World Plants: Plant List

species :

Lepisorus abbreviatus (Fée) Li Wang; Bot. J. Linn. Soc. 162(1): 35 (2010)

Distribution: Vietnam

41 VIE

Remarks: sect. 13 Belvisia

Synonyms: hidden

= Belvisia abbreviata (Fée) Hovenkamp & Franken; Blumea 37(2): 519 (1993)

= Belvisia hymenolepioides (Christ) Ching; Acta Phytotax. Sin. 10: 301 (1965)

= Belvisia hymenolepioides (Christ) V.N.Tu; Bot. Zhurn. (Moscow & Leningrad) 64(11): 1671 (1979), isonym

= Drymoglossum abbreviatum Fée; Mém. Foug. 7 (Icon. Esp. Nouv.): 26, t. 10, f. 2 (1857)

= Hymenolepis abbreviata Fée; Mém. Foug. 10 (Icon. Esp. Nouv.): 12 (1865)

= Hymenolepis hymenolepioides (Christ) Ching ex C.Chr. & Tardieu; Notul. Syst. (Paris) 8: 185 (1939)

= Lemmaphyllum abbreviatum Kuhn & C.Chr.; Dansk Bot. Ark. 6(3): 50 (1929)

= Macroplethus abbreviatus (Fée) Tagawa; Acta Phytotax. Geobot. 11: 234 (1942)

= Polypodium abbreviatum (Fée) C.Chr.; Index Filic., Suppl. Prelim.: 25 (1917)

= Polypodium hymenolepioides Christ; J. Bot. (Morot) 19: 75 (1905)

species :

Lepisorus accedens (Blume) Hosok.; Trans. Nat. Hist. Soc. Formosa 31: 477 (1941)

Distribution: peninsular Thailand; peninsular Malaysia (Terengganu, Perak, Pahang, Selangor, Negeri Sembilan, Johor); Sumatra; Borneo (Mt. Kinabalu, Sabah); Sulawesi; Philippines; Java; Lesser Sunda Isl. (Lombok, Bali); Moluccas (Seram, etc.); New Guinea (Irian Jaya, Papua New Guinea); Bismarck Arch. (Admiralty Isl., Admiralty Isl., New Ireland, New Britain); Solomon Isl. (Bougainville, Choiseul, Guadalcanal, Kolombangara, Makira, Santa Ysabel); Australia (NE-Queensland); Vanuatu; Fiji (Viti Levu, Vanua Levu, Taveuni); American Samoa (Tutuila, Ofu, Olosega, Ta´u); [Western] Samoa (Savai´i, ´Upolu); Fed. States of Micronesia (Pohnpei, Kosrae)

41 THA 42 BOR JAW LSI MLY MOL NWG PHI SUL SUM 43 BIS SOL 50 AUS 60 ASA FIJ SAM VAN 62 MIC

Remarks: sect. 11 Weatherbya

Synonyms: hidden

= Craspedaria cuspidiflora (Reinw.) Fée; Mém. Foug. 5 (Gen. Filic.): 264 (1852)

= Craspedaria javanensis Fée; Mém. Foug. 7 (Icon. Esp. Nouv.): 65 (1857)

= Drynaria acuminata Brack.; U.S. Expl. Exped., Filic. 16: 47 (1854) (non p. 41)

= Drynaria cuspidiflora (Reinw.) J.Sm.; J. Bot. (Hooker) 3: 397 (1841)

= Goniophlebium javanense (Fée) T.Moore; Index Filic. (T. Moore) 390 (1862)

= Lemmaphyllum accedens (Blume) Donk; Reinwardtia 2: 409 (1954)

= Lepidogrammitis accedens (Blume) Saiki; J. Phytogeogr. Taxon. 35(2): 80 (1987)

= Lepidogrammitis damunensis (Rosenst.) Saiki; J. Phytogeogr. Taxon. 35(2): 80 (1987) (without basionym page)

= Microsorum accedens (Blume) Copel.; Univ. Calif. Publ. Bot. 16: 111-116 (1929)

= Phymatopsis accedens (Blume) J.Sm.; Hist. Fil. 104 (1875)

= Pleopeltis accedens (Blume) T.Moore; Index Filic. (T. Moore) 77 (1857)

= Pleopeltis acuminata (Brack.) T.Moore; Index Filic. (T. Moore) 344 (1862)

= Pleopeltis damunensis Alderw.; Bull. Dép. Agric. Indes Néerl. 27: 4 (1909)

= Pleuridium cuspidiflorum (Reinw.) J.Sm.; Bonplandia 9: 261 (1861)

= Polypodium accedens Blume; Enum. Pl. Javae 2: 121 (1828)

= Polypodium cuspidiflorum Reinw.

= Polypodium damunense Rosenst.; Repert. Spec. Nov. Regni Veg. 6: 42 (1907)

= Polypodium lyciifolium Bory; Dup., Voy. Bot. 1: 260 (1828)

= Polypodium redimiens Brause; Bot. Jahrb. Syst. 56(2): 193 (1920)

= Weatherbya accedens (Blume) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.] 191, t. 6 (1947)

= Weatherbya damunensis (Rosenst.) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.] 191 (1947)

species :

Lepisorus affinis Ching; Ching & Chu H. Wang, Acta Phytotax. Sin. 8: 169, t. 22(28) (1959)

Distribution: China (Hainan)

36 CHI

Remarks: sect. 10 Lepisorus

Synonyms: hidden

= Lepisorus longifolius Ching; Ching & Chu H. Wang, Acta Phytotax. Sin. 8: 170 (1959) (non (Blume) Holttum)

= Lepisorus longus Ching; Acta Phytotax. Sin. 10: 192 (1965), nom. nov. for L. longifolius Ching

species :

Lepisorus alberti (Regel) Ching; Acta Bot. Yunnan. 5(1): 20 (1983)

Distribution: China (Gansu, Hebei, Henan, Nei Mongol, Qinghai, Shaanxi, Shanxi, Sichuan, Xinjiang, Yunnan); Tibet

36 CHI TIB

Remarks: sect. 9 Hymenophyton. Records for Taiwan refer to L. clathratus fide Knapp (Ferns of Taiwan, 2011 ff.)

Synonyms: hidden

= Lepisorus clathratus var. papakensis (Masam.) Tagawa; Acta Phytotax. Geobot. 5(2): 111 (1936)

= Lepisorus papakensis (Masam.) Ching & Y.X.Lin; Acta Bot. Yunnan. 5(1): 7 (1983)

= Lepisorus pumilus Ching & S.K.Wu; Acta Bot. Yunnan. 5(1): 16 (1983)

= Polypodium alberti Regel; Trudy Imp. S.-Peterburgsk. Bot. Sada 7. 622 (1881)

= Polypodium lineare var. alberti (Regel) C.Chr.; Index Fil. 507, 540 (1906)

= Polypodium papakense Masam.; J. Soc. Trop. Agric. Formosa 6: 569 (1934)

species :

Lepisorus amaurolepidus (Sledge) Bir & Trikha; Bir & Vasudeva, J. Bombay Nat. Hist. Soc. 68: 192 (1971)

Distribution: S-India; Sri Lanka

40 IND SRL

Remarks: sect. 17 Pseudovittaria. Included in L. contortus by Kandel & Fraser-Jenkins (2020)

Synonyms: hidden

= Lepisorus amaurolepidus (Sledge) B.K.Nayar & S.Kaur; Companion Handb. Ferns Brit. India 84 (1974), isonym

= Pleopeltis amaurolepida Sledge; Bull. Brit. Mus. (Nat. Hist.) Bot. 2: 136 (1960)

species :

Lepisorus angustus Ching; Bull. Fan Mem. Inst. Biol. Bot. 4: 86 (1933)

Distribution: China (Anhui, Chongqing, Gansu, Guangxi, Henan, Hubei, Hunan, Shaanxi, Sichuan, Yunnan, S-Zhejiang); Tibet; ?Japan

36 CHI TIB

Remarks: sect. 17 Pseudovittaria. Included in L. thunbergianus by Fraser-Jenkins et al. 2015 but kept separate on genetic basis by Zhao et al. 2019

Synonyms: hidden

= Lepisorus nanchuanensis Ching; Bull. Bot. Res., Harbin 3(4): 4 (1983)

= Lepisorus niger Ching; Fl. Xizangica 1: 312 (1983) (fide Fraser-Jenkins et al. 2015)

= Lepisorus pseudoangustus Ching; Fl. Xizangica 1: 310 (1983)

= Lepisorus thunbergianus var. angustus (Ching) Sa.Kurata; Sci. Rep. Yokosuka City Mus. 11: 39 (1965)

= Pleopeltis caudatoattenuata (Takeda) Panigrahi & Patnaik; Curr. Sci. 34: 127 (1965)

= Polypodium caudatoattenuatum (Takeda) C.Chr.; Index Filic., Suppl. Tert.: 146 (1934)

= Polypodium lineare f. caudatoattenuatum Takeda; Notes Roy. Bot. Gard. Edinburgh 8(39): 269-270 (1915)

species :

Lepisorus annamensis (C.Chr.) Li Wang; Bot. J. Linn. Soc. 162(1): 35 (2010)

Distribution: China (Hainan); Thailand; Laos; Vietnam; peninsular Malaysia (Perak); Borneo (Sabah); Sulawesi

36 CHI 41 LAO THA VIE 42 BOR MLY SUL

Remarks: sect. 13 Belvisia

Synonyms: hidden

= Belvisia annamensis (C.Chr.) Tagawa; Acta Phytotax. Geobot. 22: 107 (1967)

= Belvisia annamensis (C.Chr.) V.N.Tu; Bot. Zhurn. (Moscow & Leningrad) 64(11): 1670 (1979), isonym

= Hymenolepis annamensis C.Chr.; Dansk Bot. Ark. 6: 54-70 (1929)

= Hymenolepis callifolia var. paltonioides C.Chr.; (1929)

= Macroplethus annamensis (C.Chr.) Tagawa; Acta Phytotax. Geobot. 11: 234 (1942)

species :

Lepisorus annuifrons (Makino) Ching; Bull. Fan Mem. Inst. Biol. Bot. 4: 70 (1933)

Distribution: South Korea (incl. Cheju Isl.); Japan

38 JAP KOR

Remarks: sect. 12 Ussurienses

Synonyms: hidden

= Pleopeltis annuifrons (Makino) Nakai; (1933)

= Polypodium annuifrons Makino; Bot. Mag. (Tokyo) 13: 48 (1899)

= Polypodium japonense Makino; Bot. Mag. (Tokyo) 12: 165 (1898), nomen

species :

Lepisorus asterolepis (Baker) Ching; Fl. Jiangsuensis 1: 74 (1977)

Distribution: China (Anhui, Chongqing, Fujian, Guangxi, Guizhou, Hubei, Hunan, Jiangsu, Jiangxi, Shaanxi, Sichuan, Yunnan, Zhejiang); Tibet; ?Bhutan; NE-India (Arunachal Pradesh); Myanmar (Kachin, etc.); Vietnam

36 CHI TIB 40 IND 41 MYA VIE

Remarks: sect. 10 Lepisorus. Included in L. macrosphaerus by Fraser-Jenkins et al. 2015 but a separate species fide Zhao et al. 2019 on genetic grounds

Synonyms: hidden

= Lepisorus longipes Ching & Z.Y.Liu; Bull. Bot. Res., Harbin 4(4): 25 (1984)

= Lepisorus macrosphaerus var. asterolepis (Baker) Ching; Bull. Fan Mem. Inst. Biol. Bot. 4(3): 74 (1933)

= Lepisorus macrosphaerus var. asterolepis (Baker) R.D.Dixit; Cens. Ind. Pterid. 41 (1984), isonym

= Pleopeltis asterolepis (C.Chr.) Panigrahi; Pl. Sci. Res. (India), Abstr. 23rd. Ann. Conf. Khurda: 36 (1998)

= Pleopeltis macrosphaera var. asterolepis (Baker) Panigrahi & Patnaik; Proc. Nat. Acad. Sci. India B. 34: 482, f. 1-5 (1964)

= Polypodium asterolepis Baker; J. Bot. 18: Brit. & For. 26: 230 (1880) (non Polypodium astrolepis Liebm.)

= Polypodium excavatum var. asterolepis (Baker) C.Chr.; Index Filic. 9: 525 (1906)

= Polypodium macrosphaerum var. asterolepis (Baker) C.Chr.; (1924)

species :

Lepisorus balteiformis (Brause) Hovenkamp; Blumea 43: 110 (1998)

Distribution: New Guinea (Irian Jaya, Papua New Guinea)

42 NWG

Remarks: sect. 1 Paragramma

Synonyms: hidden

= Paragramma balteiformis (Brause) Copel.; Ann. Cryptog. Phytopathol. 5: 190 (1947)

= Polypodium balteiforme Brause; Bot. Jahrb. Syst. 66: 194 (1920)

species :

Lepisorus bicolor (Takeda) Ching; Bull. Fan Mem. Inst. Biol. Bot. 4: 66 (1933)

Distribution: China (Guizhou, Sichuan, Yunnan); Tibet; Nepal; Bhutan; N-India (Himachal Pradesh, Uttarakhand, Darjeeling, Sikkim, Arunachal Pradesh); Myanmar (Kachin, Chin, etc.); Thailand

36 CHI TIB 40 BHU IND NEP 41 MYA THA

Remarks: sect. 8 Pachyphyllon

Synonyms: hidden

= Lepisorus kashyapii (Mehra) Mehra; Bir, Res. Bull. (N. S.) Punjab Univ. 13: 24 (1962), comb. inval.

= Lepisorus kashyapii var. major Bir & Trikha; Amer. Fern J. 71(2): 55 (1981)

= Lepisorus kashyapii var. minor Bir & Trikha; Amer. Fern J. 71(2): 55 (1981)

= Pleopeltis bicolor (Takeda) Sledge; Bull. Brit. Mus. (Nat. Hist.) Bot. 2: 138 (1960)

= Pleopeltis kashyapii (Mehra) Alston & Bonner; Candollea 15: 208 (1956), nom. inval.

= Polypodium bicolor (Takeda) Alston ex Schelpe; Amer. Fern J. 44: 59 (1954) (non Mett.)

= Polypodium excavatum var. bicolor Takeda; Notes Roy. Bot. Gard. Edinburgh 8: 280 (1915)

= Polypodium kashyapii Mehra; Ferns of Mussoorie, Punjab. Univ. Publ. 24 (1939), nom. nud. (fide Fraser-Jenkins et al. 2015)

species :

Lepisorus boninensis Ching; Bull. Fan Mem. Inst. Biol. Bot. 4: 76 (1933)

Distribution: Japan (Izu Isl.); Ryukyu Isl.; Bonin Isl. (Chichijima, Anijima, Otohtojima, Hahajima); Volcano Isl. (Kita-Iwojima, Minami-Iwojima)

38 JAP KZN NNS OGA

Remarks: sect. 15 Obtusi

Conservation Status: IUCN R. NT in Japan

Synonyms: hidden

= Pleopeltis boninensis (Christ) H.Ohba; Sci. Rep. Taihoku Univ. 4, 36: 120 (1971)

= Polypodium boninense Christ; Warb., Monsunia 1: 61 (1900)

species :

Lepisorus buergerianus (Miq.) C.F.Zhao, R.Wei & X.C.Zhang; Cladistics 36(3): 253 (2019)

Distribution: China (Chongqing, Gansu, Guangxi, Hubei, Hunan, Jiangxi, Sichuan, Yunnan, Zhejiang); Japan; Ryukyu Isl.; N-Vietnam

36 CHI 38 JAP NNS 41 VIE

Remarks: sect. 7 Lepidomicrosorium

Synonyms: hidden

= Lepidomicrosorium asarifolium Ching & K.H.Shing; Bot. Res. Acad. Sin. 1983(1): 11 (1983)

= Lepidomicrosorium brevipes Ching & K.H.Shing; Bot. Res. Acad. Sin. 1983(1): 13 (1983)

= Lepidomicrosorium buergerianum (Miq.) Ching & K.H.Shing; Fl. Jiangxi 1: 322 (1993)

= Lepidomicrosorium emeicola Ching & K.H.Shing; Bot. Res. Acad. Sin. 1983(1): 7 (1983)

= Lepidomicrosorium hederaceum (Christ) Ching; Bot. Res. Acad. Sin. 1983(1): 11 (1983)

= Lepidomicrosorium lanceolatum Ching & P.S.Wang; Bot. Res. Acad. Sin. 1983(1): 8 (1983)

= Lepidomicrosorium latibasis Ching & K.H.Shing; Bot. Res. Acad. Sin. 1983(1): 6 (1983)

= Lepidomicrosorium microsorioides (W.M.Chu) Ching & W.M.Chu; Bot. Res. Acad. Sin. 1983(1): 7 (1983)

= Lepidomicrosorium subhastatum (Baker) Ching; Bot. Res. Acad. Sin. 1983(1): 12 (1983)

= Lepidomicrosorium subsessile Ching & K.H.Shing; Bot. Res. Acad. Sin. 1983(1): 6 (1983)

= Lepidomicrosorium suijiangense Ching & W.M.Chu; Bot. Res. Acad. Sin. 1983(1): 13 (1983)

= Lepidomicrosorium yiliangense Ching & K.H.Shing; Bot. Res. Acad. Sin. 1983(1): 7 (1983)

= Leptochilus buergerianus (Miq.) Bosman; Leiden Bot. Ser. 14: 114 (1991)

= Microsorum buergerianum (Miq.) Ching; Bull. Fan Mem. Inst. Biol. Bot. 4: 295-311 (1933)

= Microsorum buergerianum var. ohwianum (Tagawa) Tagawa; Acta Phytotax. Geobot. 14: 9 (1949)

= Microsorum ohwianum Tagawa; J. Jap. Bot. 12: 752 (1936)

= Microsorum subhastatum (Baker) Ching; Bull. Fan Mem. Inst. Biol. Bot. 4: 295-311 (1933)

= Microsorum subhastatum var. longifrons (Takeda) Ching; Bull. Fan Mem. Inst. Biol. Bot. 4: 298 (1933)

= Neocheiropteris buergeriana (Miq.) Fraser-Jenk.; Indian J. Forest. 42(4): 314 (2019), isonym

= Neocheiropteris buergeriana (Miq.) Nakaike; Sa. Kurata & Nakaike, Ill. Pterid. Japan 2: 5 1 8 (1981)

= Neocheiropteris subhastata (Baker) Tagawa; J. Jap. Bot. 27: 217 (1952)

= Neolepisorus microsorioides W.M.Zhu; Acta Bot. Yunnan. 1(2): 96 (1979)

= Polypodium buergerianum Miq.; Ann. Mus. Bot. Lugduno-Batavi 3: 170 (1867)

= Polypodium buergerianum var. stipitatum Takeda; Notes Roy. Bot. Gard. Edinburgh 8(39): 290-291 (1915)

= Polypodium hederaceum Christ; Bull. Acad. Géogr. Bot. Mans 215, c. fig. (1902)

= Polypodium subhastatum Baker; J. Bot. 27: 177 (1889)

= Polypodium subhastatum var. hederaceum (Christ) Takeda; Notes Roy. Bot. Gard. Edinburgh 8(39): 292 (1915)

= Polypodium subhastatum var. longifrons Takeda; Notes Roy. Bot. Gard. Edinburgh 8(39): 292 (1915)

= Polypodium superficiale var. chinense Rosenst.; Repert. Spec. Nov. Regni Veg. 13(355-358): 134 (1914)

= Tricholepidium buergerianum (Miq.) Fraser-Jenk.; Taxon. Revis. Indian Subcontinental Pteridophytes 69 (2008)

species :

Lepisorus carnosus (J.Sm.) C.F.Zhao, R.Wei & X.C.Zhang; Cladistics 36(3): 252 (2019)

Name: Penny fern

Distribution: China (Fujian, Gansu, Guangdong, Guangxi, Guizhou, Hubei, Hunan, Jiangxi, Shanxi, Sichuan, Yunnan, Zhejiang); Bhutan; NE-India (Sikkim, Meghalaya, Arunachal Pradesh); Myanmar (Kachin, etc.); Thailand; Vietnam; Philippines (Luzon)

36 CHI 40 BHU IND 41 MYA THA VIE 42 PHI

Remarks: not in Nepal or Taiwan. Sect. 4 Lemmaphyllum. Lemmaphyllum diversum and L. drymoglossoides are frequently kept separate

Synonyms: hidden

= Colysis diversa (Rosenst.) Nakai ex Nemoto; Fl. Jap. Suppl. 20 (1936)

= Drymoglossum carnosum Hook.; Gen. Fil. t. 78A (1842)

= Drymoglossum carnosum var. minus Hook.; Sp. Fil. 5: 189 (1864)

= Drymoglossum carnosum Wall. ex J.Sm.; Hook. & Baker, Gen. Fil., t. 78A (1841) (excl. var.)

= Drymoglossum subcordatum Fée; Mém. Foug., 3. Hist. Vittar.: 29 (1852)

= Elaphoglossum carnosum (Hook.) Keyserl.; Polyp. Herb. Bunge: 36 (1873)

= Goniophlebium moupinense (Franch.) Bedd.; Suppl. Ferns Brit. Ind. 91 (1892)

= Lemmaphyllum adnascens Ching; Bull. Fan Mem. Inst. Biol. Bot. 4: 101 (1933)

= Lemmaphyllum carnosum (Hook.) C.Presl; Abh. Königl. Böhm. Ges. Wiss., ser. 5, 6: 158 (1851)

= Lemmaphyllum carnosum var. drymoglossoides (Baker) X.P.Wei; J. Syst. Evol. 51(4): 493 (2013)

= Lemmaphyllum carnosum var. longifolium S.Rahman & T.Sen; Indian Fern J. 15(1-2): 182 (1998)

= Lemmaphyllum christensenii Ching; Bull. Fan Mem. Inst. Biol. Bot. 4: 98 (1933) (non Polypodium christensenianum Maxon)

= Lemmaphyllum chui Saiki; J. Phytogeogr. Taxon. 35(2): 79 (1987)

= Lemmaphyllum diversum (Rosenst.) Tagawa; Acta Phytotax. Geobot. 14: 9 (1949)

= Lemmaphyllum drymoglossoides (Baker) Ching; Bull. Fan Mem. Inst. Biol. Bot. 4: 100 (1933)

= Lemmaphyllum intermedium (Ching) Li Wang; Bot. J. Linn. Soc. 162(1): 36 (2010)

= Lepidogrammitis adnascens (Ching) Ching; Sunyatsenia 5: 258 (1940)

= Lepidogrammitis christensenii (Ching) Ching; Sunyatsenia 5: 258 (1940)

= Lepidogrammitis diversa (Rosenst.) Ching; Acta Bot. Yunnan. 1(1): 24 (1979), et Fl. Xizangica 1: 313 (1983)

= Lepidogrammitis drymoglossoides (Baker) Ching; Sunyatsenia 5: 258 (1940)

= Lepidogrammitis elongata Ching; Fl. Tsinling. 2: 232 (1974)

= Lepidogrammitis intermedia Ching; Fl. Tsinling. 2: 231 (1974)

= Lepidogrammitis kansuensis Ching; Fl. Tsinling. 2: 232 (1974)

= Notholaena carnosa Wall.; Numer. List no. 138 (1829), nomen

= Oetosis carnosa (Wall. ex J.Sm.) Kuntze; Revis. Gen. Pl. 2: 817 (1891)

= Polypodium cyclophyllum Baker; Ann. Bot. (Oxford) 5: 473 (1891)

= Polypodium diversum Rosenst.; Hedwigia 66: 346 (1915)

= Polypodium drymoglossoides Baker; J. Bot. 25: 170 (1887)

= Polypodium moupinense Franch.; Nouv. Arch. Mus. Hist. Nat., sér. 2, 10: 121 (1887)

= Polypodium wangii C.Chr.; Index Filic., Suppl. Tert.: 161 (1934)

= Pteris piloselloides Thunb.; Fl. Jap. 331 (1784) (non L. (1763))

= Taenitis carnosa (Hook.) Mett.; Fil. Hort. Bot. Lips.: 28 (1856)

= Taenitis piloselloides (Thunb.) Wall.; Numer. List no. 62 (1829), nomen

species :

Lepisorus clathratus (C.B.Clarke) Ching; Bull. Fan Mem. Inst. Biol. Bot. 4: 71 (1933)

Distribution: Siberia (Altai); Kazakhstan; Kyrgyzstan; E-Afghanistan; China (Henan, Qinghai, Shaanxi, Guangxi, Sichuan, Yunnan); Tibet; Mongolia; Japan; Ryukyu Isl.; Taiwan; Pakistan (Kurram, Hazara); Nepal; Bhutan; India (Sikkim, Darjeeling Hills, Meghalaya, Arunachal Pradesh, Himachal Pradesh, Jammu & Kashmir)

30 SIB 32 KAZ KGZ 34 AFG 36 CHI TIB 37 MON 38 JAP NNS TAI 40 BHU IND NEP PAK

Remarks: sect. 9 Hymenophyton

Synonyms: hidden

= Lepisorus clathratus var. namegatae Sa.Kurata; Hokuriko J. Bot. 5: 17 (1956)

= Lepisorus henanensis Ching & S.K.Wu; Acta Bot. Yunnan. 5(1): 13 (1983)

= Lepisorus namegatae (Sa.Kurata) Ching & Y.X.Lin; Acta Bot. Yunnan. 5(1): 16 (1983)

= Lepisorus nepalensis K.Iwats.; Fl. E. Himal. 3rd. Rep.: 198 (1975) (fide Fraser-Jenkins 2008)

= Lepisorus petiolatus Ching & Y.X.Lin; Acta Phytotax. Sin. 31(5): 475 (1993)

= Lepisorus sinuatus (Ching & S.K.Wu) S.L.Yu; Acta Phytotax. Sin. 35: 343 (1997)

= Lepisorus thaipaiensis Ching & S.K.Wu; Acta Bot. Yunnan. 5(1): 8 (1983)

= Lepisorus variabilis Ching & S.K.Wu; Fl. Xizangica 1: 308 (1983) (fide Fraser-Jenkins 1997)

= Lepisorus venosus Ching & S.K.Wu; Acta Bot. Yunnan. 5(1,- 15 (1983)

= Platygyria kongtingensis Ching & Y.X.Lin; Acta Bot. Yunnan. 5(1): 22 (1983)

= Platygyria muliensis Ching & S.K.Wu; Acta Bot. Yunnan. 5(1): 22 (1983)

= Platygyria sinuata Ching & S.K.Wu; Acta Bot. Yunnan. 2(1): 71 (1980)

= Platygyria variabilis Ching & S.K.Wu; Acta Bot. Yunnan. 5(1): 21 (1983) (fide Fraser-Jenkins 1997)

= Pleopeltis clathrata (C.B.Clarke) Bedd.; Handb. Ferns Brit. India 348 (1883)

= Pleopeltis clathrata (C.B.Clarke) Czerep.; Vasc. Pl. Russia & Adj. States (former Russia): 414 (1995), isonym

= Pleopeltis clathrata var. namegatae (Sa.Kurata) Ohwi; Fl. Japan Pterid.: 147 (1975)

= Pleopeltis variabilis (Ching & S.K.Wu) S.Singh & Panigrahi; J. Hbr. Res. 3: 18 (1990) (fide Fraser-Jenkins 1997)

= Polypodium clathratum C.B.Clarke; Trans. Linn. Soc. London, Bot. 1: 559, t. 82, f. 1 (1880)

species :

Lepisorus confluens W.M.Chu; Acta Bot. Yunnan., Suppl. 5: 55 (1992)

Distribution: China (Yunnan)

36 CHI

Remarks: sect. 17 Pseudovittaria

Synonyms: hidden

= Lepisorus iridescens Ching & Y.X.Lin; Fl. Reipubl. Popul. Sin. 6(2): 82, t. 16: 3-4 (2000)

species :

Lepisorus contortus (Christ) Ching; Bull. Fan Mem. Inst. Biol. Bot. 4: 90 (1933)

Distribution: China (Anhui, Chongqing, ?Fujian, Gansu, Henan, Hubei, Jiangxi, Shaanxi, Sichuan, Yunnan, ?Zhejiang); Taiwan; Nepal; Bhutan; India (Himachal Pradesh, Uttarakhand, Sikkim, Arunachal Pradesh, S-India); Sri Lanka; Myanmar (Kachin, etc.); N-Thailand

36 CHI 38 TAI 40 BHU IND NEP SRL 41 MYA THA

Remarks: sect. 17 Pseudovittaria

Synonyms: hidden

= Lepisorus crassirhizoma Ching & Z.Y.Liu; Bull. Bot. Res., Harbin 3(4): 3 (1983)

= Lepisorus jinfoshanensis Ching; Bull. Bot. Res., Harbin 3(4): 4 (1983)

= Lepisorus khullarii P.C.Pande & H.C.Pande; Ill. Fern Fl. Kumaon Himalaya 1: 67 (2003) (fide Fraser-Jenkins 2008)

= Lepisorus nyalamensis Ching & S.K.Wu; Fl. Xizangica 1: 311 (1983) (fide Fraser-Jenkins et al. 2015)

= Pleopeltis contorta (Christ) Alston & Bonner; Candollea 15: 209 (1956)

= Polypodium contortum (Christ) Christ; Bot. Gaz. 61: 347 (1911)

= Polypodium lineare f. contortum (Christ) Takeda; Notes Roy. Bot. Gard. Edinburgh 8: 270 (1915)

= Polypodium lineare var. contortum Christ; Nuov. Giorn. Bot. Ital. n. s. 4: 98 (1897)

= Polypodium thunbergianum var. contortum (Christ) C.Chr.; Index Filic., Suppl. Tert.: 160 (1934)

species :

Lepisorus crassipes Ching & Y.X.Lin; Acta Bot. Yunnan. 5(1): 18 (1983)

Distribution: China (Gansu, Hebei, Hubei, Qinghai, Shaanxi, Sichuan)

36 CHI

Remarks: sect. 9 Hymenophyton. Often included in L. clathratus

Synonyms: hidden

= Lepisorus hsiawutaiensis Ching & S.K.Wu; Acta Bot. Yunnan. 5(1): 6 (1983)

= Lepisorus kansuensis Ching & S.K.Wu; Acta Bot. Yunnan. 5(l): 17 (1983)

= Lepisorus ligulatus Ching & S.K.Wu; Acta Bot. Yunnan. 5(1): 6 (1983)

= Lepisorus maowenensis Ching & S.K.Wu; Acta Bot. Yunnan. 5(1): 14 (1983)

= Lepisorus patungensis Ching & S.K.Wu; Acta Bot. Yunnan. 5(1): 11 (1983)

= Lepisorus shanyangensis Ching & Y.X.Lin; Acta Bot. Yunnan. 5(1): 19 (1983)

= Lepisorus sungpanensis Ching & Y.X.Lin; Acta Bot. Yunnan. 5(1): 16 (1983)

species :

Lepisorus distans (Makino) Ching; Acta Phytotax. Sin. 10: 302 (1965)

Distribution: Russian Far East (Kuril Isl.); China (Anhui, Jiangxi, Shandong, Zhejiang); Japan

31 RFE 36 CHI 38 JAP

Remarks: sect. 12 Ussurienses. Often included as variety in L. ussuriensis

Synonyms: hidden

= Lepisorus ussuriensis var. distans (Makino) Tagawa; Acta Phytotax. Geobot. 11: 236 (1942)

= Pleopeltis distans (Makino) Vorosch.; A. K. Skvortsov (ed.), Florist. issl. v razn. raionakh SSSR: 145 (1985) (non Kaulf. (1824), nec D. Don (1825))

= Pleopeltis ussuriensis var. distans (Makino) Okuyama

= Polypodium annuifrons var. distans (Makino) Nakai; Fl. Kor. 2: 414 (1911)

= Polypodium distans (Makino) Makino; Bot. Mag. (Tokyo) 20: 31 (1906), not Kaulfuss (1824), nor D. Don (1825), nor Raddi (1825)

= Polypodium lineare var. distans Makino; Bot. Mag. (Tokyo) 15: 60 (1901)

species :

Lepisorus eilophyllus (Diels) Ching; Bull. Fan Mem. Inst. Biol. Bot. 4: 65 (1933)

Distribution: China (Gansu, Guizhou, Hubei, Sichuan, Yunnan); Tibet; NE-India (Arunachal Pradesh, Nagaland, Sikkim); Myanmar (Kachin, etc.); Thailand

36 CHI TIB 40 IND 41 MYA THA

Remarks: sect. 17 Pseudovittaria

Conservation Status: IUCN VU for India (Fraser-Jenkins 2012)

Synonyms: hidden

= Lepisorus neolewisii K.H.Shing; Acta Phytotax. Sin. 31(6): 572 (1993)

= Lepisorus pseudolewisii K.H.Shing; Vasc. Pl. Hengduan Mts. 1: 166 (1993), without latin descr. or type

= Lepisorus subconfluens Ching; Bull. Fan Mem. Inst. Biol. Bot. 4: 85 (1933)

= Pleopeltis subconfluens (Ching) Panigrahi & Patnaik; Curr. Sci. 34: 127 (1965)

= Pleopeltis subconfluens (Ching) Tagawa & K.Iwats.; (1967), isonym

= Polypodium eilophyllum Diels; Bot. Jahrb. Syst. 29: 204 (1900), based on P. involutum Baker, J. Bot. 27: 177. 1889, not Desv. (1811), nor Mett. (1856)

= Polypodium involutum Baker; J. Bot. 27: 177 (1889) (non Desv. (1811) nec Mett. (1856))

= Polypodium lewisii Christ; (non Baker)

= Polypodium lineare C.Chr.; Index Filic. 540 (1906), nom. illeg. (non Thunb.)

= Polypodium subconfluens (Ching) C.Chr.; Index Filic., Suppl. Tert.: 159 (1934)

species :

Lepisorus elegans Ching & W.M.Chu; Acta Bot. Yunnan., Suppl. 5: 55 (1992)

Distribution: China (Yunnan)

36 CHI

Remarks: sect. 17 Pseudovittaria. Sometimes included in L. contortus

species :

Lepisorus ensatus (Thunb.) C.F.Zhao, R.Wei & X.C.Zhang; Cladistics 36(3): 251 (2019)

Distribution: China (Sichuan, Guizhou, Yunnan); South Korea (Cheju Isl.); Japan (incl. Izu Isl.); Ryukyu Isl.; Taiwan; NE-India (Manipur, Meghalaya); Vietnam; Philippines (Mountain Prov.)

36 CHI 38 JAP KOR NNS TAI 40 IND 41 VIE 42 PHI

Remarks: sect. 3 Neolepisorus

Conservation Status: IUCN EN for India (Fraser-Jenkins 2012)

Synonyms: hidden

= Microsorum ensatum (Thunb.) H.Itô; J. Jap. Bot. 11: 96 (1935)

= Microsorum reticulatum Ching ex L.Shi; Acta Phytotax. Sin. 37(1): 100 (1999)

= Neocheiropteris ensata (Thunb.) Ching; Bull. Fan Mem. Inst. Biol. Bot. 4: 108-112 (1933)

= Neocheiropteris ensata f. izuensis (Sa.Kurata & Satake ex Sa.Kurata) Seriz.; J. Phytogeogr. Taxon. 29(1): 24 (1981)

= Neocheiropteris ensata f. undulatodentata Sugim.; J. Geobot. 24(3): 61 (1977)

= Neocheiropteris ensata var. izuensis Sa.Kurata & Satake; J. Geobot. 5(1): 5 (1962 (May 1962))

= Neolepisorus ensatus (Thunb.) Ching; Bull. Fan Mem. Inst. Biol. Bot. 10: 14 (1940)

= Neolepisorus ensatus var. izuensis (Sa.Kurata & Satake) Nakaike; Checkl. Lycophytes Ferns Japan 36 (2013)

= Neolepisorus tenuipes Ching & K.H.Shing; Acta Phytotax. Sin. 21(3): 273 (1983) (fide Fraser-Jenkins 2008)

= Pleopeltis ensata (Thunb.) Bedd.; Suppl. Ferns Brit. Ind. 94 (1892), isonym

= Pleopeltis ensata (Thunb.) T.Moore; Index Filic. (T. Moore) 346 (1862)

= Polypodium ensatum Thunb.; Trans. Linn. Soc. 2: 341 (1794)

= Polypodium oligolepis Baker; Bull. Misc. Inform. Kew 1898: 231 (1898)

= Polypodium phyllitidis Thunb.; Fl. Jap. 335 (1784) (non L. (1753))

species :

Lepisorus excavatus (Bory ex Willd.) Ching; Bull. Fan Mem. Inst. Biol. Bot. 4: 68 (1939)

Distribution: Guinea; Sierra Leone; Liberia; Ivory Coast; Nigeria; Cameroon; Bioko Isl. [Fernando Poo]; São Tomé; Annobon Isl.; Equatorial Guinea (Rio Muni); Gabon; Congo [Brazzaville]; D.R.Congo [Zaire]; Rwanda; Burundi; Sudan; Ethiopia; Somalia; Uganda; Kenya; Tanzania; Angola; Zambia; Malawi; E-Zimbabwe; Mozambique; South Africa (Limpopo, Mpulamanga); Madagascar; Comores (Grande Comore, Mayotte); +Mauritius; La Réunion

22 GUI IVO LBR NGA SIE 23 ANN BIO BUR CMN CON EQG GAB RWA SAO ZAI 24 ETH SOM SUD 25 KEN TAN UGA 26 ANG MLW MOZ ZAM ZIM 27 SAF 29 COM MAU MDG REU

Remarks: sect. 8 Pachyphyllon. L. bampsii, L. vesiculari-paleaceus and L. schraderi are possibly separate species.

Synonyms: hidden

= Drynaria excavata (Bory ex Willd.) Fée; Mém. Foug. 5 (Gen. Filic.): 270 (1852)

= Drynaria phlebodes (Kunze ex Mett.) Fée; Mém. Foug. 5 (Gen. Filic.): 270 (1852)

= Lepisorus bampsii (Pic.Serm.) Zink; Syst. Fern Gen. Lepisorus 36 (1993)

= Lepisorus corradii Pic.Serm.; Nuov. Giorn. Bot. Ital. ser. 2, 52: 30, t. 3 (1946)

= Lepisorus mildbraedii (Hieron.) Pic.Serm.; Nuov. Giorn. Bot. Ital. ser. 2, 52: 33 (1946)

= Lepisorus phlebodes (Kunze ex Mett.) Ching; Bull. Fan Mem. Inst. Biol. Bot. 4: 72 (1933)

= Lepisorus preussii (Hieron.) Pic.Serm.; Nuov. Giorn. Bot. Ital. ser. 2, 52: 32 (1946)

= Lepisorus preussii var. angustifolius (Hieron.) Pic.Serm.; Nuov. Giorn. Bot. Ital. ser. 2, 52: 32 (1946)

= Lepisorus preussii var. angustipaleaceus (Hieron.) Pic.Serm.; Nuov. Giorn. Bot. Ital. ser. 2, 52: 32 (1946)

= Lepisorus preussii var. ledermannii (Hieron.) Pic.Serm.; Nuov. Giorn. Bot. Ital. ser. 2, 52: 32 (1946)

= Lepisorus preussii var. winkleri (Hieron.) Pic.Serm.; Nuov. Giorn. Bot. Ital. ser. 2, 52: 32 (1946)

= Lepisorus rotundus (Bonap.) Ching; Bull. Fan Mem. Inst. Biol. Bot. 4: 51 (1933)

= Lepisorus stolzii (Hieron.) Pic.Serm.; Nuov. Giorn. Bot. Ital. ser. 2, 52: 33 (1946)

= Lepisorus vesiculari-paleaceus (Hieron.) Pic.Serm.; Nuov. Giorn. Bot. Ital. ser. 2, 62: 33 (1946)

= Phymatodes excavata (Bory ex Willd.) C.Presl; Tent. Pterid. 196 (1836)

= Phymatodes simplex (Sw.) C.Presl; Tent. Pterid. 196 (1836), nom. illeg.

= Pleopeltis bampsii Pic.Serm.; Bull. Jard. Bot. Belg. 53(1-2): 200 (1983)

= Pleopeltis corradii (Pic.Serm.) Cufod.; Phyton (Horn) 4: 189 (1952)

= Pleopeltis excavata (Bory ex Willd.) Sledge; Bull. Brit. Mus. (Nat. Hist.) Bot. 2: 138 (1960), isonym

= Pleopeltis excavata (Bory ex Willd.) T.Moore; Index Filic. (T. Moore) 347 (1862)

= Pleopeltis grevilleana (Wall.) T.Moore; Index Filic. (T. Moore) 78 (1857)

= Pleopeltis mildbraedii (Hieron.) Pic.Serm.; Webbia 23: 189 (1968)

= Pleopeltis phlebodes (Kunze ex Mett.) Pic.Serm.; Webbia 23: 189 (1968)

= Pleopeltis preussii (Hieron.) Tardieu; Mém. Inst. Fr. Afr. Noire 28: 217, t. 44(6-8) (1953)

= Pleopeltis preussii var. angustifolia (Hieron.) Tardieu; Mém. Inst. Fr. Afr. Noire 28: 219 (1953), nom. nud.

= Pleopeltis preussii var. winkleri (Hieron.) Tardieu; Mém. Inst. Fr. Afr. Noire 28: 219 (1953), nom. nud.

= Pleopeltis rotunda (Bonap.) Tardieu; Mém. Inst. Fr. Afr. Noire 28: 219, t. 44(9-11) (1953)

= Pleopeltis simplex (C.Presl) Bedd.; Handb. Ferns Brit. India: 347 (1883), nom. illeg.

= Pleopeltis stolzii (Hieron.) Pic.Serm.; Webbia 23: 189 (1968)

= Pleopeltis vesicularipaleacea (Hieron.) Pic.Serm.; Webbia 23: 190 (1968)

= Polypodium bullatum Baker; J. Linn. Soc., Bot. 15: 420 (1876)

= Polypodium excavatum Bory ex Willd.; Sp. Pl. ed. 4, 5: 158 (1810)

= Polypodium excavatum var. concolor Takeda; Notes Roy. Bot. Gard. Edinburgh 8: 280 (1915), nom. inval.

= Polypodium excavatum var. phlebodes (Mett.) C.Chr.; Index Filic. 525 (1906)

= Polypodium excavatum var. vesiculari-paleaceum (Hieron.) C.Chr.; N. B. G. B. 9: 186 (1924)

= Polypodium grevilleanum Wall.; Numer. List no. 5169 (1831), nom. nud.

= Polypodium lineare var. simplex (C.Presl) Baker; Syn. Fil., ed. 2: 354 (1874)

= Polypodium mildbraedii Hieron.; Bot. Jahrb. Syst. 46: 391 (1911)

= Polypodium phlebodes Kunze ex Mett.; Abh. Senckenberg. Naturf. Ges. 2: 92, n. 163 (1857)

= Polypodium preussii Hieron.; Bot. Jahrb. Syst. 46: 386 (1911)

= Polypodium preussii var. angustifolium Hieron.; Bot. Jahrb. Syst. 46(3): 388 (1911)

= Polypodium preussii var. angustipaleaceum Hieron.; Bot. Jahrb. Syst. 46(3): 388 (1911)

= Polypodium preussii var. ledermannii Hieron.; Bot. Jahrb. Syst. 46(3): 388 (1911)

= Polypodium preussii var. winkleri Hieron.; Bot. Jahrb. Syst. 46(3): 388 (1911)

= Polypodium rotundum Bonap.; Notes Ptéridol. 16: 47 (1924)

= Polypodium simplex Sw.; J. Bot. (Schrader) 1800(2): 19 (1801), non Burm.f. 1768

= Polypodium stolzii Hieron.; Bot. Jahrb. Syst. 46: 389 (1911)

= Polypodium vesicularipaleaceum Hieron.; Bot. Jahrb. Syst. 46: 390 (1911)

species :

Lepisorus fortuni (T.Moore) C.M.Kuo; Taiwania 30: 68 (1985)

Distribution: China (Anhui, Gansu, Guangxi, Guizhou, Hubei, Hunan, Shaanxi, Sichuan, Yunnan, Zhejiang); Tibet; Ryukyu Isl.; Bonin Isl. (Otohtojima, Hahajima); Taiwan; Bhutan; NE-India (Arunachal Pradesh, Nagaland); Myanmar; Vietnam; peninsular Malaysia (Perak)

36 CHI TIB 38 NNS OGA TAI 40 BHU IND 41 MYA VIE 42 MLY

Remarks: sect. 3 Neolepisorus. L. chinensis might be a separate species. There is confusion between the synonyms of L. fortun(e)i and L. ovatus (Neocheiropteris ovata)

Conservation Status: IUCN EN for India (Fraser-Jenkins 2012)

Synonyms: hidden

= Drynaria fortuni T.Moore; Gard. Chron. 1855: 708 (1855) ex J. Sm. in Seem. (1857)

= Lepisorus undulatus Ching & Z.Y.Liu; Bull. Bot. Res., Harbin 4(4): 23 (1984)

= Microsorum chinense (Mett. ex Kuhn) Fraser-Jenk.; Taxon. Revis. Indian Subcontinental Pteridophytes 70 (2008)

= Microsorum excelsum Ching & S.K.Wu; Fl. Xizangica 1: 327 (1983) (fide Fraser-Jenkins 2008)

= Microsorum fortuni (T.Moore) Ching; Bull. Fan Mem. Inst. Biol. Bot. 4 304 (1933)

= Microsorum henryi (Christ) C.M.Kuo; Taiwania 30: 67 (1985) (fide Fraser-Jenkins 2008)

= Microsorum takedae (Nakai) H.Itô; J. Jap. Bot. 11: 97 (1935)

= Neocheiropteris chinensis (Mett. ex Kuhn) Fraser-Jenk., Pariyar & Kandel; Ferns Fern-Allies Nepal 1: 39 (2015)

= Neocheiropteris fortuni (T.Moore) Bosman ex Fraser-Jenk., Pariyar & Kandel; Ferns Fern-Allies Nepal 1: 39 (2015)

= Neocheiropteris henryi (Christ) Fraser-Jenk.; New Species Syndr. Indian Pteridol. 179-180 (1997)

= Neolepisorus fortuni (T.Moore) Li Wang; Bot. J. Linn. Soc. 162(1): 36 (2010)

= Neolepisorus phyllomanes f. deltoideus (Baker) Ching; Bull. Fan Mem. Inst. Biol. Bot. 10: 14 (1940), based on Polypodium deltoideum Baker, J. Bot. 26: 230. 1880, not Sw. (1788), nor Liebmann (1849)

= Phymatodes takedae Nakai; Bot. Mag. (Tokyo) 43: 2 (1929)

= Polypodium bandoi C.Chr.; Index Filic., Suppl. Tert.: 145 (1934)

= Polypodium chinense Mett. ex Kuhn; J. Bot. 6: 270 (1868)

= Polypodium deltoideum Baker; J. Bot. 26: 230 (1888) (non Sw. (1788) nec Liebm (1849))

= Polypodium fortuni (T.Moore) E.J.Lowe; Ferns 1: t. 42B (1856), not Kunze ex Mettenius (1856)

= Polypodium henryi Christ; Bull. Herb. Boissier 6: 873 (1898) (non Diels (1899))

= Polypodium normale var. polysorum Baker; J. Bot. 13: 202 (1875)

= Polypodium rosthornii Diels; Bot. Jahrb. Syst. 29: 205 (1900)

= Polypodium sinense Christ; Bull. Soc. Bot. France 52, Mem. 1: 16 (1905)

= Polypodium takedae (Nakai) C.Chr.; Index Filic., Suppl. Tert.: 160 (1934)

species :

Lepisorus hachijoensis Sa.Kurata; Sci. Rep. Yokosuka City Mus. 11: 40 (1965)

Distribution: Japan (Izu Isl.)

38 JAP

Remarks: sect. 15 Obtusi

Synonyms: hidden

= Pleopeltis hachijoensis (Sa.Kurata) Á.Löve & D.Löve; Taxon 26(2-3): 324 (1977)

species :

Lepisorus henryi (Hieron. ex C.Chr.) Li Wang; Bot. J. Linn. Soc. 162(1): 35 (2010)

Distribution: China (Yunnan); Nepal; Bhutan; NE-India (Assam, Sikkim, Arunachal Pradesh, Darjeeling); Myanmar (Kachin, etc.); Thailand; Laos; Vietnam

36 CHI 40 BHU IND NEP 41 LAO MYA THA VIE

Remarks: sect. 13 Belvisia

Synonyms: hidden

= Belvisia henryi (Hieron. & C.Chr.) Raym.; Mém. Jard. Bot. Montréal 55: 32 (1962)

= Belvisia henryi (Hieron. ex C.Chr.) Tagawa; H. Hara, Fl. E. Himal. 490 (1960)

= Hymenolepis henryi Hieron. ex C.Chr.; Dansk Bot. Ark. 6(3): 67 (1929)

= Macroplethus henryi (Hieron. ex C.Chr.) Tagawa; Acta Phytotax. Geobot. 11: 234 (1942)

species :

Lepisorus heterolepis (Rosenst.) Ching; Bull. Fan Mem. Inst. Biol. Bot. 4: 86 (1933)

Distribution: China (Yunnan)

36 CHI

Remarks: sect. 17 Pseudovittaria. Included in L. contortus by Kandel & Fraser-Jenkins (2020) but kept separate on genetic grounds by Zhao et al. 2019

Synonyms: hidden

= Pleopeltis heterolepis (Rosenst.) Tagawa & K.Iwats.; Acta Phytotax. Geobot. 22: 100 (1967)

= Polypodium heterolepis (Rosenst.) C.Chr.; Index Filic., Suppl. Tert.: 150 (1934)

= Polypodium lineare var. heterolepis Rosenst.; Repert. Spec. Nov. Regni Veg. 12. 247 (1913)

= Polypodium loriforme var. heterolepis (Rosenst.) C.Chr.; Contr. U.S. Natl. Herb. 26(6): 318 (1931)

species :

Lepisorus jakonensis (Blanf.) Fraser-Jenk. & J.Krieg.; Indian J. Forest. 42(4): 310 (2019)

Distribution: China (Yunnan); Tibet; Nepal; Bhutan; India (Himachal Pradesh, Uttarakhand, Arunachal Pradesh, Sikkim, ?Darjeeling); Myanmar (Kachin)

36 CHI TIB 40 BHU IND NEP 41 MYA

Remarks: sect. 2 Pseudoclathrati

Synonyms: hidden

= Ellipinema jakonense (Blanf.) Li Bing Zhang & Liang Zhang; Molec. Phylogen. Evol. 148: 6 (2020)

= Lepisorus jakonensis (Blanf.) Ching; Acta Bot. Yunnan. 5(1): 5 (1983), comb. inval.

= Lepisorus pseudoclathratus Ching & S.K.Wu; Acta Bot. Yunnan. 5(1): 10 (1983) (tentative synonym fide Zhao et al. 2019

= Pleopeltis jakonensis (Blanf.) S.Singh & Panigrahi; J. Hill Res. 3: 17 (1990), with incorrect basionym ref.

= Polypodium clathratum var. jakonense Blanf.; J. Simla Naturalists´ Soc. 1(2): 14 (1886)

species :

Lepisorus kawakamii (Hayata) Tagawa; Acta Phytotax. Geobot. 5: 109 (1936)

Distribution: Taiwan

38 TAI

Remarks: sect. 10 Lepisorus

Synonyms: hidden

= Polypodium arisanense Rosenst.; Hedwigia 66: 347 (1915) (non Hayata (1914))

= Polypodium kawakamii Hayata; Bot. Mag. (Tokyo) 23: 77 (1909)

= Polypodium macrosorum Baker; J. Bot. 23: 106 (1885) (non Fée (1850))

species :

Lepisorus kolesnikovii (Tzvelev) Shmakov; Opred. Paporotn. Rossii (Key Ferns Russia) 32 (1999)

Distribution: Russian Far East (Khabarovsk, Primorye)

31 RFE

Synonyms: hidden

= Pleopeltis kolesnikovii Tzvelev; Novosti Sist. Vyssh. Rast. 26: 10 (1989)

species :

Lepisorus kuchenensis (Y.C.Wu) Ching; Bull. Fan Mem. Inst. Biol. Bot. 4: 69 (1933)

Distribution: China (Guangxi, Guizhou, Yunnan); +Taiwan; Vietnam

36 CHI 38 TAI 41 VIE

Remarks: sect. 10 Lepisorus

Conservation Status: very rare or extinct in Taiwan, not seen there since the 1940s

Synonyms: hidden

= Pleopeltis kuchenensis (Y.C.Wu) Á.Löve & D.Löve; Taxon 26(2-3): 324 (1977)

= Polypodium kuchenense Y.C.Wu; Y. C. Wu, K. K. Wong & Pong, Polyp. Yaoshan. 276, t. 129 (1932)

species :

Lepisorus kuratae T.Fujiw. & Seriz.; J. Plant Res. (2018)

Distribution: Japan (Honshu, Shikoku)

38 JAP

Remarks: An allotetraploid species derived from L. thunbergianus and an ancestral ¿race of L. tosaensis

species :

Lepisorus lewisii (Baker) Ching; Bull. Fan Mem. Inst. Biol. Bot. 4: 65 (1933)

Distribution: China (Anhui, Fujian, Guangdong, Guangxi, Guizhou, Hunan, Jiangxi, Sichuan, Zhejiang)

36 CHI

Remarks: sect. 17 Pseudovittaria

Synonyms: hidden

= Polypodium lewisii Baker; J. Bot. 13: 201 (1875)

species :

Lepisorus likiangensis Ching & S.K.Wu; Acta Bot. Yunnan. 5(1): 12 (1983)

Distribution: China (Gansu, Guizhou, Qinghai, Shaanxi, Sichuan)

36 CHI

Remarks: sect. 9 Hymenophyton. Often included in L. clathratus s. l.

Synonyms: hidden

= Lepisorus coaetaneus Ching & Y.X.Lin; Acta Bot. Yunnan. 5(1): 12 (1983)

= Lepisorus shansiensis Ching & Y.X.Lin; Acta Bot. Yunnan. 5(1): 18 (1983)

= Lepisorus shensiensis Ching & S.K.Wu; Acta Bot. Yunnan. 5(1): 14 (1983)

species :

Lepisorus lineariformis Ching & S.K.Wu; Fl. Xizangica 1: 309 (1983)

Distribution: SE-Tibet (Medog)

36 TIB

Remarks: sect. 17 Pseudovittaria. Included in L. contortus by Kandel & Fraser-Jenkins 2020 but kept separate on genetic basis by Zhao et al. 2019

species :

Lepisorus longifolius (Blume) Holttum; Rev. Fl. Malaya 2: 151 (1954)

Distribution: Myanmar; peninsular Thailand; Cambodia; Vietnam; peninsular Malaysia (widespread); Singapore; Sumatra; Borneo (Mt. Kinabalu, Sabah, Sarawak, Kalimantan); Sulawesi; Palawan; Philippines; Java

41 CBD MYA THA VIE 42 BOR JAW MLY PHI PLW SNG SUL SUM

Remarks: not in India. Sect. 1 Paragramma

Conservation Status: endangered (Singapore)

Synonyms: hidden

= Drynaria revoluta J.Sm.; J. Bot. (Hooker) 3: 421 (1841), nom. nud.

= Grammitis decurrens Blume; Enum. Pl. Javae 2: 119 (1828), nom. illeg., non Wall. ex Hook. & Grev. 1827

= Grammitis longifolia Blume; Enum. Pl. Javae 2: 119 (1828)

= Microsorum longifolium (Blume) Copel.; Univ. Calif. Publ. Bot. 16: 112 (1929)

= Niphobolus longifolius (Blume) Keyserl.; Polyp. Herb. Bunge 38 (1873)

= Paragramma decurrens (Blume) T.Moore; Index Filic. (T. Moore) 32 (1857), nom. illeg.

= Paragramma longifolia (Blume) T.Moore; Index Filic. (T. Moore) 32 (1857)

= Phymatodes longifolia (Blume) J.Sm.; Cat. Cult. Ferns 9 (1857)

= Phymatodes revoluta (Alderw.) Tardieu & C.Chr.; Lecomte, Fl. Indo-Chine 7(2): 468 (1941)

= Phymatosorus longifolius (Blume) Bosman & Fraser-Jenk.; New Species Syndr. Indian Pteridol. 196 (1997)

= Phymatosorus revolutus (J.Sm.) V.N.Tu; J. Biol. (Vietnam) 1(2): 20 (1979)

= Pleopeltis decurrens Blume; Enum. Pl. Javae 2: addend. (1828)

= Pleopeltis longifolia (Blume) Bedd.; Handb. Ferns Brit. India 349, t. 196 (1883)

= Pleopeltis revoluta (Alderw.) Alderw.; Bull. Dép. Agric. Indes Néerl. 27: 5 (1909)

= Polypodium contiguum Hook.; Hooker´s Icon. Pl. 6: t. 987 (1854) (nomen, non Brack. (1854))

= Polypodium decurrens (Blume) Kunze; Bot. Zeitung (Berlin) 421 (1846), nom. illeg. (non Raddi (1819))

= Polypodium longifolium (Blume) Mett.; Abh. Senckenberg. Naturf. Ges. 2: 87, n. 153, t. 1, f. 6 (1857) (non Cav. (1802) nec C. Presl (1822))

= Polypodium productum Christ; Philipp. J. Sci., Bot. 2: 178 (1907)

= Polypodium revolutum (J.Sm.) C.Chr.; Index Filic. 331 (1905), 559 (1906)

= Polypodium revolutum Alderw.; Malayan Ferns: 638 (1909)

species :

Lepisorus loriformis (Wall. ex Mett.) Ching; Bull. Fan Mem. Inst. Biol. Bot. 4: 81 (1933)

Distribution: China (Gansu, Hubei, Shaanxi, Sichuan, Yunnan); Tibet; Nepal; Bhutan; India (N-India, Uttarakhand, Sikkim, Darjeeling, Meghalaya, Arunachal Pradesh); Myanmar (Kachin)

36 CHI TIB 40 BHU IND NEP 41 MYA

Remarks: sect. 16 Sclerophyllon. L. steniste might be a separate species

Synonyms: hidden

= Drynaria loriformis (Wall. ex Mett.) J.Sm.; J. Bot. (Hooker) 4: 61 (1841)

= Lepisorus loriformis f. steniste (C.B.Clarke) Kholia; Ferns Fern-Allies Sikkim 2: 126 (2014)

= Lepisorus loriformis var. longifolius Bir & Trikha; Amer. Fern J. 64: 63 (1974)

= Lepisorus loriformis var. steniste (C.B.Clarke) Ching; Bull. Fan Mem. Inst. Biol. Bot. 4(3): 82 (1933)

= Lepisorus steniste (C.B.Clarke) Y.X.Lin; Fl. Reipubl. Popul. Sin. 6(2): 69 (2000)

= Lepisorus tenuicauda Ching; Acta Bot. Austro-Sin. 1: 22 (1983 (fide Fraser-Jenkins 1997)

= Lepisorus xiphopteris (Baker) W.M.Chu ex Y.X.Lin; Fl. Reipubl. Popul. Sin. 6(2): 68 (2000)

= Lepisorus yunnanensis Ching; Bull. Fan Mem. Inst. Biol. Bot. 11: 75 (1941)

= Phymatodes loriformis (Wall. ex Mett.) C.Presl; Tent. Pterid. 196 (1836)

= Pleopeltis linearis var. steniste (C.B.Clarke) Bedd.; Handb. Ferns Brit. India 347 (1883)

= Pleopeltis loriformis f. angustifrons (Takeda) C.Chr.; Contr. U.S. Natl. Herb. 26(6): 318 (1931)

= Pleopeltis loriformis (Wall. ex Mett.) T.Moore; Index Filic. (T. Moore) 346 (1862)

= Pleopeltis loriformis var. steniste (C.B.Clarke) Panigrahi & Patnaik; Proc. Natl. Acad. Sci. India, B 34: 481 (1964)

= Pleopeltis tenuicauda (Ching) Panigrahi; Pl. Sci. Res. (India), Abstr. 23rd. Ann. Conf. Khurda: 35 (1998)

= Polypodium excavatum var. loriforme (Wall. ex Mett.) C.Chr.; Index Filic. 9: 525 (1906)

= Polypodium lineare f. steniste (C.B.Clarke) Takeda; Notes Roy. Bot. Gard. Edinburgh 8(39): 273 (1915)

= Polypodium lineare subsp. loriforme (Wall. ex Mett.) C.Chr.; Index Filic., Suppl. (1906-1912) 1: 54 (1917)

= Polypodium lineare var. loriforme (Wall. ex Mett.) Takeda; Notes Roy. Bot. Gard. Edinburgh 8: 272 (1915)

= Polypodium lineare var. steniste C.B.Clarke; Trans. Linn. Soc. London, Bot. 1: 559 (1880)

= Polypodium loriforme f. angustifrons (Takeda) C.Chr.; Contr. U.S. Natl. Herb. 26: 318 (1931)

= Polypodium loriforme var. steniste (C.B.Clarke) C.Chr.; Contr. U.S. Natl. Herb. 26: 318 (1931)

= Polypodium loriforme Wall.; Numer. List no. 271 (1829), nomen

= Polypodium loriforme Wall. ex Mett.; Abh. Senckenberg. Naturf. Ges. 2: 92 (1856)

= Polypodium mengtzeanum Baker; Bull. Misc. Inform. Kew 1906: 14 (1906)

= Polypodium oblongisorum C.Chr.; Index Filic. 549 (1906)

= Polypodium subimmersum Baker; Bull. Misc. Inform. Kew 1895: 55 (1895)

= Polypodium subimmersum f. angustifrons Takeda; Notes Roy. Bot. Gard. Edinburgh 8(39): 276 (1915)

= Polypodium subimmersum f. mengtzeanum (Baker) Takeda; Notes Roy. Bot. Gard. Edinburgh 8(39): 276 (1915)

= Polypodium subintegrum Baker; Bull. Misc. Inform. Kew 1898: 231 (1898) (non Baker (1877))

= Polypodium xiphiopteris Baker; Bull. Misc. Inform. Kew 1906: 13 (1906)

species :

Lepisorus luchunensis Y.X.Lin; Fl. Reipubl. Popul. Sin. 6(2): 50, t. 8 (2000)

Distribution: China (Yunnan)

36 CHI

Remarks: sect. 10 Lepisorus. Status not finally confirmed

species :

Lepisorus macrosphaerus (Baker) Ching; Bull. Fan Mem. Inst. Biol. Bot. 4: 73 (1933)

Distribution: China (Guizhou, Hubei, Sichuan, Yunnan); Tibet; Bhutan

36 CHI TIB 40 BHU

Remarks: sect. 10 Lepisorus. L. asterolepis is included here by Fraser-Jenkins et al. 2015 but kept separate by Zhao et al. 2019

Synonyms: hidden

= Lepisorus macrosphaerus f. maximus (Ching) Y.X.Lin; Fl. Reipubl. Popul. Sin. 6(2): 64 (2000)

= Lepisorus macrosphaerus f. minimus (Ching) Y.X.Lin; Fl. Reipubl. Popul. Sin. 6(2): 65 (2000)

= Lepisorus macrosphaerus var. maximus Ching; Fl. Xizangica 1: 303 (1983)

= Lepisorus macrosphaerus var. minimus Ching; Fl. Xizangica 1: 304 (1983)

= Pleopeltis macrosphaera (Baker) Panigrahi & Patnaik; Proc. Nat. Acad. Sci. India 34: 481 (1964), nom. superfl.

= Pleopeltis macrosphaera (Baker) Pic.Serm.; Webbia 23(1): 189 (1968), nom. superfl.

= Polypodium intramarginale Baker; Christ, Bull. Herb. Boissier, sér. 2, 3: 509 (1903)

= Polypodium macrosphaerum Baker; Bull. Misc. Inform. Kew 1895: 55 (1895)

= Polypodium simplex var. esquirolii Christ; Bull. Acad. Int. Géogr. Bot. 15: 247 (1906)

species :

Lepisorus maculosus (Christ) C.F.Zhao, R.Wei & X.C.Zhang; Cladistics 36(3): 252 (2019)

Distribution: China (Sichuan, Yunnan, etc.); Tibet; Bhutan; India (Arunachal Pradesh, Manipur, Nagaland, Mizoram); Myanmar; Thailand; Laos; Vietnam; peninsular Malaysia; Sumatra

36 CHI TIB 40 BHU IND 41 LAO MYA THA VIE 42 MLY SUM

Remarks: sect. 5 Tricholepidium. Often included in L. normale but kept separate by Zhao et al. 2019. Neocheiropteris venosa could be a separate species but requires a new combination in Lepisorus.

Synonyms: hidden

= Lepisorus chapaensis C.Chr. & Tardieu; Notul. Syst. (Paris) 8: 186 (1939)

= Microsorum subnormale (Nakai) H.Itô; J. Jap. Bot. 11: 97 (1935)

= Neocheiropteris maculosa (Christ) Fraser-Jenk.; Indian J. Forest. 42(4): 314 (2019)

= Neocheiropteris venosa (Ching) Fraser-Jenk.; Indian J. Forest. 42(4): 314 (2019)

= Phymatodes subnormalis Nakai; Bot. Mag. (Tokyo) 43. 3 (1929)

= Pleopeltis subnormalis Alderw.; Bull. Jard. Bot. Buitenzorg, ser. 3, 2: 165 (1920)

= Polypodium maculosum Christ; Bull. Herb. Boissier 6: 872 (1896)

= Polypodium subnormale (Alderw.) C.Chr.; Index Filic., Suppl. Tert.: 159 (1934)

= Tricholepidium angustifolium Ching; Acta Phytotax. Geobot. 29(1-5): 44 (1978)

= Tricholepidium angustifolium var. falcatolineare Ching; Acta Phytotax. Geobot. 29: 44 (1978)

= Tricholepidium angustifolium var. lanceolatum (Ching & S.K.Wu) Y.X.Lin; Fl. Reipubl. Popul. Sin. 6(2): 106 (2000)

= Tricholepidium chapaense (C.Chr. & Tardieu) Ching; Acta Phytotax. Geobot. 29(1-5): 44 (1978)

= Tricholepidium intermedium Ching; Acta Phytotax. Geobot. 29(1-5): 44 (1978)

= Tricholepidium lanceolatum Ching & S.K.Wu; Acta Phytotax. Geobot. 29(1-5): 45 (1978)

= Tricholepidium maculosum (Christ) Ching; Acta Phytotax. Geobot. 29(1-5): 45 (1978)

= Tricholepidium maculosum var. subnormale (Alderw.) Ching; Acta Phytotax. Geobot. 29(1-5): 45 (1978)

= Tricholepidium subnudum Ching & S.K.Wu; Fl. Xizangica 1: 314 (1983)

= Tricholepidium tibeticum Ching & S.K.Wu; Acta Phytotax. Geobot. 29(1-5): 46 (1978)

= Tricholepidium venosum Ching; Acta Phytotax. Geobot. 29(1-5): 45 (1978)

species :

Lepisorus mamas Hovenkamp; Blumea 43: 112 (1998)

Distribution: Sumatra

42 SUM

species :

Lepisorus marginatus Ching; Fl. Tsinling. 2: 233 (1974), without type

Distribution: China (Chongqing, Gansu, Hebei, Henan, Hubei, Shaanxi, Shanxi, Sichuan)

36 CHI

Remarks: sect. 10 Lepisorus

species :

Lepisorus medioximus T.Fujiw., K.Hori & Khine; PhytoKeys 201: 26-27 (2022)

Distribution: Myanmar (Shan)

41 MYA

Remarks: sect. 17 Pseudovittaria

species :

Lepisorus medogensis Ching & Y.X.Lin; Acta Phytotax. Sin. 22(5): 401 (1984)

Distribution: SE-Tibet (Medog)

36 TIB

Remarks: sect. 17 Pseudovittaria

Synonyms: hidden

= Lepisorus zosterifolius Ching & Y.X.Lin; Acta Phytotax. Sin. 22(5): 401 (1984)

species :

Lepisorus megasorus (C.Chr.) Ching; Bull. Fan Mem. Inst. Biol. Bot. 4: 79 (1933)

Distribution: Taiwan

38 TAI

Remarks: sect. 10 Lepisorus

Synonyms: hidden

= Polypodium hypochrysum Hayata; Icon. Pl. Formosan. 6: 314, f. 127 (1915)

= Polypodium megasorum C.Chr.; Index Filic. 544 (1906), based on later homonym P. macrosorum Baker, not Fée (1850)

species :

Lepisorus mehrae Fraser-Jenk.; New Species Syndr. Indian Pteridol. 159 (1997)

Distribution: China (Yunnan, ?Sichuan); Tibet; Nepal; Bhutan; N-India (Himachal Pradesh, Uttarakhand, Sikkim, Darjeeling); Myanmar (Kachin, Chin)

36 CHI TIB 40 BHU IND NEP 41 MYA

Synonyms: hidden

= Lepisorus krameri P.C.Pande & H.C.Pande; Indian Fern J. 11(1-2): 98 (1994 publ. 1995), et Ill. Fern Fl. Kumaon Himalaya 1: 68 (2003), nom. nud. (fide Fraser-Jenkins 2015)

= Lepisorus paleparaphysus Y.X.Lin; Fl. Reipubl. Popul. Sin. 6(2): 75 (2000)

species :

Lepisorus microphyllus (C.Presl) Ebihara & T.Fujiw.; Bull. Natl. Mus. Nat. Sci., Tokyo, B. 49(3): 101 (2023)

Distribution: China (Anhui, Fujian, Guangdong, Guangxi, Hainan, Hubei, Jiangxi, Yunnan, Zhejiang); South Korea; Japan; Taiwan; NE-India (Arunachal Pradesh); Myanmar (Kachin, etc.); Vietnam

36 CHI 38 JAP KOR TAI 40 IND 41 MYA VIE

Remarks: according to Knapp (Ferns of Taiwan, 2011 ff.) var. obovatum should be included in the typical variety. The Taiwanese plants belong possibly to L. carnosum fide Knapp (Ferns of Taiwan, 2011 ff.)

Conservation Status: IUCN VU for India (Fraser-Jenkins 2012)

Synonyms: hidden

= Drymoglossum carnosum var. obovatum Harr.; J. Linn. Soc., Bot. 16: 33 (1877)

= Drymoglossum microphyllum (C.Presl) C.Chr.; Index Filic. 246 (1905)

= Drymoglossum nobukoanum Makino; J. Jap. Bot. 7: 8 (1931)

= Drymoglossum obovatum (Harr.) Christ; J. Bot. (Morot) 19: 73 (1905)

= Drymoglossum obovatum var. lutchuense Nakai; Bot. Mag. (Tokyo) 40(475): 396 (1926)

= Drymoglossum spatulatum C.Presl; Tent. Pterid. 227 (1836)

= Lemmaphyllum carnosum subsp. microphyllum (C.Presl) Fraser-Jenk. & Kandel; Indian J. Forest. 42(4): 310 (2019)

= Lemmaphyllum carnosum var. microphyllum (C.Presl) X.P.Wei; J. Syst. Evol. 51(4): 493 (2013)

= Lemmaphyllum microphyllum C.Presl; Abh. Königl. Böhm. Ges. Wiss. 5(6) (Epim. Bot.): 263 (1851)

= Lemmaphyllum microphyllum var. lutchuense (Nakai) C.Chr.; Dansk Bot. Ark. 6(3): 47, pl. 5, f. 4 (1929)

= Lemmaphyllum microphyllum var. nobukoanum (Makino) Nakaike; Enum. Pterid. Jap.: Filic. 332 (1975)

= Lemmaphyllum microphyllum var. obovatum (Harr.) C.Chr.; Dansk Bot. Ark. 6(3): 47 (1929)

= Lemmaphyllum microphyllum var. spathulatum (C.Presl) Nakaike; J. Nippon Fernist Club 3(Suppl. 2): 104 (2004)

= Lemmaphyllum minimum S.H.Fu; Acta Phytotax. Sin. 5: 75, f. 1-4 (1956)

= Lemmaphyllum nobukoanum (Makino) Ching; Icon. Filic. Sin. ad., t. 80 (1934)

= Lemmaphyllum spathulatum C.Presl; Abh. Königl. Böhm. Ges. Wiss., ser. 5, 6: 158 (1851)

= Lemmaphyllum squamosum C.Chr.; Dansk Bot. Ark. 6: no. 3, 48 (1929)

= Taenitis microphylla (C.Presl) Mett.; Kuhn, Filic. Afr. 58 (1868)

species :

Lepisorus mikawanus Sa.Kurata; Sci. Rep. Yokosuka City Mus. 11: 38 (1965)

Distribution: Japan

38 JAP

Remarks: hexaploid species in the thunbergianus complex

Conservation Status: IUCN R

species :

Lepisorus minor (W.M.Chu) C.F.Zhao, R.Wei & X.C.Zhang; Cladistics 36(3): 251 (2019)

Distribution: China (Yunnan)

36 CHI

Remarks: sect. 3 Neolepisorus. Sometimes included in L. ensatus

Synonyms: hidden

= Neolepisorus minor W.M.Chu; Acta Bot. Yunnan. 1(2): 95 (1979) (fide Fraser-Jenkins 2008)

species :

Lepisorus mishmica Benniamin & Fraser-Jenk.; Indian J. Forest. 42(4): 310 (2019)

Distribution: India (Arunachal Pradesh)

40 IND

Remarks: sect. 17 Pseudovittaria

species :

Lepisorus miyoshianus (Makino) Fraser-Jenk. & Subh.Chandra; Taxon. Revis. Indian Subcontinental Pteridophytes 37 (2008)

Distribution: China (Guangdong, Guizhou, Hubei, Hunan, Shaanxi, Sichuan, Yunnan, Zhejiang); Tibet; Japan; Taiwan; India (Arunachal Pradesh)

36 CHI TIB 38 JAP TAI 40 IND

Remarks: sect. 14 Drymotaenium

Conservation Status: IUCN CR for India (Fraser-Jenkins 2012)

Synonyms: hidden

= Drymotaenium miyoshianum (Makino) Makino; Bot. Mag. (Tokyo) 15: 109 (1901)

= Drymotaenium nakaii Hayata; Bull. Soc. Bot. France 58. 565, t. 19 (1911)

= Monogramma robusta (Christ) C.Chr.; Index Filic. 431 (1906)

= Pleurogramme robusta Christ; Bull. Herb. Boissier 6: 867 (1898)

= Taenitis miyoshiana Makino; Bot. Mag. (Tokyo) 11: 26 (1898)

species :

Lepisorus monilisorus (Hayata) Tagawa; Acta Phytotax. Geobot. 11: 303 (1942)

Distribution: Taiwan

38 TAI

Remarks: sect. 15 Obtusi

Synonyms: hidden

= Polypodium lineare var. monilisorum Hayata; Icon. Pl. Formosan. 4: 248, f. 173 (1914)

species :

Lepisorus morrisonensis (Hayata) H.Itô; J. Jap. Bot. 11: 92 (1935)

Distribution: China (Sichuan, Yunnan); Tibet; Taiwan; Nepal; Bhutan; India (Uttarakhand, Sikkim, ?Darjeeling, Meghalaya, Himachal Pradesh, Jammu & Kashmir, Arunachal Pradesh); Myanmar; N-Thailand

36 CHI TIB 38 TAI 40 BHU IND NEP 41 MYA THA

Remarks: sect. 8 Pachyphyllon

Synonyms: hidden

= Lepisorus mediocris Ching & Khullar; (1988) (nom. nud., fide Fraser-Jenkins (1997))

= Lepisorus reichsteinii Khullar; (1988) (nom. nud., fide Fraser-Jenkins (1997))

= Polypodium morrisonense Hayata; Bot. Mag. (Tokyo) 23: 77 (1909)

species :

Lepisorus mucronatus (Fée) Li Wang; Bot. J. Linn. Soc. 162(1): 35 (2010)

Name: Needle fern, Tailed fern

Distribution: China (Yunnan); Tibet; Taiwan; Sri Lanka; Myanmar (Kachin, etc.); Thailand; Laos; Cambodia; Vietnam; peninsular Malaysia (widespread); Sumatra; Borneo (Mt. Kinabalu, Sabah, Sarawak); Sulawesi; Palawan; Philippines; Java; Lesser Sunda Isl. (Bali, Lombok, Timor, etc.); Moluccas (Seram, etc.); New Guinea (Irian Jaya, Papua New Guinea); Bismarck Arch. (New Ireland, New Hanover); Solomon Isl. (Bougainville, Guadalcanal, Kolombangara, Malaita, New Georgia, Rendova, Rennell, Santa Ysabel, Vella Lavella, Makira); Australia (E-Queensland, NE-New South Wales); New Caledonia; Santa Cruz Isl.; Vanuatu; Tonga (Kao, Tafahi); Fiji (Viti Levu, Vanua Levu, Kadavu, Gau, Koro, Ovalau, Taveuni); [Western] Samoa (Savai´i); Cook Isl. (Rarotonga); Marquesas Isl. (Nuku Hiva, Ua Huka, Ua Pou, Hiva Oa, Tahuata, Fatu Hiva); Austral Isl. (Tubuai, Raivavae); Palau (Babeldaob, Koror, Malakal); Northern Marianas (Rota Isl.); Guam

36 CHI TIB 38 TAI 40 SRL 41 CBD LAO MYA THA VIE 42 BOR JAW LSI MLY MOL NWG PHI PLW SUL SUM 43 BIS SOL 50 AUS 60 FIJ NWC SAM SCZ TON VAN 61 AUI COO MRQ 62 GUM MRN PLU

Remarks: sect. 13 Belvisia

Synonyms: hidden

= Acrostichum spicatum var. schneideri F.M.Bailey; Compr. Cat. Queensland Pl. 649, 849, f. 613 (1913)

= Belvisia callifolia (Christ) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.] 192 (1947)

= Belvisia formosana (M.Ogata) Ching; Acta Phytotax. Sin. 10: 301 (1965)

= Belvisia melanesica Brownlie; Nova Hedwigia Beih. 55 (Pterid. Fl. Fiji): 369 (1977) (probably a valid species)

= Belvisia minor (Copel.) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.] 192 (1947)

= Belvisia mucronata (Fée) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.] 192 (1947)

= Belvisia vaupelii (C.Chr.) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.] 192 (1947)

= Hymenolepis callifolia Christ; Ann. Jard. Bot. Buitenz. II, 5: 128 (1905)

= Hymenolepis formosana M.Ogata; J. Jap. Bot. 11: 28, t. 1(A) (1935)

= Hymenolepis minor Copel.; Occas. Pap. Bernice Pauahi Bishop Mus. 14: 69, t. 19 (1938)

= Hymenolepis mucronata f. australiensis C.Chr.; Dansk Bot. Ark. 6(3): 64 (1929)

= Hymenolepis mucronata f. graminifolia (Rosenst.) C.Chr.; Dansk Bot. Ark. 6(3): 64 (1929)

= Hymenolepis mucronata f. helocharidioides C.Chr.; Dansk Bot. Ark. 6(3): 64 (1929)

= Hymenolepis mucronata f. latior C.Chr.; Dansk Bot. Ark. 6(3): 65 (1929)

= Hymenolepis mucronata Fée; Mém. Foug. 5 [Gen. Filic.] 82 (1851-52)

= Hymenolepis mucronata var. nigropunctata C.Chr.; Dansk Bot. Ark. 6(3): 64 (1929)

= Hymenolepis ophioglossoides Kunze; Farrnkr. 99, t. 47, f. 1 (1842) (non Kaulf.)

= Hymenolepis spicata f. longipaleacea Alderw.; Bull. Jard. Bot. Buitenzorg, sér. 2, 7: 19 (1912)

= Hymenolepis spicata f. schneideri (F.M.Bailey) Alderw.; Malay. Ferns and Fern-Allies Suppl., 432 (1917)

= Hymenolepis spicata var. bakhuizenii Alderw.; Malay. Ferns and Fern-Allies Suppl., 432, corr. 59 (1917)

= Hymenolepis spicata var. graminifolia Rosenst.; Repert. Spec. Nov. Regni Veg. 12: 530 (1913)

= Hymenolepis spicata var. novoguineensis Rosenst.; Hedwigia 56: 353 (1915)

= Hymenolepis spicata var. squamulifera Alderw.; Bull. Jard. Bot. Buitenzorg, sér. 2, 7: 19 (1912)

= Hymenolepis vaupelii Hieron. ex C.Chr.; Dansk Bot. Ark. 6(3): 65 (1929)

= Macroplethus callifolius (Christ) Tagawa; Acta Phytotax. Geobot. 11: 234 (1942)

= Macroplethus mucronatus (Fée) Tagawa; Acta Phytotax. Geobot. 11: 234 (1942)

= Macroplethus vaupelii (Hieron. ex C.Chr.) Tagawa; Acta Phytotax. Geobot. 11: 235 (1942)

species :

Lepisorus nigripes T.Fujiw. & Seriz.; J. Plant Res. (2018)

Distribution: Japan (Hokkaido, Honshu, Shikoku, Kyushu); Ryukyu Isl. (Yaku Isl.)

38 JAP NNS

Remarks: an allotetraploid species derived from L. thunbergianus and L. angustus

species :

Lepisorus normalis (D.Don) C.F.Zhao, R.Wei & X.C.Zhang; Cladistics 36(3): 252 (2019)

Distribution: China (Guangxi, Yunnan); Tibet; Japan; Nepal; Bhutan; NE-India (Arunachal Pradesh, Sikkim, Darjeeling); Myanmar (Kachin, etc.); Thailand; Laos; ?Cambodia; Vietnam; peninsular Malaysia (Pahang); Sumatra

36 CHI TIB 38 JAP 40 BHU IND NEP 41 LAO MYA THA VIE 42 MLY SUM

Remarks: sect. 5 Tricholepidium

Synonyms: hidden

= Colysis normalis (D.Don) J.Sm.; Hist. Fil. 100 (1875)

= Drynaria longifrons (Wall. ex Hook. & Grev.) J.Sm.; J. Bot. (Hooker) 3: 397 (1841)

= Drynaria normalis (D.Don) J.Sm.; J. Bot. (Hooker) 4: 61 (1841)

= Microsorum normale (D.Don) Ching; Bull. Fan Mem. Inst. Biol. Bot. 4: 295-311 (1933)

= Neocheiropteris normalis (D.Don) Tagawa; J. Jap. Bot. 27: 217 (1952)

= Neolepisorus normalis (D.Don) Ching; Bull. Fan Mem. Inst. Biol. Bot. 10: 13 (1940)

= Phymatodes normalis (D.Don) C.Presl; Tent. Pterid. 196 (1836)

= Pleopeltis longifrons (Wall. ex Hook. & Grev.) T.Moore; Index Fil. 346 (1862)

= Pleopeltis normalis (D.Don) T.Moore; Index Filic. (T. Moore) 347 (1862)

= Polypodium longifrons Wall.; Numer. List no. 274 (1829), nomen

= Polypodium longifrons Wall. ex Hook. & Grev.; Icon. Filic. 1: t. 65 (1828)

= Polypodium normale D.Don; Prodr. Fl. Nepal. 1 (1825)

= Tricholepidium mutense Ching & S.K.Wu; Acta Phytotax. Geobot. 29(1-5): 43 (1978)

= Tricholepidium normale (D.Don) Ching; Acta Phytotax. Geobot. 29(1-5): 43 (1978)

= Tricholepidium pteropodum Ching; Acta Phytotax. Geobot. 29(1-5): 43 (1978)

species :

Lepisorus novoguineensis (Rosenst.) Li Wang; Bot. J. Linn. Soc. 162(1): 35 (2010)

Distribution: Sulawesi; New Guinea (Irian Jaya, Papua New Guinea); Bismarck Arch. (New Ireland)

42 NWG SUL 43 BIS

Remarks: sect. 13 Belvisia

Synonyms: hidden

= Belvisia novoguineensis (Rosenst.) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.] 192 (1947)

= Hymenolepis novoguineensis (Rosenst.) C.Chr.; Index Filic., Suppl. Tert.: 113 (1934)

= Lemmaphyllum novoguineense (Rosenst.) C.Chr.; Dansk Bot. Ark. 6: no. 3, 51 (1929)

= Macroplethus novoguineensis (Rosenst.) Tagawa; Acta Phytotax. Geobot. 11: 234 (1942)

= Paltonium novoguineense Rosenst.; Nova Guinea 8: 729 (1912)

species :

Lepisorus nudus (Hook.) Ching; Bull. Fan Mem. Inst. Biol. Bot. 4: 83 (1933)

Distribution: China (Yunnan); Tibet; Japan; Pakistan (Hazara); Nepal; Bhutan; India (throughout, incl. Jammu & Kashmir, Uttarakhand, Himachal Pradesh, Arunachal Pradesh, Darjeeling, Sikkim); Sri Lanka; Bangladesh; Myanmar (Kachin, Chin, etc.); Thailand; Vietnam; Sumatra

36 CHI TIB 38 JAP 40 BAN BHU IND NEP PAK SRL 41 MYA THA VIE 42 SUM

Remarks: sect. 10 Lepisorus

Synonyms: hidden

= Drynaria nuda (Hook.) Fée; Mém. Foug. 5 (Gen. Filic.): 270 (1852)

= Drynaria sesquipedalis (Wall.) J.Sm.; Bot. Mag. 72: Comp. 13 (1846) (fide Fraser-Jenkins 2008)

= Drynaria zeylanica Fée; Mém. Foug. 5. (Gen. Filic.) 272 (1852)

= Lepisorus amaurolepidus var. longifolius Bir & Trikha; Amer. Fern J. 64: 63, f. 53-57 (1974) (fide Fraser-Jenkins 2008)

= Lepisorus birii Khullar; Khullar et al. 1991 (nom. nud., synonymy fide Fraser-Jenkins (1997))

= Lepisorus gyirongensis Ching & S.K.Wu; Fl. Xizangica 1: 304 (1983) (fide Fraser-Jenkins 1997)

= Lepisorus intermedius Ching & Khullar; Khullar, Ill. Fern Fl. W. Himalaya 1: 93 (1994) (fide Fraser-Jenkins 1997)

= Lepisorus leiopteris (Kunze) Bir & Trikha; Amer. Fern J. 64: 54, f. 21-26 (1974) (fide Fraser-Jenkins 2008)

= Lepisorus parvus Khullar; (1988) (nom. nud., synonymy fide Fraser-Jenkins (1997))

= Lepisorus pseudolinearis Ching & Khullar; (1988) (nom. nud., fide Fraser-Jenkins (1997))

= Lepisorus sequipedalis (Wall. ex Mett.) Fraser-Jenk.; Pakistan Syst. 5: 91 (1992), nom. superfl.

= Lepisorus tenuipes Ching & Khullar; Indian Fern J. 1(1-2): 91 (1984 publ. 1985) (fide Fraser-Jenkins 1997)

= Niphobolus sesquipedalis (Wall. ex Mett.) Keyserl.; Polyp. Herb. Bunge 39 (1873) (fide Fraser-Jenkins 2008)

= Phymatodes nuda (Hook.) J.Sm.; Bot. Voy. Herald 425 (1857)

= Pleopeltis leiopteris (Kunze) S.Singh & Panigrahi; J. Hbr. Res. 3: 18 (1990) (fide Fraser-Jenkins 2008)

= Pleopeltis linearis var. leiopteris (Kunze) C.Chr.; Index Fil. 540 (1906)

= Pleopeltis nuda Hook.; Exot. Fl. 1, t. 63 (1823)

= Pleopeltis tenuipes (Ching & Khullar) Panigrahi; Pl. Sci. Res. (India) 20: 22 (1999)

= Polypodium leiopteris Kunze; Linnaea 23: 279 319 (1859) (fide Fraser-Jenkins 2008)

= Polypodium lineare var. wightianum C.Chr.; Index Fil. 540 (1906)

= Polypodium nudiusculum Kunze; Linnaea 24: 253 (1851)

= Polypodium nudum (Hook.) Kunze; Linnaea 23: 281 (1850), nom. illeg.

= Polypodium sesquipedale f. leiopteris (Kunze) Makino; Phan. Pter. Jap. Icon. 1: t. 8 (1899)

= Polypodium sesquipedale Wall.; Numer. List no. 275 (1829), nomen

= Polypodium sesquipedale Wall. ex Mett.; Abh. Senckenb. Naturf. Ges. 2: 91 (1856)

= Polypodium wightianum Thwaites; Enum. Pl. Zeyl.: 394 (1864), nom. superfl.

= Polypodium wightianum Wall.; Numer. List no. 2222 (1829), nomen

species :

Lepisorus obscurevenulosus (Hayata) Ching; Bull. Fan Mem. Inst. Biol. Bot. 4: 76 (1933)

Distribution: China (Anhui, Chongqing, Fujian, Guangdong, Guangxi, Guizhou, Hunan, Jiangxi, Sichuan, Yunnan, Zhejiang); Taiwan; Vietnam

36 CHI 38 TAI 41 VIE

Remarks: sect. 17 Pseudovittaria. If synonymy of P. caudiceps is confirmed, this name would have priority

Synonyms: hidden

= Goniophlebium caudiceps T.Moore; Gard. Chron. n. s. 25. 234 (1886)

= Polypodium caudiceps (T.Moore) G.Nicholson; Ill. Dict. Gard. 4: 592 (1888)

= Polypodium feei var. caudiceps (T.Moore) Baker; Ann. Bot. (Oxford) 5: 469 (1891)

= Polypodium obscurevenulosum Hayata; Icon. Pl. Formosan. 6: 322, f. 134 (1915)

= Polypodium suprapunctatum Ching; Bull. Fan Mem. Inst. Biol. Bot. 4: 76 (1933)

species :

Lepisorus oligolepidus (Baker) Ching; Bull. Fan Mem. Inst. Biol. Bot. 4: 80 (1933)

Distribution: China (Anhui, Chongqing, Fujian, Guangdong, ?Guangxi, Guizhou, ?Henan, Hunan, Jiangxi, Shaanxi?, Sichuan, Yunnan, Zhejiang); ?Tibet; Japan; NE-India (Arunachal Pradesh, Manipur, Meghalaya, Tripura); Myanmar

36 CHI 38 JAP 40 IND 41 MYA

Remarks: sect. 17 Pseudovittaria. Not in India

Synonyms: hidden

= Lepisorus ellipticus Ching; Bull. Bot. Res., Harbin 2(2): 74 (1982)

= Pleopeltis oligolepida (Baker) Á.Löve & D.Löve; Taxon 26(2-3): 325 (1977)

= Polypodium lineare var. oligolepidum (Baker) Christ; Mém. Soc. Bot. France 1: 15 (1905)

= Polypodium oligolepidum Baker; Gard. Chron. n. s. 14: 594 (1880)

= Polypodium trabeculatum Copel.; Philipp. J. Sci., Bot. 3: 283 (1908)

species :

Lepisorus onoei (Franch. & Sav.) Ching; Bull. Fan Mem. Inst. Biol. Bot. 4: 87 (1933)

Distribution: South Korea (incl. Cheju Isl.); Japan; Ryukyu Isl.

38 JAP KOR NNS

Remarks: sect. 15 Obtusi

Synonyms: hidden

= Pleopeltis onoei (Franch. & Sav.) Okuyama ex Ohwi; Bull. Natl. Sci. Mus. (Tokyo), ser. 2, 3: 99 (1956)

= Polypodium lineare var. onoei (Franch. & Sav.) Makino; Bot. Mag. (Tokyo) 11: 282 (1897)

= Polypodium onoei Franch. & Sav.; Enum. Pl. Jap. 2: 246, 642 (1877)

species :

Lepisorus oosphaerus (C.Chr.) Ching; Bull. Fan Mem. Inst. Biol. Bot. 4: 73 (1933)

Distribution: N-Thailand

41 THA

Remarks: not in India

Conservation Status: VU in Thailand

Synonyms: hidden

= Pleopeltis oosphaera (C.Chr.) Panigrahi & Patnaik; Proc. Nat. Acad. Sci. India B, 34: 482 (1964)

= Polypodium oosphaerum C.Chr.; Contr. Nat. Herb. 2: 334, t. 29 (1931)

species :

Lepisorus ovatus (C.Presl) C.F.Zhao, R.Wei & X.C.Zhang; Cladistics 36(3): 251 (2019), nom. inval. (without basionym ref.)

Distribution: China (Anhui, Chongqing, Guangdong, Guangxi, Guizhou, Hubei, Hunan, Jiangsu, Jiangxi, Sichuan, Yunnan, Zhejiang); Japan; Nepal; Bhutan; India (Meghalaya, Sikkim, Arunachal Pradesh, Darjeeling); Bangladesh; Myanmar; Vietnam

36 CHI 38 JAP 40 BAN BHU IND NEP 41 MYA VIE

Remarks: sect. 3 Neolepisorus. There is confusion between the synonyms of L. ovatus and L. fortun(e)i which need to be sorted out better.

Synonyms: hidden

= Colysis ovata (C.Presl) J.Sm.; Hist. Fil. 100 (1875)

= Drynaria ovata (C.Presl) Fée; Mém. Foug. 5. (Gen. Filic.) 270 (1852)

= Microsorum ensatum var. phyllomanes (Christ) Tagawa; Acta Phytotax. Geobot. 8(3): 172 (1939)

= Microsorum ovatum (C.Presl) N.C.Nair & Bennet; Bull. Bot. Surv. India 11: 432 (1972)

= Microsorum phyllomanes (Christ) Koidz.; Acta Phytotax. Geobot. 5: 50 (1936), nom. superfl.

= Neocheiropteris ovata (C.Presl) Fraser-Jenk.; New Species Syndr. Indian Pteridol. 180 (1997), isonym

= Neocheiropteris ovata (C.Presl) B.K.Nayar & S.Kaur; Companion Handb. Ferns Brit. India: 86 (1974)

= Neocheiropteris ovata f. deltoidea (Ching) Fraser-Jenk.; Taxon. Revis. Indian Subcontinental Pteridophytes 68 (2008)

= Neocheiropteris ovata f. doryopteris (Christ) Fraser-Jenk.; Taxon. Revis. Indian Subcontinental Pteridophytes 68 (2008)

= Neocheiropteris ovatus (Wall. ex Hook. & Grev.) B.K.Nayar & Kaur; Comp. to Bedd. Handb. Ferns Brit. India 86 (1974) (comb. inval.)

= Neocheiropteris phyllomanes (Christ) Ching; Bull. Fan Mem. Inst. Biol. 4: 110 (1933), nom. superfl.

= Neocheiropteris phyllomanes f. deltoidea (Hand.-Mazz.) Ching

= Neocheiropteris phyllomanes var. doryopteris (Christ) Ching

= Neolepisorus basicordatus P.S.Wang; Acta Bot. Yunnan. 9(4): 399 (1987)

= Neolepisorus crenatus S.F.Wu; Keys Vasc. Pl. Wuling Mount. (ed. W. T. Wang et al.) 573 (1995)

= Neolepisorus cuneatus S.F.Wu; Keys Vasc. Pl. Wuling Mount. (ed. W. T. Wang et al.): 577 (1995)

= Neolepisorus dengii Ching & P.S.Wang; Acta Phytotax. Sin. 21(3): 272 (1983)

= Neolepisorus dengii f. hastatus Ching & P.S.Wang; Acta Phytotax. Sin. 21(3): 273 (1983)

= Neolepisorus emeiensis Ching & K.H.Shing; Acta Phytotax. Sin. 21(3): 271 (1983)

= Neolepisorus emeiensis f. dissectus Ching & K.H.Shing; Acta Phytotax. Sin. 21(3): 271 (1983)

= Neolepisorus lancifolius Ching & K.H.Shing; Acta Phytotax. Sin. 21(3): 271 (1983)

= Neolepisorus ovatus (C.Presl) Ching; Acta Phytotax. Sin. 9: 99 (1964)

= Neolepisorus ovatus f. deltoideus Ching; Acta Phytotax. Sin. 9: 99 (1964)

= Neolepisorus ovatus f. doryopteris (Christ) Ching; Acta Phytotax. Sin. 9: 99 (1964)

= Neolepisorus ovatus f. gracilis Ching & K.H.Shing; Acta Phytotax. Sin. 21: 269 (1983)

= Neolepisorus ovatus f. monstrosus Ching & K.H.Shing; Acta Phytotax. Sin. 21(3): 269 (1983)

= Neolepisorus ovatus f. truncatus (Ching & P.S.Wang) L.Shi & X.C.Zhang; Fl. China 2-3: 807 (2013)

= Neolepisorus phyllomanes (Christ) Ching; Bull. Fan Mem. Inst. Biol. Bot. 10: 14 (1940)

= Neolepisorus phyllomanes var. doryopteris (Christ) Ching

= Neolepisorus sinensis Ching; Acta Phytotax. Sin. 21(3): 274 (1983)

= Neolepisorus truncatus Ching & P.S.Wang; Acta Phytotax. Sin. 21(3): 270 (1983)

= Neolepisorus truncatus f. laciniatus Ching & K.H.Shing; Acta Phytotax. Sin. 21(3): 270 (1983)

= Neolepisorus tsaii Ching & K.H.Shing; Acta Phytotax. Sin. 21(3): 273 (1983)

= Phymatodes ovata C.Presl; Tent. Pterid. 197 (1836)

= Pleopeltis ovata (C.Presl) T.Moore; Index Fil. lxxviii (1857)

= Pleopeltis ovata (Wall. ex Hook. & Grev.) Bedd.; Handb. Ferns Brit. India 354 (1883), based on Polypodium ovatum Wall. ex Hook. & Grev., Icon. Filic. 1: t. 41. 1827, not Thunb. (1768)

= Polypodium hemitomum var. deltoideum (Baker) Hand.-Mazz.; Symb. Sin. 6: 44 (1929), based on Polypodium deltoideum Baker, J. Bot. 26: 230. 1880, not Sw. (1788), nor Liebmann (1849)

= Polypodium ovatum Wall. ex Hook. & Grev.; Icon. Filic. t. 41 (1827) (non Burm.f. (1768))

= Polypodium phyllomanes Christ; Bull. Acad. Geogr. Bot. Le Mans 1902. 210 c. figg. (1902)

= Polypodium phyllomanes var. doryopteris Christ; Bull. Acad. Géogr. Bot. Mans 11(153-154): 214 (1902)

= Polypodium phyllomanes var. ovatum (Wall. ex Hook. & Grev.) Christ; Bull. Acad. Int. Géogr. Bot. 11: 210 (1902), nom. inval.

species :

Lepisorus palmatopedatus (Baker) C.F.Zhao, R.Wei & X.C.Zhang; Cladistics 36(3): 253 (2019)

Distribution: China (Guizhou, Sichuan, Yunnan)

36 CHI

Remarks: sect. 6 Neocheiropteris

Conservation Status: IUCN V

Synonyms: hidden

= Cheiropteris henryi Christ; Bull. Herb. Boissier 6: 876 (1898), non Polypodium henryi (Baker) Christ 1898

= Cheiropteris palmatopedata (Baker) Christ; Bull. Herb. Boissier 7: 21, t. 1 (1899)

= Microsorum palmatopedatum (Baker) Noot.; Blumea 42(2): 343 (1997)

= Neocheiropteris palmatopedata (Baker) Christ; Bull. Soc. Bot. France 52, Mem. 1: 21 (1905)

= Polypodium palmatopedatum Baker; Bull. Misc. Inform. Kew 1898: 232 (1898)

species :

Lepisorus perrierianus (C.Chr.) Ching; Bull. Fan Mem. Inst. Biol. Bot. 4: 51 (1933)

Distribution: Madagascar

29 MDG

Remarks: sect. 8 Pachyphyllon

Synonyms: hidden

= Pleopeltis perrieriana (C.Chr.) Tardieu; Humbert, Fl. Madag. Fam. 5, 2: 112 (1960)

= Polypodium perrierianum C.Chr.; Bonap., Notes Ptéridol. 16: 193 (1925)

species :

Lepisorus platyrhynchos (Kunze) Li Wang; Bot. J. Linn. Soc. 162(1): 35 (2010)

Distribution: Philippines (Luzon, Mindoro, Agusan del Norte, Bukidnon, Davao); New Guinea (Irian Jaya, Yapen Isl.)

42 NWG PHI

Remarks: sect. 13 Belvisia

Synonyms: hidden

= Acrostichum platyrhynchos (Kunze) Hook.; Sp. 6: 280 (1864)

= Belvisia glauca (Copel.) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.] 192 (1947)

= Belvisia platyrhynchos (Kunze) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.] 192 (1947)

= Gymnopteris platyrhynchos J.Sm.; J. Bot. (Hooker) 3: 403 (1841), nomen

= Hymenolepis glauca (Copel.) C.Chr.; Dansk Bot. Ark. 6: 54-70 (1929)

= Hymenolepis platyrhynchos Kunze; Farrnkr. 1: 101 (1842)

= Hymenolepis platyrhynchos var. glauca Copel.; Leafl. Philipp. Bot. 3: 847 (1910)

= Macroplethus glaucus (Copel.) Tagawa; Acta Phytotax. Geobot. 11: 234 (1942)

= Macroplethus platyrhynchos (Kunze) C.Presl; Abh. Königl. Böhm. Ges. Wiss. 5(6) (Epim. Bot.): 142 (1851)

= Taenitis macrorhynchos Mett.; Fil. Hort. Bot. Lips. 28 (1856)

= Taenitis platyrhyncha (Kunze) Mett.; Ann. Mus. Bot. Lugduno-Batavi 4: 173 (1868)

species :

Lepisorus pseudonudus (Ching) Ching; Bull. Fan Mem. Inst. Biol. 4: 83 (1933)

Distribution: China (Yunnan, Sichuan, Gansu); Tibet

36 CHI TIB

Remarks: sect. 16 Sclerophyllon

Synonyms: hidden

= Lepisorus bilouensis Ching & Y.X.Lin ex K.H.Shing; Vasc. Pl. Hengduan Mount. 31(6): 573 (1993), without latin descr. or type, et Acta Phytotax. Sin. 1: 166 (1993)

= Pleopeltis pseudonuda (Ching) Panigrahi & Patnaik; J. Indian Bot. Soc. 43: 317 (1964)

species :

Lepisorus pseudoussuriensis Tagawa; Acta Phytotax. Geobot. 5: 110 (1936)

Distribution: Taiwan

38 TAI

Remarks: sect. 12 Ussurienses

Synonyms: hidden

= Lepisorus angustifrons Tagawa; Acta Phytotax. Geobot. 6: 110 (1936)

species :

Lepisorus pyriformis (Ching) C.F.Zhao, R.Wei & X.C.Zhang; Cladistics 36(3): 252 (2019)

Distribution: China (C-China, S-China); Japan (Honshu, Yakushima Isl.)

36 CHI 38 JAP NNS

Remarks: sect. 4 Lemmaphyllum. Often included in L. rostratus but kept separate by Zhao et al. 2019

Synonyms: hidden

= Lemmaphyllum pyriforme (Ching) Ching; Bull. Fan Mem. Inst. Biol. Bot. 4: 99 (1933)

= Lepidogrammitis pyriformis (Ching) Ching; Sunyatsenia 5: 258 (1940)

= Polypodium pyriforme Ching; Bull. Fan Mem. Inst. Biol. Bot. 2: 212, t. 29 (1931)

species :

Lepisorus rostratus (Bedd.) C.F.Zhao, R.Wei & X.C.Zhang; Cladistics 36(3): 252 (2019)

Distribution: China (Gansu, Guangdong, Guangxi, Guizhou, Hainan, Hubei, Sichuan, Yunnan, Zhejiang); S-Japan (Yakushima Isl.); Taiwan; Nepal; Bhutan; NE-India (Arunachal Pradesh, Sikkim, Darjeeling, NE-India); Myanmar (Kachin, etc.); Thailand; Laos; Cambodia; Vietnam; N-Sumatra

36 CHI 38 JAP NNS TAI 40 BHU IND NEP 41 CBD LAO MYA THA VIE 42 SUM

Remarks: sect. 4 Lemmaphyllum

Synonyms: hidden

= Lemmaphyllum carnosum var. rostratum (Bedd.) Hovenkamp; Fl. Males., Ser. 2, Pterid. 3: 76 (1998)

= Lemmaphyllum rostratum (Bedd.) Tagawa; Fl. E. Himal. 493 (1966)

= Lemmaphyllum subrostratum (C.Chr.) Ching; Bull. Fan Mem. Inst. Biol. Bot. 4: 97 (1933)

= Lepidogrammitis rostrata (Bedd.) Ching; Ching & Chu H. Wang, Acta Phytotax. Sin. 9: 372 (1964)

= Lepidogrammitis sikkimensis S.R.Ghosh; Pterid. Fl. E. India 508 (-510, fig. 154) (2004)

= Lepidogrammitis subrostrata (C.Chr.) Ching; Sunyatsenia 6: 258 (1940)

= Lepisorus rostratus (Hook.) B.K.Nayar & Kaur; Comp. to Bedd. Handb. Ferns Brit. India 84 (1974)

= Lepisorus subrostratus (C.Chr.) C.Chr. & Tardieu; Notul. Syst. (Paris) 8: 187 (1939)

= Phymatopsis rostrata (Bedd.) J.Sm.; Hist. Fil. 104 (1875)

= Pleopeltis rostrata Bedd.; Ferns Brit. India: t. 159 (1866)

= Pleopeltis subrostrata Lacaita; J. Linn. Soc., Bot. 48: 490 (1916)

= Polypodium rostratum Hook.; Hooker´s Icon. Pl. 6: t. 953 (1854), nom. illeg.

= Polypodium subrostratum C.Chr.; Index Filic. 567 (1906)

species :

Lepisorus rufofuscus T.Fujiw.; Acta Phytotax. Geobot. 73(3): 178 (2022)

Distribution: Japan (Honshu)

38 JAP

species :

Lepisorus schraderi (Mett.) Ching; Bull. Fan Mem. Inst. Biol. Bot. 4: 51 (1933)

Distribution: Rwanda; Ethiopia; Uganda; Kenya; Tanzania; Zambia; Malawi; E-Zimbabwe; Mozambique; South Africa (Limpopo, Gauteng, Mpulamanga, Free State, KwaZulu-Natal, W-Cape Prov., E-Cape Prov.); Lesotho; Eswatini [Swaziland]; Madagascar; Mauritius; La Réunion

23 RWA 24 ETH 25 KEN TAN UGA 26 MLW MOZ ZAM ZIM 27 LES SAF SWZ 29 MAU MDG REU

Remarks: sect. 8 Pachyphyllon. In South Africa L. schraderi and L. excavatus are well separated, but in East Africa the differences disappear partially.

Synonyms: hidden

= Lepisorus gueinzii (Mett.) Ching; Bull. Fan Mem. Inst. Biol. Bot. 4: 51 (1933), as ´gueintzii´

= Niphobolus schraderi (Mett.) Keyserl.; Polyp. Herb. Bunge 39 (1873)

= Phymatodes elongata (Schrad.) Pappe & Rawson; Syn. Filic. Afr. austr. 41 (1858)

= Pleopeltis schraderi (Mett.) Tardieu; Humbert, Fl. Madag. Fam. 5, 2: 110 (1960)

= Polypodium elongatum Schrad.; Gött. Gel. Anz. 918 (1818) (non Mett. (1857), nec Aiton (1789))

= Polypodium gueintzii Mett.; Abh. Senckenberg. Naturf. Ges. 2: 91 (1856)

= Polypodium lineare var. gueinzii (Mett.) C.Chr.; Index Filic. 540 (1906), as ´gueintzii´

= Polypodium lineare var. schraderi (Mett.) Sim; Ferns S. Afr. ed. 2: 276, t. 140 (1915)

= Polypodium schraderi Mett.; Abh. Senckenberg. Naturf. Ges. 2: 98, n. 179, t. 2, f. 11 (1857)

species :

Lepisorus scolopendrium (Buch.-Ham. ex Ching) Mehra & Bir; Res. Bull. (N. S.) Panjab Univ., Sci. 15: 168 (1964)

Distribution: China (Guizhou, Hainan, Sichuan, Yunnan); Tibet; N-Pakistan; Nepal; Bhutan; India (Himachal Pradesh, Uttarakhand, Sikkim, Darjeeling Hills, Jammu & Kashmir, Arunachal Pradesh); Myanmar (Chin, etc.); Thailand; Laos

36 CHI TIB 40 BHU IND NEP PAK 41 LAO MYA THA

Remarks: sect. 8 Pachyphyllon

Synonyms: hidden

= Lepisorus excavatus f. himalayensis (Bir & Trikha) Benl; Acta Bot. Barcinon. 33: 18 (1982)

= Lepisorus excavatus var. himalayensis Bir & Trikha; Amer. Fern J. 64: 58, f. 33, 37 (1974)

= Lepisorus excavatus var. mortonianus Bir & Trikha; Amer. Fern J. 64: 56, f. 25, 32 (1974)

= Lepisorus excavatus var. scolopendrium Buch.-Ham. ex Ching; Bull. Fan Mem. Inst. Biol. Bot. 4: 69 (1933) (validation of originally invalid epithet scolopendrium)

= Lepisorus himalayensis (Bir & Trikha) Khullar & Pangtey; Khullar et al., Ferns of Nainital: 165 (1991) (fide Fraser-Jenkins 2008)

= Lepisorus kumaonensis Kholia, Bhakuni, R.Punetha & Bankoti; Khuraijam (ed.), NeBIO, Imphal 3(3): 31, 34 (2012)

= Lepisorus mortonianus (Bir & Trikha) Khullar & Pangtey; Khullar et al., Ferns of Nainital: 169 (1991) (fide Fraser-Jenkins 2008)

= Lepisorus scolopendrium (Buch.-Ham. ex D.Don) Tagawa; H. Hara, Fl. E. Himal. 494 (1966), nom. superfl.

= Lepisorus scolopendrium var. himalayensis (Bir & Trikha) Pangtey & Punetha; Western Himalaya 2: 399 (1987)

= Lepisorus virescens Ching & S.K.Wu; Fl. Xizangica 1: 306 (1983)

= Pleopeltis excavata var. himalayensis (Bir & Trikha) Benl; Acta Bot. Barcinon. 33: 18 (1982)

= Pleopeltis excavata var. mortoniana (Bir & Trikha) Benl; Acta Bot. Barcinon. 33: 18 (1982)

= Pleopeltis himalayensis (Bir & Trikha) Á.Löve & D.Löve; Taxon 26: 324 (1977), nom. illeg.

= Pleopeltis mortonianus (Bir & Trikha) Á.Löve & D.Löve; Taxon 26: 324 (1977)

= Pleopeltis scolopendrium (Buch.-Ham. ex Ching) Alston & Bonner; Candollea 15: 207 (1956)

= Polypodium scolopendrium Buch.-Ham. ex D.Don; Prodr. Fl. Nepal. 1 (1825), non P. scolopendria Burm.f. (1768)

species :

Lepisorus sinensis (Christ) Ching; Bull. Fan Mem. Inst. Biol. Bot. 4: 63 (1933)

Distribution: China (SE-Yunnan); ?Bhutan; Myanmar; N-Thailand; Vietnam

36 CHI 41 MYA THA VIE

Remarks: sect. 17 Pseudovittaria

Synonyms: hidden

= Lemmaphyllum sinense (Christ) C.Chr.; Dansk Bot. Ark. 6: no. 3, 51 (1929)

= Lepisorus vittarioides Ching; Bull. Fan Mem. Inst. Biol. Bot. 4: 64 (1933)

= Neurodium sinense Christ; Bull. Herb. Boissier 6: 880 (1898)

= Paltonium sinense (Christ) C.Chr.; Index Filic. 477 (1906)

= Pleopeltis sinensis (Christ) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.] 184 (1947)

= Polypodium neurodioides C.Chr.; Contr. U.S. Natl. Herb. 26: 318 (1931)

= Polypodium neurodioides var. vittarioides (Ching) C.Chr.; Index Filic., Suppl. Tert.: 118, 154 (1934)

species :

Lepisorus sordidus (C.Chr.) Ching; Bull. Fan Mem. Inst. Biol. Bot. 4: 78 (1933)

Distribution: China (Yunnan, Sichuan); Tibet; India (Arunachal Pradesh, Nagaland, Manipur); Myanmar

36 CHI TIB 40 IND 41 MYA

Remarks: sect. 17 Pseudovittaria

Conservation Status: IUCN VU for India (Fraser-Jenkins 2012)

Synonyms: hidden

= Lepisorus sordidus f. rostratus Ching; Bull. Fan Mem. Inst. Biol. 4(3): 79 (1933)

= Pleopeltis sordida (C.Chr.) Panigrahi & Patnaik; Curr. Sci. 34: 127 (1965)

= Polypodium sordidum C.Chr.; Contr. U.S. Natl. Herb. 26: 320 (1931)

species :

Lepisorus soulieanus (Christ) Ching & S.K.Wu; Acta Bot. Yunnan. 5(1): 11 (1983), et Fl. Xizangica 1: 307 (1983)

Distribution: China; etc.

36 CHI

Remarks: sect. 9 Hymenophyton. Separate from L. clathratus fide Zhao et al. 2019. Distribution uncertain because this was for a long time included in L. clathratus.

Synonyms: hidden

= Polypodium soulieanum Christ; Bull. Soc. Bot. France 52, Mem. 1: 15 (1905)

= Platygyria soulieana (Christ) X.C.Zhang & Q.R.Liu; Acta Phytotax. Sin. 41(5): 407 (2003)

species :

Lepisorus spicatus (L.f.) Li Wang; Bot. J. Linn. Soc. 162(1): 35 (2010)

Distribution: Guinea; Liberia; Ivory Coast; Cameroon; Bioko Isl. [Fernando Poo]; São Tomé; Gabon; Congo [Brazzaville]; D.R.Congo [Zaire]; Tanzania; Malawi; Zimbabwe; Mozambique; South Africa; Madagascar; Comores; La Réunion; Seychelles; Sri Lanka; Myanmar (Kachin, etc.); SW-Thailand; Laos; Vietnam; peninsular Malaysia (Kedah, Langkawi Isl., Perak, Negeri Sembilan, Kelantan, Pahang); Sumatra; Borneo (Mt. Kinabalu, Sabah, Kalimantan); Sulawesi; Palawan; Philippines; Java; Lesser Sunda Isl. (Bali, Lombok, etc.); Moluccas; New Guinea (Irian Jaya, Papua New Guinea); Solomon Isl. (Guadalcanal); New Caledonia; Vanuatu; Fiji; Society Isl. (Tahiti, Moorea); Austral Isl. (Tubuai)

22 GUI IVO LBR 23 BIO CMN CON GAB SAO ZAI 25 TAN 26 MLW MOZ ZIM 27 SAF 29 COM MDG REU SEY 40 SRL 41 LAO MYA THA VIE 42 BOR JAW LSI MLY MOL NWG PHI PLW SUL SUM 43 SOL 60 FIJ NWC VAN 61 AUI SCI

Remarks: sect. 13 Belvisia

Synonyms: hidden

= Acrostichum spicatum L.f.; Suppl. 444 (1781)

= Belvisia revoluta (Blume) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.] 192 (1947)

= Belvisia spicata (L.f.) Mirb.; Hist. Nat. Veg. (Lam. & Mirb.) 3: 473 (1802), et ex Copel., Gen. Fil. 192 (1947)

= Gymnopteris ophioglossoides (Kaulf.) Paxton; Pocket Bot. Dict. ed. 2. Suppl.: 33 (1849)

= Gymnopteris spicata (L.f.) C.Presl; Tent. Pterid. 244, t. 11, f. 7 (1836)

= Hyalolepis ophioglossoides (Kaulf.) Kunze; Linnaea 28: 258 (1850)

= Hyalolepis revoluta (Blume) Kunze; Linnaea 23: 258 (1850)

= Hyalolepis revoluta var. planiuscula Kunze; (1850)

= Hymenolepis ophioglossoides Kaulf.; Enum. Filic. 146, t. 1, f. 9 (1824)

= Hymenolepis revoluta Blume; Enum. Pl. Javae 2: 201 (1828)

= Hymenolepis revoluta var. planiuscula (Mett.) Hieron. ex C.Chr.; Hedwigia 57: 58 (1916)

= Hymenolepis rigidissima Christ; Bull. Herb. Boissier, sér, 2, 6: 990 (1906)

= Hymenolepis spicata (L.f.) C.Presl; Abh. Königl. Böhm. Ges. Wiss. 5(6) (Epim. Bot.): 159 (1851)

= Hymenolepis spicata f. minima Alderw.; (1917)

= Hymenolepis spicata var. costulata Alderw.; (1912)

= Hymenolepis spicata var. occultivenia C.Chr.; Dansk Bot. Arch. 6(3): 57 (1929)

= Hymenolepis spicata var. tenella Alderw.; (1908)

= Hymenolepis spicata var. usambarensis C.Chr.; Dansk Bot. Arch. 6(3): 57 (1929)

= Lomaria spicata (L.f.) Willd.; Sp. Pl. ed. 4, 5: 289 (1810)

= Macroplethus revolutus (Blume) Tagawa; Acta Phytotax. Geobot. 11: 234 (1942)

= Macroplethus spicatus (L.f.) Tagawa; Acta Phytotax. Geobot. 11: 235 (1942)

= Oetosis ophioglossoides (Kaulf.) Kuntze; Revis. Gen. Pl. 2: 817 (1891)

= Onoclea spicata (L.f.) Sw.; J. Bot. (Schrader) 1801(2): 299 (1803)

= Schizaea spicata (L.f.) Sm.; Mém. Acad. Roy. Sci. (Turin) 5: 43 (1793)

= Taenitis ophioglossoides (Kaulf.) Mett.; Fil. Hort. Bot. Lips. 28 (1856)

= Taenitis revoluta (Blume) Mett.; Fil. Hort. Bot. Lips. 28 (1856)

= Taenitis revoluta var. planiuscula Mett.; Fil. Hort. Bot. Lips. 28 (1856)

= Taenitis spicata (L.f.) Mett.; Ann. Mus. Bot. Lugduno-Batavi 4: 173 (1868)

species :

Lepisorus squamatus (A.R.Sm. & X.C.Zhang) C.F.Zhao, R.Wei & X.C.Zhang; Cladistics 36(3): 252 (2019)

Distribution: China (Guangxi); N-Vietnam

36 CHI 41 VIE

Remarks: sect. 4 Lemmaphyllum

Synonyms: hidden

= Caobangia squamata A.R.Sm. & X.C.Zhang; Novon 12(4): 549 (2002)

= Lemmaphyllum squamatum (A.R.Sm. & X.C.Zhang) Li Wang; Bot. J. Linn. Soc. 162(1): 36 (2010)

species :

Lepisorus subhemionitideus (Christ) C.F.Zhao, R.Wei & X.C.Zhang; Cladistics 36(3): 253 (2019)

Distribution: China (Guangxi, Hunan, Sichuan, Yunnan); Tibet; Nepal; Bhutan; N-India; Myanmar; N-Vietnam; S-Vietnam

36 CHI TIB 40 BHU IND NEP 41 MYA VIE

Remarks: sect. 7 Lepidomicrosorium. Not in Taiwan

Synonyms: hidden

= Lepidomicrosorium caudifrons Ching & W.M.Chu; Bot. Res. Acad. Sin. 1983(1): 10 (1983)

= Lepidomicrosorium crenatum Ching & K.H.Shing; Bot. Res. Acad. Sin. 1: 11 (1983)

= Lepidomicrosorium emeiense Ching & K.H.Shing; Bot. Res. Acad. Sin. 1983(1): 10 (1983)

= Lepidomicrosorium hongchunpingense Ching & K.H.Shing; Bot. Res. Acad. Sin. 1983(1): 9 (1983)

= Lepidomicrosorium hongchunpingense var. laceratum Ching & K.H.Shing; Bot. Res. Acad. Sin. 1983(1): 9 (1983)

= Lepidomicrosorium hunanense Ching & K.H.Shing; Bot. Res. Acad. Sin. 1983(1): 6 (1983)

= Lepidomicrosorium laojunense Ching & K.H.Shing; Bot. Res. Acad. Sin. 1983(1): 10 (1983)

= Lepidomicrosorium lineare Ching & P.S.Chiu; Bot. Res. Acad. Sin. 1983(1): 5 (1983)

= Lepidomicrosorium longshengense Ching & K.H.Shing; Bot. Res. Acad. Sin. 1983(1): 9 (1983)

= Lepidomicrosorium subhemionitideum (Christ) P.S.Wang; P. S. Wang & X. Y. Wang Pterid. Fl. Guizhou 382 (2001)

= Lepidomicrosorium undulatum Ching & P.S.Chiu; Bot. Res. Acad. Sin. 1983(1): 11 (1983)

= Leptochilus subhemionitideus (Christ) Bosman; Leiden Bot. Ser. 14: 117 (1991)

= Microsorum hymenodes var. marginale (Takeda) Ching; Bull. Fan Mem. Inst. Biol. Bot. 4(10): 301 (1933)

= Microsorum jinfoshanense Ching & Z.Y.Liu; Bull. Bot. Res., Harbin 3(4): 12 (1983)

= Microsorum rubripes Ching & W.M.Chu; Bull. Bot. Res., Harbin 3(4): 11 (1983)

= Neocheiropteris sapaensis V.N.Tu; Bot. Zhurn. (Moscow & Leningrad) 65(4): 584 (1980)

= Neocheiropteris subhemionitidea (Christ) Fraser-Jenk.; New Species Syndr. Indian Pteridol. 179 (1997)

= Polypodium hymenodes var. marginale Takeda; Notes Roy. Bot. Gard. Edinburgh 8(39): 288 (1915)

= Polypodium subhemionitideum Christ; Bull. Herb. Boissier 7: 5 (1809)

= Polypodium superficiale var. attenuatum Rosenst.; Repert. Spec. Nov. Regni Veg. 13(355-358): 134 (1914)

= Polypodium superficiale var. semilineare C.B.Clarke; Trans. Linn. Soc. London, Bot. 1: 558 (1880)

= Tricholepidium subhemionitideum (Christ) Fraser-Jenk.; Taxon. Revis. Indian Subcontinental Pteridophytes 69 (2008)

species :

Lepisorus sublinearis (Baker ex Takeda) Ching; Bull. Fan Mem. Inst. Biol. Bot. 4: 78 (1933)

Distribution: China (Yunnan); Tibet; Nepal; Bhutan; N-India (Arunachal Pradesh, Sikkim, Darjeeling, Uttarakhand, NE-India); Myanmar (Kachin, Chin, etc.); N-Thailand; Laos; Vietnam

36 CHI TIB 40 BHU IND NEP 41 LAO MYA THA VIE

Remarks: sect. 16 Sclerophyllon

Synonyms: hidden

= Lepisorus cespitosus Y.X.Lin; Fl. Reipubl. Popul. Sin. 6(2): 77 (2000)

= Lepisorus lancifolius Ching; Bull. Fan Mem. Inst. Biol. Bot. 11: 75 (1941)

= Pleopeltis sublinearis (Baker ex Takeda) Tagawa & K.Iwats.; Acta Phytotax. Geobot. 22: 100 (1967)

= Polypodium sublineare Baker ex Takeda; Notes Roy. Bot. Gard. Edinburgh 8: 276 (1915)

species :

Lepisorus suboligolepidus Ching; Bull. Fan Mem. Inst. Biol. Bot. 4: 77 (1933)

Distribution: China (Hubei, Sichuan, Yunnan); Taiwan; NE-India (Meghalaya, Manipur, Tripura, Arunachal Pradesh); Myanmar; NE-Thailand

36 CHI 38 TAI 40 IND 41 MYA THA

Remarks: sect. 17 Pseudovittaria. Sometimes included in L. oligolepidus

Synonyms: hidden

= Polypodium suboligolepidum (Ching) C.Chr.; Index Filic., Suppl. Tert.: 159 (1934)

= Pleopeltis suboligolepida (Ching) Tagawa & K.Iwats.; Acta Phytotax. Geobot. 26: 172 (1975)

species :

Lepisorus subsessilis Ching & Y.X.Lin; Fl. Reipubl. Popul. Sin. 6(2): 77 (2000)

Distribution: China (Guangdong, Guangxi)

36 CHI

Remarks: sect. 10 Lepisorus

species :

Lepisorus superficialis (Blume) C.F.Zhao, R.Wei & X.C.Zhang; Cladistics 36(3): 253 (2019)

Distribution: China (Anhui, Fujian, Guangdong, Guangxi, Guizhou, Hubei, Hunan, Jiangxi, Sichuan, Yunnan, Zhejiang); Tibet; South Korea (incl. Cheju Isl.); Japan; Bonin Isl. (Hahajima); Taiwan; Bhutan; NE-India; Myanmar (Kachin, etc.); Thailand; ?Laos; Vietnam; peninsular Malaysia (Pahang); Sumatra; Sulawesi; Java; Moluccas

36 CHI TIB 38 JAP KOR OGA TAI 40 BHU IND 41 MYA THA VIE 42 JAW MLY MOL SUL SUM

Remarks: sect. 7 Lepidomicrosorium. L. ningpoense is probably a separate species fide Kuo et al. (2019)

Synonyms: hidden

= Colysis superficialis (Blume) J.Sm.; Hist. Fil. 101 (1875)

= Lepidomicrosorium angustifolium Ching & K.H.Shing; Bot. Res. Acad. Sin. 1983(1): 8 (1983)

= Lepidomicrosorium nanchuanense Ching & Z.Y.Liu; Bull. Bot. Res., Harbin 4(4): 27 (1984)

= Lepidomicrosorium ningpoense (Baker) L.Y.Kuo; Taiwania 64(4): 378 (2019)

= Lepidomicrosorium sichuanense Ching & K.H.Shing; Bot. Res. Acad. Sin. 1983(1): 8 (1983)

= Lepidomicrosorium superficiale (Blume) Li Wang; Bot. J. Linn. Soc. 162(1): 36 (2010)

= Microsorum brachylepis (Baker) Nakaike; Sa. Kurata & Nakaike, Illustr. Pterid. Japan 2: 492 (1981)

= Microsorum buergerianum f. laciniatum Ching; Bull. Fan Mem. Inst. Biol. Bot. 4(10): 303 (1933)

= Microsorum masaskei (Nakai) H.Itô; J. Jap. Bot. 11: 96 (1935)

= Microsorum ovalifolium Ching & S.K.Wu; Fl. Xizangica 1: 327 (1983) (fide Fraser-Jenkins 2008)

= Microsorum simulans Ching & Z.Y.Liu; Bull. Bot. Res., Harbin 4(4): 26 (1984)

= Microsorum superficiale (Blume) Ching; Bull. Fan Mem. Inst. Biol. Bot. 4: 295-311 (1933)

= Microsorum takhtajanii V.N.Tu; Bot. Zhurn. (Moscow & Leningrad) 64(12): 1769 (1979)

= Microsorum tibeticum (Ching & S.K.Wu) Ching & S.K.Wu; Fl. Xizangica 1: 328 (1983) (fide Fraser-Jenkins 2008)

= Neocheiropteris ningpoensis (Baker) Bosman; Leiden Bot. Ser. 14: 119 (1991)

= Neocheiropteris superficialis (Blume) Bosman; Leiden Bot. Ser. 14: 121 (1991)

= Phymatodes masaskei Nakai; Bot. Mag. (Tokyo) 43: 4 (1929)

= Pleopeltis superficialis (Blume) Bedd.; Ferns Brit. India: t. 75 (1865)

= Pleopeltis superficialis var. latifrons Bedd.; J. Bot. 31: 226 (1893)

= Polypodium brachylepis Baker; Gard. Chron. n. s. 14: 494 (1880)

= Polypodium buergerianum var. ningpoense (Baker) Takeda; Notes Roy. Bot. Gard. Edinburgh 8(39): 291 (1915)

= Polypodium masaskei (Nakai) Ogata; Icon. Filic. Jap. 8, t. 138 (1930)

= Polypodium nigrocinctum Christ; Bull. Herb. Boissier 6: 874 (1896)

= Polypodium ningpoense Baker; Ann. Bot. (Oxford) 5: 474 (1891)

= Polypodium superficiale Blume; Enum. Pl. Javae 2: 123 (1828)

= Polypodium superficiale var. anguinum Christ; Bull. Soc. Bot. France 52(Mem. 1): 16 (1905)

= Tricholepidium superficiale (Blume) Fraser-Jenk.; Taxon. Revis. Indian Subcontinental Pteridophytes 69 (2008)

= Tricholepidium superficiale (Blume) Fraser-Jenk., Kandel & Pariyar; Ferns and Fern-allies of Nepal 1: 43 (2015), isonym (isonym)

species :

Lepisorus thunbergianus (Kaulf.) Ching; Bull. Fan Mem. Inst. Biol. Bot. 4: 88 (1933), nom. cons. prop.

Distribution: Russian Far East (Kuril Isl.); China (Anhui, Fujian, Gansu, Guizhou, Hebei, Hubei, Hunan, Jiangsu, Jiangxi, Shaanxi, Sichuan, Yunnan, Zhejiang); Tibet; South Korea (Cheju Isl.); Japan; Ryukyu Isl.; Taiwan; N-Pakistan; Nepal; Bhutan; N-India (Arunachal Pradesh, Sikkim, Uttarakhand, Himachal Pradesh, Jammu & Kashmir); Myanmar; Laos; Vietnam; Philippines (N-Luzon); Hawaii (Kauai, Oahu, Molokai, Lanai, Maui, Hawaii Isl.)

31 RFE 36 CHI TIB 38 JAP KOR NNS TAI 40 BHU IND NEP PAK 41 LAO MYA VIE 42 PHI 63 HAW

Remarks: sect. 17 Pseudovittaria

Synonyms: hidden

= Drynaria elongata (Kaulf.) Brack.; U.S. Expl. Exped., Filic. 16: 42 (1854)

= Drynaria subspathulata Hook.; Hooker´s J. Bot. Kew Gard. Misc. 9: 356 (1857)

= Lepisorus atropunctatus (Keyserl.) Ching; (1933)

= Lepisorus calcifer Ching & Z.Y.Liu; Bull. Bot. Res., Harbin 4(4): 25 (1984)

= Lepisorus chingii Khullar; (1988) (nom. nud., fide Fraser-Jenkins (1997))

= Lepisorus elongatus Ching; Bull. Fan Mem. Inst. Biol. Bot. 4: 89 (1933), nom. rej. prop.

= Lepisorus linearifolius Ching & Z.Y.Liu; Bull. Bot. Res., Harbin 3(4): 5 (1983)

= Lepisorus linearis (Thunb.) Ching; (nom. illeg.)

= Lepisorus mussooriensis Khullar; (1988) (nom. nud., fide Fraser-Jenkins (1997))

= Lepisorus myriosorus Ching; Wuyi Sci. J. 1: 11 (1981)

= Lepisorus pygmaeus Ching & Z.Y.Liu; Bull. Bot. Res., Harbin 4(4): 24 (1984)

= Lepisorus simulans Ching & Z.Y.Liu; Bull. Bot. Res., Harbin 4(4): 23 (1984), et Phytotaxa 205(4): 299 (2015)

= Lepisorus stewartii Ching; Acta Bot. Austro-Sin. 1: 23 (1983) (fide Fraser-Jenkins 1997)

= Lepisorus thunbergianus var. subspathulatus (Hook.) Ching; Bull. Fan Mem. Inst. Biol. Bot. 4(3): 89 (1933)

= Lepisorus thunbergianus var. tsutaei Akasawa & E.Yokote; Bull. Kochi Women´s Univ., Ser. Nat. Sci. 27: 4 (1979)

= Microsorum lineare (Thunb.) Copel.; Univ. Calif. Publ. Bot. 16: 111-116 (1929)

= Niphobolus atropunctatus Keyserl.; Polyp. Herb. Bunge 39 (1873)

= Niphobolus linearis (Thunb.) Keyserl.; Polyp. Herb. Bunge 39 (1873)

= Phymatodes elongata (Kaulf.) C.Presl; Tent. Pterid. 196 (1836)

= Pleopeltis elongata Kaulf.; Enum. Filic. 246 (1824), nom. rej. prop.

= Pleopeltis linearis (Thunb.) T.Moore; Index Filic. (T. Moore) 346 (1862)

= Pleopeltis linearis Bedd.; Handb. Ferns Brit. India: 346 (1883), nom. illeg.

= Pleopeltis subspathulata (Hook.) T.Moore; Index Filic. (T. Moore) 348 (1862)

= Pleopeltis thunbergiana Kaulf.; Wes. d. Farrnkr. 113 (1827), based on Polypodium lineare Thunb., Syst. Veg., ed. 14: 934. 1784, not N. F. Burman (1768), nom. cons. prop.

= Pleopeltis wightiana Bedd.; Ferns S. India: t. 180 (1863), nom. superfl.

= Pleopeltis zeylanica T.Moore; Index Filic. (T. Moore) 349 (1862)

= Polypodium coraiense Christ; Repert. Spec. Nov. Regni Veg. 6: 285 (1908)

= Polypodium elongatum Goldm.; Nov. Actorum Acad. Caes. Leop.-Carol. Nat. Cur. 19(Suppl. 1): 454 (1843), nom. illeg.

= Polypodium gladiatum Wall.; Numer. List no. 279 (1829), nomen

= Polypodium lineare Thunb.; Fl. Jap. 335 (1784) (non Burm.f. (1768) nec Houtt. (1783)), et Linn. Syst. Veg. ed. 14: 934 (1784)

= Polypodium lineare var. subspathulatum (Hook.) Takeda; Notes Roy. Bot. Gard. Edinburgh 8(39): 270-271 (1915)

= Polypodium lineare var. thunbergianum (Kaulf.) Takeda; Notes Roy. Bot. Gard. Edinburgh 8(39): 268 (1915)

= Polypodium loriforme Hook.; Gard. Ferns: t. 14 (1862)

= Polypodium thunbergianum (Kaulf.) C.Chr.; Index Filic., Suppl. Tert.: 160 (1934)

species :

Lepisorus tibeticus Ching & S.K.Wu; Fl. Xizangica 1. 311 (1983)

Distribution: China (Sichuan, Yunnan); Tibet

36 CHI TIB

Remarks: sect. 17 Pseudovittaria. Included in L. thunbergianus by Fraser-Jenkins et al. 2015 but kept separate on genetic basis by Zhao et al. 2019

Synonyms: hidden

= Pleopeltis tibetica (Ching & S.K.Wu) Panigrahi; Pl. Sci. Res. (India), Abstr. 23rd Ann. Conf. Khurda: 36 (1998)

species :

Lepisorus tosaensis (Makino) H.Itô; J. Jap. Bot. 11: 93 (1935)

Distribution: China (Anhui, Chongqing, Guangdong, Guangxi, Guizhou, Hunan, Jiangsu, Jiangxi, Sichuan, Xinjiang, Yunnan, Zhejiang); Tibet; Japan; Taiwan; Vietnam

36 CHI TIB 38 JAP TAI 41 VIE

Remarks: sect. 17 Pseudovittaria

Synonyms: hidden

= Lepisorus infraplanicostalus (Hayata) Ching; Bull. Fan Mem. Inst. Biol. Bot. 4(3): 92 (1933)

= Lepisorus paohuashanensis Ching; Rec. Fl. Jiangsu (Fl. Kiang Su Prov.) 1: 467 (1977)

= Lepisorus sinicus Ching & Z.Y.Liu; Bull. Bot. Res., Harbin 3(4): 2 (1983)

= Lepisorus tosaensis f. bifidus Akasawa; Bull. Kochi Women´s Univ., Ser. Nat. Sci. 27: 4 (-5) (1979)

= Pleopeltis tosaensis (Makino) Ohwi; Bull. Natl. Sci. Mus. (Tokyo), ser. 2, 3: 99 (1956)

= Polypodium hoozanense Hayata; Icon. Pl. Formosan. 8: 152, f. 81-82 (1919)

= Polypodium infra-planicostale Hayata; Icon. Pl. Formosan. 6: 31 5, f. 128 (1915)

= Polypodium lineare var. caudatum Makino; Bot. Mag. (Tokyo) 17: 78 (1903)

= Polypodium morii Hayata; Icon. Pl. Formosan. 8: 153, f. 83-84 (1919)

= Polypodium tosaense Makino; Bot. Mag. (Tokyo) 27. 127 (1913)

species :

Lepisorus tricholepis K.H.Shing & Y.X.Lin; Fl. Reipubl. Popul. Sin. 6(2): 70 (2000)

Distribution: Tibet; Bhutan

36 TIB 40 BHU

Remarks: sect. 9 Hymenophyton

species :

Lepisorus triglossus (Baker) C.F.Zhao, R.Wei & X.C.Zhang; Cladistics 36(3): 253 (2019)

Distribution: China (Yunnan)

36 CHI

Remarks: sect. 3 Neolepisorus. Possibly a hybrid L. palmatopedatus x macrosphaerus

Conservation Status: very rare and possibly extinct

Synonyms: hidden

= Neocheiropteris triglossa (Baker) Ching; Bull. Fan Mem. Inst. Biol. Bot. 4: 108-112 (1933)

= Neolepisorus triglossus (Baker) Ching; Bull. Fan Mem. Inst. Biol. Bot. 10: 14 (1940)

= Polypodium triglossum Baker; Bull. Misc. Inform. Kew 1898: 232 (1898)

= Selliguea triphylla Christ; Bull. Herb. Boissier 6: 878 (1898)

species :

Lepisorus uchiyamae (Makino) H.Itô; J. Jap. Bot. 11: 93 (1935)

Distribution: Japan (Honshu, Shikoku, Kyushu); Ryukyu Isl.

38 JAP NNS

Remarks: sect. 12 Ussurienses

Synonyms: hidden

= Pleopeltis uchiyamae (Makino) Ohwi; Bull. Natl. Sci. Mus. (Tokyo), ser. 2, 3: 99 (1956)

= Polypodium uchiyamae Makino; Bot. Mag. (Tokyo) 20: 30 (1906)

species :

Lepisorus ussuriensis (Regel & Mack.) Ching; Bull. Fan Mem. Inst. Biol. Bot. 4: 91 (1933)

Distribution: Russian Far East (Kuril Isl., Primorye, Sakhalin); China (Anhui, Hebei, Heilongjiang, Henan, Jilin, Liaoning, Shandong); North Korea; South Korea (incl. Cheju Isl.)

31 RFE 36 CHI 38 KOR NKO

Remarks: sect. 12 Ussurienses. Not in India

Synonyms: hidden

= Pleopeltis ussuriensis Regel & Maack; Mém. Acad. Imp. Sci. St. Pétersb. 7: 44-175 (1861)

= Polypodium lineare var. ussuriense (Regel & Maack) C.Chr.; Index Filic. 9: 540 (1906)

= Polypodium lineare var. ussuriense (Regel & Maack) Christ; Index Filic. 540 (1906)

= Polypodium schraderi Milde; Fil. Eur.: 16 (1867), nom. illeg.

= Polypodium ussuriense (Regel & Maack) Regel; Acta Hort. Petrop. 7: 663 (1881)

species :

Lepisorus validinervis (Kunze) Li Wang; Bot. J. Linn. Soc. 162(1): 36 (2010)

Distribution: peninsular Malaysia (Pahang); Sumatra; Borneo (Sabah); Sulawesi; Philippines; Java; Lesser Sunda Isl. (Lombok); Moluccas (Seram); New Guinea (Irian Jaya, Papua New Guinea); Bismarck Arch. (New Ireland); Vanuatu

42 BOR JAW LSI MLY MOL NWG PHI SUL SUM 43 BIS 60 VAN

Remarks: sect. 13 Belvisia

Synonyms: hidden

= Acrostichum brachystachys (Hook.) Racib.; Pteridoph. Buitenzorg 51 (1898)

= Belvisia longissima Holttum; Blumea 14: 328 (1967)

= Belvisia squamata (Hieron. ex C.Chr.) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.] 192 (1947)

= Belvisia squamata var. borneensis (C.Chr.) Parris; Pl. Mt. Kinabalu [J. H. Beaman et al.] 98 (1992)

= Belvisia validinervis (Kunze) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.] 192 (1947)

= Belvisia validinervis var. longissima (Holttum) Hovenkamp & Franken; Blumea 37(2): 527 (1993)

= Hymenolepis brachystachys (Hook.) J.Sm.; Ferns Brit. For. 92 (1866)

= Hymenolepis brachystachys var. mirabilis Alderw.; (1918)

= Hymenolepis spicata var. brachystachys Hook.; Gard. Ferns, t. 3 (1862)

= Hymenolepis squamata Hieron.; C. Chr., Dansk Bot. Ark. 6: 54-70 (1929)

= Hymenolepis squamata var. borneensis (C.Chr.) Parris; Parris et al., Pl. Mt. Kinabalu, 1. Ferns & Fern-Allies: 98 (1992)

= Hymenolepis validinervis Kunze; Bot. Zeitung (Berlin) 6. 132 (1848)

= Macroplethus squamatus (Hieron. ex C.Chr.) Tagawa; Acta Phytotax. Geobot. 11 235 (1942)

= Macroplethus validinervis (Kunze) Tagawa; Acta Phytotax. Geobot. 11: 235 (1942)

= Taenitis validinervis (Kunze) Mett.; Ann. Mus. Bot. Lugduno-Batavi 4: 173, t. 7 (1868)

species :

Lepisorus waltonii (Ching) S.L.Yu; Acta Phytotax. Sin. 35: 343 (1997)

Distribution: Tibet

36 TIB

Remarks: sect. 9 Hymenophyton. Separate from L. clathratus fide Zhao et al. 2019

Synonyms: hidden

= Lepisorus clathratus f. waltonii (Ching) Fraser-Jenk.; Taxon. Revis. Indian Subcontinental Pteridophytes 40 (2008)

= Neocheiropteris waltonii Ching; Hooker´s Icon. Pl. t. 3158 (1932)

= Phymatopteris waltonii (Ching) Fraser-Jenk.; New Species Syndr. Indian Pteridol. 196 (1997)

= Platygyria waltonii (Ching) Ching & S.K.Wu; Acta Bot. Yunnan. 2(1): 71 (1980)

= Platygyria ×inaequibasis Ching & S.K.Wu; Acta Bot. Yunnan. 2(1): 72 (1980)

= Polypodium clathratum var. lobatum Takeda; Notes Roy. Bot. Gard. Edinburgh 8(39): 282 (1915)

= xEllipisorus inaequibasis (Ching & S.K.Wu) Li Bing Zhang & Liang Zhang; Phytotaxa 464(4): 299 (2020)

= xLepinema inaequibasis (Ching & S.K.Wu) Li Bing Zhang & Liang Zhang; Phytotaxa 455(4): 265 (2020)

species :

Lepisorus yamaokae Seriz.; Shidekobushi 3(1): 57 (2015)

Distribution: Japan (Honshu, Shikoku, Kyushu)

38 JAP

species :

Lepisorus youxingii R.H.Jiang & X.C.Zhang; Phytotaxa 437(2): 102 (2020)

Distribution: China (Guangxi)

36 CHI

species :

Lepisorus zippelii (Blume) C.F.Zhao, R.Wei & X.C.Zhang; Cladistics 36(3): 251 (2019)

Distribution: China (Guangdong, Guangxi, Guizhou, Hainan, Yunnan); Bhutan; India (N-India, Himachal Pradesh, Uttarakhand, Sikkim, Darjeeling, Meghalaya); Myanmar (Kachin, etc.); Thailand; Laos; Cambodia; Vietnam; peninsular Malaysia (Pahang, Perak, Selangor, Negeri Sembilan); Sumatra; Borneo (Mt. Kinabalu, Sabah); Sulawesi; Philippines (Luzon); Java; Lesser Sunda Isl. (Lombok); New Guinea (Papua New Guinea)

36 CHI 40 BHU IND 41 CBD LAO MYA THA VIE 42 BOR JAW LSI MLY NWG PHI SUL SUM

Remarks: sect. 3 Neolepisorus

Synonyms: hidden

= Bathmium subfalcatum (J.Sm.) Fée; Mém. Foug. 5 (Gen. Filic.): 287 (1852)

= Colysis zippelii (Blume) J.Sm.; Hist. Fil. 101 (1875)

= Drynaria subfalcata J.Sm.; J. Bot. (Hooker) 3: 397 (1841)

= Microsorum luzonicum (Copel.) Tagawa; Acta Phytotax. Geobot. 16: 51 (1955)

= Microsorum zippelii (Blume) Ching; Bull. Fan Mem. Inst. Biol. Bot. 4: 308 (1933)

= Neocheiropteris zippelii (Blume) Bosman; Leiden Bot. Ser. 14: 123 (1991)

= Neolepisorus zippelii (Blume) Li Wang; Bot. J. Linn. Soc. 162(1): 36 (2010)

= Pleopeltis heterocarpa Bedd.; Handb. Ferns Brit. India, t. 319 (1869), nom. illeg.

= Pleopeltis luzonica (Copel.) Alderw.; Bull. Dép. Agric. Indes Néerl. 27: 7 (1909)

= Pleopeltis zippelii (Blume) T.Moore; Index Filic. (T. Moore) 308 (1862)

= Polypodium heterocarpum var. zippelii (Blume) Baker; Hook. & Baker, Syn. Fil. 360 (1868)

= Polypodium luzonicum Copel.; Philipp. J. Sci. 1, Suppl. II: 162, t. 23 (1906), not C. Presl (1825)

= Polypodium oxyphyllum Kunze; Bot. Zeitung (Berlin) 116 (1848)

= Polypodium zippelii Blume; Enum. Pl. Javae 2: 172, t. 80 (1828)

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A complete, synonymic checklist of the Higher Plants of the World

Version 19.2 (April 17th, 2024)

Covering 354,696 accepted species,
8,644 hybrids, 47,758 accepted infraspecific taxa and 996,379 synonyms,
in total 1,407,477* names

[about 25,000 dubious, "unplaced" names and about 4,000 invalid names (Art. 24.3) like "typicus*" are deliberately omitted]

With complete, detailed distribution data (countries and regions) and literature references

Arranged in linear phylogenetic sequence reflecting true relationships

Fully searchable and continously updated

With 38,290 pictures of 15,532 species

Assembled through 40 years independently from other databases

In cooperation with Catalogue of Life

Contact

How to Cite:

Search

+ Family

Filter

Order

Family (28)

Genus (70)

Display options

A complete, synonymic checklist of the Higher Plants of the World

Version 19.2 (April 17th, 2024)

Covering 354,696 accepted species, 8,644 hybrids, 47,758 accepted infraspecific taxa and 996,379 synonyms, in total 1,407,477* names

[*about 25,000 dubious, "unplaced" names and about 4,000 invalid names (Art. 24.3) like "typicus" are deliberately omitted]

With complete, detailed distribution data (countries and regions) and literature references

Arranged in linear phylogenetic sequence reflecting true relationships

Fully searchable and continously updated

With 38,290 pictures of 15,532 species

Assembled through 40 years independently from other databases

In cooperation with Catalogue of Life

Covering 354,696 accepted species,
8,644 hybrids, 47,758 accepted infraspecific taxa and 996,379 synonyms,
in total 1,407,477* names

[about 25,000 dubious, "unplaced" names and about 4,000 invalid names (Art. 24.3) like "typicus*" are deliberately omitted]