, Seoul, South Korea;
Jean PELLETIER, Monnaie, France; Ralph PETERS, Hamburg, Germany; Roberto POGGI, Genova, Italy; Antonio
G. REY, Genova, Italy; Alexander RIEDEL, Karlsruhe, Germany; Giorgio SABELLA, Catania, Italy; Heinrich
SCHÖNMANN, Wien, Austria; Rudolf SCHUH, Wiener Neustadt, Austria; Ignazio SPARACIO, Païenne, Italy; Mark
G. VOLKOVTTSH, St. Petersburg, Russia; Marek WANAT, Wroclaw, Poland; Beat WERMELINGER, Zürich,
Switzerland; late Stephen L. WOOD, Provo, Utah, USA; Petr ZAHRADNIK, Trnovâ, Czech Republic.

The number of detected errors published in the previous volumes of the Catalogue demonstrates universal
difficulties in assessment of correct data in taxonomy and nomenclature. In this respect, particular
acknowledgements are due to Jan BEZDËK, Brno, Czech Republic who took over most of the work associated
with the corrections of data on Chrysomelidae, and Mikhail L. DANILEVSKY, Moscow, Russia who provided
most of the corrections given for the Cerambycidae. Oscar VORST, Utrecht, The Netherlands, detected a number
of important errors published in the Volumes 1 and 2 of the Catalogue, and Patrice BOUCHARD, Ottawa, Canada,
corrected several errors in Scarabaeoidea that are relevant to nomenclature.

In addition to the Editors and the four above mentioned colleagues, following individuals identified hitherto
overlooked errors that appeared in the previous six volumes of the Catalogue: Karl ADLBAUER, Graz, Austria;
Miguel A. ALONSO-ZARAZAGA, Madrid, Spain; Kioshi ANDO, Osaka, Japan; Ron BEENEN, Nieuwegein, The
Neuerlands; Roger BOOTH, London, UK; Hervé BOUYON, Colombe, France; Manfred DÖBERL, Abensberg,
Germany; Roland GRIMM, Tübingen, Germany; Jifi HÀJEK, Praha, Czech Republic; Jifi HÂVA, Praha, Czech
Republic; Matthias HARTMANN, Erfurt, Germany; Carolus HOLZSCHUH, Villach, Austria; Colin JOHNSON,
Glossop, UK; Horst KIPPENBERG, Herzogenaurach, Germany; Vitëzslav KUBAN, Praha, Czech Republic; Harald
LABRIQUE, Lyon, France; Meiying LIN, Beijing, China; Adriean J. MAYOR, Gatlinburg, TN, USA; Otto MERKL,
Budapest, Hungary; Yuri MIKHAILOV, Ekaterinburg, Russia; Alexey MOSEYKO, St. Petersburg, Russia; Alfred F.
NEWTON, Jr., Chicago, USA; Renato REGALIN, Milano, Italy; Gianfranco SAMA, Cesena, Italy; Matthias
SCHÖLLER Berlin, Germany; Michael SCHMITT, Greifswald, Germany; Lukas SEKERKA, Ceské Budëjovice,
18 ACKNOWLEDGEMENTS
Czech Republic; Hans SILFVERBERO, Helsinki, Finland; Eva SPRECHER-UEBERSAX, Basel, Switzerland; Dmitri
TELNOV, Riga, Latvia; Antonio VERDUGO, San Fernando, Spain, and Stefano ZOIA, Milano, Italy.

Milos TRYZNA, Krâsnâ Lipa, Czech Republic, thanks his wife Drahoslava for support and understanding, and
Barry VALENTINE thanks his wife Buena S. VALENTINE, and Susan VALENTINE-COOPER, Columbus, USA.

The realisation of the Scolytinae and Platypodinae sections (by MiloS Knizek, Forestry and Game Management
Research Institute, Czech Republic) was supported by the Research Grant No. MZe 002070203 "Stabilization of
forest fonctions in anthropically disturbed and changing environmental conditions", which is financed by the
Ministry of Agriculture of the Czech Republic.

The photograph of Lothar DIECKMANN is one of a series made kindly disposable for publication by Hoiger
DATHE, Lutz BEHNE and Christian KUTZSCHER, all from Müncheberg, Germany.

The frontispiece illustration by Olga MANSILLA, Valladolid, Spain was provided by Miguel A. ALONSO-
ZARAZAGA, Madrid, Spain.

Mr. Daniel LÖBL, Genève, Switzerland, developed the software tools needed to generate the Indexes and was
helpful in solving a number of computer problems.

THE EDITORS
Ivan Löbl Ale§ Smetana
Muséum d'histoire naturelle Agriculture and
Agri-Food Canada
Genève, Switzerland Ottawa, Canada
 19
COORDINATOR OF THIS VOLUME
Miguel A. Alonso-Zarazaga, Depto. de Biodiversidad y Biologia Evolutiva, Museo Nacional de Ciencias
Naturales (CSIC), José Gutierrez Abascal, 2, E-28006 Madrid, Spain

AUTHORS
family NEMONYCHIDAE Bedel, 1882 M. A ALONSO-ZARAZAGA
family ANTHRIBIDAE Billberg, 1820
subfamily Anthribinae Billberg, 1820 M. TRYZNA, B. D. VALENTINE
subfamily Apolectinae Lacordaire, 1865 M. TRYZNA, B. D. VALENTINE
subfamily Choraginae Kirby, 1819 M. TRYZNA, B. D. VALENTINE
subfamily Urodontinae C. G. Thomson, 1859 M. A. ALONSO-ZARAZAGA
family BELIDAE Schoenherr, 1826 M. A. ALONSO-ZARAZAGADD
family RHYNCHITTDAE Gistel, 1848 M. A. ALONSO-ZARAZAGA
family ATTELABIDAE Billberg, 1820 M. A. ALONSO-ZARAZAGA
family BRENTIDAE Billberg, 1820 A. SFORZI
family APIONIDAE Schoenherr, 1823 M. A. ALONSO-ZARAZAGA
family NANOPHYIDAE Gistel, 1848 M. A. ALONSO-ZARAZAGA
family BRACHYCERIDAE Billberg, 1820 E. COLONNELLI
family DRYOPHTHORIDAE Schoenherr, 1825 C. H. C. LYAL
family ERIRHINIDAE Schoenherr, 1825 R. CALDARA
family RAYMONDIONYMIDAE Reitter, 1913 G B. OSELLA, P. HLAVÂC,
M. MEREGALLI
family CURCULIONIDAE Latreille, 1802
subfamily Platypodinae Shuckard, 1840 M. KNIZEK
subfamily Scolytinae Latreille, 1804 M. KNIZEK
subfamily Baridinae Schoenherr, 1836 J. PRENA

MAILING AND E-MAIL ADDRESSES OF THE AUTHORS

ALONSO-ZARAZAGA Miguel A., Depto. de Biodiversidad y Biologia Evolutiva, Museo Nacional de Ciencias
Naturales (CSIC), José Gutierrez Abascal, 2, E-28006 Madrid, Spain. E-mail: zarazaga@mncn.csic.es
CALDARA Roberto, Via Lorenteggio, 37,1-20146 Milano, Italy. E-mail: roberto.caldara@gmail.com
COLONNELLI Enzo, Via delle Giunchiglie, 56,1-00172 Roma, Italy. E-mail: ecolonnelli@aliceposta.it
HLAVÄC Peter, Na doline 14, SK-040 14 Kosice, Slovakia. E-mail: phlavac@stonline.sk
KNIZEK, Milos, Forestry and Game Management Research Institute, Jilovistë-Strnady, CZ-156 04 Praha 5 -
Zbraslav, Czech Republic. E-mail: knizek@vulhm.cz
LYAL Christopher H. C, Dept. of Entomology, The Natural History Museum, Cromwell Road, London SW7
5BD, UK. E-mail: chcl@nhm.ac.uk
MEREGALLI Massimo, Dipartemento di Biologia Animale e dell'Uomo, Università di Torino, via Accademia
Albertina 17,1-10123 Torino, Italy. E-mail: massimo.meregalli@unito.it
OSELLA Giuseppe B. Dipartimento di Scienze Ambientali, Università de l'Aquila, Via Vetoio, Coppito, 1-67100
L'Aquila, Italy. E-mail: osella@univaq.it
PRENA Jens, Systematic Entomology Lab, USDA, c/o Smithsonian Institute, National Museum of Natural
History, MRC-168, CE718, PO Box 37012, Washington, DC 20013-7012, USA. E-mail:
Jens.Prena@ARS.USDA.GOV
20 AUTHORS
SFORZI Alessandra, Museo Zooiogico "La Specola", Università di Firenze, Via Romana, 17,1-50125 Firenze,
Italy. E-mail: alessandra@unifi.it
TRYZNA Milos, Bohemian Switzerland National Park Administration, Research and Nature Conservation
Department, Prazskâ 52, CZ-407 46 Krâsnâ Lipa, Czech Republic. E-mail: m.tryzna@npcs.cz
VALENTINE Barry D., 5704 Lake Breeze Court, Sarasota, Florida 34233-5015 and 2359 Eastcleft Drive,
Columbus, OH 43221, USA. E-mail: bv@nwcs.com
 21
COMMENTS OF THE EDITORS
(I) C. J. Fabricius Entomologia systematica.... Tome I. Part 2. There is a persisting uncertainty about the
publishing date of Part 2 of this Fabricius's work that affects scores of taxa described within. Evenhuis,
1997: 248 gives "4 May 1793" as the publication date of part 2, based on "Gört. Gel. Anz." (= Göttingische
Gelehrte Anzeigen). This year was accepted as correct in volume 2 of the Catalogue, while in subsequent
volumes the year 1792 is given. To alleviate the confusion around the publication date, we consider it useful
to present the following explication.
J. C. L. Hellwig published his paper "Dritte Nachricht von neuen Gattungen im entomologischen System" in
Schneider's Neuester MagazinförLiebhaber der Entomologie 1 (3): 385- 408. The cover of the issue bears
printed date "Dezember 1792". There is no reason to believe that this date is incorrect, therefore, according
to ICZN Art. 21.3. Date incompletely specified the date to be adopted is [Art. 21.3.1] the last day of the
month, when month and year, but not day, are specified or demonstrated,..., therefore the date December 31,
1792 should be accepted as correct. Further confirmation of the publication in 1792 is found in Neue
allgemeine deutsche Bibliothek ("Des zehnten Bandes zweites Stück. Fünftes bis Achtes Heft", Kiel, 1794)
on page 455, under the heading "Naturlehre und Naturgeschichte" by the following information:
"Neuestes Magazin für die Liebhaber der Entomologie, herausgeben von Dr. H. Schneider, Advokat in
Stralsund, 1791. Ersten Bandes erstes Heft, klein Octav, 126 Seiten. Zweites, drittes und viertes Heft, 1792,
jedes Heft zu 8 Bogen."
Hellwig's (or possibly Schneider's) note on p. 408 states:"Nachdem ich auch den zweiten Teil des ersten Bandes
von Fabricii Entomol. systemat. erhalten habe, bin ich belehrt, dass die beiden ersten Familien der zuletzt
angefuehrten Gattung Synchita in diesem Werke die neue Gattung Lyctus nro. 107, S. 102 ausmachen ".
Both the page and the number of the genus are cited correctly.
Hence: (1) It is a matter of fact that Fabricius's book was published before Hellwig's paper and therefore
Fabricius should be considered as the author of the competing coleopteran names. (2) As there is good
evidence that the dating of Hellwig's paper is correct, Fabricius work should be dated 1792 under the Code
provisions. [Hellwig worked in Braunschweig (Brunswick), about 200 km from Kiel, where Fabricius
worked. He was in stable contact with Fabricius and it can be assumed that he asked him to send his book as
soon as possible because of publication on the same subject. Schneider was a senator in Stralsund, even
nearer to Kiel than Braunschweig, and especially to Copenhagen, where the book was published. He was a
goodfriendof Fabricius and was also interested in his book].

(II) Voet's (1778, and subsequent editions) Catalogus systematicus coleopterorum = Catalogue systématique
des coléoptères = Systematische naamlijst van dat geslacht van insecten dat men torren noemt is not an
available publication (and it has a long and complicated publication history, see Hagen, 1857:
Entomologische Zeitung (Stettin) 18: 405- 409), as it is not "consistently applicating binominal
nomenclature" (ICZN, Art. 11.4). Hence, all Voet's names (e.g., Carabus longus on p. 160, vol. 1) are to be
removed. The same applies to Voet (1793) Voet's Beschreibungen und Abbilaungen .... This is German
translation of Voet's work by Panzer, who is also not consistently applicating binominal nomenclature. An
information about the Voet's work was briefly mentioned in the Volume 6 of the Catalogue: we consider it
useful to give more details here.

(III) Genera des coléoptères d'Europe ... Tome quatrième: references to this work are often incorrect and the
dates of issue are erroneous. Here are the data resulting from reseach of Y. Bousquet (unpublished): The first
95 pages, constituting livraisons 1-12 of the entire work, were published in 1854-1855 and were authored by
Jacquelin du Val. The remaining pages, authored by Fairmaire, constitute livraisons 121-144 and were isused
as follows: 121-127: (pp. 97-176 + pis. 35-45) 1864 (Zool. Rec.) with livraisons 121-123 recorded on
October 1864 (Ent. Soc. London), 124-127 on June 1865 (Ent. Soc. London); 128-131: (pp. 177-203) 1865
(Zool. Rec.) with livraison 128 recorded on 4 September 1865 {Ent. Soc. London), 129 on 6 November 1865
(Ent. Soc. London), 130 on 1 January 1866 (Ent. Soc. London), and 131 on 4 June 1866 (Ent. Soc. London);
132-136: (pp. 205-??) before November 1867; 137-144: (pp. ??-295) 7 December 1868 (Ent. Soc. London)
[The title page is dated 1868].
99 COMMENTS OF THE EDITORS
(TV) Latreille's work: Genera crustaceorum et insectorum secundum ordinem naturalem in familias disposita,
iconibus exemplisque plurimis explicata. Tomus secundus. Parisiis et Argentorati: A. Koenig, 280 pp. is
usually given as published in 1807 - see also the volumes 2, 3 and 4 of the Catalogue of Palaearctic
Coleoptera. The correct publication date is November 1806, according to the Journal général de la Littérature
de France, 1806 (9): 321.

(V) Lacordaire's work: Histoire naturelle des insectes. Genera des coléopères ou exposé méthodique et critique
de tous les genres proposés jusqu'ici dans cet ordre d'insectes. Tome septième contenant les familles des
curculionides (suite), scolytides, brenthides, anthribides et bruchides. Paris: Librairie Roret, 620 pp was
published in 1866 according to the printed date and is usually referred to as from 1866. Based on recent
evidence [provided by A. Davies and Y. Bouchard], it was mentioned on 12 December 1865 in The
Bookseller *S: 1016; as issued in 1865 in the Bull. bibl. Soc. Ent. Fr. (4) 5: lxxx [published May 26, 1866],
and in Cat. Libr. France 8,1865: 126.

(VT) The first edition of Redtenbacher's Fauna Austriaca is assumed to be published in 1849, as given on the
title page of the book. Besides, the preface is dated "März 1849". M. A. Alonso-Zarazaga and H. Schönmann
provided evidences that the work issued in parts. The signatures 1-10: pp. 1-160 were reported on 9-VII-
1847, in Berichte über die Mittheilungen der Freunde der Naturwissenschaften in Wien 3 (1): 58, and on 6-
r VJ.II-1847 in Leipziger Repertorium der deutschen und ausländischen Literatur 5 (3) (32): 233. The contens
of the signatures 11-20: pp. 161-320 were reported on 30-VTI-1847 in Berichte über die Mittheilungen der
Freunde der Naturwissenschaften in Wien 3(1): 111 and notices of the first two parts are in Entomologische
Zeitung (Stettin) 8 (9): 288 and Neue Jahrbücher für Philologie und Paedagogik 51 (4): 155, 1847. The
signatures 21-30, pp. 321-480 were reported on 15-XI-1847, in Berichte über die Mittheilungen der Freunde
der Naturwissenschaften in Wien 3 (4): 311. The signatures 31-40, pp. 481-640 were mentioned on 9-II-1848
in the Literarische Zeitung 16 (13): 203. Finally, the signatures 41-58, pp. i-xxviii, 641-884 are reported in
the beginning of 1849, in the Archiv für Naturgeschichte 15 (2): 127.
 23

ERRATA
The Editors are concerned about the appalling number of corrections and additions submitted by several authors
of the Catalogue and feel that there is a need to address this situation that is out of their control. Many of these
acts should have been included already in the body of the Catalogue in the first place, had the authors followed
the Instructions of the Editors and made proper use of the rules of the zoological nomenclature. The reasons for
this situation are numerous but the most common ones are as follows: 1 ) The inability to correctly assess the
availability/non-availability of names is probably the leading cause, followed closely by 2) insufficient checking
and incorrect interpretation of published information, and 3) copying of old records without proper verification.
Using incorrect date of publication is also common, but most these mistakes are eventually caught and corrected
by the Editors. Needless to say that it adds unwelcomed and unnecessary strain to the already difficult work of
the Editors.

ERRATA for VOLUME 1

p. 26, add original combination (Gyrinus) to Aulonogyrus concinnus Klug, 1833
p. 27, correct author, year and page of Gyrinus from O. F. Müller, 1764: xvii to Gyrinus Geoffroy, 1762: 193
[see ICZN, Opinion 1754]
p. 29, add original combination (Gyrinus) to Orectochilus villosus villosus O. F. Müller
p. 88, correct year and page of Nebria nivalis nivalis Paykull, from 1798: 184 to 1790: 52
p. 181, correct year of angustatus L. Redtenbacher [in synonymy with Carabus sylvestris redtenbacheri Géhin]
to 1847
p. 186, correct year of Carabus coriaceus banaticus L. Redtenbacher to 1847
p. 220, remove entry natalensis erythropyga Putzeys, 1866, and add distributional data to Clivina erythropyga
Putzeys, 1866, p. 219
p. 245, correct year of affine L. Redtenbacher [in synonymy with Bembidion geniculatum geniculatum Heer,
1837] to 1847
p. 250, correct year of nigricolle L. Redtenbacher and obscurum L. Redtenbacher [in synonymy with Bembidion
testaceum testaceum Duftschmid, 1812] to 1847
p. 264, correct year of bipustulatum L. Redtenbacher [in synonymy with Bembidion guttula guttula Fabricius,
1792] to 1847
p. 276, correct spelling of Tachys centromakulatus centromaculatus Wollaston, 1864 to Tachys centromaculatus
centromaculatus
p. 337, correct year of pallescens L. Redtenbacher [in synonymy with Trechus ovatus ovatus Putzeys, 1846] to
1847
p. 381, correct year of alpestris L. Redtenbacher [in synonymy with Harpalus rubripes Duftschmid, 1812] to
1847
p. 419, correct spelling of Dromiina to Dromiusina [see ICZN, Opinion 2149]
p. 784, correct Van de Poll N. to Neervoort van de Poll J.R.H. ['Neervoort van de Poll' is his family name]

ERRATA for VOLUME 2

p. 49, correct year for Enochrus melanocephalus (Olivier) to 1793
p. 54, correct year for Hydrophilus ater Olivier, and oblongus Olivier [in synonymy with Hydrophilus olivaceus
Fabricius, 1781] to 1793
p. 70, correct year for Acritu minutus (Herbst) to 1791, and of granulum L. Redtenbacher [in synonymy with the
latter] to 1847
p. 71, correct year for Plegaderus caesus (Herbst) to 1791
p. 72, Teretrius is of masculine gender. Correct cristatum to cristatus
24 ERRATA
p. 73, correct year for Dendrophilus punctatus (Herbst) and D. dubius (Herbst) [in synonymy with D. pygmaeus
(Linnaeus)] to 1791
p. 75, correct year for Paromalus flavicomis (Herbst) and P. parallelepipedus (Herbst) to 1791
p. 76, correct year for Haeterius Dejean to 1833 [the name appeared in the second edition of Dejean's Catalogue
which was published in separate livraisons: only the first (pp. 1-96), and the second (pp. 97-176) were
published in 1833]
p. 81, correct year for Hister sinuatus Herbst to 1791
p. 82, correct spelling of Hister desbrochers to desbrochersi
p. 83, for Margarinotus stigmosus Marseul [in synonymy with M. punctiventer (Marseul, 1854)] add {Hister)
p. 84, correct year for Margarinotus purpurascens (Herbst) to 1791
p. 87, Eblisia is of feminine gender. Change the ending of following names to feminine: abbreviata, betulina,
delata, frontalis, puncticollis, rufitla, scalpta, silvestris, liopygoformis (not lyopygofromis), taiwana, and
unica
p. 87, correct year and page of Eblisia minor (P. Rossi) to 1792: 13
p. 88, Platylister is of masculine gender. Change the ending of following names to masculine: buteanus,
gibbosus, mimicus, algirus, laevicollis, and quinquestriatus
p. 88, Platysoma is of neuter gender. Change coreana to coreanum
p. 90, correct year for Onthophilus affinis L. Redtenbacher to 1847
p. 95, correct year for Hypocaccus metalliens (Herbst) to 1791
p. 99, correct year for speculifer Latreille [in synonymy with Saprinus politus (Brahm, 1790)] to 1806
p. 102, add to Histeridae, as nomina dubia:
Attelabus cupreus Geoffroy, 1785: 17 E: FR [Attelabus sensu Geoffroy = Hister s.l.; probably a Saprinus]
Hister planus Roemer, 1789: 3 E: "Rhaetia" [Figure in pi. 2, f. 9] [it could be a synonym of Hololepta plana]
p. 122, correct year for ferrarii L. Redtenbacher [in synonymy with Nossidium pilosellum (Marsham, 1802)] to
1847
p. 123, correct entry for Ptenidium penzi to:
penzigi Flach, 1889a: 1 [= 1889b: 496] (Phenidium) E: GR A:CYTR
p. 124, correct spelling of Ptenidium sahlbergi to sahlbergii
p. 124, correct entry for Ptenidium turgitulum to:
turgidulum Flach, 1889a: 1 [= 1889b: 496] {Phenidium) E: AB ST (Kavkaz)
p. 125, remove from synonymy of Ptiliola kunzei (Heer, 1841) flammifera, and add it as validspecies:
flammifera Mtynarski, 1985: 261 (Nanoptilium) E: NL PL
p. 126, correct entry for Ptilium vexans to:
vexans Flach, 1889a: 1 [= 1889b: 504] E: FR IT SP SZ N: TU
p. 128, remove nana A. Strand, 1946 from synonymy with Acrotrichis dispar(A. Matthew, 1865)
p. 129, add in Acrotrichis Motschulsky, 1848 as valid species:
nana A. Strand, 1946a: 170 E:NLNR
p. 130, plumigera Lermina, 1792 [in synonymy with Acrotrichis grandicollis Mannerheim, 1844] seems to be a
later use or combination of:
plumigerus L'Hermina, 1791: 206 (Ips)
p. 130, correct entry for Acrotrichis soror to:
soror Flach 1889a: 2 [= 1889b: 520] (Trichopteryx) E: AB GG A: IN
p. 131, add AFR AUR to distribution of Nephanes titan (Newman, 1834)
p. 133, Apteroloma is of neuter gender. Change ending ofcoreanus to coreanum
p. 133, Pteroloma is of neuter gender. Change ending of following names to neuter: brunneum, japonicum,
magnificum, mirandum [it is from Greek pteron (wing) and loma (fringe), undoubtedly neuter]
p. 138, for Catops kirbyi Spence add (Choleva)
p. 141, correct year for Sciodrepoides fiimatus (Spence) to 1813
p. 143, correct year for testacea Latreille [in synonymy with Choleva agilis (Illiger, 1798)] to 1806
p. 144, correct year for Choleva oblonga Latreille to 1806
p. 180, correct year for Myloechus Latreille and Colon brunneum (Latreille) to 1806
p. 180, for Colon appendiculatum C. R. Sahlberg add (Catops)
p. 180, for Colon dentipes C. R. Sahlberg add (Catops)
p. 181, under genus and subgenus Agathidium Panzer, correct year for Tetratoma globosa Herbst to 1791
 ERRATA 25
p. 186, correct year for Agathidium globosum (Herbst) to 1791
p. 192, correct year for ruficollis Olivier [in synonymy with Amphicyllis globus Fabricius, 1792] to 1790
p. 192, change author, year and original combination of Anisotoma humeralis Fabricius, 1792: 79 (Sphaeridium)
to Herbst, 1791: 86 (Tetratoma) [see O. Vorst, 2010: Coleopterist's Bulletin 62: 185-188]. Anisotoma
humeralis (Fabricius, 1792) is to be treated as a junior synonym of Anisotoma humeralis (Herbst, 1791)
p. 192, correct year for Anisotoma castanea (Herbst), A. clavipes (Herbst), and ,4. orbicularis (Herbst) to 1791
p. 192, remove Anisotoma globosa (Herbst, 1791)fromsynonymy with Anisotoma humeralis. It is a synonym of
Agathidium seminulum (Linnaeus, 1758), p. 186, and is listed as such there
p. 195, for Leiodes ciliaris'W. L. E. Schmidt, 1841 add (Anisotoma)
p. 195, correct year for Leiodes armai a Herbst to 1791
p. 200, correct year and page for Colenis Erichson, 1845: 82 to 1842: 221
p. 201, for Hydnobius latifrons Curtis add (Leiodes)
p. 202, correct year for Triarthron maerkelii to 1840
p. 223, correct spelling of Euthia schaumi Kiesenwetter, 1858 to schaumii
p. 229, correct year for Silphidae and Silphinae Latreille to 1806
p. 234, add under Silphinae, as nomen dubium:
bipunctata Fabricius, 1787: 45 (Ips) E: "Europa australiore" [transferred to Silpha by De Villers, 1789; probably
a "clavicorn"]
p. 234, remove NLfromdistribution ofNicrophorus antenna tus Reitter, 1885
p. 249, remove NLfromdistribution of Olophrum assimile Paykull, 1800
p. 251, for Phyllodrepoidea crenata Ganglbauer, 1895 remove (Omalium)
p. 299, remove NL from distribution of Brachygluta xanthoptera (Reichenbach, 1816)
p. 327, correct year for Pselaphus Herbst to 1791
p. 339, delete "h" in Coprophorus
p. 359, the valid name for Aleochara kamila Likovsky, 1984 is A. diversa J. R. Sahlberg, 1876: 77 (Baryodma),
currently listed as synonym. Aleochara diversa (J. R. Sahlberg, 1876) is not a primary homonym (as it was
described in Baryodma), but a secondary homonym of Aleochara diversa Mulsant & Rey, 1853 (=
Thiasophila inquilina (Märkel, 1845). As the replacement name Aleochara kamila was (unnecessarily)
introduced after 1960 and both diversa''s are not congeneric, the name Aleochara diversa (J. Sahlberg, 1876)
is valid
p. 370, for Amischa cavifrons Sharp, 1869 [in synonymy with.4. bifoveolata (Mannerheim)] add (Homalota)
p. 374, correct year for Atheta divisa Märkel to 1844. MärkePs Beiträge zur Kenntnis der unter Ameisen
lebenden Insecten (ZeitschriftfiirEntomologie 5: 193- 271) was published before August 20, 1844 (on this
date Band 5 of ZeitschriftfiirEntomologie arrived at the library of the Stettiner Entomologischer Verein (see
Entomologische Zeitung (Stettin) 5: 505)
p. 407, remove Halobrecta algophila (Fenyes) from synonymy with HalobrectaflavipesThomson and enter the
following valid species and synonyms under Halobrecta:
algophila Fenyes, 1909: 419 (Atheta) E:FRGB AUR NAR NTR
barbarae Casey, 1910a: 18 (Atheta)
importuna Casey, 1911: 111 (Atheta)
p. 407, remove Halobrecta algae Hardy, 1851: 78 (Homalota)fromsynonymy with Halobrecta puncticeps
Thomson, 1852: 133 (Homalota), and enter the latter as its junior synonym
p. 409, remove Hydrosmecta septentrionum G. Benick from synonymy of Hydrosmecta subtilissima Kraatz and
enter the following under Hydrosmecta:
septentrionum G. Benick, 1969: 169 (Atheta) E: DE NL NR [described from "Rundhaug M. Elv"., which is
Mâlselv area in northern Norway]
p. 413, correct year for Notothecta confusa Märkel to 1844
p. 419, move Taxicera renneri G. Benick, 1982b: 127 as a synonym of T. sericophila Baudi di Selve, 1870: 379
[note: p. 414 is wrong]
p. 420, correct year for Thamiaraea hospita Märkel to 1844
p. 427, correct year for Bolitochara bella Märkel to 1844
p. 441, for Agaricochara Kraatz, correct year of type species Gyrophaena laevicollis Kraatz to 1854
p. 453, for Cypha suecica Palm remove "v" in "(Hypocvyptus)"
p. 457, correct year for Lomechusoides strumosus strumosus Fabricius to 1792
26 ERRATA
p. 460, correct year for Zyras laticollis Märkel to 1844
p. 461, correct year for Zyras similis Märkel to 1844
p. 472, for Dexiogyiaforticornis A. Strand, 1939 add {Stichoglossà)
p. 473, correct year for Haploglossa gentilis Märkel to 1844
p. 474,: genus Mniusa, correct spelling of "Mulsnat" to Mulsant
p. 476, genus Oxypoda, correct year of type species Aleochara spectabilis Märkel to 1844
p. 481, correct year for Oxypoda spectabilis Märkel to 1844
p. 484, correct year for Oxypoda vicina Kraatz to 1858b
pp. 495,496, correct year for Scaphidiinae Latreille and Scaphidiini Latreille to 1806
p. 512, genus Syntomium Curtis, correct type species to: Syntomium nigroaeneum Curtis, 1828 (= Omalium
aeneum P. W. J. Müller, 1821)
p. 537, genus Dianous Leach, correct type species to: Stenus coerulescens Gyllenhal, 1810
p. 537, for Dianous coerulescens Gyllenhal add (Stenus)
p. 638, genus Neobisnius Ganglbauer, correct type species to: Gabrius villosulus Stephens, 1833
p. 656, genus Quedius Stephens, correct type species to: Staphylinus levicollis Brullé, 1832
p. 669, for Quedius semiaeneus Stephens, 1833 add (Raphirus)
p. 671, genus Creophilus and Saphrophilus, correct year of the type species Staphylinus maxillosus Linnaeus to
1758
p. 745, add: Erichson W. F. 1842: Bericht über die wissenschaftlichen Leistungen im Gebiete der Entomologie
während des Jahres 1841. Archiv fia- Naturgeschichte 1842: 189-330 [also published separately pp. 1-142,
dated 1843]
p. 735, add: Herbst J. F. W. 1791: Natursystem aller bekannten in- und ausländischen Insecten, als eine
Fortsetzung der von Büffonschen Naturgeschichte. Der Käfer vierter Theil. Heft 1. Berlin: J. Pauli, [1] + 112
pp., pis F + xxxv-xxxix [see Coleopterist's Bulletin 62: 185-188]
p. 755, add: Flach C. 1889a: Kurze Diagnosen neuer Trichopterygier. Societas Entomologica 4: 1 -2. [issued in
April]
p. 755, correct reference for Flach, 1889: Bestimmungstabelle ... to Flach, 1889b: Bestimmungstabelle ... [issued
in December]
p. 792, Klug 1832. The work was first published separately, as follows:
Klug J. F. C. 1832: Bericht über eine auf Madagascar veranstaltete Sammlung von Insecten aus der Ordnung
Coleoptera. Königliche Akademie der Wissenschaften, Berlin, 132 pp., 5 pis. Issued later, in 1834, in the
Abhandlungen... (Physikalische Klasse) 1832, pp. 91-223, 5 pis.
p. 802, add: L'Hermina C. 1791: [new taxon]. In: Fourcroy A. F. de.: Zoologie. La médecine éclairée 1 (7): 204-
207
p. 821, in Mulsant & Rey 186 la and 1861b, add" 1 pi."
p. 821, in Mulsant & Rey 1861c, correct the title as follows: Description de quelques coléoptères nouveaux ou
peu connus et de deux genres nouveaux
p. 821, Mulsant & Rey 1872, change the year to 1871 and the title to: Histoire naturelle des Coléoptères de
France. Brévipennes Aléochariens
p. 821, Mulsant & Rey 1874a change the year to 1873c
p. 821, 822: Mulsant & Rey, 1876, 1877, 1878a, 1878b, 1878c, 1880, insert in front of each [Histoire naturelle
de Coléoptères de France]
p. 830, correct year for Olivier A. G. 1792: ... to 1793, and add Pars I, with pagination 1-368 [pp. 369-827 in
Pars II issued in 1797]
p. 840, correct year for Panzer G. W. F. 1805:... to 1803
p. 841, correct year for Panzer G. W. F. 1809:... to 1805
p. 864, add: Roemer J. J. 1789: Genera insectorum Linnaei et Fabricii iconibus illustrata a Joanne Jacobo
Roemer. Vitoduri Helvetorum: apud Henrjc. Steiner et socios, viii + 86 + [3] pp., 37 pis

ERRATA for VOLUME 3

p. 63, correct year for Platycerus Latreille [in synonymy with Ceruchus MacLeay, 1819] to 1806
p. 63, add entry: Tarandus Dejean, 1833: 174 [HN] type species Lucanus tenebrioides Fabricius, 1787 (= L.
 ERRATA 27
chrysomelinus Hochenwarth, 1785) in synonymy with Ceruchus MacLeay, 1819. Lucanus tenebrioides
Fabricius, 1787 is the originally included available species; Tarandus Dejean is a preoccupied by Tarandus
Billberg, 1827 [Mammalia]
p. 97, correct year for Amphicoma Latreille to 1806
p. 103, correct year for Aegialia Latreille to 1806
p. 106, correct year for rufitarsis Latreille [ia»ynonymywith.4/jAo<#ws luridus Fabricius, 1775] to 1806
p. 158, correct author and data of Bubas to Bubas Dejean, 1833: 143. "Bubas. Megerle." was introduced with
originally included available species: Scarabaeus bison Linnaeus, 1767 (as "Bison. Fabr."); Onitis bubalus
Olivier, 1812 as "Bubalus. Latreille."
p. 178, correct year for Sisyphus Latreille to 1806
p. 180, correct author and data of Caelodera [in synonymy with Pachypus Dejean, 1821] to Caelodera Dejean,
1833: 159. Originally included species are: Scarabaeus excavatus Fabricius, 1792 and Melolontha cornuta
Olivier, 1789. The taxon "Cornuta. Olivier." was originally listed as a synonym of "Excavata. Fabr.";
therefore the type species of the genus is Scarabaeus excavatus by monotypy
p. 186, correct year for formosa Latreille [in synonymy with Hoplia coerulea Drury, 1773] to 1806
p. 195, correct year for albida L. Redtenbacher [in synonymy with Melolontha melolontha (Linnaeus, 1758)] to
1847
p. 218, add entry: Ancylonycha Dejean, 1833: 160 type species Melolontha serrata Fabricius, 1787 in synonymy
with Holotrichia Hope, 1837. Originally included species in Ancylonycha are: Melolontha leucophthalma
Wiedemann, 1819; M. serrata Fabricius, 1787; M. fervida Fabricius, 1775; M. knochii Schönherr and
Gyllenhal, 1817 (as "Knochii. Schönherr"); M. quercina Knoch, 1801; M. ilicis Knoch, 1801; M. ilicis
Knoch, 1801. Melolontha serrata Fabricius, 1787 is the type species by subsequent designation in
Duponchel (1840: 481). Ancylonycha Blanchard, 1845 has been treated as a synonym of Phyllophaga in the
New World. Blanchard (1845: 216) provided the first description of Ancylonycha and designated "A pubera
Dej." as the type of his genus (Blanchard, 1845: 245) but this is an invalid type species designation since the
species name was not available. Reversal of Precendence (Art. 12. 23.9) cannot be used because
Ancylonycha was used as valid after 1899 (with Holotrichia as synonym) at least once by Saylor (1942: 157)
p. 266, add entry: Anisonchus Dejean, 1833: 157 type species Melolontha atriplicis Fabricius, 1787. Melolontha
atriplicis Fabricius, 1787 is the type species by monotypy. Anisonchus is junior objective synonym of
Hoplopus Laporte, 1832 in Scarabaeidae (fide Hope, 1837: 70)
p. 280, correct data for Calicnemis latreillii Laporte, 1832a to ix, pi. 7, and remove truncatifrons Laporte, 1832
from its synonymy. The latter name does not exist
p. 280, correct type species of Heteronychus Dejean, 1833 is Scarabaeus syrichtus Fabricius, 1775, by
subsequent designation in Duponchel (1845: 601). Krell (2002: 40) invalidly designated Geotrupes cricetus
Hausmann, 1807 as the type species of Heteronychus Dejean and used the same species as the type in the
Catalogue of Palearctic Coleoptera (Krell, 2006: 280). Unfortunately the valid type species for Heteronychus
Dejean, S. syrichtus Fabricius, is currently placed in the genus Syrictes Prell, 1936 (e.g., Endrödi, 1985:
685). An application should be submitted to the Commission in order to keep using the current taxonomic
concepts of Heteronychus and Syrictes. It appears that Syrictes Prell is itself a preoccupied genus name (not
Syrictes Jordan & Evermann 1927 [Pisces]) and needs a replacement name
p. 282, correct year for Pendoton quadridens Gebier to 1845
p. 284, remove entry Atropinota Heller, 1923, add the genus-group name in synonymy with Bietia Fairmaire,
1898, and placefiinkeiHeller, 1923 as valid species in Bietia, p. 301
p. 313, add under Scarabaeidae, as nomen dubium:
Scarabaeus pellucidulus Roemer, 1789: 2 E: SZ [Figure in pi. 1, f.9; mentions an unavailable name given by
Voet]
p. 314, correct year for Calyptomerus L. Redtenbacher and C. alpestris L. Redtenbacher to 1847
p. 316, correct year for Clambus pubescens L. Redtenbacher to 1847
p. 388, correct year for aurichalceiis L. Redtenbacher [in synonymy with Agrilus cuprescens cuprescens
(Ménétriés, 1832)] to 1847
p. 389, correct year for cupreus L. Redtenbacher [in synonymy with Agrilus integerrimus (Ratzeburg, 1837)] to
1847
p. 389, correct year for bicolor and quercimts L. Redtenbacher [in synonymy with Agrilus viridis viridis
(Linnaeus, 1758)] to 1847
28 ERRATA
p. 390, correct year for Agrilus convexicollis L. Redtenbacher to 1847
p. 437, correct year for dargelasi Latreille [in synonymy with Oulimnius tuberculatus (P. W. J. Müller, 1806)] to
1806
p. 443, correct year for dumerilii Latreille [in synonymy with Pomatinus substriatus (P. W. J. Müller, 1806)] to
1806
p. 451, change author ofEubria to Germar, 1818: 239; the type species remains
p. 517, correct year for Gebier A. F. von 1844: Characteristic ... to 1845
p. 560, correct year for Latreille P. A.: Genera crustaceorum et insectorum ... Tomus secundus from 1807 to
1806.
p. 574, add: Medvedev S. I. & Nikolaev G. V. 1972: Novyy vid roda Codocera Eschsch. (Coleoptera,
Scarabaeidae) iz Central'noy Azii [A new species of the genus Codocera Eschsch. (Coleoptera,
Scarabaeidae) from Central Asia]. Nasekomye Mongolii 1: 445-446.
p. 633, add: Roemer J. J. 1789: Genera insectorum Linnaei et Fabricii iconibus illustrata a Joanne Jacobo
Roemer. Vitoduri Helvetorum: apud Henrjc. Steiner et socios, viii + 86 + [3] pp., 37 pis.
p. 649, add: Solsky S. M. 1873: [new taxon]. In: Hochhut I. H. 1873: Enumeration der in den russischen
Gouvernements Kiew und Volhynien bisher aufgefundenen Käfer. Bulletin de la Société Impériale des
Naturalistes de Moscou 46 (1): 124-164.

ERRATA for VOLUME 4

p. 117, correct year for graminicola L. Redtenbacher [in synonymy with Agriotes sputator (Linnaeus, 1758)] to
1847
p. 165, correct year for Limoniscus suturalis (Gebier) to 1845
p. 190, correct year for elongatus L. Redtenbacher [in synonymy with Oedostethus tenuicornis (Germar, 1824)]
to 1847
p. 215, correct year for flavescens L. Redtenbacher [in synonymy with Erotides cosnardi (Chevrolat, 1831)] to
1847
p. 237, change author of Podabrus to Dejean, 1833: 105; the type species remains
p. 245, correct year for dilatata L. Redtenbacher [in synonymy with Cantharis quadripunctata (O. F. Müller,
1776)] to 1847
p. 245, correct year for nivalis L. Redtenbacher [in synonymy with Cratosilis denticollis (Schummel, 1846)] to
1847
p. 245, correct year for humeralis L. Redtenbacher and liturata L. Redtenbacher [in synonymy with
Metacantharis discoidea (Ahrens, 1812)] to 1847
p. 263, correct year for paludosa L. Redtenbacher [in synonymy with Rhagonycha elongata (Fallen, 1807)] to
1847
p. 263, correct year for terminalis L. Redtenbacher [in synonymy with Rhagonycha fiilva (Scopoli, 1763)] to
1847
p. 276, change type species of Silis Charpentier, 1825 to Silis rugicollis Gebier, 1833 (= Cantharis ruficollis
Fabricius, 1775). The history of the case and the consequent changes given by S. Kazantsev are as follows:
Charpentier, 1825 included originally in Silis two species, S. spinicollis Charpentier, 1825 (= Cantharis nitidida
Fabricius, 1792) and S. rubricollis Charpentier, 1825 (= Cantharis ruficollis Fabricius, 1775), not fixing the
type species. Westwood (1838) subsequently designated Cantharis ruficollis as the type species for Silis,
while Green (1966) designated C. nitidula as the type species of Silis. Green's designation, however, was
used in the Catalogue of Cantharidae of the World (Delkeskamp, 1977). Kazantsev, 1994 showed that Silis
ruficollis and S. nitidula belong to different genus-group taxa, and established a new genus, Crudosilis
Kazantsev, 1994, for the one that included S. ruficollis. Thus, the designation of Cantharis ruficollis as the
type species of Silis was overlooked by Green (1966), Delkeskamp (1977) and Kazantsev (1994). As
Crudosilis altaica Kazantsev, 1994, the type species of Crudosilis, is considered congeneric with Silis
ruficollis, Crudosilis Kazantsev, 1994, is a new synonym of Silis. For Silis auct, that includes Cantharis
nitidula Fabricius, 1792, a new genus is established:
Autos ilis Kazantsev, gen. nov.
type species Cantharis nitidula Fabricius, 1792
 ERRATA 29
Autosilis gen. nov. is distinguished from Silis by the absence of pores in the anterior half of the dorsal surface of
the pronotum, the approximately equal width of the ventrite and tergite of the genital capsule, and by the
conspicuous elongate laterophyses of the aedeagus
The resulting new combinations are as follows:
Autosilis amurensis (Kazantsev, 2008), comb. nov.
Autosilis bianchii (Barovskij, 1926), comb. nov.
Autosilis bifossicollis (Wittmer, 1963), comb. nov.
Autosilis dzungarica (Kazantsev, 1994), comb. nov.
Autosilis jacutica (Barovskij, 1926), comb. nov.
Autosilis nitidula (Fabricius, 1792), comb. nov.
Autosilis olschwangi (Kazantsev, 1994), comb. nov.
Autosilis telephoroides (Pic, 1913), comb. nov.
Autosilis triimpressa (Pic, 1926), comb. nov.
Autosilis tuvensis (Kazantsev, 2008), comb. nov.
Autosilis urjanhaica (Kazantsev, 1997), comb. nov.
Silis alaica (Kazantsev, 1998), comb. nov.
Silis altaica (Kazantsev, 1994), comb. nov.
Silis barovskii (Kazantsev, 1994), comb. nov.
Silis beringiana (Kazantsev, 1994), comb. nov.
Silis dowgailoi (Kazantsev, 1998), comb. nov.
5/7« euphrosyne Barovskij, 1928, comb. nov.
Silis ferganica (Kazantsev, 1998), comb. nov.
5/7« lomakini (Chernyshev, 1998), comb. nov.
Silis magadanica (Kazantsev, 1998), comb. nov.
Silis mingrelica (Kazantsev, 1994), comb. nov.
Silis mongolica (Kazantsev, 1998), comb. ntov.
Silis motschulskii Barovskij, 1928, comb. nov.
Silis ruficollis (Fabricius, 1775), comb. nov.
5/7« rugimana (Kazantsev, 1994), comb. nov.
5/7« sandalica (Kazantsev, 1998), comb. nov.
Silis sexdentata Mannerheim, 1853, comb. nov.
Silis sinica (Kazantsev, 1998), comb. nov.
Silis tchernyshevi (Kazantsev, 1998), comb. nov.
Silis tolli Barovskij, 1928, comb. nov.
Silis turkestanica (Kazantsev, 1998), comb. nov.
Silis wittmeri (Kazantsev, 1998), comb. nov.
Silis xanthoderus (Solsky, 1881), comb. nov.

Folowing data replace those on pp. 274 and 276, for Crudosilis and Silis:
genus Autosilis Kazantsev, 2011: 26 type species Cantharis nitidula Fabricius, 1792
amurensis Kazantsev, 2008: 337 (Silis) A: ES (Amur)
bianchii Barovskij, 1926a: 13 (5/7«) A: ES
bifossicollis Wittmer, 1963: 114 (Silis) A: "Central China"
dzungarica Kazantsev, 1994c: 102 (Silis) A: KZ
jacutica Barovskij, 1926a: 13 (Silis) A: ES FE
nitidula Fabricius, 1792a: 220 (Cantharis) E: AU CT CZ EN FR GE HU LA LT PL RO SK UK
excisa Germar, 1824: 71 (Telephorus)
spinicollis Charpentier, 1825: 194(5/7«)
olschwangi Kazantsev, 1994c: 99 (5/7«) A: WS
telephoroides Pic, 1913d: 98 (Silis) A: PA
triimpressa Pic, 1926a: 1 (5/7«) A: FE "Korea"
tuvensis Kazantsev, 2008: 338 (5/7«) A: ES (Tuva)
urjanhaica Kazantsev, 1997b: 1216(5/7«) A: ES
30 ERRATA
genus Silis Charpentier, 1825:194 type species Cantharis ruficollis Fabricius, 1775: 206
Crudosilis Kazantsev, 1994: 102 type species Crudosilis altaica Kazantsev, 1994
alaica Kazantsev, 1998a: 289 (Crudosilis) A: Kl
altaica Kazantsev, 1994c: 110 (Crudosilis) A: WS
barovskii Kazantsev, 1994c: 113 (Crudosilis) A: KZ
beringiana Kazantsev, 1994c: 110 (Crudosilis) A: FE
dowgailoi Kazantsev, 1998a: 290 (Crudosilis) A: Kl
euphrosyne Barovskij, 1928b: 219 A: TD
ferganica Kazantsev, 1998a: 292 (Crudosilis) A: Kl
lomakini Chemyshev, 1998: 240 (Crudosilis) A: ES
magadanica Kazantsev, 1998a: 286 (Crudosilis) A: FE
mingrelica Kazantsev, 1994c: 109 (Crudosilis) E: GG
mongolica Kazantsev, 1998a: 286 (Crudosilis) A: ES MG
motschulskii Barovskij, 1928b: 218 A: ES
bidentata Motschulsky, 1859d: 568 [HN]
ruficollis Fabricius, 1775: 206 (Cantharis) E: AU BE BY CT CZ DE FI FR GB GE HU IR IT LA LS LT NL
PL RO SK SV SZ UK A: KZ WS
bidentata O. F. Müller, 1776: 62 (Telephons)
gallica Pic, 1927h: 13
hungarica Pic, 1910i: 33
rubricollis Charpentier, 1825: 195
rugicollis Gebier, 1833: 279
signicollis Everts, 1920: 230
torquata Gyllenhal, 1827: 340 (Cantharis)
rugimana Kazantsev, 1994c: 107 (Crudosilis) A: FE
sandalica Kazantsev, 1998a: 292 (Crudosilis) A: Kl
sexdentata Mannerheim, 1853: 295 A: CH ES FE MG
sinica Kazantsev, 1998a: 288 (Crudosilis) A: QIN
tchernyshevi Kazantsev, 1998a: 287 (Crudosilis) A: ES
tolli Barovskij, 1928b: 220 A: FE
turkestanica Kazantsev, 1998a: 291 (Crudosilis) A: Kl
wittmeri Kazantsev, 1998a: 288 (Crudosilis) A: GAN
xanthoderus Solsky, 1881: 35 (Silotrachelus) A: Kl
p. 288, correct year for pulicarius L. Redtenbacher [in synonymy with Malthodes dimidiaticollis dimidiaticollis
(Rosenhauer, 1847)] to 1847
p. 303, correct year for Thorictus fairmairii tunisius Chobaut to 1925
p. 304, correct year for Thorictus normandi Chobaut to 1925
p. 309, correct year for atra Herbst [in synonymy with Attagenus pellio (Linnaeus, 1758)] to 1791
p. 310, correct year for Attagenus schaefferi Herbst to 1791
p. 310, correct year for brevicornis Herbst [in synonymy with Attagenus unicolor (Brahm, 1790)] to 1791
p. 316, correct year for Megatoma Herbst to 1791
p. 319, correct year for ruficornis Latreille [in synonymy with Trogoderma glabrum (Herbst, 1783)] to 1806
p. 320, correct year for Dictyalotus L. Redtenbacher [in synonymy with Endecatomus Mellié, 1847] to 1847
p. 322, correct year for dufourii Latreille [in synonymy with Lichenophanes varius (Illiger, 1801)] to 1806
p. 341, correct year for Trypopitis L. Redtenbacher [in synonymy with Priobium Motschulsky, 1845] to 1847
p. 342, correct year for Oligomerus L. Redtenbacher to 1847
p. 344, correct year for Dorcatoma Herbst to 1791
p. 350, correct year for politus L. Redtenbacher [in synonymy with Ernobius nigrimis (Sturm, 1837)] to 1847
p. 365, add under Tenebroides valid species:
ocularis Lewis, 1894: 34 A: JA
p. 366, change author ofPeltis to Kugelann, 1792: 508
p. 383, add under Cleridae, as nomen dubium:
Attelabus hungaricus Scopoli, 1772: 92 E: HU [Annus V] [probably Trichodes]
p. 387, add under Falsomelyris valid species:
priesneri Pic, 1939g: 145 N: EG
 ERRATA 31
p. 400, correct page for Allotarsus Graells, 1858 to 52
p. 400, use Allotarsus senex (Rosenhauer, 1856) as valid, and E. melaleucotrichos Graells, 1858 as its invalid
synonym
p. 408, correct page for amphicoma (Graells, 1858) [in synonymy with Enicopus scutellartis (Fabricius, 1792)]
to 50
p. 411, correct year for pini L. Redtenbacher and serratus L. Redtenbacher [in synonymy with Aplocnemus
impressus Marsham, 1802)] to 1847
p. 416, correct year for Apalochrus fiilvicollis (Gebier) to 1845
p. 417, correct data for Laius Guérin-Méneville to 1830b: pi. 2, and correct year for Laius cyaneus Guérin-
Ménevilleto 1830
p. 421, under Attalus, add [= 1900: 339] to entry atrocyaneus Uhagôn, 1898
p. 422, under Attalus, add [= 1900: 341] to entry laufferi Uhagön, 1898
p. 427, under Colotes, correct data of hispanus Uhagôn to: 1898: 41 [= 1900: 39]
p. 485, add laevis Geoffroy, 1785: 137 (Anthribus) in synonymy with Meligethes pedicularius Gyllenhal, 1808 [
it has priority, the synonymy was given in Desportes, 1820, Analyse travaux Soc. Roy. des Arts de Mans, p.
180]
p. 493, correct year for quisquiliarum L. Redtenbacher [in synonymy with Monotonia bicolor A. Villa & G. B.
Villa, 1835] to 1847
p. 494, correct year for rufa L. Redtenbacher [in synonymy with Monotonia testacea Motschulsky, 1845)] to
1847
p. 508, add bimaculatus Geoffroy, 1785: 137 {Anthribus) in synonymy with Olibrus bicolor (Fabricius, 1792) [it
has priority, the synonymy was given in Schoenherr, 1806, Syn. Ins. p. 109]
p. 510, correct year for ater Herbst [in synonymy with Phalacrus corruscus Panzer, 1797] to 1791. This name,
as that of Sphaeridium assimilis P. Rossi, 1794, should be declared nomina oblita, and P. corruscus Panzer
should be declared nomen protectum
p. 516, remove subinermis Rey, 1889 from synonymy of Cryptophagus distinguendus Sturm, 1845; it is
correctly listed under C. pallidus Sturm, 1845
p. 519, remove convexus Rey, 1889, rugicollis Rey, 1889, subhirtus Rey, 1889 and verrucifer Rey, 1889 from
synonymy of Cryptophagus pallidus Sturm, 1845; they are correctly listed under C. distinguendus Sturm,
1845
p. 521, add under Cryptophagus, nomina dubia:
argenteus Joy, 1909: lxv
p. 531, add under Cryptophagidae, as nomen dubium:
Ips cimeterius Schrank, 1798: 512 E: GE (Bavaria) [Originally stated to belong to Cryptophagus]
p. 555, change author of Sphaerosoma to Stephens, 1832: 391 with type species Sphaerosoma quercus Stephens,
1832 (= Tritoma pilosum Panzer, 1793)
p. 555, change author of Sphaerosoma quercus from Samouelle to Stephens, 1832: 391 and place it as a
synonym of pilosum Panzer, 1793
p. 559/560, correct year for Symbiotes L. Redtenbacher and S. latus L. Redtenbacher to 1847
p. 590, replace entry bimaculatus Herbst, 1792: 71 (Hydrophilus) [in synonymy with Scymnus interruptus
(Goeze, 1777)] by "bimaculatus Herbst, 1797: 340 [HN]". The former name was published in 1791, as
Sphaeridium, it is a primary homonym of Sphaeridium bimaculatum Herbst, 1783 and of unknown identity.
Sphaeridium bimaculatum Herbst, 1783 is a synonym of Tritoma bipustulata (Fabricius, 1775)
p. 603, remove tripunctata Roemer, 1789 from synonymy of Adalia bipunctata (Linnaeus, 1758) [the description
by Roemer mentions Linnaeus, Systema Naturae, so it is a misidentification]
p. 633, correct year for Gryphinus L. Redtenbacher [in synonymy with Sericoderus Stephens, 1829] to 1847
p. 642, correct year for Corticaria bella L. Redtenbacher to 1847
p. 646, correct year for trifoveolata L. Redtenbacher [in synonymy with Corticarina minuta (Fabricius, 1792)] to
1847
p. 648, add under Cucujoidea, as nomina dubia [probably Nitidulidae or Phalacridae; Geoffrey's collection is
extant]:
Anthribus connexus Geoffroy, 1785: 138 E: FR
Anthribus fulvus Geoffrey, 1785: 138 E: FR
32 ERRATA
Anthribus intersect™ Geoffroy, 1785: 139 E: FR [Phalacrus according to Desportes, 1820, Analyse travaux
Soc. Roy. des Arts de Mans, p. 180]
Anthribus nitidus Geoffroy, 1785: 138 E: FR
Anthribus pallidus Geoffroy, 1785: 139 E: FR
p. 680, correct year for Chobaut A. 1924c to 1925
p. 688, add: Delkeskamp K. 1977: Cantharidae. Pars 165. Fasc. 1. Editio secunda. In: Wilcox J. A. (ed.):
Coleopterorum Catalogus Supplementa. The Hague: W. Junk. 485 pp.
p. 716, correct year for Gebier F. A. 1844: Characteristik... to 1845
p. 723, add: Green J. W. 1966: Revision of the Nearctic species of Silis. Proceedings of the California Academy
of Sciences 32 ( 16): 447-513.
p. 723, correct title and editor for Grimmer, 1841 to: Steiermark's Coleoptern mit Einhundert sechs neu
beschriebenen Species. Grätz: C. Tanzer.
p. 726, add: Guérin-Méneville F. E. 1830b: Crustacés, Arachnides et Insectes. Pis 1, 2, 15-17. In: Duperrey L. I.
(ed.): Voyage Autour du Monde, exécuté par ordre du Roi, sur la Corvette de la Majesté, La Coquille,
pendant les années 1822, 1823, 1824 et 1825, sous le Ministère et conformément aux instructions de S. E. M.
Le Marquis de Clermont-Tonnerre, Ministre de la Marine; et publié sous les auspices de son Excellence Mgr
le Cre de Chabrol, Ministre de la Marine et des Colonies. Zoologie. Atlas. Paris: Arthus Bertrand,
p. 735, add: Herbst J. F. W. 1791: Natursystem aller bekannten in- und ausländischen Insecten, als eine
Fortsetzung der von Büffonschen Naturgeschichte. Der Käfer vierter Theil. Heft 1. Berlin: J. Pauli, [1] " 112
pp., pis F + xxxv-xxxix.
p. 735, correct for Herbst J. F. W. 1792: Natursystem ... "Heft 2". Berlin, J. Pauli, viii " 113-197 pp., pis H + xl-
xliii.
p. 749: add: Joy N. H. 1909: New British Coleoptera. The Transactions of the Entomological Society of London
1909-1910: brv.
p. 888, add volume nr. 31 to Isis von Oken in Waltl J. 1838, and 32 to Waltl J. 1839
p. 888, correct reference for Waltl J. 1838c to: Beiträge zur Kennmiss der Coleopteren der Türken. Isis von Oken
(31) 6 [1838]: 449-472.

ERRATA for VOLUME 5

p. 51, add marginalis Panzer, 1792a: 24 (Ips) in synonymy with Litargus connexus (Geoffroy, 1785)
p. 52, add. EN to distribution of Mycetophagus ater (Reitter, 1879)
p. 54, add EN to distribution of Triphyllus bicolor (Fabricius, 1777)
p. 62, correct year for dermestoides Herbst [in synonymy with Tetratoma fungorum Fabricius, 1790] to 1791
p. 65, add EN to distribution of Anisoxya fuscula (Illiger, 1798)
p. 70, add "[NP]" to Orchesia micans Panzer, 1793
p. 70, add "[NO]" to picea Herbst [in synonymy with Orchesia micans Panzer, 1793] and correct year to 1791
p. 72, add DE to distribution oîZilora obscura (Fabricius, 1794)
p. 74, correct year for Pelecotoma leachii Latreille and Pelecotoma dufourii Latreille to 1818
p. 79, add linearis Rossi, 1790: 50 (Ips) [in synonymy with Colydium elongatum (Fabricius, 1787)]; in Bertolini,
1872] '
p. 87, add under Zopheridae, as nomen dubium
Ips bifasciatus Schrank, 1798: 513 E: GE (Bavaria) [originally nr. Bitoma]
p. 91, add Ermisch, 1950 as author of Paratomoxia
p. 105, add under Mordellidae, nomen dubium:
Mordellaperlata Roemer, 1789: 8 E: SZ [pi. 7, fig. 14]
p. 124, correct year for Adesmia gebleri Gebier to 1845
p. 126, correct year for truncata Gebier [in synonymy with Cyphogenia aurita (Pallas, 1781)] to 1845
p. 144, correct distribution of Erodius bicostatus costatulus Koch, 1939 to N: AG
p. 144, add to Erodius (Dimeriseis) valid species:
chudeaui Koch, 1941:256 N: AG AFR
p. 148, correct year for Leptonychus pellucidus pellucidus Mulsant & Godart to 1866
p. 155, correct year for dayensis Mulsant & Godart [in synonymy with Pachyscelodes tuberculifera tuberculifera
 ERRATA 33
(Lucas, 1858)] to 1866
p. 155, correct year for punctata Gebier [in synonymy with Pelorocnemis punctigera (Ménétriés, 1849)] to 1845
pp. 170/171, Trachyderma is neuter: change grammatical gender of the species-group names from feminine to
neuter
p. 172, correct year for Trigonoscelis nodosa schrenkii Gebier to 1845
p. 178, correct spelling and author of Stenosis brenthoides Rossi to S. brentoides Bosc d'Antic, 1790
p. 179, remove entry canaliculata Miller, 1861 as valid taxon (the name is correctly listed as synonym of
Stenosis fidvipes Reiche & Saulcy, 1857)
p. 190, correct year for Gnathosia schrenkii Gebier to 1845
p. 204, correct year for Scythis tatarica Gebier to 1845
p. 207, correct year for laevicollis Gebier [in synonymy with Tentyria gigas Faldermann, 1836] to 1845
p. 212, correct year for nitida Gebier [in synonymy with Zophosis punctata Brullé, 1832] to 1845
p. 222, correct year for caudata Gebier [in synonymy with Blaps granulata turcomana Fischer von Waldheim,
1843] to 1845
p. 227, add to Blaps:
thibetanoides Ren, 2004: 69, 72 A: XIZ
233, correct year for brevis Gebier [in synonymy with Prosodes philacoides Fischer von Waldheim, 1844] to
1845
p. 239, add "JA" to distribution of Byrsax spiniceps Lewis, 1894
p. 240, remove entry badghysi G. S. Medvedev, 1964 from subgenus Dissonomus Jacquelin du Val, 1861; the
species is correctly listed under subgenus Paradissonomus G. S. Medvedev, 1968
p. 257, delete "SC" from distribution of Tarpela elegantula Lewis, 1894
p. 257, add "SC" to distribution of Tarpela lewisi Masumoto, 1993
p. 258, add "TAI" and "ORR" to distribution of Diphyrrhynchus shibatai Kaszab, 1964
p. 260, correct spelling of Brachydium iwojimae Masumoto, 1991 to iwojimanum
p. 264, in distribution of Gonocephalum miyakense Nakane, 1963, change "(Ryukyus)" to "(Izu Isis.)"
p. 276, correct spelling of Paloropsis irei Masumoto & Grimm, 2004 to "iriei"
p. 277, remove entry crenulatus Ménétriés, 1832 from subgenus Dandarus Dejean, 1821; the species is correctly
listed under subgenus Pandarinus Mulsant & Rey, 1854
p. 280, correct year for Heliopates batnensis Mulsant & Godart to 1866
p. 282, correct spelling oïPhylan abreviatus hamilcaris Reitter, 1904 to abbreviatus hamilcarus
p. 286, correct year for Melambius teinturieri Mulsant & Godart to 1866
p. 286, correct year for Otinia incerta incerta (Mulsant & Godart) to 1866
p. 298, correct year for Scaurus elongatus Mulsant & Godart to 1866
p. 303, add to Uloma valid species:
metogana Ren, 2004: 69, 71 A: XIZ
p. 307, correct year for populi L. Redtenbacher [in synonymy with Alphitophagus bifasciatus (Say, 1824)] to
1848
p. 307, add to Ceropria valid species:
yaoi Ren, 2004: 68, 71 A: XIZ [note: this species was described as "Coropes" (in the same paper also spelled
"Corops") that does not exist. It is obviously a misspelling of Ceropria Laporte & Brullé, 1831, because this
new species was compared to subnigra Gebien, 1925, and that is a member of Ceropria]
p. 308, delete "(Honshu)" from the distribution of Ischnodactylus parallelicornis Nakane, 1956
p. 309, delete "(Iriomote)" from the distribution of Platydema celatum Nakane, 1983
p. 309, add "JA" to distribution of Pentaphyllus philippinensis Kaszab, 1956
p. 318, correct spelling of aeneuum Panzer, 1793 [in synonymy with Scaphidema metallicum (Fabricius, 1792)
to aeneum
p. 320, remove Allecula simiola Lewis, 1895 from synonymy of Alleen la bilamellata Marseul, 1876, p. 319
p. 321, delete "JA" from distribution of Bolbostetha sauteri sauteri (Borchmann, 1925)
p. 323, Prionychus Solier, 1835 is masculine, change grammatical gender of the species-group names from
feminine to masculine (i.e., antracinus Mulsant, asiaticus Fairmaire, mauritanica Mulsant, and nitidula C.
G. Thomson)
p. 326, correct spelling of Isomira sanquinicollis Reitter, 1911 to sanguinicollis
p. 329, in Cistelina tokaraensis Nomura, 1961, change "(Cteniopus)" into "(Cteniopinus)"
34 ERRATA
pp. 330/331, add under subgenus Ctenioposomus Reitter, 1906 valid species:
drurei Pic, 1915j: 14 A: IQ (Mossoul)
infraniger Pic, 1915J: 14 A: TR
notatipennis Pic, 1915j: 14 A: SY(Alep)
testaceitarsis Pic, 1915j: 14 A: TR
p. 333, remove gracilior Fairmaire, 1871 from Heliotaurus; it is correctly listed in synonymy with Heliomophlm
scabriusculus Fairmaire, 1866, p. 332
p. 335, remove Cistella armillata Brullé, 1832 as the type species of Odontomophlus Seidlitz, 1896 [the type
species of Odontomophlus was not designated, and Cistella armillata is correctly listed as type species of
Megischina Reitter, 1906]
p. 335, remove entry betulae Küster, 1850 [in synonymy with Omophlus lepturoides Fabricius, 1787]; the name
is not available
p. 336, remove pilifer Seidlitz, 1896, it is correctly listed in synonymy with Omophlus falsarius Kirsch, 1869, p.
335
p. 336, remove entry lepturoides L. Redtenbacher, 1849 [in synonymy with Omophlus proteus Kirsch, 1869]; the
name is not available
p. 337, Paromophlus Znojko, 1950 was proposed without designated type species and is not available. Replace
the entry by:
subgenus Paromophlus Iablokoff-Khnzorian, 1983: 134 type species Cistelapicipes Fabricius, 1792
p. 337, correct year and page for Omophluspubescens (Linnaeus) to 1767: 303.
p. 337, remove entry picipes L. Redtenbacher, 1849 [in synonymy with Omophlus lividipes Muisant, 1856]; the
name is not available
p. 337, remove entry amerinae Kirsch, 1869 [in synonymy with Omophlus pubescens (Linnaeus, 1767)]; the
name is not available
p. 341, add "JA (Kyushu)" to distribution of Derosphaerus sinensis Hope, 1842
p. 348, add (Hemicera) to synonymy of Tetraphyllus scatebrae Lewis, 1894 [behind unicolorata Nakane, 1991:
4]
p. 350, correct spelling in distribution of Strongylium araii Mt. Chûjô, 1980 of (Sendaku Is.) to (Senkaku Is.)
p. 351, correct spelling of "watanabai" to "watanabei"\mder Strongylium marseuli
p. 394, correct year for Mylabris mannerheimii Gebier to 1845
p. 414, add to Pedilus valis species
signatipennis Fleischer, 1918: 200 A: KZ (Ketmen)
p. 414, correct spelling of Pedilus zubowskyi Semenov, 1900 to zubowskii
p. 419, add, below Salpingus:
genus Salpingus, nomen dubium
Rhinomacer striatus Geoffroy, 1785: 115 E: FR
p. 435, add to Cyclodinus valid species:
lesnei Pic, 1899c: 30 (Anthicus) A: PA (Karachi)
p. 449, add below Yunnanomonticola:
subfamily Anthicinae, species incertae sedis
Anthicus maindroni Pic, 1899c: 30 A: PA (Karachi)
p. 462, remove entry takeii M. Chûjô, 1957 from the subgenus Anaspis Geoffroy, 1762; the species is correctly
listed under the subgenus Nassipa Emery, 1876
p. 463, remove entry ganglbaueri Schilsky, 1899 from the subgenus Silaria Muisant, 1856; the species is
correctly listed under the subgenus Larisia Emery, 1876
p. 466, add as Tenebrionoidea, nomina dubia:
Cistela ornata Roemer, 1789: 3 E: "Rhaetia" [Figure in pi. 2, f. 8] [possibly an Anthicidae]
Ips atra Rossi, 1792: 23 E: IT [compared with /. crenata]
Ips dorsalis Fabricius, 1787: 45 E: "Europa australi"
Ipsfallax Rossi, 1792: 22 E: IT [compared with /. taxicornis, probably = /. ruficornis Ol.]
Ips hungarus Schrank, 1798: 513 E: HU
Ips nigra Fabricius, 1787: 47 E: SV
Ips pallida J. J. Roemer, 1789: 41 E: SZ [Figure in pi. 34, f. 11 ; apparently a Tenebrionidae]
Ips rufiis Schrank, 1798: 512 E: GE [originally stated = Rhyzophagus]
 ERRATA 35
Ips testacea Rossi, 1792: 22 E: IT [compared with /. crenata, N. ferruginea]
Rhinomacer schaefferi Harrer, 1784: 191 E: GE (Bavaria) [probably Mycterus tibialis]
p. 494, add: Bosc d'Antic L. A. G.1790: [new taxon]. In: Rossi P. ...[The original description states it has been
written by D'Antic: "... ad D. D'Antic, qui respondit hoc habitare etiam Monspeliae, & descriptionem
affabre factam concessit, quae sequens est."; the description follows]
p. 513, add: Fleischer A. 1918: Eine neue Pedilusart (Col. Pyrochroidae). Wiener Entomologische Zeitung 37:
200
p. 518, correct year for Gebier F. A. von 1844: Charakteristik ... to 1845
p. 570, correct year for Mulsant E. & Godart A. 1865: Descriptions ... to 1866 [issued on January 6, 1866]
p. 615, add: Ren G.-D. 2004: [new taxa]. In: Ren G.-D. & Yin H.: Coleoptera: Tencbrionidae. Pp. 67-72. In:
Yang X. (ed.): Insects of Great Yarlung Zangbo Canyon of Xizang, China. Beijing: China Science and
Technology Press (in Chinese with English summary)
p. 616, add: Roemer J. J. 1789: Genera insectorum Linnaei et Fabricii iconibus illustrata a Joanne Jacobo
Roemer. Vitoduri Helvetorum: apud Henrjc. Steiner et socios, viii + 86 + [3] pp., 37 pis

ERRATA for VOLUME 6

pp. 25/26, remove incorrect statement concerning Lepidopteryx Hope, 1840: Trogosita squamosa Gray, 1832
[not Hope, 1840] is not a junior synonym of Peltis squamulata Gebier, 1830 as indicated in the Errata to
volume 4 in volume 6 of the Palaearctic Catalogue (p. 26). It is a junior synonym of Trogosita varia
Fabricius, 1801 of Africa as pointed out by Léveillé (1888: cxciii) in the Annales de la Société
Entomologique de France (6e série), volume 8. This brings about that Lepidopteryx Hope, 1840 is not a
senior synonym of Leperina Erichson, 1844 as listed in the Errata (P. 25) but a junior synonym of
Gymnocheilis Dejean, 1835 and that Leperina Erichson, 1844 is the valid name for the genus including
Trogosita decorata Erichson, 1842 as listed in volume 4 of the Palaearctic Catalogue (p. 364)
p. 42, Microarthron komarowi (Dohrn, 1885): the original spelling of the species epithet is komaroffi, a
patronym in honour of General Komaroff. The spelling "komarowi" is not in prevailing usage, see e.g.,
Plavilstshikov, 1932, 1936; Kostin, 1973; Lobanov et al., 1981; Danilevsky, 1975, 1984; Svâcha, 1987, and
others who used the spelling "komarovf. Therefore, the original spelling komaroffi should be maintained
p. 44, Dorcadion sulcipenne argonauta Suvorov, 1913: the changed rank was previously published, see Lazarev
M. A. 2008: Zametki po spornym voprosam sistematiki i rasprostranenia zhukov-usachey (Coleoptera,
Cerambycidae) Rossii i sopredelnykh stran. Pp. 129-136. Aktualnye problemy prioritetnyh napravleniy
razvitiya estestvennyh nauk. Sbornik statey. Moskva: Prometey MPGU, 220 pp.
p. 44, Agapanthia subnigra Pic, 1890 and A. subchalydaea Reitter, 1898 were previously resurrected by Pesarini
& Sabbadini, 2004b
pp. 44/45, replace Dorcadion sareptanum kubanicum Plavilstshikov, 1934 by Dorcadion sareptanum eux inn m
Suvorov, 1915, and replace the discussion by: "According to Plavilstshikov (1958a: 181), the male syntypes
of Dorcadion euxinum Suvorov, 1915 (described from Novorossijsk) are D. sareptanum Kraatz, 1873, and at
least one type-female is D. cinerarium (Fabricius, 1787)". Consequently, Danilevsky et al., 2005 established
the synonymy of D. sareptanum euxinum Suvorov and D. kubanicum Plavilstshikov. In fact, the description
of D. euxinum Suvorov, 1915 was based on a single male, and that is the holotype subsequently studied by
Plavilstshikov (1958a), but recently (2009) not located. Two females (ZIN - one erroneously considered as a
male) were incorrectly designated by Suvorov as types of his D. euxinum, as they are not mentioned in his
original description. The female designated by Suvorov as a male cannot by the holotype, as it is much larger
(14 mm, while the holotype is given as 11.5 mm), and has many different characters. The name D. euxinum
Suvorov was several times published by Plavilstshikov (1921: 111; 1931: 64; 1958: 118 [in volume 6 of the
Catalogue]) as a synonym of D. cinerarium, because Plavilstshikov considered the erroneously designated
females as the types of D. euxinum Suvorov
p. 46, Dorcadion arietinum strandi Plavilstshikov, 1931: the synonymy was previously established by Breuning
(1962: 230) who placed "strandi Plav." as a morpha of D. phenax Jakovlev, 1900, and subsequently in
Danilevsky M. L. 2009: Species group taxa of longhorned beetles (Coleoptera, Cerambycidae) described by
N. N. Plavilstshikov and their types preserved in the Zoological Museum of the Moscow State University
and in the Zoological Institute of the Russian Academy of Sciences, St. Petersburg. Entomologicheskoe
Obozrenie 88: 630-663 [in Russian] and Entomological Review 89: 689-720
36 ERRATA
p. 46, Leptura apicalis Motschulsky, 1875: the synonymy was previously established by Lazarev, 2008 (referred
to above, see under Dorcadion sulcipenne argonauta Suvorov, 1913)
p. 46, Macrorhabdium Plavilstshikov, 1915 and Macrorhabdium ruficolle Plavilstshikov, 1915: the synonymy
was previously established by Danilevsky, 2009 (referred to above, see under Dorcadion arietinum strandi
Plavilstshikov, 1931
p. 46, Pterolophia multinotata Pic, 1931 : the synonymy was previously established by Lazarev, 2008 (referred to
above, see under Dorcadion sulcipenne argonauta Suvorov, 1913)
p. 47, correct spelling of Poecilium alni elbursense Holzschuh, 1977 to elburzense and treat it as valid; P. alni
latenigrum Pic, 1945 is infrasubspecific and therefore not a senior synonym of latter
p. 48, Judolia tibialis Marseul, 1876 is to be regarded as a senior synonym of Cortodera pallidipes pallidipes
Pic, 1898, and not a nomen dubium (p. 124).
p. 49/116, Stictoleptura slamai Danilevsky, 2010 is given as a replacement name for Stictoleptura martini
(Slâma), because of Leptura semirufula var. martini Pic, 1945 transferred to Stictoleptura. It was however
overlooked that Pic in his paper stated that the varieties he describes may be considered by other as
aberrations. Hence, the numerous new "varieties" are infrasubspecific names, and therefore it was
unnecessary to replace S. martini (Slâma, 1985)
p. 52, correct year for Phytoecia (Blepisanis) vittipennis leuthneri Ganglbauer to 1886
pp. 53/110, remove Sphenaria Pic, 1911 from synonymy of Pedostrangalia Sokolov, 1897. Sphenaria Pic, 1911
is a misspelling of Sphenalia and is unavailable
p. 56, Phytoecia subannulipes by Pic, 1910; Pic, 1910h stated "Cette espèce décrite de Syrie...", and gave a
définition; yet the name is a misspelling of P. subannularis Pic, 1901, repeated once more in Pic, 191 la: 9.
Later, Pic (1915f: 11) stated mat P. subannulipes is a Rumanian variety of P. subannularis. Thus, the name
became available in 1915 and is a synonym of P. icterica (Schaller, 1783)
p. 60, correct spelling of Dorcadion erytropteron Fischer von Waldheim, 1823 to D. erytropterum
p. 77, remove Coptocephala scopolina f. bipunctata Dépoli, 1926, f. destituta Dépoli, 1926, f. lateremaculata
Dépoli, 1926, f. scutellaris Dépoli, 1926 and f. separata Dépoli, 1926; these names are considered
infrasubspecific
p. 79, correct author of Smaragdina to Chevrolat, 1836
p. 83, Thysbe J. Thomson is currently in synonymy witii Colasposoma Laporte, 1833, not Platycorynus
Chevrolat, 1836
p. 83, correct spelling of Verna & al., 2005 to Verma & al., 2005
p. 85, remove entry Antinoe J. Thomson, 1864 from synonymy of Distenia [it is a junior homonym replaced by
Novantinoe Santos-Silva & Hovore 2007, and resurrected as valid genus]
p. 85, add [HN] to entry Thelxiope J. Thomson, 1864: 226 and add its replacement name in synonymy witii
Distenia:
Thomsonistenia Santos-Silva & Hovore, 2007: 3 [RN] type species: Thelxiope viridicyanea J. Thomson, 1864
p. 85, correct year of Distenia mellina Holzschuh,perforons Holzschuh, and rufobrunnea Holzschuh to 1995
p. 86, remove entry: caspica Faldermann, 1835 from synonymy of Parandra caspia Ménétriés, 1832; it is a
misspelling
p. 87, Aegosoma Audinet-Serville, 1832 is neuter, change grammatical gender of the species-group names from
feminine or masculine to neuter
p. 87, correct reference for Aegosoma sinicum validicorne to Gressitt, 1951b: 205
p. 90, add germari Mulsant, 1846: 291 in synonymy witii Macrotoma myardi myardi Mulsant, 1842
p. 90, remove entry: luzonum Pascoe, 1869: 666 [HN] {Macrotoma) [in synonymy with Bandar pascoei
(Lansberge, 1884)]; it is a misidentification
p. 92, add zarudnyi Plavilstshikov, 1936: 80 (Prionus) [unjustified emendation] in synonymy with Mesoprionus
zarudnii (Semenov, 1933)
p. 92, correct spelling of Microarthron komarowi to komaroffl (Dohrn, 1885) and add following unjustified
emendation in its synonymy: komarovi Semenov, 1935b: 241, 246 (Prionus) and komarowi Pic, 1898e: 33,
35 (Prionus)
p. 94, add pluschtschewskii Semenov, 1900a: 252 (Polyarthron) [unjustified emendation] in synonymy with
Pseudoprionus bienerti Heyden, 1885
p. 96, remove entry: pallescens Holzschuh, 1993a: 20 A: GAN SHA from Acanthoptera and place it in
Parastrangalis, p. 109
 ERRATA 37
p. 96, add NC NE SC to distribution of Alosterna tabacicolor erythropus Gebier, 1841
p. 96, remove fusca Matsushita, 1930 from synonymy of Alosterna tabacicolor erythropus Gebier, 1841; it is
infrasubspecific
p. 96, remove JA NE SC from distribution of Alosterna tabacicolor tabacicolor (DeGeer, 1775)
p. 96, remove entry pallescens Holzschuh, 1993a: 20 and place it under Parastrangalis Ganglbauer, p. 109
p. 97, remove planeti Pic, 1945, curierensis Pic, 1945, and circascutellaris Pic, 1945 from synonymy of
Anastrangalia dubia dubia (Scopoli, 1763); these names are infrasubspecific
p. 98, remove JA from distribution of Anoplodera rufihumerale (Tamanuki, 1938)
p. 98, add [HN] to entry Anoplodera rufipes rufipes (Schaller, 1783), and add TR to its Asian distribution; the
name is preocupied by Leptura rufipes Goeze, 1777, and both were considered congeneric after 1899
p. 98, add [HN] to entries: atra Paykull, 1800, rufiventris Tournier, 1872, and villosa Schoenherr, 1817 [in
synonymy with Anoplodera rufipes rufipes (Schaller, 1783)]
p. 98, correct year of Anoplodera tenebraria Holzschuh to 1995
p. 99, under Corennys, remove entry brevipennis prescutellaris Pic, 1947
p. 100, remove entry auriflua L. Redtenbacher, 1858fromsynonymy of Etorufuspubescens (Fabricius, 1787); it
is a numen nudum
p. 100, correct reference for Gnathostrangalia to Hayashi & Villiers, 1985a: 13
p. 101, remove femora ta Mulsant, 1863: 580fromsynonymy ofGrammoptera abdominalis Stephens, 1831 [it is
a misidentification]
p. 101, correct distribution of Grammoptera ruficornis obscuricornis Kraatz, 1886 to E: AB (Talysh) A: IN
p. 101, add CT and TR to European distribution of Grammoptera ruficornis ruficornis (Fabricius, 1781 )
p. 101, add holomelina Donisthorpe, 1905: 182 to synonymy of Grammoptera ruficornis ruficornis (Fabricius,
1781) [note: the name Grammoptera ruficornis ab. holomelina Pool, 1905fromGreat Britain is unavailable,
though is was often used as valid. It was made available by H. Donisthorpe (1905), according to ICZN Art.
12.2. Totally black forms of G. ruficornis (with all legs also black) are unknown from any other parts of the
species range (and neither in G. ruficornis obscuricornis Kraatz, 1886 from Talysh and Iran). Thus, the
eventual validity of holomelina Donisthorpe, 1905 remains to be cleared]
p. 101, add NC and NE to distribution of Grammoptera (Neoencyclops) cyanea Tamanuki, 1933
p. 102, add mediolineata Pic, 1954a: 13 to synonymy of Idiostrangalia contracta (Bates, 1884), and remove it
from Strangalia Audinet-Serville, 1835, p. 116 [note: according to N. Ohbayashi]
p. 102, remove milliati Pic, 1945 from synonymy of Judolia sexmaculata (Linnaeus, 1758); the name is
infrasubspecific
p. 102/103, transfer rufimembris Pic, 1917 listed as nomen dubium in synonymy with Judolia parallelopipeda
(Motschulsky, 1860)
p. 103, remove entry Strangaliella Hayashi, 1976: 3 type species Strangalia shikokensis Matsushita, 1935 (=
Strangalia tenuicornis Motschulsky, 1862) from synonymy of Leptura Linnaeus, 1758 and place it in
synonymy -with Parastrangalis Ganglbauer, 1889, p. 109
p. 104, correct year for unicolor Olivier [in synonymy with Leptura aethiops Poda von Neuhaus, 1761] to 1797
p. 104, add GAN SCH to distribution of Leptura gradatula Holzschuh, 2006
p. 105/110, remove entry tenuicornis Motschulsky, 1862: 20 (Strangalia) A: JA (with its three synonyms) from
Leptura Linnaeus, 1758, and place it in Parastrangalis Ganglbauer, 1889, p. 110
p. 106, remove albicornis Tamanuki, 1942 from synonymy of Metastrangalis ochraceoventra Gressitt, 1935 and
place it as valid species in Metastrangalis, occurring in TAI
p. 106, remove fukiensis Tippmann, 1955 from synonymy of Mimostrangalia lateripicta Fairmaire, 1895 and
place it as valid species in Jschnostrangalis, occuring in FUJ, p. 102
p. 107, correct entry subgenus Nivelliamorpha Boppe, 1921 to genus Nivelliamorpha
p. 107, remove medioconexa Pic, 1945 from synonymy of Nustera distigma (Charpentier, 1825); the name is not
available
p. 108, add KZ to Asian distribution of Pachytodes erraticus erraticus (Dalman, 1817)
p. 109, correct year ofParastrangalis conges ta Holzschuh and P. eucera Holzschuh to 1995
p. 110, add [HN] to entry jaegeri Fairmaire, 1866b: 279 {Leptura) [in synonymy with Pedostrangalia emmipoda
(Mulsant, 1863)]
p. Ill, remove entry brevioripennis Pic, 1955 from synonymy with Pygostrangalia vittaticollis (Pic, 1926) and
place it as a valid species, with unknown distribution
38 ERRATA
p. 113, remove entry latenigra Pic, 1915 from synonymy of Stenurella septempunctata septempunctata
(Fabricius, 1792); it is correctly listed under S. septempunctata suturata (Reiche & Saulcy, 1858), p. 114
p. 115, remove hardenbergi Bodemeyer, 1927 from synonymy of Stictoleptura fontenayi (Mulsant, 1839); the
name was introduces as an aberration and is not available
p. 115, add fulvoapicalis Plavilstshikov, 1932f: 174 (Leptura) in synonymy with Stictoleptura fulva (DeGeer,
1775)
p. 115, replace maculicornis maculicomis DeGeer, 1775 by maculicornis DeGeer, 1775
p. 115, add rubromarginata Plavilstshikov, 1932f: 174 {Leptura) in synonymy with Stictoleptura rufa dimidiata
(K. Daniel & J. Daniel, 1891)
p. 116, correct spelling oîsemirufa (Kraatz, 1880) [in synonymy with Stictoleptura trisignata (Fairmaire, 1852)]
to semirufula
p. 116, remove martini Pic, 1945 from synonymy of Stictoleptura trisignata (Fairmaire, 1852); the name is
infrasubspecific
p. 116, add BY EN LA LT JA NE NO to distribution of Stictoleptura variicornis (Dalman, 1817)
p. 117, correct year for Thrangalia Holzschuh and T. diaboliella Holzschuh to 1995
p. 117, add AB to distribution of Vadonia bitlisiensis Chevrolat, 1882
p. 117, correct page of distription of flscheri Zubkov, 1829 to 168 [in synonymy with Vadonia bipunctata
bipunctata (Fabricius, 1781)]
p. 117, add (Leptura) to entry: cribricollis Pic, 1889b: 20 [mispaginated: 5] [in synonymy with Vadonia
bitlisiensis Chevrolat, 1882]
p. 118, Vadonia unipunctata unipunctata (Fabricius, 1787) is declared nomen protectum, but this was not
supported according to ICZN Art. 23.9.1.2. The supporting references are given below by M. L. Danilevsky,
in Appendix 1
p. 118, add uninstigmata Pic, 1891b: 9 {Leptura) to synonymy of Vadonia unipunctata unipunctata (Fabricius,
1787)
p. 119, add BY and JA to distribution of Acmaeops angusticollis (Gebier, 1833)
p. 119, remove JA from distribution of Acmaeops marginatus (Fabricius, 1781)
p. 119, Acmaeops sachalinensis Tsherepanov, 1978 is listed as junior synonym of A. angusticollis (Gebier,
1833); this synonymy is to be credited to M. L. Danilevsky,. it was based on study of the type material and
was published within the Catalogue for the first time
p. 119, remove entries fulvipennis Mannerheim, 1853: 251 (Pachyta), longiceps Kirby, 1837: 187 (Leptura) and
semimarginatus Randall, 1838: 30 (Leptura) from synonymy, and add ustulatus Motschulsky, 1860: 148
(Pachyta) to synonymy of Acmaeops pratensis (Laicharting, 1784)
p. 119, add (Toxotus) to entry: dentipes Mulsant, 1842a: 209 [in synonymy with Akimerus schaefferi schaefferi
(Laicharting, 1784)]
p. 120, correct year for Brachyta bifasciata bifasciata (Olivier) t© 1797, and add MG to its distribution
p. 120, remove NMO from distribution of Brachyta bifasciata japonica and correct reference to: Matsushita,
1933b: 178
p. 120, add [= 1892b: 81] and the synonym conjuncta Rost, 1893: 344 to Brachyta caucasica caucasica Rost,
1892a
p. 120, remove [NO] from entry duodecimmaculata Fabricius, 1781: 248 (Leptura) [in synonymy with Brachyta
interrogationis (Linnaeus, 1758)]
p. 120, remove [RN] from kraatzi Ganglbauer, 1889c: 468 [in synonymy with Brachyta interrogationis
(Linnaeus, 1758)]
pp. 120/121, remove bernardinus Pic, 1915a: 41 (Evodinus) and theresae Pic, 1915a: 41 (Evodinus) from
synonymy of Brachyta interrogationis (Linnaeus, 1758); the names are infrasubspecific
pp. 120/121, correct data for immaculatus, mulsanti, multiguttatus, plavilstshikovi, and prescutellaris to
immaculatus Pic, 1934f: 28 (Evodinus), mulsanti Pic, 1934f: 31 (Evodinus), multiguttatus Pic, 1934f: 31
(Evodinus), plavilstshikovi Pic, 1934f: 31 (Evodinus), and prescutellaris Pic, 1934f: 31 (Evodinus) [all in
synonymy with Brachyta interrogationis (Linnaeus, 1758)]
p. 121, remove entry mutabilis Motschulsky, 1859a: 233 (Evodinus) from synonymy of Brachyta striolata
(Gebier, 1817), it is correctly listed in synonymy with Brachyta variabilis scapularis (Mannerheim, 1849)
p. 121, add AB to distribution of Cortodera alpina alpina (Ménétriés, 1832)
 ERRATA 39
p. 122, replace (Cartodera) by (Pachyta) for confusa Reitter, 1891b: 34 [in synonymy with Cortodera alpina
alpina (Ménétriés, 1832)]; "Cartodera" is a misspelling
p. Ï22,removeparallela Pic, 1898k: 111 from synonymy of Cortodera alpina starcki Reitter, 1888 and place it
in synonymy with C. alpina rosti Pic, 1892
p. 122,replaceentries for Cortodera alpina umbripennis Reitter by:
alpina umbripennis Reitter, 1890e: 245 E: AB AR GG ST A: IN TR
armeniaca Pic, 1898k: 114
alpina xanthoptera Pic, 1898k: 114[RN] A: TR (Angora)
flavipennis Ganglbauer, 1897a: 53 [HN]
rosinae Pic, 1902c: 8
p. 122,removeentryflavipennisGanglbauer, 1897a: 53fromsynonymy of Cortoderaflavimana(Wahl, 1838)
p. 122, add [= 1987: 615] to entry Cortodera colchica danczenkoi Danilevsky, 1985: 139
p. 122, correctreferencefor Cortodera griseipes to Pic, 1889b: 55
p. 122, correct entries for Cortodera holosericea (Fabricius, 1801 ) to:
holosericea holosericea Fabricius, 1801b: 366 (Leptura) E: AU BU HU RO SK ST UK
pilosa Pic, 1989g:50
rubripes Pic, 1898k: 114
semitestacea Pic, 1989g: 50
holosericea velutina Heyden, 1876a: 318 E: AU BH CR GR IT SL YU
birnbacheri Pic, 1898k: 114
p. 122,removethe unavailable name holosericea Gebier, 1848 from synonymy and replace data for Cortodera
analis (Gebier, 1830) by:
analis Gebier, 1830: 189 (Pachyta) A: KZ WS XIN
haemorhoidalis Pic, 1898k: 77
haemorrhoidalis Aurivillius, 1912: 197 [unjustified emendation]
hirta Gebier, 1830: \9Q (Pachyta)
ruficornis Pic, 1926d: 6
p. 122, add atropyga Pic, 1929h: 119 [DA] in synonymy with Cortodera colchica colchica Reitter, 1890
p. 122, add BU to distribution of Cortodera discolor Fairmaire, 1866
p. 123, correct entry Cortodera humeralis orientalis Adlbauer, 1988 to orientalis Adlbauer, 1988
p. 123/124, remove nomen dubium and replace entries for Cortoderapallidipes Pic by:
tibialis komarovi Danilevsky, 1996c: 63 A: KZ
tibialis rossica Danilevsky, 2001b: 7 E: UK ST
tibialis ruthena Plavilstshikov, 1936: 286 E: ST A: KZ
tibialis tibialis Marseul, 1876: cii (Judolia) E: ST
pallidipes Pic, 1898g:49
tibialis turgaica Danilevsky, 2001b: 9 E: CT A: KZ
p. 123, add TR to Asian distribution of Cortoderapseudomophlus Reitter, 1889
p. 123, correct page of description for Cortodera parfentjevi Miroshnikov, 2007 to 215
p. 123,reversevalidity for entry:
schurmanni Sama, 1997b: 107 E: GR
diversipes Pic, 1898k: 79
p. 124, correct page of description of persica Plavilstshikov, 1936 [in synonymy with Cortodera transcaspica
Plavilstshikov, 1936] to 291
p. 124, correct year for Cortodera wewalkai Holzschuh and C. wittmeri Holzschuh to 1995
p. 124, correct original combination of sylvestris Geoffroy, 1785 [in synonymy with Dinoptera collaris
(Linnaeus, 1758)] to (Stenocorus)
p. 124, correct spelling and data forfrivaldskyiKraatz, 1876b: 344 (Grammoptera) [in synonymy with C. villosa
villosa Heyden, 1876] to:frivaldszkyiKraatz, 1876b: 318 [= 1876: 344] (Grammoptera)
p. 125, correct page of description of interruptelunata G. Schmidt, 1951 [in synonymy with Evodinus borealis
(Gyllenhal, 1827)] to 11
p. 125, add flecki G. Schmidt, 1958: 77 in synonymy with Evodinus borealis (Gyllenhal, 1827)
p. 125, remove atromultiplicatus Pic, 1945, diversesignatus Pic, 1945, leprieuri Pic, 1945, morginsius Pic, 1945,
and vesubiensis Pic, 1945 from synonymy of Evodinus clathratus (Fabricius, 1792); the names are
infrasubspecific
40 ERRATA
p. 125, add (Brachyta) to entry nigritus Pic, 1891b: 6 [in synonymy with Evodinus clathratus (Fabricius, 1792)]
p. 125, remove CT from distribution of Fallacia elegans (Faldermann, 1837)
p. 125, Gaurotes glabratula Holzschuh, 1998 is valid, and G. oligothrix Chiang, 2001: 77 is an invalid RN,
correct year for the latter from 1996 to 2001
p. 126, replace entries for Gaurotes (Carilia) virginea (Linnaeus, 1758) by:
virginea aemula Mannerheim, 1852b: 306 {Pachyta) E: CT ST A: ES FE HEI HUB JIL KZ MG NMO SHX
WS
ruficollis Solsky, 1871: 403 (Pachytä)
virginea kozhevnikovi Plavilstshikov, 1915c: 105 A: FE HEI JIL NC SC
komensis Tamanuki, 1938b: 167
nigriventris Jurecek, 1921: 25
nigriventris Tamanuki, 1938b: 167 [HN]
sibirica Podany, 1962: 236
virginea thalassina Schrank, 1781a: 161 (Leptura) E: AU FR IT SL
virginea virginea Linnaeus, 1758: 398 (Leptura) E: AL AU BE BH BU BY CR CT CZ EN FI FR GE GR HU
IT LA LS LT MD NR NT PL RO SK SL SV SZ ST UK YU
notaticollis Pic, 1916a: 10
sanguinaria Pic, 1917g: 4
violacea DeGeer, 1775: 144 (Lepturä)
p. 126, original spelling of Gaurotina nigroantennata L. Chen & Chiang, 2000 is "nigroantenata"
p. 127, correct spelling of incolumnis Heyden, 1886 to incolumis [in synonymy with Pachyta bicuneata
Motschulsky, 1860]
p. 127, remove mulsanti Pic, 1945 from synonymy of Pachyta quadrimaculata (Linnaeus, 1758); the name is
infrasubspecific
p. 128, remove JA, TAI and ZHE from distribution of Pidonia debilis [Kraatz, 1879)
p. 129, add NC and SC to the distribution of Pidonia alticollis (Kraatz, 1879)
p. 129, remove JA from the distribution of Pidonia amurensis (Pic, 1900)
p. 129, correct reference for Pidonia chinensis to Hayashi & Villiers, 1985a: 17
p. 129, correct reference for rufiventris Plavilstshikov [in synonymy with Pidonia alticollis (Kraatz, 1879)] to
Plavilstshikov, 1932a: 87 [the reference is added below]
p. 130, remove suturalis Olivier from synonymy of Pidonia lurida (Fabricus, 1792); it is a misidentification of
Leptura suturalis Fabricius, 1787 and unavailable
p. 131, correct year for Pidonia palligera Holzschuh to 1995
p. 131, correct original combination and distribution of Pseudogaurotina excellens Brancsik, 1874 to (Pachyta)
and E: PL RO SK UK
p. 133, correct page of description of nubecula Bergsträsser, I778 to 26 [in synonymy with Rhagium inquisitor
inquisitor (Linnaeus, 1858)]
p. 133, under Rhagium correct spelling of qinhaiene to qinhaiense L. Chen & Chiang, 2000
p. 134, remove brevevittatus Pic, 1945 and lividipennis Pic, 1945 from synonymy of Stenocorus insitivus
insitivus Germar, 1824; the names are infrasubspecific
p. 134, remove NC and SC from distribution of Stenocorus meridianus (Linnaeus, 1758)
p. 134, correct entries for Stenocorus vittatus to:
vittatus Fischer von Waldheim, 1842: 19 (Toxotus) [in prevailing usage] A: KZ XIN
obliquus Motschulsky, 1845a: 86 (Toxotus)
suvorovi Plavilstshikov. 1936: 173 [unjustified emendation]
suworowi Reitter, 1907a: 208 (Toxotus)
turkestanicus Ganglbauer, 1889b: 280 (Toxotus)
vittattus Fischer von Waldheim, 1842: 19 (Toxotus) [original spelling]
p. 135, remove entry ambustum Heyden, 1877 from synonymy of Rhamnusium bicolor (Schrank, 1781); the
name is infrasubspecific
p. 138, remove theresae Pic, 1945 from synonymy of Asemus striatum (Linnaeus, 1758); the name is
infrasubspecific
p. 138, remove sharpi Reitter, 1913 from synonymy of Megasemus quadricostulatum Kraatz, 1879; it is
correctly listed in synonymy with Cephalallus unicolor Gahan, 1906
p. 141, correct page of description of Necydalisca Plavilstshikov, 1936 to 462
 ERRATA 41
p. 143, correct year and type species for Pufiijia Holzschuh to 1995 and to Nortia luteosignata Pu, 1991,
respectively
p. 144, remove rufotinctus Pic, 1945 from synonymy of Anaglyptus gibbosus (Fabricius, 1787); the name is
infrasubspecific
p. 147, correct reference for Aromia orientalis to Plavilstshikov, 1932e: 114
p. 151, remove semitestaceum Pic, 1945 from synonymy of Callidium aeneum aeneum (DeGeer, 1775); the
name is infrasubspecific
p. 151, correct year for Leioderes L. Redtenbacher and L. kollari L. Redtenbacher to 1848
p. 151, correct year for analis L. Redtenbacher [in synonymy with Phymatodes testaceus (Linnaeus, 1758)] to
1848
p. 152, add original combination {Poecilium) to Phymatodes magnanii Sama & Rapuzzi, 1999
p. 153, remove [HN]fromentry: Phymatoderus Reitter, 1913 [in synonymy with Poecilium Fairmaire, 1864]
p. 153, correct spelling of elbursense Holzschuh, 1977 to elburzense and use it as valid; it is not a synonym of
Poecilium alni latenigrum Pic, 1945 which is infrasubspecific
p. 153, correct year for castaneum L. Redtenbacher [in synonymy with Poecilium glabratum (Charpentier, 1825)]
to 1848
p. 153, correct year for Poecilium eximium Holzschuh to 1995
p. 155, add LT to distribution of Ropalopus macropus (Germar, 1824)
p. 156, remove XIN from distribution of Turanium johannis johannis Baeckmann, 1922
p. 156, remove Pneumida J. Thomson, 1864 from synonymy of Ceresium Newman, 1842; it was resurrected by
Holzschuh, 1995
p. 156, correct year for Ceresium fallaciosum Holzschuh to 1995
p. 157, remove entry Salpinia laosensis Gressitt & Rondon, 1970; the species is correctly listed under
Parasalpinia Hayashi, 1962
p. 158, place tripunctata Gressittt & Gordon, 1970 in synonymy with Stenodryas nigromaculata Gardner, 1942
p. 159, add SV to distribution of Cerambyx cerdo cerdo Linnaeus, 1758
p. 159, correct spelling of manderstjaernae Mulsant & Godart, 1855 to manderstjernae [in synonymy with
Cerambyx cerdo cerdo Linnaeus, 1758]
p. 159, correct distribution of Cerambyx heinzianus Demelt, 1976 to A: TR
p. 160, correct spelling of Hemadius oenochroa Fairmaire, 1889 to oenochrous, and remove "(Hemadius)" from
the entry
p. 162, correct year for Rhytidodera consona Holzschuh to 1995
p. 163, correct year for Artimpaza sausai Holzschuh, Cleomenes atricornis Holzschuh, C giganteus Holzschuh
and C. modicatus Holzschuh to 1995
p. 164, correct year for Diplothorax lucens Holzschuh to 1995
p. 164, add to Kurarua Gressitt, 1936 valis specie:
brevipes Holzschuh, 1984: 152 A: NP SD
p. 165, remove socius Gahan from synonymy of Chloropterus drciferus Chevrolat, 1863, and add to
Chloropterus, p. 166, entry:
socius Gahan, 1906a: 264 (Caloclytus) A: SD
p. 166, remove uniformis Pic, 1945 from synonymy of Chlorophorus glabromaculatus (Goeze, 1777); the name
is infrasubspecific
p. 167, use as valid name Chloropterus motschulskyi (Ganglbauer, 1887), a RN for C. latofasciatus
(Motschulsky, 1861) [the latter is a junior homonym of latifasciatus (Fischer von Waldhein, 1832), see
ICZN Art. 58.12], and correct the original combinattion to Clytus
p. 169, remove JIA from distribution of Chlorophorus varius varius (O. F. Müller, 1766)
p. 170, remove delagrangei Pic, 1945 from synonymy of Clytus ciliciensis (Chevrolat, 1863) and maritimus Pic,
1945fromsynonymy of Clytus lama Mulsant, 1847; these names are infrasubspecific
p. 172/173, correct year for Demonax jendeki Holzschuh, D. parilis Holzschuh and D. sausai Holzschuh to 1995
p. 173, add [RN] to hainanensis Gressitt & Rondon, 1970 [in synonymy with Demonax reductispinosus Gressitt,
1942]
p. 173, correct spelling of albofasciatus Pic, 1920 to albobifasciatus [in synonymy with Demonax semiluctuosus
A.White, 1855]
42 ERRATA
p. 174, remove entry subobliteratus Pic, 1918 [in synonymy with Grammographus notabilis cuneatus
(Fairmaire, 1888)]; it was proposed as a replacement name for "Chbrophorus obliterates Pic, 1902"
p. 174, correct year for Ischnodora decolorata Holzschuh to 1995
p. 177, add to Plagionotus valid subspecies:
detritus caucasicola Plavilstshikov, 1936: 435 E: AB AR GG ST A: SY TR
p. 177, remove AB, AR, GG, and ST from distribution of Plagionotus detritus s. str.
p. 177, correct reference for obscurebasalis Pic [in synonymy with Plagionotus detritus (Linnaeus, 1758)] to Pic,
1942c: 2

p. 178, replace entry for Rhaphuma Pascoe by:

genus Rhaphuma Pascoe, 1858: 240 [RN] type species Clytus quadricolor Laporte & Gory, 1836
Arcyophorus Gemminger, 1872: 2938 [unjustified emendation]
Arcyphorus Chevrolat, 1863b: 287 type species Arcyphorus histrio Chevrolat, 1863
Rhaphium A. White, 1855: 289 [HN] type species Clytus quadricolor Laporte & Gory, 1836
Raphuma J. Thomson, 1861: 221 [RN]
p. 178, correct year for Rhaphuma decora Holzschuh to 1995
p. 178, place kantiae Holzschuh, 1989 as subspecies of Rhaphuma manipurensis
p. 179, transfer Rhaphuma savioi Pic, 1924 to Chbrophorus, p. 168, and correct its distribution to: A: GUI HEB
SHA SHG SHX
p. 179, transfer curvofasciata Gressitt, 1939 [in synonymy with Rhaphuma savioi Pic, 1924] as valid species to
Demonax J. Thomson, 1861, p. 172, with distribution: A: FUJ GUA GUI HUN SCH SHA SHX ZHE
p. 180, correct type species of Sclethrus Newman, 1842 to Sclethrus amoenus Newman, 1842
p. 180, add entry: Neocollyroides Schultze, 1920: 196 type species Neocollyroides macregory Schultze, 1920 in
synonymy with Sclethrus Newman, 1842
p. 180, add original combination (Xylotrechus) to Turanoclytus ilamensis campadellii Sama & Rapuzzi, 2003
and T. raghidae Sama & Rapuzzi, 2000
p. 181/183, remove villioni Villard, 1892 with its synonym nipponicus Seki, 1935 from the subgenera Xyloclytus
Reitter and Xylotrechus Chevrolat, they are correctly listed under Ootora Niisato & Wakejima, 2008
p. 183, remove entry Xylotrechus zaisanicus Plavilstshikov, 1940 with its synonym arnoldii Kostin, 1974; they
are correctly listed under subgenus Kostiniclytus, p. 180
p. 185, remove fagniezi Pic, 1945 [original spelling: fagnezi] from synonymy of Penichroa fasciata (Stephens,
1831); the name is infrasubspecific
p. 186, correct year for Trichoferus arenbergeri Holzschuh to 1995
p. 186, add AB AR to distribution of Trichoferus campestris (Faldermann, 1835)
p. 188, remove subgenus Pakmolorchus Holzschuh, 1989 from Epania Pascoe, 1858, and place it as subgenus
under Molorchus Fabricius, 1792, p. 191
p. 190, remove sterbai Adlbauer from synonymy of Glaphyra tenuitarsis (Holzschuh, 1981) and add entry:
sterbai Adlbauer, 1988: 277 (Molorchus) A: LE
p. 190, correct year for Glaphyra unaminis Holzschuh to 1995
p. 190, add to Molorchus valid species:
abieticola Holzschuh, 2007: 218 E: TR
p. 192, remove entry aegrotum Holzschuh, 1982 from Chinobrium Gressitt, 1937 and place it under Obrium
Dejean, 1821
p. 192, add to Comusia valid species:
bengalensis Fisher, 1940: 198 (Oemospiloides) A: SD
p. 192, correct spelling of Iranobrium brancucci Holzschuh, 1993 to brancuccii
p. 193, correct year for Obrium fumigatum Holzschuh to 1995
p. 194, correct type species of Afroeme Adlbauer, 2004 to Afroeme kenyensis Adlbauer, 2004
p. 194, remove entries subgenus Hypoeschrus and indicus Gahan, 1906 Hypoeschrus J. Thomson, 1864 from
genus Hypoeschrus J. Thomson, 1864; the species is correctly listed under Noserius Pascoe, 1857
p. 194, add entry:
genus Entetraommatus Fischer, 1940: 199 type species Entetraommatus quercicola Fischer, 1940
quercicola Fischer, 1940: 199 A: UP
 ERRATA 43
p. 195, remove entries Pseiidallotraeus Pic, 1923, orientalis A. White, 1853 and rufescens Pic, 1923 from
Nysina Gahan, 1906. Pseiidallotraeus is correctly listed as subgenus of Allotraeus Bates, 1877, and the
species orientalis A. White is correctly listed in Allotraeus s. str.
p. 195, correct spelling of Prothema auratum interruptum Pic, 1927 to auratum ininterruptum
p. 196, remove simile Gressitt & Rondon, 1970fromsynonymy of Prothema cakli Heyrovsky, 1967; it is a valid,
extralimital species
p. 196, under Afghanicerus, correct spelling of dressi Tippmann, 1958 to dreesi
p. 197, remove subjuncta Pic, 1945 from synonymy of Calchaenesthes oblongomaculata (Guérin-Méneville,
1844); the name is infrasubspecific
p. 197, add Acanthoptera Latreille, 1829: 114 in synonymy with Purpuricenus Dejean, 1821, as an unnecessary
substitute name
p. 198, remove UK from distribution of Purpuricenus dalmatinus Sturm, 1843, and remove bonvouloiri Pic,
1945fromits synonymy; the latter name is infrasubspecific
p. 198, remove tournieri Pic, 1945 from synonymy of Purpuricenus kaehleri kaehleri (Linnaeus, 1758); the
name is infrasubspecific
p. 200, remove "(Pyrocalymma)"fromentry Pyrestes nigricollis Pic, 1947
p. 201, remove entry rufipes rufipes Pic, 1923fromPyrestes Pascoe; this nominal subspecies was described from
Vietnam
p. 202, add IT to distribution of Rosalia alpina alpina (Linnaeus, 1758)
p. 202, correct year for Stenhomalus clarinus Holzschuh to 1995
p. 203, remove angustissimus (Pic, 1903) from Kunbir Lameere, 1890, and place the entry under Kurarua
Gressitt, 1936, p. 164
p. 203, add Kurseongia Pic, 1930b: 15 type species Debilissa laboissierei Pic, 1930 in synonymy with Kunbir
Lameere, 1890 and add valid species:
laboissierrei Pic, 1930b: 16 (Debilissà) A: SD
p. 204, correct year for microdebilissa aethiops Holzschuh to 1995
p. 205, remove AB, AR, GG and TMfromdistribution of Stenopterus rufus rufus (Linnaeus, 1767), add AB, AR
and GG to distribution of Stenopterus rufus geniculate Kraatz, 1863, and remove INfromthe distribution of
the latter subspecies
p. 205, add to Microdebilissa valid species:
diversipes Pic, 1930b: 16 A: SD
p. 205, add to Stenopterus valid subspecies:
rufus transcaspicus Lazarev, 2008: 132 A: IN TM
p. 207, correct spelling of Neoacanthocinus [in synonymy with Acanthocinus Dejean, 1821] to Neacanthocinus
p. 207, remove European distribution of Acanthocinus carinulatus Gebier, 1833 [based on misidentified A.
griseus (Fabricius, 1792)]
p. 202, for Sybrinus sokotrensis, correct spelling of (Arasbosybrinus) to (Arabosybrinus)
p. 209, add (Liopus) to entry: japonicus Pic, 1901 v: 342 [in synonymy with Leiopus stillatus (Bates, 1884)]
p. 215, add CT to distribution of Agapanthia kirbyi (Gyllenhal, 1817)
p. 217, remove decoloripes Pic, 1945 from synonymy with Calamobius filum (Rossi, 1790); the name is
infrasubspecific
p. 221, correct year and original combination for Palimna annulata (Olivier) to 1797 and Lamia, respectively
p. 227, correct year for Sybrinus sokotrensis Téocchi et al. to 2007
p. 229, correct Asaperda agapanthina agapanthina Bates, 1873 and A. agapanthina kani Hayashi, 1976 to
Asaperda agapanthina Bates, 1873 Asaperda kani Hayashi, 1976, respectively
p. 230, under Coomanum Pic, 1927, add as synonym:
Niphosaperda Breuning, 1962d: 49 type species Niphosaperda rondoni Breuning, 1962
p. 239, place entry Diastocera J. Thomson, 1857 in synonymy with Thysia J. Tomson, 1860 and use the latter as
the valid genus-group name. Diastocera J. Thomson is a junior homonym of Diastocera Dejean, 1835
p. 239, for Thysiotes J. Thomson, 1868 add "unnecessary" to [RN]
p. 240, correct spelling of Tylophorus Blessig, 1873 and of T. wulffusi Blessig, 1873 to Thylophorus and T.
wulffiusi, respectively [in synonymy with Moechotypa J. Thomson, 1864 and M. diphysis (Pascoe, 1871 )]
p. 242, add original genus combination (Compsodorcadion) to Dorcadion tibiale Jakovlev, 1889
p. 242, add rugipenne Tippmann, 1958b: 155 in synonymy with Dorcadion fulvum fulvum (Scopoli, 1763)
44 ERRATA
p. 243, add MC to distribution oïDorcadion albosuturale Breuning, 1946
p. 243, place entry majoripenne Pic, 1926 as subspecies oïDorcadion aethiops (Scopoh, 1763) andpropinquum
Breuning, 1962 in synonymy with the latter, p. 242
pp. 243/246, remove fiorii Breuning, 1942b: 126 from synonymies of Dorcadion arenarium marsicanum
Fracassi, 1905 and D. etruscum etruscum (Rossi, 1790); it is correctly listed as etruscum fiorii Breuning,
1942 on p. 247.
p. 243, correct spelling oïDorcadion arenaroides Rabaron, 1979 to arenarioides
p. 243, place amphissae Breuning, 1974 and rugicolle Breuning, 1962 in synonymy with Dorcadion minutum
minutum Kraatz, 1873
p. 243, place atticum Kraatz, 1873 as subspecies oïDorcadion minutum Kraatz, 1873
p. 244, remove sabandum Pic, 1945 from synonymy oïDorcadion arenarium arenarium (Scopoli, 1763); the
name is infrasubspecific
pp. 244/250, place Dorcadion bravardi Pic, 1916 as subspecies of D. etruscum (Rossi, 1790)
p. 245, remove euxinum Suvorov, 1915: 119, and correct reference for perroudi to Pic, 1942b: 2 [both in
synonymy with Dorcadion cinerarium cinerarium (Fabricius, 1787)]
p. 246, 251, 254, place epirense Breuning, 1942, pindicum Breuning, 1966, and tassii Breuning, 1964 in
synonymy with Dorcadion etruscum etruscum (Rossi, 1790)
pp. 248/249 and 254, place joanninae Breuning, 1967, lianokladii Breuning, 1962, margheritae Breuning, 1964,
and wewalkai Breuning, 1974 in synonymy with Dorcadion taborskyi Heyrovsky, 1941
p. 249, correct year for Dorcadion maceki Holzschuh to 1995
p. 250, place mimarenarium Breuning, 1974 as subspecies oïDorcadion minutum Kraatz, 1873
p. 251, remove entry molitor L. Redtenbacher from synonymy of Dorcadion pédestre pédestre (Poda, 1761); the
name is not available
p. 252, replace entry D. sareptanum by:
sareptanum euxinum Suvorov, 1915 E: ST UK
kubanicum Plavilstshikov, 1934d: 120
p. 252, correct year for Dorcadion scrobicolle morulum Holzschuh to 1995
p. 255, add original genus combination (Compsodorcadion) to Dorcadion cephalotes Jakovlev, 1889
p. 262, remove albodiversum Pic, 1945 from synonymy of Iberodorcadion fuliginator pyrenaeum (Germar,
1839); the name is infrasubspecific
p. 264, correct author and data of Ibidimorphum [in synonymy with Oleocampus Chevrolat, 1835] to
Motschulsky 1860: 152
p. 264, replace Oleocampus clarus clarus Pascoe, 1859 by O. clarus Pascoe, 1859
p. 267, correct page of description of Lamia textor Linnaeus, 1758 to 392
p. 270, correct type species of Choeromorpha Chevrolat, 1843 to Choeromorpha pigra Chevrolat, 1843 [by
monotypy; not Aurivillius, 1920]
p. 270, correct year ïor fuscus Olivier and villicus Olivier [in synonymy with Coptops aedificator (Fabricius,
1792)] to 1797
p. 272, correct data of subobliterata Pic, 1902: 62 [in synonymy with Mesosa longipennis Bates, 1873 to
subobliterata Pic, 1901m: 62
p. 279, replace entry Blepephaeus undulatus Pu, 1999 by:
puae Lin, nom. nov. A: YUN
undulatus Pu, 1999: 79 (Perihammus) [nee Blepephaeus undulatus (Pic, 1930) transferred from Psaromaia by Breuning,
1961]
p. 279, correct year and page of description for Cereopsius Pascoe and C. exoletus to Pascoe, 1857b: 105 and
Pascoe, 1857, respectively
p. 279, correct spelling of Falsapriona [in synonymy with Cerosterna Dejean, 1835] to Falsopriona Pic, 1925
p. 281, correct spelling oïHoplothrix Gahan, 1888 to Haplothrix Gahan, 1888
p. 281, correct year and original combination of Macrochenus tigrinus (Olivier) to 1797 and Lamia, respectively
p. 281, correct spelling of Mimothetus Pic, 1935 and Mimosthetus Pic, 1935 to Mimothestus
p. 282, correct spelling of szechenyanus Frivaldszky, 1892 to szechenyianus [in synonymy with Monochamus
millegranus (Bates, 1891)]
p. 290, remove schrammi Pic, 1945 from synonymy oïParmena cruciata Pic, 1945; the name is infrasubspecific
 ERRATA 45
p. 296, remove planeti Pic, 1945 and sinuatesignata Pic, 1945 from synonymy of Oberea erythrocephala
erythrocephala (Schrank, 1776); the names are infrasubspecific
p. 299, remove semijuncta Pic, 1945 from synonymy of Oberea maculicollis P. H. Lucas, 1842; the name is
infrasubspecific
p. 299, add nigromaculicollis Breuning, 1960: 35 in synonymy with Oberea nigriceps (A. White, 1844)
p. 299, add kwangtitngensis Breuning, 1960: 37 and rufoantennata Breuning, 1960: 37 in synonymy with
Oberea notata Pic, 1936
p. 300, add SD to distribution of Oberea walkeri Gahan, 1894, and add atrosternalis Breuning, 1960: 38,
nigrobasicollis Breuning, I960: 38, and sikkimensis Breuning, 1960: 38 in its synonymy
p. 303, change entry for Phytoecia circumdata circumdata Kraatz, 1882 and P. circumdata pilosicollis
Holzschuh, 1981 to P. circumdata Kraatz, 1882 and P. pilosicollis Holzschuh, 1981, respectively
p. 303, change entry for Phytoecia armeniaca armeniaca Frivaldszky, 1878 to P. armeniaca Frivaldszky, 1878,
and entry for P. armeniaca testaceovittata Pic, 1934 modify as follows:
testaceovittata nataliLobanov, 1994a: 105 E: AB
testaceovittata testaceovittata Pic, 1934c: 18 (Musaria) A: IN
iranica Villiers, 1960b: 99
p. 304, add CR to distribution of Phytoecia cephalotes Küster, 1846
p. 306, change data for annulifera Pic [in synonymy with Phytoecia croceipes Reiche & Saulcy, 1858] to
annulifera T. Pic, 1900b: 67
p. 307, add subannulipes Pic, 1915: 11 to synonymy of Phytoecia icterica (Schaller, 1783)
p. 308, correct entry for Pilemia annulata (Hampe, 1852) to:
anmdata annulata Hampe, 1852b: 315 (Phytoecia) E: AB A: IN TR
annulata wawerkana Reitter, 1905b: 239 A: TR
angorensis Pic, 1952a: 2
p. 310,removesubbicolor Breuning, 1958fromsynonymy with Exocentrus diversiceps Pic, 1931; it is correctly
listed as valid species on p. 311
p. 324, correct spelling of Eutetrapha viridis Pu & Jin, 1991 to E. virides Pu & Jin, 1991
p. 328, place yuasai Gressitt, 1935 in synonymy with Menés ia sulphurata (Gebier, 1825)
p. 330, remove mesmini Pic, 1910 from synonymy of Saperda perforata (Pallas, 1773), the name is
infrasubspecific
p. 331, remove Saperda subobliterata Pic, 1910; it is not available [introduced as a variety in a paper, in which
also subspecies are used], and use mandschukuoensis Breuning, 1943 as valid
p. 334, add, under Lamiinae, as nomen dubium:
Lamia aspera Roemer, 1789: 46 [pi. 5] E: IT
p. 335, correct year for rufotestacea Kraatz [in synonymy with Zeugophora turneri Power, 1863] to 1872c
p. 336, add original combination (Clytraxeloma) to Temnaspis nigropunctata (Pic,1896)
p. 337, add anatolica Pic, 1916a: 10 in synonymy with Orsodacne humeralis humeralis Latreille, 1804
p. 339, correct "family" Bruchinae to "subfamily"
p. 340, add to Acanthoscelides valid species:
monophthalmus Schaeffer, 1907: 300 (Bruchus) Ei: GR (Cyprus) Ai: JA (Bonin Isl.) TAI YUN AFRi NTR
ORRi
p. 347, add under Bruchidius, as species incertae sedis:
gagliardii Pic, 1933d: 14 E: IT
p. 350, add pilifer Olivier, 1791: 562 (Curculio) [RN] in synonymy with Bruchus atomarius (Linnaeus, 1760)
p. 353, add under Bruchinae, nomen dubium:
Bruchus bipunctatus Roemer, 1789: 6 [pi. 4] E: "Rhaetia"
p. 356, under Donacia thalassina intermedia, correct spelling of semenowi to semenovi
p. 360, add augustoduni Pic, 1916a: 6 and semijuncta Pic, 1916a: 10 in synonymy with Crioceris
duodecimpunctata (Linnaeus, 1758)
p. 360, add (Lenta) to entry crucifera Tenkl, 1929 [in synonymy with Crioceris macilenta Weise, 1881]
p. 360, add [HN] to entry weisei Heyede, 1906 [in synonymy with Crioceris macilenta Weise, 1881]
46 ERRATA
p. 361, add donceeli Pic, 1916a: 6, montandoni Pic, 1916a: 6, russica Pic, 1916a: 6, semireducta Pic, 1916a: 6
and turcica Pic, 1916a: 6 in synonymy with Crioceris quatuordecimpunctata (Scopoli, 1763)
p. 361, entry epipleuralis Takizawa, 1983 is unverified
p. 365, add to Lilioceris valid species, and synonym:
dohertyi Jacoby, 1908: 79 (Crioceris) A: AP
glabra Jakob, 1961: 61 A: YUN
jakobi R. E. White, 1981: 31 [RN] A: FUJ JIA
minima Jakob, 1961: 61 [HN]
laeviuscula Weise, 1882: 69 (Crioceris) E: SP
p. 366, add to Lilioceris valid species:
semicostata Jacoby, 1908: 77 (Crioceris) A: SD ORR
semihirsuta Jacoby, 1908: 75 (Crioceris) A: UP ORR
p. 366, add FR SZ to distribution of Lilioceris tibialis (A. Villa & G. B. Villa, 1838)
p. 366, correct year for alpina L. Redtenbacher [in synonymy with Lilioceris tibialis (A. Villa & G. B. Villa,
1838)] to 1848, and change the original combination to Lema
p. 367, add SV to distribution of Oulema septentrionis Weise, 1880
p. 368, add [RN] to Anisodera guerinii Baly, 1858, and the following synonyms:
ferruginea Guérin-Méneville, 1840c: 333 [HN]
foveolata Pic, 1924b: 29
sinuaticollis Pic, 1926b: 10
p. 368, add barbicornis Weise, 1897c: 119 in synonymy with Anisodera propinqua Baly, 1888
p. 368, add following synonyms to Estigmena chinensis (Hope, 1840):
atricollis Pic, 1924b: 29
latior Pic, 1924b: 30
nigripennis Pic, 1924b: 29
ruficolor Pic, 1924b: 30
p. 371, add following synonyms to Callispa dimidiatipennis Baly, 1858:
latithorax Pic, 1927b: 25
vicina Pic, 1927b: 25
p. 371, add curtipennis Pic, 1926j: 324 in synonymy with Callispa pallida Gestro, 1888
p. 371, add to Callispa valid species:
procédera Uhmann, 1939: 1 A: YUN ORR
p. 378, add to Cassida valid species:
tectiformis Normand, 1949: 95 N: TU
p. 378, add SP to distribution of Cassida viridis (Linnaeus, 1758)
p. 379, remove entry paliji Matis, 1970: 31 from Cassida weisei Jakobson, 1894. The species is correctly listed
as valid on p. 376
p. 380, add Falsocassidula Pic, 1939g: 149 in synonymy with Nabathaea Spaeth, 1911
p. 380, add rabinovitchi Pic, 1939g: 149 (Cassida) in synonymy with Nabathaeapygmaea Spaeth, 1911
p. 381, add atripennis Pic, 1926a: 10 in synonymy with Agonita chinensis (Weise, 1922)
p. 382, add annamita Pic, 1930b: 4 in synonymy with Gonophorapulchella Gestro, 1888
p. 382, add to Micrispa as valid species:
dentatithorax Pic, 1924a: 4 (Gonophora) A: YUN ORR and place yunnanica (Chen & Sun, 1962) in its
synonymy
p. 382, add to Downesia valid species and synonyms:
auberti Fairmaire, 1888d: 375 (Hanoia) A: FUJ ORR
basipennis Pic, 1924a: 5
diversepunctata Pic, 1935b: 172
latenigra Pic, 1924a: 5
nigra Pic, 1934b: 10
nigritarsis Pic, 1924a: 4
p. 383, add following synonyms to Sinagonia maculigera (Gestro, 1888):
juncta Pic, 1924a: 5
multimaculata Pic, 1924a: 5
p. 383, add to Asamangulia valid species:
yonakuni Kimoto & Gressitt, 1966: 570 A: JA (Ryukyu)
 ERRATA 47
p. 383/384, transfer Dactylispa soror Weise, 1897c as synonym of Dactylispa nigripennis Motschulsky, 1860
p. 384, add to Dactylispa valid species:
filiola Weise, 1897c: 135 A: FLU GUA GUX HAI JIA ORR
p. 384, add (Hispa) to entry Dactylispa nigromaculata Motschulsky, 1860
p. 385, correct spelling of Dactylispa sjoestedti Uhmann, 1928 to D. sjostedti
p. 385, remove entry Dactylispa xanthospila (Gestro, 1890). It is correctly listed as synonym of D.
nigromaculata (Motschulsky, 1860), p. 384
p. 385, add NC SC under Asian distribution of Dactylispa (Platypriella) excisa excisa (Kraatz, 1879)
p. 385, add SC under Asian distribution of Dactylispa {Platypriella) subquadrata subquadrata (Baly, 1874)
p. 385, add rufescens Shirôzu, 1957: 55 in synonymy with Dactylispa (Triplispa) angulosa (Solsky, 1872)
p. 386, add to Dactylispa (Triplispa) valid species:
koreana An, Kwon & Lee, 1985: 5 A: SC
p. 387, add SC to Asian distribution of Hispellimis chinensis Gressitt, 1950
p. 387, add SC to Asian distribution of Hispellinus morens (Baly, 1874)
p. 388, add atripes Pic, 1925a: 16 in synonymy with Leptispa abdominalis Baly, 1858
p. 388, add ORR to distribution of Leptispa abdominalis Baly, 1858
p. 388, add to Leptispa valid species:
bengalensis Takizawa, 1989b: 330 A: SD
p. 389, add to Leptispa valid species:
parallelaparallela Gestro, 1890: 220 (Downesia) A: SD
p. 389, add vietnamica L. N. Medvedev, 1992f: 154 in synonymy of Notosacantha sauteri (Spaeth, 1914)
p. 389, add HAI to Asian distribution of Oncocephala atrangula Gressitt, 1938
p. 390, correct year of Oncocephala tuberculata (Olivier) to 1797
p. 390, add EN and LT to distribution of Chrysomela collaris Linnaeus, 1758
p. 390, add EN and LT to distribution of Chrysomela cuprea Fabricius, 1775
p. 390, add EN and LT to distribution of Chrysomela lapponica Linnaeus, 1758
p. 391, add EN and LT to distribution of Chrysomela populi Linnaeus, 1758
p. 391, add EN, LA and LT to distribution of Chrysomela saliceti saliceti (Weise, 1884)
p. 391, add EN and LT to distribution of Chrysomela tremula tremula Fabricius, 1787
p. 391, add EN and LT to distribution of Chrysomela vigintipunctata vigintipunctata (Scopoli, 1763)
p. 392, add EN and LT to distribution of Plagiodera versicolora (Laicharting, 1781)
p. 392, add to Plagiodera valid species:
borealis Gressitt & Kimoto, 1963: 344 A: LIA
p. 393, for Plagiosterna aenea aenea (Linnaeus, 1758), add to distribution: CT DE EN FI LT NR NT SV
p. 393, change spelling of purpureatincta to purpureotincta, under Plagiosterna aenea aenea (Linnaeus, 1758)
p. 393, add EN and LT to distribution of Gastrophysa (Gastrophysa) polygoni polygoni (Linnaeus, 1758)
p. 393/394, add LT to distribution of Gastrophysa (Gastrophysa) viridula viridula (DeGeer, 1775)
p. 394, under Gastrophysa viridula viridula DeGeer, 1775, add original combination (Gastroidea) to cyanescens
Weise, 1898
p. 394, add LA and LT to distribution of Phratora (Phratora) vulgatissima (Linnaeus, 1758)
p. 395, add EN and LT to distribution of Phratora (Phyllodecta) atrovirens (Cornelius, 1857)
p. 395, add EN and LT to distribution of Phratora (Phyllodecta) laticollis (Suffrian, 1851)
p. 395, add EN and LT to distribution of Phratora (Phyllodecta) tibialis tibialis (Suffrian, 1851)
p. 395, add EN and LT to distribution of Phratora (Phyllodecta) vitellinae (Linnaeus, 1758)
p. 396, add SP to distribution of Neophaedon pseudopyritosus (Codina Padilla, 1963)
p. 396, add IN to Asian distribution of Neophaedon pyritosus (Rossi, 1792)
p. 396, add SP to distribution of Neophaedon pseudopyritosus (Codina Padilla, 1963)
p. 396, add IN to distribution of Neophaedon pyritosus (Rossi, 1792)
p. 396, add EN and LT to distribution of Phaedon (Phaedon) armoraciae (Linnaeus, 1758)
p. 396-397, add EN and LT to distribution of Phaedon (Phaedon) cochleariae cochleariae (Fabricius, 1792)
p. 397, add EN to distribution of Phaedon (Phaedon) concinnus concinnus Stephens, 1834
p. 397, add LT to distribution of Phaedon (Phaedon) laevigatus laevigatus (Duftschmid, 1825)
p. 397, add AU to distribution of Phaedon (Phaedon) salicinus (Heer, 1845)
p. 396, add to Neophaedon valid species:
48 ERRATA
sichiianicus Lopatin, 2006c: 597 A: SCH
p. 397, add violaceus Weise, 1884e: 543 in synonymy with Phaedon laevigatus laevigatus (Duftschmid, 1825)
p. 398, add EN and LT to distribution of Prasocuris (Hydrothassa) glabra (Herbst, 1783)
p. 398, add EN and LT to distribution of Prasocuris (Hydrothassa) hannoveriana (Fabncius, 1775)
p. 398, add EN and: LT to distribution of Prasocuris {Hydrothassa) marginella marginella (Linnaeus, 1758)
p. 398, add EN and LT to distribution of Prasocuris (Prasocuris) junci (Brahm, 1790)
p. 398, add EN and LT to distribution of Prasocuris (Prasocuris) phellandrii (Linnaeus, 1758)
p. 399, remove LA from distribution of Chrysolina (Allochrysolina) fuliginosa galii (Weise, 1884)
p. 400, correct type species of subgenus Anopachys Motschulsky, 1860 to Chrysomela asclepiadis A. Villa & G.
B. Villa, 1833
p. 400, add BY, LA and LT to distribution of Chrysolina (Anopachys) aurichalcea (Mannerheim, 1825)
p. 402, add LT to distribution of Chrysolina (Chalcoidea) analis (Linnaeus, 1767)
p. 402, add EN and LT to distribution of Chrysolina (Chalcoidea) carnifex carnifex (Fabricius, 1792)
p. 403, correct year for Chrysolina (Chalcoidea) marginata dierythra (Rottenberg) to 1872a
p. 403, add EN to distribution of Chrysolina (Chalcoidea) marginata marginata (Linnaeus, 1758)
p. 403, add concolor Demaison, 1896: 12 (Chrysomela) in synonymy with Chrysolina (Chalcoidea) sellata
(Weise, 1894)
p. 404, add to synonymy of Chrysocrosita Bechyné, 1950:
Lithocrosita L. N. Medvedev, 1982: 79 type species Chrysomela rugulosa auct. [nee Gebier, 1841] (= Crosita
concinna (Weise, 1894)
p. 405, remove LA from the distribution of Chrysolina (Chrysolina) rufa squalida (Suffrian, 1851 )
p. 405, add in synonymy with Chrysolina (Chrysolina) rufa squalida (Suffrian, 1851) entry:
dahli Suffrian, 1851a: 28 (Chrysomela)
p. 405, add DE EN FI NR to distribution of Chrysolina (Chrysolina) staphylaea staphylaea (Linnaeus, 1758)
p. 405, add EN to distribution of Chrysolina (Chrysomorpha) cerealis cerealis (Linnaeus, 1767)
p. 406, remove entry aerea L. Redtenbacher from synonymy of Chrysolina globosa globosa (Panzer, 1803); the
name is not available
p. 407, under Chrysolina (Colaphosoma) sturmi Westhoff, 1882: 268, remove entry vulgatissima Schrank,
1781a: 69 [HN]: it is a misidentification
p. 408, add EN to distribution of Chrysolina (Erythrochrysa) politapolita (Linnaeus, 1758)
p. 408, add EN and LT to distribution of Chrysolina (Euchrysolina) graminis graminis (Linnaeus, 1758)
p. 408, correct page of description of taupini Mallet, 1924 from 74 to 77 [under Chrysolina (Euchrysolina)
graminis graminis Linnaeus, 1758]
p. 408, add EN and LT to distribution of Chrysolina (Fastuolina)fastuosafastuosa (Scopoli, 1763)
p. 408, add TD to distribution of Chrysolina (Gnathomela) dolens (Kraatz, 1885)
p. 408, add chrysomeliformis Bogatchev, 1961: 112 (Lobodera) in synonymy with Chrysolina (Gnathomela)
dolens (Kraatz, 1885)
p. 409, add EN and LT to distribution of Chrysolina (Hypericia) geminata (Paykull, 1799)
p. 409, add EN and LT to distribution of Chrysolina (Hypericia) hyperici hyperici (Forster, 1771)
p. 413, add [HN] to daccordii Lopatin, 2001: 135 [in synonymy with Chrysolina (Pezocrositd) juldusana
Lopatin, 1962]
p. 414, add to Semenowia Weise, 1889 valid species:
daccordii L. N. Medvedev & Sprecher-Uebersax, 1999: 366 (Semenovia) A: NP
p. 414, change original combination (Chrysomela) to (Semenowia), in Chrysolina (Semenowia) chalcea Weise,
1889
p. 414, add EN to distribution of Chrysolina (Sphaeromela) varions (Schaller, 1783)
p. 415, change original combination (Oreina) to (Chrysochloa) in Chrysolina (Stichoptera) colasi CQhos,-1952
p. 415, add EN and LT to distribution of Chrysolina (Stichoptera) gypsophilae (Küster, 1845)
p. 415, add EN and LT to distribution of Chrysolina (Stichoptera) sanguinolenta (Linnaeus 1758)
p. 416, add LA and LT to distribution of Chrysolina (Sulcicollis) oricalcia (O. F. Müller, 1776)
p. 416, for Chrysolina (Synerga) coerulans angelica Reiche & Saulcy, 1858, change E: IN IS SY to A: IN IS SY
p. 416, add LA to distribution of Chrysolina (Synerga) coerulans coerulans (L. G. Scriba, 1791)
p. 417, add original combination (Chrysomela) to yanonigra Jolivet, 1943 [in synonymy with Chrysolina
(Synerga) herbacea herbacea (Duftschmid, 1825)]
 ERRATA 49
p. 419, under Chrysolina incertae sedis, remove entry concolor Demaison, 1896: 12 (Chrysomela)
p. 419, under Chrysolina incertae sedis, lepida Brullé, 1832 described from Greece was replaced by brullei
Poitevin, 1934. Modify the entry as follows:
brullei Portevin, 1934: 232 {Chrysomela) [RN] fi: GR
lepida Brullé, 1832: 270 (Chrysomela) [HN]
p. 420, remove entry betulae Linnaeus, 1758: 369 (Chrysomela) E: SV
p. 420, under Crosita Motschulsky, 1860, remove entry: Lithocrosita L. N. Medvedev, 1982: 79 type species
Chrysomela rugulosa Gebier, 1841
p. 422, add BY and GR to distribution of Oreina (Allorina) caerulea (Olivier, 1790)
p. 423, add flammea Jakob, 1964: 11 (Chrysochloa) in synonymy with Oreina (Chyrsochloa) speciosissima
fuscoaenea Schummel, 1844
p. 423, under Oreina speciosissima fuscoaenea Schummel, 1844, add [RN] to letzneri Weise, 1883a: 214
(Orinà)
p. 424, add GR to distribution of Oreina (Oreina) alpestris balcanica (Weise, 1883)
p. 425, under Oreina speciosa speciosa, change original combination of illustris G. Müller, 1948 from
(Chrysomela) to (Chrysochloa)
p. 425, correct distributrion of Oreina sulcata basilea (Gebier, 1823) to A: KZ WS
p. 425, correct distributrion of Oreina sulcata sulcata Gebier, 1823) to A: ES
p. 425, add SP to distribution of Oreina (Oreina) viridis viridis (Duftschmid, 1825)
p. 426, add GR to distribution of Oreina (Virgulatorina) virgulata ljubetensis (Apfelbeck, 1912)
p. 427, add tibialis L. Redtenbacher, 1850: 50 (Colaphus) in synonymy with Colaphellus apicalis (Ménétriés,
1849)
p. 427, add to Colaphellus valid subspecies:
pulchellus arabicus L. N. Medvedev, 1996: 249 (Colaphus) A: SA
p. 427, add LA and LT to distribution of Colaphellus sophiae sophiae (Schaller, 1783)
p. 427, add implagatum Achard, 1923c: 128 in synonymy with Colaspidema (Colaphomega) signatipenne
Guérin-Méneville, 1844: 296
p. 427, add original combination (Chrysomela) to viennensis Schrank, 1781 [in synonymy with Colaphellus
sophiae sophiae (Schaller, 1783)]
p. 427, add original combination (Colaphus) to Colaphellus tenuipes Weise, 1883
p. 427, add implagatum Achard, 1923c: 128 in synonymy with Colaspidema (Colaphomega) signatipenne
Guérin-Méneville, 1844
p. 428, add "Persia" to distribution of Cystocnemis concolor
p. 428, add original combination (Oreomela) to mohri Daccordi, 1975 [in synonymy with Cystocnemis concolor
(Kraatz, 1879)]
p. 428, replace LA by LT in distribution of Entomoscelis adonidis (Pallas, 1771)
p. 428, remove detersa Bedel, 1915 and viberti Bedel, 1915 from synonymy of Entomoloscelis rumicis
(Fabricius, 1787); both are unavailable
p. 428, add valid genus and species:
genus Falsocolaphellus Pic, 1928c: 10 type species Falsocolaphellus syriacus Pic, 1928
syriacus Pic, 1928c: 10 A: S Y
p. 431, add to Suinzona valid species:
monticola S.-H. Chen, 1961: 433 A: SCH
p. 431, correct spelling of transsylvanicus Fuss, 1861 to transsilvanicus Fuss, 1861 and add original combination
(Phaedon) [in synonymy with Sclerophaedon carniolicus (Germar, 1824)]
p. 432, remove all entries under Acacicola Lea, 1902; see below, under Peltoschema Reitter
p. 433, add EN and LT to distribution of Gonioctena (Gonioctena) decemnotata (Marsham, 1802)
p. 433, add EN and LT to distribution of Gonioctena (Gonioctena) linnaeana linnaeana (Schrank, 1781)
p. 434, add LT to distribution of Gonioctena (Gonioctena) viminalis viminalis (Linnaeus, 1758)
p. 434, correct year for bicolor Kraatz, 1979 to 1879a [in synonymy with Gonioctena viminalis viminalis
(Linnaeus, 1758)]
p. 434, remove haemorrhoidalis (Scopoli, 1763)fromsynonymy with Gonioctena viminalis viminalis (Linnaeus,
1758), it is a misidentification, and not available
50 ERRATA
p. 435, correct spelling of notâtihorax (Pic, 1924) to notatithorax (Pic, 1924) [in synonymy with Gonioctena
viminalis viminalis (Linnaeus, 1758)]
p. 435, correct original combination (Chrysomela) to (Coccinella) in signata (Scopoli, 1763) [in synonymy with
Gonioctena viminalis viminalis (Linnaeus, 1758)]
p. 435, add EN to distribution of Gonioctena (Goniomena) intermedia (Helliesen, 1913)
p. 435, add EN to distribution of Gonioctena (Goniomena) pallida (Linnaeus, 1758)
p. 435, add EN to distribution of Gonioctena (Goniomena) quinquepunctata quinquepunctata (Fabricius, 1787)
p. 436, change original combination in type species of subgenus Spartomena Reitter, 1913 from Phytodecta
fornicata to Gonioctena fornicata Brüggemann, 1873
p. 436, add berytensis Pie, 1916a: 10, binotaticollis Pic, 1916a: 10 and obliquemaculata Pic, 1916a: 10
(Phytodecta) in synonymy with Gonioctena (Spartomena) fornicata Brüggemann, 1873
437, add to Paropsides Motschulsky, 1860 valid species:
nigropunctata Jacoby, 1892b: 918 A: NP ORR
p. 437, add after Paropsides Motschulsky, 1860 following entries:
genus Peltoschema Reitter, 1880: 4 type species Peltoschemafilicornis Reitter, 1880
Acacicola Lea, 1902: 392 type species Acacicola tristis Lea, 1902
Pyrgo Weise, 1901c: 166 [HN]
Pyrgoides Kelly & Reid 1999:264 [RN] type species Paropsis hamadryas Stâl, 1860
subgenus Asiparopsis S.-H. Chen, 1934c: 69 type species Paropsidespardalis Jacoby, 1892
convexa Weise, 1902i: 368 (Pyrgo) A: GUX ORR
mausonensis Achard, 1914b: 72 (Pyrgo) [RN]
unicolor S.-H. Chen, 1934c: 70 (Asiparopsis)
pp. 437/438, correc t author of Timarcha from Latreille, 1829 to Samouelle, 1819: 213 and change the type
species from Tenebrio laevigatus Linnaeus, 1767 to Chrysomela tenebricosa Fabricius, 1775. Samouelle
included two species in Timarcha: Chrysomela tenebricosa Fabricius, 1775 (p. 213) and Chrysomela
coriaria Laicharting, 1781 (p. 445). Chrysomela tenebricosa was designated as the type species of Timarcha
by Chevrolat (1843: 655). The change of the type species does not affect the current taxonomic division of
Timarcha
p. 451, remove entry: genus Lochmaeina L. N. Medvedev, 1956 with the species rosea L. N. Medvedev, 1956 [it
is a synonym of Sangariolapunctatostriata Motsch. (Alticinae) (p. 558)]
p. 464, removepicta S.-H. Chen, 1942 from Sphenoraia; it is correctly placed in Aplosonyx (p. 456)
p. 470, remove discolor Faldermann, 1837 from Calomicrus; it is correctly placed in Luperus (p. 479)
p. 476, change Hoplasoma minor to Hoplasoma minus; the gender of Hoplasoma is neuter
p. 479, correct year for aetnensis Rottenberg [in synonymy with Luperus biraghii Ragusa, 1871 ] to 1872a
p. 480, remove moorii Baly, 1874 from Luperus; it is correctly placed in Stenoluperus (p. 560)
p. 483, add to Monolepta valid species:
gestroi Jacoby, 1892b: 983 A: UP ORR
p. 483, remove Monolepta madrasensis Wilcox, 1973 and its synonyms duvivieri Jacoby, 1904 and lineata
Weise, 1915; all are correctly placed in synonymy with Monolepta sexlineata Chûjô, 1938 (p. 484)
p. 484, add to Monolepta valid species:
picturata Jacoby, 1896c: 292 A: UP ORR
subflavipennis Kimoto, 1989: 154 A: GAN ORR
p. 485, remove costata Takizawa & Basu, 1987 as valid species in Paleosepharia- this taxon is correctly placed
in synonymy witii Paleosepharia pokharensis Kimoto, 1982 (p. 486)
p. 486, add SD to distribution of Paleosepharia pokharensis Kimoto, 1982
p. 486, transfer from Japonitata to Paraplotes valid species:
indica Takizawa & Basu, 1987: 279 A: SD
pp. 488/489, Charaea Baly, 1878 has priority over Taphinellina Maulik, 1936; use Charaea as valid
p. 491, add under Acrocrypta Baly, 1862, synonyms:
Pseudocrypta L. N. Medvedev, 1997g: 78 type species Pseudocrypta nigripennis L. N. Medvedev, 1996
Sebaethiella L. N. Medvedev, 1994b: 16 type species Sebaethiella mindanaica L. N. Medvedev, 1993
p. 491, add to Acrocrypta Baly, 1862 valid species:
assamensis Jacoby, 1893b: 151 A: GUA SCH YUN ORR
intermedia Jacoby, 1892b: 925 (Sphaerometopa) A: YUN ORR
 ERRATA 51
p. 493, coTKCtßlipendulae Chashchina, 2006: 858 E: CT (Chelyabinskya oblast') as valid species, not synonym
of engstromi (J. R. Sahlberg, 1893) and remove "CT (Chelyabinskya oblast')" from distribution of latter
species
p. 494, add A: TR to distribution of Altica longicollis Allard, 1860
p. 495, add A: TR to distribution of Altica talyshana Konstantinov, 1995
p. 495, add in synonymy with Amphimela Chapuis, 1875:
Throscoryssa Maulik, 1928: 159 type species Throscoryssa citri Maulik, 1928
p. 495, add to Amphimela Chapuis, 1875 valid species:
citri Maulik, 1928: 160 (Throscoryssa) A: GUA ORR
p. 495, remove Amphimela shirahatai Chûjô, 1957; it is correctly placed as Podagricomela shirahatai Chûjô,
1957,p. 551
p. 495, remove entry: genus Amphimeloides Jacoby, 1887 and the species bistripunctata S.-H. Chen, 1933 and
sexmaculata Kimoto & Takizawa, 2002 [Amphimeloides was synonymized with the genus Ivalia]
p. 495, correct distribution of Angulaphthona latipennis (Pic, 1921) to A: SA YE N: EG
p. 496, add albinea C. G. Thomson, 1866: 182 (Teinodactyla) in synonymy with Aphthona abdominalis
(Duftschmid, 1825)
p. 496, add A: TR to distribution of Aphthona atrocaerulea (Stephens, 1831)
p. 496, add SK to distribution of Aphthona beckeri Jakobson, 1896
p. 497, add nigella Allard, 1861: 332 in synonymy with Aphthona carbonaria Rosenhauer, 1856
p. 497, under Aphthona czwalinai Weise, 1888, change 1889e to 18891 in coerulescens Weise, 1889
p. 498, under Aphthona gerhardi, add [HN] to nepalensis Scherer, 1989: 42 (Bhutajana)
p. 498, add SD to distribution of Aphthona indica Jacoby, 1900
p. 500, remove entry: plenifrons Wollaston, 1864: 408 (Haltica) from synonymy of Aphthona nonstriata and
place it as valid species in Heyrovskya (p. 519)
p. 502, add cyrenaica Heikertinger, 1944: 80 in synonymy with Aphthona vaulogeri Pic, 1910
p. 503, add crassina Gistel, 1857a: 80 in synonymy with Apteropeda globosa (Illiger, 1794)
p. 503, add entries:
genus Argopistoides Jacoby, 1892b: 931 type species Argopistoides septempunctata Jacoby, 1892
Torodera Weise, 1902: 163 type species Torodera octomaculata Weise, 1902
septempunctata Jacoby, 1892b: 932 A: FUJ GUA ORR
sexpunctata Scherer, 1987: 69 (Torodera) A: BT NP SD
p. 504, add LA to distribution of Argopus nigritarsis (Gebier, 1823)
p. 504, add entries:
genus Asiophrida Medvedev, 2000c: 176,180 type species Haltica marmorea Wiedemann, 1819
subgenus Asiophrida Medvedev, 2000c: 176, 180 type species Haltica marmorea Wiedemann, 1819
scaphoides Baly, 1865g: 430 (Podontid) A: FUJ GAN GUI HUB JIA NP SCH TAI XIZ YUN ZHE
binduta Maulik, 1926: 233 (Ophrida)
spectabilis Baly, 1862c: 452 (Podontid) A: FUJ GUI HUB JIA SCH TAI XIZ YUN ZHE "Korea"
rufoflava Fairmaire, 1889a: 73 (Podontid)
xanthospilota Baly, 1881: 54 (Blepharida) A: HEB SCH
p. 504, add IN to distribution of Batophila olexai Krai, 1964
p. 505, correct year for Himalalta Medvedev from 1940 to 1990 [in synonymy with Benedictus Scherer, 1969]
p. 505, add HKG to distribution of Bikasha collaris (Baly, 1877)
p. 506, change authorship of aeneicolor to Pic, 1915a: 5 [in synonymy with Chaetocnema angustula
(Rosenhauer, 1847)]
p. 506, add IN to Asian distribution of Chaetocnema confusa (Boheman, 1851)
p. 507, add CZ to distribution of Chaetocnemaprocerula (Rosenhauer, 1856)
p. 507, add E: CT and A: TR to distribution of Chaetocnema sahlbergii (Gyllenhal, 1827)
p. 508, add SK to distribution of Chaetocnema breviuscula (Faldermann, 1837)
p. 510, add subunicolor Pic, 1918d: 18 (Chalcoides) in synonymy with Crepidodera aurata (Marsham, 1802)
p. 510, add purpurascens Pic, 1918d: 18 (Chalcoides) in synonymy with Crepidodera aurea (Geoffroy, 1785)
p. 512, add A: TR to distribution of Crepidodera nigricoxis Allard, 1878
p. 512, delete SV and add IS to distribution of Dibolia depressiuscula Letzner, 1847
p. 512, add A: TR to distribution of Dibolia numidica Doguet, 1971
52 ERRATA
p. 514, add HKG to distribution of Euphitrea gressitti (Chûjô, 1965)
p. 515, add entry:
genus Eutrea Baly, 1875a: 24 type species Eutrea bowringii Baly, 1875
bowringii Baly, 1875a: 25 A: HKG
p. 516, add to Hemipyxis valid species:
monticola Scherer, 1969: 195 {Sebaethe) A: SD
p. 517, add to Hespera valid species:
bengalensis Scherer, 1969: 40 A: SD
p. 518, add HKG to distribution of Hespera lomasa Maulik, 1926
p. 518, add albopilosa Chûjô, 1957b: 7 in synonymy with Hespera lomasa Maulik, 1926
p. 519, remove genus Himalalta Medvedev, 1990 with species brevlcornis Medvedev, 1990, and striata
Medvedev, 1990. They are correctly placed in synonymy witii Benedictus Scherer, 1969 (p. 505)
p. 520, add HKG to distribution of Hypnosis parvula Jacoby, 1884
p. 520, add Amphimeloides Jacoby, 1887a: 96 type species Amphimeloides dorsalis Jacoby, 1887 and
Ancyloscelis Ogloblin, 1930: 100 type species Mniophila ruflcollis Motschulsky, 1866 in synonymy with
Ivalia Jacoby, 1887
p. 520, add to Ivalia valid species:
bistripunctata S.-H. Chen, 1933e: 188 {Amphimeloides) A: JA (Honshu, Kyushu)
sexmaculata Kimoto & Takizawa, 2002: 143 {Amphimeloides) A: NP
p. 522, add KA to distribution of Longitarsus almorae Maulik, 1926
p. 523, add caligans Krai, 1945: 31 and septorum Buysson, 1908: 29 {Thyamis) in synonymy with Longitarsus
ballotae Marsham, 1802
p. 523, add SD UP and AFRi to distribution of Longitarsus belgaumensis Jacoby, 1896
p. 524, add arbustorum Gistel, 1857a: 9 {Crepidodera) [NO] in synonymy with Longitarsus curtus (Allard,
1861)
p. 524, add SD to distribution of Longitarsus cyanipennis Bryant, 1924
p. 526, add RO to distribution of Longitarsus gracilis Kutschera, 1864
p. 526, add A: TR to distribution of Longitarsus helvolus Kutschera, 1864
p. 526, add SK to distribution of Longitarsus juncicola (Foudras, 1860)
p. 527, change 1889e to 18891 in Longitarsus longiseta Weise
p. 529, add scutellata Gistel, 1857a: 24, 37 {Thyamis) in synonymy with Longitarsus ochroleucus ochroleucus
(Marsham, 1802)
p. 529, correct spelling of Longitarsus ozbek to ozbeki
p. 530, addperplexus Dollman, 1912: 188 in synonymy with Longitarsusplantagomaritimus Dollman, 1912
p. 531, add eryngii Gistel, 1831: 303 {Haltica) in synonymy with Longitarsus rutilus (Illiger, 1807)
p. 531, add SK to distribution of Longitarsus scrobipennis Heikertinger, 1913
p. 531, add SK and IN to distribution of Longitarsus scutellaris Mulsant & Rey, 1874
p. 531, add A: TR to distribution of Longitarsus {Longitarsus) stragulatus stragulatus Foudras, 1860
p. 533, add A: TR to distribution of Longitarsus {Testergus) corynthius corynthius Reiche & Saulcy, 1858
p. 533, add A: TR to distribution of Longitarsus {Testergus) onosmae Peyerimhoff, 1912
p. 534, add ST (North Caucasus) to distribution of Longitarsus {Testergus) truncatellus Weise, 1890
p. 535, correct spelling of Lypnae Baly, 1876 to Lypnea Baly, 1876
p. 535, add to Manobia valid species:
coomani Chen, 1934f: 385 A: GUX ORR
p. 537, Microcrepis is masculinum, change endings of species to apterus, brunneus, castaneus and politus
p. 538, add bryozoa Gistel, 1857a: 24 {Teinodactyla) in synonymy with Mniophila muscorum muscorum (Koch,
1803)
p. 538, add agrostidis Gistel, 1857a: 5 {Aphthona) [NO] in synonymy with Neocrepidodera cyanescens concolor
(K. Daniel, 1900)
p. 538, add fugax Gistel, 1857a: 88 {Haltica) in synonymy with Neocrepidodera cyanescens cyanescens
(Duftschmid, 1825)
p. 540, add to Luperomorpha valid species:
costipennis S.-Y. Wang, 2002b: 684, 693 A: FUJ
p. 540, add to Nisotra valid species and subspecies:
 ERRATA 53
apicefulva Bryant, 1941b: 410 A: NP ORR
cardoni naini Scherer, 1969: 146 A: UP
p. 540, add SD to distribution of Nisotra gemella (Erichson, 1834)
p. 540, remove TAI from distribution of Nonarthra coreana Chûjô, 1935
p. 541, remove TAI from distribution of Nonarthra tibialis (Jacoby, 1885)
p. 541, add CZ to distribution of Ochrosis ventralis (Illiger, 180^
p. 542, remove Ophrida scaphoides (Baly, 1865) and its synonym binduta Maulik, 1926; Ophrida spectabilis
(Baly, 1862) and its synonym rufoflava (Fairmaire, 1889); and Ophrida xanthospilota (Baly, 1881). They
were transferred to the genus Asiophrida Medvedev, 2000
p. 542, add styriae Gistel, 1857a: 18 in synonymy with Orestia alpina (Germar, 1824)
p. 545, add SV to distribution of Phyllotreta astrachanica Lopatin, 1977
p. 545, add IN to distribution of Phyllotreta beschkentica Lopatin, 1961
p. 546, remove Phyllotreta brevistriata Kimoto, 1966 and place it in synonymy with Phyllotreta chujoe Madar,
1959
p. 546, add A: TR to distribution of Phyllotreta bulgarica Gruev, 1977
p. 546, remove SVfromdistribution of Phyllotreta diademata Foudras, 1860
p. 547, add A: TR to distribution of Phyllotreta fornuseki Cizek, 2003
p. 547, add A: TR to distribution of Phyllotreta ganglbaueri Heikertinger, 1909
p. 547, add A: TR to distribution of Phyllotreta lorestanica Warchalowski, 1973
p. 547, add IN to distribution of Phyllotreta hermonensis Fürth, 1979
p. 548, add IN to distribution of Phyllotreta ozbeki Gruev & Asian, 1998
p. 548, add IN to distribution of Phyllotreta parfentjevi Shapiro, 1958
p. 549, add to Phyllotreta valid species and its synonym:
subtilior CsM, 1940: 66 [RN] A: HP
subtilis Weise, 1922:122 [HN]
p. 549, add AFRi to distribution of Podagricafuscicornis (Linnaeus, 1767)
p. 551, change spelling of Podontia quatordecimpunctata Linnaeus, 1767 and its synonym quatordecimpunctata
L. Redtenbacher, 1844 to quatuordecimpunctata
p. 551, change the year of description of Polyclada benti Gahan, 1896a to 1895
p. 552, move Psylliodes {Psylliodes) aemulans Har. Lindberg, 1953 and Psylliodes (Psylliodes) amplicollis
Wollaston, 1865 to subgenus Eupus Wollaston, 1854
p. 552, add to Eupus Wollaston, 1854 valid species:
stolida Wollaston, 1860a: 11 N: CI MR
p. 552, add araratica lablokoff-Khnzorian, 1968: 265 in synonymy with Psylliodes aerea Foudras, 1860 and add
A: AR to distribution of the latter species
p. 552, add to Psylliodes s. str. valid species:
agropyri Paliy, 1961: 390 E: CT A: KZ
p. 552, add AFRi to distribution of Psylliodes affinis (Paykull, 1799)
p. 552,removeentry: araratica lablokoff-Khnzorian, 1968: 265 A: AR
p. 553, add caerulea Kral, 1947:21 in synonymy with Psylliodes attenuata Koch, 1803
p. 553, delete entry balyi Jacoby, 1884a: 30 A: CH JA (Ryukyus) TAI ORR, and place the name in synonymy
with Psylliodes chlorophana Erichson, 1842
p. 553, add brachybiotes Gistel, 1857a: 27 and eubiotes Gistel, 1857a: 13 in synonymy with Psylliodes
chrysocephala chiysocephala (Linnaeus, 1758)
p. 553, correct year for catinensis Rottenberg [in synonymy with Psylliodes chrysocephala chrysocephala
(Linnaeus, 1758)] to 1872a
p. 554, modify Psylliodes cucullata cucullata (Illiger, 1807) to Psylliodes cucullata (Illiger, 1807)
p. 554, add augustalisi Pic, 1909: 146 [DA] in synonymy with Psylliodes cucullata (Illiger, 1807)
p. 554, remove entry Psylliodes cucullata heydeni Weise, 1888c: 785 E: FR (Pyrénées) PT SP
p. 554, add A: TR to distribution of Psylliodes drusei Fürth, 1983
p. 554, add AU HU to distribution of Psylliodes frivaldszkyi Weise, 1888
p. 554, modify Psylliodes gibbosa gibbosa Allard, 1860 to Psylliodes gibbosa Allard, 1860
p. 554, add valligenus Gistel, 1857a: 87 (Longitarsus) in synonymy with Psylliodes glabra (Duftschmid, 1825),
and add CZ HU RO SK to its distribution
54 ERRATA
p. 555, add to Psylliodes s. str. valid species:
heydeni Weise, 1888c: 785 E: FR (Pyrénées) PT SP
p. 555, add AU CZ to distribution of Psylliodes illyrica Leonardi & Gruev, 1993
p. 555, correct year for ventricosa Rottenberg [in synonymy with Psylliodes inflata Reiche & Saulcy, 1858] to
1872]
p. 555, add JO LE SY to distribution of Psylliodes instabilis Foudras, 1860
p. 555, add FR (Corse) to distribution of Psylliodes kiesenwetteri Kutschera, 1864
p. 555, add SA to distribution of Psylliodes maculatipes Pic, 1924
p. 556, remove A: AE SA from distribution of Psylliodes moricandiae Peyerimhoff, 1925
p. 556, add IS to distribution of Psylliodespeyerimhoffi Heikertinger, 1916
p. 557, remove Psylliodes stolida Wollaston, 1860 (transferred to subgenus Eupus, p. 552)
p. 557, add AU to distribution of Psylliodes sturanyi Apfelbeck, 1906
p. 557, add A: TR to distribution of Psylliodes toelgi Heikertinger, 1914
p. 557, add nigricapilla Krai, 1945: 31 in synonymy with Psylliodes tricolor Weise, 1888c
p. 557, remove normandi Heikertinger, 1916: 34 from the synonymy with Psylliodes vehemens and put it as
valid subspecies of vehemens. Distribution of vehemens normandi: E: AZ SP (Malaga) N: AG CI MO MR
TU and of vehemens vehemens: E: PT (Madeira)
p. 557, remove Psylliodes agropyri Paliy, 1961: 390 E: CT A: KZ from Psylliodes sbg. Semicnema and place it
to Psylliodes (s. str.) (p. 552)
p. 558, add Lochmaeina L. N. Medvedev, 1956a: 157 in synonymy with Sangriola Jakobson, 1922, and rosea L.
N. Medvedev, 1956a: 157 (Lochmaeina) in synonymy with punctatostriata Motschulsky, 1861
p. 559, add to Sphaeroderma valid species:
gressitti Scherer, 1969: 205 A: SD
himalayense Scherer, 1969: 208 A: SD ORR
nigriceps Scherer, 1969: 210 A: SD
p. 560, correct spelling of Stenoluperus moori (Baly, 1874) to moorii, and add "Korea" to its distribution
p. 561, remove entry: genus Torodera Weise, 1902 with its synonym Argopistoides Jacoby, 1892 and species
septempunctata Jacoby, 1892b: 932 and sexpunctata Scherer, 1987: 69 [genus Argopistoides has priority
over Torodera thus all taxa are moved to page 503]
p. 562, add to Trachytetra valid species:
nigrocyanea Scherer, 1969: 83 (Zipangia) A: SD
p. 565, add humeralis Pic, 1928b: 25 in synonymy with Aspidolopha buqueti egregia (Boheman, 1859)
p. 565, add instigmata Pic, 1933a: 3 in synonymy with Aspidolopha melanophtalma (Lacordaire, 1848)
p. 565, add infasciata Pic, 1927a: 26, medioobliterata Pic, 1927c: 7 and obliterata Pic, 1927a: 26 in synonymy
with Aspidolopha thoracica Jacoby, 1892
p. 565, add imbasalis Pic, 1943b: 16, obliterata Pic, 1943d: 8, paulofasciata Pic, 1943d: 8 and unireducta Pic,
1943d: 8 in synonymy with Ceratobasis koenigii (Fabricius, 1775)
p. 565, add original combination (Chilotoma) to Cheilotoma erythrostoma italica (Leoni, 1906)
p. 565, add baudii Leoni, 1906: 147 (Chilotoma) in synonymy with Cheilotoma erythrostoma italica (Leoni,
1906)
p. 565, add leosinii Leoni, 1906: 146 (Chilotoma) [DA] and orientalis Jakobson, 1901b: 107 (Chilotoma) in
synonymy with Cheilotoma musciformis musciformis Goeze, 1777
p. 566, add fiiscitarsis Lacordaire, 1848: 298 (Clythra) [see New Acts] and virens Lacordaire, 1848: 298
(Clythra) [see New Acts] in synonymy with Chilotomina nigritarsis (Lacordaire, 1848)
p. 566, remove Clytrella L. N. Medvedev, 1961 from synonymy with Clytra Laicharting, 1871; it is correctly
listed as synonym of Clytraria Semenov, 1903
p. 566, add original combination (Clythra) to Clytra aliéna Weise, 1897, and to the following invalid names:
bella Baly, 1865, bicoloriceps Pic, 1933, connexa Fricken, 1888, appendicina Lacordaire, 1848, and
succincta Lacordaire, 1848
p. 566, add antistita Weise, 1889m: 563 [see New Acts] and ehnbergi Jakobson, 1901b- 110 in synonymy with
Clytra (Clytra) arida Weise, 1889
p. 566, add inapicalis Pic, 1927a: 31 in synonymy with Clytra (Clytra) duodecimmaculata duodecimmaculata
(Fabricius, 1775)
 ERRATA 55
p. 566, add dissimilis Weise, 1889m: 563 in synonymy with Clytra (Clytra) quadripunctata quadripunctata
(Linnaeus, 1758)
p. 566, add inhumeralis Pic, 1943d: 14 in synonymy with Clytra (Clytra) subfasciata Lacordaire, 1848
p. 567, add angulata Weise, 1900c: 273, conjuncta Heyden, 1891b: 368, croceicollis Weise, 1900c: 273,
lacordairei Weise, 1900c: 273, marginicollis Weise, 1900c: 273, milliati Pic, 1942d: 7 and sierrana J.
Daniel, 1903b: 253 in synonymy with Clytra (Clytraria) atraphaxidis atraphaxidis (Pallas, 1773)
p. 567, add original combination (Clythra) to Clytra (Clytraria) atraphaxidis maculifrons Zubkov, 1833
p. 567, add ashabadensis Brancsik, 1899: 103, flexuosa Weise, 1900c: 274, nigromaculata Pic, 1897e: 84,
quinquemaculata Pic, 1920c: 7 [HN] and vulgaris Weise, 1900c: 274 in synonymy with Clytra (Clytraria)
atraphaxidis maculifrons Zubkov, 1833
p. 567, add notâtithorax Pic, 1915a: 5 in synonymy with Clytra (Clytraria) jacobsoni Semenov, 1903
p. 567, add original combination (Clythra) to the invalid names aleppensis L. Redtenbacher, 1843, elegans
Faldermann, 1837, diss im ilithorax Desbrochers des Loges, 1870, and unifasciata Ménétriés, 1836
p. 567, add juncta Pic, 1920c: 7 and signicollis Weise, 1900c: 272 in synonymy with Clytra (Clytraria)
novempunctata Olivier, 1808
p. 567, add defecta Jakobson, 1901b: 110 [RN] [see New Acts] and deflciens Weise, 1900c: 274 [HN] [see New
Acts] in synonymy with Clytra (Clytraria) ruflna (Solsky, 1881)
p. 567, add drurei Pic, 1920c: 7 and subjuncta Pic, 1920c: 7 in synonymy with Clytra (Clytraria) Valeriana
Valeriana Ménétriés, 1832
p. 567, add original combination (Clythra) to Clytra (Ovoclytra) nigrocincta nigrocincta Lacordaire, 1848 and
ovataovata Lacordaire, 1848
p. 567, add cyprica Pic, 1918d: 11 [DA] [see New Acts], deficiens Heyden, 1891b: 368 [DA] [see New Acts],
graeca Pic, 1915a: 5 [DA], multipunctata Pic, 1920c: 7, semireducta Pic, 1918d: 15 and subinterrupta Pic,
1920c: 7 in synonymy with Clytra (Ovoclytra) nigrocincta nigrocincta Lacordaire, 1848
p. 567, add octomaculatum Pic, 1932c: 35 in synonymy with Clytrasoma conforme (Lacordaire, 1848)
p. 567, add trifasciata L. Redtenbacher, 1844: pi. 28 (Clythra) in synonymy with Clytrasoma dispar (L.
Redtenbacher, 1844)
p. 567, add reductum Pic, 1932c: 35, rufithorax Pic, 1932c: 35 and subobliteratum Pic, 1932c: 35 in synonymy
with Clytrasoma palliatum (Fabricius, 1801)
p. 568, add biinterrupta Pic, 1918d: 23, biscrensis Pic, 1918d: 23, bistrijuncta Pic, 1905j: 156 and trimaculata
Pic, 1918d: 23 in synonymy with Coptocephala aeneopicta (Fairmaire, 1864)
p. 568, add TU to North African distribution of Coptocephala plagiocephala (Fabricius, 1792)
p. 568, add kocheri Pic, 1953d: 62 [DA] [see New Acts] and leprieuri Pic, 1897j: 198 in synonymy with
Coptocephala crassipes crassipes Lefèvre, 1876
p. 568, add Coptocephala quadrimaculata Fabricius, 1792: 57 (Cryptocephalus) [HN] in synonymy with C.
gebleri Gebier, 1841
p. 568, add latenotata Pic, 1949a: 4 [see New Acts] in synonymy with Coptocephala destinoi Fairmaire, 1884
p. 568, add impressiceps Pic, 1918d: 18 [DA] [see New Acts] and tambei Pic, 1942d: 7 [see New Acts] in
synonymy with Coptocephala fallaciosa Fairmaire, 1884
p. 568, add vitalei Pic, 1913c: 114 in synonymy with Coptocephala fossulata Lefèvre, 1872
p. 568, add amasiensis Weise, 1889e: 128 and inundata Reitter, 1905c: 251 in synonymy with Coptocephala
gebleri Gebier, 1841
p. 568, change spelling of Coptocephala kabakowi to kabakovi
p. 568, add rubriceps Pic, 1916a: 10 in synonymy with Coptocephala kerimii Fairmaire, 1875
p. 568, add theryi Pic, 1936e: 26 [HN] in synonymy with Coptocephala metalliconotata Pic, 1933
p. 568, add andalusiaca Pic, 1918d: 17, espanoli Pic, 1933d: 15, externepunctata Pic, 1895f: 89, theryi Pic,
1918d: 17 and tunisea Pic, 1901q: 80 in synonymy with Coptocephala plagiocephala (Fabricius, 1792)
p. 568, add fabianii Burlini, 1946: 78 [see New Acts] in synonymy with Coptocephala raffrayi (Desbrochers des
Loges, 1870)
p. 569, add IT SZ to distribution of Coptocephala rubicunda rubicunda (Laicharting, 1781)
p. 569, remove cyanocephala Olivier, 1791: 35 (Clythra) [DA] from synonym of Coptocephala scopolina
scopolina (Linnaeus, 1767); it is correctly listed as nomen dubium
p. 569, add inhumeralis Fuente, 1910b: 445 and subfasciata Pic, 18971: 206 in synonymy with Coptocephala
scopolina floralis (Olivier, 1791)
56 ERRATA
p. 569, add bipimctata Dépoli, 1926b: 92 [DA] [see New Acts], destituta Dépoli, 1926b: 92 [DA] [see New
Acts], lateremaculata Dépoli, 1926b: 92 [DA] [see New Acts], scutellaris Dépoli, 1926b: 92 [DA] [see New
Acts] and separata Dépoli, 1926b: 92 [DA] [see New Acts] in synonymy with Coptocephala scopolina
Scopolina (Linnaeus, 1767)
p. 569, add aenescens Pic, 1932d: 23 in synonymy witii Coptocephala unicolor(P. H. Lucas, 1845)
p. 569, add to Coptocephala valid subspecies:
unifasciata babylonica Lopatin, 1995b: 100 A: IQ
p. 569, add/a//ax Weise, 1887a: 56, maculiceps Kraatz, 1872b: 223, phenax Jakobson, 1896c: 106 [DA] [see
New Acts], reducta G. Müller, 1948: 80, rufweps Weise, 1890f: 363 [DA] [see New Acts] and triornata
Roubal, 1948: 33 in synonymy with Coptocephala unifasciata unifasciata Scopoli, 1763
p. 569, add quinquemaculata Lacordaire, 1848: 255 (Clythra) [see New Acts] in synonymy with Diapromorpha
dejeani dejeani (Lacordaire, 1848)
p. 569, add vitalisi Pic, 1928b: 24 (Ceratobasis) and postbinotata Pic, 1928b: 24 (Ceratobasis) in synonymy
with Diapromorpha pinguis (Lacordaire, 1848)
p. 570, in Labidostomis s. str. remove entry centrisculpta afghanica Lopatin & Nesterova, 2007 [a primary junior
homonym of L. nuristanica afghanica L. N. Medvedev, 1978] and replace it by:
centrisculpta kqfiristanica Regalin & L. N. Medvedev, nom. nov. A: AF
afghanica Lopatin & Nesterova, 2007b: 430
p. 5 70, add damryi Weise, 1881: 95 in synonymy with Labidostomis (Chlorostola) centromaculata Gene, 1839
p. 570, add violaceipennis Pic, 1932d: 23 in synonymy with Labidostomis (Chlorostola) guerini lejeunei
Fairmaire, 1866
p. 570, add reymondi Kocher, 1959: 7 in synonymy with Labidostomis (Chlorostola) rufomarginata Vaulonger
de Beaupré, 1895
p. 570, add obliterata Pic, 1920c: 6» lineata Pic, 1920c: 6 and suturella Pic, 1920c: 6 in synonymy with
Labidostomis (Chlorostola) syriaca (Lacordaire, 1848)
p. 570, add fallacissima Jakobson, 1901b: 106 in synonymy with Labidostomis (Labidostomis) amurensis
amurensis Heyden, 1884
p. 570, add arisi Pic, 1920c: 7 in synonymy with Labidostomis (Labidostomis) arcuata Pic, 1920
p. 570, add taurica Lacordaire, 1848: 63 (Clythra) in synonymy with Labidostomis (Labidostomis) brevipennis
Faldermann, 1837
p. 570, add fulvipennis Lacordaire, 1848: 54 (Clythra) in synonymy with Labidostomis (Labidostomis)
cyanicornis (Germar, 1822)
p. 570, under Labidostomis (Labidostomis) diversifrons Lefèvre, 1872, change 1897k to 18971 in attenuata Pic,
1897
p. 571, add bijuncta Pic, 1912a: 17, bisbijuncta Pic, 1912a: 17 and posticejuncta Pic, 1912a: 17 in synonymy
witii Labidostomis (Labidostomis) hebraea (Lacordaire, 1848)
p. 571, add atlasica Kocher, 1959: 7 in synonymy with Labidostomis (Labidostomis) hybrida P. H. Lucas, 1845
p. 571, add inhumeralis Pic, 1920c: 6 in synonymy with Labidostomis (Labidostomis) luristanica Pic, 1920
p. 571, add pyrenaea Kraatz, 1872b: 197 and semideficiens Pic, 1906a: 19 in synonymy with Labidostomis
lusitanica (Germar, 1824)
p. 571, add subinterrupta Pic, 1932d: 22 in synonymy with Labidostomis (Labidostomis) mairei Peyerimhoff,
1922
p. 571, add nitidicollis Lefèvre, 1872: 85 in synonymy with Labidostomis (Labidostomis) metallica metallica
Lefèvre, 1872
p. 572, add elongata Lacordaire, 1848: 51 (Clythra) in synonymy with Labidostomis (Labidostomis)
pallidipennis (Gebier, 1830)
p. 572, change Labidostomis propinquapropinqua Faldermann, 1837 to L. propinqua Faldermann, 1837
p. 572, change 1901a to 1901b in Labidostomis (Labidostomis) transitoria Jakobson, 1901
p. 573, correct spelling of Labidostomis (Labidostomis) uygur to ujgur
p. 573, correct spelling of Welleschmiedia to Wellschmiedia
p. 573, add nigronotata Lacordaire, 1848: 167 (Clythra) [see New Acts] in synonymy with Lachnaia
(Barathraea) cerealis (Olivier, 1808)
p. 573, remove (Lachnaia) from Lachnaia (Barathraea) padillai Tomov, 1982
 ERRATA 57
p. 573, add original combination (Lachnaea) to pardoi Cobos, 1956, L. (Barathraea) separata Pic, 1897, to L.
(Lachnaia) italica italica Weise, 1881, L. (Lachnaia) italica occidentals Grasso, 1963, L. (Lachnaia)
lucidipennis Peyerimhoff, 1926, L. (Lachnaia) orientalis Weise, 1881, L.(Lachnaia) peyerimhoffl Kocher,
1953, L. (Lachnaia) pseudobarathraea K. Daniel & J. Daniel, 1898, insidiosa Cobos, 1954, and curtipennis
Pic, 1936
p. 573, add bistigmata Pic, 1912J: 90 (Lachnaea), jurjurensis Pic, 1913c: 129 (Lachnaea) and kabyliana Pic,
1912j : 90 (Lachnaea) in synonymy with Lachnaia (Lachnaia) paradoxa (Olivier, 1808)
p. 573, add original combination (Clytra) to Lachnaia (Lachnaia) pubescens Dufour, 1820
p. 573, add subfasciata Pic, 1897J: 197 (Lachnaea) in synonymy with Lachnaia (Lachnaia) pubescens Dufour,
1820
p. 573, add uniustigmata Pic, 1898J: 74 (Lachnaea) in synonymy with Lachnaia (Lachnaia) puncticollis
Chevrolat, 1840
p. 573, add pontica Weise, 1884b: 157 (Lachnaea) in synonymy with Lachnaia (Lachnaia) sexpunctata
(Scopoli, 1763)
p. 573, add triguttata Lacordaire, 1848: 184 (Clythra) in synonymy with Lachnaia (Lachnaia) tristigma
(Lacordaire, 1848)
p. 574, add binotaticollis Pic, 1916a: 6, confluens Weise, 1893b: 1118, gallica Schulz, 1909: 595, immaculata
Schulz, 1909: 594, latemaculata Pic, 1897h: 165, quadrimaculata Schulz, 1909: 595 and trimaculata Schulz,
1909: 595 in synonymy with Macrolenes dentipes (Olivier, 1808)
p. 574, add se/ta Lacordaire, 1848: 378 (Clythra) in synonymy with Melitonoma sobrinum (Lacordaire, 1848)
p. 574, add bistrimaculata Pic, 1927a: 29 (Cyaniris), disjuncta Pic, 1927a: 29 (Cyaniris), subeincta Pic, 1927a:
29 (Cyaniris) and trinotaticollis Pic, 1927a: 29 (Cyaniris) in synonymy with Miochira montana thibetana
Pic, 1927
p. 574, add carnerii Pic, 1920e: 21 (Cyaniris) and latecyanescens Pic, 1946a: 3 in synonymy wiüi Otiocephala
opaca (Rosenhauer, 1856)
p. 575, change (= Clythra gambiensis Lacordaire, 1848) to (= Clythra signata Lacordaire, 1848) as the type
species of Paraclytra L. N. Medvedev, 1971 [replacing the homonym Clythra notata Klug, 1835]
p. 575, remove IS OM AFR from distribution of Paraclytra crocata (Lacordaire, 1848)
p. 575, change Paraclytra signata sennariensis (Lacordaire, 1848) to Paraclytra sennariensis (Lacordaire, 1848)
and add SY to its distribution
p. 575, add binotata Pic, 1927g: 7', fokienensis Pic, 1927g: 7, octonotata Chûjô, 1952b: 89 (Coptocephald) and
uninotata Pic, 1927g: 7 in synonymy with Physauchenia pollens (Lacordaire, 1848)
p. 575, add atrobasalis Pic, 1932b: 13 (Cyaniris), basidisjuncta Pic, 1934d: 20 (Cyaniris), formosana Chûjô,
1934e: 284 (Cyaniris), immaculata Heyden, 1887e: 295 (Coptocephald), kiangsuna Pic, 1938c: 17
(Cyaniris) and latereducta Pic, 1934d: 20 (Cyaniris) in synonymy with Physosmaragdina nigrifrons (Hope,
1843)
p. 575, for Medvedevella Özdikmen, 2008, replace [RN] by [unnecessary RN]
p. 575, for Necyomantes Gistel, 1848, replace [unnecessary substitute name] by [RN]
p. 576, add dissimilis Lefèvre, 1872: 339 (Gynandrophlhalma) in synonymy wiüi Smaragdina aurita aurita
(Linnaeus, 1767)
p. 576, add banatica Weise, 1881: 123 (Gynandrophthalma) in synonymy with Smaragdina chloris chloris
(Lacordaire, 1848)
p. 576, add picticollis Weise, 1889e: 128 (Gynandrophthalma) in synonymy with Smaragdina flavicollis
(Charpentier, 1825)
p. 577, add ilicis Lacordaire, 1848: 289 (Clythra) [see New Acts] in synonymy with Smaragdina hypoerita
(Lacordaire, 1848)
p. 577, add to Smaragdina valid species:
minuta Jacoby, 1908: 115 (Gynandrophthalma) A: SD ORR
p. 577, add kuatunensis Pic, 1954b: 56 (Cyaniris) in synonymy with Smaragdina laevicollis (Jacoby, 1890)
p. 578, add melania Chûjô, 1952b: 82 (Gynandrophthalma) in synonymy with Smaragdina octomaculata Chûjô,
1952
p. 578, add lucidipes Pic, 18971: 206 (Chilotoma) in synonymy with Smaragdina reyi (C.F.N. Brisout de
Barneville, 1866)
p. 578, add to Smaragdina valid species:
58 ERRATA
scheren Lopatin, 2006c: 593 A: SCH
p. 583, change spelling of Cryptocephalus mniczechi to mniszechi
p. 578, add latemaculata Pic, 18971: 205 (Gynandrophthalmd) in synonymy with Smaragdina scutellaris
(Lefèvre, 1872)
p. 578, add bijuncta Pic, 1906e: 42 (Coptocephala), latejuncta Pic, 1914f: 18 (Cyaniris) and subjuncta Pic,
1914i: 139 (Cyaniris) in synonymy with Smaragdina thoracica thoracica (Fischer von Waldheim, 1842)
p. 578, add kraatzi Clavareau, 1906: 44 (Gynandrophthalmd) [HN] and liturata Weise, 1881: 128
{Gynandrophthalmd) [see New Acts] in synonymy with Smaragdina tibialis (Brullé, 1832)
p. 578, add aegyptiaca Lefèvre, 1872: 337 (Gynandrophthalmd) and venusta Lefèvre, 1872: 337
(Gynandrophthalma) in synonymy with Smaragdina unipunctata (Olivier, 1808)
p. 579, add thoracica Küster, 1846b: nr. 99 (Cyaniris) [HN] in synonymy with Smaragdina xanthaspis Germar,
1824: 547
p. 579, add donceeli Pic, 1929b: 14 (Antipa), lacordairei Pic, 1929b: 14 (Antipd) and palaestina Pic, 1929b: 14
(Antipa) in synonymy with Tituboea arabica (Olivier, 1808)
p. 579, add flavipennis Pic, 1950a: 16 (Antipa), legionis Kocher, 1959: 12 and vagenotata Pic, 1939c: 18 in
synonymy with Tituboea biguttata (Olivier, 1791)
p. 579, add semijuncta Pic, 1918d: 18 in synonymy with Tituboea chobauti (Pic, 1896)
p. 579, add binotaticollis Pic, 1914f: 1, infasciata Kocher, 1952: 117 (Antipa), obliterata Pic, 1897h: 164 and
semiconjuncta Pic, 1914f: 1 in synonymy with Tituboea fasciata Lefèvre, 1872
p. 579, change spelling of Tituboea fasciatipes to fasciaticeps
p. 579, add leprieuri Pic, 1895f: 89, nigricollis Heyden, 1886g: 49 and semiundulata Pic, 1939c: 18 in synonymy
with Tituboea illigeri (Lacordaire, 1848)
p. 579, add subjuncta Pic, 1903g: 114 in synonymy witii Tituboea laticollis (Olivier, 1808)
p. 579, add armeniaca Pic, 1918d: 18 in synonymy with Tituboea macropus (Illiger, 1800)
p. 579, add atriceps Pic, 1924c: 22 and decemmaculata Pic, 1937c: 13 (Antipa) in synonymy with Tituboea
minor Fairmaire, 1894
p. 580, add infasciata Pic, 1912a: 73 in synonymy with Tituboea nigriventris Lefèvre, 1872
p. 580, add mecheriensis Pic, 1895a: 13, kocheri Pic, 1949a: 16, siciliensis Pic, 1912a: 74 [DA] [see New Acts]
and unipunctata Pic, 1897h: 165 in synonymy with Tituboea octopunctata (Fabricius, 1787)
p. 580, add octopunctata Lacordaire, 1848: 162 [HN] (Clythra) [see New Acts] in synonymy with Tituboea
paykulli (Lacordaire, 1848)
p. 580, add akbesiana Pic, 1897e: 82 and humerlifera Pic, 1900u: 91 in synonymy with Tituboea sexmaculata
(Fabricius, 1781)
p. 580, add bisbinotata Pic, 1912j: 90, notaticeps Pic, 1912a: 73 and robustior Pic, 1912a: 73 in synonymy with
Tituboea subabbreviata Pic, 1912
p. 583, correct spelling of Cryptocephalus mniczechi Tappes, 1869 to mniszechi
p. 583, under Cryptocephalus sarafschanensis duvivieri Weise, 1892, add [RN] to solskyanus Jakobson, 1901b:
112
p. 590, correct spelling of Cryptocephalus bhutanicus Lopatin, 1975 to bhutanensis
p. 591, correct year for Cryptocephalus chikatunovi Lopatin, 2007b to 2008d
p. 594, add semipurpureus Pic, 1914f: 12 in synonymy with Cryptocephalus globicollis Suffrian, 1847
p. 595, under Cryptocephalus hypochoeridis Linnaeus, 1758: 370, add [RN] to frigidus Jakobson, 1901b: 113
p. 595, delete Cryptocephalus infraniger Pic, 1915a: 14 A: TR (Tokat) - t h e description of C. infraniger refer to
the genus Cteniopus (Tenebrionidae: Alleculinae) [see Errata for vol. 5]
p. 595, add berladensis Pic, 1915a: 13 in synonymy with Cryptocephalus laetus Fabricius, 1792
p. 597, add to Cryptocephalus valid species:
nigroflavus Lopatin, 2004c: 614 A: SCH
p. 597, delete [RN] in Cryptocephalus octomaculatus Rossi, 1790
p. 598, under Cryptocephalus octopunctatus octopunctatus, change spelling of colmariensis to colmarensis
p. 601, add purpureicolor Pic, 1914f: 11 and semiviridescens Pic, 1914f: 12 in synonymy with Cryptocephalus
sericeus (Linnaeus, 1758)
p. 602, delete Cryptocephalus testaceitarsis Pic, 1915a: 14 A: TR (Tokat) - the description of C. testaceitarsis
refer to the genus Cteniopus (Tenebrionidae: Alleculinae) [see Errata for vol. 5]
 ERRATA 59
p. 603, under Cryptocephalus tristigma tristigma Charpentier, 1825, change 1953o;4« 1953d in marocanus Pic,
1953
p. 604, add to Cryptocephalus valid species:
zinicus Lopatin, 2004c: 614 A: SCH
p. 604, add atroscutellaris Pic, 1912a: 74 in synonymy with Cryptocephalus {Heterichnus) carinthiacus
Suffrian, 1848
p. 605, correct year for princeps Rottenberg [in synonymy with Cryptocephalus stragula Rossi, 1794] to 1872b
p. 606, add to Melixanthus valid species:
atricillus Suffrian, 1854a: 80 A: HP ORR
kuluensis Lopatin, 1979b: 436
p. 607, add to Melixanthus valid species:
puncticollis Lopatin, 2005d: 873 A: SCH
p. 608, add kabakovi L. N. Medvedev & Samoderzhenkov, 1987: 23 in synonymy with Adiscus laetus Weise,
1904
p. 608, add to Adiscus valid species:
weisei L. N. Medvedev, 2008: 203 A: SD ORR
p. 608, add to Coenobius valid species:
azureus Schöller, 1999: 74 [RN] A: SD
cyaneus Lopatin, 1979b: 437 [HN]
burmanicus Jacoby, 1892b: 889 A: SD ORR
p. 609, change spelling of Coenobius pakistanus to pakistanicus
p. 609, add Thelyterotarsus {Thelylankus) Lopatin, 1987b: 778 type species Thelyterotarsus {Thelylankus)
insularis Lopatin, 1987b in synonymy with Acolastus
p. 610, add to Acolastus {Acolastus) valid species:
mitjaevi Lopatin, 2008c: 37 A: UZ
p. 615, under Pachybrachis pradensis Marseul, 1875, in incallidus change 1897a: 2 to 1897J: 196
p. 616, under Pachybrachis tesselatus tesselatus (Olivier, 1791) change the year and page of description of
melanopygus G. Müller from 1952: 352 to 1948: 84
p. 617, addferrantei Pic, 1912: 75 in synonymy with Stylosomus niloticus Suffrian, 1857
p. 625, add [HN] to entry Macrocoma impressa Berti & Rapilly, 1973
p. 626, correct year for Macrocoma vanharteni Lopatin, 2007b to 2008d
p. 626 add under Macrocoma valid species:
substriata Weise, 1904f: 100 {Pseudocolaspis) A: IN TR
p. 627, correct entry under Mecistes Chapuis, 1874 to:
seriatus Lefêvre, 1885a: 122 A: SA AFR
saudica L. N. Medvedev, 1997f: 321
p. 627, correct spelling of {Cyptocephalus) to {Cryptocephalus), for Pachnephorus pilosus (Rossi, 1790)
p. 629, add to Trichochrysea valid subspecies and its synonym:
nitidissima scutellaris Pic, 1926b: 11 A: NP ORR
donckieri Pic, 1926b: 11
p. 634, add bonneuli Pic, 1934b: 6 {Corynodes) in synonymy with Platycorynus peregrinus (Herbst, 1783)
p. 637, under Basilepta balyi (Harold, 1877) correct authorship of otsukae to Matsumura in Otsuka, 1911
p. 638, add to Basilepta valid species:
pretiosa Jacoby, 1908: 304 {Nodostoma) A: XIZ ORR
p. 639, add mouhoti Baly, 1864a: 11 and laosensis Pic, 1928b: 28 in synonymy with Callisina quadripustulata
Baly, 1864
p. 640, correct year for Chloropterus lefevrei arabicus Lopatin, 2007b to 2008d
p. 640, under Chloropterus lefevrei lefevrei Reitter, 1890, add in synonymy: bipunctatus Reitter, 1897c: 225
and bucharicus Reitter, 1897c: 226
p. 640, correct distribution of Chloropterus pallidus Chobaut, 1898 to: N: AG
p. 641, add to Microeurydemus valid species:
wraniki Lopatin, 1994: 526 A: YE
p. 643, use Floricola Gistel, 1848 as valid, and Eupales Lefèvre, 1885 as its invalid synonym
60 ERRATA
p. 647 add: An S. L., Kwon Y. J. & Lee. S-M. 1985: Classification of the leafbeetles from Korea Part I.
Subfamily Hispinae (Coleoptera: Chrysomelidae). Insecta Koreana 5: 1-9.
p. 662, add: Bogatchev A. V. 1961: Novye vidy zhukov-chernotelok (Tenebrionidae) iz Tadzhikskoy SSR,
drugikh respublik Sredney Azii i sopredel'nykh stran. Trudy Institute Zoologii i Parazitologii Akademii
Nauk Tadzhikskoy SSR 20: 107-121.
p. 684, add: Chiang (=Jiang) S.-N. 2001: [New name]. In: Jiang S.-N. & Chen Li: Fauna Sinica, Insecta. Vol.
21, Coleoptera, Cerambycidae Lepturinae. Beijing: Science Press, 296 pp.
p. 698, add: Delobel A. 2007: [New taxon]. In: Kergoat G. J., Delobel P. & Delobel A.: Phylogenetic
relationships of a new species of seed-beetle infesting Cercis siliquastrum L. in China and in Europe
(Coleoptera: Chrysomelidae: Bruchinae: Bruchini). Annales de la Société Entomologique France (N. S.) 43:
265-271.
p. 702, add: Donisthorpe H. 1905: Gramoptera holomelina, Pool, a good species. Entomologist's Record and
Journal of Variation 17: 182-183.
p. 716, correct year for Gahan, 1896a to 1895
p. 716, correct pagination of Gahan C. J. 1896b to: 448-461
p. 726, correct title and editor for Grimmer, 1841 to: Steiermark's Coleoptern mit Einhundert sechs neu
beschriebenen Species. Grätz: C. Tanzer
p. 730, remove entry: Hammarström R. 1893:...
p. 733, add: Hayashi M. 1982: The Cerambycidae of Japan (Col.) (13). The Entomological Review of Japan 37:
141-152.
p. 734, correct year for Hayashi M. & Villiers A. 1985b to 1987
p. 738, add: Heyden L. F. D. J. von 1886g: Zusammenstellung der von Herrn Dr. Med. W. Kobelt von seiner
Reise in den Provinzen Alger und Constantine, sowie von Tunis mitgebrachten Coleopteren. Bericht über die
Senckenbergische naturforschende Gesellschaft in Frankfurt am Main 1886: 35-57.
p. 738, add: Heyden L. F. D. J. von 1887e: Zweiter Beitrag zur Coleopteren-Fauna von Pecking in Nord-China.
Deutsche Entomologische Zeitschrift 31: 293-295.
p. 756, add: Kelly P. G. & Reid C. A. M. 1999: Designation of type species for me genera of Australian
paropsine beeties (Coleoptera, Chrysomelinae). Memoirs of Museum Victoria 57: 263-266.
p. 759, add: Kocher L. 1952: Antipa Paykulli Lac et fasciata Lef (Col. Chrysom.). Comptes Rendus des Séances
Mensuelles de la Société de Sciences Naturelles et Physique du Maroc 18: 117-118.
p. 763, correct year for Kraatz G. 1871e to 1872c
p. 763, add: Kraatz G. 1876: [note, p. 318]. In: Heyden L. F. J. D. von.: Die Cortodera- (Muls.) und
Grammoptera- (Serv.) Arten. Deutsche Entomologische Zeitschrift 20: 317-320.
p. 764, add: Krâl J. 1945: Novi nebo mâlo znâmi dfepôici ôeské fauny. De novis vel minus cognitis halticinarum
speciebus formisque faunae Bohemiae. (Coleoptera, Chrysomelidae). Casopis Ceskoslovenské Spolecnosti
Entomologické 42: 31-33.
p. 764, add: Krâl J. 1947: Pfispëvek k poznâni dïepôikù fauny Ceskoslovenska. (3.). Ad halticidarum faunae
Cechoslovakiae cognitionem additamentum III. (Coleoptera, Phytophaga). Casopis Ceskoslovenské
Spolecnosti Entomologické 44: 21 -26.
p. 765: correct pagination for Krynicki J. 1832: Enumeratio ... to [3] + 68-179
p. 765: correct title for Krynicki J. 1834 ... to Addenda et nonnula synonyma Rossiae meridionalis
Coleopterorum
p. 771, correct volume in Lameere A. 1912a: ... Mémoires de la Société Entomologique de Belgique to 21
p. 772, add: Lazarev M. A. 2008: Zametki po spornym voprosam sistematiki i rasprostranenia zhukov-usachey
(Coleoptera, Cerambycidae) Rossii i sopredelnykh stran. Pp. 129-136. In: Aktualnye problemy prioritetnyh
napravleniy razvitiya estestvennyh nauk. Sbornik statey. Moskva: Izdatelstvo «Prometey» MPGU, 220 pp.
p. 772: complete title of Lazarev M. A. 2009 to: Armenian Dorcadion (Coleoptera: Cerambycidae) of
"cinerarium-group"
p. 780, correct reference for Lopatin I. K. 1979c: Novyy rod... to Entomologicheskoe Obozrenie 58:...
p. 781, add: Lopatin I. K. 1987b: Novye podrod i vid roda Thelyterotarsus (Coleoptera, Chrysomelidae) iz Shri-
Lanki [New subgenus and species of the genus Thelyterotarsus (Coleoptera, Chrysomelidae) from Sri
Lanka]. Entomologicheskoe Obozrenie 66: 778-780.
p. 783, correct year for Lopatin I. K. 2007b to 2008d
 ERRATA 61
p. 784, add: Lopatin I. K. 2008c: Acolastus mitjaevi sp. n. - novyy vid zhukov-listoedov iz Sredney Azii i ego
polozhenie v sisteme roda (Coleoptera, Chrysomelidae, Cryptocephalinae). [Acolastus mitjaevi sp. n. - new
species of chrysomelids from Middle Asia and its status in genus system (Coleoptera, Chrysomelidae,
Cryptocephalinae)]. Tethys Entomological Research 16: 37-38.
p. 787, add: Mallet P.-M. 1933: Le mélanisme chez Chrysomela grammis L. [sic!] (Col. Chrysomeidae). Bulletin
de l'Association des Naturalistes de la Vallée du Loing 16: 74-75.
p. 793 add: Medvedev L. N. 1992f: Reviziya zhukov-listoyedov podsemeystva Hispinae (Coleoptera,
Chrysomelidae) fauny Vietnama. Chast I. Pp. 127-156. In: Medvedev L. N. (ed.): Sistematyka i ekologia
nasekomykh Vietnama. Moskva, 264 pp.
p. 795, add: Medvedev L. N. & Samoderzhenkov E. V. 1987: Fauna Cryptocephalinae (Coleoptera,
Chrysomelidae) Vietnama [Cryptocephalinae of Vietnam]. Pp. 69-80. In: Medvedev L. N. (ed.):
Entomofauna Vietnama [Entomofauna of Vietnam]. Nauka, Moskva, 192 pp. (in Russian),
p. 801, correct title in Müller J. 1907 to Cerambycidae Dalmatiae.
p. 802, correct pagination in Mulsant E. 1847d to 515-521
p. 808, in Normand (1949), change in title (fascicule 3) to (fascicule 4) and pagination from 65-104 to 79-104
p. 812, replace references to Olivier A. G. 1792a and 1792b by:
Olivier A. G. 1797: Encyclopédie méthodique,... Tome septième. Pars II. Paris: Panckoucke, pp. 369-827. [pars
I, pp. 1-368 of me vol. 7 was published in 1793]
p. 819, correct pagination for Pic M. 1890e: to clxxv-clxxvi
p. 823, correct title to Pic M. 1898v: to Diagnose d'une ...
p. 828 add: Pic M. 1912: Coléoptères égyptiens nouveaux ou plus ou moins rares. Bulletin de la Société Royale
Entomologique d'Egypte 4 [ 1911 ] : 71 -76.
p. 831, correct pagination for Pic M. 1924c to 17-18,...
p. 837, add: Plavilstshikov N. N. 1932e: Zhuki-drovoseki vrediteli drevesiny. Moskva-Leningrad:
Gosudarstevennoe Lesnoe Tekhnicheskoe IzdatePstvo, 200 pp., and remove reference to Plavilstshikov N.
N.1933a
p. 837, add: Plavilstshikov N. N. 1932f: Lepturinen-Studien (Col., Cerambycidae). I. Casopis Ceskoslovenské
Spolecnosti Entomologické 29: 87-88, 174-175.
p. 848, add: Roemer J. J. 1789: Genera insectorum Linnaei et Fabricii iconibus illustrata a Joanne Jacobo
Roemer. Vitoduri Helvetorum: apud Henrjc. Steiner et socios, viii + 86 + [3] pp., 37 pis.
p. 848, add: Rost C. 1892b: Brachyta bifasciata Ol. var. caucasica Rost. Entomologische Nachrichten 18 (6): 81
p. 848, correct year for Rottenberg A. L. A. M. 1871a and 1871b to 1872a and 1872b, respectively
p. 849, add: Roubal J. 1948: Systematickâ, zoogeografickâ, bibliografickâ studie o éervenych druzich rodu
Coptocephala Dejean (Chrysomelidae). [De chrysomelidarum génère Coptocephala Dejean adnotationes
systematicae, zoogeographicae, necnon bibliographicae]. Entomologické Listy 11: 33-34.
p. 854, add: Santos-Silva A. & Hovore F. T. 2007: Divisào do gênero Distenia Lepeletier & Audinet-Serville,
notas sobre a venaçao alar em Disteniini, homonimias, sinonimia e redescriçôes (Coleoptera, Cerambycidae,
Disteniinae). Papéis Avulsos de Zoologia 47: 1-29.
p. 855, add: Schaeffer C. 1907: New Bruchidae wim notes on known species and list of species known to occur
at Brownsville, Texas, and in the Huachuca mountains, Arizona. Bulletin of the Brooklyn Institute of Arts
and Sciences. Museum 1: 291-306.
p. 857, add: Schultze W. 1920: Eights contribution to the Coleoptera fauna of the Philippines. Philippine Journal
of Science 16: 191-203, 2pls.
p. 881, add: Wang S.-Y. 2002b: [new taxon]. In: Wang S.-Y. & Yu P.-Y.: Chrysomelidae: Alticinae. Pp. 633-
693. In: Huang P.-K. (ed.): Fauna of Insects of Fujian Province of China. Vol. 6. Fujian Science &
Technology Press, 894 pp.
p. 883, correct pagination for Waterhouse C. O. 1889: Coleoptera to: Pp. 122-131/
p. 892, add: Yen Chia-Hsien 1932: A new species of Cerambycidae from Kwangsi. Peking Natural History
Bulletin 7(2): 165-166.
62 ERRATA
Appendix 1
M. L. Danilevsky
Supporting references for the preservation of Vadonia unipunctata unipuncata (Fabricius, 1787) over Vadonia
pilosa (Förster, 1771) under ICZN, Art. 23.9. The name/»/tea Forster has not been used as valid after 1899
to my knowledge.

Bartenev A. F. 2004: Obzor vidov zhukov-usachey (Coleoptera: Cerambycidae) fauny Ukrainy. [A review of the
long-horned beetles species (Coleoptera: Cerambycidae) of the fauna of Ukraine]. Izvestiya Kharkovskogo
Entomologicheskogo Obshchestva [The Kharkov Entomological Society Gazette] 2003(2004) 11 (1-2): 24-
43.
Bartenev A. F. 2009: Zhuki-usachi Levoberezhnoy Ukrainy i Kryma. Kharkov: Kharkovskiy Natzionalnyy
Universitet. [Longicorn-beetles of Left-Bank Ukraine and Crimea. Kharkov: Kharkov National University],
405 pp.
Danilevsky M. L. 2009: Species group taxa of longhorned beetles (Coleoptera, Cerambycidae) described by N.
N. Plavilstshikov and their types preserved in the Zoological Museum of the Moscow State University and in
the Zoological Institute of the Russian Academy of Sciences, St. Petersburg. Entomological Review 89 (6):
689-720.
Danilevsky M. L. & Miroshnikov A. X 1985: Zhuki-drovoseki Kavkaza {Coleoptera, Cerambycidae).
Opredelitel. Krasnodar: Kubanskiy Selskokhozyaistvennyy Institut, 417 pp., 10 pis.
Esjunin S. L. & Kozminyh V. O. 1992: Materialy k faune zhestkokrylykh zakaznika "Troitzkiy". In:
Chlenistonogie okhranyaemykh territoriy Cheliabinskoy oblasti. Sverdlovsk. UrO AN SSSR. [Materials on
the Coleoptera fauna of "Troitzky" reservation.- Arthropoda of preserved territories of Cheljabinsk region.
Sverdlovsk. Ural Branch of Ac. of Sc of the USSR]: 59-67.
Isaev A. Yu. & Magdeev D. V. 2003: Fauna zhukov-usachey (Coleoptera, Cerambycidae) lesostepnoy zony
Povolzhya. In: Issledovaniya v oblasti biologii i metodiki ee prepodavaniya. Mezhvuzovskiy sbornik
nauchnykh trudov k 100-Ietiyu D. N. Florova, vyp. 3, ch. 1. [Fauna of longicorn-beetles (Coleoptera,
Cerambycidae) of forest-steppe zone of Volga valley.- Investigations in the field of biology and methods of
its teaching. Inter-institutes collection of transactions to the 100 anniversary of D. N. Flerov, 3(1).] Samara:
290-300.
Isaev A. Yu., Egorov L. V. & Egorov K. A. 2004: Zhestkokryfye (Insecta, Coleoptera) lesostepi Srednego
Povolzhya. Katalog. [Beetles (Insecta, Coleoptera) of forest-steppe of Middle Volga. Catalogue.] Ulianovsk,
72pp.
Isaev A. Yu. 2007: Opredelitel zhestkokrylykh Srednego Povolzhya (chast III. Polyphaga, Phytophaga). Seriya
"Priroda Ulyanovskoy oblasti" [A key for beetles of Middle Volga (part III. Polyphaga, Phytophaga). Serie
"Nature of Ulianovsk Region"], N 14. Ulianovsk. Publishing house "Vektor - C", 1-225pp.
Kaliuzhnaja N. S., Komarov E. V. & Cherezova L. B. 2000: Zhestkokryfye nasekomye Nizhnego Povolzhya.
[Coleoptera of lower Volga river.] Volgograd, 204 pp.
Kostin I. A. 1968: Rasprostranenie drovosekov (Coleoptera, Cerambycidae) listvennykh porod v Kazakhstane.
In: Nasekomye - vrediteli selskogo i lesnogo khozyaistva Kazakhstana. Trudy Instituta Zoologii, Akademiya
nauk Kaz. SSR [Distribution of Cerambycidae (Coleoptera) in the leaf-bearing woods in Kazakhstan.- In:
Pest Isects of Agriculture and Forests of Kazakhstan. Proceedings of the Institute of Zoology, Academy of
Sciences of Kazakhstan SSR] 30: 184-197.
Kostin I. A. 1973: Zhuki-dendrofagi Kazakhstana {koreedy, drovoseki, zlatki). Alma-Ata: Akademiya Nauk
Kazakhskoy SSR, 288 pp.
Kryzhanovsky O. L. 1974: 35. Sem. Cerambycidae - drovoseki ili usachi. In: Nasekomye i kleshchi - vrediteli
selskokhozyaistvennykh kultur. [35. Fam. Cerambycidae - timber beetlest or longicorn beetles. In: Insects
and ticks - pests of agricultural crops.] Leningrad: Nauka, 139-157.
Lobanov A. L., Danilevsky M. L. & Murzin S. V. 1981: Sistematicheskiy spisok usachei (Coleoptera,
Cerambycidae) fauny SSSR. I. Entomologicheskoe Obozrenie 60: 784-803.
Magdeev D. V. 1996: Zhuki-usachi (Cerambycidae, Coleoptera). In: Flora i fauna zapovednikov. Vyp. 61.
Bespozvonochnye Zhigulevskogo zapovednika. Moscow: 39-47.
Magdeev D. V. 2003: Fauna zhukov-usachey (Coleoptera, Cerambycidae) Samarskoy oblasti. In:
Kraevedchiskie zapiski, XI. Samarskiy oblastnoy istoriko-kraevedchiskiy muzey. [Fauna of Longicorn-
 ERRATA 63
Beetles (Coleoptera, Cerambycidae) of Samara Region.-;/Local history notes, XI. Samara Regional Local
History Museum.] "Fine Design", Samara: 202-208.
Martynov V. V. & Pisarenko T. A. 2004: Ecologo-faunisticheskiy obzor zhukov-usachey (Coleoptera:
Cerambycidae) yugo-vostochnoy Ukrainy. Izvestiya Kharkovskogo Entomologicheskogo Obshchestva [A
review of the fauna and ecology of the long-homed beetles (Coleoptera: Cerambycidae) of southeast
Ukraine.- The Kharkov Entomological Society Gazette] [2003] 11: 44-69.
Mikhailov Yu. E. 1999: Nasekomye muzeya-zapovednika "Arkaim". Zhestkokrylye: vidovoy sostav i zametki
po structure populyatziy. In: Prirodnye sistemy Yuzhnogo Urala. Cheliabinsk. Cheliabinskiy gos. universitet.
[Insects of "Arkaim" museum-natural reserve. Beetles: species composition and notes on the structure of
populations.- Natural systems of South Urals. Cheljabinsk: Cheljabinsl University: 221-248.
Miroshnikov A. I. 1998: Novaya klassifikacia zhukov-drovosekov kompieksa Anoplodera triby Lepturini
(Coleoptera, Cerambycidae) fauny golarktiki. I. Entomologicheskoe Obozrenie 11: 384-421.
Naumov R. V. 1994: Zhuki-usachi (Coleoptera, Cerambycidae) Ulyanovskoy oblasti. Ser. "Priroda Ulyanovskoy
oblasti". [Longicorn-beetles (Coleoptera, Cerambycidae) of Ulianovsk region. Ser. "The Nature of Ulianovsk
Region"], 5. Ulianovsk: 43-46.
Negrobov S. O., Tzurikov M. N., Logvinovsky V. D., Fomichev A. I., Prokin A. A. & Gilmutdinov K. S. 2005:
Otryad Coleoptera. In: Kadastr bespozvonochnykh Voronezhskoy oblasti. [Order Coleoptera.- Cadastre of
Invertebrata of Voronezh region.] Voronezh: 534-673.
Pisarenko T. A. 1999: Osoblivosti fauni rodini vysachiv (Coleoptera, Cerambycidae) promislovikh i zapovidnikh
rayoniv pivdenno-skhidnoy Ukraini. Naukoviy Visnik. Suchasna ekologia i problemi stalogo rozvitku
suspibtva. Vipusk 9.7, Lvov: 149-155.
Plavilstshikov N. N. 1965: 75. Sem. Cerambycidae - zhuki-drovoseki, usachi [75-th Fam. Cerambycidae -
Timber Beetles, Longicornes]. Pp. 389-419. In: Opredelitel nasekomykh Evropeyskoy chasti SSSR, t. 2,
zhestkokrylye i veerokrylye. Moscow-Leningrad: Nauka.
Romadina K. G. 1954: Drevogryzushchie lichinki Zhukov-usachey (Cerambycidae) doliny reki Urala
[Woodboring Longicorn (Cerambycidae) beetle-larvae of Ural river valley]. Trudy Zoologicheskogo Institu ta
akademii nauk SSSR 16:211 -228.
Shapovalov A. M., Nemkov V. A., Rusakov A. V. & Shovkun D. F. 2006: Zhuki-usachi (Coleoptera,
Cerambycidae) Orenburgskoy oblasti [Longicorn-beetles (Coleoptera, Cerambycidae) of Orenburg Region].
Vestnik Orenburgskogo Gosudarstvennogo Universiteta 4 (54): 105-109.
Zagaikevich I. K. 1979: Nekotorye aspekty khorologii usachey (Coleoptera, Cerambycidae) fauny Ukrainy. In:
VU Mezhdunarodniy simpozium po entomofaune Sredney Evropy. Materialy. Leningrad, 19-24 sentiabria
1977: 195-197.
64
NEW NOMENCLATURAL and TAXONOMIC ACTS, and COMMENTS
Comments to classification
M. A. Alonso-Zarazaga
The systematic arrangement here followed departs from that found in Lawrence & Newton (1995), expressly
declared as being that given in Kuschel (1995). The merits and demerits of this and other classification
systems for Curculionoidea have been the subject of different criticisms (e.g., Alonso-Zarazaga & Lyal,
1999; Oberprieler et al., 2007; among others). While Alonso-Zarazaga & Lyal (I.e.) inclined to the use of an
eclectic classification since their main target was a nomenclatural catalogue, Oberprieler et al. (I.e.) made
an approach that incorporated recent acquisitions in the knowledge of the phylogenetic relationships of the
weevil taxa. However, neither of these two, nor any other classification, has won a general acceptance. The
reasons are obvious. In general, the consensus is that the main taxa of primitive weevils are more or less
well delimited, while their relationships are not yet too clear; in the case of the advanced weevils, neither
the relationships, nor the group limits are clear yet. The delimitation of the family Curculionidae as
including (or not) Dryophthoridae (as Dryophtiiorinae) and Platypodidae (as Platypodinae) is just a matter
of personal liking.
The resulting classification used for Curculionoidea in vols. 7 and 8 of this catalogue is eclectic again: I have
accepted the placement of Oxycorynidae as a subfamily Oxycoryninae of Belidae, and probably the best
solution for Attelabidae and Rhynchitidae would have been to merge both in a single family, since the latter
seems to be paraphyletic with respect to the former. The big family Brentidae is not supported by my
unpublished molecular analyses, which makes of this "brentoid complex" a grade rather than a clade, so I
have opted for keeping the arrangement in Alonso-Zarazaga & Lyal (1999), since the three families
involved (Brentidae s. str., Apionidae, Nanophyidae) are easily differentiated on morphological grounds
alone. Similarly, I have kept separated the Brachyceridae and the Dryophthoridae, and the Curculionidae s.
str. start with the Platypodinae, related by some analyses to the Dryophthoridae. Some other analyses plead
that there could be a large family Brachyceridae, uniting all the monocot feeders at the base of
Curculionidae, including here also Dryophthoridae. We have nothing certain here as well, and I prefer to
keep the familiar groups already used in Alonso-Zarazaga & Lyal (1999), a catalogue that has served its
purpose and many others, even if the monophyly of several subfamilies is difficult to invoke (e.g.,
Molytinae, Cryptorhynchinae, etc.).

Curculionoidea
M. A. Alonso-Zarazaga
General comments
The synonymies of the species published by Gravenhorst (1807) have followed, when available, his own
manuscript notes in his own copy of this book, conserved in the library of the University of Wroclaw
(Poland).
As a general rule, colour varieties whose distribution was not separated from that of the main species in me
original description (e.g., by giving a procedence or locality) have been deemed to be infrasubspecific
(Arts. 45.6.1 and 45.6.4) on the ground that "the content of the work unambiguously reveals mat die name
was proposed for an infrasubspecific entity" due to the absence of data which may allow considering them
"geographical subspecies".

Anthribidae
Milos Tryzna & Barry D. Valentine
Comments
The classifications of Anthribidae are imperfectly supported by data on morphology. Consequently, the present
arrangement is tentative, based also on unpublished information. Changes in family- and genus-group taxa
are given as new.
 NEW ACTS and COMMENTS 65
New assignments
Allcmdrus LeConte, 1876 is transferred from Stenocerini to Allandrini
Autotroph Jordan, 1924 is transferred from Zygaenodini to Discotenini
Apatenia Pascoe, 1859 is transferred from Corrhecerini to Platyrhinini
Aphaulimia Morimoto, 1972 is transferred from Corrhecerini to Platyrhinini
Asemorhinus Sharp, 1891 is transferred from Sintorini to Tophoderini
Baseocolpus Jordan, 1949 is transferred from Corrhecerini to Platyrhinini
Bothrus Wolfrum, 1956 is transferred from Tropiderini to Phloeophilini
Botriessa Jordan, 1928 is transferred from Zygaenodini to Piesocorynini
Cleorisintor Jordan, 1923 is transferred from Corrhecerini to Mauiini
Dei "isemias Jordan, 1928 is transferred from Mauiini to Tophoderini
Disphaerona Jordan, 1902 is transferred from Corrhecerini to Ecelonerini
Dolichocera Gray, 1832 is transferred from Mecocerini to Tophoderini
Enedreytes Schoenherr, 1839 is transferred from Stenocerini to Discotenini
Eusphyrus LeConte, 1876 is transferred from Ormiscini to Zygaenodini
Exillis Pascoe, 1860 is transferred from Platystomini to Zygaenodini
Gibber Jordan, 1895 is transferred from Corrhecerini to Zygaenodini
Hypseus Pascoe, 1860 is transferred from Corrhecerini to Platyrhinini
Ulis Jordan, 1936 is transferred from Mauriini to Zygaenodini
Mecotropis Lacordaire, 1865 is transferred from Mecocerini to Ptychoderini
Nessiara Pascoe, 1860 is transferred from Corrhecerini to Platyrhinini
Oxyderes Jordan, 1928 is transferred from Corrhecerini to Platyrhinini
Penestica Pascoe, 1859 is transferred from Platystomini to Corrhecerini
Phaeochrotes Pascoe, 1860 is transferred from Stenocerini to Allandrini
Phaulimia Pascoe, 1859 is transferred from Corrhecerini to Platyrhinini
Plintheria Pascoe, 1859 is transferred from Stenocerini to Allandrini
Pseudeuparius Jordan, 1914 is transferred from Cratoparini to Discotenini
Phloeophilus Schoenherr, 1833 is transferred from Mecocerini to Phloeophilini
Phaenotheriolum Ganglbauer, 1903 is transferred from Stenocerini to Piesocorynini
Phaenotherion Frivaldszky, 1877 is transferred from Stenocerini to Piesocorynini
Phaenotheriosoma Frieser, 1978 is transferred from Stenocerini to Piesocorynini
Rawasia Roelofs, 1880 is transferred from Ecelonerini to Tophoderini
Streneoderma Blackburn, 1900 is transferred from Anthribini to Zygaenodini
Ulorhinus Sharp, 1891 is transferred from Corrhecerini to Platyrhinini

Resurrected taxa
Allandrini Pierce, 1930 is resurrected from synonymy with Stenocerini Kolbe, 1895
Eusphyrus LeConte, 1876 is resurrected from synonymy with Ormiscus G. R. Waterhouse, 1845
Hypseus Pascoe, 1860 is resurrected from synonymy with Phaulimia Pascoe, 1859

Changes in rank
Apolectini Lacordaire, 1865 are raised to subfamily rank, Apolectinae

New synonyms
Dasycorynus Lacordaire, 1865, syn. nov. ofXylinada Berthold, 1827
Japanthribus Shibata, 1978, syn. nov. of Jordanthribus Zimmerman, 1938
Vaienfriesiini Aionso-Zarazaga & Lyal, 1999, syn. nov. of Araecerini Lacordaire, 1865

New combination
Pseudeuparius sepicola (Fabricius, 1792), comb. nov.fromEnedreytes

New taxon
Stenorhinini Valentine, tribe nov. Type genus: Stenorhis Jordan, 1928
Rostrum absent; mouthparts projecting, but very reduced and articulating on venter of head capsule; eyes
66 NEW ACTS and COMMENTS
rounded, dorsolateral, finely faceted; transverse pronotal carina basal; elytra with ten striae, the usual short
scutellar stria absent, and the ninth stria fused with tenth above the junction of metasternum and abdomen.
This tribe includes only the type genus. Distribution: Japan to India, east to Philippines and Samoa.

Type species designation

Pachygenia Motschulsky, 1874, currently in synonymy with Ceaus Pascoe, 1860, was established for two new
species, P. impluviata Motschulsky, 1874 and P. guttula Motschulsky, 1874. A type species was not
designated. We designate here Pachygenia impluviata Motschulsky, 1874 as the type species of Pachygenia
Motschulsky.

Misidentification
Phaeochrotes pudens (Gyllenhal, 1833) is frequently misidentified as "cinctus Paykull".

Anthribidae: Urodontinae
M. A. Alonso-Zarazaga
Designation of type species
Exurodon Schilsky, 1912 originally included Urodon fulvipes Reitter, 1897 and U. arisi sp. n. Urodon arisi
Schilsky, 1912 is hereby designated as type species of Exurodon.

New combination
Bruchela rosinae (Pic, 1904), comb. nov. from Urodon

Species incertae sedis (following name was omitted in currently used references)
Bruchus undatus Fabricius, 1787a: 41 from N: "Africa" was placed in Macrocephalus by Manuel (1797: 606),
now in synonymy of Platystomos

Nomina dubia (following names were omitted in currently used references)

Anthribus discifer Schrank, 1798: 474 E: GE
Anthribus ephippiger Gistel, 1831: 305 E: GE
Anthribus lividus Olivier, 1789a: 161 E: FR
Anthribus pallinosus Anonymus, 1826: 338 E: NL
Anthribus striatus Olivier, 1789a: 161 "Europe"
Anthribus unicolor Olivier, 1789a: 160 E: FR
Anthribus vittatus Geoffroy, 1785: 138 E: FR

Rhynchitidae
M. A. Alonso-Zarazaga
Comments
The systematics and nomenclature of this family have recently been entangled by numerous publications by A.
A. Legalov. Severe criticisms against Legalov's proposals have been raised by different authors: Wanat &
Mokrzycki (2005), Riedel (2006, 2009), Setliff (2007), Oberprieler et al. (2007), Xie & Liang (2008).
Legalov has reached in some groups the heigth of describing in a piecework fashion almost one genus or
subgenus per species, based solely on poor characters like colour or brightness, obscuring the phylogenetic
relationships. My own intents to reproduce his phylogenetic results with his own matrices had no success,
and widely different phylogenies appear when using either cladistic analysis programs or a bayesian
approach with them, which are, at the same time, regarded as swollen with characters useless for a sound
phylogenetic analysis. Moreover, this author is prone to linguistic, geographic and nomenclatural mistakes,
and his keys and descriptions in many cases do not allow a clear separation of taxa. Consequently,
Legalov's (2003c, 2007a, etc.) works should be considered with the greatest caution. Inconsistent wording
(and in some cases uninteligible English translations) make it very difficult to understand them. The graphic
material supporting his descriptions and diagnoses is in many cases minute (almost specimen size), blurred,
fuzzy or carelessly repeated from other species. His nomenclatural acts are also very dubious and lack
 NEW ACTS and COMMENTS 67
understanding of the Code (cf. for example, Probyctiscidius below), using undescribed genera in 2002c
(Parasynatops or Thompsonirhinus) for describing species, when these genera were not made available
until one year later (2003c). These works are overrun by persistent misspellings of taxa (e.g., Apoderus
gemnosus [sic!] vs. correct Apoderus gemmosus Jekel, or Attelabus curculionides vs. correct
curculionoides, or Deporaus septentrionalis vs. original - although incorrectly latinized - D.
septemtrionalis, or Mechoris and Mecoris instead of the correct Mecorhis, just to mention a few) and
authors' names (Kâno instead of Kôno), by incorrect authorships and dates (e.g., Eomesauletes politus
attributed wrongly to "Boheman, 1828" [in fact 1829] when it was described by Lepeletier & Serville,
1825), by confused etymologies (e.g., Japanodeporaus "is formed from the words «Japanese» and
«eugnamptus»"), by recognition of taxa that are unavailable according to the Code (e.g., Clitostylini Voss,
1926), by taxa lacking descriptions, by using species-groups whose diagnoses are only known to the author,
by ambiguous diagnoses for taxa, by combinations given as new when they have been already proposed or
in combinations proposed with a genus not being treated (e.g., Maculinvolvulus setosellus under
Squamrhynchites; Legalov, 2007a: 147), by synonymies given as new when they have already been
proposed by others and overlooked (e.g., Arodepus Voss placed in synonymy of Arodepus Heller as new
(2007a: 120) when this synonymy was already given by Alonso-Zarazaga & Lyal (1999: 43), by
erroneously recorded original genera, by apparently new distributional records not referred to specimens in
collections or incomplete distributional data, or absent taxa, or in assigning incorrectly genders (e.g.,
Mecorhis is feminine, not masculine) or overcorrecting nouns in apposition as if they were adjectives
(many names ending in —cola, like Rhynchites penangicola, converted into Opacoinvolvulus penangicolus,
ck Involvulus gemma, converted into an incorrect /. gemmus, or Apoderus sissu converted into an incorrect
sissus), etc.
Legalov modifies geographic data: Byctiscus complanatus Voss, 1933 was described from a locality
Shanhaikwan which is in the Chinese province Hebei. The place, now called Shanhaiguan is some 20 km
NE of Qinhuangdao. Although this is recognized in Legalov (2003b), he suddenly changes it in his
treatment of Aspidobyctiscus (Taiwanobyctiscus) complanatus (Legalov, 2007a: 207) to "Shang-hai-Kwan"
and places it in Shanghai province, where no locality with this or a similar name occurs. Similar changes
happen everywhere in his work, so that his geographical data are subject to caution and not all have been
incorporated in this Catalogue because of these inherent problems, or because he mentions a single studied
specimen for a species, while its distribution covers several geographical units. Consequently, some of his
general geographic data have not been used here, lacking a published support by specimens.
I have not followed the disputable substitution of extant tribal names with tiiose of tribes known only from
fossils which are extremely difficult to identify, the relationships between them and identities proposed by
Legalov being still more debatable. Consequently, I have returned to the use of the common name
Deporaini Voss, 1929, leaving Isotheini Scudder, 1893 for the fossil taxa only, whose placement in my
opinion is very doubtful. In my opinion (and in that of other authors, cf. Riedel (2006), Setliff (2007) all the
suitable ranks have been upgraded in many cases by one or two levels, without justification. A good point
for Legalov's work is that it has called attention towards a neglected group that needs a deeper work devoid
of superficial descriptions and ambiguous key couplets. I am not confident that the high number of new
monospecific genera and subgenera, or monogeneric subtribes or tribes, will serve to clarify the
relationships of the species involved; as in many other cases, they seem to have been named because of the
narrow taxonomic concepts of the author at all levels and the so called "mihi" itch. Several groups of
species have been separated based on a very limited analysis of a few weak (gradational) characters, and
given generic status, when they most likely qualify only as subgenera or species groups. Valuable
characters, such as shape of penis or armature of endophallus, are used only secondarily, and mentioned,
but not thoroughly described, while elytral colour or length of vestiture are used for the creation of genera.
Consequently, in the treatment here adopted, I have been compelled to downgrade several taxa and to
synonymize others. The subtribes of Rhynchitini, excepting perhaps Anisomerinina, characterized by a
peculiar shortening of the first ventrite, are of little value and are based on weak characters. I have not used
them here.
Another conundrum: In a key, Voss (1938c) described two Rhynchites comosellus (p. 159 (no. 164) from
"Indien" and p. 164 (no. 202) from "Hinterindien"). In 1969, Voss treated the first as a valid species in
Involvulus, as /. comosellus (p. 245) from "Indien: Sikkim" and the second as a homonym species (p. 290),
renamed /. commutatus, from "Birma: Ruby Mts.". In Legalov's (2003b) treatment, both species appear
68 NEW ACTS and COMMENTS
under Thompsonirhinus sensu stricto, but the first comosellus has dissapeared, confused with the second,
which is in synonymy of T. consimilis Voss, 1938, a species described from Kumaon (India, UP), while the
Birmanian species is erroneously spelled T. "commulatatus" and its replaced name, the second comosellus,
is absent (since it is under T. consimilis). Moreover, the first comosellus, which should be under consimilis,
is treated as an incertae sedis species under Parinvolvulus in Legalov's (2003b) treatment and transferred to
a new genus Orrhynchites in 2007a. This nomenclatural problem is corrected here, since Legalov (2007a)
has not done it. The correct name for the Birmanian species should be Mecorhis (Thompsonirhinus)
commutata (Voss, 1969b: 290), comb, nov., according to the system here proposed. On the other hand, I
give here as new the synonymy Rhynchites consimilis Voss, 1938 = Rhynchites comosellus Voss, 1938 (p.
159), syn. nov., since that given in Legalov points to a different homonym R. comosellus, and combine it
with Orrhynchites, as O. consimilis (Voss, 1938), comb. nov.
Voss's revisions use subspecies and "forms" simultaneously, so the latter are deemed to be infrasubspecific and
are not given in the catalogue (ICZN, Art. 45.6.4), unless they have been raised to specific or subspecific
rank by a subsequent author before 1985 (Art. 45.6.4.1).
Deporaus uniformis Kôno, 1930a: 14, currently listed in synonymy of Deporaus (Roelofsideporaus) unicolor
(Roelofs, 1874) by variuos catalogues, is a misspelling of the former. Consequently, it has no status in
nomenclature and is not recorded in the text.

Changes in rank
Arodepus Heller, 1922, stat dem. to subgenus of Caenorhinus C. G. Thomson, 1859
Biblarodepus Voss, 1924, stat. dem. to subgenus of Capylarodepus Voss, 1922
Bicolorhynchus Legalov, 2003, stat. dem. to subgenus of Ecnomonychus Voss, 1933
Capylarodepopsis Legalov, 2003, stat. dem. to subgenus of Capylarodepus Voss, 1922
Cateugnamptus Legalov, 2007, stat. dem. to subgenus of Eugnamptobius Voss, 1922
Chinadeporaus Legalov, 2007, stat. dem. to subgenus of Deporaus Samouelle, 1819
Chryseugnamptus Legalov, 2003, stat. dem. to subgenus of Eugnamptobius Voss, 1922
Coccygorhynchites Prell, 1926, stat. dem. to subgenus of Lasiorhynchites Jekel, 1860
Convexodeporaus Legalov, 2007, stat. dem. to subgenus of Caenorhinus C. G. Thomson, 1859
Cyaneugnamptus Legalov, 2003, stat. dem. to subgenus of Eugnamptobius Voss, 1922
Epirhynchites Voss, 1953, stat. dem. to subgenus of Rhynchites D. H. Schneider, 1791
Eugnamptellus Legalov, 2003, stat. dem. to subgenus of Eugnamptobius Voss, 1922
Flavorhynchites Legalov, 2003, stat. dem. to subgenus of Ecnomonychus Voss, 1933
Haplorhynchites Voss, 1924, stat dem. to subgenus of Involvulus Schrank, 1798
Heterorhynchites Voss, 1932, stat. dem. to subgenus of Mecorhis Billberg, 1820
Himalayrhynchites Legalov, 2003, stat. dem. to subgenus of Ecnomonychus Voss, 1933
Indinvolvulus Legalov, 2003, stat. dem. to subgenus of Ecnomonychus Voss, 1933
Macu Iinvolvulus Legalov, 2003, stat. dem. to subgenus of Ecnomonychus Voss, 1933
Maculorhinus Legalov, 2003, stat. dem. to subgenus of Ecnomonychus Voss, 1933
Megalarodepus Voss, 1922, stat. dem. to subgenus of Capylarodepus Voss, 1922
Metinvolvulus Legalov, 2003, stat. dem. to subgenus of Cneminvolvulus Voss, 1960
Mirabrhynchites Legalov, 2003, stat. dem. to subgenus of Mecorhis Billberg, 1820
Nelasiorhynchites Legalov, 2003, stat. dem. to subgenus of Lasiorhynchites Jekel, 1860
Neoarodepus Legalov, 2003, stat. dem. to subgenus of Caenorhinus C. G. Thomson, 1859
Neocoenorhinidius Legalov, 2003, stat dem. to subgenus of Neocoenorrhinus Voss, 1952
Nepalorhynchites Legalov, 2007, stat. dem. to subgenus of Involvulus Schrank, 1798
Nigroarodepus Legalov, 2003, stat. dem. to subgenus of Caenorhinus C. G. Thomson, 1859
Opacoinvolvulus Legalov, 2003, stat. dem. to subgenus of Cneminvolvulus Voss, 1960
Oreugnamptus Voss, 1941, stat. dem. to subgenus of Eugnamptobius Voss, 1922
Paleodeporaus Legalov, 2003, stat. dem. to subgenus of Deporaus Samouelle, 1819
Paratemnocerus Legalov, 2003, stat. dem. to subgenus of Temnocerus Thunberg, 1815
Parinvolvulus Legalov, 2003, stat. dem. to subgenus of Involvulus Schrank, 1798
Parvodeporaus Legalov, 2007, stat. dem. to subgenus of Deporaus Samouelle, 1819
Philorectus Voss, 1924, stat dem. to subgenus of Mecorhis Billberg, 1820
Pilosoinvolvulus Legalov, 2003, stat dem. to subgenus of Cneminvolvulus Voss, 1960
 NEW ACTS and COMMENTS 69
Platyrhynchites Voss, 1922, stat. dem. to subgenus of Deporaus Samouelle, 1819
Platyrhynchitoides Legalov, 2007, stat. dem. to subgenus of Caenorhinus C. G. Thomson, 1859
Proteugnamptellus Legalov, 2003, stat. dem. to subgenus of Neoeugnamptus Le galov, 2003
Pseudarodepus Legalov, 2003, stat dem. to subgenus of Caenorhinus C. G. Thomson, 1859
Pseudodepasophilus Voss, 1942, stat dem. to subgenus of Caenorhinus C. G. Thomson, 1859
Pseudomechoris Legalov, 2003, stat. dem. to subgenus of Mecorhis Billberg, 1820
Pseudodeporaus Voss, 1922, stat. dem. to subgenus of Capylarodepus Voss, 1922
Pseudvulns Legalov, 2003, stat dem. to subgenus of Cneminvolvulus Voss, 1960
Rubrinvolvulus Legalov, 2003, stat dem. to subgenus of Ecnomonychus Voss, 1933
Schoenitemnus Legalov, 2003, stat dem. to subgenus of Neocoenorrhinus Voss, 1952
Stenorhynchites Voss, 1932, stat dem. to subgenus of Lasiorhynchites Jekel, 1860
Teretriorhynchites Voss, 1938, stat dem. to subgenus of Involvulus Schrank, 1798
Thompsonirhinus Legalov, 2003, stat dem. to subgenus of Mecorhis Billberg, 1820
Vietnamodeporaus Legalov, 2007, stat. dem. to subgenus of Caenorhinus C. G. Thomson, 1859

Infrasubspecific names
The following names (and many others) were clearly proposed as infrasubspecific names under Art. 45.6.2 or
45.6.4 of the Code, and have been omitted from the list. In these cases, Art. 45.6.4.1 cannot be applied.
Auletes wagenblasti f. aenea Voss, 1939 [Art. 45.6.4, raised to species as Auletes aeneus Legalov, 2002].
Byctiscus impressus f. angusticollis Voss, 1930 [Art. 45.6.4, raised to species as Byctiscus angusticollis
Legalov, 2007].
Byctiscus impressus f. thibetana Voss, 1933 [Art. 45.6.4, raised to species as Byctiscus thibetanus Legalov,
2007].
Art. 45.6.4.1 is to be applied here:
Involvulus (Involvulus) pilosotessellatus [sic!] f. continentalis Voss, 1958, an infrasubspecific name, raised to
subspecies by Voss, 1969b: 264, is deemed to be available from its original description. Now in synonymy
of Mecorhis {Thompsonirhinus) mandschurica (Voss, 1939).

New combinations
Caenorhinus (Arodepus) marginellus Faust, 1898, comb. nov. from Eugnamptus
Caenorhinus (Flavodeporaus) klapperichi (Voss, 1941), comb. nov. from Deporaus
Caenorhinus (Nigroarodepus) biounctatus (Legalov, 2007), comb. nov. from Nigroarodepus
Capylarodepus medvedevi (Legalov, 2007), comb. nov. from Capylarodepopsis
Cneminvolvulus (Metinvolvulus) haradai (Kôno, 1940), comb. nov. from Rhynchites
Cneminvolvulus (Metinvolvulus) sundukovi (Legalov, 2007), comb. nov. from Metinvolvulus
Cneminvolvulus (Opacoinvolvulus) aterrimus (Voss, 1938), comb. nov. from Rhynchites
Cneminvolvulus (Opacoinvolvulus) cognatus (Voss, 1958), comb. nov. from Involvulus
Cneminvolvulus (Opacoinvolvulus) gensanensis (Voss, 1920), comb. nov. from Rhynchites
Cneminvolvulus (Opacoinvolvulus) lepidus (Voss, 1930), comb. nov. from Rhynchites
Cneminvolvulus (Opacoinvolvulus) pseudocivicus (Legalov, 2007), comb. nov. from Opacoinvolvulus
Cneminvolvulus (Pilosoinvolvulus) decipiens (Voss, 1938), comb. nov. from Rhynchites
Cneminvolvulus (Pilosoinvolvulus) densatus (Voss, 1938), comb. nov. from Rhynchites
Cneminvolvulus (Pilosoinvolvulus) siwalikensis (Voss, 1956), comb. nov. from Involvulus
Cneminvolvulus (Pilosoinvolvulus) solutus (Faust, 1886), comb. nov.fromRhynchites
Cneminvolvulus (Pilosoinvolvulus) yunnanicus (Voss, 1938), comb. nov.fromRhynchites
Cneminvolvulus (Pseudvulus) meyeri (Voss, 1937), comb. nov.fromRhynchites
Cneminvolvulus (Pseudvulus) nepalensis (Voss, 1937), comb. nov. from Pseudvulus
Deporaus (Chinadeporaus) minuroides (Voss, 1958), comb. nov.fromDepasophilus
Deporaus (Chinadeporaus) nepalensis (Legalov, 2007), comb. nov.fromChinadeporaus
Deporaus (Chinadeporaus) sichuanensis (Legalov, 2007), comb. nov.fromChinadeporaus
Deporaus (Chinadeporaus) strigellus (Voss, 1958), comb. nov.fromDepasophilus
Deporaus (Chinadeporaus) tschungseni (Voss, 1958), comb. nov.fromDepasophilus
Deporaus (Chinadeporaus) yunnanicus (Legalov, 2007), comb. nov.fromChinadeporaus
Deporaus (Paleodeporaus) daliensis (Legalov, 2007), comb. nov.fromPaleodeporaus
70 NEW ACTS and COMMENTS
Deporaus {Paleodeporans) jiriemis (Legalov, 2007), comb. nov. from Paleodeporaus
Deporaus (Paleodeporaus) kathmanduensis (Legalov, 2007), comb. nov. from Paleodeporaus
Deporaus (Paleodeporaus) xishamensis (Legalov, 2007), comb. nov. from Paleodeporaus
Deporaus (Parvodeporaus) gaolingensis (Legalov, 2007), comb. nov. from Parvodeporam
Ecnomonychus (Himalayrhynchites) canus (Faust, 1898), comb. nov. from Rhynchites
Ecnomonychus (Maculinvolvulus) fasciculosus (Voss, 2007), comb. nov. from Rhynchites
Ecnomonychus (Maculinvolvulus) merkli (Legalov, 1942), comb. nov. from Maculinvolvulus
Ecnomonychus (Maculinvolvulus) singularis (Roelofs, 1874), comb. nov. from Rhynchites
Eugnamptobius (Chryseugnamptus) caeruleoflavus (Legalov, 2007), comb. nov. from Chryseugnamptus
Eugnamptobius (Chryseugnamptus) chengduensis (Legalov, 2007), comb. nov. from Chryseugnamptus
Eugnamptobius (Chryseugnamptus) desbrochersi (Legalov, 2007), comb. nov. from Chryseugnamptus
Eugnamptobius (Cyaneugnamptus) lunlinensis (Legalov, 2003), comb. nov. from Cyaneugnamptus
Eugnamptobius (Cyaneugnamptus) pilosellus (Voss, 1941), comb. nov. from Eugnamptus
Eugnamptobius (Cyaneugnamptus) pseudosauteri (Legalov, 2007), comb. nov. from Cyaneugnamptus
Eugnamptobius (Cyaneugnamptus) sauteri (Voss, 1922), comb. nov. from Eugnamptus
Eugnamptobius (Eugnamptobius) subcoeruleifrons (Voss, 1939), comb. nov. from Eugnamptus
Eugnamptobius (Oreugnamptus) lacunosus (Voss, 1949), comb. nov. from Eugnamptus
Eugnamptobius (Oreugnamptus) tessellatus (Voss, 1920), comb. nov. from Eugnamptus
Eugnamptus kubani (Legalov, 2007), comb. nov. from Chineugnamptus
Involvulus (Parinvolvulus) berezovskyi (Legalov, 2007), comb. nov. from Parinvolvulus
Involvulus (Parinvolvulus) hauseri (Wagner, 1907), comb. nov. from Rhynchites
Involvulus (Parinvolvulus) przhevalskyi (Legalov, 2007), comb. nov. from Parinvolvulus
Involvulus (Parinvolvulus) schochrini (Legalov, 2007), comb. nov. from Parinvolvulus
Involvulus (Teretriorhynchites) caeruleus iranensis (Legalov, 2004), comb. nov. from Teretriorhynchites
Involvulus (Teretriorhynchites) dundai (Legalov, 2007), comb. nov. from Teretriorhynchites
Involvulus (Teretriorhynchites) kangdingensis (Legalov, 2007), comb. nov. from Teretriorhynchites
Involvulus (Teretriorhynchites) kozlovi (Legalov, 2007), comb. nov. from Teretriorhynchites
Involvulus (Teretriorhynchites) nigrocyaneus (Legalov, 2007), comb. nov. from Teretriorhynchites
Involvulus (Teretriorhynchites)potanini (Legalov, 2007), comb. nov. from Teretriorhynchites
Involvulus (Teretriorhynchites) xiahensis (Legalov, 2007), comb. nov. from Teretriorhynchites
Involvulus (Teretriorhynchites) zhondiensis (Legalov, 2007), comb. nov. from Teretriorhynchites
Mecorhis (Heterorhynchites) alcyonea (Pascoe, 1875), comb. nov. from Rhynchites
Mecorhis (Heterorhynchites) arguta (Faust, 1882), comb. nov. from Rhynchites
Mecorhis (Heterorhynchites) azurea (Olivier, 1807), comb. nov. from Rhynchites
Mecorhis (Heterorhynchites) bhamoensis (Faust, 1895), comb. nov. from Rhynchites
Mecorhis (Heterorhynchites) copiosa (Voss, 1938), comb. nov. from Rhynchites
Mecorhis (Heterorhynchites) satrapa (Voss, 1938), comb. nov. from Rhynchites
Mecorhis (Heterorhynchites) semenovi (Legalov, 2007), comb. nov. from Heterorhynchites
Mecorhis (Heterorhynchites) pristis (Marshall, 1924), comb. nov. from Rhynchites
Mecorhis (Mirabrhynchites) mirifica (Legalov, 2007), comb. nov. from Mirabrhynchites
Mecorhis (Philorectus) insolita (Voss, 1924), comb. nov. from Philorectus
Mecorhis (Philorectus) sjaomonjanica (Legalov, 2004), comb. nov. from Philorectus
Mecorhis (Pseudomechoris) aethiops (Bach, 1854), comb. nov. from Rhynchites
Mecorhis (Thompsonirhinus) amicta (Voss, 1938), comb. nov. from Rhynchites
Mecorhis (Thompsonirhinus) annapurnae (Legalov, 2007), comb. nov. from Thompsonirhinus
Mecorhis (Thompsonirhinus) coemlescens (Voss, 1938), comb. nov. from Rhynchites
Mecorhis (Thompsonirhinus) coemlescens tsinpinensis (Legalov, 2007), comb. nov. from Thompsonirhinus
Mecorhis (Thompsonirhinus) commutata (Voss, 1969), comb. nov. from Involvulus
Mecorhis (Thompsonirhinus) cyanescens (Voss, 1939), comb. nov. from Rhynchites
Mecorhis (Thompsonirhinus) eduardi (Legalov, 2002), comb. nov. from "Thompsonirhinus"
Mecorhis (Thompsonirhinus) gentilis (Voss, 1930), comb. nov. from Rhynchites
Mecorhis (Thompsonirhinus) gracilis (Voss, 1938), comb. nov. from Rhynchites
Mecorhis (Thompsonirhinus) indubia (Voss, 1930), comb. nov. from Rhynchites
Mecorhis (Thompsonirhinus) kundensis (Legalov, 2007), comb. nov. from Thompsonirhinus
 NEW ACTS and COMMENTS 71
Mecorhis {Thompsonirhinus) leucoscutellata (Voss, 1932), comb. nov.fromRhynchites
Mecorhis {Thompsonirhinus) mandschurica (Voss, 1939), comb. nov.fromRhynchites
Mecorhis {Thompsonirhinus) obsita (Voss, 1930), comb. nov.fromRhynchites
Mecorhis {Thompsonirhinus) pilositessellata (Voss, 1922), comb. nov.fromRhynchites
Mecorhis {Thompsonirhinus)plumbea (Roelofs, 1874), comb. nov. from Rhynchites
Mecorhis {Thompsonirhinus) sichuanensis (Legalov, 2007), comb. nov.fromThompsonirhinus
Mecorhis {Thompsonirhinus) simulons (Voss, 1924), comb. nov.fromRhynchites
Neocoenorrhinus {Neocoenorhinidius) cuprinus (Chevrolat, 1860), comb. nov.fromRhynchites
Neoeugnamptus {Proteugnamptellus) bimaculosus (Voss, 1939), comb. nov.fromEugnamptus
Neoeugnamptus {Proteugnamptellus) da mon I uns is (Legalov, 2003), comb. nov.fromProteugnamptellus
Neoeugnamptus {Proteugnamptellus) indicus (Legalov, 2003), comb. nov.fromProteugnamptellus
Neoeugnamptus {Proteugnamptellus) julonxueshanicus (Legalov, 2007), comb. nov.fromProteugnamptellus
Orrhynchites consimilis (Voss, 1938), comb. nov.fromRhynchites
Rhynchites {Epirhynchites)fiiedmani (Legalov, 2007), comb. nov.fromEpirhynchites
Rhynchites {Epirhynchites) zherichini (Legalov, 2004), comb. nov.fromEpirhynchites

New locality
Neocoenorrhinus {Neocoenorrhinus) germanicus: SP: NAVARRA: 1 ex., Roncesvalles, 29-VI-[19]76 (T.
Yélamos) (Col. Alonso-Zarazaga, MNCN)

New replacement name

Involvulus legalovi Alonso-Zarazaga, nom. nov. for Involvulus {Parinvolvulus) ussuriensis (Legalov, 2003), a
secondary homonym [non Involvulus {Teretriorhynchites) caeruleus ussuriensis (Voss, 1930)]

New synonyms
Aphlorhynchites Sawada, 1993, syn. nov. of Teretriorhynchites Voss, 1938
A thompsonirhinus Legalov, 2003, syn. nov. of Thompsonirhinus Legalov, 2003
Chineugnamptus Legalov, 2007, syn. nov. of Eugnamptus Schoenherr, 1839
Chinorhynchites Legalov, 2003, syn. nov. of Teretriorhynchites Voss, 1938
Chinorhynchitoides Legalov, 2007, syn. nov. of Teretriorhynchites Voss, 1938
Colonnellinius Legalov, 2003, syn. nov. of Epirhynchites Voss, 1953
Eorhinchus Legalov, 2007, syn. nov. of Thompsonirhinus Legalov, 2003
Eosawadaia Legalov, 2004, syn. nov. of Heterorhynchites Voss, 1932
Fujarhynchites Legalov, 2007, syn. nov. of Parinvolvulus Legalov, 2003
Japonodeporaus Legalov, 2007, syn. nov. of Roelofsideporaus Legalov, 2003
Maculinvoles Legalov, 2003, syn. nov. of Thompsonirhinus Legalov, 2003
Neorhynchites Voss, 1969, syn. nov. of Epirhynchites Voss, 1953
Nigroinvolvulus Legalov, 2003, syn. nov. of Parinvolvulus Legalov, 2003
Paleodeporaoides Legalov, 2007, syn. nov. of Paleodeporaus Legalov, 2003
Parinvolvoides Legalov, 2003, syn. nov. of Parinvolvulus Legalov, 2003
Probyctiscidius Legalov, 2007, syn. nov. of Byctiscidius Voss, 1923, objective synonyms. Byctiscidius
insularis Voss, 1923 is the type species of the former genus by original designation and of the latter by
monotypy. The correct type species is mentioned by Alonso-Zarazaga & Lyal (1999) and by Legalov
(2003b), so it is not easily understandable why Legalov (2007a: 199) gives as type species of Byctiscidius
the species Rhynchites parcus Voss, 1920, which was not placed in this genus until Voss (1931b: 219)
made an invalid subsequent designation of this species as type species of his genus, overlooking the extant
monotypy. For the time being, R. parcus is kept in Byctiscidius sensu lato.
Protoessodius Legalov, 2003, syn. nov. of Essodius Sharp, 1889
Pseudorhinoides Legalov, 2007, syn. nov. of Thompsonirhinus Legalov, 2003
Pseudorhinus Legalov, 2003, syn. nov. of Heterorhynchites Voss, 1932
Pyrorhynchites Legalov, 2003, syn. nov. of Epirhynchites Voss, 1953
Sawadaia Alonso-Zarazaga & Lyal, 1999, syn. nov. of Heterorhynchites Voss, 1932
Terminassianaeus Legalov, 2003, syn. nov. of Epirhynchites Voss, 1953
Tshernyshevinius Legalov, 2003, syn. nov. of Epirhynchites Voss, 1953
72 NEW ACTS and COMMENTS
Vietnamorhynchites Legalov, 2003, syn. nov. of Heterorhynchites Voss, 1932
Bydiscus princeps regalis (Roelofs, 1874), syn. nov. of Byctiscus princeps (Solsky, 1872)
Eugnamptus bimaculosus Voss, 1941, syn. nov. of Neoeugnamptus (Proteugnamptellus) bimaculosus (Voss,
1939)
Eugnamptus subcoeruleifrons Voss, 1941, syn. nov. of Eugnamptobius {Eugnamptobius) subcoeruleifrons
(Voss, 1939)
Involvulus aethiops juraensis Voss, 1969, syn. nov. of Mecorhis (Pseudomechoris) aethiops (Bach, 1854)
Rhinomacer testaceus Harrer, 1784, syn. nov. of Lasiorhynchites (Stenorhynchites) caeruleocephalus (Schaller,
1783)
Rhynchites comosellus Voss, 1938 (p. 159), syn. nov. of Orrhynchites consimilis (Voss, 1938)

Priority
Depending upon authors' likings, priority of Jekel (1860a) over Motschulsky (1860) has been reversed or
grossly ignored, even with contradicting uses in different species. I confirm here this priority, which had
already been sanctioned by Sharp (1889: 61) and Faust (1890b: 428), and consequently modified very
recently by incorrect usages.

Spelling
Aspidobyctiscus (Taiwanobyctiscus) cyanocupenis Legalov & Liu, 2005 is an "incorrect original spelling"
according to Art. 32.5.1, and its correction is mandatory, since the authors mention in the 'Etymology' the
correct Latin adjective cupreus as the origin for their name. It has been corrected here to cyanocupreus.
However, I have refrained from correcting the incorrect first component of the composite epithet to cyanei-,
since incorrect latinization must not be corrected.
Auletobius habashanensis Legalov, 2007a is spelled on p. 38 both as habashanensis and hanashanensis. Since
the type locality is correctly named Habashan, habashanensis is here selected as the valid original spelling
(First Revisor action, Art. 24.2.3).
Deporaus septemtrionalis Sawada, 1993, although an incorrect latinization, is used consistently by this author
throughout his text. The spelling septentrionalis in Legalov (2003) is an incorrect subsequent spelling.
Eugnamptus ixigerum Voss, 1941 is an incorrect original spelling, since the genus is masculine in gender, and
is here transcribed as ixiger. The selected epithet is a bad latinization (it should have been exigerum) and is
not corrected here.
Elautobius yunnanicus Legalov, 2003 is spelled yannanicus on p. 237 and yunnanicus in the captions of fig.
683-698 on p. 233, being "alternative original spellings". I hereby, acting as the First Reviser, select the
spelling yunnanicus as valid, so that the spelling yannanicus is an incorrect original spelling (Art. 24.2.3).
Involvulus gemma (Semenov & Ter-Minasian, 1937): The spelling gemmus used by Legalov (2003) does not
meet the requirements of the Code and is an unavailable incorrect subsequent spelling. The word is a Latin
substantive of the 1st declension, meaning "jewel, precious stone"; it cannot be declined for gender.
Mecorhis Billberg, 1820 is the correct spelling of this genus, and its proper Latin rendering (from Greek mekôs,
long, and rhis, nose). Legalov (2003), apparently following Voss, used the incorrect subsequent spelling
Mechoris constructed for the first time in synonymy by Stephens (1833). The correct spelling has been used
as valid by Alonso-Zarazaga & Lyal (1999). Legalov (/. c.) erroneously treated this genus as masculine,
even if Alonso-Zarazaga & Lyal (1999) showed it is feminine. Legalov (2007a) made a new incorrect
subsequent spelling using only Mecoris.
Mecorhis (Thompsonirhinus) satrapa Voss, 1938: The spelling satrapus used by Legalov (2003c, 2007a) does
not meet the requirements of the Code and is an unavailable incorrect subsequent spelling, the word is a
Latin substantive of the 1st declension, coming from a Greek word, meaning "governor of a Persian
province", and it cannot be declined for gender.
Nigroinvolvulus Legalov, 2003c: 298 is here deemed, acting as the First Revisor (Art. 24.2.3), to be the valid
spelling, and the alternative original spelling Nigroinvlovulus (same page) is here rejected, becoming
unavailable.
Opacoinvolvulus pseudocivinus Legalov, 2007a: 163 is here deemed to be an incorrect original spelling, since
the author states that "this new species is very close to O. civinus" and that "the name is formed by addition
of the prefix «pseudo-» to «civinus»", but the species meant and treated close to it is O. civicus. It is
corrected to O. pseudocivicus under Art. 32.5.1.
 NEW ACTS and COMMENTS 73
Pseudocartomynchites Legalov, 2007a: 146 is here deemed to be an incorrect original spelling, based on the
fact that the author, in the Etymology, states that the name "is formed by addition of the prefix «pseudo-» to
«cartorhynchites». It is here corrected to Pseudocartorhynchites, under Art. 32.5.1.
Sawadaeuops sichuanensis Legalov & Liu, 2005 (p. 121, description) is here selected as the correct original
spelling over the spelling S. sitchuanensis used on p. 105 (figs. 31, 32), p. 109 (fig. 65) and p. 121 (last line
of key), acting as First Reviser (Art. 24.2.3).

Unavailable names
Aspidobyctiscus (Taiwanobyctiscus) marinus was the combination proposed by Legalov (2003c: 328) for
Byctiscus paviei f. marina Voss, 1922, an unavailable infrasubspecific name. Legalov failed to fulfill the
requirements of Art. 16.1, so Aspidobyctiscus (Taiwanobyctiscus) marinus is unavailable as well. Its valid
name is Aspidobyctiscus (Taiwanobyctiscus) separandus (Voss, 1930).
Eumetopon flavomaculatus eduardi Legalov, 2003, proposed as a RN for Auletobius flavomaculatus f.
chinensis Voss, 1939 in synonymy of Eumetoponflavomaculatum(Voss, 1922) is unavailable (Art. 11.6.3).
Parinvolvulus (Parinvolvulus) carinaticollis was the combination proposed by Legalov (2007a: 185) for
Involvulus (Involvulus) planiusculus f. carinaticollis Voss, 1958, an unavailable infrasubspecific name.
Legalov failed to fulfill the requirements of Art. 16.1, so Parinvolvulus (Parinvolvulus) carinaticollis is
unavailable as well. Its valid name is Involvulus (Parinvolvulus) immixtus Voss, 1969.

Attelabidae
M. A. Alonso-Zarazaga
Comments
The comments made under Rhynchitidae on Legalov's influence in the systematics and nomenclature of the
taxa of this family are also valid here, as well as many others, since both families have been treated as a
single one for a long time. The phylogenetic analysis of the family conducted by Legalov (2003) ended in a
classification where a paraphyletic subfamily Attelabinae was accepted (in the classic sense). In the present
treatment only the tribes presented by this author as monophyletic entities will be used. The general
upgrading of taxa one or two ranks without justification has also been corrected, following the
abovementioned criticisms. Many of these taxa will probably not withstand a serious, in-depth analysis, as
already pointed by Riedel (2009).
Legalov (2003b, 2007a) is applying incorrectly the Code, by considering that Voss (1933c: 117) made available
a subgenus Charops, with type species by monotypy Euops armipes Voss, 1933. After 31 December 1930,
any new genus-group taxon needed to comply with Art. 13.1 to be available, or, in the case of combined
descriptions, needed to be "marked by "gen. nov., sp. nov." or an equivalent expression" (Art. 13.4). This is
not the case wim this use of Charops placed between the genus and species name in the description of
Euops (Charops) armipes Voss, 1933, without any further mention or description. It is also not
understandable why the available name Charops Riedel, 1998 (type species Euops paradoxus Voss, 1935
by original designation), a junior homonym of Charops Holmgren, 1858) has been replaced by the
unavailable Charops Voss, 1933, and why Riedeliops Alonso-Zarazaga & Lyal, 2002, a replacement name
for Riedel 's nominal taxon, has been placed as a replacement name of the unavailable Charops Voss. This
has produced also undesirable changes in adscription of specific taxa. Consequently, some changes are
needed (see below).
Legalov (2003b: 481; 2007a: 301) records as type species for Pseudoplapoderus Voss, 1932 the species
Centrocorynus [sic!] charlottae Voss, 1932, by monotypy. However, as it has been already pointed out by
Alonso-Zarazaga & Lyal (1999: 46) this nominal taxon is unavailable, since it was published after 1930 and
had no original type species designation, the mention of it by Legalov being this author's invention. The
original description included three species, namely, Paracentrocorynus charlottae sp. n. (and not
Centrocorynusl), Centrocorynus inspersus Voss, 1929 (currently placed in Maculphrysus) and
Hoplapoderus nepalensis Voss, 1926 (currently in synonymy of//, echinatus (Gyllenhal, 1833)). This
inclusion is demonstrated by the use of plural in this adscription sentence: "Von den verwandten Arten
haben sich die genannten, von denen die vorliegende die ausgezeichnetste darstellt, durch ihre
charakteristischen Merkmale scharf ab, es wird daher für diese [my emphasis] die Untergattung
Pseudoplapoderus vorgeschlagen." Being a genus-group nominal taxon without an original type species
74 NEW ACTS and COMMENTS
designation, it is unavailable (Art. 13.3). The type species designations and descriptions made by Legalov
(2003b, 2007a) do not make it available either, since they fail to comply with Art. 16.1. Consequently,
Pseudoplapoderus Voss, 1932 is not included in this Catalogue. The species, being similar to others in the
genus Maculphrysus, is here included in the latter. To confuse things further, there is a subgenus
Pseudplapoderus Legalov, 2003 of genus Paroplapoderus Voss, 1926, differring in just one letter, and
being a bad Greek formation.
Legalov (2007a: 329) found that the two syntypes of Attelabus gemmatus Thunberg, 1784 represented two
different species, one being currently known as Hoplapoderus gemmatus (Thunberg, 1784) and the other
known as Hoplapoderus gemmosus (Jekel, 1860) (and not gemnosus, as he uses to write!). Both species
were easily discernable even if one of the specimens was "strongly destroyed". He selected the specimen
belonging to Hoplapoderus gemmosus as the lectotype of Attelabus gemmatus, making thus both taxa
subjective synonyms, and leaving one taxon without name, which he described as a new species with a few
lines on the next page. With this action, he threw the constant concept and use of a species for 223 years
overboard. Since this action is contrary to the Code aims of stability and universality, I do not take
Legalov's action in consideration and I will apply to the International Commission of Zoological
Nomenclature for its suppression. In the meantime, I keep the current usage as it was before Legalov's
action.
"Apoderus nitens" Kôno, 1930c: 66, currently placed in synonymy of Compsapoderus erythrogaster (Snellen
van Vollenhoven, 1865) by several catalogues, is a misidentification of Apoderus nitens Roelofs, 1874, and
is nomenclaturally unavailable. It has not been recorded in the main text.

Changes in rank
Alexsandricorynus Legalov, 2003, stat dem. to subgenus of Physapoderus Jekel, 1860
Allolabus Voss, 1925, stat. dem. to subgenus of Henicolabus Voss, 1925
Asuniops Legalov, 2007, stat dem. to subgenus of Euops Schoenherr, 1839
Centrocorynidius Legalov, 2003, stat. dem. to subgenus of Physapoderus Jekel, 1860
Chinolabus Legalov, 2003, stat. dem. to subgenus of Henicolabus Voss, 1925
Eocentrocorynus Legalov, 2003, stat. dem. to subgenus of Physapoderus Jekel, 1860
Helleriapoderus Legalov, 2003, stat dem. to subgenus of Strigapoderus Jekel, 1860
Henicolaboides Legalov, 2007, stat dem. to subgenus of Henicolabus Voss, 1925
Heterapoderites Legalov, 2003, stat. dem. to subgenus of Heterapoderus Voss, 1927
Heterapoderopsis Legalov, 2003, stat. dem. to subgenus of Heterapoderus Voss, 1927
Korotyaevirhinus Legalov, 2003, stat dem. to subgenus of Hamiltonius Alonso-Zarazaga & Lyal, 1999
Macroderites Legalov, 2003, stat dem. to subgenus of Heterapoderus Voss, 1927
Macrosynaptopsis Legalov, 2005, stat. dem. to subgenus of Euops Schoenherr, 1839
Metocalolabus Legalov, 2003, stat dem. to subgenus of Calolabus Voss, 1925
Microcorynus Legalov, 2003, stat dem. to subgenus of Heterapoderus Voss, 1927
Paracompsus Legalov, 2003, stat dem. to subgenus of Compsapoderus Voss, 1927
Parasuniops Legalov, 2003, stat dem. to subgenus of Euops Schoenherr, 1839
Parasynatops Legalov, 2003, stat dem. to subgenus of Euops Schoenherr, 1839
Philippinotrachelus Legalov, 2003, stat. dem. to subgenus of Physapoderus Jekel, 1860
Pseudoeuops Legalov, 2003, stat dem. to subgenus of Euops Schoenherr, 1839
Suniops Voss, 1928, stat dem. to subgenus of Euops Schoenherr, 1839
Suniopsidius Legalov, 2005, stat. dem. to subgenus of Euops Schoenherr, 1839

Infrasubspecific names
The following names (and many others) were clearly proposed as infrasubspecific names under Art. 45.6.2 or
45.6.4 of the Code, and have been omitted from the list. In these cases, Art. 45.6.4.1 cannot be applied.
Euops lespedezae f. koreanus Voss, 1924 [Art. 45.6.4, raised to subspecies as Euops lespedezae koreanus
Sawada & Morimoto, 1985]
Euops splendida f. nigra Kôno, 1927 [Art. 45.6.4, raised to species as Parasynaptopsis nigra Legalov, 2003
(incorrectly rendered as nigrum, gender is masculine)]. Currently a synonym of Euops {Parasynaptopsis)
splendidus Voss, 1930.
Art. 45.6.4.1 is to be applied here:
 NEW ACTS and COMMENTS 75
Apoderus rubidus f. thibetana Voss, 1927, an infrasubspecific name, raised to subspecies by Ter-Minassian,
1950: 210, is deemed to be available from its original description. Now a valid species in Leptapoderus
Jekel, 1860.

New combinations
Calolabus (Metocalolabus) chujoi (Kôno, 1939), comb. nov.fromHimatolabus
Catalabus (Catalaboides) marieae (Kresl, 2007), comb. nov.fromParamecolabus
Compsapodenis (Paracompsus) afghanus (Legalov, 2003), comb. nov.fromParacompsus
Compsapoderus {Paracompsus) bistriolatus (Faust, 1898), comb. nov.fromApoderus
Compsapoderus (Paracompsus) corallinoides (Legalov, 2007), comb. nov. from Paracompsus
Compsapoderus (Paracompsus) lepidulus (Voss, 1927), comb. nov.fromApoderus
Compsapoderus (Paracompsus) lunanfensis (Legalov, 2007), comb. nov.fromParacompsus
Compsapoderus (Paracompsus) luteibasis (Voss, 1935), comb. nov.fromApoderus
Compsapoderus (Paracompsus) micros (Legalov, 2007), comb. nov.fromParacompsus
Compsapoderus (Paracompsus) nigrofasciatus (Pajni, Haq & Gandhi, 1987), comb. nov.fromApoderus
Compsapoderus (Paracompsus) nigroscutellaris (Legalov, 2007), comb. nov.fromParacompsus
Compsapoderus (Paracompsus) pygidialis (Voss, 1927), comb. nov.fromParacompsus
Compsapoderus (Paracompsus) seclusus (Faust, 1898), comb. nov.fromApoderus
Compsapoderus (Paracompsus) varions (Voss, 1928), comb. nov.fromApoderus
Euops (Asuniops) opacus (Legalov, 2007), comb. nov.fromAsuniops
Euops (Chinoeuops) davidiani (Legalov, 2003), comb. nov.fromSawadaeuops
Euops (Chinoeuops) hubeiensis (Legalov & Liu, 2005), comb. nov.fromSawadaeuops
Euops (Chinoeuops) nielamus (Legalov & Liu, 2005), comb. nov.fromSawadaeuops
Euops (Kobusynaptops) verrucosus (Legalov & X. Zhang, 2007), comb. nov.fromKobusynaptops
Euops (Levoeuops) asiaticus (Legalov & Liu, 2005), comb. nov.fromRiedeliops
Euops (Levoeuops) asiaticus uttaranchalicus (Legalov, 2007), comb. nov.fromRiedeliops
Euops (Levoeuops) nepalensis (Legalov, 2003), comb. nov.fromRiedeliops
Euops (Macrosynaptopsis) zhangi (Legalov & Liu, 2005), comb. nov.fromMacrosynaptopsis
Euops (Parasuniops) fochaiensis (Legalov, 2003), comb. nov.fromParasuniops
Euops (Parasynaptopsis) carinatus (Legalov & X. Zhang, 2007), comb. nov.fromMorphoeuops
Euops (Parasynaptopsis) pseudostriatus (Legalov, 2007), comb. nov.fromParasynaptopsis
Euops (Parasynatops) berezovskyi (Legalov, 2003), comb. nov.fromParasynatops
Euops (Parasynatops) moanus (Legalov, 2003), comb. nov.fromParasynatops
Euops (Parasynatops) pseudopolitus (Legalov, 2003), comb. nov.fromParasynatops
Euops (Parasynatops) tzinpinensis (Legalov & Liu, 2005), comb. nov.fromParasynatops
Euops (Pseudoeuops) bicoloroides (Legalov, 2003), comb. nov.fromParasynatops
Euops (Pseudoeuops) lushuensis (Legalov & Liu, 2005), comb. nov.fromParasynatops
Euops (Pseudoeuops) pyralis (Legalov & Liu, 2005), comb. nov.fromParasynatops
Euops (Pseudoeuops) tibetanus (Legalov & Liu, 2005), comb. nov.fromParasynatops
Euops (Pseudoeuops) viridis (Legalov, 2003), comb. nov.fromParasynatops
Euops (Riedeliops) beijingensis (Legalov, 2003), comb. nov.fromParasynatops
Euops (Riedeliops) centralchinensis (Legalov & Liu, 2005), comb. nov.fromSawadaeuops
Euops (Riedeliops) meghalaicus (Legalov, 2007), comb. nov.fromOrienteuops
Euops (Riedeliops) potanini (Legalov, 2003), comb. nov.fromParasynatops
Euops (Riedeliops) sichuanensis (Legalov & Liu, 2005), comb. nov.fromSawadaeuops
Euops (Sawadaeuopsis) punctatus (Legalov & Liu, 2005), comb. nov. from Sawadaeuops
Euops (Suniopsidius) multicoloratus (Legalov & Liu, 2005), comb. nov.fromSuniopsidius
Euops (Vietsuniops) menghunsis (Legalov & Liu, 2005), comb. nov.fromSuniops
Hamiltonius (Korotyaevirhinus) notatus (Fabricius, 1792), comb. nov. from Korotyaevirhinus
Henicolabus (Chinolabus) bicolor (Legalov & Liu, 2005), comb. nov.fromChinolabus
Henicolabus (Chinolabus) gansuanus (Legalov, 2007), comb. nov. from Chinolabus
Henicolabus (Chinolabus) ningxianus (Legalov & Liu, 2005), comb. nov.fromChinolabus
Henicolabus (Henicolaboides) nanlingensis (Legalov, 2008), comb. nov. from Henicolaboides
Henicolabus (Henicolaboides) oberthueri (Legalov, 2007), comb. nov.fromHenicolaboides
76 NEW ACTS and COMMENTS
Henicolabus (Henicolaboides) potanini (Legalov, 2007), comb. nov. from Henicolaboides
Heterapodenis (Helleriapodems) subvariolaris (Voss, 1935), comb. nov. from Apoderus
Heterapodenis (Macroderites) nepalensis (Legalov, 2003), comb. nov. from Macroderites
Heterapoderus (Macroderites) orchinensis (Legalov, 2007), comb. nov. from Macroderites
Heterapodenis (Microcorynus) blandus (Faust, 1895), comb. nov. from Apoderus
Heterapoderus (Microcorynus) blandus blandoloides (Legalov, 2007), comb. nov. from Microcorynus
Heterapoderus (Microcorynus) parunicus (Legalov, 2003), comb. nov. from Microcorynus
Heterapoderus (Strigapodems) scabrosus (Voss, 1928), comb. nov. from Apoderus
Maculphrysus charlottae (Voss, 1932), comb. nov. from Paracentrocorynus
Physapodenis (Alexsandricorynus) assamensis (Boheman, 1845), comb. nov. from Apoderus
Physapodenis (Eocentrocorynus) antennalis (Legalov, 2007), comb. nov. from Eocentrocorynus
Physapodenis (Eocentrocorynus) breviclavus (Legalov, 2007), comb. nov. from Eocentrocorynus
Physapodenis (Eocentrocorynus) fosculus (Voss, 1929), comb. nov. from Centrocorynus
Physapodenis (Eocentrocorynus) fusculus yunnanicus (Legalov, 2003), comb. nov. from Eocentrocorynus
Physapodenis (Eocentrocorynus) gracilicornis (Voss, 1929), comb. nov. from Centrocorynus
Physapodenis (Eocentrocorynus) huangi (Legalov, 2007), comb. nov. from Eocentrocorynus
Physapodenis (Eocentrocorynus) pokharensis (Legalov, 2003), comb. nov. from Eocentrocorynus
Physapodenis (Eocentrocorynus) ruficlavis (Voss, 1929), comb. nov. from Centrocorynus
Physapodenis (Eocentrocorynus) sichuanensis (Legalov, 2003), comb. nov. from Eocentrocorynus
Pseudallapoderus ateroides (Legalov, 2002), comb. nov. from Strigapodems
Tomapodems himalayensis (Legalov, 2007), comb. nov. from Tomapoderopsis
Tomapodems orientalis (Legalov, 2007), comb. nov. from Tomapoderopsis
Tomapoderus pici (Legalov, 2007), comb. nov. from Tomapoderopsis

New placements
Riedeliops (Levoeuops) Legalov, 2007 is transferred to Euops (Levoeuops)
Riedeliops (Riedeliopsis) Legalov, 2007 is transferred to Euops (Riedeliopsis)
Sawadaeuops (Chinoeuops) Legalov, 2003 is transferred to Euops (Chinoeuops)
Sawadaeuops (Sawadaeuopsis) Legalov, 2005 is transferred to Euops (Sawadaeuopsis)
Suniops (Vietsuniops) Legalov, 2007 is transferred to Euops (Vietsuniops)

New replacement name

Homapoderus (Homapoderus) gyllenhali Alonso-Zarazaga, nom. nov. for Homapoderus (Homapoderus)
cyaneus Gyllenhal, 1839 (Apoderus), non Hope, 1831

New synonymies
Aheterapoderus Legalov, 2007, syn. nov. of Heterapoderus Voss, 1927
Ahumerilabus Legalov, 2007, syn. nov. of Humerilabus Legalov, 2003
Algirolabus Legalov, 2003, syn. nov. of Attelabus Linnaeus, 1758
Amacroderites Legalov, 2007, syn. nov. of Macroderites Legalov, 2003
Aphrysapoderus Legalov, 2007, syn. nov. of Physapodenis Jekel, 1860
Compsapoderopsis Legalov, 2003, syn. nov. of Compsapodems Voss, 1927
Cyanotrachelophorus Legalov, 2003, syn. nov. of Paratrachelophorus Voss, 1924
Eoallolabus Legalov, 2003, syn. nov. of Allolabus Voss, 1925
Eoheterapodems Legalov, 2003, syn. nov. of Heterapoderus Voss, 1927
Eophrysapoderus Legalov, 2007, syn. nov. of Physapodenis Jekel, 1860
Euscelophiloides Legalov, 2007, syn. nov. of Euscelophilidius Legalov, 2003
Formosusorynus Legalov, 2003, syn. nov. of Eocentrocorynus Legalov, 2003
Granulapoderus Legalov, 2003, syn. nov. of Apodems Olivier, 1807
Jekelilabus Legalov, 2003, syn. nov. of Allolabus Voss, 1925
Konoapodems Legalov, 2003, syn. nov. of Apodems Olivier, 1807
Morphotrachelophorus Legalov, 2003, syn. nov. of Paratrachelophoms Voss, 1924
Neheterapodems Legalov, 2003, syn. nov. of Heterapoderus Voss, 1927
Neocorynidius Legalov, 2003, syn. nov. of Eocentrocorynus Legalov, 2003
 NEW ACTS and COMMENTS 77
Neparasynatops Legalov, 2007, syn. nov. of Parasynatops Legalov, 2003
Parahamiltonius Legalov, 2007, syn. nov. of Hamiltonius Alonso-Zarazaga & Lyal, 1999
Parmaculphrysus Legalov, 2003, syn. nov. of Maculphrysus Legalov, 2003
Proattelabus Legalov, 2003, syn. nov. of Attelabus Linnaeus, 1758
Pseudcycnolodes Legalov, 2003, syn. nov. of Cycnotrachelodes Voss, 1955
Pseudmadaranus Legalov, 2003, syn. nov. of Agomadaranus Voss, 1958
Pseudoheterapoderus Legalov, 2003, syn. nov. of Heterapoderus Voss, 1927
Pseudoparaphorus Legalov, 2003, syn. nov. of Paratrachelophorus Voss, 1924
Tomapoderopsis Legalov, 2003, syn. nov. of Tomapoderus Voss, 1926

Spelling
Agomadaranus sticticus yannanicus Legalov, 2004 is here deemed to be an incorrect original spelling for
yunnanicus, since in the original description the origin is mentioned as Yunnan, under Art. 32.5.1. This
corrected spelling is the only one used later by the same author (Legalov, 2007a)
Attelabus octomaulatus, spelling insistently used by Legalov (2007a), is an incorrect subsequent spelling of
Attelabus octomaculatus Jekel, 1860
Centrocorynidius Legalov, 2003, etymologically correct, appears in Legalov (2003: p. 559) as an alternative
original spelling and, acting as a First Reviser (Art. 24.2.3), it is here selected as valid; the spelling
Centrocorinidius is here rejected as invalid
Cycnotrachelodes camphoricola (Voss, 1924) is the valid spelling for this species, since the specific epithet
camphoricola is a substantive in apposition ("the one who lives in camphor-trees"). C. camphoricolus, as
used by Legalov, is an incorrect subsequent spelling, and is unavailable
Euopini Voss, 1925 is the correct name of the tribe based on the genus Euops Schönherr, 1839. The spelling
Euopsini is contrary to the Code, since the stem of Euops is Euop-, but this is the incorrect spelling
insistently used by Legalov
Euops pseudlespedezae Z.-Q. Li is here deemed to be an incorrect alternative original spelling and Euops
pseudolespedezae, etymologically correct, is deemed to be valid. Action of First Reviser (Art. 24.2.3).
Leptapoderus hymalayensis is here deemed to be an incorrect original spelling, thence unavailable, since the
author states in its etymology: "from Himalaya - «hymalayensis»"..It is corrected to himalayensis (Art.
32.5.1)
Physapoderus Jekel, 1860 is incorrectly written Phrysapoderus in Legalov's work, insistently (incorrect
subsequent spelling)
Tomapoderopsis hymalayensis Legalov, 2007 is here deemed to be an incorrect original spelling, thence
unavailable, since its author states in its etymology: "from Himalaya - «hymalayensis»". It is corrected to
himalayensis (Art. 32.5.1)

Brentidae
M. A. Alonso-Zarazaga
Spelling
Baryrhynchus minisculus R.-Z. Zhang, 1993 (p. 161 & 172, keys) is here selected as the correct original spelling
over the spelling B. minisulus used on p. 168, acting as First Reviser (ICZN, Art. 24.2.3). This name is a
probable misspelling of minusculus that cannot be corrected.
Baryrhynchus setulosus R.-Z. Zhang, 1993 (p. 160, 166 & 167) is here selected as the correct original spelling
over the spelling B. setolosus used on p. 171, acting as First Reviser (ICZN, Art. 24.2.3).

Apionidae
M. A. Alonso-Zarazaga
Comments
Some species have been mentioned in past Palaearctic catalogues in error: "Apion" periscelis Gyllenhal, 1839,
from Persia (correctly from Peru, probably a misreading of the label); Coelopterapion subulirostre
(Gyllenhal, 1833), a Brazilian species (redescribed from Japan under its synonym Apion daimio Sharp,
1891, cf. Kissinger (1989)).
78 NEW ACTS and COMMENTS
Apion parvulum Mulsant & Rey, 1858 is a primary homonym of A. parvulum Gerstäcker, 1854 (now in
Coelocephalapion), an American species. This species was described from "Lyon, Beaujolais", from an
unspecified number of specimens of both sexes. The Muséum d'Histoire naturelle de Lyon sent for study a
series of 4 specimens kept in the Coll. Claude Rey as types. They are: 1) one male, rectangular label: $;
length 1.215 mm (without rostrum), more than apical half of prorostrum bright, ratio lenght of
rostrum/length of pronotum (Lrp): 1.03; 2) one male, with a round label, length 1.243 (without rostrum),
rostrum bright in apical third, Lrp: 0.91; 3) one female, rectangular label: ? , immature with testaceous
appendages, length 1.436 mm (without rostrum), rostrum with bright apex only, Lrp: 1.36; 4) one male, no
label, length 1.381 mm (without rostrum), more than half the prorostrum bright Lrp: 0.92. They fit the
description of the species currently known as Apion hoffinanni Wagner, 1930 as exposed by Dieckmann
(1977). The first specimen is here designated as lectotype, and the three other as paralectotypes, of Apion
parvulum Mulsant & Rey, 1858. This species must be known as Squamapion serpyllicola (Wencker, 1864)
as shown below.
Apion schilskyi Wagner, 1906 is currently placed in Pseudopiezotrachelus, but it has never been found again
and Wagner himself stated that the species had an exotic appearance, reminiscent of Central-American
species. It is possible that the label the holotype carries is false and the species does not belong to the
Palaearctic fauna, as it has happened before (see above).
Apion serpyllicola Wencker, 1864 was originally mentioned as a synonym of the concept that Wencker had of
Apion parvulum Mulsant & Rey, 1858. The latter concept is coincident with the original, as the extant types
show, despite the opposing comments of many authors who did not study the types and based their
judgements just on the fact that Lyon was a "very southern locality". According to Art. 11.6.1 of the Code,
Apion serpyllicola Wencker, 1864 is an available name, having been used as such with its original author
and date for the original concept of Wencker, for example, by Planet (1918: 55) or by Hustäche (1931:
178). Unfortunately, Wagner (1930: 1391) selected an opposite and incorrect view, and created the
unnecessary homonym Apion hoffmanni as a replacement name for Apion parvulum Mulsant & Rey, in
which he was followed unquestioningly by later authors. The decisions taken here according to the Code
are meant to stabilize the nomenclature of these two sibling species as soon as possible. I hereby designate
as lectotype of A. serpyllicola Wencker one female, in bad state and very dirty, near a bottom label stating:
"A. serpyllicola / Wencker / Parvulum ? Muls. et Rey / Coll. Wencker France" and carrying me following
labels: 1) rectangular, small, white: Ç; 2) rectangular, small, green; 3) rectangular, small, white: C; 4)
rectangular, large, manuscript: parvulum / Wencker; 5) rectangular, large, manuscript: parvulum / Wnk. /
serpyllicola; 6) rectangular, large, printed: TYPE (red); 7) rectangular, large, printed: MUSÉUM PARIS /
ANCIENNE COLLECTION (Coll. Générale ex Coll. Wencker). This specimen measures 1.577 mm
(without rostrum), shows the characters listed for A. parvulum by Wencker (I.e.) and for A. hoffmanni
Wagner by Dieckmann (1977), in particular the matt rostrum except at the apex, and the ratio length of
rostrum/length of pronotum is 1.5. This specimen is supposed to come from "Vosges" as stated in me
original description. In Wencker's collection there is a male, after a green label stating: "serpyllicola /
Wencker / Parvulum / Mul. et Rey ?" carrying the following labels: 1) rectangular, small, printed: 101; 2)
rectangular, narrow, small: Francfort. This being one of the originally quoted localities, this specimen is
deemed to be a paralectotype. Both species (S. minutissimum and S. serpyllicola) show an overlapping
distribution in France, that would merit a thorough study.
Apion squamosum Faust, 1884b: 455. Bajtenov (1981c: 640) designated as a lectotype a male from Derbent,
housed in the Museum für Tierkunde, Dresden. This lectotype designation was overlooked by Schoen
(1988a: 40) who designated as lectotype the other female syntype in the same museum, stating that the male
syntype specimen from Derbent (Bajtenov's lectotype) was not conspecific with his (invalid) lectotype.
Consequently, this species may conceal two different taxa. Egypt and Sinai records are probably due to
misidentifications.
Curculio obscurus Marsham, 1802, described from England, based on two specimens, was transferred to the
genus Apion by Kirby (1808) and subsequently mentioned as a British species several times, either as a
variety of Apion fulvirostre or of A. ruflrostre. The study of the two syntypes reveals that this'species is a
synonym of the Nearctic species Trichapion rostrum (Say, 1826) and that its mention from "Anglia" was
erroneous. Consequently, this species is not included in the present catalogue. Marsham's nominal species
is a homonym of Curculio obscurus Fabricius, 1775 (now Barynotus), so that the name of the American
species does not change. This species was described again from "Anglia" as Apion scrobicolle Gyllenhal,
 NEW ACTS and COMMENTS 79
1839. It would be interesting to know whether these are coincidences, or perhaps a population of this
species was present in England at those times.
Oryxolaemus croceifemoratus has been incorrectly cited from MC based on a locality (Keretschkoi) which is
mentioned by the collector himself to be in the Greek Chalkidiki Peninsula (Schatzmayr, 1909, Wiener
Entomologische Zeitung 28(2): 39). Its record from Sladun in Bulgaria (Tracy & Robins, 2009, Zootaxa
2011: 46) is a misidentification. The locality corresponds to present-day Exochi 10 km east of Thessaloniki,
at 40°37'48"N, 23°2'37"E, near the village of Chortiatis (Dr. Anastâsios Legâkis comm. pers.).
Oxystoma cerdo var. dimidiatum Desbrochers des Loges, 1897 was described from an unspecified number of
specimens from "Corse" and "Sicile". According to Dieckmann (1971: 93) its types are lost. In the same
paper this author described Apion pseudocerdo, without recognising the former species. Having studied the
type material, which is not lost, I had the opportunity of using O. dimidiatum instead of O. pseudocerdo
(Alonso-Zarazaga, 1990a: 142) and some other authors followed this practice, even if I had not given a
formal synonymy. I have studied 3 specimens of Oxystoma cerdo var. dimidiatum, conserved in the Coll.
Desbrochers des Loges (Muséum National d'Histoire Naturelle, Paris): lectotype, one male, with labels:
handwritten: Sicile (on the back of the original board) / dimidiatum 1. / Ex Musaeo Desbrochers 1914 /
cerdo; two other specimens, with similar labels, are paralectotypes. These specimens are identical with
Apion pseudocerdo Dieckmann, 1971. The synonymy is formally established below.
In the treatment of Pseudopiezotrachelus collaris (Schilsky, 1906), the synonymies proposed by Korotyaev in
1999 are accepted for the time being. This is probably a species complex and minute but constant
differences can be found across the whole distributional area assigned to this "species", both in external and
internal morphological characters.
Stenopterapion {Cobosiotherium) scutellare (Kirby, 1811) has several records from N: AG TU A: TR SY, but
they have never been confirmed and are suspected to be misidentifications. The species is known only from
the mid-Atlantic coastal strip of Europe.

Changes in rank
Rhodapion Alonso-Zarazaga, 1990, stat. dem. to subgenus of Melanapion Wagner, 1930. This taxon was
upgraded to genus by Colonnelli (2003) for unknown reasons, it is here downgraded again to its original
status.

Homonyms
Apion elongatum v. torquatum Wencker, 1864: 183 {Apion) is here considered to be a junior homonym of
Apionflavimanumv. torquatum Wencker, 1864: 148. First Reviser's Action (Art. 24.2).
Apion subulatum Kirby, 1808 is a primary homonym of Apion subulatum II liger, 1807, and both are placed in
the genus Oxystoma Duméril, 1805, the former as a valid species, me latter as a synonym of O. pomonae
(Fabricius, 1798). Illiger's name has never been used as valid, but I have been unable to fulfill the
requirements of Art. 23.9.1.2 for O. subulatum (Kirby). I keep here the current usage until an application is
presented to the International Commission on Zoological Nomenclature.
Curculio laevigatus Paykull, 1792 was transferred by Schoenherr (1833, p. 309) to the genus Apion as a
synonym of the junior synonym Apion sorbi (Fabricius, 1792), latter replaced with the senior^, laevigatum
(Paykull, 1792). Apion laevigatum Kirby, 1808 was described as a different species. Both species were at
the same time in Apion since 1833 and, being homonyms, A. laevigatum Kirby was replaced by its junior
synonym A. brunnipes Boheman, 1839. Except for a few uses as a valid name, last time in 1863. Apion
brunnipes has been in use as a valid name in the genus Apion since its original description and later in
Acentrotypus, Ceratapion or Taphrotopium. Alonso-Zarazaga ( 1990b) placed it in Acentrotypus and, except
for few very conservative treatments, it has never returned to Apion. In the same paper, Paykull's species
was transferred to Omphalapion, as O. laevigatum (Paykull, 1792). Colonnelli (2003) used Acentrotypus
laevigatus (Kirby) as valid, since the two species were no more congeneric and the secondary homonymy
had ceased to exist. This nomenclatural act is contrary to the Code (Art. 59.3), since Apion laevigatum
Kirby, 1808 is a permanently invalid name, having been replaced before 1961, and its substitute name
{Apion brunnipes) according to Art. 60.1, is in wide use.
Curculio ochropus Gmelin, 1790: 1757 (currently in synonymy of Protapion apricans (Herbst, 1797)) is here
considered to be a junior homonym of Curculio ochropus Gmelin, 1790: 1805 (currently in synonymy of
Polydrususflavipes(DeGeer, 1775)). First Reviser's Action (Art. 24.2).
80 NEW ACTS and COMMENTS
Infrasubspecific names
The following names (and many others) were clearly proposed as infrasubspecific names under Art. 45.6.2 or
45.6.4 of the Code, and have been omitted from the Catalogue. In these cases, Art. 45.6.4.1 cannot be
applied:
Apion aesliviim ab. nificroides Schatzmayr, 1925 [Art. 45.6.2, raised to species as Apion rußcroides
Dieckmann, 1973]. Now a valid species in Protapion Schilsky, 1908.
Apion candidum var. rutae Hustache, 1931 [Art. 45.6.4, raised to subspecies as Metapion candidum rutae
Ehret, 1997].
Apion tubiferum var. sicanum Wencker, 1864 [Art. 45.6.4, raised to subspecies as Phrissotrichum
(Phrissotrichum) sicanum Ehret, 1997].
Apion varium var. melichari Wagner, 1908 [Art. 45.6.4, raised to species as Piezotrachelus melichari Wanat,
1990].
Apion (Perapion) defensum var. nuristanicola Voss, 1959 [Art.45.6.4, raised to species as Perapion
nuristanicola Wanat, 1995]. Synonymized here with Perapion himalayense Bhateja & Pajni, 1989 (see
below).
Art. 45.6.4.1 is to be applied here:
Apion (Ceratapion) sulcifrons var. cuprifulgens Schilsky, 1906, an infrasubspecific name, raised to species by
Schoen (1983: 43), is deemed to be available from its original description. Now a valid species in
Taphrotopium Reitter, 1916.
Apion plicatum var. badakschanicum Voss, 1959, an infrasubspecific name, raised to species by Bajtenov
(1977e: 26), is here deemed to be available from its original description. Now a valid species in
Hemitrichapion Voss, 1959.
Apion rufescens var. not a turn Wagner, 1912, an infrasubspecific name, raised to subspecies by Balfour-Browne
(1944b: 149), is here deemed to be available. Currently a synonym of Taeniapion rufescens (Gyllenhal,
1833).
Apion rufulum var. rolphi and var. zoufali Wagner, 1912, infrasubspecific names, raised to subspecies by
Hustache (1931: 114, 118), are here deemed to be available. Currently synonyms of Taeniapion rufulum
(Wencker, 1864).

New combinations
Aspidapion mountanus (Pajni, Bhateja & Kumar, 1991), comb. nov. from Apion
Holotrichapion splendididum (Desbrochers des Loges, 1875), comb. nov. from Apion
Hoplopodapion depressum (Bhateja & Pajni, 1989), comb. nov. from Onychapion
Hoplopodapion dumonti (Peyerimhoff, 1929), comb. nov. from Apion
Hoplopodapion eremita (Peyerimhoff, 1920), comb. nov. from Apion
Hoplopodapion ferreri (Alonso-Zarazaga, 1983), comb. nov. from Onychapion
Hoplopodapion kirschi (Desbrochers des Loges, 1870), comb. nov. from Apion
Hoplopodapion koestlini (Schoen, 1993), comb. nov. from Onychapion
Hoplopodapion lopatini (Ter-Minassian, 1963), comb. nov. from Apion
Hoplopodapion medvedevi (Korotyaev, 1984), comb. nov. from Onychapion
Hoplopodapion poupillieri (Wencker, 1864), comb. nov. from Apion
Hoplopodapionpumilio (Desbrochers des Loges, 1893), comb. nov. from Apion
Hoplopodapion pyripenne (Reitter, 1908), comb. nov. from Apion
Hoplopodapion reaumuriae (Ter-Minassian, 1975), comb. nov. from Apion
Hoplopodapion spinitarsatum (Bhateja & Pajni, 1989), comb. nov. from Onychapion
Ischnopterapion (Jschnopterapion) plumbeomicans pericarti (Ehret, 1991), comb. nov. from Apion
Ischnopterapion (Jschnopterapion) tempereanum (Ehret, 1991), comb. nov. from Apion
Metapion sojunovii (Bajtenov & Sojunov, 1990), comb. nov. from Apion
Onychapion fasciculosum (Motschulsky, 1851), comb. nov. from Apion
Perapion jelineki (Bajtenov & Fremum, 1981), comb. nov. from Apion
Protapion porrectum (Schrank, 1781), comb. nov. from Curculio, based on description. Identity within genus
doubtful.
 NEW ACTS and COMMENTS 81
Protapion temisiense (Palagesiu, 1988), comb, nov.,fromApion, according to the original description; spelling
corrected to meet the requirements of Art. 31.2. A probable synonym of Protapion varipes (Germar, 1817);
holotype requested for study without success
Pseudaspidapion striopunctatum (Pajni, Bhateja & Kumar, 1991 ), comb. nov.fromApion
Pseudopiezotrachelns placidus (Faust, 1887), comb, nov.,fromApion. However, placement is doubtful
Pseudopiezotrachelus uruppensis (Kôno, 1935), comb, nov., from Apion. However, placement is doubtful
Stenopterapion (Cobosiotherium) subsquamosum (Desbrochers des Loges, 1891), comb. nov.fromApion
Stenopterapion {Stenopterapion) volkovitshi (Korotyaev, 1988), comb. nov.fromLeptapion
Harpapion ceylonicum (Gerstaecker, 1854), comb, nov.,fromApion (Taeniapion)

New localities
Apion rubens: PT: FARO: 1 ex., Castro Marim, 11-VI-1981, #7273 (Serrano) (Col. Artur Serrano)
Aspidapion soror. SP: many specimens, Gibraltar (Walker) (NHM, London)
Exapion lemovicinum: SP: ASTURIAS: Oviedo: 1 f., [Las] Arenas de Cabrales (R. Frey) / 784 / MZH
Holotrichapion {Holotrichapion) umbrinum: SP: all specimens mentioned as ononisfromMalaga province in
Alonso-Zarazaga ( 1980)

New placement
Necatapion Friedman & Freidberg, 2007, described as a genus incertae sedis in Apionidae, is here placed in
subtribe Catapi ina, whose main characters it shares

New replacement names

Perapion {Perapion) barkleyi Alonso-Zarazaga, nom. nov. for Apion smreczynski Bajtenov, 1979 (non Apion
smreczynskii Hoffmann, 1934) (homonyms under Art. 58.14). Cordially dedicated to my good colleague
andfriendMaxwell Barkley (NHM, London).
Taeniapion tyrrhenicum Alonso-Zarazaga, nom. nov. for Apion distinctirostre Desbrochers des Loges, 1888
(non Apion distinctirostre Desbrochers des Loges, 1869).

New synonymies
Apion barbirostre Hoffmann, 1953, syn. nov. of Holotrichapion {Holotrichapion) umbrinum (Wollaston,
1864). Based on absence of distinguishing characters
Apion cretaceum torretassoi Schatzmayr, 1933, syn. nov. of Lepidapion {Hydryocneme) cretaceum
(Rosenhauer, 1856). I have been unable to find stable differences across the distribution of the species.
Specimens from SP are identical to thosefromIS or TU
ApionflavicrusDesbrochers des Loges, 1901, syn. nov. of Exapion squamulatum (Gyllenhal, 1833), based on
comparison of types. Gyllenhal's species was incorrectly recorded originally from "Cap. Bonae Spei", but
in fact it is a common Moroccan species
Apion limonii limoniastri Flach, 1908, syn. nov. of Pseudaplemonus limonii (Kirby, 1808). Based on absence
of distinguishing characters in locotypic specimens
Apion ononis saturnium Normand, 1937, syn. nov. of Holotrichapion {Holotrichapion) umbrinum (Wollaston,
1864). Based on absence of distinguishing characters
Apion parvulum Mulsant & Rey, 1858 (non Gerstäcker, 1854), syn. nov. of Squamapion serpyllicola (Wencker,
1864). Based on study of types
Apion pilosellum Gravenhorst, 1807, syn. nov. of Eutrichapion {Cnemapion) vorax (Herbst, 1797). Based on
MS synonymy by Gravenhorst
Apion pseudocerdo Dieckmann, 1971, syn. nov. (formally) of Oxystoma dimidiatum (Desbrochers des Loges,
1897). Based on study of types
Apion rubens Stephens, 1839, syn. nov. of Apion rubens Walton, 1837. Objective synonymy
Apion serpyllicola Gozis, 1881b, syn. nov. of Squamapion serpyllicola (Wencker, 1864). This name is a
homonym and an unnecessary replacement name for Apion parvulum Mulsant & Rey, 1858
Apion ulicis Dufour, 1843, syn. nov. of Exapion ulicis (Forster, 1771). Based on the original description.
Perapion nuristanicola Wanat, 1995, syn. nov. of Perapion himalayense Bhateja & Pajni, 1989. Based on the
original description
82 NEW ACTS and COMMENTS
Pseudapiemonus limonii salarius Ehret, 1997, syn. nov. of Pseudaplemonus limonii (Kirby, 1808). Based on
absence of distinguishing characters in the original description

Priority
Catapion burdigalense (Wencker, 1859) is a junior synonym of C. semicyaneum (Mulsant & Rey, 1858). The
synonymy of the two names was established by Desbrochers des Loges (1898b: 35). However, the senior
name was not used as valid. Since the species is rarely mentioned, the conditions of Art. 23.9.1.2 are not
met, and I have decided to follow the Principle of Priority.
Curculio astralagi Herbst, 1795 is a senior synonym of Exapion genistae (Kirby, 1811). It is an incorrect
latinization based on Herbst's incorrect usage of the plant genus Astragalus as Astralagus. However, it has
never been used since its inception. I refrain from doing it here and a reversal of the precedence under Art.
23.9 will be done elsewhere.
Curculio aterrimus Linnaeus, 1758 is a senior synonym of Perapion marchicum (Herbst, 1797). However, it
has not been used for a valid species since mid 19th century. A formal reversal of the precedence under Art.
23.9 will be done elsewhere.

Resurrected names
Holotrichapion {Holotrichapion) umbrinum (Wollaston, 1864), stat. res., from synonymy under H. (H.) ononis
(Kirby, 1808), based on the study of type material.
Phrissotrichum {Schilskyapiori) moroderi (Desbrochers des Loges, 1907), s t a t res., from synonymy under P.
(5.) grenieri Desbrochers des Loges, 1875, based on the study of type material

Spellings
Giustiapion Legalov, 2003 (p. 67) is here selected as the valid spelling over Giustianapion on p. 68 (key), here
rejected as an incorrect alternative original spelling (First Reviser action, Art. 24.2.3).
Apion compactem pseudinexpertum Schatzmayr, 1925 (p. 38) is here selected as the valid spelling over A.
pseudoinexpertum on p. 37, here rejected as an incorrect alternative original spelling (First Reviser action,
Art. 24.2.3).
Apion galloecianum Desbrochers des Loges, 1894e: 29 is the original spelling of this nominal species, but it has
only been used in the original description. Desbrochers des Loges (1894f: 36) used the spelling
gallaecianum and this has been used as the original one since then, it is used here following Art. 33.3.1 of
the Code, attributed to the same author and date.
Apion hoffinanni Miller, 1857: 23 [conditional name, available, Art. 11.5.1] is here selected as the valid spelling
over A. hofmanni on p. 22, here rejected as an incorrect alternative original spelling since the collector's
name was Hoffmann (First Reviser's Action, Art. 24.2.3).
Apion kiienburgi Reitter, 1898 is an incorrect original spelling, being named after the Count and Countess von
Küenburg (Art. 31.1.2). Consequently, it had to be corrected to Euprotapion kueenburgorum, conserving
authorship and date (Art. 32.4) as Alonso-Zarazaga (1990b: 83) did.
Apion mountanus Pajni, Bhateja & Kumar, 1991 has as specific epithet a pseudo-Latin word, based on a
misconstructed English frame and, according to the Code, it cannot be amended. It is here considered to be
a noun in apposition and, thence, invariable.
Apion peyrimhoffi Desbrochers des Loges, 1902 is the original spelling of this nominal species, correctly
written A. peyerimhoffi in the Index to the same work (p. 190), which is 5 months later. I consider this to be
a clear evidence of an inadvertent error (Art. 32.5.1) which makes the original spelling an incorrect one.
Moreover, the corrected spelling is in use since its publication, attributed to the original reference.
Apion schoennherri Boheman, 1839 is the original spelling of this nominal species, but it has only been used in
the original description. A subsequent spelling Apion schoenherri, attributed to the same author and date, is
in current usage and is deemed here to be valid (Art. 33.3.1).
Conapium (Conapium) cupreum Bhateja & Pajni, 1989 (originally as Conapion, incorrect subsequent spelling)
is here corrected under Art. 32.5 to meet the requirements of Art. 31.2 (originally as cupreus).
Hemitrichapion tschegitunense Legalov, 2001 is here corrected under Art. 32.5 to meet the requirements of Art.
31.2 (originally as tschegitunensis).
Hemitrichapion tschernovi nanschanicum Legalov, 2001 is here corrected under Art. 32.5 to meet the
requirements of Art. 31.2 (originally as nanschanicus).
 NEW ACTS and COMMENTS 83
Hemitrichapion tschernovi terektinense Legalov, 2001 is here corrected under Art. 32.5 to meet the
requirements of Art. 31.2 (originally as terektinensis).
Hemitrichapion tschernovi tuvense Legalov, 2001 is here corrected under Art. 32.5 to meet the requirements of
Art. 31.2 (originally as tuvensis).
Hoplopodapion depressum Bhateja & Pajni, 1989 is here corrected under Art. 32.5 to meet the requirements of
Art. 31.2 (originally as Onychapion depressus).
Hoplopodapion spinitarsatum Bhateja & Pajni, 1989 is here corrected under Art. 32.5 to meet the requirements
of Art. 31.2 (originally as Onychapion spinitarsatus).
Perapion himalay ense Bhateja & Pajni, 1989 is here corrected under Art. 32.5 to meet the requirements of Art.
31.2 (originally as himalayensis).
Phrissotrichum wenckeri (C. N. F. Brisout de Barneville, 1863) is here deemed to be the specific spelling in
prevaling usage, being originally described as Apion wenkeri (Art. 33.2.3.1).

Nanophyidae
M. A. Alonso-Zarazaga
Type species designation
Sphaerula Kiesenwetter, 1864 is currently placed in synonymy with Nanophyes Schoenherr, 1838. It was
established for a group of species having fused claws. Curculio lythri Fabricius, 1787, one of the originally
included species, is here designated as the type species of Sphaerula Kiesenwetter, 1864.

Homonyms
Nanophyes nitidulus var. atrolineatus Pic, 1919d: 13 (from Spain, Algeria and Morocco) is here given
precedence over Nanophyes nitidulus var. atrolineatus Pic, 1919d: 14 (from Rumania); First Reviser's
Action (Art. 24.2).

New combinations
Allomalia gemmaria (Faust, 1887), comb. nov. from Nanophyes
Allomalia piffli (Voss, 1964), comb. nov. from Corimalia
Hypophyes exiguus (Faust, 1890), comb. nov. from Nanophyes
Hypophyes exiguus bulganensis (Zherikhin, 1976), comb. nov. from Corimalia
Nanomimus borisi (Zherikhin, 1996), comb. nov. from Nanophyes
Nanophyes triptolemus (Gistel, 1857), comb. nov. from Nanodes
Nanophyes villaticus (Gistel, 1857), comb. nov.fromNanodes

New replacement name

Nanophyes helenae Alonso-Zarazaga, nom. nov. for Nanophyes atrolineatus Pic, 1924 (non Nanophyes
nitidulus var. atrolineatus Pic, 1919). Cordially named after Mlle Hélène Perrin (MNCN, Paris).

New synonyms
Curculio salicariae Fabricius, 1781, syn. nov. of Nanophyes marmoratus (Goeze, 1777). Type in coll. Banks
seen.
Nanophyes dispersenotatus Pic, 1916, syn. nov. of Nanophyes marmoratus (Goeze, 1777). Type dissected
(Coll. Pic, MNHN, Paris). Original locality is Cuba in Portugal, not the West Indies island.
Nanophyes pectoralis Pic, 1929, syn. nov. of Corimalia latifrons (Pic, 1897). Types compared.
Nanophyes helveticus Tournier, 1868, syn. nov. of Dieckmanniellus chevrieri (Boheman, 1845). Type of the
former lost, that of the second checked.
Rhynchaenus pulicarius Gravenhorst, 1807: 204, syn. nov. of Nanophyes marmoratus (Goeze, 1777). Based on
MS synonymy by Gravenhorst.

Resurrected names
Nanophyes chevrieri Boheman, 1845, resurrected, found to be incorrectly placed in synonymy with
Dieckmanniellus nitidulus (Gyllenhal, 1838), after study of the type material. It is here considered a valid
species in the genus Dieckmanniellus.
84 NEW ACTS and COMMENTS
Spellings
Nanophyes marmoratus miguelangeli Zherikhin, 1996, p. 240, is here selected as the valid spelling over N.
marmoratus meguelangeli in the caption of fig. 105, here rejected as an incorrect alternative original
spelling (First Reviser's Action, Art. 24.2.3).

Brachyceridae
E. Colonnelli
New synonymy
Brachycerus callosus Schoenherr, 1833, unnecessary replacement name for Brachycerus variolosus Thunberg,
1799 (from South Africa) is considered synonym of Brachycerus chevrolati Fâhraeus, 1840.

Type species designations

Desmidophorus senex Boheman, 1845, one of the two species originally included in Trichosomus by Chevrolat
(1881), is here designated as the type species of Trichosomus Chevrolat, 1881. The latter name is a synonym
of Desmidophorus Dejean, 1835 and a homonym of Trichosomus Swainson, 1839.
Desmidophorus aureolus Gyllenhal, 1837 is here designated as the type species Desmidophorinus Hubenthal,
1917, originally proposed as a subgenus, and presently placed in synonymy with Desmidophorus Dejean,
1835. Desmidophorinus originally consisted of 13 species, all from me Oriental region and New Guinea.

Comments
Ocladius aegyptiacus Toumier, 1875, usually cited from Egypt, is distributed only in Erytrea and Ethiopia tiiat
were part of lower Egypt at me time of its description (Meregalli & Colonnelli 2006). Records of Üüs
species, as well as tiiose of O. barani (and of O. sharpi) by Alfieri (1976), are erroneous and refer to other
species in Meregalli & Colonnelli (2006).

Dryophthoridae
C. H. C. Lyal
New synonyms
Prosipalinus Voss, 1962, syn. nov. ofSipalinus Marshall, 1943
Diocalandra elongata form nigromaculata Voss, 1956, syn. nov. of Diocalandra elongata Roelofs, 1875
Cyrtotrachelus rufopectinipes subnotatus Voss, 1931, syn. nov. of Cyrtotrachelus rufopectinipes Chevrolat,
1883
Sphenophorus striatus maderensis Hoffmann, 1938, syn. nov. of Cosmopolites sordidus Chevrolat, 1885
Rhabdocnemis maculatus carinicollis Voss, 1937, syn. nov. of Rhabdoscelus maculatus Gyllenhal, 1838

Changes in rank and assignments

Otidognathus pygidialis Jordan, 1894, raised from subspecies of Otidognathus areolatus Fairmaire, 1900
Otidognathus pygidialis areolatus, Fairmaire, 1900, downgraded from Otidognathus areolatus Fairmaire, 1900
Otidognathus pygidialis intermedius Günmer, 1938, transferred from Otidognathus areolatus intermedins
Günther, 1938
Otidognathus rubriceps cantonensis Günther, 1938, comb. nov. and downgraded from Cyrtotrachelus
cantonensis Günther, 1938 (sensu Li-zhong, 2002: 316)
Otidognathus rubriceps notatus Voss, 1932, downgraded from Otidognathus 'notabilis' [sic] sensu Li-Zhong,
2002:316
Type species designations
Omotemnus Chevrolat, 1883: The name was established by Chevrolat, 1883: 559 for the species Calandra
serrirostris Fabricius, 1801 (incorrectly attributed to Olivier) (with varieties seriatus Fabricius, 1801 and
raumuri Gyllenhal, 1838), Omotemnus rhinoceros Chevrolat, 1883 and Omotemnus miniatocrinitus
Chevrolat, 1883; no type species was designated. Calandra serrirostris Fabricius, 1801. is here designated as
the type species.
 NEW ACTS and COMMENTS 85
Hilipomorphus Desbrochers des Loges, 1891: The name was established by Desbrochers des Loges, 1891: cclx
for the species H. cardoni Desbrochers des Loges, 1891 and H. subfasciatus Desbrochers des Loges, 1891.
No type species was designated. Hilipomorphus subfasciatus Desbrochers des Loges, 1891 (= Nassophasis
morreni Roelofs, 1885) is here designated as the type species.

Dexipeus Pascoes, 1885: The name was established by Pascoe, 1885: 311 for the new species Dexipeus costatus
and D. fumosus. No type species was designated. Dexipeus fumosus is here designated as the type species.

Erirhinidae
R. Caldara
Designations of type species
Apachiscelus Desbrochers des Loges, 1875 originally included Apachiscelus pellitus Desbrochers des Loges,
1875 and lethierryi Chevrolat, 1860. Apachiscelus pellitus Desbrochers des Loges, 1875 is hereby
designated as the type species.
Jekelia Tournier, 1874, originally included Tychius ephippiatus Fairmaire, 1870 and Jekelia depressipennis
Tournier, 1874, the latter subsequently transferred to Echinocnemus. Therefore, Tychius ephippiatus
Fairmaire, 1870 is hereby designated as the type species.
Erirhinus Stephens, 1829 [non Schoenherr, 1825] originally included Curculio festucae Herbst, 1795, C. nereis
Paykull, 1800, C. inquisitor Herbst, 1797, and Erirhinus arundineti Stephens (nomen nudum; taxon
described only in 1831). Curculio festucae Herbst, 1795 is hereby designated as the type species.

New synonymy
Curculio inquisitor Herbst, 1797 is considered synonym of Thryogenes festucae (Herbst, 1795) and not of T.
nereis (Paykull, 1800) (Caldara & Winkelman, in press)
Curculio scirpi Rossi, 1790, syn. nov. of Notaris scirpi (Fabricius, 1792), based on the description. See also
below

Priorities
Thompson (2006) considered the name Rhynchaenus steveni Krynicki, 1832 as a nomen oblitum and used
Erirhinus steveni Gyllenhal, 1835 as the valid name for Procas picipes steveni. However, although the
name steveni Krynicki meets the conditions of Art. 23.9.1.1, since it was never used as a valid name after
1899, the name steveni Gyllenhal does not meet the conditions of Art. 23.9.1.2, since it was never used in
the immediately preceding 50 years. Therefore, it is not possible to apply the reversal of precedence.
Notaris scirpi (Fabricius, 1792) is declared nomen protectum and Curculio rhamni Herbst, 1784 and Curculio
scirpi Rossi, 1790 are nomina oblita (Caldara & Winkelman, in press).

Species incertae sedis

Procas rasus Desbrochers des Loges, 1897and P. sibiricus Pic, 1904 are reported as incertae sedis by
Thompson (2006)

Erroneous assignment
Ochodontus Desbrochers des Loges, 1897: I have examined a syntype of the type species Ochodontus
rubrirostris Desbrochers des Loges, 1897. It belongs to Molytinae Acicnemidini and is presently studied by
Korotyaev & Savitski.
Tsherepanovia Korotyaev, 1992 does not belong to Erirhinidae but its placement is presently doubtful
(Korotyaev pers. com.); for pratical reasons it is left within Erirhinidae.

Doubtful assignment
The two species of Absoloniella Formânek, 1913 are known only by original description and therefore the
assignment of this genus to Erirhinidae is doubtful.
Hypoglyptus Gerstaecker, 1855 and Ruffodytes Osella, 1973 have genitalia as in Curculionidae and therefore
their assignment to Erirhinidae is doubtful.
Siraton Mustache, 1934 is known onlyfromone female and therefore its assignment to Erirhinidae is doubtful.
86 NEW ACTS and COMMENTS
Comments
Glaridorhimis Kolenati, 1858: The type species of the monotypic genus is G. khuenburgii Kolenati, 1858,
probably Neotropical (see Alonso-Zarazaga & Lyal, 1999).

Erirhinidae
M. A. Alonso-Zarazaga
Species ineertae sedis
Erirhinus indistinctus Motsehulsky, 1845 was not mentioned after its description, which is uninformative for
placing it.

Raymondionymidae
G. Osella, P. Hlavâô & M. Meregalli
New synonymy
Ferreria taurinorum (Osella, 1977), syn. nov. of Ferreria marqueti marqueti (Aube, 1863)

New combination
Ferreria doriai (Osella, 1977), comb. nov. from Raymondionymus

Curculionidae: Scolytinae
M. Knizek
New synonyms
Hylastes imitator Reitter, 1900, syn nov. of Hylurgops longipillus Reitter, 1895
Bonips Cognato, 2001, syn. nov. of Ips DeGeer, 1775
Emarips Cognato, 2001, syn. nov. of Ips DeGeer, 1775
Granips Cognato, 2001, syn. nov. of Ips DeGeer, 1775
Cyrtogenius kabei Murayama, 1955, syn. nov. of Cyrtogenius kumamotoensis Niisima, 1917
Xyleborus californiens Wood, 1975, syn. nov. of Cyclorhipidion bodoanum (Reitter, 1913)
Xyleborus quercus Kurentsov, 1948, syn. nov. of Cyclorhipidion pelliculosum (Eichhoff, 1878)
Xyleborus starki Nunberg, 1956, syn. nov. of Cyclorhipidion pelliculosum (Eichhoff, 1878)
Xyleborus khinganensis Murayama, 1943, syn. nov. of Anisandrus dispar (Fabricius, 1792)
Xyleborus alni Niisima, 1909, syn. nov. of Xyleborinus attenuatus (Blandford, 1894)
Stephanoderes myrmidon Eichhoff, 1878, syn. nov. of Hypothenemus eruditus (Westwood, 1834)

Resurrections
Cisurgus pusillus (Wollaston, 1860) from synonymy with Crypturgus wollastonii Eichhoff, 1878, proposed as
an unnecessary replaceent name because of supposed hymonymy with Crypturgus pusillus (Gyllenhal,
1813)
Coleobothrus Enderlein, 1929 from synonymy with Aphanarthrum Wollaston, 1854
Xyleborinus angustatus (Eichhoff, 1866) from synonymy with Xyleborinus saxesenii (Ratzeburg, 1837)

New combinations
Dryocoetes cornivorus (Murayama, 1950), comb. nov. from Xyleborus
Dryocoetiops dinoderoides (Blandford, 1894), comb. nov. from Taphrorychus
Dryocoetiops moestus (Blandford, 1894), comb. nov. from Taphrorychus
Cyclorhipidion pelliculosum (Eichhoff, 1878), comb. nov. from Xyleborus
Cyclorhipidion seiryorense (Murayama, 1930), comb. nov. from Xyleborus

New replacement name

Hypothenemus nobuchii Knizek, nom. nov. for Hypothenemus oblongus (Nobuchi, 1981) [nee Hypothenemus
oblongus (Niisima, 1910)]
 NEW ACTS and COMMENTS 87
Reversal of precedence
Cryphalus saltuarius Weise, 1891 is a well known, widely distributed western Palaearctic species. Its senior
synonym, Cryphalus scriba Gozis, 1886, has not been used as valid since 1899 and is here declared nomen
oblitum, while Cryphalus saltuarius Weise, 1891 is declared nomen protectum, according to ICZN Art.
23.9.1.2.
Hylurgops glabratus (Zetterstedt, 1828) is a well known, widely distributed western Palaearctic species. Its
senior synonym, Hylurgops paykullii (Duftschmid, 1825), has not been used as valid since 1899 and is here
declared nomen oblitum, while Hylurgops glabratus (Zetterstedt, 1828) is declared nomen protectum,
according to ICZN Art. 23.9.1.2.
Hylurgops palliatus (Gyllenhal, 1813) is a well known, widely distributed Palaearctic species. Its senior
synonyms, Hylurgops piceus (Marsham, 1802) and Hylurgops rufus (Marsham, 1802), have not been used as
valid since 1899 and are here declared nomina oblita, while Hylurgops palliatus (Gyllenhal, 1813) is
declared nomen protectum, according to ICZN Art. 23.9.1.2.
The use of these three names, Cryphalus saltuarius Weise, 1891, Hylurgops glabratus (Zetterstedt, 1828), and
Hylurgops palliatus (Gyllenhal, 1813), is supported by the following publications (see chapter References
for complete citation):
Alexandrovitch et al. 1996, Bovey 1987, Bright & Skidmore 1997, 2002, Burakowski et al. 1992, Heliövaara et
al. 1998, Izhevskiy et al. 2005, Knizek 2004, Krivolutskaya 1996, Kfistek & Urban 2004, Lekander 1968,
Lekanderetal. 1977, Lundberg 1995, Masutti 1995, Michalski &Mazur 1999, Nunberg 1981, Pfeffer 1989,
1995, Pfeffer & Knizek 1993, Razowski 1991, Schnaider 1991, Silfverberg 1992, Voolma et al. 2000, Wood
& Bright 1992, Yin et al. 1984, and others.

Spelling
Phloeosinus thuyae Perris, 1855 is the original spelling. All subsequent authors used the spelling "thujae"
which is used in the Catalogue. See also Bulletin zool. nomencl., Opinion 1167.
Scolytogenes darwini Eichhoff, 1878, corrected original spelling from "darvini" [subsequently spelled darwinii,
in Eichhoff, 1878].
Scolytus transcaspicus Eggers, 1922, a junior synonym of S. schevyrewi Semenov, 1902, was originaly
misspelled "transcapicus".

Priority
Kissophagus hederae (Schmitt, 1843) is a name currently used for a well known, widely distributed western
Palaearctic species. However, its senior synonym, Kissophagus vicinus (Comolli, 1837), has also been used
as valid after 1899 and cannot be declared nomen oblitum (ICZN Art. 23.9.1.1). Therefore, Kissophagus
hederae (Schmitt, 1843) is here placed in synonymy, and K vicinus (Comolli, 1837) used as valid.

Rejected names
Blastophagus Eichhoff, 1864: 25 with type species Dermestes piniperda Linnaeus, 1758, and Myelophilus
Eichhoff, 1878c: 400 with type species Dermestes piniperda Linnaeus, 1758, are currently placed as
synonyms of Tomicus Latreille, 1802. They were rejected in Opinion 670.

Comments
Xyleborus volvulus (Fabricius, 1794) was subsequently cited as described in Bostrichus by Fabricius, 1775:
454, but his work from 1775 does not provide the description of that species. Bostrichus volvulus is
described by Fabricius, 1794: 454, in the Appendix specierum nuper detectarum. Fabricius described
previously Apate volvulus (Fabricius 1792: 363). The description of the latter species doesn't fit the
description of Bostrichus volvulus Fabricius, 1794, and its identity remains unknown.

Curculionidae: Scolytinae
M. A. Alonso-Zarazaga
Nomina dubia (not referred to in currently used publications)
Bostrichus carinatus Waltl, 1839: 223 E: GE
Bostrichus monocerus Gravenhorst, 1807: 193 E: GE
88 NEW ACTS and COMMENTS
Bostrichuspilosus Gistel, 1831: 306 E: GE
Bostrichas pumilio Gravenhorst, 1807: 193 "Patria ignota"
Ips ater Marsham, 1802: 59 E: GB
Ips inermis Marsham, 1802: 54 E: GB
Ips sulcatus Marsham, 1802: 59 E: GB [= Hylesinus crenatus?]
Scolytuspusillus Olivier, 1795: (78) 14 E: FR
Scofytus varius Olivier, 1795: (78) 11 E: FR

Curculionidae: Baridinae
J. Prena
Type species designation
Lissoharis Faust, 1881 included four species-group names and neither was designated as the type species.
Because the name had been used already in the Carabidae by Chaudoir (1838), Reitter (1913) replaced it
with Elasmobaris. Korotyaev (2002) designated Baridius alboguttatus H. Brisout de Bameville, 1870a as the
type species for Lissotarsus Faust, but this action is invalid because the name originally was not included in
Lissoharis. Lissoharis balassogloi Faust, 1881, currently considered a synonym of B. alboguttatus, is
designated here as the type species of Lissoharis Faust; it automatically becomes the type species of
Elasmobaris.

Changes in rank
Turkmenobaris Zaslavskij, 1956, as genus (based on published morphological and ecological evidence)

New combinations and other assignments

Anthinobaris Morimoto & Yoshihara, 1996, from incertae sedis to Baridini, Baridina
Arganthonius Alonso-Zarazaga, 2005, from incertae sedis to near Athesapeuta in Madarini, Leptoschoinina
Batobaris Morimoto & Yoshihara, 1996, from incertae sedis to Baridini, Baridina
Caenobaris Nasreddinov, 1980, from incertae sedis to near Eumycterus in Neosharpiini
Cucubaris Alonso-Zarazaga, 2005, from incertae sedis to near Acythopeus in Madarini, Leptoschoinina
Formosobaris Morimoto & Yoshihara, 1996, from incertae sedis to Baridini, Baridina
Maurobaris Alonso-Zarazaga, 2005, from incertae sedis to Baridini, Baridina
Moreobaris Morimoto & Yoshihara, 1996, from incertae sedis to Baridini, Baridina
Nespilobaris Morimoto & Yoshihara, 1996, from incertae sedis to Baridini, Baridina
Pharcidobaris Morimoto & Yoshihara, 1996, from incertae sedis to Baridini, Baridina
Pteridobaris Morimoto & Yoshihara, 1996, from incertae sedis to Baridini, Baridina
Taiwanobaris Morimoto & Yoshihara, 1996, from incertae sedis to Baridini, Baridina
Aulacobaris atlasica (Hustache, 1923), comb, nov., from Baris (based on description)
Aulacobaris berthieri (Pic, 1901), comb, nov., from Baris (based on description)
Aulacobaris concinna (Boheman, 1844), comb, nov., from Baris (type material studied)
Aulacobaris deflexa (Desbrochers des Loges, 1900), comb, nov., from Baris (based on description)
Aulacobaris devillei (F. Solari, 1952), comb, nov., from Baris (based on description)
Aulacobaris edoughensis (Desbrochers des Loges, 1892), comb, nov., from Baris (based on description)
Aulacobaris fallaciosa (Hoffmann, 1960), comb, nov., from Baris (based on description)
Aulacobaris holdereri (Hartmann, 1900), comb, nov., from Baris (based on description)
Aulacobaris kaszabi (Korotyaev, 1994), comb, nov., from Baris (based on description)
Aulacobaris kiesenwetteri (Faust, 1891), comb, nov., from Baris (type material studied)
Aulacobaris licens (Reitter, 1895), comb, nov., from Baris (based on description)
Aulacobaris mecinoides (Desbrochers des Loges, 1892), comb, nov., from Baris (based on description)
Aulacobaris nobilis (Desbrochers des Loges, 1896), comb, nov., from Baris (type material studied)
Aulacobaris ochsi (Hoffmann, 1950), comb, nov., from Baris (based on description)
Aulacobaris perdura (Reitter, 1895), comb, nov., from Baris (type material studied)
Aulacobaris raisae (Korotyaev, 1987), comb, nov., from Baris (type material studied)
Aulacobaris ruteri (Ferragu, 1962), comb, nov., from Baris (based on description)
Aulacobaris sareptana (Desbrochers des Loges, 1900), comb, nov., from Baris (based on description)
 NEW ACTS and COMMENTS 89
Aulacobaris violaceomicans (A. Solari & F. Solari, 1905), comb, nov., from Baris (type material studied)
Cosmobaris discolor (Boheman, 1836), comb, nov., from Baris (type material studied)
Labiaticola araxicola (Reitter, 1895), comb, nov., from Baris (type material studied)
Labiaticola cariniventris (A. Solari & F. Solari, 1909), comb, nov., from Baris (based on description)
Labiaticola crassipes (Faust, 1884), comb, nov., from Baris (type material studied)
Labiaticola rugulipennis (Reitter, 1895), comb, nov., from Baris (type material studied)
Labiaticola teheranicus (Hoffmann, 1956), comb, nov., from Baris (based on description)
Orchidophilus hisamatsui (Kojima & Yoshihara, 2009), comb, nov., from Omobaris (based on description)

New synonyms
Baridius kirschi Desbrochers des Loges, 1892, syn. nov. of Baris kirschi Faust, 1882
Baridius soricinus Fairmaire, 1884, syn. nov. of Cosmobaris discolor (Boheman, 1836)
Baridius squamulatus Pic, 1896, syn. nov. of Cosmobaris discolor (Boheman, 1836)
Baris alboseriata Reitter, 1908, syn. nov. of Cosmobaris discolor (Boheman, 1836)
Baris borkhsenii Zaslavskij, 1956, syn. nov. of Cosmobaris scolopacea (Germar, 1819)
Baris limbata nigra Reitter, 1895, syn. nov. of Baridius limbatus H. Brisout de BameviUe, 1870 and Baridius
limbatus niger Desbrochers des Loges, 1892
Baris lorata Marshall, 1911, syn. nov. of Aegyptobaris arctithorax (Pic, 1899)
Baris orientalis Roelofs, 1875, syn. nov. of Cosmobaris scolopacea (Germar, 1819)
Baris rhagodiae Rheinheimer, 1992, syn. nov. of Cosmobaris discolor (Boheman, 1836)
Baris vicina H. Brisout de BameviUe 1870, syn. nov. of Cosmobaris discolor (Boheman, 1836)
Baris violacea Grimmer, 1841, syn. nov. of Aulacobaris picicornis (Marsham, 1802)
Curculio maculatus Townson, 1797, syn. nov. of Baris analis (Olivier, 1791)

Alternative original spellings

Baridius suffriani Hochhuth, 1847: 566 [121 in reprint] is selected here as the correct original spelling and
Baridius suffrianii Hochhuth, 1847: 586 [141 in reprint] is suppressed as the incorrect original spelling.
90
CATALOGUE

superfamily CURCULIONOIDEA Latreille, 1802

family NEMONYCHIDAE Bedel, 1882
subfamily Cimberidinae Gozis, 1882
tribe Cimberidini Gozis, 1882
genus Cimberis Gozis, 1881a: 150 type species Rhinomacer attelaboides Fabricius, 1787
subgenus Cimberis Gozis, 1881a: 150 type species Rhinomacer attelaboides Fabricius, 1787
attelaboides Fabricius, 1787a: 123 {Rhinomacer) E: AU BU CT DE FI FR GB GE HU IT LA LT LU NL NR
NT PL SP ST SV SZ N: AG A: ES FE KZ NC TR WS
canescens Semenov, 1900: 129
rhinomacer Thunberg, 1791: 115 (Curculio)
rhinomacer Paykull, 1792: 126 (Curculio) [HN]

tribe Doydirhynchini Pierce, 1916

genus Doydirhynchus Dejean, 1821: 79 type species Rhynchites austhacus Olivier, 1807
Diodyrhynchus Schoenherr, 1833: 240 [unjustified emendation]
Doedycorhynchus Labram & Imhoff, 1843: no. 4 [unnecessary RN]
Doedycorrhynchus Bedel, 1883: 18 [unnecessary RN]
austriacus Olivier, 1807: 27 {Rhynchites) E: AU BH CR CZ FR GE GR HU IT LA LU NL NR NT PL PT SK
SP SV SZ UK A: TR
castaneus Germar, 1833: 242
cilicicus K. Daniel & J. Daniel, 1903: 327
karamani Stierlin, 1886a: 229
lutesc ens Schi I sky, 1903: 92
mêlas Herbst, 1786: 166 (Curculio)
pallidicolor Pic, 1905e: 190
bicolor Pic, 1905d: 137 N: AG MO TU

subfamily Nemonychinae Bedel, 1882

genus Nemonyx L. Redtenbacher, 1845: 96 type species Rhinomacer lepturoides Fabricius, 1801
Nematonyx Agassiz, 1846: 247 [unjustified emendation]
canescens Solsky, 1881: 263 A: KZ TM UZ
lepturoides Fabricius, 1801: 429 (Rhinomacer) E: AB AR AU BU CT CZ FR GE GR HU IT KZ MD PL RO
SK ST SZ UK A: LE TR
grisescens Reitter, 1899b: 209
scutellatus Abeille de Perrin, 1901: 234 N: TU
semirufus Pic, 1898c: 74 N: AG
variicolor Abeille de Perrin, 1898: 258

family Nemonychidae, nomina dubia

Rhinomacer gibbus O. F. Müller, 1776: 90 E: DE
Rhinomacer nemoralis O. F. Müller, 1776: 90 E: DE
Rhinomacerpolitus O. F. Müller, 1776: 90 E: DE
Rhinomacer similis O. F. Müller, 1776: 90 E: DE

family ANTHRIBIDAE Billberg, 1820

subfamily Anthribinae Billberg, 1820
tribe Allandrini Pierce, 1930
genus AUandrus LeConte, 1876: 396 type species Allandrus bifasciatus LeConte, 1876
Tropiderinus Reitter, 1916a: 5 type species Tropideres munieri Bedel, 1882
 ANTHRIBIDAE 91
fiiscipennis Guillebeau, 1891: 199 {Tropideres) E: CZ FR PL SK UK
rufipennis Rey, 1893a: 49 {Tropideres)
interrupt™ Reitter, 1889d: 286 {Tropideres) E: AB AR A: IN IS JO TM
iriei Morimoto, 1980: 18 A: ES FE JA
munieri Bedel, 1882: 8 {Tropideres) E: GR SP N: AG
therondi Tempère, 1954: 48 {Enedreytes) E: FR
undulatus Panzer, 1795: 294 {Anthribus) E: AU BE CZ DE EN FI FR GE HU IT LU NL NR NT PL SK SP SV
SZ N: "N Africa"
cinctus Paykull, 1792: 123 {Curculio) [HN]
edgreni Schoenherr, 1839: 211 {Tropideres)

genus Phaeochrotes Pascoe, 1860b: 42 type species Phaeochrotes porcellus Pascoe, 1860
porcellus Pascoe, 1860b: 42 A: YUN ORR
pudens Gyllenhal, 1833: 153 {Tropideres) E: AU BE BH BU CR CZ DE FR GE GR HU IT LU PL RO SK ST
SZTRUKYU A:FEMGNPSYTR
inornatus Bach, 1854: 168 {Tropideres)

genus Plintheria Pascoe, 1859b: 435 type species Plintheria luctuosa Pascoe, 1859
Alloderes Jordan, 1944: 141 type species Alloderes vitiensis Jordan, 1944
caliginosa Shibata, 1980b: 39 A: JA (Ogasawara)
diversa Shibata, 1979: 97 A: TAI
sparsa Boheman, 1833: 151 {Tropideres) A: HAI ORR
variolosa Shibata, 1969: 26 A: JA

tribe Anthribini Billberg, 1820

genus Anthribus Geoffroy, 1762: 306 type species Anthribus fasciatus Forster, 1770
Anthotribus J. J. Hoffmann, 1803: 110 [unjustified emendation]
Antribus Illiger, 1801: 127 [unjustified emendation]
Antriptus Illiger, 1801: 127 [unjustified emendation]
Brachytarsus Schoenherr, 1823: 1135 type species Anthribus varius Fabricius, 1787 (= Anthribus nebulosus Forster, 1770)
Paropes Schoenherr, 1823: 1135 type species Anthribus varius Fabricius, 1787 {^Anthribus nebulosus Forster, 1770)
Paropus Fischer von Waldheim, 1829: 84 type species Anthribus varius Fabricius, 1787 (= Anthribus nebulosus Forster,
1770)
Pseudobrachytarsus Pierce, 1930: 29 type species Anthribus fasciatus Forster, 1770
fasciatus Forster, 1770: 5 E: AL AU BE BH BU CR CZ DE EN FI FR GB GE GR HU IT LA LU NL PL PT RO
SK SP SV SZ TR UK YU N: AG MO NARi
marmoratus Geoffroy, 1785: 136
rubripes Rey, 1893a: 49 {Brachytarsus)
rufipes Motschulsky, 1860: 153 {Brachytarsus)
scabrosus Fabricius, 1775: 64 {Bruchus)
ventralis Rey, 1893a: 49 {Brachytarsus)
kuwanai Yuasa, 1931: 21 {Brachytarsus) A: CH ES FE JA SC
vandykei Jordan, 1933b: 383
nebulosus Forster, 1770: 5 E: AU BE BH BU BY CR CZ DE EN FI FR GB GR HU IT LA LT LU NL NR NT
PL PT RO SK SP SV SZ UK YU A: FE NARi
capsularius Schrank, 1789: 70 {Bruchus)
clathratus Herbst, 1786: 158 {Bruchus)
gavoyi Chobaut, 1922: 88
küsteri Reitter, 1916a: 9 {Brachytarsus)
nebulosus Küster, 1859: {Brachytarsus) [HN]
variegatus Geoffroy, 1785: 136
varius Fabricius, 1787: 42 {Bruchus)
niveovariegatus Roelofs, 1879b: lv {Brachytarsus) A: BEI FE GUI HUN JA JIA SC SCH YUN XIN
lajievorus Chao, 1976: 339 {Brachytarsus)
nigrovariegatus Roelofs, 1880a: 29 {Brachytarsus)
scapularis Gebier, 1833: 291 E: BU BY CZ DE EN FI GE LA LT NR NT PL SV TR N: AG
constrictus Stierlin, 1867: 224 {Brachytarsus)
92 ANTHRIBIDAE
gebleri Reitter, 1916a: 9 (Brachytarsus)
roseus Reitter, 1916a: 9 (Brachytarsus)
subroseus Reitter, 1916a: 9 E: FR GR '
clopi Chobaut, 1922:87

tribe Basitropidini Lacordaire, 1865

genus Basitropis Jekel, 1855: 90 type species Basitropis nitidicutis Jekel, 1855
Baseotropis Gemminger & Harold, 1872: 2742 [unjustified emendation]
affinis Jordan, 1903b: 432 A: HAI ORR
bimaculata Senoh & Matoba, 2004: 229 A: JA (Ryukyus)
lutosa lutosa Jordan, 1895a: 194 A: HAI ORR
nitidicutis nitidicutis Jekel, 1855: 92 A: FE HAI HKG JA NP TAI UP ORR
brevis Jordan, 1897: 642
dispar Sharp, 1891:320
orientalis Motschulsky, 1875: 231 (Eugomts)
persimilis Jordan, 1916:363 A: HAI TAI ORR
seinoi Morimoto, 1978a: 43 A: JA (Ogasawara)

genus Gynandrocerus Lacordaire, 1865: 568 type species Gynandrocerus antennalis Lacordaire, 1865
praecox Kolbe, 1895: 390 N: MO (S. of Casablanca) AFR

tribe Corrhecerini Lacordaire, 1865

genus Aulodina Jordan, 1903a: 128 type species Aulodina unicolor Jordan, 1903
alternata Frieser, 1977: 809 A: "Arabia" AFR

genus Cylindroides Fairmaire, 1886: 455 type species Cylindroides alboplagiatus Fairmaire, 1886
Aulodes Quedenfeldt, 1887: 321 type species Aulodes bifasciatus Quedenfeldt, 1887
arabicus Frieser, 1981c: 437 A: SA
bifasciatus Quedenfeldt, 1887: 323 {Aulodes) A: IS SA

genus Habrissus Pascoe, 1859b: 432 type species Habrissus pilicornis Pascoe, 1859
Habrocerastes Lacordaire, 1865: 548 type species Habrocerastes sellifer Lacordaire, 1865
analis analis Morimoto, 1981: 98 A: JA SC SCH
anal is chinensis Frieser, 1995a: 16 A: SCH
cylindricus Sharp, 1891: 311 (Tropideres) A: FE (Sakhalin) JA
formosanus Jordan, 1912a: 143 A: ES JA (Ryukyus) TAI
kejvali Frieser, 1995a: 17 A: SCH
longipes Sharp, 1891: 312 {Tropideres) A: FE JA TAI
nigronotatus Morimoto, 1981: 100 A: JA (Ryukyus)
pardalis Sharp, 1891: 313 (Tropideres) A: FE (Kurdes, Sakhalin) JA
unciferoides Nakane, 1963b: 31 (Hypseus) A: JA

genus Penestica Pascoe, 1859a: 332 type species Penestica inepta Pascoe, 1859
brevis albescens Shibata, 1963a: 47 (Paraphloeobius) A: JA
brevis brevis Jordan, 1912a: 145 (Paraphloeobius) A: TAI

tribe Cratoparini LeConte, 1876

genus Euparius Schoenherr, 1823: 1135 type species Anthribus lunatus Fabricius, 1801 (= Macrocephalus
marmoreus Olivier, 1795)
Caccorhinus Sharp, 1891: 321 type species Caccorhinus oculatus Sharp, 1891
Cratoparis Dejean, 1834: 235 [unnecessary RN]
boninensis Morimoto, 1978a: 38 A: JA (Ogasawara)
concolor Morimoto, 1978a: 36 A: JA (Okinawa, Amami, Ryukyus)
koltzei Reitter, 1895c: 159 (Cratoparis) A: FE
koreanus Park & Morimoto, 1999: 97 A: SC
 ANTHRIBIDAE 93
lateralis patruelis Frieser, 2001: 27 A: NP
modicus modicus Jordan, 1933b: 382 A: JA (Ryukyus) TAI ORR
modicus yaeyamanus Morimoto, 1978a: 36 A: JA (Ryukyus) TAI
oculatus formosanus Jordan, 1912a: 144 A: FUJ JA (Ryukyus) TAI
oculatus oculatus Sharp, 1891: 321 (Caccorhinus) A: FE JA SC SHA TAI
towu/Nakane, 1963b: 31 A: JA

tribe Discotenini Lacordaire, 1865

genus Autotroph Jordan, 1924a: 241 type species Autotropis modesta Jordan, 1924
basipennis Sharp, 1891: 310 (Tropideres) A: ES FE JA SC
distinguenda Sharp, 1891: 310 {Tropideres) A: ES FE JA SC TAI
fitlvescens Shibata, 1984a: 33 A: TAI
modesta conspersa Jordan, 1928b: 93 A: TAI ORR
modesta rugulosa Jordan, 1924a: 242 A: TAI ORR
montana montana Wolfrum, 1948: 141 A: FE FUJ NP TAI YUN ORR
montana sibirica Frieser, 1981a: 54 A: FE
montana taiwana Frieser, 1981a: 54 A: TAI
okinawana Morimoto, 1981: 57 A: JA (Okinawa)

genus Enedreytes Schoenherr, 1839: 215 type species Enedreytes hilaris Fâhraeus, 1839
Enedreutes Agassiz, 1846: 138 [unjustified emendation]
Enedreutes Lacordaire, 1865: 536 [HN, unjustified emendation]
Enedreutes Gemminger & Harold, 1872: 2735 [HN, unjustified emendation]
gotoi Shibata, 1969: 28 A: FE JA SC
hilaris Fâhraeus, 1839: 216 E: CR FR GE GR HU IT SP TR N: AG MO A: TR

genus Pseudeuparius Jordan, 1914: 344 type species Cratoparis monoceros Fâhraeus, 1839
centromaculatus Gyllenhal, 1833: 143 (Euparius) E: GR IT TR YU N: AG MO A: IS TR
ceroderes Boheman, 1845: 352 (Cratoparis)
targionii Piccioli, 1877: 214 (Cratoparis)
sepicola Fabricius, 1792: 378 (Anthribus) E: AU BE BU CR CZ DE FR GB GE GG HU IT LU NL PL PT SK
SP ST SV SZ TR UK YU
ephippium Boheman, 1833: 152 (Tropideres)
fascirostris Clairville, 1798: 116 (Platyrhinus)
combraliensis Gozis, 1882: 69 (Tropideres)

tribe Ecelonerini Lacordaire, 1865

gams Dendropemon Schoenherr, 1839: 161 type species Anthribus perfolicornis Fabricius, 1801
Dendrotrogus Jekel, 1855: 82 type species Dendrotrogus hypocrita Jekel, 1855
Ecelonerus Schoenherr, 1839: 163 type species Ecelonerus subfasciatus Fâhraeus, 1839
Icelonirus Agassiz, 1846: 193 [unnecessary RN]
Icelonirus Gemminger & Harold, 1872: 2742 [HN, unjustified emendation]
Xenotropis Fairmaire, 1895: cclxxxi type species Xenotropis rugicollis Fairmaire, 1895 (= Dendrotrogus colligens
Walker, 1859)
angustipennis Jordan, 1895a: 191 (Dendrotrogus) A: HAI TAI ORR
colligens Walker, 1859a: 261 (Dendrotrogus) A: HAI TAI ORR
colligendus Bovie, 1906: 290 (Dendrotrogus)
rugicollis Fairmaire, 1895: 282 (Xenotropis)
japonicus Morimoto, 1979a: 15 (Dendrotrogus) A: JA TAI
nagaoi Morimoto, 1979a: 17 (Dendrotrogus) A: JA
ohkurai Shibata, 1978c: 97 (Dendrotrogus) A: JA (Ogasawara)
perfolicornis Fabricius, 1801:407 (Anthribus) A: HAI YUN ORR

genus Disphaerona Jordan, 1902: 77 type species Disphaeronapunctata Jordan, 1902

chinensis Frieser, 1995a: 15 A: SCH
94 ANTHRIBIDAE
genus Eucorynus Schoenherr, 1823:1135 type species Anthribus crassicornis Fabricius, 1801
crassicornis Fabricius, 1801: 407 {Anthribus) A: ES FE FUJ GUX HAI HKG JA NC NP PA SC SHA TAI UP
YUN AFR ORR
clavator Fairmaire, 1903:43
colligendus Walker, 1859a: 261
setosulus Pascoe, 1859b: 434

tribe Jordanthribini Morimoto, 1978

genus Jordanthribus Zimmerman, 1938: 236 type species Jordanthribus planifacietus Zimmerman, 1938
Japanthribus Shibata, 1978c: 98 type species Japanthribus kitsuii Shibata, 1978
kusuii Shibata, 1978c: 100 {Japanthribus) A: JA (Ogasawara)

tribe Mauiini Valentine, 1989

genus Cleorisintor Jordan, 1923a: 90 type species Cleorisintor glaucus Jordan, 1923
glaucus Jordan, 1923a: 91 A: GUA HAI ORR

genus Mauia Blackburn, 1885: 194 type species Mauia satelles Blackburn, 1885 (= Araeocerus subnotatus
Boheman, 1859)
Contexta Jordan, 1902: 78 type species Contexta murina Jordan, 1902 (= Araeocerus subnotatus Boheman, 1859)
subnotata Boheman, 1859: 116 {Araeocerus) A:CHJANP ORR AFR AUR
murina Jordan, 1902: 78 {Contexta)
satelles Blackburn, 1885: 195

genus Protaedus Pascoe, 1860b: 39 type species Protaedus moerens Pascoe, 1860
Telphes Wolfram, 1924: 35 type species Telphes mannulus Wolfram, 1924
brevicornis Senoh, 1986b: 708 A: JA (Ryukyus)

tribe Mecocerini Lacordaire, 1865

genus Acanthothorax Gaede, 1832: Classe IX, pi. 15 type species Acanthothorax longicornis Gaede, 1832
Mecocerus Schoenherr, 1833: 115 type species Mecocerus gazella Gyllenhal, 1833 (= Acanthothorax longicornis Gaede
1832)
allectus indochinensis Jordan, 1916: 360 {Mecocerus) A: YUN ORR
asmenus asmenus Jordan, 1913a: 204 {Mecocerus) A: CH "NE India" ORR
callosus Jordan, 1904b: 231 {Mecocerus) A: YUN ORR

tribe Mycteini Morimoto, 1972

genus Mycteis Pascoe, 1860b: 44 type species Mycteis marginicollis Pascoe, 1860
foveolatus Wolfram, 1948: 139 A: FUJ

genus Sympaector Kirsch, 1875: 52 type species Sympaector vittifrons Kirsch, 1875
rugirostris Sharp, 1891: 302 {Tropideres) A: FE JA SC
ussuriensis Egorov, 1996b: 184 A: FE

tribe Ozotomerini Morimoto, 1972

genus Ozotomerus Perroud, 1853: 406 type species Ozotomerus maculosus Perroud, 1853
Dipieza Pascoe, 1859a: 331 type species Dipieza waterhousei Pascoe, 1859 (= Oedecerus bipunctatus Montrouzier, 1855)
Oedecerus Montrouzier, 1855: 46 type species Oedecerus bipunctatus Montrouzier, 1855
amamianus Morimoto, 1978a: 40 A: JA (Ryukyus) TAI
deletus Frieser, 1995a: 30 A: YUN ORR
japonicus japonicus Sharp, 1891: 320 A: JA TAI
japonicus laferi Egorov, 1986: 15 A: ANH FE SC SHA
nigromaculatus Morimoto, 1978a: 41 A: JA TAI
patruelis Frieser, 1995a: 31 A: YUN ORR
 ANTHRIBIDAE 95

tribe Phloeophilini Lacordaire, 1865

genus Bothrus Wolfrum, 1956:112 type species Bothrus stirophorus Wolfrum, 1956
taiwanus taiwanus Shibata, 1978a: 23 (Physopterus) A: TAI

genus Morimotanthribus Senoh & Tryzna, 2006: 397 type species Morimotanthribus chinensis Senoh &
Tryzna, 2006
chinensis Senoh & Tryzna, 2006: 399 A: HUB

genus Phloeophilus Schoenherr, 1833: 156 type species Phloeophilus agrestis Boheman, 1833
Cappadox Alonso-Zarazaga & Lyal, 1999: 29 type species Phloeophilus sulcifrons Fâhraeus, 1839
Lemmatophilus Rye, 1874: 299 [RN]
Lemmophilus Rye, 1872: 86 [unnecessary RN]
Physopterus Lacordaire, 1865: 498 type species Mecocerus gibbosus Guérin-Méneville, 1843
Platynorhynchus Schoenherr, 1839: 194 type species Phloeophilus sulcifrons Fâhraeus, 1839
agrestis Boheman, 1833: 157 A: NP SD UP
mirabilis Frieser, 1996c: 29 (Physopterus) A: JIX YUN
oculatus Jordan, 1904b: 231 A: HUN ORR

tribe Piesocorynini Valentine, 1960

genus Botriessa Jordan, 1928c: 127 type species Botriessa sepidiopsis Jordan, 1928
cylindricollis Frieser, 1999: 30 A: NP

genus Phaenotheriolum Ganglbauer, 1903b: 215 type species Phaenotherion steindachneri Frivaldszky, 1883
africanum Vauloger, 1901: 233 (Phaenotherion) N: AG
espagnoli Gonzalez, 1969: 47 (Phaenotherium) E: SP
hispidum Frieser, 1978: 296 E: TR
steindachneri Frivaldszky, 1883: 36 (Phaenotherion) E: TR A: IS LE
villosum Frieser, 1978: 296 E: TR

genus Phaenotherion Frivaldszky, 1877: 330 type species Phaenotherion pulszkyi Frivaldszky, 1877
Phaenotherium Ganglbauer, 1903 b: 215 [unjustified emendation]
bodemeyeri Wolfrum, 1956: 110 E: TR
fasciculatum fasciculatum Reitter, 1891c: 248 E: IT
fasciculatum kuchtae Frieser, 1955: 451 E: CR
ganglbaueri Apfelbeck, 1918: 100 (Phaenotherium) E: ALGR
graecum Frieser, 1995b: 37 E: GR
knirschi Breit, 1912:151 E: UK (Krim)
pulszkyi Frivaldszky, 1877: 331 E: HU RO ST
innotatum Pic, 1908a: 43
strejceki Frieser, 1995b: 38 E: GG ST
zellichi Ganglbauer, 1903b: 217 E: CR YU

genus Phaenotheriosoma Frieser, 1978: 297 type species Phaenotheriosoma besucheti Frieser, 1978
adductum Frieser, 1980: 76 E: TR
besucheti Frieser, 1978: 297 A: LE
brachati Frieser, 1995b: 39 E: TR
complanatum Frieser, 1980: 76 E: TR
latifolium Frieser, 1980: 77 E: TR
loebli Frieser, 1978: 298 A: IS LE
pselaphoides Frieser, 1980: 76 E: TR
simplicorne Frieser, 1995b: 40 E: TR
96 ANTHRIBIDAE
tribe Platyrhinini Imhoff, 1856
genus Apatenia Pascoe, 1859b: 434 type species Apatenia viduata Pascoe, 1859
infcms Jordan, 1912a: 141 A: TAI

genus Aphaulimia Morimoto, 1972: 35 type species Tropideres debilis Sharp, 1891
debilis Sharp, 1891: 311 {Tropideres) A: ES FE JA SC
grammica grammica Jordan, 1912a: 142 {Phaulimia) A: JA TAI

genus Baseocolpus Jordan, 1949: 6 type species Baseocolpus clivosus Jordan, 1949
abundans Frieser, 1997: 39 {Pantorhaenas) A: NP SD
foveatus Frieser, 1999: 30 A: YUN

genus Hypseus Pascoe, 1860b: 39 type species Hypseus fascicularis Pascoe, 1860
angulatus Shibata, 1980a: 26 {Phaulimid) A: TAI
areifer Frieser, 1985: 89 {Apatenia) A: TAI
tuberosus Senoh, 1995: 1 {Phaulimid) A: JA (Ryukyus)

genus Nessiara Pascoe, 1860a: 60 type species Nessia didyma Pascoe, 1859
Nessia Pascoe, 1859a: 329 [HN] type species Nessia didyma Pascoe, 1859
Straboscopus Lacordaire, 1865: 533 type species Straboscopus riehlü Lacordaire, 1865
mosonica Jordan, 1904b: 234 A: TAI ORR

genus Oxyderes Jordan, 1928c: 114 type species Hypseus frenatus Jordan, 1897
fastigatus Jordan, 1924a: 236 {Apatenia) A: ES FE HUB JA SC TAI
obsolescens Wolfrum, 1948: 138 A: FUJ

genus Phaulimia Pascoe, 1859b: 437 type species Phaulimia ephippiata Pascoe, 1859
aberrans Sharp, 1891: 309 A: FE JA
angusta Morimoto, 1981: 83 A: JA (Ryukyus)
annulipes Morimoto, 1981: 86 A: JA (Ogasawara)
confmis Sharp, 1891: 309 {Tropideres) A: ES FE JA SC
decorata Shibata, 1963b: 1 {Hypseus) A: JA (Izu, Ryukyus)
incerta Shibata, 1963b: 2 {Hypseus) A: JA (Ryukyus) TAI
kurosawai Senoh, 1986a: 317 A: JA (Ryukyus)
maculithorax Senoh, 1986a: 314 A: JA (Ryukyus)
minor Shibata, 1980b: 43 A: JA (Ogasawara)
rufobasis Morimoto, 1981: 88 A: FE JA
vicina Shibata, 1980a: 27 A: TAI

genus Platyrhinus Clairville, 1798: 112 type species Platyrhinus costirostris Clairville, 1798 (= Curculio
resinosus Scopoli, 1763)
Anthhbus Fabricius, 1790: 219 [HN] type species Curculio latirostris Fabricius, 1775 (= Curculio resinosus Scopoli
1763)
resinosus Scopoli, 1763: 24 {Curculio) E: AB AL AU BE BH BU CR CT CZ DE EN FI FR GB GE GR HU IT
LA LT LU MC MD NL NR NT PL RO SK SL SP SV SZ TR UK YU "Caucasus" N: AG MO A: IN IS
ater Geoffroy, 1785: 137 (Anthribus)
costirostris Clairville, 1798: 114
flavifrons Fuessly, 1775: 6 {Silpha)
latirostris Fabricius, 1775: 151 {Curculio)
oblongus Sulzer, 1776: 28 {Silpha)
striatus O. F. Müller, 1776: 85 {Crioceris)
striatus O. F. Müller, 1776: 86 {Curculio) [HN]
terlineatus Gmelin, 1790: 1803 {Curculio) [unjustified RN]
 ANTHRIBIDAE 97
genus Ulorhinus Sharp, 1891: 300 type species UlorhinusfiinebrisSharp, 1891
akitai Senoh, 1989: 235 A: JA (Tsushima)
bilineatus Germar, 1819: 119 (Anthribus) E: AU BH BU CR CZ HU IT MC RO SK SL ST SZ TR UK YU
bisignatus Fàhraeus, 1839: 213 (Tropideres)
fiinebris Sharp, 1891: 301 A: FE (Kuriles) JA
gokani Morimoto, 1981: 91 A: FE (Kuriles) JA
saitoi Senoh, 2003:285 A: JA

tribe Platystomini Pierce, 1916

genus Caenophloeobius Morimoto, 1979a: 5 type species Phloeomimus inconspicuus Wolfrum, 1948
inconspicuus Wolfrum, 1948: 146 (Phloeomimus) A: FUJ JA TAI
planatus Shibata, 1963a: 46 (Phloeobius)

genus Phloeobius Schoenherr, 1823: 1135 type species Anthribus griseus Fabricius, 1792 (= Ptinus gigas
Fabricius, 1775)
Branconymus A. Hoffmann, 1959: 341 type species Branconymus vayssierei A. Hoffmann, 1959 (= Phloeobius
hypoxanthus Jordan, 1911)
albescens Jordan, 1895a: 198 A: HAI TAI ORR
alternons Wiedemann, 1819: 172 (Anthribus) A: CH J A TAI ORR
apicalis Walker, 1859a: 262
gibbosus Roelofs, 1879b: lv [= 1880a: 26] A: GAN JA SCH SHA SHX TAI ORR
gigas nigroungulatus Gyllenhal, 1833: 133 (Anthribus) [HN] A: GUI GUX HKG TAI YUN ORR
laetus Jordan, 1923a: 103 A: SD YUN ORR
lepticornis Jordan, 1912b: 141 A: AP SCH TAI XIZ ORR
mimes Sharp, 1891: 319 A: JA
stenus Jordan, 1923b: 173 A: JA SHG
stenoides Shibata, 1963a: 45
triarrhenus R.-Z. Zhang,Wang & Shuai, 1994: 9 A: HUN
genus Platystomos D. H. Schneider, 1791: 21 type species Curculio albinus Linnaeus, 1758
Anthotribus Gistel, 1856: 375 [HN]
Anthrodus Megerle, 1826: 32 type species Anthribus albinus sensu auct.
Macrocephalus Olivier, 1789: 36 [HN] type species Curculio albinus Linnaeus, 1758
Platystomos Hellwig, 1792: 393 [HN] type species Curculio albinus Linnaeus, 1758
albinus Linnaeus, 1758: 385 (Curculio) E: AL AU BE BH BU BY CR CZ DE EN FI FR GB GE GG HU IT LA
LT LU MC NL NR NT PL RO SK SL SP ST SV SZ UK YU A: ES FE HEB IN KZ SC WS
desertus Schilsky, 1907: 79 (Platystomus)
thierriati Viturat, 1895: 110 (Anthribus)
uniformis Reitter, 1916a: 7 (Platystomus)
albisignatus Senoh, 1996: 164 A: YUN ORR
macrocerus macrocerus Jordan, 1904b: 235 (Anthribus) A: SD ORR
sellatus asteromaculatus Oda, 1978: 27 A: JA (Ryukyus)
sellatus longicrus Park, Hong, Woo & Kwon, 2001: 182 A: HUB SC SHA
sellatus sellatus Roelofs, 1879a: 302 (Anthribus) A: FE (Kuriles) JA TAI
daimio Sharp, 1891: 319 (Anthribus)
genus Platystomos, species incertae sedis
undatus Fabricius, 1787a: 41 (Bruchus) N: "Africa"
tribe Ptychoderini Jekel, 1855
genus Mecotropis Lacordaire, 1865: 495 type species Mecotropis bipunctata Lacordaire, 1865
cylindrica Jordan, 1904b: 230 A: HAI ORR
hieroglyphica Hope, 1831: 26 (Macrocephalus) A: NP ORR
kyushuensis Nakane, 1963b: 31 (Mecocerus) A: HUN JA
ogasawarai Morimoto, 1979a: 9 A: JA (Ryukyus)
punctulifera Frieser, 2005: 5 A: JIX ORR
98 ANTHRIBIDAE
reginalis Frieser, 2001: 5 A: BT
taiwana Shibata, 1982: 25 A: TAI
unoi Shibata, 1969: 10 A: JA (Ryukyus) TAI

genus Phloeopemon Schoenherr, 1839: 159 type species Anthribus acuticornis Fabricius, 1801
continentalis Jordan, 1923b: 167 A: HAIYUN ORR

tribe Sintorini Lacordaire, 1865

genus Sintor Schoenherr, 1839:148 type species Sintor quadrilineatus Fâhraeus, 1839
Blabirhimis Sharp, 1891: 299 type species Blabirhinus dorsalis Sharp, 1891
Rhinanthribus Motschulsky, 1875: 241 type species Rhinanthribus dispar Motschulsky, 1875
biplaga Jordan, 1903b: 416 A: SD YUN ORR
bipunctatus Shibata, 1969: 14 A: JA (Yakushima, Ryukyus)
dorsalis dorsalis Sharp, 1891: 300 {Blabirhinus) A: FUJ JA SC
dorsalis intermedius Shibata, 1969: 14 A: JA (Tsushima) TAI
fasciatus Jordan, 1903b: 417 A: TAI ORR
floridus Jordan, 1931:296 A: "NE India" ORR
Jukiensisfiikiensis Wolfrum, 1948: 134 A: FUJ GUX
melanipus Jordan, 1923a: 89 A: HAI SCH ORR
pustulatus Shibata, 1978a: 25 A: TAI
sulcifrons Frieser, 1996c: 31 A: NP
suturalis Jordan, 1895c: 374 A: NP ORR
yamawakii Morimoto, 1980: 13 A: JA (Ryukyus)

tribe Tophoderini Lacordaire, 1865

genus Asemorhinus Sharp, 1891: 298 type species Asemorhinus nebulosus Sharp, 1891
nebulosus indochinensis Jordan, 1923a: 91 A: HAI ORR
nebulosus nebulosus Sharp, 1891: 299 A: JA TAI
nebulosus nepalicus Frieser, 1989: 29 A: NP

genus Derisemias Jordan, 1928b: 86 type species Derisemias picticollis Jordan, 1928
coronatus Frieser, 1989: 30 A: UP
taiwanus Frieser, 1981b: 208 A: TAI

genus Dolichocera Gray, 1832: 65 type species Dolichocera childreni Gray, 1832
Eugigas J. Thomson, 1858a: 434 [unnecessary RN]
Meganthribus Jordan, 1913b: 263 [unnecessary RN]
harmandi tessellata Jordan, 1895c: 369 {Eugigas) A: CH SD ORR

genus Rawasia Roelofs, 1880b: 203 type species Rawasia ritsemae Roelofs, 1880
annulipes Jordan, 1895a: 188 A:NPYUN ORR
ritsemae Roelofs, 1880b: 204 A: FUJ J A (Ryukyus) TAI ORR
flavescens Nakane, 1963b: 31 {Eucorynus)

tribe Trigonorhinini Valentine, 1999

genus Opanthribus Schilsky, 1907: Z type species Anthribus tessellatus Boheman, 1829
Kimenus Wolfrum, 1961: 319 type species Kimenus submetallicus Wolfrum, 1961
Paramesus Fâhraeus, 1871: 443 [HN] type species Paramesus lituratus Fâhraeus, 1871
tessellatus Boheman, 1829: 119 {Anthribus) E: AU CR CZ FR GE HU IT LU PL SK SP SZ UK N- MO TU A:
ES FE JA SC SCH SHA
brunneipennis Reitter, 1916a: 8
fallax Perris, 1874: 13 {Brachytarsus)
 ANTHRIBIDAE 99
genus Tngonorhinus Wollaston, 1861a: 102 type species Trigonorhinus pardalis Wollaston, 1861
(=Brachytarsus areolatus Boheman, 1845)
Anthribulus LeConte, 1876:405,406 type species Anthribulus rotundatus LeConte, 1876
Brachylarsoides Pierce, 1930: 29, 30 type species Brachytar, sus griseus LeConte, 1876
Trigonorrhinus Gemminger & Harold, 1872: 2745 [unjustified emendation]
areolatus Boheman, 1845: 344 (Brachytarsus) E: IT (Sicilia) SP N: AG TU
baguenai Cobos, 1954:42 (Tropideres)
pantherinus P. H. Lucas, 1847:406 (Brachytarsus)
pardalis Wollaston, 1861a: 102
dolgovi Korotyaev, 1977: 373 {Brachytarsus) A: ES FE MG
zeae Wolfrum, 1931: 75 {Brachytarsus) N: MR A: JA NTR
japonic us Morimoto, 1999: 61
lindbergi A. Hoffmann, 1963a: 128 (Tropideres)
trimaculatus Senoh, 1986b: 706 (Opanthribus)

tribe Tropiderini Lacordaire, 1865

genus Acorynus Schoenherr, 1833: 123 type species Acorynus sulcirostris Boheman, 1833
amaurus Wolfrum, 1948: 135 A: FUJ
anchis anchis Jordan, 1912a: 137 A: SCH TAI YUN
apical is Jordan, 1894b: 623 A: SD
apicinotus Wolfrum, 1948: 136 A: FUJ SHA
asanoi asanoi Nakane, 1963b: 33 {Tropideres) A: JA
asanoi mikuraensis Nakane, 1978: 80 A: JA (Mikura)
latirostris Sharp, 1891: 303 (Tropideres) A: FE JA
luzonicus lanhsuanus Shibata, 1982: 26 A: TAI
okinawanus Morimoto, 1980: 30 A: JA (Ryukyus)
poecilus Shibata, 1963b: 4 A: JA (Ryukyus)
similis Jordan, 1894b: 620 A: SD
singularis Shibata, 1963b: 4 A: JA (Ryukyus)

genus Androceras Jordan, 1928b: 83 type species Mucronianus khasianus Jordan, 1903
flabellicorne Sharp, 1891: 305 {Tropideres) A: FE HUB JA SC SHA
gerrhus Jordan, 1923c: 217 {Mucronianus) A: NP ORR
laticorne Jordan, 1928b: 84 A: YUN ORR
pulcherrimum Jordan, 1932: 310 A: SD NP

genus Cedus Pascoe, 1860b: 37 type species Cedus tuberculatus Pascoe, 1860
Byastus Pascoe, 1860b: 38 type species Byastus cephalotes Pascoe, 1860
Pachygenia Motschulsky, 1875: 233 type species Pachygenia impluviata Motschulsky, 1875
amabilis Shibata, 1979: 100 A: TAI
diversus Jordan, 1911:111 A: TAI
insignis Shibata, 1980b: 41 A: JA (Ogasawara)
japonicus Shibata, 1963a: 43 A: JA (Ryukyus)
prominens Shibata, 1979: 98 A: TAI

genus Eurymycter LeConte, 1876: 394 type species Macrocephalus fasciatus Olivier, 1795
lacteocaudatus Fairmaire, 1889: 57 {Tropideres) A: GAN SCH SHA

genus Gonotropis LeConte, 1876: 393 type species Gonotropis gibbosa LeConte, 1876
Agonotropis Egorov, 1987: 105 type species Agonotropis terminassianae Egorov, 1987
Basarukinia Egorov, 1996b: 190 type species Gonotropis insignis Morimoto, 1980
crassicornis Sharp, 1891: 306 {Tropideres) A: FE JA
dorsalis Gyllenhal, 1813: 5 {Anthribus) E: AU BY CZ DE EN FI FR GE IT LA LT NR NT PL SV A:
"Sibirien" NAR
gibbosa LeConte, 1876: 394 E: CZ GE HU SK SV A: SC NAR
100 ANTHRIBIDAE
ins ignis Morimoto, 1980: 36 A: FE JA
murakamii Oda, 1979: 111 A: JA
terminassianae Egorov, 1987: 107 (Agonotropis) A: FE

genus Hucus Pascoe, 1859b: 436 type species Hucus melanostoma Pascoe, 1859
hiatus Jordan, 1936a: 314 A:NP ORR

genus Litocerus Schoenherr, 1833: 125 type species Litocerus histrio Gyllenhal, 1833
adelphus Wolfrum, 1948: 137 A: FUJ
ambustus Wolfrum, 1948: 136 A: FUJ
bicuspis Jordan, 1912a: 138 A: TAI
communis amamianus Shibata, 1969: 16 A: JA (Ryukyus)
communis communis Jordan, 1912a: 139 A: JA (Ryukyus) TAI
dysallus Jordan, 1912a: 138 A: TAI
histrio histrio Gyllenhal, 1833: 126 A: TAI ORR
kimurai Shibata, 1969: 19 A: JA
multiguttatus Nakane, 1963b: 33 (Tropideres) A: JA
propinquus Wolfrum, 1948: 138 A: FUJ
sticticus Jordan, 1904b: 233 A: TAI
striatifrons Senoh, 1985a: 75 A: JA (Ogasawara)
tokarensis insensibilis Shibata, 1969: 18 A: JA (Yaku) ORR
tokarensis ogasawaranus Shibata, 1980b: 41 A: JA (Ogasawara)
tokarensis tokarensis Nakane, 1963b: 33 (Tropideres) A: JA (Tokara)
tokarensis yoshimii Shibata, 1982:28 A: TAI
tuberculatus Shibata, 1982: 27 A: TAI
vallestris Frieser, 2001: 11 A: NP

genus Merarius Fairmaire, 1889: 56 type species Merarius davidis Fairmaire, 1889
davidis Fairmaire, 1889: 56 A: XIZ YUN ORR
holzschuhi Frieser, 2001: 12 A: BT
quadrituberculatus Hope, 1831:27 (Anthribus) A: NP SD
glabriceps Jordan, 1911: 106

genus Mucronianus Jordan, 1894b: 627 type species Mucronianus ruflpes Jordan, 1894
takemurai Nakane, 1963b: 32 (Litocerus) A: JA

genus Nessiodocus Heller, 1925: 106 type species Nessiodocus jordani Heller, 1925
antennalis Morimoto, 1980: 23 A: JA
confluens Frieser, 2001: 12 A:NP
propinquus Shibata, 1980a: 24 A: JA (Ryukyus) TAI
répandus répandus Jordan, 1912a: 141 (Mucronianus) A: FE JA TAI
latefasciatus Nakane, 1963b: 33 (Tropideres)
trifasciatus Shibata, 1980a: 23 A: TAI
triodes Jordan, 1912a: 140 (Mucronianus) A: FE HAI JA TAI

genus Sphinctotropis Kolbe, 1895: 379 type species Sphinctotropis albofasciata Kolbe, 1895
Spathorrhamphus T. A. Marshall, 1902: 210 type species Spathorrhamphus corsicus T. A. Marshall, 1902
Corsica T. A. Marshall, 1902: 211 (Spathorrhamphus) E: FR (Corse)
laxa Sharp, 1891: 304 (Tropideres) A: ES FE FUJ GUA GUI GUX HUB JA JIX SC SHA
notabilis Jordan, 1928b: 82 (Tropideres) A: YUN ORR
paviei Lesne, 1891: xci (Litocerus) A: NP TAI UP ORR
scabrosa Frieser, 1983: 51 A: YUN
verrucosa Karsch, 1882: 388 (Tropideres) A: BT NP SCH ORR
 ANTHRIBIDAE 101
genus Tropideres Schoenherr, 1823: 1135 type species Curculio albirostris Herbst, 1784 (= Curculio
albirostris Schaller, 1783)
Tropidoderes Gistei, 1856: 375 type species Curculio albirostris Herbst, 1784 (= Curculio albirostris Schaller, 1783)
Tropidoderes Gemminger & Harold, 1872: 2733 [HN, unjustified emendation]
albirostris Schaller, 1783: 287 {Curculio) E: AU BE BH BU BY CR CZ DE FR GE IT LU NL PL PT RO SK
SP ST SV SZ UK N: MO A: KI UZ "Sibiria"
albirostris Herbst, 1784a: 80 (Curculio) [HN]
planirostris Piller & Mitterpacher, 1783: 84 (Curculio)
quercinus Reitter, 1916a: 5
cyaneotergum Oda, 1979: 115 A: FE JA SC
instructus Frieser, 1983: 51 A: NP
insularis Shibata, 1963b: 7 A: JA (Ryukyus)
japonicusjaponicus Roelofs, 1879b: lv [= 1880a: 27] (Litocerus) A: FE JA TAI ORR
japonicus lutulentus Frieser, 1996a: 20 A: GUT JTX
japonicus odai Morimoto, 1980: 39 A: JA TAI
laevicollis Frieser, 1996a: 19 A: YUN
lateralis Motschulsky, 1875: 235 A: NP UP YUN "E India" ORR
laenatus Jordan, 1938: 140 (Acorynus)
luteago fuscatus Frieser, 1996a: 21 A: YUN ORR
luteago luteago Jordan, 1930: 143 A: NP "N India" ORR
marmoreus marmoreus Jordan, 1894b: 628 A: CH "NE India" ORR
naevulus Faust, 1887b: 162 A: ES FE GAN HEB JA NMO SC SHA
germanus Sharp, 1891: 304
vi/is Sharp, 1891:305
yezoenzis Oda, 1979: 121
nanus Shibata, 1982: 29 A: TAI
rectirostris Frieser, 1996a: 19 A: NP
roelofsi carens Wolfirum, 1948: 140 A: FUJ GUI SHA ORR
roelofsi poecilus Jordan, 1912a: 139 A: TAI
roelofsi roelofsi Lewis, 1879: 465 (Litocerus) A: JA
scitus Jordan, 1933b: 372 A: "NE India" ORR
securus Boheman, 1839: 207 A: FE FUJ GUA HAI HEI JA JTX NP SC TAI ORR
arma Jordan, 1903 b: 424 (Litocerus)
rufescens Roelofs, 1879b: lv [= 1880a: 28] (Litocerus)
signellus divisus Frieser, 2002b: 48 A: AP SHA ORR
signellus signellus Jordan, 1912a: 140 A:FUJJIXTAI ORR

tribe Xenocerini Lacordaire, 1865

genus Peribathys Jordan, 1937b: 333 type species Xenocerus everetti Jordan, 1894
klapperichi Wolfrum, 1948: 143 A: FUJ
okinawanus Senoh, 1984a: 33 A: JA (Okinawa)
shinonagai Senoh, 1982: 63 A: JA (Ryukyus)
uenoi Senoh, 1982: 67 A: JA TAI

genus Xenocerus Schoenherr, 1833:117 type species Xenocerus saperdoides Gyllenhal, 1833
khasianus dives Jordan, 1923a: 100 A:GUXYUN ORR

tribe Xylinadini Lacordaire, 1865

genus Stiboderes Jordan, 1925: 243 type species Xylinades chevrolati Ritsema, 1883
impressus impressus Jordan, 1895b: 253 (Xylinades) A: SD ORR
impressus stibinus Jordan, 1912a: 143 A: JA TAI

genus Xylinada Berthold, 1827:380 type species Anthribus nodicornis Weber, 1801
Dasycorynus Lacordaire, 1865: 560 type species Dasycorynus riehlii Lacordaire, 1865
Xylinades Latreille, 1828: 590 type species Xylinades westermanni Gyllenhal, 1833
102 ANTHRIBIDAE
annulipes Jordan, 1895b: 255 (Xylinades) A: AP JA TAI XIZ ORR
aspericollis Jordan, 1895b: 258 (Xylinades) A: YUN ORR
continua Wolfrum, 1948: 145 (Xylinades) A: FUJ SCH YUN
japonica chinensis Wolfram, 1948: 144 (Xylinades) A: FUJ
japonicajaponica Sharp, 1891: 317 (Xylinades) A: JA TAI
oshimai Morimoto, 1979a: 12 A: JA (Ryukyus)
parumsignata Jordan, 1932: 306 (Xylinades) A: BT ORR
phycas Jordan, 1912a: 144 (Xylinades) A: TAI ORR
plagiata Jordan, 1895b: 257 (Xylinades) A: HAI NP YUN ORR
rugiceps Jordan, 1895b: 259 (Xylinades) A: AP YUN ORR
striatifrons Jordan, 1895b: 260 (Xylinades) A: JA TAI ORR

tribe Zygaenodini Lacordaire, 1865

genus Atinellia Jordan, 1925: 252 type species Atinellia senex Jordan, 1925
acuticollis Wolfram, 1948: 142 A: FUJ HEN HUB SCH YUN
affinis Shibata, 1979: 103 A: TAI
signata Shibata, 1979: 105 A: TAI
tessellata Shibata, 1979: 104 A: TAI
variegata Shibata, 1979: 102 A: TAI

genus Dissoleucas Jordan, 1925: 252 type species Anthribus niveirostris Fabricius, 1798
maenamii Senoh, 1986b: 710 A: JA
niveirostris Fabricius, 1798: 160 (Anthribus) E: AR AU BE BH BU BY CR CZ DE EN FI FR GB GE HU IT
LA LT LU NL NR NT PL RO SK SP SV SZ TR UK YU
dubius Ponza, 1805: 65 (Curculio) [HN]

genus Epiplaterus Jordan, 1949: 7 type species Epiplaterus modicus Jordan, 1949
contaminatus Frieser, 1999: 27 A: JIA ORR

genus Eusphyrus LeConte, 1876: 399 type species Eusphyrus walshii LeConte, 1876
Opisthotropis Hoffmann & Tempere, 1954: 66 [HN] type specie» Tropideres vasconicus Hoffmann & Tempère, 1954
vasconicus Hoffmann & Tempère, 1954: 66 (Tropideres) E: FR IT SP

genus Exechesops Schoenherr, 1847: 4 type species Exechesops quadrituberculatus Schoenherr, 1847
Zygaenodes Pascoe, 1859a: 328 type species Zygaenodes wollastoni Pascoe, 1859
acaulus Wolfrum, 1953: 315 A: SCH
alces Frieser, 1995a: 22 A: NP
antiallus Jordan, 1912c: 602 (Zygaenodes) A: NP ORR
becvari Frieser, 1995a: 27 A: SCH SHA YUN
discoidalis Wolfrum, 1922: 58 (Zygaenodes) A: TAJ
foliatus Frieser, 1995a: 28 A: CH FE SC
elenae Egorov, 1996b: 194
fornicatus Frieser, 1995a: 22 A: NP SD
frontalis Wolfrum, 1922: 59 (Zygaenodes) A: TAI
holzschuhi Frieser, 1995a: 21 A: NP ORR
leucopis Jordan, 1928b: 91 (Zygaenodes) A: CH FE JA SC ORR
signatellus Frieser, 1995a: 23 A: NP
sobrinus Frieser, 1995a: 24 A: NP ORR
validus Frieser, 1998: 6 A: NP

genus ExUlis Pascoe, 1860b: 43 type species Exillis longicornis Pascoe 1860
japonicola Nakane, 1963b: 32 A: HAI JA TAI
longicornis Pascoe, 1860b: 43 A: TAI ORR
 ANTHRIBIDAE 103
genus Gibber Jordan, 1895a: 176 type species Gibber tuberculatus Jordan, 1895
bicornis Frieser, 1995a: 18 A: NP "India" ORR
brevirostris Sharp, 1891: 307 (Tropideres) A: ES FE JA
callistus Jordan, 1924a: 238 A: NP ORR
incisus Sharp, 1891: 308 (Tropideres) A: FE JA TAI
invenus tus Frieser, 1991: 63 A: NP ORR
minor Wolfrum, 1948: 139 A:FUJTAI ORR
nodulosus Sharp, 1891: 308 (Tropideres) A: FE JA NC SC
ogasawarensis Morimoto, 1981: 79 A: JA (Ogasawara)
tuberculatus Jordan, 1895a: 177 A: TAI ORR

genus Ulis Jordan, 1936a: 319 type species Ulis rusia Jordan, 1936
arma Shibata, 1969: 24 A: JA (Ishigaki, Izu, Tanegashima, Yakushima, Ryukyus) TAI
elisa Shibata, 1980a: 21 A: TAI

genus Nerthomma Pascoe, 1860b: 42 type species Nerthomma sticticum Pascoe, 1860
aplotum Jordan, 1912a: 142 A: JA (Ryukyus) TAI ORR

genus Noxius Jordan, 1936b: 328 type species Blaberus fallax Fâhraeus, 1839
Blaberus Schoenherr, 1839: 248 [HN] type species Blaberus fallax Fâhraeus, 1839
curtirostris Mulsant & Rey, 1861a: 338 (Tropideres) E: AU BU CR CZ FR GE GR HU IT SK SP SZ TR UK
YU N: AG MO TU A: CY IN IS TR YE
japonicus Morimoto, 1981: 54 A: JA
montanus Frieser, 1996b: 50 A: NP UP

genus Rhaphitropis Reitter, 1916a: 5 type species Anthribus marchicus Herbst, 1797
angustula Frieser, 1995a: 20 A: NP UP
antennata Wolfrum, 1948: 141 A: FUJ
carbo Jordan, 1936a: 316 A: NP ORR
carinata Shibata, 1978b: 94 A: TAI
caroli Frieser, 1997: 46 A: NP
communis Shibata, 1978b: 93 A: JA SCH TAI YUN
cosmia Jordan, 1937a: 200 A: "N India" ORR
cylindrica Shibata, 1978b: 97 A: TAI
discus Jordan, 1936a: 315 A: NP UP ORR
elusa Jordan, 1928b: 93 A: JIX ORR
flavipes Morimoto, 1981: 76 A: JA (Ryukyus)
guttifer guttifer Sharp, 1891: 314 (Tropideres) A: FE TAI "Korea"
concolor Sharp, 1891: 314 (Tropideres)
guttifer taiwana Shibata, 1978b: 99 A: TAI
imperfecta Sharp, 1891: 316 (Tropideres) A: JA
incana Joràsa, 1932: 311 A: NP ORR
indica indica Jordan, 1925: 248 A: "NW India" ORR
japonica Shibata, 1978c: 95 A: ES FE JA
malefida Frieser, 1997: 47 A: SD ORR
manca Frieser, 1996c: 33 [HN]
marchica Herbst, 1797: 167 (Anthribus) E: AR AU BE BH BU CZ DE FR GE GG HU IT LA LU NL NR PL
SK SP SZ UK YU
cincta Bach, 1854: 168 (Tropideres) [HN]
maculosa Mulsant, 1858: 259 (Tropideres)
reyi Gozis, 1882: 67 (Tropideres)
scripta Thunberg, 1815: 122 (Amblycerus)
midori Shibata, 1978c: 93 A: JA
iriei Morimoto, 1981: 74
nigra Morimoto, 1981: 73 A: JA
 104 ANTHRIBIDAE
mgromaculata Morimoto, 1981: 71 A: JA (Ryukyus) SC
oxyacanthae C. N. F. Brisout de Bameville, 1863: 95 (Enedreutes) E: AU BU FR GE HU IT SK SP SZ
perspicua Frieser, 1996b: 48 A: NP
pigmaea Shibata, 1978b: 96 A: TAI
satoiSenoh, 2007: 222 A: JA
stevensi Jordan, 1925: 250 A: NP SD
truncatoides Morimoto, 1981: 72 A: ES FE JA SC

genus Streneoderma Blackburn, 1900b: 157 type species Streneodermaplanatum Blackburn, 1900
insigne Wolfrum, 1934: 6 A: NP TAI ORR

genus Uncifer Jordan, 1904a: 88 type species Uncifer sticticus Jordan, 1904
akashii Morimoto, 1981: 60 A: FE (Sakhalin) JA
angulatus Park & Morimoto, 1999: 100 A: SC
bruchoides Sharp, 1891: 316 (Tropideres) A: JA
difficilis Sharp, 1891:317 (Tropideres) A: ES FE JA SC
discrepans Shibata, 1981: 23 A: JA (Ryukyus)
fenestratus Frieser, 2001: 17 A:NP
hispidus Shibata, 1981: 26 A: JA (Yakushima, Ryukyus)
monticulatus Frieser, 2001: 20 A: NP
myodes Jordan, 1937a: 201 A: "N India"
pectoralis Sharp, 1891: 315 {Tropideres) A: ES FE JA
pissodes Jordan, 1933a: 3 A: "N India"
sakoi Shibata, 1981: 24 A: JA (Tsushima, Yakushima)
stigma Frieser, 2001: 16 A: NP
stigmosus Jordan, 1936a: 319 A: "N India"
triangulus Park, Hong, Woo & Kwon, 2001: 193 A: SC
truncatus Sharp, 1891: 315 (Tropideres) A: FE (Sakhalin) JA SC

genus Unciferina Morimoto, 1981: 65 type species Unciferina japonica Morimoto, 1981
brevifrons Morimoto, 1981: 67 A: JA
japonica Morimoto, 1981: 66 A: JA
persimilis Morimoto, 1981: 68 A: JA (Ryukyus)

subfamily Apolectinae Lacordaire, 1865

tribe Apolectini Lacordaire, 1865
genus Apolecta Pascoe, 1859b: 431 type species Mecocerus parvulus J. Thomson, 1858
lewisi Sharp, 1891: 318 A: FE JA
nodicornis Frieser, 1996a: 25 A: YUN
paraplesia Jordan, 1912a: 145 A: TAI
tonkiniana Jordan, 1904b: 236 A: TAI ORR

subfamily Choraginae Kirby, 1819

tribe Araecerini Lacordaire, 1865
genus Araecerus Schoenherr, 1823: 1135 type species Anthribus coffeae Fabricius, 1801 (= Curculio
fasciculatus DeGeer, 1775)
Araeocerus Schoenherr, 1839: 273 [HN, emendation]
Arrhaecerus Germar, 1829: 357 type species Anthribus coffeae Fabricius, 1801 (= Curculio fasciculatus DeGeer, 1775)
fasciculatus DeGeer, 1775: 276 (Curculio) E: AU BE CZ DE FI FR GB GE GR HU IT LU NL NR PL SP SV
SZ TR A: ANH FUJ GAN GUA GUI GUX HEB HEN HKG HUB HUN IS JA JIA JIX LIA NMO QIN SC
SCH SHA SHN TAI TR YUN ZHE COS
alternons Germar, 1824: \15 (Anthribus)
cacao Fabricius, 1775: 64 (Bruchus)
capsinicola Fabricius, 1798: 159 (Bruchus)
 ANTHRIBIDAE 105
coffeae Fabricius, 1801: 411 (Anthribus)
griseus Stephens, 1831: 211 (Phloeobita) ;
japonicus Thunberg, 1815: 122 (Amblycerus)
mateui Cobos, 1954:41 (Tropideres)
parvirostris J. Thomson, 1858b: 113 (Cratoparis) [DA]
peregrinus Herbst, 1797: 16S (Bruckus)
seminarius Chevrolat, 1871: 7
ishigakiensis Morimoto, 1978a: 27 A: JA (Ryukyus)
levipennis Jordan, 1924a: 248 A: CH JA (Ogasawara, Ryukyus) TAI ORR
mutabilis Wolfrum, 1948: 147 (Misthosima) A: FUJ ORR
quadridens Tricser, 2001:29 A: NP ORR
siliquae Frieser, 1981c: 438 A: TAI
suturalis Boheman, 1839: 273 A: NP ORR
tarsalis Sharp, 1891: 323 A: HUB JA SC SHA TAI ORR
varions Jordan, 1946: 520 A: JA (Ogasawara) NAR

genus Deropygus Sharp, 1891: 326 type species Deropygus histrio Sharp, 1891
cavus Shibata, 1986: 49 A: JA (Ryukyus)
grammicus Senoh, 1985b: 70 A: TAI
histrio Sharp, 1891: 326 A: JA SC TAI
maedai Shibata, 1986: 50 A: JA
morimotoi Senoh, 1984b: 27 A: JA (Ryukyus)
niger Senoh, 1985b: 72 A: TAI
nipponicus Senoh, 1984b: 29 A: JA
uedai Shibata, 1986: 53 A: JA

genus Doticus Pascoe, 1882: 27 type species Doticuspalmaris Pascoe, 1882

alternatus Jordan, 1895c: 400 A: YUN ORR
alternons Jordan, 1904a: 90 (Araecerus)

genus Megatermis Jordan, 1937c: 338 type species Megatermis mameti Jordan, 1937
weigeli Frieser, 1998: 12 A: NP (Kathmandu)

genus Misthosima Pascoe, 1859b: 434 type species Misthosima mera Pascoe, 1859
brevitarsis Senoh, 1990: 261 A: JA (Ryukyus)
shoi Senoh, 1990:263 A: JA

genus Valenfriesia Alonso-Zarazaga & Lyal, 1999: 38 [RN] type species Notioxenus bewickii Wollaston, 1861
Notioxenus Wollaston, 1861b: 212 [HN] type species Notioxenus bewickii Wollaston, 1861
deropygoides Senoh, 1984d: 576 (Notioxenus) A: TAI
nakanei Morimoto, 1978a: 19 (Notioxenus) A: JA (Ogasawara)
nitida Morimoto, 1978a: 20 (Notioxenus) A: JA (Ryukyus)
tomicoides Sharp, 1891: 328 (Notioxenus) A: JA
wollastoni Sharp, 1891: 327 (Notioxenus) A: FE (Kuriles) JA

genus Xanthoderopygus Senoh, 1984b: 31 type species Deropygus flavicollis Morimoto, 1978
didymus Jordan, 1924a: 251 (Deropygus) A: JA (Ryukyus) TAI ORR
flavicollis Morimoto, 1978a: 25 (Deropygus) A: JA (Ryukyus)
jocosus Sharp, 1891: 327 (Deropygus) A: JA TAI
kasaharai Senoh & Matoba, 2002: 359 A: JA (Honshu)
matobai Senoh, 1984b: 37 A: JA
ogasawarensis Senoh, 1984b: 34 A: JA (Ogasawara)
watanabei Senoh, 1984b: 32 A: JA (Tsushima) SC
106 ANTHRIBIDAE

tribe Choragini Kirby, 1819

genus Choragus Kirby, 1819: 447 type species Choragus sheppardi Kirby, 1819
Alticopus A. Villa & G. B. Villa, 1833: 35 type species Alticopus galeazzii A. Villa & G. B. Villa, 1833 (= Choragus
sheppardi Kirby, 1819)
Halticopus Agassiz, 1846: 171 [unjustified emendation]
anobioides Sharp, 1891: 325 A: FE (Kuriles, Sakhalin) JA
aurolineatus Abeille de Perrin, 1893: 73 E: FR IT N: AG TU
brevior Frieser, 1981c: 443 A: TAI
cissoides Sharp, 1891: 325 A: FE (Kuriles) JA
compactes Sharp, 1891: 323 A: JA SC
cryphaloides Sharp, 1891: 325 A: FE (Kuriles, Sakhalin) JA SC
cryptocephalus Sharp, 1891: 324 A: FE (Kuriles, Sakhalin) JA
dentipes Frieser, 1981c: 440 A: NP SD
distendens Frieser, 1981c: 442 A: TAI
horni Wolfrum, 1930: 89 E: AU CZ DE GE GG HU LA NR PL SK ST SV
konishii Shibata, 1987: 49 A: JA
malus Johraku, 1953: 9, 12 A: JA
monticulatus Frieser, 2001: 46 A: NP
mundulus Sharp, 1891: 324 A: JA
nepalicus Frieser, 2001:46 A: NP
quadridens Frieser, 1981c: 441 A: BT
rogei Frieser, 2002a: 43 E: FR (Aude)
sheppardi Kirby, 1819: 448 E: AU BE BU CR CZ DE FI FR GB GE GG HU IR IT LU NL NT PL RO SK SP
SV SZ TR UK
bostrichoides P. W. J. Müller, 1821: 188 (Anthribus)
caucasicus Motschulsky, 1875: 229 (Alticopus)
galeazzii A. Villa & G. B. Villa, 1833: 35 (Alticopus)
niger Stephens, 1830: 343
nitidipennis Gerhardt, 1901: 158
pygmaeus Robert, 1832: pi. 16 (Anthribus)
subsulcatus Rey, 1893a: 49
subarmatus Frieser, 1981c: 442 A: SD
theryi Abeille de Perrin, 1893: 73 A: SP N: AG TU
velutinus Johraku, 1953: 7, 11 A: JA
vittatus Reitter, 1885a: 276 E: TR "S Caucasus"

genus Citacalus Johraku, 1953:13 type species Citacaluspygidialis Johraku, 1953

pygidialis Johraku, 1953: 13 A: JA

genus Melanopsacus Jordan, 1924b: 608 type species Melanopsacus fortis Jordan, 1924
anthracinus Jordan, 1924b: 612 A: BTNP SD
bhutanensis Frieser, 2001: 41 A: BT
elongatus Senoh, 1984c: 82 A: JA
gagates Jordan, 1924b: 613 A: SD
grenieri C. N. F. Brisout de Barneville, 1867: 185 (Choragus) E: BU FI FR HU IT SK SP N: AG MO
inaequalis Shibata, 1984b: 31 A: JA (Ryukyus)
kinke Morimoto, 1978a: 33 A: JA (Ryukyus) TAI
makiharai Morimoto, 1978a: 32 A: JA (Ryukyus)
minutus Jordan, 1924b: 613 A: SD
punctatus Senoh, 1984c: 80 A: JA
subfascial Jordan, 1924b: 614 A: SD ORR

genus Pseudochoragus Petri, 1912: x type species Pseudochoragus brachycerus Petri, 1912 (= Choragus
piceus Schaum, 1845)
piceus Schaum, 1845: 88 (Choragus) E: AU CZ DE FR GE HU PL SK SP SV SZ TR
 ANTHRIBIDAE 107
brachycents Petri, 1912: x

tribe Cisanthribini Zimmerman, 1994

genus Cisanthribus Zimmerman, 1938: 247 type species Cisanthribus convexus Zimmerman, 1938
nakcmei Morimoto, 1986: 273 A: JA (Ryukyus)

tribe Stenorhinini Valentine, tribe nov. [see New Acts]

genus Stenorhis Jordan, 1928a: 167 type species Stenorhis ampedus Jordan, 1928
bifoveolata Shibata, 1984a: 33 A: TAI
cylindrata Frieser, 1981c: 443 A: TAI
hirashimai Morimoto, 1978a: 22 A: JA (Ryukyus)

tribe Xenorchestini Lacordaire, 1865

genus Glabrinotropis Frieser, 1981b: 215 type species Glabrinotropis chinensis Frieser, 1981
chinemis Frieser, 1981b: 215 A: HKG

genus Xenorchestes Wollaston, 1854: 417 type species Xenorchestes saltitans Wollaston, 1854
saltitans Wollaston, 1854: 418 N: MR

subfamily Urodontinae C. G. Thomson, 1859

genus Bruchela Dejean, 1821: 78 type species Bruchus rufipes Olivier, 1790
Arodum Gistel, 1856: 375 type species Bruchus rufipes Olivier, 1790
Exurodon Schilsky, 1912: 96 type species Urodon arisi Schilsky, 1912
Urodon Schoenherr, 1823: 1134 type species Bruchus rufipes Olivier, 1790
albescens Reitter, 1897d: 245 (Urodon) A: TM UZ
albida Küster, 1848a: 82 (Urodon) E: SP PT N: MO
albosuturata Reitter, 1897d: 245 (Urodon) E: AB
anatolica Pic, 1904b: 40 (Urodon) E: AL BU GR IT RO ST "Yugoslavia" A: TR
angelovi Strejcek & Cocev, 1994: 150 (Urodon) E: BU
arisi Schilsky, 1912: 97 (Urodon) A: KZ TM
austrina Peyerimhoff, 1949: 295 (Urodon) N: MO
baudii baudii Desbrochers des Loges, 1893c: 14 (Urodon) N: AG MO A: QA
granulata Schilsky, 1904: 78 (Urodon)
baudiifarsetiae Peyerimhoff, 1931a: 117 (Urodon) N: AG
cana Küster, 1848a: 83 (Urodon) E: FR GR HU IT (Sardegna, Sicilia) MA PT SP N: MO
carpetana K. Daniel, 1906: 181 (Urodon) E: SP
ciliata Schilsky, 1904: 78 (Urodon) A: UZ
concolor Fâhraeus, 1839: 142 (Urodon) E: BU CT GR ST A: TR UZ "Caucasus"
allardi Jekel, 1876: 536 (Urodon) [RN]
villosa Allard, 1869: [21] (Urodon) [RN]
villosa Allard, 1870a: 326 (Urodon) [RN, HN]
conformis Gyllenhal, 1833: 114 (Urodon) E: CZ FR GE HU NL SK SP ST ÜK N: AG MO
sericata Motschulsky, 1875a: 228 (Urodon)
cretacea K. Daniel & J. Daniel, 1903: 327 (Urodon) A: TR
demaisoni Peyerimhoff, 1926b: 195 (Urodon) N: AG MO
densata Reitter, 1897d: 245 (Urodon) A: TR
exigua Motschulsky, 1875a: 228 (Urodon) E: ST UK
ferrantei Reitter, 1908a: 51 (Urodon) N: EG A: SI
testaceomarginalis Pic, 1939: 148 (Urodon)
flavescens Küster, 1850a: 88 (Urodon) E: AU IT FR MC PT SP N: AG MO A: IS
armata Baudi di Selve, 1887a: 132 [= 1887b: 491] (Urodon)
fortirostris Korotyaev, 1988b: 122 E: AR A: INTR
fulvipes Reitter, 1897d: 246 (Urodon) A: TM UZ
hesperidis Iablokoff-Khnzorian, 1957: 172 (Urodon) E: AR
humeralts Motschulsky, 1875a: 228 (Urodon) N: AG
108 ANTHRIBIDAE
kasparyani Korotyaev, 1988b: 130 E: ABOG
kaszabi Strejcek, 1973: 233 (Urodon) E: AB AR AU BU CT CZ GE GG GR HU RO SK ST TR UK IN. AG A:
ES IN KZ MG SY TR UZ WS
Orientalis Strejcek, 1982: 135 (Urodon)
tchitensis Egorov, 1996a: 166 (Urodon)
korbi Schilsky, 1904: 78 (Urodon) E: BU IT RO ST A: TR
longa AMaid, 1868: 101 (Urodon) N: AG MO TU
spinicollis Perris, 1869: 32 (Urodon)
luteae Peyerimhoff, 1929c: 145 (Urodon) N: AG
maura K. Daniel, 1912: 150 (Urodon) N: MO
medvedevi Korotyaev, 1988b: 126 E: KZ A: KZ TM UZ
musculus K. Daniel & J. Daniel, 1903: 328 (Urodon) E: AU BU RO ST UK "Transcaucasia" A: TR
parallela Küster, 1848a: 84 (Urodon) E: FR SP N: AG MO
parvula Motschulsky, 1875a: 228 (Urodon) E: ST UK
picta Motschulsky, 1875a: 228 (Urodon) N: AG
pygmaea Gyllenhal, 1833: 114 (Urodon) E: FR IT PT SP N: AG MO TU
alensis Gredler, 1866: 299 (Urodon)
meridionalis Jacquet, 1888: 31 (Urodon)
pus illa Baudi di Selve, 1887a: 131 [= 1887b: 490] (Urodon)
randoniae Peyerimhoff, 1926a: 105 (Urodon) N: AG
reymondiA. Hoffmann, 1953b: 189 (Urodon) N: MO
rosinae Pic, 1904b: 40 (Urodon) A: TR
rufipes nigritarsis Reitter, 1916b: 126 (Urodon) E: AU CZ FR GE PL SK ST
prieferti A. Hoffmann, 1961: 38 (Urodon)
ruteriA. Hoffmann, 1950b: 193 (Urodon)
rufipes rufipes Olivier, 1790: 202 (Bruchus) E: AU BU CZ FR GB GE GR HU IT LU MC NL PL RO SK SP
ST SV SZ N: MO
argentata Küster, 1847a: 91 (Urodon)
rufipes Fabricius, 1792: 373 (Bruchus) [HN]
sericea Fabricius, 1801:411 (Anthribus)
scabra Baudi di Selve, 1890a: 213 [= 1890b: 345] (Urodon) A: SY
schouwiae Peyerimhoff, 1929c: 146 (Urodon) N: AG
schusteri Schilsky, 1912: 85 (Urodon) E: AU BU CZ FR GR HU MC RO SK A: KZ TR
semisuturalis Reitter, 1908a: 50 (Urodon) N: EG
sinaitica K. Daniel, 1909: 327 (Urodon) A: SI
sugonyaevi Korotyaev, 1988b: 124 E: AR
suturalis Fabricius, 1792: 372 (Bruchus) E: AR AU BU CZ FR GE GR HU IT LU NL PL RO SK SP ST SZ
UK N: AG MO TU A: IN SY TR WS
angularis Allard, 1868: 101 (Urodon)
tibialis Baudi di Selve, 1887a: 125 [= 1887b: 483] (Urodon)
testaceipes Reiche, 1861a: 91 (Urodon) E: SP N: AG MO TU
luteipes Reitter, 1897d: 246 [incorrect subsequent spelling]
verae Korotyaev, 1988b: 128 E: AB
genus Cercomorphus Perris & Jacquelin du Val, 1864: 293 type species Cercomorphus duvalii Perris, 1864
Microctilodes Reitter, 1875a: 27 type species Microctilodes ragusae Reitter, 1875
abbreviatus Reitter, 1896: 234 E: SP
bicolor Abeille de Perrin, 1895: cccxxvi A: IT (Sardegna) N: AG TU
abdominalis Pic, 1896a: 30
obscurior Pic, 1896a: 30
bos Abeille de Perrin, 1895: cccxxvi A: IS
demoflysi Normand, 1940: 52 N: TU
duvalii Perris, 1864: 295 E: SP
pectinator Peyerimhoff, 1924: 63 N: AG
ragusae Reitter, 1875a: 30 (Microctilodes) E: IT (Sicilia) MA
sterilis Peyerimhoff, 1919b: 233 N: AG
 BELIDAE 109
syriacus Pic, 1925a: 10 A: SY
syrticus Peyerimhoff, 1925a: 17 N: TU
titschacki Uyttenboogaart, 1934: 164 N: CI (Lanzarote)

family Anthribidae, nomina dubia

Anthribus discifer Schrank, 1798: 474 E: GE
Anthribus ephippiger Gistel, 1831: 305 E: GE
Anthribus lividus Olivier, 1789: 161 E: FR
AnthribuspallinosusAnonymus, 1826:338 E:NL
Anthribus striatus Olivier, 1789: 161 "Europe"
Anthribus unicolor Olivier, 1789: 160 E: FR
Anthribus vittatus Geoffroy, 1785: 138 E: FR

family BELIDAE Schoenherr, 1826

subfamily Oxycoryninae Schoenherr, 1840
tribe Aglycyderini Wollaston, 1864
genus Aglycyderes Westwood, 1864:179 type species Aglycyderes setifer Westwood, 1864
setifer Westwood, 1864: 179 N: CI
tavakiliani Menier, 1974: 14 N: MO

family RHYNCHITIDAE Gistel, 1848

subfamily Rhynchitinae Gistel, 1848
tribe Auletini Desbrochers des Loges, 1908
subtribe Auletina Desbrochers des Loges, 1908
genus Auletes Schoenherr, 1826: 46 type species Auletes tubicen Schoenherr, 1826
Tubicenus Latreille, 1829: 591 [no species included]
aeneus Legalov, 2002e: 178 N: LB
graecus Legalov, 2002e: 179 E: GR
tubicen Schoenherr, 1826: 47 E: CR FR GR IT SP TR N: AG MO TU A: TR
meridionalis Jacquelin du Val, 1854: 8
tubicen Boheman, 1829: 125
wagenblasti Uyttenboogaart, 1935b: 247 E: SP (Baléares)

subtribe Auletobiina Legalov, 2001

genus Auletobius Desbrochers des Loges, 1869c: 396 type species Auletes basilaris Gyllenhal, 1839 (=
Involvulus sanguisorbae Schrank, 1798)
subgenus Auletobius Desbrochers des Loges, 1869c: 396 type species Auletes basilaris Gyllenhal, 1839 (=
Involvulus sanguisorbae Schrank, 1798)
Auletulus Voss, 1922d: 23 type species Auletulus kamerunensis Voss, 1922
Metopon G.R. Waterhouse, 1842: 68 [HN] type species Metopon suturalis G.R. Waterhouse, 1842
Metopum Agassiz, 1846: 232 [RN] type species Metopon suturalis G.R. Waterhouse, 1842
Parauletes Sawada, 1993: 6 type species Auletobius monticola Voss, 1937
arunachalensis Legalov, 2007a: 36 A: AP
bengalensis Legalov, 2003b: 95 A: SD
berberidis Voss, 1933c: 135 A: SD UP
calvus Sharp, 1889: 72 {Auletes) A: JA
egorovi Legalov, 2006c: 16 A: FE HEI NC
emeishanicus Legalov, 2007a: 37 A: SCH
flavus Legalov, 2007a: 37 A: TAI
fochaensis Legalov, 2003b: 95 A: YUN
fukienensis Voss, 1941a: 239 A: FUJ
fumigatus Roelofs, 1874: 151 (Auletes) A: FE FUJ JA NC SC
110 RHYNCHITIDAE
japonicus Voss, 1922d: 41
testaceus Roelofs, 1874: 152 (Auletes)
ghumensis Legalov, 2007a: 38 A: SD
habashcmensis Legalov, 2007a: 38 A: YUN
hunanicus Legalov, 2007a: 38 A: HUN
irkutensis Faust, 1893b: 201 A: BEI ES FE JIL LIA MG NC
lumlensis Legalov, 2003b: 97 A: NP
nitens Kôno, 1930a: 32 A: TAI
obscurethoracalis Legalov, 2007a: 40 A: NP
planifrons Sawada, 1993: 28 A: JA (Ryukyu)
puberulus Faust, 1882a: 283 {Auletes) A: FE NC SC
ruber Legalov, 2007a: 41 A: YUN
rufus Legalov, 2007a: 41 A: NP
sanguisorbae Schrank, 1798: 476 (Involvulus) E: AU BU CR CT CZ GE GR HU IT PL RO SK SL ST SZ UK
A: ANH ES FE HEI JA KZ MG NC NMO UZ WS NAR
basilaris Gyllenhal, 1839: 346 (Auletes)
congruus Walker, 1866: 331 (Rhynchites)
nigrocyaneus Waltl, 1839: 222 {Rhynchites)
subcoeruleus LeConte, 1876: 4 (Auletes)
schoeni Legalov, 2001b: 41 A: SCH
shanpingensis Legalov, 2007a: 42 A: SCH
subbasalis Voss, 1933c: 119 A: FUJ SHN SHX ORR
taiwanensis Legalov, 2007a: 42 A: TAI
volkovitshi Legalov, 2003b: 102 A: NP
subgenus Canarauletes Legalov, 2007a: 44 type species Auletes convexifrons Wollaston, 1864
anceps Wollaston, 1864: 305 (Auletes) N: CI
freyi Uyttenboogaart, 1940: 667
convexifrons Wollaston, 1864: 305 (Auletes) N: CI (Gran Canaria)
cylindricollis Wollaston, 1864: 304 (Auletes) N: CI
maderensis Wollaston, 1854: 416 (Auletes) N: MR (Madeira)
maroccanus A. Hoffmann, 1953b: 193 E: SP N: MO
subgenus Pseudometopum Legalov, 2003b: 105 type species Auletobius subgranulatus Voss, 1933
subgranulatus Voss, 1933c: 120 A: YUN ORR
submaculatus Sharp, 1889: 73 (Auletes) A: FE JA
subgenus Rostauletes Legalov, 2007a: 46 type species Auletobius longirostris Legalov, 2007
longirostris Legalov, 2007a: 46 A: TAI

subtribe Pseudomesauletina Legalov, 2003

genus Aletinus Desbrochers des Loges, 1908a: 11 type species Auletes maculipennis Jacquelin du Val, 1854
subgenus Aletinus Desbrochers des Loges, 1908a: 11 type species Auletes maculipennis Jacquelin du Val, 1854
maculipennis Jacquelin du Val, 1854: 8 (Auletes) E: FR (Corse) GR IT N: AG MO TU
concolor Desbrochers des Loges, 1869c: 406 (Auletobius)
subgenus Heterauletes Voss, 1933c: 116 type species Auletes constrictus Reitter, 1891
akinini Faust, 1885a: 161 (Auletes) A: AF GAN KI KZ NMO TD TM UZ
beckeri Desbrochers des Loges, 1875b: 210 (Auletes) E: AB AR ST
constrictus Reitter, 1891a: 32 (Auletes) E: AB AR GG A: TM TR UZ
mongolicus Ter-Minasian, 1984: 302 (Auletobius) A: MG

genus Dicranognathus L. Redtenbacher, 1844: 538 type species Dicranognathus nebulosus L Redtenbacher,
1844
bipunctatus Legalov, 2007a: 67 A: SCH
hunanicus Legalov, 2007a: 67 A: HUN
nebulosus L. Redtenbacher, 1844: 538 A: KA UP
obscurofasciatus Voss, 1936b: 280 A: YUN
 RHYNCHITIDAE 111
genus Eomesauletes Legalov, 2001b: 52 type species Rhynchites politus Boheman, 1829 (= Rhynchites politus
Lepeletier & Audinet-Serville, 1825)
politus Lepeletier & Audinet-Serville, 1825: 304 (Rhynchites) E: AR BH BU CR FR GR IT MC PL PT SP ST
TR UK N: MO A: CY IS JO LE SY TR
cremienVoss, 1922d:61 (Auletobius)
emgei Stierlin, 1888: 60 (Auletes)
ilicis Gene, 1839a: 36 [= 1839b: 76] (Rhynchites)
politus Boheman, 1829: 124 (Rhynchites) [HN]
tessoni Mulsant & Godait, 1868a: 407 (Auletes)

genus Eumetopon Voss, 1922d: 30 type species Auletobius flavimaculatus Voss, 1922
subgenus Eumetopon Voss, 1922d: 30 type species Auletobius flavimaculatus Voss, 1922
flavimaculatum Voss, 1922d: 89 (Auletobius) A: ZHE ORR
chinense Voss, 1939d: 608 (Auletobius) [HN]

genus Eurostauletes Voss, 1933c: 116 type species Auletes rubrorufus Solsky, 1881
longimanus Gebier, 1830: 145 (Rhynchites) A: ES FE KZ MG WS
longipes Gyllenhal, 1839: 321 (Rhynchites)
kaszabi Ter-Minasian, 1971a: 264 (Auletobius)
procerus Reitter, 1901a: 83 (Auletes) A: AF KI TD UZ
rubrorufus Solsky, 1881: 261 (Auletes) A: KI KZ TD UZ
turkestanensis Voss, 1933e: 134 (Auletobius)

genus Mesauletobius Legalov, 2001b: 51 type species Auletes pubescens Kiesenwetter, 1852
pubescens Kiesenwetter, 1852: 627 (Auletes) E: BU CR FR GR IT PT SP TR N: AG MO A: TR
cisticola Fairmaire, 1859b: clxiii (Auletes)
subplumbeus Chevrolat, 1860b: 129 (Auletes)

genus Pseudomesauletes Legalov, 2001b: 53 type species Auletes uniformis Roelofs, 1874
subgenus Ovauletes Legalov, 2007a: 59 type species Pseudomesauletes latus Legalov, 2007
brevis Legalov, 2007a: 59 A: NP
indiens Legalov, 2007a: 59 A: UP
latus Legalov, 2007a: 60 A: FUJ
sitschuanensis Legalov, 2001b: 55 (Auletobius) A: SCH
yunnanicus Legalov, 2007a: 60 A: YUN
subgenus Pseudomesauletes Legalov, 2001b: 53 type species Auletes uniformis Roelofs, 1874
fasciatus Legalov, 2007a: 61 A: YUN
formosanus Voss, 1922c: 277 (Auletobius) A: TAI
hirtellus Voss, 1941a: 240 (Auletobius) A: FUJ GUA GUX HEN HUB ZHE ORR
longicollis Faust, 1898: 299 (Auletobius) A: HP ORR
nigrinus Voss, 1920: 161 (Auletobius) A : H P U P
nuristanensis Voss, 1959a: 68 (Auletobius) A: AF
subpicescens Voss, 1933c: 125 (Auletobius) A: UP ORR
subtuberculatus Voss, 1922c: 278 (Auletobius) A: TAI
tuberculatus Voss, 1922c: 278 (Auletobius) A: TAI
uniformis Roelofs, 1874: 152 (Auletes) A: FE FUJ JA NC SC ZHE
okinawaensis Voss, 1971b: 43 (Auletobius)
subgenus Rubrauletes Legalov, 2003b: 123 type species Auletobius consimilis Voss, 1930
binbyanicus Legalov, 2007a: 65 A: YUN
chinensis Voss, 1933c: 128 (Auletobius) A: SHN
consimilis Voss, 1930b: 66 (Auletobius) A: NP UP YUN ORR
dundai Legalov, 2007a: 65 A: SCH YUN
huitongensis Legalov, 2007a: 66 A: HUN
klapperichiVow, 1941a: 241 (Auletobius) A: FUJ YUN
kryzhanovskyi Legalov, 2009b: 62 A: FUJ YUN
112 RHYNCHITIDAE
nanpingensis Legalov, 2007a: 66 A: SCH
nepalensis Voss, 1974: 44 (Auletobius) A: NP
perturbants Voss, 1930b: 65 (Auletobius) A: SCH
poirasi Legalov, 2009b: 64 A: AP NP UP
sanguineus Voss, 1920: 162 (Auletobius) A: HP UP
thompsoni Legalov, 2009a: 30 A: YUN
vereshaginae Legalov, 2007a: 66 A: YUN

genus Pseudomesauletes, species incertae sedis

reichei Desbrochers des Loges, 1869c: 407 (Auletobius) [DA] A:"Arabie"

genus Pseudodicranognathus Legalov, 2001b: 64 type species Pseudodicranognathus motschulskyi Legalov,

2001
motschulskyi Legalov, 2001b: 64 A: "Northeastern India"

genus Yunnanuletes Legalov, 2007a: 58 type species Yunnanuletes heishuensis Legalov, 2007
heishuensis Legalov, 2007a: 59 A: SCH YUN
murzini Legalov, 2007a: 59 A: YUN

tribe Byctiscini Voss, 1923

subtribe Byctiscina Voss, 1923
genus Aspidobyctiscus Schilsky, 1903: U type species Rhynchites lacunipennis Jekel, 1860
subgenus Aspidobyctiscus Schilsky, 1903: U type species Rhynchites lacunipennis Jekel, 1860
lacunipennis Jekel, 1860: 225 (Rhynchites) A: ANH BEI ES FE FUJ GAN GUA GUX HEB HEI HEN HKG
HUB HUN JA JIA JIL JIX LIA NC NP SC SCH SHA SHG SHN TAI WS ORR
amurensis Voss, 1930c: 199 (Byctiscus)
cicatricosus Motschulsky, 1860: 173 (Attelabus)
collaris Voss, 1930c: 199 (Byctiscus)
subtilis Voss, 1930c: 199 (Byctiscus)
marshalli Legalov, 2009a: 35 A:NPUP ORR
nanpingensis Legalov, 2009a: 35 A: SCH
yunnanicus Voss, 1930c: 199 (Byctiscus) A: YUN
subgenus Chinobyctiscus Legalov, 2005: 110 type species Aspidobyctiscus mirabilis Legalov & Liu, 2005
mirabilis Legalov & Liu, 2005: 111 A: HUB SCH
subgenus Eobyctiscus Legalov, 2005: 104 type species Byctiscus coerulans Voss, 1929
coerulans Voss, 1929a: 28 (Byctiscus) A: BEI FUJ HUB JIA JIX SHG TAI ZHE
pseudocoerulans Legalov & Liu, 2005a: 107 A: GUX
shiguensis Legalov, 2007a: 206 A: YUN
vossi Legalov & Liu, 2005: 104 A: ZHE
zhejiangensis Legalov & Liu, 2005: 106 A: ZHE
subgenus Nepalobyctiscus Legalov, 2003b: 332 type species Aspidobyctiscus giganteus Legalov, 2003
giganteus Legalov, 2003b: 332 A: NP
sculpntrants Pascoe, 1875: 394 (Rhynchites) A: UP
subgenus Parabyctiscus Legalov, 2003b: 330 type species Aspidobyctiscus kazantsevi Legalov, 2003
kazantsevi Legalov, 2003b: 330 A: SCH
konoi Legalov & Liu, 2005: 112 A: YUN
niger Legalov, 2003b: 332 A: XIZ
subgenus Taiwanobyctiscus Kôno, 1929a: 128 type species Byctiscus paviei Aurivillius, 1891
complanants Voss, 1933d: 111 (Byctiscus) A: HEB
cyanocupreus Legalov & Liu, 2005: 110 A: YUN
nigrocyaneus Legalov & Liu, 2005: 108 A: YUN
paviei Aurivillius, 1891: 207 (Byctiscus) A: ANH FUJ GUA GUX HEB HEI HUB JIA JIL JIX LIA SCH SHA
TAI YUN "Korea" ORR
separandus Voss, 1930c: 200 (Byctiscus) A: TAI
 RHYNCHITIDAE 113
sitchuanensis Legalov, 2003b: 328 A: SCH

genus Byctisculus Legalov, 2003b: 324 type species Byctiscidiw ? griseus Voss, 1931
griseoides Legalov, 2007a: 200 A: YUN
griseus Voss, 1930c: 231 {Byctiscidius) A: YUN ORR
genus Byctiscus C. G. Thomson, 1859: 130 type species Curculio populi Linnaeus, 1758
Bystictus Desbrochers des Loges, 1908c: 80 [unjustified RN]
angusticollis Legalov, 2007a: 201 A: FUJ JIX
betulae Linnaeus, 1758: 381 {Curculio) E: AB AR AU BE BH BU BY CR CT CZ DE EN FI FR GB GE GG
GR HU IT LA LT LU MC MD ME NL NR NT PL PT RO SB SK SL SP ST SV SZ UK A: ES FE GUI HEI
HEN IS JIL JIX KZ LIA SY SW TM TR WS "Korea
alniO. F. Müller, 1776: 91 {Rhinomacer) [RN]
auratus Geoffroy, 1785: 113 {Rhinomacer
betuleti Fabricius, 1792: 387 {Attelabus) [RN]
bispinus O. F. Müller, 1776: 91 {Rhinomacer
destitutes Voss, 1930c: 197
inermis O. F. Müller, 1776: 91 {Rhinomacer)
intermedins Voss, 1930c: 197
nitens Marsham, 1802: 242 {Curculio) [HN]
unispinus O. F. Müller, 1776: 91 {Rhinomacer)
bilineatoides Legalov, 2007a: 202 A: SCH SHX YUN
bilineatus Legalov, 2003b: 326 A: HEB ZHE
congener Jekel, 1860: 224 {Rhynchites) A: ES FE FUJ HEB HEN HUB JA LIA NC SC SCH SHG SHX ZH
motschoulskyi Sharp, 1889: 58 [RN]
motschulskyi Lewis, 1879: 22 {Rhynchites) [unnecessary RN]
motschulskyi Faust, 1890e: 429 [HN]
puberulus Motschulsky, 1860: 170 {Rhynchites)
subauratus Voss, 1943b: 23
tsherskii Suvorov, 1915: 345
davidiani Legalov, 2003b: 325 A: HUB SCH
davidis Fairmaire, 1878: 130 {Rhynchites) A: SCH
fausti Sharp, 1889: 60 A: FE JA NC SC SC
gibbirostris Schilsky, 1906a: 85
ßkienensis Voss, 1949: 158 A: FUJ
fulminans Voss, 1930c: 199 A: HEN JIX SHN ZHE
lucidus Voss, 1930c: 199
himalayaensis Legalov, 2007a: 203 A: "Himalaya"
impressus formosanus Voss, 1922c: 285 A: TAI
impressus impressus Fairmaire, 1900: 636 {Rhynchites) A: ANH FUJ GAN GUA GUI HUB JIX SCH SHA
SHG ZHE
chinensis Formanek, 1911: 208
subpectitus Voss, 1943c: 232
kresli Legalov, 2003b: 327 A: SCH
macros Legalov, 2004a: 70 A: SHA SCH
minimus Legalov & Liu, 2005: 102 A: HEB
moupinensis Legalov, 2007a: 204 A: SCH
mutator Faust, 1890e: 429 A: GAN YUN
populi Linnaeus, 1758: 381 {Curculio) E: AU BE BH BU BY CR CT CZ DE EN FI FR GB GE GR HU IT LA
LT LU MC MD NL NR NT PL RO SB SK SL SP SV SZ UK A: ES FE IN JA KI KZ MG NE NMO QIN
SCH SHX TD TR UZ WS
nigritutus Schilsky, 1889: 340
parvulus Sharp, 1889: 60
potanini Legalov, 2004a: 70 A: SCH
princeps Solsky, 1872: 284 {Rhynchites) A: BEI ES FE GAN HEB HEI HEN JA JIL LIA NC SC SCH SHG
SHX XIZ
regalis Roelofs, 1874: 142 {Rhynchites)
qingensis Legalov, 2009a: 34 A: SHA
114 RHYNCH1TIDAE
regularis Voss, 1930b: 79 A: ANH FE HEB HEI LIA NC SC
rugosus Gebier, 1830: 146 (Rhynchites) A: BEI ES FE FUJ GAN HEI HUB JA JIL KZ MG NC NMO SC SCH
SHX WS XIN ZHE ORR
diversicolor Kolbe, 1886: 217 (Rhynchites)
omissus Voss, 1920: 169
similaris Voss, 1920: 170 A: HUB ZHE
thibetanus Legalov, 2007a: 206 A: XIZ
venustus Pascoe, 1875: 393 {Rhynchites) A: FE HUB JA NC SC
haroldi Roelofe, 1879b: liii (Rhynchites)
Ai'meKôno, 1929b: 53
reversus Sharp, 1889:60
genus Byctiscus, species incertae sedis
auronitens Gistel, 1857: 534 (Rhynchites) E: GE
bicolor Gistel, 1857: 534 (Rhynchites) E: GE

genus Neobycüscidius Legalov, 2007a: 201 type species Neobycüscidius hanungensis Legalov, 2007
weibaoshanensis Legalov, 2008a: 50 A: YUN

subtribe Listrobyctiscina Legalov, 2003

genus Nelistrobycüscus Legalov, 2003b: 338 type species Rhynchites gagates Pascoe, 1875
subgenus Listrobyctiscidius Legalov, 2003b: 339 type species Byctiscuspatruelis Voss, 1922
patruelis Voss, 1922c: 283 (Byctiscus) A: TAI
pseudopatruelis Legalov & Liu, 2005: 112 A: LIA
sichuanensis Legalov, 2007a: 213 A: SCH

subtribe Svetlanaebyctiscina Legalov, 2003

genus Baikovius Legalov, 2005:100 type species Baikovius unicus Legalov & Liu, 2005
unicus Legalov & Liu, 2005: 100 A: YUN

genus Byctiscidius Voss, 1923: 510 type species Byctiscidius insularis Voss, 1923
Probyctiscidius Legalov, 2007a: 199 type species Byctiscidius insularis Voss, 1923
parais Voss, 1920: 168 (Rhynchites) A: LIA

genus ByctiscophUus Voss, 1930c: 193 type species Byctiscophilus championi Voss, 1931
championi Voss, 1931b: 232 A: NP UP

genus Probyctisculus Legalov, 2007a: 199 type species Probyctisculus mustangensis Legalov, 2007
mustangensis Legalov, 2007a: 200 A: NP
nanitalensis Legalov, 2007a: 200 A: UP

genus Svetlanaebyctiscus Legalov, 2001a: 341 type species Caenorhinus vitis Ter-Minasian, 1959
dundai Legalov, 2008a: 49 A: YUN
vitis Ter-Minasian, 1959: 361 (Caenorhinus) A: FE

tribe Deporaini Voss, 1929

subtribe Chonostropheina Morimoto, 1962
genus Chonostropheus Prell, 1924: 162 type species Attelabus tristis Fabricius, 1794
Rhinchitobius Kôno, 1928: 177 type species Attelabus tristis Fabricius, 1794
chinensis Voss, 1939d: 613 A: ZHE
chujoi Voss, 1956a: 13 A: JA
seminiger Reitter, 1881: 516 {Rhynchites) E: AB AR AU FR GE GG IT ST SZ A: TR
bavariensis Voss, 1939d: 615
tristis Fabricius, 1794: 454 (Attelabus) E: AL AU BE BH BU CR CZ FR GE GR HU IT MD NL PL RO SB SK
SL ST SZ UK
 RHYNCHITIDAE 115
subtribe Deporaina Voss, 1929
genus Caenorhinus C. G. Thomson, 1859: 130 type species Rhynchites megacephalus Germar, 1824 (=
Rhynchites mannerheimii Hummel, 1823)
subgenus Acaenorhinus Legalov, 2007a: 106 type species Deporaus impressipennis Voss, 1922
nigriceps Voss, 1922e: 394 (Deporaus) A: FUJ ORR
subgenus Apoderites Sawada, 1987: 654 type species Apoderites commodus Sawada, 1987
commodus Sawada, 1987: 654 (Apoderites) A: JA
subgenus Arodepus Heller, 1922b: 551 type species Eugnamptus marginellus Faust, 1898
Arodepus Voss, 1922e: 387 [HN] type species Deporauspenangensis Voss, 1922
marginellus Faust, 1898: 299 (Eugnamptus) A: NP ORR
marginatus Faust, 1895b: 168 {Deporaus) [HN]
semirufiis Faust, 1898: 299 (Deporaus)
subgenus Asiodeporaus Legalov, 2007a: 109 type species Caenorhinus indochinensis Legalov, 2007
bicoloripes Legalov, 2007a: 109 A: YUN
subgenus Caenorhinus C. G. Thomson, 1859: 130 type species Rhynchites megacephalus Germar, 1824 (=
Rhynchites mannerheimii Hummel, 1823)
brevis Legalov, 2007a: 116 A: YUN
dundai Legalov, 2007a: 116 A: SCH
eumegacephalus Sawada, 1993: 80 (Deporaus) A: FE JA
mannerheimii Hummel, 1823: 45 (Rhynchites) E: AU BE CT CZ DE EN FI FR GB GE HU IT LA LT LU MC
NL NR NT PL SB SP SV A: ES FE FUJ HEB HEI JA MG NC SC SCH SHX UP WS YUN
constrictus Gyllenhal, 1839: 335 (Rhynchites)
cyaneopennis Stephens, 1831: 199 (Rhynchites)
laevicollis Stephens, 1831: 199 (Rhynchites)
megacephalus Germar, 1824: 187 (Rhynchites)
planipennis Roelofs, 1874: 150 (Rhynchites)
potanini Legalov, 2007a: 117 A: SCH
rasuwanus Legalov, 2007a: 117 A: NP
rusticapitus Legalov, 2007a: 117 A: SCH
yundianensis Legalov, 2007a: 117 A: YUN
subgenus Convexodeporaus Legalov, 2007a: 104 type species Deporaus parvicollis Voss, 1942
parvicollis Voss, 1942b: 98 (Deporaus) A: FUJ
subgenus Flavodeporaus Legalov, 2007a: 109 type species Deporaus nigricornis Heller, 1922
atrorufus Voss, 1942b: 97 (Deporaus) A: FUJ YUN
bifasciatus Legalov, 2007a: 110 A: YUN
eduardi Legalov, 2007a: 111 A: YUN
femoratus Voss, 1942b: 97 (Deporaus) A: FUJ
fiikienensis Voss, 1941a: 245 (Deporaus) A: FUJ
fuscipennis Sharp, 1889: 71 (Deporaus) A: FE JA (Hokkaido, Honshu)
flavipes Schilsky, 1906a: 80 (Rhynchites) [HN]
gilviventris Voss, 1938b: 105 (Deporaus) A: FUJ ORR
insularis Sawada, 1993: 76 (Deporaus) A: JA (Ryukyu) NC
klapperichi Voss, 1941a: 244 (Deporaus) A: FUJ
mansiensis Legalov, 2007a: 113 A: YUN
nigroapicalis Legalov, 2007a: 113 A: YUN ZHE
ohdaisanus Nakane, 1963b: 33 (Deporaus) A: JA (Honshu) NC
phohaensis Legalov, 2007a: 114 A: YUN
pseudofemoratus Legalov, 2007a: 114 A: FUJ
tigris Sawada, 1993: 72 (Deporaus) A: JA (Kyushu)
vossi Sawada, 1993: 78 (Deporaus) [RN] A: FE JA (Honshu, Shikoku) SC
pallidiventris Voss, 1957: 33 (Deporaus) [HN]
subgenus Hypodeporaus Voss, 1922e: 388 type species Deporaus conicirostris Voss, 1922
pokharensis Legalov, 2007a: 106 A: NP
subgenus Metallarodepus Legalov, 2003b: 195 type species Deporaus flaviventris Voss, 1924
davidiani Legalov, 2007a: 117 A: SCH
116 RHYNCHITIDAE
flaviventris Voss, 1924b: 39 {Deporaus) A: SCH SD YUN OR
flaviventris Voss, 1938b: 106 (Deporaus) [HN]
flaviventroides Legalov, 2007a: 118 A: YUN
orlovi Legalov, 2007a: 118 A: NP
socius Faust, 1882a: 286 (Rhynchites) A: SD UP ORR
subgenus Neoarodepus Legalov, 2003b: 183 type species Deporaus subcoarctatus Voss, 1938
rufiventris Voss, 1922c: 280 (Deporaus) A: TAI
subcoarctatus Voss, 1938b: 104 (Deporaus) A: FUJ NP UP
subgenus Nigroarodepus Legalov, 2003b: 185 type species Caenorhinus zherichini Legalov, 2003
biounctatus Legalov, 2007a: 104 (Nigroarodepus) A: YUN
subgenus Orientarodepus Legalov, 2003b: 186 type species Deporaus minimus Kôno, 1928
minimus koreanus Voss, 1929a: 28 (Depasophilus) A: BEI FE FUJ HEB HEI NC SC
pauculus Voss, 1941b: 117 (Deporaus)
minimus minimus Kôno, 1928: 175 (Deporaus) A: JA
pacatoides Voss, 1937b: 207 (Depasophilus)
subgenus Platyrhynchites Voss, 1922e: 387 type species Deporaus basalis Voss, 1922
robertsi Voss, 1935d: 95 A: PA
robertsi Voss, 1938b: 105 [HN]
subgenus Platyrhynchitoides Legalov, 2007a: 105 type species Deporaus indicus Voss, 1924
indicus Voss, 1924b: 39 A: NP SD ORR
subgenus Pseudodepasophilus Voss, 1942b: 96 type species Depasophilus blandulus Voss, 1942
blandulus Voss, 1942b: 95 (Depasophilus) A: FUJ

genus Caenorhinus, species incertae sedis

disjunctus Voss, 1938b: 100 (Deporaus) "India" ORR
incertus Voss, 1958: 12 (Deporaus) A: FUJ
marginatus Pascoe, 1883a: 123 (Eugnamptus) A:GUAGUXYUN ORR
SK&rt/is Voss, 1938b: 105 (Deporaus) A: FUJ

genus Capylarodepus Voss, 1922e: 387 type species Deporaus galerucoides Heller, 1922
subgenus Capylarodepopsis Legalov, 2003b: 179 type species Capylarodepopsis medvedevi Legalov, 2003
medvedevi Legalov, 2007a: 101 (Capylarodepopsis) A: YUN ORR

genus Chokkirius Kôno, 1929a: 122 type species Rhynchites rosti Schilsky, 1906 (= Rhynchites truncatus
Sharp, 1889)
truncatus Sharp, 1889: 67 (Rhynchites) A: FE JA NC SC TAI
rosti Schilsky, 1906a: 79 (Rhynchites)

genus Deporaus Samouelle, 1819: 201 [NP] type species Attelabus betulae Linnaeus, 1758
subgenus Caeruleodeporaus Legalov, 2007a: 94 type species Deporaus lizipingensis Legalov, 2007
brunneoclavus Legalov, 2007a: 94 A: YUN
cangshauensis Legalov, 2007a: 95 A: YUN
flaviclavus Legalov, 2007a: 95 A: SCH YUN
gunlaishanensis Legalov, 2007a: 95 A: YUN
hingensis Legalov, 2007a: 95 A: YUN
kangdingensis Legalov, 2007a: 95 A: SCH
lizipingensis Legalov, 2007a: 95 A: SCH
mugezoensis Legalov, 2007a: 95 A: SCH
nanlingensis Legalov, 2007a: 95 A: GUA
weishanensis Legalov, 2007a: 96 A: YUN
subgenus Chinadeporaus Legalov, 2007a: 91 type species Deporaus bicolor Voss 1938
bicolor Voss, 1938b: 106 A: FUJ GAN JIA SHG ZHE ORR
minuroides Voss, 1958: 10 (Depasophilus) A: FUJ
nepalensis Legalov, 2007a: 92 (Chinadeporaus) A: NP
ruber Legalov & Liu, 2005: 99 A: YUN
 RHYNCHITIDAE 117
sichuanensis Legalov, 2007a: 92 (Chinadeporaus) A: SCH
strigellus Voss, 1958: 10 (Depasophilus) A: FUJ
tschungseni Voss, 1958: 9 (Depasophilus) A: FUJ
yunnanicus Legalov, 2007a: 92 (Chinadeporaus) A: YUN
subgenus Deporaus Samouelle, 1819: 201 [NP] type species Attelabus betulae Linnaeus, 1758
Platyrynchus Thunberg, 1815: 110 [NO] type species Attelabus betulae Linnaeus, 1758
betulae Linnaeus, 1758: 387 (Attelabus) E: AB AU BE BH BU BY CR CT CZ DE EN FI FR GB GE GG HU
IR IT LA LT LU MC MD NL NR NT PL RO SB SK SL SP ST SV SZ UK N: AG A: BEI ES FE HEB HEI
HUB JA (Hokkaido) KZ MG NC TR WS ZHE
excoriatoniger DeGeer, 1775: 259 (Curculio)
femoratus Olivier, 1789: 280 (Attelabus)
fidliginosus Gmelin, 1790: 1758(CwrcM//o)
niger Geoffroy, 1785: 114 (Rhinomacer)
nigrostriatus Goeze, 1777: 380 (Curculio)
nigrostriatus Petagna, 1792: 221 (Curculio) [HN]
populi Scopoli, 1763: 25 (Curculio) [HN]
populneus Gmelin, 1790: 1801 (Curculio) [RN]
podager Desbrochers des Loges, 1889a: xxxv (Rhynchites) A: IN SY TR
asiaticus Voss, 1974: 45 (Deporaus)
subgenus Neodeporaus Kôno, 1928: 177 type species Neodeporaus femoralis Kôno, 1928
femoralis Kôno, 1928: 176 (Neodeporaus) A: TAI
subgenus Paleodeporaus Legalov, 2003b: 174 type species Deporaus rhynchitoides Sawada, 1993
Paleodeporaoides Legalov, 2007a: 93 type species Paleodeporaus daliensis Legalov, 2007
amplicollis Voss, 1942b: 99 (Deporaus) A: FUJ ZHE
daliensis Legalov, 2007a: 94 (Paleodeporaus) A: YUN
jiriensis Legalov, 2007a: 92 (Paleodeporaus) A: NP
kathmanduensis Legalov, 2007a: 93 (Paleodeporaus) A: NP
luctuosus Voss, 1938b: 107 A: UP
reitteri Voss, 1935c: 62 A: SCH
rhynchitoides Sawada, 1993: 85 A: FE JA (Kyushu)
ussuriensis Legalov, 2006a: 3 A: BEI FE GAN HEI HUB
xishamensis Legalov, 2007a: 93 (Paleodeporaus) A: YUN
subgenus Parvodeporaus Legalov, 2007a: 94 type species Parvodeporaus gaoligongiensis Legalov, 2007
gaoligongiensis Legalov, 2007a: 94 (Parvodeporaus) A: YUN
subgenus Pseudapoderites Legalov, 2003b: 173 type species Rhynchites pacatus Faust, 1882
azarovae Legalov, 2006a: 2 A: FE (Sakhalin)
lewisi Legalov, 2007a: 96 A: "China"
pacatus Faust, 1882a: 285 (Rhynchites) A: FE HEI JA NC SC
pseudopacatus Legalov, 2007a: 96 A: YUN
septemtrionalis Sawada, 1993: 84 A: FE JA
subgenus Roelofsideporaus Legalov, 2003b: 174 type species Deporaus qffectatus Faust, 1887
Japonodeporaus Legalov, 2007a: 96 type species Deporaus hartmanni Voss, 1929
affectatus Faust, 1887b: 163 A: BEI ES FE HEB HEI JA (Hokkaido) JIL NC SHA
hartmanni Voss, 1929a: 28 A: FE JA (Honshu, Kyushu)
nidifiais Sawada & Lee, 1986: 672 A: FE JA (Honshu, Kyushu)
unicolor Roelofs, 1874: 149 (Rhynchites) A: FE FUJ HEB HEI JA JIL NC SC YUN ORR
yunnanicus Voss, 1930b: 80 (Deporaus)

genus Eusproda Sawada, 1987: 657 type species Rhynchites proximus Faust, 1882
fengshuensis Legalov, 2009a: 31 A: FUJ
proximo Faust, 1882a: 287 (Rhynchites) A: ES FE JA (Tsushima) NC SC NE ZHE
illibatus Voss, 1920: 166 (Rhynchites)
sichotana Ter-Minasian, 1950: 150 (Depasophilus)
tumida Voss, 1938b: 107 (Deporaus) A: SD
118 RHYNCHITIDAE
genus Exrhynchites Voss, 1930b: 80 type species Deporaus puberulus Faust, 1895
himachalensis Legalov, 2007a: 90 A: HP
major Voss, 1932b: 72 {Deporaus) A: YUN
mechtensis Legalov, 2007a: 91 A: NP
minor Voss, 1937a: 278 {Deporaus) A: YUN ZHE
puberulus Faust, 1895b: 166 {Deporaus) A: FUJ GUA GUI GUX HUB YUN ORR
subplanus Voss, 1920: 166 {Rhynchites) A: HP UP

genus Paradeporaus Kôno, 1927b: 60 type species Paradeporaus parasiticus Kôno, 1927 (= Rhynchites
depressus Faust, 1882)
Adeporaus Voss, 194Id: 117 type species Deporauspunctatissimus Reitter, 1899 (= Rhynchites depressus Faust, 1882)
depressus Faust, 1882a: 287 {Rhynchites) A: BEI FE HEI HUB JA NC
parasiticus Kôno, 1927b: 60
punctatissimus Reitter, 1899d: 306 {Deporaus)

tribe Eugnamptini Voss, 1930

genus Aderorhinus Sharp, 1889: 68 type species Rhynchites crioceroides Roelofs, 1874
bicolor Legalov, 2007a: 71 A: AP NP
crioceroides Roelofs, 1874: 147 {Rhynchites) A: FE JA SC ORR
formosanus Legalov, 2007a: 72 A: TAI
nepalensis Legalov, 2007a: 72 A: NP
pedicellaris Voss, 1930b: 67 A: FUJ HUN SCH YUN ZHE
variabilis Voss, 1941c: 124 A: NP YUN ORR
vossi Legalov, 2007a: 72 A: NP ORR

genus Eugnamptobius Voss, 1922e: 412 type species Eugnamptus insularis Voss, 1922
subgenus Cateugnamptus Legalov, 2007a: 72 type species Eugnamptus affinis Voss, 1924
affinis Voss, 1924b: 35 {Eugnamptus) A: SD ORR
congestus Voss, 1941c: 191 {Eugnamptus) A: FUJ GUA GUI HAI SCH SD YUN ORR
venus tus Kôno, 1930a: 29 {Aderorhinus) A: FE JA TAI "Korea"
subgenus Chryseugnamptus Legalov, 2003b: 150 type species Rhynchites flavirostris Desbrochers des Loges,
1890
caeruleoflavus Legalov, 2007a: 78 {Chryseugnamptus) A: ZHE
chengduensis Legalov, 2007a: 78 {Chryseugnamptus) A: SCH
desbrochersi Legalov, 2007a: 78 {Chryseugnamptus) A: YUN
subgenus Cyaneugnamptus Legalov, 2003b: 147 type species Cyaneugnamptus tenuicolloides Legalov, 1903
lunlinensis Legalov, 2003b: 149 {Cyaneugnamptus) A: YUN
pilosellus Voss, 1941c: 192 {Eugnamptus) A: SD ORR
pseudosauteri Legalov, 2007a: 78 {Cyaneugnamptus) A: TAI
sauteri Voss, 1922c: 279 {Eugnamptus) A: JA (Ryukyu) TAI
subgenus Eugnamptobius Voss, 1922e: 412 type species Eugnamptus insularis Voss, 1922
Eugnamptinus Legalov, 2003b: 143 type species Eugnamptus beiger Voss, 1941
aurifrons Roelofs, 1874: 151 {Eugnamptus) A: JA
cervinus Voss, 1930b: 67 {Eugnamptus) A: SCH YUN
distinetus Kôno, 1930a: 30 {Eugnamptus) A: TAI
diversus Voss, 1949: 156 {Eugnamptus) A: FUJ
dubius Voss, 1949: 155 {Eugnamptus) A: FUJ
flavipes Sharp, 1889: 70 {Eugnamptus) A: FE JA
fukienensis Voss, 1942b: 90 {Eugnamptus) A: FUJ
kuatunensis Voss, 1949: 153 {Eugnamptus) A: FUJ
morimotoi Nakane, 1963b: 34 {Eugnamptus) A: JA (Honshu)
nepalensis Legalov, 2003b: 145 {Eugnamptinus) A: NP
pedes tris Voss, 1949: 155 {Eugnamptus) A: FUJ
pelletieri Legalov, 2007a: 75 A: GUX
 RHYNCHITIDAE 119
sanguinolentes Voss, 1939d: 609 {Eugnamptus) A: FUJZHE
santaysanensis Legalov, 2007a: 76 A: YUN
subcoeruleifrons Voss, 1939d: 610 {Eugnamptus) A: ZHE ORR
subcoeruleifrons Voss, 1941c: 204 {Eugnamptus) [HN]
tsindunensis Legalov, 2007a: 76 A: YUN
weishanensis Legalov, 2007a: 77 A: YUN
subgenus Oreugnamptus Voss, 1941c: 208 type species Eugnamptus tessellatus Voss, 1920
lacunosus Voss, 1949: 157 {Eugnamptus) A: FUJ
tessellatus Voss, 1920: 171 {Eugnamptus) A: HP NP SD UP

genus Eugnamptus Schoenherr, 1839: 339 type species Anthribus collaris Fabricius, 1801 (= Rhynchites
angustatus Herbst, 1797)
Chineugnamptus Legalov, 2007a: 83 type species Chineugnamptus kubani Legalov, 2007
dibaphus Voss, 1942a: 26 A: FUJ YUN ORR
kazantsevi Legalov, 2003b: 156 A: ANH
kubani Legalov, 2007a: 83 {Chineugnamptus) A: YUN
manshinsis Legalov, 2003b: 157 A: YUN
monchadskii Legalov, 2007a: 86 A: YUN
palleolus Voss, 1939d: 610 A: HKG
palleolus Voss, 1941c: 169 [HN]
rudis Legalov, 2007a: 87 A: YUN
rusticus Legalov, 2003b: 160 A: FUJ GUA HUB SCH YUN ZHE
sitshuanensis Legalov, 2003b: 160 A: HUB SCH YUN
zhejiangensis Legalov, 2003b: 161 A: ANH FUJ HUB SHA ZHE

genus Neoeugnamptus Legalov, 2003b: 152 type species Rhynchites amurensis Faust, 1882
subgenus Neoeugnamptus Legalov, 2003b: 152 type species Rhynchites amurensis Faust, 1882
amurensis Faust, 1882a: 285 {Rhynchites) A: BEI FE FUJ GUI HEI JA JIL NC SC SHA ZHE
fragilis Sharp, 1889: 69 {Eugnamptus)
gracilicornis Schilsky, 1906a: 81 {Rhynchites)
austrochinensis Legalov, 2007a: 79 A: ZHE
cangshanensis Legalov, 2007a: 79 A: YUN
cyaneus Legalov, 2007a: 79 A: YUN
dundai Legalov, 2007a: 80 A: SCH
habashanensis Legalov, 2007a: 80 A: YUN
instabilis Voss, 1941c: 180 {Eugnamptus) A: YUN
konstantinovi Legalov, 2007a: 80 A: YUN
lijiangensis Legalov, 2007a: 80 A: YUN
linanensis Legalov, 2003b: 152 A: ZHE
nepalicus Legalov, 2003b: 153 A: NP
panfilovi Legalov, 2003b: 153 A: HAI YUN
parvulus Voss, 1941c: 182 {Eugnamptus) A: ZHE
potanini Legalov, 2003b: 153 A: SCH
taihorinensis Voss, 1922c: 280 {Eugnamptus) A: TAI
tarokoensis Legalov, 2007a: 81 A: TAI
volkovitshi Legalov, 2003b: 153 A: YUN
weibaoshanensis Legalov, 2007a: 81 A: YUN
yunnanensis Voss, 1941c: 180 {Eugnamptus) A: YUN
subgenus Proteugnamptellus Legalov, 2003b: 139 type species Eugnamptus bimaculosus Voss, 1941 (=
Eugnamptus bimaculosus Voss, 1939)
bimaculosus Voss, 1939d: 610 {Eugnamptus) A: NP UP
bimaculosus Voss, 1941c: 166 {Eugnamptus) [HN]
damonlunsis Legalov, 2003b: 140 {Proteugnamptellus) A: YUN
indicus Legalov, 2003b: 141 {Proteugnamptellus) A: SD
julonxueshanicus Legalov, 2007a: 71 {Proteugnamptellus) A,. YUN
120 RHYNCHITIDAE
tribe Rhynchitini Gistel, 1848
genus Auletomorphinus Legalov, 2007a: 134 type species Auletomorphinus limbourgi Legalov, 2007
yunnanensis Legalov, 2003b: 241 A: YUN

genus Auletomorphus Voss, 1923: 489 type species Rhynchites cupido Pascoe, 1875
subgenus Auletomorphus Voss, 1923: 489 type species Rhynchites cupido Pascoe, 1875
cupido Pascoe, 1875: 394 A: FUJ SD UP ORR
bicuspis Voss, 1924b: 42 (Rhynchites)
montanus Legalov, 2003b: 240 A: FUJ GUA NP SCH YUN
tonkinensis Voss, 1924b: 41 (Rhynchites) A: FUJ JIX SHN YUN ORR
monticola Voss, 1930b: 68 (Rhynchites)

genus Bicolorhynchus Legalov, 2003b: 284 type species Bicolorhynchus longiclavus Legalov, 2003
longiclavoides Legalov, 2007a: 151 A: YUN

genus Cartorhynchites Voss, 1958: 5 type species Rhynchites pallicollis Voss, 1932
subgenus Cartorhynchoides Legalov, 2003b: 276 type species Rhynchites apertus Sharp, 1889
aerosus Voss, 1958: 6 (Involvulus) A: FUJ
amamiensis Voss, 1971b: 44 (Involvulus) A: FE JA (Ryukyu)
apertus Sharp, 1889: 66 (Rhynchites) A: FE JA
crassiusculus Voss, 1938c: 145 (Rhynchites) A: "India"
flavus Sawada, 1993: 52 (Involvulus) A: JA (Ryukyu)
nantouensus Legalov, 2007a: 144 A: TAI
subtilis Sawada, 1993: 56 (Involvulus) A: FE JA (Ryukyu)
subgenus Hyperinvolvulus Legalov, 2003b: 277 type species Cartorhynchites angusticlavus Legalov, 2003
coarctus Voss, 1938d: 164 (Rhynchites) A: SD ORR
motschulskyi Legalov, 2007a: 145 A: NP
subgenus Pseudocartorhynchites Legalov, 2007a: 146 type species Cartorhynchites beijinensis Legalov, 2007
beijinensis Legalov, 2007a: 146 A: BEI

genus Cneminvolvulus Voss, 1960c: 413 type species Rhynchites rugosicollis Voss, 1920
subgenus Cneminvolvulus Voss, 1960c: 413 type species Rhynchites rugosicollis Voss, 1920
rugosicollis Voss, 1920: 165 (Rhynchites) A: FE FUJ HEI JA (Tsushima) LIA NC SC SHX
foochowensis Voss, 1960c: 414 (Involvulus)
mandli Voss, 1931c: 192 (Rhynchites)
subgenus Eocnemvulus Legalov, 2003b: 249 type species Cneminvolvulus cuprinus Legalov, 2003
desbrochersi Legalov, 2007a: 160 A: "Ind[ia]. Bor[ealis]."
longehirtus Fairmaire, 1882b: 67 (Rhynchites) A: "Inde boréale"
phohaensis Legalov, 2007a: 160 A: YUN
rasuwanus Legalov, 2007a: 160 A: NP
subgenus Metinvolvulus Legalov, 2003b: 249 type species Rhynchites haradai Kôno, 1940
haradai Kôno, 1940a: 67 (Rhynchites) A: FE HUB JA
lupulus Sawada, 1993: 63 (Involvulus) A: JA (Kyushu, Yakushima) NC
sundukovi Legalov, 2007a: 155 (Metinvolvulus) A: FE
subgenus Nigrorhynchites Legalov, 2007a: 158 type species Rhynchites celebicus Voss, 1938
aes Sawada, 1993: 62 (Involvulus) A: JA (Ryukyu)
comix Sawada, 1993: 61 (Involvulus) A: FE JA
subgenus Opacoinvolvulus Legalov, 2003b: 250 type species Rhynchites gensanensis Voss 1920
aterrimus Voss, 1938d: 156 (Rhynchites) A: FUJ YUN
cognatus Voss, 1958: 7 (Involvulus) A: FUJ ZHE
gensanensis Voss, 1920: 164 (Rhynchites) A: FE HEI LIA NC SC
atrocyaneus Ter-Minasian, 1944b: 15 (Rhynchites)
lepidus Voss, 1930b: 69 (Rhynchites) A: FUJ SCH
lepidus Voss, 1938d: 155 (Rhynchites) [HN]
planiusculus Voss, 1935b: 108 (Rhynchites) A: FUJ ORR
 RHYNCHITIDAE 121
pseudocivicus Legalov, 2007a: 163 (Opacoinvolvulus) A: SD
subgenus Pilosoinvolvulus Legalov, 2003b: 243 type species Rhynchites solutus Faust, 1886
decipiens Voss, 1938c: 149 {Rhynchites) A: YUN "China, Himalaya"
densatus Voss, 1938c: 149 (Rhynchites) A: AF KA
siwalikensis Voss, 1956d: 415 (Involvulus) A: UP
solutus Faust, 1886: 152 (Rhynchites) A: HP KA PA UP XIN
yunnanicus Voss, 1938c: 149 (Rhynchites) A: YUN
subgenus Protocnemvulus Legalov, 2007a: 161 type species Cneminvolvulus weibaoshanensis Legalov, 2007
weibaoshanensis Legalov, 2007a: 161 A: YUN
subgenus Pseudocnemvulus Legalov, 2007a: 161 type species Cneminvolvulus nanlingensis Legalov, 2007
nanlingensis Legalov, 2007a: 161 A: GUA
subgenus Pseudvulus Legalov, 2003 b: 246 type species Involvulus schusteri Formânek, 1912
conjunctulus Voss, 1959a: 69 (Involvulus) A: AF
liesenfeldti Voss, 1956d: 416 (Involvulus) A: UP "China"
meyeri Voss, 1937a: 275 (Rhynchites) A: YUN
nepalensis Legalov, 2007a: 157 (Pseudvulus) A: NP
schusteri Formânek, 1912: 51 (Rhynchites) A: XIZ

genus Cneminvolvulus, species incertae sedis

pudens Voss, 1938d: 157 (Rhynchites) A: UP

genus CyUorhynchites Voss, 1930b: 73 type species Rhynchites ursulus Roelofs, 1874
subgenus CyUorhynchites Voss, 1930b: 73 type species Rhynchites ursulus Roelofs, 1874
auricapillus Voss, 1930b: 77 (Rhynchites) A: CHQ HEB JIA JIX SCH SHN
homalinus Voss, 1930b: 76 (Rhynchites) A: YUN
hunanicus Legalov, 2007a: 170 A: HUN JIA
ursulus afghanus Legalov, 2003b: 256 A: AF PA
ursulus quercuphillus Legalov, 2007a: 170 A: FE NC SC
ursulus rostralis Voss, 1930b: 78 (Rhynchites) A: FUJ GUA HEB HEN HUN JIA LIA SCH XIN YUN ORR
ursulus ursulus Roelofs, 1874: 142 (Rhynchites) A: JA
subgenus Hypocyllorhynus Legalov, 2003b: 255 type species CyUorhynchites indicus Legalov, 2003
indicia Legalov, 2003b: 255 A: SD XIZ
subgenus Hyporhynchites Voss, 1935b: 101 type species Rhynchites lauraceae Voss, 1935
cheliensis Legalov, 2007a: 167 A: YUN
subgenus Pseudocyllorhynus Legalov, 2003b: 255 type species Rhynchites subcumulatus Voss, 1930
andrewesi Voss, 1930b: 75 (Rhynchites) A: SCH YUN ORR
cumulatus Voss, 1930b: 74 (Rhynchites) A: FUJ HEB HEI HEN JIA SCH SHA SHN YUN
nankingensis Legalov, 2007a: 169 A: JIA
shaanxiensis Legalov, 2007a: 169 A: SHA
spinicollis Voss, 1930b: 75 (Rhynchites) A: BTNP SD
subcumulatus Voss, 1930b: 76 (Rhynchites) A: YUN

genus Ecnomonychus Voss, 1933b: 55 type species Ecnomonychus usambaricus Voss, 1933
subgenus Himalayrhynchites Legalov, 2003b: 281 type species Rhynchites canus Faust, 1898
canus Faust, 1898: 297 (Rhynchites) A: HP KA SD UP
seriatopilosus Schilsky, 1906b: 26 (Rhynchites)
subgenus Maculinvolvulus Legalov, 2003b: 282 type species Rhynchites singularis Roelofs, 1874
fasciculosus Voss, 1942b: 91 (Rhynchites) A: FUJ
merkli Legalov, 2007a: 147 (Maculinvolvulus) A: TAI
singularis Roelofs, 1874: 147 (Rhynchites) A: FE FUJ JA

genus Eduardirhinus Legalov, 2003b: 252 type species Rhynchites tenuicornis Voss, 1938
tenuicornis Voss, 1938d: 154 (Rhynchites) A: SD
 122 RHYNCHITIDAE
genus Involvulus Schrank, 1798: 360 type species Involvulus metallicus Schrank, 1798 (= Curculio cupreus
Linnaeus, 1758)
subgenus Involvulus Schrank, 1798: 360 type species Involvulus metallicus Schrank, 1798 (= Curculio cupreus
Linnaeus, 1758)
Euvolvulus Reitter, 1916a: 264 type species Curculio cupreus Linnaeus, 1758
baojiensis Legalov, 2007a: 187 A: SHA
cupreus Linnaeus, 1758: 379 {Curculio) E: AU BE BU BY CR CTXZ DE EN FI FR GB GE HU IT LA LT LU
MCMDNLNRNTPLPTROSKSLSPSVSZUK N: AG A: ES FE HEI HUB IN JA KZ LIA MG NC
SCWS
dybofskyi Faust, 1882a: 289 (Rhynchites)
metallicus Schrank, 1798: 477
punctatus Herbst, 1797: 131 {Rhynchites)
purpureas Linnaeus, 1758: 378 (Curculio)
cylindricollis Schilsky, 1906a: 82 {Rhynchites) A: FE JA
purpureoviolaceus Voss, 1920: 164 {Rhynchites)
gemma Semenov & Ter-Minasian, 1937: 64 {Rhynchites) A: BEI SCH SHA YUN
yanmensis Legalov, 2009c: 221 A: YUN
subgenus Nepalorhynchites Legalov, 2007a: 187 type species Rhynchites ferox Faust, 1898
ferox Faust, 1898: 298 {Rhynchites) A: HP KA NP UP
pilosistriatus Voss, 1920: 163 {Rhynchites)
subgenus Parinvolvulus Legalov, 2003b: 297 type species Rhynchitespilosus Roelofs, 1874
Fujarhynchites Legalov, 2007a: 186 type species Rhynchites carinulatus Voss, 1942
Nigroinvolvulus Legalov, 2003b: 298 type species Rhynchites apionoides Sharp, 1889
Parinvolvoides Legalov, 2003b: 299 type species Parinvolvulus ussuriensis Legalov, 2003 (= Involvulus legalovi Alonso
Zarazaga, nom. nov.)
apionoides Sharp, 1889: 67 {Rhynchites) A: FE FUJ JA (Honshu)
berezovskyi Legalov, 2007a: 185 {Parinvolvulus) A: GAN
carinulatus Voss, 1942b: 93 {Rhynchites) A: FUJ
egenus Voss, 1933d: 109 {Rhynchites) A: ZHE
hauseri Wagner, 1907b: 323 {Rhynchites) A: QIN
immixtus Voss, 1969b: 287 [RN] A: FUJ
planiusculus Voss, 1941a: 243 {Rhynchites) [HN]
legalovi Alonso-Zarazaga, nom. nov. [see New Acts] A: FE
ussuriensis Legalov, 2003b: 300 (Parinvolvulus) [HN]
pilosus Roelofs, 1874: 145 (Rhynchites) A: FE JA NC SC "China"
placidus Sharp, 1889: 63 (Rhynchites) A: FE JA
przehvalskyi Legalov, 2007a: 186 {Parinvolvulus) A: GAN
schochrini Legalov, 2007a: 187 {Parinvolvulus) A: SC
zherichini Legalov, 2004b: 85 A: FE
subgenus Teretriorhynchites Voss, 1938c: 135 type species Curculio caemleus DeGeer, 1775
Aphlorhynchites Sawada, 1993: 50 type species Rhynchites laevior Faust, 1882 (= Rhynchites amabilis Roelofs, 1874)
Chinorhynchites Legalov, 2003b: 297 type species RhynchitesfitlvihirtusVoss, 1938
Chinorhynchitoides Legalov, 2007a: 182 type species Teretriorhynchites kangdingensis Legalov, 2007
amabilis Roelofs, 1874: 145 (Rhynchites) A: ANH BEI ES FE HEB HEI HEN JA JIL LIA MG NC SC SHA
SHN SHX WS
laevior Faust, 1882a: 284 (Rhynchites)
nudipennis Voss, 1929a: 25 (Rhynchites)
vossi Ter-Minasian, 1971a: 265 (Caenorhinus)
caemleus caemleus DeGeer, 1775: 251 {Curculio) E: AB AU BE BH BU BY CR CT CZ DE EN FR GB GE
GG HU IT LA LT LU MD NL NT PL RO SB SK SL SP ST SV SZ UK A: KZ TR
confinis Stephens, 1831: 202 (Rhynchites)
conicus Illiger, 1807a: 306 (Rhynchites)
caemleus iranensis Legalov, 2004a: 69 {Teretriorhynchites) A: IN
caemleus ussuriensis Voss, 1930b: 69 (Rhynchites) A: ANH BEI ES FE HEI NC LIA SC
dundai Legalov, 2007a: 184 {Teretriorhynchites) A: YUN
fitlvihirtus Voss, 1938c: 143 (Rhynchites) A: XIN
 RHYNCHITIDAE 123
funebris Sharp, 1889: 64 {Rhynchites) A: HUB JA
torticollis Faust, 1882a: 283 {Rhynchites) A: FE SC
kangdingensis Legalov, 2007a: 182 {Teretriorhynchites) A: SCH
kozlovi Legalov, 2007a: 184 {Teretriorhynchites) A: YUN
nigrocyaneus Legalov, 2007a: 183 {Teretriorhynchites) A: SCH
potanini Legalov, 2007a: 184 {Teretriorhynchites) A: SCH
pubescens Fabricius, 1775: 131 {Curculio) E: AR AU BE BH BU BY CR CT CZ DE FR GE GG HU IT KZ
MD PL PT RO SK SL SP ST SZ UK A: ES FE IN KI KZ TR WS
byctiscoidiceps Voss, 1938c: 143 {Rhynchites)
fallax Starck, 1889: 55 {Rhynchites)
flectirostris Pic, 1926a: 9 {Rhynchites)
multipunctatus Bach, 1858: 372 {Rhynchites)
parellinus Gyllenhal, 1833: 224 {Rhynchites)
virescens Thunberg, 1815:117 {Rhynchites) [DA]
szechuanensis Voss, 1956d: 414 A: SCH
xiahensis Legalov, 2007a: 184 {Teretriorhynchites) A: YUN
zhondiensis Legalov, 2007a: 185 {Teretriorhynchites) A: YUN

genus Japonorhynchites Legalov, 2003b: 222 type species Rhynchites sanguinipennis Roelofs, 1874
bicoloroides Legalov, 2007a: 132 A: YUN
bisulcatus Voss, 1922c: 283 {Rhynchites) A: TAI
caeligenus Sawada, 1993: 47 {Notocyrtus) A: JA (Kyushu)
sanguinipennis Roelofs, 1874: 144 {Rhynchites) A: FE JA SC
zhejiangensis Legalov, 2007a: 133 A: ZHE

genus Lasiorhynchites Jekel, 1860: 227 type species Rhynchites pubescens Fabricius sensu Herbst, 1797 (=
Rhynchites cavifrons Gyllenhal, 1833)
subgenus Coccygorrhynchites Prell, 1926: 288 type species Rhynchites sericeus Herbst, 1797
sericeus Herbst, 1797: 135 {Rhynchites) E: AB AR AU BE BU CR CZ DE FR GB GE GG HU IT LU MD NL
PL PT RO SK SL SP ST SV SZ N: AG A: SY TR
ophthalmicus Stephens, 1831: 200 {Rhynchites)
similis Curtis, 1837: 642 {Rhynchites)
splendidulus Kiesenwetter, 1852a: 626 {Rhynchites)
subgenus Lasiorhynchites Jekel, 1860: 227 type species Rhynchites pubescens Fabricius sensu Herbst, 1797 (=
Rhynchites cavifrons Gyllenhal, 1833)
cavifrons Gyllenhal, 1833: 226 {Rhynchites) [NP] E: AB AL AR AU BE BU CR CZ DE FR GB GE GG GR
HU IT LU MC MD NL PL RO SK SL SP ST SV SZ UK N: AG A: TR
cyanicolor Gyllenhal, 1833: 229 {Rhynchites)
viridis Geoffroy, 1785: 113 {Rhinomacer) [NO]
freidbergi Legalov & Friedman, 2007: 187 A: IS TR
subgenus Nelasiorhynchites Legalov, 2003b: 205 type species Rhynchites syriacus Desbrochers des Loges, 1869
brevirostris Roelofs, 1874: 148 {Rhynchites) A: FE JA NC
olivaceus Gyllenhal, 1833: 228 {Rhynchites) E: AU BE BH BU CR CT CZ DE FR GB GE HU IR IT LU MD
NL NR NT PL RO SK SL SP SV SZ UK N: AG
comatus Gyllenhal, 1833: 229 {Rhynchites)
pauciseta Wasmann, 1884: 183 {Rhynchites)
praeustus Boheman, 1845: 362 {Rhynchites) E: AU BH BU CR CZ FR GR HU IT MA ME PL RO SK SL TR
UK N: AG TU A: IS JO LE SY TR
caligatus Haliday, 1872: 125 {Rhynchites)
graecus Voss, 1933e: 136
lesinensis Voss, 1933e: 137
lividus Vitale, 1905:85
htridus Boheman, 1845: 363 {Rhynchites)
nigripennis Baudi di Selve, 1894:10 {Rhynchites)
semiruber Boheman, 1845: 362 {Rhynchites)
xanthomelas Voss, 1933e: 137
124 RHYNCHITIDAE
syriacus Desbrochers des Loges, 1869c: 388 (Rhynchites) E: GR A: IS SY
kindermanni Desbrochers des Loges, 1869c: 389 (Rhynchites)
ussuriensis Legalov, 2002d: 1523 A: FE SC SHA
vaucheri Desbrochers des Loges, 1897a: 177 (Rhynchites) E: SP N: MO
subgenus Stenorhynchites Voss, 1932a: 157 type species Curculio caeruleocephalus Schaller, 1783
caeruleocephalus Schaller, 1783: 282 (Curculio) E: AU CZ FR GE GR HU IT LT LU NT PL PT SK SP ST SV
SZ UK "Yugoslavia" N: AG MO
cyanocephalus Herbst, 1783: 72 (Curculio)
testaceus Harter, 1784: 188 (Rhinomacer)

genus Mecorhis Billberg, 1820: 39 type species Curculio ungaricus Herbst, 1783
subgenus Heterorhynchites Voss, 1932c: 171 type species Caenorhinus incertus Voss, 1932
Eosawadaia Legalov, 2004a: 66 type species Rhynchites subtectus Vos, 1938
Pseudorhinus Legalov, 2003b: 269 [HN] type species Rhynchites carinensis Faust, 1895
Sawadaia Alonso-Zarazaga & Lyal, 1999: 41 type species Rhynchites azureus Olivier, 1807
Vietnamorhynchites Legalov, 2003b: 272 type species Heterorhynchites medvedevi Legalov, 2003
alcyonea Pascoe, 1875: 391 (Rhynchites) A: "India"
arguta Faust, 1882a: 288 (Rhynchites) A: SD
bhamoensis Faust, 1895b: 164 (Rhynchites) A: YUN ORR
copiosa Voss, 1938d: 167 (Rhynchites) A: SDUP
satrapa Voss, 1938d: 163 (Rhynchites) A: "China"
semenovi Legalov, 2007a: 196 (Heterorhynchites) A: SCH
pristis G. A. K. Marshall, 1924: 283 (Rhynchites) A: UP ORR
subgenus Mecorhis Billberg, 1820: 39 type species Curculio ungaricus Herbst, 1783
Homalorhynchites Voss, 1938c: 136 type species Curculio ungaricus Herbst, 1783
ungarica Herbst, 1783: 71 (Curculio) E: AB AL AR AU BH BU BY CR CZ GE GG GR HU IT MC MD PL
SK SL ST TR UK A: IN IQ IS KZ SY TR WS
marginata Schrank, 1798:476 (Involvulus)
ursus Gebier, 1833: 292 (Rhynchites)
subgenus Mirabrhynchites Legalov, 2003b: 273 type species Mirabrhynchites kabakovi Legalov, 2003
mirifica Legalov, 2007a: 155 (Mirabrhynchites) A: YUN
subgenus Philorectus Voss, 1924b: 40 type species Philorectus insolitus Voss, 1924
Mirabinvolvulus Legalov, 2003b: 274 type species Mirabinvolvulus orlovi Legalov, 2003
insolita Voss, 1924b: 40 (Philorectus) A: SD XIZ ORR
sjaomonjanica Legalov, 2004a: 68 (Philorectus) A: YUN
subgenus Pseudomechoris Legalov, 2003a: 71 type species Rhynchites aethiops Bach, 1854
aethiops Bach, 1854: 172 (Rhynchites) [RN] E: AB AL AR AU BH BU CR CT CZ FR GE HU IT MC MD PL
RO SK SL ST SZ TR UK A: TR
crassicornis Gmelin, 1790: 1759 (Curculio)
juraensis Voss, 1969b: 250 (Involvulus)
planirostris Gyllenhal, 1833: 216 (Rhynchites)
subgenus Thompsonirhinus Legalov, 2003b: 267 type species Rhynchitesplumbeus Roelofs, 1874
Athompsonirhinus Legalov, 2003b: 270 type species Rhynchites subplumbeus Voss, 1938
Eorhinchus Legalov, 2007a: 194 type species Rhynchites simulons Voss, 1924
Maculinvoles Legalov, 2003b: 270 type species Rhynchites kiritshenkoi Ter-Minasian, 1944
Pseudorhinoides Legalov, 2007a: 192 type species Thompsonirhinus sichuanensis Legalov, 2007
amicta Voss, 1938c: 146 (Rhynchites) A: SD ORR
annapurnae Legalov, 2007a: 192 (Thompsonirhinus) A: NP
coerulescens coerulescens Voss, 1938d: 163 (Rhynchites) A: HKG
coerulescens tsinpinensis Legalov, 2007a: 192 (Thompsonirhinus) A: YUN
cyanescens Voss, 1939a: 71 (Rhynchites) [RN] A: SD
cyanicolor Voss, 1938d: 166 (Rhynchites) [HN]
eduardi Legalov, 2002c: 91 (Thompsonirhinus) A: JIX YUN XIN
obscurus Voss, 1938d: 166 (Rhynchites) [HN]
gentilis Voss, 1930b: 72 (Rhynchites) A: UP JLX YUN ZHE
gracilis Voss, 1938d: 165 (Rhynchites) A: UP
 RHYNCHITIDAE 125
indubia Voss, 1930b: 71 (Rhynchites) A: YUN ZHE
kundensis Legalov, 2007a: 192 (Thompsonirhinus) A: GAN
leucoscutellata Voss, 1932b: 71 {Rhynchites) A: YUN
mandschurica Voss, 1939c: 396 (Rhynchites) A: FE FUJ HEI JIL LIA YUN
kiritschenkoi Ter-Minasian, 1944c: 26 (Rhynchites)
continentalis Voss, 1958: 8 (Invotvulus)
obsita Voss, 1930b: 70 (Rhynchites) A: FUJ YUN
pilositessellata Voss, 1922c: 281 (Rhynchites) A: TAI
plumbea Roelofs, 1874: 143 (Rhynchites) A: FE HUB JA JIX NC NE SC SCH ZHE
sichuanensis Legalov, 2007a: 192 (Thompsonirhinus) A: SCH
simulons Voss, 1924b: 43 (Rhynchites) A: YUN

genus Metallorhynchites Legalov, 2007a: 164 type species Rhynchites aereus Voss, 1938
chalceus Voss, 1938c: 149 (Rhynchites) A: HP UP

genus Metarhynchites Voss, 1923: 489 type species Rhynchites arduus Voss, 1923
subgenus Metarhynchites Voss, 1923: 489 type species Rhynchites arduus Voss, 1923
fukienensis Voss, 1941a: 242 (Rhynchites) A: FUJ
schenklingi Voss, 1922c: 282 (Rhynchites) A: TAI
suborichalceus Voss, 1932d: 300 (Rhynchites) A: YUN

genus Montaninvolvulus Legalov, 2003b: 259 type species Rhynchites bucklandiae G. A. K. Marshall, 1924
subgenus Montaninvolvulus Legalov, 2003b: 259 type species Rhynchites bucklandiae G. A. K. Marshall, 1924
aeneoniger Faust, 1890e: 426 (Rhynchites) A: GAN GUX NE
bucklandiae G. A. K. Marshall, 1924: 282 (Rhynchites) A: SD
gatlangensis Legalov, 2007a: 165 A:NP
melancholicus Voss, 1938d: 154 (Rhynchites) A: SD ORR
liveus Voss, 1938d: 165 (Rhynchites)
potanini Legalov, 2007a: 165 A: GAN
xilingensis Legalov, 2007a: 165 A: SCH

genus Neocoenorrhinus Voss, 1952: 182 type species Rhynchites germanicus Herbst, 1797
subgenus Neocoenorhinidius Legalov, 2003b: 216 type species Rhynchites pauxillus Germar, 1824
abeillei Desbrochers des Loges, 1869c: 420 (Rhynchites) E: FR GR IT
assimilis Roelofs, 1874: 146 (Rhynchites) A: FE JA NC
cribrum Desbrochers des Loges, 1875b: 210 (Rhynchites) E: GR TR A: IS LE SY TR
cyaneus Formânek, 1911: 206 (Rhynchites)
stierlini Voss, 1938a: 64 (Caenorhinus)
cuprinus Chevrolat, 1860b: 128 (Rhynchites) N: AG
interpunctatus Stephens, 1831: 201 (Rhynchites) E: AU BE BU CR CT CZ DE EN FR GB GE HU IT LU MD
NL PL PT SB SK SL SP ST SV SZ TR UK "Transcaucasia" N: AG
confiisus Desbrochers des Loges, 1869c: 363 (Rhynchites)
longirostris Stephens, 1831: 202 (Rhynchites)
interruptus Voss, 1920: 168 (Rhynchites) A: FE JA (Hokkaido, Honshu)
lukjanovitschi Ter-Minasian, 1944c: 25 (Coenorrhinus)
italicus Voss, 1929a: 24 (Caenorhinus) E: BH FR GR IT
mekongensis Legalov, 2007a: 128 A: YUN
pauxillus Germar, 1824: 186 (Rhynchites) [NP] E: AB AR AU BE BH BU CR CT CZ DE EN FR GB GE GG
HU IT LA LT LU MD ME NL NT PL RO SB SK SL SP ST SV SZ TR UK A: IN KZ TM TR WS ORR
atrocaeruleus Stephens, 1831: 201 (Rhynchites)
caeruleus Geoffroy, 1785: 114 (Rhinomacer) [NO]
livescens Voss, 1929a: 25 (Caenorhinus)
pauculus Voss, 1933e: 134 (Caenorhinus)
persicus Gyllenhal, 1837:164 (Rhynchites)
persicus Gyllenhal, 1839: 330 (Rhynchites) [HN]
tulcidorsum Schrank, 1798: 477 (Involvulus) [DA]
126 RHYNCHITIDAE

pseudocribrum graecus Legalov, 2007a: 129 A: GR

pseudocribrum pseudocribrum Legalov, 2002a: 217 (Temnocerus) E: AB TR A: IN TR
thomsoni Faust, 1884a: 191 (Rhynchites) A: FE H EI LIA MG SHA
subgenus Neocoenorrhinus Voss, 1952: 182 type species Rhynchites germanicus Herbst, 1797
germanicus Herbst, 1797: 134 (Rhynchites) E: AB AL AU BE BH BU BY CR CT CZ DE EN FI FR GB GE
GG GR HU IR IT LA LT LU MD NL NR NT PL RO SB SK SL SP ST SV SZ TR UK A: ES FE KZ MG
TR WS
rugipennis Stephens, 1831: 202 (Rhynchites)
subgenus Schoenitemnus Legalov, 2003a: 69 type species Rhynchites minutus Herbst, 1797
minutus Herbst, 1797: 135 (Rhynchites) E: AB AU BE BH BU BY CR CT CZ DE EN FR GB GE GG GR HU
I T L U M C M D M E N L N R P L P T R O S K S L S P S T S V S Z U K N: AG A : I N T R
aeneovirens Marsham, 1802: 239 (Curculio)
fragariae Gyllenhal, 1833: 233 (Rhynchites)
longirostris Bach, 1854: 175 (Rhynchites) [HN]
obscurus Gyllenhal, 1833: 215 (Rhynchites)
smeraldinus O. G. Costa, 1839: 11 (Attelabus)

genus Orrhynchites Legalov, 2007a: 154 type species Rhynchites comosellus Voss, 1938 (p. 159) (= Rhynchites
consimilis Voss, 1938)
consimilis Voss, 1938d: 166 (Rhynchites) A: NP SD UP YUN
comosellus Voss, 1938d: 159 (Rhynchites)

genus Perrhynchites Voss, 1953: 45 type species Rhynchites aereipennis Desbrochers des Loges, 1895
aereipennis Desbrochers des Loges, 1895d: 94 (Rhynchites) A: LE SY TR
lodosi Voss, 1973: 37 (Rhynchites)
bellus Fremuth & Legalov, 2002: 215 E: TR A: TR

genus Perrhynchitoides Legalov, 2007a: 131 type species Caenorhinus brunneus Voss, 1932
brunneus Voss, 1932c: 176 (Caenorhinus) A: "China bor."
pelliceus Faust, 1890e: 427 (Rhynchites) A: GAN SCH

genus Proelautobius Legalov, 2007a: 140 type species Involvulus dentatus Voss, 1953
dentatus Voss, 1953: 42 (Involvulus) A: FUJ
erythropterus Voss, 1922c: 282 (Rhynchites) A: TAI
fujianicus Legalov, 2007a: 140 A: FUJ
hunanicus Lergalov, 2007a: 140 A: HUN
indochinensis Voss, 1939a: 61 (Rhynchites) [RN] A: "China" ORR
gracilicornis Voss, 1938c: 138 (Rhynchites) [HN]
jianxiensis Legalov, 2007a: 141 A: JTX
shaowuensis Legalov, 2007a: 141 A: FUJ
yunnanicus Legalov, 2003b: 237 (Elautobius) A: YUN

genus Pustulorhimis Legalov, 2003b: 275 type species Pustulorhinus yunnanicus Legalov, 2003
yunnanicus Legalov, 2003b: 275 A: YUN

genus Rhodocyrtus Alonso-Zarazaga & Lyal, 1999: 42 [RN] type species Rhynchites cribripennis Desbrochers
des Loges, 1869
Notocyrtus Desbrochers des Loges, 1908c: 78 [HN] type species Rhynchites cribripennis Desbrochers des Loges, 1869
cribripennis Desbrochers des Loges, 1869c: 355 (Rhynchites) E: BU CR FR (Corse) GR IT ST TR A: CY IS
LE SY TR

genus Rhynchitallus Voss, 1960c: 415 type species Rhynchitallus cyclops Voss, 1960
cyclops Voss, 1960c: 416 A: HUB JIA
 RHYNCHITIDAE 127
genus Rhynchites D. H. Schneider, 1791: 83 type species Curculio bacchus Linnaeus, 1758
subgenus Rhynchites D. H. Schneider, 1791: 83 type species Curculio bacchus Linnaeus, 1758
Prorhynchites Voss, 1973: 40 type species Rhynchites purpureipennis Voss, 1973 (= Rhynchites slovenicus Purkynë,
1954)
bacchus Linnaeus, 1758: 381 {Curculio) E: AB AL AR AU BE BH BU BY CR CT CZ FR GE GG GR HU IT
LT LU MC MD ME NL NR NT PL PT RO SB SK SL SP ST SV SZ TR UK N: AG A: IN IS KZ TM TR
UZWSXTN
circassicus Voss, 1938d: 170
cupreatus Voss, 1929a: 26
jekeli Desbrochers des Loges, 1869c: 348
laetus Germar, 1824: 186
splendidus Krynicki, 1832: 144
Julgidus Faldermann, 1835: 420 A: BEI ES FE FUJ GAN GUI HEB HEI HKG HUB HUN HL JLX LIA MG
NIN NMO SCH SHA SHN SHX ZHE
bacchoides Voss, 1934: 82
confragosicollis Voss, 193 3d: 110
faldermanni Schoenherr, 1839: 326 [RN]
kozlovi Suvorov, 1915: 345
tygosanensis Voss, 1938d: 171
lenaeus Faust, 1891c: 386 E: AB AR BU GG GR MC SK ST TR A: IS SY TR
asiaticus Desbrochers des Loges, 1908b: 65
slovenicus Purkynë, 1954: 168 E: AR SL TR A: IS TR
lenaeiformis Ter-Minasian, 1966b: 855
purpureipennis Voss, 1973: 39
subgenus Epirhynchites Voss, 1969b: 348 type species Rhynchites heros Roelofs, 1874
Colonnellinius Legalov, 2003b: 289 type species Rhynchites smymensis Desbrochers des Loges, 1869
Neorhynchites Voss, 1969b: 342 type species Rhynchites velatus LeConte, 1880
Pyrorhynchites Legalov, 2003a: 72 type species Rhynchites giganteus Krynicki, 1832 (= Rhynchites giganteus
Schoenherr, 1832)
Terminassianaeus Legalov, 2003b: 287 type species Rhynchites lopatini Ter-Minasian, 1968
Tshernyshevinius Legalov, 2003 a: 72 type species Rhynchites zaitzevi Kieseritzky, 1926
amygdali Iablokoff-Khnzorian, 1954: 147 {Rhynchites) E: AR
auratus Scopoli, 1763: 26 {Curculio) E: AB AR AU BE BH BU BY CR CT CZ EN FR GE GG GR HU IT LA
LT LU MC MD NL NT PL PT RO SK SL SP ST SZ TR UK A: AF ES IN IQ KI KZ TD TM TR UZ XIN
WS
angorensis Pic, 1926a: 10
armeniacus Zajcev, 1937: 66
aurifer Olivier, 1791a: 564 {Curculio) [HN, RN]
chamaecerasi Ter-Minasian, 1950: 113
ferghanensis Nevskij, 1929:445
microcarpae Ter-Minasian, 1944a: 53 {Rhynchites)
friedmani Legalov, 2007b: 197 {Epirhynchites) A: IS
giganteus Schoenherr, 1832: 214 E: AB AR AU BU CR CT CZ FR GG GR HU IT MD NT RO SK SL ST TR
UK A: IN KZ TR UZ "China"
giganteus Krynicki, 1832: 144 [HN]
giganteus Gyllenhal, 1833: 218 [HN]
emgei Stierlin, 1890:165
pyri Motschulsky, 1853: 80
rectirostris Gyllenhal, 1833: 220
versicolor O. G. Costa, 1839: 10 {Attelabus)
vestitus Rey, 1893b: 73
hajastanicus Ter-Minasian, 1965: 380 E: AR ST A: TR
heros Roelofs, 1874: 141 A: BEI ES FE FUJ GUA GUI GUX HEB HEI HEN HUB HUN JA JIA JIL JIX LIA
MG NC NIN NMO SC SCH SHA SHN SHX XIN YUN ZHE
foveipennis Fairmaire, 1888: 136
ignitus Voss, 1953: 43
koreanus Kôno, 1926a: 89
mongoliens Voss, 1930b: 78
128 RHYNCHITIDAE
sumptuosus Roelofs, 1875b: cxxxii
lopatini Ter-Minasian, 1968: 1865 A: TM UZ
pathcius Voss, 1959a: 70 A: AF
sarafschanicus Voss, 1929a: 27 A: AF TD TM UZ
nuristanicus Voss, 1959a: 71
smyrnensis Desbrochers des Loges, 1869c: 342 A: IN IS JO SY TM TR
trojanus Gyllenhal, 1839: 324 E: GR TR A: CY IS JO TR
maximus Desbrochers des Loges, 1869c: 338
turcicus Legalov, 2003a: 73 A: TR
zaitzevi Kieseritzky, 1926: 148 A: IN
zherichini Legalov, 2004a: 68 {Epirhynchites) A: ES
genus Rhynchites, species incertae sedis
alliariae Linnaeus, 1758: 378 {Curculio) E: SV
icosandriae Scopoli, 1763: 28 {Curculio) E: SL
purpureus Linnaeus, 1758: 378 {Curculio) E: "Europa"
rhedi Schrank, 1781: 106 {Curculio) E: AU
rugosus Gistel, 1857: 584 [HN] E: GE

genus Rubrrhynchites Legalov, 2007a: 130 type species Rubrrhynchites curvirostris Legalov, 2007
curvirostris Legalov, 2007a: 130 A: YUN

genus Taiwanorhynchites Legalov, 2007a: 154 type species Rhynchites impressicollis Voss, 1938
impressicollis Voss, 1938c: 138 {Rhynchites) A: TAI

genus Tatianaerhynchhes Legalov, 2002b: 87 type species Curculio aequatus Linnaeus, 1767
aequatus Linnaeus, 1767: 607 {Curculio) E: AB AL AR AU BE BH BU CR CT CZ DE EN FI FR GB GE GG
GR HU IT KZ LA LT LU MC MD ME NL NR NT PL PT RO SB SK SL SP ST SV SZ TR UK A: CY IN
IS JO KZ SY TM TR
bicolor Rossi, 1792: 44 {Attelabus) [HN]
creticus Voss, 1955: 271 {Merhynchites)
interstitialis Desbrochers des Loges, 1869c: 354 {Rhynchites)
phryganophilus Iablokoff-Khnzorian, 1956: 223 {Caenorrhinus)
ruber Geoffroy, 1785: 113 (Rhinomacer)
ruber Fairmaire, 1859a: c {Rhynchites)
semiruber Stierlin, 1864:492 {Rhynchites)

genus Temnocerus Thunberg, 1815: 110 type species Attelabus planirostris Fabricius, 1801 = Curculio nanus
Paykull, 1792
Paratemnocerus Legalov, 2003b: 209 type species Rhynchites subglaber Desbrochers des Loges, 1897
Pselaphorhynchites Schilsky, 1903: F type species Curculio nanus Paykull, 1792
coeruleus Fabricius, 1798: 163 {Attelabus) E: AB AN AU BE BH BU BY CR CT CZ DE EN FI FR GB GE GG
GR HU IR IT KZ LA LT LU MD NL NR NT PL PT RO SB SK SL SP ST SV SZ TR UK N: AG A: ES FE
KI KZ TR WS
tomentosus Gyllenhal, 1839: 335 {Rhynchites)
uncinatus C. G. Thomson, 1865: 36 {Rhynchites)
daliangensis Legalov, 2009a: 31 A: SCH
dundai Legalov, 2006b: 7 A: YUN
elaeagni Korotyaev & Legalov, 2006: 126 A: KZ
japonicus Morimoto, 1958b: 97 {Pselaphorhynchites) A: ES FE JA (Hokkaido, Honshu) NE
longiceps C. G. Thomson, 1888: 1203 {Rhynchites) E: AU BE CT CZ DE FI FR GB GE IT LA LT LU NL NR
NT PL SK SV SZ UK A: ES KZ WS
harwoodi Joy, 1911: 270 {Rhynchites)
morimotoi Sawada, 1997: 117 {Pselaphorhynchites) A: JA (Kyushu)
terminassianae Legalov, 2003c: 409
nanus Paykull, 1792: 136 {Curculio) E: AN AU BE BH BU BY CR CT CZ DE EN FI FR GB GE HU IT LA
L T L U N L N R N T P L P T R O S L S P S T S V S Z U K N: AG A: ES KI KZ MG TM TR WS
 ATTELABIDAE 129
cylindricus Stephens, 1831: 199 (Rhvnchites)
planirostris Fabricius, 1801: 425 (Attelabus)
rubripes Reitter, 1916a: 263 (Rhynchites) A: ES FE KZ LIA MG SHX
mongolianus Voss, 1967: 253 (Pselaphorhynchites)
yasumatsui Kôno & Morimoto, 1958: 26 (Pselaphorhynchites)
semicyaneus Bedel, 1884b: 208 (Rhynchites) N: AG
sibiricus Legalov, 2006c: 18 A: ES
subglaber Desbrochers des Loges, 1897a: 177 (Rhynchites) E: CT ST A: ES KI KZ MG WS
angustifrons Voss, 1932c: 179 (Rhynchites)
tibialis Voss, 1967: 255 (Pselaphorhynchites)
yunnanicus Legalov, 2003b: 212 A: YUN

family Rhynchitidae, nomina dubia

Curculio icosandriae Scopoli, 1763: 28 E: SL
Curculio rhedi Schrank, 1781: 106 E: AU
Curculio violascens Schrank, 1781: 105 E: AU
Rhinomacerfitlgidus Geoffroy, 1785: 116 E: FR
Rhynchites rugosus Gistel, 1857: 584 [HN] E: GE

family ATTELABIDAE Billberg, 1820

tribe Apoderini Jekel, 1860
genus Anisonychus Voss, 1927: 41 type species Apoderus interstitialis Voss, 1922
subgenus Anisonychidius Legalov, 2007a: 361 type species Anisonychus mirabilis Legalov, 2003
mirabilis Legalov, 2003b: 557 A: SCH YUN ORR
rugosus Voss, 1927: 42 (Apoderus) A: HAI

genus Apoderus Olivier, 1807:12 type species Attelabus coryli Linnaeus, 1758
Granulapoderus Legalov, 2003b: 553 type species Apoderus rugicollis Schilsky, 1906
Konoapoderus Legalov, 2003b: 551 type species Apoderus jekelii Roelofs, 1874
coryli Linnaeus, 1758: 387 (Attelabus) E: AL AU BE BH BU BY CR CT CZ DE EN FI FR GB GE GR HU IT
LA LT LU MC MD NL NR NT PL PT RO SK SL SP ST SV SZ UK A: BEI ES FE FUJ GAN HEB HEI JA
JIA JIL JIX LIA MG NC NE NMO SC SCH SHA SHX TAI YUN XIN WS
avellanae Linnaeus, 1767: 619 (Attelabus)
collaris Scopoli, 1763: 25 (Curculio)
dauricus Laxmann, 1770: 595 (Attelabus)
denigratus Gmelin, 1790: 1809 (Attelabus)
excoriatoruber DeGeer, 1775: 257 (Curculio)
gibbicollis Faust, 1882a: 295
homalinus Voss, 1927: 28
kamtschaticus Motschulsky, 1845b: 380
morio Bonelli, 1812: 175
wgerGortani & Grandi, 1904: 171 (Attelabus) [HN]
nigricollis Faust, 1882a: 295 [HN]
nigrifrons Delta Beffa, 1912: 1
superans Faust, 1882a: 295
fidus Faust, 1890e: 421 A: BEI FUJ GAN HEB JIX SCH
horribilis Legalov, 2003b: 553 A: SCH
jekelii Roelofs, 1874: 132 A: BEI ES FE HEN HUB JA JIL NC SC
extensus Voss, 1927: 33
hiratai Uno, 1939:578
okamotonis Kôno, 1930b: 58
rufithorax Kolbe, 1886: 215
sharpi Voss, 1922b: 174 [RN]
uniformis Sharp, 1889: 51 [HN]
kresli Legalov, 2003b: 549 A: SHA
ludyi Reitter, 1890b: 174 E: IT A: SY
130 ATTELABIDAE
a/m Chinaglia, 1908:29
carnicus Gortani & Grandi, 1904: 171 (Attelabus)
pseudofidus Legalov, 2003b: 551 A: SCH
rugicollis Schilsky, 1906a: 76 A: BEI FUJ GUI GUX HUB SCH SD SHA YUN
kuatunamis Voss, 1949: 160
tuberculimerus Voss, 1930b: 84
volkovitshi Legalov, 2003b: 551 A: YUN

genus Centrocorynus Jekel, 1860: 167 type species Apoderus scutellaris Gyllenhal, 1833
subgenus Centrocorynus Jekel, 1860: 167 type species Apoderus scutellaris Gyllenhal, 1833
propinquus Voss, 1929b: 96 A: SHA ORR
rujulus Voss, 1929b: 97 A: SD ORR
scutellaris Gyllenhal, 1833: 191 (Apoderus) A: SD SHA TAI YUN ZHE ORR

genus Compsapoderus Voss, 1927: 62 type species Attelabus erythropterus Gmelin, 1790
subgenus Compsapoderus Voss, 1927: 62 type species Attelabus erythropterus Gmelin, 1790
Compsapoderopsis Legalov, 2003b: 528 type species Apoderus dimidiatus Faust, 1890
continental Legalov, 2007a: 338 A: ANH FUJ GUI HUN JIA JIX NC SCH
dimidiatus Faust, 1890e: 421 (Apoderus) A: GAN HUB NO SCH
dispar Voss, 1928: 117 (Apoderus) A: "India"
erythrogaster Snellen van Vollenhoven, 1865: 165 (Apoderus) A: HUB JA NC SC YUN
rufiventris Roelofs, 1874: 135 (Apoderus)
shimauchiensis Kôno, 1930b: 67 (Apoderus)
erythropterus Gmelin, 1790: 1809 (Attelabus) E: AB AU BY CR CT CZ FR GE HU IT LA LT NT PL SK SV
SZ UK A: ANH BEI ES FE GAN HEB HEI HUB JA JIA JIL KZ LIA MG NC NMO SC SCH SHG SHN
WS ZHE
atricolor Faust, 1887a: 28 (Apoderus)
bicolor L. Redtenbacher, 1868: 161 (Apoderus)
frontalis Faust, 1882a: 294 (Apoderus)
intermedius Illiger, 1794: 615 (Attelabus)
politus Gebier, 1825: 50 (Apoderus)
foveipennis Suffrian, 1870: 228 (Apoderus) A: JA
niterts Roelofs, 1874: 133 (Apoderus)
geminus Sharp, 1889: 50 (Apoderus) A: FE GUI HUB HUN JA (Hokkaido, Honshu) NC SC SCH YUN
minimus Roelofs, 1874: 136 (Apoderus) A: ANH FUJ GUI HUN JA JIA JTX NC SCH
mushanus Kôno, 1930b: 59 (Apoderus) A: TAI
subgenus Paracompsus Legalov, 2003b: 537 type species Apoderus lepidulus Voss, 1927
afghanus Legalov, 2003b: 537 (Paracompsus) A: AF
bistriolatus Faust, 1898: 296 (Apoderus) A: AF HP PA SD UP ORR
incanus Stebbing, 1903: 189 (Apoderus)
nigrimaculatus Voss, 1920: 174 (Apoderus)
corallinoides Legalov, 2007a: 347 (Paracompsus) A: SD ORR
lepidulus Voss, 1927: 74 (Apoderus) A: SD XIZ
lunanfensis Legalov, 2007a: 347 (Paracompsus) A: SCH
luteibasis Voss, 1935d: 102 (Apoderus) A: NP UP ORR
micros Legalov, 2007a: 348 (Paracompsus) A: YUN
nigrofasciatus Pajni, Haq & Gandhi, 1987: 10 (Apoderus) A: AP SCH YUN
nigroscutellaris Legalov, 2007a: 348 (Paracompsus) A: NP
pygidialis Voss, 1927: 87 (Apoderus) A: "Himalaya"
seclusus Faust, 1898: 296 (Apoderus) A: XIZ
varions Voss, 1928: 118 (Apoderus) A: HP

genus Cycnotrachelodes Voss, 1955: 273 type species Apoderus roelofsi Harold, 1877
Pseudcycnolodes Legalov, 2003b: 571 type species Apoderus coeruleatus Faust, 1895
camphoricola Voss, 1924b: 45 (Cycnotrachelus) A: ANH FUJ GUA GUX HAI HEN JIA SHX YUN ORR
coeruleatus Faust, 1895b: 159 (Apoderus) A: BT FUJ YUN ORR
 ATTELABIDAE 131
subcoeruleus Voss, 1929b: 117 (Cycnotrachelus)
cyanopterus Motschulsky, 1861: 22 {Apoderus) A: BEI ES FE FUJ HEB HEI JA JIA JIL LIA NC SC SHX
YUNZHE
coloratus Faust, 1882a: 292 {Apoderus)
formosanus Voss, 1924c: 281 {Cycnotrachelus) A: TAI
roelofsi Harold, 1877: 358 {Apoderus) A: ES FE JA (Honshu, Shikoku) NC NE SC
sitchuanensis Legalov, 2003b: 571 A: SCH YUN ZHE
ussuriensis Voss, 1931c: 194 {Cycnotrachelus) A: FE HEI HUN LIA NC NE SC ZHE

genus Cycnotrachelus Jekel, 1860: 160 type species Apoderus flavotuberosus Jekel, 1860
flavoguttatus Voss, 1929b: 128 A: FUJ YUN ORR
flavonotatus Voss, 1935a: 516 A: YUN ORR
flavotuberosus Jekel, 1860: 163 {Apoderus) A: SD TAI YUN ORR
diversenotatus Pic, 1928a: 30 (Apoderus)

genus Hamiltonius Alonso-Zarazaga & Lyal, 1999: 46 type species Apoderus sexguttatus Voss, 1927
subgenus Hamiltonius Alonso-Zarazaga & Lyal, 1999: 46 type species Apoderus sexguttatus Voss, 1927
Parahamiltonius Legalov, 2007a: 349 type species Apoderus trinotatus Faust, 1883
sexguttatus Voss, 1927: 40 {Apoderus) A: FUJ GUA GUX YUN ORR
subgenus Korotyaevirhinus Legalov, 2003b: 538 type species Attelabus notatus Fabricius, 1792
notatus Fabricius, 1792: 385 {Attelabus) A: GUA GUX HKG HUN SCH YUN ORR
constans Faust, 1883a: 466 {Apoderus)

genus Heterapoderus Voss, 1927: 52 type species Apoderus sulcicollis Jekel, 1860
subgenus Heterapoderopsis Legalov, 2003b: 542 type species Apoderus pauperulus Voss, 1927
bicallosicollis Voss, 1932b: 74 {Apoderus) A: FUJ GUI HEB HEN JIX SCH YUN
cantonensis Voss, 1927: 55 {Apoderus) A: GUA JIA
pauperulus Voss, 1927: 54 {Apoderus) A: FUJ GUA GUI GUX HUN JIA SCH YUN ZHE ORR
subgenus Heterapoderus Voss, 1927: 52 type species Apoderus sulcicollis Jekel, 1860
Aheterapoderus Legalov, 2007a: 356 type species Apoderus brachialis Voss, 1924
Eoheterapoderus Legalov, 2003b: 547 type species Apoderus geniculatus Jekel, 1860
Neheterapoderus Legalov, 2003b: 543 type species Apoderus macropus Pascoe, 1883
Pseudoheterapoderus Legalov, 2003b: 546 type species Apoderus crenatus Jekel, 1860
ageniculatus Legalov, 2007a: 356 A: GUA FUJ SCH
brachialis Voss, 1924c: 282 {Apoderus) A: TAI
crenatus Jekel, 1860: 173 {Apoderus) A: GUX YUN ORR
franzi Voss, 1974: 47 {Apoderus) A: NP
geniculatus Jekel, 1860: 174 {Apoderus) A: FUJ GUA GUI GUX HEB HEN HUN JIA JIX SCH SHG YUN
ZHE ORR
monchadskii kathmanduensis Legalov, 2007a: 356 A: NP
monchadskii monchadskii Legalov, 2007a: 356 A: YUN
piceus Voss, 1927: 54 (Apoderus) A: FUJ GUA GUX HAI HKG ZHE
sulcicollis Jekel, 1860: 174 (Apoderus) A: FUJ GUA GUI GUX HEI HUN JIA SCH SHA SHG YUN ORR
subgenus Macroderites Legalov, 2003b: 539 type species Macroderites nepalensis Legalov, 2003
Amacroderites Legalov, 2007a: 350 type species Macroderites orchinensis Legalov, 2007
nepalensis Legalov, 2003b: 539 (Macroderites) A: NP
orchinensis Legalov, 2007a: 351 (Macroderites) A: FUJ
subgenus Microcorynus Legalov, 2003b: 539 type species Apoderus blandus Faust, 1895
blandus blandoloides Legalov, 2007a: 351 {Microcorynus) A: YUN
parunicus Legalov, 2003b: 540 {Microcorynus) A: YUN

genus Heterapoderus, species incertae sedis

theresae Pic, 1929a: 23 [DA] A: YUN
132 ATTELABIDAE
genus Leptapodents Jekel, 1860:169 type species Apoderus quadripunctatus Gyllenhal, 1833 (- Apoderus
pectoralis Thunberg, 1815) . . . », .
subgenus Leptapoderidius Legalov, 2007a: 341 type species Apoderus rubidus Motschulsky, 1860
bimaculatus Voss, 1929d: 372 {Apoderus) A: SHN
corallinus Legalov, 2007a: 342 A: YUN
frater Voss, 1927: 73 {Apoderus) A: FUJ HEN JIA SCH SHN ZHE
i mi talus Voss, 1941a: 248 {Apoderus)
ganlanbebsis Legalov, 2007a: 342 A: SCH YUN
himalayensis Legalov, 2007a: 342 A: NP
kansuensis Legalov, 2007a: 343 A: GAN
nigricans Legalov, 2007a: 343 A: SCH
nigroapicalus Jekel, 1860: 175 {Apoderus) A: FUJ GUA GUI GUX HUB HUN JIA JIX SCH SD SHA SHN
TAI YUN ZHE ORR
apicalis Faust, 1890c: 257 {Apoderus)
papei Voss, 1927: 62 {Apoderus)
omeishanicus Legalov, 2007a: 343 A: SCH
pseudocrucifer Legalov, 2007a: 343 A: YUN
rubidus Motschulsky, 1860: 172 {Apoderus) A: ES FE FUJ GUX HEB HEI HUB JA JIA NC SC SCH SHN
ZHE
rubidus Faust, 1882a: 294 {Apoderus) [HN]
rufescens Roelofs, 1874: 133 {Apoderus)
sejugatus Voss, 1924c: 283 {Apoderus) A: TAI
siaomonianensis Legalov, 2007a: 344 A: YUN
thibetanus Voss, 1927: 73 {Apoderus) A: SCH
tianmuensis Legalov, 2007a: 344 A: ANH ZHE
vossi Biondi, 2001: 225 {Apoderus) [RN] A: HUB SCH XIZ YUN ZHE
pedestris Voss, 1930b: 87 {Apoderus) [HN]
pedestroides Legalov, 2002c: 92 [RN]
yunlingensis Legalov, 2007a: 344 A: YUN
subgenus Leptapodents Jekel, 1860: 169 type species Apoderus quadripunctatus Gyllenhal, 1833 {= Apoderus
pectoralis Thunberg, 1815)
qffinis Schilsky, 1906a: 77 {Apoderus) A: BEI FE HUB JA NC SC SCH SHA YUN
cinctipectoralis Voss, 1930b: 86 {Apoderus)
subgenus Maculapoderus Legalov, 2003b: 535 type species Apoderus submaculatus Voss, 1927
submaculatus Voss, 1927: 82 {Apoderus) A: YUN ORR
subgenus Paraleptapoderus Legalov, 2003b: 534 type species Apoderus carbonicolor Motschulsky, 1860
carbonicolor Motschulsky, 1860: 171 {Apoderus) A: FE HEB HEI LIA NC SC
humerosus Motschulsky, 1860: 171 {Apoderus)
maurus Motschulsky, 1860: 171 {Apoderus)
praecellens Sharp, 1889: 46 {Apoderus) A: FE (Kuriles) HUB GUI JA (Honshu, Shikoku) NC SC SCH
subgenus Pseudapoderopsis Legalov, 2003b: 530 type species Leptapodents tamdaoensis Legalov, 2003
proprius Legalov, 2007a: 340 A: SCH
signa tus Voss, 1928: 118 {Apodems) A: SHG
subgenus Pseudoleptapoderus Legalov, 2003b: 535 type species Apoderus balteatus Roelofs, 1874
balteatus Roelofs, 1874: 135 {Apoderus) A: FE FUJ GUI HEB HEN HUB HUN JA NC SC SCH ZHE
dubatolovi Legalov, 2003b: 536 A: SCH
friedrichi Voss, 1956d: 422 {Apodems) A: HUB SCH TAI
inbalteatus Legalov, 2007a: 346 A: SCH YUN
nigroflavus Fairmaire, 1878: 129 {Apoderus) A: SHX
sinicus Voss, 1927: 77 {Apoderus) A: FUJ HEN JIA
subfasciatus Voss, 1929d: 373 {Apodems) A: YUN

genus Leptapodents, species incertae sedis

atronitidus Pic, 1929a: 24 {Apodems) A: JA "China"
atrovittatus Pic, 1929a: 24 {Apodems) A: "China"
 ATTELABIDAE 133
donckieri Pic, 1929a: 24 (Apoderus) A: "China"
luteoplicatus Pic, 1929a: 23 {Apoderus) A: "China"
proximus Voss, 1929b: 99 (Centrocorynus) A: SD
subcinctus Pic, 1929a: 24 (Apoderus) A: "China"
thoracicus Voss, 1958: 17 (Apoderus) A: FUJ

genus Mordkovitshirhinus Legalov, 2003b: 559 type species Mordkovitshirinus yunnanicus Legalov, 2003
yunnanicus Legalov, 2003b: 559 A: YUN

genus Paracycnotrachelus Voss, 1924b: 45 type species Attelabus cygneus Fabricius, 1801
breviusculus Voss, 1929b: 146 A: FUJ JIA SC SCH ORR
chinensis Jekel, 1860: 164 (Apoderus) A: ANH BEI ES FE FUJ GUA HAI HEB HEI HEN HKG HUB JA
(Honshu) JIA JIL JIX LIA NC NE QIN SC SCH SHA SHG SHN SHX TAI YUN ZHE
longiceps Motschulsky, 1860: 173 (Apoderus)
consimilis Voss, 1929b: 145 A: ANH BEI GAN HAI HEB HUB HUN JIA SHN ZHE
cygneus Fabricius, 1801: 417 (Attelabus) A: GAN GUA GUX JIA YUN ORR
siamensis Voss, 1929b: 137
montanus Jekel, 1860: 165 (Apoderus) A: FUJ HAI HKG HUB JIA YUN ORR

genus Paratrachelophorus Voss, 1924b: 44 type species Paratrachelophorus brachmanus Voss, 1924
Cyanotrachelophorus Legalov, 2007a: 375 type species CycnotrachelusfiikienensisVoss, 1949
Morphotrachelophorus Legalov, 2003b: 577 type species Paratrachelophorus marsi Legalov, 2003
Pseudoparaphorus Legalov, 2003b: 575 type species Apoderus potanini Faust, 1890
belokobylskü Legalov, 2007a: 373 A: ZHE
brachmanus Voss, 1924b: 44 A: NP SD XIZ ORR
daliensis Legalov, 2003b: 583 A: YUN
foveostriatus Voss, 1930b: 88 (Paracycnotrachelus) A: HAI YUN ZHE
fiikienensis Voss, 1949: 161 (Cycnotrachelus) A: FUJ
fiilvus Roelofs, 1874: 130 (Apoderus) A: JA
gigas Legalov, 2003b: 581 A: YUN ORR
kresli Legalov, 2003b: 581 A: NP
longicornis Roelofs, 1874: 131 (Apoderus) A: FE GUI HEB HUB JA NC NE SC TIA
marsi Legalov, 2003b: 581 A: YUN ORR
nodicornis Voss, 1924c: 280 A: FUJ TAI YUN
katonis Kôno, 1930b: 76
vossi Kôno, 1930b: 75
nodicornoides Legalov, 2003b: 583 A: FUJ GUA JIX ORR
potanini Faust, 1890e: 423 (Apoderus) A: FUJ GAN SCH SHA SHX ORR
sikkimensis Legalov, 2007a: 374 A: SD

genus Physapoderus Jekel, 1860: 170 type species Attelabus biguttatus Fabricius, 1801
subgenus Alexsandricorynus Legalov, 2003b: 561 type species Apoderus assamensis Boheman, 1845
assamensis Boheman, 1845: 354 (Apoderus) A: SCH YUN ORR
bilineatus Faust, 1883a: 464 (Apoderus)
subgenus Eocentrocorynus Legalov, 2003b: 561 type species Apoderus aemulus Faust, 1895
Formosusorynus Legalov, 2003b: 566 type species Centrocorynus gracilicornis Voss, 1929
Neocorynidius Legalov, 2003b: 567 type species Centrocorynus ruficlavis Voss, 1929
antennalis Legalov, 2007a: 367 (Eocentrocorynus) A: YUN
breviclavus Legalov, 2003b: 565 (Eocentrocorynus) A: YUN
fiisculusfusculus Voss, 1929b: 100 (Centrocorynus) A: YUN ORR
fusculus yunnanicus Legalov, 2003b: 566 (Eocentrocorynus) A: YUN
gracilicornis Voss, 1929b: 98 (Centrocorynus) A: FUJ GUA GUX HUB JIA HAI YUN ZHE ORR
huangi Legalov, 2007a: 367 (Eocentrocorynus) A: YUN
pokharensis Legalov, 2003b: 566 (Eocentrocorynus) A: NP
ruficlavis Voss, 1929b: 99 (Centrocorynus) A: FUJ HUB HUN YUN ORR
134 ATTELABIDAE

sichuanensis Legalov, 2007a: 367 (Eocentrocorynus) A: SCH

subgenus Physapoderus Jekel, 1860: 170 type species Attelabus biguttatus Fabncms, 1801
Aphrysapoderus Legalov, 2007a: 365 type species Apoderus pulchellus Pascoe, 1883
Eophrysapoderus Legalov, 2007a: 365 type species Apoderus crucifer Heller, 1922
Paracentrocorynus Voss, 1929b: 97 type species Attelabus biguttatus Fabricius, 1801
crucifer Heller, 1922a: 13 {Apoderus) A: FUJ GUA GUI GVX HAI QIN YUN ZHE ORR
langanus Pic, 1929a: 25 (Leptapoderus)
maculipennis Voss, 1929b: 108 (Centrocorynus)
genus Strigapoderus Jekel, 1860:169 type species Apoderus tranquebaricus Fabricius, 1798
subgenus Helleriapoderus Legalov, 2007a: 353 type species Apoderus nelligrinus Heller, 1908
subvariolaris Voss, 1935d: 100 (Apoderus) A: "India"
subgenus Strigapoderus Jekel, 1860: 169 type species Apoderus tranquebaricus Fabricius, 1798
scabrosus Voss, 1928: 116 (Apoderus) A: "China" ORR
tribe Apoderini, nomina dubia
Apoderus cyaneus Hope, 1831: 27 A: NP
Apoderus dauricus Motschulsky, 1860: 173 [HN] A: ES

tribe Attelabini Billberg, 1820

genus Attelabus Linnaeus, 1758: 387 type species Attelabus curculionoides Linnaeus, 1767 (= Curculio niten
Scopoli, 1763)
Algirolabus Legalov, 2003b: 436 type species Attelabus variolosus Fabricius, 1801
Chyphus Thunberg, 1815: 110 type species Attelabus curculionoides Linnaeus, 1767 (= Curculio nitens Scopoli, 1763)
Cyphus Bedel, 1883: 23 [HN] type species Attelabus curculionoides Linnaeus, 1767 (= Curculio nitens Scopoli, 1763)
Proattelabus Legalov, 2007a: 276 type species Cyphus sulcifrons Atgod-Vallon, 1895
Rhinomacer O. F. Müller, 1764: xiii [HN] type species Rhinomacer coryli O. F. Müller, 1776 (= Curculio nitens Scopoli,
1763)
chalybaeus K. Daniel & J. Daniel, 1898: 72 E: AB A: IN TM
cyanellus Voss, 1925b: 258 [RN] A: ES FE MG "China"
cyaneus Boheman, 1845: 358 [HN]
nitensiformis Kôno, 1927a: 37
subcyaneus Voss, 1931c: 192
metallicus X.-C. Zhang, 1995a: 208 A: NIN
nitens Scopoli, 1763: 25 (Curculio) E: AB AL AR AU BE BH BU BY CR CT CZ DE EN FI FR GB GE GG
GR HU IT LA LT LU MC MD ME NL NT PL PT RO SB SK SL SP ST SV SZ TR UK A: IN IS SY TM
TR
atricornis Mulsant & Guillebeau, 1855: 159
balcanicus K. Daniel & J. Daniel, 1898: 73
coccineus Geoffroy, 1785: 115 (Rhinomacer)
curculioniformis Schrank, 1798: 475 (Involvulus) [RN]
curculionoides Linnaeus, 1767: 619 (Attelabus)
fascicollis Reitter, 1916a: 267
hispanicus Jekel, 1860: 199
maculipes Küster, 1845b: 64
marginalis Pic, 1898a: 4
obscurior Pic, 1898a: 4
pulvinicollis Jekel, 1860: 199
rubricollis Reitter, 1916a: 267
sichuanensis Legalov & Liu, 2005: 126 A: SCH
sulcifrons Argod-Vallon, 1895: xxviii (Cyphus) E: AR BU GG GR MC A- TR SY
suturalis Jekel, 1860: 200 A: CY SY TR
variolosus Fabricius, 1801: 420 E: SP N: AG MO
foveipennis Jekel, 1860: 201

genus Attelabus, species incertae sedis

bicolor Gmelin, 1790: IS09 (Attelabus) E: "Europa"
 ATTELABIDAE 135
calibatus Townson, 1797: 473 {Attelabus) E: "Hungary"
efferans Townson, 1797: 473 {Attelabus) E: "Hungary"
ilicis O. G. Costa, 1839: 10 {Attelabus) E: FT:»
duodecimpunctatus Scopoli, 1763: 36 {Attelabus) E: "In Austriaco Littorali" (now CR IT SL)
quattuordecimpunctatus Scopoli, 1763: 37 {Attelabus) E: "In Austriaco Littorali" (now CR IT SL)

genus Catalabus Voss, 1925b: 214 type species Attelabus sexplagiatus Heller, 1922
subgenus Catalaboides Legalov, 2003b: 447 type species Paramecolabus kazantsevi Legalov, 2003
elegans Voss, 1933d: 113 {Paramecolabus) A: UP
kazantsevi Legalov, 2003b: 447 (Paramecolabus) A: SCH
nigrosuturalis Voss, 1930b: 80 {Paramecolabus} A: SCH
marieae Kresl, 2007: 127 {Paramecolabus) A: NP SD
pallidipennis Voss, 1925b: 220 (Paramecolabus) A: SD TAI ORR
rasuwanus Legalov, 2007a: 281 A: NP SD
simulatus G. A. K. Marshall, 1923: 283 (Attelabus) A: SD ORR
subgenus Catalabus Voss, 1925b: 214 type species Attelabus sexplagiatus Heller, 1922
quadriplagiatus Voss, 1953: 49 (Paramecolabus) A: FUJ HAI YUN

genus Cyrtolabus Voss, 1925b: 241 type species Attelabus christophi Faust, 1884
amitinus Voss, 1932b: 72 (Attelabus) A: YUN
chinensis Legalov, 2004a: 83 A: JLX
christophi Faust, 1884b: 471 (Attelabus) A: ES FE HEN JIX MG NE SC
mutus Faust, 1890e: 425 (Attelabus) A: FE GAN HEB HEI HUB JIA JIX SC SCH SHA SHX YUN ZHE
nanpingensis Legalov, 2007a: 275 A: SCH
ouhouensis Legalov, 2007a: 275 A: ANH

genus Enoplolabus Voss, 1925b: 214 type species Attelabus gestroi Faust, 1895
gestroi Faust, 1895b: 161 (Attelabus) A: YUN ORR

genus Henicolabus Voss, 1925b: 224 type species Attelabus giganteus Faust, 1882
subgenus Allolabus Voss, 1925b: 214 type species Attelabus lewisii Sharp, 1889
Eoallolabus Legalov, 2003b: 442 type species Henicolabus simplex Voss, 1925
JekeUlabus Legalov, 2003b: 441 type species Attelabus octospilotus Jekel, 1860 (= Attelabus octomaculatus Jekel, 1860)
kresli Legalov, 2004a: 83 A: NP
lewisii Sharp, 1889: 53 (Attelabus) A: FUJ HEI HUB JA NC SC ZHE
octomaculatus Jekel, 1860: 190 (Attelabus) A: TAI ORR
octospilotus Jekel, 1860: 201 (Attelabus)
subgenus Chinolabus Legalov, 2005: 129 type species Chinolabus ningxianus Legalov & Liu, 2005
bicolor Legalov & Liu, 2005: 129 (Chinolabus) A: QIN
gansuanus Legalov, 2007a: 282 (Chinolabus) A: GAN
ningxianus Legalov & Liu, 2005: 130 (Chinolabus) A: NIN
subgenus Henicolaboides Legalov, 2007a: 283 type species Henicolabus kuitchauensis Legalov, 2003
gigantinus Legalov & Liu, 2005: 127 A: BT YUN ORR
haematideus Voss, 1930b: 82 A: SCH
hypomelas Fairmaire, 1878: 130 (Attelabus) A: SCH
nanlingensis Legalov, 2008c: 208 (Henicolaboides) A: GUA
oberthueri Legalov, 2007a: 283 (Henicolaboides) A: SCH
potanini Legalov, 2007a: 284 (Henicolaboides) A: SCH YUN
ruflceps Voss, 1949: 159 A: FUJ HUN
spinipes Schilsky, 1906a: 88 (Attelabus) A: BEI FUJ GUA GUI GUX HUN JIA JIX SCH TAI YUN ZHE
nigrocapitus Legalov, 2007a: 283 (Henicolaboides)
yunnanicus Legalov, 2003b: 441 A: YUN
subgenus Henicolabus Voss, 1925b: 224 type species Attelabus giganteus Faust, 1882
giganteus Faust, 1882a: 291 (Attelabus) A: BEI FE FUJ HEB HEI JIL LIA NC SC SCH
136 ATTELABIDAE
genus Humerilabus Legalov, 2003b: 442 type species Henicolabusfausti Voss, 1925
chinensis Legalov, 2003b: 445 A: YUN
fausti Voss, 1925b: 226 (Henicolabus) [RN] A: NP HAI YUN ORR
humerosus Faust, 1895b: 163 (Attelabm) [HN]
longulus Legalov & Liu, 2005: 128 A: YUN
nepalensis Legalov, 2003b: 446 A: NP

genus holabus Voss, 1925b: 214 type species Attelabus caeruleus Jekel, 1860 (= Isolabus jekeli Legalov, 2002)
elongaticeps Voss, 1924c: 279 {Attelabus) A: TAI "mainland of China"
intrusus Voss, 1956d: 419 A: SCH
jekeli Legalov, 2002c: 92 [RN] A: FUJ JA JTA JIX HKG SHG ZHE
caeruleus Jekel, 1860: 202 (Attelabus) [HN]
magnus Voss, 1925b: 215 A: FUJ GUI GUX HEN HUB HUN JIA JIX SHA SCH SHN XIZ ZHE ORR
longicollis Fairmaire, 1894: 222 (Attelabus) [HN]

genus Lamprolabus Jekel, 1860: 189 type species Attelabus bispinosus Gyllenhal, 1833
anjiensis Legalov, 2007a: 277 A: ZHE
bihastatus Frivaldszky, 1892: 117 (Attelabus) A: FUJ GUA HAI HUB HUN JIX YUN
bispinosus Gyllenhal, 1833: 204 (Attelabus) A: JIX TAI ORR
tabangensis Voss, 1961b: 244
fochaensis Legalov, 2003b: 438 A: YUN
pseudobispinosus Legalov & Liu, 2005: 126 A YUN
sitchuanensis Legalov, 2003b: 439 A: GUA SCH
spiculatus Boheman, 1845: 359 (Attelabus) A: FUJ HAI YUN ORR
corallipes Pascoe, 1883b: 90 (Attelabus)
latispinosus Voss, 1929c: 209
tibetanus Legalov & Liu, 2005: 127 A: XIZ

genus Paramecolabus Jekel, 1860:190 type species Attelabus discolor Fâhraeus, 1839
subgenus Paramecolaboides Legalov, 2007a: 280 type species Attelabus feae Faust, 1895
castaneicolor Jekel, 1860: 190 (Attelabus) A: FUJ GAN HUB HUN SHG XIN

genus Phialodes Roelofs, 1874: 137 type species Phialodes rufipennis Roelofs, 1874
subgenus Phialodes Roelofs, 1874: 137 type species Phialodes rufipennis Roelofs, 1874
rufipennis Roelofs, 1874: 138 A: FE JA SC
distinctus Roelofs, 1874: 138
rubripennis Chevrolat, 1878: xxxi (Attelabus)
subgenus Chinphialodes Legalov, 2008c: 204 type species Phialodes hubeiensis Legalov, 2008
hubeiensis Legalov, 2008c: 204 A: HUB
tumidus X.-C. Zhang, 1995a: 207 (Attelabus) A: BEI GAN HEB LIA NIN SCH SHX

tribe Clitostylini Voss, 1929

genus Allapoderus Voss, 1927:15 type species Apoderus dentipes Faust, 1883
subgenus Allapoderus Voss, 1927: 15 type species Apoderus dentipes Faust, 1883
dentipes Faust, 1883a: 462 (Apoderus) A: HAI YUN ORR
manaliensis Voss, 1920: 172 (Apoderus) A: HP NP
symaoensis Legalov, 2003b: 474 A: YUN

genus Indotrachelus Legalov, 2007a: 298 type species Cycnotrachelus himalavensis Paini Haa & Gandhi,
1989 '
himalayensis Pajni, Haq & Gandhi, 1989: 520 (Cycnotrachelus) A: NP UP

genus Maculphrysus Legalov, 2003b: 479 type species Apoderus quadrimaculatus Faldermann, 1835
Parmaculphrysus Legalov, 2003b: 480 type species Centrocorynus inspersus Voss 1929
charlottae Voss, 1932b: 76 (Paracentrocorynus) A: YUN
 ATTELABIDAE 137
nigrostictus Voss, 1929b: 112 {Centrocorynus) A: NP
quadrimaculatus Faldermann, 1835: 419 {Apoderus) A: BEI FE MG NC SC
yunnanicus Legalov, 2003b: 480 A: FUJ GUA GUX YUN

genus Micrapoderus Legalov, 2003b: 476 type species Apoderus minutissimus Voss, 1920
bengalensis Legalov, 2003b: 477 A: SD
minutissimus Voss, 1920: 173 {Apoderus) A: HP KA NP UP
staudingeri Voss, 1920: 172 {Apoderus) A: LIA
uttaranchalensis Legalov, 2007a: 298 A: UP
yunpinensis Legalov, 2003b: 477 A: YUN

genus Morphocorynus Legalov, 2003b: 479 type species Apoderus nigricollis Roelofs, 1874
nigricollis Roelofs, 1874: 130 {Apoderus) A: FE HEN HUB JA (Honshu, Shikoku) NC SC
montanus Roelofs, 1874: 134 {Apoderus) [HN]

genus Pseudallapoderus Legalov, 2003b: 477 type species Apoderus sissu G. A. K. Marshall, 1913
ateroides Legalov, 2002c: 92 {Strigapoderus) [RN] A: YUN
ater Voss, 1932b: 73 {Apoderus) [HN]
fimebris Voss, 1930b: 85 {Apoderus) A: FUJ HUB ZHE ORR
kruegeri Voss, 1927: 61 {Apoderus) A: KA NP
pseudosissus Legalov, 2003b: 478 A: YUN
sikkimensis Legalov, 2007a: 299 A: SD
sissu G. A. K. Marshall, 1913: 227 {Apoderus) A: AF HP NP UP YUN ORR

genus Pseudophrysus Legalov, 2003b: 479 type species Cycnotrachelus parvulus Voss, 1928
parvulus Voss, 1928: 119 {Cycnotrachelus) A: HUB SCH YUN

tribe Euopini Voss, 1925

genus Euops Schoenherr, 1839: 318 type species Euops australasiae Fâhraeus, 1839 (= Attelabus falcatus
Guérin-Méneville, 1833)
subgenus Asuniops Legalov, 2007a: 219 type species Euops subopacus Voss, 1924
opacus Legalov, 2007a: 219 {Asuniops) A: JA
subgenus Chinoeuops Legalov, 2003b: 399 type species Sawadaeuops davidiani Legalov, 2003
davidiani Legalov, 2003b: 399 {Sawadaeuops) A: HUB SCH
hubeiensis Legalov & Liu, 2005: 123 {Sawadaeuops) A: HUB
nielamus Legalov & Liu, 2005: 124 {Sawadaeuops) A: XIZ
nitidicollis Voss, 1942b: 101 A: FUJ GAN HUB HUN
yunnanensis Liang, 2005: 265 A: GUI YUN
australis Legalov & Liu, 2005: 124 {Sawadaeuops)
subgenus Kobusynaptops Kôno, 1927a: 40 type species Euops pustulosus Sharp, 1889
pustulosus Sharp, 1889: 56 A: JA
verrucosus Legalov & X. Zhang, 2007: 231 {Kobusynaptops) A: YUN
subgenus Levoeuops Legalov, 2007a: 233 type species Riedeliops vietnamensis Legalov, 2003
asiaticus asiaticus Legalov & Liu, 2005: 119 {Riedeliops) A: HUB XIZ YUN
asiaticus uttaranchalicus Legalov, 2007a: 234 {Riedeliops) A: LTP
nepalensis Legalov, 2003b: 385 {Riedeliops) A: NP
yingjiangensis Liang, 2005: 265 A: YUN
terminassianae Legalov & Liu, 2005: 119 {Riedeliops)
subgenus Macrosynaptopsis Legalov, 2005: 115 type species Macrosynaptopsis zhangi Legalov & Liu, 2005
zhangi Legalov & Liu, 2005: 115 {Macrosynaptopsis) A: FUJ
subgenus Parasuniops Legalov, 2003b: 367 type species Parasuniops fochaiensis Legalov, 2003
fochaiensis Legalov, 2003b: 368 {Parasuniops) A: YUN
subgenus Parasynaptopsis Legalov, 2003b: 377 type species Euops chinensis Voss, 1922
Indoeuops Legalov, 2007a: 236 type species Indoeuops ammattiensis Legalov, 2007 (= Euops andrewesi Voss, 1935)
138 ATTELABIDAE
Macrodentipes Liang & Li, 2005: 266 type species Euops hefengensis Liang, 2005 (= Morphoeuops yunnanicus Legalov
2003)
Morphoeuops Legalov, 2003b: 377 type species Morphoeuops yunnanicus Legalov, 2003
carinatus Legalov & X. Zhang, 2007: 228 (Morphoeuops) A: XIZ
chinensis Voss, 1922a: 166 A: FUJ GUA GUI GUX HUB HUN JIA JIX TAI ZHE
cuprifiilgens Voss, 1942b: 102 A: FUJ GUA
lespedezae koreanus Sawada & Morimoto, 1985: 191 A: FE JA (Tsushima) JIL NC SC
lespedezae lespedezae Sharp, 1889: 55 A: HEI JA
pseudolespedezae Z.-Q. Li, 2008: 69 A: JIL
pseudostriatus Legalov, 2007a: 237 (Parasynaptopsis) A: GUA
splendidus Voss, 1930a: 35 [RN] A: FE JA NC SC ZHE
niger Legalov, 2003b: 378 (Parasynaptopsis)
splendens Roelofs, 1874: 139 (Attelabus) [HN]
sthatus Voss, 1924c: 279 A: TAI
taiwanensis Legalov, 2007a: 237 (Parasynaptopsis) A: TAI
yunnanicus Legalov, 2003b: 377 (Morphoeuops) A: HUB YUN
hefengensis Liang, 2005: 266
subgenus Parasynatops Legalov, 2003b: 378 type species Parasynatops politoides Legalov, 2003 (= Attelabus
politus Roelofs, 1874)
Neparasynatops Legalov, 2007a: 240 type species Parasynatops moanus Legalov, 2003
berezovskyi Legalov, 2003b: 379 (Parasynatops) A: SCH
championi Voss, 1929c: 217 A: FUJ GUI GUX HAI TAI UP ZHE ORR
kharsu Voss, 1933d: 116 A: UP
konoi Sawada & Morimoto, 1985: 185 A: FE (Kuriles) JA
moanus Legalov, 2003b: 382 (Parasynatops) A: SCH
politus Roelofs, 1874: 140 (Attelabus) A: JA
politoides Legalov, 2002c: 92 ("'Parasynatops") [RN]
pseudopolitus Legalov, 2003b: 380 (Parasynatops) A: HUB SCH YUN
tzinpinensis Legalov & Liu, 2005: 116 (Parasynatops) A: YUN
viridicollis Voss, 1924a: 64 A: SD
subgenus Pseudoeuops Legalov, 2003b: 380 type species Parasynatops bicoloroides Legalov, 2003
bicoloroides Legalov, 2003b: 381 (Parasynatops) A: FUJ
clarus Sawada & Morimoto, 1985: 188 A: TAI
guizhouensis Liang, 2005:258 A: GUI
lushuensis Legalov & Liu, 2005: 118 (Parasynatops) A: YUN
pyralis Legalov & Liu, 2005: 117 (Parasynatops) A: YUN
tibetanus Legalov & Liu, 2005: 117 (Parasynatops) A: XIZ
viridis Legalov, 2003b: 383 (Parasynatops) A: SCH YUN
subgenus Riedeliops Alonso-Zarazaga & Lyal, 2002: 10 [RN] type species Euops paradoxus Voss, 1935
Asynaptops Legalov, 2007a: 225 type species Euops keiseri Voss, 1957
Asynaptopsis Legalov, 2007a: 225 type species Asynaptops colombensis Legalov, 2007
Charops Riedel, 1998: 100 [HN] type species Euops paradoxus Voss, 1935
Nigroeuops Legalov, 2003b: 403 type species Sawadaeuops ovalis Legalov, 2003
Orienteuops Legalov, 2003b: 388 type species Euops tonkinensis Voss, 1933
Orienteuopsidius Legalov, 2008b: 215 type species Riedeliops rasuwanus Legalov, 2003
Sawadaeuops Legalov, 2003b: 398 type species Attelabus punctatostriatus Motschulsky, 1861
beijingensis Legalov, 2003b: 379 (Parasynatops) A: HEB
bowringii Jekel, 1860:221 A: UP ORR
armipes Voss, 1933d: 117
colombensis Legalov, 2007a: 225 (Asynaptops)
submetallicus Voss, I935d: 98
centralchinensis Legalov & Liu, 2005: 122 (Sawadaeuops) A: HUB SHA
fugongensis Wang & Liang, 2008: 178 A: YUN
gardneri G. A. K. Marshall, 1931: 263 A: SD
indigena Voss, 1924c: 278 A: TAI
matsumuranus KÔDO, 1927a: 42
 ATTELABIDAE 139
jingguensis Liang & Sakurai, 2005: 263 A: YUN
mirabilis Legalov & Liu, 2005: 120 (Orienteuops)
jiuzhaiensis Liang, 2005: 261 A: SCH YUN
subelongatus Legalov & Liu, 2005: 123 (Sawadaemps)
meghalaicus Legalov, 2007a: 232 {Orienteuops) A: SD ORR
ovalis Legalov, 2003b: 403 (Sawadaeuops) A: HUN ORR
potanini Legalov, 2003b: 380 (Parasynatops) A: GAN
punctatostriatus Motschulsky, 1861: 22 (Attelabus) A: FE JA SC
aceri Kôno, 1926b: 223
phaedonius Sharp, 1889: 55
puncticollis Schilsky, 1906a: 92 [HN]
schilskyi Voss, 1922b: 174 [RN]
rasuwanus Legalov, 2003b: 386 (Riedeliops) A: NP SD XIZ
bengalensis Legalov, 2007a: 239 {Parasynatops)
sichuanensis Legalov & Liu, 2005: 121 {Sawadaeuops) A: SCH
subgenus Sawadaeuopsis Legalov, 2005: 125 type species Sawadaeuops punctatus Legalov & Liu, 2005
punctatus Legalov & Liu, 2005: 125 {Sawadaeuops) A: YUN
subgenus Suniopsidius Legalov, 2005: 113 type species Suniopsidius multicoloratus Legalov & Liu, 2005
multicoloratus Legalov & Liu, 2005: 114 {Suniopsidius) A: YUN
subgenus Vietsuniops Legalov, 2007a: 223 type species Suniops gorochovi Legalov, 2003
menghunsis Legalov & Liu, 2005: 113 {Suniops) A: YUN

tribe Euscelophilini Voss, 1925

genus Calolabus Voss, 1925b: 241 type species Attelabus cupreus Roelofs, 1874
subgenus Calolabus Voss, 1925b: 241 type species Attelabus cupreus Roelofs, 1874
cupreus Roelofs, 1874: 139 (Attelabus) A: JA
subgenus Metocalolabus Legalov, 2003b: 433 type species Metocalolabus orientalis Legalov, 2003
chujoi Kôno, 1939: 143 {Himatolabus) A: TAI

genus Euscelophilus Voss, 1925a: 30 type species Euscelus chinensis Schilsky, 1906
Cupreuscelophilus Legalov, 2005: 130 type species Cupreuscelophilus mayongi Legalov & Liu, 2005
Euscelophilidius Legalov, 2003b: 463 type species Euscelus gibbicollis Schilsky, 1906
Euscelophiloides Legalov, 2007a: 292 type species Euscelophilus rugulosus X.-C. Zhang, 1995
burmanus G. A. K. Marshall, 1948: 411 A: FUJ SCH YUN ORR
tianchiensis Liang, 1994: 489
camelus Voss, 1937a: 280 A: YUN
caperratus Xie & Liang, 2008: 39 A: GAN
chinensis Schilsky, 1906a: 89 {Euscelus) A: BEI FUJ GUA GUI GUX HAI HEN HUB NC SC SCH SHN
ORR
denticulatus X.-C. Zhang, 1995b: 481 A: BEI FUJ GUA GUI GUX HAI HUB SCH ZHE
dimidiatus Voss, 1956d: 418 A: HEN SCH
fasciatus X.-C. Zhang, 1993: 199 (Himatolabus) A: XIZ
gaoligongensis Xie & Liang, 2008: 36 A: YUN
gibbicollis Schilsky, 1906a: 90 (Euscelus) A: BEI GUI HEB HUB NC SC SCH YUN
hidakai Liang, 1994: 493
jingpingensis Liang, 1994: 487 A: YUN
kunmingensis Liang, 1994: 490 A: YUN
longispinus Xie & Liang, 2008: 35 A: CHQ GUI GUX
mayongi Legalov & Liu, 2005: 131 (Cupreuscelophilus) A: ZHE
niger Liang, 1994: 484 A: HUB
rugulosus X.-C. Zhang, 1995b: 480 A: YUN
vitalisi Heller, 1922a: 13 {Trachelolabus) A: YUN ORR
yongshanensis Liang, 1994: 492 A: SCH
140 ATTELABIDAE
genus Trachelolabus Jekel, 1860: 217 type species Trachelolabus whitei Jekel, 1860
floridus X.-C. Zhang, 1993: 198 (Himatolabus) A: XIZ YUN
qinni Liang, 1994: 488 (Euscelophilns)
ghumensis Legalov, 2007a: 291 A: SD
longispinus X.-C. Zhang, 1993: 197 {Himatolabus) A: XIZ YUN
whitei Jekel, 1860: 218 A: XIZ ORR

tribe Hoplapoderini Voss, 1926

genus Agomadaranus Voss, 1958: 15 type species Apoderus pardalis Snellen van Vollenhoven, 1865
Pseudmadaranus Legalov, 2003b: 508 type species Paroplapoderus fasciatus Voss, 1929 (= Paroplapoderus
multicostatus Pic, 1928)
armatus Voss, 1926a: 55 {Paroplapoderus) A: SD ORR
bihumeratus Jekel, 1860: 180 {Apoderus) A: NP SCH SD YUN ORR
breviceps Voss, 1926a: 53 {Paroplapoderus)
bistrispinosus Faust, 1895b: 157 {Apoderus) A: GUI SCH XIZ YUN ORR
coniceps Voss, 1926a: 53 {Paroplapoderus) A: BEI FUJ SCH YUN XIZ
jekeli Legalov, 2007a: 325 A: NP SD
kresli Legalov, 2004a: 85 A: NP
kryzhanovskyi Legalov, 2003b: 506 {Paroplapoderus) A: YUN
melanostictoides Legalov, 2003b: 502 {Paroplapoderus) A: ZHE
melanostictus Fairmaire, 1878: 129 {Apoderus) A: BEI FUJ HEB HEN HUB HUN SCH SHN SHX YUN
hauseri Voss, 1930b: 83 {Paroplapoderus)
multicostatus Pic, 1928c: 3 {Paroplapoderus) A: YUN ORR
fasciatus Voss, 1929d: 363 {Paroplapoderus)
parbihumeratus Legalov, 2003b: 504 {Paroplapoderus) A: SCH YUN
pardalis Snellen van Vollenhoven, 1865: 164 {Apoderus) A: FE FUJ GUA GUI HAI HUN JA JIA JIX SCH
YUN
tigrinus Roelofs, 1874: 136 {Apoderus)
pardaloides Voss, 1924c: 284 {Hoplapoderus) A: TAI
semiannulatus Jekel, 1860: 181 {Apoderus) A: FUJ GUA GUI HEB HEN HUB HUN JIA SCH SHG SHX ZHE
shirakii Kôno, 1930b: 44 {Paroplapoderus) A: HAI TAI ORR
sticticus sticticus Voss, 1926a: 43 {Paroplapoderus) A: HAI ORR
sticticus yunnanicus Legalov, 2004a: 85 A: FUJ SCH YUN
xilingensis Legalov, 2007a: 327 A: SCH

genus Architomapoderus Legalov, 2007a: 318 type species Parapoderus nilgiriensis Voss, 1935
vittatus Voss, 1935d: 99 {Parapoderus) A: "India"

genus Hoplapoderus Jekel, 1860: 171 type species Attelabus gemmatus Thanberg, 1784
caliginosus Faust, 1895b: 157 {Apoderus) A: BT JA SCH TAI YUN ORR
echinatoides Legalov, 2003b: 511 A: BEI FE HAI HUB HUN YUN ZHE ORR
echinatus Gyllenhal, 1833: 195 {Apoderus) A: FE GUA HEN HP NP SD YUN ORR
ceylonicus Janczyk, 1960: 49 {Paroplapoderus)
nepalensis Voss, 1926a: 38
gemmatus Thunberg, 1784b: 68 {Attelabus) A: NP UP ORR
jekeli Legalov, 2007a: 330 [See New Acts]
gemmosus Jekel, 1860: 182 {Apoderus) [See New Acts] A: FE GUA GUX HAI HKG HUN JA (Kyushu) JIA
NC SD SHG TAI YUN ORR
chevrolati Faust, 1891a: 284 {Apoderus)
minutituberosus Haq, Pajni & Gandhi, 1988: 284 {Hoplapoderus)
genus Paroplapoderus Voss, 1926a: 41 type species Apoderus fallax Gyllenhal, 1839
subgenus Erycapoderus Voss, 1926a: 69 type species Apoderus angulipennis Kolbe, 1886
angulipennis angulipennis Kolbe, 1886: 215 {Apoderus) A: NC NE SC SHN
angulipennis shaanxinsis Legalov, 2004a: 84 A: SHA
 ATTELABIDAE 141
validus Voss, 1926a: 70 A: ANH BEI FUJ GAN HEN JIA SCH SHN TAI ZHE ORR
nigroguttatus Kôno, 1930b: 51
subgenus Gomadaranus Kôno, 1930b: 48 type species Apoderus vitticeps Jekel, 1860
bihiberculatus Voss, 1926a: 66 A: SD
kuanchenensis Legalov, 2007a: 321 A: HEB
nigroflavus Legalov, 2006c: 20 A: ANH SHN
obtusus Voss, 1926a: 62 A: GUA
punctatus Pic, 1928d: 247 A: XIZ
rasuwanus Legalov, 2004a: 84 A: NP
shiguensis Legalov, 2007a: 322 A: YUN
tristoides Voss, 1924c: 284 (Hoplapoderus) A: TAI
turbidus Voss, 1926a: 63 A: BEI JIA LIA NC SC SHA SHN SHX ZHE
vanvolxemi Roelofs, 1875b: cxxxi {Hoplapoderus) A: FE HUB HUN JA SC
japonicus Voss, 1926a: 62
tuberculatus Harold, 1878: 85 {Apoderus)
ulmi Kôno, 1929b: 49
vitticeps Jekel, 1860: 184 {Apoderus) A: BEI FUJ GUI GUX HUB JIA JIX LIA SCH SHG SHN TAI ZHE
subgenus Paroplapoderus Voss, 1926a: 41 type species Apoderus fallax Gyllenhal, 1839
fallax Gyllenhal, 1839: 287 {Apoderus) A: BEI ES FE HEB HEI LIA MG NP SHX ZHE
subgenus Pseudplapoderus Legalov, 2003b: 501 type species Apoderus tentator Faust, 1895
allem Legalov, 2008c: 212 A: BT
nigriceps Legalov, 2003b: 498 A: YUN
tentator annapurnae Legalov, 2007a: 323 A: NP
tentator tentator Faust, 1895b: 157 {Apoderus) A: FUJ GUA XIZ YUN ORR
tzinpinensis Legalov, 2003b: 501 A: YUN

genus Phymatapoderus Voss, 1926a: 71 type species Apoderus latipennis Jekel, 1860
elongatipes Voss, 1926a: 72 A: GUI GUX HUB SCH YUN ZHE ORR
euflavimanus Legalov, 2003b: 494 A: SCH
flavimanus Motschulsky, 1860: 171 {Apoderus) A: ES FE FUJ HEI JA JIX LIA NC SC SCH YUN ZHE
pavens Voss, 1926a: 74
latipennis Jekel, 1860: 179 {Apoderus) A: FUJ GAN GUA GUI HUB HUN JA JIA JIX MG NC SC SCH TAI
YUN ZHE ORR
monticola Voss, 1926a: 75
yunnanicus Voss, 1929c: 227
monstrosus Pic, 1928c: 3 A: SHG
parelongatipes Legalov, 2003b: 494 A: SCH YUN
sitchuanensis Legalov, 2003b: 495 A: SCH
subornatus Sharp, 1889: 47 {Apoderus) A: JA
taiwanensis Legalov, 2003b: 495 A: TAI

genus Tomapoderus Voss, 1926a: 76 type species Attelabus ruficollis Fabricius, 1781
Tomapoderopsis Legalov, 2003b: 495 type species Apoderus cyclops Faust, 1895
coeruleipennis Schilsky, 1903: 75 (Apoderus) A: FUJ HKG HUB HUN YUN TAI "N China"
cyclops Faust, 1895b: 155 (Apoderus) A: BT YUN ORR
flaviceps Desbrochers des Loges, 1890: 213 (Apoderus) A: HP NP SD TAI ORR
flavirostris Voss, 1926a: 81 A: NP ORR
himalayensis Legalov, 2007a: 320 (Tomapoderopsis) A: NP
melli Voss, 1926a: 82 A: GUA HUN
orientalis Legalov, 2007a: 320 (Tomapoderopsis) A: SCH YUN ORR
pici Legalov, 2007a: 321 (Tomapoderopsis) A: SCH
ruficollis Fabricius, 1781:200 (Attelabus) A: ANH BEI ES FE GUI HEB HEI JA JIL LIA MG NC NMO SC
SHA SHN SHX TAI
testaceimembris Pic, 1928c: 3 A: FUJ GUA GUI HUB HUN SCH YUN ORR
142 BRENTIDAE
family BRENTIDAE Billberg, 1820
subfamily Cyladinae Schoenherr, 1823
genus Cylas Latreille, 1802:196 type species Brentus brunneus Olivier, 1790
Protocylas Pierce, 1941: 219 type species Cylas laevicollis Boheman, 1833
formicarius Fabricius, 1798: 174 {Brentus) A: AEi FUJ GUA GUI GUX HAI HEN HKG HUN JAi JIA JIX PA
QA SA SCi SCH SHN TAJ UP YUN ZHE AFRi AURi NARi NTRi ORR
formicarius Fabricius, 1798: 163 (Attelabus) [HN]
turcipennis Boheman, 1833: 369
submetallicus Desbrochers des Loges, 1890: 214 A: SD ORR

subfamily Brentinae Billberg, 1820

tribe Arrhenodini Lacordaire, 1865
genus Agriorrhynchus Power, 1878a: 241 type species Agriorrhynchus borrei Power, 1878
borrei Power, 1878a: 241 A: SD ORR
consonus Kleine, 1923a: 410 (Prosorychodes)
cynicus Damoiseau, 1966b: 452 A: GUA
klapperichi Kleine, 1950: 24 A: FUJ
quadrituberculatus Senna, 1892b: 470 A: SD ORR

genus Ananesiotes Kleine, 1922b: 132 type species Ananesiotes insularis Kleine, 1922
insularis Kleine, 1922b: 133 A: TAI

genus Anepsiotes Kleine, 1917b: 318 type species Anepsiotes schenklingi Kleine, 1917
commendabilis Kleine, 1925a: 159 A: UP ORR
schenklingi Kleine, 1917b: 321 A: YUN ORR

genus Baryrhynchus Lacordaire, 1865: 428 type species Arrhenodes latirostris Gyllenhal, 1833
subgenus Baryrhynchus Lacordaire, 1865: 428 type species Arrhenodes latirostris Gyllenhal, 1833
angulatus R.-Z. Zhang, 1993: 161 A: YUN SCH
bengalensis Kleine, 1937: 203 A: SD
concretus R.-Z. Zhang, 1993: 164 A: YUN
convexus R.-Z. Zhang, 1993: 168 A: HAI
cratus R.-Z. Zhang, 1993: 165 A: YUN
dehiscens Gyllenhal, 1833: 324 (Arrhenodes) A: YUN ORR
latirostris Gyllenhal, 1833: 323 (Arrhenodes) A: SD ORR
merocephalus Kleine, 1916b: 178 A: FUJ SD YUN ORR
miles Boheman, 1845: 372 (Arrhenodes) A: HAI NP SD SHG TAI YUN ORR
approximans F. Walker, 1859a: 262 (Arrhenodes)
minisculus R.-Z. Zhang, 1993: 168 A: YUN
nitidus R.-Z. Zhang, 1993: 169 A: HAI
odontus R.-Z. Zhang, 1993: 163 A: SCH
phaeus R.-Z. Zhang, 1993: 162 A: FUJ ZHE
planus R.-Z. Zhang, 1993: 163 A: YUN
setulosus R.-Z. Zhang, 1993: 166 A: YUN
speciosissimus Kleine, 1916b: 180 A: FUJ SCH YUN ORR
yaeyamensis Morimoto, 1979b: 27 A: JA
subgenus Eupsalomimus Kleine, 1916b: 151 type species Baryrrhynchus lineicollis Power, 1879
poweri Roelofs, 1880a: 26 (Baryrrhynchus) A: FUJ GUA GUX HAI HUB HUN JA JIX FE FUJ SCH TAI ZHE
AUR ORR
tokarensis Ohbayashi & Satô, 1966: 1 A: JA

genus Blysmia Pascoe, 1872:323 type species Blysmia ruficollis Pascoe, 1872
Ypselogonia Kleine, 1922c: 157 type species Ypselogonia peregrina Kleine, 1922
guttata Kleine, 1928: 102 (Ypselogonia) A: TAI ORR
 BRENTIDAE 143
peregrina Kleine, 1922c: 158 (Ypselogonia) A: TAI ORR

genus Hemiorychodes Kleine, 1921b: 83 type species Hemiorychodes cambodje nsis Kleine, 1921
cambodjensis Kleine, 1921b: 84 A: CH ORR
modestus Kleine, 1921b: 85 A: YUN ORR

genus Orfilaia Haedo Rossi, 1955: 63 type species Arrhenodes vulsellatus Gyllenhal, 1833
arabica Muizon, 1954: 327 (Eupsalis) A: YE
reichet Fairmaire, 1859: clxiv (Arrhenodes) E: GR (Kriti) N: TU A: CY IS SY

genus Orychodes Pascoe, 1862: 389 type species Brentus serrirostris Lund, 1800
Caenorychodes Kleine, 1921b: 87 type species Brentus serrirostris Lund, 1800
im*« Kirsch, 1875: 51 A: TAI ORR
octoguttatus Nakane, 1963b: 34 (Cerobates) A: JA
planicollis F. Walker, 1859a: 262 (Nemocephalus) A: JA SD TAI YUN ORR
serrirostris Lund, 1800: 86 (Brentus) A: HAI TAI ORR
femoratus L. W. Schaufuss, 1885: 206 (Orychodes)
striolatus Kirsch, 1875: 51 (Orychodes)
sinensis Fairmaire, 1888: 138 A: BEI

genus Parorychodes Kleine, 1921b: 80 type species Orychodes degener Senna, 1892
cereus Kleine, 1925a: 151 A: YUN ORR

genus Prophthalmus Lacordaire, 1865: 427 type species Brentus tridentatus Lund, 1800
heikertingeri Kleine, 1916c: 255 A: SD ORR
longirostris Gyllenhal, 1833: 323 (Prophthalmus) A: HAI ORR
assimilis Kleine, 1916c: 248
potens potens Lacordaire, 1865:428 A:NPSDYUN ORR
amethystinus Kleine, 1916c: 242
sanguinalis Pascoe, 1872: 322
pugnator Power, 1878b: xliv A: TAI ORR
tridentatus Lund, 1800: 90 (Brentus) A: TAI ORR
wichmanni Kleine, 1916c: 263 A: JA SD TAI YUN ORR

genus Pseudorychodes Senna, 1894a: 375 type species Orychodes insignis Lewis, 1883
crassus Senna, 1894a: 378 A: SD ORR
insignis Lewis, 1883: 301 (Orychodes) A: JA FE TAI "Korea"
mandschuricus Kleine, 1939b: 318 A: HEI
sauteri Kleine, 1928: 100 A: TAI

genus Suborychodes Kleine, 1917b: 328 type species Suborychodes intermedius Kleine, 1917
intermedius Kleine, 1917b: 330 A: UP ORR

genus Synorychodes Kleine, 1921a: 9 type species Synorychodes opacus Kleine, 1921
glaber Kleine, 1937: 204 A: SD

tribe Eremoxenini Semenov, 1892

genus Amorphocephala Damoiseau, 1966a: 307 type species Brentus coronatus Germar, 1817
Amorphocephalus Schoenherr, 1840: 485 [HN] type species Brentus coronatus Germar, 1817
arabica Damoiseau, 1964a: 428 (Amorphocephalus) A: SA
coronata Germar, 1817a: 247 (Brentus) E: AL BH BU CR FR GR IT MC PT SP TR YU N: AG MO A: CY
IN IS RU SY "Caucasus"
delicata Kleine, 1925a: 144 (Amorphocephalus) A: PA
deserticola Kleine, 1944: 153 (Amorphocephalus) N: MO
excantator Damoiseau, 1964a: 425 (Amorphocephalus) A: JO
144 BRENTIDAE
jickelii L. W. Schaufuss, 1876: 402 (Amorphocephalus) N: EG AFR
scotti Muizon, 1954: 325 {Amorphocephalus) A: YE
senegalensis Power, 1879: 486 (Amorphocephalus) N:AGEG AFR
princeps Kleine, 1918b: 101 (Amorphocephalus)
sulcata Calabresi, 1920: 64 N: EG LB A: SA AFR

genus Cobalocephalus Morimoto, 1982: 33 type species Amorphocephalus gyotokui Nakane, 1963
gyotokui Nakane, 1963c: 35 (Amorphocephalus) A: JA
mizobei Morimoto, 2009: 287 A: JA (Iriomote Is;)

genus Eremoxenus Semenov, 1892: 438 type species Eremoxenus chan Semenov, 1892
Acramorphocephaloides Bartolozzi, 1988: 155 type species Acramorphocephaloides algirus Bartolozzi, 1988
algirus Bartolozzi, 1988: 157 (Acramorphocephaloides) N: AG
chan Semenov, 1892: 440 A: IN KZ TM UZ

genus Leptamorphocephalus Kleine, 1918b: 132 type species Leptamorphocephalus laborator Kleine, 1918
laborator Kleine, 1918b: 132 A: TAI ORR
laevis Power, 1879: 486 (Amorphocephalus) A: SD ORR
cupidus Kleine, 1925a: 143 (Leptamorphocephalus)

genus Paramorphocephalus Kleine, 1920b: 236 type species Paramorphocephalus diabolus Kleine, 1920
fumosus Morimoto, 1976: 279 (Leptamorphocephalus) A: JA

genus Perisymmorphocerus Kleine, 1919: 13 type species Perisymmorphocerus gracilis Kleine, 1919 (=
Cordus latirostris Power, 1879)
latirostris Power, 1879: 484 (Cordus) A: YE AFR
gracilis Kleine, 1919: 15 (Perisymmorphocerus)

genus Symmorphocerus Schoenherr, 1847: 8 type species Symmorphocerus monticola Schoenherr, 1847
beloni Power, 1879: 488 A: IQTR
cardoni Senna, 1894b: 385 A: PA ORR
consequens Kleine, 1926b: 366 A: IQ KZ
piochardi Bedel, 1878: clxxxiv (Amorphocephalus) E: GR A: JO LE SY TR
cretensis Hertel, 1964: 167 (Amorphocephalus)
muchei Hertel, 1961: 117 (Amorphocephalus)
piliger Desbrochers des Loges, 1895: 99 (Amorphocephalus)
saalasii. R. Sahlberg, 1913a: 61 (Symmorphocerus)
unionis J. R. Sahlberg, 1913b: 201 (Amorphocephalus)

subfamily Cyphagoginae.Kolbe, 1892

tribe Cyphagogini Kolbe, 1892
genus Allaeometrus Senna, 1903: 157 type species Allaeometrus breviceps Senna, 1903
Subdysmorphorhynchus Kleine, 1917b: 323 type species Subdysmorphorhynchus carinatus Kleine, 1917 (= Allaeomet
breviceps Senna, 1903)
breviceps Senna, 1903: 157 A: JLXTAI ORR
carinatus Kleine, 1917b: 326 (Subdysmorphorhynchus)

genus Callipareius Senna, 1892b: 444 type species Callipareius feae Senna, 1892
subgenus Callipareius Senna, 1892b: 444 type species Callipareius feae Senna, 1892
Asaphepterum Kleine, 1916a: 85 type species Asaphepterumformosanum Kleine, 1916
Diastrophus Perroud, 1865: 141 [HN] type species Diastrophus planitarsis Perroud, 1865
Pseudosebasius Kleine, 1944: 150 type species Pseudosebasius rubrocastaneus Kleine 1944
bistriatus Kleine, 1935: 73 (Opysthenoxys) A: BT FUJ SD ORR
lineatosignatus Kleine, 1939a: 1 (Eterozemus)
feae Senna, 1892b: 445 A: SD YUN ORR
japonicus Nakane, 1963b: 34 (Asaphepterum) A: JA
 BRENTIDAE 145
formosanus Kleine, 1916a: 87 (Asaphepterum) A: JA TAI ORR
foveatus Senna, 1894b: 384 A: SD ORR
mediochs Kleine, 1935: 74 (Pittodes)
miyakawai Morimoto, 1979b: 25 A: JA
subgenus Metacidotes Schedl, 1961b: 193 type species Metacidotes pulvifrons Schedl, 1961
Coomania Kleine, 1941: 226 [HN] type species Coomania ponderosa Kleine, 1941
kojimai Morimoto, 1982: 31 A: JA TAI "Korea"
ponderosus Kleine, 1941: 227 (Coomania) A: HAI ORR

genus Calodromus Guérin-Méneville, 1832: CL IX., pL 34 type species Calodromus mellyi Guérin-Méneville
&Gory, 1832
Alloeodromus Senna, 1895a: 179 type species Alloeodromus insignis Senna, 1895
Callidromus Agassiz, 1846: 60 [unjustified RN]
horishanus Kôno, 1930: 154 A: TAI
ito Heller, 1910: 180 A: TAI
mellyi Guérin-Méneville & Gory, 1832: CL. IX, pi. 34 A: YUN ORR
sarawakensis Heller, 1910: 179 (Calodromus)

genus Chelorhinus Kleine, 1922a: 92 type species Chelorhirms propheti Kleine, 1922
Epigogus Kleine, 1923b: 159 type species EpigogusflexibilisKleine, 1923
Pseudopisthenoxys Kleine, 1942 type species Pseudopisthenoxys insculptus Kleine, 1942
propheti Kleine, 1922a: 93 A: TAI

genus Cyphagogus Parry, 1849:182 type species Cyphagogus westwoodi Parry, 1849
beesoni Damoiseau, 1964b: 484 A: UP
boninensis Morimoto & Kojima, 2005: 126 A: JA
confertulus Kleine, 1925a: 125 A: BT YUN ORR
fragosus Kleine, 1925a: 126
corporaali Kleine, 1918c: 177 A: UP ORR
cyrtotrachelus J. Thomson, 1857: 119 (Calodromus) A: UP ORR
buccatus Kleine, 1916a: 9 (Cyphagogus)
incisus Senna, 1903: 153 A: SD ORR
obconiceps Senna, 1903: 154 (Cyphagogus)
iwatensis Morimoto, 1976: 268 A: JA
longulus Senna, 1898a: 52 A: BT ORR
simulator Senna, 1902: 155
modiglianii Senna, 1893c: 258 A: TAI AUR ORR
planifrons Kirsch, 1875: 46 A: BT YUN ORR
gladiator Kleine, 1921c: 307
signipes Lewis, 1883: 297 A: HUN JA TAI
westwoodi Parry, 1849: 182 A: BT ORR

genus Eterozemus Senna, 1903: 160 type species Zemiosespubens Senna, 1892
Dictyotopterus Kleine, 1916a: 75 type species Dictyotoptenis splendens Kleine, 1916
Orthopareia Kleine, 1925b: 591 type species Orthopareia idonea Kleine, 1925 (= Dictyotopterus philippinensis Kleine,
1921)
Sparganophasma Kleine, 1926a: 11 type species Sparganophasma carum Kleine, 1926
celtis Lewis, 1883: 296 (Zemioses) A: JA
pubens Senna, 1892b: 439 (Zemioses) A: TAI ORR
coomani Kleine, 1944: 151 (Allaeometrus)
splendens Kleine, 1916a: 77 (Dictyopterus) A: TAI

genus Mesoderes Senna, 1898b: 65 type species Mesoderes sexnotatus Senna, 1898
Dysmorphorhynchus Kleine, 1916a: 75 type species Dysmorphorhynchus amabilis Kleine, 1916 (=Mesoderes sexnotatus
Senna, 1898)
Thrasycephalus Kleine, 1916a: 53 type species Thrasycephalus guttatus Kleine, 1916
146 BRENTIDAE
latecostatus Kleine, 1916a: 58 (Isomorphus) A: JA TAI

genus Microsebus Kolbe, 1892:168 type species Microsebus adelphus Kolbe, 1892
Eusebus Kleine, 1922: 173 type species Microsebus adelphus Kolbe, 1892
adelphus Kolbe, 1892:169 A: UP ORR
bhutanicola Damoiseau, 1979: 30 A: BT
trifasciatus Kleine, 1916a: 57 {Isomorphus) A: TAI YUN ORR

genus Opisthenoxys Kleine, 1922: 26 type species Opisthenoxys ochraceus Kleine, 1922
Barathrodes Damoiseau, 1961: 11 type species Barathrodes binotatus Damoiseau, 1961 (= Opisthenoxys ochraceu
Kleine, 1922)
famulus Kleine, 1925: 128 A: SD ORR

tribe Hoplopisthiini Senna & Calabresi, 1919

genus Carcinopisthius Kolbe, 1892:174 type species Carcinopisthius fruhstorferi Kolbe, 1892
Pseudotaphroderes Bolkay, 1911: 263 type species Pseudotaphroderes forficatus Bolkay, 1911 (= Carcinopisthius kolbe
Senna, 1892)
Stratiopisthius Senna & Calabresi, 1919: 76 type species Hoplopisthius doriae Senna, 1892
oberthueri Senna, 1893a: 255 {Hoplopisthius) A: TAI ORR
felschei Kolbe, 1892: \15 (Carcinopisthius)
maculatus Senna, 1903: 163 (Carcinopisthius)
formosanus Bolkay, 1911: 265 (Pseudotaphroderes)

genus Hoplopisthius Senna, 1892b: 451 type species Hoplopisthius trichemerus Senna, 1892
trichemerus Senna, 1892b: 452 A: TAI YUN ORR
javanus Kolbe, 1892: 173 (Hoplopisthius)

tribe Stereodennini Sharp, 1895

genus Cerobates Schoenherr, 1840: 487 type species Brentus tristriatus Lund, 1800
subgenus Cerobates Schoenherr, 1840: 487 type species Brentus tristriatus Lund, 1800
angustipennis Senna, 1895a: 182 A: SD ORR
adustus Senna, 1895a: 184 (Cerobates)
birmanicus Senna, 1894a: 364 A: TAI ORR
canaliculars Motschulsky, 1858: 96 A: JA SD TAI YUN ORR
formosanus Schonfeldt, 1911: 190 A: JA TAI ORR
laevipennis Senna, 1896:219 A: NP ORR
planicollis Morimoto, 1976: 276 A: JA
sexsulcatus Motschulsky, 1858:95 A: YUN ORR
glaberrimus Senna, 1892b: 450
sumatranus Senna, 1893d: 306 A: SDTAI ORR
andamanicus Senna, 1896: 223
tristriatus Lund, 1800 (Brentus) A: UP ORR
elytralis Senna, 1897: 545
longirostris Senna, 1898d: 227
turgidulus Damoiseau, 1987: 78 A: TAI
subgenus lonthocerus Lacordaire, 1865: 415 type species lonthocerus crematus Lacordaire, 1865
asiaticus Kleine, 1920a: 47 (lonthocerus) A: JA NP TAI YUN ORR
angustifrons Kleine, 1922d: 148 (lonthocerus)
bicolor Heller, 1916: 297 (lonthocerus) A: TAI ORR
carinensis Senna, 1892b: 450 (Cerobates) A: SD YUN ORR
caesius Kleine, 1944: 151 (lonthocerus)
crematus Lacordaire, 1865: 416 A: HAI ORR
concisus Kleine, 1925a: 130 (Cerobates)
laticostatis Kleine, 1920a: 38 (lonthocerus) A: TAI
nigripes Lewis, 1883: 298 (lonthocerus) A: JA TAI "Korea"
sondaicus Senna, 1893d: 302 (lonthocerus) A: TAI ORR
 BRENTIDAE 147

angulaticeps Senna, 1898a: 53 (Ionthocerus)

borneensis Senna, 1898c: 190 (Ionthocerus)

subfamily Trachelizinae Lacordaire, 1865

tribe Ithystenini Lacordaire, 1865
genus Ceocephalus Guérin-Méneville, 1833: pi. 36 type species Ceocephalus furcillatus Guérin-Méneville,
1833
Diurus Dejean, 1834: 244 type species Ceocephalus furcillatus Gyllenhal, 1833 (= Ceocephalus furcillatus Guérin-
Méneville, 1833)
Diurus Pascoe, 1862: 392 [HN] type species Ceocepalus furcillatus Gyllenhal, 1833 (= Ceocephalus furcillatus Guérin-
Méneville, 1833)
Fornicaria Gistel, 1848: xi [unjustified RN]
antennatus Ritsema, 1882: 214 (Diurus) A: TAI ORR
sphacelatus Pascoe, 1887b: 19 (Diurus)

genus Eterodiurus Senna, 1911: 51 type species Eterodiurus singularis Senna, 1911
singularis Senna, 1911: 52 A: YUN ORR
compendarius Kleine, 1925a: 162 (Diurus)

genus Ischnomerus Labram & Imhoff, 1838: fasc. 2 type species Ischnomerus erythroderes Labram & Imhoff,
1838
Cediocera Pascoe, 1887b: 20 type species Cediocera longicornis Pascoe, 1887
Heteroplites Lacordaire, 1865 type species Teramocerus erythroderes Bolieman, 1840 (= Ischnomerus erythroderes
Labram & Imhoff, 1838)
longicornis Pascoe, 1887b: 20 (Cediocera) A: YUN ORR
tristis Senna, 1892a: 181 (Cediocera) A: CH ORR

tribe Microtrachelizini Zimmerman, 1994

genus Aneorhachis Kleine, 1923d: 135 type species Aneorhachis astricta Kleine, 1923
monticola Damoiseau, 1987: 61 A: BT

genus Higonius Lewis, 1883: 299 type species Higonius poweri Lewis, 1883
subgenus Higonodes Zimmerman, 1994: 184 type species Higonius novenarius Damoiseau, 1989
philippinensis Mantilleri, 2009: 914 A: TAI ORR
subgenus Higonius Lewis, 1883: 299 type species Higonius poweri Lewis, 1883
cilo Lewis, 1883: 300 A: AP BT HAI JA SD TAI ORR
crux Olliff, 1883: 300 A: NP ORR
hirsutus Senna, 1893b: 117 A: TAI ORR
bilobicollis Senna, 1898d: 235
ma layanus Mantilleri, 2007: 5
subgenus Taphrocomister Senna, 1895a: 185 type species Taphrocomister singularis Senna, 1895
singularis Senna, 1895a: 185 (Taphrocomister) A: SD ORR
abruptus Kleine, 1935: 75

genus Microtrachelizus Senna, 1893b: 315 type species Trachelizus lyratus Perroud, 1864
Ceunonus Kleine, 1922e: 138 type species Ceunonus minutus Kleine, 1922 (= Microtrachelizus aethiopicus Calabresi,
1920)
apertus Kleine, 1925a: 140 A: SD YUN ORR
attritus Kleine, 1923d: 136 A: UP
beneficus Kleine, 1925a: 139 A: UP ORR
contiguus Senna, 1893c: 266 (Trachelizus) A: UP ORR
cylindricornis Power, 1880: 187 (Trachelizus) A: YUN ORR

tribe Pseudoceocephalini Kleine, 1922

genus Hormocerus Schoenherr, 1823:1137 type species Brentus reticulatus Lund, 1800
Apterorrhinus Senna, 1895b: 59 type species Apterorrhinus compressitarsus Senna, 1895
148 APIONIDAE
reticulata Lund, 1800: 81 {Brentus) A: HAI TAI YUN ORR
annulipes L. W. Schaufuss, 1887: 138 (Hormocerus)
dehanii Oyllenhal, 1833: 360 (Ceocephalus)
scobricollis Boheman, 1845: 373 (Ceocephalus)

genus Peraprophthalmus Kleine, 1923c: 49 type species Peraprophthalmus applicatus Kleine, 1923
applicants Kleine, 1923c: 51 A: UP ORR

genus Rhinopteryx Lacordaire, 1865:449 type species Ceocephalus foveipennis J. Thomson, 1858
foveipennis J. Thomson, 1858: 119 {Ceocephalus) A: SA AFR
carinaerostris Kleine, 1914: 190
errans Kolbe, 1883: 36

genus Schizotrachelus Lacordaire, 1865: 454 type species Schizotrachelus brevicaudatus Lacordaire, 1865
Opisthenoplus Kleine, 1922b: 139 type species Opisthenoplus fascinatus Kleine, 1922
cavus Walker, 1859a: 262 {Ceocephalus) A: SD UP ORR
fascinatus Kleine, 1922b: 140 {Opisthenoplus) A: UP TAI ORR
masculinus Goossens, 2005: 54 A: SD ORR

tribe Trachelizini Lacordaire, 1865

genus Hypomiolispa Kleine, 1918a: 76 type species Miolispa ceylonica Desbrochers des Loges, 1890
gardneri Kleine, 1931: 94 A: SD
incerta Kleine, 1937: 204 A: SD
mikagei Morimoto, 1976: 271 A: JA TAI

genus Miolispa Pascoe, 1862: 393 type species Miolispa suturalis Pascoe, 1862
Cacotrachelus Sharp, 1900: 386 type species Cacotrachelus sculptipennis Sharp, 1900 (= Miolispa salomonensis Senna
1894)
crassifemoralis Kleine, 1917a: 292 A: TAI AUR ORR
cruciata Senna, 1898b: 69 A: JA TAI ORR
discors Senna, 1895c: 358 A: TAI ORR

genus Trachelizus Dejean, 1834: 243 type species Brentus bisulcatus Lund, 1800
bisulcatus Lund, 1800: 67 {Brentus) A: HAI JA UP TAI YUN AUR ORR
bicanaliculatus L. W. Schaufuss, 1885: 206 (Miolispa)
exophthalmus Lea, 1899: 636 (Ceocephalus)
semivelatus L. W. Schaufuss, 1885: 206 (Miolispa)
dividuus Kleine, 1925a: 131 A: SD ORR
japonicus Morimoto, 1976: 272 A: JA
makiharai Morimoto, 1976: 274 A: JA

family APIONIDAE Schoenherr, 1823

subfamily Apioninae Schoenherr, 1823
tribe Apionini Schoenherr, 1823
genus Apion Herbst, 1797: 100 type species Curculio frumentarius Linnaeus, 1758
Apiolum Kirby, 1808: 10 [RN] type species CurculiofrumentariusLinnaeus, 1758
/4/«'oMi/.sRafinesque, 1814: 29 [RN] type species CurculiofrumentariusLinnaeus, 1758
Apium Agassiz, 1846: 29 [RN] type species CurculiofrumentariusLinnaeus, 1758
Apius Billberg, 1820: 40 [RN, HN] type species CurculiofrumentariusLinnaeus, 1758
Erythrapion Schilsky, 1906b: vi type species Apion haematodes Kirby, 1808
Oxystomum Gistel, 1856: 260 type species CurculiofrumentariusLinnaeus, 1758
Oxeostomum Gistel, 1856: 374 type species Curculio frumentarius Linnaeus, 1758
anarchajense Ter-Minasian, 1971b: 659 A: KZ
arcticum Korotyaev, 1988a: 144 A: FE
 APIONIDAE 149
cnientatum Walton, 1844:452 E: AR AU BH BU CR CT CZ DE EN FA FI FR GB GE GR HU IT LA LT LU
NL NR PL PT SB SK SP SV N: AG MO A: CY ES SY WS
desideratum Sharp, 1918a: 4 (Erythrapion)
dellabeffae Schatzmayr, 1922a: 109 E: IT
distincticolle Desbrochers des Loges, 1870a: 197 E: SP
frumentarium Linnaeus, 1758: 378 (Curculio) E: AB AR AU BH BU CR CT CZ DE EN FR GB GE GG GR
HU IR IT LA LT LU NL NR PL PT RO SB SK SP ST SV SZ UK N: AG A: AF CY IN IS JO SY TR
miniatum Germar, 1833: 282
sanguineum DeGeer, 1775: 251 (Curculio)
gallicola Ter-Minasian, 1971b: 659 A:TM
graecum Desbrochers des Loges, 1897c: 4 E: BU CR GG GR ST
kasparyam; Korotyaev, 1988b: 132 (Apion)
haematodes haematodes Kirby, 1808: 76 E: AU BH BU CR CT CZ DE EN FI FR GB GE GG GR HU IR IT
LA LT LU NL NR NT PL PT RO SB SK SL SP SV SZ UK N: MO MR A: SY
brachypterum Sharp, 1918a: 7 (Erythrapion)
coccineum Gmelin, 1790: 1742 (Curculio)
fraudator Sharp, 1918a: 5 (Erythrapion)
haematodes occultum Faust, 1885b: 244 N: AG MO TU
henoni Abeille de Perrin, 1894: 94 N: AG TU
oranense Desbrochers des Loges, 1898b: 35
longithorax Desbrochers des Loges, 1889a: xxxv E: AL IT N: AG MO TU
rubens Walton, 1837: 17 E: AU BU CT CZ DE EN FI FR GB GE IT LA LU NL NR NT PL PT SK SP ST SV
SZ "Caucasus" N: AG A: WS
algiricum Everts, 1879: 59 [DA]
podolicum Rybinski, 1902: 12
rubens Stephens, 1839: 256 [HN]
rubiginosum Grill, 1893: 253 E: AB AR AU BH BU CT CZ DE EN FI FR GB GE HU IT LA LT LU NL NR
NT PL PT SK SP ST SV SZ UK "Caucasus" N: AG A: IN TM SY WS
rufiim A. Solan & F. Solan, 1905: 100
rubroflavum Bajtenov, 1981a: 29 A: KZ
strigipenne Schilsky, 1902: 25 E: AB AR A: TD TM
tribe Aplemonini Kissinger, 1968
genus Aizobius Alonso-Zarazaga, 1990a: 102 type species Apion sedi Germar, 1818
pseudosedi Bajtenov, 1975b: 411 (Apion) A: FE JIA KZ WS
robustirostris Desbrochers des Loges, 1870a: 184 (Apion) E: FR PT SP N: AG MO TU
schaeferi A. Hoffmann, 1960a: 22 (Apion)
sedi Germar, 1818: 49 (Apion) E: AB AU CT CZ DE EN FI FR GB GE GG HU IT LA LT LU NL NR NT PL
RO SK ST SV UK N: AG A: KZ SY WS
maassi Flach, 1908: 130 (Apion)
tumidicolle Bach, 1854: 201 (Apion)

genus Aplemonus Schoenherr, 1847: 5 type species Aplemonus gibbipennis Schoenherr, 1847
arabicus Wagner, 1909a: 27 {Apion) N: EG A: IN OM QA AFR
superbus Tournier, 1868a: 147 (Apion) N: EG "Arabia" AFR
cupreomicans Wagner, 1912b: 36 (Apion)
ennediensis A. Hoffmann, 1964: 1217 (Apion)
stigma G. A. K. Marshall, 1930: 562 (Apion)
tuberculoma Bhateja & Pajni, 1989a: 50 A: UP ORR

genus Cistapion Wagner, 1924: 134 type species Apion cyanescens Gyllenhal, 1833
cyanescens Gyllenhal, 1833: 306 {Apion) E: FR IT PT SP N: AG MO
capiomonti Wencker, 1858b: cv (Apion)
rectinasus Desbrochers des Loges, 1904a: 56 (Apion)
150 APIONIDAE
genus Helianthemapion Wagner, 1930:1386 type species Apion aciculare Germar, 1817
aciculare Germar, 1817b: 245 (Apion) E: AU BU CZ FR GB GE IT LU PL(?) SK SP SZ N: AG MO
lasuretim Schilsky, 1901b: 60 (Apion)
pulchellum Miller, 1857: 15 (Apion)
genuense Osella, 1973a: 198 (Apion) E: IT
kefense Normand, 1921: 78 (Apion) E: SP N: TU
velatum Gerstaecker, 1854b: 278 {Apion) E: AU BE BU CZ FR GG HU IT NL SK SP SZ N: LB MO A: CY IS
SI SY TR
aerugineum Kirsch, 1878: 304 (Apion)
helianthemi Bedel, 1887c: cliv (Apion)

genus Hoplopodapion Solari, 1933: 256 type species Apion kirschi Desbrochers des Loges, 1870
Aphoplopodapion Ehret, 1997a: 205 type species Apion poupillieri Wencker, 1864
depressum Bhateja & Pajni, 1989a: 43 (Onychapiori) A: PA
dumonti Peyerimhoff, 1929a: 15 (Apion) N: AG LB TU A: IS AFR
eremita Peyerimhoff, 1920: 45 (Apion) N: AG LB TU
ferreri Alonso-Zarazaga, 1983a: 18 (Onychapion) E: SP
kirschi Desbrochers des Loges, 1870a: 202 (Apion) N: AG EG LB MO TU A: IS
edmundi Györffy, 1923: 90 (Apion)
rufopurpureum Reitter, 1908a: 54 (Apion)
vincenti Desbrochers des Loges, 1904a: 54 (Apion)
koestlini Schoen, 1993: 64 (Onychapion) N: AG TU
lopatini Ter-Minasian, 1963: 649 (Apion) A: MG TD TM UZ
medvedevi Korotyaev, 1984: 312 (Onychapion) A: TM UZ
poupillieri Wencker, 1864: 127 (Apion) E: SP N: AG EG LB MO TU A: IS SA SI AFR
pumilio Desbrochers des Loges, 1893a: 17 (Apion) E: AB
pyripenne Reitter, 1908a: 55 (Apion) A: IN KZ "Transcaspia"
betpakdalense Bajtenov, 1981a: 27 (Apion)
reaumuriae Ter-Minasian, 1975: 249 (Apion) A: MG
spinitarsatum Bhateja & Pajni, 1989a: 45 (Onychapion) A: PA
tibesticola Alonso-Zarazaga, 1983b: 627 (Onychapion) [RN] N: AG A: AE QA AFR
grande A. Hoffmann, 1962b: 438 (Apion) [HN]

genus Margaritapion Korotyaev, 1990b: 220 type species Apion nitrariae Ter-Minasian, 1970
nitrariae Ter-Minasian, 1970: 633 (Apion) A: KZ MG TR

genus Onychapion Schilsky, 1901b: E type species Apion tamarisci Gyllenhal, 1839
fasciculosum Motschulsky, 1851: 657 (Apion) N: EG
tamarisci Gyllenhal, 1839: 388 (Apion) E: AB FR IT (Sardegna, Sicilia) PT SP N: AG MO EG TU A: AF JO
SI ORR
gautardi Tournier, 1868a: 146 (Apion)
tamarixus Menon & Parshad, 1961: 85 (Apion)

genus Osellaeus Alonso-Zarazaga, 1990a: 93 type species Apion bonvouloirii C. N. F. Brisout de Barneville,
1880
bonvouloirii baldensis Bellô, Meregalli & Osella, 1981: 519 (Apion) E: IT
bonvouloirii bonvouloirii C. N. F. Brisout de Barneville, 1880: 232 (Apion) E: FR IT SZ
georgeli A. Hoffmann, 1929a: 43 (Apion)

genus Perapion Wagner, 1907c: 259 type species Apion curtirostre Germar, 1817
subgenus Eroosapion Ehret, 1994: 20 type species Apion lemoroi C. N. F. Brisout de Barneville, 1880
lemoroi C. N. F. Brisout de Barneville, 1880: 231 (Apion) E: AB CR FR GE GB HU IT SB SK SP ST UK N:
AG TU A: FE KZ MG SY TM
subgenus Hemiperapion Wagner, 1930: 1385 type species Apion horvathi Schilsky, 1901
altnemelicum Bajtenov, 1973: 60 (Apion) A: KZ
 APIONIDAE 151
centrasiaticum Bajtenov, 1981b: 35 {Apion) A: MG
chioneum Iablokoff-Khnzorian, 1957: 174 {Apion) E: AB AR A: KZ MG
turanicum Bajtenov, 1973: 60 {Apion)
ehreti Legalov, 2000: 93 A: KZ
emeljanovi Korotyaev, 1984: 312 A: MG
horvathi Schilsky, 1901a: 153 (Apion) E: AR A: TR
calligonicola Iablokoff-Khnzorian, 1957: 173 (Apion)
jacobsoni Wagner, 1910a: 153 (Apion) E: ST A: KI KZ TM UZ
mesasiaticum Ter-Minasian, 1966a: 100 (Apion)
terminassianae Legalov, 2000: 91 A: MG
subgenus Perapion Wagner, 1907: 259 type species Apion curtirostre Germar, 1817
affine Kirby, 1808: 68 (Apion) E: AU BU CZ DE EN FI FR GB GE HU IT LA LT LU NL NR NT PL SB SK
ST SV "Caucasus" N: AG A: SY
arenarium Bajtenov, 1978b: 73 (Apion) A: KZ
barkleyi Alonso-Zarazaga, nom. nov. [see New Acts] A: TM
smreczynski Bajtenov, 1979c: 369 (Apion) [HN]
connexion Schilsky, 1902: 28 (Apion) E: CT MD ST UK A: KZ UZ WS
arcuatum Bajtenov, 1977a: 15 (Apion)
curtirostre Germar, 1817b: 230 (Apion) E: AB AR AU CR CT CZ DE EN FI FR GB GE GG HU IR IT LA LT
LU NL NR NT PL RO SB SK ST SV SZ UK A: JA "Siberia" NARi
aquilinum Boheman, 1839: 440 (Apion)
humile Germar, 1817b: 232 (Apion)
medianum C. G. Thomson, 1865:48 (Apion)
tenellum C. R. Sahlberg, 1834: 18 (Apion)
dealbatum Bajtenov, 1980b: 129 (Apion) A: KZ
eretzisrael Friedman & Freidberg, 2007: 106 A: IS
fallax Wollaston, 1864: 313 (Apion) N: CI MO TU
neofallax Warner, 1958: 348 (Apion) [unnecessary RN]
hartmannianum Wagner, 1910a: 153 (Apion) A: TIA
hydrolapathi Marsham, 1802: 249 (Curculio) E: AU BE CR DE FR GB GE GR IR IT NL PT SP ST SV N: AG
MO TU A: SY
caeruleopenne Stephens, 1831: 173 (Apion)
lapathorum Kirby, 1808: 67 (Apion)
ilvense Wagner, 1905: 38 (Apion) E: FR GB GR IT PT SP N: AG MO TU A: SY TR
jelineki Bajtenov & Fremuth, 1981: 247 (Apion) A: IN
kuraense Bajtenov, 1982b: 69 (Apion) E: GG
marchicum Herbst, 1797: 118(" 128") (Apion) E: AU BH CR CT CZ DE EN FI FR GB GE GG GR HU IR IT
L A L T L U N L N R N T P L P T S B S K S P S V U K N: AG A:SYWS
laevithorax Gyllenhal, 1833: 288 (Apion)
rumicis Kirby, 1808: 67 (Apion)
spartii Kirby, 1808: 56 (Apion)
violaceum Gyllenhal, 1813: 50 [HN] (Apion)
marseuli Wencker, 1864: 261 (Apion) N: AG LB MO TU A: AE IN IS QA SA
ormusense Bajtenov & Fremuth, 1981: 248 (Apion)
myochroum Schilsky, 1902: 29 (Apion) A: AF KZ MG UZ
oblongum Gyllenhal, 1839: 421 (Apion) E: AB AR AU BH BU CT CZ GE GG HU LA LT PL RO SB SK ST
UK A: ES HEI KI MG SY TR UZ WS
sibiricum Boheman, 1839:442 (Apion)
violaceum lopezi A. Hoffmann, 1957a: 58 (Apion) E: SP (Mallorca)
violaceum violaceum Kirby, 1808: 65 (Apion) E: AU BH BU CR CT CZ DE EN FI FR GB GE GR HU IR IT
LA LT LU ME NL NR NT PL PT RO SB SK SP ST SV UK "Caucasus" N: AG LB MO TU A: CY ES IS
NC SC JA SY TR WS
alpinum Wencker, 1864: 254 (Apion)
harcyniae Hubenthai, 1911: 192 (Apion)
kaufinanni Bokor, 1923: 100 (Apion)
obscurum Gerhardt, 1898: 335(Apion) [HN]
152 APIONIDAE
viilpecula Schubert, 1940: 39 {Apion) A: TM
subgenus Rhaphidoplectron Alonso-Zarazaga, 1990a: 104 type species Apion defensum Faust, 1887
defensum Faust, 1887e: 177 (Apion) A: AF IN KZ TD UZ
himalayense Bhateja & Pajni, 1989a: 46 A: AF HP KA PA UP ORR
miristanicola Wanat, 1995b: 487

genus Phrissotrichum Schilsky, 1901b: 4 type species Apion tubiferum Gyllenhal, 1833
subgenus Phrissotrichum Schilsky, 1901b: 4 type species Apion tubiferum Gyllenhal, 1833
boiteli Normand, 1939a: 52 {Apion) E: IT SP N: AG LB
joannium Ehret, 1997b: 109 E_PTSP N: MO
sicanum Ehret, 1997b: 108 E: FR GR IT PT SP N: AG LB MO TU A: TR
tubiferum Gyllenhal, 1833: 284 (Apion) E: BU CR FR GG GR IT MC PT SP N: AG LB MO MR TU A: CY
IS SY TR
tubuliferum Wollaston, 1864: 312 (Apion) N: CI
wenckeri C. N. F. Brisout de Barneville, 1863: 96 (Apion) E: FR PT SP
subgenus Schilskyapion Alonso-Zarazaga, 1990a: 90 type species Apion perrisii Wencker, 1859
apenninicola Osella, Zuppa & Altea, 1993: 56 E: IT
brevipile Desbrochers des Loges, 1895d: 83 (Apion) E: IT N: AG MO TU
delphinense Hustache, 1913: 408 (Apion) E: FR IT
grenieri Desbrochers des Loges, 1875a: 31 (Apion) E: FR IT
leonhardi Reitter, 1903: 178 (Apion) E: BH
moroderi Desbrochers des Loges, 1907: 85 (Apion) E: SP
osellai Sacco & Colonnelli, 1985: 441 (Apion) E: IT
perrisii Wencker, 1859b: ccxxxviii (Apion) E: FR PT SP
revelieri Perris, 1869: 24 (Apion) E: FR (Corse) IT (Sardegne) N: AG
diversum Desbrochers des Loges, 1870a: 186 (Apion)
rugicolle Germar, 1817b: 201 (Apion) E: AU BH CZ FR GE HU IT SK SZ
hirsutum A. Villa & G. B. Villa, 1835: 49 (Apion)
setiferum Gyllenhal, 1833: 266 (Apion)

genus Pseudaplemonus Wagner, 1930: 1385 type species Apion limonii Kirby, 1808
Cypriapion Ehret, 1997a: 204 type species Apion chevrolati Gyllenhal, 1833
aeneicollis Gerstaecker, 1854b: 277 (Apion) N: AB GG ST A: IN IQ IS MG SA SY TM TR UZ "Siberia"
aereirostre Desbrochers des Loges, 1901a: 77 (Apion)
laudabile Faust, 1891c: 415 (Apion)
albanicus Formânek, 1925: 27 (Apion) E: AL
artemisiae Moravitz, 1861: 292 (Apion) E: ST "Caucasus" A: WS
bermani Korotyaev, 1997: 285 A: FE
chevrolati Gyllenhal, 1833: 260 (Apion) E: FR IT PT SP SZ N: MO
carbonarium Everts, 1879: 60 {Apion)
externepunctatus Desbrochers des Loges, 1875a: 30 (Apion) E: CT RO ST
limonii Kirby, 1808: 78 (Apion) E: BE FR GB GE NL PT SP N: AG MO
limoniastri Flach, 1908: 130 (Apion)
salarius Ehret, 1997a: 199
martjanovi Faust, 1891c: 413 (Apion) A: ES MG WS
prasolovi Korotyaev, 1991: 875 (Perapiori) A: KZ
genus Pseudoperapion Wagner, 1930:1385 type species Apion brevirostre Herbst, 1797
brevirostre Herbst, 1797: 120("130") (Apion) E: AU BU CR CT CZ DE FR GE HU IT LU NL PL PT RO SB
SK SP ST SV SZ "Caucasus" A: IS SY TR "Siberia"
interstitielle Boheman, 1839: 443 (Apion)
liebmanni Schubert, 1958: 32 (Apion) N: AG TU
laportei A. Hoffmann, 1960b: 169 (Apion)
 APIONIDAE 153
genus Pseudostenapion Wagner, 1930:1386 type species Apion simum Germar, 1817
simum Germar, 1817b: 235 {Apion) E: AU CR CT CZ DE EN FI FR GE HU IT LA LT LU NL PL RO SB SK
SP ST SV UK "Caucasus" N: AG

tribe Aspidapiini Alonso-Zarazaga, 1990

genus Alocentron Schilsky, 1901b: G type species Apion curvirostre Gyllenhal, 1833
subgenus Alocentron Schilsky, 1901b: G type species Apion curvirostre Gyllenhal, 1833
curvirostre Gyllenhal, 1833: 264 (Apion) E: AB AR AU BU CZ GE GG HU RO SB SK SL ST N: AG A: IS
SY TR WS
nonveilleri A. Hoffmann, 1953b: 194 {Apion)
subgenus Bulborhinapion Schatzmayr, 1926: 143 type species Apionpachyrrhynchum Gemminger, 1871
pachyrrhynchum Gemminger, 1871a: 123 (Apion) [RN] A: ES FE JA SC
bulbinasum Sharp, 1891: 295 (Apion)
crassirostre Motschulsky, 1860: 169 (Apion) [HN]
schrencki Schilsky, 1906b: cxviii {Apion) [RN]
subgenus Nearctalox Alonso-Zarazaga, 1990a: 54 type species Apion hibisci Fall, 1918
tschukotkiamim Bajtenov, 1980a: 111 (Apion) A: FE

genus Aspidapion Schilsky, 1901b: G type species Apion validum Germar, 1817
subgenus Aspidapion Schilsky, 1901b: G type species Apion validum Germar, 1817
Euapion Iablokoff-Khnzorian, 1959a: 75 type species Apion uncinatum Iablokoff-Khnzorian, 1959
acerifoliae Suppantschitsch, 1996: 227 N: CI
caprai Giusto, 1993: 551 E: IT
inarmatum Korotyaev, 1985: 143 A: SD YUN ORR
mountanus Pajni, Bhateja & Kumar, 1991: 121 (Apion) A: HP KA
panfilovi Korotyaev, 1985: 134 A: YUN ORR
radio lus chalybeipenne Wollaston, 1854: 413 (Apion) E: AZ MR N: CI
albescens Wagner, 1906a: 20 (Apion)
radiolus radiolus Marsham, 1802: 247 (Curculio) E: AB AR AU AZ BU CT CR CZ DE EN FI FR GB GE GG
GR HU IR IT LA LT LU NL NR NT PL PT RO SB SK SP ST SV SZ UK N: AG LB MO TU A: AF CY
IQ IS JO KZ LE SY TD TR UZ WS AFR
ferruginipes Wencker, 1864: 166 (Apion)
nigrescens Stephens, 1831: 176 (Apion)
oxurum Kirby, 1808: 73 (Apion)
rougeti Wencker, 1858b: cvi (Apion)
soror Rey, 1895c: 74 (Apion) E: CR FR GB GR IT SP A: TR N: AG TU
foveatoscutellatum Wagner, 1906a: 19 (Apion)
uncinatum Iablokoff-Khnzorian, 1959a: 74 (Apion) E: AR
validum Germar, 1817b: 246 (Apion) E: AB AR AU CZ GE GG HU IT KZ PL RO SK SL ST SZ UK N: AG
A: IN KI KZ SY TD TM TR UZ WS
grimmii Hochhuth, 1847: 462 (Apion)
subgenus Koestlinia Alonso-Zarazaga, 1990a: 53 type species Attelabus aeneus Fabricius, 1775
aeneum Fabricius, 1775: 131 (Curculio) E: AB AR AU BU CR CZ DE FR GB GE GG GR HU IT LU MC NL
PL PT SB SK SP ST SV SZ UK N: AG MO TU A: AF CY IN IS KI KZ LE SY TM UZ
chalceus Marsham, 1802:243 (Curculio)
motschulskyi Hochhuth, 1847: 461 (Apion)
oblongus Goeze, 1777: 380 (Curculio) [HN]
oblongus Geoffroy, 1785: 114 (ßhinomacer)
teres Gmelin, 1790: 1758 (Curculio)

genus Flavopodapion Korotyaev, 1987b: 103 type species Apion morosum Faust, 1898
gilvipes Gemminger, 1871a: 123 (Apion) [RN] A: FUJ NP SD TAI YUN ORR
flavimanum Motschulsky, 1858: 94 (Apion) [HN]
pictum Wagner, 1909: 26 (Apion)
shaowuanum Voss, 1958: 19 (Apion)
154 APIONIDAE
genus Harpapion Voss, 1966: 211 type species Apion considerandum Fâhraeus, 1871
ceylonicum Gerstaecker, 1854a: 258 (Apion) [DA] A: YUN ORR
cingalense Walker, 1859a: 263 (Apion)
singhalense Heller, 1902:341 (Apion)
dongollanum Wagner, 1910a: \54 (Apion) A: SA AFR

genus Miniapion Korotyaev, 1992a: 40 type species Miniapion potanini Korotyaev, 1992
potanini Korotyaev, 1992a: 41 A: SCH
rubroplagiatum Voss, 1958: 132 (Apion) A: FUJ
sulcirostre Sharp, 1891: 295 (Apion) A: FE JA (Honshu)

genus Pseudaspidapion Wanat, 1990: 58 type species Apion spadiceum Wagner, 1908
brunneorufum Balfour-Browne, 1945: 55 (Apion) A: YE AFR ORR
coelebs Korotyaev, 1987b: 100 (Aspidapion) A: YUN
kryzhanovskii Korotyaev, 1987b: 95 (Aspidapion) A: YUN
rufopiceum Wagner, 1909a: 26 (Apion) A: TAI
scorteccii Voss, 1971c: 541 (Apiotherium) A: YE AFR
striopunctatum Pajni, Bhateja & Kumar, 1991: 125 (Apion) A: HP ORR
yunnanicum Korotyaev, 1985: 139 (Aspidapion) A: YUN
zagulajevi Korotyaev, 1987b: 97 (Aspidapion) A: YUN

tribe Ceratapiini Alonso-Zarazaga, 1990

genus Acentrotypus Alonso-Zarazaga, 1990a: 51 type species Apion brunnipes Boheman, 1839
brunnipes Boheman, 1839: 386 (Apion) E: BE BH BU CR CZ FR GB GE GG HU IT NL PL PT SP SZ N: AG
MO TU A: IN TR
laevigatum Kirby, 1808: 70 (Apion) [permanently invalid name]

genus Ceratapion Schilsky, 1901b: F type species Apion carduorum Kirby, 1808
subgenus Acanephodus Alonso-Zarazaga, 1990a: 46 type species Apion onopordi Kirby, 1808
onopordi onopordi Kirby, 1808: 71 (Apion) E: AB AL AR AU BE BH BU CR CT CZ DE EN FI FR GB GE
GG GR HU IT LA LT LU MC MD ME NL NR NT PL PT RO SB SK SL SP ST SV SZ TR UK N: AG TU
A: CH ES FE IN KI KZ TM TR UZ WS
frater Desbrochers des Loges, 1870a: 200 (Apion)
hipponense Desbrochers des Loges, 1894c: 87 (Apion)
jablokovkhnzoriani Bajtenov, 1982b: 69 (Apion)
rugipenne Hochhuth, 1851 : II (Apion)
onopordi parviclava Desbrochers des Loges, 1897c: 8 (Apion) E: AR GG GR N: EG A: CY IN IS JO LE SY
TR
cavatum Desbrochers des Loges, 1907: 89 (Apion)
chenoeephalum Desbrochers des Loges, 1902c: 160 (Apion)
quadricostatum Schilsky, 1906a: 8 (Apion)
parens Desbrochers des Loges, 1870a: 161 (Apion) E: PT SP N: AG MO TU
rectipes Desbrochers des Loges, 189Id: lvi [=189lb: 83] (Apion)
subgenus Angustapion Wanat, 1995a: 299 type species Apion akbesianum Desbrochers des Loges, 1897
aegyptiacum Desbrochers des Loges, 1870a: 201 (Apion) N: EG
akbesianum Desbrochers des Loges, 1897c: 10 (Apion) A: TR
austriacum Wagner, 1904: 374 (Apion) E: AU BE BH CZ DE EN FR GE HU LA LU MD PL RO SK SZ UK
beckeri Desbrochers des Loges, 1875a: 27 (Apion) E: AB AR BU GG HU IT MC RO SB ST TR UK A: AF IN
IS LE TD TM TR UZ
angulirostre Schilsky, 1901b: 14 (Apion)
biseriatum Desbrochers des Loges, 1875a: 26 (Apion)
sublaevithorax Desbrochers des Loges, 1894c: 90 (Apion)
boehmi Wagner, 1911: 48 (Apion) N: EG
bokharanum Györfry, 1923: 91 (Apion) A: UZ
 APIONIDAE 155
cylindricolle Gyllenhal, 1839: 380 {Apion) E: BU FR GG HU IT RO SB SK ST UK A: TR
longiceps Schilsky, 1906a: 5 {Apion)
montandoni Desbrochers des Loges, 1897c: 13 {Apion)
decolor Desbrochers des Loges, 1875a: 27 {Apion) E: AB AR BU CR CT FR GR ME RO SL N: EG A: IN IS
JO LE SY TR WS
brevithorax Desbrochers des Loges, 1897c: 12 {Apion)
efratense Bajtenov & Lodos, 1978b: 147 {Apion)
parcius Voss, 1965: 333 {Apion)
fremuthi Wanat, 1995a: 365 E: AB A: IN TR
gibbiceps Desbrochers des Loges, 1894b: 66 {Apion) A: IN
kasbekianum Gerstaecker, 1854a: 238 {Apion) E: AB AR GG A: TR
angustissimum Desbrochers des Loges, 1889a: xxxv {Apion)
Uapperichi Wanat, 1995a: 311 A: IS JO
lancirostre Chevrolat, 1859: 385 {Apion) N: AG EG MO TU
lanciferum Desbrochers des Loges, 1897c: 9 {Apion) [unnecessary RN]
libicum Wanat, 1995a: 378 N: LB
longiclava Desbrochers des Loges, 1897c: 12 (Apion) E: FR IT SP N: AG MO TU
clavatum Schilsky, 1906a: 3 (Apion)
macrorrhynchum Eppelsheim, 1888: 381 (Apion) E: BU GR MC SP A: IN TR
mlderae G. A. K. Marshall, 1938: 167 (Apion) A: HP
opacinum Faust, 1887b: 179 (Apion) A: FE JA (Honshu) NC WS
peninsulae Wanat, 1990: 66 A: YE
perlongum Faust, 1891c: 410 (Apion) E: RO ST
poggii Wanat, 1995a: 319 N: LB
ramoni Friedman & Freidberg, 2007: 78 A: IS
sefrense Desbrochers des Loges, 1897c: 10 (Apion) N: AG TU
angustius Desbrochers des Loges, 1898b: 35 (Apion)
sejugum Desbrochers des Loges, 1893b: 10 (Apion) E: "Caucasus" A: IS SY TR
transsylvanicum Schilsky, 1906a: 10 (Apion) E: HU RO UK
subgenus Ceratapion Schilsky, 1901b: F type species Apion carduorum Kirby, 1808
armatum Gerstaecker, 1854a: 237 (Apion) E: AR AU BE BH BY CR CZ DE EN FR GB GE HU IT LA NL PL
RO SK SL ST SV SZ
barnevillei Wencker, 1864: 133 (Apion)
calcaratum Wollaston, 1864: 310 (Apion) N: CI
carduorum Kirby, 1808: 72 (Apion) E: AL AR AU BE BH BU CR DE FR GB GE GG GR HU IR IT LU MC
NL PL PT RO SB SK SL SP ST SV SZ UK N: AG MO TU A: AF ES SY TR
ferdinandi Schatzmayr, 1925: 96 (Apion)
galactidis Wencker, 1858a: xxii (Apion)
lacertense Tottenham, 1941: 14 (Apion)
magyaricum Györffy, 1923: 93(Apion)
meridianum Wencker, 1864: 130 (Apion)
dalmatinum Györffy, 1923: 92 (Apion) E: CR IT
damryi Desbrochers des Loges, 1894c: 91 (Apion) E: BH BU CR FR GR IT MC PT RO SL SP SZ N: AG LB
MO TU A: IS TR
gridellii Solan, 1940: 70 (Apion)
gibbifrons Hustache, 1932b: 136 (Apion) A: IN LE SY TR
cystocephalum Bajtenov & Lodos, 1978b: 146 (Apion)
gibbirostre Gyllenhal, 1813: 52 (Apion) E: AB AR AU BE BH BU CR CT CZ DE EN FI FR GB GE GG GR
HU IR IT LA LU MA MC MD ME NL NR NT PL PT RO SB SK SL SP ST SV SZ UK N: AG EG MO TU
' A: CY ES FE IN IS KZ LE SY TR WS
conforme Desbrochers des Loges, 1875a: 27 (Apion)
cyaneus DeGeer, 1775: 252 (Curculio) [HN]
kenedii Bokor, 1923: 96 (Apion)
russicum Desbrochers des Loges, 1870a: 179 (Apion)
tumidum Stephens, 1835: 420 (Apion)
kazakhstanicum Ter-Minasian, 1970: 634 (Apion) E: GG A: KZ WS
156 APIONIDAE
secundum Ter-Minasian, 1975: 252 (Apion) E: ST A: ES KZ MG TR
dentirostre Ter-Minasian, 1972: 241 (Apion) [HN]
subgenus Clementiellus Alonso-Zarazaga, 1990a: 46 type species Apion robusticorne Desbrochers des Loges,
1866
orientale Gerstaecker, 1854a: 237 (Apion) E. AU BH BU CR GR HU IT RO SK ST UK A: SY TR
henschi Reitter, 190le: 226 (Apion)
similans Schilsky, 1901b: 21 (Apion)
robusticorne Desbrochers des Loges, 1866: 44 (Apion) E: IT MA PT SP N: AG MO TU
bipartitum Desbrochers des Loges, 1902c: 160 (Apion)
insolitum Desbrochers des Loges, 1870a: 196 (Apion)
subgenus Echinostroma Alonso-Zarazaga, 1990b: 450 type species Apion penetrans Germar, 1817
basicorne Illiger, 1807a: 307 (Apion) E: AB AL AR AU BE BH BU CR CT CZ DE FR GE GG GR HU IT LA
NL NT PL RO SB SK SP ST SV SZ UK A: CY IN IS LE TR
atripenne Desbrochers des Loges, 1902c: 159 (Apion)
intermedium Desbrochers des Loges, 1908e: 97 (Apion) [HN]
simillimum Desbrochers des Loges, 1891e: lvii [= 1891c: 83] (Apion)
spathula Desbrochers des Loges, 1894c: 98 (Apion)
subcavifrons Desbrochers des Loges, 1894c: 87 (Apion)
subdentirostre Desbrochers des Loges, 1875a: 27 (Apion)
tauricum Desbrochers des Loges, 1902c: 159 (Apion)
curtii Wagner, 1920: 196 (Apion) E: CR FR GR IT
dentirostre Gerstaecker, 1854a: 236 (Apion) E: SP N: MO
penetrans caullei Wencker, 1858a: xxi (Apion) E: BE FR GE NL SZ
penetrans penetrans Germar, 1817b: 244 (Apion) E: AL AU BH BU BY CR CT CZ DE EN GE GR HU IT LA
LT LU MD ME PL RO SB SK SL ST SV SZ UK A: KZ TR
disions Desbrochers des Loges, 1889a: xxxiii (Apion)
ovipenne Hochhuth, 1851: 9 (Apion)
subconicicolle Desbrochers des Loges, 1870a: 199 (Apion)
penetrans wanati Alonso-Zarazaga, 1993b: 233 [= 1993a: 123] E: PT SP N: AG MO
rhopalorrhynchum Iablokoff-Khnzorian, 1967: 88 (Apion) E: AR A: TR
scalptum scalptum Mulsant & Rey, 1858: 267 (Apion) E: AU BH CR FR GE IT MC PT SP UK N: AG TU
scalptum caviceps Desbrochers des Loges, 1870a: 201 (Apion) E: AB AR BU GG GR ST TR UK A: AF CY
IN IS KI KZ LE PA SY TD TM TR UZ
pilicorne Desbrochers des Loges, 1875a: 26 (Apion)
uniseriatum Faust, 1885a: 185 (Apion) A: AF IN KI KZ SY TD TM UZ
insculpticolle Desbrochers des Loges, 1902c: 160 (Apion)
remaudierei A. Hoffmann, 1956: 248 (Apion)
sculpticolle Desbrochers des Loges, 1897c: 8 (Apion)

genus Ceratapion, species incertae sedis

bacanasicum Bajtenov, 1973: 61 (Apion) A: KZ
brevitibia Bajtenov, 1977a: 16 (Apion) A: TR
edentatum Desbrochers des Loges, 1891d: lvi [= 1891c: 83, 1894d: 87] (Apion) N: AG
fallaciosum Desbrochers des Loges, 1892a: 107 (Apion) E: IT N: AG MO TU
nurense Bajtenov, 1977a: 16 (Apion) A: KZ
proximum Györfry, 1923: 92 (Apion) A: UZ

genus Diplapion Reitter, 1916a: 241 type species Apion stolidum Germar, 1817
confluens Kirby, 1808: 62 (Apion) E: AB AL AR AU BU BY CR CT CZ DE EN FI FR GB GE GG GR HU IR
ITKZLALTLUMCMDNLNRNTPLPTROSBSKSPSTSVSZTRUK N:AGTU A:CYINISJO
SYTR
asiaticum Desbrochers des Loges, 1894d: 116 (Apion)
inapertum Desbrochers des Loges, 1897c: 13 (Apion)
roelofsi Everts, 1879: 58 (Apion)
detritum Mulsant & Rey, 1858: 261 (Apion) E: AB BH BU CR CT CZ DE FR GE GG GR HU IT KZ LA MA
MC MD ME PL RO SB SK SP ST UK N: AG EG MO TU A: CY IN SI SY TR
catenulatum Wagner, 1910b: 944 (Apion)
 APIONIDAE 157
crenulatum Desbrochers des Loges, 1894d: 115 (Apion)
ragusae Everts, 1879: 58 (Apion)
rumaniacum Wagner, 1910b: 944 (Apion)
subcrenulatum Desbrochers des Loges, 1901a: 79 (Apion)
viridicoeruleum Everts, 1879: 59 (Apion)
hamatum Wagner, 1906a: 22 (Apion) E: AB AR A: IN
niiens Schilsky, 1901b: 24 (Apion) E: IT(Sicilia) SP N: AG MO TU
sareptanum Desbrochers des Loges, 1867b: 216 (Apion) E: RO ST A: TR
curtipenne Desbrochers des Loges, 1870a: 179 (Apion)
saudiarabicum Wanat, 1995a: 189 A: SA
squamans Desbrochers des Loges, 1907: 89 (Apion) E: AR N: EG A: IS JO SY
squamuliferum Desbrochers des Loges, 1891c: 323 (Apion) E: IT PT SP N: AG LB MO MR TU
subsquamiferum Desbrochers des Loges, 1894d: 115 (Apion)
stolidum Germar, 1817b: 218 (Apion) E: AB AR AU BH BU BY CR CT CZ DE EN FI FR GB GE GR HU IR
IT KZ LA LT LU MC MD NL NR NT PL PT RO SB SK SL SP ST SV SZ UK N: AG MO A: CY KZ WS
westwoodi Wollaston, 1864: 311 (Apion) N: CI

genus Omphalapion Schilsky, 1901b: G type species Curculio laevigatus Paykull, 1792
beuthini An. Hoffmann, 1874: 208 (Apion) E: FR GB SP N: MO
buddebergi Bedel, 1887b: 363 (Apion) E: BU GE LA PL SP ST A: IN
extinctum Kraatz, 1888: 174 (Apion)
concinnum Schilsky, 1906b: cxviii (Apion) [RN] A: IN IS LE
puncticolle Schilsky, 1906a: 27 (Apion) [HN]
dispar Germar, 1817b: 181 (Apion) E: AR AU BE BH BU BY CR CZ DE EN FR GE GR HU IT LA MA NL
PL RO SK SL ST SV SZ UK A: TR
brisouti Bedel, 1887b: 363 (Apion)
corcyraeum Schilsky, 1906b: 2 (Apion)
hookerorum Kirby, 1808: 69 (Apion) E: AR AU BU BY CT CZ DE EN FI FR GB GE GR HU IT LA LT LU
MD NL NR NT PL PT SK SP ST SV SZ UK N: AG LB MO A: CY IN LE SY TR WS
laevigatum Paykull, 1792: 141 (Curculio) E: AL AU BH BU BY CR CT CZ DE EN FI FR GB GE GG GR HU
IT KZ MD NL NR NT PL PT SB SK SL SP ST SV SZ UK A: IN KZ TR
carbonarium Germar, 1817b: 176 (Apion)
sahlbergi Gyllenhal, 1825: 61 (Apion)
sorbi Fabricius, 1792: 390 (Attelabus)
viridescens Marsham, 1802: 249 (Curculio)
fossicolle Desbrochers des Loges, 1889a: xxxiv (Apion) A: SY TR
fossulatum Desbrochers des Loges, 1897c: 15 (Apion)
pseudodispar Wanat, 1995a: 146 E: AU BH BU CZ HU IT MC MD TR UK A: CY IN TR
rhodopense Angelov, 1962: 199 (Apion) E : B U G R

genus Protoceratapion Wanat, 1995a: 76 type species Apion deletum Schilsky, 1906
Protosynapion Legalov, 1997b: 468 type species Apion deletum Schilsky, 1906
deletum Schilsky, 1906a: 38 (Apion) A: KI KZ UZ WS
egregium Wagner, 1906b: 204 (Apion)
tibetanum Balfour-Browne, 1944a: 8 (Apion) A: XIZ

genus Taphrotopium Reitter, 1916a: 241 type species Apion sulcifrons Herbst, 1797
subgenus Omphatopium Wanat, 1995a: 106 type species Apion irkutense Faust, 1888
irkutense Faust, 1888b: 174 (Apion) A: ES KZ WS
subgenus Taphrotopium Reitter, 1916a: 241 type species Apion sulcifrons Herbst, 1797
cuprijulgens Schilsky, 1906b: XI (Apion) E: AB AR A: IN KI KZ TD TM UZ
karatavicum Bajtenov, 1973: 61 (Apion)
steveni Gyllenhal, 1839: 393 (Apion) E: AB AR ST A: IN KI KZ MG TM UZ WS XIN
aterrimum Schilsky, 1906b: XI (Apion) (incorrect subsequent spelling)
nigerrimum Faust, 1894a: 65 (Apion)
158 APIONIDAE
sulcifrons Herbst, 1797: 122("132") (Apion) E: AU BY CT CZ DE EN FR GE GG HU LA LT MD NT PL RO
SK SP ST SV UK A: KZ WS SY

tribe Exapiini Alonso-Zarazaga, 1990

genus Exapion Bedel, 1887a: 360 type species Curculio fuscirostris Fabricius, 1775
Ulapion Ehret, 1997a: 208 type species Curculio ulicis Forster, 1771
approximation Desbrochers des Loges, 1904a: 55 (Apion) N: MO
batnense Desbrochers des Loges, 1894e: 30 (Apion) N: AG
chobauti Schilsky, 1906a: 19 (Apion)
breviusculum Rosenhauer, 1856: 239 (Apion) E: PT SP
canescens Desbrochers des Loges, 1889a: xxxiv (Apion) E: GR TR A: CY IS JO LE SY TR
Judaicum Schilsky, 1906a: 18 (Apion)
compactum asturiense Wagner, 1914a: 27 (Apion) E: PT SP
compactum circovichi Schatzmayr, 1921: 83 (Apion) E: IT SL
compactum compactum Desbrochers des Loges, 1888: cxciii (Apion) E: AU BE BH CR CT CZ DE FR GE HU
IT LU NL PT SB SK SL SP SZ N: AG A: WS
compactum monticola Schilsky, 1906a: 15 (Apion) E: AU BH IT
compactum pseudinexpertum Schatzmayr, 1925: 38 (Apion) E: SZ
confusum Desbrochers des Loges, 1891e: lvii [= 1891b: 83] (Apion) N: AG TU
corniculatum Germar, 1817b: 129 (Apion) E: AU BE BH BU CR CT CZ FR GE HU IT NL PL RO SB SK SL
UK
hungaricum Desbrochers des Loges, 1894f: 44 (Apion)
crassiusculum Desbrochers des Loges, 1894e: 32 (Apion) E: FR
difficile Herbst, 1797: 114 ("124") (Apion) E: AU CZ DE FR GB GE HU IT LU NL PL RO SB SK SL A: TR
germanicum Desbrochers des Loges, 1894e: 32 (Apion)
kiesenwetteri Desbrochers des Loges, 1870a: 204 (Apion)
toelgi Wagner, 1914a: 25 (Apion)
elongatissimum Desbrochers des Loges, 1870a: 198 (Apion) E: PT SP
elongatulum Desbrochers des Loges, 1891c: 324 (Apion) [RN] E: AU BH BU CR CT CZ FR GE HU IT PL RO
SB SK ST SZ UK
longulum Desbrochers des Loges, 1875a: 30 (Apion) [HN]
vittigerum Wagner, 1910b: 947 (Apion)
fasciolatum Wagner, 1912c: 88 (Apion) A: FR SP
heikertingeri Wagner, 1918: 97 (Apion) [RN]
valentinorum A. Hoffmann, 1929c: 314 (Apion) [unjustified emendation]
valentinum Clermont, 1919: 316 (Apion)
flavinasus Desbrochers des Loges, 1907: 86 (Apion) E: ST
formaneki Wagner, 1929b: 166 (Apion) E: AU BE BH BU CR CZ FR GE HU IT LU PL RO SB SK SL SZ YU
comoni A. Hoffmann, 1958a: 1556 (Apion)
fiilvum Desbrochers des Loges, 1895a: 52 (Apion) E: FR (Corse) IT (Sardegne) N: TU
fuscirostreflachi Wagner, 1906b: 204 (Apion) E: PT
fuscirostre fuscirostre Fabricius, 1775: 131 (Curculio) E: AU CZ DE FR GB GE HU IT LU NL PL PT SK SP
SV N: AG A: SY NARi
albovittatum Herbst, 1784a: 74 (Curculio)
curvirostre Herbst, 1784a: 78 (Curculio) [DA]
melanopus Marsham, 1802:248 (Curculio)
venus tum Herbst, 1795: 163 (Curculio) [HN, RN]
gattefossei Hustache, 1937: 15 (Apion) N: MO
genistae Kirby, 1811: 347 (Apion) E: FR GB GE GR IT PL PT SP N: AG
astragali Gozis, 1881b: 181 (Apion) [HN] [unjustified emendation]
astralagi Herbst, 1795: 98 (Curculio) [see New Acts]
bivittatum Gerstaecker, 1854a: 259 (Apion)
funiculare Mulsant & Rey, 1858:269 (Apion)
hilft deubeli Wagner, 1912a: 19 (Apion) E: RO
hilfi hilfl Wagner, 1912a: 17 (Apion) E: BU TR
inexpertum Wagner, 1906b: 207(Apion) [RN] E: AU FR GE IT SZ
 APIONIDAE 159
eppelsheimi Desbrochers des Loges, 1897c: 17 (Apion) [HN]
laufferi laufferi Schilsky, 1906a: 20 {Apion) E: SP
laufferi paganettii Wagner, 1914a: 26 (Apion) E: SP
lemovicinum A. Hoffmann, 1929c: 312 (Apion) E: FR SP
liguricum A. Solan & F. Solari, 1905: 101 (Apion) E: IT
mendax Desbrochers des Loges, 1907: 87 (Apion) N: AG TU
oblongulum Desbrochers des Loges, 1875a: 31 (Apion) E: PT SP
putonii C. N. F. Brisout de Barneville, 1866: 386 (Apion) E: PT SP
reyi Desbrochers des Loges, 1897c: 19 (Apion) E: FR SP
sarotae Peyerimhoff, 1926c: 382 (Apion) N: AG
schatzmayri Wagner, 1923: 97 E: IT
squamulatum Gyllenhal, 1833: 275 (Apion) N: MO
flavicrus Desbrochers des Loges, 1901a: 80 (Apion)
sublineatum Schilsky, 1906a: 21 (Apion) "Armenien oder Turkestan"
subparallelum Desbrochers des Loges, 1888: cxciii (Apion) E: FR IT PT SP N: AG MO TU
subrectirostre Desbrochers des Loges, 1895a: 51 (Apion)
uliciperda Pandellé, 1867: 184 (Apion) E : F R P T S P N: AG
ulicis Forster, 1771: 31 (Curculio) E: CZ FR GB GE IR IT PL PT SP SZ N: AG MO AURi NARi
nigrirostris Fabricius, 1775: 132 (Curculio)
sarothamni Gradl, 1882: 331 (Apion)
ulicis Dufour, 1843: 77 (Apion) [HN]
wagnerianum Schatzmayr, 1925: 24 E: SP
winkleri Wagner, 1912a: 19 (Apion) E: GR
zangezuricum Iablokoff-Khnzorian, 1977: 303 (Apion) E: AR

genus Lepidapion Schilsky, 1906b: v type species Apion argentatum Gerstaecker, 1854
subgenus Hidryocneme Alonso-Zarazaga, 1990a: 76 type species Apion cretaceum Rosenhauer, 1856
bremondiA. Hoffmann, 1938b: 100 (Apion) N: MO
canariense Wagner, 1914a: 23 (Apion) N: CI (Tenerife)
cretaceum Rosenhauer, 1856: 240 (Apion) E: FR PT SP N: AG EG LB MO TU A: IS JO SI
alfierii Pic, 1915c: 33 (Apion)
cobosi Solari, 1952: 41 (Apion)
torretassoi Schatzmayr, 1933: 169 (Apion)
curvipilosum Wagner, 1908c: 102 (Apion) N: CI (Tenerife)
senex Wollaston, 1864: 306 (Apion) N: CI (Gomera, La Palma)
spartocytisi G. A. K. Marshall, 1928a: 115 (Apion) N: CI (Gran Canada)
subgenus Lepidapion Schilsky, 1906b: vtype species Apion argentatum Gerstaecker, 1854
acuminatum Schilsky, 1906a: 31 (Apion) E: SP
argentatum argentatum Gerstaecker, 1854a: 258 (Apion) E: FR IT PT SP N: AG MO TU A: TR
argentatum majolae Schatzmayr, 1933: 169 (Apion) N: LB
argentatum sardoum Schatzmayr, 1933: 169 (Apion) E: IT (Sardegna)
bletoni Hustache, 1939: 198 (Apion) N: MO
gallaecianum Desbrochers des Loges, 1894e: 29 (Apion) E: SP
nimbatum Normand, 1937: 234 (Apion) N: AG
plagiatum Desbrochers des Loges, 1891c: 324 (Apion) E: SP
pseudogallaecianum A. Hoffmann, 1929b: 132 (Apion) E: FR
squamidorsum Desbrochers des Loges, 1908d: 83 (Apion) E: SP N: AG MO MR
squamigerum Jacquelin du Val, 1854: 9 (Apion) E: FR IT (Sicilia) PT SP N: AG MO

tribe Ixapiini Alonso-Zarazaga, 1990

genus Ixapion Roudier & Tempère, 1973: 80 [RN] type species Apion variegatum Wencker, 1864
Ixias Sainte Claire-Deville, 1924: 122 [HN] type species Apion variegatum Wencker, 1864
variegatum Wencker, 1864: 188 (Apion) E: AU CZ FR GB GE HU IT LU PL SK SP SZ
bicolor Gredler, 1857: 153 (Apion) [HN]
160 APIONIDAE
genus Trichopterapion Wagner, 1930: 1387 type species Apion holosericeum Gyllenhal, 1833
holosericeum Gyllenhal, 1833: 268 (Apion) E: AB AR AU BH BU BY CR CZ FR GE GG HU IT LA MC MD
RO SB SK SL ST SZ UK A: TR
cuneatum Hochhuth, 1847: 464 (Apion)
hiemale Hampe, 1861: 67 (Apion)

tribe Kalcapiini Alonso-Zarazaga, 1990

genus Afrothymapion Wanat, 1990: 64 type species Apion tanganum Hartmann, 1897
tanganum Hartmann, 1897: 83 (Apion) A: SA YE AFR

genus Hypuranius Korotyaev, 1996: 877 type species Apion borchmanni Voss, 1958
borchmanni Voss, 1958: 131 (Apion) A: FUJ SD TAI ORR

genus Kalcapion Schilsky, 1906b: iv type species Apion pallipes Kirby, 1808
pallipes Kirby, 1808: 38 (Apion) E: AU BE BH BU CZ DE EN FI FR GB GE HU IT LA LU NL NT PL RO SK
SL SP (Mallorca) ST SV SZ UK A: IS
geniculatum Germar, 1817b: 115 (Apion)
pallidipes Bedel, 1886: 208 (Apion) [RN]
pallipes Gyllenhal, 1813: 34 (Apion) [HN]
semivittatitm fortunatum Roudier, 1963: 149 (Apion) N: CI
semivittatum sagittiferum Wollaston, 1854: 410 (Apion) N: MR
semivittatum semivittatum Gyllenhal, 1833: 271 (Apion) E: AL AU AZ BU CR FR GB GE GG GR HU IT LU
MA ME NL PT RO SB SP ST SZ UK N: AG CI LB MO TU A: CY IS JO LE SY TR
albopilosum P. H. Lucas, 1847: 408 (Apion)
germari Walton, 1844: 456 (Apion)
pallidactylum Gyllenhal, 1839: 394 (Apion)
syriacum Desbrochers des Loges, 1897c: 6 (Apion) [DA]
tonsuratum Betta, 1847: 31 (Apion)
separandum Aube, 1866: 163 (Apion) E: FR SP
terminasianae Alonso-Zarazaga, 1986: 198 (Apion) [RN] A: AF
rubidum Ter-Minasian, 1971b: 658 (Apion) [HN]
genus Melanapion Wagner, 1930: 1390 type species Apion minimum Herbst, 1797
subgenus Melanapion Wagner, 1930: 1390 type species Apion minimum Herbst, 1797
ferganicum Bajtenov, 1977d: 228 (Apion) A: UZ
mandli Schubert, 1959: 30 (Apion) A: ES FE NC SC
wintert Korotyaev & Egorov, 1996: 859
minimum Herbst, 1797: 115 ("125") (Apion) E: AB AR AU BU BY CT CZ DE EN FI FR GB GE GG HU IT
LA LT LU MD NL NR NT PL PT SB SK SP ST SV UK N: AG A: ES FE KZ MG UZ WS
denominandum Jacquelin du Val, 1855: 205 (Apion) [RN]
foraminosum Gyllenhal, 1833: 289 (Apion)
trifolii Gyllenhal, 1833: 285 (Apion)
velox Kirby, 1811: 349 (Apion)
naga Nakane, 1963a: 385 (Apion) A: FE HUB JA NC
subgenus Rhodapion Alonso-Zarazaga, 1990a: 56 type species Apion hauseri Wagner, 1906
hauseri Wagner, 1906b: 203 (Apion) A: AF KZ MG TD TM UZ
subrubripes Ter-Minasian, 1972: 240 (Apion)

genus Sergiola Korotyaev, 1996: 862 type species Apion siljverbergi Bajtenov, 1983
subgenus Golovninia Korotyaev & Egorov, 1996: 873 type species Apion hilleri Schilsky, 1902
hilleri Schilsky, 1902: 67 (Apion) A: JA
ussuricola Korotyaev & Egorov, 1996: 876 A: FE
subgenus Sergiola Korotyaev, 1996: 862 type species Apion siljverbergi Bajtenov, 1983
gressitti Korotyaev, 1996: 867 A: GUA
griseopubescens Roelofs, 1874: 129 (Apion) A: JA SC
vasta Schilsky, 1902: 64 (Apion)
 APIONIDAE 161
praecaria Faust, 1889b: 225 {Apion) A: FE GUA JA NC SC
rasnitsyni Korotyaev & Egorov, 1996: 868 A: FE
silfverbergi Bajtenov, 1983: 59 {Apion) A: FE

genus Squamapion Bokor, 1923: 98 type species Apion vicinum Kirby, 1808
Thymapion Sainte Claire-Deville, 1924: 135 type species ApionflavimanumGyllenhal, 1833
Squamapion Wagner, 1929a: 253 [HN] type species ApionflavimanumGyllenhal, 1833
Teuchocnemapion Voss, 1966: 234 type species Apion jordanianum Voss, 1965
atomarium Kirby, 1808: 59 {Apion) E: AR AU CR CZ DE EN FI FR GB GE GG GR HU IT LA LU NL NT PL
PT RO SB SK SP ST SV SZ UK N: AG TU A: CY MG TR
acium Gyllenhal, 1833: 257 {Apion)
atomarium Gyllenhal, 1813: 60 {Apion) [HN]
pusillum Germar, 1817b: 209 {Apion)
bifarium Balfour-Browne, 1944b: 149 {Apion) A: CY
cineraceum Wencker, 1864: 146 (Apion) E: AU BH CR CZ EN FI FR GB GE GG HU IT LT LU NT PL SK SP
ST SV SZ UK
consors consors Desbrochers des Loges, 1875a: 31 (Apion) E: FR (Corse) IT(Sardegne, Sicilia)
consors solariorum Wagner, 1908c: 105 (Apion) N: AG MO
delagrangei Desbrochers des Loges, 1895d: 83 (Apion) E: GR A: IS TR
tavriense Bajtenov & Lodos, 1980: 145 (Apion)
elongatum Germar, 1817b: 214 (Apion) E: AB AU CZ FR GE GG GR HU IT LA LT LU PL RO SK SP ST SZ
UK "Caucasus" N: AG A: KZ WS
arcirostre Desbrochers des Loges, 1895g: 139 (Apion) [DA]
cylindricum Desbrochers des Loges, 1896a: 6 (Apion)
ineanum Boheman, 1839: 414 (Apion)
longitubus Desbrochers des Loges, 1889a: xxxiv (Apion)
milium Gyllenhal, 1833: 277 (Apion)
torquatum Wencker, 1864: 183 (Apion) [HN]
fissile Faust, 1888b: 175 (Apion) A: FE
flavimanum Gyllenhal, 1833: 276 (Apion) E: AU BH BU CT CZ DE EN FR GB GE GG GR HU IT LA LU NL
PL RO SB SK SP ST SV SZ UK A: TR WS
annulipes Wencker, 1864: 145 (Apion) [DA]
latithorax Desbrochers des Loges, 1895g: 140 (Apion) [DA]
neresheimeri Wagner, 1929a: 252 (Apion)
picicorne Stephens, 1839: 257 (Apion)
torquatum Wencker, 1864: 148 (Apion) [DA]
uhagonis Desbrochers des Loges, 1870a: 195 (Apion)
fukienense Voss, 1958: 131 (Apion) A: FUJ GUX
gracilitubus Desbrochers des Loges, 1896b: 71 (Apion) E: AR A: TR
griseopilosum Wagner, 1908a: 27 (Apion) A: YE AFR
kandarense Hustache, 1946: 81 (Apion) N: MO
latesquamatum Wanat, 1997: 293 A: IN TR
leucophaeatum Wencker, 1864: 183 (Apion) E: CR FR IT SL SP N: AG MO TU
angustipenne Desbrochers des Loges, 1896a: 12 (Apion)
emendatum Wagner, 1930: 1390 (Apion) [RN]
errabundum Schilsky, 1906c: 699 (Apion) [HN]
obtusipenne Desbrochers des Loges, 1896a: 11 (Apion)
obtusiusculum Desbrochers des Loges, 1896a: 9 (Apion)
tenuirostre Desbrochers des Loges, 1895h: 149 (Apion)
lukjanovitshi Korotyaev, 1988b: 136 (Thymapion) A: KZ WS
medium Desbrochers des Loges, 1891e: lviii [= 1981b: 83] (Apion) N: AG MO TU
megatoma Korotyaev, 1996: 880 A: FE FUJ SC
sergii Korotyaev, 1996: 881
minutissimum Rosenhauer, 1856: 242 (Apion) E: FR GR IT SP N: AG MO TU
tunicense Desbrochers des Loges, 1895a: 56 (Apion)
mroczkowskii Wanat, 1997: 287 E: AU PL UK
oblivium Schilsky, 1902: 54 (Apion) E: AU BH CZ DE FR GE HU IT LA PL SK SV SZ UK A: TR
162 APIONIDAE
oculatum Wanat, 1997: 289 E: AR A: TR
origani Planet, 1918: 155 {Apion) E: FR LU NL SP SV
ottonis Wagner, 1914a: 23 (Apion) E: BU A: TR
phocopus Eppelsheim, 1888: 380 (Apion) E: GR A: IS JO LE TR
jordanianum Voss, 1965: 337 {Apion)
samarense Faust, 1891c: 411 (Apion) E: BU CT FR HU IT PL RO ST UK "Caucasus" A: KZ MG TM WS
dilatipes Desbrochers des Loges, 1896a: 10 (Apion)
doderoi Schatzmayr, 1922: 50 (Apion)
humerosum Desbrochers des Loges, 1896a: 10 (Apion) [HN]
pannonicum Györffy, I955: 267 (Apion)
pseudelongatum Wagner, 1909b: 766 (Apion) [RN]
serpyllicola Wencker, 1864: 150 (Apion) E: AU CZ FI FR GE HU LU PL SB SK SV SZ
hoffinanni Wagner, 1930: 1391 {Apion) [HN]
parvulum Mulsant & Rey, 1858: 263 {Apion) [HN]
serpyllicola Gozis, 1881b: 181 (Apion) [RN] [HN]
smreczynskii A. Hoffmann, 1934: 235 (Apion)
solarii Wagner, 1908b: 69 (Apion) A: SA AFR
subconsors Bajtenov & Lodos, 1978a: 93 (Apion) A: TR
subrecticolle Voss, 1961a: 419 (Apion) A: SA AFR
terraesanctae Friedman & Freidberg, 2007: 90 A: IS
vemtstulum Wencker, 1864: 148 (Apion) E: CR GR IT RO ST
zuberi Desbrochers des Loges, 1870a: 199 (Apion)
vicinum Kirby, 1808: 25 (Apion) E: AU BU CT CZ DE EN FI FR GB GE HU IT LU NL NR NT PL RO SB SK
SP ST SV UK "Caucasus" N: AG A: FE SY WS
incrassatum Germar, 1817b: 140 (Apion)
sulcithorax Desbrochers des Loges, 1901b: 82 (Apion)

genus Taeniapion Schilsky, 1906b: iv type species Curculio urticarius Herbst, 1784
delicatulum Wollaston, 1857: 120 (Apion) E: MR N: CI
diverserostratum Uyttenboogaart, 1935a: 13 (Apion)
fasciatum Desbrochers des Loges, 1891c: 325 (Apion)
reitterianum Wagner, 1912c: 84 (Apion) N: EG A: IS
nifescens Gyllenhal, 1833: 273 (Apion) E: AU BH CR FR GR HU IT MC PT SP SZ N: AG EG LB MO TU
A: CY IS JO LE SY TR
notatum Wagner, 1912c: 84 (Apion)
pallidulum Gyllenhal, 1839: 400 {Apion)
rufulum Wencker, 1864: 162 (Apion) E: CR CZ FR GE GR HU IT MC NL PL PT RO SK SP ST N: AG LB
MO TU A: LE SY TR
rolphi Wagner, 1912c: 85 (Apion)
semirufiim Rey, 1888:4 (Apion)
zoufali Wagner, 1912c: 86 (Apion)
tyrrhenicum Alonso-Zarazaga, nom. nov. [see New Acts] E: FR (Corse) IT (Sardegne)
distinctirostre Desbrochers des Loges, 1888: cxciii [= 1894a: 40 ] (Apion) [HN]
urticarium atlanticum Uyttenboogaart, 1935a: 14 (Apion) E: MR. N: CI
urticarium urticarium Herbst, 1784a: 74 (Curculio) E: AB AR AU BH BU CR CT CZ DE EN FI FR GB GE
GG GR HU IT LA LU MD NL NR NT PL RO SB SK SP ST SV SZ UK N: AG A: FE JA TR WS
concinnum Marsham, 1802: 248 (Curculio)
fasciatum Olivier, 1789: 283 {Attelabus) [HN]
maximi Krausse, 1916: 109 (Apion)
scalptor Herbst, 1795: 105 (Curculio)
turcicum Desbrochers des Loges, 1897c: 7 (Apion)
vernale Paykull, 1792: 138 (Curculio)
wolffi Krausse, 1916: \09 (Apion)
tribe Malvapiini Alonso-Zarazaga, 1990
genus Malvapion A. Hoffmann, 1958: 1471 type species Curculio malvae Fabricius, 1775
Malvapion Voss, 1959a: 74 [HN] type species Curculio malvae Fabricius, 1775
 APIONIDAE 163
afghanum Voss, 1963: 403 {Apion) A: AF
malvae Fabncius, 1775: 132 {Curculio) E: AB AR AU BE BU CR CZ FR GB GE GG GR HU IR IT LU MR
NL PL PT RO SB SK SP ST SV SZ UK N: AG EG LB MO MR TU A: CY IQ IN IS JO SI SY TD TR
flavescens A. Villa & G. B. Villa, 1835: 49 {Apion)
herbarum Aube, 1850: 338 {Apion)

genus Pseudapion Schilsky, 1906b: vi type species Apion fiilvirostre Gyllenhal, 1833
ebrium Giusto, 2002: 57 E: IT
fiilvirostre Gyllenhal, 1833: 274 {Apion) E: AB AR AU BH BU CR FR GR HU IT PT RO SB SK SP ST SZ
UK N: AG A: ES KI SY TR XIZ
atritarse Gyllenhal, 1833: 277 {Apion)
fortipubens Reitter, 190le: 226 {Apion)
moschatae Hofmann, 1938a: 381 {Apion) E: AU FR GE IT LU NL PT SP SZ
rufirostre Fabncius, 1775: 132 {Curculio) E: AB AU CR CZ DE FR GB GE GG HU IR IT LU MC NL PL PT
RO SB SK SP ST SV N: AG MO TU A: SY
destefanii Vitale, 1890: 148 {Apion)
malvarum Kirby, 1808: 33 {Apion)

genus Rhopalapion Schilsky, 1906b: v type species Apion longirostre Olivier, 1807
longirostre Olivier, 1807: 35 {Apion) E: AB AL AR AU BU CR CZ FR GE GG GR HU IT LU MC ME NL PL
RO SB SK SP ST SZ TR UK N: AG A: AF CY IN IS JO LE SY TD TM TR UZ NARi

tribe Metapiini Alonso-Zarazaga, 1990

genus Metapion Schilsky, 1906b: v type species Apion candidum Wencker, 1864
Pseudometapion Wagner, 1930: 1387 type species Apion gaudiale Faust, 1885
Paronychapion Voss, 1961c: 181 type species Apion causticum Faust, 1885
candidum candidum Wencker, 1864: 142 {Apion) E: FR IT PT SP SZ
candidum rutae Ehret, 1997d: 32 E: FR N: EG TU
causticum Faust, 1885a: 187 {Apion) A: AF KI KZ TM TD UZ
cylindronotum Ter-Minasian, 1972: 239 {Apion) A: MG
densesquamatum Pic, 1908c: 58 {Apion) N: EG
emmrichi Bajtenov, 1981a: 27 {Apion) A: KZ
ermischi Voss, 1969a: 53 {Apiotherium) E: AU BU MC UK A: TR
gaudiale Faust, 1885a: 189 {Apion) E: "Caucasus" A: AF IN KI KZ TD TM UZ
korbi Schilsky, 1901b: 99 {Apion)
otiosum Faust, 1891c: 414 (Apion)
subfarinosum Desbrochers des Loges, 1897c: 23 (Apion)
gelidum Faust, 1885a: 188 (Apion) E: AR A: AF ES IN KI KZ SY TD TM UZ
rimmae Bajtenov, 1981d: 124 (Apion)
hartmanni Desbrochers des Loges, 1897c: 17 (Apion) E: SP
kysilkumicum Bajtenov, 1979a: 73 (Apion) A: UZ
lepidopterum Wagner, 1912c: 86 (Apion) A: AF IN TM
merale Faust, 1885a: 188 (Apion) A: AF IN KI KZ TD TM UZ
schiwanum Voss, 1959a: 76 (Apion)
nasreddinovi Korotyaev, 1988a: 142 A: TD UZ
korotyaevi Schoen, 1988: 36 (Apion)
normandi Desbrochers des Loges, 1898b: 36 {Apion) N: LB TU A: IS TR
obtectum Schilsky, 1902: 74 (Apion) A: KI KZ
gladiator Bajtenov, 1977a: 15 (Apion)
oculare Gyllenhal, 1833: 257 (Apion) E: AL AR BU GG MC RO SB TR UK
breiti Wagner, 1910b: 944 (Apion)
kolenatii Schoenherr, 1858: 151 (Apion)
sojunovii Bajtenov & Sojunov, 1990: 72 (Apion) A: TM
squamosum Faust, 1884b: 455 {Apion) E: AB AR GG ST A: KZ TD TM UZ
squarrosum Normand, 1937: 233 (Apion) N: TU
subcandidum Schilsky, 1902: 69 (Apion) E: SP
164 APIONIDAE
tribe Oxystomatini Alonso-Zarazaga, 1990
subtribe Catapiina Alonso-Zarazaga, 1990
genus Catapion Schilsky, 1906b: vi type species Apion seniculus Kirby, 1808
Pseudocatapion Wagner, 1930: 1392 type species Apion burdigalense Wencker, 1859 (= Apion semicyaneum Mulsant &
Rey, 1858)
Aceratapion Voss, 1969a: 55 type species Ceratapion kabulense Voss, 1969 (= Apion gemulum Faust, 1885)
acanthyllidis Normand, 1921: 77 (Apion) N: AG MO TU
amudariense Bajtenov, 1981c: 85 (Apion) A: UZ
austrinum Wollaston, 1864: 312 (Apion) N: CI (Gomera)
austroturcum Bajtenov & Lodos, 1977b: 315 (Apion) A: TR
calabricum A. Solan & F. Solari, 1922b: 28 (Apion) E: IT
caucasicum Hochhuth, 1847: 463 (Apion) E: "Caucasus" A: IN UZ
acuticlava Desbrochers des Loges, 1902c: 159 (Apion) [DA]
corsicum Desbrochers des Loges, 1888: cxciii (Apion) E: BU FR (Corse) GR IT "Yugoslavia" A: TR
curtisii Stephens, 1831: 169 (Apion) E: BH BU FR GB GE GG GR IT NL PT SP N: AG A: IS LE SY TR
curtulum Desbrochers des Loges, 1870a: 203 (Apion)
frontale Desbrochers des Loges, 1895g: 142 (Apion)
frontale Desbrochers des Loges, 1901b: 83 (Apion) [HN]
cyanellum Desbrochers des Loges, 1896e: 114 (Apion) E: AR A: TR
gemulum Faust, 1885a: 186 (Apion) A: AF IN KZ TM UZ
doriae Desbrochers des Loges, 1893b: 11 (Apion)
kabulense Voss, 1969a: 55 (Ceratapion)
mus Desbrochers des Loges, 1897c: 25 (Apion)
soricinum Desbrochers des Loges, 1892a: 108 (Apion)
halperini Friedman & Freidberg, 2007: 120 A: IS
jaffense Desbrochers des Loges, 1896c: 76 (Apion) E: AB BU CZ HU IT PL SB SK A: IS
ononiphagum Schatzmayr, 1920: 42 (Apion)
koestlini Dieckmann, 1989: 249 (Apion) E: FR GE IT LU PL
lodosiA. Hoffmann, 1954b: 284 (Apion) A: TR
meieri Desbrochers des Loges, 1901b: 82 (Apion) E: AU BE CR CT CZ DE FI FR GE GG IT LA LU MD NR
NT PL PT SK SV
natricis Planet, 1918: 154 (Apion) E: FR IT SP N: MO
pubescens Kirby, 1811: 350 (Apion) E: AU CR CZ DE EN FR GB GE GG GR HU IT LU NL PL SB SK SP ST
SV UK N: AG CI MO TU A: CY IS JO SY
ceuthorhynchoides Wollaston, 1864: 314 (Apion)
civicum Germar, 1817b: 234 (Apion)
Salicis Gyllenhal, 1833: 286 (Apion)
schneiden Tournier, 1878: 306 (Apion) E: GG
koenigi Desbrochers des Loges, 1897c: 26 (Apion)
semicyanescens Desbrochers des Loges, 1904a: 54 (Apion) A: IN "Tianshan"
semicyaneum Mulsant & Rey, 1858: 265 (Apion) E: AU BU CR FR GR IT SP ST "Caucasus" N: AG MO TU
A:ISJOKZSYTR
burdigalense Wencker, 1859b: ccxxxvii (Apion)
fesense Hustache, 1946: 81 (Apion)
talpa Desbrochers des Loges, 1870a: 185 (Apion)
seniculus Kirby, 1808: 61 (Apion) E: AL AR AU BE BU CR CT CZ DE EN FI FR GB GE GG GR HU IR IT
LA LT LU MD NL NRNT PL PT RO SB SK SP ST SV SZ UK N: AG MO TU A: ES HEI IS JO KZ LE
SY TM TR WS
angustiforme J. R. Sahlberg, 1921: 34 (Apion)
dudichi Györfry, 1924: 27 (Apion) [DA]
murinum Everts, 1882: 252 (Apion) [HN] [DA]
palpebratum Gyllenhal, 1833: 258 (Apion)
plebejum Germar, 1817b: 215 (Apion)
tenuitubus Desbrochers des Loges, 1908d: 83 (Apion)
tenuius Gyllenhal, 1813: 57 (Apion)
seriatosetosulum Wencker, 1864: 179 (Apion) E: AU BU CR FR HU IT PT SL SP N: AG MO TU
neapolitanum Desbrochers des Loges, 1870a: 203 (Apion)
 APIONIDAE 165
setosum Wencker, 1864: 180 (Apion) A: TR
subpubescens Wencker, 1864: 176 (Apion) N: AG TU
mendosum Normand, 1937:236 (Apion) [DA]
turkestanicum Desbrochers des Loges, 1892a: 107 (Apion) A: AF KZ
vulnerariae Dieckmann & Wanat, 1991: 353 (Apion) E: AU

genus Necatapion Friedman & Freidberg, 2007: 155 type species Apion bruleriei Desbrochers des Loges,
1875
bruleriei Desbrochers des Loges, 1875a: 30 (Apion) A: CY IS JO LE TR
schuberhAlonso-Zarazaga, 1986: 198 (Apion) [RN]
tauricum Schubert, 1963: 81 (Apion) [HN]

subtribe Oxystomatina Alonso-Zarazaga, 1990

genus Cyanapion Bokor, 1923:100 type species Apion alcyoneum Germar, 1817
subgenus Bothryorrhynchapion Bokor, 1923: 100 type species Apion gyllenhalii Kirby, 1808
Coelorhinapion Ehret, 1990: 253 type species Apion platalea Germar, 1817
afer Gyllenhal, 1833: 291 (Apion) E: AL AU BH BU CR CZ EN FI FR GB GE HU IT LA LU MC NR NT PL
RO SB SK SP SV SZ UK "Caucasus" A: ES WS
jurvum C. R. Sahlberg, 1834: 17 (Apion)
puncticolle Stephens, 1839: 259 (Apion)
validirostre Gyllenhal, 1833: 301 (Apion)
gnarum Faust, 1890b: 106 (Apion) E: NT PL A: ES WS
gyllenhalii Kirby, 1808: 63 (Apion) E: AU CT CZ DE EN FI FR GB GE GG HU IT LA LT LU NL NR NT PL
RO SB SK SP ST SV UK A: ES FE MG WS
punctigerum Thunberg, 1815: 118 (Apion) [HN]
unicotor Kirby, 1808: 58 (Apion)
offensum Faust, 1891c: 413 (Apion) E: UK
platalea Germar, 1817b: 143 (Apion) E: AU BH CR CZ FR GE GG HU IT LU NL PL RO SB SK SP ST SZ
UK A: WS
protractum Sharp, 1891: 294 (Apion) A: JA (Honshu) SC
subgenus Cyanapion Bokor, 1923: 100 type species Apion alcyoneum Germar, 1817
Cyanapion Ehret, 1990: 250 [HN] type species Apion spencii Kirby, 1808
alcyoneum Germar, 1817b: 205 (Apion) E: AU FR GE GR HU IT PL PT SK SP A: KZ WS
hispanicum Wencker, 1864: 232 (Apion)
hoffmanni Miller, 1857: 22 (Apion)
burakowskii Wanat, 1995b: 482 A: TR
columbinum Germar, 1817b: 185 (Apion) E: AU BU CR CZ DE FI FR GE HU IT LU NL RO SB SK SP ST SV
SZ UK N: AG A: KI KZ SY TR WS
pedemontanum Desbrochers des Loges, 1870a: 197 (Apion)
coniceps Desbrochers des Loges, 1897c: 27 (Apion) E: "Caucasus"
eniseiense Bajtenov, 1977a: IS (Apion) A: ES
pseudarrogans Reitter, 1901e: 227 (Apion) A: KZ UZ
rudicolle Hochhuth, 1851: 12 (Apion) [DA] E: UK "Caucasus"
spencii Kirby, 1808: 57 (Apion) E: AU CR CT CZ DE EN FI FR GÖ GE HU IT KZ LA LT LU NL NR NT PL
SB SK ST SV SZ UK N: AG A: ES KZ WS
foveolatum Kirby, 1808:48 (Apion)
intrusum Gyllenhal, 1827: 550 (Apion)
nigrum Gerhardt, 1898: 335 (Apion) [HN]
genus Eutrichapion Reitter, 1916a: 243 type species Attelabus viciae Paykull, 1800
subgenus Cnemapion Bokor, 1923: 100 type species Apion vorax Herbst, 1797
gribodoi Desbrochers des Loges, 1896b: 69 (Apion) E: AL BH CR CZ HU IT RO SB SK TR
ochroceras Desbrochers des Loges, 1907: 87 (Apion)
vorax Herbst, 1797: 119(" 129") (Apion) E: AU BU CR CT CZ DE EN FI FR GB GE GR HU IR IT LT LU NL
PL PT RO SB SK SP ST SV SZ UK N: AG CI MO TU A: CY IS LE SY "Siberia"
desbrochersi Kirsch, 1880: 13 (Apion)
166 APIONIDAE
fiiscicome Marsham, 1802: 244 (Curculio)
longipes Wollaston, 1864: 315 (Apion)
pallicorne Gyllenhal, 1833: 302 (Apion)
pilosellum Gravenhorst, 1807: 198 (Apion)
simplicipes Desbrochers des Loges, 1891e: Ivii [=189lb: 83] (Apion)
villosulum Marsham, 1802: 250 (Curculio) [HN]
subgenus Eutrichapion Reitter, 1916a: 243 type species Attelabus viciae Paykull, 1800
Pseudotrichapion Bokor, 1923: 101 type species Attelabus viciae Paykull, 1800
ervi Kirby, 1808: 23 (Apion) E: AB AR AU BU CT CZ DE EN FI FR GB GE GG GR HU IR IT LA LU NL
NR NT PL SB SK SP ST SV SZ UK N: AG A: ES FE JA MG SY WS
amurense Bajtenov, 1983a: 19 (Apion)
kihige Nakane, 1963a: 358 (Apion)
lathyri Kirby, 1808: 24 (Apion)
nigroclavatum Gerhardt, 1911: 338 (Apion)
opthalmicum Desbrochers des Loges, 1896b: 71 (Apion)
hingstoni Balfour-Browne, 1944a: 9 (Apion) A: XIZ
meditabundum Faust, 1890b: 104 (Apion) A: ES FE MG WS
subaequalicum Voss, 1967: 260 (Apion)
mystriophorum Alonso-Zarazaga, 1994: 72 E: PT SP
rectirostre Schilsky, 1906a: 58 (Apion) A: ES FE HEI WS
tomiense Bajtenov, 1979b: 86 (Apion) A: WS
tomiense Bajtenov, 1982a: 48 (Apion) [HN]
viciae Paykull, 1800: 181 (Attelabus) E: AB AR AU BE BU CR CT CZ DE EN FI FR GB GE GG GR HU IR
I T L A L T L U N L N R N T P L R O S B S K S P S T S V S Z U K N: AG A: CY ES FE JA KZ MG NC SC SY
TR WS NAR
alaskanum Fall, 1926: 205 (Apion)
griesbachii Stephens, 1831: 179 (Apion)
kaszabi Voss, 1967: 256 (Apiotherium)
rufinasus Desbrochers des Loges, 1891c: 326 (Apion)
sapporense Kôno, 1927c: 241 (Apion)
voisini Ehret, 1997c: 130 (Oxystoma)
subgenus Phalacrolobus Alonso-Zarazaga, 1990a: 146 type species Apion melancholicum Wencker, 1864
hydropicum Wencker, 1864: 174 (Apion) E: FR IT SZ
melancholicum Wencker, 1864: 217 (Apion) E: AU CT CZ DE FI FR GE HU IT LA LU PL RO SB SK ST SV
UK A: TR WS
hadrops C. G. Thomson, 1865: 72 (Apion)
italicum Desbrochers des Loges, 1904a: 57 (Apion)
provinciale Desbrochers des Loges, 1889a: xxxiv (Apion)
subgenus Psilocalymma Alonso-Zarazaga, 1990a: 146 type species Curculiopunctiger Paykull, 1792
Pseudotrichapion Ehret, 1990: 257 [HN] type species Curculio punctiger Paykull, 1792
arrogans Wencker, 1858b: cvi (Apion) A: IS JO S Y TR
cyaneomicans Desbrochers des Loges, 1875a: 28 (Apion)
crassiforme Bajtenov, 1977a: 17 (Apion) A: ES
facetum Gyllenhal, 1839: 431 (Apion) E: AR AU BH BU CR CT CZ DE EN FI FR GE GR HU IT LU NR NT
PL SK ST SV UK N: AG A: ES KZ MG TR WS
perspicax Wencker, 1864:221 (Apion)
sundevalli Boheman, 1839: 435 (Apion)
punctiger Paykull, 1792: 141 (Curculio) E: AR AU BU CR CT CZ DE EN FI FR GB GE GG GR HU IT KZ
LA LT LU NL NR PL PT RO SB SK ST SV SZ UK N: AG A: SY TR WS
paykulli Gozis, 1875: 75 (Apion) [unnecessary RN]
rhomboidale Desbrochers des Loges, 1870a: 182 (Apion) E: AU BH HU IT SB SL A: ES WS
stierlini Desbrochers des Loges, 1870a: 181 (Apion)
genus Hemitrichapion Voss, 1959a: 80 type species Apion plicatum Faust, 1887
subgenus Dimesomyops Alonso-Zarazaga, 1990a: 131 type species Apionpavidum Germai, 1817
andalusiacum Desbrochers des Loges, 1889a: xxxiv (Apion) E: FR SP N: AG MO
ciliare Desbrochers des Loges, 1889: ccxvi (Apion)
 APIONIDAE 167
pavidum Germar, 1817b: 203 (Apion) E: AR AU BU CR CZ EN FR GE GG GR HU IT LA PL RO SB SK SP
ST SZ UK N: AG TU A: IS SY TR
orbitale Boheman, 1839:434 (Apion)
plumbeum Gyllenhal, 1833: 301 (Apion)
schilskyi Gerhardt, 1911: 338 (Apion) [HN]
rapulum Wencker, 1864: 175 (Apion) E: AU FR GG IT RO SP SZ N: AG
subgenus Hemitrichapion Voss, 1959a: 80 type species Apion plicatum Faust, 1887
badakschanicum Voss, 1959a: 80 (Apion) A: AJF
perspicillum Faust, 1887c: 186 (Apion) A: IN KI KZ TD TM UZ
plicatum Faust, 1887c: 185 (Apion) A: KI KZ UZ XIN
subgenus Lotapion Alonso-Zarazaga, 1990a: 130 type species Apion wagneri Flach, 1906
sandovali Ferragu, 1957: 145 (Apion) E: SP
wagneri Flach, 1906: 121 (Apion) E: PT SP N: MO
subgenus Microtinocyba Legalov, 2001c: 255 type species Apion tschernovi Ter-Minasian, 1973
reitteri Desbrochers des Loges, 1892a: 108 (Apion) A: KI KZ UZ
tschernovi nanschanicum Legalov, 2001c: 257 A: GAN
tschernovi terektinense Legalov, 2001c: 257 A: WS
tschernovi tschernovi Ter-Minasian, 1973: 615 (Apion) A: ES QIN
tschernovi tuvense Legalov, 2001c: 257 A: ES
subgenus Tinocyba Alonso-Zarazaga, 1990a: 131 type species Apion waltoni Stephens, 1839
alexsandri Legalov, 2001c: 248 A: KZ WS
amguemae Korotyaev, 1991: 878 (Eutrichapion) A: FE
armeniacum Desbrochers des Loges, 1896c: 85 (Apion) E: AB AR GG A: TR
araxicola Wagner, 1912c: 90 (Apion)
derelictum Desbrochers des Loges, 1907: 85 (Apion) E: AR
filicorne Wencker, 1864: 235 (Apion) E: CR FR GR IT SP ST TR N: AG MO TU A: IS LE
decorum Wencker, 1864: 238 (Apion)
insidiosum Desbrochers des Loges, 1875a: 28 (Apion) A: IN KZ LE SY TD UZ "Caucasus"
persicum Desbrochers des Loges, 1893b: 11 (Apion)
juniperi Boheman, 1839: 433 (Apion) E: AU FR GE GG GR HU IT LS SK SP SZ
marqueü Wencker, 1859b: ccxxxvii (Apion)
montanum Miller, 1862: 348 (Apion)
kastecum Bajtenov, 1977c: 31 (Apion) A: KZ
korotyaevi Legalov, 2001c: 253 A: WS
lanigerum Gemminger, 1871a: 123 (Apion) [RN] E: AR AU FR GE GG GR IT LU SP SZ
lanuginosum Wencker, 1864: 236 (Apion) [HN]
lenense Schilsky, 1906a: 67 (Apion) A: ES
lethierryi Desbrochers des Loges, 1870a: 188 (Apion) E: BU FR HU IT A: AG
gavoyi Desbrochers des Loges, 1904a: 53 (Apion)
reflexum Gyllenhal, 1833: 290 (Apion) E: AB AR AU BU CZ FR GB GE GG GR HU IT LU NL PL SB SK SP
ST SZ UK N: AG A: ES IN MG TR WS
differens Desbrochers des Loges, 1875a: 28 (Apion) [DA]
foveatum Desbrochers des Loges, 1904a: 55 (Apion)
livescerum Gyllenhal, 1833: 298 (Apion)
translaticium Gyllenhal, 1839: 427 (Apion)
romani Legalov, 2007a: 400 A: WS
strictum Desbrochers des Loges, 1896b: 72 (Apion) E: "Caucasus"
suppantschitschi Legalov, 2002f: 285 A: WS
tschegitunense Legalov, 2001c: 254 A: FE
waltoni Stephens, 1839: 259 (Apion) E: AU CR FR GB GE HU IT LU SK SP SZ

genus Hemitrichapion, species incertae sedis [formerly Metatrichapion s.l.]

petryszaki Bajtenov, 1979d: 649 (Apion) E: AB
tigrense Bajtenov & Lodos, 1977a: 7 (Apion) A: TR
zagrossicum Bajtenov & Fremuth, 1981: 250 (Apion) A: IN
168 APIONIDAE
genus Holotrichapion Györffy, 1956: 45 type species Apion ononis Kirby, 1808
subgenus Apiops Alonso-Zarazaga, 1990a: 127 type species Attelabus pist F abricius, 1801
kcimdeschense Voss, 1959a: 79 (Apion) A: AF
pisi Fabricius, 1801: 425 (Attelabus) E: AU BU CR CZ DE EN FI FR GB GE GG GR HU IR IT LA LT LU
MC NL PL PT RO SB SK SP ST SV TR UK N: AG MO TU A: TR SY AURi
aeratum Stephens, 1831: 185 (Apion)
amplipenne Gyllenhal, 1833: 308 (Apion)
costipenne Fauvel, 1867: 193 (Apion)
cyanipenne Gyllenhal, 1833: 307 (Apion)
gravidum Olivier, 1807: 31 (Apion)
indistinctum Motschulsky, 1849: 144 (Apion)
pasticum Germar, 1817b: 187 (Apion)
punctifrons Kirby, 1808: 50 (Apion)
sulciferum Hubenthai, 1912: 74 (Apion)
pul/um Gyllenhal, 1833: 299 (Apion) E: AB AR AU BU CZ GE GG GR HU LA LT PL RO SB SK ST UK A:
IN IS SY TR
aestimatum Faust, 1891c: 410 (Apion)
subgenus Holotrichapion Györffy, 1956: 45 type species Apion ononis Kirby, 1808
Holotrichapion Ehret, 1990: 253 [HN] type species Apion ononis Kirby, 1808
ononis Kirby, 1808: 25 (Apion) E: AU CZ DE FR GB GE GG GR HU IT LA LT LU NL PL RO SB SK SP SV
UK A: CY IS SY
einerascens Germar, 1817b: 138 (Apion)
glaucinum Gyllenhal, 1833: 255 (Apion)
mecops Boheman, 1839: 413 (Apion)
ononidis Bedel, 1886: 215 (Apion) [RN, HN]
perplexum Gyllenhal, 1833: 293 (Apion)
umbrinum Wollaston, 1864: 314 (Apion) E: IT SP N: AG CI LB MO TU A: IN
barbirostre A. Hoffmann, 1953a: 143 (Apion)
saturnium Normand, 1937: 238 (Apion)
semisericeum Wagner, 1920: 199 (Apion) A: FE JA SC
subgenus Legaricapion Ehret, 1990: 257 type species Apion aethiops Herbst, 1797
Schoenius Alonso-Zarazaga, 1990a: 127 type species Apion aethiops Herbst, 1797
aethiops Herbst, 1797: 120("130") (Apion) E: AB AU CZ DE EN FI FR GB GE GG GR HU IR IT LA LU NL
N R N T P L P T S K S P S T S V U K N: AG MO TU A:SYWS
lucidithorax Desbrochers des Loges, 1896e: 117 (Apion)
subcaeruleum Stephens, 1831: 183 (Apion)
subsulcatum Marsham, 1802: 249 (Curculio)
stenoeephalum Penis, 1858: 136 (Apion)
antennale Desbrochers des Loges, 1896e: 118 (Apion) E: FR (Corse) GR IT N: TU A: IS
gracilicolle Gyllenhal, 1839: 428 (Apion) E: AU CR CZ FR GE GG GR HU IT PT SP ST N- AG MO TU A:
ISSY
leptocephalum Aube, 1850: 339 (Apion)
subtrapezicolle Desbrochers des Loges, 1897c: 27 (Apion)
subgenus Nesiapion Alonso-Zarazaga, 1990a: 127 type species Apion wollastoni Chevrolet 1852
wollastoni Chevrolat, 1852: 579 (Apion) N: CI MR
rotundipenne Wollaston, 1854: 415 (Apion)

genus Holotrichapion, species incertae sedis

splendidulum Desbrochers des Loges, 1875a: 29 (Apion) A: IS JO SY TR
damascenum Desbrochers des Loges, 1875a: 29 (Apion)

genus Loborhynchapion Gyôrffy, 1956: 52 type species Apion amethystinum Miller, 1857
Loborhynchapion Smreczynski, 1965: 52 [HN] type species Apion amethystinum Miller 1857
Loborhynchapion Osella, 1973a: 206 [HN] type species Apion amethystinum Miller 1857
amethystinum Miller, 1857: 23 (Apion) E: AU CZ GE HU SK ST UK A: ES FE IN MG SY TR UZ WS "Tian-
Shan"
boudali Normand, 1937: 237 (Apion) [DA] N: TU
 APIONIDAE 169
brundini Wagner, 1943: 160 (Apion) E: EINRNT SV
lobirostre Reitter, 1901e: 227 (Apion) A: AF KI KZ TD TM UZ WS
obtusum Desbrochers des Loges, 1866: 47 (Apion) E: FR IT SZ

genus Mesotrichapion Györfly, 1956: 54 type species Apion punctirostre Gyllenhal, 1839
Mesotrichapion Ehret, 1990: 256 [HN] type species Apion punctirostre Gyllenhal, 1839
conocephalum Desbrochers des Loges, 1875a: 28 (Apion) E: AB AR ST A: SY
araxellum Reitter, 1916a: 255 (Apion)
dauricum Faust, 1888b: 176 (Apion) A: ES WS
dudkorum Legalov, 1997b: 469 A: WS
punctirostre Gyllenhal, 1839: 425 (Apion) E: AR AU BU CR CT CZ FR GE GG HU IT PL RO SB SK SP ST
SZ UK A: GAN KZ MG WS
fossulifrons Desbrochers des Loges, 1896h: 152 (Apion)
schmidtii Bach, 1854: 206 (Apion)
schmidtii Miller, 1857: 24 (Apion) [HN]
scandinavicum Dieckmann, 1977: 112 (Apion) E: FI NR SV
subinsidiosum Bajtenov, 1974b: 38 (Apion) A: KI KZ
wrangelianum Korotyaev, 1976: 61 (Apion) A: FE

genus Oryxolaemus Alonso-Zarazaga, 1990a: 144 type species Apion scabiosum Weise, 1889
croceifemoratus Gyllenhal, 1839: 396 (Apion) E: FR GR IT N: TU A: CY IS LE TR
boops Gyllenhal, 1839: 396 (Apion)
flavifemoratus Herbst, 1797: 115("125") (Apion) E: AU BU CR CZ DE FR GE HU IT LU PL RO SK SL SZ
A:TR
scabiosus Weise, 1889: 184 (Apion) E: AU CR CZ FR GR IT PT SK SP N: AG MO TU A: IS
viridulus Solari, 1955: 49 (Apion)
viridimicans Desbrochers des Loges, 1884: 161 (Apion) N: AG MO TU

genus Oxystoma Duméril, 1805: 226 type species Attelabus pomonae Fabricius, 1798
Oxyrinus Rafinesque, 1815: 115 [RN] type species Attelabus pomonae Fabricius, 1798
Neoxystoma Bedel, 1912: 275 [RN] type species Attelabus pomonae Fabricius, 1798
abruptum Sharp, 1891: 293 (Apion) A: FE JA (Hokkaido, Honshu) SCH
bipartirostre Desbrochers des Loges, 1889a: xxxiii (Apion) E: ST
cerdo Gerstaecker, 1854a: 235 (Apion) E: AB AU BE BU CR CT CZ DE EN FI FR GB GE GG HU IR IT LA
LT LU NL NR NT PL RO SB SK SL ST SV SZ UK N: AG A: ES FE KI KZ MG TR WS XIN
consanguineum Desbrochers des Loges, 1870a: 187 (Apion)
craccae Linnaeus, 1767: 606 (Curculio) E: AU BU CT CZ DE EN FI FR GB GE GG HU IR IT LA LT LU NL
PL PT RO SB SK SP ST SV UK N: AG MO A: FE SY TR WS
ruficorne Herbst, 1797: 110 (Apion)
subulatum Illiger, 1807a: 307 (Apion) [RN]
viciae DeGeer, 1775: 253 (Curculio)
dimidiatum Desbrochers des Loges, 1897c: 2 E: AU BH BU CR CZ FR (Corse) GE HU IT PL RO SB SK SP
UK
pseudocerdo Dieckmann, 1971: 95 (Apion)
fausti Desbrochers des Loges, 1889a: xxxiii (Apion) E: AB ST A: TR
insignicolle Desbrochers des Loges, 189Id: lvi [= 1891b: 83] (Apion) E: UK A: TR
cilicicum Schubert, 1969: 103 (Apion)
laeviusculum Desbrochers des Loges, 1904a: 58 N: AG
ochropus Germar, 1818: 46 (Apion) E: AB AR AU BU CZ FR GE GG GR HU IT LU NL PL RO SB SK SP ST
SZ UK N: AG A: CY IN IS SY TR "Siberia"
opeticum Bach, 1854: 188 (Apion) E: AR AU BE CT CZ DE EN FI FR GE GG HU IT LT NL NT PL SK SV
SZ A: ES GAN KZ MG WS
angulosum Motschulsky, 1860: 170 (Apion)
dietrichi Dietrich, 1857: 133 (Apion)
nigricorne Motschulsky, 1860: \69 (Apion)
170 APIONIDAE
pomonae Fabricius, 1798: 164 (Attelabus) E: AB AR AU BU CR CZ DE FR GE GB GG GR HU IT LT LU NL
PL PT RO SB SK SP ST SV UK N: AG A: IN SY "Siberia"
breviatum Desbrochers des Loges, 1875a: 32
caerulescens Marsham, 1802: 245 (Curculio)
conspicuum Desbrochers des Loges, 1867a: 42 (Apion)
glabrum Marsham, 1802: 245 (Curculiö) [HN]
subulatum IIliger, 1807a: 307 (Apion)
subulatum Kirby, 1808: 28 (Apion) [HN] E: AU BU CT CZ DE EN FI FR GB GE GG HU IR IT LA LT LU NL
NR NT PL SB SK SP ST SV UK N: AG A: ES FE MG SY WS
marshami Stephens, 1831: 168 (Apion)
selousi Newbery, 1913: 154 (Apion)

genus Pirapion Reitter, 1916a: 243 type species Apion immune Kirby, 1808
immune Kirby, 1808: 52 (Apion) E: AU CZ DE FR GB GE HU IR IT LU NL PT SK SP N: AG "Caucasus"
betulae Gyllenhal, 1833:260 (Apion)
cribricolle Perris, 1858: U6 (Apion) [HN]
redemptum Schatzmayr, 1920: 42 (Apion) E: AU BH HU IT RO SL

genus Tatyanapion Legalov, 1997a: 284 type species Apion laticeps Desbrochers des Loges, 1870
laticeps Desbrochers des Loges, 1870a: 205 (Apion) E: CT FI NT A: KZ WS
lindbergi Wagner, 1943: 158 (Apion)

subtribe Synapiina Alonso-Zarazaga, 1990

genus Ischnopterapion Bokor, 1923:101 type species Apion loti Kirby, 1808
subgenus Chlorapion Györfiy, 1956: 56 type species Apion virens Herbst, 1797
Chlorapion Voss, 1959a: 74 [HN] type species Apion virens Herbst, 1797
Chlorapion Ehret, 1990: 256 [HN]..type species Apion virens Herbst, 1797
cognatum Hochhuth, 1851: 14 (Apion) E: AR BH BU FR GG IT RO SP UK N: AG MO TU A: SY TM TR
arragonicum Everts, 1884: 219 (Apion)
semicaerulans Desbrochers des Loges, 1889b: ccxvi (Apion)
cyrenaicum Solari, 1940: 70 (Apion) N: LB
impressidorsum Desbrochers des Loges, 1896g: 114 [=1907: S6](Apion) A: TR
neorelictum Bajtenov & Lodos, 1977c: 407 (Apion)
leprieuri Wencker, 1864: 214 (Apion) N: AG TU
peyerimhoffi Desbrochers des Loges, 1902a: 109 (Apion) N: AG
subglabrum Desbrochers des Loges, 1870a: 183 (Apion) E: AU GR SL ST A: IS SY TR
glabratum Kiesenwetter, 1864: 289 (Apion) [HN]
glabrum Gemminger, 1871a: 123 (Apion) [RN]
virens Herbst, 1797: 118 (" 128") (Apion) E: AU BU CT CZ DE EN FI FR GB GE HU IR IT LA LT LU NL NR
NT PL RO SB SK SP SV UK N: AG CI MO TU A: SY "Siberia" NARi
aeneocephalum Gyllenhal, 1813: 49 (Apion)
subgenus Ischnopterapion Bokor, 1923: 101 type species Apion loti Kirby, 1808
Leptapion Ehret, 1990: 247 type species Apion loti Kirby, 1808
aeneomicans aeneomicans Wencker, 1864: 242 (Apion) E: AU BU CR CZ FR GE HU IT RO SK SP
rubripes Desbrochers des Loges, 1896d: 90 (Apion)
aeneomicans nothum Desbrochers des Loges, 1897c: 24 (Apion) [DA] N: AG TU
fairmairei Wencker, 1864: 241 (Apion) E: SP N: MO
fallens Marseul, 1888: 452 (Apion) [RN] E: AU BH BU CR CZ FR GE GR HU IT RO SK ST SZ N- AG
fallax Wencker, 1864: 242 (Apion) [HN]
helenae Schatzmayr, 1933: 171 (Apion) N: TU
loti Kirby, 1808: 58 (Apion) E: AN AU BU CR CT CZ DE EN FI FR GB GE GG HU IR IT LA LT LU NL NR
NT PL PT RO SB SK SP ST SV SZ UK N: MO AG A: IN KZ MG SY WS
angustatum Kirby, 1811: 352 (Apion)
angustatum Gyllenhal, 1813: 56 (Apion) [HN]
derelictum Desbrochers des Loges, 1907: 90 (Apion) [HN]
glabratum Germar, 1818: 47 (Apion)
helveticum Desbrochers des Loges, 1907: 88 (Apion)
 APIONIDAE 171
languidum Gyllenhal, 1833: 292 (Apion)
loti Gyllenhal, 1813: 60 (Apion) [HN]
maroccanum Desbrochers des Loges, 1908d: 84 (Apion)
tenuirostre Desbrochers des Loges, 1904b: 109 (Apion) [HN]
modestum Germar, 1817b: 234 (Apion) E: BU CZ DE FR GB GE GRIR IT LU MC ME PL PT SP SV SZ A:
AG MO
plumbeomicam pericarti Ehret, 1991: \16 (Apion) E:FRSP
plumbeomicansplumbeomicam Rosenhauer, 1856: 243 (Apion) E: CR FR IT (Sardegna) PT SP N: AG MO TU
A: SA
sicardi Desbrochers des Loges, 1893b: 9 (Apion)
subconiceps Desbrochers des Loges, 1904a: 57 (Apion) [DA]
tempereanum Ehret, 1991: 173 (Apion) E: FR

genus Ischnopterapion, incertae sedis

distinctirostre Desbrochers des Loges, 1869a: [2] (Apion) "Patria ignota: Europe et confins"

genus Protopirapion Alonso-Zarazaga, 1990a: 123 type species Apion atratulum Germar, 1817
atratulum Germar, 1817b: 192 (Apion) E: AU BU CZ DE FR GB GE HU IR IT LU NL PL PT SK SP SZ N:
AG
striatum Marsham, 1802: 249 (Curculio) [HN]
kraatzii Wencker, 1859a: 273 (Apion) E. FR IT PT SP N: AG TU
crassinassus Desbrochers des Loges, 1896e: 112 (Apion)
parvithorax Desbrochers des Loges, 1891e: lvü [= 1891b: 83] (Apion)

genus Stenopterapion Bokor, 1923: 101 type species Apion tenue Kirby, 1808
subgenus Cobosiotherium Alonso-Zarazaga, 1990a: 118 type species Apion dubium Desbrochers des Loges,
1896
cantabricum Desbrochers des Loges, 1869b: [1] (Apion) [RN] E: FR SP
cantabricum Desbrochers des Loges, 1870b: 160 (Apion) [HN]
heydeni Desbrochers des Loges, 1869a: [2] (Apion) [HN]
dubium Desbrochers des Loges, 1896e: 112 (Apion) E: SP N: MO
scutellare Kirby, 1811:353 (Apion) E: AU FR GB IR PT SP
kirbii Stephens, 1831: 191 (Apion)
kirbyi Germar, 1818: 50 (Apion)
ulicicola Penis, 1840: 90 (Apion)
subsquamosum Desbrochers des Loges, 1891e: lvii [= 1891b: 83] (Apion) E: PT
subgenus Stenopterapion Bokor, 1923: 101 type species Apion tenue Kirby, 1808
argamani Friedman & Freidberg, 2007: 138 A: IS SY
intermedium Eppelsheim, 1875: 76 (Apion) E: AB AR AU BU CR CZ FR GB GE GG HU IT LU PL SK SP ST
SZ UK A: IN ES KZ TR WS
amphibolum Faust, 1890b: 105 (Apion)
margelanicum Wagner, 1912c: 90 (Apion) A: UZ
meliloti Kirby, 1808: 64 (Apion) E: AU BU CR CT CZ DE EN FI FR GB GE HU IT LA LU NL NR NT PL PT
RO SB SP SK ST SV UK N: AG A: ES KZ IN MG SY UZ WS
aeneovirens Schilsky, 1902: 86 (Apion)
bifoveolatum Stephens, 1831: 176 (Apion)
interstitiale Desbrochers des Loges, 1891c: 326 (Apion) [HN]
subviolaceum Desbrochers des Loges, 1908d: 82 (Apion) [DA] E: or A: "Russia"
tenue Kirby, 1808: 61 (Apion) E: AR AU BU CR CT CZ DE EN FR GB GE GG HU IT LA LT LU NL NR PL
RO SB SK SP ST SV TR UK N: AG MO TU A: CY ES IS TR WS
volkovitshi Korotyaev, 1988a: 144 (Leptapion) A: KZ UZ
genus Synapion Schilsky, 1902:13a type species Apion ebeninum Kirby, 1808
subgenus Giustiapion Legalov, 2003b: 67 type species Apion falzonii Schatzmayr, 1922
uliiatlant is Antoine, 1936: 243 (Apion) N: MO
falzonii Schatzmayr, 1922b: 162 (Apion) E: IT
172 APIONIDAE
perraudieri Desbrochers des Loges, 1884: 159 {Apion) N: AG
subgenus Parasynapion Legalov, 2003b: 67 type species Synapion alexandri Korotyaev, 1988
alexcmdri Korotyaev, 1988b: 136 A: ES
kaulbacki Balfour-Browne, 1944a: 7 (Apion) A: XIZ
kerzhneri Ter-Minasian, 1971b: 658 (Apion) A: MG
pistillum Faust, 1894b: 146 (Apion) A: AFKI
substriatum Schilsky, 1902: \1 (Apion)
sobolevae Korotyaev, 1988b: 134 A: TD
subgenus Synapion Schilsky, 1902: 13a type species Apion ebeninum Kirby, 1808
brutum Peyerimhoff, 1925c: 284 (Apion) N: MO
corvinum Faust, 1889b: 224 (Apion) A: JA (Honshu)
ebeninum Kiiby, 1808: 55 (Apion) E: AU CR CZ DE EN FI FR GB GE HU IR IT LU NL NR NT PL SK SV
SZ UK N: AG A: WS
ebeninum Gyllenhal, 1813: 48 (Apion) [HN]
kunzei Boheman, 1839: 419 (Apion)
kozlovi Korotyaev, 1988b: 136 A: QIN

subtribe Trichapiina Alonso-Zarazaga, 1990

genus Betulapion Ehret, 1994:17 type species Apion simile Kirby, 1811
balchaschense Bajtenov, 1978a: 56 (Apion) A: KZ
simile simile Kirby, 1811:351 (Apion) E: AU CT CZ DE EN FI FR GB GE GR HU IT LA LT LU NL NR NT
PL SK SP SV N: AG A: ES FE MG TR WS
eppelsheimi Faust, 1887b: 179 (Apion)
pedicellare C. G. Thomson, 1865: 78 (Apion)
superciliosum Gyllenhal, 1813: 58 (Apion)
triste Germar, 1817b: 233 (Apion)

tribe Piezotrachelini Voss, 1959

genus Conapium Motschulsky, 1866: 430 type species Apion gracile Gerstaecker, 1854
subgenus Conapium Motschulsky, 1866: 430 type species Apion gracile Gerstaecker, 1854
araneiforme Wagner, 1909a: 27 (Apion) A: TAI
benignum Faust, 1898: 292 A: HP UP ORR
bomaense Faust, 1899: 431 (Apion) A: YE AFR
clavipes Gerstaecker, 1854b: 272 (Apion) A: HKG HP UP ORR
constrictum Hartmann, 1897: 84 (Apion) A: SA AFR
cupreum Bhateja & Pajni, 1989b: 183 A: UP ORR
formosanum Wagner, 1914b: 25 (Apion) A: TAI
gagatinum Motschulsky, 1858: 92 (Apion) A: HP UP ORR
martensi Korotyaev, 1987b: 118 A: NP
schenklingi Wagner, 1914b: 23 (Apion) A: TAI ORR
subgenus Piezoconapion Korotyaev, 1990a: 136 type species Apion jekeli Faust, 1892
medvedevi Korotyaev, 1990a: 136 A: YUN ORR

genus Euprotapion Wagner, 1927: 66 type species Apion kueenburgorum Reitter, 1898
kueenburgorum Reitter, 1898a: 125 (Apion) E: AR A: KZ TD TR UZ

genus Fremuthiella Alonso-Zarazaga, 1990a: 83 type species Apion iranicum Bajtenov & Fremuth, 1981
alaica Bajtenov, 1977e: 307 (Apion) A: UZ
aspericollis Wagner, 1908d: 306 (Apion) A: TR
auletoides Reitter, 190le: 228 (Apion) E: AB
avida Faust, 1891c: 412 (Apion) E: CT ST
interruptostriata Desbrochers des Loges, 1870a: 180 (Apion) E: CT ST
iranica Bajtenov & Fremuth, 1981: 250 (Apion) A: IN
kurdaica Bajtenov, 1981f: 408 (Apion) A: KZ
pamiroalaica Bajtenov, 1975a: 74 (Apion) A : TD UZ
 APIONIDAE 173
transversa Bajtenov & Lodos, 1977a: 5 (Apion) A: TR
vossi Ter-Minasian, 1972: 242 (Apion) A: ES MG

genus Microconapion Korotyaev, 1992a: 57 type species Apionpallidirostre Roelofs, 1882

formosicola Wagner, 1914b: 28 (Apion) A: TAI
pallidirostre Roelofs, 1874: 128 (Apion) A: FE JA NE "Korea"
dorsale Schilsky, 1902: 77 (Apion)

genus Piezotrachelus Schoenherr, 1839: 365 type species Piezotrachelus ? germarii Gyllenhal, 1839
subgenus Mepiezotrachelus Voss, 1961a: 425 type species Apion angustithorax Wagner, 1908
yemenicus Wanat, 1990: 76 A: YE
subgenus Piezotrachelus Schoenherr, 1839: 365 type species Piezotrachelus germarii Gyllenhal, 1839
caudatus Korotyaev, 1987a: 90 A: SD
curvirostris Korotyaev, 1987a: 93 A: YUN ORR
fitliginosus Wagner, 1908a: 42 (Apion) A: YE AFR
japonicus Roelofs, 1874: 128 (Apion) A: FUJ HEB JA JIA JIX NC SC TAI YUN ZHE ORR
tschungseni Voss, 1958: 19
kuatunensis Voss, 1958: 19 A: FUJ GUA
lindemannae Voss, 1960a: 3 A: HP PA UP ORR
maraquoensis Voss, 1961a: 429 A: YE AFR
melichari Wanat, 1990: 74 A: YE AFR
congruus Wagner, 1908a: 40 (Apion) [HN]
occidentalis Wagner, 1912b: 44 (Apion) [HN]
ovalipennis Korotyaev, 1987a: 89 A: YUN ORR
sauteri Wagner, 1909a: 27 (Apion) A: GAN JIA TAJ YUN ORR
sinensis Korotyaev, 1987a: 87 A: YUN
sulcatulicollis Voss, 1956c: 187 A: AF HP KA NP PA UP ORR
subgenus Sympiezotrachelus Voss, 1959b: 84 type species Apion helleri Wagner, 1908
punctisternus Balfour-Browne, 1942: 799 A: YE AFR
cephalotes Voss, 1971c: 544
genus Protapion Schilsky, 1908: 608 |RN] type species Apion apricans Herbst, 1797 ]
Podapion Schilsky, 1906b: vii [HN] type species Apion apricans Herbst, 1797
angusticolle Gyllenhal, 1833: 282 (Apion) E: BU CR FR GR IT MC UK N: AG MO A: CY IN IS LE SY
cyprium Balfour-Browne, 1944b: 153 (Apion)
hirticorne Desbrochers des Loges, 1895i: 169 (Apion)
holdhausi Wagner, 1905: 37 (Apion)
longimanum Mulsant & Rey, 1858: 273 (Apion)
rotundicolle Desbrochers des Loges, 1895i: 168 (Apion)
tereticolle Desbrochers des Loges, 1897c: 22 (Apion)
apricans Herbst, 1797: 117("127") (Apion) E: AB AR AU BE CR CT CZ DE EN FI FR GB GE GG GR HU IR
IT LA LT LU NL NR NT PL PT RO SB SK SP ST SV SZ UK N: AG MO A: ES FE IN SY WS
algiricum Desbrochers des Loges, 1895J: 182 (Apion) [HN] [DA]
curvipes Desbrochers des Loges, 1895i: 164 (Apion)
encans tum Wencker, 1864: 201 (Apion)
fuscipes Desbrochers des Loges, 1891c: 320 (Apion)
ochropus Gmelin, 1790: 1757 (Curculio) [HN]
assimile assimile Kirby, 1808: 42 (Apion) E: AB AR AU BE BU CR CT CZ DE EN FI FR GB GE GG GR HU
IR IT LA LT LU NL NR NT PL PT SB SK SP ST SV SZ UK N: AG MO TU A: ES IS SY TR WS
incertum Desbrochers des Loges, 1895i: 164 (Apion) [RN]
méridionale Wencker, 1864: 203 (Apion) [HN]
assimile ryei Blackburn, 1874: 128 (Apion) E: GB (Outer Hebrides, Orkney, Zetland)
brenskei Desbrochers des Loges, 1895i: 167 (Apion) E: GR MC A: IS ®Y TR
burgodionum Schubert, 1958: 33 (Apion) [RN] E: FR IT
delphinense Schubert, 1954: 689 (Apion) [HN]
rosai Pesarini, 1977a: 97 (Apion)
174 APIONIDAE

caiffense Desbrochers des Loges, 1895J: MS (Apion) A: IS

curviscelis Desbrochers des Loges, 1908d: 81 (Apion) E: FR
dentipes Gerstaecker, 1854a: 260 (Apion) E: BH BU CR FR (Corse) GR IT MC PT SB SP A: IS SY TR
armiferum Wencker, 1864: 190 (Apion) [DA]
difforme Germar, 1818: 46 (Apion) E: AU BE BU CR FR GB GE GR IT LU ME NL UK N: AG TU A: SY
TR
tibiale Desbrochers des Loges, 1867a: 40 (Apion)
dissimile Germar, 1817b: 171 (Apion) E: AB AU BE CR CT CZ DE EN FR GB GE GG HU IT LA LU NL PL
RO SB SK SP SV UK N: AG TU A: IS TR "Central Asia"
adjectum Desbrochers des Loges, 1893b: 10 (Apion)
heterocerum C. G. Thomson, 1865: 62 (Apion)
filirostre Kirby, 1808: 26 (Apion) E: AB AR AU BU CR CT CZ DE EN FR GB GE GG HU IT LA LT LU NL
PL RO SB SK ST SV SZ UK "Caucasus" A: FE TR WS
brevicorne Schilsky, 1902: 3 (Apion) [HN]
morio Germar, 1817b: 178 (Apion)
cantianum Wagner, 1906a: 33 (Apion) [RN]
julvipesjulvipes Geoffroy, 1785: 114 (Rhinomacer) E: AB AR AU BE BH BU CR CT CZ DE EN FI FR GB
GE GG GR HU IR IT LA LT LU NL NRNT PL PT RO SB SK SP ST SV SZ UK N: AG MO A: CY ES
FE KZ SY WS
bergrothi Desbrochers des Loges, 1895j: 194 (Apion)
coxale Desbrochers des Loges, 1895i: 170 (Apion)
dichroum Bedel, 1886: 209 (Apion) [RN]
maculicoxis Desbrochers des Loges, 1897c: 23 (Apion)
julvipes lederi Kirsch, 1878: 307 (Apion) E: GG A: TR
gracilipes Dietrich, 1857: 134 (Apion) E: AU CZ FI FR GE HU LT LU PL RO SK SL SV SZ UK A: ES
interjectum boreum Gonget, 1997: 129 (Apion) E: DE FI NR SV
interjection interjectum Desbrochers des Loges, 1895i: 168 (Apion) E: AN AU BH BU CR CT CZ FR GE GR
HU IT LA MA PL PT RO SB SK SL SP SZ UK N: AG MO TU A: IS SY
moczarskii Wagner, 1904: 378 (Apion)
laevicolle Kirby, 1811: 348 (Apion) E: BU CR CZ FR GB GR HU IT MC PT SK SP TR N: AG MO TU A:
SYTR
lesnei Schilsky, 1906a: 28 (Apion) A: FE HEI
sichotealinicum Bajtenov, 1982b: 70 (Apion)
libanicum Desbrochers des Loges, 1895i: 174 (Apion) A: LE
nigritarse Kirby, 1808: 36 (Apion) E: AB AR AU BE CR CZ DE EN FR GB GE GG GR HU IR IT LA LU NL
PL PT RO PT SB SK SP ST SV SZ N: AG MO A: CY ES SY TR
concoloripes Desbrochers des Loges, 1908d: 81 (Apion)
waterhousei Boheman, 1839:408 (Apion)
obesulum Desbrochers des Loges, 1895i: 174 (Apion) A: S Y
ocularium Desbrochers des Loges, 1895j: 187 (Apion) E: GR IT SP N: AG MO TU
mediterraneum Wagner, 1908d: 311 (Apion) [RN]
oblitum Desbrochers des Loges, 1895 i: 168 (Apion) [HN]
ononidis Gyllenhal, 1827: 539 (Apion) E: AU BU CR CT CZ DE EN FR GB GE GG GR HU IT LA LU NL NR
NT PL RO SB SK SP ST SV UK N: AG MO TU A: CY IN IS JO SY TR
bohemanii Boheman, 1839: 405 (Apion)
ononicola Bach, 1854: 195 (Apion) [RN]
pédale Mulsant & Rey, 1858: 271 (Apion) E: FR IT SP
politum Desbrochers des Loges, 1875a: 29 (Apion) A: IS LE JO SY TR
ruficroides Dieckmann, 1973: 87 (Apion) E: AU FR HU IT RO SP SZ
ruficrus Germar, 1817b: 171 (Apion) E: AU BH BU BY CT CZ EN FR GE GG HU IT LA LT MD NT PL RO
SK UK A: WS
nigricrus Desbrochers des Loges, 1895i: 168 (Apion)
schoenherri Boheman, 1839: 406 (Apion) E: AU BH BU CR FR GB GG GR HU IT RO SB SK SP ST UK A:
SYTR
spinicoxale Wagner, 1908c: 103 (Apion) A: INTR
propinquum Desbrochers des Loges, 1908d: 81 (Apion) [DA] [HN]
 APIONIDAE 175
temisiense Palagesiu, 1988: 140 {Apion) E: RO
trifolii Linnaeus, 1768: 224 (Curculio) E: AB AR AU BU CT CR CZ DE EN FI FR GB GE GG GR HU IR IT
LA LT LU NL NR NT PL PT RO SB SK SP ST SV SZ UK N: AG TU A: CY IQ IS JO SY TR
aestivum Germar, 1817b: 169 (Apion)
graecoinsulare Wagner, 1926: 140 (Apion)
hipponense Desbrochers des Loges, 1895J: 186 (Apion) [HN]
leachii Stephens, 1831: 189 (Apion)
linden Wencker, 1858a: xxiii (Apion)
truquii Reiche & Saulcy, 1858: 651 (Apion) E: BU GR A: CY IS JO LE SY TR
varipes Germar, 1817b: 173 (Apion) E: AB AR AU BU CR CT CZ DE EN FR GB GE GG GR HU IT LT LU
NL NR NT PL RO SB SP ST SV UK N: AG A: IN SY TR WS
flavipes Fabricius, 1775: 133 (Curculio)
woerzi Wagner, 1904: 377 (Apion) E: AL GR ME

genus Protapion, species dubiae

porrectum Schrank, 1781: 127 (Curculio) E: AU
tubicen Wencker, 1864: 200 (Apion) E: IT (Sicilia) SP "Pirineos"

genus Pseudoconapion Voss, 1959b: 61 type species Conapium sparsum Faust, 1899
griseosetulosum Desbrochers des Loges, 1875a: 29 (Apion) N: EG A: SA YE
indigoferae Wanat, 1990: 70 A: YE
koreanum Korotyaev, 1987b: 110 A: NC SC
pakistanum Voss, 1960a: 1 (Conapiori) A: PA
niloticum Desbrochers des Loges, 1895d: 83 (Apion) N: EG LB A: YE AFR
mateuiA. Hoffmann, 1962b: 436 [=1963b: 218] (Apion)
mirei A. Hoffmann, 1962b: 436 [=1964: 1216] (Apion)
segne Faust, 1895a: 232 (Apion) A: SA YE AFR
subplumbeum Desbrochers des Loges, 1904b: 108 (Apion) [DA]

genus Pseudopiezotrachelus Wagner, 1907a: 277 type species Apion probum Faust, 1899
brevirostris Korotyaev, 1990a: 147 A: YUN
collaris Schilsky, 1906a: nr. 58 (Apion) [RN] A: AF BEI FE FUJ HP HUB JA JIA JLX KA NC NP SC SCH SD
SHA SHN UP ZHE AUS ORR
conicicollis Schilsky, 1902: 32 (Apion) [HN]
cyrton Alonso-Zarazaga, 1986: 198 (Apion) [RN]
frieseri Alonso-Zarazaga, 1989b: 167
remaudierei A. Hoffmann, 1962c: 664 (Apion) [HN]
silvanus Alonso-Zarazaga, 1986: 199 [RN]
tumidum Gerstaecker, 1854b: 271 (Apion) [HN]
unicolor Roelofs, 1874: 129 (Apion) [HN]
montshadskii Korotyaev, 1990a: 140 A: YUN
placidus Faust, 1887b: 180 (Apion) [DA] A: FE HUB JA NC SC
lugubris Faust, 1889b: 224 (Apion)
ussuriensis Bajtenov, 1977a: 17 (Apion)
simplicirostris Korotyaev, 1987b: 117 A: YUN
schilskyi Wagner, 1906a: 33 (Apion) [RN] [DA] A: UZ
gibbosus Faust, 1887g: 303 (Apion) [HN]
subtilirostris Korotyaev, 1985: 150 A: FUJ GUA JIX YUN ORR
triangulicollis Motschulsky, 1858: 92 (Apion) A: SCH YUN ORR
uruppensis Kôno, 1935: 105 (Apion) A: FE

genus Pseudoprotapion Ehret, 1990: 258 type species Attelabus astragali Paykull, 1800
Wagnerium Alonso-Zarazaga, 1990a: 84 type species Attelabus astragali Paykull, 1800
ammodendroni Bajtenov, 1976: 185 (Apion) A: KZ
androssowi Wagner, 1914a: 28 (Apion) A: KZ
176 APIONIDAE
astragali astragali Paykull, 1800: 180 (Attelabus) E: AR AU BH CR CZ DE EN FR GB GE GG GR HU IT LA
L U N L N R P L P T R O S B S K S P S T S V U K N: AG MO TU A: ES KZ LE MG SY TR UZ WS
saeculare Gozis, 1881b: 181 (Apion) [unnecessary RN]
astragali dwneei A. Hoffmann, 1957a: 58 (Apion) N: MO
coeleste Faust, 1886: 151 (Apion) A: KA HP
elegantulum Germar, 1818: 48 (Apion) E: AB AR AU BU CR CT CZ FR GE GG GR HU IT LA NL RO SK SP
ST SZ UK N: AG MO A: ES SY WS
coracinum Gyllenhal, 1833: 299 (Apion)
incisum Boheman, 1839: 428 (Apion)
laticolle Penis, 1858: 135 (Apion) [HN]
neglectum Gyllenhal, 1833: 253 (Apion)
pineae Rosenhauer, 1856: 244 (Apion)
tricarinatum Waltl, 1835: 76 (Apion)
ergenense Becker, 1864: 482 (Apion) E: BU CZ HU IT PL SK ST UK A: TR
externestriatum Desbrochers des Loges, 1896e: 118 (Apion) E: "Caucasus" A: IN UZ
kobdense Bajtenov, 1977a: 18 (Apion) A: MG
omiense Bajtenov, 1977a: 18 (Apion) A: WS
paracoeleste Balfour-Browne, 1944a: 6 (Apion) A: SD XIZ
phrygium Wagner, 1908d: 307 (Apion) A: TR
pseudelegantulum Wagner, 1912c: 83 (Apion) A: IQ KZ "Transcaspia"
tianschanicum Bajtenov, 198le: 177 (Apion) A: UZ
unguiculare Schilsky, 1902: 36 (Apion) A: IN UZ

genus Scapapion Korotyaev, 1992a: 46 type species Scapapion brunnicorne Korotyaev, 1992
brunnicorne Korotyaev, 1992a: 48 A: YUN ORR
horticola Korotyaev, 1992a: 51 A: YUN ORR
ruficorne Korotyaev, 1992a: 50 A: YUN ORR
uniforme Korotyaev, 1992a: 53 A: YUN

genus Trichoconapion Korotyaev, 1985:153 type species Trichoconapion hirticorne Korotyaev, 1985
hirticorne Korotyaev, 1985: 154 A: SD YUN ORR

tribe Podapiini Wanat, 2001

genus Podapion Riley, 1883: 62 type species Podapion gallicola Riley, 1883
baenai Alonso-Zarazaga, 1998: 134 E: SP
mariae Formanek, 1911: 205 (Magdalis) E: BH

subfamily Apioninae, species incertae sedis

Apion agyrtinum Gistel, 1857: 541 E: GE
Apion astragali Thunberg, 1815: 118 E: SV
Apion atratum Gravenhorst, 1807: 198 E: "Patria ignota"
Apion chloris Gravenhorst, 1807: 199 E: "Patria ignota"
Apion coeruleum Herbst, 1797: 113 ("123") A: GE
Apion curtisii Curtis, 1840: 281 [HN] E: GB
Apion excellens Wagner, 1906b: 205 A: UZ
Apion litigiosum Wencker, 1864: 189 A: TR
Apion longirostre Gravenhorst, 1807: 198 [HN] E: "Patria ignota"
Apion nigrocyaneum Beguin-Billecocq, 1909a: 456 Ei: SP Ni: MO NTR
Apionparpanense Rühl, 1885: 100 E: SZ
Apion strobilanthi Desbrochers des Loges, 1890: 215 A: SD

family Apionidae, nomina dubia

Curculio pulex Goeze, 1777: 380 E: FR
mimt lus Geoffroy, 1785: 115 (Rhinomacer)
pulex Gmelin, 1790: 1758 (Curculio) [HN]
 NANOPHYIDAE 177
family NANOPHYIDAE Gistel, 1848
subfamily Corimaliinae Alonso-Zarazaga, 1989
genus Allomalia Alonso-Zarazaga, 1989c: 228 type species Nanophyes quadrivirgatus A. Costa, 1863
gemmaria Faust, 1887d: 210 {Nanophyes) A: SY
piffli Voss, 1964b: 709 (Corimalia) A: PA
quadrivirgata A. Costa, 1863: 23 (Nanophyes) E: AB BU CR FR GG GR HU IT SP (Baléares) UK N: AG EG
LB MO TU A: CY IS KZ SY UZ
innotatithoraxPk, 1919a: 11 (Nanophyes)
larnacana Pic, 1925b: 13 (Nanophyes)
semiobliterata Pic, 1919a: 11 (Nanophyes)
septemmaculata Pic, 1919a: 11 (Nanophyes)
sexpunctata Kiesenwetter, 1864: 286 (Nanophyes)
trilineata H. Brisout de Barneville, 1869: 346 (Nanophyes)
setulosa Tournier, 1868b: 569 (Nanophyes) E: ST N: AG EG LB TU A: IS LE QA SI AFR
biskrensis H. Brisout de Barneville, 1869: 347 (Nanophyes)
juncta Pic, 1919a: 11 (Nanophyes)
ruficolor Pic, 1935b: 12 (Nanophyes)
separata Pic, 1925b: 14 (Nanophyes)
thoracica Vic, 1925b: 14 (Nanophyes)

genus Corimalia Gozis, 1885: 129 type species Nanophyes tamarisci Gyllenhal, 1838
Pseudocorimalia Voss, 1960b: 141 type species Corimalia exsanguis Voss, 1960
aliéna Faust, 1890b: 103 (Nanophyes) A: ES KZ MG WS
auliensis Pic, 1901d: 90 (Nanophyes)
atritarsis Pic, 1925b: 13 (Nanophyes) N: AG EG A: SI AFR
mutica Peyerimhoff, 1929d: 180 (Nanophyes)
bilineata Tournier, 1868b: 568 (Nanophyes) E: ST N: EG
brunneonotata Pic, 1913b: 177 (Nanophyes) A: IQ QA S Y
alepensis Pic, 1925b: 13 (Nanophyes)
chinensis Faust, 1890e: 474 (Nanophyes) A: GAN
convexipennis Pic, 1925b: 13 (Nanophyes) A: IQ
exsanguis kaszabi Zherikhin, 1976: 427 A: MG
exsanguis exsanguis Voss, 1960b: 141 A: AF IN
faustifausti Reitter, 1890a: 161 (Nanophyes) E: AB BU ST Â: TR
fausti orientalis Zherikhin, 1972: 560 A: MG
gurjevae Zherikhin, 1972: 562 A: MG
gyrata Peyerimhoff, 1929d: 182 (Nanophyes) N: AG EG AFR
helenae Korotyaev & Zherikhin, 1996: 150 E: AB AR ST A: IN IQ KI KZ MG S Y TD TM UZ
inconspicua H. Brisout de Barneville, 1869: 345 (Nanophyes) N: AG
latifrons Pic, 1897a: 28 (Nanophyes) N: AG EG LB TU A: IS JO SI
pectoralis Pic, 1929b: 10 (Nanophyes)
letourneuxi Pic, 1919a: 11 (Nanophyes) N: EG
lukjanovitshi Zherikhin, 1991: 907 A: TM UZ
lunula ta Wollaston, 1863: 218 (Nanophyes) N: CI (Gran Canaria)
maculaticeps Pic, 1929b: 10 (Nanophyes) E: HU
martini H. Brisout de Barneville, 1883: 29 (Nanophyes) N: AG EG TU A: IS AFR
henoni Pic, 1897a: 28 (Nanophyes)
mongolica Faust, 1890e: 475 (Nanophyes) A: MG
multilineata Peyerimhoff, 1930a: 206 (Nanophyes) E: GG N: AG EG A: SY TR AFR
obscuriceps Pic, 1913a: 169 (Nanophyes) A: IQ
obscurithorax Pic, 1897a: 29 (Nanophyes) N: EG
pallida Olivier, 1807: 113 (Cionus) E: CR FR GR IT PT SP N: AG EG LB TU A: IS LE SY
caesifrons H. Brisout de Barneville, 1869: 341 (Nanophyes)
centromaculata A. Costa, 1863: 24 (Nanophyes)
impunctata Rey, 1893c: 83 (Nanophyes)
notatipennis Pic, 1897a: 28 (Nanophyes)
178 NANOPHYIDAE
quinqiiestigma Pic, 1912: 50 {Nanophyes)
oliverii Desbrochers des Loges, 1866: 49 (Nanophyes)
stigmatica Kiesenwetter, 1852: 649 (Nanophyes)
uninotata Pic, 1919a: 11 (Nanophyes)
unipunctata Rey, 1893c: 83 (Nanophyes)
pantherina Zherikhin, 1991: 909 A: TM
pilosella Voss, 1960b: 142 A: AF
postica Gyllenhal, 1838: 787 (Nanophyes) E: FR IT SP N: AG EG TU
chakouri Pic, 1930: 14 (Nanophyes)
impostica Pic, 1919e: 22 (Nanophyes)
obliterata Pic, 1925b: 14 (Nanophyes)
tristigma Rottenberg, 1872: 235 (Nanophyes)
pruinosa Faust, 1885c: 188 (Nanophyes) A: UZ
reaumuriae Zherikhin, 1984: 306 A: ES KI MG
rufosuturalis Pic, 1938: 11 (Nanophyes) E: "Macédoine"
schatzmayri Giordani-Soika, 1937: 20 A: IS
tamarisci Gyllenhal, 1838: 786 (Nanophyes) E: FR IT PT SP N: AG LB MO TU
languida Boheman, 1845: 195 (Nanophyes)
tetrastigma Aube, 1863a: 113 (Nanophyes) E: FR IT PT SP N: AG TU AFR
rubens Aube, 1864: 327 (Nanophyes)
torretassoi Giordani-Soika, 1937: 25 A: IS

genus Hypophyes Reitter, 1916a: 239 type species Rhynchaenus pallidulus Gravenhorst, 1807
aphyllae Peyerimhoff, 1929d: 184 (Nanophyes) N: AG EG A: QA SI TR
exiguus bulganensis Zherikhin, 1976: 427 (Corimalia) A: MG
exiguus exiguus Faust, 1890e: 475 (Nanophyes) A: MG
hyalinus Zherikhin, 1972: 565 (Corimalia) E: ST A: KZ MG WS
betpakdalensis Bajtenov, 1982a: 47 (Corimalia)
minutissimus Tournier, 1868b: 570 (Nanophyes) E: SP ST "Caucasus" N: AG EG LB TU A: AF IQ SI TM
AFR
alfierii Pic, 1926b: 229 (Nanophyes)
externenotatus Pic, 1935b: 12 (Nanophyes)
fayoumensis Pic, 1935b: 12 (Nanophyes)
liliputanus H. Brisout de Barneville, 1869: 349 (Nanophyes)
maculithorax Pic, 1897c: 299 (Nanophyes)
maculatus Tournier, 1868b: 569 (Nanophyes)
modicenotatus Pic, 1901a: 19 (Nanophyes)
scapularis Pic, 1925b: 14 (Nanophyes)
unidentatus Pic, 1929b: 10 (Nanophyes)
tepensis Pic, 1929b: 10 (Nanophyes)
pallidulus Gravenhorst, 1807: 203 (Rhynchaenus) E: FR GB IT PT SP SZ N: AG TU
doriae H. Brisout de Barneville, 1873: 10 (Nanophyes)

genus Titanomalia Alonso-Zarazaga, 1989c: 229 [RN] type species Nanophyes auletoides Reitter, 1911 (=
Nanophyes komaroffi Faust, 1877)
Hadrophyes Reitter, 1911: 246 [HN] type species Nanophyes auletoides Reitter, 1911 (= Nanophyes komaroffi Faust
1877)
Hadrophyes Reitter, 1916a: 239 [HN] type species Nanophyes auletoides Reitter, 1911 (= Nanophyes komaroffi Faust
1877)
komaroffi Faust, 1877: 41 (Nanophyes) E: AB ST
auletoides Reitter, 1911: 246 (Nanophyes)
valida Formânek & Melichar, 1916: 76 (Nanophyes) A: TD

subfamily Nanophyinae Gistel, 1848

genus Alonsiellus Zherikhin, 1996: 237 type species Nanophyes pubescens Roelofs, 1874
pubescens Roelofs, 1874: 172 (Nanophyes) A: FE JA SC
 NANOPHYIDAE 179
genus Ctenomeropsis Voss, 1939e: 64 type species Orobitis nigra Waltl, 1835
nigra Waltl, 1835: 77 (Orobitis) E: CR FR GR IT PT SP UK N: AG MO TU
ericetorum Dufour, 1843: 87 (Nanodes)
sicula Boheman, 1845: 191 (Nanophyes)
subrufescens Pic, 1909b: 122 (Nanophyes)

genus Dieckmanniellus Alonso-Zarazaga, 1989c: 241 type species Nanophyes nitidulus Gyllenhal, 1838
chevrieri Boheman, 1845: 193 (Nanophyes) E: AB AL AR AU BU CZ FR GE GR HU IT MC PT RO SB SK
SP ST SZ N: EG TU A: IS
helveticus Tournier, 1868b: 567 (Nanophyes)
nigromaculatus Pic, 1897c: 299 (Nanophyes)
spretus Jacquelin du Val, 1855: 66 (Nanophyes)
gracilis L. Redtenbacher, 1847: 370 (Nanophyes) E: AU BH BU CR CZ FR GB GE GR HU IT PL SK SP UK
N:AGTU
geniculatus Aube, 1864: 327 (Nanophyes)
geniculatus Kiesenwetter, 1864: 285 (Nanophyes) [HN]
nigritarsis Aube, 1862a: 72 (Nanophyes) E: FR IT (Sardegna, Sicilia)
morulus Vitale, 1906: 135 (Nanophyes)
nitidulus Gyllenhal, 1838: 785 (Nanophyes) E: AB AL AR AU BU CR CZ FR GE GR HU IT MA PL PT RO
SK SP SZ UK N: AG CI EG MO TU A: ES IS KZ SY TR UZ WS
atrolineatus Pic, 1919d: 13 (Nanophyes)
atrolineatus Pic, 1919d: 14 (Nanophyes) [HN]
bonvouloiri Pic, 1919d: 13 (Nanophyes)
dalmatinus Pic, 1919d: 13 (Nanophyes)
difficilis Toumier, 1868b: 567 (Nanophyes)
diversenotatus Pic, 1919d: 13 (Nanophyes)
longulus Woüaston, 1864: 299 (Nanophyes)
margellanicus Pic, 1919d: 13 (Nanophyes)
metallicus Vitale, 1906: l'if» (Nanophyes)
rufithorax Pic, 1919d: 13 (Nanophyes)
rufonotatus Pic, 1919d: 13 (Nanophyes)

genus Microon Alonso-Zarazaga, 1989c: 245 type species Rhynchaenus sahlbergi C. R. Sahlberg, 1835
sahlbergi C. R. Sahlberg, 1835: 51 (Rhynchaenus) E: AU CT CZ DE EN FI FR GE HU IT (Sardegna) NT PL
SK SP ST SV A: TR WS
brevicollis C. N. F. Brisout de Barneville, 1863: 112 (Nanophyes)
castaneidorsis Pic, 1928b: 6 (Nanophyes)
laterale Rosenhauer, 1847: 57 (Nanophyes)
pallens Bettinger, 1935: 2 (Nanophyes)
sparnacensis Bettinger, 1935: 3 (Nanophyes)

genus Nanodiscus Kiesenwetter, 1864: 284 type species Nanophyes transversus Aube, 1850
transversus Aube, 1850: 346 (Nanophyes) E: FR GR IT SP N: AG CI LB MO TU
aureolus Penis, 1864: 299 (Nanophyes)
cuneatus Kiesenwetter, 1852: 650 (Nanophyes)
juniperi Chevrolat, 1860c: 210 (Anthonomus)
micaceus Rey, 1893c: 85 (Nanophyes)

genus Nanomimus Alonso-Zarazaga, 1989c: 248 type species Cionus hemisphaericus Olivier, 1807
anulatus Aragona, 1830: 24 (Nanodes) E: AU CR FR GR IT SB SZ N: AG
annulatus Gyllenhal, 1838: 782 (Nanophyes) [HN]
nigritious Gredler, 1886: 365 (Nanophyes)
borisi Zherikhin, 1996:431 (Nanophyes) A: FE
circumscriptus Aube, 1864: 326 (Nanophyes) E: AR AU CT CZ EN FI FR GE HU KZ LA PL SK ST SV SZ
UK A: WS
latemaculatus Pic, 1900b: 65 (Nanophyes)
obliteratus Pic, 1900b: 65 (Nanophyes)
 180 NANOPHYIDAE
hemisphaericus Olivier, 1807: 113 (Cionus) E: AB AU BH BU CR CZ FR GE GG GR HU IT MA PL PT SB
SK SL SP SZ TR UK N: AG EG MO TU A: KZ LE TR
algericus Pic, 1919d: 15 (Nanophyes)
bivitlatus Fuente, 1912:43 (Nanophyes)
hipponensis Pic, 19}5b: 9 (Nanophyes)
lineatithorax Pic, 1901a: 18 (Nanophyes)
maculipes Rey, 1893c: 85 (Nanophyes
merkli Pic, I919d: 15 (Nanophyes)
postsuturalis Pic, 1915b: 9 (Nanophyes)
quadhpunctatus Pic, 1919d: 15 (Nanophyes)
tibialis Pic, 1919d: 14 (Nanophyes)
turkestanicus Pic, 1901a: 18 (Nanophyes)
ulmi Germar, 1821:314 (Cionus)
smreczynskii Dieckmann, 1963: 192 (Nanophyes) E: BU A: TR
suturalis Pic, 1907a: 127 (Nanophyes) A: FE JA SC
yvonnae A. Hoffmann, 1932: 47 (Nanophyes) E: FR

genus Nanophyes Schoenherr, 1838: 170 [RN] type species Curculio lylhri Fabncius, 1787 (= Curculio
tnarmoratits Goeze, 1777)
Nanodes Schoenherr, 1825: 587 type species Curculio lylhri Fabncius, 1787 (= Curculio marmoratus Goeze, 1777)
Sphaerula Stephens, 1829: 12 [UN] type species Curculio lylhri Fabricius, 1787 (= Curculio marmoratus Goeze, 1777)
Sphaerula Kiesenwetter, 1864: 284 [HN] type species Curculio Ivthri Fabncius, 1787 (= Curculio marmoratus Goeze,
1777)
aegyptiacus Pic, 1900a: 7 N: EG
albovittatus Roelofs, 1874: 173 A: JA SC
teslaceipes Pic, 1907a: 127
basilineatus Heller, 1931: 101 A: TAI
brevis bleusei Pic, 1900a: 8 E: PT SP N: AG
brevis brevis Boheman, 1845: 195 E: AB AU BE BH BU CR CT CZ FR GB GE GR HU IT LU PL RO SB SK
SZUK N:EG A: IN TR
gradli Voss, 1939e: 64
theresae Pic, 1900c: 79
brevis fallen Rey, 1893c: 83 E: FR IT
marie/ A. Hoffmann, 1938c: 127
brevis öbscurus Zherikhin, 1981: 58 A: FE
caucasiens Pic, 1898b: 36 E: "Caucasus" A: TR
zurcheri Pic, 1908b: 51
chibizo Kôno, 1930c: 152 A: TAI
dimorphus Morimoto, 1964: 82 A: JA (Ryukyu)
clonckieri Pic, 1907a: 128 A: YUN
flaveolus Formânek & Melichar, 1916: 70 A: SA TR YE AFR
fiiscomaculatus Pajnt & Bhateja, 1982: 455 A: HP UP ORR
globiformis Kiesenwetter, 1864: 284 E: AU BH BU CT CZ FR GE GR HU IT PL PT RO SB SK SP ST SZ UK
N: MO A: FE
galliots Bedel, 1887d: clxii
globulus Germar, 1821: 315 (Cionus) E: AU BH CR CT CZ FR GE HU IT NL PL RO SK SP SV SZ UK A:
FE
jacqueli Pic, 1935b: 12
pygmaeus Herbst, 1795:142 (Curculio) (HN]
stramineus Bach, 1854: 386
helenae Alonso-Zarazaga, nom. nov. [see New Acts] A: "China"
atrolineatus Pic, 1924: 14 [HN]
impafiformis Voss, 1960b: 140 A: AF
japonicus Roelofs, 1879b: liv A: FE JA SC
leslaceus Roelofs, 1879a: 297
kM-anglsehensis Voss, 1958: 99 A: FUJ
longipilis Pic, 1919c: 13 E: "Caucasus"
 NANOPHYIDAE 181
marmoratus marmoratus Goeze, 1777:413 (Curculio) E: AB AU BU CR CT CZ DE EN FI FR GB GE GG HU
IR IT LA LT LU MD NL NR NT PT RO SB SK SP ST SV SZ UK A: ES KZ LE SC SY TR WS
angustipennis Bach, 1854: 386
anteapicalis Pic, 1919b: 11
apicalis Pic, 1919b: 10
atromaculatus Pic, 1919b: 11
berytensis Pic, 1919c: 13
bifasciatus Pic, 1919b: 11
circassicus Pic, 1919b: 11
circumcinctus Pic, 1919b: 11
discoideus Pic, 1919b: 10
dispersenotatus Pic, 1916: 14
latepallidus Pic, 1919b: 10
leucozonius Gmelin, 1790: 1805 {Curculio)
leukoranus Pic, 1919b: II
leprieuri Pic, 1919b: 11
longemaculatus Pic, 1919b: 11
fy/Än' Fabricius, 1787a: 102 {Curculio) [HN]
fy/Arz Piller & Mitterpacher, 1783: 68 {Curculio)
marmoratus Geoffroy, 1785: 125 {Curculio) [HN]
marmoratus Gmelin, 1790: 1805 {Curculio) [HN]
martial is Pic, 1919b: 11
moldaviensis Pic, 1919b: 11
montandoni Pic, 1919b: 11
muelleri Reitter, 1902a: 196
multinotatus Pic, 1919b: 11
nigrobasalis Pic, 1919b: 11
pulicarius Gravenhorst, 1807: 204 {Rhynchaenus)
rufescens Pic, 1919b: 11
rufipes Tourniez", 1868b: 566
salicariae Fabricius, 1781: 167 {Curculio)
subattenuatus Pic, 1919c: 13
syriacus H. Brisout de Barneville, 1869: 337
tournieri Pic, 1919b: 10
transversus Olivier, 1791: 497 {Curculio) [RN]
trinotatus Pic, 1919b: 12
vz'ffafttf Geoffroy, 1785: 124 {Curculio) [HN]
marmoratus miguelangeli Zherikhin, 1996: 240 A: FE JA
mihokoae Morimoto, 1964: 83 A: JA
nigriceps Boheman, 1859: 147 A: HKG
pallidipes Pic, 1897c: 298 E: "Caucasus"
/>a//*>es Roelofs, 1874: 173 A: JA
poecilopteruspoecilopterus H. Brisout de Barneville, 1869: 334 E: SP N: AG AFR
proles Heller, 1915: 25 A: AP FUJ GUA JA (Ryukyus) NP SD YUN ORR
rubricus Rosenhauer, 1856: 298 E: FR GR IT SP N: AG MO MR TU
fossularum Buysson, 1915: 58
rufipes Motschulsky, 1866: 444 A: FUJ ORR
san is harem is Zherikhin, 1992: 3 A: NP
shaowuensis Voss, 1958: 100 A: FUJ
suavis Gistel, 1857: 529 E: GE
triptolemus Gistel, 1857: 546 (Nanodes) E: GE
usuironis Kôno, 1930c: 152 A: JA
villaücus Gistel, 1857: 546 {Nanodes) E: GE
vulgaris Pajni & Bhateja, 1982: 462 A: NP UP ORR

genus Pericartiellus Alonso-Zarazaga, 1989c: 238 type species Nanophyes flavidus Aube, 1850
durieui P. H. Lucas, 1849: 460 {Nanophyes) E: FR PT SP N: AG TU
lucasi Pic, 1900c: 79 {Nanophyes)
182 BRACHYCERIDAE
flavidus Aube, 1850: 345 (Nanophyes) E: FR OE SP ST SZ A: TR
nicodi Pic, 1915a: 2 (Nanophyes)
vernetensis Pic, 1928b: 5 (Nanophyes)
luteonotatus Pic, 1919a: 11 (Nanophyes) E: "Caucasus"
palaestinus Pic, 1900a: 7 (Nanophyes) A: IS
te/epA/ï Bedel, 1900: 304 (Nanophyes) E: CZ FR SK ST N: AG A: KZ WS
buyssoni Pic, 1928b: 5 (Nanophyes)
reductenotatus Pic, 1915b: 9 (Nanophyes)
zinovjevi Zherikhin, 1981: 59 (Nanophyes) A: FE

genus Pseudorobitis L. Redtenbacher, 1868:162 type species Pseudorobitis gibbus L. Redtenbacher, 1868
Temnalysis Alonso-Zarazaga, 1989a: 125 type species Temnalysis axeli Alonso-Zarazaga, 1989
axeli Alonso-Zarazaga, 1989a: 127 (Temnalysis) A: JA (Ryukyu) TAI
gibbus L. Redtenbacher, 1868: 163 A: SHG
miwai Kôno, 1930c: 154 (Nanophyes) [DA] A: TAI

genus Shiva Pajni & Bhateja, 1982: 467 type species Shiva dimorphorostris Pajni & Bhateja, 1982
taiwanus Kantoh & Kojima, 2009: 165 A: TAI

genus Zherikhinia Alonso-Zarazaga, 1999: 62 [RN] type species Nanophyes formosensis Kôno, 1930: 150
P i a Alonso-Zarazaga, 1989c: 232 [HN] type species Nanophyes formosensis Kôno, 1930: 150
formosensis Kôno, 1930c: 150 (Nanophyes) A: JA TAI

subfamily Nanophyinae, species incertae sedis

Curculio salicariae Piller & Mitterpacher, 1783: 69 [HN] E: CR

family BRACHYCERIDAE Billberg, 1820

subfamily Brachycerinae Billberg, 1820
tribe Brachycerini Billberg, 1820
genus Brachycerus Olivier, 1789: 36 type species Curculio barbarus Linnaeus, 1758
Dyerocera Pascoe, 1887: 334 type species Dyerocera gravida Pœcoe, 1887
Brachycerinus Bovie, 1909: 11 type species Brachycerus dorsalis Gyllenhal, 1833
Brachyceromorphus Bovie, 1909: 11 type species Curculio tauriculus Sparrman, 1785
aegyptiacus Olivier, 1807: 51 E: GR (Kriti) N: EG A: IS SY
incertus Desbrochers des Loges, 1871: 367
olivieri Desbrochers des Loges, 1871: 367
albidentatus Gyllenhal, 1840: 654 E: CR FR (Corse) IT MA
argillaceus Reiche & Saulcy, 1858: 654 E: GR TR A: CY IS LB TR
nodulosus Reiche & Saulcy, 1858: 658
balearicus Bedel, 1874: 171 E: SP (Baléares)
barbarus Linnaeus, 1758: 386 (Curculio) E: FR IT SP N: MO TU
curtulus Desbrochers dés Loges, 1871: 366
fluctiger Gyllenhal, 1840: 651
interntptus Rey, 1894: 63
lateralis Gyllenhal, 1833: 407
latro Gyllenhal, 1833:412
libertinus Fâhraeus, 1840: 650
paradoxus Rey, 1894: 63
ramosus L. W. Schaufuss, 1882: 542
chevrolati Fâhraeus, 1840: 657 E: IT (Sicilia) PT SP N: AG MO TU
callosus Fâhraeus, 1840: 657 [HN, unnecessary RN]
parens Desbrochers des Loges, 1871: 366
peninsularis Chevrolat, 1866: 105
raffrayi Desbrochers des Loges, 1871: 365
 BRACHYCERIDAE 183
semituberculatus P. H. Lucas, 1847: 411
cinereus Olivier, 1807: 56 E: AR GR RO N: EG MO A: LB SI
pulverulentus Olivier, 1807: 56
cribrarius Olivier, 1807: 58 E: BU A: IS LB TR
fimbriates Desbrochers des Loges, 1871: 365
ornatus Reiche & Saulcy, 1858: 658
crispatus Fabricius, 1792: 382 N: AG TU
clathratus Desbrochers des Loges, 1871: 362
rectecostatus Desbrochers des Loges, 1871: 362
cylindripes Bedel, 1874: 171 N: MO
desertus A. Hoffmann, 1950a: 91 N: MO
foveicollis Gyllenhal, 1833:419 E: AU BU HU MD RO SK ST TR A: SY TR
muricatus Fabricius, 1792: 383 [HN]
ventralis Desbrochers des Loges, 1871: 366
foveifrons Bedel, 1874: 194 A: IS SY TR
freyi Zumpt, 1937b: 359 A: TR
gibbosus Haaf, 1957: 25 N: EG A: SA YE AFR
graecus Zumpt, 1937c: 402 E: GR A: TR
hustachei Zumpt, 1937b: 357 N: MO
junix Lichtenstein, 1796: 55 E: AR CR GR IT MC RO A: CY IN IS LB SY TM TR
siculus Gyllenhal, 1833: 414
tauricus Desbrochers des Loges, 1871: 367
kabylianus Desbrochers des Loges, 1871: 364 N: AG TU
velutinus Desbrochers des Loges, 1871: 364
kubanicus Arzanov, 2005: 69 E: ST
lutulentus Gyllenhal, 1833: 418 E: UK
lutosus Gyllenhal, 1833: 417 E: BU BH FR GR HU IT ST TR A: TR
superciliosus Gyllenhal, 1833:422
mlokosevitschi Arzanov, 2005: 69 E: GG
muricatus Olivier, 1790: 136 E: BH CR FR GR HU IT MA SP TR N: AG MO TU A: TR
algirus Fabricius, 1787: 129 (Curculio) [HN]
attenuates Vitale, 1903: 42
cannula Rey, 1894: 63
cirrosus Gyllenhal, 1840: 660
exemptes Rey, 1894:63
perodiosus Gyllenhal, 1840: 661
planirostris Gyllenhal, 1833: 417
pygmaeus Vitale, 1903: 5
semiaeneus Desbrochers des Loges, 1871: 365
sulcifrons Gyllenhal, 1833: 420
normandi Desbrochers des Loges, 1898a: 11 N: EG LB MO TU
orbipennis Reiche & Saulcy, 1858: 656 A: IS SY
perlatus A. Solari & F. Solari, 1922a: 85 N: EG LB MO TU
persicus Zumpt, 1937b: 357 A: IN
plicatus Gyllenhal, 1833: 415 E: CR GR (Rodos) IT (Sardegna, Sicilia) PT SP N: AG CI LB MO TU A: CY
TR
clathratus Desbrochers des Loges, 1871: 362
insignis Miller, 1861:206
opacus Wollaston, 1864: 334
scutipennis Desbrochers des Loges, 1871: 363
tetanicus P. H. Lucas, 1847: 414
vespertilio Desbrochers des Loges, 1871: 363
pradieri atlasicus A. Hoffmann, 1962a: 121 N: MO
pradieripradieri Fairmaire, 1857: 536 E: FR SP
quadrisulcatus Fischer von Waldheim, 1830: 185 E: AB AR A: TR
difformis Faldermann, 1837: 16
repertus Roudier, 1958: 59 [RN] E: SP
184 BRACHYCERIDAE
hypocrita Bedel, 1874: 174 [HIM]
riguus Erichson, 1841: 185 A: AG
rotundicollis Escalera, 1918: 415 E: SP
rufipes Zumpt, 1937b: 353 E: GR (Kriti)
schatzmayri Zumpt, 1937b: 367 E: IT (Lampedusa Is.) N: TU
scutellaris P. H. Lucas, 1847: 413 N: AG LB TU
similaris Zumpt, 1937b: 360 N: AG
sinuatus Olivier, 1807: 58 E: AB AR BU GR IT (Sicilia) MC RO UK A: CY TR
bessert Krynicki, 1832: 145
spinicollis Bedel, 1874: 205 A: IS LE N: EG
trcmsversus Olivier, 1807: 57 N: AG MO
turkmenicus Arzanov, 2005: 69 A: TM
undatus Fabricius, 1798: 162 E: BH BU CR FR GR HU IT MA SP TR N: AG TU
capensis Schoenherr, 1833: 427
cornifrons Desbrochers des Loges, 1871: 363
corrosus Gyllenhal, 1833:409
europaeus Thunberg, 1799: 25
incultus Gyllenhal, 1833: 411
hispidus Desbrochers des Loges, 1871: 361
lacunatus Latreille, 1806: 253
fyrae Vitale, 1905: 206
mauritanicus Olivier, 1807: 52
nubilus Desbrochers des Loges, 1871: 361
ovatus Brullé, 1832:235
pterygomalis Gyllenhal, 1833: 406
sericeus Desbrochers des Loges, 1871: 361
serratus Olivier, 1807: 51
subvariolatus Desbrochers des Loges, 1871: 360
zaninii F. Solari, 1949: 94 N: MO

subfamily Ocladiinae Lacordaire, 1865

tribe Ocladiini Lacordaire, 1865
genus Ocladius Schoenherr, 1825: 587 type species Rhynchaenus salicorniae Olivier, 1807
abdeldayemi Meregalli & Colonnelli, 2006: 275 N: EG A: SI
aidhabensis Meregalli & Colonnelli, 2006: 267 A: SI
balkis Meregalli & Colonnelli, 2006: 293 A: YE
barani Pascoe, 1874: 35 A: SA YE
sharpi Tournier, 1875a: 476
bifasciatus Tournier, 1875a: 475 N: EG A: SA YE AFR
setipes Ancey, 1881: 372
bufo Vauloger, 1899: 405 N: AG TU
engelhardi Pic, 1894c: 611 A: TM UZ
eremorum Meregalli & Colonnelli, 2006: 282 A: AE OM SA YE
grandii Osella & Meregalli, 1986: 110 E: SP (Cabo de Gata)
holomelas Fairmaire, 1877: 98 N: AG MO
paucisquamis Meregalli & Colonnelli, 2006: 277 A: IS JO
rufithorax Pic, 1894b: 588 A: SA YE AFR
russelli Meregalli, Borovec & Colonnelli, 2001: 147 N: MO
salicorniae Olivier, 1807: 111 {Rhynchaenus) A: IN IQ KZ TD TM UZ
trinotatus Fischer von Waldheim, 1821: 14 (Curculid) [HN]
solarii Caldara, 1976: 300 N: EG LB A: SI
subelongatus Pic, 1905b: 122 A: SI
walkeri Cameron, 1905: 179 A: YE (Perim Is.)
 DRYOPHTHORIDAE 185

tribe Desmidophorini Morimoto, 1962

genus Desmidophorus Dejean, 1835: 296 type species Curculio hebes Fabricius, 1781
Botrobatys Chevrolat, 1842: 671 type species Curculio fascicularis Olivier, 1791
Desmidophorinus Hubenthal, 1917a: 111 type species Desmidophorus aureolus Gyllenhal, 1837
Pseudotrichosomus Hustache, 1925b: 386 [unjustified RN]
Trichosomus Chevrolat, 1881:91 [HN] type species Curculio senex Boheman, 1845
brachmanus Faust, 1895b: 263 A: SD
confucii Boheman, 1837: 362 A: S W ORR
crassus Hubenthal, 1917b: 200 A: JA TAI
hebes Fabricius, 1781: 174 {Curculio) A: CE SD ORR
aterrimus Aurivillius, 1891: 220
morphosus Pascoe, 1888: 416
helleri Hubenthal, 1917b: 205 A:NP ORR

family DRYOPHTHORIDAE Schoenherr, 1825

subfamily C r y p t o d e r m a t i n a e Bovie, 1908
genus Cryptoderma Ritsema, 1885:54 [RN] type-species Calandra discors Fabricius, 1801
Nosoxylon Gistel, 1848: x [unused RN]
Oxyrhynchus Schoenherr, 1823: 1137 [HN] type species Calandra discors Fabricius, 1801
formosense Kôno, 1934b: 279 A: TAI
fortunei G. R. Waterhouse, 1853:172 {Oxyrhynchus) A: FUJ GUI GUX HUB HUN JA JLX SCH TAI ZHE
"Korea"
grande Fairmaire, 1891: xxiv {Oxyrhynchus) A: CH
kuniyoshii Morimoto, 1978b: 106 A: JA

subfamily Dryophthorinae Schoenherr, 1825

genus Dryophthorus Germar, 1824: 302 type species Curculio lymexylon Fabricius, 1792 (= Curculio
corticalis Paykull, 1792)
Bulbifer Dejean, 1821: 99 [supressed] type species Curculio lymexylon Fabricius, 1792 (= Curculio corticalis Paykull,
1792)
Dryophora Berthold, 1827: 391 [unnecessary RN] type species Curculio lymexylon Fabricius, 1792 (= Curculio corticalis
Paykull, 1792)
Tetraspartus Pascoe, 1885: 309 type species Tetraspartus bagoides Pascoe, 1885
Tetratemnus Wollaston, 1873a: 9 type species Tetratemnus sculpturatus Wollaston, 1873
corticalis Paykull, 1792: 41 {Curculio) E: AU BU CZ DE EN FI FR GB GE GG GR HU IT LA LT NL NR PL
SK SL SP ST SZ UK A: ES FE JA AFR AUS NAR
lymexylon Fabricius, 1792: 420 {Curculio)
sculpturatus Kôno, 1938: 144 {Tetratemnus) [HN]
japonicus Konishi, 1963: 126 A: JA
konishii Morimoto, 1985: 67 A: JA
kusuii Morimoto, 1985: 71 A: JA
ogasawarensis Morimoto, 1985: 69 A: JA
sculpturatus Wollaston, 1873a: 11 {Tetratemnus) A: ES FUJ JA

genus Stenommatus Wollaston, 1873b: 442 type species Stenommatus fiyi Wollaston, 1873
nanus Champion, 1925: 270 A: UP
ocularis Konishi, 1963: 128 {Dryophthorus) A: JA

subfamily Orthognathinae Lacordaire, 1865

tribe Orthognathini Lacordaire, 1865
genus Sipalinus G. A. K. Marshall, 1943: 119 typespecies Calandra granulata Fabricius, 1801 (= Curculio
gigasFabricius, 1775)
Hyposipalus Voss, 1962: 356 type species Curculio guineensis Fabricius, 1798
Prosipalinus Voss, 1962:357 type species Hyposipalus fallaciosus Voss, 1962
186 DRYOPHTHORIDAE
Sipalus Schoenherr, 1825: 587 [HN] type species Calandra granulata Fabricius, 1801 (= Curculio gigas Fabricius, 1775)
gigas gigas Fabricius, 1775: 127 (Curculio) A: AP FE FUJ GUA GUI GUX HKG HUB HUN IN JA JIA JIX
MG NC SC SCH SD TAI UP YUN "Himalayas" ORR
chinensis Fairmaire, 1887: \30 (Sipalus)
formosanus Kôno, 1934a: 7 (Sipalus)
granulalus Fabricius, 1801: 432 (Calandra)
hypocrita Boheman, 1845: 209 (Sipalus)
tinctus Walker, 1859b: 218 (Sipalus)
yunnanensis Vaurie, 1971: 40 A: YUN

subfamily Rhynchophorinae Schoenherr, 1833

tribe Diocalandrini Zimmerman, 1993
genus Diocalandra Faust, 1895b: 353 type species Calandra frumenti Fabricius, 1801
elongata Roelofs, 1875a: 187 (Calandra) A: FUJ JA NARi
nigromaculata Voss, 1956b: 44
frumenti Fabricius, 1801: 438 (Calandra) Ei: FR Ni: CI A: HKGJAiTAI AFR AUR ORR
bifasciata Boisduval, 1835: 445 (Calandra)
crucigera Motschulsky, 1858: 69 (Sphaenophorus)
punctigera Pascoe, 1885: 305(Calandra)
sechellarum Kolbe, 1910: 46 (Calandra)
stigmaticollis Gyllenhal, 1838: 972 (Sitophilus)
subfasciata Boheman, 1838: 971 (Sitophilus)
subsignataBoheman, 1838: 911(Sitophilus)
kamiyai Morimoto, 1978b: 109 A: JA
sasa Morimoto, 1978b: 108 A: JA

genus Myocalandra Faust, 1895b: 353 type species Myocalandra discors Faust, 1895
Paracalendra Chûjô & Morimoto, 1959a: 26 type species Paracalendra saccharivora Chûjô & Morimoto, 1959 (=
Sitophilus exaratus Boheman, 1838)
exarata Boheman, 1838: 970 (Sitophilus) A: ES JA TAI ORR
exquisita Walker, 1859b: 218 (Sphenophorus)
porcata Pascoe, 1885: 306 (Calandra)
saccharivora Chûjô & Morimoto, 1959a: 26 (Paracalendra)

tribe Litosomini Lacordaire, 1865

genus Autonopis Pascoe, 1874: 75 type species Autonopis lineata Pascoe, 1874
agrana Heller, 1927: 12 A: UP
tenuicornis Heller, 1927: 12 A: TAI

genus Fursovia Alonso-Zarazaga & Lyal, 1999: 65 [RN] type species Calandrellapashini Fursov, 1935
Calandrella Fursov, 1935: 440 [HN] type species Calandrella pashini Fursov, 1935
pashini Fursov, 1935: 440 (Calandrella) A: TD

genus Laogenia Pascoe, 1874: 75 type species Laogenia sorex Pascoe, 1874
formosana Heller, 1926: 185 A: JA TAI

genus Sitophilus Schoenherr, 1838: 967 type species Curculio oryzae Linnaeus, 1763
Calandra Gistel, 1848: 136 [HN] [unnecessary RN] type species Curculio oryzae Linnaeus, 1763
glandium G. A. K. Marshall, 1920: 277 (Calandra) A: UP
gotschi Hochhuth, 1847: 579 E: AR GG "Caucasus" "Transcaucasus"
granarius Linnaeus, 1758: 378 (Curculio) E: AU BU BY CR CZ DE EN FI FR GB GE GG GR HU IC IT LA
LT MA NL NR PL PT RO SP ST SV SZ UK YU N: AG MO A: AF GAN HKG HUB HUN JIA KI SA
TAI WS XIN ZHE COS
contractus Geoffroy, 1785: 126 (Curculio)
laevicosta R. A. Philippi & F. Philippi, 1864b: 374 (Calandra)
pulicarius Panzer, 1798: 54 (Curculio) [HN]
 DRYOPHTHORIDAE 187
remotepunctatus Gyllenhal, 1838: 979 (Sitophilus)
segetis Linnaeus, 1758: 381 (Curculio)
unicolor Marsham, 1802: 275 (Curculio)
linearis Herbst, 1795: 5 (Rhynchophorus) E: AB AR AU IT PL ST TR N: CI A: HAI IN IQ IS JO LE SA SI
SY UP YUN AFR AUR NAR NTR ORR
striates Thunberg, 1815:112 (Cordyle)
tamarindi Christy, 1834: 36 (Calandra)
oryzae Linnaeus, 1763: 395 (Curculio) E: AB AR AU BU BY CR CZ DE EN FI FR GB GG GR HU IC IT LA
MA NL NR PL PT SP ST SV SZ TR UK N: CI EG MO A: CY FE FUJ GUA GUI GUX HAI HKG HUB
HUN IN IQ IS JA JIA JO LE NMO SCH SHX SD SI SY TAI WS XIZ YE YUN COS
frugilegus DeGeer, 1775: 273 (Curculio)
funebris Rey, 1895b: 50 (Calandra)
granarius Strem, 1783: 56 (Curculio) [HN]
minor Sasaki, 1899: 202 (Calandra)
orizae Masters, 1886: 684 (Calandra)
quadriguttatus Montrouzier, 1861a: 910 (Sphenophorus)
sasakii Takahashi, 1928: 164 (Calandra) [RN]
rugicollis Casey, 1892: 687 (Calandra) A: UP AFR NAR
rugosicollis Hustache, 1921b: 191, 192 (Calandra)
shoreae G. A. K. Marshall, 1920: 276 (Calandra)
sculpturatus Gyllenhal, 1838: 974 (Sitophilus) A: CY AFR ORR
zeamais Motschulsky, 1855: 77 (Sitophilus) E: AB AL AN AR AU AZ CZ DE FI GR GB IT PL SP ST SV TR
N: CI EG MR A: BT CY GUI GUX HEI HEN HKG HUB HUN JIX IN IQ IS JA JO LE NP SD SY COS
chilensis R. A. Philippi & F. Philippi, 1864a: 274 (Calandra)
platensis Zacher, 1922: 56 (Calandra)
quadrimaculus Walker, 1859b: 219 (Cossonus)
genus Tryphetus Faust, 1895b: 355 type species Sitophilus incarnatus Gyllenhal, 1838
nigromaculatus Voss, 1958: 127 A: FUJ

tribe Ommatolampini Lacordaire, 1865

genus Ommatolampes Schoenherr, 1838: 837 type species Calandra haemorrhoidalis Wiedemann, 1819
Ommatolampus Pascoe, 1887a: 374 [unjustified emendation] type species Calandra haemorrhoidalis Wiedemann, 1819
paratasioides Heller, 1897:245 (Ommatolampus) A: TAIi ORR

tribe Polytini Zimmerman, 1993

genus Polytus Faust, 1895b: 353 type species Sitophilus mellerborgii Boheman, 1838
mellerborgii Boheman, 1838: 976 (Sitophilus) A; J^. TAI UP AFR AUR ORR
musaecola Fairmaire, 1898b: 489 (Sphenophorus)
remotus Sharp, 1885: 183 (Calandra)

tribe Rhynchophorini Schoenherr, 1833

genus Cyrtotrachelus Schoenherr, 1838: 833 type species Curculio longipes Fabricius, 1781 (= Cyrtotrachelus
thompsoni Alonso-Zarazaga & Lyal, 1999)
Roelofsia Ritsema, 1891: 148 type species Cyrtotrachelus buquetii Guérin-Méneville, 1844
bipartitus Hartmann, 1899a: 29 A: NP SD ORR
buquetii borealis Jordan, 1894a: 493 (Roelofsia) A: JA SCH SHG ORR
buquetii buquetii Guérin-Méneville, 1844: 176 A: FUJ GUA GUI GUX JA JIA SCH SD ORR
dux Boheman, 1845:221
nigrocinctus Faust, 1895b: 320
rex Chevrolat, 1883:555
davidis Fairmaire, 1878: 127 A: "Central China"
himalayanus Heller, 1923: 155 A: SD
rufopectinipes Chevrolat, 1883: 556 A: HUB JIX SD YUN ORR
birmanicus Faust, 1895b: 321
montanus Heller, 1923: 154
subnotalus Voss, 1931a: 37
 188 DRYOPHTHORIDAE
thompsoni Aionso-Zarazaga & Lyal, 1999: 64 [RN] A: FUJ GUA GUI GUX HEN HKG HUB HUN JA JIA JIX
SCH SD SHA TAI YUN ZHE AFR ORR
longimamis Fabricius, 1775: 822 (Curcuiio) [HN]
longipes Fabricius, 1781: 162 (Curcuiio) [HN]

genus Macrocheirus Schoenherr, 1838: 831 type species Macrocheirus praetor Gyllenhal, 1838
Macrochirus Agassiz, 1846: 220 [UN, unjustified emendation] type species Macrocheirus praetor Gyllenhal, 1838
Macrochirus Gemminger, 1871b: 2640 [HN] type species Macrocheirus praetor Gyllenhal, 1838
longipes Drury, 1773: 61 (Curcuiio) A: CH ORR

genus Omotemnus Chevrolat, 1883: 559 type species Calandra serrirostris Fabricius, 1801
carnifex Faust, 1891b: 344 A: CH
caelirostris Heller, 1894: 98 A: NP ORR
compressirostris Jordan, 1894a: 492
fleutiauxi Faust, 1891b: 342 A: HAI
rhinocerus Chevrolat, 1883: 560 A: CH

genus Otidognathus Lacordaire, 1865: 273 [RN] type species Litorhynchus westermanni Boheman, 1845
Litorhynchus Schoenherr, 1845: 222 [HN] type species Litorhynchus westermanni Boheman, 1845
aphones Günther, 1934: 454 A: GUA
tonkinensis Günther, 1934: 455
bifasciatus Chevrolat, 1882a: cxi (Litorrhynchus) A: SD ORR
decemstriatus Chevrolat, 1883: 557
robustus Faust, 1895b: 324 (Litorrhynchus)
subfasciatus Chevrolat, 1882b: cxi (Litorrhynchus)
complus Pascoe, 1887a: 373 A: HKG ORR
davidis badins Günther, 1938: 85 A: JIA JIX ZHE
davidis davidis Fairmaire, 1878: 127 (Cyrtotrachelus) A: AHN FUJ GUA GUX HEN HUB HUN JA JIA JIX
SCH SHG ZHE
extraordinarius Günther, 1938: 80 A: SD
incertus Günther, 1938: 86 A: FUJ
obscurior Voss, 1958: 120
jansoni Roelofs, 1875a: 186 A: FUJ GUA GUI HAI HEN HUB HUN JA JIA JIX SCH ZHE "Korea"
maculipennis Voss, 1931a: 38
nigropictus Fairmaire, 1878: 128
satteloides Voss, 1958: 120
nemorivagus Günther, 1935: 154 A: BT
pygidialis areolatus Fairmaire, 1900: 636 (Cyrtotrachelus) A: FUJ SCH
pygidialis intermedius Günther, 1938: 87 A: FUJ
pygidialis pygidialis Jordan, 1894a: 495 A: FUJ JA JLX JIA XIN
quadrimaculatus Buquet, 1844: 177 (Cyrtotrachelus) A: FUJ GUA NP SD ORR
assamensis Chevrolat, 1882c: cxxxviii [RN]
simplex Voss, 1958: 119
rants Günther, 1935: 154 A: SHG ORR
rubriceps cantonensis Günther, 1938: 83 A: GUA
rubriceps melli Günther, 1938: 83 A: SCH
rubriceps notatus Voss, 1932d: 299 A: YUN
rubriceps rubriceps Chevrolat, 1882b: cxii {Litorhynchus) A: FUJ SCH SD ORR
celatus Pascoe, 1887a: 374
westermanni Boheman, 1845: 223 (Litorhynchus) A: SD ORR
genus Protocerius Schoenherr, 1838: 828 type species Curcuiio colossus Olivier, 1791
fervidus Pascoe, 1871a: 216 A: SDUP ORR
grandis Guérin-Méneville, 1844: 174 A: CH ORR
 DRYOPHTHORIDAE 189
genus Rhynchophorus Herbst, 1795: pl. LX type species Curculio palmarum Linnaeus, 1758
Cordyle Thunberg, 1797:44 type species Curculio palmarum Linnaeus, 1758
ferrugineus Olivier, 1791: 473 (Curculio) Ei: FR GG IT PT SP TR Ni: EG MO A: AEi BAi GUA HAI INi IQ
ISi JAi JOi KUi NP OMi PAi QAi SAi TAI XIA YUN AUR NTRi ORR
cinctus Faust, 1893c: 520
dimidiatus Faust, 1895b: 330
glabrirostris Schaufuss, 1855: 204
hemipterus Sulzer, 1776b: 39 (Curculio)
indostanus Chevrolat, 1883: 562
pascha Boheman, 1845: 218
papuanus Kirsch, 1877b: 156
seminiger Faust, 1895b: 330
signaticollis Chevrolat, 1883: 562
schach Fabricius, 1801: 433 (Calandra)
sexmaculatus Thunberg, 1797: 46 (Cordyle)
tenuirostris Chevrolat, 1883: 561
vulneratus Panzer, 1798: [x] (Curculio)

tribe Sphenophorini Lacordaire, 1865

genus Adapanetus Günther, 1936a: 77 type species Adapanetus sericoclava Günther, 1936
sericoclava fukienensis Voss, 1956b: 44 A: FUJ

genus Anathymus Pascoe, 1885: 299 type species Anathymus singularis Pascoe, 1885
tricolor Heller, 1931: 110 A: TAI

genus Aplotes Chevrolat, 1885b: 100 type species Aphtes aliéna Chevrolat, 1885
roelofsi Chevrolat, 1882d: clix (Sphenophorus) [RN] A FUJ HAI JA "Korea"
trapezicollis Voss, 1956b: 42 A: FUJ

genus Cactophagus LeConte, 1876: 331 type species Sphenophorus validus LeConte, 1858
Cactophagoides Champion, 1910: 89 type species Cactophagoides verrucosus Champion, 1910
Eucactophagus Champion, 1910: 96 type species Calandra aurofasciata Brème, 1844
Paradiaphorus Chevrolat, 1885c: 287 type species Sphenophorus crenatus Gyllenhal, 1838
Phyllerythrurus Chevrolat, 1885a: 92 type species Curculio sanguinolentus Olivier, 1791
crenatus Gyllenhal, 1838: 876 (Sphenophorus) N: Cli NTR

genus Cercidocerus Guérin-Méneville, 1833: pl. 39bis type species Cercidocerus nigrolateralis Guérin-
Méneville, 1833
distinctus Faust, 1890a: 79 A: SD
erubescens Heller, 1931: 111 A: TAI
lateralis Fâhraeus, 1845: 232 A: SD ORR
rufipes Günther, 1935: 166 A: SD ORR
schoenherri Guérin-Méneville, 1844: 179 A: SD ORR

genus Cosmopolites Chevrolat, 1885c: 289 type species Calandra sordida Germar, 1824
sordidus Germar, 1824: 299 (Calandra) E: SPi N: MRi A: FUJ GUA GUI GUX GUI HKG HAI JA NP SD
TAI YUN AFR AUR NAR NTRi ORR
cribricollis Walker, 1859b: 218 (Sphenophorus)
maderensis A. Hoffmann, 1938a: 48 (Sphenophorus)
striatus Fâhraeus, 1845:251 (Sphenophorus)

genus Metaprodioctes Günther, 1937: 281 type species Prodioctes dux Faust, 1895
formosanus Heller, 1931: 110 (Prodioctes) A: TAI
trilineatus Hope, 1831: 27 (Calandra) A: NP SD
tristis Faust, 1895b: 337 (Prodioctes) A: SD
190 DRYOPHTHORIDAE
genus Nassophasis C. O. Waterhouse, 1879:17 type species Nassophasis foveata C. O. Waterhouse, 1879
Hilipomorphus Desbrochers des Loges, 1891f: ccclx type species Hilipomorphus subfasciatus Desbrochers des Loges,
1891 (=» Nassophasis morreni Roelofs, 1885)
aspericollis Heller, 1941: 160 A: JAi ORR
cardoni Desbrochers des Loges, 189If: ccclx {Hilipomorphus) A: SD ORR
klapperichi Voss, 1956b: 42 A: FUJ
subverrucosa Voss, 1958: 123 A: FUJ

genus Odoiporus Chevrolat, 1885c: 288 type species Calandra longicollis Olivier, 1807
longicollis Olivier, 1807: 86 {Calandra) A: FUJ GUA GUI GUX HKG IN JA NP SD TAI UP YUN XIZ ORR
glabricollis Gyllenhal, 1838: 913 {Sphenophorus)
glabridiscus Walker, 1859b: 218 {Sphenophorus)
major Heller, 1899:33
planipennis Gyllenhal, 1838: 911 {Sphenophorus)

genus Prodioctes Pascoe, 1874: 66 type species Prodioctes quinarius Pascoe, 1874
subgenus Prodioctes Pascoe, 1874: 66 type species Prodioctes quinarius Pascoe, 1874
tristis Faust, 1895b: 337 A: SD
subgenus Paraprodioctes Voss, 1958: 121 type species Prodioctes chinensis Voss, 1958
chinensis Voss, 1958: 120 A: FUJ

genus Rhabdoscelus G. A. K. Marshall, 1943:119 [RN] type species Sphenophorus maculatus Gyllenhal, 1838
Rhabdocnemis Faust, 1893a: 150 [HN] type species Sphenophorus maculatus Gyllenhal, 1838
fissicaudus Chevrolat, 1885b: 104 {Trochorhopalon) A: JA ORR
pygidialis Faust, 1895b: 348 {Rhabdocnemis)
maculatus Gyllenhal, 1838: 881 {Sphenophorus) A: YUN ORR
carinicollis Voss, 1937a: 273 {Rhabdocnemis)
eucnemis Heller, 1899: 33 {Rhabdocnemis)
obscurus Boisduval, 1835: 448 {Calandra) A: JA JIX TAI AUR ORR
beccarii Pascoe, 1885: 301 {Sphenophorus)
insularis Boheman, 1859: 148 {Sphenophorus)
interruptecostatus L. W. Schaufuss, 1885: 204 {Sphenophorus)
maculatus Matsumura, 1915: 222, pi. 23 {Sphenophorus) [HN]
nudicollis Kirsch, 1877b: 156 (Sphenophorus)
promissus Pascoe, 1885: 300 (Sphenophorus)
tincturatus Pascoe, 1885: 301 (Sphenophorus)
similis Chevrolat, 1883: 573 (Cercidecerus) A: JAi ORR
lineaticollis Heller, 1912: 395 (Rhabdocnemis)

genus Scyphophorus Schoenherr, 1838: 855 type species Scyphophorus acupunctatus Gyllenhal, 1838
acupunctatus Gyllenhal, 1838 Ei: IT SP NAR NTR
anthracinus Gyllenhal, 1838: 858
asperulus LeConte, 1857: 58 (Rhynchophorus)
interstitialis Gyllenhal, 1838: 856
robustior Horn, 1873:409

genus Sipalomimus Voss, 1958: 125 type species Sipalomimus singularis Voss, 1958
singularis Voss, 1958: 126 A: FUJ TAI

genus Sphenocorynes Schoenherr, 1838: 866 type species Curculio quadripunctatus Weber, 1801 (=
Rhynchophorus cinereus Illiger, 1800)
cinereus Illiger, 1800: 115 (Rhynchophorus) A: TAI ORR
quadripunctatus Weber, 1801: 93 (Curculio) [HN]
grandis Günther, 1936b: 107 A: MAC
kosempoensis Janczyk, 1964: 435 (Sphenocorynus) A: TAI
ocellatus Pascoe, 1887a: 376 (Sphenocorynus) A: JA TAI
perelegans Fairmaire, 1898a: 15 (Sphenocorynus)
 DRYOPHTHORIDAE 191

quattuordecimmaculatus Hope, 1831: 27 (Calandra) A: NP SD ORR

rubricosus Faust, 1895b: 335 (Prodioctes) A: SD ORR

genus Sphenophorus Schoenherr, 1838: 874 type species Curciilio abbreviates Fabricius, 1787
Calandra Clairville, 1798: pi. 2 [supressed] type species Curculio abbreviatus Fabricius, 1787
Merothricus Chevrolat, 1885c: 290 type species Sphenophorus rusticus Gyllenhal, 1838
Nesorthognathus Voss, 1943d: 234 type species Nesorthognathus pedestris Voss, 1943 (= Sphenophorus crassus
Blanchard, 1847)
Sitonobia Gistel, 1856: 369 type species Curculio decurtatus Gmelin, 1790 (= Curculio abbreviatus Fabricius, 1787)
Trichischius LeConte, 1876: 426 type species Trichischius crenatus LeConte, 1876
abbreviatus Fabricius, 1787: 99 (Curculio) E: AL AU BU CR CZ FR GG GR HU IT MA PT RO SP ST SZ UK
YU "Caucasus" A: CY KZ TM WS
brachypteros Gmelin, 1790: 1741 (Curculio)
decurtatus Gmelin, 1790: 1747 (Curculio)
denominates ChevroM, 1885b: 106 [RN}
elegans Geoffroy, 1785: 118 (Curculio)
inaequalis Allard, 1870b: 208 [HN]
paludicola Waltl, 1835: 81 (Calandra)
porculus Fabricius, 1798: 166 (Curculio)
ragusae Stierlin, 1882b: 399
scotina Germar, 1824: 298 (Calandra)
kuatunensis Voss, 1958: 122 A: FUJ
kuznetzovi Egorov, 1979b: 153 (Calandra) A: FE
meridionalis Gyllenhal, 1838: 934 E: AL FR GR IT SP N: AG
grandini Marseul, 1870: lvii [RN]
pumilus Allard, 1870b: 210 [HN]
sanguinipennis Chevrolat, 1885b: 107
uniseriatus Stierlin, 1882b: 401
parumpunctatus Gyllenhal, 1838: 931 E: AU BU FR GR IT PöT SP N: AG A: CY
opacus Gyllenhal, 1838: 932
siculus Stierlin, 1876: 476
piceus Pallas, 1771: 464 (Curculio) E: BU CR FR GE GR HU IT MA NL PL PT RO SP ST SZ TR UK YU
"Transcaucasus" N: AG A: CY WS "Turkestan"
alaiensis Faust, 1887j : 318
eliconensis Vitale, 1906: 133 (Calandra)
nitens Allard, 1870b: 207
opacus Stierlin, 1882b: 399 [HN]
rubidus Rey, 1895b: 50
striatopunctatus Reitter, 1883: 234 [HN]
striatopunctatus Goeze, 1777: 410 (Curculio) E: AU BU BY FR GE HU IT NL PT RO SP SZ UK YU A: CY
RU
abbreviatus Herbst, 1795: 11 (Rhynchophorus)
ardesia Olivier, 1807: 92 (Calandra)
elegans Geoffroy, 1785: 118 (Curculio)
fimbriatus Gmelin, 1790: 1804 (Curculio)
helveticus Stierlin, 1882b: 400
mutilatus Laicharting, 1781: 216 (Curculio)
venants venatus Say, 1831: 22 (Rhynchophorus) Ni: CI NARi (Hawaii)
confusus Gyllenhal, 1838: 944
fallax Boheman, 1845: 256
immunis Say, 1831:23 (Rhynchophorus)
placidus Say, 1831: 23 (Rhynchophorus)
venatus vestitus Chittenden, 1904: 134 Ei: FR SP Ni: MO Ai: ES JA QA TAI NAR

genus Tetratopos Chevrolat, 1883: 569 type species Calandra sericans Wiedemann, 1823
Tetratopus Csiki, 1936: 29 [unjustified emendation]
longicollis Faust, 1895b: 330 A: SD
192 ERIRHINIDAE
genus Trochorhopalus Kirsch, 1877b: 156 type species Sphenophorus strangulates Gyllenhal, 1838
Trochorrhopalus Rye, 1879: 74 [unnecessary emendation]
Trochorrhopalus Champion, 1914: 492 [HN, unnecessary emendation]
humeralis Chevrolat, 1885b: 103 {Trochorhopalus) A: YUN ORR

genus Trymatoderus Fair maire, 1889: 55 type species Trymatoderus spongiicollis Fairmaire, 1889
carinicollis Günther, 1936a: 76 A: JIX ORR
spongiicollis Fairmaire, 1889: 55 A: CH

subfamily Stromboscerinae Lacordaire, 1865

genus Allaeotes Pascoe, 1885: 312 type species Allaeotes griseus Pascoe, 1885
niger He, Zhang & Pelsue, 2003b: 127 A: FUJ GUA JIA ZHE

genus Dexipeus Pascoe, 1885: 311 type species Dexipeus fiimosus Pascoe, 1885
uenoi Morimoto, 1978b: 105 A: JA

genus Dryophthoroides Roelofs, 1879b: liv type species Dryophthoroides sulcatus Roelofs, 1879
Elatticus Pascoe, 1885: 310 type species Elatticus beccarii Pascoe, 1885
Pembertonia Zimmerman, 1944: 202 type species Pembertonia seftoni Zimmerman, 1944
parvungulis G. A. K. Marshall, 1931: 270 A: SD
sulcatus Roelofs, 1879b: liv A: FUJ JA

genus Nephius Pascoe, 1885: 308 type species Nephius gestroi Pascoe, 1885
Anius Pascoe, 1885: 312 type species Aniuspauperatus Pascoe, 1885
salebrosus Voss, 1958: 129 (Anius) A: FUJ

genus Orthosinus Motschulsky, 1863: 525 type species Orthosinus velatus Motchsulky, 1863
foveatus Voss, 1953: 81 A: FUJ
japonicus Morimoto & Miyakawa, 1985: 82 A: JA
subulirostris G. A. K. Marshall, 1931: 271 A: SD

genus Synommatoides Morimoto, 1978b: 104 type species Synommatoides shirouzi Morimoto, 1978
scutellatus He, Zhang & Pelsue, 2003a: 123 A: ZHE
shirozui Morimoto, 1978b: 104 A: JA SC

genus Synommatus Wollaston, 1873b: 444 type species Synommatus confluens Wollaston, 1873
interrupts Pascoe, 1885: 311 A: FUJ JA ORR

genus Tetrasynommatus Morimoto, 1985: 73 type species Tetrasynommatus thompsoni Morimoto, 1985
thompsoni Morimoto, 1985: 75 A: J A

genus Xerodermia Lacordaire, 1865: 307 type species Xerodermus porcellus Lacordaire, 1865 (= Orthosinus
sculpticollis Motschulsky, 1863)
himalayanus G. A. K. Marshall, 1931: 272 A: UP

family Dryophthoridae, species incertae sedis

Calandra quadrimaculata Hope, 1931: 27 A: NP
Calandra goliathus Hope, 1931:27 A: NP

family ERIRHINIDAE Schoenherr, 1825

subfamily Erirhininae Schoenherr, 1825
tribe Arthrostenini Reitter, 1913
genus Afghanocryptus Voss, 1961c: 247 type species Afghanocryptus amseli Voss, 1961
amseli Voss, 1961c: 247 A: AF TM
 ERIRHINIDAE 193
genus Aralkumia Bajtenov, 1974a: 234 type species Aralkumia deserti Bajtenov, 1974
deserti Bajtenov, 1974a: 234 A: KZ

genus Arthrostenus Schoenherr, 1826: 287 type species Arthrostenus fullo Boheman, 1836
Graphicotera Tournier, 1873: 183 type species Graphicotera excelsa Toumier, 1873
adanensis Pic, 1914: 51 A: TR
alternons Kirsch, 1871b: 393 N: EG
cinereus Boheman, 1836: 535 E: "Caucasus"
excelsus Tournier, 1873: 184 (Graphicotera)
fullo Boheman, 1836: 534 E: AR GR RU UK A: SY TM
lineatus Pic, 1940: 5
far Faust, 1885c: 179 A: UZ
ignoratus Faust, 1884: 453 E: AR RU A: TD
rotroui Pic, 1940: 5 N: AG A: TR
argentatus Pic, 1940: 5
spadiceus Boheman, 1836: 534 E: "Caucasus"
syriacus Pic, 1907b: 138 A: SY

genus Thecorhinus Faust, 1887e: 175 type species Thecorhinus modestus Faust, 1887
modestus Faust, 1887e: 176 A: KZ TM UZ

genus Trichocaulus Fairmaire, 1877: 98 type species Trichocaulus longipilis Fairmaire, 1877
subgenus Trichocaulus Fairmaire, 1877
brevipilis Pelletier & Borovec, 2003: 89 N: AG TU
longipilis Fairmaire, 1877: 98 N: AG

genus Turanocryptus Korotyaev, 1988b: 139 type species Turanocryptus belovi Korotyaev, 1987
belovi Korotyaev, 1988b: 139 A: TM

tribe Erirhinini Schoenherr, 1825

genus Absoloniella Formanek, 1913: 135 [DA] type species Absoloniella cylindrica Formanek, 1913
cylindrica Formanek, 1913: 135 E: BH
reitteriJ. Müller, 1912: 303 (Caulomorphus) E: IT

genus Apachiscelus Desbrochers des Loges, 1875a: 18 type species Apachiscelus pellitus Desbrochers des
Loges, 1875
lethierryi Chevrolat, 1860a: 77 (Procas) N: AG
pellitus Desbrochers des Loges, 1875a: 18
rufescens Pic, 1905c: 131 N: AG

genus Bagoopsis Faust, 1881: 317 type species Bagoopsis volgensis Faust, 1881
globicollis Fairmaire, 1863: 112 (Erirrhinus) E: BU CZ GR HU IT MC ME RO SB SK TR
confiisa Faust, 1887f: 187 (Echinocnemus)
dalmatina Stierlin, 1892: 365 (Notaris)
pugnax Faust, 1881: 321
volgensis Faust, 1881: 319 E: MD RU A: KZ

genus Caenosilapillus Chûjô & Morimoto, 1959b: 147 type species Caenosilapillus babai Chûjô & Morimoto,
1959
babai Chûjô & Morimoto, 1959b: 148 A: JA

genus Echinocnemus Schoenherr, 1843: 315 type species Erirhinus squameus Boheman, 1835
Colchis Toumier, 1874a: 73 type species Colchis tibialis Toumier, 1874
Lostlanus Desbrochers des Loges, 1900a: 9 type species Lostianus sardous Desbrochers des Loges, 1900
194 ERIRHINIDAE
aegyptiacus Desbrochers des Loges, 1896e: 103 (Bagous) N: EG
carinirostris Tournier, 1874a: 75 (Colchis) E: GG
depressipennis Tournier, 1874b: 459 (Jekelia) N: AG
dorsalis Chevrolat, 1879: 117 A: XIZ
margelanicus Faust, 1887f: 190 A: AF PA TD TM UZ
phytonomoides Voss, 1953: 80 (Hydronomus) A: JA NE SE TAI
reiften Schilsky, 1907: 32 N: EG
sardous Desbrochers des Loges, 1900a: 10 (Lostianus) E: IT (Sardegna)
squameus Boheman, 1835: 311 (Erirhinus) A: CH ES FE JA NC SC ORR
bipunctatus Roelofs, 1874: 123
tibialis Tournier, 1874a: 74 (Colchis) N: EG
truncatus Chevrolat, 1879: 117 A: NO

genus Grypus Germar, 1817c: 340 type species Curculio equiseti Fabricius, 1775
Aplopus Dejean, 1821: 86 type species Curculio equiseti Fabricius, 1775
Grypidius Schoenherr, 1826:231 [RN] type species Curculio equiseti Fabricius, 1775
brunnirostris Fabricius, 1792: 404 (Curculio) E: AU BE BU BY CZ DE EN FR GE HU IT LA LT NL PL SK
SVSZ NAR
oblitérants Herbst, 1795: 273 (Curculio)
equiseti Fabricius, 1775: 130 (Curculio) E: AL AU BE BH BU BY CR CZ DE EN FI FR GE GB HU IR IT LA
LT ME NL NR PL RO RU SB SK SL SV SZ UK A: ES FE KZ WS NAR
atrirostris Fabricius, 1781: 499 (Curculio)
gibbosus Gmelin, 1790: 1756 (Curculio) [unjustified RN]
insquamipennis Delahon, 1914: 622
nigrogibbosus Degeer, 1775: 224 (Curculio)
kaschmirensis Voss, 1960a: I I A : KA
mannerheimi Faust, 1881: 312 A: ES FE JA
rugicollis Voss, 1934: 81 A: SCH

genus Hypoglyptus Gerstaecker, 1855:172 [DA] type species Hypoglyptuspictus Gerstaecker, 1855
Acentroides Leonhard, 1912:341 type species Acentroides conspersus Leonhard, 1912
conspersus Leonhard, 1912: 341 (Acentroides) E: GR
elegans Brülle, 1832: 245 (Tychius) E: GR
gracilis Kiesenwetter, 1864: 274 E: GR
graecus Pic, 1902: 67 (Hylobius) E: GR
heydeni Faust, 1889c: 226 E: AR MC A: TR
pictus Gerstaecker, 1855: 173 E: GR (Corfu)

genus leans Tournier, 1874a: 93 type species Erirhinus sparganii Gyllenhal, 1835
Prionochelus Desbrochers des Loges, 1875a: 21 type species Prionochelus squamosus Desbrochers des Loges, 1875 (=
Erirhinus sparganii cinereus Miller, 1861 )
sparganii cinereus Miller, 1861: 209 (Erirhinus) A: TR S Y
squamosus Desbrochers des Loges, 1875a: 21 (Prionochelus)
sparganiipertinax Gyllenhal, 1835: 309 (Erirhirms) E: RU A: HEI KI KZ TD TM UZ WS
subaureus Reitter, 1900: 58 (Echinocnemus)
sparganii sparganii Gyllenhal, 1835: 310 (Erirhinus) E: AU BH BU BY CR HU IT RU SK UK A: FE IN MG
NCSW
efferus Faust, 1887f: 188
pilipes Desbrochers des Loges, 1896e: 102 (Bagous)

genus Jekelia Tournier, 1874b: 459 type species Tychius ephippiatus Fairmaire, 1871 (= Aubeonymus notatus
Mulsant & Godart, 1868)
Prolobodontus Desbrochers des Loges, 1875a: 19 type species Prolobodontus ahenus Desbrochers des Loges, 1875 (=
Aubeonymus notatus Mulsant & Godart, 1868)
notata Mulsant & Godart, 1868b: 412 (Aubeonymus) E: IT A: MO AG TU
ahena Desbrochers des Loges, 1875a: 20 (Prolobodontus)
 ERIRHINIDAE 195
ephippiata Fairmaire, 1871:401 (Tychius)

genus Lepidonotaris Zumpt, 1929: 217 type species Dorytomuspetax C. R. Sahlberg, 1823
petax C. R. Sahlberg, 1823:41 (Dorytomus) E: BU MD ST A: FE HEI KZ MG WS

genus Neoicaris A. Hoffmann, 1968: 150 type species Neoicaris descampsi A. Hoffmann, 1968
descampsiA. Hoffmann, 1968: 151 A: IN

genus Notaris Germar, 1817c: 340 type species Curculio acridulus Linnaeus, 1758
Erirhinus Schoenherr, 1825: 582 type species Curculio aethiops Fabricius, 1792
Erycus Tournier, 1874a: 93 type species Curculio scirpi Fabricius, 1792
acridulus acridulus Linnaeus, 1758: 378 (Curculio) E: AU BE BH BU BY CR CZ DE EN FI FR GB GE HU
IC IR IT LA MD NL NR PL RO RU SB SK SL SP SV SZ UK A: ES FE JA KI KZ TD UZ WS
anularis Faust, 1882b: 161
merkli Stierlin, 1882a: 252 (Erirhinus)
montana Faust, 1882b: 161
punctum Fabricius, 1792: 401 (Curculio)
rigida Marsham, 1802: 270 (Curculio)
aethiops Fabricius, 1792: 405 (Curculio) E: AU BH BU BY CR CZ DE EN FI FR GB GE HU IC IR IT LA LT
NR PL RO RU SB SK SL SU SZ UK A: ES FE MG WS NAR
holomelana Herbst, 1795: 271 (Curculio)
lapponica Faust, 1882b: 165
morio Mannerheim, 1853: 240 (Erirhinus)
rufipes Motschulsky, 1845b: 377
aterrima Hampe, 1850: 356 (Phloeophagus) E: AU BH BU BY CR CZ FR GE HU IT PL RO RU SB SK SL
SZUK
brancsiki Tournier, 1874a: 95
gerhardti Letzner, 1872: 8
daurica Faust, 1882b: 155 A: ES
discreta Faust, 1882b: 167 A: KI KZ TD UZ
disions Faust, 1890e: 470 A: ES XIZ
eversmanni Faust, 1882b: 169 A: ES FE
illibata Faust, 1882b: 163 A: ES FE
imprudens Faust, 1885c: 170 A: KI KZ TD UZ
maerkeli Boheman, 1843: 164 (Erirhinus) E: AU BY CZ EN HU IT LT PL RO RU SK SL SZ
mandschurica Voss, 1940: 51 A: ES FE HEI
lederi Faust, 1882b: 174 E: ST "Caucasus"
nivalis Faust, 1882b: 167 A: KI KZ
oberti altaica Legalov, 1997: 471 (Erirhinus) A: KZ WS
oberti oberti Faust, 1885c: 171 A: TD "Pamir"
oryzae Ishida, 1902: 90 (Erirhinus) A: ES FE JA NC NE SC SE SW
Jùnebris Kôno, 1930: 158
scirpi Fabricius, 1792: 405 (Curculio) [NP] E: AL AU BE BH BU BY CR CZ.DE EN FI FR GB GE GR HU IR
IT LA NL NR PL RO RU SB SK SL SP SV SZ UK A: ES FE JA KZ WS
rhamni Herbst, 1784a: 78 (Curculio) [NO]
rotundicollis Motschulsky, 1860: 157
scirpi Rossi, 1790: 118 (Curculio) [NO]
genus Notaroides Korotyaev, 1979: 166 type species Notaroides brevirostris Korotyaev, 1979
brevirostris Korotyaev, 1979: 167 A: XIN

genus Notodermus Desbrochers des Loges, 1875a: 19 type species Notodermus bruleriei Desbrochers des
Loges, 1875
Pseudypera Voss, 1936a: 57 type species Pseudypera subtessellata Voss, 1936
bruleriei Desbrochers des Loges, 1875a: 19 A: SY
macula lus Pic, 1915a: 1 (Procas) A: LE
196 ERIRHINIDAE
saulcyi Reiche & Saulcy, 1858: 677 (Procas) A: CY IN
steinen Voss, 1936a: 59 (JPseudypera) A: TR
subtessellatus Voss, 1936a: 58 (Pseudypera) A: TR

genus Ochodontus Desbrochers des Loges, 1897b: 18 [EA] type species Ochodontus rubrirostris Desbrochers
des Loges, 1897
rubrirostris Desbrochers des Loges, 1897b: 19 E: GG

genus Picia Tournier, 1895: cclxxx [RN] type species Faustia ephimeropoides Tournier, 1889 (= Hydronomus
sinuatocollis Faust, 1885)
Faustia Tournier, 1889: 150 [HN] type species Faustia ephimeropoides Tournier, 1889 (= Hydronomus sinuatocollis
Faust, 1885)
mesopotamica Tournier, 1889: 152 (Faustia) E: AB GG A: IN IQ SA TM TR
caucasica Tournier, 1889: 152 (Faustia)
distinctipennis Pic, 1901c: 58
flaveola Reitter, 1888: 125 (Ephimeropus)
millingi Tournier, 1889: 153 (Faustia)
sinuatocollis Faust, 1885c: 178 (Hydronomus) E: AB AR GG RU N: EG A: AF IN IQ KZ TD TM TR UZ
"Punjab" AFR
alfierii Pic, 1914:49
drurei Pic, 1917: 10
ephimeropoides Tournier, 1889: 153 (Faustia)
fenestrata Reitter, 1888: 125 (Ephimeropus)
punjabensis Pajni & Kamal Tewari, 1985: 610 (Hydronomidius)
syriaca Reitter, 1888: 125 (Ephimeropus) E: GR A: CY IS TR SY

genus Picianus Zumpt, 1929: 217 type species Echinocnemus sieversi Faust, 1884
sieversi Faust, 1884: 452 (Echinocnemus) E: AR GG A: KZ SY TD TM UZ
squamulosus Pic, 1907b: 138 (Erirhinus)

genus Procas Stephens, 1831: 90 type species Curculio picipes Marsham, 1802
armillatus Fabricius, 1801: 444 (Rhynchaenus) E: AZ IT FR GR MA MR SP N: AG CI LB MO TU
atomarius Gene, 1839: 37 (Erirhinus)
cottyi Perris, 1864:298
biguttatus Faust, 1882a: 275 A: ES FE JA NC NE SC
okunii Kôno, 1930: 159
granulicollis Walton, 1848: 166 E: GB SP
michaelis Thompson, 2006: 24 E: SP
picipes levantinus Thompson, 2006: 23 A: CY IS JO TR
picipes picipes Marsham, 1802: 272 (Curculio) E: BE FR GB GE IT NL NR PL SP
minutus Desbrochers des Loges, 1893b: 9
moestus Bach, 1854: 296 (Erirhinus)
picipes steveni Krynicki, 1832: 147 (Rhynchaenus) E: AB BU MD RU UK A: TR
steveni Gyllenhal, 1835: 287 (Erirhinus)
genus Procas, species incertae sedis
rasus Desbrochers des Loges, 1897b: 2 E: SP
Sibiriern Pic, 1904a: 27 A: ES WS

genus Ruffodytes Osella, 1973b: 491 [DA] type species Ruffodytes hellenicus Osella 1973
hellenieus Osella, 1973b: 492 E: GR
italicus Osella, 1976: 200 E: IT
nitidipennis Osella, 1989: 451 E: GR
pacei Osella, 1976: 197 E: IT
servadeii Osella, 1982: 273 E: IT
 ERIRHINIDAE 197
genus Siraton Hustache, 1934: 250 [DA] type species Siraton devillei Hustache, 1934
devillei Hustache, 1934: 250 E: IT (Elba)

genus Theanellus Reitter, 1913a: 77 type species Theanellus bagoides Reitter, 1913
Syrdariella Ter-Minasian, 1977: 68 type species Syrdariella grisea Ter-Minasian, 1977
alepensis Pic, 1915a: 1 (Procas) A: S Y
alternons Faust, 1885c: 173 (Apachiscelus) A: UZ
antoinei Hustache, 1932a: 48 (Procas) N: MO
bagoides Reitter, 1913a: 77 A: AF KZ TD TM UZ
fastidiosus Pic, 1904c: 93 (Procas) N: TU
griseus Ter-Minasian, 1977:68 (Syrdariella) A: UZ
kocheriA. Hoffmann, 1953a: 140 N: MO
plumipilis Hustache, 1931b: 79 A: SY
indentatus Pic, 1934: 17
putoni Tournier, 1875b: ccix (Procas) N: AG
testaceus Bajtenov, 1974a: 37 (Procas) A: KZ

genus Thryogenes Bedel, 1884a: 108 type species Curculio festucae Herbst, 1795
Eurhinus Stephens, 1829: 14 [HN] type species Curculio festucae Herbst, 1795
festucae Herbst, 1795: 327 (Curculio) E: AL AU BE BH BU BY CR CZ DE EN FI FR GB GE GR HU IR IT
LA LT MD ME NL NR PL RO RU SB SK SL SP SV SZ UK A: ES FE KI KZ TD TM UZ WS
carecti Hoppe, 1795: 59 (Curculio)
caricis Thunberg, 1798: 111 (Curculio)
inquisitor Herbst, 1797: 42 (Curculio)
florii Zumpt, 1928: 277 E: DE FR GB GE HU IT PL
atrirostris Lohse, 1992: 370
nereis Paykull, 1800: 240 (Curculio) E: AU BY CZ DE EN FI FR GB GE HU IR IT LA LT NR PL SK SV SZ
UK A: ES FE MG WS
arundineti Stephens, 1831: 80 (Erirhinus)
palustris Gyllenhal, 1835: 313 (Erirhinus)
typhae Ahrens, 1812: 15 (Rhynchaenus)
scirrhosus Gyllenhal, 1835: 312 (Erirhinus) E: AU BE BH BU BY CZ DE EN FI FR GB GE HU IT LA NL
NO PL RO RU SB SK SL SV UK
ussurensis Egorov, 1979a: 87 A: FE MG SW

genus Tournotaris Alonso-Zarazaga & Lyal, 1999: 70 type species Curculio bimaculatus Fabricius, 1787
bimaculata Fabricius, 1787: 98 (Curculio) E: AL AU BE BH BU BY CR CZ DE EN FI FR GB GE GR HU IR
IT LA MD ME NL NO PL RO RU SB SK SL SV SZ UK A: ES FE JA KI KZ NE TD TM UZ WS NAR
frivaldszkyi Tournier, 1874a: 94 (Notaris)
salaria O. Schneider, 1898: 75 (Notaris)
subcostata Motschulsky, 1845: 378 (Erirrhinus)
wyomingensis Chittenden, 1906:115 (Notaris)
granulipennis Tournier, 1874a: 94 (Notaris) E: HU PL RO SK TR UK
ochotica Korotyaev, 1984: 387 (Notaris) A: FE MG

genus Tychiophaenus Voss, 1960b: 128 [RN] type species Tychiosoma bodemeyeri Voss, 1943
Tychiosoma Voss, 1943c: 227 [HN] type species Tychiosoma bodemeyeri Voss, 1943
bodemeyeri Voss, 1943c: 228 (Tychiosoma) A: KZ

genus Tyrodomus Voss, 1971a: 3 type species Tyrodomus patruelis Voss, 1971
patruelis Voss, 1971a: 4 A: IN

genus Xeronoma lablokoff-Khnzorian, 1964: 173 type species Xeronoma latirostris Iablokoff-Khnzorian,
1964
latirostris Iablokoff-Khnzorian, 1964: 174 E: AR
198 RAYMONDIONYMIDAE

tribe Himasthlophallini Zherikhin, 1991

genus Himasthlophallus Egorov & Zherikhin, 1991: 32 type species Himasthlophallus flagellifer Egorov &
Zherikhin, 1991
flagellifer Egorov & Zherikhin, 1991: 33 A: FE

tribe Stenopelmini LeConte, 1876

genus Lissorhoptrus LeConte, 1876: 183 type species Bagous simplex Say, 1831
subgenus Lissorhoptrus LeConte, 1876 type species Bagous simplex Say, 1831
oryzophilus Kuschel, 1952: 46 Ei: IT Ai: CH FE JA NC SC NAR NTRi
pseudoryzophilus Guan, Huang & Lu, 1995: 212

genus Stenopelmus Schoenherr, 1835: 468 type species Stenopelmus rufinasus Gyllenhal, 1835
Degorsia Bedel, 1902: 359 type species Degorsia champenoisi Bedel, 1902 (= Stenopelmus rufinasus Gyllenhal, 1835)
Monius Schoenherr, 1845: 501 [RN] type species Stenopelmus rufinasus Gyllenhal, 1835
Panscopus Schoenherr, 1843: 351 [RN, HN] type species Stenopelmus rufinasus Gyllenhal, 1835
rufinasus Gyllenhal, 1835: 469 Ei: BE FR GB GE IR IT NL SP UK Ai: JA AFRi NAR
champenoisi Bedel, 1902: 359 {Degorsia)

tribe Tanysphyrini Gistel, 1848

genus Brachygyius G. A. K. Marshall, 1939: 582 [RN] type species Brachypus lixoides Schoenherr, 1826
Brachyppa E. Strand, 1943: 96 [RN] type species Brachypus lixoides Schoenherr, 1826
Brachypus Schoenherr, 1826: 217 [HN] type species Brachypus lixoides Schoenherr, 1826
cylindricus Hochhuth, 1847: 536 {Brachypus) E: "Caucasus"
lixoides Gyllenhal, 1836: 103 (Brachypus) A: "Asia"
lixoides Gyllenhal, 1835: 103 {Brachypus) [HN]

genus Glaridorhinus Kolenati, 1858: 343 type species Glaridorhinus khuenburgii Kolenati, 1858
khuenburgii Kolenati, 1858: 343 E:AU

genus Tanysphyrus Germar, 1817c: 340 type species Curculio lemnae Paykull, 1792
Tmysphyroides Egorov, 1996c: 431 type species Tanysphyrus ussurensis Egorov, 1996 (^Tanysphyrus brevipennis Voss,
1953)
ater Blatchley, 1928: 244 E: AU CZ FI GE PL SL SV UK NARi NTRi
makolskii Smreczynski, 1957: 123
brevipennis Voss, 1953: 79 A: FE JA NC SC SE
ussuriensis Egorov, 1996c: 433
khancaensis Egorov, 1996c: 434 A: FE JA NC SC
lemnae Paykull, 1792: 78 (Curculio) E: AU BE BU BY DE EN FI FR GB GE IR IT LA LT MD NL NR NT PL
RU SV SZ UK A: WS NARi NTRi
callae Voss, 1943a: 17
inspectatus Herbst, 1795: 301 (Curculio)
major Roelofs, 1874: 125 A: FE JA SC SE NC ORR
family Erirhinidae, taxa incertae sedis
Erirhinus indistinctusMotschulsky, 1845a: 101 E : L A N T

genus Tsherepanovia Korotyaev, 1992b: 888 [EA] type species Tsherepanovia globosa Korotyaev 1992
globosa Korotyaev, 1992b: 889 A: TM

family RAYMONDIONYMIDAE Reitter, 1913

genus Alaocephala Ganglbauer, 1906: 141 type species Raymondia delarouzei C N F Brisout de Barneville,
1862
delarouzei catalonica Osella, 1977: 23 E: SP (Barcelona, Gerona)
delarouzei coiffaiti A. Hoffmann, 1958: 1749 E: FR (Pyrénées-Orientales)
 RAYMONDIONYMIDAE 199
delarouzei delarouzei C. N. F. Brisout de Barneville, 1862: 603 {Raymondia) E: FR (Pyrénées-Orientales)

genus Alaocyba Perris, 1869: 31 type species Alaocyba carinulata Perris, 1869
carinulata Perris, 1869: 31 E: FR (Corse) IT (Sardegna)
coniceps Normand, 1914: 303 N: TU
elongatula Normand, 1914: 302 N: TU
lampedusae Dodero, 1916: 352 E: IT (Lampedusa Is.)
marcuzzii Osella, 1977: 15 E: IT(Abruzzo, Basilicata)
melitensis Magnano & Mifsud, 1998: 176 E: MA
separanda Dodero, 1916: 352 E: IT (Pantelleria Is.)
theryi Meyer, 1897: 207 N: AG

genus Coiffaitiella Osella, 1977:30 type species Raymondia benjamini Marquet, 1875
benjamini aurouxi Gonzalez, 1965: 100 (Raymondionymus) E: SP (Pirineos or.)
benjamini benjamini Marquet, 1875: 28 (Raymondia) E: FR (Pyrénées-Orientales) SP (Pirineos or.)
carinirostris Osella, 1971: 80 {Raymondionymus) E: SP (Pyrénées-Orientales)
hispanica Osella, 1971: 77 (Raymondionymus) E: FR (Pyrénées-Orientales) SP (Pirineos or.)
lusitanica Osella, 1971: 75 (Raymondionymus) E: PT (Monchique)
pyrenaica Osella, 1971: 82 (Raymondionymus) E: SP (Pirineos or.)
remilleti Osella, 1971: 73 (Raymondionymus) E: PT SP (Cadiz)

genus Derosasius Ganglbauer, 1996: 143 type species Raymondia damryi Perris, 1874
damryi aritzensis Osella, 1977: 90 E: TT (Sardegna)
damryi damryi Perris, 1874: 12 (Raymondia) E: FR (Corse)
damryi sardous Osella, 1977: 91 E: IT (Sardegna)

genus Ferreria Alonso-Zarazaga & Lyal, 1999: 72 type species Raymondia marqueti Aube, 1863
doriai Osella, 1977: 77 (Raymondionymus) E: IT (Liguria)
marqueti appenina Dieck, 1870: 354 (Raymondia) E: IT (Toscana)
hispalensis Desbrochers des Loges, 1902a: 106 (Raymondia)
vallebrosae Desbrochers des Loges, 1902a: 106 (Raymondia)
marqueti marqueti Aube, 1863: 130 (Raymondia) E: FR GBi SZ
curvinasus Abeille de Perrin, 1872: xliv (Raymondia)
taurinorum Osella, 1977: 81 (Raymondionymus)
marqueti mauritanica Normand, 1939b: 514 (Raymondionymus) N: TU
marqueti veneta Osella, 1977: 79 (Raymondionymus) E: CR (Istria, Dalmazia) IT (Veneto)

genus Raymondiellus Ganglbauer, 1906:159 type species Raymondia sicula Rottenberg, 1872
casalei Meregalli, 1983: 131 N: AG (Petite Kabylie)
doderoi Ganglbauer, 1906: 165 (Raymondionymus) E: IT (Sardegna)
hipponensis Peyerimhoff, 1919a: 73 (Raymondionymus) N: AG
inopinatus Osella, 1977: 120 E: IT(Abruzzo, Molise)
kabylianus Pic, 1896b: 114 (Raymondia) N: AG
auripes Chobaut, 1909: 182 (Raymondia)
lagrecai Osella, 1977: 97 E: IT(Sicilia)
latialis Osella, 1977: 112 E: IT (Lazio)
pacei Osella, 1977: 117 E: IT (Calabria, Puglia)
sbordonii Osella, 1977: 114 E: IT (Ponza Is., Zannone Is.)
sardous insularis Osella, 1977: 104 E: IT (Sardegna)
sardous sardous Perris, 1869: 30 E: IT (Sardegna)
sericatus Normand, 1937: 255 (Raymondionymus) N: AG
siculus Rottenberg, 1872: 240 (Raymondia) E: IT (Sicilia)
solarii Ganglbauer, 1906: 162 (Raymondionymus) E: IT (Sardegna)
theresae sparsepunctatus Normand, 1937: 254 (Raymondionymus) N: AG
theresae theresae Pic, 1900d: 90 (Alaocyba) N: TU
200 RAYMONDIONYMIDAE
vignai Osella, 1977: 109 E: IT (Sardegna)
genus Raymondionymus Wollaston, 1873b: 531 [RN] type species Raymondia fossor Aube, 1861
Raymondia Aube, 1861: 195 [HN] type species Raymondia fossor Aube, 1861
Pararaymondionymus Osella, 1977: 40 type species Alaocyba stricticollis Reitter, 1894
andreinii Osella, 1977: 46 {Pararaymondionymus) E: IT (Toscana)
bartolii Osella, 1977: 49 {Pararaymondionymus) E: IT (Prealpi Liguri)
burlei Secq & Secq, 1996: 16 {Pararaymondionymus) E: FR (Ariège)
curtii Meregalli, Osella & Zuppa, 2006: 105 E: IT (Piemonte)
eximius Meregalli & Osella, 2006: 38 E: IT (Latium)
fossor hübe, 1861: 195 {Raymondia) E: FR(Var)
gardinii Osella & Giusto, 1983: 430 {Pararaymondionymus) E: IT (Alpi Liguri)
hoffmanni Hervé, 1949: 133 E: FR (Var)
laevithorax Perris, 1876: 11 {Raymondia) E: FR (Corse)
laneyriei Hervé, 1949: 137 E: FR (Var)
lavagnei Mayet, 1898: 87 E: FR (Hérault)
longicollis Perris, 1869: 29 E: FR (Corse)
magnificus Osella, 1977: 52 {Pararaymondionymus) E: IT (Campania)
meggiolaroi Osella, 1977: 50 {Pararaymondionymus) E: IT (Alpi Liguri, Alpi Piemontesi)
mingazzinii Osella & Abbazzi, 1985: 475 {Pararaymondionymus) E: IT (Emilia-Romagna)
mirabilis Osella, 1977: 45 {Pararaymondionymus) E: IT (Calabria)
ochsi Hervé, 1949: 136 E: FR (Alpes Maritimes)
orientalis Hervé, 1953: 133 {Raymondionymus) E: FR (Alpes-Maritimes)
perrisi Grenier, 1864: 137 {Raymondia) E: FR (Lot-et-Garonne, Haute-Garonne)
aubei Marquet, 1864: 372 {Raymondia)
problematicus Hervé, 1949: 133 E: FR (Var) IT (Liguria)
pulcherrimus Magrini, Bastianini & Abbazzi, 2009: 340 E: IT (Lazio)
sanfilippoi Osella & Giusto, 1983: 432 {Pararaymondionymus) E: FR (Alpes-Maritimes) IT (Alpi Liguri, Alpi
Piemontesi)
stricticollispicenus Osella, 1977: 41 {Pararaymondionymus) E: IT (Marche)
stricticollis stricticollis Reitter, 1894d: 306 {Alaocyba) E: IT (Umbria)
vincenti Secq & Secq, 1996: 16 {Pararaymondionymus) E: FR (Haute-Garonne, Ariège)
zoiai Osella & Giusto, 1983: 434 {Pararaymondionymus) E: IT (Alpi Piemontesi)
genus Tarattostichus Ganglbauer, 1906:166 type species Alaocyba {Raymondia) stussineri Reitter, 1891
irregularis Kodric, 1939: 361 {Raymondionymus) E: CR IT (Friuli-Venezia Giulia) SL
stussineri Reitter, 189Id: 260 {Alaocyba) E: CR (Istria) IT (Friuli-Venezia Giulia) SL

genus Ubychia Rost, 1893: 343 type species Ubychia stygia Rost, 1893
elipsoidalis Osella & Nonveiller, 1982: 23 E: SB
ganglbaueri Reitter, 1912b: 318 {Raymondionymus) E: CR (Dalmacia) SB
holdhausi Ganglbaueri, 1903: 184 {Alaocyba) E: BH SB
icari Osella, 1980: 816 E: GR (Ikaria)
leonhardi leonhardi Reitter, 1914: 82 E: IT (Prealpi di Lombardia, Veneto)
leonhardi ticinensis Osella, 1977: 140 E: SZ(Ticino)
mingrelica iranica Osella, 1986: 5 A: IN (Mäzandarän)
mingrelica mingrelica Reitter, 1894c: 246 {Alaocyba) E: GG A: IN TR
reitteri Ganglbauer, 1906: 169 {Raymondionymus) E: GR (Iperos, Kerlrira)
salpingoides Kraatz, 1881: 226 {Raymondia) E: BH CR ME
stygia Rost, 1893: 343 E: GG A: TR
 PLATYPODINAE 201
family CURCULIONIDAE Latreilie, 1802
subfamily Platypodinae Shuckard, 1840
supertribe Platypoditae Shuckard, 1840
tribe Platypodini Shuckard, 1840
genus Baiocis Browne, 1962c: 651 type species Crossotarsus pernanulus Schedl, 1935
pernanulus Schedl, 1935e: 482 (Crossotarsus) A: SD AUR ORR

genus Carchesiopygus Schedl, 1939d: 402,403 type species Crossotarsus wollastoni Chapuis, 1865
impariporus Beeson, 1937: 62 (Crossotarsus) A: SD ORR
lobacanthus Schedl, 1969b: 59 A: UP
oculatus Beeson, 1937: 62 (Crossotarsus) A: UP
psilacanthurus Beeson, 1937: 60 (Crossotarsus) A: SD

genus Crossotarsus Chapuis, 1865: 23,44 type species Platypus wallacei J. Thomson, 1857
Crossotarsinulus Schedl, 1972a: 84, 85 type species Crossotarsus sauteri Strohmeyer, 1913
adustipennis Motschulsky, 1861: 19 (Genyocerus) A: JA
bonvouloiri Chapuis, 1865: 55 A: SD ORR
cinnamomi Beeson, 1937: 97 A: SD UP ORR
concinnus Blandford, 1894c: 578 [RN] A: JA TAI
chapuisi Blandford, 1894a: 129 [HN]
coniferae Stebbing, 1906: 411 A: HP KA UP YUN ORR
emancipatus Murayama, 1934c: 138 A: JA TAI
externedentatus Fairmaire, 1850: 51 (Platypus) A: JA SD TAI UP AFR AUR ORR
formosanus Strohmeyer, 1912a: 41
heritierae Stebbing, 1906: 420 (Diapus)
nilgiricus Beeson, 1937: 73
posticus Broun, 1910: 301 (Platypus)
soundersi Chapuis, 1865: 80
submontanus Beeson, 1937: 72
usambariensis Strohmeyer, 1911b: 182
fairmairei Chapuis, 1865: 79 A: JA UP ORR
flavomaculatus Strohmeyer, 1912a: 40 A: JA TAI ORR
machili Beeson, 1937: 76 A: SD
mussooriensis Beeson, 1937: 84 A: UP
nepalensis Schedl, 1973c: 212 A: BT NP
niponicus Blandford, 1894a: 130 A: JA TAI
piceae Stebbing, 1906: 413 A: UP
piceus Chapuis, 1865: 56 A: TAI AUR ORR
saltatorinus Schedl, 1954c: 157 (Platypus) A: FUJ ORR
sauteri Strohmeyer, 1913: 164 A: TAI
rengetensis Niisima & Murayama, 1925: 208
simplex Murayama, 1925: 231 A: JA SC TAI
siva Beeson, 1937:85 A: SD
squamulatus Chapuis, 1865: 87 A: PA SD YUN ORR
fragmentus Sampson, 1912: 249
minor Sampson, 1928: 2
squamuloides Beeson, 1937: 79
sundri Beeson, 1937: 79
terminatus Chapuis, 1865:83 A: TAI UP AUR ORR
nicobaricus Beeson, 1937: 86
sedulus Schedl, 1972c: 66
venustus Chapuis, 1865: 88
venustulus Beeson, 1937: 85
«raw Schedl, 1935f: 640 A: SD ORR
wallacei J. Thomson, 1857: 343 (Platypus) A: TAI YUN AUR ORR
wilmoti Stebbing, 1911: 105 (Platypus) A: KA UP
202 PLATYPODINAE
genus Dinoplatypus Wood, 1993: 273 type species Platypus cupulatus Chapuis, 1865
cmthocephali Schedl, 1969b: (to (Platypus) A: SD
biuncus Blandford, 1896b: 194 {Platypus) A: UP ORR
calamus Blandford, 1894a: 137 (Platypus) A: GUA FUJ JA SC TAI
cavus Strohmeyer, 1913: 162 (Platypus) A: BT PA ORR
cupulifer Wichmann, 1914b: 413 (Platypus)
cupulatus Chapuis, 1865: 278 (Platypus) A: BT HAI SD AUR ORR
blumi Schedl, 1979a: 158 (Platypus)
decern Sampson, 1922: 147 (Platypus) A: FUJ NP ORR
falcatus Strohmeyer, 1908b: 73 (Platypus) A: NP UP ORR
flectus Niisima & Murayama, 1931: 197 (Platypus) A: FUJ GUA JA TAI
formosanus Niisima & Murayama, 1925: 214 (Platypus) [HN]
kusukusensis Murayama, 1956b: 13 (Platypus)
murayamaensis Schedl, 1941a: 43 (Platypus)
hamatus Blandford, 1894a: 138 (Platypus) A: JA
lepidus Chapuis, 1865: 282 (Platypus) A: TAI AFR AUR ORR
/wrt/ger Motschulsky, 1863: 510 (Platypus) A: JA TAI YUN AFR AUR ORR
caliculus Chapuis, 1865: 280 (Platypus)
schultzei Strohmeyer, 191 la: 26 (Platypus)
pseudocupulatus Schedl, 1935f: 635 (Platypus) A: "China" AUR ORR
artecavus Schedl, 194Id: 422 (Platypus)
sundaensis Schedl, 1941d: 421 (Platypus)
tenuis Murayama, 1925: 233 A: JA ORR

genus Doliopygus Browne, 1962c: 648 type species Crossotarsus bohemani Chapuis, 1865
Doliopygus Schedl, 1972a: 143 [HN] type species Crossotarsus bohemani Chapuis, 1865
crinitus Chapuis, 1865: 90 (Crossotarsus) N: EG AFR

genus Megaplatypus Wood, 1993: 275 type species Platypus dentatus Dalman, 1823
mutatus Chapuis, 1865: 136 (Platypus) Ei: IT NTR
plicatus Brèthes, 1909:225 (Platypus)
su lea tus Chapuis, 1865: 137 (Platypus)

genus Peroplatypus Wood, 1993: 272 type species Platypus truncatipennis Schedl, 1964
abruptus Sampson, 1923: 285 (Platypus) A: UP

genus Platypus Herbst, 1794a: 128 type species Bostrichus cylindrus Fabricius, 1792
Cylindra Illiger, 1802: 303 [unjustified RN]
Stenoplatypus Strohmeyer, 1914a: 35 type species Crossotarsus spinulosus Strohmeyer, 1912
arisannensisMurayama, 1934c: 135 A.TAI
beaveri Browne, 1975: 306 A: TAI ORR
formosanus Niisima & Murayama, 1925: 217 (Diapus) [HN]
keelungensis Browne, 1985:294
biformis Chapuis, 1865: 255 A: SD PA UP ORR
contaminatus Blandford, 1894a: 131 (Crossotarsus) A: FUJ JA TAI "Himalaya"
curtatus Sampson, 1923: 286 A: SD ORR
curtus Chapuis, 1865: 261 A: FUJ TAI AUR ORR
artecurtus Schedl, 1960a: 50
cylindrus Fabricius, 1792: 364 (Bostrichus) E: AU BE BU BY CR CZ DE FR GB GE GR HU IT LA MC NL
NR PL PT RO SK SL SP ST "Caucasus" SV SZ UK YU N: AG EG LB MO TU A: IN TR
bimaculatus Duftschmid, 1825: 88 (Cylindra)
cylindriformis Reiner, 1895b: 94
flavicornis Kugelann, 1792: 496 (Bostrichus) [HN]
platypus Duftschmid, 1825: 87 (Cylindra)
darjeelingensis Schedl, 1969b: 61 A: SD
dasycauda Browne, 1980a: 376 A: TAI
 PLATYPODINAE 203
errans Sampson, 1922: 145 {Crossotarsus) A: UP SD ORR
eugeniae Schedl, 1971b: 390 A: UP
formosanus Niisima & Murayama, 1925: 215 A: JA TAI
taiwansis Schedl, 1960b: 111
horishensis Murayama, 1928: 284 A: TAI
hybridus Schedl, 1935b: 395 A: PA AUR ORR
indicus Strohmeyer, 1910: 131 A: SD TAI UP ORR
jansoni Chapuis, 1865: 244 A: TAI AFR AUR ORR
kiushuensis Murayama, 1936: 145 A: GUI JA TAI
klapperichi Schedl, 1941 a: 44 (Stenoplatypus) A:FUJGUA
koryoensis Murayama, 1930: 24 {Crossotarsus) A: FE JA SC TAI
lewisi Blandford, 1894a: 134 A: BT FUJ GUI JA SC SD TAI ORR
lopchuensis Schedl, 1969b: 68 A: SD
lunatulus Browne, 1980d: 497 A: TAI
modestus Blandford, 1894a: 133 A: JA TAI
niijimai Murayama, 1931b: 197 A: TAI
obtusipennis Schedl, 1940a: 2 A: TAI ORR
octodentatus Browne, 1985: 294 A: TAI
quadriporus Beeson, 1937: 96 (Crossotarsus) A: BT SD ORR
querci Browne, 1980c: 488 A: TAI
quercivorus Murayama, 1925: 229 (Crossotarsus) A: FE JA SD TAI AUR ORR
sexfenestratus Beeson, 1937: 94 (Crossotarsus)
secretus Sampson, 1921: 25 A: BT SD ORR
eruditus Schedl, 1942b: 212
sexualis Beeson, 1937: 95 (Crossotarsus) A: UP
simulons Schedl, 1941c: 413 A: SY TR
sinensis Schedl, 1941a: 43 A: FUJ GUA
stenoplicatus Schedl, 1940a: 3 A: FUJ ORR
suffodiens Sampson, 1913: 447 A: SD ORR
taiheizanensis Murayama, 1932: 485 (Crossotarsus) A: TAI

genus Treptoplatypiis Schedl, 1972a: 243 type species Crossotarsus trepanatus Chapuis, 1865
oxyurus Dufour, 1843: 92 (Piatypus) E: FR GE GR IT SK SP A: IN TR
quadriporus Schedl, 1971b: 397 A: FUJ
severini Blandford, 1894a: 136 (Piatypus) A: FE JA SC SE TAI
solidus Walker, 1859a: 261 (Piatypus) A: IS JA NP PA SC SD TAI UP YUN AUR ORR
caudal us Motschulsky, 1863: 509 (Piatypus)
cordatus Motschulsky, 1863: 510 (Piatypus)
exilis Chapuis, 1865: 268 (Piatypus)
obtusus Chapuis, 1865: 268 (Piatypus)
pilifrons Chapuis, 1865: 265 (Piatypus)
rudis Chapuis, 1865: 268 (Piatypus)
xylographus Schedl, 1969a: 216 (Platypus) A: TAI

supertribe Tesserocerinitae Blandford, 1896

tribe Diapodini Strohmeyer, 1914
genus Diapus Chapuis, 1865: 43,329 type species Diapus quadrispinatus Chapuis, 1865
aculeatus Blandford, 1894a: 139 A: BT JA TAI UP ORR
aulacus Beaver & Liu, 2008: 112 A: TAI
minor Schedl, 1954c: 158 A: SD ORR
molossus Chapuis, 1865:333 A: UP ORR
cap Hiatus Sampson, 1913: 449
impressus O. E. Janson, 1893: 74
quadrispinatus Chapuis, 1865: 332 A: UP YUN ORR
quinquespinatus Chapuis, 1865: 334 A: BT JA TAI UP XIZ YUN AFR AUR ORR
204 SCOLYTINAE
tonkinensis Schedl, 1972d: 201
truncatus Niisima & Murayama, 1934: 143 A: SD TAI UP ORR
spatulifer Browne, 1977: 101

genus Genyocems Motschulsky, 1858: 68 type species Genyocerus albipennis Motschulsky, 1858
Diacavus Schedl, 1939c: 363 type species DiapusfrontalisStrohmeyer, 1914
albipennis Motschulsky, 1858: 68 A: JA ORR
irregularis Browne, 1970: 582 {Diacavus)
diaphanus Schedl, 1939c: 364 (Diacavus) A: JAi UP ORR
abdominalis Schedl, 1942b: 218 (Diacavus)
assamensis Browne, 1964: 760 (Diacavus)
philippinensis Schedl, 1942b: 216 (Diacavus)
sexporus Schedl, 1942: 217 (Diacavus)
talurae Stebbing, 1906: 418 (Diapus) A: PA UP ORR
dipterocarpi Browne, 1977: 98
furtivus Sampson, 1913:450 (Diapus)
minis Sampson, 1913: 452 (Diapus)

subfamily Scolytinae Latreille, 1804

supertribe Hylesinitae Erichson, 1836
tribe Diamerini Hagedorn, 1909
genus Diamerus Erichson, 1836: 57 type species Hylesinus hispidus Klug, 1833
Acanthurus Eichhoff, 1886: 24 type species Acanthunis spinipennis Eichhoff, 1886 (= Hylesinus curvifer Walker, 1859)
Lissoclastus C. F. C. Schaufuss, 1905b: 71 type species Lissoclastus pimelioides C. F. C. Schaufuss, 1905
ater Hagedorn, 1909: 735 A: YUN ORR
curvifer Walker, 1859a: 261 (Hylesinus) A: HAI YUN HP SD UP AUR ORR
dissimilis Hagedorn, 1909: 735
spinipennis Eichhoff, 1886: 24 (Acanthurus)
fici Blandford, 1898: 426 A: SDYUNXIZ ORR
matangi Sampson, 1919: 108 A: XIZ ORR
nigrisetosus Eggers, 1936a: 79 A: SD ORR
striatus Eggers, 1927e: 67 A: SD YUN ORR
variegatus Schedl, 1959a: 41 A: UP ORR

genus Sphaerotrypes Blandford, 1894a: 61 type species Sphaerotrypes pila Blandford, 1894
bengalensis Wood, 1988a: 35 A: SD
coimbatorensis Stebbing, 1906: 395 A: ANH BEI HEB HEI HEN HUN SCH SHA SHX ORR
globulus Blandford, 1894a: 63 A: UP ORR
jugransi Tsai & Yin, 1966: 236, 239 A: ANH SCH SHA SHX
juglansis Krivolutskaya, 1970: 210 A: FE
magnus Tsai & Yin, 1966: 234,239 A: SCH YUN
pila Blandford, 1894a: 62 A: HEB JA LIA SC SCH TAI XIZ YUN
carpini Eggers, 1926a: 134
imitans Eggers, 1926a: 134
pyri Tsai & Yin, 1966: 237, 240 A: HAI SCH SHX YUN
querci Stebbing, 1908: 5 A: ANH HEB SCH SHX UP YUN ORR
tectus Beeson, 1921: 516 [RN]
siwalikensis Stebbing, 1906: 389 A: NP PA SD UP ORR
assamensis Stebbing, 1907a: 23
tsugae Tsai & Yin, 1966: 236,240 A: SCH SHA YUN
ulmi Tsai & Yin, 1966: 237, 240 A: SCH SHA SHX
yunnanensis Tsai & Yin, 1966: 238, 240 A: YUN

tribe Hylastini LeConte, 1876

genus Hylastes Erichson, 1836: 47 type species Bostrichus ater Paykull, 1800
Ipsocossonus Oke, 1934: 250 type species Ipsocossonus anomalus Oke, 1934'(= Bostrichus ater Paykull 1800)
 SCOLYTINAE 205
ambiguus Blandford, 1894a: 57 A: JA
angustatus Herbst, 1794a: 111 (Bostrichus) E: AU BE BU BY CR CT CZ DE EN FI FR GB GE GR HU IT LA
LT MC MD NR NT PL PT SK SL SP ST SV SZ TR UK YU A: TR AFRi
scandinavicus Lekander, 1965: 199
ater Paykull, 1800: 153 {Bostrichus) E: AU AZ BE BH BU BY CR CT CZ DE FI FR GB GE GR IT LA LT
MC NL NR PL PT SK SL SP ST SV SZ UK YU A: ES FE FUJ JA KZ SC TR WS "Manchuria" AURi
s
NTRi
angusticollis Eggers, 1929a: 9
anomalus Oke, 1934: 251 (Ipsocossonus)
chloropus Duftschmid, 1825: 102 (Hylesinus)
pinicola Bedel, 1888: 390
attenuatus Erichson, 1836: 50 E: AU AZ BE BH BY CR CT CZ EN FI FR GB GE GR IT LA LT MC NR NT
PL PT SK SL SP ST SV SZ UK YU N: MR A: HEI JA LIA SC TAI TR
batnensis anatolicus Knfëek & Pfeffer, 1995: 74 A: TR
batnensis batnensis C. N. F. Brisout de Barneville, 1883: 146 E: IT N: AG MO
brunneus Erichson, 1836: 48 (Hylesinus) E: AU AZ BE BU CT CZ DE EN FI GB GE GR IR IT LA MC NR
NT PL SK SL ST SV SZ A: ES FE HP MG NC NE SC TR UP WS "Manchuria"
aterrimus Eggers, 1933a: 3
rotundicollis Reitter, 1895b: 60
cunicularius Erichson, 1836: 49 E: AU BE BH BU BY CR CT CZ DE EN FI FR GB GE GR HU IR IT LA LS
LT MC NR NT PL PT SK SL SV ST SZ UK YU N: AG A: JA ES FE HUB LIA NC SHA SCH SY TR
WSXINYUN
starki Eggers, 1933a: 1 (Hylurgops)
gergeri Eggers, 191 lb: 119 E: AU CZ HU IT RO SK SZ
fallax Wichmann, 1911a: 100
linearis Erichson, 1836: 49 E: BE CR CZ FR GB GE GR HU IT MC PL PT SK SL SP SZ UK YU N: AG CI
MO MR TU A: CY IS SY TR AFRi
clavus Wollaston, 1854: 305
corticiperda Erichson, 1836: 50
flavicornis Lindberg, 1950: 19
vaholosus Perris, 1852: 181
/owe/Paiva, 1861:211 N: CI
opacus Erichson, 1836: 51 E: AU BE BU BY CT CZ DE EN FI FR GB GE GR HU IR IT LA LS LT MC NR
NT PL SK SL ST SV SZ YU A: ES FE JA KZ MG NC SC WS "Manchuria" NARi
simplex Rey, 1892:30
parallelus Chapuis, 1875: 196 A: ES FE GAN HEI JA JIL LIA QIN SC SCH SHA TAI
plumbeus Blandford, 1894a: 57 E: CT FI NT PL SV A ES FE HEI JA JIL LIA NC SC SCH SHA SHN TAI
WS
fiishunensis Murayama, 1940: 235 (Hylurgops)
obscurus Chapuis, 1876: 197 [HN]
septentrionalis Eggers, 1923b: 135
substriatus Strohmeyer, 1914b: 7 A: KI KZ TD TR "China"
techangensis Tsai & Hwang, 1964a: 232, 233 A: SCH XIZ YUN
genus Hylurgops LeConte, 1876: 389 type species Hylastes pinifex Fitch, 1858
bonvouloiri Chapuis, 1869: 22 (Hylastes) N: AG
eusulcatus Tsai & Hwang, 1964b: 239, 241 A: SCH SHA YUN
glabratus Zetterstedt, 1828: 343 (Hylurgus) [NP] E: AU BU BH BY CT CZ DE EN FI FR GE GR HU IT LA
NR NT PL SK SL SP SV SZ A: ES FE HEI JA JIL KZ MG NC SC TAI TR WS
decumanus Erichson, 1836: 51 (Hylastes)
paykullii Duftschmid, 1825: 99 (Hylesinus) [NO]
tenebrosus C. R. Sahlberg, 1836a: 139 (Hylesinus)
inouyei Nobuchi, 1959a: 10 A: JA
interstitialis Chapuis, 1875: 196 (Hylastes) A: FE GAN HEI JA JIL NC LIA SC
niponicus Murayama, 1936: 142
junnanicus Sokanovskiy, 1959a: 93, 94 A: YUN
206 SCOLYTINAE
longipillus Reitter, 1895b: 63 A: FE GAN GUX HEI JA NC LIA SCH SHA SHX YUN
imitator Reitter, 1900: 59 (Hylastes)
likiangensis Tsai & Hwang, 1964b: 237, 240
major Eggers, 1944a: 67 A: FUJ HUN XIZ YUN
palliatus Gyllenhal, 1813: 340 (Hylesinus) [NP] E: AU BE BH BU BY CR CT CZ DE EN FI FR GB GE HU
IR IT LA LS LT MC NL NR NT PL SK SL SP SV ST SZ UK YU N: AG A: ES FE HEI JA KZ NC SC
SHA TR WS NARi
abietiperda Bechstein, 1818: 74 (Bostrichus)
fuscus Duftschmid, 1825: 105 (Hylesinus)
helfen A. Villa & G. B. Villa, 1835: 49 (Hylurgus)
marginatus Duftschmid, 1825: 104 (Hylesinus)
parvus Eggers, 1933a: 2
piceus Marsham, 1802: 58 (lps) [NO]
rufescens Stephens, 1830: 364 (Hylurgus)
rufus Marsham, 1802: 57 (Ips) [NO]
spessiwzeffi Eggers, 1914b: 187 A: ES FE JA NC NE SC TAI
modestus Murayama, 1937: 367
squamosus Murayama, 1942: 56
sulcatus Eggers, 1933f: 97 A: SCH YUN
transbaicalicus Eggers, 1941b: 119 A: FE JA
tuberculatus Eggers, 1933e: 98 A: SCH

tribe Hylesinini Erichson, 1836

genus Alniphagus Swaine, 1918: 73 type species Hylesinus aspericollis LeConte, 1876
Hylastinoides Spessivtsev, 1919: 249 type species Hylastes alni Niisima, 1909 (= Hylesinus costatus Blandford, 1894)
costatus Blandford, 1894a: 63 (Hylesinus) A: FE JA JJL NC SC TAI
alni Niisima, 1909: 137 (Hylastes)
imitator Sokanovskiy, 1958: 38

genus Dactylipalpus Chapuis, 1869: 12 type species Dactylipalpus transversus Chapuis, 1869
Dactylopselaphus Gemminger & Harold, 1872: 2678 [RN] type species Dactylipalpus transversus Chapuis, 1869
Ethadopselaphus Blandford, 1896d: 321 type species Ethadopselaphus cicatricosus Blandford, 1896
transversus Chapuis, 1869: 12 A: TAI AUR ORR
quadratocollis Chapuis, 1869: 12

genus Ficicis Lea, 1910:147 type species Ficicis varions Lea, 1910
Ficiphagus Murayama, 1958: 930 type species Phloeosinus goliathoides Murayama, 1955 (= Hylesinus porcatus Chapuis,
1869)
porcatus Chapuis, 1869: 31 (Hylesinus) A: HAI JA SD TAI UP YUN AUR ORR
crassus Beeson, 1929: 220 (Hylesinus)
goliathoides Murayama, 1955: 94 (Phloeosinus)
insularum Beeson, 1940: 192 (Hylesinus)
philippinensis Eggers, 1923a: 137 (Hylesinus)
koebelei Lea, 1910: 147
subcostatus Eggers, 1923a: 137 (Hylesinus)
subopacus Eggers, 1930a: 10 (Hylesinus)
javanus Eggers, 1923a: 135 (Hylesinus) A: UP ORR

genus Hylastinus Bedel, 1888: 388 type species Ips obscurus Marsham 1802
achillei Reitter, 1895b: 54 N: AG A:SY
fankhauseri Reitter, 1895b: 54 E: AU CR FR IT SL SZ
kroaticus Fuchs, 1912: 49 E: BU CR GR
obscurus Marsham, 1802: 57 (Ips) E: AU BE CR CZ DE FR GB GE GR IR IT LA NL PL PT SK SL ST SV SP
SZ UK YU N: AG CI MO MR NARi NTRi
crenatulus Duftschmid, 1825: 104 (Hylesinus)
crenatus Olivier, 1795: 12 (Scolytus)
fuscescens Stephens, 1830: 365 (Hylurgus)
 SCOLYTINAE 207
pilosus Eggers, 1944c: 140
trifolii P. W. J. Müller, 1803: 47 (Dermestes)
tiliae Semenov, 1902: 271 E: GG ST

genus Hylesinopsis Eggers, 1920a: 40 type species Hylesinopsis dubia Eggers, 1920
Chilodendron Schedl, 1953b: 74 type species Chilodendron planicolle Schedl, 1953
Trypographus Schedl, 1950c: 213 type species Trypographus joveri Schedl, 1950
arabiae Schedl, 1975c: 353 A: YE
saudiarabiae Schedl, 1971c: 434 {Chilodendron) A: SA

genus Hylesinus Eabricius, 1801: 390 type species Bostrichus crenatus Fabricius, 1787
Leperisinus Reitter, 1913b: 41 type species BostrichusfraxiniPanzer, 1799 (= Bostrichus varius Fabricius, 1775)
botscharnikovi Stark, 1931: 81, 83 E: AB ST
cholodkovskyi Berger, 1917: 246 A: FEHEI
cingulatus Blandford, 1894a: 67 A: FE HEI JA JIL NC SC
crenatus Fabricius, 1787: 37 (Bostrichus) E: AU BE BY CR CT CZ DE EN FI FR GB GE GR HU IR IT LA
LT LU MC NR NT PL RO SK SL SP SV ST SZ UK YU N: AG MO A: KI TR
crenatus Gravenhorst, 1807: 196 (Anthribus) [HN]
prutenskyi Sokanovskiy, 1959b: 276
eos Spessivtsev, 1919: 248 A: FE HEI JA NC NE "Central China"
laticollis Blandford, 1894a: 65 A: FE HEI JA NC SC
lubarskii Stark, 1936a: 153
striatus Eggers, 1933a: 4
macmahoni Stebbing, 1909: 16 (Sphaerotrypes) A: KA PA ORR
alternons Schedl, 1959c: 172
fraxinoides Schedl, 1959a: 39 {Leperisinus)
mandshuricus Eggers, 1922a: 15 A: "Manchuria"
nobilis Blandford, 1894a: 64 A: FE HEI JA
shabliovskyi Kurentsov, 1941: 113,229
wachtli orni Fuchs, 1906: 51 E: AU BE BU CR CZ DE FI FR GB GE GR HU IT MC NL NR PL RO SK SL SP
SVSZUK
wachtli wachtli Reitter, 1887: 193 E: FR
pravdini Stark, 1936a: 153 A: FE NC
toranio D'Anthoine, 1788: 270 (Byrrhus) E: AU BE BU CR CZ DE FR GB GE GR HU IT LA MC MD NR PL
SK SL SP SV ST SZ UK YU N: AG MO A: IS JAi LE TR NTRi
antipodus Schedl, 1952a: 17
bicolor Brülle, 1832: 250 (Hylesinus)
asai/Gredler, 1863:370
oleiperda Fabricius, 1792: 366 (Bostrichus)
scaber Marsham, 1802: 56 (Ips)
suturalis W. Redtenbacher, 1842: 21
tristis Blandford, 1894a: 66 A: JA SC
tupolevi Stark, 1936a: 151 E: AB A: YUN KI TD
denticulosus Sokanovskiy, 1956: 44
tupolevi Eggers, 1942: 28 (Leperisinus)
varius Fabricius, 1775: 60 (Bostrichus) E: AU BE BU BY CR CT CZ DE EN FI FR GB GE GR HU IR IT LA
LS LT LU MA MC NR NT PL PT RO SK SL SP SV ST SZ UK YU N: AG MO TU A: HEI IN JIL LIA
SHATR
griseus Marsham, 1802: 55 (Ips)
fraxini Panzer, 1799b: 13 (Bostrichus)
haemorrhoidalis Marsham, 1802: 56 (Ips)
henscheli Knotek, 1892: 234
melanocephalus Fabricius, 1792: 368 (Bostrichus)
minutus Fabricius, 1777:211 (Bostrichus)
picipennis Stephens, 1830: 359
pubescens Fabricius, 1798: 161 (Anthribus)
rufescens Marsham, 1802: 55 (Ips)
208 SCOLYTINAE
verae Petrov, 2002: 121 A: TM

genus Kissophagus Chapuis, 1869: 34 type species Hylesinus hederae Schmitt, 1843 (= Hylesinus vicinus
Comolli, 1837)
erinacellus Wichmann, 1916: 18 E: CR GR
vicinus Comolli, 1837: 36 (Hylesinus) E: AU BE CR FR GB GE GR HU IT MA NL PL SP ST SZ UK A: TR
N:AG
hederae Schmitt, 1843: 108 (Hylesinus)
nuesslini Reitter, 1913b: 44
novaki Reitter, 1894a: 45 E: AU CR CZ FR GR IT SK ST SZ N: AG
binodus Reitter, 1913b: 45

genus Longulus Krivolutskaya, 1968: 50 type species Hylesinus elatus Niisima, 1913
elatus Niisima, 1913: 2 (Hylesinus) A: FE (Kuriles) JA

genus Neopteleobius Nobuchi, 1971a: 125 type species Hylesinus scutulatus Blandford, 1894
scutulatus Blandford, 1894a: 67 (Hylesinus) A: JA LIA NC SC
trepanatus Wichmann, 1914a: 137 (Pteleobius)

genus Pteleobius Bede), 1888: 392,393 type species Bostrichus vittatus Fabricius, 1792
kraatzii Eichhoff, 1864: 30 (Hylesinus) E: AU BE BU BY CR CT CZ FR GE GR HU IT MC NL PL SK SP ST
SZ UK YU N: AG CI A: TR
putonii Eichhoff, 1868d: 403 (Hylesinus)
vittatus Fabricius, 1792: 368 (Bostrichus) E: AU BE BU BY CR CT CZ FR GB GE GR HU IR IT LA LT MC
NL PL RO SK SP ST SZ UK YU A: IS TR
coadunatus Marsham, 1802: 58 (Ips)
furcatus Marsham, 1802: 55 (Ips)
sericeus Marsham, 1802: 55 (Ips)

tribe Hylurgini Gistel, 1848

genus Chaetoptelius Fuchs, 1913b: 43 [RN] type species Hylesinus vestitus Mulsant & Rey, 1861
Acrantus Broun, 1882:409 [RN] type species Homarus mundulus Broun, 1881
Chaetophorus Fuchs, 1912: 46 [HN] type species Hylesinus vestitus Mulsant & Rey, 1861
Homarus Broun, 1881: 740 [HN] type species Homarus mundulus Broun, 1881
vestitus Mulsant & Rey, 1861a: 340 [= 1861b: 87] (Hylesinus) E: BU CR FR GR HU IT MC SP ST UK N: AG
CI MO TU A: IN IQ IS SY TR UP NTR ORR
indigenus Wollaston, 1864: 267 (Hylesinus)

genus Dendroctonus Erichson, 1836: 52 type species Bostrichus micans Kugelann, 1794
armandi Tsai & Li, 1959: 80 A: GAN GUI HEI HEN HUB HUN SCH SHA SHX YUN
prosorovi Kurentsov & Kononov, 1966: 29
micans Kugelann, 1794: 523 (Bostrichus) E: AU BE BU BY CR CT CZ DE EN FI FR GB GE GR HU IT LA
LT LU NL NR NT PL SK SL SP SV SZ UK YU A: ES FE GAN HEI JA JIL KZ LIA MG NMO QIN SCH
TR WS XIN XIZ
abietinus Fabricius, 1792: 367 (Bostrichus) [invalid, see Opinion 670]
Valens LeConte, 1857: 59 Ai: HEB HEN SHA SHX NAR
beckeri Thatcher, 1954: 4

genus Hylurgus Latreille, 1806:274 type species Bostrichus ligniperda Fabricius 1787
indicus Wood, 1985: 273 A: UP
ligniperda Fabricius, 1787: 37 (Bostrichus) E: AU AZ BE BH BU BY CR CT CZ DE EN FI FR GE GR HU IT
LALTMCMDNLNRNTPLPTROSKSLSPSTSVSZUKYU N:AGCIMOMRTU ACYHEI
JA TR WS AFRi AURi NARi
elongatus Herbst, 1794a: 117 (Bostrichus)
flavipes Panzer, 1799a: 9 (Bostrichus)
longulus Kolenati, 1846:38
 SCOLYTINAE 209
micklitzi Wachtl, 1881: 227 E: CR FR GE GR HU IT MA SP ST N: AG EG LB MO TU A: IS TR
nomen dubium
rufipes Stephens, 1835: 419 E: GB

genus Pseudohylesinus Swaine, 1917: 11 type species Pseudohylesinus grandis Swaine, 1917 (= Hylurgus
sericeus Mannerheim, 1843)
nebulosus nebulosus LeConte, 1860: 285 {Hylesinus) A: TR NAR
sericeus Mannerheim, 1843: 296 (Hylurgus) A: JAi NAR
grandis Swaine, 1917: 13
yasumatsui Nobuchi, 1971b: 160

genus Pseudoxylechinus Wood & Huang, 1986: 465 type species Pseudoxylechinus uniformis Wood & Huang,
1986
indicus Wood, 1986b: 468 A: SD
piceae Chen & Yin, 2001: 336, 338 A: YUN
rugatus Wood & Huang, 1986: 467 A: YUN
sinensis Wood & Huang, 1986: 467 A: YUN
tibetensis Wood & Huang, 1986: 466 A: XIZ
tiliae Niisima, 1910: 2 (Kissophagus) A: JA
uniformis Wood & Huang, 1986: 465 A: SCH SHA
variegatus Wood & Huang, 1986: 466 A: SHX

genus Tomicus Latreille, 1802: 203 type species Dermestes piniperda Linnaeus, 1758
brevipilosus Eggers, 1929c: 103 (ßlastophagus) A: FUJ JA SC YUN
khasianus Murayama, 1959: 75 (Blastophagus)
multisetosus Murayama, 1963a: 37 (Blastophagus)
destruens Wollaston, 1865: 45 (Hylurgus) E: CR FR GR IT PT SP ST UK N: AG MR A: CY IS TR
minor Hartig, 1834: 413 (Hylesinus) E: AU BE BH BU BY CR CT CZ DE EN FI FR GB GE GR HU IT LA
LT LU MC NL NR NT PL PT RO SK SL SP ST SV SZ TR UK YU A: ANH ES FE CY FUJ GAN GUI
GUX HEB HEI HEN HUB HUN JA JIA JIL JIX KZ LIA MG NC NE NMO SC SCH SHA SHX SCH TR
WS YUN ZHE
corsicus Eggers, 1911a: 75 (Myelophilus)
pilifer Spessivtsev, 1919: 250 (Myelophilus) A: BEI FE HEB HEI HUB JIL NC NMO QIN SCH SHA SHX
XIZ YUN
piniperda Linnaeus, 1758: 355 (Dermestes) E: AU BE BH BU BY CR CT CZ DE EN FI FR GB GE GR HU IR
IT LA LS LT LU MC NL NR NT PL PT RO SK SL SP ST SV SZ UK YU N: AG MO MR TU A: ANH
CY ES FE FUJ GAN GUI HEB HEI HEN HUB HUN IS JA JIA JIL JIX KZ LIA MG NC NMO QIN SC
SCH SHA SHN SHX SCH TAI TR WS YUN ZHE NARi ORR
analogus LeConte, 1868: 172 (Hylurgus)
major Eggers, 1943a: 50 (Blastophagus)
testaceus Fabricius, 1787: 37 (Bostrichus)
puellus Reitter, 1895b: 53 (Myelophilus) A: FE JA NC
orientalis Krivolutskaya, 1956: 828 (Blastophagus)
starki Eggers, 1929c: 104 (Blastophagus)
yunnanensis Kirkendall & Faccoli, 2008: 32 A: YUN

genus Xylechinus Chapuis, 1869: 36 type species Hylesinuspilosus Ratzeburg, 1837

Pruniphagus Murayama, 1958: 930 type species Priniphagus gummensis Murayama, 1958
Squamosinus Nunberg, 1964: 431 type species Squamosinus chiliensis Nunberg, 1964
Xylechinops Browne, 1973: 283 type species Xylechinus australis Schedl, 1957
arisanus Eggers, 1939b: 114 A: TAI
bergeri Spessivtsev, 1919: 249 A: FE HEI NC
darjeelingensis Schedl, 1971b: 371 A: NP SD
gummensis Murayama, 1958: 931 (Pruniphagus) A: JA
obscurus Eggers, 1941c: 222 A: FUJ
210 SCOLYTINAE
ougeiniae Wood, 1988a: 37 A: UP
padus Wood, 1988a: 38 A: HP KA UP YUN
pilosus Ratzeburg, 1837: 178 {Hylesinus) E: AU BE CT CZ DE EN FI FR GB GE GR HU IT LA NR NT PL
RO SK SL SV ST SZ UK A: ES FE HEI JA JIL KZ MG NC SHA WS XIN

tribe Hyorrhynchini Hopkins, 1915

genus Hyorrhynchus Blandford, 1894a: 58 type species Hyorrhynchus lewisi Blandford, 1894
ebianensis Huang & Yin, 1983: 339 A: SCH
kalimpongensis Maiti & Saha, 1989: 41 A: SD
lewisi Blandford, 1894a: 60 A: FE (Kuriles) JA TAI ORR
drescheri Eggers, 1936a: 82
gramdatus Eggers, 1936a: 80
sensarmai Maiti & Saha, 1989: 44 A: SD
shiva Maiti & Saha, 1989: 46 A: SD ORR
tuberopectus Huang & Yin, 1983: 340 A: SCH
unicornis Nobuchi, 1966: 51 A: JA TAI

genus Pseudohyorrhynchus Murayama, 1950b: 61 type species Pseudohyorrhynchus wadai Murayama, 1950
blandfordi Sampson, 1913: 446 {Hyorrhynchus) A: SD YUN XIZ ORR
flavopannus Huang & Yin, 1983: 338 {Hyorrhynchus) A: SCH
wadai kurilensis Krivolutskaya, 1996: 331 A: FE (Kuriles)
wadai wadai Murayama, 1950b: 61 A: JA TAI

genus Sueus Murayama, 1951: 1 type species Sueus sphaerotrypoides Murayama, 1951 (= Hyorrhynchus
niisimai Eggers, 1926)
Parasphaerotrypes Murayama, 1958: 933 type species Sphaerotrypes controversae Murayama, 1950 (= Hyorrhynchus
niisimai Eggers, 1926)
niisimai Eggers, 1926a: 133 {Hyorrhynchus) A: HAI JA SD TAI AUR ORR
controversae Murayama, 1950b: 62 {Sphaerotrypes)
pilosus Eggers, 1936a: 81 {Hyorrhynchus)
sphaerotrypoides Murayama, 1951: 2

tribe Hypoborini Nüsslin, 1911

genus Hypoborus Erichson, 1836: 62 type species Hypoborus ficus Erichson, 1836
ficus Erichson, 1836: 62 E: AB AU AZ BH BU CR FR GR HU IT MA MC PT SL SP ST SZ UK YU N: AG
CI EG MO MR TU A: CY IN IQ IS JO TR
siculus Ferrari, 1867: 18

genus Liparthrum Wollaston, 1854: 294 type species Liparthrum bituberculatum Wollaston, 1854
Dacryophthorus Schedl, 1971a: 281 type species Dacryophthorus brincki Schedl, 1971
Erineosinus Blackman, 1920: 53 type species Erineosinus squamosus Blackman, 1920
Phloeochilus Schedl, 1953d: 292 type species Phloeochiluspalaquius Schedl, 1953
Phloeotrypetus Wood, 1960a: 16 type species Phloeotrypetus palauensis Wood, 1960
Trypanophellos Bright, 1982: 166 type species Trypanophellos necopinus Bright, 1982
arnoldi Semenov, 1902: 272 E: AB BU CR GG RO ST A: TR -
babadjanidis Eggers, 1910b: 558
artemisiae Wollaston, 1854: 299 N: CI MR
bartschti Mühl, 1891: 202 E: AU HU SK ST "Caucasus" A: TR
bicaudatum Wollaston, 1865: 44 N: CI
bituberculatum Wollaston, 1854: 297 N: CI MR
canum Israelson, 1990: 6 N: CI
Colchicum Semenov, 1903: 79 E: CR SP ST N: AG
corsicum Eichhoff, 1878a: 383 [=1878b: 110] E: FR (Corse)
curtum Wollaston, 1854: 298 E: AZ N: CI MR
ciliatum Eggers, 1928c: 283
 SCOLYTINAE 211
genistae genistae Aube, 1862b: 388 {Hypoborus) E; FR IT SP N: AG MO TU A: IS TR
peyrimhoffi Pfeffer, 1941a: 26
genistae georgi Knotek, 1895: 89 E: BH CR GR ST UK (Krim) "Caucasus" A: CY
albidum Wichmann, 1916: 21
cytisi Eggers, 1927b: 121
inarmatum Wollaston, 1860b: 364 N: CI MO MR
laurivorum Schedl, 1968a: 23 A: TR
longifolia Stebbing, 1903: 267 {Cryphalus) A: HP UP ORR
/owe/Wollaston, 1862a: 174 N: AZCI
mandibulare Wollaston, 1854: 295 E: AZ GB (Guernsey) SP N: CI MR
mori Aube, 1862b: 387 {Hypoborus) E: CR FR GR HU IT MA UK N: AG MO TU
dalmatinum Eggers, 1944c: 140
nigrescens Wollaston, 1865: 44 N: CI MO A: TR
degener Lindberg, 1953: 18
semidegener Israelson, 1990: 7 N: MR
tribe Phloeosinini Nüsslin, 1912
genus Hyledius Sampson, 1921: 35 type species Hyledius asper Sampson, 1921 (= Olonthogaster nitidicollis
Motschulsky, 1866)
Holonthogaster Gemminger & Harold, 1872: 2676 [unjustified emendation] type species Olonthogaster nitidicollis
Motschulsky, 1866
Hylurgulus Eggers, 1927d: 392 type species Hylurgulus sumatranus Eggers, 1927 (= Phloeosinus hylurgulus Eggers,
1940)
Olonthogaster Motschulsky, 1866: 401 [suppressed for priority, ICZN, Opinion 1167] type species Olonthogaster
nitidicollis Motschulsky, 1866
Phloeosinopsis Schedl, 1936a: 23 type species Phloeosinopsis armata Schedl, 1936
ceylonicus Schedl, 1959d: 473 {Phloeosinus) A: YUN ORR
cribratus Blandford, 1896b: 198 {Phloeosinus) A: HAI SCH YUN ORR
malayensis Schedl, 1936a: 22 {Phloeosinus)
genus Phloeocranus Schedl, 1942a: 7 type species Phloeocranus bruchoides Schedl, 1942
Diamerides Browne, 1949: 893 type species Diamerides litseae Browne, 1949 (= Phloeocranus bruchoides Schedl, 1942)
bruchoides Schedl, 1942a: 8 A: SD ORR
litseae Browne, 1949: 894 {Diamerides)

genus Phloeosinopsioides Schedl, 1964e: 317 [RN] type species Phloeosinopsis triseriata Schedl, 1964
Phloeosinopsis Schedl, 1964b: 297 [HN] type species Phloeosinopsis triseriata Schedl, 1964
formosanus Schedl, 1935e: 479(Ay/ecAi«us) A: TAI ORR

genus Phloeosinus Chapuis, 1869: 37 type species Hylesinus thujae Perris, 1855
abietis Tsai & Yin, 1964: 92, 95 A: HUB YUN
acatayi Schedl, 1958a: 33 A: SY TR
libani Balachowsky, 1969: 649
arisanus Niisima, 1942: 73 A: TAI
armatus Reitter, 1887: 192 E: GR IT A: CY IS JO LE S Y TR "Palestina" NARi
andresi Eggers, 1927b: 120
aubei Perris, 1855: Ixviii {Hylesinus) E: AL AR AU BE BH BU CR CZ FR GB GE GR HU IT MC NL PL PT
RO SK SP ST SZ UK YU N: AG CI BO LB MO TU A: ANH BEI CY GAN GUI HEB HEN HUN IN IS
JLA NC QIN SHA SHN SCH SHX SY TAI TM TR YUN
hercegovinensis Eggers, 1922b: 120 /
praenotatus Gredler, 1866:370 {Phloeophthorus)
schumensis Eggers, 1922c: 166
transcaspicus Semenov, 1903: 79
camphoratus Tsai & Yin, 1964: 93, 96 A: YUN ORR
cedri C. N. F. Brisout de Barneville, 1883: 146 E: SP N: AG MO
maura Balachowsky, 1969: 649
212 SCOLYTINAE
gifiiensis Murayama, 1954: 190 A: JA
gifensis Murayama, 1954: 190 [incorrect original spelling]
gillerforsi Bright, 1987: 3 E: AZ N: CI
henschi Reitter, 190Id: 201 E: BH CR GR MC UK A: LE SY
inermis Nunberg, 1948: 14
krimaeus Eggers, 1937: 334
stoeckleini Schedl, 1935a: 241
hopehi Schedl, 1953a: 23 A: BEI HEB HUN NIN SC SCH SHA SHX
jubatus Sampson, 1919: 112 A: UP
kiushuensis Murayama, 1955: 93 A: JA
dubius Blandford, 1894a: 70
izuensis Nobuchi, 1959a: 9
kumamotoensis Murayama, 1955: 91 A: JA
lewisi Chapuis, 1876: 198 A: GUI JA JIL TAI
minutus Blandford, 1894a: 71
machilus Schedl, 1959c: 173 (Hylesinus) A: FUJ HUN UP
cinnamomi Tsai & Yin, 1964: 94, 96
osumiensis Murayama, 1955: 90 A: JA
perlatus Chapuis, 1876: 198 A: FUJ GUI HEN HUN JA NC SC SCH SHX TAI YUN
pertuberculatus Eggers, 1939b: 115 A: TAI
pfefferi Knfzek, 1994: 124 A: CY
phoebe Wood, 1988c: 199 A: UP
pulchellus Blandford, 1894a: 69 A: JA
rudis Blandford, 1894a: 73 Ei: FR NL A: JA SC
shotoensis Murayama, 1955: 88
sannohensis Murayama, 1954: 190 A: GUI JA
seriates Blandford, 1894a: 72 A: JA
shensi Tsai & Yin, 1964: 90, 95 A: SHA SHX YUN
sinensis Schedl, 1953a: 23 A: ANH FUJ GUA GUI GUX HEN HUB HUN JIA JIX SCH SHA SHX YUN ZHE
squamulatus Chapuis, 1869: 39 A: UP "Himalaya"
thujae Perris, 1855: lxvii {Hylesinus) E: AL AU BE BH BU CR CZ FR GB GE GR HU IT MA MC NL PL PT
RO SK SL SP ST SZ UK YU N: AG CI EG LB MO TU A: TR
impressus Olivier, 1795: 78: 12 (Scofytus) [invalid, see Opinion, 1167]
juniperi Döbner, 1860: 261 (Dendroctonus)
prostratus Peyerimhoff, 1918: 259
serrifer Wichmann, 1916: 17
turkestanicus Semenov, 1902: 269 A: KI KZ TD UZ

tribe Phloeotribini Chapuis, 1869

genus Phloeotribus Latreille, 1797: 50 type species Bostrichus oleae Fabricius, 1792 (= Scolytus scarabaeoides
Bernard, 1788)
Comesiella Del Guercio, 1925b: 213 type species Comesiella sicula Del Guercio, 1925 (= Phloeophthorus pubifrons
Guillebeau, 1893)
Elzearius Guillebeau, 1893: 64 type species Elzearius crenatus Guillebeau, 1893
Eulytocerus Blandford, 1897: 161 type species Eulytocerus championi Blandford, 1897
Phloeophthorus Wollaston, 1854: 299 type species Phloeophthorusperfoliatus Wollaston, 1854
Phthorophloeus Rey, 1883: 128 type species Phloeophthorus spinulosus Rey, 1883
brevicollis Kolenati, 1846: 38 (Hylesinus) E: CR FR (Corse) HU ST UK N: MO A: TR
caucasicus Reitter, 1891a: 32 E: AB AU CZ FR GE ST UK YU A: CY KI TM
cristatus Fauvel, 1889: 71 (Phloeophthorus) E: BH CR FR IT SP SZ N: AG CI MR A: TR
geschwindi Seitner, 1920: 283 (Phloeophthorus)
helveticus Guillebeau, 1893:60 (Phloeophthorus)
latus Wichmann, 1916: 14 (Phloeophthorus)
ovalis Eggers, 1941b: 120 (Phloeophthorus)
pectinicornis Balachowsky, 1949a,: 116 (Phloeophthorus)
fraxini Eggers, 1913: 239 (Phloeophthorus) N: AG
 SCOLYTINAE 213
hercegovinensis Seitner, 1920: 284 (Phloeophthorus) E: BH BU CR GR MC SP
liminaris Harris, 1852: 78 (Tomicus) Ei: IT NAR
mississippiensis Blackman, 1921:4 (Phthorophloeus)
maroccanus Guillebeau, 1896: 152 (Phloeophthorus) E: FR N: AG MO
mayeti Guillebeau, 1893: 62 (Phloeophthorus) N: AG A: TR
muricatus Eggers, 1929a: 9 (Phloeophthorus) E: BU HU RO ST UK A: TR
perfoliatus Wollaston, 1854: 301 (Phloeophthorus) E: FR IT SP N: MR
abeillei Guillebeau, 1893: 58 (Phloeophthorus)
peyerimhoffi Eggers, 1913: 239 (Phloeophthorus) E: GR SP N; AG MO
pseudocristatus Pfeffer, 1972: 40 (Phloeophthorus) E: GR A: TR
pubifrons Guillebeau, 1893: 59 (Phloeophthorus) E: AL CR FR GR IT SP N: AG A: CY TR
corsicus Guillebeau, 1893: 60 (Phloeophthorus)
guillebeaui Reitter, 1913b: 35 (Phloeophthorus)
lineigera Guillebeau, 1893: 60 (Phloeophthorus)
sicula Del Guercio, 1925b: 210 (Comesiella)
rhododactylus Marsham, 1802: 58 (Ips) E: AU BE CR CZ DE FI FR GB GE GR HU IR IT LA MC NL NR PL
PT RO SK SP SV SZ N: AG EG LB MO MR TU A: TR
austriacus Guillebeau, 1893: 58 (Phloeophthorus)
retamae Perris, 1864: 300 (Hylesinus)
spartiiNördlinger, 1847: 217 (Hylesinus)
tarsalis Förster, 1849: 383 (Hylesinus)
scarabaeoides Bernard, 1788: 270 (Scolytus) E: BU CR FR GR HU IT MA PT SP SZ UK N: AG CI EG LB
MO TU A: CY IQ IS JO LE SY TR
oleae Fabricius, 1792: 366 (Bostrichus)
oleiphilus Del Guercio, 1925a: 196
sharpi Guillebeau, 1893: 62 (Phloeophthorus) E: FR N: AG
spinulosus Rey, 1883: 127 (Phloeophthorus) E: AU BE BY CT CZ DE EN FI FR GE HU IT LA LS NL NR NT
PL RO SK SP SV SZ UK "Yugoslavia" A: ES FE JA KZ MG WS
chapuisii Blandford, 1891: 213 (Phloeophthorus)
crenatus Guillebeau, 1893: 64 (Elzearius)
perrisi Guillebeau, 1893: 62 (Phloeophthorus)
vinogradowi Semenov, 1902: 269 (Phloeophthorus) E: BU GR ST

tribe Polygraphini Chapuis, 1869

genus Carphoborus Eichhoff, 1864: 27 type species Bostrichus minimus Fabricius, 1798
Estenoborus Reitter, 1913b: 58 type species Hylesinus perrisi Chapuis, 1869
bonnairei C. N. F. Brisout de Barneville, 1884: lii N: AG MO TU A: CY
cholodkovskyi Spessivtsev, 1916: 64 E: CT EN FI NR NT PL SV A: ES FE KZ MG WS
borealis Karpinski, 1933: 28
costatus Wichmann, 1915a: 106 A: HP KA UP "NW Himalayas"
henscheli Reitter, 1887: 192 E: GG A: CY IS SY TR
jurinskii Eggers, 1910a: 36 A: ES MG
marani Pfeffer, 1941b: 177 E: GR HU ST
minimus Fabricius, 1798: 158 (Bostrichus) E: AL AU BH BU BY CR CT CZ EN FI FR GE GG GR HU IT MC
NL NT PL RO SK SL SP ST SZ UK YU N: AG EG LB MO TU A: IS NE TR "SW Siberia"
balgensis Murayama, 1943: 99
squamulatus L. Redtenbacher, 1874: 370
perrisi Chapuis, 1869: 31 (Hylesinus) E: BH BU CR FR GR HU IT RO SP ST UK YU N: AG TU A: CY IN
IQ IS SY TD TM TR UP
abachidsei Stark, 1952: 232
kushkensis Sokanovskiy, 1954: 16
pini Eichhoff, 1881: 38, 131 E: BH CR FR GR HU IT SP N: AG TU A: CY TR
attritus Peyerimhoff, 1932: 274
rossicus Semenov, 1902: 272 E: AU CT FI NT SV A: ES FE WS
taireiensis Murayama, 1943: 98 A: NE
teplouchovi Spessivtsev, 1916: 65 E:CT FI NT SV A: ES FE GAN HEI LIA MG WS
214 SCOLYTINAE
zhobi Stebbing, 1909: 18 (Phloeosinus) A: HP PA

g e n u s Polygraphus Erichson, 1836: 57 type species Bostrichus pubescens Fabricius, 1792 (= Dermestes
poiigraphus Linnaeus, 1758)
Lepisomus Kirby, 1837: 193 type species Apate rufipennis Kirby, 1837
Nipponopolygraphus Nobuchi, 1981a: 12 type species Nipponopolygraphus kaimochii Nobuchi, 1981
Ozophagus Eggers, 1920b: 234 type species Ozophagus camerunus Eggers, 1920 (= Polygraphus primus Wichman, 1915)
Pseudopolygraphus Seitner, 1911: 105 type species Polygraphus grandiclava C. G. Thomson, 1886
Spongotarsus Hagedorn, 1908:372 type species Spongotarsus quadrioculatus Hagedorn, 1908
abietis Kurentsov, 1941: 131, 229 A: FE N E N C
amplifolius Yin & Huang, 1 9 9 6 : 3 4 7 , 3 5 3 A: HUN
angustus Tsai & Yin, 1965: 329, 335 A: SCH
aterrimus Strohmeyer, 1908a: 69 A: HP UP ORR
difficilis Wood, 1988b: 194 A: H P PA UP O R R
formosanus Nobuchi, 1967: 16 A: TAI
fulvipennis Niisima, 1941: 131 A: JA
gracilis Niisima, 1909: 136 A : F E J A J I L
grandiclava C. G. Thomson, 1886b: lxxii E: AU BE BH B U CR C T C Z F R G E H U IT M D N L PL R O SK SL
ST SZ U K Y U
cembrae Seitner, I911: 109
horyurensis Murayama, 1937: 368 A: FE JA N C N E SC TAI "Manchuria"
japonicus Nunberg, 1956: 208 [RN] A: JA
granulatus Niisima, 1941: 125 [HN]
uchimappensis Murayama, 1956a: 282
jezoensis Niisima, 1909: 135 A: FE JA N C
junnanicus Sokanovskiy, 1959a: 93, 95 A: HAI SCH XIZ Y U N O R R
kaimochii Nobuchi, 1981a: 13 {Nipponopolygraphus) A: JA O R R
querci Wood, 1988b: 195
kisoensis Niisima, 1941: 131 A: FE (Sakhalin) JA
longifolia Stebbing, 1903: 255 A: HP UP
himalayensis Stebbing, 1908: 8
major Stebbing, 1903: 234 A: B T HP KA N P PA U P XIZ O R R
meakanensis Niisima, 1935: 2 A: JA
nigrielytris Niisima, 1 9 1 3 : 2 A: FE (Kamchatka, Kuriles, Sakhalin) JA
nobuchii Choo & W o o , 1989: 57 A: SC
parvulus Murayama, 1956a: 279, 283, 291 A: JA
pini Stebbing, 1914: 522 A: H P KA PA UP
minor Stebbing, 1903: 239 [HN]
poiigraphus Linnaeus, 1758: 355 (Dermestes) E: AU BE BH B U B Y C R C T C Z D E E N FI FR G B GE GR HU
IT LA LS LT LU M D NR N T PL RO SK ST SV SZ U K YU A: ES FE G A N JA K Z M G N M O QIN SCH
SHA SHX TR W S XIZ Y U N AFRi
griseus Eggers, 1923b: 136
pubescens Fabricius, 1792: 368 (Bostrichus)
proximus Blandford, 1894a: 75 Ei: CT N T A: ES FE HEI JA JIL N C SC
laticollis Eggers, 1926a: 135
magnus Murayama, 1956a: 279,282,290
miser Blandford, 1894a: 76
nigricans Kurentsov, 1948: 50
oblongus Blandford, 1894a: 75
pterocaryi Yin & Huang, 1996: 345, 352 A: A N H FUJ SHN
punctifrons C. G. Thomson, 1886a: xi E: CT CZ EN FI LA N R N T PL SK SV A: E S F E M G W S
bicolor Sokanovskiy, 1960: 675
krivolutzkianus Stark, 1956: 830
seriatus Reitter, 1913b: 56
rudis hexiensis Yin & Huang, 1996: 348 A: G A N NIN QIN SHA
rudis rudis Eggers, 1933e: 99 A: LIA GAN GUI QIN SCH XIZ Y U N
 SCOLYTINAE 215
likiangensis Tsai & Yin, 1965: 328, 334
retiventriculus Tsai & Yin, 1965: 329, 334
squameus Yin & Huang, 1981: 558, 570
zhungdianensis Tsai & Yin, 1965: 327,334
setosus Schedl, 1979b: 128 A: HP UP
shariensis Niisima, 1941: 127 A: FE (Kuriles, Sakhalin) JA
sinensis Eggers, 1933e: 100 A: SCH SHA SHX
squamosus Schedl, 1975d: 384 (Blastophagus) A: BT
squamulatus Niisima, 1941: 129 A: FE (Sakhalin) JA
ssiori Niisima, 1909: 132 A: FE (Kuriles, Sakhalin) JA LIA
subopacus C. G. Thomson, 1871: 393 E: AU BU CT CZ EN FI FR GE HU LA NR NT PL SK SV SZ YU A:
ES FE HEI JA JIL MG NC SC WS
fontinalis Kurentsov, 1941: 136
minor Lindemann, 1875b: 242
nanus Schedl, 1955a: 22
sachalinensis Eggers, 1926a: 135
szemaoensis Tsai & Yin, 1965: 330, 335 A: GUI SCH YUN ORR
taiwanensis Schedl, 1967a: 128 A: TAI
trenchi Stebbing, 1905: 6 A: PA
verrucifrons Tsai & Yin, 1965: 331, 335 A: SCH YUN XIZ
yunnanensis Bright, 2002: 44 [RN] A: YUN
querci Yin & Huang, 1996: 346, 353 [HN]

supertribe Scolytitae Latreille, 1804

tribe Corthylini LeConte, 1876
subtribe Corthylina LeConte, 1876
genus Gnatharus Wood & Yin, 1986: 463 type species Gnatharus tibetensis Wood & Yin, 1986
tibetensis Wood & Yin, 1986: 463 A: XIZ ORR

genus Gnathotrichus Eichhoff, 1869b: 275 type species Gnathotrychus corthyloides Eichhoff, 1869 (= Tomicus
materiarius Fitch, 1858)
Ancyloderes Blackman, 1938: 205 type species Cryphaluspilosulus LeConte, 1868
Paraxyleborus A. Hoffmann, 1942: 72 type species Xyleborus duprezi A. Hoffmann, 1936 (= Tomicus materiarius Fitch,
1858)
Prognathotrichus Bright, 1972b: 1678 type species Prognathotrichus primus Bright, 1972
materiarius Fitch, 1858: 726 {Tomicus) Ei: AU BE CZ FI FR GE IT NL PL SP SV SZ NAR
corthyloides Eichhoff, 1869b: 275
duprezi A. Hoffmann, 1936: 43 {Xyleborus)

genus Monarthrum Kirsch, 1866: 213 type species Monarthrum chapuisi Kirsch, 1866
Corthylomimus Ferrari, 1867: 48 type species Bostrichus fasciatus Say, 1826
Cosmocorynus Ferrari, 1867: 62 type species Cosmocorynus cristatus Ferrari, 1867
Pterocyclon Eichhoff, 1869b: 276 type species Pterocyclon laterale Eichhoff, 1869
Anchonocerus Eichhoff, 1878b: 67,431 type species Anchonocerus ruflpes Eichhoff, 1878 (= Pterocyclon ingens
Eichhoff, 1869)
Phthorius Eichhoff, 1878b: 67,433 type species Pterocyclon ingens Eichhoff, 1869
Trypocranus Eichhoff, 1878b: 67,435 type species Trypocranus cincinnatus Eichhoff, 1878
Eupteroxylon Eggers, 1936b: 392 type species Eupteroxylon comatum Eggers, 1936 (= Pterocyclon scrobiceps Eichhoff,
1878)
Xyleborips Reitter, 1913b: 111 type species Xyleborus meuseli Reitter, 1905
malt Fitch, 1856: 326 (Tomicus) Ei: IT NAR
longulum Eichhoff, 1869b: 278 (Pterocyclon)
meuseli Reitter, 1905: 249 (Xyleborus) A: ES

subtribe Pityophthorina Eichhoff, 1878

genus Aruptus Eichhoff, 1872a: 136 type species Araptus rufopalliatus Eichhoff, 1872
Brachydendrulus Schedl, 1951c: 114 type species Brachydendrulus eggersi Schedl, 1951
216 SCOLYTINAE
Breviophthorus Schedl, 1938c: 176 type species Breviophthorus brasiliensis Schedl, 1938 (= Neodryocoetes gracilentus
Schedl, 1972)
Gnathoborus Schedl, 1970a: 93 type species Breviophthorus argentiniae Schedl, 1958
Gnathocranus Schedl, 1951c: 116 type species Gnathocranus novateutonicus Schedl, 1951
Neodryocoetes Eggers, 1933b: 9 type species Neodryocoetes hymenaeae Eggers, 1933
Sphenoceros Schedl, 1939f: 565 type species Sphenoceros Umax Schedl, 1939
Thamnophthorus Blackman, 1942: 178 type species Thamnophthorus volastos Schedl, 1938
micrographus Schedl, 1972b: 60 {Breviophthorus) E: LA NTR
micrograptinus Wood, 1989: 179 [RN] (Pityophthorus)

genus Mimiocurus Schedl, 1957b: 72 type species Mimiocurus acuminatus Schedl, 1957
Micracidendron Schedl, 1957b: 71 type species Micracidendron montanum Schedl, 1957
Mimiophthorus Schedl, 1957b: 77 type species Brachydendrulus congonus Schedl, 1952 [as montamis, lapsus, corrected
in Schedl 1963 and 1964]
oleanderi Schedl, 1961a: 187 {Neodryocoetes) Ei: TR

genus Pityophthorus Eichhoff, 1864: 39 type species Bostrichus lichtensteinii Ratzeburg, 1837
Cladoborus Sawamoto, 1942: 165 type species Cladoborus arakii Sawamoto, 1942
Conophthocranulus Schedl, 1935d: 343 type species Conophthocranulus blackmani Schedl, 1935
Ctenyophthorus Schedl, 1955b: 26 type species Ctenyophthorus glabratulus Schedl, 1955
Gnathophorus Schedl, 1935d: 342 [HN] type species Gnathophorus sparsepilosus Schedl, 1935
Gnathophthorus Wood, 1962: 76 [RN] type species Gnathophorus sparsepilosus Schedl, 1935
Hagedornus R. Lucas, 1920: 683 [RN] type species Trigonogenius fallax Hagedorn, 1912
Hypopityophthorus Bright, 1981: 14 type species Pityophthorus inops Wood, 1976
Myeloborus Blackman, 1928: 16 type species Pityophthorus ramiperda Swaine, 1917
Neomips Schedl, 1954a: 37 type species Neomips brasiliensis Schedl, 1954 (^Pityophthorus dimorphus Schedl, 1959)
Pityophthoroides Blackman, 1942: 199 type species Pityophthoroides pudens Blackman, 1942
Trigonogenius Hagedorn, 1912c: 354 [HN] type species Trigonogenius fallax Hagedom, 1912
abietinus Wood, 1989: 179 [RN] A: FE
abietis Kurentsov, 1941: 179, 234 [HN]
kurentzovi Krivolutskaya, 1996: 370 [RN]
sibiricus Nunberg, 1956: 208 [HN]
arakii Sawamoto, 1942: 167 (Cladoborus) A: JA
balcanicus Pfeffer, 1940b: 123 E: AL BH BU MC RO SK YU
buyssoni angeri Pfeffer, 1927: 111 E: FR (Corse)
buyssoni buyssoni Reitter, 1901b: 101 E: BU FR GR IT SP SZ "Yugoslavia"
carniolicus Wichmann, 1910: 145 E: AU CR CZ FR GE HU IT PL SK SL SZ
cedri Wood, 1989: 182 A: HP KA
cephalonicae Pfeffer, 1940b: 119 E: GR PL
polonicus Karpinski, 1949: 125
chilgoza Wood, 1989: 183 A: HP
deodara Stebbing, 1903: 274 (Cryphalus) A: HP KA UP
himalayensis Stebbing, 1914: 540 (Cryphalus)
sampsoni Stebbing, 1914: 551
exsculptus Ratzeburg, 1837: 162 (Bostrichus) E: AU BH BU CZ FR GE HU IT PL SK SZ
macrographus Eichhoff, 1881: 47,200
flavimaculatus Murayama, 1963b: 392 A: "Tairei in North Manchuria"
glabratus Eichhoff, 1878b: 179 E: AU BU BY CR CT CZ DE EN FI FR GE GR HU IT LT LU NO NR NT PL
SK SP ST SV SZ UK YU N: EG A: NO WS
henscheli Seitner, 1887: 44 E: AU BH BU FR GE GR HU IT RO SZ UK
senex Wichmann, 1913b: 143
jucurtdus Blandford, 1894a: 87 A: FE (Sakhalin) JA NC NE SC
kirgisicus Pyatnitskiy, 1931: 167 A: KI KZ
knoteki Reitter, 1898: 356 E: AU BU GE IT PL SZ UK A: YUN
robustus Pfeffer, 1940b: 118
lapponicus Kurentsov, 1941: 177 E: NT A: FE
lapponicus Stark, 1952: 356
 SCOLYTINAE 217
lichtensteinii Ratzeburg, 1837: 162 {Bostrichus) E: AU BE BH BU CT CZ DE EN FI FR GB GE GR HU IT LA
LT LU MC NL NR NT PL SK SL SP ST SV SZ UK YU A: ES FE (Kuriles) FUJ KZ MG NE NO
rossicus Eggers, 1915a: 13
mauretcmicus Peyerimhoff, 1930b: 259 E: FR N: AG EG LB MO TU
micrographus micrographus Linnaeus, 1758: 355 {Dermestes) E: AU BU BY CR CT CZ DE EN FI FR GB GE
GR HU IT LA LT LU NR NT LU PL RO SP SV SZ YU "Caucasus" A: KZ TR WS
curtulus Sokanovskiy, 1954: 19
fennicus Eggers, 1914b: 183
longulus Sokanovskiy, 1954: 19
micrographus sibiricus Stark, 1952: 344 A: ES FE MG NO
morosovi Spessivtsev, 1926a: 48 E: AU CT CZ EN FI LA NT PL SV A: ES HEI LIA WS
parfentievi Pyatnitskiy, 1931: 169 A : KI KZ
schrenkianae Pyatnitskiy, 1931:171
pini Kurentsov, 1941: 176,233 A: ES FE HEI NC "Ural"
pinsapo Pfeffer, 1982: 154 E: SP
pityographus cribratus Pfeffer, 1940b: 116 E: GR ST A: TR
maritimus Stark, 1952: 344
pityographus pityographus Ratzeburg, 1837: 162 {Bostrichus) E: AU BE BH BU CR CZ DE EN FR GE GR
HU IT MC PL RO SK SL SP SV SZ UK YU "Caucasus"
bibractensis Balachowsky, 1949a: 236
pubescens Marsham, 1802: 58 {Ips) E: AU BE CR CZ DE FR GB GE GR HU IR IT NL NR PL SK SL SP SV
SZ N: AG EG LB MO TU A: CY IS TR
ramulorum Penis, 1856: 191 {Tomicus)
sachalinensis Krivolutskaya, 1956: 838 A: FE (Sakhalin)
seiryuensis Murayama, 1963b: 393 A: "Seiryu in South Manchuria"
sichotensis Kurentsov, 1941: 175, 233 A: ES FE
traegardhi Spessivtsev, 1921: 219 E: AU CT EN FI NR NT PL SV A: ES FE MG WS
genus Pityotrichus Wood, 1962: 76 [RN] type species Pityophilus barbatus Blackman, 1928
Pityophilus Blackman, 1928: 147 [HN] type species Pityophilus barbatus Blackman, 1928
turkmenicus Mandelshtam & Petrov, 2006: 214 A: TM

genus Pseudopityophthorus Swaine, 1918: 93 type species Crypturgus minutissimus Zimmermann, 1868
Xenophthorus Wood & Yin, 1986: 462 type species Pseudopityophthorus peregrinus Wood & Yin, 1986
peregrinus Wood & Yin, 1986: 462 A: XIZ

tribe Cryphalini Lindemann, 1876

genus Allernoporus Kurentsov, 1941: 159,230 type species Allernoporus euonymi Kurentsov, 1941
euonymi Kurentsov, 1941a: 159, 230 A: FE NC SC

genus Coriacephilus Schedl, 1939c: 339 type species Stephanoderes coriaceus Eichhoff, 1878
coriaceus Eichhoff, 1878b: 494 {Stephanoderes) A: TAI ORR
birmanus Eggers, 1925: 156 {Cryphalus)

genus Cosmoderes Eichhoff, 1878a: 387 [= 1878b: 495] type species Cosmoderes monilicollis Eichhoff, 1878
Dendriops Schedl, 1953c: 125 type species Dendriops granulicollis Schedl, 1953 (= Cosmoderes monilicollis Eichhoff,
1878)
Erioschidias Wood, 1960a: 21 type species Cryphalus setistriatus Lea, 1910
Pseudocosmoderes Nobuchi, 1981a: 16 type species Pseudocosmoderes attenuates Nobuchi, 1981
attenuatus Nobuchi, 1981a: 17 {Pseudocosmoderes) A: TAI ORR
consobrinus Blandford, 1894a: 86 A: JA TAI
monilicollis Eichhoff, 1878a: 387 [=1878b: 496] A: GUI SD TAI ORR
coriaceus Schedl, 1959d: 474 {Erioschidias)
granulicollis Schedl, 1953c: 125 {Dendriops)
218 SCOLYTINAE
genus Cryphalus Erichson, 1836: 61 type species Bosfrichus asperatus Gylienhal, 1813
Allarthrum Hagedorn, 1912c: 355 type species Allarlhrum kolbei Hagedorn, 1912
Cryptarthrum Blandford, 1896b: 200 type species Cryptarthrum walked Blandford, 1896
Ericryphalus Hopkins, 1915a: 38 type species Ericryphalus henshawi Hopkins, 1915 (= Hypothenemus sylvicola Perkins,
1900)
Ernocryphalus Murayama, 1958: 934 type species Ernocryphalus birosimensis Murayama, 1958
Piperius Hopkins, 1915a: 39 type species Piperiuspini Hopkins, 1915 (= Hypothenemus sylvicola Perkins, 1900)
Pseudocryphalus Ferrari, 1869: 252 type species Bostrichus sidneyanus Nördlinger, 1856
Taenioglyptes Bedel, 1888: 398 type species Bostrichuspiceae Ratzeburg, 1837
abbreviates Schedl, 1943: 35 A: JA
aquilonius Nobuchi, 1975: 52 {Taenioglyptes) A: JA
asperatus Gylienhal, 1813: 368 (Bostrichus) E: AU BE BH BU BY CT CZ CR DE EN FI FR GB GE GR HU
IR IT LA LT LU MC NL NR NT PL RO SK SL SP ST SV SZ YU "Caucasus" N: AG MO A: ES FE JA
NC TR
abietis Ratzeburg, 1837: 163 (Bostrichus)
babaiMurayama, 1961:29 A.JA
bicarinatus Nobuchi, 1975: 53 (Taenioglyptus) A: JA
birosimensis Murayama, 1958: 935 (Ernocryphalus) A: JA
capucinus Schedl, 1938e: 425 A: "China"
carpini Berger, 1917: 234 A: FE JA SC
carpinivorus Murayama, 1930: 14 A: SC
chamaecipariae Niisima, 1910: 10 A: JA
chinlingensis Tsai & Li, 1963: 612, 623 A: SCH SHA SHX
coryli Stark, 1936b: 144 A: FE NC
cryptomeriae Niisima, 1908: 91 A: JA
dissimilis Nobuchi, 1975: 57 (Taenioglyptes) A: JA
dorsalis Motschulsky, 1866: 403 (Hypoborus) A: HAI ORR
indicus Eichhoff, 1878a: 384 [= 1878b: 489]
nebulosus Motschulsky, 1866: 403 (Hypoborus)
sericeus Motschulsky, 1866: 402 (Hylesinus)
exiguus Blandford, 1894a: 82 A: ANH BEI FE (Kuriles) GUI HEB JA JLA NC SC SCH SHA SHN TAI ZHE
pilosus Sasaki, 1899:238
felis Wood, 1989: 180 A: UP
ficivorus Murayama, 1958: 933 A: JA
formosanus Schedl, 1942b: 175 A: TAI
fulmineus Wood, 1989: 180 A: HP SCH UP
fulvus Niisima, 1908: 92 A: JA LIA NC NE NO SC SHN TAI
pini Eggers, 1921: 39
garciniae Nobuchi, 1959b: 24 A: JA (Ryukus)
hirsutus Nobuchi, 1975: 52 (Taenioglyptes) A: JA
intermedius Ferrari, 1867: 79 E: AU CZ GE HU IT LS PL RO SK SL SZ UK
jeholensis Murayama, 1939: 143 A: HEB JA SC "Manchuria"
jezoensis Inouye & Nobuchi, 1957:49 A: JA
jugransiNiisima, 1913: 3 A: JA
kagoshimensis Nobuchi, 1975: 56 (Taenioglyptes) A: JA
kurenzovi Stark, 1936b: 150 A: FE NC
punctulatus Eggers, 1942: 29
ussuriensis Eggers, 1942: 29
kyotoensis Nobuchi, 1966: 53 A: JA
laricis Niisima, 1909: 142 A: FE (Sakhalin) JA SC
latus Eggers, 1929a: 10 A: ES FE HEI JIL NC
premayaensis Murayama, 1943: 97
lepocrinus Tsai & Li, 1963: 619, 624 A: SCH SHA YUN
lipingensis Tsai & Li, 1959: 90 A: GUI HAI SCH SHA SHX YUN
longisetosus Nobuchi, 1975: 57 (Taenioglyptes) A: JA
longus Eggers, 1926a: 136 (Ernoporus) A: FE JA
 SCOLYTINAE 219
alni Krivolutskaya, 1958: 145
major Stebbing, 1903: 270 A: HP UP ORR
morinda Stebbing, 1903: 265
malus Niisima, 1909: 144 A: FE HEI JA LIA NC SC
padi Krivolutskaya, 1958: 144
mandschuricus Eggers, 1929a: 10 A: FE HEI NC
markangensis Tsai & Li, 1963: 618, 624 A: HUB JIA SCH YUN
massonianus Tsai & Li, 1963: 613, 623 A: HEN HUN JIA SCH YUN
meridionalis Nobuchi, 1975: 55 (Taeniogfyptes) A: GUI HUN JA
miyalopiceus Tsai & Li, 1963: 608, 622 A: SCH XIZ YUN
montanus Nobuchi, 1964: 129 A: JA
niponensis Inouye & Nobuchi, 1957: 51 A: JA
numidicus Eichhoff, 1878a: 385 [=1878b: 487] E: BU FR GR IT SP SZ N: AG EG LB MO TU A: TR
parvulus Niisima, 1910: 8 A: JA
piceae Ratzeburg, 1837: 163 (Bostrichus) E: AU BH BU BY CR CT CZ FR GE GR HU IT LA LT MC PL RO
SK SL SP ST SZ UK YU N: AG A: FE JA NC NE SC TR "Manchuria"
hattorii Kôno, 1938:67
orientalis Eggers, 1911b: 122
subdepressus Eggers, 1940f: 37
piceus Eggers, 1926a: 136 A: FE HEI JANE "Korea"
pilosus Tsai & Li, 1963: 617, 624 A: SCH YUN
pruni Eggers, 1929a: 11 A: FE NC YUN
pseudochinlingensis Tsai & Li, 1963: 610, 623 A: SHA SHX
pseudotabulaeformis Tsai & Li, 1963: 613, 624 A: BEI HEB SHA SCH
pulchellus Nobuchi, 1975: 54 (Taeniogfyptes) A: JA
redikorzevi Berger, 1917: 232 A: FE HUB NC SC SCH YUN
rhusii Niisima, 1909: 145 A: FE (Kuriles) JA NC
kurilensis Krivolutskaya, 1968: 53
saltuarius Weise, 1891: 336 [RN NP] E: AU BU BY CT CZ DE EN FI GE HU IT LS NR NT PL SK SV SZ
YU "Caucasus" A: ES FE GUX SCH WS
scriba Gozis, 1886: 31 [RN NO]
asperatus Ratzeburg, 1837: 163 [HN] [Bostrichus)
samoensis Beeson, 1929: 224 A: HAI ORR
sawadai Nobuchi & Takahashi, 1965: 3 A: JA
scopiger Berger, 1917: 228 A: FE HEI JA NC SC
sichotensis Kurentsov, 1941: 152, 230 A: FE JA
sinoabietis Tsai & Li, 1963: 606, 622 A: GAN SCH YUN
opienensis Tsai & Li, 1963: 607, 622
sordidus Nobuchi, 1975: 51 (Taeniogfyptes) A: JA
strohmeyeri Stebbing, 1914: 538 A: KA SCH SD UP XL2 YUN
indicus Stebbing, 1906: 403
substriatus Schedl, 1942b: 174 A: YUN ORR
szechuanensis Tsai & Li, 1963: 614, 624 A: SCH
tehchangensis Tsai & Li, 1963: 615, 624
tabulaeformis Tsai & Li, 1963: 609, 623 A: GUI HEB LIA SCH SHA SHX YUN ZHE
chienzhuangensis Tsai & Li, 1963: 610, 623
taiwanus Schedl, 1970c: 359 A: TAI
viburni Stark, 1936b: 151 A: FE SHA SHX
viburni Eggers, 1942:30
yamaguchii Inouye & Nobuchi, 1957: 55 A: JA

genus Eidophelus Eichhoff, 1876: 200 type species Eidophelus imitons Eichhoff, 1876
Phellodendrophagus Krivolutskaya, 1958: 107,150 type species Phellodendrophagus elegans Krivolutskaya, 1958 (=
Eidophelus imitans Eichhoff, 1876)
Idophelus Rye, 1877: 362 [unjustified emendation]
imitans Eichhoff, 1876: 201 A: FE JA SC TAI ORR
220 SCOLYTINAE
elegans Krivolutskaya, 1958: 150 (Phellodendrophagus)
nitidus Schedl, 1959d: 475 (Ptilopodius)
minutus Blandford, 1894a: 88 A: JA

genus Ernocladius Wood, 1980: 93 type species Cryphalus corpulentus Sampson, 1919
corpulent™ Sampson, 1919: 113 (Cryphalus) A: JA ORR
sundri Schedl, 1969b: 48 (Margadillius)

genus Ernoporicus Berger, 1917: 242 type species Ernoporicus spessivtzevi Berger, 1917
Eocryphalus Kurentsov, 1941: 161,231 type species Eocryphalus semenovi Kurentsov, 1941
Ernopocerus Wood, 1954: 986 type species Ernoporus caucasicus Lindemann, 1876
alniphagus Nobuchi, 1975: 43 A: JA
ater Nobuchi, 1975: 42 A: JA
caucasicus Lindemann, 1876a: 373 (Ernoporus) E: AB AU BE CZ DE FI FR GB GE GR HU IT NR PL RO SK
SL SP ST SV SZ UK A: TR
schreineri Eichhoff, 1881: 46, 185 (Cryphalus)
corni Kurentsov, 1941: 162, 232 (Hypothenemus) A: FE NC
fagi Fabricius, 1798: 157 (Apate) E: AL AU BE BU CR CT CZ DE FR GB GE GR HU IT MC NL NR PL RO
SK SP ST SV SZ UK YU A: TR
serratus Panzer, 1795: 288 (Bostrichus) [HN]
thomsoni Ferrari, 1867: 14 (Cryphalus)
insularum Krivolutskaya, 1968: 56 (Hypothenemus) A: FE (Kuriles) JA
elongatus Nobuchi, 1975: 42 (Ericryphalus)
krivolutskayae Wood, 1992: 79 (Hypothenemus) [unnecessary RN]
semenovi Kurentsov, 1941: 161,231 (Eocryphalus) A: FE
spessivtzevi Berger, 1917: 243 A: FE
takahashii Nobuchi, 1975: 43 A: JA
zachvatkini Krivolutskaya, 1958: 149 A: FE

genus Ernoporus C. G. Thomson, 1859:147 type species Apate tiliae Panzer, 1793
Cryphalops Reitter, 1889b: 94 type species Cryphalus lederi Reitter, 1889 (= Apate tiliae Panzer, 1793)
Euptilius Schedl, 1940e: 589 type species Ernoporus concentralis Eggers, 1936
Stephanorhopalus Hopkins, 1915a: 35 type species Stephanorhopalus nulodori Hopkins, 1915 (= Stephanorhopalus
melodori Hopkins, 1915)
acanthopanaxi Niisima, 1913: 4 (Cryphalus) A: J A
formosanus Browne, 1981a: 129 (Ptilopodi™) A: TAI
japonic™ Nobuchi, 1966: 52 A: JA
shimanensis Murayama, 1953a: 36 A: JA
tiliae Panzer, 1793: 14 (Apate) E: AU BE BU BY CR CT CZ DE EN FI FR GB GE GR HU IT LA NR NT PL
RO SK SL SP ST SV SZ UK YU "Caucasus" A: FE NC SC TR
eggersi Kurentsov, 1941: 155
lederi Reitter, 1889b: 93 (Cryphalus)
ratzeburgi Ferrari, 1867: 11 (Craphalus)
starki Eggers, 1942: 31

genus Hypocryphalus Hopkins, 1915a: 41 type species Hypocryphalus rotundus Hopkins, 1915
Dacryphalus Hopkins, 1915a: 42 type species Dacryphalus obesus Hopkins, 1915
formosanus Schedl, 1952b: 62 A: TAI
mangiferae Stebbing, 1914: 542 (Cryphalus) A: TAI UP AFR AUR NAR NTR ORR
griseus Blackburn, 1885: 194 (Hypothenemus)
inops Eichhoff, 1872a: 131 (Cryphalus)
mangiferae Eggers, 1928a: 85
mimicus Schedl, 1942a: XT {Cryphalus)
opacus Schedl, 1942a: 20
robustus Eichhoff, 1872a: 131 (Cryphalus)
subcylindricus Schedl, 1942a: 16 (Cryphalus)
perminimus Schedl, 1942a: 13 (Cryphalus) A: JA (Ryukyus)
 SCOLYTINAE 221
constrictus Schedl, 1942c: 174 (Cryphalus)
constrictus Schedl, 1942a: 22
froggatti Nunberg, 1961: 611
scabricollis Eichhoff, 1878b: 491 {Cryphalus) Ei: MA A: HAI UP YUN ORR
brevisetosus Schedl, 1943: 36 {Cryphalus)
dilutus Eichhoff, 1878a: 384 [=1878b: 490] {Cryphalus)
discretus Eichhoff, 1878a: 385 [= 1878b: 490] {Cryphalus)

genus Hypothenemiis Westwood, 1834: 35 type species Tomtcus eruditus Westwood, 1834
Adiaeretus Hagedorn, 1909: 744 type species Adiaeretus spinosus Hagedorn, 1909 (= Stephanoderes elephas Eichhoff,
1872)
Archeophalus Schedl, 1941c: 392 type species Archeophalus natalensis Schedl, 1941
Chondronoderes Schedl, 1940e: 589 type species Stephanoderes magnus Eggers, 1924
Ernophloeus Nunberg, 1958: 484 type species Emophloeus costalimai Nunberg, 1958 (= Stephanoderes fitscicollis
Eichhoff, 1878)
Homoeocryphalus Lindemann, 1876b: 168 type species Stephanoderes ehlersi Lindemann, 1876 (= Tomicus eruditus
Westwood, 1834)
Lepiceroides Schedl, 1957b: 59 type species Lepiceroides aterrimus Schedl, 1957 (= Hypothenemus aterrimulus Wood,
1989)
Macrocryphalus Nobuchi, 1981a: 14 type species Macrocryphalus oblongus Nobuchi, 1981
Pachynoderes Schedl, 1941c: 393 type species Pachynoderes deprecator Schedl, 1941
Stephanoderes Eichhoff, 1872a: 132 type species Stephanoderes chapuisii Eichhoff, 1872 (= Crypturgus dissimilis
Zimmermann, 1868)
Stylotentus Schedl, 1963: 448 type species Hypothenemus concolor Hagedorn, 1909
Triarmocerus Eichhoff, 1878a: 383[=1878b: 119] type species Triarmocerus cryphaloides Eichhoff, 1878
amakusanus Murayama, 1934b: 287 {Stephanoderes) A: JA
areccae Hornung, 1842: 117 {Bostrichus) A: TAI U P Y U N A F R A U R N A R N T R O R R
bambesanus Eggers, 1940c: 232 {Stephanoderes)
bauhaniae Schedl, 1950a: 19
capital is Beeson, 1935a: 102
depressus Eichhoff 1878b: 155 [RN] {Stephanoderes)
eupolyphagus Beeson, 1940: 193
fimgicola Eggers, 1908d: 216 (Stephanoderes)
heterolepis Costa Lima, 1928: 117
hispidus Eggers, 1925: 156 {Stephanoderes)
martiniquensis Eggers, 1941a: 99 {Stephanoderes)
oahuensis Schedl, 1941b: 110
obscurus Eichhoff 1872a: 133 [HN] {Stephanoderes)
occidentalis Schedl, 1954b: 76 (Stephanoderes)
polyphagus Eggers, 1924: 104 (Stephanoderes)
subglabratus Schedl, 1942c: 174
subvestitus Eggers, 1940c: 232 (Stephanoderes)
vafer Blandford, 1896c: 241
arundinis Eichhoff, 1878a: 386 [=1878b: 157] {Stephanoderes) E: IT
bezazlani Peyerimhoff, 1935: 192 N: A G
birmanus Eichhoff, 1878a: 384 [=1878b: 486] {Triarmocerus) A: GUI G U X HKG H U N JA PA SCH TAI U P
YUN A U R N A R N T R O R R
alter Eggers, 1923a: 219 (Stephanoderes)
ampliatus Eggers, 1936c: 627 (Stephanoderes)
castaneus Wood, 1954: 1027 (Stephanoderes)
farinosus Blandford, 1896c: 241
maculicollis Sharp, 1879: 101
nibarani Beeson, 1933: 10 (Stephanoderes)
pacificus Beeson, 1940: 197 (Stephanoderes)
peritus Blandford, 1894a: 84
perkinsi Hopkins, 1915a: 31 (Stephanoderes)
psidii Hopkins, 1915a: 32 (Stephanoderes)
sterculiae Hopkins, 1915a: 32 (Stephanoderes)
uter Eggers, 1923a: 219 (Stephanoderes)
valens Sampson, 1914: 385
222 SCOLYTINAE
californiens Hopkins, 1915a: 19 A: IS SC AFR NAR NTR
thoracicus Hopkins, 1916: 598
tritici Hopkins, 1915a: 19
zeae Schedl, 1973b: 169 (Stephanoderes)
crudiae Panzer, 1791: 37 (Bostrichus) E: AZ A: FUJ GUI SCH SD TAI YUN AFR NAR NTR ORR
brasiliensis Hopkins, 1915a: 26 (Stephanoderes)
differens Hopkins, 1915a: 25 {Stephanoderes)
fallax Costa Lima, 1924:316 (Stephanoderes)
guatemalensis Hopkins, 1915a: 26 (Stephanoderes)
hispidulus LeConte, 1868: 156 (Cryphalus)
hivaoea Beeson, 1935a: 105 (Stephanoderes)
largipennis Toleda Piza, 1924b: 354 (Stephanoderes)
lebronneci Beeson, 1935a: 104 (Stephanoderes)
lecontei Hopkins, 1915a: 27 (Stephanoderes)
mucronifer Wollaston, 1867: 116 (Cryphalus)
nanus Hagedorn, 1909: 744
paraguayensis Hopkins, 1915a: 26 (Stephanoderes)
polyphagus Costa Lima, 1924: 316 (Stephanoderes)
trinitatis Hopkins, 1915a: 28 (Stephanoderes)
uniseriatus Eggers, 1924: 103 (Stephanoderes)
dolichocola Hopkins, 1915a: 19 A: GUA
erectus LeConte, 1876: 356 A: ANH GAN GUI HEB HEN HUB JIA LIA SCH SHA SHN SHX TR YUN
AFR NAR
brunneicollis Hopkins, 1915a: 33 (Stephanoderes)
cubensis Hopkins, 1915a: 32 (Stephanoderes)
discedens Schedl, 1950a: 23 (Stephanoderes)
puncticollis Hopkins, 1915a: 32 (Stephanoderes)
validus Blandford, 1904: 228
eruditus Westwood, 1834: 34 (Tomicus) E: AB AZ CR FR IT MA SP ST TR UK N: AG CI EG MO MR A:
FUJ GUA GUI GUX HEB HUN IN IS JA SC SCH SD SHN TAI TR UP YUN AFR AUS NAR NTR
ORR
argentinensis Schedl, 1939e: 408
asaroriensis Beeson, 1940: 195
asiminae Hopkins, 1915a: 16
aspericollis Wollaston, 1860b: 365 (Cryphalus)
basjoo Niisima, 1910: 9 (Cryphalus)
bicolor Eggers, 1920b: 241
bicolor Schedl, 1939a: 32 [HN]
boieldieui Perroud, 1864: 188 (Bostrichus)
bradfordi Hopkins, 1915a: 15
citri Ebeling, 1935:21
communis C. F. C. Schaufuss, 1891: 11 (Stephanoderes)
cylindricus Schedl, 1939e: 409
cylindripennis Schedl, 1957b: 51
dubiosus Schedl, 1940b: 207
ealaensis Eggers, 1944b: 94 (Archeophalus)
ehlersi Lindemann, 1876b: 168 (Homoeocryphalus)
elongatus Hopkins, 1915a: 25 (Stephanoderes)
erythrinae Eggers, 1936c: 628 (Stephanoderes)
evonymi Hopkins, 1915a: 26 (Stephanoderes)
ferrugineus Hopkins, 1915a: 20
flavicollis Hopkins, 1915a: 24 (Stephanoderes)
flavipes Hopkins, 1915a: 18
flavosquamosus Hopkins, 1915a: 15
germari Eichhoff, 1878a: 386 [=1878b: 159] (Stephanoderes)
glabratellus Schedl, 1953d: 292
glabratus Schedl, 1942b: 175
gracilis Eggers, 1929b: 51 [RN] (Stephanoderes)
hamamelidis Hopkins, 1915a: 16
heathi Hopkins, 1915a: 20
 SCOLYTINAE 223
insularis Perkins, 1900: \%\
intersetosus Eggers, 1928a: 85 (Stephanoderes)
juglandis Blackman, 1922: 88
koebelei Hopkins, 1915a: 17
lezhavai Pyatnitskiy, 1929: 8
lineatifrons Hopkins, 1915a: 17
mali Hopkins, 1915a: 17
m aniens is Schedl, 1941b: 110
mesoleius Schedl, 1959d: 480
minutulus Schedl, 1972e: 225
minutus Schedl, 1978b: 299 {Cryphalus)
myristicae Hopkins, 1915a: 16
myrmedon Eichhoff, 1878b: 160 (Stephanoderes)
myrmidon Eichhoff, 1878a: 386 (Stephanoderes)
nanulus Schedl, 1949b: 263 (Stephanoderes)
nigricollis Hopkins, 1915a: 16
nigripennis Hopkins, 1915a: 19
obscuriceps Schedl, 1952e: 449
obscurus Ferrari, 1867: 17 [HN] (Cryphalus)
parcius Schedl, 1957b: 49
parilis Schedl, 1951c: 100
parvus Hopkins, 1915a: 17
pruni Hopkins, 1915a: 16
punctifrons Hopkins, 1915a: 18
punctipennis Hopkins, 1915a: 20
pusillus Eggers, 1927c: 173
pygmaeus Hopkins, 1915a: 24 (Stephanoderes)
rotroui Peyerimhoff, 1919b: 255
rumseyi Hopkins, 1915a: 16
sacchari Hopkins, 1915a: 17
schwarzi Hopkins, 1915a: 11 (Cosmoderes)
striatopunctatus Lea, 1910: 142 (Cryphalus)
subconcentralis Hopkins, 1915a: 25 (Stephanoderes)
subcylindricus Schedl, 1940c: 233 (Stephanoderes)
tantillus Lea, 1910: 142 (Cryphalus)
tectonae Stebbing, 1903: 263 (Cryphalus)
tenuis Hopkins, 1915a: 16
tigrensis Schedl, 1952e: 452 (Stephanoderes)
tuberculosus Hagedorn, 1912b: 339
unicolor Hopkins, 1915a: 25 (Stephanoderes)
vianai Schedl, 1958b: 42
webbi Hopkins, 1915a: 17
furukawai Murayama, 1934a: 59 (Cryphalus) A: JA SC
fuscicollis Eichhoff, 1878a: 386 [=1878b: 148] (Stephanoderes) A: JA TAI AFR NAR NTR ORR
aequaliclavatus Schedl, 1939a: 33
costalimai Nunberg, 1958: 486 (Ernophloeus)
ghanaensis Schedl, 1962a: 67
sundaensis Eggers, 1927d: 396 (Stephanoderes)
glabripennis Hopkins, 1915a: 32 (Stephanoderes) A: GUX ORR
hampei Ferrari, 1867: 11, 12 (Cryphalus) A: IN TAI AFR AUR NAR NTR ORR
cofeicola Campos Novaes, 1922: 67 (Xyleborus)
coffeae Hagedorn, 1910b: 1 (Stephanoderes)
coffeivorus Weele, 1910: 308 (Xyleborus)
cooki Hopkins, 1915a: 27 (Stephanoderes)
glabellus Schedl, 1952e: 452 (Stephanoderes)
punctatus Eggers, 1924: 101 (Stephanoderes)
ingens Schedl, 1942a: 18 (Stephanoderes) A: GUA GUI GUX YUN ORR
grandis Schedl, 1969b: 49 (Cryphalomorphus)
japonicus Niisima, 1910: 10 (Cryphalus) A: JA
224 SCOLYTTNAE
javanus Eggers, 1908d: 216 {Stephanoderes) A: GUI GUX HAI HUB HUN SCH TAI YUN AFR NAR NTR
ORR
bananensis Eggers, 1922c: 167 (Stephanoderes)
kalshoveni Schedl, 1939a: 35 (Stephanoderes)
ohesus Hopkins, 1915a: 30 (Stephanoderes)
philippinensis Hopkins, 1915a: 31 (Stephanoderes)
pistor Schedl, 1951c: 102 (Stephanoderes)
prosper Schedl, 1951c: 103 (Stephanoderes)
subagnatus Eggers, 1940b: 101 (Stephanoderes)
kraunhiae Murayama, 1950a: 50 (Cryphalus) A: JA
leprieurii Perris, 1866: 194 (Dryocoetes) E: IT MA N: AG A: CY IS SY TR
albipilis Reitter, 1887: 195 (Stephanoderes)
kraussei Wichmarm, 191 Ib: 210
mateui Schedl, 1965a: 198 (Stephanoderes) A: IS
oblongus Niisima, 1910: 9 (Cryphalus) A: JA
nobuchü Knízek, nom. nov. [see New Acts] A: JA
oblongus Nobuchi, 1981a: 14 (Macrocryphalus) [HN]
sapporoensis Niisima, 1910: 3 (Cryphalus) A: JA
schedli Browne, 1963: 54 [RN] A: TAI
pubescens Schedl, 1942a: 18 [HN] (Stephanoderes)
seoulensis Choo & Woo, 1989: 58 A: SC
seriatus Eichhoff, 1872a: 133 (Stephanoderes) N: EG A: TAI TR AFR AUR NAR NTR ORR
andersoni Wood, 1954: 1045 (Stephanoderes)
asperatus Schedl, 1967b: 226 (Stephanoderes)
cassavaensis Schedl, 1938d: 453
darwinensis Schedl, 1942c: 178 (Stephanoderes)
ficus Hopkins, 1915a: 28 (Stephanoderes)
fiebrigi Hopkins, 1915a: 27 (Stephanoderes)
floridensis Hopkins, 1915a: 27 (Stephanoderes)
georgiae Hopkins, 1915a: 26 (Stephanoderes)
hawaiiensis Schedl, 1941b: 112 (Stephanoderes)
liquidambaraeWooá, 1954: 1046 (Stephanoderes)
lucasi Hopkins, 1915a: 28 (Stephanoderes)
marovoayi Schedl, 1953b: 81
minutus Hopkins, 1915a: 26 (Stephanoderes)
niger Hopkins, 1915a: 31 (Stephanoderes)
nitidipennis Hopkins, 1915a: 29 (Stephanoderes)
nitidulus Hopkins, 1915a: 29 (Stephanoderes)
pecanis Hopkins, 1915a: 29 (Stephanoderes)
pini Hopkins, 1915a: 27 (Stephanoderes)
pulverulentas Eichhoff, 1872a: 133 (Stephanoderes)
robustus Blackman, 1922: 88
salicis Hopkins, 1915a: 27 (Stephanoderes)
soltaui Hopkins, 1915a: 28 (Stephanoderes)
striatulus Schedl, 1942a: 12
subopacicollis Hopkins, 1915a: 30 (Stephanoderes)
tamarindi Hopkins, 1915a: 27 (Stephanoderes)
texanus Hopkins, 1915a: 26 (Stephanoderes)
virentis Hopkins, 1915a: 28 (Stephanoderes)
vulgaris C. F. C. Schaufuss, 1898b: 209 (Stephanoderes)
setosus Eichhoff, 1868b: 391 (Hypoborus) A: TAI AFR NAR NTR ORR
congonus Hagedorn, 1912b: 337 (Stephanoderes)
simoni Reitter, 1887: 194 (Stephanoderes) A: IS SY
taihokuensis Schedl, 1952b: 63 (Stephanoderes) A: JA TAI
cosmoderoides Murayama, 1961: 30
tredli Reitter, 1908a: 55 (Cryphalus) N: EG
tristis Eichhoff, 1876: 200 (Stephanoderes) A: JA
apatoides Eichhoff, 1876: 201 (Dryocoetes)
 SCOLYTINAE 225
%
genus Procryphalus Hopkins, 1915a: 33 type species Procryphalus populi Hopkins, 1915 (= Cryphalus
mucronatus LeConte, 1879)
fraxini Berger, 1917: 238 (Emoporus) A: FE HEI JIL NC SC

genus Scolytogenes Eichhoff, 1878a: 387 [=1878b: 497] type species Scolytogenes darwini Eichhoff, 1878
Cryphalomorphus C. F. C. Schaufuss, 1891: 12 type species Cryphalomorphus communis C. F. C. Schaufuss, 1891
Cryphalophilus Schedl, 1970c: 358 type species Cryphalophilus afer Schedl, 1970
Ernoporides Hopkins, 1915a: 34 type species ErnoporidesfloridensisHopkins, 1915 (= Bostrichus jalappae Letzner,
1849)
Hypothenoides Hopkins, 1915a: 11 type species Hypothenoides parvus Hopkins, 1915
Lepicerinus Hinton, 1936: 473 [RN] type species Lepicerus aspericollis Eichhoff, 1878
Lepicerus Eichhoff, 1878a: 388 [= 1878b: 501] [HN] type species Lepicerus aspericollis Eichhoff, 1878
Lepidocerus Rye, 1880: 103 type species Lepicerus aspericollis Eichhoff, 1878
Letznerella Reitter, 1913b: 68 type species Bostrichus jalappae Letzner, 1849
Negritus Eggers, 1923a: 141 type species Negritus ater Eggers, 1923
Neocryphalus Eggers, 1922c: 169 type species Neocryphalus usagaricus Eggers, 1922
Scolytogenes Eichhoff, 1878b: 497 type species Scolytogenes darwinii Eichhoff, 1878b: 497 (= Scolytogenes darwini
Eichhoff, 1878)
Xylocryptus Schedl, 1975b: 352 type species Xylocryptuspapuanus Schedl, 1975 (= Scolytogenespapuensis Wood, 1989)
aspericollis Eichhoff, 1878a: 388 [=1878b: 501] (Lepicerus) A: SD AFR ORR
stierlini Eggers, 1911b: 121 (Cryphalus)
badins Nobuchi, 1975: 44 (Cryphalomorphus) A: JA
camelliae Nobuchi, 1975: 45 (Cryphalomorphus) A: JA
candidus Nobuchi, 1975: 47 (Cryphalomorphus) A: JA
darwini Eichhoff, 1878a: 387 A: UP YUN ORR
darwinii Eichhoff, 1878b: 497
cryptolepis Schedl, 1951 b: 55
major Eggers, 1927e: 69 (Negritus)
similis Eggers, 1923a: 142 (Negritus)
exilis Yin, 2001: 327 A: GUA
expers Blandford, 1894a: 85 (Hypothenemus) A: JA
fujisanus Nobuchi, 1975: 46 (Cryphalomorphus) A: JA
fulvipennis Nobuchi, 1975: 47 (Cryphalomorphus) A: JA TAI
glabratus Yin, 2001: 330 A: HUB YUN
incultus Yin, 2001: 325 A: JIA
indiens Wood, 1989: 184 A: KA UP ORR
insularis Nobuchi, 1975: 48 (Cryphalomorphus) A: JA (Mikura Is.)
jalappae Letzner, 1849: 99 (Bostrichus) Ai: JA NAR NTR
knabi Hopkins, 1915a: 34 (Ernoporides)
alienus Schedl, 1976: 65 (Cryphalomorphus)
caraibicus Schedl, 1951c: 96 (Cryphalomorphus)
floridensis Hopkins, 1915a: 34 (Ernoporides)
minutissimus Schedl, 1951c: 97 (Cryphalomorphus)
ritchiei Sampson, 1918: 295 (Hypothenemus)
subtriatus Schedl, 1952d: 360 (Cryphalomorphus)
magnocularis Yin, 2001: 326 A: YUN
polisquamosus Yin, 2001: 329 A: FUJ
puerarae Choo & Woo, 1989: 58 A: SC
punctatulus Nobuchi, 1976: 72 [RN] A: JA
nobuchii Schedl, 1980a: 118 [RN] (Cryphalomorphus)
punctatus Nobuchi, 1975: 49 [HN] (Cryphalomorphus)
punctatus Schedl, 1951b: 55 (Cryphalomorphus) A: SD
scolytomimoides Nobuchi, 1975: 50 (Cryphalomorphus) A: JA
venustus Yin, 2001: 328 A: GUA
yunnanensis Yin, 2001: 324 A: YUN
226 SCOLYTINAE
genus Trischidias Hopkins, 1915a: 12 type species Trischidias georgiae Hopkins, 1915
atomus Hopkins, 1915a: 15 (Hypothenemus) A: TAI NTR
impressifrons Hopkins, 1915a: 15
marylandicae Hopkins, 1915a: 15
robiniae Hopkins, 1915a: 15
toxicodendri Hopkins, 1915a: 15

genus Trypophloeus Fairmaire, 1864: 105 type species Bostrichus binodulus Ratzeburg, 1837
Glvptoderes Eichhoff, 1878b: 34 type species Bostrichus binodulus Ratzeburg, 1837
alni Lindemann, 1875a: 136 (Cryphalus) E: AU BY CT EN FI GE IT NR NT PL SZ UK A: SCH WS XIN
holdhausi Wichmann, 1912: 186
binodulus Ratzeburg, 1837: 163 {Bostrichus) E: AU BE BH BU BY CR CT CZ DE EN FI FR GB GE HU IR
IT LA MC NL NR NT PL RO SK SL SP SV SZ UK YU N: AG MO A: FE JA NC WS
berezinae Stark, 1952: 285
grothii Hagedorn, 1904b: 232 (Cryphalus)
kurenzovi Nunberg, 1956: 208 [RN]
kurenzowi Schedl, 1959b: 42
populi Kurentsov, 1941: 164,231 [HN]
spiculatus Eggers, 1927b: 122
bispinulus Eggers, 1927b: 121 E: CT EN FI LA NR NT SV A: ES FE MG WS
dejevi Stark, 1936b: 152 A : ES FE MG SV WS
dejevi Eggers, 1942: 31 [HN]
grandis Schedl, 1964a: 99 N: AG MO
granulatus Ratzeburg, 1837: 164 (Bostrichus) E: BE BU CR CT CZ DE EN FI FR GE GR HU IT LA LT NR
PL SK SP ST SV SZ UK YU N: EG
tredlii Hagedom, 1904b: 232
klimeschi Eggers, 1915c: 188 A: KI TD TM UZ
niger Stark, 1936b: 152 A: FE
palmi Hansen, 1956: 183 E: EN FI LA NT SV
rybinskii corsicus Eggers, 1912b: 113 E: FR (Corse)
rybinskii rybinskii Reitter, 1895b: 72 E: AU CZ FR GE HU IT PL SK SP ST UK
Salicis Stark, 1952: 283
striatulus Mannerheim, 1853: 235 (Cryphalus) E: HUi NAR
nitidus Swaine, 1912: 349
punctipennis Hopkins, 1915a: 37
tremulae Stark, 1952: 287 E: BU GG SK ST UK

tribe Crypturgini LeConte, 1876

genus Aphanarthrum Wollaston, 1854: 292 type species Aphanarthrum euphorbiae Wollaston, 1854
aeonii Lindberg, 1953: 18 N: CI
affine Wollaston, 1860a: 166 N: CI MO
armatum Wollaston, 1862a: 167 N: CI
monodi Paulian & Villiers, 1941: 102
bicinctum bicinctum Wollaston, 1860a: 165 N: CI MO
bicinctum obsitum Wollaston, 1865: 43 N: CI
bicinctum vestitum Wollaston, 1865: 43 N: CI
bicolor Wollaston, 1860a: 165 N: CI MR
canariense canariense Wollaston, 1860a: 164 N: CI MR
longipes Israelson, 1972: 256
canariense neglectum Schedl, 1964a: 99 N: CI
canescens canescens Wollaston, 1865: 41 N: CI
canescens polyspiniger Israelson, 1972: 256 N: CI
canescens simplex Wollaston, 1865: 41 N: CI
euphorbiae Wollaston, 1854: 293 N: MR
glabrum glabrum Wollaston, 1860a: 167 N: CI
glabrum nudum Israelson, 1972: 251 N: CI
 SCOLYTINAE 227
jubaejubae Wollaston, 1860a: 164 N: CI
jubae tuberculatum Wollaston, 1865: 40 N: CI
mairei Peyerimhoff, 1923: 53 N: CI MO AFR
duongi Villiers, 1946: 140
goniomma Enderlein, 1929: 142
saturatum Peyerimhoff, 1925b: 12 :
piscatorium Wollaston, 1860a: 166 N: CI MR
pygmaeum Wollaston, 1865: 42 N: CI
laticollis Wollaston, 1865: 42
reticulatum Wood, 1988b: 191 A: UP ORR
royaleanum Wood, 1988b: 191 A: UP ORR
subglabrum Israelson, 1972: 251 N: CI
wollastoni Israelson, 1972: 251 N: CI

genus Cisurgus Reitter, 1895b: 65 type species Crypturgus filum Reitter, 1889
ferulae Pfeffer, 1983: 294 A: UZ
filum Reitter, 1889c: 126 {Crypturgus) A: TM
mourus Eggers, 1910b: 559 E: CR N: AG TU A: IS
karamani Reitter, 1913b: 64
occidentalis Peyerimhoff, 1923: 53 N: AG MO TU
resiniferae Peyerimhoff, 1925b: 13
ragusae Reitter, 1906b: 241 E: CR IT N: TU
hystrix Abeille de Perrin, 1894: 94 {Crypturgus)
seselii Coffin & Téocchi, 1991: 157 E: FR
pusillus Wollaston, 1860a: 167 {Aphanarthrum) N: AG CI
wollastonii Eichhoff, 1878b: 77 [unnecessary RN] {Crypturgus)

genus Coleobothrus Enderlein, 1929: 144 type species Coleobothrus jandiacus Enderlein, 1929 (=
Aphanarthrum alluaudi Peyerimhoff, 1923)
alluaudi Peyerimhoff, 1923: 52 {Aphanarthrum) N: CI MO AFR
jandiacus Enderlein, 1929: 144
luridus Wollaston, 1860a: 163 {Aphanarthrum) N: CI
annulicollis Enderlein, 1929: 146
flavicollis Enderlein, 1929: 146

genus Crypturgus Erichson, 1836: 60 type species Bostrichus pusillus Gyllenhal, 1813
beesoni Eggers, 1936c: 627 A: KA UP
cedri Eichhoff, 1868d: 403 N: AG
cinereus Herbst, 1794a: 116 {Bostrichus) E: AU BE BU BY CR CT CZ DE EN FI FR GE GR HU IT LA LS
LT LU MC NR NT PL RO SK SL SP ST SV SZ UK YU A: ES FE HEI JIL MG NC TR WS
apfelbecki Eggers, 1940f: 36
atticus Eggers, 1911b: 120
corsicus Eggers, 1923b: 135
tenerrimus C. R. Sahlberg, 1836a: 140 {Hylesinus)
concolor Wollaston, 1864: 263 {Aphanarthrum) N: CI
cribrellus Reitter, 1895b: 64 E: BU CR FR IT MC PT SL SP UK YU
cylindricollis Eggers, 1940f: 37 E: BH BU CR GR MA A: TR
dubius Eichhoff, 1872b: 139 E: "Pyreneien" A:TR
hispidulus C. G. Thomson, 1870: 338 E: AU BU BY CT CZ DE EN FI GE HU IT LA LT NL NR NT PL SK
SV SZ UK A: ES FE HEI LIA NC WS
maulei Roubal, 1910:203
mediterraneus Eichhoff, 1869a: 41 E: CR FR GR IT PT SP ST N: AG MO TU A: CY IS TR
mediterranem Eichhoff, 1872b: 139 [HN]
numidicus Ferrari, 1867: 6 E: BH BU CR FR GR HU IT MA SP ST N: AG TU A: TR
abbreviatus Eggers, 1911b: 123
barbeyi Strohmeyer, 1929: 182
228 SCOLYTINAE
brevipennis Reitter, 1913b: 63
parallelocollis Eichhoff, 1878b: 73 E: GR A: TR
gaunersdoifeh Reitter, 1885b: 389
pusillus Gyllenhal, 1813:371 (Bostrichus) E: AU BE BH BY CR CT CZ DE EN FI FR GE GR HU IT LA LT
LU MC NL NR NT PL PT RO SK SL SP ST SV SZ UK YU N: AG EG LB MO TU A: ES FE HEI HP JA
JIL KA NC NP PA SC SCH TR UP WS XIZ YUN "NW Himalaya" NARi
aphodioides A. Villa & G. B. Villa, 1833: 36 (Bostrichus)
atomus LeConte, 1868: 152
danicus Eggers, 1932b: 80
minimus Stebbing, 1903: 252 (Polvgraphus)
subcribrosus Eggers, 1933a: 5 E: AU CT CZ DE EN FI GE GB BH BY FI LA LT NR NT PL SK SL SV SZ A:
ES FE WS
tuberosus Niisima, 1909: 139 A: FE JA TAI

genus Deropria Enderlein, 1929: 143 type species Aphanarthrum elongatum Eggers, 1927
elongata Eggers, 1927a: 39 (Aphanarthrum) N: CI

tribe Dryocoetini Lindemann, 1876

genus Coccotrypes Eichhoff, 1878a: 391 type species Bostrichus dactyliperda Fabricius, 1801
Cryphaloides Formânek, 1908: 91 type species Cryphaloides donisthorpei Formânek, 1908 (= Bostrichus carpophagus
Hornung, 1842)
Dendrurgus Eggers, 1923a: 144 type species Dendrurgus sundaensis Eggers, 1923 (= Thamnurgides myristicae Roepke,
1919)
Poecilips C. F. C. Schaufuss, 1898a: 110 type species Poecilips sannio C. F. C. Schaufuss, 1898
Spermatoplex Hopkins, 1915a: 48 type species Spermatoplex rhizophorae Hopkins, 1915
Thamnurgides Hopkins, 1915a: 45 type species Thamnurgidespersicae Hopkins, 1915 (= Coccotrypes advena Blandford,
1894)
advena Blandford, 1894a: 100 A: JA SD UP AUR NARi NTR ORR
cubanus Eggers, 1934b: 79 (Thamnurgides)
minor Eggers, 1923a: 150 (Dendrurgus)
niger Schedl, 1939c: 345 (Poecilips)
miciferus Schedl, 1938a: 10 (Poecilips)
persicae Hopkins, 1915a: 45 (Thamnurgides)
philippinensis Eggers, 1923a: 145 (Dendrurgus)
Philippinern is Schedl, 1933a: 104
setosus Beeson, 1929: 228 (Thamnurgides)
subnitidus Schedl, 1954c: 147 (Poecilips)
ternatensis Eggers, 1923a: 146 (Dendrurgus)
vicarius Beeson, 1939: 285 (Thamnurgides)
babai Murayama, 1961: 29 (Thamnurgus) A: JA
barbatus Schedl, 1934: 90 (Thamnurgides) A: AP ORR
ater Eggers, 1936a: 84 (Thamnurgides)
bambusae Beeson, 1939: 289 (Thamnurgides)
dipterocarpi Beeson, 1939: 288 (Thamnurgides)
cardamomi C. F. C. Schaufuss, 1905a: 8 A: JA N P SE TAI
carpophagus Hornung, 1842: 116 (Bostrichus) E: A Z FR N : CI M R A: JA SC SD TAI U P A F R A U R NAR
NTR ORR
anonae Hopkins, 1915a: 46
bakeri Hopkins, 1915a: 46
canariensis Eggers, 1928b: 117
ceylonicus Schedl, 1949a: 119
donisthorpei Formânek, 1908: 91 (Cryphaloides)
exasperatus Schedl, 1975a: 455
grisseopuberulus Schedl, 1972b: 59
hubbardi Hopkins, 1915a: 46
integer Eichhoff, 1878a: 391 [=1878b: 311]
libériens is Hopkins, 1915a: 47
nanus Eggers, 1920a: 33
 SCOLYTINAE 229
phoenicola Beeson, 1939: 281
pi los u I its Sc lied 1, 1949a: 118
punclulatus Eggers, 1951: 151
pygmaeus Eichhoff, 1878a: 391 [= 1878b: 310]
rolliniae Hopkins, 1915a: 47
thrinacis Hopkins, 1915a: 46
trevori Beeson, 1939: 282
cyperi Beeson, 1929: 230 (Thamnurgides) A: FUJ SD TAI XIZ YUN AUR NAR NTR ORR
caraibicus Schedl, 1952d: 345 (Poecilips)
conspiciens Schedl, 1936c: 110 (Xyleborus)
eggersi Schedl, 1952d: 347 (Poecilips)
indiens Eggers, 1936c: 631 (Thamnurgides)
insularis Eggers, 1940a: 127 (Dryocoetes)
insularis Eggers, 1940a: 129
pilifrons Browne, 1970: 568 (Poecilips)
subaplanatus Schedl, 1942a: 23 (Poecilips)
subimpressus Eggers, 1940a: 127 (Dryocoetes)
dactyliperda Fabricius, 1801: 387 (Bostrichus) E: FR GR HU ICi IT MA PT SP SZ N: CI EG LB MO MR A:
HEBi IS JA JO UP AFR AUR NAR NTR ORR
bassiaevorus Hopkins, 1915a: 47
borassi Beeson, 1939: 283
eggersii Hagedorn, 1904d: 449
elaeocarpi Beeson, 1939: 284
laboulbenei Decaux, 1890: 953
moreirai Eggers, 1928a: 86
obscurus Rey, 1892: 30
palmicola Hornung, 1842: 116 (Bostrichus)
tanganus Eggers, 1935b: 307
tropicus Eichhoff, 1878a: 391 [= 1878b: 312]
fagi'HobM.chi, 1959a: 12{Poecilips) A: JA
graniceps Eichhoff, 1877b: 120 (Dryocoetes) A: JA SW TAI
graniceps Eichhoff, 1878b: 314 [HN]
japonicus Eggers, 1926b: 145 (Poecilips) A: JA
klapperichi Schedl, 1953a: 26 {Poecilips) A: JA FUJ
longior Eggers, 1927e: 83 {Poecilips) A: FUJ JA SD TAI UP XIZ YUN ORR
apicatus Schedl, 1971b: 372 (Poecilips)
linearis Eggers, 1941c: 222 (Poecilips)
naidaijinensis Murayama, 1957b: 605 (Dryocoetes)
nitidipennis Schedl, 1950d: 896 (Poecilips)
oblongus Eggers, 1927e: 83 (Poecilips)
monoceros Beeson, 1939: 291 {Thamnurgides) A: AP ORR
norimasanus Murayama, 1954: 192 {Dryocoetes) A: JA
mbilus Blandford, 1894a: 95 (Dryocoetes) A: JA SC SCH SD UP XIZ ORR
brevipilosus Beeson, 1939: 298 (Thamnurgides)
corticis Beeson, 1939: 298 (Thamnurgides)
himalayensis Beeson, 1939: 299 (Thamnurgides)
mauritianus Browne, 1970: 569 (Poecilips)
parvus Beeson, 1939:297 (Thamnurgides) [HN]
papuanus Eggers, 1923a: 148 {Dendrurgus) A: SD TAI UP ORR
decipiens Browne, 1972: 21 (Poecilips)
glandis Beeson, 1939: 287 (Thamnurgides)
rubidus Beeson, 1939: 290 (Thamnurgides)
perditor Blandford, 1894a: 99 A: JA "China"
rugicollis Eggers, 1925: 157 (Thamnurgides) A: FUJ ORR
salakensis Schedl, 1939a: 38 (Poecilips) A: SD "China" ORR
acuminatus Schedl, 1966a: 34 (Poecilips)
opacifions Beeson, 1939: 294 (Thamnurgides)
punctalus Eggers, 1927e: 85 (Poecilips)
vulgaris Eggers, 1923a: 151 (Dendrurgus) A: SD TAI UP NARi ORR
230 SCOLYTINAE
brevior Eggers, 1927e: 84 (Poecilips)

genus Cynanchophagus Aksentev, 1987:150 type species Cynanchophagus cornutus Aksentev, 1987
cornutus Aksentev, 1987: 151 A: TD

genus Cyrtogenius Strohmeyer, 1910: 127 type species Cyrtogenius bicolor Strohmeyer, 1910
Artepityophthorus Schedl, 1969c: 157 type species Artepityophthorus aries Schedl, 1969
Carpophloeus Schedl, 1958d: 143 type species Carpophloeus rugipennis Schedl, 1958
Carposinus Hopkins, 1915a: 9,47 type species Carposinus pini Hopkins, 1815 (= Dryocoetes luteus Blandford, 1894)
Eulepiops Schedl, 1939c: 344 type species Eulepiops glaber Schedl, 1939
Metahylastes Eggers, 1922c: 165 type species Metahylastes africanus Eggers, 1922 (= Cyrtogenius africus Wood, 1988)
Mimidendrulus Schedl, 1957b: 68 type species Mimidendrulus movoliae Schedl, 1957
Orosiotes Niisima, 1917: 1 type species Orosiotes kumamotoensis Niisima, 1917
Ozodendron Schedl, 1957b: 13 type species Pelicerus grandis Beeson, 1929
Ozodendron Schedl, 1964c: 243 [HN] type species Pelicerus grandis Beeson, 1929
Pelicerus Eggers, 1923a: 216 type species Lepicerus nitidus Hagedorn, 1910
Taphroborus Nunberg, 1961: 617 type species Taphroborus vaticae Nunberg, 1961
brevior Eggers, 1927e: 86 (Pelicerus) A: JA SC ORR
kumamotoensis Niisima, 1917: 2 (Orosiotes) A: JA
kabei Murayama, 1955: 87
luteus Blandford, 1894a: 94 (Dryocoetes) A: ANH FUJ GUA GUI GUX HAI HEN HUB HUN JA JIA JDC SC
SCH SHA SHX TAI YUN ZHE ORR
formosanus Schedl, 1952b: 63 (Oriosiotes)
nitidus Hagedorn, 1910a: 1 (Lepicerus) A: FUJ AUR ORR
orientalis Eggers, 1923a: 217 (Pelicerus)
perakensis Schedl, 1936a: 9 (Dryocoetes) A: HKG ORR
philippinensis Eggers, 1927e: 87 (Pelicerus) A: JA (Ryukyus) ORR

genus Dactylotrypes Eggers, 1927a: 37 type species Dactylotrypes uyttenboogaarti Eggers, 1927 (=Xyloterus
longicollis Wollaston, 1864)
longicollis Wollaston, 1864: 256 (Xyloterus) Ei: CR FR IT SP SZ N: CI MRi
draconis Enderlein, 1929: 148
uyttenboogaarti Eggers, 1927a: 38

genus Dryocoetes Eichhoff, 1864: 38 type species Bostrichus autographus Ratzeburg, 1837
Anodius Motschulsky, 1860: 155 type species Bostrichus autographus Ratzeburg, 1837
Dryocoetinus Balachowsky, 1949a: 180 type species Bostrichus villosus Fabricius, 1792
aceris Krivolutskaya, 1968: 58 A: FE "Kuriles"
affinis Blandford, 1894a: 93 A: JA
alni Georg, 1856: 59 (Bostrichus) E: AU BE BU BY CT CZ DE EN FI FR GB GE GR HU IT LA LS MD NL
NR NT PL RO SK SL ST SV SZ UK "Caucasus" A: ES TR WS
leonhardi Eggers, 1912a: 49
marshami Rye, 1868: 188 (Tomicus)
similis Eggers, 1911b: 121
autographus Ratzeburg, 1837: 160 (Bostrichus) E: AU BE BH BU BY CR CT CZ DE EN FI FR GB GE GR
HU IR IT LA LS LT MD NL NR NT PL RO SK SL SP ST SV SZ UK YU "Caucasus" N- AG MR A: ES
FE GAN HAI HEI JA LIA MG NC SC SHA SHX TAI WS "Manchuria" NAR NTRi
alternons Eggers, 1931: 30
americanus Hopkins, 1915a: 51
artepunctatus Eggers, 1941b: 122
brasiliensis Schedl, 1940b: 207
longicollis Eggers, 1941b: 121
pseudotsugae Swaine, 1915: 360
sachalinensis Sokanovskiy, 1960: 675
semicastaneus Mannerheim, 1852: 358 (Bostrichus)
septentrionis Mannerheim, 1843: 298 (Bostrichus)
suecicus Eggers, 1923b: 136
victoris Mulsant & Rey, 1853: 91 (Bostrichus)
 SCOLYTINAE 231
villosus Herbst, 1794a: 121 (Bostrichus) [HN]
baikalicus Reitter, 1899c: 286 E: CT NT A: ES FE HEI HUB JA KZ MG NC NMO WS
ursus Sokanovskiy, 1960: 675
brevipilosus Murayama, 1957b: 604 A: JA
carpinivorus Choo & Woo, 1989: 58 A: SC
cerasi Eggers, 1942: 32 A: FE
cerasi Stark, 1950:229
cornivorus Murayama, 1950b: 63 (Xyleborus) A: JA
cristatus Inouye & Nobuchi, 1959: 122 A: JA
hectographus Reitter, 1913b: 76 E: BE BU BY CT CZ EN FI FR GE HU IT LA LT NR NT PL SK SL SV SZ
A: ES FE GAN HEI JA JIL KZ LIA MG NC QIN SCH SHA SHX TAI WS XIZ YUN
ater Eggers, 1933e: 101 (Ozopemon)
formosanus Nobuchi, 1967: 18
himalayensis Strohmeyer, 1908c: 161 Ei: FR SZ A: KA UP
indicia Stebbing, 1914: 549 A: HP KA NP UP
infiiscatus Murayama, 1937: 370 A: ES FE JA MG NC NE SC
orientalis Kurentsov, 1941: 169,233
pilosiusculus Kurentsov, 1948: 51
italus Eggers, 1940e: 44 E: IT
karamatsu Sawamoto, 1940: 97, 102 A: JA
kepongi Schedl, 1953d: 296 (Dryocoetes) A: TAI ORR
krivolutzkajae Mandelshtam, 2001b: 11 A: FE
niijimai Nobuchi, 1959a: 11 A.JA
padi Kurentsov, 1941: 170 A: FE
padi Stark, 1952:335
pilosus Blandford, 1894a: 92 A: JA
pini Niisima, 1909: 152 A: FE (Sakhalin) JA LIA SC
quadrisulcatus Strohmeyer, 1908b: 72 A: KA UP
rugicollis Eggers, 1926a: 137 A: FE JA JIL NC NE SHX
striates Eggers, 1933a: 8 A: FE HEI HUB JA
abietinus Kôno & Tamanuki, 1939: 90
uniseriatus Eggers, 1926b: 145 A: FE (Sakhalin) JA SHA SHX
ussuriensis Eggers, 1933a: 8 A: FE WS
rugulosus Eggers, 1933a: 9
villosus minor Eggers, 1908a: 122 E: CR FR (Corse) GR IT SL N: CI A: TR
sardus Strohmeyer, 1912b: 57
villosus villosus Fabricius, 1792: 367 (Bostrichus) E: AU BE BU BY CR CZ DE EN FR GB GE GR IR IT
(Sardegne, Sicilia) LA LT MC NR PL PT SK SL SP ST SV SZ UK YU N: AG MR TU A: TR
eichhoffi Ferrari, 1867: 29
histerinus Dufour, 1843: 91 (Bostrichus)
starhoni Reitter, 1913b: 77
genus Dryocoetiops Schedl, 1957b: 13 type species Ozopemon laevis Strohmeyer, 1911
coffeae Eggers, 1923a: 161 (Dryocoetes) A: JA SD TAI "China" ORR
javanus Eggers, 1936a: 87 (Dryocoetes)
dinoderoides Blandford, 1894a: 97 (Dryocoetes) A: JA
kepongi Schedl, 1953d: 296 (Dryocoetes) A: TAI ORR
moestus Blandford, 1894a: 96 (Dryocoetes) A: JA

genus Lymantor Levendal, 1889: 25,68 type species Lymantor sepicola Levendal, 1889 (= Tomicus coryli
Penis, 1855)
aceris aceris Lindemann, 1875a: 140 (Dryocoetes) E: AB AU CZ EN FR GE HU LA LT PL SK ST SZ UK YU
A: ES MG
aceris schablioviskii Stark, 1936b: 153 A: ES FE MG
coryli Penis, 1855: lxviii (Tomicus) E: AU BE BU CR CT CZ DE EN FI FR GB GE GR HU IT LA LT NR NT
PL RO SK SL SP ST SV SZ UK YU "Caucasus" A: ES FE TR
232 SCOLYTINAE
sepicola Lovendal, 1889b: 25, 69

genus Pseudothamnurgus Eggers, 1912b: 115 type species Dryocoetes mediterraneus Eggers, 1910
mediterraneus Eggers, 1910b: 560 {Dryocoetes) E: FR IT N: AG
nitidicollis Reitter, 1887: 197 {Dryocoetes) N: MO
normandi Eggers, 1910a: 37 {Thamnurgus) E: FR N: AG TU AFR
elegans Wichmann, 1913a: 118
scrutator Pandellé, 1883: 136 (Thammirgus) E : B E F R S P A: "Palestina"

genus Taphronurgus Reitter, 1913b: 90 type species Thamnurgus exul Reitter, 1891
exul Reitter, 1891b: 199 {Thamnurgus) A: Kl KZ

genus Taphrorychus Eichhoff, 1878b: 49,204 type species Bostrichus bicolor Herbst, 1794
Pseudopoecilips Murayama, 1957b: 614 type species Pseudopoecilips mikuniyamensis Murayama, 1957
Saliciphilus Sokanovskiy, 1954: 20 type species Hypothenemus machnovskii Sokanovskiy, 1952
Taphrocoetes Pfeffer, 1987: 22 type species Taphrorychus hirtellus Eichhoff, 1878
Taphroterus Schedl, 1965b: 341 type species Taphroterus primitus Schedl, 1965
alni Pfeffer, 1940a: 53 E: FR
betulae Schedl, 1974: 87 A: PA
bicolor Herbst, 1794a: 116 {Bostrichus) E: AU BE BH BU CR CZ DE FR GB GE HU IT LS LT MC NL NR
PL PT RO SK SP SV SZ UK YU "Caucasus" A: NC SC TR
fiiscus Marsham, 1802: 53 {Ips)
carpini Kurentsov, 1941: 167 {Dryocoetes) A: FE
carpini Eggers, 1942: 32 {Dryocoetes) [HN]
carpini Stark, 1952: 329 {Dryocoetes) [HN]
ceratoniae Peyerimhoff, 1926c: 388 N: AG MO
cribripennis Eggers, 1944c: 142
hewetti Stebbing, 1908: 11 {Dryocoetes) A: HP UP
hirtellus Eichhoff, 1878b: 208 E: BH BU CR CZ HU MC RO SK YU N: AG A: TR
mecedamts Reitter, 1913b: 95
lenkoranus Reitter, 1913b: 96 E: AB AR GG ST
machnovskii Sokanovskiy, 1952: 113 {Hypothenemus) A: KI
mikuniyamensis Murayama, 1957b: 616 {Pseudopoecilips) A: JA
minor Eggers, 1923b: 137 E: IT N: AG
picipennis Eggers, 1926a: 138 {Dryocoetes) A: HUB JA SCH
pilosus Murayama, 1957b: 619 {Pseudopoecilips) A: JA
brevipilosus Murayama, 1957b: 621 {Pseudopoecilips)
ramicola Reitter, 1895b: 94 E: BU ST A: S Y TR
pusillus Eggers, 1933a: 7
siculus Eggers, 1908a: 121 E: BH IT (Sicilia) SZ
striatus Nobuchi, 1966: 52 A: JA
taradakensis Murayama, 1957b: 618 {Pseudopoecilips) A: JA
villifrons Dufour, 1843: 91 {Bostrichus) E: AR AU BE BU CR CT CZ FR GB GE HU IT MC NT PT SK SP ST
SZ UK YU N: AG EG LB MO TU A: IN TR
bulmerincqii Kolenati, 1846: 39 {Bostrichus)
capronatus Perris, 1866: 193 {Dryocoetes)
coronatus Eggers, 1944c: 141
roubali Pfeffer, 1940a: 53
schimitscheki Eggers, 1940f: 38
splendens Sokanovskiy, 1958: 39

genus Thamnurgus Eichhoff, 1864: 40 type species Bostrichus euphorbiae Küster, 1845
armeniacus Reitter, 1897c: 244 E: AB AR
posticepunctatus Eggers, 1937: 334
brylinskyi Reitter, 1889a: 40 E: AR FR A: TM TR
caucasicus Reitter, 1887: 195 E: AR BU CT ST UK A: TM
 SCOLYTINAE 233
pegani Eggers, 1933a: 6
characiae Rosenhauer, 1878: 162 E: FR GR IT MA SP N: AG TU
sardus Eggers, 1912b: 114
delphinii Rosenhauer, 1856: 302 (Bostrychus) E: FR GR IT MC SP ST UK N: AG MO TU A: TM TR
holtzi Strohmeyer, 1907:6
robustus Eggers, 1908a: 122
semirufiis Reitter, 1906a: 36
euphorbiae Küster, 1845a: 39 (Bostrichus) E: BH CR FR GE GR IT SP UK YU N: AG A: SY
siculus Eggers, 1912b: 115
jemeniae Schedl, 1975c: 353 A: YE
kaltenbachii Bach, 1849: 199 E: AU BE BH CR FR GE HU IT PL SP SZ N: AG
csikii Endrödi, 1957: 307
declivis Reitter, 1897c: 244
mairei Peyerimhoff, 1949: 302 N: AG MO
orientalis Schedl, 1978a: 38 A: IS
petzi Reitter, 1901c: 182 E: AU HU MD
rossicus Alekseev, 1957: 161 (Thamnurgus) A: ST
varipes Eichhoff, 1878a: 390 [=1878b: 212] E: AU BH BU CZ FR GE GR HU PL RO SK ST YU A: TR
genus Triotemnus Wollaston, 1864: 264 type species Triotemnus subretusus Wollaston, 1864
Cladoctoporcus Schedl, 1975a: 454 type species Cladoctoporcus scrofa Schedl, 1975
antoinei Peyerimhoff, 1949: 299 N: MO
grangeri Peyerimhoff, 1919b: 253 (Lymantor) N: AG
lepineyi Balachowsky, 1949b: 98 N: MO
longicollis Peyerimhoff, 1925b: U N : MO
scrofa Schedl, 1975a: 454 {Cladoctoporcus) A: UP
subretusus Wollaston, 1864: 265 E: SP N: CI
ulianai Gatti & Pennacchio, 2004: 104 E: IT

genus Xylocleptes Ferrari, 1867: 37 type species Bostrichus bispinus Duftschmid, 1825
Hylonius Nunberg, 1973: 16 type species Hylonius brunneus Nunberg, 1973
Xestips Hagedorn, 1912c: 353 type species Xestips marginatus Hagedorn, 1912
bicuspis Reitter, 1887: 196 A: SY
bispinus Duftschmid, 1825: 92 (Bostrichus) E: AU BE BU CR CZ DE FR GB GE GR HU IT MC NL PL RO
SK SL SP ST SZ UK YU N: AG EG LB MO TU A: TR
relus us Olivier, 1795: 10 (Scolytus)
biuncus Reitter, 1894a: 45 E: CR IT FR SL SP N: AG MO A: TR

tribe Ipini Bedel, 1888

genus Acanthotomicus Blandford, 1894a: 89 type species Acanthotomicus spinosus Blandford, 1894
Mimips Eggers, 1932a: 33 type species Ips pilosus Eggers, 1924
perexiguus Blandford, 1896b: 201 (Tomicus) A: SD YUN ORR
denticulus Eggers, 1923a: 163 (Ips)
philippinensis Eggers, 1927e: 78 (Ips)
spinosus Blandford, 1894a: 90 A: JA TAI YUN

genus Ips DeGeer, 1775: 190 type species Dermestes typographus Linnaeus, 1758
Bonips Cognato, 2001: 779 type species Tomicus bonanseai Hopkins, 1906
Cumatotomicus Ferrari, 1867: 44 type species Bostrichus stenographus Duftschmid, 1825 (= Dermestes sexdentatus
Boerner, 1766)
Cyrtotomicus Ferrari, 1867:44 type species Bostrichus acuminatus Gyllenhal, 1827
Emarips Cognato, 2001: 779 type species Tomicus emarginatus LeConte, 1876
Granips Cognato, 2001: 780 type species Tomicus grandicollis Eichhoff, 1868
acuminatus Gyllenhal, 1827: 620 (Bostrichus) E: AU BE BH BU BY CR CT CZ DE EN FI FR GB GE GR HU
IR IT LA LT LU MC MD NL NR NT PL RO SK SL SP ST SV SZ UK YU "Caucasus" A: ES FE FUJ
234 SCOLYTINAE
GAN HEB HEI HEN HEI HUN JA JIL KZ LIA MG NC NMO QIN SC SCH SHA SHN SHX SY TAI TR
WSXINYUN ORR
geminatus Zetterstedt, 1828: 345 (Boslrichus)
heydeni Eichhoff, 1884: 298 (Tomicus) ""'5
amitinus Eichhoff, 1872b: 138 (Tomicus) E: AU BE BH BU CR CZ DE EN FI FR GE GR HU IT LA LT MC
NL NT PL RO SK SZ YU A: TR
montana Fuchs, 1913a: 80
cembrae Heer, 1836: 28 (Bostrichus) E: AU CT CZ DE FR GB GE GR HU IT LS NL NT PL SK SL SZ A: FE
HEI JA JIL KZ MG SC WS
engadinensis Fuchs, 1913a: 82
fallax Eggers, 1915b: 96
shinanoensis Yano, 1924: 2
chinensis Kurentsov & Kononov, 1966: 31 A: YUN
duplicates C. R. Sahlberg, 1836a: 144 (Bostrichus) E: AU BY CT CZ EN FI FR GE HU LA LT NR NT PL SK
SV A: ES FE HEI JA JIL KZ LIA MG NC NMO SCH WS XIN
infucatus Eichhoff, 1877a: 392 (Tomicus)
infitcatus Eichhoff, 1878b: 247 [HN] (Tomicus)
judeichii Kirsch, 1871a: 388 (Bostrichus)
rectangulus Eichhoff, 1867: 83 (Tomicus)
hauseri Reitter, 1895b: 81 A: ES FE JIL KI KZ TD TR WS XIN
ussuriensis Reitter, 1913b: 107
longifolia Stebbing, 1909: 26 (Tomicus) A: BT HP NP PA UP XIZ "W Himalaya"
nitidus Eggers, 1933f: 101 A: GAN GUI HEI NMO QIN SCH XIN XIZ YUN
schmutzenhoferi Holzschuh, 1988: 482 A: BT
sexdentatus Boerner, 1766: 78 (Dermestes) E: AU BE BH BU BY CR CT CZ DE EN FI FR GB GE GR HU IT
LA LT LU MC MD NL NR NT PL PT RO SK SL SP ST SV SZ UK YU "Caucasus" A: ES FE GAN HEB
HEI HEN HUB JA JIL KZ LIA MG NC NMO SC SCH SHA SHX TR WS YUN ORR
junnanicus Sokanovskiy, 1959a: 94, 95
pinastri Bechstein, 1818: 74 (Bostrichus)
stenographus Duftschmid, 1825: 88 (Bostrichus)
shangrila Cognato & Sun, 2007: 8 A: GAN QIN SCH SHA XIZ YUN
stebbingi Strohmeyer, 1908a: 69 A: AF BT HP KA NP PA UP XIZ
blandfordi Stebbing, 1909: 27 (Tomicus)
ribbentropi Stebbing, 1909: 25 (Tomicus)
subelongatus Motschulsky, 1860: 155 (Tomicus) Ei: NT A: ES FE HEI HEN JA JIL LIA MG NC NMO SHA
SHN SHX TAI WS XIN
typographus Linnaeus, 1758: 355 (Dermestes) E: AU BE BH BU BY CR CT CZ DE EN FI FR GB GE GR HU
ICi IR IT LA LS LT LU MD NR NT PL PT RO SK SL SP ST SV SZ UK YU N: AG A: ES FE GAN HEI
HEN JA JIL KZ MG NC NMO QIN SC SCH SHA TR WS XIN
japonicus Niisima, 1909: 147
octodentatus Paykull, 1800: 146 (Bostrichus)
genus Orthotomicus Ferrari, 1867: 44 type species Bostrichus laricis Fabricius, 1792
Neotomicus Fuchs, 1911: 33 type species Bostrichus laricis Fabricius, 1792
angulatus Eichhoff, 1876: 200 (Tomicus) A: CE FE FUJ JA JIA NC NO SC TAI YUN AURi ORR
erosus Wollaston, 1857: 95 (Tomicus) E: AZ AU BU CR CT FR GB GR IT MA MC NL PL PT SL SP ST SZ
UK YU N: AG EG LB MO MR TU A: AF ANH FUJ GUA GUI GUX HEN HUB HUN IN IS JIA JIX JO
LIA SCH SHA SHN SHX SY TR UZ YUN ZHE "Korea" AFRi NARi
rectangulus Ferrari, 1867: 83 (Tomicus)
golovjankoi Pyatnitskiy, 1930: 179 A: ES FE HEI JA JIL NC NE SC
kuniyoshü Nobuchi, 1959b: 23 (Ips) A: JA (Ryukyus) TAI
laricis Fabricius, 1792: 365 (Bostrichus) E: AU BE BH BU BY CR CT CZ DE EN FI FR GB GE GR HU IR IT
LA LS LT LU MC MD NL NR NT PL PT RO SK SL SP ST SV SZ UK YU "Caucasus" N AG MO A: ES
FE HEB HEI JA KZ MG NC SC SHA SHX TR WS
chalcographus Olivier, 1795: 7 (Scolytus)
 SCOLYTINAE 235
longicollis Gyllenhal, 1827: 621 (Bostrichus) E: BH BU BY<3T CZ DE EN FI FR GE GR HU IT LA MC NR
NT PL SK SL SP ST SV SZ UK YU A: SY TR
oblitus Perns, 1862: 218 (Tomicus)
mannsfeldi Wachtl, 1880: 51 (Tomicus) I:.AU BH BU CR FR GR HU IT MC PL RO SP YU A: TR
multidentatus Murayama, 1953a: 34 (Ips) A: JA
nankinensis Kurentsov & Kononov, 1966: 32 A: JIA
nobilis Wollaston. 1862b: 441 (Tomicus) N: CI
pinivorus Schedl, 1961a: 187 A: TR
proximus Eichhoff, 1868d: 403 (Tomicus) E: AU BE BH BU BY CR CT CZ DE EN FI FR GE GR HU IT LA
LT MD NL NR NT PL PT SK SL SP ST SV SZ UK YU "Caucasus" N: AG A: ES FE FUJ IQ JA KZ MG
NC NO SC TR WS "Manchuria" AFRi ORR
fejferi Keler, 1925: 191 (Ips)
omissus Eichhoff, 1872b: 138 (Tomicus)
robustus Knotek, 1899: 296 (Tomicus) E: AU BH BU CZ GR HU SK N: AG A: SY TR
melanurus Reitter, 1913b: 109
starki Spessivtsev, 1926b: 217 E: BY CT LT NT PL ST A: ES FE HEI JIL QIN SCH SHA SHX WS XIN YUN
suturalis Gyllenhal, 1827: 622 (Bostrichus) E: AB AU BE BH BU BY CT CZ DE EN FI FR GB GE GR HU ICi
IT LA LT MC MD NL NR NT PL PT RO SC SK SL SP ST SV SZ UK YU "Caucasus" A: ES FE JA JIL
KZ LIA MG NC QIN SC SCH SHA SHX YUN TR WS
nigritus Gyllenhal, 1827: 623 (Bostrichus)
tosaensis Murayama, 1950a: 52 (Ips) A: JA "Korea"
tridentatus Eggers, 1921: 41 E: AU A: TR

genus Pityogenes Bedel, 1888: 397 type species Dermestes chalcographus Linnaeus, 1760
Eggersia Lebedev, 1926: 121 type species Bostrichus bidentatus Herbst, 1784
Pityoceragenes Balachowsky, 1947:44 type species Bostrichus quadridens Hartig, 1834
bidentatus Herbst, 1784a: 24 (Bostrichus) E: AU BE BH BU BY CR CT CZ DE EN FI FR GB GE GR HU IR
IT LA LT LU MC MD NL NR NT PL PT RO SK SL SP ST SV SZ UK YU "Caucasus" A: ES FE IS JA
KZ MG TR WS AFRi NARi
ater Fabricius, 1792: 368 (Bostrichus)
bidens Fabricius, 1792: 368 (Bostrichus)
bispinus Guyon, 1855: 4815 (Bostrichus)
carniolica Fuchs, 1911: 35
obtusus Eggers, 1932b: 81
bistridentatus Eichhoff, 1878b: 282 (Tomicus) E: AL AU BH BU CR CZ FR GE GR HU IT MC PL RO SK SL
SP ST SZ UK YU A: CY JA NE TR
albanicus Eggers, 1911b: 122
pilidens Reitter, 1895b: 79
calcaratus Eichhoff, 1878b: 281 (Tomicus) E: CR FR GR HU IT MA PT SP ST UK N: AG MO TU A: IS SY
TR
herbellae Strohmeyer, 1929: 181
lipperti Hen sehe I, 1885: 242 (Tomicus)
opaeifrons Reitter, 1913b: 99
chalcographus Linnaeus, 1760: 143 (Dermestes) E: AU BE BH BU BY CR CT CZ DE EN FI FR GB GE GR
HU ICi IRi IT LA LS LT MC MD NR NT PL SK SL SP ST SV SZ YU "Caucasus" A: ES FE HEI JA JIL
LIA MG NC NMO SC SCH SHA TR WS XIN NTRi
sexdentatus Olivier, 1795: 11 (Scolytus)
spinosus DeGeer, 1775: 197 (Ips)
xylographus C. R. Sahlberg, 1836b: 148 (Bostrichus)
conjunetus Reitter, 1887: 196 E: AU BU CZ FR GE HU IT RO SK SL SZ UK A: ES FE HEB JA LIA MG TR
WS "Mandschurei"
baicalicus Eggers, 1933c: 49
foveolatus Eggers, 1926a: 137 A: ES FE JA NC
irkutensis irkutensis Eggers, 1910a: 38 E: CT FI NT SV A: ES FE MG TR WS
irkutensis monacensis Fuchs, 1911:3 E: AU BU CZ GE HU LT PL RO SZ UK YU "Caucasus"
bialowiezensis Karpinski, 1931: 34
236 SCOLYTINAE
japonicus Nobuchi, 1974: 39 A: GUI JA SCH SHA SHX XIZ YUN
pennidens Reitter, 1889e: 374 (Tomicus) E: GR ST A: CY IS SY TR
porifrons Eggers, 1933c: 50 E: GR A: CY
quadridens Hartig, 1834: 109 (Bostrichus) E: AU BE BH BU BY CR CT CZ DE EN FI FR GB GE GR HU IR
IT LA LS LT MC MD NR NT PL SK SL SP ST SV SZ UK YU "Caucasus" A: ES FE KZ TR WS
rudnevi Sokanovskiy, 1959b: 278 A: FE
saalasi Eggers, 1914b: 188 E: FI NR NT PL SV A: ES GAN LIA MG QIN WS XIN
niger Sokanovskiy, 1960: 676
scitus Blandford, 1893a: 63 A: HP KA NP PA UP XIZ YUN ORR
coniferae Stebbing, 1909:30
seirindewis Murayama, 1929: 26, 30 A: FE HEI JA LIA NC SC SCH SHA
aizawai Kôno, 1938: 69
nitidus Eggers, 1941b: 121
spessivtsevi Lebedev, 1926: 120 A: HP KA KI KZ TD UP XIN
trepanatus Nördlinger, 1848: 239 (Bostrichus) E: AU BE BH BU BY CT CZ DE EN FI FR GB GE HU IT LA
NL NR NT PL PT RO SK SP ST SV SZ UK YU
austriacus Wachtl, 1887: 320 (Tomicus)
elongatus Lavendal, 1889: 61 (Tomicus)

genus Pityokteines Fuchs, 1911: 33 type species Ips curvidens Germar, 1824
Orthotomides Wood, 1951: 32 type species Orthotomicus lasiocarpi Swaine, 1916
Pityoctines C. F. C. Schaufuss, 1916: 1246 type species Ips curvidens Germar, 1824
curvidens Germar, 1824: 462 (Tomicus) E: AU BE BH BU BY CR CT CZ FR GE GR HU IT LA LT LU MC
NT PL PT RO SK SL SP ST SZ UK YU A: ES JA TR AFRi NTRi
calligraphus Duftschmid, 1825: 91 (Bostrichus)
orthographus Duftschmid, 1825: 91 (Bostrichus)
psilonotus Germar, 1824: 463 (Tomicus)
marketae Knizek, 1998: 189 A: TR
spinidens Reitter, 1895b: 85 (Ips) E: AU BH BU CR CZ FR GE GR HU IT LS NT PL RO SK SP ST SZ YU
A: SC TR
heterodon Wachtl, 1895: 15 (Tomicus)
vorontzowi Jakobson, 1896: 521 (Tomicus) E: AU BH BU CR CZ FR GE GR HU IT MC PL RO SK SL SP ST
SZ UK YU A: TR

genus Pseudips Cognato, 2000: 365 type species Bostrichus concinnus Mannerheim, 1852
orientalis Wood & Yin, 1986: 461 (Ips) A: QIN SCH XIZ

tribe Micracidini LeConte, 1876

genus Pseudothysanoes Blackman, 1920: 46 type species Pseudothysanoes drakei Blackman, 1920 (=
Cryphalus rigidus LeConte, 1876)
Aphanocleptus Wood, 1960b: 63 type species Aphanocleptus coniferae Wood, 1960
Bostrichips Schedl, 1952a: 21 type species Bostrichips spinatus Schedl, 1952
Chalcohyus Blackman, 1943b: 363 type species Chalcohyus securigerus Blackman, 1943
Cryptocleptes Blackman, 1920: 51 [HN] type species Cryptocleptes dislocatus Blackman, 1920
Cryptulocleptus Wood, 1962: 76 [RN] type species Cryptocleptes dislocatus Blackman, 1920
Gretschkinia Sokanovskiy, 1959b: 276 type species Gretschkinia mongolica Sokanovskiy, 1959 (= Cryphalus modestus
Murayama, 1940)
Neoglostatus Schedl, 1978b: 300 type species Neoglostatus squamosus Schedl, 1978 (= Cryptocleptes abbreviatus Sched
1954)
modestus Murayama, 1940: 236 (Cryphalus) A: ES GAN HEB HEI JA LIA MG NC NE SHX
mongolica Sokanovskiy, 1959b: 277 (Gretschkinia)

tribe Premnobiini Browne, 1962

genus Premnobius Eichhoff, 1878b: 65,404 type species Premnobius cavipennis Eichhoff, 1878
Premnophilus Browne, 1962a: 79 type species Xyleborus joveri Schedl, 1951 (= Premnobius quadrispinosus Schedl,
1938)
 SCOLYTINAE 237
ambitiosus C. F. C. Schaufuss, 1898a: 109 (Xylocleptes) N: CI AFR NTR
brasiliensis Nunberg, 1958: 490
spinosus Hagedorn, 1908: 376

tribe Scolytini Latreille, 1804

genus Cnemonyx Eichhoff, 1868a: 150 type species Cnemonyx galeritus Eichhoff, 1868
Ceratolepis Chapuis, 1869: 52 type species Ceratolepis jucunda Chapuis, 1869
Loganius Chapuis, 1869: 52 type species LoganiusflavicornisChapuis, 1869
Minulus Eggers, 1912c: 206 type species Minulus barbatus Eggers, 1912 (= Cnemonyx galeritus Eichhoff, 1868)
Coptodryas Schedl, 1949b: 262 [HN] type species Coptodryas hylurgoides Schedl, 1949
Coptosomus Schedl, 1952d: 363 [RN] type species Coptodryas hylurgoides Schedl, 1949
galeritus Eichhoff, 1868a: 150 E: GRi NTR
barbatus Eggers, 1912c: 207 (Minulus)

genus Scolytus Geoffroy, 1762: 309 type species Bostrichus scolytus Fabricius, 1775
Archaeoscolytus Butovitsch, 1929: 21,23 type species Scolytus claviger Blandford, 1894
Confusoscolytus Tsai & Hwang, 1962: 3, 13 type species Eccoptogaster confusus Eggers, 1922 (= Scolytus japonicus
Chapuis, 1876)
Coptogaster Illiger, 1804: 108 type species Bostrichus scolytus Fabricius, 1775
Eccoptogaster Gyllenhal, 1813: 346 type species Bostrichus scolytus Fabricius, 1775
Ekkoptogaster Herbst, 1794a: 124 type species Bostrichus scolytus Fabricius, 1775
Pinetoscolytus Butovitsch, 1929: 22,48 type species Scolytus morawitzi Semenov, 1902
Pygmaeoscolytus Butovitsch, 1929: 21, 28 type species Bostrichus pygmaeus Fabricius, 1787
Ruguloscolytus Butovitsch, 1929: 20,47 type species Bostrichus rugulosus P. W. J. Müller, 1818
Scolytochelus Reitter, 1913b: 23 type species Ips multistriatus Marsham, 1802
Spinuloscolytus Butovitsch, 1929: 21, 24 type species Ips multistriatus Marsham, 1802
Tubuloscolytus Butovitsch, 1929: 21, 33 type species Eccoptogaster intricatus Ratzeburg, 1837
abaensis Tsai & Yin, 1962: 10, 16 A: NIN SCH SHA SHX
amygdali Guérin-Méneville, 1847: xlvii E: AU BU CR FR GG GR IT MA MC SP ST UK N: CI EG MO TU
A: CY IS JO PA SY TR ORR
aegyptiacus Pic, 1920: 55
anatolicus Eggers, 1911a: 74 (Eccoptogaster)
mailleri Eggers, 1912a: 49
rufipennis Brancsik, 1874: 135
aomoriensis Nobuchi, 1973: 16 A: JA
aratus Blandford, 1894a: 79 A: FE HEI JA JIL NC SC
aequipunctatus Niisima, 1905: 71
brevipennis Kurentsov, 1941: 100,227
intermedius Kurentsov, 1941: 82
azerbaidzhanicus Michalski, 1964: 662 E: AB
betulae Niisima, 1943: 141 A: JA
butovitschi Stark, 1936b: 153 A: BEI ES FE HEB HEI MG NMO SHA MG
butovitschi Eggers, 1942: 35 [HN]
carpini Ratzeburg, 1837: 187 (Eccoptogaster) E: AU BE BH BU CR CT CZ EN FR GE GR HU IT LA LU NL
PL RO SK SL SP ST SV SZ UK YU "Kaukasus" A: TR
balcanicus Eggers, 191 la: 75 (Eccoptogaster)
peregrinus Eggers, 1908d: 215 (Eccoptogaster)
chelogaster Schedl, 1958e: 168 A: AP ORR
chikisanii Niisima, 1905: 69 A: FE HEN JA NC NE SHX
curviventralis Niisima, 1905: 70
mandschuricus Schedl, 1941a: 42
claviger Blandford, 1894a: 80 A: FE JA NC SC "China"
platystylus Wichmann, 1915b: 213 (Eccoptogaster)
dahuricus Chapuis, 1869: 60 A: FE ES HEI JA JIL NC
agnatus Blandford, 1894a: 78
possyeti'Stark, 1938: 129
ecksteini Butovitsch, 1929: 26 E: AB KZ ST "Kaukasus" A: IN TM
eichhoffi Reitter, 1895b: 40 E: AB BU A: IN
238 SCOLYTINAE
iranicus Eggers, 1941b: 123
ellipticus Murayama, 1958: 929 A: JA
ensifer Eichhoff, 1881: 41, 163 E: AU AZ BH BU CT CZ FR GE GR HU IT PL RO SK ST SZ UK YU A: IN
TR
esuriem Blandford, 1894a: 77 A: FE HEB HEI JA JIL SHA
sachalinensis Michalski, 1964: 666
frontalis Blandford, 1894a: 79 A: JA NE SC TAI
formosanus Eggers, 1939b: 115
gretschkini Sokanovskiy, 1956: 43 A: TD
intricatus Ratzeburg, 1837: 186 (Eccoptogaster) E: AB AU BE BH BU BY CT CR CZ DE EN FI FR GB GE
GR HU IT KZ LA LT LU MC NL NR NT PL RO SK SL SP ST SV SZ YU "Caucasus" N: AG MO TU A:
IN NC TR
lenkoramis Eggers, 1942: 34
penicillatus Reitter, 1913b: 21
picicolor Stephens, 1830: 362
simmeli Eggers, 1923b: 133 (Eccoptogaster)
jacobsoni Spessivtsev, 1919: 246 (Eccoptogaster) A: FE HEI JA LIA NC
montana Kurentsov, 1935: 22
rimskii Kurentsov, 1941: 103,228
japonicus Chapuis, 1876: 199 A: BEI ES FE HEB HEI JA JIA JIL LIA MG NC NMO SC SCH SHA SHG
SHXTAI
confusus Eggers, 1922a: 13 (Eccoptogaster)
mandli Eggers, 1922a: 13 (Eccoptogaster)
starki Kurentsov, 1941: 101, 225
subconfusus Eggers, 1941b: 123
ussuriensis Kurentsov, 1941: 102, 226
jaroschewskii Schevyrew, 1893: 90 [RN] E: AB AR GG ST UK A: KZ TD UZ
granulifer Reitter, 1913b: 19
kostini Sokanovskiy, 1954: 15
rufostriatus Sokanovskiy, 1956: 44
tauricus Eggers, 1914b: 185 (Eccoptogaster)
unispinosus Schevyrew, 1890c: 470 [HN]
kashmirensis Schedl, 1958e: 168 A: KA
kirschiifasciatus Reitter, 1890c: 395 E: FR IT (Sicilia) SP ST N: AG A: TM TR UZ
demaisoni Eggers, 1912a: 47 (Eccoptogaster)
kirschii kirschii Skalitzky, 1876: 110 E: AU BU CR CZ FR GE HU IT MC PL PT SK SP ST SZ UK YU
"Caucasus" N: AG A: IQ TR WS AFRi
ruguloides Sokanovskiy, 1954: 14
koenigi Schevyrew, 1890: 471 E: AB AU BH BU CR FR HU IT MD RO ST UK "Caucasus" N: AG MO A:
ES IQ TM
aceris Knotek, 1892: 235
belokanicus Stark, 1941: 302
siculus Eggers, 1908c: 193 (Eccoptogaster)
koltzei Reitter, 1894b: 128 A: FE NC "Manchuria"
vexator Reitter, 1913b: 23
kozikowskü Michalski, 1964: 663 E: UK
laevis Chapuis, 1869: 54 E: AU BU BY CR CT CZ DE EN FR GB GE GG GR HU IT LA LS NR NT PL SK
SL SP ST SV SZ UK YU "Kaukasus"
pomacearum Butovitsch, 1929: 44
loevendali Eggers, 1912c: 203 (Eccoptogaster)
major Stebbing, 1903: 203 A: AF HP KA PA UP
deodara Stebbing, 1903: 220
minor Stebbing, 1903: 207
mali Bechstein, 1805: 882 (Bostrichus) E: AU BE BH BU BY CR CT CZ DE EN FI FR GB GE GR HU IR IT
LA LT LU MA MC NL NR NT PL RO SK SL SP ST SV SZ UK YU "Caucasus" N- AG EG MO A: ES
FE IN JIL LIA TR WS NARi
bicallosus Eggers, 1933d: 75
 SCOLYTINAE 239
castaneus Ratzeburg, 1837: 187 (Eccoplogaster)
nitiduhis Chapuis, 1869: 59
pruni Ratzeburg, 1837: 186 {Eccoptogaster)
pyri Ratzeburg, 1837: 186 (Eccoptogaster)
s trig! latus Reitter, 1908b: 23
sulcatus LeConte, 1868: 167
morawitzi Semenov, 1902: 267 E: CT NT A: ES FE HEI LIA MG NC WS
pini Eggers, 1942: 33
multistriatus Marsham, 1802: 54 [lps) E: AL AU BE BU BY CR CT CZ DE EN FR GB GE GR HU IR IT LA
LT LU MC NL NR NT PL PT RO SK SL SP ST SV SZ YU N: AG EG MO TU A: IN TR UZ AURi
NARi NTRi
abhorrens Wichmann, 1913c: 210 {Eccoptogaster)
flavicornis Chevrolat, 1835: pl. 40
javanus Chapuis, 1869: 56
nodifer Reitter, 1913b: 24
papuanus Schedl, 1936a: 8
therondi A. Hoffmann, 1939: 36
triomal us Eichhoff, 1881: 41, 160
ulmi L. Redtenbacher, 1847: 361
nakanei Nobuchi, 1967: 16 A: TAI
nitidus Schedl, 1936a: 8 A: HP KA PA UP XIZ
numidicus C. N. F. Brisout de Barneville, 1883: 147 N: AG MO
nunbergi Michalski, 1964: 665 A: FE
orientalis Eggers, 1910b: 557 (Eccoptogaster) E: BU RO ST UK A: TM
affinis Eggers, 1914a: 108 (Eccoptogaster)
parviclaviger Yin & Huang, 1980: 49, 52 A: JIL SHX
pilosus Yin & Huang, 1980: 49, 52 A: SCH
pomi Yin & Huang, 1980: 50, 53 A: YUN XIZ
pubescens Stark, 1936b: 154 A: FE NC SC
pubescens Eggers, 1942: 34 [HN]
pygmaeus Fabricius, 1787: 37 (Bostrichus) E: AB AU BE BU CR CT CZ FI FR GB GE GG GR HU IT LT MC
NL PL PT RO SK SL SP ST SV SZ UK YU A: TR
armatus Comolli, 1837: 37
noxius Ratzeburg, 1837: 187 (Eccoptogaster)
querci Yin & Huang, 1980: 50, 53 A: SCH YUN
ratzeburgii E. W. Janson, 1856: 86 E: AU BE BU BY CR CT CZ DE EN FI FR GB GE GR HU IT LA LT LU
NL NR NT PL RO SK SL SP ST SV SZ YU "Caucasus" A: ES FE HEI IN JA JIL KZ MG NC SC TR WS
amurensis Eggers, 1908b: 144 (Eccoptogaster)
bituberculatus Puzyr, 1951: 46
lineatus Kurentsov, 1941: 104, 226
sahlbergi Eggers, 1912c: 204 (Eccoptogaster)
sibiricus Eggers, 1922a: 14 (Eccoptogaster)
rugulosus P. W. J. Müller, 1818: 247 (Bostrichus) E: AB AL AU AZ BE BH BU BY CR CT CZ DE EN FI FR
GB GE GR HU IR IT LA LT LU MA MC NL NR NT PL PT RO SK SL SP ST SV SZ UK YU "Caucasus"
N: AG EG MO TU A: CY ES GAN IN IQ IS KZ LE MG PA SA SY TR WS XIN NARi NTRi ORR
assimilis Boheman, 1858: 88 (Eccoptogaster)
baluchistani Schedl, 1958er 165
caucasicus Butovitsch, 1929: 54
fauveli Reitter, 1895b: 43
haemorrhous Schmidberger, 1837: 270
intermedius Sokanovskiy, 1960: 675
manglissiensis Lezhava, 1940: 71
mediterraneus Eggers, 1922b: 121 (Eccoptogaster)
punctatus Ratzeburg, 1837: 187 (Eccoptogaster)
samarkandicus Butovitsch, 1929: 56
sanctaluciae A. Hoffmann, 1935: 84, 87
similis Butovitsch, 1929: 54
taxicola Lezhava, 1943: 193
240 SCOLYTINAE
schevyrewi Semenov, 1902: 265 E: ST A: BEI ES FE GAN GUI HEB HEI HEN JIA JIL KI KZ LIA MG NC
NE NIN NMO QIN SC SCH SHA SHN SHX TD TM UZ XIN NARi
emarginatus Wichmann, 1915b: 215 {Eccoptogaster)
frankei Wichmann, 1915b: 214 {Eccoptogaster)
seulensis Murayama, 1930: 5
sinensis Eggers, 1910a: 35
transcaspicus Eggers, 1922b: 116 {Eccoptogaster)
scolytus Fabricius, 1775: 59 {Bostrichus) E: AU BE BU BY CR CT CZ DE EN FI FR GB GE GR HU IR IT LA
LT LU NL NT PL PT RO SL SK SP ST SV SZ UK YU "Caucasus" A: ES IN KZ TR UZ WS
bostrichus Tigny, 1801: 79
californicus LeConte, 1868: 166
destructor Olivier, 1795: 5
fuchsi Reitter, 1913b: 15
geoffroi Goeze, 1777: 143 {Dermestes)
niger Geoffroy, 1785: 139
punctatus O. F. Müller, 1776: 57
scolythus Sulzer, 1776a: 21 {Dermestes)
variabilis Sokanovskiy, 1958: 37
semenovi Spessivtsev, 1919: 247 {Eccoptogaster) A: BEI ES FE HEB HEI JUi MG NC SC SHA SHX
kononovi Kurentsov, 1941: 98, 228
shanhaiensis Yin & Huang, 1980: 50, 53 A: JIL LIA
sinopiceus Tsai, 1962: 9, 14 A: GAN HEB HEI NIN NMO QIN SCH XIZ YUN
squamosus Yin & Huang, 1980: 49, 52 A: YUN
sulcifrons Rey, 1892: 30 E: BU CR CT FR GR HU IT KZ MA MC PL ST UK
leonii Eggers, 1908c: 194 {Eccoptogaster)
tadzhikistanicus Stark, 1941: 302 A: TD
triarmatus Eggers, 1912c: 205 {Eccoptogaster) E: AU DE EN FR IT NR SK ST SV SZ "Yugoslavia"
trispinosus Strohmeyer, 1908a: 69 A: FE JA NC NE
grandis Kurentsov, 1941: 104,227
varshalovitchi Michalski, 1973: 55 E: AB
ventrosus Schevyrew, 1890: 470 A: FE NE
yablonianus Murayama, 1943: 96 A: "Manchukuo"
zaitzevi Butovitsch, 1929: 31 E: GG UK

tribe Scolytoplatypodini Blandford, 1893

genus Scolytoplatypus C. F. C. Schaufuss, 1891: 31 type species Scolytoplatypus permirus C. F. C. Schaufuss,
1891
Spongocerus Blandford, 1893b: 431 type species Scolytoplatypus tycon Blandford, 1893
Strophionocerus Sampson, 1921: 36 [RN] type species Scolytoplatypus mikado Blandford, 1893
Taeniocerus Blandford, 1893b: 437 [HN] type species Scolytoplatypus mikado Blandford, 1893
blandfordi Gebhardt, 2006: 162 A: NP TAI ORR
brahma Blandford, 1898: 425 A: SD ORR
h a mat its Hagedorn, 1904c: 260
hirsutus Blackman, 1943a: 224
paucegranulatus Eggers, 1935a: 242
calvus Beaver & Liu, 2007: 227 A: TAI
daimio Blandford, 1893b: 433 A: BT FE HP JA KA NC NP SC UP
kunala Strohmeyer, 1908c: 161
muticus Hagedorn, 1904a: 124
siomio Blandford, 1893b: 436
darjeelingi Stebbing, 1914: 607 A: HP HUN NP SCH SD UP XIZ YUN ORR
gardneri Maiti & Saha, 2009: 87 A: SD
lopchuensis Maiti & Saha, 2009: 90 A: SD
mikado Blandford, 1893b: 437 A: FUJ GUX HAI JA NC SC SCH SHA TAI ORR
sinensis Tsai & Hwang, 1965: 121, 123
minimus Hagedorn, 1904a: 125 A: SD UP ORR
nitidicollis Eggers, 1935a: 241 A: SD ORR
 SCOLYTINAE 241
pubescens Hagedorn, 1904a: 123 A: NP SD TAI UP ORR
kabakovi Aksentev, 1992: 192
raja Blandford, 1893b: 440 A: HP HUN KA NP PA SCH SD SHA TAI UP XIZ YUN ORR
himalayensis Stebbing, 1914: 604
ruficauda Eggers, 1939a: 9 A: NP ORR
samsinghensis Maiti & Saha, 2009: 101 A: SD
shogun Blandford, 1894a: 126 A: JA
supercilliosus Tsai & Hwang, 1965: 121, 124 A: SCH XIZ
tycon Blandford, 1893b: 432 A: FE HEI JA NC SC TAI
ussuriensis Berger & Kholodkovskiy, 1916:4
zahradniki Knizek, 2008: 120 A: BEI SCH SHA

tribe Xyleborini LeConte, 1876

genus Amasa Lea, 1894: 322 type species Amasa thoracica Lea, 1894 (= Tomicus truncatus Erichson, 1842)
Anaxyleborus Wood, 1980: 90 type species Tomicus truncatus Erichson, 1842
Pseudoxyleborus Eggers, 1930b: 206 type species Pseudoxyleborus beesoni Eggers, 1930
calamoides Murayama, 1934b: 291 (Xyleborus) A: JA
concitata Schedl, 1969a: 214 {Xyleborus) A: TAI
eugeniae Eggers, 1930b: 183 (Xyleborus) A: SD UP ORR
geminata Hagedorn, 1904a: 126 (Xyleborus) A: SD
schlichii Stebbing, 1914: 592 (Xyleborus) A: SD ORR
brevipennis Schedl, 1971b: 378 (Xyleborus)
glabra Eggers, 1930b: 185 (Xyleborus)
tnmcata Stebbing, 1907: 40 (Acanthotomicus)
uniseriata Eggers, 1936a: 89 (Xyleborus)
verax Schedl, 1939a: 43 (Xyleborus)

genus Ambrosiodmus Hopkins, 1915a: 55 type species Xyleborus tachygraphus Zimmermann, 1868
Brownia Nunberg, 1963: 37 type species Xyleborus illepidus Schedl, 1941 (= Pityophthorus obliquus LeConte, 1878)
Phloeotrogus Motschulsky, 1863: 512 type species Phloeotrogus obliquecauda Motschulsky, 1863
asperatus Blandford, 1895: 321 (Xyleborus) A: GUI HAI JA SD TAI XIZ ORR
citri Beeson, 1930: 215 (Xyleborus)
cristatuloides Schedl, 1971a: 284 (Xyleborus)
nepotulomorphus Eggers, 1936a: 88 (Xyleborus)
nepotulus Eggers, 1923a: 179 (Xyleborus)
cristatulus Schedl, 1953d: 300 (Xyleborus) A: FUJ ORR
dihingensis Eggers, 1930b: 189 (Xyleborus) A: SD ORR
lantanae Eggers, 1930b: 180 (Xyleborus) A: SD ORR
lewisi Blandford, 1894a: 104 (Xyleborus) A: GUA GUI GUX HAI JA SC SD SCH TAI XIZ YUN NARi
ORR
lewekianus Eggers, 1923a: 181 (Xyleborus)
tegalensis Eggers, 1923a: 181 (Xyleborus)
tuberculatus Strohmeyer, 1912a: 38 (Ozopemori)
minor Stebbing, 1909: 20 (Phloeosinus) A : B T N P T A I U P ORR
crassus Hagedorn, 1910a: 8 (Xyleborus)
nodulosus Eggers, 1941c: 223 (Xyleborus) A: FUJ
pernodulus Schedl, 1957b: 86
obliquecauda Motschulsky, 1863: 513 (Phloeotrogus) A: JA ORR
carinipennis Eichhoff, 1868a: 152 {Xyleborus)
semirufus Schedl, 1959d: 499 (Xyleborus)
permarginatus Schedl, 1933b: 200 (Xyleborus) A: JA (Ryukyu) ORR
rubricollis Eichhoff, 1876: 202 (Xyleborus) Ei: IT A: ANH BEI FUJ GUI HEB HEI HUN IT JA NC SC SCH
SHA SHN SHX TAI XIZ YUN ZHE AURi NARi ORR
strohmeyeri Schedl, 1975a: 457 (Xyleborus)
taboensis Schedl, 1952b: 65 (Xyleborus)
242 SCOLYTINAE
genus Ambrosiophilus Hulcr & Cognato, 2009: 21 type species Xyleborus restrictus Schedl, 1939
atratus Eichhoff, 1876: 201 (Xyleborus) Ei: IT A: FUJ IT JA NC SC SHX TAI NARi
hunariensis Browne, 1983a: 33 {Xyleborus) A: GUI HUN TAI YUN ORR
metanepotulus Eggers, 1939b: 119 {Xyleborus) A: TAI
satoi Schedl, 1966b: 39 {Xyleborus) A: BT TAI
subnepotulus Eggers, 1930b: 178 {Xyleborus) A.TAI ORR

genus Anisandrus Ferrari, 1867: 24 type species Apate dispar Fabricius, 1792
apicalis Blandford, 1894a: 105 {Xyleborus) A: ANH BT FE GUX HAI JA NC SD SC SCH SHX TAI XIZ
YUN ORR
cristatus Hagedorn, 1908: 377 [HN] {Xyleborus)
fabricii Schedl, 1964d: 217 [RN] (Xyleborus)
dispar Fabricius, 1792: 363 {Apate) E: AB AU BE BH BU BY CR CT CZ DE EN FI FR GB GE GR HU IT LA
LT MC MD NL NR NT PL SK SL SP ST SV SZ UK YU "Caucasus" A: ES FE HEI IN JA KZ MG NC SC
SHATRWS NARi ORR
aequalis Reitter, 1913b: 81
brevis Panzer, 1796: 20 (Bostrichus)
cerasi Eggers, 1937: 335 {Xyleborus)
khinganensis Murayama, 1943: 100 {Xyleborus)
pyri Peck, 1817: 207 (Scolytus)
ratzeburgii Kolenati, 1846: 39 {Bostrichus)
rugulosus Eggers, 1922a: 17 {Xyleborus)
swainei Drake, 1921: 203
tachygraphus C. R. Sahlberg, 1836b: 152 {Bostrichus)
thoracicus Panzer, 1796: 18 {Bostrichus)
hirtus Hagedorn, 1904a: 126 {Xyleborus) A: FUJ SD TAI ORR
hirtuosus Beeson, 1930: 217 {Xyleborus)
taiwanensis Browne, 1980b: 386 {Xyleborus)
maiche Eggers, 1942: 36 {Xyleborus) E: UK A: ES FE HEI NC NARi
ursinus Hagedorn, 1908: 381 {Xyleborus) A: SD ORR
ursulus Eggers, 1923a: 173 {Xyleborus) A: FUJ SD ORR

genus Arixyleborus Hopkins, 1915a: 59 type species Arixyleborus rugosipes Hopkins, 1915
Xyleboricus Eggers, 1923a: 212 type species Xyleboricus canaliculatus Eggers, 1923
malayensis Schedl, 1954c: 150 {Xyleboricus) A: FUJ JA SD SCH XIZ YUN ORR
yakushimanus Murayama, 1955: 83 {Xyleborus)
moestus Eggers, 1930b: 189 {Xyleborus) A: SD ORR
rugosipes Hopkins, 1915a: 59 A: TAI ORR
médius Eggers, 1927e: 104 {Webbia)
camphorae Eggers, 1936c: 634 {Webbia)

genus Beaverium Hulcr & Cognato, 2009: 25 type species Xyleborus insulindicus Eggers 1923
chujoi Schedl, 1951b: 73 A: TAI

genus Cnestus Sampson, 1911: 383 type species Cnestus magnus Sampson, 1911
Tosaxyleborus Murayama, 1950a: 49 type species Tosaxyleborus pallidipennis Murayama, 1950 (= Cnestus murayamai
Schedl, 1962)
ater Eggers, 1923a: 210 {Xyleborus) A: FUJ ORR
retusiformis Schedl, 1936a: 31 {Xyleborus)
bicornioides Schedl, 1952d: 368 {Xyleborus) A: SD ORR
gravidus Blandford, 1898: 427 (Xyleborus) A: HAI SD XIZ ORR
klapperichi Schedl, 1955c: 46 {Xyleborus) A: FUJ
maculatus Browne, 1983a: 33 A: HAI HUB HUN SCH XIZ YUN
murayamai Schedl, 1962c: 207 A: JA SC TAI
murayamai Browne, 1963: 54 [RN]
pallidipennis Murayama, 1950a: 49 (Tosaxyleborus) [HN]
 SCOLYTINAE 243
mutilatus Blandford, 1894a: 103 {Xyleborus) A: ANH FUJ GUI HAI JA SC SCH SHA TAI YUN ZHE AUR
NARi ORR
banjoewangi Schedl, 1939a: 41 {Xyleborus)
sampsoni Eggers, 1930b: 184 {Xyleborus)
taitonus Eggers, 1939b: 118 {Xyleborus)
nitidipennis Schedl, 1951b: 88 {Xyleborus) A: SD TAI ORR
testudo Eggers, 1939b: 116 {Xyleborus) A: TAI ORR

genus Coptodryas Hopkins, 1915a: 54 type species Coptodryas confusa Hopkins, 1915
chrysophylli Eggers, 1930b: 205 {Xyleborus) A: SD ORR
cruralis Schedl, 1975a: 456 {Xyleborus) A: SD XIZ ORR
izuensis Murayama, 1952: 16 {Xyleborus) A: JA
mus Eggers, 1930b: 203 {Xyleborus) A: GUI SD ORR
reddens Sampson, 1923: 287 {Xyleborus) A: GUI ORR
crassitarsus Schedl, 1936a: 28 {Xyleborus)
minusculus Eggers, 1923a: 212 {Xyleborus)
minutissimus Eggers, 1930b: 204 {Xyleborus)
tuberculosa Browne, 1981b: 602 {Xyleborus)

genus Cyclorhipidion Hagedorn, 1912c: 355 type species Cydorhipidion pelliculosum Hagedorn, 1912 (=
Xyleborus perlaetus Schedl, 1960)
Kelantanius Nunberg, 1961: 621 type species Xyleborus punctatopilosus Schedl, 1936
Terminalinus Hopkins, 1915a: 57 type species Terminalinus terminaliae Hopkins, 1915
bodoanum Reitter, 1913b: 82 {Xyleborus) Ei: AU BE CR FR GE IT NL SV SZ A: FE NC NE (Maoershan)
NARi
californicum Wood, 1975: 399 {Xyleborus)
punctulatum Kurentsov, 1948: 52 {Xyleborus)
distinguendum Eggers, 1930b: 205 {Xyleborus) A: UP
eggersi Beeson, 1930: 215 {Xyleborus) A: SD
formosae Wood, 1992: 80 {Xyleborus) [RN] A: TAI
formosanum Browne, 1981a: 131 {Xyleborus) [HN]
fiikiense Eggers, 1941c: 225 {Xyleborus) A: FUJ NP TAI UP
tenuigraphum Schedl, 1953a: 29 {Xyleborus)
improbum Sampson, 1913: 444 {Xyleborus) A: SD XIZ ORR
inarmatum Eggers, 1923a: 209 {Xyleborus) A: HP SD ORR
lineatum Eggers, 1930b: 177 {Xyleborus) A: SD UP ORR
melancranis Beeson, 1930: 218 {Xyleborus)
mussooriense Eggers, 1930b: 179 {Xyleborus) A: UP
ohnoi Browne, 1980a: 375 A: TAI
pelliculosum Eichhoff, 1878a: 392 [=1878b: 336] {Xyleborus) A: FE JA NC SC SCH SHX NARi
quercus Kurentsov, 1948: 51 [HN] {Xyleborus)
seiryorense Murayama, 1930: 21 {Xyleborus)
starki Nunberg, 1956: 209 [RN] {Xyleborus)
perpilosellum Schedl, 1935b: 402 {Xyleborus) A: HAI AUR ORR
punctatopilosum Schedl, 1936b: 532 {Xyleborus)
sulcatum Eggers, 1930b: 180 {Xyleborus) A: FUJ
umbratum Eggers, 1941c: 223 {Xyleborus) A: FUJ
xyloteroides Eggers, 1939b: 120 A: TAI

genus Diuncus Hulcr & Cognato, 2009: 28 type species Xyleborus papatrae Schedl, 1972
ciliatoformis Schedl, 1953f: 81 {Xyleborus) A: TAI ORR
corpulentus Eggers, 1930b: 198 {Xyleborus) A: NP SD TAI XIZ ORR
haberkorni Eggers, 1920a: 43 {Xyleborus) A: HAI JA SD TAI UP AFRi ORR
taichuensis Schedl, 1952b: 64 {Xyleborus)
mucronatulus Eggers, 1930b: 199 {Xyleborus) A: SD
mucronatus Eggers, 1923a: 191 {Xyleborus) A: JA ORR
244 SCOLYTINAE
genus Dryoxylon Bright & Rabaglia, 1999: 333 type species Xyleborus onoharaensis Murayama, 1934
onoharaense Murayama, 1934b: 293 (Xyleborus) A: JA NARi

genus Eccoptopterus Motschulsky, 1863: 515 type species Eccoptopterus sexspinosus Motschulsky, 1863 (=
Scolytus spinosus Olivier, 1795)
Eurydactylus Hagedorn, 1909: 733 [RN] type species Platydactylus gracilipes Eichhoff, 1886
Platydactylus Eichhoff, 1886: 25 [HN] type species Platydactylus gracilipes Eichhoff, 1886
spinosus Olivier, 1795: 9 (Scolytus) A: JA SD TAI AFR AUR ORR
abnormis Eichhoff, 1869c: 282 (Xyleborus)
eccoptopterus Schedl, 195Id: 154 (Xyleborus)
sexspinosus Motschulsky, 1863:515

genus Euwallacea Hopkins, 1915a: 54 type species Xyleborus wallacei Blandford, 1896
andamanensis Blandford, 1896b: 222 (Xyleborus) A: SD UP ORR
burmanicus Beeson, 1930: 210 (Xyleborus)
granulipennis Eggers, 1930b: 194 (Xyleborus)
intextus Beeson, 1930: 211 (Xyleborus)
noxius Sampson, 1913: 445 (Xyleborus)
senchalensis Beeson, 1930: 212 (Xyleborus)
siobanus Eggers, 1923a: 186 (Xyleborus)
barbatomorphus Schedl, 1951b: 72 (Xyleborus) A: TAI
bicolor Blandford, 1894a: 113 (Xyleborus) A: JA SD UP TAI YUN ORR
artelaevis Schedl, 1942b: 196 (Xyleborus)
filiformis Schedl, 1975b: 364 (Xyleborus)
glabratulus Browne, 1983b: 560 (Xyleborus)
laevis Eggers, 1923a: 201 (Xyleborus)
privatus Beeson, 1930: 213 (Xyleborus)
rameus Schedl, 1940d: 441 (Xyleborus)
rodgeri Beeson, 1930: 213 (Xyleborus)
unimodus Beeson, 1929: 238 (Xyleborus)
tumidus Schedl, 1975b: 371 (Xyleborus)
destruens Blandford, 1896b: 221 (Xyleborus) A: TAI YUN AUR ORR
barbatulus Schedl, 1934: 86 (Xyleborus)
barbatus Hagedorn, 1910a: 11 (Xyleborus)
nandarivatus Schedl, 1950b: 52 (Xyleborus)
pseudobarbatus Schedl, 1942b: 193 (Xyleborus)
fornicatus Eichhoff, 1868a: 151 (Xyleborus) A: GUA GUI GUX HAI JA (Bonin, Ryukyus) SCH SD UP TAI
YUN XIZ AFRi NARi ORR
fornicatior Eggers, 1923a: 184 (Xyleborus)
perbrevis Schedl, 1951b: 59 (Xyleborus)
schultzei Schedl, 1951b: 68 (Xyleborus)
tapatapaoensis Schedl, 195Id: 152 (Xyleborus)
whitfordiodendrus Schedl, 1942b: 189 (Xyleborus)
fiilvus Murayama, 1936: 142 (Xyleborus) A: JA
fiinereus Lea, 1910: 139 (Xyleborus) A: TAI AUR ORR
galeatus Blandford, 1894a: 123 (Xyleborus) A: JA
interjectus Blandford, 1894b: 576 (Xyleborus) A: ANH FUJ GAN GUA GUI HAI HUB HUN JA NP SCH SD
TAI UP YUN XIZ ORR
pseudovalidus Eggers, 1925: 159 (Xyleborus)
malloti Eggers, 1930b: 192 (Xyleborus) A: SDUP ORR
piceus Motschulsky, 1863: 512 (Anodius) A:SDTAI AFR ORR
imitans Eggers, 1927d: 404 (Xyleborus)
indicus Eichhoff, 1878a: 392 [= 1878b: 354] (Xyleborus)
samoensis Beeson, 1929: 237 (Xyleborus)
subcoriaceus Eggers, 1927e: 92 (Xyleborus)
sibsagaricus Eggers, 1930b: 196 (Xyleborus) A: SD ORR
dalbergiae Eggers, 1930b: \96 (Xyleborus)
tristis Eggers, 1930d: 194 (Xyleborus) A: SD ORR
 SCOLYTINAE 245
validus Eichhoff, 1876: 202 {Xyleborus) A: ANH FUJ HUN JA SC TAI YUN NARi ORR
velatus Sampson, 1913: 443 (Xyleborus) A: SD UP XIZ ORR
asperipennis Eggers, 1934a: 27 (Xyleborus)
assamensis Eggers, 1930b: 195 (Xyleborus)
wallacei Blandford, 1896b: 220 (Xyleborus) A: SD ORR
confinis Eggers, 1923a: 200 (Xyleborus)
gravelyi Wichmann, 1914b: 411 (Xyleborus)
indocorus Schedl, 1979c: 162 (Xyleborus)
ovalicollis Eggers, 1930b: 193 (Xyleborus)
perakensis Schedl, 1942b: 194 (Xyleborus)
siporanus Hagedom, 1910a: 11 (Xyleborus)

genus Hadrodemius Wood, 1980: 94 type species Xyleborus globus Blandford, 1896
comans Sampson, 1919: 109 (Xyleborus) A: FUJ GUX HAI HUN JA JLX SCH SD TAI XIZ ZHE ORR
amorphus Eggers, 1926b: 147 (Xyleborus)
metacomans Eggers, 1930b: 188 (Xyleborus)
globus Blandford, 1896b: 208 (Xyleborus) A: TAI ORR
fitscus Eggers, 1923a: 174 (Xyleborus)
tomentosus Eggers, 1939: 10 (Xyleborus)
ursus Eggers, 1923a: 173 (Xyleborus)
pseudocomans Eggers, 1930b: 187 (Xyleborus) A: FUJ GUI HAI HUN SD XIZ ORR
artecomans Schedl, 1953a: 24 (Xyleborus)
genus Leptoxyleborus Wood, 1980: 94 type species Phloeotrogus sordicauda Motschulsky, 1863
concisus Blandford, 1894a: 107 (Xyleborus) A: JA SD TAI ORR
incurvus Eggers, 1930b: 197 (Xyleborus)
marginatus Eggers, 1927e: 91 (Xyleborus)
peguensis Eggers, 1930b: 198 (Xyleborus)
sordicaudulus Eggers, 1927e: 91 (Xyleborus)

genus Microperus Wood, 1980: 94 type species Xyleborus theae Eggers, 1940 (= Xyleborus diversicolor
Eggers, 1923)
alpha Beeson, 1929: 239 (Xyleborus) A: GUI TAI ORR
corporaali Eggers, 1923a: 210 (Xyleborus) A: BT ORR
julvulus Schedl, 1942a: 35 (Xyleborus) [RN]
fulvus Schedl, 1939a: 48 (Xyleborus) [HN]
huangi Browne, 1983a: 34 (Xyleborus) A: FUJ HUN
kirishimanus Murayama, 1955: 85 (Xyleborus) A: JA TAI
kadoyamaensis Murayama, 1934b: 290 (Xyleborus) A: JA SC TAI
perparvus Sampson, 1922: 151 (Xyleborus) A: FUJ GUI HUN JA SD TAI XIZ ORR
isukubanus Murayama, 1954: 195 (Xyleborus)

genus Webbia Hopkins, 1915b: 222 type species Webbia dipterocarpi Hopkins, 1915
Prowebbia Browne, 1962b: 208 type species Prowebbia subuculae Browne, 1962
Xelyborus Schedl, 1939c: 349 type species Xelyborus bicomis Schedl, 1939
pabo Sampson, 1922: 150 A: UP XIZ YUN ORR

genus Xyleborinus Reitter, 1913b: 83 type species Bostrichus saxesenii Ratzeburg, 1837
andrewesi Blandford, 1896b: 227 (Xyleborus) A: JA NP SD TAI UP YUN AFR AUR NARi ORR
gracilior Browne, 1984: 101 (Cryptoxyleborus)
insolitus Bright, 1972a: 77 (Xyleborus)
persphenos Schedl, 1970b: 219 (Xyleborus)
angustatus Eichhoff, 1866: 278 (Xyleborus) E: UK
artestriatus Eichhoff, 1878b: 507 (Xyleborus) A: SD TAI UP "China" AUR ORR
beaveri Browne, 1978: 603 (Xyleborus)
laticollls Blandford, 1896b: 226 (Xyleborus)
rugipennis Schedl, 1953d: 303 (Xyleborus)
246 SCOLYTINAE
a/to„,«te Blandford, 1894a: 114 (Xyleborus) Ei: AU CTCZ GE NL NT P L S K S P SV SZ UK A: FE JA SC
TAI NARi
a/m Niisima, 1909: 160 {Xyleborus)
exiguus Walker, 1859a: 260 (Bostrichus) A:NP AUR NARi ORR
muriceus Eichhoff, 1878a: 392 [= 1878b: 506] (Xyleborus)
gracilis Eichhoff, 1868c: 145 {Xyleborus) Ei: AZ NAR NTR
aspericauda Eggers, 1941a: 106 (Xyleborus)
neogracilis Schedl, 1954a: 46 (Xyleborus)
schoenherri Schedl, 1981: 5 (Xyleborus)
octiesdentatus Murayama, 1931a: 46 (Xyleborus) A: JA SC SCH NARi
saxesenii Ratzeburg, 1837: 167 (Bostrichus) E: AB AL AU AZ BE BU BY CR CT CZ DE EN FR GB GE GR
HU IT LA LT LU MA MC MD NL NR NT PL PT RO SK SL SP ST SV SZ UK YU "Caucasus" N: AG CI
EG LB MO MR TU A: ANH ES FE FUJ GUI GUX HEB HEI HUN IN IS JA JIA HL JLX KA KI KZ MG
NC NE NIN SC SCH SD SHA SHX SY TAI TD TM TR UP WS XIZ YUN ZHE AFR AUR NAR NTR
ORR
aesculi Ferrari, 1867: 22 (Xyleborus)
arbuti Hopkins, 1915a: 64 (Xyleborus)
cinctipennis Schedl, 1980b: 186 (Xyleborus)
decolor Boieldieu, 1859: 473 (Tomicus)
dohrnii Wollaston, 1854: 290 (Tomicus)
floridensis Hopkins, 1915a: 63 (Xyleborus)
frigidus Blackburn, 1885: 193 (Xyleborus)
librocedri Swaine, 1934: 205
opimulus Schedl, 1976: 77 (Xyleborus)
paraguayensis Schedl, 1949b: 276 (Xyleborus)
pecanis Hopkins, 1915a: 63 (Xyleborus)
peregrinus Eggers, 1944c: 142 (Xyleborus)
pseudoangustatus Schedl, 1948: 28 (Xyleborus)
pseudogracilis Schedl, 1937: 169 (Xyleborus)
quercus Hopkins, 1915a: 63 (Xyleborus)
retrusus Schedl, 1940b: 208 (Xyleborus)
sobrinus Eichhoff, 1876: 202 (Xyleborus)
subdepressus Rey, 1883: 142 (Xyleborus)
subspinosus Eggers, 1930b: 203 (Xyleborus)
tsugae Swaine, 1934: 204
schaufussi Blandibrd, 1894a: 117 (Xyleborus) A: JA TAI
speciosus Schedl, 1975a: 457 (Xyleborus) A: SD
subgranulatus Eggers, 1930b: 202 (Xyleborus) A: SD ORR

genus Xyleborus Eichhoff, 1864: 37 type species Bostrichus monographus Fabricius, 1792
Anaeretus Dugès, 1887: 141 type species Xyleborus guanajuatensis Dugès, 1887 (= Bostrichus volvulus Fabricius, 1794)
Boroxylon Hopkins, 1915a: 58 type species Boroxylon stephegynis Hopkins, 1915 (= Phloeotrogus bidentatus
Motschulsky, 1863)
Heteroborips Reitter, 1913b: 82 type species Bostrichus cryptographus Ratzeburg, 1837
Mesoscolytus Broun, 1904:125 type species Apate inurbana Broun, 1880
Notoxyleborus Schedl, 1934: 84 type species Notoxyleborus kalshoveni Schedl, 1934
Progenius Blandford, 1896a: 20 type species ProgeniusfleutiauxiBlandford, 1896 (= Xyleborus subcostatus Eichhoff,
1869)
affinis Eichhoff, 1868c: 401 Ei: AU HU A: IS TAI AFR AUR NAR NTR ORR
fuscobrunneus Eichhoff, 1878b: 372
mascarensis Eichhoff, 1878b: 372
parvus Eichhoff, 1878b: 372
proximus Eggers, 1943b: 66
sacchari Hopkins, 1915a: 64
societatis Beeson, 1935b: 120
subaffin is Eggers, 1933b: 36
aquilus Blandford, 1894a: 109 A: FUJ HUN JA SC SCH
armiger Schedl, 1953a: 28 A: FUJ SCH YUN
 SCOLYTINAE 247
armipennis Schedl, 1953a: 27 A: FUJ YUN
ashuensis Murayama, 1954: 193 A: JA
canus Niisima, 1909: 161 A: JA
ciliatus Eggers, 1940d: 141 A: FUJ
cognatus Blandford, 1896a: 19 A: UP AUR ORR
collis Niisima, 1910: 12 A: JA
conditus Schedl, 1971b: 379 A: NP UP
cryptograph™ Ratzeburg, 1837: 160 (Bostrichus) E: AU BE BY CR CT CZ DE EN FI FR GE GR HU IT LA
MC MD NR NT PL SK SL SP ST SV SZ UK YU "Caucasus" A: ES FE KZ WS
villosus Ratzeburg, 1837: 160 (Bostrichus)
cuneiformis Schedl, 1958c: 104 A.TAI ORR
defensus Blandford, 1894a: 118 A: JA SC
denseseriatus Eggers, 1941c: 225 A: FUJ
diversepilosus Eggers, 1941c: 224 (Xyleborus) A: FUJ TAI
dryographus Ratzeburg, 1837: 167 (Bostrichus) E: AB AU BE BH BU CR CZ FR GB GE GR HU IT LU MC
PL PT SK SL SP ST SZ UK YU N: AG EG LB MO TU A: IN IQ JA TR
donisthorpi Schedl, 1951b: 51
flavus Stephens, 1830: 356 (Tomicus) [HN]
linearis Schedl, 1949b: 273
sampsoni Donisthorpe, 1940: 6
emarginatus Eichhoff, 1878a: 392 [=1878b: 510] A: FUJ GUI HUB HUN JA SCH SHA SHX TAI XIZ YUN
ORR
cinchonae Veen, 1897: 135 (Tomicus)
cordatus Hagedom, 1910a: 12
exesus Blandford, 1894a: 119
palmeri Hopkins, 1915b: 54 (Coptoborus)
semicircularis Schedl, 1973a: 92
terminaliae Hopkins, 1915a: 54 (Coptoborus)
eurygraphus Ratzeburg, 1837: 168 (Bostrichus) E: AU BH CR CT CZ FR GE GR HU IT LU MC MD NL PL
PT SK SP ST SZ UK YU N: AG EG LB MO TU A: TR
fallax Eichhoff, 1878a: 392 [= 1878b: 508] A: NP ORR
amphicranulus Eggers, 1923a: 204
assamensis Stebbing, 1909: 29 (Tomicus)
fallax Stebbing, 1914: 582 [HN]
fastigatus Schedl, 1935b: 402
shoreae Stebbing, 1909: 28 (Tomicus)
ferrugineus Fabricius, 1801: 388 (Bostrichus) A: TAI YE AFR AUR NAR NTR
amplicollis Eichhoff, 1869c: 280
argentinensis Schedl, 1931: 345
conjusus Eichhoff, 1868c: 401
fuscatus Eichhoff, 1868c: 400
hopkinsi Beeson, 1929: 246
insularis Sharp, 1885: 193
nesianus Beeson, 1940: 200
notatus Eggers, 1941a: 107
nyssae Hopkins, 1915a: 66
retusicollis Zimmermann, 1868: 146
rufopiceus Eggers, 1932c: 303
schedli Eggers, 1934b: 83
soltaui Hopkins, 1915a: 66
subitus Schedl, 1949b: 280
tanganus Hagedorn, 1910a: 8
trypanaeoides Wollaston, 1867: 114 (Tomicus)
festivus Eichhoff, 1876: 202 A: JA
ganshoensis Murayama, 1952: 16 A: JA
glabratus Eichhoff, 1877b: 127 A: JA SD TAI NARi ORR
ishidai Niisima, 1909: 156 A: JA
japonicus Nobuchi, 1981b: 150 A: JA
248 SCOLYTINAE
kaimochii Nobuchi, 1981b: 143 A: JA
kojimai Murayama, 1936: 143 A: JA
kraunhiae Niisima, 1910: 14 A: JA
kwnamotoensis Murayama, 1934b: 288 A: JA
kumanoensis Nobuchi, 1985: 25 [RN] A: JA
bispinus Nobuchi, 1981b: 147 [HN]
laetus Niisima, 1909: 159 A: JA
lignographus Schedl, 1953a: 28 A: FUJ
longipilus Eggers, 1926b: 146 A: JA "Korea"
machili Niisima, 1910: 14 A: JA
magnns Niisima, 1910:11 A: JA
metacuneolus Eggers, 1940(1: 150 A: TAI ORR
minutus Blandford, 1894a: 116 A: JA SC TAI ORR
breviusculus Schedl, 1942b: 196
pernitidus Schedl, 1954c: 152
misatoensis Nobuchi, 1981b: 146 A: JA
miyazakiensis Murayama, 1936: 144 A: JA
monographus Fabricius, 1792: 365 (Bostrichus) E: AB AL AU BE BU BY CR CZ DE EN FR GB GE GR HU
IT LA LU MC NL NR NT PL PT RO SK SL SP ST SV SZ UK YU N: AG MO A: IQ SC TR
nitidipennis Roubal, 1937: 67
tuberculosa Herbst, 1794a: 113 (Bostrichus)
montanus Niisima, 1910a: 13 A: JA
muticus Blandford, 1894a: 112 A: JA
nagaoensis Murayama, 1934b: 294 A: JA
nameranus Murayama, 1954: 194 A: JA
ohtoensis Nobuchi, 1981b: 149 A: JA
okinosenensis Murayama, 1961: 31 A: JA
osumiensis Murayama, 1934b: 292 A: JA
percristatus Eggers, 1939a: 12 A: SCH YUN
perforons Wollaston, 1857: 96 (Tomicus) Ei: AZ N: CI MR A: JA GUX SD SHX TAI YUN AFR AUR
ORR
apertus Schedl, 1939c: 355
criticus Schedl, 1950d: 899
cylindrus Schedl, 1951b: 94
denticulus Motschulsky, 1863: 512 (Anodius)
duponti Montrouzier, 1861b: 265 (Bostrichus)
hirsutus Lea, 1894: 321 (Xylopertha)
immaturus Blackburn, 1885: 193
kraatzii Eichhoff, 1868a: 152
philippinensis Eichhoff, 1878b: 374
minimus Schedl, 1955d: 305
testaceus Walker, 1859a: 260 (Bostrichus)
tuberculatus Motschulsky, 1863: 511 (Anodius)
whitteni Beeson, 1935a: 113
pfeilii Ratzeburg, 1837: 168 (Bostrichus) E: AU BE BU CR CT CZ FR GE GR HU IT PL RO SK SL SP ST SZ
UK YU N: AG MO A: FUJ HUN JA SC SCH TR YUN NARi
adumbratus Blandford, 1894a: 115
alni Mulsant & Rey, 1856: 111 (Bostrichus)
vicarius Eichhoff, 1876:203
pinicola Eggers, 1930b: 206 A: TAI ORR
pinivorus Browne, 1980a: 374
praevius Blandford, 1894a: 110 A: JA SC
pumilus Eggers, 1923a: 209 A: SD XIZ ORR
cylindricus Eggers, 1927e: 94
ipidia Schedl, 1972c: 63
planodeclivis Browne, 1974: 70
quercicola Eggers, 1926b: 146 A: JA
 SCOLYTINAE 249
septentrional is Niisima, 1909: 162 A: JA
seriatus Blandford, 1894a: 111 A: FE JA NC SC SCH SHA SHX NARi
aceris Kurentsov, 1941: 188
kalopanacis Kurentsov, 1941: 187
Orientalis Eggers, 1933c: 54
perorientalis Schedl, 1957b: 85
shionomisakiensis Murayama, 1951: 3 A: JA
similis Ferrari, 1867: 23 A: GUA JA (Bonin) JO NP PA SD TAI UP YUN AFR AUR NARi ORR
bucco C. F. C. Schaufuss, 1898b: 212
capita C. F. C. Schaufuss, 1898b: 215
dilatatulus Schedl, 1953c: 127
dilatatus Eichhoff, 1878b: 393
ferrugineus Boheman, 1858: 88 (Bostrichus)
novaguineanus Schedl, 1936b: 530
parvulus Eichhoff, 1868a: 1S2
submarginatus Blandford, 1896b: 223
sinensis Eggers, 1941c: 224 A: FUJ
takinoyensis Murayama, 1953b: 109 A: JA
todo Kôno, 1938:71 A: JA
volvulus Fabricius, 1794: 454 (Bostrichus) A: JA SC TAI AFR AUR NAR NTR ORR
alternons Eichhoff, 1869c: 280
badius Eichhoff, 1869c: 280
granulans Schedl, 1950d: 898
grenadensis Hopkins, 1915a: 65
guanajuatensis Dugès, 1887: 140
hubbardi Hopkins, 1915a: 65
interstitialis Eichhoff, 1878b: 375
rileyi Hopkins, 1915a: 65
schwärzt Hopkins, 1915a: 65
si 1res tris Beeson, 1929: 241
torquatus Eichhoff, 1868a: 146
vagabundus Schedl, 1949b: 277
wakayamensis Nobuchi, 1981b: 144 A: JA
xylographus Say, 1826: 256 (Bostrichus) N: CI A: TR NAR
canadensis Swaine, 1917: 24
inermis Eichhoff, 1868c: 401
genus Xylosandrus Reitter, 1913b: 83 type species Xyleborus morigerus Blandford, 1894
Apoxyleborus Wood, 1980: 90 type species Xyleborus mancus Blandford, 1898
amputates Blandford, 1894b: 575 (Xyleborus) A: FUJ HUN JA SC SCH TAI
melli Schedl, 1938f: 463 (Xyleborus)
beesoni Sana, Maiti &Chakraborti, 1993: 11 A: SD
borealis Nobuchi, 1981c: 34 A: JA SC
brevis Eichhoff, 1877b: 121 (Xyleborus) A: JA NP SC TAI XJZ YUN ORR
cucullatus Blandford, 1894a: 121 (Xyleborus)
compactus Eichhoff, 1876: 201 (Xyleborus) A: FUJ GUA GUI GUX HAI HUB HUN JA SCH TAI YUN ZHE
AFR AUR NARi NTR ORR
morstatti Hagedorn, 1912a: 37 (Xyleborus)
crassiusculus Motschulsky, 1866: 403 (Phloeotrogus) Ei: IT A: ANH BT FUJ GUA GUI HKG HAI HEB
HUB HUN HP JA NC NP SC SCH SD SHA SHN TAI UP XIZ YUN ZHE AFR AUR NARi NTR ORR
bengalensis Stebbing, 1908: 12 (Dryocoetes)
declivigranulatus Schedl, 1936a: 30 (Xyleborus)
ebriosus Niisima, 1909: 154 (Xyleborus)
mascarenus Hagedorn, 1908:379 (Xyleborus)
okoumeensis Schedl, 1935c: 271 (Xyleborus)
semigranosus Blandford, 1896b: 211 (Xyleborus)
semiopacus Eichhoff, 1878b: 334 (Xyleborus)
250 SCOLYTINAE
discolor Blandford, 1898: 429 (Xyleborus) A: FUJ GUA HAI JA (Ryukyus) SCH SD TAI UP YUN AUS
ORR
posticestriatus Eggers, 1939b: 119 {Xyleborus)
diversepilosus Eggers, 1941c: 224 (Xyleborus) A: FUJ TA
germanus Blandford, 1894a: 106 (Xyleborus) Ei: AU BE CR CZ FR GE GG HU IT NL SP ST SZ A: ANH FE
FUJ GUA GUI GUX HAI HEN HUB HUN JA NC SC SCH SHA SHX TAI TR XIZ YUN ZHE NARi
ORR
orbatus Blandford, 1894a: 123 (Xyleborus)
jaintianus Schedl, 1967: 161 (Xyleborus) A: NP OR
mancus Blandford, 1898: 428 (Xyleborus) A: GAN HAI JA TAI XIZ AFR ORR
abruptus Sampson, 1914: 388 (Xyleborus)
formosanus Eggers, 1930b: 186 (Xyleborus)
mesuae Eggers, 1930b: 182 (Xyleborus) A: SD UP ORR
morigerus Blandford, 1894b: 264 (Xyleborus) Ei: AU FR IT A: JO LE SD TAI ORR AFR AUR NARi
NTRi
abruptoides Schedl, 1955d: 298 (Xyleborus)
coffeae Wurth, 1908b: 64 [= 1908a: 4] (Xyleborus)
difficilis Eggers, 1923a: 174 (Xyleborus)
luzonicus Eggers, 1923a: 174 (Xyleborus)
subsimilis Eggers, 1930b: \S6 (Xyleborus) A: HAI SD ORR

tribe Xyloctonini Eichhoff, 1878

genus Scolytomimus Blandford, 1895: 319 type species Scolytomimus dilutus Blandford, 1895
Neoxyloctonus Eggers, 1923a: 143 type species Neoxyloctonus philippinensis Eggers, 1923
Scolytocleptes Schedl, 1962d: 490 type species Scolytomimus maculatus Beeson, 1929
dilutus Blandford, 1895: 320 A: UP

tribe Xyloterini LeConte, 1876

genus Indocryphalus Eggers, 1939a: 6 type species Indocryphalus malaisei Eggers, 1939 (=Xyloterus
intermedius Sampson, 1913)
aceris Niisima, 1910:4 (Xyloterus) A: FE JA NC SC
dainichiensis Murayama, 1954: 191 (Xyloterus)
intermedius Sampson, 1913: 445 (Xyloterus) A: BT HP NP SD XIZ YUN ORR
machili Wood, 1988c: 197 A: UP ORR
major Eggers, 1926b: 148 (Trypodendron) A: JA
ashuensis Murayama, 1950a: 51 (Xyloterus)
pubipennis Blandford, 1894a: 125 (Trypodendron) A: FE JA JIL NC SC TAI
sordidus Blandford, 1894b: 577 (Trypodendron) A: JA
sinensis Eggers, 1941c: 225 (Trypodendron) A: FUJ

genus Trypodendron Stephens, 1830: 353 type species Dermestes domesticus Linnaeus, 1758
Xyloterus Erichson, 1836: 60 type species Bostrichus lineatus Olivier, 1795
Trypodendrum Agassiz, 1846: 380 type species Dermestes domesticus Linnaeus, 1758
Xylotrophus Gistel, 1848: [4] type species Dermestes domesticus Linnaeus, 1758
Trypodendrum Gistel, 1856: 368 type species Dermestes domesticus Linnaeus, 1758
domesticum Linnaeus, 1758: 356 (Dermestes) E: AU BE BH BU BY CR CT CZ DE EN FI FR GB GE GR HU
IR IT LA LT LU MC NL NR NT PL RO SK SL SP ST SV SZ UK YU A: TR NARi
limbatum Fabricius, 1787: 33 (Apate)
dorjitenzingi Schmutzenhofer, 1988: 487 A: BT
gaimaense Murayama, 1937: 369 (Xyloterus) A: JA NC SC
granulatum Eggers, 1933c: 51 A: ES FE MG "Central Asia"
laeve Eggers, 1939b: 122 E: AU CT CZ EN FI GE LA NR NT PL SK SV SZ A- JA "China"
piceumk. Strand, 1946: 172
lineatum Olivier, 1795: 18 (Bostrichus) E: AU BE BH BU BY CR CT CZ DE EN FI FR GB GE GR HU IR IT
LA LS LT LU MC NL NR NT PL RO SK SL SP ST SV SZ YU "Caucasus" N: AG EG LB MO TU A: ES
FE GAN HEI IS JA JIL KZ MG NC NE NMO QIN SC SCH SHA SHN TR WS XIN "Central Asia" NARi
 BARIDINAE 251
bivittatum Kirby, 1837: 192 (Apate)
boréale Swaine, 1917:21
cavifrons Mannerheim, 1843: 297 (Bostrichus)
méridionale Eggers, 1940f: 38
vittiger Eichhoff, 1881: 298
niponicum Blandford, 1894a: 124 A: ES FE JA MG NC NE SC
proximum Niisima, 1909: 165 (Xyloterus) A: FE HEI JA J1L NC NE SC SCH
pulchellum Murayama, 1957a: 585 (Xyloterus) A: JA
signatum Fabricius, 1792: 363 (Apate) E: AB AU BE BU BY CR CT CZ DE EN FI FR GB GE GR HU IT LA
LT MC NR NT PL SK SL SP ST SV SZ UK YU "Caucasus" A: ES FE GAN HEI IN JA MG NC SC SCH
TR WS YUN
obtusum Eggers, 1939b: 121
quereus Eichhoff, 1865: 381 (Xyloterus)
quinquelineatum Adams, 1817:312 (Bostrichus)
suturale Eggers, 1933c: 52
waringii Curtis, 1840: 279 (Bostrichus)

subfamily Scolytinae, nomina dubia

Apate volvulus Fabricius, 1792: 363 E: FR
Bostrichus fagi Nördlinger, 1848: 242 E: GE
Bostrichus carinatus Waltl, 1839: 223 (Bostrychus) E: GE
Bostrichus monocerus Gravenhorst, 1807: 193 E: GE
Bostrichus pilosus Gistel, 1831: 306 (Bostrychus) E: GE
Bostrichus pumilio Gravenhorst, 1807: 193 "Patria ignota"
Ips ater Marsham, 1802: 59 E: GB
Ips inermis Marsham, 1802: 54 E: GB
Ips sulcatus Marsham, 1802: 59 E: GB
Scolytuspusillus Olivier, 1795: (78): 14 E: FR
Scolytus varius Olivier, 1795: (78): 11 E: FR

subfamily Baridinae Schoenherr, 1836

tribe Apostasimerini Schoenherr, 1844
subtribe Madopterina Lacordaire, 1865
genus Centrinopsis Roelofs, 1875a: 185 type species Centrinopsis nitens Roelofs, 1875
Idogenia Pascoe, 1885: 287 type species Idogenia croeeicornis Pascoe, 1885
nitens Roelofs, 1875a: 185 A: FE FUJ JA SC

subtribe Zygobaridina Pierce, 1907

genus Limnobaris Bedel, 1885:183 type species Curculio t-album Linnaeus, 1758
babai Chûjô & Morimoto, 1959b: 153 A: FUJ JA
bedeli Reitter, 1888: 274 (Limnobaris) E: AB
crocopelma Gyllenhal, 1836: 720 (Baridius) E: UK
dolorosa Goeze, 1777: 411 (Curculio) E: AU BE BU CR CT CZ DE FI FR GB GE HU IT NL PL RO SK ST
SV SZ A: TR UZ
barbiellinii Leoni, 1907: 196
dolorosa Gmelin, 1790: 1804 (Curculio) [HN]
junerea Geoffroy, 1785: 121 (Curculio)
funerea Herbst, 1795: 164 (Curculio) [HN]
koltzei Faust, 1892a: 333
pilislriata Stephens, 1831: 10 (Baris)
sculpturata Faust, 1885c: 201 (Baris)
japonica Yoshihara & Morimoto, 1994a: 447 A: JA
jueunda Reitter, 1910: 202 A: FE HEI JA NE SC
sahlbergi Reitter, 1901a: 82 (Limnobaris) N: EG A: KI
scutellaris Reitter, 1888: 273 (Limnobaris) E: ST
252 BARIDINAE
l-album Linnaeus, 1758: 379 (Curculio) E: AU BE BU CZ DE FR GB GE HU IR IT NL NR PL RO SK SP SV
SZ
hypoleuca Marsham, 1802: 274 {Curculio)
martula J. R. Sahlberg, 1892: 223 (Baridius)
nigrina Herbst, 1795: 60 (Curculio)
pusio Boheman, 1844: 173 (Baridius)
reitteri Munster, 1928:281

genus Linogeraeus Casey, 1920: 390 type species Centrinus lineellus LeConte, 1860
punctatissimus Boheman, 1836: 750 (Centrinus) Ei: SP NTR

genus Pertorcus Voss, 1953: 75 type species Pertorcus tibialis Voss, 1953
tibialis basalis Voss, 1958: 76 A: FUJ
tibialispilifer Voss, 1958: 76 A: FUJ
tibialis tibialis Voss, 1953: 76 A: FUJ

tribe Baridini Schoenherr, 1836

subtribe Baridina Schoenherr, 1836
genus Acythogaster Heller, 1940: 111 type species Acythopeus babaulti Hustache, 1928
babaulti Hustache, 1928: 17 (Acythopeus) A: HKG

genus Aegyptobaris Pic, 1899:140 type species Baris arctithorax Pic, 1899
arctithorax Pic, 1899: 140 (Baris) N: EG AFR
lorata G. A. K. Marshall, 1911: 207 (Baris)

genus Anthinobaris Morimoto & Yoshihara, 1996: 31 type species Baridius dispilotus Solsky, 1870
albisquama Zaslavskij, 1956: 349 (Baris) A: NO
dispilota chinensis Frivaldszky, 1892: 117 (Baris) A: LIA
dispilota davidis Fairmaire, 1878: 127 (Baridius) A: CE SCH
reinii Roelofs, 1879b: liv (Baris)
dispilota dispilota Solsky, 1870: 312 (Baridius) A: FE FUJ JA NC NE SC
coreana Kolbe, 1886: 218 (Baris)
freyi Zumpt, 1937a: 28 (Baris) A: JIX
flavonotata Voss, 1941d: 894 (Baris) A: ZHE
fukienensis Voss, 1956b: 40 (Baris) A: SE
kiboshi ihai Nakane, 1963d: 40 (Baris) A: JA
kiboshi kiboshi Nakane, 1963d: 40 (Baris) A: JA
Ijaodunensis Zaslavskij, 1956: 350 (Baris) A: NE
shirozui Morimoto, 1965: 45 (Baris) A: TAI
tainanica Voss, 1932b: 68 (Baris) A: TAI
virgatoidas Voss, 1932b: 66 (Baris) A: YUN
yunnanica Voss, 1932b: 67 (Baris) A: YUN

genus Aulacobaris Desbrochers des Loges, 1892b: 59 type species Baridius corinthius Fairmaire, 1892
Orthobaris Casey, 1920: 345 type species Curculio cuprirostris Fabricius, 1787
andalusiaca H. Brisout de Barneville, 1870a: 42 (Baridius) E: IT SP
angusta angusta Brülle, 1832: 248 (Baridius) E: AU BH BU CR GR HU IT RO SP SK ST A* SY
violacea Gyllenhal, 1836: 708 (Baridius)
angusta occidentalis A. Hoffmann, 1954a: 54 (Baris) N: MO
atlasica Hustache, 1923: 74 (Baris) E: MO
berthieri Pic, 1901b: 29 (Baridius) A: LE
binaghii Solan, 1952: 52 (Baris) E: IT
chlorizans Germar, 1824: 201 (Baris) [NP] E: AU BE CZ GB GE FR HU IT NL PL RO SK SP ST SZ N: AG
MOTU
 BARIDINAE 253
brassicae Du four, 1851: 345 (Baris)
eelfis Gredler, 1866: 355 (Baridius)
chloris Panzer, 1794: 8 (Curculio) [NO]
luczotii Boheman, 1844: 146 (Baridius)
coerulescens chlorodia Boheman, 1844: 162 (Baridius) E: IT
coerulescens coerulescens Scopoli, 1763: 29 (Curculio) E: AU BU CR CZ FR GE GR HU IT PL RO SK SL SP
ST SZ N: AG MO TU A: AP IN ST SY TR
chalybeus Gmelin, 1790: 1759 (Curculio)
coerulescens pulchella P. H. Lucas, 1849: 452 (Baridius) E: IT FR N: AG
coerulescens trapezicollis Desbrochers des Loges, 1892b: 53 (Baridius) N: AG
concinna Boheman, 1844: 164 (Baridius) E: AB ST UK
corinthia Desbrochers des Loges, 1892b: 31 (Baridius) E: IT MA N: AG EG
corsicana Schultze, 1904: 36 (Baris) E: FR IT SP
cribellata Kiesenwetter, 1864: 292 (Baridius) E. GR IT A: CY IS SY TR
heydeni Desbrochers des Loges, 1900b: 41 (Baridius)
stricticollis Desbrochers des Loges, 1892b: 30 (Baridius)
cuprirostris cuprirostris Fabricius, 1787: 100 (Curculio) [NP] E: AU BE BH BU CZ GE FR HU IT NL PL PT
RO SK SP SZ N: AG MO TU
graminea Gmelin, 1790: 1804 (Curculio)
intermedia Hustache, 1930: 192 (Baris)
paris i na Thunberg, 1784a: 16 (Curculio) [NO]
virens Gmelin, 1790: \159 (Curculio)
viridis Geoffroy, 1785: 120 (Curculio) [HN]
viridisericea Goeze, 1777: 410 (Curculio) [NO]
cuprirostris nitidula H. Brisout de Barneville, 1871: 302 (Baridius) E: [without country]
cuprirostris sicula Boheman, 1844: 161 (Baridius) E: IT
cuprirostris yvoniae Pic, 1909a: 114 (Baridius) E: FR
deflexa Desbrochers des Loges, 1900b: 41 (Baridius) E: GR
devillei F. Solari, 1952: 54 (Baris) E: FR
edoughensis Desbrochers des Loges, 1892b: 28 (Baridius) N: AG
fallaciosa A. Hoffmann, 1960b: 167 (Baris) N: MO
fallax H. Brisout de Barneville, 1870a: 42 (Baridius) E: AU BU CZ FR GE IT SK SP SZ
gudenusi gudenusi Schultze, 1901: 212 (Baris) E: AU BH CZ FR GR HU IT SK SP ST A: TD
gudenusipurpurascens Schultze, 1901: 213 (Baris) E: AU
holdereri Hartmann, 1900: 163 (Baris) A: NW
ignifer chloroptera Fuente, 1906: 285 (Baris) E: SP
purpurea Reitter, 1906c: 379 (Baris)
ignifer ignifer Fuente, 1901: 134 (Baris) E: SP
janthina Boheman, 1836: 708 (Baridius) E: AB BU CR GR IT SP ST A: IN IS SY
johanni Korotyaev, 1988b: 148 (Baris) E: ST A : I N K Z T D U Z
squamipes Faust, 1888a: 43 (Baris) [HN]
kaszabi Korotyaev, 1994: 883 A: ES MG
kaufmanni Reitter, 1897b: 220 (Baris) E: AU BH FR IT RO
monlicola A. Solari & F. Solari, 1909: 281 (Baris)
submonticola Hustache, 1938a: 80 (Baris) [RN]
kiesenwetteri Faust, 1890d: 326 (Baris) E: GR A: SY TR
triplagiata Desbrochers des Loges, 1892b: 31 (Baridius)
lepidii Germar, 1824: 200 (Baris) E: AU BE CZ DE GB GE FR HU IT NL PL RO SK SP ST SV SZ UK A:
KZUZ NARi
impunctata Stephens, 1831: 11 (Baris)
nitens Gmelin, 1790: 1758 (Curculio) [HN]
nitidulus O. F. Müller, 1776: 88 (Curculio) [HN]
licens Reitter, 1895a: 27 (Baris) E: AB
malachitica Chevrolat, 1860d: 459 (Baridius) E: SP N: AG MO
mecinoides Desbrochers des Loges, 1892b: 32 (Baridius) N: AG TU
nivalis H. Brisout de Barneville, 1870a: 41 (Baridius) E: BU FR IT SP
254 BARIDINAE
nobilis Desbrochers des Loges, 1896f: 105.2 (Baidius [sic]) N: AG
ochsiA. Hoffmann, 1950b: 197 (Baris) E: FR
perdura Reitter, 1895a: 18 (Baris) A: AB
pertusa Kiesenwetter, 1864: 291 (Baridius) E: BU GR IT SP A: TR
picicornis melillaensis A. Hoffmann, 1954a: 54 (Baris) E: MO
picicornis picicornis Marsham, 1802: 276 (Curculiö) E: AR AU BU CZ DE FR GB GE GR HU IT PL RO SK
SP ST SV SZ A: SY TR
abrotani Germar, 1824: 202 (Baris)
punctata Gyllenhal, 1836: 711 (Baridius)
violacea Grimmer, 1841: 43 (Baris)
picicornis virescens Brülle, 1832: 249 (Baridius) E: GR
prasina alcaidei A. Hoffmann, 1957: 82 (Baris) E: SP
prasina algirica Desbrochers des Loges, 1892b: 61 (Baridius) N: AG
prasinella Hustache, 1938a: 85 (Baris) [RN]
prasina prasina Boheman, 1836: 707 (Baridius) E: IT SP N: AG MO TU
raisae Korotyaev, 1988a: 146 (Baris) E: ST
ruteri Ferragu, 1962: 32 (Baris) A: TR
sareptana Desbrochers des Loges, 1900b: 42 (Baridius) E: ST
siciliensis Desbrochers des Loges, 1892b: 31 (Baridius) E: IT
stierlini Tournier, 1873: 190 (Baris) E: IT
violaceomicansperamoena Reitter, 1907: 296 (Baris) E: KZ
violaceomicans violaceomicans A. Solan & F. Solari, 1905: 534 (Baris) E: MD RO ST
viridipennis Rosenhauer, 1856: 284 (Baridius) E: IT MA SP N: MO

genus Barinomorphoides Morimoto, 1962: 379 type species Barinomorphoides similaris Morimoto, 1962
similaris Morimoto, 1962: 379 A: JA

genus Barinomorphus Morimoto, 1962: 377 type species Barinomorphus antennatus Morimoto, 1962
antennatus Morimoto, 1962: 377 A: JA SC

genus Baris Germar, 1817c: 340 type species Baris artemisiae Herbst, 1795 (=Baris artemisiae Panzer, 1794)
Cyphirhinus Schoenherr, 1826: 276 type species Baridius uncinatus Schoenherr, 1826
abeillei Desbrochers des Loges, 1892b: 27 [DA] A: SY
amanicola Pic, 1905a: 98 [DA] A: SY
analis analis Olivier, 1791: 489 (Curculio) E: AU BE BU CZ FR GB GE HU IT KZ NL PL RO SK SP ST A:
CY
maculatus Townson, 1797:476 (Curculio)
analis insularis Desbrochers des Loges, 1892b: 23 (Baridius) E: FR IT
artemisiae Panzer, 1794: 10 (Curculio) E: AU BE CZ FI FR GE HU IT NL NR PL RO SK ST SV SZ A: ES
FE JA SC
artemisiae Herbst, 1795: 101 (Curculiö)
absinthii Lepeletier & Audinet-Serville, 1825: 296 (Rhynchaenus)
rufitarsis Motschulsky, 1860: 157 (Baridius)
auliensis Reitter, 1901a: 81 [EA] A: KZ
aurifasciata Escalera, 1914: 481 (Baridius) [EA] N: MO
beauprei Hustache, 1938b: 37 [EA] N: AG
behnei Korotyaev, 1988b: 148 [DA] A: TD
belovi Korotyaev, 1988b: 148 [DA] A: UZ
bohemani Schultze, 1897a: 189 [*1897c: 499] [DA] E: BH YU
brevissima Roubal, 1927: 93 [DA] E: ST
cataphracta Peyerimhoff, 1925c: 282 [DA] N: MO
charbinensis Zaslavskij, 1956: 368 [DA] A: HEI,.
chevrolati Coye, 1869: 378 (Baridius) [EA] A: LE
confinis LeConte, 1869: 362 (Baridius) [DA] Ei: FR
consulta Faust, 1886: 156 [EA] A: KA ORR
 BARIDINAE 255
convexicollis Boheman, 1836: 690 (Bariclius) [EA] E: RO ST UK A: IN KZ UZ
copiosepunctata Zaslavskij, 1956: 374 [DA] A: TM
corvina Voss, 1934: 80 A: NE
crambephaga Korotyaev & Gültekin, 1999: 141 [DA] A: TR
dimidiata Reitter, 1897b: 224 [DA] E: AB
dzhungarica Korotyaev, 1988b: 147 [DA] A: KZ
edmundi Korotyaev, 1988b: 145 [DA] A: KZ
egypta Kirsch, 1880: 13 (Baridius) [EA] A: EG
eumicteroides A. Hoffmann, 1962c: 657 [EA] A: IN
ezoana Kôno, 1940b: 78 A: JA SC
fimbriata Faust, 1885c: 202 [EA] A: UZ
foveolatopunctata Motschulsky, 1849: 137 {Baridius) [DA] E: UK
goekseli Korotyaev & Gültekin, 2003b: 752 [EA] A: TR
grandicollis Schultze, 1905: 281 [DA] A: TR
inornata Voss, 1937a: 270 [DA] A: YUN
kiritshenkoi Zaslavskij, 1956: 365 [EA] A: MG
kirschi Faust, 1882c: 435 [EA] E: ST A: IN KZ TM
kirschi Desbrochers des Loges, 1892b: 38 (Baridius) [HN]
kisilkumia Bajtenov, 1974b: 36 [EA] A: KZ
krausei Korotyaev, 1988b: 145 [DA] A: KZ
libanicola Pic, 1900d: 90 (Baridius) [DA] A: LE
limbata H. Brisout de Barneville, 1870a: 56 (Baridius) E: AB KZ ST UK
nigra Desbrochers des Loges, 1892b: 40 (Baridius)
nigra Reitter, 1895a: 14 [HN]
subpolita Zaslavskij, 1956: 368
longicollis Faust, 1890e: 473 [DA] A: MG
maracandica Zaslavskij, 1956: 372 [DA] A: UZ
mauretanica Schultze, 1904: 37 [DA] N: AG
melanoleuca Iablokoff-Khnzorian, 1964: 175 [DA] A: TD
memnonia memnonia Boheman, 1836: 657 (Baridius) [EA] E: IT ST A: CT CY IN IS KI KZ TD TM UZ
intercalaris Faust, 1885c: 200
schwarzenbergii Hochhuth, 1847: 553 [108 in reprint] (Baridius)
memnonia turgaica Zaslavskij, 1956: 365 [EA] A: KZ
menthae Kôno, 1937: 316 [DA] A: JA SC
nesapia Faust, 1887g: 303 E: AB AU BU ES GE GG PL ST N: EG A: KI KZ TD TM TR
nigritarsis Boheman, 1844: 156 (Baridius) [DA] E: "Caucasus"
noaeae Becker, 1875: 138 (Baridius) [EA] E: AB ST A: IN
brisouti Faust, 1882c: 433
globicollis Reitter, 1912a: 98 (Neobaris)
parumpunctata Fairmaire, 1875: 535 (Baridius) [DA] E: IT N: TU
pedemontana Desbrochers des Loges, 1908e: 92 [DA] N: IT
perlucida Reitter, 1902b: 192 [DA] E: AB
piliventris Zaslavskij, 1956: 366 [DA] A: FE NC NE
pilosa Roelofs, 1875a: 182 [DA] A: JA SC
praeambula Faust, 1897: 89 [EA] A: TM
propinqua Desbrochers des Loges, 1900b: 43 (Baridius) [DA] N: AG
przewalskyi Zaslavskij, 1956: 365 [EA] A: NE
pseudospitzyi Zaslavskij, 1956: 367 [DA] A: FE
pygidialis Voss, 1937a: 271 [EA] A: SCH
quinquecarinata Zaslavskij, 1956: 351 [EA] A: BEI
rotroui Peyerimhoff, 1949: 298 [EA] N: MO
rufa Boheman, 1844: 166 (Baridius) [DA] E: BU FR IT SP
semistriata Boheman, 1836: 695 (Baridius) [DA] E: AB BU MO ST UK A: CY KI TR
gimmerthalii Hochhuth, 1847: 560 [115 in reprint] (Baridius)
picitarsis Boheman, 1836: 694 (Baridius)
256 BARIDINAE
striatopunctata Reitter, 1895a: 20
shaowuensis Voss, 1953: 76 [EA] A: TAI
serbica Schultze, 1897a: 189 [= 1897c: 500] [DA] E: YU
serraticollis Hustache, 1923: 73 [DA] N: MO
spitzyi Hochhuth, 1847: 564 [119 in reprint] E: AR BU RO ST
steppensis Roubal, 1935: 208 A: AU SK
subanalis Pic, 1900d: 90 {Baridius) [DA] A: SY
subferruginea Reitter, 1910: 201 [DA] E: IT
subtilina Reitter, 1899b: 208 [DA] E: GG
suffriani Hochhuth, 1847: 566 [121 in reprint] {Baridius) [EA] E: GG
sulcata Bohemaa, 1836: 673 {Baridius) [EA] E: HU KZ RO ST UK A: WS
tadzhica Korotyaev, 1988b: 147 [DA] A: TD
ussuriensis chinganensis Zaslavskij, 1956: 367 [DA] A: NE
ussuriensis ussuriensis Zaslavskij, 1956: 367 [DA] A: FE NC
volgensis Zaslavskij, 1956: 373 [DA] E: ST
v-signum Voss, 1937a: 270 [DA] A: TAI YUN

genus Bariscelis Reitter, 1895a: 6 type species Bariscelis fausti Reitter, 1895
fausti Reitter, 1895a: 6 A: TM

genus Batobaris Morimoto & Yoshihara, 1996: 34 type species Batobaris rubi Morimoto & Yoshihara, 1996
fasciata Voss, 1958: 86 {Acythopheus) A: FUJ
rubi Morimoto & Yoshihara, 1996: 35 A: JA

genus Calyptopygus G. A. K. Marshall, 1948: 466 type species Calyptopygus ellipticus G. A. K. Marshall,
1948
albosparsus Reitter, 1910: 203 {Limnobaris) [DA] A: FE JA SC
kumei Yoshihara & Morimoto, 1997: 1 A: JA TAI

genus Chelonebarus G. A. K. Marshall, 1940: 67 type species Chelonebarus partimpunctatus G. A. K.

Marshall, 1940
burmanus G. A. K. Marshall, 1948: 460 A: TAI

genus Cosmobaris Casey, 1920: 344 type species Cosmobaris americana Casey, 1920 (= Baris scolopacea
Germar, 1819)
discolor Boheman, 1836: 703 {Baridius) N: AG EG A: CY IS JO SY AFR AURi NARi PACi
alboseriata Reitter, 1908: 54 {Baris)
rhagodiae Rheinheimer, 1992: 191 {Baris)
soricina Fairmaire, 1884: brix {Baridius)
squamulata Pic, 1896b: 115 {Baridius)
vicina H. Brisout de Barneville, 1870a: 39 {Baris)
scolopacea Germar, 1819: 132 {Baris) E: AR AU BU CZ FR GB GR HU IT NL RO SK SP ST SZ UK N: EG
TN A: BEI ES FE HKG IN JA KZ NC SC TD TM TR UZ NARi
americana Casey, 1920: 344
borkhsenii Zaslavskij, 1956: 357 {Baris)
carnerii Pic, 1922: 30 {Baridius)
colorata Boheman, 1836: 700 {Baridius)
lebedevi Roubal, 1929: 56 {Baris)
orientalis Roelofs, 1875a: 184 {Baris)
pallidicornis Boheman, 1836: 702 {Baridius)
parvula Boheman, 1836: 701 {Baridius)
sionilli Hayes, 1936:28
squamiger Hayes, 1936: 27
vestita Perris, 1858: 143 {Baridius)
 BARIDINAE 257
genus Elasmobaris Reitter, 1913a: 72 type species Lissotarsus balassogloi Faust, 1881 (= Baridius
alboguttatus Brisout)
Lissotarsus Faust, 1881: 327 [HN] type species Lissotarsus balassogloi Faust, 1881 (= Baridius alboguttatus Brisout)
aegyptiaca Desbrochers des Loges, 1891a: 31 {Lissotarsus) N: EG
demaisoni Desbrochers des Loges, 1892b: 64 (Baridius) [RN]
alboguttata alboguttata H. Brisout de Bameville, 1870a: 40 (Baridius) E: AB N: AG EG MO A: AF IN IQ
MG RU QA SA SY TD UZ
balassogloi Faust, 1881: 328 (Lissotarsus)
alboguttata subelongata Pic, 1921: 10 (Lissotarsus)
annularis Faust, 1881: 331 (Lissotarsus) [EA] A: TM
bedeli Faust, 1891b: 334 (Lissotarsus) [EA] N: AG TU
biskrensis Chobaut, 1897: 39 (Lissotarsus) [EA] A: AG MO
cretacea Reitter, 1895a: 7 (Lissotarsus) N: EG
nigropicta Reitter, 1895a: 7 (Lissotarsus) E: AB
signifera Faust, 1881: 331 (Lissotarsus) [EA] A: KI

genus Eremobaris Zaslavskij, 1956: 355 type species Baridiuspicturatus Ménétriés, 1849
albopicta Reitter, 1895a: 22 (Baris) E: AB
picturata Ménétriés, 1849: 263 (Baridius) E: BU FR GR IT SK SP ST N: AG EG MO A: ES IN
opiparis Jacquelin du Val, 1853: 715 (Baridius)

genus Formosobaris Morimoto & Yoshihara, 1996: 26 type species Formosobaris conicotylosa Morimoto &
Yoshihara, 1996
conicotylosa Morimoto & Yoshihara, 1996: 27 A: TAI

genus Hollisiella Alonso-Zarazaga & Lyal, 1999: 90 [RN] type species Barisoma carbo Faust, 1888
Barisoma Faust, 1888c: 309 [HN] type species Barisoma carbo Faust, 1888
carbo Faust, 1888c: 310 A: HKG

genus Labiaticola Alonso-Zarazaga & Lyal, 1999: 90 type species Baris teruelensis Hustache, 1927
araxicola Reitter, 1895a: 12 (Baris) E: AB
atricolor Boheman, 1844: 91 (Baridius) E: AR AU BU GR HU IT PL RO ST YU A: NW SY TR
landgrebei Hochhuth, 1847: 558 (Baridius) [unjustified emendation]
landgrebii Hochhuth, 1847: 558 [113 in reprint] [original spelling]
jonicus Miller, 1862: 280 (Baris)
turcicus H. Brisout de Bameville, 1870a: 51 (Baridius)
cariniventris cariniventris A. Solari & F. Solari, 1909: 279 (Baris) N: AG MO
cariniventris joffrei Hustache, 1926: 318 (Baris) E: FR
clandestinus Faust, 1887i: 306 (Baris) A: TM UZ
crassipes Faust, 1884: 462 (Baris) A: TM
despicatus Faust, 1889a: 207 (Baris) E: AB AR A: IN TR
korbi Schultze, 1900: 383 (Baris) E: SP
mêlas Boheman, 1836: 705 (Baridius) E: IT SP ST A: KZ
litigiosus Desbrochers des Loges, 1892b: 27 (Baridius)
phlomidis Becker, 1892: 70 (Baris)
renardii Hochhuth, 1847: 562 [117 in reprint] (Baridius)
ocellatus Desbrochers des Loges, 1900b: 43 (Baridius) E: IT
rectirostris rectirostris Faust, 1892b: 63 (Baris) A: NW TD TM
rectirostris subcurvatus Zaslavskij, 1956: 359 (Baris) A: "Central Asia"
rugulipennis Reitter, 1895a: 12 (Baris) E: AB
sibiricus Faust, 1890b: 102 (Baris) E: ST A: ES TM
syriacus Faust, 1897: 90 (Baris) A: IN SY TR
teheranicus A. Hoffmann, 1956: 245 (Baris) A: IN
teruelensis Hustache, 1927: 26 (Baris) E: SP
258 BAR1DINAE
genus Malvaevora Zaslavskij, 1956: 358 type species Curculio limidus Rossi, 1792
timida polita Reitter, 1895a: 20 (Baris) A: "Caucasus" E: GR N: AG MO
timida timida Rossi, 1792: 37 {Curculio) E: AU BU FR HU IT KZ RO SK SL SP ST SZ IN: AG MO TU A:
CY IN KZ TM TR

genus Maurobaris Alonso-Zarazaga, 2005: 91 type species Baridius spoliatus Boheman, 1836
spoliata Boheman, 1836: 692 {Baridius) E: FR GR IT SP N: AG MO TU

genus Melaleucas Chevrolat, 1882a: xc type species Melaleucus x-littera Chevrolat, 1882
sellatus Boheman, 1844: \7A (Baridius) E: IT SP N: AG CI MO
x-signum Reitter, 1895a: 23 (Baris) N: EG
zarudnyi; Zaslavskij, 1956: 356 (Baris) A: IN

genus Melitobaris Peyerimhoff, 1929b: 40 type species Melitobaris dumonti Peyerimhoff, 1929
dumonti Peyerimhoff, 1929b: 40 N: TU

genus Melanobaris Alonso-Zarazaga & Lyal, 1999: 90 type species Baridius morio Boheman, 1844
atramentaria Boheman, 1836: 696 (Baridius) [DA] E: AB AU BU CZ HU PL SK SP
atronitens algirica Desbrochers des Loges, 1892b: 25 (Baridius) [DA] N: AG
atronitens atronitens Chevrolat, 1861: 121 (Baridius) [DA] E: FR SP N: AG MO TU
atronitens gallica Desbrochers des Loges, 1892b: 25 (Baridius) [DA] N: FR
balassogloi Faust, 1885c: 200 (Baris) [DA] A: TD TM UZ
carbonaria Boheman, 1836: 676 (Baridius) [DA] E: AU CZ HU KZ RO SK ST UK A: AF
dalmatina H. Brisout de Barneville, 1870a: 53 (Baridius) [DA] E: BU CR IT SK ST UK
elevata Reitter, 1899a: 160 (Baris) [DA] E: SP
erysimi erysimi Chobaut, 1917: 109 (Baris) [DA] E: FR SP
erysimi erysimioides A. Hoffmann, 1942: 17 (Baris) [DA] E: FR SP
gloriae Korotyaev & Gültekin, 2003a: 97 [DA] A: TR
hochhuthi Faust, 1882c: 435 (Baris) [DA] E: AB AR GG ST UK A: IN TM
laticollis Marsham, 1802: 276 (Curculio) [NP, DA] E: BE BU CZ GB GE HU IT PL PT RO SK SP N: AG MR
A:SY
absinthii Panzer, 1794: 9 (Curculio) [emendation] [NO]
apsinthii Panzer, 1794: 9 (Curculio) [original spelling] [NO]
atriplicis Fabricius, 1777: 225 (Curculio) [NO]
glabra Herbst, 1784a: 77 (Curculio) [NO]
nitens Fabricius, 1792:403 (Curculio) [HN]
picina Germai, 1824: \99 (Baris)
subtarsalis Rey, 1895a: 26 (Baris)
morio Boheman, 1844: 174 (Baridius) E: AU BE CZ FR GE GR HU IT SP N: AG EG MO TU
lethierryi Desbrochers des Loges, 1892b: 24 (Baridius)
resedae Bach, 1856: 243 {Baris)
pyrenaicaA. Hoffmann, 1933: 151 (Baris) [DA] E: SP
quadraticollis caucasica Schultze, 1897b: 397 (Baris) [DA] E: AB
quadraticollis quadraticollis Boheman, 1836: 693 (Baridius) [DA] E: BU FR IT SP N: AG MO TU
sulcicollis Chevrolat, 1866: 106 (Baridius)
quadraticollis semirubra Bedel, 1903: 370 (Baris) [DA] N: MO
semiopaca Reitter, 1895a: 20 (Baris) [DA] E: AB
sinapis galliae Tempère, 1961: 95 (Baris) [DA] E: FR
sinapis sinapis Normand, 1937: 261 (Baris) [DA] N: TU
genus Moreobaris Morimoto & Yoshihara, 1996: 40 type species Baris deplanata Roelofs, 1875
castanea Morimoto & Miyakawa, 1985: 34 (Baris) A: JA
deplanata Roelofs, 1875a: 183 (Baris) A: JA JIA SC ZHE
kozuensis Morimoto & Miyakawa, 1985: 36 (Baris) A: JA
repandirostris Zaslavskij, 1956: 351 (Baris) A: FE
 BARIDINAE 259
rubricata Hustache, 1921a: 94 (Baris) A: JA

genus Neobaris Reitter, 1895a: 5 type species Baridius duplicatus Schoenherr, 1832
duplicata Schoenherr, 1832: 223 {Baridius) A: IN ST
logunovi Legalov, 1999: 376 [DA] A: BS

genus Nespilobaris Morimoto & Yoshihara, 1996: 41 type species Acythopeus parabasimaculatus Morimoto
& Lee, 1992
accidirostris Zaslavskij, 1956: 352 (Baris) A: FE
basimaculata Voss, 1958: 85 (Acythopeus) A: SE
nipponica Kôno, 1928: 174 (Baris) A: JA SC
parabasimaculata Morimoto & Lee, 1992: 10 (Acythopeus) A: JA SC

genus Omobaris G. A. K. Marshall, 1927: 217 type species Omobaris calanthes G. A. K. Marshall, 1927
chloranthae Kojima & Yoshihara, 2009: 174 [DA] A: JA TAI
parvula Kojima & Yoshihara, 2009: 177 [DA] A: JA

genus Pharcidobaris Morimoto & Yoshihara, 1996: 29 type species Pharcidobaris miyamotoi Morimoto &
Yoshihara, 1996
miyamotoi Morimoto & Yoshihara, 1996: 30 A: JA
suvorovi Reitter, 1910: 201 (Baris) A: ES FE NC NE

genus Phrissoderes G. A. K. Marshall, 1948: 459 type species Phrissoderes costalis G. A. K. Marshall, 1948
Centrinoplesius Voss, 1958: 77 type species Centrinoplesius exsculptus Voss, 1958
Pseudorhyssematus Morimoto, 1962: 381 type species Rhyssomatus rufitarsis Roelofs, 1875
exsculptus Voss, 1958: 77 (Centrinoplesius) A: FUJ
rufitarsis Roelofs, 1875a: 159 (Rhyssomatus) A: JA

genus Psilarthroides Morimoto & Miyakawa, 1985: 41 type species Psilarthroides humuli Morimoto &
Miyakawa, 1985
czerskyi Zaslavskij, 1956: 366 (Baris) A: FE JA NE SC
humuli Morimoto & Miyakawa, 1985: 42
lafèri Egorov, 1978: 158 (Baris)

genus Pteridobaris Morimoto & Yoshihara, 1996: 27 type species Baris maritima Roelofs, 1875
maritima Roelofs, 1875a: 181 (Baris) A: JA

genus Taiwanobaris Morimoto & Yoshihara, 1996: 18 type species Taiwanobaris femorata Morimoto &
Yoshihara, 1996
femorata Morimoto & Yoshihara, 1996: 18 A: TAI

genus Theryus Pic, 1897b: 29 type species Theryus longicollis Pic, 1897
longicollis Pic, 1897b: 29 N: AG
albus Abeille de Perrin, 1898: 257 (Baridius)

genus Turkmenobaris Zaslavskij, 1956:357 type species Baris kryzhanovskii Zaslavskij, 1956
kryzhanovskii Zaslavskij, 1956: 357 (Baris) A: TM
mirifica Iablokoff-Khnzorian, 1959: 79 E: AR

genus Ulobaris Reitter, 1895a: 5 type species Baridius loricatus Boheman, 1836
kuchenbeisseri Hartmann, 1899b: 51 (Baris) A: NO
loricata Boheman, 1836: 716 (Baridius) E: BU RO ST A: KI KZ MG TD TM UZ
crinipes H. Brisout de Barneville, 1870a: 45 (Baridius)
260 BARIDINAE
tribe Baridini, species incertae sedis
Calandra uniseriatus Dufour, 1843: 8 E: FR

tribe Madarini Jekel, 1865

subtribe Leptoschoinina Lacordaire, 1865
genus Acythopeus Pascoe, 1874: 61 type species Acythopeus tristis Pascoe, 1874
Amyclides Hustache, 1956: 162 type species Baris perrieri Fairmaire, 1901
subgenus Carpobaris Zaslavskij, 1956: 355 type species Baris granulipennis Tournier, 1873
Parathesapeuta Voss, 1964a: 588 type species Athesapeuta colocynthae Voss, 1964 (=Baridius granulipennis Tournier,
1873)
curvirostris curvirostris Boheman, 1844: 190 {Baridius) N: AG AFR
traegardhi Aurivillius, 1905: M (Baris)
curvirostris granulipennis Tournier, 1873: 190 (Baridius) N: EG A: AE IQ IS JO SA
colocynthae Voss, 1964a: 588 (Athesapeuta)
cucumidis Voss, 1969a: 63 (Athesapeuta)
globulicollis Voss, 1969a: 62 (Athesapeuta)
curvirostrispersicus Thompson, 1973: 34 A: AF IN
inflatirostris Voss, 194Id: 893 A: ZHE

genus Arganthonius Alonso-Zarazaga, 2005: 90 type species Baris novaki Penecke, 1936
albomaculatus H. Brisout de Barneville, 1870a: 40 (Baris) E: SP
novaki Penecke, 1936: 220 (Baris) E: PT SP N: MO

genus Athesapeuta Faust, 1895b: 313 type species Athesapeuta motschulskyi Voss, 1958 (= Baridius
subsignatus Motschulsky, 1866)
amoenaVoss, 1958: 80 A: FUJ
chinensis Faust, 1895b: 316 A: SHG
latefasciata Voss, 1958: 81 A: FUJ
sculpticollis Voss, 1958: 79 A: FUJ

genus Cucubaris Alonso-Zarazaga, 2005: 91 type species Baris villae Comolli, 1837
villae Comolli, 1837: 35 (Baris) E: AU HU IT RO SK ST SZ A: AF

genus Dendrobaris Egorov, 1976: 64 type species Baris tatyanae Egorov, 1976
Spilobaris Morimoto & Yoshihara, 1996: 37 type species Acythopeus insularis Morimoto & Miyakawa, 1985
flavosignata Roelofs, 1879a: 301 (Baris) A: JA
insularis Morimoto & Miyakawa, 1985: 39 (Acythopeus) A: JA
kurentzovi Egorov, 1976: 67 (Baris) A: FE
maculata Roelofs, 1879a: 299 (Baris) A: JA SC TAI
tatyanae Egorov, 1976: 67 (Baris) A: FE

genus Lepidomyctides Yoshihara & Morimoto, 1994b: 723 type species Lepidomyctides nagaii Yoshihara &
Morimoto, 1994
nagaii Yoshihara & Morimoto, 1994b: 725 A: JA TAI

genus Paracythopeus Voss, 1958: 86 type species Paracythopeus perrugosus Heller 1940
collaris Voss, 1958: 86 A: FUJ

genus Parallelodemas Faust, 1895b: 306 type species Parallelodemas perfectum Faust 1895
impar Voss, 194Id: 895 A: ZHE

genus Pellobaris Morimoto & Yoshihara, 1996:16 type species Baris melancholka Roelofs 1875
melancholica Roelofs, 1875a: 181 A: FE JA SC
 BARIDINAE 261

subtribe Madarina Jekel, 1865

genus Orchidophilus Buchanan, 1935: 45 type species Orchidophilus peregrinator Buchanan, 1935
aterrimus C. O. Waterhouse, 1874: 226 (Baridius) Ei: FR GB GE NL SV Ai: JA AFRi NARi NTRi ORR
e6wn/er Pascoe, 1887c: 359 (Baris) Ei: GB ORR
gilvonotatus Barber, 1918: 17 (Acythopeus)
epidendri Murray, 1869: 1279 (Centrinus) Ei: GB GE AFRi AUSi NARi ORR
genuinus Pascoe, 1887c: 359 (Acythopeus)
orchidearum Kolbe, 1906: 4 (Apotomorhinus)
orchivora Blackburn, 1900a: 61 (Baris)
hisamatsui Kojima & Yoshihara, 2009: 179 (Omobaris) A: JA
ran Morimoto, 1994: 236 A: JA SC TAI

subtribe Tonesiina Alonso-Zarazaga & Lyal, 1999

genus Manilabaris Zimmerman, 1940: 314 type species Manilabaris Cucurbitae Zimmerman, 1940
armipes Roelofs, 1875a: 183 (Baris) A: JA

tribe Neosharpiini A. Hoffmann, 1956

genus Caenobaris Nasreddinov, 1980: 476 type species Caenobaris zaslavskii Nasreddinov, 1980
zaslavskii Nasreddinov, 1980: 477 A: TD

genus Eumycterus Schoenherr, 1838: 1083 type species Eumycterus albosquamulatus Boheman, 1838
albosquamulatus Boheman, 1837: 1084 E: GR N: AG EG MO A: AF CY IN IS SY TM TR UZ AFR
tenuirostris H. Brisout de Barneville, 1870a: 39 (Baridius)
gracilis Voss, 1958: 78 (Eumycterus) [DA] A: FE FUJ NC
laodioides Nakane, 1963d: 40 A: JA

genus Neosharpia A. Hoffmann, 1956: 247 type species Neosharpia artemisiae A. Hoffmann, 1956
artemisiae A. Hoffmann, 1956: 247 (Neosharpia) A: IN
capucina Faust, 1881: 332 (Lissotarsus) A: KZ
262

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Schedl K. E. 1952e: Fauna Argentinensis V. 96. Contribution to the morphology and taxonomy of the
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Schedl K. E. 1953c: Bark and ambrosia - beetles from Indochina. 127 Contribution to the morphology and
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Voss E. 1936a: Über unbeschriebene Curculioniden der palaearktischen Region (Col., Cure). (61. Beitrag zur
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Voss E. 1936b: Monographie der Rhynchitinen-Tribus Auletini. III. Teil der Monographie der Rhynchitinae-
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97: 279-289.
Voss E. 1937a: Über ostasiatische Curculioniden (Col., Cure). (70. Beitrag zur Kenntnis der Curculioniden.).
Senckenbergiana 19: 226-282.
Voss E. 1937b: Über Arten und Gattungen der Unterfamilien Belinae, Rhynchitinae und Attelabinae (Cure,
Col.). (69. Beitrag zur Kenntnis der Curculioniden). Stettiner Entomologische Zeitung 98: 199-209.
Voss E. 1938a: Bemerkenswerte Ergebnisse einer Revision der Rhynchitinae der Sammlung Stierlin. (71.
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Berlin-Dahlem 5: 62-65.
Voss E. 1938b: Monographie der Rhynchitinen-Tribus Deporaini sowie der Unterfamilien Pterocolinae -
Oxycoryninae (Allocorynini). VII. Teil der Monographie der Rhynchitinae-Pterocolinae. (73. Beitrag zur
Kenntnis der Curculioniden.). Stettiner Entomologische Zeitung 99: 59-117.
Voss E. 1938c: Monographie der Rhynchitinen-Tribus Rhynchitini. 2. Gattungsgruppe: Rhynchitina. V. 2. Teil
der Monographie der Rhynchitinae - Pterocolinae. (45. Beitrag zur Kenntnis der Curculioniden.).
Koleopterologische Rundschau 24: 129-152.
Voss E. 1938d: Monographie der Rhynchitinen-Tribus Rhynchitini. 2. Gattungsgruppe: Rhynchitina. V. 2. Teil
der Monographie der Rhynchitinae - Pterocolinae. (45. Beitrag zur Kenntnis der Curculioniden.). (Cont.).
Koleopterologische Rundschau 24: 153-171.
Voss E. 1939a: [new taxa]. In: Dalla Torre K. W. & Voss E.: Pars 167. Curculionidae III Rhynchitinae II,
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Voss E. 1939b: [new taxa]. In: Koch C : Die Käfer der libyschen Ausbeute des Herrn Georg Frey. Mitteilungen
der Münchnneer Entomologischen Gesellschaft 29: 216-293, pis VIII-DC.
Voss E. 1939c: Eine unbeschriebene Rhynchites-Art aus der Mandschurei (Col. Cure). 78. Beitrag zur
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Voss E. 1939d: Über einige ostasiatische Rhynchitinen, Attelabinen und Apoderinen (Col. Cure). (81. Beitrag
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Voss E. 1939e: Über vorwiegend unbeschriebene Rüssler der paiäarktischen Region (Col. Cure) (84. Beitrag zur
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Voss E. 1940: Einige Rüsslerarten aus Turkestan und Mandschukuo. (Col. Cure) (85. Beitrag zur Kenntnis der
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Voss E. 1941b: Bemerkenswerte Rüsselkäfer aus Manschukuo. (Coleoptera: Curculionidae). (90. Beitrag zur
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Voss E. 1941c: Monographie der Rhynchitinen-Tribus Rhinocartini sowie der Gattungsgruppe Eugnamptina der
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Voss E. 1943d: Einige neue Rüslerarten aus Argentina und Paraguya (Coleoptera: Curculionidae). Arbeiten über
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Voss E. 1949: Über einige in Fukien (China) gesammelte Rüssler. III. (Col. Cure). 113. Beitrag zur Kenntnis
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Voss E. 1955: Bemerkenswerte Ergebnisse einer Revision der Attelabiden des Ungarischen
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358

INDEX
Family-group names

Aglycyderini Wollaston, 1864 109 Corrhecerini Lacordaire, 1865 92

Allandrini Pierce, 1930 90 Corthylina LeConte, 1876 215
ANTHRIBIDAE Billberg, 1820 90 Corthylini LeConte, 1876 215
Anthribinae Billberg, 1820 90 Cratoparini LeConte, 1876 92
Anthribini Billberg, 1820 91 Cryphalini Lindemann, 1876 217
APIONIDAE Schoenherr, 1823 148 Cryptodermatinae Bovie, 1908 185
Apioninae Schoenherr, 1823 148 Crypturgini LeConte, 1876 226
Apionini Schoenherr, 1823 148 CURCULIONIDAE Latreille, 1802 201
Aplemonini Kissinger, 1968 149 CURCULIONOIDEA Latreille, 1802 90
Apoderini Jekel, 1860 129 Cyladinae Schoenherr, 1823 142
Apolectinae Lacordaire, 1865 104 Cyphagoginae Kolbe, 1892 144
Apolectini Lacordaire, 1865 104 Cyphagogini Kolbe, 1892 144
Apostasimerini Schoenherr, 1844 251 Deporaina Voss, 1929 115
Araecerini Lacordaire, 1865 104 Deporaini Voss, 1929 114
Arrhenodini Lacordaire, 1865 142 Desmidophorini Morimoto, 1962 185
Arthrostenini Reitter, 1913 192 Diamerini Hagedorn, 1909 204
Aspidapiini Alonso-Zarazaga, 1990 153 Diapodini Strohmeyer, 1914 203
ATTELABIDAE Billberg, 1820 129 Diocalandrini Zimmerman, 1993 186
Attelabini Billberg, 1820 134 Discotenini Lacordaire, 1865 93
Auletina Desbrochers des Loges, 1908 109 Doydirhynchini Pierce, 1916 90
Auletini Desbrochers des Loges, 1908 109 Dryocoetini Lindemann, 1876 228
Auletobiina Legalov, 2001 109 DRYOPHTHORIDAE Schoenherr, 182?
Baridina Schoenherr, 1836 252 185
Baridinae Schoenherr, 1836 251 Dryophthorinae Schoenherr, 1825 185
Baridini Schoenherr, 1836 252 Ecelonerini Lacordaire, 1865 93
Basitropidini Lacordaire, 1865 92 Eremoxenini Semenov, 1892 143
BELIDAE Schoenherr, 1826 109 ERIRHINIDAE Schoenherr, 1825 192
BRACHYCERIDAE Billberg, 1820 182 Erirhininae Schoenherr, 1825 192
Brachycerinae Billberg, 1820 182 Erirhinini Schoenherr, 1825 193
Brachycerini Billberg, 1820 182 Eugnamptini Voss, 1930 118
BRENTIDAE Billberg, 1820 142 Euopini Voss, 1925 137
Brentinae Billberg, 1820 142 Euscelophilini Voss, 1925 139
Byctiscina Voss, 1923 112 Exapiini Alonso-Zarazaga, 1990 158
Byctiscini Voss, 1923 112 Himasthlophallini Zherikhin, 1991 198
Catapiina Alonso-Zarazaga, 1990 164 Hoplapoderini Voss, 1926 140
Ceratapiini Alonso-Zarazaga, 1990 154 Hoplopisthiini Senna & Calabresi, 1919 146
Chonostropheina Morimoto, 1962 114 Hylastim* LeConte, 1876 204
Choraginae Kirby, 1819 104 Hylesinini Erichson, 1836 206
Choragini Kirby, 1819 106 Hylesinitae Erichson, 1836 204
Cimberidinae Gozis, 1882 90 Hylurgini Gistel, 1848 208
Cimberidini Gozis, 1882 90 Hyorrhynchini Hopkins, 1915 210
Cisanthribini Zimmerman, 1994 107 Hypoborini Ntisslin, 1911 210
Clitostylini Voss, 1929 136 IpiniBedel, 1888 233
Corimaliinae Alonso-Zarazaga, 1989 177 Ithystenini Lacordaire, 1865 147
 INDEX 359
Ixapiini Alonso-Zarazaga, 1990 159 RAYMONDIONYMIDAE Reitter, 1913
Jordanthribini Morimoto, 1978 94 198
Kalcapiini Alonso-Zarazaga, 1990 160 RHYNCHITIDAE Gistel, 1848 109
Leptoschoinina Lacordaire, 1865 260 Rhynchitinae Gistel, 1848 109
Listrobyctiscina Legalov, 2003 114 Rhynchitini Gistel, 1848 120
Litosomini Lacordaire, 1865 186 Rhynchophorinae Schoenherr, 1833 186
Madarina Jekel, 1865 261 Rhynchophorini Schoenherr, 1833 187
Madarini Jekel, 1865 260 Scolytinae Latreille, 1804 204
M adopterina Lacordaire, 1865 251 Scolytini Latreille, 1804 237
Malvapiini Alonso-Zarazaga, 1990 162 Scolytitae Latreille, 1804 215
Mauiini Valentine, 1989 94 Scolytoplatypodini Blandford, 1893 240
Mecocerini Lacordaire, 1865 94 S intorini Lacordaire, 1865 98
Metapiini Alonso-Zarazaga, 1990 163 Sphenophorini Lacordaire, 1865 189
Micracidini LeConte, 1876 236 Stenopelmini LeConte, 1876 198
Microtrachelizini Zimmerman, 1994 147 Stenorhinini Valentine, tribe nov. 107
Mycteini Morimoto, 1972 94 Stereodermini Sharp, 1895 146
NANOPHYIDAE Gistel, 1848 177 Stromboscerinae Lacordaire, 1865 192
Nanophyinae Gistel, 1848 178 Svetlanaebyctiscina Legalov, 2003 114
NEMONYCHIDAE Bedel, 1882 90 Synapiina Alonso-Zarazaga, 1990 170
Nemonychinae Bedel, 1882 90 Tanysphyrini Gistel, 1848 198
Neosharpiini A. Hoffmann, 1956 261 Tesserocerinitae Blandford, 1896 203
Ocladiinae Lacordaire, 1865 184 Tonesiina Alonso-Zarazaga & Lyal, 1999
Ocladiini Lacordaire, 1865 184 261
Ommatolampini Lacordaire, 1865 187 Tophoderini Lacordaire, 1865 98
Orthognathinae Lacordaire, 1865 185 Trachelizinae Lacordaire, 1865 147
Orthognathini Lacordaire, 1865 185 Trachelizini Lacordaire, 1865 148
Oxycoryninae Schoenherr, 1840 109 Trichapiina Alonso-Zarazaga, 1990 172
Oxystomatina Alonso-Zarazaga, 1990 165 Trigonorhinini Valentine, 1999 98
Oxystomatini Alonso-Zarazaga, 1990 164 Tropiderini Lacordaire, 1865 99
Ozotomerini Morimoto, 1972 94 Urodontinae C G . Thomson, 1859 107
Phloeophilini Lacordaire, 1865 95 Xenocerini Lacordaire, 1865 101
Phloeosinini Nüsslin, 1912 211 Xenorchestini Lacordaire, 1865 107
Phloeotribini Chapuis, 1869 212 Xyleborini LeConte, 1876 241
Piesocorynini Valentine, 1960 95 Xylinadini Lacordaire, 1865 101
Piezotrachelini Voss, 1959 172 Xyloctonini Eichhoff, 1878 250
Pityophthorina Eichhoff, 1878 215 Xyloterini LeConte, 1876 250
Platypodinae Shuckard, 1840 201 Zygaenodini Lacordaire, 1865 102
Platypodia Shuckard, 1840 201 Zygobaridina Pierce, 1907 251
Platypoditae Shuckard, 1840 201
Platyrhinini Imhoff, 1856 96
Platystomini Pierce, 1916 97
Podapiini Wanat, 2001 176
Polygraphini Chapuis, 1869 213
Polytini Zimmerman, 1993 187
Premnobiini Browne, 1962 236
Pseudoceocephalini Kleine, 1922 147
Pseudomesauletina Legalov, 2003 110
Ptychoderini Jekel, 1855 97
360
INDEX
Genus-group names
(synonyms are in italics)

Absoloniella Formânek, 1913 193 Alocentron Schilsky, 1901 153

Acaenorhinus Legalov, 2007 115 Alonsiellus Zherikhin, 1996 178
Acanephodus Alonso-Zarazaga, 1990 154 Alticopus A. Villa & G. B. Villa, 1833 106
Acanthothorax Gaede, 1832 94 Amacroderites Legalov, 2007 131
Acanthotomicus Blandford, 1894 233 Amasa Lea, 1894 241
Acanthurus Eichhoff, 1886 204 Ambrosiodmus Hopkins, 1915 241
Acentroides Leonhard, 1912 194 Ambrosiophilus Hulcr & Cognato, 2009 242
Acentrotypus Alonso-Zarazaga, 1990 154 Amorphocephala Damoiseau, 1966 143
Aceratapion Voss, 1969 164 Amorphocephalus Schoenherr, 1840 143
Acorynus Schoenherr, 1833 99 Amyctides Hustache, 1956 260
Acramorphocephaloides Bartolozzi, 198É> Anaeretus Dugès, 1887 246
144 Ananesiotes Kleine, 1922 142
Acrantus Broun, 1882 208 Anathymus Pascoe, 1885 189
Acythogaster Heller, 1940 252 Anaxyleborus Wood, 1980 241
Acythopeus Pascoe, 1874 260 Anchonocerus Eichhoff, 1878 215
Adapanetus Günther, 1936 189 Ancyloderes Blackman, 1938 215
Adeporaus Voss, 1941 118 Androceras Jordan, 1928 99
Aderorhinus Sharp, 1889 118 Aneorhachis Kleine, 1923 147
Adiaeretus Hagedorn, 1909 221 Anepsiotes Kleine, 1917 142
Aegyptobaris Pic, 1899 252 Angustapion Wanat, 1995 154
Afghanocryptus Voss, 1961 192 Anisandrus Ferrari, 1867 242
Afrothymapion Wanat, 1990 160 Anisonychidius Legalov, 2007 129
Aglycyderes Westwood, 1864 109 Anisonychus Voss, 1927 129
Agomadaranus Voss, 1958 140 Anius Pascoe, 1885 192
Agonotropis Egorov, 1987 99 Anodius Motschulsky, 1860 230
Agriorrhynchus Power, 1878 142 Anthinobaris Morimoto & Yoshihara, 1996
Aheterapoderus Legalov, 2007 131 252
Aizobius Alonso-Zarazaga, 1990 149 Anthotribus Gistel, 1856 97
Alaocephala Ganglbauer, 1906 198 Anthotribus J. J. Hoffmann, 1803 91
Alaocyba Perris, 1869 199 Anthribulus LeConte, 1876 99
Aletinus Desbrochers des Loges, 1908 110 Anthribus Fabricius, 1790 96
Alexsandricorynus Legalov, 2003 133 Anthribus Geoffroy, 1762 91
Algirolabus Legalov, 2003 134 Anthrodus Megerle, 1826 97
Allaeometrus Senna, 1903 144 Antribus llliger, 1801 91
Allaeotes Pascoe, 1885 192 Antriptus Illiger, 1801 91
Allandrus LeConte, 1876 90 Apachiscelus Desbrochers des Loges, 1875
Allapoderus Voss, 1927 136 193
Allarthrum Hagedorn, 1912 218 Apatenia Pascoe, 1859 96
Allernoporus Kurentsov, 1941 217 Aphanarthrum Wollaston, 1854 226
Alloder es Jordan, 1944 91 Aphanocleptus Wood, 1960 236
Alloeodromus Senna, 1895 145 Aphaulimia Morimoto, 1972 96
Allolabus Voss, 1925 135 Aphlorhynchites Sawada, 1993 122
Allomalia Alonso-Zarazaga, 1989 177 Aphoplopodapion Ehret, 1997 150
Alniphagus Swaine, 1918 206 Aphrysapoderus Legalov, 2007 134
 INDEX 361
Apiolum Kirby, 1808 148 Auletulus Voss, 1922 109
Apion Herbst, 1797 148 Aulodes Quedenfeldt, 1887 92
ApionusRafinesque, 1814 148 Aulodina Jordan, 1903 92
Apiops Alonso-Zarazaga, 1990 168 Autonopis Pascoe, 1874 186
Apium Agassiz, 1846 148 Autotropis Jordan, 1924 93
Apius Billberg, 1820 148 Bagoopsis Faust, 1881 193
Aplemonus Schoenherr, 1847 149 Baikovius Legalov, 2005 114
Aplopus Dejean, 1821 194 Baiocis Browne, 1962 201
Aplotes Chevrolat, 1885 189 Barathrodes Damoiseau, 1961 146
Apoderites Sawada, 1987 115 Barinomorphoides Morimoto, 1962 254
Apoderus Olivier, 1807 129 Barinomorphus Morimoto, 1962 254
Apolecta Pascoe, 1859 104 Baris Germar, 1817 254
Apoxyleborus Wood, 1980 249 Bariscelis Reitter, 1895 256
Apterorrhinus Senna, 1895 147 Barisoma Faust, 1888 257
Araecerus Schoenherr, 1823 104 Baryrhynchus Lacordaire, 1865 142
Araeocerus Schoenherr, 1839 104 Basarukinia Egorov, 1996 99
Aralkumia Bajtenov, 1974 193 Baseocolpus Jordan, 1949 96
Araptus Eichhoff, 1872 215 Baseotropis Gemminger & Harold, 1872 92
Archaeoscolytus Butovitsch, 1929 237 Basitropis Jekel, 1855 92
Archeophalus Schedl, 1941 221 Batobaris Morimoto & Yoshihara, 1996 256
Architomapoderus Legalov, 2007 140 Beaverium Hulcr & Cognato, 2009 242
Arganthonius Alonso-Zarazaga, 2005 260 Betulapion Ehret, 1994 172
Arixyleborus Hopkins, 1915 242 Bicolorhynchus Legalov, 2003 120
Arodepus Heller, 1922 115 Blaberus Schoenherr, 1839 103
Arodepus Voss, 1922 115 Blabirhinus Sharp, 1891 98
A rodum Gistel, 1856 107 Blysmia Pascoe, 1872 142
Arrhaecerus Germar, 1829 104 Bonips Cognato, 2001 233
Artepityophthorus Schedl, 1969 230 Boroxylon Hopkins, 1915 246
Arthrostenus Schoenherr, 1826 193 Bostrichips Schedl, 1952 236
Asaphepterum Kleine, 1916 144 Bothrus Wolfrum, 1956 95
Asemorhinus Sharp, 1891 98 Bothryorrhynchapion Bokor, 1923 165
Asiodeporaus Legalov, 2007 115 Botriessa Jordan, 1928 95
Aspidapion Schilsky, 1901 153 Botrobatys Chevrolat, 1842 185
Aspidapion Schilsky, 1901 153 Brachycerinus Bovie, 1909 182
Aspidobyctiscus Schilsky, 1903 112 Brachyceromorphus Bovie, 1909 182
Asuniops Legalov, 2007 B7 Brachycerus Olivier, 1789 182
Asynaptops Legalov, 2007 138 Brachydendrulus Schedl, 1951 215
Asynaptopsis Legalov, 2007 138 Brachygyius G. A. K. Marshall, 1939 198
Athesapeuta Faust, 1895 260 Brachyppa E. Strand, 1943 198
Athompsonirhinus Legalov, 2003 124 Brachypus Schoenherr, 1826 198
Atinellia Jordan, 1925 102 Brachytarsoides Pierce, 1930 99
Attelabus Linnaeus, 1758 134 Brachytarsus Schoenherr, 1823 91
Aulacobaris Desbrochers des Loges, 189^» Branconymus A. Hoffmann, 1959 97
252 Breviophthorus Schedl, 1938 216
Auletes Schoenherr, 1826 109 Brownia Nunberg, 1963 241
Auletobius Desbrochers des Loges, 1869 109 Bruchela Dejean, 1821 107
Auletomorphinus Legalov, 2007 120 Bulbifer Dejean, 1821 185
Auletomorphus Voss, 1923 120 Bulborhinapion Schatzmayr, 1926 153
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Coccygorrhynchites Prell, 1926 123 Cyaneugnamptus Legalov, 2003 118
Coelorhinapion Ehret, 1990 165 Cyanotrachelophorus Legalov, 2007 133
Coiffaitiella Osella, 1977 199 Cyclorhipidion Hagedorn, 1912 243
Colchis Tournier, 1874 193 Cycnotrachelodes Voss, 1955 130
Coleobothrus Enderlein, 1929 227 Cycnotrachelus Jekel, 1860 131
Colonnellinius Legalov, 2003 127 Cylas Latreille, 1802 142
Comesiella Del Guercio, 1925 212 Cylindra Illiger, 1802 202
Compsapoderopsis Legalov, 2003 130 Cylindroides Fairmaire, 1886 92
Compsapoderus Voss, 1927 130 Cyllorhynchites Voss, 1930 121
Conapium Motschulsky, 1866 172 Cyllorhynchites Voss, 1930 121
Confiisoscolytus Tsai & Hwang, 1962 237 Cynanchophagus Aksentev, 1987 230
Conophthocranulus Schedl, 1935 216 Cyphagogus Parry, 1849 145
Contexta Jordan, 1902 94 Cyphirhinus Schoenherr, 1826 254
Convexodeporaus Legalov, 2007 115 Cyphus Bede\, 1883 134
Coomania Kleine, 1941 145 Cypriapion Ehret, 1997 152
Coptodryas Hopkins, 1915 243 Cyrtogenius Strohmeyer, 1910 230
Coptodryas Schedl, 1949 237 Cyrtolabus Voss, 1925 135
Coptogaster Illiger, 1804 237 Cyrtotomicus Ferrari, 1867 233
Coptosomus Schedl, 1952 237 Cyrtotrachelus Schoenherr, 1838 187
Cordyle Thunberg, 1797 189 Dacryophthorus Schedl, 1971 210
Coriacephilus Schedl, 1939 217 Dacryphalus Hopkins, 1915 220
Corimalia Gozis, 1885 177 Dactylipalpus Chapuis, 1869 206
Corthylomimus Ferrari, 1867 215 Dactylopselaphus Gemminger & Harold,
Cosmobaris Casey, 1920 256 1872 206
Cosmocorynus Ferrari, 1867 215 Dactylotrypes Eggers, 1927 230
Cosmoderes Eichhoff, 1878 217 Dasycorynus Lacordaire, 1865 101
Cosmopolites Chevrolat, 1885 189 Degorsia Bedel, 1902 198
Cratoparis Dejean, 1834 92 Dendriops Schedl, 1953 217
Crossotarsinulus Schedl, 1972 201 Dendrobaris Egorov, 1976 260
Crossotarsus Chapuis, 1865 201 Dendroctonus Erichson, 1836 208
Cryphaloides Formânek, 1908 228 Dendropemon Schoenherr, 1839 93
Cryphalomorphus C. F. C. Schaufuss, 1891 Dendrotrogus Jekel, 1855 93
225 Dendrurgus Eggers, 1923 228
Cryphalophilus Schedl, 1970 225 Deporaus Samouelle, 1819 116
Cryphalops Reitter, 1889 220 Deporaus Samouelle, 1819 117
Cryphalus Erichson, 1836 218 Derisemias Jordan, 1928 98
Cryptarthrum Blandford, 1896 218 Deropria Enderlein, 1929 228
Cryptocleptes Blackman, 1920 236 Deropygus Sharp, 1891 105
Cryptoderma Ritsema, 1885 185 Derosasius Ganglbauer, 1906 199
Cryptulocleptus Wood, 1962 236 Desmidophorinus Hubenthai, 1917 185
Crypturgus Erichson, 1836 227 Desmidophorus Dejean, 1835 185
Ctenomeropsis Voss, 1939 179 Dexipeus Pascoe, 1885 192
Ctenyophthorus Schedl, 1955 216 Diacavus Schedl, 1939 204
Cucubaris Alonso-Zarazaga, 2005 260 Diamerides Browne, 1949 211
Cumatotomicus Ferrari, 1867 233 Diamerus Erichson, 1836 204
Cupreuscelophilus Legalov, 2005 139 Diapus Chapuis, 1865 203
Cyanapion Bokor, 1923 165 Diastrophus Perroud, 1865 144
Cyanapion Ehret, 1990 165 Dicranognathus L. Redtenbacher, 1844 HO
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Euparius Schoenherr, 1823 92 Golovninia Korotyaev & Egorov, 1996 160
Euprotapion Wagner, 1927 172 Gomadaranus Kôno, 1930 141
Eupsalomimus Kleine, 1916 142 Gonotropis LeConte, 1876 99
Eupteroxylon Eggers, 1936 215 Granips Cognato, 2001 233
Euptilius Schedl, 1940 220 Granulapoderus Legalov, 2003 129
Eurostauletes Voss, 1933 111 Graphicotera Tournier, 1873 193
Eurydactylus Hagedorn, 1909 244 Gretschkinia Sokanovskiy, 1959 236
Eurymycter LeConte, 1876 99 Grypidius Schoenherr, 1826 194
Euscelophilidius Legalov, 2003 139 Grypus Germar, 1817 194
Euscelophiloides Legalov, 2007 139 Gynandrocerus Lacordaire, 1865 92
Euscelophilus Voss, 1925 139 Habrissus Pascoe, 1859 92
Eusebus Kleine, 1922 146 Habrocerastes Lacordaire, 1865 92
Eusphyrus LeConte, 1876 102 Hadrodemius Wood, 1980 245
Eusproda Sawada, 1987 117 Hadrophyes Reitter, 1911 178
Eutrichapion Reitter, 1916 165 Hadrophyes Reitter, 1916 178
Eutrichapion Reitter, 1916 166 Hagedornus R. Lucas, 1920 216
Euvolvulus Reitter, 1916 122 Halticopus Agassiz, 1846 106
Euwallacea Hopkins, 1915 244 Hamiltonius Alonso-Zarazaga & Lyal, 1999
Exapion Bedel, 1887 158 131
Exechesops Schoenherr, 1847 102 Harpapion Voss, 1966 154
Exillis Pascoe, 1860 102 Helianthemapion Wagner, 1930 150
Exrhynchites Voss, 1930 118 Helleriapoderus Legalov, 2007 134
Exurodon Schilsky, 1912 107 Hemiorychodes Kleine, 1921 143
Faustia Tournier, 1889 196 Hemiperapion Wagner, 1930 150
Ferreria Alonso-Zarazaga & Lyal, 1999 199 Hemitrichapion Voss, 1959 166
Ficicis Lea, 1910 206 Hemitrichapion Voss, 1959 167
Ficiphagus Murayama, 1958 206 Henicolaboides Legalov, 2007 135
Flavodeporaus Legalov, 2007 115 Henicolabus Voss, 1925 135
Flavopodapion Korotyaev, 1987 153 Heterapoderopsis Legalov, 2003 131
Formosobaris Morimoto & Yoshihara, 1996 Heterapoderus Voss, 1927 131
257 Heterauletes Voss, 1933 110
Formosusorynus Legalov, 2003 133 Heteroborips Reitter, 1913 246
Fornicaria Gistel, 1848 147 Heterorhynchites Voss, 1932 124
Fremuthiella Alonso-Zarazaga, 1990 172 Hidryocneme Alonso-Zarazaga, 1990 159
Fujarhynchites Legalov, 2007 122 Higonius Lewis, 1883 147
Fursovia Alonso-Zarazaga & Lyal, 1999 186 Higonodes Zimmerman, 1994 147
Genyocerus Motschulsky, 1858 204 Hilipomorphus Desbrochers des Loges, 1891
Gibber Jordan, 1895 103 190
Giustiapion Legalov, 2003 171 Himalayrhynchites Legalov, 2003 121
Glabrinotropis Frieser, 1981 107 Himasthlophallus Egorov & Zherikhin, 1991
Glaridorhinus Kolenati, 1858 198 198
Glyptoderes Eichhoff, 1878 226 Hollisiella Alonso-Zarazaga & Lyal, 1999
Gnatharus Wood & Yin, 1986 215 257
Gnathoborus Schedl, 1970 216 Holonthogaster Gemminger & Harold, 1872
Gnathocranus Schedl, 1951 216 211
Gnathophorus Schedl, 1935 216 Holotrichapion Ehret, 1990 168
Gnathophthorus Wood, 1962 216 Holotrichapion Györffy, 1956 168
Gnathotrichus Eichhoff, 1869 215 Homalorhynchites Voss, 1938 124
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 INDEX 367
Lissorhoptrus LeConte, 1876 198 Mesoderes Senna, 1898 145
Lissotarsus Faust, 1881 257 Mesoscolytus Broun, 1904 246
Listrobyctiscidius Legalov, 2003 114 Mesotrichapion Ehret, 1990 169
Litocerus Schoenherr, 1833 100 Mesotrichapion Györffy, 1956 169
Litorhynchus Schoenherr, 1845 188 Metacidotes Schedl, 1961 145
Loborhynchapion Györffy, 1956 168 Metahylastes Eggers, 1922 230
Loborhynchapion Osella, 1973 168 Metallarodepus Legalov, 2003 115
Loborhynchapion Smreczynski, 1965 168 Metallorhynchites Legalov, 2007 125
Loganius Chapuis, 1869 237 Metapion Schilsky, 1906 163
Longulus Krivolutskaya, 1968 208 Metaprodioctes Günther, 1937 189
Lostianus Desbrochers des Loges, 1900 193 Metarhynchites Voss, 1923 125
Lotapion Alonso-Zarazaga, 1990 167 Metinvolvulus Legalov, 2003 120
Lymantor Lovendal, 1889 231 Metocalolabus Legalov, 2003 139
Macrocephalus Olivier, 1789 97 Metopon G.R. Waterhouse, 1842 109
Macrocheirus Schoenherr, 1838 188 Metopum Agassiz, 1846 109
Macrochirus Agassiz, 1846 188 Micracidendron Schedl, 1957 216
Macrochirus Gemminger, 1871 188 Micrapoderus Legalov, 2003 137
Macrocryphalus Nobuchi, 1981 221 Microconapion Korotyaev, 1992 173
Macrodentipes Liang & Li, 2005 138 Microcorynus Legalov, 2003 131
Macroderites Legalov, 2003 131 Microctilodes Reitter, 1875 108
Macrosynaptopsis Legalov, 2005 137 Microon Alonso-Zarazaga, 1989 179
Maculapoderus Legalov, 2003 132 Microperus Wood, 1980 245
Maculinvoles Legalov, 2003 124 Microsebus Kolbe, 1892 146
Maculinvolvulus Legalov, 2003 121 Microtinocyba Legalov, 2001 167
Maculphrysus Legalov, 2003 136 Microtrachelizus Senna, 1893 147
Malvaevora Zaslavskij, 1956 258 Mimidendrulus Schedl, 1957 230
Malvapion A. Hoffmann, 1958 162 Mimiocurus Schedl, 1957 216
Malvapion Voss, 1959 162 Mimiophthorus Schedl, 1957 216
Manilabaris Zimmerman, 1940 261 Mimips Eggers, 1932 233
Margaritapion Korotyaev, 1990 150 Miniapion Korotyaev, 1992 154
Mauia Blackburn, 1885 94 Minulus Eggers, 1912 » 237
Maurobaris Alonso-Zarazaga, 2005 258 Miolispa Pascoe, 1862 148
Mecocems Schoenherr, 1833 94 Mirabinvolvulus Legalov, 2003 124
Mecorhis Billberg, 1820 124 Mirabrhynchites Legalov, 2003 124
Mecotropis Lacordaire, 1865 97 Misthosima Pascoe, 1859 105
Meganthribus Jordan, 1913 98 Monarthrum Kirsch, 1866 215
Megaplatypus Wood, 1993 202 Monius Schoenherr, 1845 198
Megatermis Jordan, 1937 105 Montaninvolvulus Legalov, 2003 125
Melaleucus Chevrolat, 1882 258 Mordkovitshirhinus Legalov, 2003 133
Melanapion Wagner, 1930 160 Moreobaris Morimoto & Yoshihara, \99t i
Melanobaris Alonso-Zarazaga & Lyal, 1999 258
258 Morimotanthribus Senoh & Tryzna, 2006 95
Melanopsacus Jordan, 1924 106 Morphocorynus Legalov, 2003 137
Melitobaris Peyerimhoff, 1929 258 Morphoeuops Legalov, 2003 138
Mepiezotrachelus Voss, 1961 173 Morphotrachelophorus Legalov, 2003 133
Merarius Fairmaire, 1889 100 Mucronianus Jordan, 1894 100
Merothricus Chevrolat, 1885 191 Mycteis Pascoe, 1860 94
Mesauletobius Legalov, 2001 111 Myeloborus Blackman, 1928 216
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 INDEX 369
Osellaeus Alonso-Zarazaga, 1990 150 Paropes Schoenherr, 1823 91
Otidognathus Lacordaire, 1865 188 Paroplapoderus Voss, 1926 140
Ovauletes Legalov, 2007 111 Paroplapoderus Voss, 1926 141
Oxeostomum Gistel, 1856 148 Paropus Fischer von Waldheim, 1829 91
Oxyderes Jordan, 1928 96 Parorychodes Kleine, 1921 143
Oxyrhynchus Schoenherr, 1823 185 Parvodeporaus Legalov, 2007 117
Oxyrinus Rafinesque, 1815 169 Pelicerus Eggers, 1923 230
Oxystoma Duméril, 1805 169 Pellobaris Morimoto & Yoshihara, 1996 260
Oxystomum Gistel, 1856 148 Pembertonia Zimmerman, 1944 192
Ozodendron Schedl, 1957 230 Penestica Pascoe, 1859 92
Ozophagus Eggers, 1920 214 Perapion Wagner, 1907 150
Ozotomerus Perroud, 1853 94 Perapion Wagner, 1907 151
Pachygenia Motschulsky, 1875 99 Peraprophthalmus Kleine, 1923 148
Pachynoderes Schedl, 1941 221 Peribathys Jordan, 1937 101
Paleodeporaoides Legalov, 2007 117 Pericartiellus Alonso-Zarazaga, 1989 181
Paleodeporaus Legalov, 2003 117 Perisymmorphocerus Kleine, 1919 144
Panscopus Schoenherr, 1843 198 Peroplatypus Wood, 1993 202
Parabyctiscus Legalov, 2003 112 Perrhynchites Voss, 1953 126
Paracalendra Chûjô & Morimoto, 1959 186 Perrhynchitoides Legalov, 2007 126
Paracentrocorynus Voss, 1929 134 Pertorcus Voss, 1953 252
Paracompsus Legalov, 2003 130 Phaenotheriolum Ganglbauer, 1903 95
Paracycnotrachelus Voss, 1924 133 Phaenotherion Frivaldszky, 1877 95
Paracythopeus Voss, 1958 260 Phaenotheriosoma Frieser, 1978 95
Paradeporaus Kôno, 1927 118 Phaenotherium Ganglbauer, 1903 95
Paradiaphorus Chevrolat, 1885 189 Phaeochrotes Pascoe, 1860 91
Parahamiltonius Legalov, 2007 131 Phalacrolobus Alonso-Zarazaga, 1990 166
Paraleptapoderus Legalov, 2003 132 Pharcidobaris Morimoto & Yoshihara, 1996
Parallelodemas Faust, 1895 260 259
Paramecolaboides Legalov, 2007 136 Phaulimia Pascoe, 1859 96
Paramecolabus Jekel, 1860 136 Phellodendrophagus Krivolutskaya, 195 S
Paramesus Fâhraeus, 1871 98 219
Paramorphocephalus Kleine, 1920 144 Phialodes Roelofs, 1874 136
Paraprodioctes Voss, 1958 190 Philorectus Voss, 1924 124
Pararaymondionymus Osella, 1977 200 Phloeobius Schoenherr, 1823 97
Parasphaerotrypes Murayama, 1958 210 Phloeochilus Schedl, 1953 210
Parasuniops Legalov, 2003 137 Phloeocranus Schedl, 1942 211
Parasynapion Legalov, 2003 172 Phloeopemon Schoenherr, 1839 98
Parasynaptopsis Legalov, 2003 137 Phloeophilus Schoenherr, 1833 95
Parasynatops Legalov, 2003 138 Phloeophthorus Wollaston, 1854 212
Paratemnocerus Legalov, 2003 128 Phloeosinopsioides Schedl, 1964 211
Parathesapeuta Voss, 1964 260 Phloeosinopsis Schedl, 1936 211
Paratrachelophorus Voss, 1924 133 Phloeosinopsis Schedl, 1964 211
Parauletes Sawada, 1993 109 Phloeosinus Chapuis, 1869 211
Paraxyleborus A. Hoffmann, 1942 215 Phloeotribus Latreille, 1797 212
Parinvolvoides Legalov, 2003 122 Phloeotrogus Motschulsky, 1863 241
Parinvolvulus Legalov, 2003 122 Phloeotrypetus Wood, 1960 210
Parmaculphrysus Legalov, 2003 136 Phrissoderes G. A. K. Marshall, 1948 259
Paronychapion Voss, 1961 163 Phrissotrichum Schilsky, 1901 152
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Pseudomesauletes Legalov, 2001 111 Rhaphitropis Reitter, 1916 103
Pseudometapion Wagner, 1930 163 Rhinanthribus Motschulsky, 1875 98
Pseudometopum Legalov, 2003 110 RhinchitobinsKôno, 1928 114
Pseudoparaphorus Legalov, 2003 133 RhinomacerO. F. Müller, 1764 134
Pseudoperapion Wagner, 1930 152 Rhinopteryx Lacordaire, 1865 148
Pseudophrysus Legalov, 2003 137 Rhodapion Alonso-Zarazaga, 1990 160
Pseudopiezotrachelus Wagner, 1907 175 Rhodocyrtus Alonso-Zarazaga & Lyal, 1999
Pseudopityophthorus Swaine, 1918 217 126
Pseudopoecilips Murayama, 1957 232 Rhopalapion Schilsky, 1906 163
Pseudopolygraphus Seitner, 1911 214 Rbynchitallus Voss, 1960 126
Pseudoprotapion Ehret, 1990 175 Rhynchites D. H. Schneider, 1791 127
Pseudorhinoides Legalov, 2007 124 Rhynchophorus Herbst, 1795 189
Pseudorhinus Legalov, 2003 124 Riedeliops Alonso-Zarazaga & Lyal, 2002
Pseudorhyssematus Morimoto, 1962 259 138
Pseudorobitis L. Redtenbacher, 1868 182 Roelofsia Ritsema, 1891 187
Pseudorychodes Senna, 1894 143 Roelofsideporaus Legalov, 2003 117
Pseudosebasius Kleine, 1944 144 Rostauletes Legalov, 2007 110
Pseudostenapion Wagner, 1930 153 Rubrauletes Legalov, 2003 111
Pseudotaphroderes Bolkay, 1911 146 Rubrrhynchites Legalov, 2007 128
Pseudothamnurgus Eggers, 1912 232 Ruffodytes Osella, 1973 196
Pseudothysanoes Blackman, 1920 236 Ruguloscolytus Butovitsch, 1929 237
Pseudotrichapion Bokor, 1923 166 Saliciphilus Sokanovskiy, 1954 232
Pseudotrichapion Ehret, 1990 166 Sawadaeuops Legalov, 2003 138
Pseudotrichosomus Hustache, 1925 185 Sawadaeuopsis Legalov, 2005 139
Pseudoxyleborus Eggers, 1930 241 Sawadaia Alonso-Zarazaga & Lyal, 1999 124
Pseudoxylechinus Wood & Huang, 1986 209 Scapapion Korotyaev, 1992 176
Pseudplapoderus Legalov, 2003 141 Schilskyapion Alonso-Zarazaga, 1990 152
Pseudvulus Legalov, 2003 121 Schizotrachelus Lacordaire, 1865 148
Pseudypera Voss, 1936 195 Schoenitemnus Legalov, 2003 126
Psilarthroides Morimoto & Miyakawa, 1985 Schoenius Alonso-Zarazaga, 1990 168
259 Scolytochelus Reitter, 1913 237
Psilocalymma Alonso-Zarazaga, 1990 166 Scolytocleptes Schedl, 1962 250
Psix Alonso-Zarazaga, 1989 182 Scolytogenes Eichhoff, 1878 225
Pteleobius Bedel, 1888 208 Scolytomimus Blandford, 1895 250
Pteridobaris Morimoto & Yoshihara, 1996 Scolytoplatypus C. F. C. Schaufuss, 1891 240
259 Scolytus Geoffroy, 1762 237
Pterocyclon Eichhoff, 1869 215 Scyphophorus Schoenherr, 1838 190
Pustulorhinus Legalov, 2003 126 Sergiola Korotyaev, 1996 160
Pygmaeoscolytus Butovitsch, 1929 237 Shiva Pajni & Bhateja, 1982 182
Pyrorhynchites Legalov, 2003 127 Sintor Schoenherr, 1839 98
Rawasia Roelofs, 1880 98 Sipalinus G. A. K. Marshall, 1943 185
Raymondia Aube, 1861 200 Sipalomimus Voss, 1958 190
Raymondiellus Ganglbauer, 1906 199 Sipalus Schoenherr, 1825 186
Raymondionymus Wollaston, 1873 200 Siraton Hustache, 1934 197
Rhabdocnemis Faust, 1893 190 Sitonobia Gistel, 1856 191
Rhabdoscelus G. A. K. Marshall, 1943 190 Sitophilus Schoenherr, 1838 186
Rhaphidoplectron Alonso-Zarazaga, 199C1 Sparganophasma Kleine, 1926 145
152 Spathorrhamphus T. A. Marshall, 1902 100
372 INDEX

Spermafoplex Hopkins, 1915 228 259

Sphaerotrypes Blandford, 1894 204 Taiwanobyctiscus Kôno, 1929 112
Sphaerula Kiesenwetter, 1864 180 Taiwanorhynchites Legalov, 2007 128
Sphaerula Stephens, 1829 180 Tanysphyroides Egorov, 1996 198
Sphenoceros Schedl, 1939 216 Tanysphyrus Germar, 1817 198
Sphenocorynes Schoenherr, 1838 190 Taphroborus Nunberg, 1961 230
Sphenophorus Schoenherr, 1838 191 Taphrocoetes Pfeffer, 1987 232
Sphinctotropis Kolbe, 1895 100 Taphrocomister Senna, 1895 147
Spilobahs Morimoto & Yoshihara, 19966 260 Taphronurgus Reitter, 1913 232
Spinuloscolytus Butovitsch, 1929 237 Taphrorychus Eichhoff, 1878 232
Spongocerus Blandford, 1893 240 Taphroterus Schedl, 1965 232
Spongotarsus Hagedorn, 1908 214 Taphrotopium Reitter, 1916 157
Squamapion Bokor, 1923 161 Tarattostichus Ganglbauer, 1906 200
Squamapion Wagner, 1929 161 Tatianaerhynchites Legalov, 2002 128
Squamosinus Nunberg, 1964 209 Tatyanapion Legalov, 1997 170
Stenommatus Wollaston, 1873 185 Telphes Wolfrum, 1924 94
Stenopelmus Schoenherr, 1835 198 Temnalysis Alonso-Zarazaga, 1989 182
Stenopiatypus Strohmeyer, 1914 202 Temnocerus Thunberg, 1815 128
Stenopterapion Bokor, 1923 171 Teretriorhynchites Voss, 1938 122
Stenorhis Jordan, 1928 107 Terminalinus Hopkins, 1915 243
Stenorhynchites Voss, 1932 124 Terminassianaeus Legalov, 2003 127
Stephanoderes Eichhoff, 1872 221 Tetraspartus Pascoe, 1885 185
Stephanorhopalus Hopkins, 1915 220 Tetrasynommatus Morimoto, 1985 192
Stiboderes Jordan, 1925 101 Tetratemnus Wollaston, 1873 185
Straboscopus Lacordaire, 1865 96 Tetratopos Chevrolat, 1883 191
Stratiopisthius Senna & Calabresi, 1919» 146 Tetratopus Csiki, 1936 191
Streneoderma Blackburn, 1900 104 Teuchocnemapion Voss, 1966 161
Strigapoderus Jekel, 1860
134 Thamnophthorus Blackman, 1942 216
Strophionocerus Sampson, 1921
240 Thamnurgides Hopkins, 1915 228
Stylotentus Schedl, 1963
221 Thamnurgus Eichhoff, 1864 232
Subdysmorphorhynchus Kleine, 1917
144 Theanellus Reitter, 1913 197
Suborychodes Kleine, 1917
143 Thecorhinus Faust, 1887 193
Sueus Murayama, 1951
210 TheryusPic, 1897 259
Suniopsidius Legalov, 2005
139 Thompsonirhinus Legalov, 2003 124
Svetlanaebyctiscus Legalov, 2001
Symmorphocerus Schoenherr, 1847 114 Thrasycephalus Kleine, 1916 145
Sympaector Kirsch, 1875 144 Thryogenes Bedel, 1884 197
Sympiezotrachelus Voss, 1959 94 Thymapion Sainte Claire-Deville, 1924 161
Synapion Schilsky, 1902 173 Tinocyba Alonso-Zarazaga, 1990 167
Synapion Schilsky, 1902 171 Titanomalia Alonso-Zarazaga, 1989 178
Synommatoides Morimoto, 1978 172 Tomapoderopsis Legalov, 2003 141
Synommatus Wollaston, 1873 192 Tomapoderus Voss, 1926 141
Synorychodes Kleine, 1921 192 Tomicus Latreille, 1802 209
Syrdariella Ter-Minasian, 1977 143 Tosaxyleborus Murayama, 1950 242
Taeniapion Schilsky, 1906 197 Tournotaris Alonso-Zarazaga & Lyal, 1999
Taeniocerus Blandford, 1893 162 197
Taenioglyptes Bedel, 1888 240 Trachelizus Dejean, 1834 148
Taiwanobaris Morimoto & Yoshihara, 1 218 Trachelolabus Jekel, I860 140
996 Treptoplatypus Schedl, 1972 203
 INDEX 373
Triarmocerus Eichhoff, 1878 221 Vietsuniops Legalov, 2007 139
Trichischius LeConte, 1876 191 Wagnerium Alonso-Zarazaga, 1990 175
Trichocaulus Fairmaire, 1877 193 Webbia Hopkins, 1915 245
Trichoconapion Korotyaev, 1985 176 Xanthoderopygus Senoh, 1984 105
Trichopterapion Wagner, 1930 160 Xelyborus Schedl, 1939 245
Trichosomus Chevrolat, 1881 185 Xenocerus Schoenherr, 1833 101
Trigonogenius Hagedorn, 1912 216 Xenophthorus Wood & Yin, 1986 217
Trigonorhinus Wollaston, 1861 99 Xenorchestes Wollaston, 1854 107
Trigonorrhinus Gemminger & Harold, 1872 Xenotropis Fairmaire, 1895 93
99 Xerodermus Lacordaire, 1865 192
Triotemnus Wollaston, 1864 233 Xeronoma Iablokoff-Khnzorian, 1964 197
Trischidias Hopkins, 1915 226 Xestips Hagedorn, 1912 233
Trochorhopalus Kirsch, 1877 192 Xyleboricus Eggers, 1923 242
Trochorrhopalus Champion, 1914 192 Xyleborinus Reitter, 1913 245
Trochorrhopalus Rye, 1879 192 Xyleborips Reitter, 1913 215
Tropideres Schoenherr, 1823 101 Xyleborus Eichhoff, 1864 246
Tropiderinus Reitter, 1916 90 Xylechinops Browne, 1973 209
Tropidoderes Gemminger & Harold, 1872 Xylechinus Chapuis, 1869 209
101 Xylinada Berthold, 1827 101
Tropidoderes Gistel, 1856 101 Xylinades Latreille, 1828 101
Trymatoderus Fairmaire, 1889 192 Xylocleptes Ferrari, 1867 233
Trypanophellos Bright, 1982 210 Xylocryptus Schedl, 1975 225
Tryphetus Faust, 1895 187 Xylosandrus Reitter, 1913 249
Trypocranus Eichhoff, 1878 215 Xyloterus Erichson, 1836 250
Trypodendron Stephens, 1830 250 Xylotrophus Gistel, 1848 250
Trypodendrum Agassiz, 1846 250 Ypselogonia Kleine, 1922 142
Trypodendrum Gistel, 1856 250 Yunnanuletes Legalov, 2007 112
Trypographus Schedl, 1950 207 Zherikhinia Alonso-Zarazaga, 1999 182
Trypophloeus Fairmaire, 1864 226 Zygaenodes Pascoe, 1859 102
Tsherepanovia Korotyaev, 1992 198
Tshernyshevinius Legalov, 2003 127
Tubicenus Latreille, 1829 109
Tubuloscolytus Butovitsch, 1929 237
Turanocryptus Korotyaev, 1988 193
Turkmenobaris Zaslavskij, 1956 259
Tychiophaenus Voss, 1960 197
Tychiosoma Voss, 1943 197
Tyrodomus Voss, 1971 197
UbychiaRost, 1893 200
Ulapion Ehret, 1997 158
Ulobaris Reitter, 1895 259
Ulorhinus Sharp, 1891 97
Uncifer Jordan, 1904 104
Unciferina Morimoto, 1981 104
Urodon Schoenherr, 1823 107
Valenfriesia Alonso-Zarazaga & Lyal, 1999
105
Vietnamorhynchites Legalov, 2003 124
 Catalogue of Palaearctic Coleoptera
Structured knowledge of life, as it evolved and is present on our planet, requires
cataloguing. Catalogues summarize the work in systematics, they enable assessment
of faunal diversity and classification, and provide information on past taxonomic
acts. They are also essential in achieving a universal language in taxonomy that
assures unambiguous transfer of biological information. Erroneous information due
to incorrect use of taxonomic names has an impact in many fields, including
genetics, physiology, ecology, plant protection, nature conservation, and legislation.

Over the years, the Palaearctic Coleoptera have undoubtedly been studied by more
students that those of other parts of the world, and more than representatives of
most other orders of insects. It is therefore remarkable that an overview of the
present knowledge of the about 1 0 0 , 0 0 0 species of the Palaearctic Coleoptera is
badly lagging that of, e.g., the Nearctic Coleoptera, or Palaearctic Diptera.

The aim of Catalogue of Palaearctic Coleoptera is to provide a tool that addresses

the most urgent needs: 1) a list of available names, both valid and invalid, of taxa
occurring in the Palaearctic Region, in their verified original orthography and with
the correct publication dates, 2) a complete list of verified references to primary
descriptions, and 3) informative distributions of the species and subspecies.
Thus, the catalogue is expected to respond to questions related to biodiversity,
and to increase the badly needed nomenclatorial stability.

Catalogue of Palaearctic Coleoptera is a collective work involving many authors

and will be published in eight volumes. Each volume will group taxa with respect
to the adopted classification, and will be "independent", having its own
reference section and Index.

Previously published:
Volume 1: Archostemata, Myxophaga, Adephaga. 2 0 0 3 .
Volume 2: Hydrophiloidea, Histeroidea, Staphylinoidea. 2 0 0 4 .
Volume 3 : Scarabaeoidea, Scirtoidea, Daseilloidea,
Buprestoidea, Byrrhoidea. 2 0 0 6 .
Volume 4 : Elateroidea, Derodontoidea, Bostrichoidea
Lymexyloidea, Cleroidea, Cucujoidea. 2 0 0 7 .
Volume 5: Tenebrionoidea. 2 0 0 8 .
Volume 6: Chrysomeloidea. 2 0 1 0 .

ISBN 978-87-88757-93-4 (Vol. 7)

'757934 ISBN 978-87-88757-71-2 (Vols 1-8)