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An Illustrated Checklist of the Flora of Qatar - Unesco

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UNESCO OFFICE IN DOHA<br />

<strong>An</strong> <strong>Illustrated</strong> <strong>Checklist</strong><br />

<strong>of</strong> <strong>the</strong> <strong>Flora</strong> <strong>of</strong> <strong>Qatar</strong><br />

John Norton, Sara Abdul Majid, Debbie Allan,<br />

Mohammed Al Safran, Benno Böer & Renee Richer


<strong>An</strong> <strong>Illustrated</strong> <strong>Checklist</strong><br />

<strong>of</strong> <strong>the</strong> <strong>Flora</strong> <strong>of</strong> <strong>Qatar</strong><br />

John Norton, Sara Abdul Majid, Debbie Allan,<br />

Mohammed Al Safran, Benno Böer & Renee Richer<br />

2009<br />

UNESCO OFFICE IN DOHA


First published in 2009 by Browndown Publications, Gosport, UK.<br />

ISBN 978-0-9563961-0-5<br />

Text copyright © J.A. Norton, S. Abdul Majid, D.R. Allan, M. Al Safran, B. Böer & R. Richer, 2009.<br />

Photographs copyright © J.A. Norton, except Plates 45 & 66 © S. Abdul Majid; 3, 17, 19, 20 & 109<br />

© S.J. Aspinall; 1, 4, 8, 11, 14, 28, 31, 32, 34, 35, 37, 40, 46, 51, 65, 67, 69 & 77 © R. Richer; 2, 64<br />

& 74 © M. Al Safran.<br />

All rights reserved. No part <strong>of</strong> this publication may be reproduced without prior permission <strong>of</strong> <strong>the</strong><br />

publisher.<br />

The designations employed and <strong>the</strong> presentation <strong>of</strong> <strong>the</strong> material throughout this publication do not<br />

imply <strong>the</strong> expression <strong>of</strong> any opinion whatsoever on <strong>the</strong> part <strong>of</strong> UNESCO concerning legal status <strong>of</strong><br />

any country, territory, city or area <strong>of</strong> its authorities, or concerning <strong>the</strong> delimitation <strong>of</strong> its frontiers<br />

or boundaries. The views expressed in this publication are those <strong>of</strong> <strong>the</strong> respective authors and do<br />

not necessarily reflect <strong>the</strong> views <strong>of</strong> UNESCO.<br />

The production <strong>of</strong> this book was generously supported by UNESCO Doha, Maersk Oil <strong>Qatar</strong> and<br />

<strong>Qatar</strong> Foundation.<br />

The authors would be pleased to receive comments or corrections on <strong>the</strong> text, fur<strong>the</strong>r information<br />

on <strong>the</strong> occurrence <strong>of</strong> species in <strong>Qatar</strong> and good quality digital photographs <strong>of</strong> plants growing in<br />

<strong>Qatar</strong>. Correspondence should be addressed to: John Norton, Browndown Publications, 215 Forton<br />

Road, Gosport, Hampshire PO12 3HB, UK (e-mail: books@jnecology.com)<br />

Printed by Ashford Colour Press Ltd, Gosport, UK.<br />

Cover photograph: a sandy wadi with Acacia tortilis and Lycium shawii, within an area <strong>of</strong> gently undulating<br />

limestone ‘hamada’ – <strong>the</strong> characteristic landscape <strong>of</strong> <strong>Qatar</strong> (south-west <strong>Qatar</strong>, March 2007). Insets: Convolvulus<br />

cephalodpodus, <strong>An</strong>astatica hierochuntica, Centaurea sinaica and Cynomorium coccineum.<br />

Contents<br />

Authors ii<br />

Foreword iii<br />

Preface iv<br />

Acknowledgements v<br />

Introduction vi<br />

Selection <strong>of</strong> Species vi<br />

Sources <strong>of</strong> Information vi<br />

Overview <strong>of</strong> Habitats and Vegetation vii<br />

Notes on Layout and Terminology x<br />

Map <strong>of</strong> <strong>the</strong> State <strong>of</strong> <strong>Qatar</strong> xiv<br />

SPECIES ACCOUNTS 1-84<br />

Pteridophyta 1<br />

Gymnospermae 1<br />

<strong>An</strong>giospermae: Dicotyledoneae 1<br />

<strong>An</strong>giospermae: Monocotyledoneae 69<br />

References and Bibliography 85<br />

Internet Resources 88<br />

Index to Families and Species 89


Authors<br />

John Norton is an independent ecological consultant and botanist, based in Hampshire, UK.<br />

He has spent more than two and a half years in total working in <strong>the</strong> Arabian Peninsula, including<br />

spells in nor<strong>the</strong>rn Saudi Arabia and <strong>the</strong> United Arab Emirates to conduct research on <strong>the</strong> Houbara<br />

Bustard. Since 1998 he has made several trips to <strong>Qatar</strong> to undertake wildlife surveys and impact<br />

assessments, including a detailed survey <strong>of</strong> <strong>the</strong> Dukhan area in 2007. He has a keen interest in <strong>the</strong><br />

wildlife and ecology <strong>of</strong> desert habitats and has built up an extensive collection <strong>of</strong> photographs and<br />

herbarium specimens. In <strong>the</strong> UK he is a committee member <strong>of</strong> <strong>the</strong> Hampshire <strong>Flora</strong> Group, which<br />

organises field meetings, and encourage people to develop botanical identification skills.<br />

Benno Böer is a botanist and ecologist working at <strong>the</strong> UNESCO Doha Office as <strong>the</strong> Ecological<br />

Sciences Advisor for <strong>the</strong> Arab Region. He has been involved in several international research<br />

programmes in ecosystem study, vegetation mapping and o<strong>the</strong>r forms <strong>of</strong> botanical research in<br />

Saudi Arabia, <strong>Qatar</strong>, Bahrain, Oman, Kuwait and <strong>the</strong> United Arab Emirates and has authored<br />

or contributed to several scientific publications and books including <strong>the</strong> two most recent floras<br />

covering <strong>the</strong> UAE (Jongbloed, et al. 2000, 2003). He has a special interest in <strong>the</strong> rehabilitation <strong>of</strong><br />

coastal and desert ecosystems and through UNESCO helped to set up <strong>the</strong> first biosphere reserve<br />

for <strong>the</strong> country at Al Reem in 2007 and is helping to establish <strong>the</strong> Quranic Botanic Garden in Doha.<br />

Sara Abdul Majid is an environmental scientist and dedicated conservationist. Born and raised in<br />

<strong>Qatar</strong>, she has developed a passion for <strong>the</strong> unique desert environment <strong>of</strong> <strong>the</strong> country. She worked at<br />

an environmental consulting firm (URS Doha <strong>of</strong>fice) before pursuing her Masters in Conservation<br />

at University College London. During her time with URS, Sara has been involved in conducting<br />

terrestrial and marine field surveys including <strong>the</strong> study <strong>of</strong> baseline flora, fauna, soil/sediment and<br />

groundwater conditions. She has recently conducted research at Al Reem Biosphere Reserve,<br />

including <strong>the</strong> characterization and mapping <strong>of</strong> vegetation communities, in collaboration with<br />

UNESCO and <strong>the</strong> Ministry <strong>of</strong> Environment (<strong>Qatar</strong>).<br />

Debbie Allan is a self-taught botanist and naturalist who has studied <strong>the</strong> flora <strong>of</strong> her local area <strong>of</strong><br />

Gosport in Hampshire for more than 15 years. Toge<strong>the</strong>r with John Norton she is working on a<br />

flora <strong>of</strong> <strong>the</strong> area, which lies on a floristically rich coastal peninsula, to be published in 2010. Past<br />

work has included five years spent indexing scientific publications and 15 months (to April 2006)<br />

computerising botanical records for <strong>the</strong> Hampshire Rare Plant Register, a project funded by <strong>the</strong><br />

Botanical Society <strong>of</strong> <strong>the</strong> British Isles and <strong>the</strong> Hampshire Wildlife Trust. She presently works as an<br />

assistant ecologist with John Norton.<br />

Mohammed Al Safran was born in <strong>Qatar</strong> and has developed a special interest in its flora and plant<br />

ecology. He studied at <strong>Qatar</strong> University and subsequently obtained a Masters degree in Agricultural<br />

and Environmental Sciences at Newcastle University in 2008 and is presently completing his<br />

PhD on <strong>the</strong> ecology and status <strong>of</strong> Prosopis cineraria and Acacia tortilis in <strong>Qatar</strong>. He is also currently<br />

teaching at <strong>Qatar</strong> University Department <strong>of</strong> Biology & Environmental Sciences and is supervising<br />

<strong>the</strong> management <strong>of</strong> <strong>the</strong> ‘Biology Field’ at <strong>the</strong> university campus, helping to involve students in <strong>the</strong><br />

study <strong>of</strong> flora, fauna and soil relationships.<br />

Renee Richer is visiting assistant pr<strong>of</strong>essor at Weill Cornell Medical College in <strong>Qatar</strong>. Trained as a<br />

botanist and ecologist at Harvard University, she worked in sou<strong>the</strong>rn Africa from 1994-2002. While<br />

assistant pr<strong>of</strong>essor and director <strong>of</strong> <strong>the</strong> Environmental Conservation and Research Center at <strong>the</strong><br />

American University <strong>of</strong> Armenia, she worked closely with government, NGOs and communities in<br />

fur<strong>the</strong>ring environmental development in <strong>the</strong> country, culminating in <strong>the</strong> Whitley award in 2007.<br />

With a keen interest in environmental science and sustainable development in arid regions, she<br />

has authored scientific publications and co-authored a book on environmental course design. She<br />

is currently studying <strong>the</strong> flora <strong>of</strong> <strong>Qatar</strong>.<br />

Foreword<br />

This illustrated plant species <strong>Checklist</strong> <strong>of</strong> <strong>the</strong> <strong>Flora</strong> <strong>of</strong> <strong>Qatar</strong> has been produced in <strong>the</strong><br />

overall framework <strong>of</strong> a major project linking biodiversity conservation with cultural<br />

heritage, namely <strong>the</strong> Quranic Botanic Garden (QBG) in <strong>Qatar</strong>. The guidelines for <strong>the</strong> QBG<br />

have been developed under <strong>the</strong> guidance <strong>of</strong> <strong>Qatar</strong> Foundation and with <strong>the</strong> international<br />

and inter-disciplinary support and expert advice <strong>of</strong> Mr. Abdulaziz Al Midfa (United Arab<br />

Emirates), Pr<strong>of</strong>. Dr. Abdulrahman Al Muftah (<strong>Qatar</strong>), Pr<strong>of</strong>. Dr. Kamal Batanouny (<strong>Qatar</strong><br />

Foundation), Dr. Benno Böer (UNESCO), Dr. Gary Brown (Kuwait), Dr. Uwe Herpin<br />

(Brazil), Pr<strong>of</strong>. Dr. Fareed Krupp (Germany), Dr. Guido Licciardi (UNESCO) and Ms.<br />

Narjes Yedder (Tunisia). They have developed <strong>the</strong> over-arching vision to ‘Provide <strong>the</strong> people<br />

<strong>of</strong> <strong>Qatar</strong> and <strong>the</strong> Gulf Region with a Centre <strong>of</strong> Excellence for Research and Education, promoting<br />

<strong>the</strong> conservation <strong>of</strong> natural and cultural heritage, emphasising <strong>the</strong> teachings <strong>of</strong> <strong>the</strong> Holy Qur’an and<br />

<strong>the</strong> Hadith. This will also function as a prime recreational destination’. I am highly thankful to<br />

<strong>the</strong> above experts.<br />

A lot has been achieved in a short time, since October 2007, when UNESCO and <strong>Qatar</strong><br />

Foundation embarked on this joint initiative, which received generous support from<br />

Maersk Oil <strong>Qatar</strong>, and which was assisted by Ralf Oliver <strong>of</strong> London and Botanic Gardens<br />

Conservation International. The International Advisory Committee met, guidelines were<br />

developed based on expert recommendations, <strong>the</strong> master plan was developed, reviewed,<br />

revised and produced, and an external evaluation report was compiled. These were all<br />

essential steps towards <strong>the</strong> establishment <strong>of</strong> <strong>the</strong> Quranic Botanic Garden.<br />

The flora <strong>of</strong> <strong>Qatar</strong> and <strong>the</strong> Arabian Peninsula has always been important for <strong>the</strong> Arab<br />

people, as a source for livestock grazing, construction material, firewood and food. It is<br />

important in our modern times that we study <strong>the</strong> natural vegetation, learn to understand<br />

vegetation ecology, and document and conserve our national and regional flora. The<br />

indigenous plants have undoubtedly a large number <strong>of</strong> medicinal, pharmaceutical,<br />

landscaping, scientific and economic values that are yet to be discovered. This volume is a<br />

new and essential contribution, because and it contains a comprehensive listing <strong>of</strong> species<br />

that occur in <strong>the</strong> country, as well as providing important eco-geographical information<br />

about <strong>the</strong>se species.<br />

It is with great joy that I can witness and provide <strong>the</strong> foreword for this milestone as a<br />

supporting element <strong>of</strong> <strong>the</strong> QBG. During <strong>the</strong>ir research <strong>the</strong> authors faced many deficits in<br />

<strong>the</strong> existing literature, namely a lack <strong>of</strong> reliable, accurate and up-to-date botanical data.<br />

I congratulate <strong>the</strong>m for <strong>the</strong>ir achievement, including those elements <strong>of</strong> real capacity<br />

building. I highly recommend this volume as an important source <strong>of</strong> information about,<br />

and education into, <strong>the</strong> understanding <strong>of</strong> <strong>the</strong> flora <strong>of</strong> <strong>Qatar</strong>.<br />

Saif Al Hajari<br />

Vice-Chairperson <strong>of</strong> <strong>Qatar</strong> Foundation<br />

ii iii


Preface<br />

My interest in Arabian plant life started in 1988 when I travelled to <strong>the</strong> Harrat al Harrah area in<br />

north-west Saudi Arabia on <strong>the</strong> first <strong>of</strong> five trips over a period <strong>of</strong> two years to study <strong>the</strong> habitat <strong>of</strong><br />

<strong>the</strong> Houbara Bustard, a traditional quarry <strong>of</strong> <strong>the</strong> Arab falconer. Although a dry, rocky area, <strong>the</strong><br />

flora here is relatively rich, especially in <strong>the</strong> sandy wadis and silt pans which retain some moisture<br />

during <strong>the</strong> summer months. My work <strong>the</strong>n took me to Abu Dhabi during <strong>the</strong> winter <strong>of</strong> 1991–92,<br />

where although working as an ornithologist I visited all parts <strong>of</strong> <strong>the</strong> Emirate and quickly became<br />

familiar with <strong>the</strong> range <strong>of</strong> plant life <strong>of</strong> this predominantly sandy area. Travels to <strong>the</strong> Al Liwa oasis,<br />

at <strong>the</strong> edge <strong>of</strong> <strong>the</strong> Empty Quarter, were particularly memorable during this time.<br />

In November 1994 I returned to Abu Dhabi to work for six months for one <strong>of</strong> <strong>the</strong> predecessors <strong>of</strong><br />

<strong>the</strong> Abu Dhabi Environment Agency (EAD), where I carried out both bird and plant-related work.<br />

This is where I first met Benno Böer, who had already been employed as an ecologist <strong>the</strong>re for<br />

several years – having previously spent time in <strong>the</strong> Eastern Province <strong>of</strong> Saudi Arabia as well as Abu<br />

Dhabi, studying <strong>the</strong> ecology <strong>of</strong> mangroves, saltmarshes and desert vegetation. During <strong>the</strong> spring <strong>of</strong><br />

1995 a good rainy spell allowed us <strong>the</strong> opportunity to study <strong>the</strong> germination and development <strong>of</strong><br />

native desert plants, particularly in <strong>the</strong> confines <strong>of</strong> our research station at Sweihan, a 1.5 km x 1.5 km<br />

square enclosure (Böer & Norton 1996a, 1996b). Whilst <strong>the</strong> flora inside <strong>the</strong> station flourished, <strong>the</strong><br />

plants outside soon disappeared due to <strong>the</strong> attentions <strong>of</strong> <strong>the</strong> local herd <strong>of</strong> camels. Encouragingly,<br />

we also noticed that parts <strong>of</strong> an adjacent area had been fenced <strong>of</strong>f to prevent grazing. This may<br />

have been a measure to protect <strong>the</strong> wintering grounds <strong>of</strong> <strong>the</strong> Houbara, or could have been a now<br />

rare example <strong>of</strong> <strong>the</strong> traditional Bedouin hema system <strong>of</strong> grazing protection.<br />

From this background, we subsequently both went on to develop a keen interest in <strong>the</strong> habitats and<br />

flora <strong>of</strong> <strong>the</strong> region and <strong>of</strong> <strong>Qatar</strong> in particular. After publishing an important account <strong>of</strong> his Abu<br />

Dhabi work on <strong>the</strong> anthropogenic impacts on desert ecosystems (Böer 1999), Benno moved to Doha<br />

to work for UNESCO, where he has gone on to promote <strong>the</strong> restoration and wise management <strong>of</strong><br />

desert habitats. Since 2002 he has been working to set up <strong>the</strong> first biosphere reserve for <strong>the</strong> country<br />

at Al Reem, which was finally established in 2007. He was also instrumental in establishing <strong>the</strong><br />

Quranic Botanic Garden in Doha.<br />

I carried on working in <strong>the</strong> UK as an ecological consultant, but in 1998, 2003 and 2004 I made<br />

brief visits to <strong>Qatar</strong> to carry out ecological surveys for environmental impact assessments at <strong>the</strong><br />

two main industrial ‘cities’ <strong>of</strong> Mesaieed and Ras Laffan. More recently, during February and March<br />

2007 I organised a large-scale flora and fauna survey <strong>of</strong> <strong>Qatar</strong> Petroleum’s Dukhan Concession<br />

Area, on <strong>the</strong> west coast. This followed <strong>the</strong> wettest winter for 10 years in <strong>the</strong> Gulf, which resulted<br />

in remarkable green swa<strong>the</strong>s <strong>of</strong> annual vegetation across large parts <strong>of</strong> <strong>Qatar</strong>. I was joined on this<br />

survey by Simon Aspinall, an old friend and <strong>the</strong>n long-term former resident <strong>of</strong> Abu Dhabi, who<br />

besides being a renowned ornithologist is also one <strong>of</strong> <strong>the</strong> most experienced botanists working in<br />

<strong>the</strong> region. He has contributed several <strong>of</strong> <strong>the</strong> photographs included in this book and made more<br />

than his fair share <strong>of</strong> noteworthy discoveries.<br />

The flora <strong>of</strong> <strong>the</strong> Arabian Peninsula has principally been studied and documented by visiting or<br />

expatriate Egyptian, Pakistani, Sudanese and western botanists, most notably Abdel Bari, Boulos,<br />

Batanouny, Chaudhary, Collenette, Jongbloed and Mandaville. Although some <strong>of</strong> <strong>the</strong>se have<br />

investigated <strong>the</strong> flora in a pr<strong>of</strong>essional role through specific research programmes, o<strong>the</strong>rs have<br />

pursued <strong>the</strong>ir studies in <strong>the</strong>ir own time in an amateur capacity, particularly <strong>the</strong> Western botanists.<br />

In <strong>Qatar</strong>, most <strong>of</strong> <strong>the</strong> research has been carried out through <strong>the</strong> University <strong>of</strong> <strong>Qatar</strong>, which was<br />

founded in 1977. Pr<strong>of</strong>essor Batanouny published <strong>the</strong> first and only definitive account <strong>of</strong> <strong>the</strong> flora<br />

in his Ecology and <strong>Flora</strong> <strong>of</strong> <strong>Qatar</strong> soon after in 1981. This was followed by a short, but useful account<br />

<strong>of</strong> <strong>the</strong> flora by El Amin in Wild Plants <strong>of</strong> <strong>Qatar</strong>, published by <strong>the</strong> Arab Organization for Agricultural<br />

Development in 1983, <strong>the</strong> result <strong>of</strong> a 14-month collecting trip. No fur<strong>the</strong>r comprehensive works<br />

on <strong>the</strong> flora <strong>of</strong> <strong>Qatar</strong> have been published since this time, apart from a few journal papers dealing<br />

largely with saline habitats and a detailed report by <strong>the</strong> University on vegetation types and soil<br />

relationships (Abulfatih et al. 2001).<br />

Unfortunately, <strong>the</strong> scientific value <strong>of</strong> <strong>the</strong> books by Batanouny and El Amin is now diminishing<br />

because plant names have changed and new species have been discovered. The flora has changed<br />

in many areas due to increased grazing pressure, <strong>the</strong> recreational use <strong>of</strong> desert areas and particularly<br />

through <strong>the</strong> increase in irrigation schemes in farms, towns and cities. Benno and I realised that a<br />

new flora <strong>of</strong> <strong>Qatar</strong> was urgently needed, but despite our reasonable familiarity with <strong>the</strong> country we<br />

also knew that nei<strong>the</strong>r <strong>of</strong> us had enough knowledge to be able to write a full scientific flora. This will<br />

hopefully follow in a few years time after we (or o<strong>the</strong>rs) have carried out more detailed research and<br />

have visited some <strong>of</strong> <strong>the</strong> lesser-explored areas. In <strong>the</strong> meantime, we have compiled this checklist<br />

with <strong>the</strong> intention <strong>of</strong> providing an up-to-date scientific list <strong>of</strong> <strong>the</strong> plants that are known to occur,<br />

including brief details where known <strong>of</strong> <strong>the</strong>ir status, distribution, habitat and uses.<br />

We were lucky to have found two local <strong>Qatar</strong>is, Sara Abdul Majid and Mohammed Al Safran, both<br />

with a keen interest in <strong>the</strong> country’s natural heritage, to assist us in compiling this book. Renee<br />

Richer who is currently resident in <strong>Qatar</strong> also provided useful information on <strong>the</strong> distribution<br />

<strong>of</strong> species and medicinal uses. The remaining author, Debbie Allan, shared with me much <strong>of</strong> <strong>the</strong><br />

general research and compilation <strong>of</strong> information from o<strong>the</strong>r floras. Whilst <strong>the</strong> checklist is not<br />

intended to be an identification guide and does not contain any keys or detailed descriptions, we<br />

hope that it will provide a useful source <strong>of</strong> information for ecologists and o<strong>the</strong>r scientists carrying<br />

out surveys and o<strong>the</strong>r research. However, our main aim in publishing it is to stimulate interest and<br />

understanding <strong>of</strong> <strong>the</strong> natural habitats, vegetation and flora <strong>of</strong> <strong>Qatar</strong>, which in contrast to <strong>the</strong> rest<br />

<strong>of</strong> <strong>the</strong> Gulf states have been poorly studied and under-valued.<br />

The production <strong>of</strong> <strong>the</strong> checklist has been kindly funded by UNESCO and will be used to promote<br />

and inform <strong>the</strong>ir Quranic Botanic Gardens project, being run in conjunction with <strong>the</strong> <strong>Qatar</strong><br />

Foundation and supported by Maersk Oil <strong>Qatar</strong>. The aims <strong>of</strong> <strong>the</strong> project include <strong>the</strong> establishment<br />

<strong>of</strong> a living collection <strong>of</strong> all native plant species occurring in <strong>the</strong> State <strong>of</strong> <strong>Qatar</strong>, as well as plants from<br />

<strong>the</strong> Arabian Peninsula and those mentioned in <strong>the</strong> Holy Quran and <strong>the</strong> Hadith. It also aims to<br />

build up a knowledge-base <strong>of</strong> plant taxonomy and vegetation ecology amongst <strong>Qatar</strong>i botanists. In<br />

this regard we have great hopes that young <strong>Qatar</strong>i men and women will discover <strong>the</strong> beauty <strong>of</strong> <strong>the</strong><br />

national flora, develop a passion for wildlife and come to understand <strong>the</strong> importance <strong>of</strong> botanical<br />

and ecological scientific research into <strong>the</strong> natural world.<br />

John Norton<br />

Gosport, UK, October 2009<br />

Acknowledgements<br />

The authors would like to thank Her Highness Sheikha Mozah bint Nasser al Misned, UNESCO<br />

Special Envoy for Higher and Basic Education for commitment towards botanical education in<br />

<strong>Qatar</strong>. We also express our thanks to Dr Saif Al Hajiri, Vice-Chairperson <strong>of</strong> <strong>Qatar</strong> Foundation<br />

for his interest. Fur<strong>the</strong>rmore, we sincerely thank Jacob Thomasen, Saad Al Mohammadi, Peter<br />

Christensen and Maersk Oil <strong>Qatar</strong> for <strong>the</strong>ir support, making this publication possible. We also<br />

express our thanks to UNESCO Representative in <strong>the</strong> Arab States <strong>of</strong> <strong>the</strong> Gulf and Director <strong>of</strong> <strong>the</strong><br />

Doha Office, Dr Hamed Al-Hammami, and all <strong>of</strong> his staff, for <strong>the</strong>ir support.<br />

We are also indebted to Eric Clement (Gosport, UK) for providing advice and help on identification<br />

and taxonomic issues, for pro<strong>of</strong>-reading <strong>the</strong> text and making many useful suggestions, and also for<br />

checking plant specimens and for <strong>the</strong> extended loan <strong>of</strong> books on <strong>the</strong> Arabian flora. Thanks are also<br />

due to Simon Aspinall (Norfolk, UK) for permission to reproduce photographs.<br />

iv v


Selection <strong>of</strong> Species<br />

Introduction<br />

This publication lists all species <strong>of</strong> vascular plants (flowering plants and ferns) recorded growing<br />

in <strong>the</strong> wild in <strong>the</strong> State <strong>of</strong> <strong>Qatar</strong>. These comprise all species known to be native, but also many that<br />

have been introduced intentionally or accidentally by man and are now naturalised, <strong>of</strong>ten as weeds<br />

<strong>of</strong> cultivation. Several commonly-planted species are also included that are not yet known to be<br />

widely naturalised, but could become so in <strong>the</strong> future. They include a few salt-tolerant trees and<br />

shrubs that are becoming increasingly important for urban landscaping and ornamental use in<br />

<strong>the</strong> Gulf. Such species are widely planted to provide shelter from wind and sun, visual screening <strong>of</strong><br />

industrial buildings and for <strong>the</strong>ir general aes<strong>the</strong>tic value in ‘greening’ projects. The total number<br />

<strong>of</strong> species included in <strong>the</strong> checklist is nearly 400 <strong>of</strong> which about 270 are likely to be truly native.<br />

Naturalised species are <strong>of</strong> interest to botanists and ecologists as <strong>the</strong>y <strong>of</strong>ten become established very<br />

quickly when introduced into suitable habitats in places away from <strong>the</strong>ir country <strong>of</strong> origin. Thus<br />

in some cases <strong>the</strong>y may compete with native species for water and o<strong>the</strong>r resources. It is important<br />

in publications such as this to document <strong>the</strong>se species and publicise <strong>the</strong>ir presence so that <strong>the</strong>ir<br />

status can be monitored. In <strong>Qatar</strong> and elsewhere in <strong>the</strong> Gulf one tree in particular, <strong>the</strong> Mesquite<br />

Prosopis juliflora, originally from central America, has now become an undesirable invasive species.<br />

Although primarily found in and around inhabited areas, it is increasingly seen growing in more<br />

remote desert localities. In contrast, <strong>the</strong> native Prosopis cineraria or ghaf, which is on <strong>the</strong> edge <strong>of</strong><br />

its range in <strong>Qatar</strong>, is undergoing a serious decline. The vast majority <strong>of</strong> <strong>the</strong> naturalised species<br />

included in this checklist are annual or perennial ‘mesophytic’ plants that are dependent on<br />

artificial irrigation. Many, however are also found in <strong>the</strong> numerous natural depressions in <strong>Qatar</strong>,<br />

particularly after winter rains, when soils may remain damp for a considerable time. It is prudent<br />

to monitor such species to see if <strong>the</strong>y are having any impact on <strong>the</strong> native flora, which also abound<br />

in <strong>the</strong>se depressions.<br />

Batanouny and El Amin visited cultivated areas as part <strong>of</strong> <strong>the</strong>ir research and so recorded <strong>the</strong>se<br />

naturalised and ‘escaped’ species in some detail. Regrettably, <strong>the</strong> present authors have not been<br />

able to revisit many farm areas; instead we have concentrated out efforts on natural habitats. We<br />

have <strong>the</strong>refore not been able to confirm <strong>the</strong> current status <strong>of</strong> many <strong>of</strong> <strong>the</strong> species mentioned in<br />

<strong>the</strong> earlier publications. Several species recorded in <strong>Qatar</strong> have not been fully confirmed through<br />

expert determination <strong>of</strong> specimens. The two published volumes <strong>of</strong> <strong>the</strong> <strong>Flora</strong> <strong>of</strong> <strong>the</strong> Arabian<br />

Peninsula and Socotra (Miller & Cope 1996, Cope 2007) only list species as occurring in <strong>Qatar</strong> (and<br />

o<strong>the</strong>r countries) where <strong>the</strong> authors have seen specimens. In this checklist unconfirmed species have<br />

been noted in <strong>the</strong> text and include, for example, those recorded by El Amin but not previously by<br />

Batanouny, and a few old records by Obeid (1975). Where <strong>the</strong> identification is thought very likely<br />

to have been in error <strong>the</strong> species name has been shown in square brackets.<br />

Sources <strong>of</strong> Information<br />

This checklist is partly a syn<strong>the</strong>sis <strong>of</strong> historically published information, especially from <strong>the</strong> previous<br />

accounts <strong>of</strong> <strong>the</strong> <strong>Qatar</strong> flora by Batanouny (1981) and El Amin (1983), as already mentioned. It was<br />

felt important to summarise <strong>the</strong> information included by <strong>the</strong>se authors, as <strong>the</strong>ir works include much<br />

useful data on status, distribution and uses <strong>of</strong> species. More recent information was obtained from<br />

a list <strong>of</strong> newly recorded species for <strong>Qatar</strong> published by Abdel Bari (1997). O<strong>the</strong>r principal sources<br />

include <strong>the</strong> aforementioned Miller & Cope 1996 and Cope 2007, along with Boulos’s <strong>Flora</strong> <strong>of</strong> Egypt<br />

(Boulos 1999-2005), Jongbloed’s Wild Flowers <strong>of</strong> <strong>the</strong> UAE (Jongbloed et al. 2003) and Mandaville’s<br />

<strong>Flora</strong> <strong>of</strong> Eastern Saudi Arabia (1990). O<strong>the</strong>r recent floras covering Gulf states were also consulted,<br />

including those for Bahrain: Phillips (1988) and Cornes & Cornes (1989) and for Kuwait: Daoud<br />

& Al-Rawi (1985) and Shuaib (1995). Specific comments made by <strong>the</strong>se authors have been cited in<br />

<strong>the</strong> text, but to save space <strong>the</strong> dates <strong>of</strong> publication have not usually been repeated.<br />

Much information for this checklist has, however, been derived from <strong>the</strong> authors’ knowledge<br />

and experience <strong>of</strong> <strong>the</strong> flora both in <strong>Qatar</strong> and o<strong>the</strong>r parts <strong>of</strong> <strong>the</strong> Arabian Gulf. None<strong>the</strong>less, <strong>the</strong><br />

occurrence and distribution <strong>of</strong> many species over much <strong>of</strong> <strong>Qatar</strong> is poorly known and so some <strong>of</strong><br />

our remarks are ei<strong>the</strong>r brief or generalised. Fur<strong>the</strong>r research and a mapping survey <strong>of</strong> <strong>the</strong> whole<br />

country is badly needed. The least explored parts <strong>of</strong> <strong>the</strong> state include <strong>the</strong> extreme north coast, <strong>the</strong><br />

sou<strong>the</strong>rn borderlands with Saudi Arabia and <strong>the</strong> <strong>of</strong>fshore islands that lie within <strong>Qatar</strong>i territory. It<br />

is probable that such areas could support a number <strong>of</strong> hi<strong>the</strong>rto unrecorded native species. It is also<br />

likely that a large number <strong>of</strong> non-native species will be added to <strong>the</strong> list in future years as more are<br />

introduced through importation <strong>of</strong> food, forestry plants and livestock. Fur<strong>the</strong>rmore, little is known<br />

about <strong>the</strong> species <strong>of</strong> permanently wet areas, such as ponds and lagoons created through irrigation<br />

run-<strong>of</strong>f and water treatment schemes.<br />

Photographs and line-drawings are a valuable means <strong>of</strong> checking <strong>the</strong> identity <strong>of</strong> plant species, so<br />

in addition to providing a selection <strong>of</strong> our own photographs in this checklist, we have also carefully<br />

referenced published photographs and illustrations from <strong>the</strong> floras and o<strong>the</strong>r guides covering <strong>the</strong><br />

Gulf region. In addition to those mentioned above, <strong>the</strong>se comprise Western (1989) and <strong>the</strong> two most<br />

recent publications covering <strong>the</strong> Sultanate <strong>of</strong> Oman (Ghazanfar 2003, Pickering & Patzelt 2008),<br />

which contain a small but useful selection <strong>of</strong> plants found in <strong>Qatar</strong>.<br />

Much <strong>of</strong> <strong>the</strong> time spent compiling <strong>the</strong> checklist was taken up with checking taxonomy and recent<br />

changes to nomenclature. Names <strong>of</strong> plants unfortunately change constantly as new information<br />

from field-work or herbarium research is published, particularly now that DNA studies are widely<br />

carried out. We have tried to use <strong>the</strong> most up to date name available, if we believe <strong>the</strong> change has<br />

been or will be accepted by <strong>the</strong> scientific community. In some cases <strong>the</strong> use <strong>of</strong> a scientific name <strong>of</strong><br />

a plant species comes down to personal choice. Therefore we have largely followed Miller & Cope<br />

(1996), Cope (2007), Jongbloed et al. (2003) and Boulos (1999-2005); but have also made use <strong>of</strong><br />

internet resources such as ePIC (electronic Plant Information Centre administered by <strong>the</strong> Royal<br />

Botanic Gardens, Kew), IPNI (International Plant Names Index), GRIN (Germplasm Resources<br />

Information Network, United States Department <strong>of</strong> Agriculture) and for Asteraceae and some<br />

o<strong>the</strong>r plants <strong>the</strong> Euro+Med PlantBase. Commonly-used synonyms have been given for each species<br />

where relevant and any taxonomic uncertainties have been noted in <strong>the</strong> species accounts.<br />

Arabic names have been compiled from first-hand knowledge <strong>of</strong> <strong>the</strong> authors, partially supplemented<br />

by literature searches. Names have been listed where well known in <strong>Qatar</strong> or in widespread use in<br />

<strong>the</strong> Gulf or o<strong>the</strong>r parts <strong>of</strong> <strong>the</strong> Arab-speaking world. Most <strong>the</strong> publications listed above also have<br />

useful lists <strong>of</strong> Arabic names, but Mandaville (1990) is recommended as he gives <strong>the</strong> sources <strong>of</strong> most<br />

<strong>of</strong> his names and provides a comprehensive Arabic index.<br />

Overview <strong>of</strong> Habitats and Vegetation<br />

<strong>Qatar</strong> is largely an arid, desert area with a hot, sunny climate, receiving minimal rainfall, which is<br />

<strong>of</strong>ten sporadic in occurrence. Fogs occur in winter and spring and contribute a more regular source<br />

<strong>of</strong> water to <strong>the</strong> flora. The majority <strong>of</strong> <strong>the</strong> land surface consists <strong>of</strong> flat to gently undulating calcareous<br />

rocks, with sand formations occurring in <strong>the</strong> south-east. The country is largely low-lying, but <strong>the</strong>re<br />

are some small jebels in <strong>the</strong> south (rising to just over 100 metres at <strong>the</strong> highest point) and a rocky<br />

Miocene ridge extending from Dukhan to near <strong>the</strong> sou<strong>the</strong>rn border on <strong>the</strong> west coast. Away from<br />

<strong>the</strong> deeper sand areas, <strong>the</strong> soils are generally shallow, consisting <strong>of</strong> sands, silts and gravels, but <strong>the</strong><br />

north <strong>of</strong> <strong>the</strong> country also supports numerous natural depressions known as ‘rodat’, <strong>of</strong> various sizes,<br />

many <strong>of</strong> which have much deeper, richer soils, and have <strong>the</strong>refore been converted to farmland to<br />

grow crops. The reader is recommended to refer to Batanouny (1983) and Abulfatih et al. (2001)<br />

for fur<strong>the</strong>r information on geology, soils and climate.<br />

From a plant ecologist’s point <strong>of</strong> view <strong>the</strong> main edaphic (soil) factors affecting choice <strong>of</strong> habitat<br />

<strong>of</strong> desert species in areas such as <strong>the</strong> Gulf are <strong>the</strong> degree <strong>of</strong> retention <strong>of</strong> soil moisture, <strong>the</strong> level<br />

vi vii


<strong>of</strong> salinity, soil particle size and <strong>the</strong> degree <strong>of</strong> soil compaction. These are <strong>of</strong>ten inter-related. The<br />

mineral and nutrient content <strong>of</strong> soils may be important secondary factors affecting plant growth<br />

in some areas. Thus, <strong>the</strong> flora <strong>of</strong> <strong>Qatar</strong> can usually be placed into one <strong>of</strong> five main habitat groups:<br />

(a) strongly ‘xerophytic’ species <strong>of</strong> rock and gravel deserts where soil depth is limited and<br />

conditions are very dry;<br />

(b) ‘halophytic’ species tolerant <strong>of</strong> saline areas, such as saltmarshes, coastal sands, sabkha edge<br />

and inland oolitic sands;<br />

(c) species that grow in natural silt and sand depressions where water retention is higher than<br />

for (a);<br />

(d) species adapted to grow in deeper sand, where water is available under <strong>the</strong> surface;<br />

(e) species associated with man-made and man-influenced sites, particularly those receiving<br />

artificial irrigation, such as roadsides, farms, gardens and sewage ponds.<br />

Species that are more catholic in <strong>the</strong>ir requirements may be found in more than one group.<br />

<strong>An</strong>nual species mainly fall in groups (a), (c) and (e) and can grow in almost any situation or soil<br />

type, provided <strong>the</strong>re is at least a shallow surface covering <strong>of</strong> sand or silt. Group (e) includes many<br />

naturalised species that are weeds <strong>of</strong> cultivation. However, a number <strong>of</strong> true desert species also grow<br />

in irrigated habitats; <strong>of</strong>ten more luxuriantly due to <strong>the</strong> increased supply <strong>of</strong> water. Some <strong>of</strong> <strong>the</strong>se are<br />

adapted to spread quickly in times <strong>of</strong> good rainfall.<br />

The main habitat types and associated vegetation in <strong>Qatar</strong> can be summarised as follows:<br />

Dunes and deep sand areas are strongly undulating or form chains <strong>of</strong> <strong>the</strong> familiar crescentshaped<br />

‘barchan’ dunes; <strong>the</strong>y occur mainly in <strong>the</strong> south and south-east and are moderately saline,<br />

characteristically supporting <strong>the</strong> mound-forming shrub Seidlitzia rosmarinus.<br />

Sand sheets are shallower areas <strong>of</strong> sand forming flat or gently undulating terrain; <strong>the</strong>y may be<br />

stabilised and firm, or sometimes quite s<strong>of</strong>t. They occur mainly in <strong>the</strong> south-west and are <strong>of</strong>ten<br />

dominated by sand-binding grasses, especially Panicum turgidum, or locally by <strong>the</strong> shrum Haloxylon<br />

salicornicum.<br />

White sand <strong>of</strong> marine origin occurs inland in SW <strong>Qatar</strong>. It is moderately saline and supports a<br />

distinctive vegetation dominated by Cyperus conglomeratus and Tetraena qatarensis. Agriophyllum minus<br />

is a distinctive associate.<br />

Coastal sand on beaches is usually coarse-textured and calcareous due to abundant shell fragments.<br />

It is <strong>of</strong>ten grey, or sometimes pinkish in hue. Halophytic shrubs such as Cornulaca spp. usually<br />

dominate, but <strong>the</strong> habitat <strong>of</strong>ten supports diverse mixed grass and shrub vegetation. Naturalised<br />

date palms (Phoenix dactylifera) occur in this habitat at Umm Bab.<br />

Wadis and runnels are linear features formed in <strong>the</strong> landscape, which may flow with water during<br />

times <strong>of</strong> heavy rainfall. In <strong>Qatar</strong> <strong>the</strong>y are generally not well defined in terms <strong>of</strong> depth or gradient, but<br />

are characterised by linear or oval patches <strong>of</strong> accumulated sand, <strong>of</strong>ten within o<strong>the</strong>rwise featureless<br />

hamada or gravel desert areas. They support distinctive types <strong>of</strong> vegetation, composed <strong>of</strong> certain<br />

grasses, perennial and annual herbs and also <strong>of</strong>ten support stands <strong>of</strong> Lycium shawii. The lily Dipcadi<br />

erythraeum is especially characteristic <strong>of</strong> small sandy runnels.<br />

Sand and silt pans and larger natural depressions (‘rodat’) support moderately compact to<br />

very compact soils composed <strong>of</strong> fine sand, silt or loam. They <strong>of</strong>ten become flooded during rains<br />

and may retain moisture for some time afterwards. Soils <strong>of</strong>ten have higher quantity <strong>of</strong> organic<br />

matter than o<strong>the</strong>r desert areas. They are usually characterised by stands <strong>of</strong> trees and shrubs<br />

comprising Acacia spp., Prosopis juliflora, Ziziphus nummularia and Lycium shawii; and are <strong>of</strong>ten rich<br />

in annual and perennial herbs and grasses. Characteristic species <strong>of</strong> silt pans include Cymbopogon<br />

commutatus, Pulicaria undulata, <strong>An</strong>drachne telephioides, Astragalus eremophilus, Althaea ludwigii and<br />

Corchorus depressus. In <strong>the</strong> south, depressions are mostly small, circular features a few metres to a<br />

few hundred metres across, but in <strong>the</strong> north and central parts <strong>of</strong> <strong>Qatar</strong> <strong>the</strong>y may be irregular or<br />

more linear in shape and can reach 2-3 kilometres in length. These larger rodat have <strong>of</strong>ten been<br />

converted to farmland and are irrigated and cultivated to grow crops. For fur<strong>the</strong>r information see<br />

Batanouny (1981) and Babikir (1986).<br />

‘Sabkha’ refers to usually low-lying areas <strong>of</strong> sandy desert which have been inundated by seawater<br />

or rainwater that has dried out leaving a salt-encrusted surface. They occur mainly on or near <strong>the</strong><br />

coast in <strong>Qatar</strong>, with larger ones occurring east <strong>of</strong> Dukhan and at Mesaieed. Most are too salty to<br />

support any vegetation, but many have a marginal zone (referred to as ‘sabkha edge’) that is <strong>of</strong>ten<br />

dominated by halophytic species such as Limonium axillare, Salsola spp. or Tetraena qatarensis. Native<br />

species <strong>of</strong> Tamarix may also occur.<br />

Saltmarsh occurs in coastal bays and creeks, <strong>of</strong>ten in muddy areas, and is locally dominated by<br />

mangrove Avicennia marina, with dense stands <strong>of</strong> Arthrocnemum macrostachyum and o<strong>the</strong>r halophytes.<br />

Gypsum desert occurs in <strong>the</strong> west and north-west, particularly in <strong>the</strong> Dukhan area. Soils are<br />

composed <strong>of</strong> a high proportion <strong>of</strong> calcium carbonate and are strongly calcareous. They vary from<br />

s<strong>of</strong>t to moderately compact and are also mixed with sand and silt in places. They support a variety<br />

<strong>of</strong> vegetation types that also occur in o<strong>the</strong>r sandy and gravelly habitats, but a few species such as<br />

Helian<strong>the</strong>mum kahiricum and Erodium glaucophyllum are particularly characteristic.<br />

Gravel plain is used to describe flat desert areas with a surface layer <strong>of</strong> gravel or stones. Several<br />

variations <strong>of</strong> this habitat occur in <strong>Qatar</strong>, with soils varying from s<strong>of</strong>t to compact and fine to coarsetextured.<br />

Most <strong>of</strong> <strong>the</strong>se are characterised by relatively poor cover <strong>of</strong> vegetation, <strong>of</strong>ten <strong>of</strong> lowgrowing<br />

or stunted perennial herbs, grasses and shrubs. Typical species include Helian<strong>the</strong>mum lippii,<br />

Polycarpaea repens and Stipagrostis spp.<br />

‘Hamada’ is flat or gently undulating desert with stones and rocks on <strong>the</strong> surface or embedded in<br />

<strong>the</strong> surface, usually with compact soils and only small quantities <strong>of</strong> wind-blown sand and silt on <strong>the</strong><br />

surface. In <strong>Qatar</strong> this habitat is formed from <strong>the</strong> limestone bedrock and is a feature <strong>of</strong> large areas<br />

<strong>of</strong> <strong>the</strong> interior <strong>of</strong> <strong>the</strong> country. The vegetation is usually dominated by Lycium shawii and Tetraena<br />

qatarensis, with <strong>the</strong> annual grass Stipa capensis turning <strong>the</strong> landscape green after winter rains.<br />

Rock outcrops occur in <strong>the</strong> west and south-west <strong>of</strong> <strong>the</strong> country and occasionally elsewhere. The<br />

summits are <strong>of</strong>ten ra<strong>the</strong>r bare <strong>of</strong> vegetation due to <strong>the</strong> lack <strong>of</strong> soil particles for growth, but <strong>the</strong>ir<br />

sides accumulate pockets <strong>of</strong> wind-blown sand and can be quite rich in vegetation. After winter rains<br />

<strong>the</strong> low-growing annual Sclerocephalus arabicus may be locally abundant.<br />

Cultivated and irrigated soils are damp periodically or all year round and <strong>of</strong>ten have higher levels<br />

<strong>of</strong> organic matter and nutrients. Some may be moderately saline. Depending on <strong>the</strong> purpose for<br />

which <strong>the</strong>y are used <strong>the</strong>y can support various kinds <strong>of</strong> vegetation, <strong>of</strong>ten <strong>of</strong> non-native, naturalised<br />

species.<br />

Wetland areas include pools and channels that take waste water from irrigation and water treatment<br />

schemes and remain wet or waterlogged all year round. Such sites usually support dense reedbeds<br />

<strong>of</strong> Phragmites australis around <strong>the</strong> margins, <strong>of</strong>ten mixed in with Tamarix. O<strong>the</strong>r tall emergent plants<br />

sometimes occur. The edge zones <strong>of</strong>ten support lower-growing wetland grasses such as Polypogon<br />

monspeliensis and Aeluropus lagopoides. <strong>An</strong> extensive lake system to <strong>the</strong> west <strong>of</strong> Doha has been formed<br />

in recent years from waste water sources.<br />

viii ix


Notes on Layout and Terminology<br />

Abbreviations Used in <strong>the</strong> Text<br />

To make this checklist easy to read and understand, abbreviations and codes have not been used,<br />

except to denote approximate geographical areas <strong>of</strong> <strong>Qatar</strong>. The country can be roughly divided<br />

into equal thirds: north (N), central (C) and east (E), with <strong>the</strong> nor<strong>the</strong>rn and sou<strong>the</strong>rn parts each also<br />

conveniently divided into NW and NE, SW and SE. This appears to be <strong>the</strong> system used by earlier<br />

authors. Main localities are shown on <strong>the</strong> map (p. xiv).<br />

Shortened author names are used in <strong>the</strong> lists <strong>of</strong> photographs and illustrations (see below). Initials<br />

<strong>of</strong> <strong>the</strong> present authors (BB, SAM, JN, MS & RR) are used in <strong>the</strong> text when attributing plant records.<br />

Square brackets around a species name indicates that <strong>the</strong>re is a strong possibility it was misidentified<br />

by previous authors and has not occurred in <strong>Qatar</strong>.<br />

Order <strong>of</strong> Species<br />

The checklist is arranged in <strong>the</strong> traditional order <strong>of</strong> ferns (Pteridophyta), cone-bearing flowering<br />

plants (Gymnospermae), ‘double seed-leaved’ flowering plants (Dicotyledoneae) and ‘single seedleaved’<br />

flowering plants (Monocotyledoneae). Families and species are arranged in alphabetical<br />

order within <strong>the</strong>se. Only a single fern (Ophioglossum polyphyllum) and gymnosperm (Ephedra foliata)<br />

are presently known to occur in <strong>Qatar</strong>.<br />

Family Names<br />

Family names are mostly those recommended by <strong>the</strong> <strong>An</strong>giosperm Phylogeny Group (2003), with<br />

<strong>the</strong> modern names being used for Apiaceae, Asteraceae, Brassicaceae, Fabaceae, Lamiaceae and<br />

o<strong>the</strong>rs (now less <strong>of</strong>ten referred to as Umbelliferae, Compositae, Cruciferae, Leguminosae and<br />

Labiatae). Tradition has been upheld by recognising Capparaceae, but including it within it those<br />

species <strong>of</strong>ten separated into <strong>the</strong> Cleomaceae.<br />

Scientific Names<br />

The currently accepted scientific name <strong>of</strong> each species is given in bold type, with <strong>the</strong> author citation<br />

in standard format. Synonyms in common or recent use are given in italics with <strong>the</strong>ir authors below<br />

<strong>the</strong> species entry.<br />

Vernacular Names<br />

Vernacular (or common) names in English are listed only for widely occurring and familiar species<br />

or genera. Some authors <strong>of</strong> regional floras have invented <strong>the</strong>ir own English names, but this practice<br />

serves no scientific use and can lead to confusion, so for a large number <strong>of</strong> species no English<br />

vernacular name has been given. In some cases <strong>the</strong> names given are local to <strong>the</strong> place <strong>of</strong> origin,<br />

but have become more widely used (e.g. some Indian names <strong>of</strong> species naturalised in <strong>the</strong> Gulf).<br />

The Arabic vernacular names are shown in Arabic script and as English transliterations. Words <strong>of</strong><br />

similar root but pronounced differently are separated by a slash symbol (/). This usually reflects a<br />

slightly different dialect or different derivations <strong>of</strong> <strong>the</strong> same word, but not all variations have been<br />

given. Single or double uppercase letters in <strong>the</strong> transliterations are used to denote <strong>the</strong> emphatic<br />

consonants. The Arabic letter ’ain has usually been denoted by a single apostrophe, and <strong>the</strong> letter qaf<br />

by <strong>the</strong> English ‘q’, even though <strong>of</strong>ten pronounced as a hard ‘g’ in <strong>the</strong> Gulf and usually spelt as such.<br />

Growth Form & Flowering Period<br />

The species’ growth form or habit, and height is summarised using general terms (see Box 2).<br />

The likely flowering period in <strong>Qatar</strong> is shown as abbreviated month ranges, e.g. ‘Feb–Mar’, or ‘all<br />

year’ for species that may flower at any time <strong>of</strong> <strong>the</strong> year. It should be borne in mind that flowering<br />

periods may be variable for many species, as <strong>the</strong>y are <strong>of</strong>ten determined by <strong>the</strong> timing <strong>of</strong> winter or<br />

spring rains.<br />

Status<br />

Each species is stated as being ei<strong>the</strong>r native (occurring in <strong>Qatar</strong> within part <strong>of</strong> its natural range) or<br />

introduced (occurring as a result <strong>of</strong> human action). A preceding question mark [?] indicates that <strong>the</strong><br />

status is uncertain. What has been termed here as <strong>the</strong> ‘rarity status’ or overall occurrence <strong>of</strong> species<br />

in <strong>Qatar</strong> is categorised as ‘very common’, ‘common’, ‘local’ or ‘rare’ (see Box 1). This provides<br />

a broad assessment <strong>of</strong> <strong>the</strong> frequency <strong>of</strong> species, regardless <strong>of</strong> habitat type or local abundance.<br />

These classifications could be used as an initial stage in selecting species <strong>of</strong> conservation priority;<br />

for example those native species that are <strong>of</strong> local or rare occurrence. A preceding question mark<br />

indicates that <strong>the</strong> rarity status is uncertain, but for a small number <strong>of</strong> planted or introduced species<br />

<strong>the</strong> occurrence is unknown and this category has been omitted.<br />

For species not recorded by Batanouny or El Amin, a source for <strong>the</strong> first published record is given, or<br />

where published for <strong>the</strong> first time in this checklist, details <strong>of</strong> <strong>the</strong> record are given, with <strong>the</strong> author’s<br />

name or initials. Specimens <strong>of</strong> most <strong>of</strong> <strong>the</strong>se species have been retained by JN and identification<br />

confirmed by E. Clement; <strong>the</strong>y will deposited at Kew herbarium in due course.<br />

<strong>Qatar</strong> has no endemic species, but mention is made <strong>of</strong> a few ‘near-endemics’, for example species<br />

with distributions restricted to parts <strong>of</strong> <strong>the</strong> Gulf. A list <strong>of</strong> o<strong>the</strong>r Gulf countries in which <strong>the</strong> species<br />

has been recorded is given at <strong>the</strong> end <strong>of</strong> <strong>the</strong> status section in alphabetical order. Oman has been<br />

excluded from <strong>the</strong>se lists as most <strong>of</strong> it lies outside <strong>the</strong> Arabian Gulf area. For Saudi Arabia only<br />

species included in Mandaville (1990) are noted, in addition to a few additional ones mapped by<br />

Miller & Cope (1996) and Cope (2007), though <strong>the</strong>se sources were not exhaustively checked.<br />

Abundance, Habitat & Distribution<br />

The local abundance, habitat preferences and distribution over <strong>Qatar</strong> are summarised in this<br />

section. Local or ‘relative’ abundance is <strong>the</strong> frequency and abundance <strong>of</strong> species relative to <strong>the</strong><br />

area or habitat in which <strong>the</strong>y are growing (see Box 3). Information on habitat preferences and<br />

distribution has usually been combined, ra<strong>the</strong>r than listing separately, since in many cases <strong>the</strong>se are<br />

inter-related. General information on habitats is given in <strong>the</strong> Introduction. Distribution is given in<br />

terms <strong>of</strong> <strong>the</strong> main geographical areas <strong>of</strong> <strong>Qatar</strong> as described above or by reference to more specific<br />

localities or areas. Localities are mentioned mainly for <strong>the</strong> more localised and rare species, where<br />

necessary repeating details given by previous authors. Most are included on <strong>the</strong> map (page xiv).<br />

Box 1. Categories used to summarise occurrence in <strong>Qatar</strong><br />

very common Species distributed over most parts <strong>of</strong> <strong>the</strong> country and occurring in most<br />

habitat types.<br />

common Species with a wide range, but may be absent from some geographical<br />

areas or from certain types <strong>of</strong> habitat.<br />

local Species <strong>of</strong> scattered distribution, i.e. occurring only in a small number <strong>of</strong><br />

<strong>of</strong>ten widely spaced sites. Invariably this will apply to species dependent<br />

on certain types <strong>of</strong> habitat or habitat conditions which <strong>the</strong>mselves are<br />

localised.<br />

rare Species <strong>of</strong> very limited distribution in <strong>Qatar</strong>; usually restricted to about<br />

five or fewer localities (based on published information and current<br />

knowledge).<br />

x xi


Box 2. Main terms used to describe growth form and habit<br />

tree Long-lived, woody plant with clearly defined trunk; growing to 3 m or<br />

more in height, but <strong>of</strong>ten shorter if browsed by livestock.<br />

shrub/small shrub Long-lived woody plant usually without clearly defined trunk; usually<br />

attaining a height <strong>of</strong> 0.3–3 m; ‘small shrub’ has been used to refer to<br />

shrubs that do not usually exceed 60 cm in height.<br />

shrublet Perennial plants with <strong>the</strong> appearance <strong>of</strong> herbs, but with tough, woody<br />

lower stems; usually less than 30 cm in height, but <strong>of</strong>ten much shorter<br />

(may be spreading over <strong>the</strong> ground).<br />

annual herb Herbaceous, i.e. non-woody, plants usually completing a life cycle <strong>of</strong><br />

germination to seed production within a 12-month period. In desert<br />

conditions annual herbs rarely grow taller than 30 cm and are usually<br />

much shorter.<br />

biennial/perennial herb Herbaceous plants (o<strong>the</strong>r than grasses, rushes and sedges) living longer<br />

than a year. Biennial herbs normally complete a life cycle in two years;<br />

‘tall herb’ is sometimes used to refer to herbs normally exceeding 1 m in<br />

height.<br />

annual/perennial grass Grasses are members <strong>of</strong> <strong>the</strong> Poaceae family. These share <strong>the</strong> characteristics<br />

<strong>of</strong> having usually hollow stems with simple linear flat or rolled leaves and<br />

specialised flowers without coloured petals.<br />

sedge Members <strong>of</strong> <strong>the</strong> Cyperaceae family. Like grasses <strong>the</strong>y have specialised and<br />

distinctive flowering structures, but mostly have solid stems.<br />

rush Members <strong>of</strong> <strong>the</strong> Juncaceae family (one species in <strong>Qatar</strong>).<br />

climber Plants but which rely on o<strong>the</strong>r plants for support. They <strong>of</strong>ten take <strong>the</strong> form<br />

<strong>of</strong> long stems winding around <strong>the</strong> stems <strong>of</strong> o<strong>the</strong>r plants in spiral fashion.<br />

parasitic plant Plants lacking chlorophyll, and <strong>the</strong>refore having now green parts; usually<br />

growing on and obtaining nutrients from roots or stems <strong>of</strong> o<strong>the</strong>r plants.<br />

Box 3. Terms used to describe local abundance<br />

dominant Species forming <strong>the</strong> greatest proportion <strong>of</strong> biomass <strong>of</strong> <strong>the</strong> vegetation <strong>of</strong> a<br />

localised area or habitat and usually also <strong>the</strong> highest percentage cover <strong>of</strong><br />

all species present. Such species are <strong>of</strong>ten used to define vegetation types.<br />

The term is not used here to refer to <strong>the</strong> most numerous or highest cover<br />

species in sparse desert vegetation, as this can be misleading.<br />

abundant Species that are very numerous over an area or habitat and are <strong>of</strong>ten<br />

characteristic <strong>of</strong> it, usually contributing a high proportion <strong>of</strong> <strong>the</strong> plant<br />

biomass and/or vegetation cover.<br />

frequent Species that are numerous in a particular area or habitat type and <strong>of</strong>ten<br />

characteristic <strong>of</strong> it, though not normally contributing a high proportion<br />

<strong>of</strong> <strong>the</strong> total biomass or vegetation cover.<br />

occasional Species occurring at low frequency wherever <strong>the</strong>y occur, contributing very<br />

little to <strong>the</strong> total biomass <strong>of</strong> <strong>the</strong> vegetation or to <strong>the</strong> cover.<br />

Uses<br />

This section summarises <strong>the</strong> main cultural and economic uses <strong>of</strong> plants, ei<strong>the</strong>r recorded historically<br />

and traditionally or during <strong>the</strong> present-day, though <strong>the</strong> sources consulted <strong>of</strong>ten do not make a<br />

clear distinction between past and present. Mention is also made <strong>of</strong> <strong>the</strong> main desert species <strong>of</strong><br />

importance for grazing and browsing animals, i.e. camels, sheep and goats (now ubiquitous over<br />

<strong>the</strong> Gulf and representing <strong>the</strong> biggest threat to <strong>the</strong> native flora). Poisonous species have also been<br />

highlighted in <strong>the</strong> accounts.<br />

Most <strong>of</strong> <strong>the</strong> information on medicinal uses has been summarised from <strong>the</strong> earlier floras or from<br />

Ghazanfar (1994) and Rizk & El-Ghazaly (1995). These have been cited in <strong>the</strong> text but <strong>the</strong> dates<br />

have not been repeated to save space. Only <strong>the</strong> well known or general uses have been mentioned<br />

here, so <strong>the</strong>se publications should be consulted for more detailed information. A wealth <strong>of</strong> data<br />

is also available on <strong>the</strong> internet, due to <strong>the</strong> current widespread interest in herbal medicine. The<br />

authors <strong>of</strong> this checklist do not accept liability for any incorrect or misleading information given.<br />

Notes<br />

Additional taxonomic information and o<strong>the</strong>r notes are included here.<br />

Photographs and Illustrations (‘Photos’)<br />

This entry lists <strong>the</strong> page or plate numbers <strong>of</strong> photographs and o<strong>the</strong>r illustrations published in<br />

<strong>the</strong> floras covering <strong>the</strong> Gulf region. Line drawings and coloured illustrations are indicated by a<br />

superscript ‘L’ after <strong>the</strong> number. Line drawings in Miller & Cope (1996) and Cope (2007) have not<br />

been included. The publications have been referred to using abbreviated authors’ names as follows:<br />

El Amin El Amin, H.M (Arab Organization for Agricultural Development) (1983). Wild plants <strong>of</strong><br />

<strong>Qatar</strong>.<br />

Bat Batanouny, K.H. (1981). Ecology and <strong>Flora</strong> <strong>of</strong> <strong>Qatar</strong>.<br />

Coll Collenette, S. (1999). Wildflowers <strong>of</strong> Saudi Arabia.<br />

Corn Cornes, M.D. & Cornes, C.D. (1989). The wild flowering plants <strong>of</strong> Bahrain.<br />

Daoud Daoud, H.S. & Al-Rawi, A. (1985). <strong>Flora</strong> <strong>of</strong> Kuwait. Volume 1: Dicotyledoneae.<br />

Ghaz Ghazanfar, S.A. (2003). <strong>Flora</strong> <strong>of</strong> Oman, Volume 1, Piperaceae–Primulacaea. [CD with book]<br />

Jong Jongbloed, M. et al. (2003). The comprehensive guide to <strong>the</strong> wildflowers <strong>of</strong> <strong>the</strong> United Arab<br />

Emirates.<br />

Mand Mandaville, J.P. (1990). <strong>Flora</strong> <strong>of</strong> Eastern Saudi Arabia.<br />

Phil Phillips, D.C. (1988). Wild flowers <strong>of</strong> Bahrain. A field guide to herbs, shrubs and trees.<br />

Pick Pickering, H. & Patzelt, A. (2008). Field guide to <strong>the</strong> wild plants <strong>of</strong> Oman.<br />

Shuaib Shuaib, L. (1995). Wild flowers <strong>of</strong> Kuwait.<br />

West Western, A.R. (1989). The flora <strong>of</strong> <strong>the</strong> United Arab Emirates: an introduction.<br />

Although not referenced here, Sheila Collenette’s earlier book, <strong>An</strong> illustrated guide to <strong>the</strong> flowers <strong>of</strong><br />

Saudi Arabia (Collenette 1985), is also a valuable resource, as it contains a different selection <strong>of</strong><br />

photographs to those shown in her later publication (Collenette 1990). Plate numbers for species<br />

illustrated in this checklist are given to <strong>the</strong> right <strong>of</strong> <strong>the</strong> scientific name.<br />

xii xiii


Map <strong>of</strong> <strong>the</strong> State <strong>of</strong> <strong>Qatar</strong><br />

Map <strong>of</strong> <strong>the</strong> State <strong>of</strong> <strong>Qatar</strong>, showing main roads and principal localities mentioned in <strong>the</strong> text.<br />

Coordinates are shown in <strong>Qatar</strong> National Grid.<br />

PTERIDOPHyTA<br />

OPHIOGLOSSACEAE<br />

Ophioglossum polyphyllum A. Braun Plate 3<br />

Vernacular names: adder’s-tongue fern<br />

Growth form: Fern. Fl. Feb–Mar.<br />

status: Native. Local. First published record: Abdel Bari (1995); see also Abdel Bari (1997). Also<br />

recorded: UAE.<br />

habitat & distribution: Occasional, N and S <strong>Qatar</strong>, sandy areas.<br />

uses: Used to be eaten as salad greens (Jongbloed et al.).<br />

Photos: Coll 629; Jong 7.<br />

GyMNOSPERMAE<br />

EPHEDRACEAE<br />

Ephedra foliata Boiss.<br />

Ephedra ciliata Fischer & C.A. Mey.; Ephedra peduncularis Boiss.<br />

Vernacular names: shrubby horsetail, ’alanda<br />

ةدنلع<br />

Growth form: Climber. Fl. Apr–May.<br />

status: Native. Local. Also recorded: Bahrain, E Saudi Arabia, UAE.<br />

habitat & distribution: Occasional in sandy depressions, with Ziziphus or Acacia, N and C <strong>Qatar</strong>.<br />

uses: Batanouny mentions that protruding branches are usually ‘trimmed’ by camels. ‘Certain<br />

plants <strong>of</strong> this genus yield ephedrine, used in <strong>the</strong> treatment <strong>of</strong> colds, asthma and hay fever’ (Cornes<br />

& Cornes). ‘The crushed boiled plant is used for tanning’ (El Amin).<br />

Photos: Bat 17; Coll 297; Corn 50; Jong 9; Phil 87; West 30.<br />

ANGIOSPERMAE: DICOTyLEDONEAE<br />

ACANTHACEAE<br />

OPHIOGLOSSACEAE / EPHEDRACEAE / ACANTHACEAE<br />

Avicennia marina (Forssk.) Vierh. Plates 1, 4<br />

Avicennia <strong>of</strong>ficinalis L.<br />

ةروش ,مرق<br />

Vernacular names: dwarf mangrove, qurm, shourah<br />

Growth form: Tree. Fl. Mar.<br />

status: Native. Local. Also recorded: Bahrain, E Saudi Arabia, UAE.<br />

habitat & distribution: Sheltered bays and creeks, <strong>of</strong>ten along <strong>the</strong> tide line; usually dominant<br />

where it occurs. Main surviving areas are at Umm Tais, Ras Laffan, Al Dhakhira, Al Khor, all in NE<br />

<strong>Qatar</strong> and Al Wakra. Abulfatih et al. report finding a stunted specimen in a muddy inlet on <strong>the</strong> W<br />

coast and suggested <strong>the</strong> conditions were too saline <strong>the</strong>re. A Government programme to restore<br />

mangrove vegetation has been in place since 1988, with successful work being carried out at Ras<br />

Laffan, but recently a large area has been cleared at Al Wakra and some <strong>of</strong> <strong>the</strong> area at Al Khor has<br />

also been destroyed. Areas at Umm Tais and Al Dhakhira are protected as natural reserves.<br />

uses: Probably not very palatable to animals. ‘Sometimes browsed by camels when o<strong>the</strong>r plants are<br />

not available’ (Mandaville). ‘White mangrove is a source <strong>of</strong> tannin’ (Cornes & Cornes). The bark,<br />

roots and seeds have various medicinal uses.<br />

Photos: El Amin 2; Coll 745; Corn 36, 196; Jong 196, 197; Mand 174; Phil 28, 29; Pick 225; West 125.<br />

xiv 1


ACANTHACEAE / AIZOACEAE<br />

Blepharis ciliaris (L.) B.L. Burtt Plate 2<br />

Blepharis edulis auct.<br />

Vernacular names: eyelash plant, shawk al-Dub, naqi’, niqeyl<br />

Growth form: Perennial herb. Fl. Mar–Apr.<br />

status: Native. Local. Also recorded: E Saudi Arabia, UAE.<br />

ليقن ,يقن ,بضلا كوش<br />

habitat & distribution: Occasional, on compact stony and gravelly soils; mainly in C <strong>Qatar</strong>; rare in<br />

<strong>the</strong> west. El Amin recorded along <strong>the</strong> road from Al Sheehaniyah (where it still common) to Dukhan;<br />

it also occurs along <strong>the</strong> road to Al Zubarah (RR).<br />

uses: Grazed by camels but not by sheep or goats (El Amin). The roots are ground to make a<br />

powder (kohl) and used to treat eye inflammation and cataracts. The seeds have healing and antiinflammatory<br />

properties (Rizk & El-Ghazaly).<br />

Photos: Bat 93; Coll 5; Jong 106; Mand 196; Pick 15; West 135.<br />

AIZOACEAE<br />

Aizoon canariense L. Plate 6<br />

قدح ,ةنفج<br />

Vernacular names: jafnah, hadaq<br />

Growth form: Perennial herb. Fl. Mar–Apr.<br />

status: Native. Very common. Also recorded: Bahrain, Kuwait, E Saudi Arabia, UAE. habitat &<br />

distribution: Frequent; usually on harder substrates with shallow deposits <strong>of</strong> sand. uses: Eaten by<br />

camels; leaves are eaten in salads.<br />

Photos: El Amin 1; Bat 19; Coll 12; Corn 5, 53; Daoud 151; Jong 109; Mand 14; Phil 59, 60; Pick 91; Shuaib<br />

86; West 35.<br />

Aizoon hispanicum L.<br />

Vernacular names: jafnah<br />

Growth form: <strong>An</strong>nual herb. Fl. Mar–May.<br />

status: Native. Local. Also recorded: Bahrain, Kuwait, E Saudi Arabia.<br />

habitat & distribution: Coarse gravels, sandy depressions and runnels; C <strong>Qatar</strong>.<br />

Photos: Coll 12; Corn 53; Daoud 149, 150; Mand 15; Shuaib 87.<br />

Mesembryan<strong>the</strong>mum cryptanthum Hook. f. Plate 5<br />

Mesembryan<strong>the</strong>mum forskahlii Hochst. ex Boiss.; Opophytum forskahlii (Hochst. ex Boiss.) N.E. Br.<br />

لوساغ ,حمس<br />

Vernacular names: samH, ghasool<br />

Growth form: <strong>An</strong>nual herb. Fl. Apr.<br />

status: Native. Local. Also recorded: Bahrain, E Saudi Arabia.<br />

habitat & distribution: Noted in two places in western <strong>Qatar</strong>, 2007 (JN), including one site on<br />

<strong>the</strong> outskirts <strong>of</strong> Dukhan township and ano<strong>the</strong>r south <strong>of</strong> Umm Bab; both on compact, calcareous<br />

soils. Also Al Khor island (RR). Batanouny cited a record by Boulos (1978).<br />

uses: Ground seeds are used in making bread and o<strong>the</strong>r foodstuffs.<br />

Photos: Coll 13; Corn 54; Mand 16; Phil 61.<br />

Mesembryan<strong>the</strong>mum nodiflorum L. Plate 7<br />

Cryophytum nodiflorum (L.) L. Bolus<br />

لوساغ ,حمس ,رح<br />

Vernacular names: Egyptian fig marigold, hurr, samH, ghasool<br />

Growth form: <strong>An</strong>nual herb. Fl. Mar–Apr.<br />

ةنفج<br />

status: Native. Common. Also recorded: Bahrain, Kuwait, E Saudi Arabia, UAE.<br />

habitat & distribution: Frequent in dried out saline areas, such as <strong>the</strong> edges <strong>of</strong> irrigated fields<br />

and along beaches, on compacted sand or o<strong>the</strong>r soils; mainly NE <strong>Qatar</strong>.<br />

Photos: Coll 13; Corn 54; Daoud 152, 153; Jong 110; Phil 62; Shuaib 87.<br />

Zaleya pentandra (L.) C. Jeffrey<br />

Trian<strong>the</strong>ma pentandra L.<br />

Vernacular names: lami<br />

Growth form: <strong>An</strong>nual or perennial herb. Fl. Dec–Mar.<br />

status: Introduced. Rare. First published record: Abdel Bari (1997). Also recorded: UAE.<br />

habitat & distribution: A weed <strong>of</strong> waste ground.<br />

Photos: Jong 113; West 36.<br />

AMARANTHACEAE<br />

Aerva javanica (Burm. f.) Juss. ex Schult.<br />

Aerva persica (Burm. f.) Merr.; Aerva tomentosa Forssk.<br />

Vernacular names: tuwaim, Tirf, ra’<br />

ءار ,فرط ,ميوت<br />

Growth form: Small shrub. Fl. Feb–Mar.<br />

status: Native. Common. Also recorded: Bahrain, E Saudi Arabia, UAE.<br />

habitat & distribution: Occasional to frequent on rocky substrates with shallow sand, or in silty<br />

and sandy depressions.<br />

uses: Several authors note that <strong>the</strong> flower heads were formerly used by <strong>the</strong> bedouin for stuffing<br />

saddles, pillows and cushions. Used as a remedy for toothache and dermatitis, and an infusion<br />

is used to reduce swelling (Rizk & El-Ghazaly). Flowers are used as a wound dressing and to stop<br />

bleeding; liquid from <strong>the</strong> roots is used to treat eye diseases in cattle in Saudi Arabia (Ghazanfar).<br />

Photos: El Amin 3; Bat 31; Coll 33; Corn 58; Ghaz 54; Jong 115; Mand 52; Phil 26, 27; Pick 149; West 53, 54.<br />

Amaranthus graecizans L.<br />

Vernacular names: quTayf, sindar<br />

ردنس ,فيطق<br />

Growth form: <strong>An</strong>nual herb. Fl. Mar–Apr.<br />

status: Introduced. Common. Also recorded: Bahrain, E Saudi Arabia, UAE.<br />

habitat & distribution: Occasional in cultivated and disturbed areas, including farms and gardens.<br />

uses: Leaves are used as an emollient; crushed leaves are applied to scorpion stings, snake bites,<br />

irritating or itchy rashes (Ghazanfar).<br />

Photos: Bat 32; Coll 35; Corn 61; Jong 118; Pick 200; West 54.<br />

Amaranthus hybridus L.<br />

Amaranthus chlorostachys Willd.<br />

Vernacular names: rowaf<br />

Growth form: <strong>An</strong>nual herb. Fl. Mar–Apr.<br />

status: Introduced. Local. Also recorded: E Saudi Arabia, UAE.<br />

habitat & distribution: A weed <strong>of</strong> cultivated fields and gardens.<br />

Photos: Bat 32; Coll 35; Jong 119.<br />

AIZOACEAE / AMARANTHACEAE<br />

2 3<br />

يملا<br />

فاور


AMARANTHACEAE / APIACEAE<br />

Amaranthus viridis L.<br />

Amaranthus gracilis Desf.<br />

Growth form: <strong>An</strong>nual herb. Fl. Mar–Apr.<br />

status: Introduced. Common. Also recorded: Bahrain, E Saudi Arabia, UAE.<br />

habitat & distribution: Occasional in cultivated and disturbed areas, including farms and gardens.<br />

uses: Used in <strong>the</strong> treatment in a wide variety <strong>of</strong> stomach complaints and as a laxative (Rizk & El-<br />

Ghazaly).<br />

Photos: El Amin 1; Bat 32; Coll 37; Corn 61; Jong 120.<br />

APIACEAE<br />

Ammi majus L.<br />

Vernacular names: Bishop’s weed, khillah, nayniya<br />

Growth form: <strong>An</strong>nual herb. Fl. Apr–May.<br />

status: Introduced. Also recorded: Kuwait, UAE.<br />

habitat & distribution: Cultivated areas.<br />

ةينين ,ةلخ<br />

uses: Used in <strong>the</strong> Middle East for centuries as a treatment for particular skin diseases (Rizk & El-<br />

Ghazaly).<br />

Photos: Coll 722; Jong 531; Pick 151; Shuaib 59; West 106.<br />

<strong>An</strong>ethum graveolens L.<br />

Vernacular names: dill<br />

Growth form: <strong>An</strong>nual herb. Fl. Feb–Apr.<br />

status: Introduced. Also recorded: E Saudi Arabia, UAE.<br />

habitat & distribution: Cultivated areas.<br />

uses: Leaves and seeds used as salad, also used as a herb and spice in cooking for its aniseed-like<br />

flavouring, e.g. to flavour rice dishes. Numerous medicinal uses are described by Rizk & El-Ghazaly.<br />

Photos: Bat 75; Coll 722; Jong 532; West 106.<br />

Bupleurum semicompositum L.<br />

Growth form: <strong>An</strong>nual herb. Fl. Mar.<br />

status: Native. Rare. Also recorded: Bahrain, Kuwait, E Saudi Arabia.<br />

habitat & distribution: Very rare on sandy soils (Batanouny). Mainly a coastal species in <strong>the</strong> Gulf.<br />

Photos: Coll 724; Daoud 172 L ; Shuaib 58.<br />

Foeniculum vulgare (L.) Mill.<br />

Vernacular names: fennel, shamar<br />

رمش<br />

Growth form: Biennial or perennial herb. Fl. Mar–May.<br />

status: Introduced. Local. Not in o<strong>the</strong>r Gulf floras.<br />

habitat & distribution: <strong>An</strong> occasional escape, e.g. along irrigated roadsides.<br />

uses: Cultivated for its aniseed-flavoured seeds, which are widely used in Asian cooking. Numerous<br />

medicinal uses are described by Rizk & El-Ghazaly.<br />

Photos: Coll 730.<br />

Torilis nodosa (L.) Gaertn.<br />

Growth form: <strong>An</strong>nual herb. Fl. Feb–Apr.<br />

status: Introduced. Rare. First published record: Abdel Bari (1997). Also recorded: UAE.<br />

habitat & distribution: Details not noted by Abdel Bari, but recorded from wadis and plantations<br />

in <strong>the</strong> UAE (Jongbloed).<br />

Photos: Jong 537.<br />

APOCYNACEAE<br />

Nerium oleander L.<br />

Nerium mascatense DC.<br />

Vernacular names: common oleander, Haban<br />

Growth form: Tree. Fl. All year.<br />

status: Introduced. Also recorded: UAE.<br />

habitat & distribution: Not known but widespread throughout <strong>the</strong> Gulf; native to parts <strong>of</strong> UAE<br />

and Oman.<br />

uses: Widely planted in towns for shade and landscaping. The bitter roots are used medicinally,<br />

including as a diuretic and emetic. Ghazanfar states that <strong>the</strong> leaves are used for <strong>the</strong> treatment <strong>of</strong><br />

bronchitis and coughs in Arabia.<br />

Photos: Jong 125; Pick 47; West 110.<br />

ASCLEPIADACEAE<br />

APIACEAE / APOCyNACEAE / ASCLEPIADACEAE<br />

Calotropis procera (Aiton) W.T. Aiton Plate 8<br />

راخشأ ,راشع<br />

Vernacular names: Sodom’s apple, ’ushar, ashkhar<br />

Growth form: Shrub or tree. Fl. May–Jul.<br />

status: Native. Local. Also recorded: Bahrain, Kuwait, UAE.<br />

habitat & distribution: In <strong>Qatar</strong> mainly in disturbed areas where <strong>the</strong>re is run-<strong>of</strong>f <strong>of</strong> rainwater<br />

or irrigation, such as including farms, roadsides and gardens. Rarely away from habitation. Not<br />

included in <strong>the</strong> accounts by Batanouny or El Amin, but was probably less common in <strong>the</strong> past.<br />

uses: Not edible and probably poisonous. Mandaville discusses <strong>the</strong> toxicity in some detail. He also<br />

notes that it is ‘well known to many bedouin who remember <strong>the</strong> shrub as <strong>the</strong> source <strong>of</strong> wood used<br />

in preparation <strong>of</strong> <strong>the</strong> best charcoal for <strong>the</strong> manufacture <strong>of</strong> black gunpowder’. Leaves and latex are<br />

used for treating wounds, pain, scorpion stings and for streng<strong>the</strong>ning muscles affected by paralysis<br />

(Ghazanfar).<br />

Photos: Coll 52; Corn 37; Jong 128, 129; Phil 133; Pick 21; Shuaib 83; West 111.<br />

Glossonema varians (Stocks) Benth. ex Hook. f. Plates 9, 10<br />

Glossonema edule N.E. Br.<br />

)ةرمثلا( ةوارج ,)هتبنلا( رتع<br />

Vernacular names: ’itr (plant), jarawah (fruit)<br />

Growth form: Perennial herb. Fl. Mar–Apr.<br />

status: Native. Common. Also recorded: Bahrain, E Saudi Arabia, UAE.<br />

habitat & distribution: Frequent to abundant, or locally dominant in shallow sandy runnels and<br />

depressions on gravel plains and hamada; mostly S <strong>Qatar</strong>.<br />

uses: ‘Whole plant is good fodder; fruit eaten by animals’ (El Amin); Mandaville also states that<br />

<strong>the</strong> very young leaves and raw young fruits are edible.<br />

Photos: El Amin 1; Bat 77 L ; Coll 65; Corn 162; Jong 132; Mand 155; Phil 134; Pick 94; West 112.<br />

4 5<br />

نابح


ASCLEPIADACEAE / ASTERACEAE<br />

Leptadenia pyrotechnica (Forssk.) Decne. Plate 11<br />

Vernacular names: burning bush, desert broom, markh<br />

خرم<br />

Growth form: Shrub or tree. Fl. Mar–Apr.<br />

status: Native. Local. Also recorded: Bahrain, E Saudi Arabia, UAE.<br />

habitat & distribution: Occasional in sandy habitats. Although a dominant component <strong>of</strong> <strong>the</strong><br />

coastal vegetation over much <strong>of</strong> <strong>the</strong> Gulf, it prefers non-saline, fairly deep sand and <strong>the</strong>refore is<br />

restricted in <strong>Qatar</strong>, mainly to western coastal areas. In NE <strong>Qatar</strong> it is rare near <strong>the</strong> coast at Dukhan<br />

(JN) and locally frequent at Ras Abrouq (SAM); grazed, stunted specimens have been seen at Al<br />

Kharrarah (RR).<br />

uses: Eaten by camels (El Amin). Young flowers and fruits ‘considered edible by sou<strong>the</strong>rn tribesmen’<br />

(Mandaville). Pickering & Patzelt (2008) note that <strong>the</strong> buds are edible and that dried hair from<br />

<strong>the</strong> seeds is used as kindling. <strong>An</strong> infusion made from <strong>the</strong> stems is taken as a diuretic (Ghazanfar).<br />

Photos: El Amin 1; Bat 78; Coll 70; Corn 165; Jong 133; Phil 132, 135; Pick 95; West 113.<br />

ASTERACEAE<br />

Aaronsohnia factorovskyi Warb. & Eig Plate 13<br />

Growth form: <strong>An</strong>nual herb. Fl. Mar–Apr.<br />

status: Native. Local. Also recorded: Kuwait, E Saudi Arabia.<br />

habitat & distribution: Occasional or locally frequent in small sandy or silty depressions; C and<br />

S <strong>Qatar</strong>.<br />

uses: ‘The plant is sometimes eaten raw by bedouin, who may also use it in <strong>the</strong> preparation <strong>of</strong> iqt<br />

(dried sour milk cakes)’ (Mandaville citing Dickson 1955).<br />

Photos: Bat 95; Coll 166; Mand 210, 211; Shuaib 111.<br />

Artemisia inculta Delile<br />

Artemisia sieberi Besser; Artemisia herba-alba Asso<br />

Vernacular names: shiH<br />

حيش<br />

Growth form: Perennial herb. Fl. May.<br />

status: Native. Rare. Not in o<strong>the</strong>r Gulf floras.<br />

habitat & distribution: Recorded by Batanouny at a single location along <strong>the</strong> road N <strong>of</strong> Doha,<br />

in a small depression.<br />

uses: The plant is known to have anti-microbial properties and is used as an antiseptic and an<br />

insecticide. Rizk & Ghazaly list many medicinal uses, including treatment for stomach complaints,<br />

coughs, bronchitis and to cure nervous troubles.<br />

Photos: None.<br />

Atractylis carduus (Forssk.) C. Chr. Plate 14<br />

Atractylis flava Desf.<br />

خيوش ,ينوميل ,ةولج<br />

Vernacular names: white thistle, jalwa, laymoony, shuwwaykh<br />

Growth form: <strong>An</strong>nual or perennial herb. Fl. Mar–Apr.<br />

status: Native. ?Common. Also recorded: Bahrain, Kuwait, E Saudi Arabia, UAE.<br />

habitat & distribution: Occasional in sandy habitats. Previously recorded from Al Karaanah, Al<br />

Wakra and Wadi Al Banat and recently from Al Kharrarah and o<strong>the</strong>r parts <strong>of</strong> W <strong>Qatar</strong>.<br />

uses: El Amin reported that it is not liked by animals but eaten by camels when nothing else is<br />

available.<br />

Photos: El Amin 7; Bat 96; Coll 174; Corn 204; Jong 143; Phil 67; Shuaib 115.<br />

Calendula arvensis (Vaill.) L.<br />

Vernacular names: field marigold, Hanwa<br />

Growth form: <strong>An</strong>nual herb. Fl. Feb–Apr.<br />

status: Introduced. Common. Also recorded: Bahrain, Kuwait, E Saudi Arabia, UAE.<br />

habitat & distribution: Occasional as a weed <strong>of</strong> disturbed areas and silty depressions.<br />

Photos: El Amin 38 L ; Bat 97; Coll 175; Jong 144; Phil 68; Shuaib 107.<br />

Calendula tripterocarpa Rupr.<br />

Growth form: <strong>An</strong>nual herb. Fl. Apr.<br />

status: Native. Rare. Also recorded: E Saudi Arabia.<br />

habitat & distribution: Sandy and gravelly soils in rodat; C <strong>Qatar</strong>.<br />

Photos: Coll 175; Mand 214.<br />

Carthamnus eriocephalus (Boiss.) Greuter<br />

Carduncellus eriocephalus Boiss.<br />

Vernacular names: kharshooT, lowmeya<br />

ةيمول ,طوشرخ<br />

Growth form: Perennial herb. Fl. Mar–Apr.<br />

status: Native. Local. Not in o<strong>the</strong>r Gulf floras.<br />

habitat & distribution: Occasional in rocky areas. El Amin recorded from Jebel Dukhan and<br />

Wadi Al Banat.<br />

Photos: None.<br />

Centaurea sinaica DC. Plate 15<br />

Centaurea pseudosinaica Mouterde, non Czerep.<br />

Vernacular names: birkan, murrar<br />

رارم ,ناكرب<br />

Growth form: <strong>An</strong>nual or biennial herb. Fl. Mar–Jul.<br />

status: Native. Common. Also recorded: Kuwait, E Saudi Arabia, UAE.<br />

habitat & distribution: Occasional in shallow sandy deposits.<br />

Photos: El Amin 7; Bat 97, 101; Coll 178; Jong 148; Mand 224, 225; Pick 99; Shuaib 116; West 141.<br />

Cichorium pumilum Jacq.<br />

Growth form: <strong>An</strong>nual herb. Fl. Summer.<br />

status: Introduced. Rare. Not in o<strong>the</strong>r Gulf floras.<br />

habitat & distribution: A weed in alfalfa (Medicago sativa) fields (Batanouny).<br />

Photos: None.<br />

ASTERACEAE<br />

Eclipta prostrata (L.) L.<br />

Eclipta alba (L.) Hassk.<br />

Vernacular names: false daisy<br />

Growth form: <strong>An</strong>nual herb. Fl. Apr–May.<br />

status: Introduced. Also recorded: Bahrain, E Saudi Arabia, UAE.<br />

habitat & distribution: Recorded from a single location by Batanouny, but occurs in irrigated<br />

lawns and o<strong>the</strong>r damp places elsewhere in <strong>the</strong> Gulf, so may be more widespread now in <strong>Qatar</strong>.<br />

uses: Rizk & El-Ghazaly list numerous cosmetic and medicinal uses.<br />

Photos: Coll 191; Corn 207; Jong 155; Mand 207; Pick 156; West 142.<br />

6 7<br />

ةونح


ASTERACEAE<br />

Erigeron bonariensis L.<br />

Conyza bonariensis (L.) Cronquist<br />

Growth form: <strong>An</strong>nual herb. Fl. Mar–Sep.<br />

status: Introduced. Local. Also recorded: Kuwait, E Saudi Arabia, UAE.<br />

habitat & distribution: Occasional, in gardens and cultivated areas.<br />

uses: Flowering branches are used as an antirheumatic and diuretic; <strong>the</strong> aerial part is used in <strong>the</strong><br />

treatment <strong>of</strong> liver and urinary diseases, and stomach ulcers (Rizk & El-Ghazaly).<br />

Photos: El Amin 7; Bat 97; Coll 182; Jong 151; Shuaib 108.<br />

Filago desertorum Pomel Plate 12<br />

Filago spathulata C. Presl forma desertorum (Pomel) Pamp.<br />

Vernacular names: quTeyna<br />

Growth form: <strong>An</strong>nual herb. Fl. Mar.<br />

status: Native. Common. Also recorded: Bahrain, E Saudi Arabia, UAE.<br />

habitat & distribution: Occasional in shallow sand, especially depressions and runnels.<br />

uses: El Amin noted that <strong>the</strong> plant is eaten by ‘rabbits’ (i.e. Cape Hare Lepus capensis).<br />

Photos: El Amin 40 L ; Bat 98; Coll 192; Corn 208; Jong 156; Phil 69; West 143.<br />

[Filago prolifera Pomel]<br />

Growth form: <strong>An</strong>nual herb. Fl. Mar.<br />

status: First published record: Obeid (1975). Not in o<strong>the</strong>r Gulf floras.<br />

notes: Not confirmed for <strong>Qatar</strong>. Batanouny and El Amin did not see this species, and it does<br />

not seem to have been recorded by o<strong>the</strong>r botanists in Arabia, so <strong>the</strong> record may be in error for F.<br />

desertorum.<br />

Photos: none.<br />

Flaveria trinervia (Spreng.) C. Mohr<br />

Growth form: <strong>An</strong>nual herb. Fl. Apr.<br />

status: Introduced. Local. Also recorded: Bahrain, E Saudi Arabia, UAE.<br />

habitat & distribution: Frequent in irrigated, cultivated areas. Noted as a ‘serious weed’ in some<br />

regions by Cornes & Cornes.<br />

Photos: Coll 193; Corn 208; Jong 157; West 143.<br />

Glebionis coronaria (L.) Spach<br />

Chrysan<strong>the</strong>mum coronarium L.<br />

Vernacular names: crown daisy<br />

Growth form: <strong>An</strong>nual herb. Fl. Mar–Apr.<br />

status: Introduced. Rare. Also recorded: E Saudi Arabia.<br />

habitat & distribution: Recorded from irrigated lawns in Doha b Batanouny.<br />

uses: Medicinal uses are listed by Rizk & El-Ghazaly.<br />

Photos: Coll 180.<br />

Gymnarrhena micrantha Desf. Plate 17<br />

Vernacular names: dog’s paw, camel’s eye<br />

Growth form: <strong>An</strong>nual herb. Fl. Mar–Apr.<br />

ةنيطق<br />

بلكلا فك<br />

status: Native. Common. Also recorded: Kuwait, E Saudi Arabia, UAE.<br />

habitat & distribution: Occasional on gravel plains, stony or rocky areas, growing in thin sandy<br />

deposits.<br />

uses: Poisonous and animals avoid it (El Amin).<br />

Photos: El Amin 8; Coll 193; Jong 158; Mand 201; Shuaib 113.<br />

Ifloga spicata (Forssk.) Sch. Bip. Plate 18<br />

Vernacular names: Hasaj, shajarat al ’anz<br />

Growth form: <strong>An</strong>nual herb. Fl. Mar–Apr.<br />

status: Native. Very common. Also recorded: Bahrain, Kuwait, E Saudi Arabia, UAE.<br />

زنعلا ةرجش ,جسح<br />

habitat & distribution: Frequent, on gravel plains, sand sheets, coastal sand and o<strong>the</strong>r areas with<br />

firm sandy soils; over most <strong>of</strong> <strong>Qatar</strong>.<br />

Photos: El Amin 40 L ; Coll 196; Corn 215; Jong 160; Mand 200; Phil 70; Shuaib 114; West 145.<br />

Koelpinia linearis Pall. Plate 19<br />

Vernacular names: goat’s beard, lahiat al-tais<br />

سيتلا ةيهلا<br />

Growth form: <strong>An</strong>nual herb. Fl. Mar–Apr.<br />

status: Native. ?Local. First published record: Abdel Bari (1997). Also recorded: Bahrain, Kuwait,<br />

UAE.<br />

habitat & distribution: A plant in fruit was photographed by S. Aspinall from SW <strong>Qatar</strong> in March<br />

2007 (see Plate).<br />

Photos: Coll 197; Corn 215; Jong 163; Phil 71; Shuaib 112.<br />

Laphangium luteoalbum (L.) Tzvelev<br />

Gnaphalium luteoalbum L.<br />

Growth form: <strong>An</strong>nual herb. Fl. Not known.<br />

status: Introduced. ?Rare. First published record: Abdel Bari (1997). Not in o<strong>the</strong>r Gulf floras.<br />

habitat & distribution: Cultivated areas.<br />

Photos: None.<br />

Launaea capitata (Spreng.) Dandy<br />

Launaea glomerata (Cass.) Hook. f.<br />

Vernacular names: Hawa/Huwa<br />

Growth form: Biennial herb. Fl. Feb–Apr.<br />

status: Native. Very common. Also recorded: Kuwait, E Saudi Arabia, UAE.<br />

habitat & distribution: Frequent in sandy and silty soils.<br />

uses: Edible to man and animals (El Amin).<br />

Photos: El Amin 8, 32 L ; Bat 100; Coll 200; Jong 165; Shuaib 109; West 145.<br />

Launaea goraeensis H<strong>of</strong>fm.<br />

Growth form: <strong>An</strong>nual herb. Fl. Mar.<br />

status: ?Native. ?Rare. Not confirmed for <strong>Qatar</strong>. Not in o<strong>the</strong>r Gulf floras.<br />

habitat & distribution: Sandy soils in SE <strong>Qatar</strong> and Umm Bab (El Amin).<br />

Photos: None.<br />

ASTERACEAE<br />

8 9<br />

ةوح


ASTERACEAE<br />

Launaea mucronata (Forssk.) Muschl.<br />

Launaea cassiniana (Jaub. & Spach) Kuntze; Sonchus cassinianus Jaub. & Spach<br />

Vernacular names: Hawa/Huwa, ’a<strong>the</strong>ed, yamroor, Hajaid, Safara<br />

Growth form: Perennial herb. Fl. Mar–Apr.<br />

status: Native. Local. Also recorded: Bahrain, Kuwait, E Saudi Arabia, UAE.<br />

ةرفص ,ديجح ,رورمي ,ديثع ,ةوح<br />

habitat & distribution: Occasional on sandy or silty soils; <strong>of</strong>ten in disturbed areas around<br />

habitation. Occurs along parts <strong>of</strong> <strong>the</strong> NE coast and in sandy habitats in <strong>the</strong> south (MS).<br />

notes: Subspp. L m. mucronata and L.m. cassiana both occur in <strong>the</strong> Gulf, <strong>the</strong> latter <strong>of</strong>ten being<br />

treated as a separate species.<br />

Photos: El Amin 32 L ; Coll 201; Jong 167, 168; Mand 232; Phil 73; Shuaib 109; West 146.<br />

Launaea nudicaulis (L.) Hook. f.<br />

Vernacular names: Hawa/Huwa, Huwa ghanam, ghazzal<br />

لازغ ,منغ ةوح ,ةوح<br />

Growth form: Biennial or perennial herb. Fl. Feb–Apr.<br />

status: Native. Very common. Also recorded: Bahrain, Kuwait, E Saudi Arabia, UAE.<br />

habitat & distribution: Frequent in sandy habitats.<br />

uses: ‘The leaves are used to treat fevers and to stop excessive bleeding after childbirth’ (Ghazanfar).<br />

Rizk & El-Ghazaly also state that <strong>the</strong>y are put on <strong>the</strong> heads <strong>of</strong> children to cure fever.<br />

Photos: El Amin 32 L ; Bat 102; Coll 202; Corn 216; Jong 169; Mand 233; Phil 74; Shuaib 110; West 147.<br />

Launaea procumbens (Roxb.) Ramayya & Rojagopal<br />

Launaea fallax (Jaub. & Spach) O. Kuntze<br />

Vernacular names: Hawa/Huwa<br />

Growth form: Perennial herb. Fl. Mar–Apr.<br />

status: Native. Common. Also recorded: Bahrain, E Saudi Arabia, UAE.<br />

habitat & distribution: Frequent in sandy habitats, including rodat, beaches and disturbed areas<br />

around habitation.<br />

Photos: El Amin 32 L ; Bat 100; Coll 202; Jong 170; Phil 75; West 147.<br />

Matricaria aurea (Loefl.) Sch. Bip.<br />

Growth form: <strong>An</strong>nual herb. Fl. Feb–Apr.<br />

status: Native. Rare. Also recorded: E Saudi Arabia, UAE.<br />

habitat & distribution: Silty depressions.<br />

uses: Chamomile tea is brewed from flower heads, ‘known to both bedouin and villagers’<br />

(Mandaville) and is used for general stomach problems such as colic, cramps and stomach aches<br />

(Ghazanfar).<br />

Photos: Coll 204; Jong 172.<br />

Matricaria chamomilla L.<br />

Matricaria recutita L.<br />

Growth form: <strong>An</strong>nual herb. Fl. Not known.<br />

status: Introduced. Rare. First published record: Abdel Bari (1997). Not in o<strong>the</strong>r Gulf floras.<br />

habitat & distribution: Recorded at Al Sheehaniyah and Doha.<br />

Photos: None.<br />

ةوح<br />

Pallenis hierichuntica (Michon) Greuter<br />

Asteriscus hierichunticus (Michon) Wiklund; Asteriscus pygmaeus (DC.) Coss. & Durieu; Odontospermum<br />

pygmaeum (DC.) O. H<strong>of</strong>fm.<br />

Vernacular names: jahwayan<br />

نايوهج<br />

Growth form: <strong>An</strong>nual herb. Fl. Feb–Mar.<br />

status: Native. Local. Also recorded: Kuwait, E Saudi Arabia, UAE.<br />

habitat & distribution: Silty depressions, N and C <strong>Qatar</strong> (El Amin).<br />

uses: El Amin mentions that it is used by locals to clean and perfume <strong>the</strong>ir teeth.<br />

Photos: El Amin 6; Bat 95; Coll 173; Jong 141; Mand 206; Shuaib 113.<br />

Pentanema divaricatum Cass.<br />

Vicoa pentanema Aitch. & Hemsl.<br />

Vernacular names: zmiem al bar<br />

Growth form: <strong>An</strong>nual herb. Fl. Mar–Apr.<br />

status: Native. Local. Also recorded: Bahrain, E Saudi Arabia, UAE.<br />

10 11<br />

ربلا ميمز<br />

habitat & distribution: Occasional in sandy and silty depressions with Ziziphus trees. In Saudi<br />

Arabia Mandaville recorded it as a weed <strong>of</strong> disturbed ground around farms and roadsides.<br />

uses: The plant has antifungal and antibacterial properties.<br />

Photos: El Amin 40 L ; Bat 107; Jong 174; West 152.<br />

Picris aspleniodes L.<br />

Picris radicata (Forssk.) Less.<br />

Vernacular names: Hawdhan<br />

Growth form: <strong>An</strong>nual herb. Fl. Mar–Apr.<br />

status: Native. Common. Not in o<strong>the</strong>r Gulf floras.<br />

habitat & distribution: Occasional in sandy depressions in S <strong>Qatar</strong>.<br />

Photos: El Amin 8; Bat 104; Coll 208.<br />

Picris cyanocarpa Boiss.<br />

Growth form: <strong>An</strong>nual herb. Fl. Mar–Apr.<br />

status: Native. Local. Also recorded: E Saudi Arabia.<br />

habitat & distribution: Occasional in saline sandy habitats.<br />

Photos: Coll 207.<br />

Pluchea dioscoridis (L.) DC.<br />

Conyza dioscoridis (L.) Desf.<br />

ASTERACEAE<br />

Growth form: Perennial herb or shrub. Fl. Jan–Dec.<br />

status: Introduced. Local. Also recorded: E Saudi Arabia, UAE.<br />

habitat & distribution: Recorded only once by Batanouny, but <strong>the</strong> species is now widespread<br />

(Abdel Bari 1997). Plentiful in Phragmites ponds at Dukhan water treatment works in 2007 (JN).<br />

notes: There is a discussion on <strong>the</strong> taxonomy <strong>of</strong> this species in Mandaville (1990, p. 289).<br />

Photos: Coll 209; Jong 177; Pick 54.<br />

ناذوح


ASTERACEAE<br />

Pulicaria gnaphalodes (Vent.) Boiss.<br />

Pulicaria undulata Kostel.<br />

Vernacular names: jithjath, nafayj<br />

Growth form: Perennial herb or small shrub. Fl. Mar–May.<br />

status: Native. Common. Also recorded: Bahrain.<br />

جيفن ,ثاجثج<br />

habitat & distribution: Occasional at Ras Laffan and Mesaieed (JN), but absent from <strong>the</strong> west<br />

coast. Also listed by Abdel Bari as ‘widespread’. Appears to prefer compact or disturbed gravelly or<br />

silty soils, particularly in depressions and wadis.<br />

uses: ‘A plant unpalatable to grazing animals and flourishing where o<strong>the</strong>rs are regularly grazed<br />

bare’ (Cornes & Cornes).<br />

notes: This is <strong>the</strong> species recorded by Batanouny as P. undulata, <strong>the</strong> name now given to Pulicaria<br />

crispa. At <strong>the</strong> time his flora was published P. crispa was known as Francoeuria crispa.<br />

Photos: El Amin 8; Bat 103; Corn 212; Phil 78.<br />

[Pulicaria sicula (L.) Moris]<br />

Growth form: <strong>An</strong>nual herb. Fl. Not known.<br />

status: First published record: Obeid (1975). Not in o<strong>the</strong>r Gulf floras.<br />

notes: A North African species not recorded by o<strong>the</strong>r botanists in Arabia. Obeid’s record could<br />

relate to a casual occurrence <strong>of</strong> an introduced plant or be in error for ano<strong>the</strong>r species.<br />

Photos: none.<br />

Pulicaria undulata (L.) C.A. Mey. Plate 16<br />

Francoeuria crispa (Forssk.) Cass.; Pulicaria crispa (Forssk.) Oliv. See note under P. gnaphalodes.<br />

لبجلا ياش ,ثاجثج<br />

Vernacular names: jithjath (and variations), shay el-jebel<br />

Growth form: Perennial herb. Fl. Mar–Jun.<br />

status: Native. Very common. Also recorded: Bahrain, Kuwait, E Saudi Arabia, UAE.<br />

habitat & distribution: Often abundant in small silty depressions. Scattered plants may occur<br />

anywhere in sandy or silty habitats.<br />

uses: Various authors report its use in traditional medicine, including use as a diuretic.<br />

Photos: Bat 99 L , 103; Coll 210; Corn 211; Jong 183; Phil 77; Shuaib 111; West 149.<br />

Reichardia tingitana (L.) Roth Plate 20<br />

Reichardia orientalis (L.) Hochr.<br />

ناذوح ,ريرم<br />

Vernacular names: mureer, Hawdhan<br />

Growth form: <strong>An</strong>nual herb. Fl. Mar–May.<br />

status: Native. Very common. Also recorded: Bahrain, Kuwait, E Saudi Arabia, UAE.<br />

habitat & distribution: Occasional in shallow sandy deposits, sandy depressions and cultivated<br />

areas.<br />

uses: Those listed by Ghazanfar include treatment <strong>of</strong> colic, constipation, swollen and inflamed<br />

eyes, using <strong>the</strong> leaves.<br />

Photos: El Amin 9, 38 L ; Coll 212; Corn 34, 220; Jong 184; Mand 234; Phil 79; Pick 107; Shuaib 107; West 149.<br />

Rhanterium epapposum Oliv. Plate 21<br />

Vernacular names: ’arfaj<br />

جفرع<br />

Growth form: Small shrub. Fl. Mar–Jul.<br />

status: Native. Local. Also recorded: Bahrain, Kuwait, E Saudi Arabia, UAE.<br />

habitat & distribution: Being a favoured grazing species is rarely seen in quantity. Frequent in <strong>the</strong><br />

western part <strong>of</strong> Ras Laffan Industrial City (JN 2004) where protected from grazing; very scattered<br />

in western <strong>Qatar</strong> in 2007. Batanouny and El Amin recorded mainly from S <strong>Qatar</strong>, where may still<br />

be locally abundant. Usually on thin, sandy or loamy deposits; <strong>of</strong>ten in rocky areas.<br />

uses: Widely documented as an important grazing plant. Several authors also mention its use for<br />

firewood. Used in <strong>the</strong> treatment <strong>of</strong> digestive disorders and for its antimicrobial properties (Rizk &<br />

El-Ghazaly).<br />

Photos: El Amin 38 L ; Bat 105 L , 106; Coll 213; Jong 185; Mand 203, 204; Phil 80; Shuaib 111; West 150.<br />

Senecio glaucus L.<br />

Senecio coronopifolius Desf.; Senecio desfontainei Druce<br />

Vernacular names: mureer<br />

ريرم<br />

Growth form: <strong>An</strong>nual herb. Fl. Jan–Mar.<br />

status: Native. Local. Also recorded: Bahrain, Kuwait, E Saudi Arabia, UAE.<br />

habitat & distribution: Occasional or locally frequent on sandy beaches and in cultivated<br />

depressions (Batanouny). Occurs at Al Wakra coast (El Amin), Ras Laffan (JN) and Umm Tais (RR).<br />

uses: Rizk & El-Ghazaly state that <strong>the</strong> plant contains alkaloids and is toxic to animals and humans.<br />

notes: The plant in <strong>the</strong> Gulf is S. glaucus subsp. coronopifolius (Maire) C. Alexander.<br />

Photos: El Amin 9; Coll 217; Corn 223; Jong 188; Mand 212, 213; Phil 66, 81; Shuaib 107; West 150.<br />

Senecio vulgaris L.<br />

Vernacular names: groundsel<br />

Growth form: <strong>An</strong>nual herb. Fl. Mar–Apr.<br />

status: Introduced. Local. Not in o<strong>the</strong>r Gulf floras.<br />

habitat & distribution: A weed <strong>of</strong> irrigated, cultivated places.<br />

uses: Medicinal uses are described by Rizk & El-Ghazaly.<br />

Photos: Coll 218.<br />

Sonchus asper (L.) Hill<br />

Vernacular names: prickly sowthistle<br />

Growth form: <strong>An</strong>nual herb. Fl. Mar–May.<br />

status: Introduced. Rare. Also recorded: Bahrain, E Saudi Arabia.<br />

habitat & distribution: <strong>An</strong> occasional weed <strong>of</strong> gardens and o<strong>the</strong>r cultivated places.<br />

Photos: Coll 219; Corn 224.<br />

Sonchus maritimus L.<br />

ASTERACEAE<br />

Growth form: Perennial herb. Fl. Apr–May.<br />

status: Introduced. Rare. Not in o<strong>the</strong>r Gulf floras.<br />

habitat & distribution: Reported from a farm near Al Khor by El Amin.<br />

notes: A typographical error in El Amin’s book (p. 41) inadvertently put this species into <strong>the</strong> genus<br />

Senecio, but <strong>the</strong> record in any case seems doubtful as this species is not recorded elsewhere in <strong>the</strong><br />

Gulf.<br />

Photos: None.<br />

12 13


ASTERACEAE<br />

Sonchus oleraceus L.<br />

Vernacular names: smooth sow-thistle, wild lettuce, aldheid<br />

Growth form: <strong>An</strong>nual or biennial herb. Fl. Feb–Apr.<br />

status: Introduced. Common. Also recorded: Bahrain, Kuwait, E Saudi Arabia, UAE.<br />

habitat & distribution: <strong>An</strong> occasional weed <strong>of</strong> cultivation.<br />

uses: The leaves and sap have numerous medicinal properties.<br />

Photos: Bat 106; Coll 219; Corn 224; Jong 190; Phil 82; Shuaib 109; West 151.<br />

Sonchus tenerrimus L.<br />

Growth form: <strong>An</strong>nual or biennial herb. Fl. Apr–May.<br />

status: Introduced. Rare. Also recorded: UAE.<br />

habitat & distribution: A weed <strong>of</strong> cultivation.<br />

Photos: Coll 220; Jong 191.<br />

Symphyotrichum squamatum (Spreng.) G.L. Nesom<br />

Aster squamatus (Spreng.) Hieron.<br />

نايوهج<br />

Growth form: Perennial herb. Fl. Mar–May.<br />

status: Introduced. Local. Also recorded: Bahrain, E Saudi Arabia.<br />

habitat & distribution: Frequent. Batanouny noted that it grew pr<strong>of</strong>usely as a weed in some<br />

irrigated rodat in N <strong>Qatar</strong>. <strong>An</strong> increasing species in <strong>the</strong> Middle East.<br />

notes: Shown on Plate 7 <strong>of</strong> El Amin, where labelled ‘Conyza maritima’, but <strong>the</strong>re is no text entry.<br />

Photos: El Amin 7; Bat 96; Coll 173; Corn 203.<br />

Vernacular names: jahwayan<br />

Tripleurospermum auriculatum (Boiss.) Rech. f.<br />

Matricaria auriculata (Boiss.) Muschl.<br />

Vernacular names: zafira, daqiqa, qaraiS<br />

Growth form: <strong>An</strong>nual herb. Fl. Apr.<br />

status: Native. Local. Also recorded: Bahrain, E Saudi Arabia.<br />

ديذلا<br />

صيرق ,ةقيقد ,هريفز<br />

habitat & distribution: Occasional in silty rodat in N and C <strong>Qatar</strong>, <strong>of</strong>ten under Acacia and Ziziphus.<br />

uses: ‘Said to be eaten with stews’ (El Amin).<br />

Photos: El Amin 9; Bat 107; Coll 222; Corn 218.<br />

Urospermum picroides (L.) F.W. Schmidt<br />

Growth form: <strong>An</strong>nual herb. Fl. Mar–Apr.<br />

status: Native. ?Rare. First published record: Abulfatih et al. (2001) (included in a list <strong>of</strong> species<br />

recorded). Also recorded: Bahrain, Kuwait, UAE.<br />

habitat & distribution: A plant <strong>of</strong> rocky areas.<br />

Photos: Coll 222; Jong 192; Shuaib 108; West 151.<br />

Xanthium spinosum L.<br />

Growth form: <strong>An</strong>nual herb. Fl. Jun.<br />

status: Introduced. Rare. Not in o<strong>the</strong>r Gulf floras.<br />

habitat & distribution: Cultivated fields; N <strong>Qatar</strong>.<br />

uses: The plant is used in <strong>the</strong> treatment <strong>of</strong> diabetes, rabies and o<strong>the</strong>r diseases and medical<br />

conditions; <strong>the</strong> leaves have diuretic properties. Poisoning <strong>of</strong> cattle from <strong>the</strong> plant has been reported<br />

(Rizk & El-Ghazaly).<br />

Photos: Coll 225.<br />

BORAGINACEAE<br />

<strong>An</strong>chusa hispida Forssk. Plate 24<br />

Gastrocotyle hispida (Forssk.) Bunge<br />

سيسبنع ,سمر<br />

Vernacular names: rims, ’anbasees<br />

Growth form: <strong>An</strong>nual herb. Fl. Mar–Apr.<br />

status: Native. Common. Also recorded: Bahrain, Kuwait, E Saudi Arabia, UAE.<br />

habitat & distribution: Occasional in sandy areas and silty depressions; especially N and C <strong>Qatar</strong>.<br />

Photos: Bat 84; Coll 88; Corn 173; Daoud 243; Jong 200; Phil 48; Shuaib 66; West 121.<br />

Arnebia decumbens (Vent.) Coss. & Kralik<br />

ASTERACEAE / BORAGINACEAE<br />

Vernacular names: Arabian primrose, kaHal<br />

لحاك<br />

Growth form: <strong>An</strong>nual herb. Fl. Feb–Apr.<br />

status: Native. Local. Also recorded: Bahrain, Kuwait, E Saudi Arabia, UAE.<br />

habitat & distribution: Occasional in C and W <strong>Qatar</strong>. Previously recorded from Salwa and Jebel<br />

Dukhan, on saline, sandy soils.<br />

uses: The deep red root is used by bedouin women as a cosmetic (rouge).<br />

Photos: El Amin 2; Coll 82; Corn 170; Daoud 237; Jong 201; Phil 45; Shuaib 66.<br />

Arnebia hispidissima (Lehm.) DC. Plate 22<br />

بنرلأا ةشيشح ,حيلم ,لحاك<br />

Vernacular names: Arabian primrose, Prophet flower,<br />

kaHil, melleiH, Hasheshat al ’arneb<br />

Growth form: <strong>An</strong>nual or biennial herb. Fl. Mar–Apr.<br />

status: Native. Very common. Also recorded: Bahrain, E Saudi Arabia, UAE.<br />

habitat & distribution: Abundant on shallow sandy deposits in all habitat types except saline<br />

areas. Perhaps more abundant in S and SW <strong>Qatar</strong>.<br />

uses: ‘Women formerly used <strong>the</strong> powdery red dust, that rubs <strong>of</strong>f <strong>the</strong> thick dark-red roots <strong>of</strong> <strong>the</strong>se<br />

plants, as a cosmetic’ (Cornes & Cornes). Root strongly red and probably used as a dye (Mandaville).<br />

The cosmetic use also recorded for <strong>the</strong> UAE. The whole plant is used for fevers including malaria<br />

(Ghazanfar).<br />

Photos: El Amin 2; Bat 83; Coll 83; Corn 170; Jong 202; Mand 171; Phil 46; Pick 108; West 120.<br />

Echiochilon jugatum I.M. Johnst. Plate 23<br />

Echiochilon kotschyi (Boiss. & Hohen.) I.M. Johnst.; Lithospermum kotschyi (Boiss. & Hohen.) I.M.<br />

Johnst.<br />

Vernacular names: stoneseed, najmat al bar<br />

ربلا ةمجن<br />

Growth form: Shrublet or small shrub. Fl. Feb–Apr.<br />

status: Native. Local. Also recorded: Bahrain, E Saudi Arabia, UAE.<br />

habitat & distribution: Occasional on compact soils at Ras Laffan and o<strong>the</strong>r coastal areas in <strong>the</strong><br />

NE (JN, MS).<br />

Photos: Bat 84; Coll 87; Corn 173; Jong 206; Phil 47; West 121.<br />

14 15


BORAGINACEAE<br />

Echium horridum Batt.<br />

Growth form: <strong>An</strong>nual herb. Fl. Mar–Apr.<br />

status: Native. Rare. Also recorded: E Saudi Arabia.<br />

habitat & distribution: Previously recorded from SW <strong>Qatar</strong> in sandy habitats.<br />

uses: Reported by El Amin to help urine complaints and to give relief from fever.<br />

Photos: None.<br />

Heliotropium bacciferum Forssk. Plate 25<br />

Heliotropium crispum Desf.; Heliotropium kotschyi (Bunge) Gürke<br />

Vernacular names: turnsole, heliotrope, ramram, dhanab al-’aqrab<br />

برقعلا بنذ ,مارمر<br />

Growth form: Shrub. Fl. Mar–Apr.<br />

status: Native. Very common. Also recorded: Bahrain, Kuwait, E Saudi Arabia, UAE.<br />

habitat & distribution: Frequent to abundant or sometimes dominant in sandy, <strong>of</strong>ten compacted<br />

soils, including coastal beach sand. Also in disturbed areas by roadsides and around towns and<br />

villages.<br />

uses: Dried powdered leaves are used as a poultice for abscesses, boils, sprains and swellings (Rizk<br />

& El-Ghazaly). Also used for treating ulcers, mouth blisters and snake bites (Ghazanfar).<br />

Photos: El Amin 2; Bat 85; Coll 89; Corn 174; Daoud 240; Jong 210; Mand 167; Phil 44, 51; Pick 160; Shuaib<br />

67; West 123.<br />

Heliotropium ramosissimum (Lehm.) Sieb.<br />

Vernacular names: ramram<br />

مارمر<br />

Growth form: <strong>An</strong>nual herb. Fl. Apr.<br />

status: Native. Local. Also recorded: Bahrain, E Saudi Arabia, UAE.<br />

habitat & distribution: Occasional on gravelly or sandy soils. Prevously recorded for N <strong>Qatar</strong>,<br />

but also found in <strong>the</strong> south.<br />

uses: Medicinal uses are similar to those recorded for H. bacciferum.<br />

Photos: Coll 91; Corn 177; Phil 52.<br />

Heliotropium zeylanicum (Burm. f.) Lam.<br />

Growth form: Herb. Fl. Not known.<br />

status: Introduced. ?Rare. First published record: Abdel Bari (1997). The salt-tolerant plant H.<br />

curassavicum L. is also very likely to occur in <strong>Qatar</strong> in gardens and farms. Not in o<strong>the</strong>r Gulf floras.<br />

habitat & distribution: A weed <strong>of</strong> cultivation.<br />

Photos: None.<br />

Lappula spinocarpos (Forssk.) Asch. ex Kuntze<br />

Growth form: <strong>An</strong>nual herb. Fl. Mar–Apr.<br />

status: Native. Local. Also recorded: Kuwait, E Saudi Arabia, UAE.<br />

habitat & distribution: Occasional in silty depressions and gravelly soils.<br />

Photos: Bat 86; Coll 93; Daoud 241, 242; Jong 217; Mand 169; Shuaib 68.<br />

Moltkiopsis ciliata (Forssk.) I.M. Johnst.<br />

Lithospermum callosum Vahl<br />

Plate 27<br />

Vernacular names: Halam/Halamah, ghabshah<br />

هشبغ ,ةملح<br />

Growth form: Shrublet or small shrub. Fl. Mar–Apr.<br />

status: Native. Common. Also recorded: Bahrain, Kuwait, E Saudi Arabia, UAE.<br />

habitat & distribution: Frequent on sand sheets, depressions and o<strong>the</strong>r sandy areas.<br />

Photos: Bat 86; Coll 93; Corn 178; Daoud 239; Jong 218; Phil 53; Shuaib 68; West 123.<br />

Ogastemma pusillum (Coss. & Durand ex Bonnet & Baratte) Brummitt Plate 26<br />

Growth form: <strong>An</strong>nual herb. Fl. Mar–Apr.<br />

status: Native. Common. Also recorded: Kuwait, E Saudi Arabia, UAE.<br />

habitat & distribution: Occasional to frequent on hard ground, including hamada and o<strong>the</strong>r<br />

rocky areas. Not mentioned by earlier authors and thought to have been discovered new to <strong>Qatar</strong> in<br />

2007; however it is probable that <strong>the</strong>re has been confusion with <strong>the</strong> very similar Lappula spinocarpos.<br />

Photos: Coll 94; Jong 219; Shuaib 68.<br />

BRASSICACEAE<br />

<strong>An</strong>astatica hierochuntica L. Plates 28, 29<br />

ةئيفق ,ناقرب ,ةمطاف عيمج ,ءارذعلا فك ,ميرم فك<br />

Vernacular names: rose <strong>of</strong> Jericho, virgin’s hand<br />

kaf Maryam, kaf al athra, jumay’ faTimah, birkan, qufay’ah<br />

Growth form: <strong>An</strong>nual herb. Fl. Mar–Apr.<br />

status: Native. Very common. Also recorded: Bahrain, Kuwait, E Saudi Arabia, UAE.<br />

habitat & distribution: Occasional to abundant in shallow sandy deposits, including runnels and<br />

small depressions. The plant is extremely common just west <strong>of</strong> Kharrarah (RR).<br />

uses: Its use as a charm and herbal aid in childbirth is widely reported. Batanouny mentions that<br />

it is sold in markets <strong>of</strong> <strong>Qatar</strong> as well as o<strong>the</strong>r Arabian countries. The dried plant is soaked in water<br />

and when it has unfurled <strong>the</strong> water is drunk by <strong>the</strong> expectant mo<strong>the</strong>r. Fur<strong>the</strong>r information is given<br />

in Phillips and Rizk & El-Ghazaly.<br />

Photos: El Amin 11; Bat 36; Coll 246; Corn 98; Daoud 87 L ; Ghaz 184, 185; Jong 223; Mand 84, 85; Phil<br />

142, 143; West 60.<br />

Brassica rapa L.<br />

Vernacular names: turnip<br />

Growth form: <strong>An</strong>nual herb. Fl. Mar–Apr.<br />

status: Introduced. Also recorded: E Saudi Arabia.<br />

habitat & distribution: Farms and cultivated areas.<br />

uses: A widely cultivated vegetable. Has been used in <strong>the</strong> treatment <strong>of</strong> various medical conditions<br />

including arthritis, chest-colds, dysentery, fever, mastitis, rheumatism, scurvy and skin ailments<br />

(Rizk & El-Ghazaly).<br />

Photos: Coll 248.<br />

Brassica tournefortii Gouan<br />

Vernacular names: qarraS, shiltam<br />

Growth form: <strong>An</strong>nual herb. Fl. Feb–Mar.<br />

status: ?Native. Also recorded: Kuwait, E Saudi Arabia, UAE.<br />

habitat & distribution: A weed <strong>of</strong> cultivation.<br />

Photos: Coll 249; Daoud 83, 84; Jong 224; Shuaib 60.<br />

BORAGINACEAE / BRASSICACEAE<br />

16 17<br />

متلش ,صارق


BRASSICACEAE<br />

Carrichtera annua (L.) DC.<br />

Growth form: <strong>An</strong>nual herb. Fl. Feb–Mar.<br />

status: Native. Rare. First record: SW <strong>Qatar</strong>, March 2007 (JN). Also recorded: Kuwait, E Saudi<br />

Arabia.<br />

habitat & distribution: A single plant in fruit growing in thin soil on a rocky summit.<br />

Photos: Coll 250; Daoud 94, 95, 96; Shuaib 63.<br />

Eremobium aegyptiacum (Spreng.) Asch. & Sweinf. ex Boiss. Plate 30<br />

Eremobium lineare (Delile) Boiss.<br />

Growth form: <strong>An</strong>nual herb. Fl. Feb–Mar.<br />

status: Native. Local. First record?: SW <strong>Qatar</strong>, March 2007 (JN & S. Aspinall). Miller & Cope<br />

mention a record for which <strong>the</strong>re is no supporting specimen. Also recorded: E Saudi Arabia, UAE.<br />

habitat & distribution: Occasional, mainly on compact, gravelly soils in SW <strong>Qatar</strong> (found in 2007<br />

at several scattered localities following winter rains). Elsewhere in <strong>the</strong> Gulf area it is a widespread<br />

plant <strong>of</strong> deeper sand, becoming locally abundant after rains. Gravel and sand forms have been<br />

noted by Collenette (see Miller & Cope).<br />

Photos: Coll 253; Jong 231; Mand 95; West 61.<br />

Eruca vesicaria (L.) Cav.<br />

Eruca sativa Mill.<br />

Vernacular names: rocket, jirjeer<br />

ريجرج<br />

Growth form: <strong>An</strong>nual or biennial herb. Fl. Feb–Apr.<br />

status: Introduced. Local. Also recorded: Bahrain, Kuwait, E Saudi Arabia, UAE.<br />

habitat & distribution: A weed in gardens and o<strong>the</strong>r cultivated areas.<br />

uses: Eaten in salads. The whole plant is considered an aphrodisiac and considered a cure for a<br />

wide range <strong>of</strong> medical conditions, including inflammations, ulcers, stomach-ache, epilepsy and<br />

toothache (Rizk & El-Ghazaly).<br />

Photos: Coll 253; Daoud 89; Jong 232; Shuaib 61.<br />

[Erucaria crassifolia (Forssk.) Delile]<br />

Growth form: <strong>An</strong>nual herb. Fl. Mar–Apr.<br />

status: Not confirmed for <strong>Qatar</strong>. Not in o<strong>the</strong>r Gulf floras.<br />

notes: Miller & Cope note that it is recorded from <strong>Qatar</strong> by Batanouny (1981) and <strong>the</strong> UAE by<br />

Western. However, <strong>the</strong>y remark that no verified material has been seen and <strong>the</strong> occurrence <strong>of</strong> <strong>the</strong><br />

species in Arabia needs confirmation. Batanouny reported it as growing ‘especially in rodat in N<br />

<strong>Qatar</strong>’.<br />

Photos: The plant identified as E. crassifolia is shown in Bat 37 and West 61.<br />

Erucaria hispanica (L.) Druce<br />

Erucaria lineariloba Boiss.<br />

Vernacular names: pink mustard<br />

Growth form: <strong>An</strong>nual herb. Fl. Mar–Apr.<br />

status: Native. Local. Also recorded: Bahrain, E Saudi Arabia, UAE.<br />

habitat & distribution: Recorded by El Amin from rodat in N <strong>Qatar</strong>.<br />

Photos: Coll 254; Jong 233; Phil 144; West 62.<br />

Farsetia heliophila Bunge ex Cosson Plate 31<br />

Farsetia hamiltonii Royle; Farsetia arabica Boulos<br />

Growth form: Perennial herb or shrublet. Fl. Mar–May.<br />

status: Native. Common. Also recorded: Bahrain, E Saudi Arabia, UAE.<br />

habitat & distribution: Occasional in thin sandy or silty deposits in stony or rocky areas. Frequent<br />

at Abu Nakhla sewage ponds (RR).<br />

notes: Records <strong>of</strong> Farsetia linearis Decne. ex Boiss. in <strong>the</strong> Gulf are in error for F. heliophila according<br />

to Miller & Cope.<br />

Photos: Corn 101; Jong 234; Phil 145.<br />

Lepidium aucheri Boiss.<br />

Vernacular names: rashad bari<br />

Growth form: <strong>An</strong>nual herb. Fl. Mar–Apr.<br />

status: Native. Rare. Also recorded: Kuwait, E Saudi Arabia.<br />

habitat & distribution: A species <strong>of</strong> silty depressions which have held rainwater.<br />

Photos: Coll 257; Daoud 84 L .<br />

18 19<br />

يرب داشر<br />

Lepidium didymum L.<br />

Coronopus didymus (L.) Sm.<br />

Vernacular names: swine-cress<br />

Growth form: <strong>An</strong>nual herb. Fl. All year.<br />

status: Introduced. Local. First published record: Abdel Bari (1997). Also recorded: E Saudi<br />

Arabia, UAE.<br />

habitat & distribution: This species is a recent colonist in <strong>the</strong> Gulf; it is only included in <strong>the</strong> most<br />

recent flora for <strong>the</strong> region (Jongbloed et al.). In <strong>Qatar</strong> it is likely to occur in irrigated lawns and<br />

roadsides.<br />

Photos: Jong 229.<br />

Lepidium sativum L.<br />

Vernacular names: garden cress<br />

Growth form: <strong>An</strong>nual herb. Fl. Jan–Mar.<br />

status: Introduced. ?Rare. First published record: Abdel Bari (1997). Also recorded: Kuwait.<br />

habitat & distribution: Recorded from a roadside.<br />

uses: A salad herb. Various medicinal uses are reported.<br />

Photos: Coll 258; Shuaib 61.<br />

Matthiola longipetala (Vent.) DC.<br />

Growth form: <strong>An</strong>nual herb. Fl. Feb–Mar.<br />

status: Native. Rare. Also recorded: Kuwait, E Saudi Arabia.<br />

habitat & distribution: Previously recorded from Al Wakra and Salwa.<br />

Photos: El Amin 50 L ; Coll 261; Daoud 108, 109, 11; Shuaib 62.<br />

Raphanus sativus L.<br />

Vernacular names: radish, fijl<br />

Growth form: <strong>An</strong>nual or biennial herb. Fl. Mar–Apr.<br />

BRASSICACEAE<br />

لجف


BRASSICACEAE<br />

status: Introduced. Also recorded: Bahrain, E Saudi Arabia.<br />

habitat & distribution: <strong>An</strong> escape from cultivation in farm areas.<br />

uses: Widely grown as a salad vegetable. A wide variety <strong>of</strong> medicinal benefits and uses are described.<br />

Photos: Coll 264; Corn 102.<br />

Savignya parviflora (Delile) Webb Plate 32<br />

Vernacular names: jirjees, kuljuman, jiljilan<br />

Growth form: <strong>An</strong>nual herb. Fl. Feb–Mar.<br />

status: Native. Common. Also recorded: Bahrain, Kuwait, E Saudi Arabia, UAE.<br />

نلاجلج ,نامجلك ,سيجرج<br />

habitat & distribution: Occasional to frequent on rocky ground and shallow, compacted sand;<br />

especially SW <strong>Qatar</strong>.<br />

uses: Rizk & El-Ghazaly report that in Bahrain <strong>the</strong> leaves are used as purgative and hypoglycaemic.<br />

Photos: El Amin 50 L ; Bat 37; Coll 264; Corn 105; Daoud 97, 98,; Jong 241; Mand 79; Phil 146; Shuaib 62;<br />

West 64.<br />

Schimpera arabica Hochst. & Steud. ex Steud.<br />

Schimpera persica Boiss.<br />

Growth form: <strong>An</strong>nual herb. Fl. Feb–Mar.<br />

status: Native. Rare. Also recorded: Kuwait, E Saudi Arabia, UAE.<br />

habitat & distribution: Usually occurs on silty sand; recorded from S <strong>Qatar</strong>.<br />

Photos: Coll 264; Daoud 102, 103; Mand 86, 87; Shuaib 63; West 65.<br />

Sinapis arvensis L.<br />

Vernacular names: wild mustard, khardal<br />

Growth form: <strong>An</strong>nual herb. Fl. Mar–Apr.<br />

status: Introduced. Rare. Also recorded: E Saudi Arabia, UAE.<br />

habitat & distribution: A weed <strong>of</strong> cultivated and irrigated areas.<br />

Photos: Coll 265; Jong 242.<br />

Sisymbrium erysimoides Desf.<br />

Vernacular names: thuwaT<br />

Growth form: <strong>An</strong>nual herb. Fl. Feb–Apr.<br />

status: Native. Local. Also recorded: E Saudi Arabia, UAE.<br />

habitat & distribution: Recorded from N and C <strong>Qatar</strong> in silty or sandy rodat; <strong>of</strong>ten associated<br />

with Ziziphus nummularia trees.<br />

Photos: El Amin 52 L ; Coll 265; Jong 243; Pick 110; West 65.<br />

Sisymbrium irio L.<br />

Growth form: <strong>An</strong>nual herb. Fl. Feb–Apr.<br />

status: Introduced. Also recorded: Bahrain, Kuwait, E Saudi Arabia, UAE.<br />

habitat & distribution: A weed <strong>of</strong> cultivation, irrigated soils and waste ground.<br />

Photos: Coll 266; Daoud 117; Jong 244; Shuaib 60.<br />

Sisymbrium orientale L.<br />

Growth form: <strong>An</strong>nual herb. Fl. Mar–May.<br />

لدرخ<br />

طاوث<br />

status: Introduced. Rare. Also recorded: UAE.<br />

habitat & distribution: A weed <strong>of</strong> lawns (Batanouny) and waste ground (Abdel Bari).<br />

Photos: El Amin 11; Bat 37; Coll 266.<br />

Zilla spinosa (L.) Prantl<br />

Vernacular names: shaja, silla, shubrum<br />

مربش ,لايس ,اجش<br />

Growth form: Small shrub. Fl. Mar–Apr.<br />

status: Native. Local. Also recorded: Kuwait, E Saudi Arabia, UAE.<br />

habitat & distribution: Occasional or frequent in SW <strong>Qatar</strong> in sandy and silty depressions.<br />

The village <strong>of</strong> Umm al Shubrum is named after <strong>the</strong> plant, which occurs in <strong>the</strong> area. Also found at<br />

Mekainis, Al Kharrarah and Umm Slal Mohammed (MS).<br />

uses: ‘A useful remedy in <strong>the</strong> treatment <strong>of</strong> ailments such as kidney stones’ (Rizk & El-Ghazaly).<br />

Photos: Coll 267; Daoud 93; Jong 245; Mand 76; West 66.<br />

CAPPARACEAE<br />

Capparis spinosa L.<br />

Vernacular names: common caper, shafallaH<br />

Growth form: Shrub. Fl. Mar.<br />

status: Native. Common. Also recorded: Bahrain, E Saudi Arabia, UAE.<br />

habitat & distribution: Occasional in stony and rocky areas and compact silty soils in depressions<br />

and by roads; mainly in NE <strong>Qatar</strong>.<br />

uses: Buds are cooked and pickled and used as flavouring in cooking (El Amin). The leaves <strong>of</strong> <strong>the</strong><br />

plant are used for treating earache, coughs, expelling stomach worms and for diabetes. (Ghazanfar).<br />

Rizk & El-Ghazaly also list numerous o<strong>the</strong>r medicinal uses.<br />

notes: There are two widespread varieties in Arabia: var. spinosa (C. aegyptiaca Lam., C.s. var.<br />

aegyptia) and var. mucronifolia (Boiss.) Hedge & Lamond. For <strong>Qatar</strong> earlier authors recorded var.<br />

spinosa (<strong>the</strong> form <strong>of</strong>ten seen by roadsides), but Miller & Cope also list var. mucronifolia (<strong>of</strong> rocky<br />

areas).<br />

Photos: El Amin 3; Bat 33 L ; Coll 108; Corn 106; Ghaz 165; Jong 248; Mand 69, 70; Phil 57, 58; West 57.<br />

Cleome amblyocarpa Barratte & Murb.<br />

Cleome africana Botsch.<br />

Growth form: <strong>An</strong>nual herb. Fl. Apr.<br />

status: Native. Local. Also recorded: E Saudi Arabia, UAE.<br />

habitat & distribution: Occasional in sandy saline depressions.<br />

uses: Jongbloed et al. note its use to treat abdominal and rheumatic pains.<br />

Photos: Coll 109; Jong 249.<br />

Cleome brachycarpa Vahl ex DC.<br />

Growth form: <strong>An</strong>nual or short-lived perennial. Fl. Dec–Mar.<br />

status: Native. Rare. Also recorded: UAE.<br />

habitat & distribution: Recorded from Al Khor by Batanouny.<br />

Photos: Bat 34; Coll 110; Ghaz 170; Jong 251; West 57.<br />

BRASSICACEAE / CAPPARACEAE<br />

20 21<br />

حلفش


CAPPARACEAE / CARyOPHyLLACEAE<br />

Cleome noeana Boiss.<br />

Cleome dolichostyla Jafri<br />

Growth form: <strong>An</strong>nual or perennial herb, or small shrub. Fl. May–Aug.<br />

status: Native. Rare. First published record: Miller & Cope (1996). Also recorded: UAE.<br />

habitat & distribution: Gravel desert on <strong>the</strong> north coast (Miller & Cope).<br />

notes: Only subsp. noeana occurs in <strong>Qatar</strong>; subsp. brachystyla (Deflers) Chamberlain & Lamond is<br />

found elsewhere in Arabia. The plant named as C. dolichostyla Jafri in Arabia is included within C.<br />

noeana by Miller & Cope.<br />

Photos: Jong 253, West 58.<br />

Cleome scaposa DC.<br />

Vernacular names: DHefra, zefra<br />

Growth form: <strong>An</strong>nual or short-lived perennial. Fl. Apr.<br />

status: Native. Local. Also recorded: UAE.<br />

ةرفز ,ةرفظ<br />

habitat & distribution: Rocky places; previously recorded N <strong>of</strong> Doha by Batanouny and recently<br />

in a few sites in NE <strong>Qatar</strong> (MS).<br />

Photos: Bat 35; Coll 112; Ghaz 169; Jong 255.<br />

Dipterygium glaucum Decne. Plate 33<br />

Vernacular names: ’alqa<br />

ةقلع<br />

Growth form: Perennial herb or shrublet. Fl. Mar.<br />

status: Native. Local. First record: SW <strong>Qatar</strong>, March 2007 (JN and S. Aspinall). Also recorded: E<br />

Saudi Arabia, UAE.<br />

habitat & distribution: In moderately firm sand mixed with gravel, and similar habitats. A species<br />

also <strong>of</strong> stable sand dunes in some o<strong>the</strong>r parts <strong>of</strong> <strong>the</strong> Gulf.<br />

uses: ‘Of some importance for grazing in more remote regions’ (Mandaville).<br />

Photos: Coll 113; Ghaz 176-8; Jong 256; Pick 117; West 59.<br />

CARYOPHYLLACEAE<br />

Herniaria hemistemon J. Gay Plate 34<br />

Vernacular names: um waja’ alkabid, ghebayra<br />

ةريبغ ,دبكلا عجو مأ<br />

Growth form: Perennial herb. Fl. Feb–Apr.<br />

status: Native. Very common. Also recorded: Bahrain, Kuwait, E Saudi Arabia, UAE.<br />

habitat & distribution: Frequent on compact or thin soils over most <strong>of</strong> <strong>Qatar</strong>.<br />

uses: ‘Good fodder’ (El Amin). The scientific name refers to its use in treating hernias (mentioned<br />

for Europe by Jongbloed et al.). It is used in Bahrain as a diuretic and purgative (Rizk & El-Ghazaly).<br />

Photos: Bat 20; Coll 119; Corn 62; Daoud 146; Jong 264; Mand 24; Phil 150; Shuaib 95; West 39.<br />

Herniaria hirsuta L.<br />

Vernacular names: ’eysh shawlah<br />

Growth form: <strong>An</strong>nual herb. Fl. Mar.<br />

status: Native. Rare. Also recorded: Bahrain, Kuwait, E Saudi Arabia, UAE.<br />

habitat & distribution: Recorded only once on Jebel Dukhan by El Amin.<br />

هلوش شيع<br />

uses: Used as a diuretic and astringent (Batanouny); a powerful narcotic and stomach irritant (Rizk<br />

& El-Ghazaly).<br />

Photos: Coll 120; Shuaib 95.<br />

Paronychia arabica (L.) DC. Plate 35<br />

Growth form: <strong>An</strong>nual herb. Fl. Mar–Apr.<br />

status: Native. Common. Also recorded: Bahrain, Kuwait, E Saudi Arabia, UAE.<br />

habitat & distribution: Occasional on shallow sandy soils.<br />

uses: ‘The entire plant is used as a stimulant and aphrodisiac’ (Rizk & El-Ghazaly).<br />

Photos: Bat 21; Coll 122; Corn 62; Daoud 145; Jong 265; Phil 151; Shuaib 95; West 39.<br />

Polycarpaea repens (Forssk.) Asch. & Schweinf. Plate 36<br />

Vernacular names: kameela, makhor<br />

روخام ,ةليمك<br />

Growth form: Perennial herb. Fl. Mar–Apr.<br />

status: Native. Very common. Also recorded: Bahrain, Kuwait, E Saudi Arabia, UAE.<br />

habitat & distribution: Frequent or abundant on sand sheets, gravel plains and o<strong>the</strong>r sandy<br />

habitats.<br />

uses: Mandaville remarks that <strong>the</strong> plant is traditionally used by one <strong>of</strong> <strong>the</strong> bedouin tribes to treat<br />

mange <strong>of</strong> camels. Ghazanfar records its use as an antidote for snake bite.<br />

Photos: Coll 123; Corn 65; Daoud 142, 143; Ghaz 67; Jong 266; Phil 152; Shuaib 96; West 40.<br />

Polycarpaea robbairea (Kuntze) Greuter & Burdet<br />

Robbairea delileana Milne-Redh.<br />

Growth form: Perennial herb. Fl. Feb–Apr.<br />

status: Native. Local. Also recorded: Kuwait, E Saudi Arabia, UAE.<br />

habitat & distribution: El Amin recorded from Umm Slal Ali and Al Kharrarah on gravelly and<br />

stony soils.<br />

Photos: Bat 22; Coll 123; Daoud 144; Ghaz 69; Jong 267; Shuaib 96.<br />

Polycarpaea spicata Wight ex Arn.<br />

Vernacular names: daqeeqa<br />

Growth form: <strong>An</strong>nual herb. Fl. Mar–Apr.<br />

status: Native. Local. Also recorded: Bahrain, UAE.<br />

habitat & distribution: Occasional on sandy and gravelly soils, including disturbed soils around<br />

towns.<br />

Photos: Bat 21; Coll 124; Ghaz 68; Jong 268.<br />

Polycarpon tetraphyllum (L.) L.<br />

Growth form: <strong>An</strong>nual herb. Fl. Feb–Mar.<br />

status: Introduced. Rare. Also recorded: Kuwait, E Saudi Arabia, UAE.<br />

habitat & distribution: A weed <strong>of</strong> gardens, usually on sandy soils.<br />

Photos: Coll 124; Daoud 140, 141; Jong 269; Shuaib 96.<br />

CARyOPHyLLACEAE<br />

22 23<br />

ةقيقد


CARyOPHyLLACEAE<br />

Sclerocephalus arabicus Boiss. Plate 38<br />

Vernacular names: knucklehead weed, thrais, Haras<br />

سرح ,سيرث<br />

Growth form: <strong>An</strong>nual herb. Fl. Mar.<br />

status: Native. Very common. Also recorded: Bahrain, Kuwait, E Saudi Arabia, UAE.<br />

habitat & distribution: Abundant, <strong>of</strong>ten locally dominant. Always on hard surfaces, including<br />

gravel plains, stony and and rocky areas, where <strong>the</strong>re is a thin covering <strong>of</strong> silt or sand.<br />

uses: El Amin remarks that it is not liked by animals due to its bristles.<br />

Photos: Bat 22; Coll 125; Corn 65; Daoud 147; Jong 270; Mand 25; Phil 153; Shuaib 97; West 40.<br />

Silene arabica Boiss.<br />

Growth form: <strong>An</strong>nual herb. Fl. Mar–Apr.<br />

status: Native. Rare. Also recorded: Kuwait, E Saudi Arabia.<br />

habitat & distribution: Sandy habitats; N <strong>Qatar</strong>.<br />

Photos: Coll 126; Daoud 134, 135; Mand 18; Shuaib 98.<br />

Silene conica L.<br />

Growth form: <strong>An</strong>nual herb. Fl. Not known.<br />

status: Native. Rare. First published record: Abdel Bari (1997). Also recorded: Kuwait.<br />

habitat & distribution: Recorded along <strong>the</strong> road to Umm Bab; also Dukhan and C <strong>Qatar</strong>.<br />

Photos: None.<br />

Silene villosa Forssk. Plate 39<br />

Vernacular names: desert campion, Turbah<br />

ةبرط<br />

Growth form: <strong>An</strong>nual herb. Fl. Feb–Mar.<br />

status: Native. Very common. Also recorded: Bahrain, Kuwait, E Saudi Arabia, UAE.<br />

habitat & distribution: Occasional to frequent in sandy habitats. Not rare as noted by Batanouny,<br />

but only numerous after rains.<br />

notes: Miller and Cope (1996, p. 219) distinguish two forms ‘A’ and ‘B’ in Arabia, <strong>of</strong> which only<br />

form B is recorded for <strong>Qatar</strong>.<br />

Photos: El Amin 3; Bat 23; Coll 130; Daoud 133; Ghaz 72; Jong 274; Mand 20; Phil 154; Shuaib 99; West 41.<br />

Spergula fallax (Lowe) E.H.L. Krause Plate 37<br />

Vernacular names: daqeeqa<br />

Growth form: <strong>An</strong>nual herb. Fl. Mar–Apr.<br />

status: Native. Common. Also recorded: Bahrain, Kuwait, E Saudi Arabia, UAE.<br />

habitat & distribution: Frequent or abundant in sandy or silty depressions, particularly under<br />

shade <strong>of</strong> trees, but also in o<strong>the</strong>r habitats.<br />

Photos: Bat 23; Coll 132; Daoud 116 L ; Ghaz 70; Jong 275; Mand 21; Pick 167; West 42.<br />

Spergularia bocconei (Scheele) Asch. & Graebn.<br />

Growth form: <strong>An</strong>nual or biennial herb. Fl. Mar–May.<br />

status: Native. Common. Also recorded: Bahrain, UAE.<br />

habitat & distribution: Frequent in sandy or silty depressions, particularly under shade <strong>of</strong> trees.<br />

Photos: Coll 132; Corn 66.<br />

ةقيقد<br />

Spergularia diandra (Guss.) Boiss.<br />

Vernacular names: umm threib, qaleiqah<br />

ةقيلق ,بيرث مأ<br />

Growth form: <strong>An</strong>nual herb. Fl. Feb–Mar.<br />

status: Native. Very common. Also recorded: Bahrain, Kuwait, E Saudi Arabia, UAE.<br />

habitat & distribution: Abundant on thin sandy deposits on hamada and o<strong>the</strong>r rocky areas; also<br />

in o<strong>the</strong>r sandy habitats, gardens and cultivated areas.<br />

Photos: Coll 132; Daoud 138, 139; Jong 276; Mand 22; Shuaib 97.<br />

Stellaria media (L.) Vill.<br />

Vernacular names: chickweed<br />

Growth form: <strong>An</strong>nual herb. Fl. Mar–Apr.<br />

status: Introduced. Rare. Also recorded: E Saudi Arabia, UAE.<br />

habitat & distribution: A weed <strong>of</strong> gardens and o<strong>the</strong>r irrigated, cultivated areas.<br />

uses: Rizk & El-Ghazaly list numerous medicinal uses.<br />

Photos: Coll 133; Jong 279.<br />

Stellaria pallida (Dumort.) Murb.<br />

Vernacular names: lesser chickweed<br />

Growth form: <strong>An</strong>nual herb. Fl. Jan–Mar.<br />

status: ?Native. Uncertain. Also recorded: Kuwait.<br />

habitat & distribution: A species <strong>of</strong> moist ground.<br />

notes: The only Stellaria species listed for <strong>Qatar</strong> by Miller & Cope, who comment on <strong>the</strong> difficulty<br />

in identifying this and S. media with certainty.<br />

Photos: None.<br />

Vaccaria hispanica (Mill.) Rauschert<br />

Vaccaria pyramidata Medik.<br />

Growth form: <strong>An</strong>nual herb. Fl. Mar–Apr.<br />

status: Introduced. Rare. Also recorded: Kuwait, E Saudi Arabia.<br />

habitat & distribution: Recorded from Al Khor by El Amin.<br />

Photos: Coll 134; Shuaib 98.<br />

CASUARINACEAE<br />

CARyOPHyLLACEAE / CASUARINACEAE<br />

Casuarina equisetifolia L.<br />

Vernacular names: Australian pine, she-oak<br />

Growth form: Tree. Fl. Not known.<br />

status: Introduced. Common. Not in o<strong>the</strong>r Gulf floras.<br />

habitat & distribution: Mainly gardens and roadsides in towns and cities.<br />

uses: Widely planted for ornament and shade.<br />

notes: Several species are planted in <strong>the</strong> Gulf (Miller & Cope); C. equisetifolia L. probably being <strong>the</strong><br />

more common and widespread. C. cunninghamiana Miq. is listed by El Amin (p. 149) as planted in<br />

<strong>Qatar</strong> and <strong>the</strong> hybrid with C. equisetifolia has been recorded in Kuwait.<br />

Photos: None.<br />

24 25


CHENOPODIACEAE<br />

CHENOPODIACEAE<br />

Agathophora alopecuroides (Delile) Fenzl. ex Bunge<br />

Agathophora iraqensis Botsch.; Halogeton alopecuroides (Delile) Moq.; Salsola alopecuroides Delile<br />

Growth form: Small shrub. Fl. Mar–Apr.<br />

status: Native. Rare. Also recorded: E Saudi Arabia.<br />

habitat & distribution: Recorded at Al Khor and Umm Slal Ali.<br />

notes: Two varieties occur in Arabia, var. alopecuriodes and var. papillosa (Maire) Boulos; both may<br />

grow in <strong>Qatar</strong>.<br />

Photos: Bat 26; Mand 49.<br />

Agriophyllum minus Fisch. & Mey. Plate 43<br />

Agriophyllum montasiri El-Gazzar<br />

Growth form: <strong>An</strong>nual herb. Fl. Apr.<br />

status: Native. Local. Also recorded: E Saudi Arabia, UAE.<br />

habitat & distribution: Occasional in saline sandy areas on <strong>the</strong> coast and in ‘white sand’ areas<br />

inland. So far recorded only from SW <strong>Qatar</strong>.<br />

Photos: El Amin 16 L ; Coll 139; Jong 281.<br />

<strong>An</strong>abasis setifera Moq. Plate 40<br />

Vernacular names: hamD al ’arnab, sha’ran<br />

ناريعش ,بنرلأا ضمح<br />

Growth form: Small shrub. Fl. Aug–Nov.<br />

status: Native. Common. Also recorded: Bahrain, Kuwait, E Saudi Arabia, UAE.<br />

habitat & distribution: Widespread on harder substrates including compacted, moderately<br />

saline, sandy soils including sabkha edge.<br />

uses: El Amin states that it is known for good salt grazing for camels and o<strong>the</strong>r animals.<br />

Photos: El Amin 3; Bat 24; Coll 140; Corn 70; Daoud 191, 192, 19; Ghaz 53; Jong 282; Pick 204; Shuaib<br />

75; West 45.<br />

Arthrocnemum macrostachyum (Moric.) K. Koch<br />

Arthrocnemum glaucum (Delile) Ung.-Sternb.<br />

Vernacular names: shinan, ’ajram, HamaD<br />

ضمح ,مرجع ,نانش<br />

Growth form: Shrub. Fl. Mar–May.<br />

status: Native. Common. Also recorded: Bahrain, E Saudi Arabia, UAE.<br />

habitat & distribution: Abundant in saltmarsh along coasts at Al Khor, Ras Eshairiq, Umm Bab<br />

and Dukhan. Also locally dominant on coastal sabkha at Ras Abrouq.<br />

uses: Dry stems are used as firewood.<br />

Photos: El Amin 4; Bat 24; Coll 141; Corn 82; Ghaz 37, 38; Jong 283; Pick 205; West 45.<br />

Atriplex leucoclada Boiss. Plate 41<br />

Vernacular names: orache, rughl<br />

لغر<br />

Growth form: Shrub. Fl. Oct–Nov.<br />

status: Native. Local. Also recorded: Bahrain, Kuwait, E Saudi Arabia, UAE.<br />

habitat & distribution: Occasional in coastal areas.<br />

uses: Grazed by sheep and camels and is valuable salt grazing (El Amin). Cornes & Cornes note<br />

that ‘Atriplex species are a good source <strong>of</strong> fodder, with high protein content’.<br />

notes: Two forms are recorded for Arabia: var inamoena (Aellen) Zoh. and var. turcomanica (Moq.)<br />

Zoh. Only <strong>the</strong> latter is confirmed for <strong>Qatar</strong> by Miller & Cope.<br />

Photos: El Amin 4; Bat 25; Coll 142; Corn 73; Daoud 162, 163; Ghaz 33; Jong 284; Mand 28; Phil 96; Shuaib<br />

78; West 46.<br />

Bassia eriophora (Schrad.) Asch.<br />

Vernacular names: quTn<br />

Growth form: <strong>An</strong>nual herb. Fl. Apr–May.<br />

status: Native. Rare. Also recorded: Bahrain, Kuwait, E Saudi Arabia.<br />

habitat & distribution: El Amin recorded it near Al Khor.<br />

uses: El Amin remarks that <strong>the</strong> white fibres are used as cotton stuffings.<br />

Photos: El Amin 4; Coll 143; Corn 74; Daoud 165, 166, 16; Mand 29; Shuaib 78.<br />

Bassia muricata (L.) Asch.<br />

Vernacular names: jebira, quTn<br />

Growth form: <strong>An</strong>nual herb. Fl. Mar–Apr.<br />

status: Native. Rare. Also recorded: Kuwait, E Saudi Arabia, UAE.<br />

26 27<br />

نطق<br />

نطق ,ةريبج<br />

habitat & distribution: Occasional in nor<strong>the</strong>rn <strong>Qatar</strong>. Occurs in <strong>the</strong> Dukhan area on compacted<br />

gypsum soils (JN).<br />

uses: Grazed and liked by camels, sheep and o<strong>the</strong>r livestock.<br />

Photos: Coll 144; Daoud 139 L ; Jong 285; Shuaib 78.<br />

Beta vulgaris L.<br />

CHENOPODIACEAE<br />

Vernacular names: beet, salaq, shamandar, bari<br />

يرب ,ردنمش ,قلس<br />

Growth form: Perennial herb. Fl. Mar.<br />

status: Introduced. ?Common. Also recorded: E Saudi Arabia, UAE.<br />

habitat & distribution: <strong>An</strong> escape from cultivation; <strong>the</strong> wild form occurs in saltmarshes but has<br />

not been recorded from <strong>Qatar</strong>.<br />

uses: Cultivated as a vegetable. The plant has diuretic, aphrodisiac and various o<strong>the</strong>r medicinal<br />

properties (Rizk & El-Ghazaly).<br />

notes: Beta vulgaris subsp. vulgaris is <strong>the</strong> cultivated garden beet and B. vulgaris subsp. maritima (L.)<br />

Arcang. is <strong>the</strong> wild sea beet.<br />

Photos: Coll 144; Jong 286.<br />

Bienertia cycloptera Bunge ex Boiss.<br />

Growth form: Perennial herb. Fl. Sep–Oct.<br />

status: Native. ?Rare. First record: Mesaieed, November 1998 (JN). Also recorded: Kuwait, E Saudi<br />

Arabia, UAE.<br />

habitat & distribution: A species <strong>of</strong> coastal saltmarshes.<br />

notes: A specimen collected from <strong>the</strong> coast at Mesaieed was provisionally determined as this<br />

species by E. Clement, but dried specimens <strong>of</strong> fleshy-leaved Chenopodiaceae are very difficult to<br />

identify. It is not known to have been previously recorded from <strong>Qatar</strong>, but is widespread elsewhere<br />

along <strong>the</strong> Gulf coast and may have been overlooked at o<strong>the</strong>r sites in <strong>Qatar</strong>.<br />

Photos: Coll 144; Daoud 175,176,17; Jong 287; Mand 35; Shuaib 76.


CHENOPODIACEAE<br />

Chenopodium album L.<br />

Vernacular names: shawlah, siamjat al reiH, weraq, zarbekh<br />

خيبرز ,قارو ,حيرلا تاجميس ,هلوش<br />

fat hen, white goosefoot<br />

Growth form: <strong>An</strong>nual herb. Fl. Apr–May.<br />

status: Introduced. Common. Also recorded: Kuwait, E Saudi Arabia, UAE.<br />

habitat & distribution: A frequent weed <strong>of</strong> cultivated and irrigated areas.<br />

uses: Various medicinal uses have been documented.<br />

Photos: Bat 25; Coll 145; Jong 288; Shuaib 78; West 47.<br />

Chenopodium murale L.<br />

Vernacular names: zarbeeH, khaisa, weraq, abu ’affeyn<br />

نيفع وبأ ,قارو ,ةسياخ ,حيبرز<br />

fat hen, white goosefoot<br />

Growth form: <strong>An</strong>nual herb. Fl. Feb–Apr.<br />

status: Introduced. Common. Also recorded: Bahrain, Kuwait, E Saudi Arabia, UAE.<br />

habitat & distribution: Occasional in disturbed areas, especially around towns and villages.<br />

Photos: El Amin 4; Bat 25; Coll 146; Corn 77; Daoud 161; Ghaz 31; Jong 289; Phil 97; Pick 207; Shuaib<br />

78; West 47.<br />

Cornulaca aucheri Moq.<br />

Vernacular names: hadh, saley<br />

Growth form: <strong>An</strong>nual herb. Fl. Oct–Nov.<br />

status: Native. Common. Also recorded: Bahrain, Kuwait, E Saudi Arabia, UAE.<br />

habitat & distribution: Coastal areas, including Umm Bab, Salwa and Al Khor.<br />

Photos: Bat 26; Coll 147; Corn 77; Ghaz 44; Jong 291; Shuaib 77.<br />

Cornulaca monacantha Delile<br />

Vernacular names: thallaj, hadh<br />

Growth form: Small shrub. Fl. Oct–Nov.<br />

status: Native. Common. Also recorded: Bahrain, E Saudi Arabia, UAE.<br />

habitat & distribution: Frequent in coastal areas and in S <strong>Qatar</strong> on deeper saline sand.<br />

يلس ,ذاح<br />

ذاح ,جلاث<br />

uses: Reported by Rizk & El-Ghazaly to have some degree <strong>of</strong> antimicrobial activity when ingested.<br />

Leaves are used to treat jaundice in <strong>Qatar</strong> (Ghazanfar).<br />

Photos: El Amin 5; Bat 26; Coll 148; Corn 77; Ghaz 45; Jong 280 L , 292; West 48.<br />

Halocnemum strobilaceum (Pall.) M. Bieb.<br />

Vernacular names: sabat, hadhadi<br />

يداذح ,تبس<br />

Growth form: Shrub. Fl. Sep–Oct.<br />

status: Native. Local. Also recorded: Bahrain, Kuwait, E Saudi Arabia, UAE.<br />

habitat & distribution: Recorded by Batanouny from saltmarsh at Al Khor and Abu Samrah.<br />

uses: Cornes & Cornes report that <strong>the</strong> plant is intensively grazed by camels in sabkha areas and also<br />

by gazelles; Mandaville (citing an earlier author) mentions a herdsmen’s belief that overfeeding on<br />

this plant is a cause <strong>of</strong> lung disease in camels.<br />

Photos: Coll 149; Corn 85; Daoud 168, 169; Jong 290, 293; Mand 30; Phil 98; Shuaib 75; West 48.<br />

Halopeplis perfoliata (Forssk.) Bunge ex Schweinf. & Asch. Plate 42<br />

زيرخ<br />

Vernacular names: khurreyz (string <strong>of</strong> beads)<br />

Growth form: Shrub. Fl. Mar.<br />

status: Native. Local. Also recorded: Bahrain, E Saudi Arabia, UAE.<br />

habitat & distribution: Locally abundant in saltmarsh and sabkha edge, or occasionally on saline<br />

sand; mainly N <strong>Qatar</strong> and <strong>the</strong> E coast.<br />

Photos: El Amin 5; Bat 27; Coll 149; Corn 38, 85; Ghaz 35, 36; Jong 280 L , 290, 294; Phil 99; Pick 57; West 49.<br />

Haloxylon persicum Bunge<br />

Haloxylon ammodendron (C.A. Meyer) Bunge<br />

CHENOPODIACEAE<br />

Vernacular names: ghada, qadha, rimth<br />

ثمر ,ىذق<br />

Growth form: Shrub or small tree. Fl. Feb–May.<br />

status: Native. Rare. Also recorded: Bahrain, E Saudi Arabia, UAE.<br />

habitat & distribution: A characteristic species <strong>of</strong> deep, <strong>of</strong>ten drifting sand. Recorded only by El<br />

Amin, but, confusingly, he refers to this species as Haloxylon salicornicum (and what is now known<br />

as H. salicornicum under <strong>the</strong> older name <strong>of</strong> Hammada elegans). He recorded it east <strong>of</strong> Salwa at Mah<br />

Sobia hill (<strong>the</strong> area now known as Al Mashbiya), near <strong>the</strong> Saudi Arabian border.<br />

uses: Grazed by camels; highly valued as firewood by bedouin and desert villagers (Mandaville).<br />

Photos: El Amin 5; Coll 150; Jong 296; West 49.<br />

Haloxylon salicornicum (Moq.) Bunge ex Boiss. Plate 44<br />

Hammada elegans (Bunge) Botsch.; Hammada salicornica (Moq.) Iljin<br />

ثمر<br />

Vernacular names: rimth<br />

Growth form: Shrub. Fl. Sep–Oct.<br />

status: Native. Local. Also recorded: Bahrain, Kuwait, E Saudi Arabia, UAE.<br />

habitat & distribution: Abundant in parts <strong>of</strong> SW <strong>Qatar</strong> in sandy areas between Abu Samrah and<br />

Al Karaanah. Also found at Khor Al Adaid, but local elsewhere. <strong>An</strong> area has recently developed at<br />

<strong>the</strong> Al Sheehaniyah racecourse in C <strong>Qatar</strong>.<br />

uses: Good fodder for camels. O<strong>the</strong>r animals graze on it when nothing else is around’ (El Amin).<br />

Mandaville notes that it is important camel grazing when annuals and grasses are not available<br />

but remarks: ‘excessive saltbush grazing is recognised to be physiologically damaging’. A source<br />

<strong>of</strong> firewood. A relatively small number <strong>of</strong> medicinal uses are documented; stems are used to treat<br />

hypoglycaemia (Ghazanfar).<br />

Photos: El Amin 5; Bat 29; Coll 151; Corn 78; Daoud 188, 189, 19; Ghaz 46, 47; Jong 297; Phil 100; Shuaib<br />

74; West 50.<br />

Salicornia europaea L. Plate 45<br />

Salicornia maritima Wolff & Jefferies; Salicornia herbacea L.<br />

Vernacular names: glasswort<br />

Growth form: <strong>An</strong>nual herb. Fl. Nov.<br />

status: Native. Rare. First published record: Abdel Bari (1997). Also recorded: Bahrain, Kuwait.<br />

habitat & distribution: Locally abundant at Al Khor; also at Umm Tais (RR).<br />

notes: Batanouny undoubtedly recorded Salsola soda L. in error for this species.<br />

Photos: Daoud 170, 171; Phil 101; Shuaib 76.<br />

28 29


CHENOPODIACEAE<br />

Salsola cyclophylla Baker<br />

Growth form: Small shrub. Fl. Oct–Dec.<br />

status: Native. Rare. Also recorded: Bahrain, Kuwait, E Saudi Arabia, UAE.<br />

habitat & distribution: Recorded by Batanouny in S <strong>Qatar</strong> ‘where it dominates a community on<br />

<strong>the</strong> road to <strong>the</strong> UAE’. Typical habitats in Saudi Arabia include sand dunes, rocky slopes, limestone<br />

and sandstone plateaux (Miller & Cope).<br />

Photos: Coll 153; Jong 298; Mand 40.<br />

Salsola drummondii Ulbr.<br />

Growth form: Shrub. Fl. Oct–Dec.<br />

status: Native. ?Rare. First published record: Miller & Cope (1996). Also recorded: E Saudi Arabia,<br />

UAE.<br />

habitat & distribution: In o<strong>the</strong>r parts <strong>of</strong> <strong>the</strong> Gulf this is a species <strong>of</strong> saline sandy areas, including<br />

sabkha; usually near <strong>the</strong> coast.<br />

Photos: Jong 299; Pick 58.<br />

Salsola imbricata Forssk. Plate 46<br />

Salsola baryosma (Roem. & Schult.) Dandy; Salsola foetida Delile ex Spreng.<br />

طيرخ<br />

Vernacular names: saltwort, khareeT<br />

Growth form: Small shrub. Fl. Jun–Sep.<br />

status: Native. Common. Also recorded: Bahrain, Kuwait, E Saudi Arabia, UAE.<br />

habitat & distribution: Frequent or abundant in coastal saline areas, including beaches and<br />

sabkha, occasionally in o<strong>the</strong>r habitats.<br />

uses: ‘In Bahrain, <strong>the</strong> flowers are used as a diuretic and anti-inflammatory’ Rizk & El-Ghazaly.<br />

notes: Plate 46 shows a Salsola species, possibly S. imbricata.<br />

Photos: El Amin 6; Coll 153; Corn 39, 74, 78; Daoud 180, 181; Ghaz 51; Jong 280 L , 300; Phil 102; Shuaib<br />

77; West 50.<br />

Salsola schweinfurthii Solms-Laub.<br />

Growth form: Small shrub. Fl. Not known.<br />

status: Native. ?Local. Also recorded: E Saudi Arabia, UAE.<br />

habitat & distribution: Occasional in saline areas in S <strong>Qatar</strong> according to Batanouny.<br />

notes: Needs confirmation as not listed for <strong>Qatar</strong> by Miller & Cope.<br />

Photos: Coll 154.<br />

Salsola villosa Schultes<br />

Salsola mandavillei Botsch.; Salsola vermiculata sensu Mandaville, non L.; Salsola vermiculata var.<br />

villosa (Schultes) Moq.<br />

Growth form: Small shrub. Fl. Mar.<br />

status: Native. ?Local. Also recorded: Bahrain, E Saudi Arabia.<br />

habitat & distribution: Recorded by Batanouny from C and N <strong>Qatar</strong> in calcareous stony soils<br />

and saline areas.<br />

uses: Favourite grazing for camels (Mandaville). Salt grazing (El Amin). Often used as firewood<br />

(Mandaville).<br />

notes: Needs confirmation as not listed for <strong>Qatar</strong> by Miller & Cope.<br />

Photos: Mand 41.<br />

Seidlitzia rosmarinus Bunge ex Boiss. Plate 47<br />

ناملج ,نانش<br />

Vernacular names: shinan/ushnan, julman<br />

Growth form: Shrub. Fl. Sep–Oct.<br />

status: Native. Local. Also recorded: Bahrain, Kuwait, E Saudi Arabia, UAE.<br />

habitat & distribution: Abundant and <strong>of</strong>ten dominant on undulating sand, sabkha edge and over<br />

sand dunes from Mesaieed southwards.<br />

uses: The leaves are used as a cleansing agent (Ghazanfar).<br />

Photos: Bat 30; Coll 155; Corn 86; Daoud 179; Jong 302; Mand 37; Shuaib 75.<br />

Suaeda aegyptiaca (Hasselq.) Zohary Plate 48<br />

Chenopodium aegyptiacum Hasselq.; Schanginia aegyptiaca (Hasselq.) Aellen; Schanginia baccata<br />

(Forssk.) Moq.<br />

Vernacular names: seablite, suweid, Hatallus, julman<br />

ناملج ,سلتح ,ديوس<br />

Growth form: Perennial herb. Fl. Sep–Oct.<br />

status: Native. Common. Also recorded: Bahrain, Kuwait, E Saudi Arabia, UAE.<br />

habitat & distribution: Occasional in coastal areas and wherever saline soils occur, including<br />

irrigated farmland.<br />

uses: El Amin remarks that fresh leaves are sold in markets in <strong>Qatar</strong> for eating in salads. Stems and<br />

leaves are also used to treat tooth and gum infections. Ghazanfar mentions that <strong>the</strong> plant is used<br />

as snuff to alleviate dizziness, headaches and o<strong>the</strong>r ailments.<br />

Photos: Bat 29; Coll 156; Corn 81; Daoud 173, 174; Jong 303; Phil 103; Shuaib 80; West 52.<br />

Suaeda vermiculata Forssk. ex J.F. Gmel. Plate 49<br />

Suaeda fruticosa Forssk. ex J.F. Gmel.<br />

Vernacular names: seablite, suweid<br />

ديوس<br />

Growth form: Shrub. Fl. Oct.<br />

status: Native. Common. Also recorded: Bahrain, Kuwait, E Saudi Arabia, UAE.<br />

habitat & distribution: Occasional or frequent in coastal saline habitats.<br />

uses: Jong et al. note that <strong>the</strong> plant is suitable for saline irrigation and a promising landscape plant,<br />

and is also used to treat asthma.<br />

Photos: El Amin 6; Bat 30; Coll 157; Corn 81; Daoud 172; Ghaz 41; Jong 280 L , 304; Phil 104; Pick 59;<br />

Shuaib 80; West 53.<br />

Traganum nudatum Delile<br />

Growth form: Shrub. Fl. Mar.<br />

status: Native. ?Rare. First published record: Miller & Cope (1996). Also recorded: Kuwait, E Saudi<br />

Arabia, UAE.<br />

habitat & distribution: A plant <strong>of</strong> rocky places over Arabia generally.<br />

Photos: Coll 158; Daoud 178; Shuaib 80.<br />

CISTACEAE<br />

Helian<strong>the</strong>mum kahiricum Delile Plate 50<br />

قورقر<br />

Vernacular names: raqrouq<br />

Growth form: Shrublet. Fl. Mar.<br />

status: Native. Local. Also recorded: Bahrain, Kuwait, E Saudi Arabia, UAE.<br />

CHENOPODIACEAE / CISTACEAE<br />

30 31


CISTACEAE / COMBRETACEAE / CONVOLVULACEAE<br />

habitat & distribution: Mainly on harder, calcareous substrates in W <strong>Qatar</strong>. Frequent in <strong>the</strong><br />

Dukhan area (JN).<br />

Photos: El Amin 25 L ; Bat 71; Coll 159; Corn 97; Daoud 35; Ghaz 133-4; Jong 305; Phil 160; West 101.<br />

Helian<strong>the</strong>mum lippii (L.) Dum.-Cours. Plate 51<br />

Vernacular names: raqrouq<br />

Growth form: Shrublet. Fl. Mar–Apr.<br />

status: Native. Very common. Also recorded: Bahrain, Kuwait, E Saudi Arabia, UAE.<br />

habitat & distribution: Frequent to abundant on hamada, gravel plains and sandy soils.<br />

uses: Rizk & El-Ghazaly report that <strong>the</strong> plant possesses antimicrobial activity.<br />

Photos: El Amin 6, 25 L ; Bat 71, 72 L ; Coll 159; Daoud 36; Jong 306; Phil 161; Shuaib 106; West 102.<br />

COMBRETACEAE<br />

Conocarpus lancifolius Engl.<br />

Vernacular names: damas<br />

Growth form: Tree. Fl. Not known.<br />

status: Introduced. Also recorded: E Saudi Arabia.<br />

قورقر<br />

habitat & distribution: Planted at Mesaieed Industrial City (JN), but may be more widespread.<br />

uses: A saline-tolerant tree producing dense foliage, making it suitable for use as shade and<br />

screening.<br />

Photos: None.<br />

CONVOLVULACEAE<br />

Convolvulus arvensis L.<br />

Vernacular names: bindweed, ’ullayq<br />

قيلع<br />

Growth form: Perennial herb. Fl. Mar–Apr.<br />

status: Introduced. Common. Also recorded: Bahrain, Kuwait, E Saudi Arabia, UAE.<br />

habitat & distribution: Occasional or frequent in lawns, gardens and o<strong>the</strong>r cultivated areas.<br />

uses: The plant is poisonous if eaten in any quantity (Rizk & El-Ghazaly); roots and leaves are used<br />

as an anti-haemorrhagic (Ghazanfar).<br />

Photos: El Amin 9; Coll 226; Corn 181; Jong 310; Phil 139; Shuaib 54; West 117.<br />

Convolvulus cephalopodus Boiss. Plate 52<br />

Convolvulus buschiricus Bornm.<br />

Vernacular names: khatme<br />

Growth form: Perennial herb. Fl. Mar–Apr.<br />

status: Native. Local. Also recorded: Kuwait, E Saudi Arabia, UAE.<br />

habitat & distribution: Occasional in sandy areas; particularly W <strong>Qatar</strong>.<br />

Photos: Bat 79; Daoud 21, 215, 216; Jong 311; Pick 168; Shuaib 55; West 117.<br />

Convolvulus fatmensis Kunze<br />

Growth form: <strong>An</strong>nual herb. Fl. Mar.<br />

status: Native. Rare. Also recorded: E Saudi Arabia.<br />

يمتاخ<br />

habitat & distribution: Recorded by Batanouny from silty soils in rodat.<br />

Photos: Bat 80; Coll 229.<br />

Convolvulus glomeratus Choisy<br />

Vernacular names: ’ullayq<br />

Growth form: Perennial herb. Fl. Mar.<br />

status: Native. Rare. Also recorded: UAE.<br />

habitat & distribution: Recorded from cultivated rodat, in light or sandy soils.<br />

uses: Taken as a purgative (Rizk & El-Ghazaly).<br />

Photos: El Amin 10; Bat 80; Coll 229; Jong 312.<br />

Convolvulus pilosellifolius Desr. Plate 53<br />

Vernacular names: melbow, haTHmy, rukheima<br />

ةميخر ,يمثه ,وبليم<br />

Growth form: Perennial herb. Fl. Mar–May.<br />

status: Native. Common. Also recorded: Bahrain, Kuwait, E Saudi Arabia, UAE.<br />

habitat & distribution: Locally abundant in silty depressions and rodat in C <strong>Qatar</strong>.<br />

uses: Eaten as salad or cooked with rice to add flavour (El Amin, Jongbloed et al.). Used as a<br />

purgative in Bahrain (Rizk & El-Ghazaly).<br />

Photos: El Amin 10, 44 L ; Bat 82; Coll 230; Corn 182; Daoud 185 L ; Jong 313; Phil 140; Shuaib 54; West 118.<br />

Convolvulus prostratus Forssk.<br />

Convolvulus deserti Hochst. & Steud.; Convolvulus microphyllus (Roth) Spreng.<br />

قيلع<br />

Growth form: Perennial herb. Fl. Mar–May.<br />

status: Native. Common. Also recorded: Bahrain, E Saudi Arabia, UAE.<br />

habitat & distribution: Occasional in silt pans and o<strong>the</strong>r depressions; also on gravelly soils and<br />

in runnels.<br />

notes: C. prostratus and C. deserti are two closely related taxa for which <strong>the</strong> taxonomy is poorly<br />

understood. Batanouny listed both species, but it is probable that only one variable entity occurs.<br />

Photos: El Amin 10; Bat 79, 81; Coll 230; Jong 314; West 118.<br />

Vernacular names: ’ullayq<br />

Cressa cretica L.<br />

Vernacular names: alkali weed, rosin weed, nedewah<br />

ةويدن<br />

Growth form: Perennial herb. Fl. Mar–May.<br />

status: Native. Local. Also recorded: Bahrain, Kuwait, E Saudi Arabia, UAE.<br />

habitat & distribution: Occasional on compact saline soils around <strong>the</strong> coasts; especially N <strong>Qatar</strong>.<br />

Recorded from Ras Laffan (JN), just west <strong>of</strong> Doha (RR) and is common at Al Zubarah (RR).<br />

uses: In Bahrain, <strong>the</strong> leaves are used as an expectorant (Rizk & El-Ghazaly); also used as an<br />

aphrodisiac (Ghazanfar).<br />

Photos: El Amin 10; Bat 82; Coll 233; Corn 182; Daoud 212, 213; Jong 317; Phil 141; Shuaib 55; West 119.<br />

Cuscuta chinensis Lam.<br />

Vernacular names: Chinese dodder<br />

Growth form: <strong>An</strong>nual parasitic herb. Fl. Jan–Dec.<br />

status: Introduced. ?Local. Needs confirmation for <strong>Qatar</strong>. Not in o<strong>the</strong>r Gulf floras.<br />

CONVOLVULACEAE<br />

32 33<br />

قيلع


CONVOLVULACEAE / CUCURBITACEAE<br />

habitat & distribution: Batanouny mentions that this species is a fairly common parasite <strong>of</strong><br />

cultivated alfalfa (Medicago sativa) and sweet basil (Ocimum basilicum), as well as Ziziphus nummularia.<br />

uses: Rizk & El-Ghazaly list a large number <strong>of</strong> medicinal uses.<br />

Photos: None.<br />

Cuscuta pedicellata Ledeb.<br />

Vernacular names: dodder<br />

Growth form: Parasitic herb. Fl. Apr.<br />

status: Introduced. Rare. Also recorded: E Saudi Arabia.<br />

habitat & distribution: Recorded from cultivated areas on many different hosts including crops<br />

such as alfalfa (Medicago sativa) and sweet basil (Ocimum basilicum); also on trees such as Prosopis and<br />

Ziziphus, and also on Chenopodium, Fagonia and Trigonella spp.<br />

Photos: Coll 234.<br />

Cuscuta pentagona Engelm.<br />

Vernacular names: dodder<br />

Growth form: Parasitic herb. Fl. Not known.<br />

status: Introduced. Rare. First published record: Abdel Bari (1997). Not in o<strong>the</strong>r Gulf floras.<br />

habitat & distribution: Abdel Bari records it as a ‘common parasite on ornamental annuals’.<br />

Photos: None.<br />

CUCURBITACEAE<br />

Citrullus colocynthis (L.) Schrad. Plate 54<br />

Cucumis colocynthis L.<br />

Vernacular names: hanDHal, shary (fruit), Hadaj<br />

desert squash, bitter gourd/squash<br />

Growth form: Perennial herb. Fl. Apr–Jul.<br />

status: Native. Common. Also recorded: Bahrain, Kuwait, E Saudi Arabia, UAE.<br />

جدح ,)ةرمث( يرش ,لظنح<br />

habitat & distribution: Occasional in sandy areas, particularly in depressions and runnels; though<br />

single plants may cover a large area.<br />

uses: El Amin reports that <strong>the</strong> ripe fruit were used to prepare gunpowder. The plant is well known<br />

to have strong laxative properties; o<strong>the</strong>r medicinal uses recorded by Ghazanfar include treatment<br />

<strong>of</strong> dog, insect and snake bites; to relieve pain in joints and as a hair dye.<br />

Photos: El Amin 16; Bat 73, 74 L ; Coll 268; Daoud 4, 46, 47, 48; Ghaz 151; Jong 323; Mand 67, 68; Phil 105;<br />

Pick 170; Shuaib 55; West 104.<br />

Cucumis prophetarum L.<br />

Vernacular names: shary, haneiDalan, Hadak<br />

Growth form: Perennial herb. Fl. Apr–May.<br />

status: Introduced. ?Rare. Also recorded: E Saudi Arabia, UAE.<br />

habitat & distribution: Previously recorded from Al Karaanah.<br />

uses: The plant has purgative properties (Rizk & El-Ghazaly).<br />

Photos: El Amin 16; Bat 73, 74; Coll 270; Ghaz 147-9; Jong 325; Pick 122; West 105.<br />

قدح ,نلاضينه ,يرش<br />

CYNOMORIACEAE<br />

Cynomorium coccineum L. Plate 55<br />

ثوثرط<br />

Vernacular names: red thumb, desert thumb, Tarthuth<br />

Growth form: Parasitic perennial herb. Fl. Mar.<br />

status: Native. Local. Also recorded: Bahrain, Kuwait, E Saudi Arabia, UAE.<br />

habitat & distribution: On damp, saline sand, including beaches, thus mainly with a coastal<br />

distribution. Frequent at Dukhan water treatment works, growing on Tetraena qatarensis and nearby<br />

on <strong>the</strong> west coast (JN); also occurs at Ras Laffan (RR), Mesaieed and Khor Al Adaid (MS). O<strong>the</strong>r<br />

host plants include species <strong>of</strong> Chenopodiaceae.<br />

uses: The roots are edible and were sold in earlier times as a vegetable. Flowers were used as a dye<br />

in Saudi Arabia (Ghazanfar); used as a aphrodisiac in Bahrain (Rizk & El-Ghazaly).<br />

Photos: Bat 75; Coll 276; Corn 33, 145; Daoud 70; Jong 326; Mand 134; Phil 64, 65; Shuaib 51; West 105.<br />

EUPHORBIACEAE<br />

CyNOMORIACEAE / EUPHORBIACEAE<br />

<strong>An</strong>drachne telephioides L. Plate 56<br />

هظميوح<br />

Vernacular names: HuweimDHa<br />

Growth form: Perennial herb. Fl. Mar–May.<br />

status: Native. Common. Also recorded: Bahrain, Kuwait, E Saudi Arabia, UAE.<br />

habitat & distribution: Frequent in silty depressions.<br />

uses: The plant is poisonous and used as an insecticide (Rizk & El-Ghazaly).<br />

Photos: Bat 63; Coll 302; Corn 120; Daoud 64, 65; Jong 330; Phil 168; Shuaib 84; West 92.<br />

Chamaesyce arabica (Hochst. & Steud. ex Boiss.) Soják<br />

Euphorbia arabica <strong>An</strong>derson<br />

Growth form: <strong>An</strong>nual herb. Fl. Feb–Apr.<br />

status: Native. Rare. First published record: Abdel Bari (1997). Also recorded: UAE.<br />

habitat & distribution: Recorded near Abu Samrah on a ‘mud plain’.<br />

Photos: Jong 334; West 93.<br />

Chamaesyce granulata Forssk.<br />

Euphorbia granulata (Forssk.) Soják<br />

Vernacular names: labeinah<br />

ةنيبل<br />

Growth form: <strong>An</strong>nual or biennial herb. Fl. Mar–May.<br />

status: Introduced. Common. Also recorded: Kuwait, E Saudi Arabia, UAE.<br />

habitat & distribution: Occurs on various soils and a weed in urban areas, but may have been<br />

frequently mis-recorded for C. serpens.<br />

uses: Sap is applied to poisonous bites (Jongbloed et al.).<br />

notes: One <strong>of</strong> several very similar prostrate Euphorbia species, <strong>of</strong>ten treated as Chamaesyce, that<br />

occur in damp, disturbed habitats, particularly in lawns, gardens and plantations.<br />

Photos: El Amin 11; Bat 63; Coll 315; Daoud 67; Jong 335; Shuaib 85; West 95.<br />

34 35


EUPHORBIACEAE<br />

Chamaesyce hirta (L.) Millsp.<br />

Euphorbia hirta L.<br />

Vernacular names: labeinah<br />

Growth form: <strong>An</strong>nual herb. Fl. Apr–Sep.<br />

status: Introduced. Rare. Also recorded: Kuwait, E Saudi Arabia, UAE.<br />

habitat & distribution: A weed <strong>of</strong> cultivated land (Batanouny).<br />

uses: The sap is used to treat cuts and wounds (Rizk & El-Ghazaly), a number <strong>of</strong> o<strong>the</strong>r medicinal<br />

uses are also listed.<br />

Photos: Coll 317; Jong 338; Shuaib 86.<br />

Chamaesyce prostrata (Aiton) Small<br />

Euphorbia prostrata Aiton<br />

Growth form: <strong>An</strong>nual herb. Fl. Mar.<br />

status: Introduced. Common. Also recorded: UAE.<br />

habitat & distribution: Frequent in irrigated areas in C <strong>Qatar</strong>, especially in rodat.<br />

uses: Amongst those listed by Rizk & El-Ghazaly are to treat flu, dysentery, to purify blood and<br />

improve <strong>the</strong> milk <strong>of</strong> nursing mo<strong>the</strong>rs. It is also used as a diabetes remedy and <strong>the</strong> sap applied to<br />

sores and used as a snake bite remedy.<br />

Photos: El Amin 12; Bat 64; Jong 342.<br />

Chamaesyce serpens (Kunth) Small<br />

Euphorbia serpens Kunth<br />

Growth form: <strong>An</strong>nual herb. Fl. Dec–May.<br />

status: Introduced. Not confirmed for <strong>Qatar</strong>, but very likely to occur. Also recorded: Bahrain, E<br />

Saudi Arabia, UAE.<br />

habitat & distribution: Likely to occur in similar habitats to C. granulata.<br />

Photos: Coll 321; Corn 122; Jong 343; West 94.<br />

Chrozophora oblongifolia (Delile) A. Juss. ex Spreng.<br />

Chrozophora verbascifolia (Willd.) A. Juss.<br />

Vernacular names: Tanoom<br />

مونط<br />

Growth form: Perennial herb or small shrub. Fl. Dec–May.<br />

status: Native. Local. Also recorded: Kuwait, E Saudi Arabia, UAE.<br />

habitat & distribution: Occasional in cultivated depressions, field margins and disturbed areas<br />

in C <strong>Qatar</strong>.<br />

uses: Chrozophora species have been traditionally used in Arabia to produce dyes. Fur<strong>the</strong>r details<br />

are given by Mandaville.<br />

Photos: Coll 302; Daoud 66; Jong 331; Pick 209; West 92.<br />

Chrozophora tinctoria (L.) Raf.<br />

Chrozophora obliqua (Vahl.) A. Juss. ex Spreng.<br />

Vernacular names: Tanoom, zerraij<br />

جيرز ,مونط<br />

Growth form: <strong>An</strong>nual (occasionally perennial) herb. Fl. Apr–Jun.<br />

status: Native. Local. Also recorded: Kuwait, E Saudi Arabia, UAE.<br />

habitat & distribution: Occasional in cultivated rodat, field margins and disturbed areas; C <strong>Qatar</strong>.<br />

ةنيبل<br />

uses: Like C. oblongifolia, used to produce dyes; however for C. tinctoria Mandaville states that<br />

although long used in sou<strong>the</strong>rn Europe and <strong>the</strong> Mediterranean <strong>the</strong>re do not appear to be any<br />

records <strong>of</strong> this use in eastern Arabia.<br />

Photos: El Amin 11, 55 L ; Coll 302; Mand 135; Shuaib 85.<br />

Euphorbia dracunculoides Lam.<br />

Growth form: <strong>An</strong>nual herb. Fl. Mar.<br />

status: Introduced. ?Rare. Also recorded: E Saudi Arabia.<br />

habitat & distribution: Recorded by El Amin from disturbed soils <strong>of</strong> rodat; C <strong>Qatar</strong>.<br />

Photos: Coll 312.<br />

Euphorbia heterophylla L.<br />

Euphorbia geniculata Ortega<br />

EUPHORBIACEAE<br />

Vernacular names: labeinah<br />

ةنيبل<br />

Growth form: <strong>An</strong>nual herb. Fl. Jan–Apr.<br />

status: Introduced. Common. Also recorded: UAE.<br />

habitat & distribution: Recorded by Batanouny as ‘common’ at Al Wabrah where naturalised,<br />

but may be more widespread now.<br />

Photos: Coll 317; Jong 336.<br />

Euphorbia peplus L.<br />

Vernacular names: petty spurge<br />

Growth form: <strong>An</strong>nual herb. Fl. Mar.<br />

status: Introduced. Rare. Also recorded: Bahrain, E Saudi Arabia, UAE.<br />

habitat & distribution: Occasional as a roadside weed in Dukhan township in 2007 (JN).<br />

uses: The plant is good for asthma and catarrh; <strong>the</strong> milky juice is used to remove corns, warts and<br />

in <strong>the</strong> treatment <strong>of</strong> cancer <strong>of</strong> <strong>the</strong> stomach, liver and uterus (Rizk & El-Ghazaly).<br />

Photos: Bat 64; Coll 320; Corn 122; Jong 341.<br />

Euphorbia retusa Forssk. Plate 57<br />

Euphorbia cornuta Pers.<br />

Growth form: <strong>An</strong>nual (occasionally perennial) herb. Fl. Feb–Apr.<br />

status: Native. Rare. Also recorded: E Saudi Arabia.<br />

habitat & distribution: Recorded on a silt pan NE <strong>of</strong> Zikreet in 2007 (JN).<br />

Photos: Coll 320.<br />

Mercurialis annua L.<br />

Growth form: <strong>An</strong>nual herb. Fl. All year.<br />

status: Introduced. Not in o<strong>the</strong>r Gulf floras.<br />

habitat & distribution: Recorded by Batanouny but no information given on status or distribution;<br />

likely to occur in damp, cultivated areas, as in o<strong>the</strong>r parts <strong>of</strong> <strong>the</strong> world.<br />

Photos: Coll 326.<br />

36 37


EUPHORBIACEAE / FABACEAE<br />

Phyllanthus niruri L.<br />

Vernacular names: stone-breaker<br />

Growth form: <strong>An</strong>nual herb. Fl. Not known.<br />

status: Introduced. ?Local. First published record: Abdel Bari (1997). Not in o<strong>the</strong>r Gulf floras.<br />

habitat & distribution: Recorded as ‘widespread’ in gardens.<br />

uses: As its English name indicates, well-known as a cure for kidney stones and also used to treat<br />

liver diseases. Dried leaves and extracts <strong>of</strong> <strong>the</strong> plant are sold commercially around <strong>the</strong> world. O<strong>the</strong>r<br />

reported uses include to aid digestion, relieve pain and to treat malaria and o<strong>the</strong>r fevers.<br />

notes: P. amarus Schumach. is closely-related and has similar uses (this is much more common as<br />

a naturalised species than P. niruri).<br />

Photos: None.<br />

Ricinus communis L.<br />

Vernacular names: castor oil plant, ’arash<br />

شارع<br />

Growth form: Shrub. Fl. Jan–May.<br />

status: Introduced. Also recorded: E Saudi Arabia, UAE.<br />

habitat & distribution: In <strong>the</strong> Gulf generally it is widely planted in shelterbelts, but also escapes<br />

into areas around gardens and farms.<br />

uses: It is well known that <strong>the</strong> oil is extracted for medicinal uses, particularly as a laxative, and<br />

that <strong>the</strong> seeds contain a deadly poison, <strong>the</strong> alkaloid ‘ricinine’. The oil is also used in <strong>the</strong> chemical<br />

industry and as a lubricant. Ghazanfar also gives various o<strong>the</strong>r uses, including <strong>the</strong> treatment <strong>of</strong> bad<br />

breath, blisters, ulcers, toothache, inflamed eyes and rheumatic pain.<br />

Photos: Coll 327; Jong 346; Pick 175.<br />

FABACEAE<br />

Acacia ehrenbergiana Hayne<br />

Acacia flava (Forssk.) Schweinf.<br />

Plate 58<br />

Vernacular names: salam<br />

ملس<br />

Growth form: Shrub or tree. Fl. Apr–May.<br />

status: Native. Common. Also recorded: E Saudi Arabia, UAE.<br />

habitat & distribution: Frequent in rodat and o<strong>the</strong>r sandy areas, <strong>of</strong>ten with o<strong>the</strong>r large shrubs<br />

and trees. Less less widely distributed than A. tortilis, but it is locally dominant in C and SE <strong>Qatar</strong>,<br />

especially at Al Sheehaniyah. Mandaville states that it is <strong>the</strong> commonest Acacia <strong>of</strong> <strong>the</strong> Eastern<br />

Province <strong>of</strong> Saudi Arabia.<br />

uses: <strong>An</strong>imals graze on leaves and pods; stems are used as firewood and <strong>the</strong> species is a good<br />

sand stabiliser (El Amin). The plant has antimicrobial activity and extracts are used to produce a<br />

molluscicide to kill snails (Rizk & El-Ghazaly).<br />

Photos: El Amin 13; Bat 41, 42 L ; Coll 471; Jong 423; Mand 109, 110; Pick 237.<br />

Acacia nilotica (L.) Delile<br />

Acacia arabica Willd.<br />

Vernacular names: Arabian gum tree, Sunt, jard (fruit)<br />

Growth form: Tree. Fl. Mar–Nov.<br />

status: Introduced. Also recorded: E Saudi Arabia, UAE.<br />

)ةرمث( درج ,تنص<br />

habitat & distribution: Subsp. indica is widely planted in <strong>the</strong> Gulf and presumably in <strong>Qatar</strong>; it is<br />

native to parts <strong>of</strong> Saudi Arabia and Oman.<br />

uses: The wood has various construction uses and can used as firewood and for producing charcoal.<br />

The bark is a source <strong>of</strong> tannin and dye. Gum Arabic is obtained from this and o<strong>the</strong>r Acacia species<br />

and used commercially as stabiliser in <strong>the</strong> food industry; also as an ingredient in o<strong>the</strong>r products<br />

such as glues, paints and cosmetics.<br />

Photos: Jong 424; Pick 239; West 69.<br />

Acacia tortilis (Forssk.) Hayne<br />

Acacia raddiana Savi; Acacia spirocarpa Hochst. ex A. Rich.<br />

Plate 59<br />

Vernacular names: samr<br />

رمس<br />

Growth form: Shrub or tree. Fl. May–Jun.<br />

status: Native. Very common. Also recorded: E Saudi Arabia, UAE.<br />

habitat & distribution: Locally dominant in sandy depressions, wadis and larger runnels, <strong>of</strong>ten<br />

preferring <strong>the</strong> slightly elevated ground at <strong>the</strong> edges <strong>of</strong> such features.<br />

uses: Grazed by livestock and used for fodder and firewood. The gum is <strong>of</strong> economic importance<br />

in some regions. The wood is used for building camel folds (Rizk & El-Ghazaly).<br />

Photos: El Amin 13; Bat 41, 43 L ; Coll 476; Jong 422, 425; Mand 112, 113; Pick 236; West 69.<br />

Alhagi maurorum Medik.<br />

Alhagi camelorum Fisch.; Alhagi graecorum Boiss.<br />

Vernacular names: camelthorn, ’aaqool<br />

لوقاع<br />

Growth form: Small shrub. Fl. Apr–May.<br />

status: Introduced. Local. Also recorded: Bahrain, Kuwait, E Saudi Arabia, UAE.<br />

habitat & distribution: Occurs in <strong>the</strong> NW <strong>of</strong> Al Khor town (MS) and seen by MAS at two locations<br />

in W <strong>Qatar</strong>. Batanouny recorded it at Abu Samrah and Umm Bab. Usually in saline or disturbed<br />

sandy soils, including farms and coastal areas.<br />

uses: Grazed by camels. The plant exudes a sugary sap that dries into ‘manna’ (Jongbloed et<br />

al. citing an earlier author). Ghazanfar states that <strong>the</strong> whole plant is used for treating cataracts,<br />

migraine, jaundice, migraine, painful joints and as an aphrodisiac.<br />

notes: The distinction between A. maurorum and A. graecorum is very slight and probably not<br />

taxonomically significant.<br />

Photos: Corn 126; Daoud 34; Jong 347; Phil 31; Shuaib 89; West 70.<br />

Argyrolobium arabicum (Decne.) Jaub. & Spach<br />

Argyrolobium abyssinicum Jaub. & Spach<br />

Growth form: <strong>An</strong>nual or short-lived perennial herb. Fl. Not known.<br />

status: ?Native. ?Rare. First published record: Obeid (1975). Not in o<strong>the</strong>r Gulf floras.<br />

habitat & distribution: In Egypt <strong>the</strong> plant grows is sandy and rocky hillsides and in wadis (Boulos).<br />

notes: Similar to Lotus so <strong>the</strong>re may be some doubt over <strong>the</strong> identification <strong>of</strong> this record; Boulos<br />

gives <strong>the</strong> distribution as extending to Arabia, but <strong>the</strong> species is not shown in Collenette.<br />

Photos: None.<br />

Astragalus annularis Forssk. Plate 60<br />

Vernacular names: Halaq<br />

قلح<br />

Growth form: <strong>An</strong>nual herb. Fl. Feb–Apr.<br />

status: Native. Local. Also recorded: Bahrain, Kuwait, E Saudi Arabia, UAE.<br />

FABACEAE<br />

38 39


FABACEAE<br />

habitat & distribution: Occasional to frequent in sandy depressions. Abundant in <strong>the</strong> Dukhan to<br />

Umm Bab area after rains in 2007 (JN).<br />

Photos: Coll 480; Corn 129; Daoud 25, 26; Jong 349; Mand 120; Phil 32; Shuaib 92; West 71.<br />

Astragalus corrugatus Bertol.<br />

Astragalus tenuirugis Boiss.<br />

Vernacular names: Halaq, khawatim al bar<br />

Growth form: <strong>An</strong>nual herb. Fl. Mar–Apr.<br />

status: Native. Local. Also recorded: Bahrain, Kuwait, E Saudi Arabia, UAE.<br />

ربلا متاوخ ,قلح<br />

habitat & distribution: Recorded in S <strong>Qatar</strong> at Al Karaanah and some areas in <strong>the</strong> north (MS).<br />

Grows in silty and sandy depressions and o<strong>the</strong>r areas with fine, shallow soils.<br />

Photos: El Amin 70 L ; Bat 44; Corn 129; Daoud 20, 21; Jong 350; Mand 122; Phil 33; Shuaib 93.<br />

Astragalus eremophilus Boiss. Plate 61<br />

Vernacular names: Halaq<br />

Growth form: <strong>An</strong>nual herb. Fl. Mar–Apr.<br />

status: Native. Rare. Also recorded: E Saudi Arabia, UAE.<br />

habitat & distribution: Recorded by Batanouny from depressions and a wadi in S <strong>Qatar</strong> near Al<br />

Karaanah and also from a sandy area in W <strong>Qatar</strong> near Umm Bab (JN).<br />

Photos: El Amin 70 L ; Bat 44; Coll 483; Jong 351; Mand 123.<br />

Astragalus hamosus L.<br />

Vernacular names: Halaq<br />

Growth form: <strong>An</strong>nual herb. Fl. Mar–Apr.<br />

status: Native. Rare. Also recorded: Kuwait, E Saudi Arabia, UAE.<br />

habitat & distribution: Previously recorded from Al Wabrah.<br />

Photos: El Amin 13, 70 L ; Bat 45; Coll 484.<br />

Astragalus hauarensis Boiss.<br />

Astragalus gyzensis Delile<br />

قلح<br />

Growth form: <strong>An</strong>nual herb. Fl. Mar–May.<br />

status: Native. Rare. First record: near Umm Bab, March 2007 (S. Aspinall). Also recorded:<br />

Bahrain, Kuwait, E Saudi Arabia, UAE.<br />

habitat & distribution: Mandaville describes it as ‘perhaps <strong>the</strong> most common Astragalus <strong>of</strong> sandy<br />

habitats’, so it may occur elsewhere in sou<strong>the</strong>rn <strong>Qatar</strong>.<br />

Photos: Daoud 22, 23, 24; Jong 353; Mand 124; Shuaib 93.<br />

Vernacular names: Halaq<br />

Astragalus schimperi Boiss.<br />

Vernacular names: Halaq<br />

Growth form: <strong>An</strong>nual herb. Fl. Mar–Apr.<br />

status: Native. Rare. Also recorded: Bahrain, Kuwait, E Saudi Arabia, UAE.<br />

habitat & distribution: Recorded at Jebel Dukhan.<br />

Photos: Coll 486; Daoud 18, 19; Shuaib 93.<br />

قلح<br />

قلح<br />

قلح<br />

Astragalus sieberi DC.<br />

Astragalus zubairensis Eig<br />

Vernacular names: khanaSir al ’aroos<br />

Growth form: Shrublet. Fl. Mar–Apr.<br />

status: Native. Rare. Also recorded: Kuwait, E Saudi Arabia.<br />

سورعلا رصانخ<br />

habitat & distribution: Recorded in SW <strong>Qatar</strong>, west <strong>of</strong> Al Karaanah and Wadi Al Ireig. Recently<br />

seen in Mesameer area (MS).<br />

Photos: El Amin 14, 70 L ; Bat 45; Coll 486; Shuaib 94.<br />

Astragalus tribuloides Delile<br />

Vernacular names: rakhami<br />

40 41<br />

يماخر<br />

Growth form: <strong>An</strong>nual herb. Fl. Mar–Apr.<br />

status: Native. Common. Also recorded: Bahrain, Kuwait, E Saudi Arabia, UAE.<br />

habitat & distribution: Occasional to frequent in sandy depressions and runnels; mainly C <strong>Qatar</strong>.<br />

Photos: El Amin 14, 70 L ; Bat 46; Coll 488; Corn 129; Daoud 16, 17; Jong 355; Shuaib 93.<br />

Cullen plicata (Delile) C. H. Stirt.<br />

Psoralea plicata Delile<br />

Vernacular names: Hama<br />

Growth form: Small shrub. Fl. Mar–Apr.<br />

status: Introduced. Rare. Also recorded: E Saudi Arabia.<br />

habitat & distribution: Recorded by Batanouny from roadside runnel at Al-Amiriyah and a<br />

protected rodah in Trainah on waste land near irrigation.<br />

uses: <strong>An</strong> infusion <strong>of</strong> <strong>the</strong> leaves is drunk for respiratory and intestinal ailments and <strong>the</strong> fruits are<br />

used in <strong>the</strong> treatment <strong>of</strong> gastric ulcers (Rizk & El-Ghazaly).<br />

Photos: Bat 51; Coll 517.<br />

Hippocrepis areolata Desv.<br />

Hippocrepis bicontorta Loisel.<br />

Vernacular names: umm al qurain<br />

Growth form: <strong>An</strong>nual herb. Fl. Mar–Apr.<br />

status: Native. Common. Also recorded: Bahrain, Kuwait, E Saudi Arabia, UAE.<br />

habitat & distribution: Occasional to frequent in sandy areas; especially W <strong>Qatar</strong>.<br />

امح<br />

نيرقلا مأ<br />

Photos: El Amin 14, 73 L ; Bat 47; Coll 498; Corn 130; Daoud 30, 31; Jong 361; Mand 131; Phil 35; Shuaib<br />

91; West 73.<br />

Hippocrepis constricta Kunze<br />

Vernacular names: qileiqlan<br />

Growth form: <strong>An</strong>nual herb. Fl. Mar–Apr.<br />

status: Native. ?Local. Also recorded: E Saudi Arabia, UAE.<br />

FABACEAE<br />

habitat & distribution: Recorded by Batanouny from sandy soils in rodat, but he did not give<br />

details <strong>of</strong> abundance.<br />

Photos: El Amin 73 L ; Bat 48; Coll 498; Jong 362; West 74.<br />

نلاقيلق


FABACEAE<br />

Hippocrepis multisiliquosa L.<br />

Vernacular names: qileiqlan<br />

Growth form: <strong>An</strong>nual herb. Fl. Mar.<br />

status: Native. Rare. Not in o<strong>the</strong>r Gulf floras.<br />

habitat & distribution: Depressions and runnels.<br />

Photos: El Amin 14, 73 L ; Bat 48.<br />

Hippocrepis unisiliquosa L.<br />

Growth form: <strong>An</strong>nual herb. Fl. Feb–Apr.<br />

status: Native. ?Rare. Also recorded: Bahrain, E Saudi Arabia, UAE.<br />

habitat & distribution: Batanouny recorded as ‘fairly common’ in depressions, but it is not<br />

mentioned by El Amin; Mandaville states that it appears to be very rare in Saudi Arabia.<br />

Photos: Coll 499; Jong 363.<br />

Hymenocarpos circinnatus (L.) Savi<br />

Growth form: <strong>An</strong>nual herb. Fl. Apr.<br />

status: Native. Rare. Not in o<strong>the</strong>r Gulf floras.<br />

habitat & distribution: Recorded by Batanouny from Jebel Al Wakra.<br />

Photos: None.<br />

Indig<strong>of</strong>era intricata Boiss. Plate 62<br />

Vernacular names: Hameira<br />

ةريمح<br />

Growth form: Shrublet. Fl. Jan–Jun.<br />

status: Native. Common. First published record: Mandaville (1990, p.181). Also recorded: E Saudi<br />

Arabia, UAE.<br />

habitat & distribution: Occasional in sandy habitats. It appears to be relatively frequent in<br />

western coastal areas (JN, SAM) and is also common across a large part <strong>of</strong> <strong>the</strong> UAE (Jongbloed et al.).<br />

uses: The roots and leaves are used as a tonic (Rizk & El-Ghazaly).<br />

notes: Probably mis-named as I. articulata Gouan by earlier authors.<br />

Photos: El Amin 15; Bat 49; Jong 367; Pick 66.<br />

Indig<strong>of</strong>era oblongifolia Forssk.<br />

Growth form: Shrublet. Fl. Not known.<br />

status: ?Introduced. Rare. First published record: Abdel Bari (1997). Also recorded: UAE.<br />

habitat & distribution: Abdel Bari found this species near Doha central market, suggesting that<br />

it has been introduced by human activity.<br />

uses: Ghazanfar states <strong>the</strong> roots are used for an analgesic and as an anti-inflammatory; leaves are<br />

used as a hair wash.<br />

Photos: Jong 369; West 75.<br />

Lathyrus inconspicuus L.<br />

Growth form: <strong>An</strong>nual herb. Fl. Feb–Mar.<br />

status: Introduced. Rare. First record: SW <strong>Qatar</strong>, March 2007 (S. Aspinall). Not in o<strong>the</strong>r Gulf floras.<br />

نلاقيلق<br />

habitat & distribution: A weed <strong>of</strong> crops such as chick-pea, so <strong>the</strong> record may refer to a plant that<br />

had escaped from a cultivated area. Not recorded elsewhere in <strong>the</strong> Gulf and possibly not from <strong>the</strong><br />

Arabian Peninsula, but known from <strong>the</strong> Mediterranean region.<br />

Photos: None.<br />

Lotononis platycarpa (Viv.) Pic.Serm.<br />

Lotononis dichotoma (Delile) Boiss.; Lotus platycarpa Viv.<br />

Growth form: <strong>An</strong>nual herb. Fl. Mar.<br />

status: Native. Common. Also recorded: E Saudi Arabia, UAE.<br />

habitat & distribution: Occasional in sandy habitats in S <strong>Qatar</strong>. Previously also recorded from Al<br />

Wakra and Wadi Al Banat.<br />

Photos: El Amin 15, 76 L ; Bat 53; Coll 506; Jong 370; Mand 114.<br />

Lotus garcinii DC.<br />

Growth form: Small shrub. Fl. Dec–Jun.<br />

status: Native. Rare. Also recorded: Bahrain, E Saudi Arabia, UAE.<br />

habitat & distribution: A species <strong>of</strong> coastal, slightly saline sand. Recorded from NW <strong>Qatar</strong> by<br />

Batanouny.<br />

Photos: Coll 507; Corn 133; Jong 371; Pick 67; West 76.<br />

Lotus glinoides Delile<br />

Lotus schimperi Steud. ex Boiss.<br />

Vernacular names: Hawrbeith<br />

Growth form: <strong>An</strong>nual herb. Fl. Apr–May.<br />

status: Native. Local. Also recorded: Bahrain, UAE.<br />

habitat & distribution: Occasional in sandy areas; W and S <strong>Qatar</strong>.<br />

42 43<br />

ثيبروح<br />

notes: L. glinoides and L. schimperi were formerly regarded as distinct species, and are probably best<br />

amalgamated, but not all authors agree.<br />

Photos: Coll 507; Jong 372; West 76.<br />

Lotus halophilus Boiss. & Spruner Plate 63<br />

Lotus pusillus Viv.; Lotus villosus Forssk., non Burm. f.<br />

لازغلا نرق ,ثيبروح<br />

Vernacular names: gazelle’s horn, Hawrbeith, qarn al ghazal<br />

Growth form: <strong>An</strong>nual herb. Fl. Feb–Mar.<br />

status: Native. Very common. Also recorded: Bahrain, Kuwait, E Saudi Arabia, UAE.<br />

habitat & distribution: Abundant in sandy habitats, especially after winter rains. Depressions in<br />

<strong>the</strong> Dukhan area in 2007 turned green with this species. May be less common in E <strong>Qatar</strong>.<br />

uses: Grazed by sheep and goats (El Amin). Due to its abundance and presumed high protein<br />

content <strong>of</strong> <strong>the</strong> pods this species would presumably have been an important seasonal grazing species<br />

in <strong>the</strong> region.<br />

Photos: El Amin 76 L ; Coll 508; Corn 133; Daoud 14, 15; Jong 373; Phil 36; Shuaib 91.<br />

Medicago laciniata (L.) Mill.<br />

Medicago aschersoniana Urb.<br />

Vernacular names: medick, nafal, jut bari, barseem bari, abu Hassak<br />

Growth form: <strong>An</strong>nual herb. Fl. Feb–Mar.<br />

FABACEAE<br />

كسح وبأ ,يرب ميسرب ,يرب تج ,لفن


FABACEAE<br />

status: Native. Very common. Also recorded: Bahrain, Kuwait, E Saudi Arabia, UAE.<br />

habitat & distribution: Occasional to frequent in sandy habitats.<br />

uses: Said to be ‘palatable grazing’ by Batanouny, although El Amin says it is grazed by animals<br />

but not much liked because <strong>of</strong> <strong>the</strong> bristly fruit.<br />

notes: A very variable species that was formerly separated into two species or varieties (see<br />

Mandaville).<br />

Photos: El Amin 78 L ; Bat 50; Coll 509; Corn 134; Daoud 10, 11; Jong 374; Mand 117; Phil 37; Shuaib 89;<br />

West 77.<br />

Medicago polymorpha L.<br />

Vernacular names: nafal, jut bari<br />

Growth form: <strong>An</strong>nual herb. Fl. Mar.<br />

status: Introduced. Local. Also recorded: E Saudi Arabia, UAE.<br />

habitat & distribution: Occasional in rodat in C <strong>Qatar</strong> and in gardens.<br />

Photos: Coll 511.<br />

Medicago sativa L.<br />

Vernacular names: alfalfa, lucerne, burseem, jatt<br />

Growth form: Perennial herb. Fl. May–Jul.<br />

status: Introduced. ?Common. Also recorded: Kuwait, E Saudi Arabia.<br />

habitat & distribution: May occur as an escape from cultivation.<br />

يرب ميسرب ,يرب تج<br />

تج ,ميسرب<br />

uses: This is a very common and widely grown fodder crop throughout <strong>the</strong> Arabian peninsula.<br />

Ghazanfar states <strong>the</strong> leaves are used to treat bruises, fractures and to stop nose-bleeds. Seeds are<br />

used as an aphrodisiac.<br />

Photos: Bat 50.<br />

Melilotus albus Medik.<br />

Vernacular names: sweet clover, Handaquq, nafal<br />

Growth form: <strong>An</strong>nual or biennial herb. Fl. Mar–Apr.<br />

status: Introduced. Common. Also recorded: E Saudi Arabia, UAE.<br />

habitat & distribution: Occasional in irrigated places, including gardens.<br />

لفن ,قوقدنح<br />

uses: Grazed by animals (El Amin), but widely known to have toxic effects on livestock. Used to<br />

treat rheumatic pain and various o<strong>the</strong>r medical conditions and has narcotic action (Jongbloed et<br />

al., Rizk & El-Ghazaly).<br />

Photos: El Amin 80 L ; Bat 50; Coll 512; Jong 375.<br />

Melilotus indicus (L.) All.<br />

Vernacular names: scented trefoil, indian melilot, Handaquq, nafal<br />

Growth form: <strong>An</strong>nual herb. Fl. Mar–Apr.<br />

status: Introduced. Common. Also recorded: Bahrain, Kuwait, E Saudi Arabia, UAE.<br />

habitat & distribution: Occasional in irrigated places, including gardens.<br />

لفن ,قوقدنح<br />

uses: Rizk & El-Ghazaly list various medicinal uses including treatment <strong>of</strong> rashes, abdominal<br />

cramps and genital diseases.<br />

Photos: El Amin 15; Coll 512; Corn 134; Daoud 12, 13; Jong 376; Mand 118; Phil 38; Pick 128; Shuaib 89;<br />

West 77.<br />

Ononis reclinata L.<br />

Vernacular names: small restharrow<br />

Growth form: <strong>An</strong>nual herb. Fl. Mar.<br />

status: Native. Rare. Also recorded: Bahrain, UAE.<br />

habitat & distribution: Reported as rare by Batanouny and seen only once recently near Jebel<br />

Dukhan (JN).<br />

Photos: Coll 514.<br />

Ononis sicula Guss.<br />

Growth form: <strong>An</strong>nual herb. Fl. Mar.<br />

status: Native. Rare. Also recorded: UAE.<br />

habitat & distribution: Recorded from sandy soils in cultivated rodat.<br />

Photos: Coll 515; Jong 378.<br />

Parkinsonia aculeata L.<br />

Vernacular names: ’aqrab<br />

برقع<br />

Growth form: Tree. Fl. Not known.<br />

status: Introduced. Not in o<strong>the</strong>r Gulf floras.<br />

habitat & distribution: Presumably widespread in urban areas.<br />

uses: Planted as an ornamental tree, for erosion control and a source <strong>of</strong> firewood and charcoal.<br />

Photos: Bat 16.<br />

Prosopis cineraria (L.) Druce Plate 64<br />

فاغ<br />

Vernacular names: ghaf<br />

Growth form: Tree. Fl. May–Aug.<br />

status: Native. Rare. Also recorded: E Saudi Arabia, UAE.<br />

habitat & distribution: A survey by MS confirmed that <strong>the</strong>re are two main areas <strong>of</strong> distribution in<br />

C and N <strong>Qatar</strong>. At <strong>the</strong> ‘Al Ghafat’ area at Rawdat Rashed <strong>the</strong>re are eight trees and at <strong>the</strong> ‘Al Ghaf ’<br />

area near Medinat Al Shamal <strong>the</strong>re are 13 trees. All are mature specimens, which according to Abdel<br />

Bari et al. (2007) have an estimated age <strong>of</strong> 90-150 years or more. Over <strong>the</strong> country as a whole <strong>the</strong>re<br />

are likely to be no more than 50 native trees surviving. No young or regenerating trees have been<br />

seen and it is clear that <strong>the</strong> species is declining. <strong>Qatar</strong> lies at <strong>the</strong> western edge <strong>of</strong> <strong>the</strong> distribution <strong>of</strong><br />

<strong>the</strong> species in <strong>the</strong> Gulf, with <strong>the</strong> main native range centred on Dubai (where <strong>the</strong> species is also under<br />

threat from habitat destruction) and <strong>the</strong> eastern Emirates, extending into Oman. It only just reaches<br />

Saudi Arabia at <strong>the</strong> edges <strong>of</strong> <strong>the</strong> Rub’ Al Khali (Mandaville). Worldwide it occurs in <strong>the</strong> deserts <strong>of</strong><br />

nor<strong>the</strong>rn India, parts <strong>of</strong> sou<strong>the</strong>rn India, Pakistan, Iran and Afghanistan. It is also planted in <strong>Qatar</strong>.<br />

uses: A very important tree in <strong>the</strong> Arabian Gulf. Bedouin used to eat young leaves and seed pods,<br />

and <strong>the</strong>y are also consumed by animals. The tree provides good firewood and is used in afforestation<br />

projects (El Amin). Many medicinal uses are recorded (see also under P. juliflora).<br />

Photos: Jong 426, 427; Pick 241; West 78.<br />

Prosopis juliflora (Sw.) DC. Plate 65<br />

فيوغ<br />

Vernacular names: mesquite, ghuweif<br />

Growth form: Tree. Fl. Oct–Jan, Mar–May.<br />

status: Introduced. Very common. Also recorded: Bahrain, E Saudi Arabia, UAE.<br />

FABACEAE<br />

44 45


FABACEAE<br />

habitat & distribution: Occasional to abundant or dominant in sandy areas, disturbed habitats<br />

and cultivated areas. Originally planted in and around towns and villages but now naturalised and<br />

has spread into desert areas. Several authors note that it is an invasive species and may compete<br />

with native trees. Many municipalities in <strong>the</strong> region have tried to eradicate <strong>the</strong> species with poison,<br />

burning and uprooting, but this is unlikely to be effective since it is now well established in various<br />

habitats and produces large quantities <strong>of</strong> seeds (see also Uses). A prominent individual specimen<br />

stands on <strong>the</strong> island in Doha Bay, formerly known as Palm Tree Island. Since August 2005 it has<br />

not been irrigated but seemingly survives in good condition.<br />

uses: Jongbloed et al. note that <strong>the</strong> ‘pods are nutritious and valuable as fodder’, although Cornes<br />

& Cornes say that <strong>the</strong> foliage is apparently unpalatable as it is never browsed. This species may<br />

be best managed by using for charcoal production, which would have environmental benefits by<br />

reducing <strong>the</strong> need to import charcoal originating from tropical sources. There have been claims<br />

that <strong>the</strong> species causes allergies, but <strong>the</strong>re seems to be a lack <strong>of</strong> substantial scientific evidence for<br />

this. The plant is a traditional remedy for catarrh, colds, dysentery, inflammation, stomach-ache,<br />

sore throats and wounds; <strong>the</strong> root has diuretic properties (Rizk & El-Ghazaly).<br />

Photos: Coll 516; Corn 14, 30, 38; Jong 429; Phil 30 L ; Pick 242.<br />

Rhynchosia minima (L.) DC.<br />

Growth form: Perennial herb. Fl. Mar–Apr.<br />

status: Native. Rare. Also recorded: UAE.<br />

habitat & distribution: Recorded from Al Magdah, N <strong>Qatar</strong>.<br />

uses: Rizk & Ghazaly report that it used in <strong>the</strong> treatment <strong>of</strong> skin diseases.<br />

Photos: Coll 519; Jong 381.<br />

Scorpiurus muricatus L.<br />

Growth form: <strong>An</strong>nual herb. Fl. Mar–Apr.<br />

status: ?Introduced. Rare. Also recorded: Kuwait, E Saudi Arabia, UAE.<br />

habitat & distribution: Recorded from cultivated rodat in N <strong>Qatar</strong>.<br />

uses: ‘Good fodder’ (El Amin).<br />

Photos: Bat 52; Coll 522; Shuaib 89; West 80.<br />

Senna alexandrina Mill.<br />

Cassia senna L.<br />

Vernacular names: senna makki, helul<br />

لولح ,يكم ةنس<br />

Growth form: Shrub. Fl. Mar–Dec.<br />

status: Introduced. ?Local. First published record: Abdel Bari (1997). Also recorded: UAE.<br />

habitat & distribution: A garden weed.<br />

uses: Leaves are used for constipation and stomach cramps (Ghazanfar).<br />

Photos: Jong 382.<br />

Senna italica Mill. Plate 66<br />

Cassia italica (Mill.) Spreng.; Cassia obovata Collad.<br />

لولح ,ةنس ,قرشع<br />

Vernacular names: senna, ’ishriq, senna, helul<br />

Growth form: Shrublet or small shrub. Fl. Mar–May.<br />

status: Native. Common. Also recorded: Bahrain, E Saudi Arabia, UAE.<br />

habitat & distribution: Occasional, particularly on more compacted soils on gravelly and rocky<br />

areas, and also in disturbed areas near to habitation.<br />

uses: Grazed by sheep and goats (El Amin), but Mandaville remarks that some bedouin consider<br />

it toxic to livestock. It is used as a ‘purgative and stimulant and sold in markets for that use’<br />

(Batanouny), but Mandaville says (<strong>of</strong> eastern Arabia) that <strong>the</strong>re is ‘apparently no record <strong>of</strong> such<br />

medicinal use’. Ghazanfar also notes that <strong>the</strong> leaves and seeds are used to treat constipation and<br />

stomach cramps.<br />

Photos: El Amin 12; Bat 47; Coll 523; Corn 1, 39, 125; Jong 383; Mand 105, 106; Phil 34; West 72.<br />

Senna occidentalis (L.) Link<br />

Cassia occidentalis L.<br />

Growth form: <strong>An</strong>nual or short-lived perennial herb. Fl. Not known.<br />

status: Introduced. ?Local. First published record: Abdel Bari (1997). Also recorded: Kuwait.<br />

habitat & distribution: A garden weed.<br />

Photos: Coll 524.<br />

Sesbania sesban (L.) Merr.<br />

Sesbania aegyptiaca (Poir.) Pers.<br />

Vernacular names: sesban<br />

Growth form: Shrub or tree. Fl. Not known.<br />

status: Introduced. Not in o<strong>the</strong>r Gulf floras.<br />

habitat & distribution: Likely to be found planted or naturalised under shade <strong>of</strong> trees along<br />

roadsides or in date plantations. Mandaville comments that in eastern Saudi Arabia it is selfpropogating<br />

in <strong>the</strong> larger oases.<br />

uses: Eaten by man and used for animal fodder; planted for ornamental use and as a soil improver;<br />

saline-tolerant. A species <strong>of</strong> high economic importance in some countries. Leaves are used to treat<br />

skin rashes and wounds (Ghazanfar).<br />

Photos: None.<br />

Taverniera aegyptiaca Boiss.<br />

Growth form: Perennial herb or small shrub. Fl. Mar-Apr.<br />

status: Native. Rare. Also recorded: Bahrain.<br />

habitat & distribution: ‘Long runnels dissecting <strong>the</strong> Miocene country in S <strong>Qatar</strong>’ (Batanouny).<br />

notes: There is some confusion over <strong>the</strong> recording <strong>of</strong> this and <strong>the</strong> next species. Most Gulf floras<br />

only record T. spartea; <strong>the</strong> two are similar in appearance and so Batanouny’s identification may be<br />

in doubt.<br />

Photos: Bat 52; Coll 525; Corn 137; Phil 42.<br />

Taverniera spartea (Burm. f.) DC.<br />

Taverniera aegyptiaca Boiss.<br />

Vernacular names: ’aelijaan, dahseer<br />

Growth form: Shrub. Fl. Mar–Apr.<br />

status: Native. Local. Also recorded: E Saudi Arabia, UAE.<br />

FABACEAE<br />

46 47<br />

ريسهد ,ناجلع<br />

habitat & distribution: Mainly in NE <strong>Qatar</strong>, where locally frequent on more compact soils. Wellgrown<br />

bushes occur at Ras Laffan Industrial City (JN) and <strong>the</strong> species is also found at Al Dhakira,<br />

Fuwairit and Simaismah (MS).<br />

Notes: See under T. aegyptiaca. Plants in <strong>the</strong> NE are presumed to be this species, but have not been<br />

collected or confirmed by an expert.<br />

Photos: Coll 527; Jong 385; Mand 133; West 81.


FABACEAE<br />

Trifolium resupinatum L.<br />

Growth form: <strong>An</strong>nual herb. Fl. Mar–Apr.<br />

status: Introduced. Rare. Also recorded: E Saudi Arabia, UAE.<br />

habitat & distribution: A weed <strong>of</strong> irrigated farmland.<br />

uses: Used as fodder (El Amin).<br />

Photos: West 82.<br />

Trigonella anguina Delile<br />

Vernacular names: nafal<br />

Growth form: <strong>An</strong>nual herb. Fl. Mar–Apr.<br />

status: Native. Common. Also recorded: Bahrain, Kuwait, E Saudi Arabia.<br />

habitat & distribution: Occasional or frequent in silty depressions; mainly N <strong>Qatar</strong>.<br />

uses: Noted as palatable to livestock by Batanouny and El Amin.<br />

Photos: El Amin 84 L ; Bat 53; Coll 532; Daoud 8, 9; Shuaib 90.<br />

Trigonella glabra Thunb.<br />

Trigonella hamosa L., nom. rej.<br />

Vernacular names: Egyptian fenugreek, nafal, quTeifah<br />

Growth form: <strong>An</strong>nual herb. Fl. Mar–Apr.<br />

status: Introduced. Common. Also recorded: Bahrain, Kuwait, E Saudi Arabia, UAE.<br />

habitat & distribution: Frequent or abundant in farms and gardens.<br />

uses: Used as fodder (El Amin). Eaten raw by bedouin as a salad herb (Mandaville).<br />

Photos: El Amin 15, 84 L ; Bat 53; Coll 533; Corn 138; Daoud 4, 5; Jong 390; Shuaib 90; West 83.<br />

Trigonella monantha C.A. Mey.<br />

Vernacular names: nafal<br />

Growth form: <strong>An</strong>nual herb. Fl. Mar–Apr.<br />

status: Introduced. Rare. Also recorded: E Saudi Arabia.<br />

habitat & distribution: Cultivated rodat in C <strong>Qatar</strong>.<br />

Photos: None.<br />

48<br />

لفن<br />

ةفيطق ,لفن<br />

Trigonella stellata Forssk. Plate 67<br />

Vernacular names: star fenugreek, nafal, quTeifah, Hilba bariyah<br />

ةيرب ةبلح ,ةفيطق ,لفن<br />

Growth form: <strong>An</strong>nual herb. Fl. Mar–Apr.<br />

status: Native. Very common. Also recorded: Bahrain, Kuwait, E Saudi Arabia, UAE.<br />

habitat & distribution: Frequent in wadis, runnels, depressions and o<strong>the</strong>r sandy areas.<br />

uses: Batanouny notes that it is palatable to sheep & goats, but Mandaville states that excessive<br />

grazing is reported to cause bloat in livestock. Used by Shammar bedouin women as sweet-scented<br />

hairdressing (Mandaville quoting Dickson 1955).<br />

Photos: El Amin 84 L ; Bat 54; Coll 533; Corn 138; Daoud 6, 7; Jong 391; Mand 116; Phil 43; Shuaib 90.<br />

Vicia monantha Retz.<br />

Growth form: <strong>An</strong>nual herb. Fl. Apr–May.<br />

status: ?Introduced. Rare. Also recorded: Bahrain, E Saudi Arabia, UAE.<br />

لفن<br />

PLATES<br />

1: Avicennia marina 2: Blepharis ciliaris<br />

3: Ophioglossum polyphyllum 4: Avicennia marina 5: Mesembryan<strong>the</strong>mum<br />

cryptanthum<br />

6: Aizoon canariense<br />

7: Mesembryan<strong>the</strong>mum nodiflorum


8: Calotropis procera<br />

9: Glossonema varians<br />

10: Glossonema varians 11: Leptadenia pyrotechnica 12: Filago desertorum<br />

13: Aaronsohnia factorovskyi 14: Atractylis carduus<br />

15: Centaurea sinaica<br />

17: Gymnarrhena micrantha<br />

20: Reichardia tingitana<br />

18: Ifloga spicata<br />

16: Pulicaria undulata<br />

21: Rhanterium epapposum<br />

19: Koelpinia linearis


22: Arnebia hispidissima 23: Echiocaulon jugatum<br />

24: <strong>An</strong>chusa hispida<br />

25: Heliotropium bacciferum 26: Ogastemma pusillum<br />

27: Moltkiopsis ciliata 28: <strong>An</strong>astatica hierochuntica<br />

29: <strong>An</strong>astatica hierochuntica 30: Eremobium aegyptiacum 31: Farsetia heliophila<br />

32: Savignya parviflora 33: Dipterygium glaucum<br />

35: Paronychia arabica<br />

34: Herniaria hemistemon<br />

36: Polycarpaea repens 37: Spergula fallax


38: Sclerocephalus arabicus 39: Silene villosa<br />

40: <strong>An</strong>abasis setifera<br />

41: Atriplex leucoclada 42: Halopeplis perfoliata<br />

43: Agriophyllum minus 44: Haloxylon salicornicum<br />

45: Salicornia europaea 46: Salsola sp.<br />

47: Seidlitzia rosmarinus<br />

50: Helian<strong>the</strong>mum kahiricum<br />

48: Suaeda aegyptiaca 49: Suaeda vermiculata<br />

51: Helian<strong>the</strong>mum lippii


52: Convolvulus cephalopodus<br />

54: Citrullus colocynthis<br />

56: <strong>An</strong>drachne telephioides<br />

53: Convolvulus pilosellifolius 58: Acacia ehrenbergiana 59: Acacia tortilis<br />

55: Cynomorium coccineum<br />

57: Euphorbia retusa<br />

60: Astragalus annularis 61: Astragalus eremophilus<br />

62: Indig<strong>of</strong>era intricata 63: Lotus halophilus


64: Prosopis cineraria<br />

68: Erodium glaucophyllum<br />

65: Prosopis juliflora<br />

66: Senna italica 67: Trigonella stellata<br />

69: Erodium laciniatum 70: Salvia aegyptiaca<br />

71: Frankenia pulverulenta 72: Monsonia nivea<br />

73: Teucrium polium<br />

75: Neurada procumbens<br />

74: Cocculus pendulus<br />

76: Cistanche tubulosa


77: Plantago ciliata<br />

78: Emex spinosa<br />

79: Limonium axillare 80: Scrophularia deserti 81: Fagonia indica<br />

82: Rumex vesicarius 83: Ziziphus nummularia<br />

84: Lycium shawii 85: Corchorus depressus<br />

86: Fagonia ovalifolia 87: Seetzenia lanata 88: Tetraena qatarensis<br />

89: Tetraena simplex 90: Phoenix dactylifera


91: Cyperus conglomeratus<br />

93: Dipcadi erythraeum<br />

95: Chrysopogon plumulosus<br />

92: Cenchrus ciliaris<br />

94: Dipcadi erythraeum<br />

96: Cutandia memphitica<br />

97: Dichanthium foveolatum 98: Stipa capensis (left) and Lasiurus scindicus<br />

99: Aeluropus lagopoides 100: Centropodia forsskalii 101: Chrysopogon plumulosus<br />

102: Lasiurus scindicus 103: Ochthochloa compressa


104: Panicum turgidum 105: Pennisetum divisum<br />

106: Polypogon monspeliensis 107: Sporobolus ioclados 108: Stipa capensis<br />

109: Schismus arabicus<br />

110: Stipagrostis plumosa<br />

habitat & distribution: Recorded as a weed <strong>of</strong> irrigated land.<br />

Photos: Coll 534; West 83.<br />

FRANKENIACEAE<br />

Frankenia pulverulenta L. Plate 71<br />

Vernacular names: moleiH<br />

Growth form: <strong>An</strong>nual herb. Fl. Feb–May.<br />

status: Native. Common. Also recorded: Bahrain, Kuwait, E Saudi Arabia, UAE.<br />

habitat & distribution: Occasional to abundant on saline ground, including cultivated areas and<br />

coastal sand.<br />

uses: Rizk & El-Ghazaly report it to be used as a diuretic.<br />

Photos: El Amin 58 L ; Bat 73; Coll 329; Corn 113; Daoud 41, 42, 43; Jong 393, 394; Phil 163; Shuaib 87;<br />

West 103.<br />

GENTIANACEAE<br />

Centaurium pulchellum (Sw.) Druce<br />

Erythraea pulchella (Sw.) Fries<br />

Vernacular names: lesser centaury<br />

Growth form: <strong>An</strong>nual herb. Fl. Feb–May.<br />

status: Introduced. Rare. Also recorded: Bahrain, E Saudi Arabia, UAE.<br />

habitat & distribution: Recorded by Batanouny in Al Wabrah. Regarded mainly as a naturalised<br />

weed in Arabia.<br />

uses: Various medicinal uses are reported by Rizk & El-Ghazaly.<br />

Photos: Coll 330; Corn 166; Jong 396; Mand 153; Phil 89; West 110.<br />

Enicostema verticillatum (L.) Engl. ex Gilg<br />

Enicostema axillare (Lam.) A. Raynal comb. illeg.<br />

Growth form: Perennial herb. Fl. Not known.<br />

status: Introduced. Rare. First published record: Abdel Bari (1997). Not in o<strong>the</strong>r Gulf floras.<br />

habitat & distribution: Recorded from <strong>Qatar</strong> University car park.<br />

Photos: None.<br />

GERANIACEAE<br />

FABACEAE / FRANKENIACEAE / GENTIANACEAE / GERANIACEAE<br />

Erodium glaucophyllum (L.) L’Hér. Plate 68<br />

Vernacular names: kebaisha, timmair<br />

ريمت ,ةشيبك<br />

Growth form: Perennial herb. Fl. Mar–Apr.<br />

status: Native. Local. Also recorded: Bahrain, Kuwait, E Saudi Arabia.<br />

habitat & distribution: Occasional in gypsum desert, shallow or compacted silty soils. Particularly<br />

numerous at Dukhan (JN); also recorded Ras Laffan (JN), Al Wakra, Al Karaanah (earlier authors).<br />

uses: El Amin notes that it is grazed by animals, although Mandaville says that some bedouin report<br />

that overgrazing on this species can lead to bloat.<br />

Photos: Bat 55; Coll 332; Corn 146; Daoud 227; Mand 149; Phil 90, 91; Shuaib 72.<br />

حيلم<br />

49


GERANIACEAE / LAMIACEAE<br />

Erodium laciniatum (Cav.) Willd. Plate 69<br />

Erodium pulverulentum (Cav.) Willd.<br />

Vernacular names: cut-leaved crane’s-bill, Hambaz, qarnwah<br />

ةونرق ,زابمح<br />

Growth form: <strong>An</strong>nual herb. Fl. Mar–Apr.<br />

status: Native. Common. Also recorded: Bahrain, Kuwait, E Saudi Arabia, UAE.<br />

habitat & distribution: Frequent to abundant in sandy and silty depressions; mainly C and N<br />

<strong>Qatar</strong>.<br />

Photos: Bat 55; Coll 333; Corn 149; Daoud 23, 232, 233; Jong 397; Phil 92; Shuaib 72.<br />

Erodium oxyrrhynchum M. Bieb.<br />

Erodium bryoniifolium Boiss.<br />

Growth form: <strong>An</strong>nual (occasionally perennial) herb. Fl. Mar–Apr.<br />

status: Native. Rare. Also recorded: Kuwait, E Saudi Arabia.<br />

habitat & distribution: Recorded from rodat in N <strong>Qatar</strong>.<br />

Photos: Coll 335; Daoud 228, 229.<br />

Geranium molle L.<br />

Growth form: <strong>An</strong>nual herb. Fl. Mar–Apr.<br />

status: Introduced. Local. Not in o<strong>the</strong>r Gulf floras.<br />

habitat & distribution: Occasional in N and S <strong>Qatar</strong> in rodat and gardens, usually in shade <strong>of</strong><br />

trees.<br />

Photos: Bat 56; Coll 337.<br />

Monsonia heliotropioides (Cav.) Boiss.<br />

Growth form: Perennial herb. Fl. Feb–May.<br />

status: Native. ?Local. Also recorded: E Saudi Arabia, UAE.<br />

habitat & distribution: Reported by Batanouny as common; S and W <strong>Qatar</strong>. El Amin also states<br />

that it occurs at Al Wakra, Al Karaanah and Mekainis.<br />

Photos: Bat 56; Coll 338.<br />

Monsonia nivea (Decne.) Webb Plate 72<br />

Vernacular names: qarnow, dahmah<br />

ةمهد ,ونرق<br />

Growth form: <strong>An</strong>nual or biennial herb. Fl. Mar–Apr.<br />

status: Native. Common. Also recorded: Bahrain, Kuwait, E Saudi Arabia, UAE.<br />

habitat & distribution: Frequent on gravel and stone plains, compacted sand sheets and similar<br />

habitats.<br />

uses: El Amin mentions that some local people boil it to cure fevers.<br />

Photos: Bat 56; Coll 339; Corn 149; Daoud 235; Jong 399; Phil 93; Shuaib 72; West 86.<br />

LAMIACEAE<br />

Salvia aegyptiaca L. Plate 70<br />

Vernacular names: na’aim/na’eem, ra’al<br />

Growth form: Shrublet or small shrub. Fl. Mar–May.<br />

status: Native. Common. Also recorded: Bahrain, Kuwait, E Saudi Arabia, UAE.<br />

لعر ,ميعن<br />

habitat & distribution: Occasional in shallow silty or sandy soils, in rocky areas, also in depressions<br />

with coarse soils.<br />

uses: The plant is used in <strong>the</strong> treatment <strong>of</strong> diarrhoea, gonorrhoea, haemorrhoids and eye diseases<br />

(Rizk & El-Ghazaly, Ghazanfar).<br />

Photos: El Amin 12; Bat 87; Coll 460; Corn 199; Daoud 248; Jong 405; Phil 137; Pick 32; West 127.<br />

Teucrium polium L. Plate 73<br />

Vernacular names: ja’ad/yaad<br />

Growth form: Perennial herb. Fl. Apr–May.<br />

status: Native. Local. Also recorded: Bahrain, Kuwait, E Saudi Arabia, UAE.<br />

habitat & distribution: Stony runnels and silty soils; C and N <strong>Qatar</strong>. Occasional at Ras Laffan<br />

(JN) and mentioned for Jebel Fuwairit by earlier authors.<br />

uses: Many reputed medicinal uses, e.g. for fevers and cholera (El Amin), malaria and insect bites<br />

(Jongbloed et al., Mandaville). O<strong>the</strong>r medicinal uses are listed by Rizk & El-Ghazaly.<br />

Photos: El Amin 12, 64 L ; Bat 88, 89 L ; Coll 467; Corn 200; Daoud 244; Mand 176; Phil 138; Shuaib 69.<br />

LINACEAE<br />

Linum strictum L.<br />

Growth form: <strong>An</strong>nual herb. Fl. Mar–Apr.<br />

status: Native. ?Rare. Also recorded: Bahrain.<br />

habitat & distribution: Recorded by El Amin from ‘C <strong>Qatar</strong>, Wakra hills’ (presumably Jebel Al<br />

Wakra, E <strong>Qatar</strong>). Habitat is given as light soils on depressions.<br />

uses: The seeds yield an oil similar to linseed oil (El Amin).<br />

Photos: El Amin 87 L .<br />

LORANTHACEAE<br />

Plicosepalus acaciae (Zucc.) Wiens & Polhill<br />

Loranthus acaciae Zucc.<br />

لدح<br />

Growth form: Climbing, parasitic shrub. Fl. Sep–Oct.<br />

status: Native. Local. First record: Al-Madheed (2004). Not in o<strong>the</strong>r Gulf floras.<br />

habitat & distribution: Occasional, growing on Acacia and Lycium shawii; especially rodat in C &<br />

N <strong>Qatar</strong>.<br />

Photos: Pick 70.<br />

Vernacular names: Hadal<br />

LYTHRACEAE<br />

Lawsonia inermis L.<br />

Lawsonia alba Lam.<br />

Vernacular names: henna, Henna/Hinna<br />

Growth form: Tree. Fl. Sep–Feb.<br />

status: Introduced. Also recorded: E Saudi Arabia, UAE.<br />

LAMIACEAE / LINACEAE / LORANTHACEAE / LyTHRACEAE<br />

habitat & distribution: Planted in urban areas in <strong>the</strong> Gulf, but distribution in <strong>Qatar</strong> uncertain.<br />

50 51<br />

دعج<br />

انح


LyTHRACEAE / MALVACEAE<br />

uses: Widely used in <strong>the</strong> Middle East as a dye for painting hands and feet and worldwide for colouring<br />

hair and beards. The leaves are used for fevers, as a local anaes<strong>the</strong>tic and anti-inflammatory and for<br />

treating mouth ulcers (Ghazanfar). Jongbloed et al. remark that it is one <strong>of</strong> <strong>the</strong> traditional ‘plants<br />

<strong>of</strong> paradise’ which should not be cut for fuel.<br />

Photos: Bat 16; Jong 441; Pick 179.<br />

MALVACEAE<br />

Abutilon fruticosum Guill. & Perr.<br />

Vernacular names: lowaq, ja’ja’an<br />

Growth form: Shrub. Fl. Jan–Apr.<br />

status: Introduced. ?Local. Also recorded: UAE.<br />

habitat & distribution: Occasional in rodat; C and N <strong>Qatar</strong>.<br />

Photos: Bat 70; Coll 549; Ghaz 129.<br />

Abutilon pannosum (G. Forst.) Schltdl.<br />

Vernacular names: lowaq<br />

Growth form: Shrub. Fl. Apr–Jun.<br />

status: Introduced. ?Local. Also recorded: E Saudi Arabia, UAE.<br />

habitat & distribution: Occasional on rodat in C <strong>Qatar</strong>.<br />

ناعجعج ,قاول<br />

notes: Abutilon hirtum (Lam.) Sweet is listed as a synonym by Jongbloed, though usually regarded<br />

as a separate species.<br />

Photos: Bat 70; Coll 551; Jong 413; Pick 132; West 99.<br />

Althaea ludwigii L.<br />

Vernacular names: khatma<br />

Growth form: <strong>An</strong>nual or biennial herb. Fl. Mar–Apr.<br />

status: Native. Common. Also recorded: Kuwait, E Saudi Arabia, UAE.<br />

habitat & distribution: Occasional in silty, loamy and gravelly soils; mainly in depressions in C<br />

<strong>Qatar</strong>.<br />

Photos: El Amin 89 L ; Bat 69; Coll 552; Daoud 52; Jong 414; Shuaib 56.<br />

Malva nicaeensis All.<br />

Vernacular names: khubeizah<br />

Growth form: <strong>An</strong>nual or biennial herb. Fl. Mar–Apr.<br />

status: Introduced. Rare. Not in o<strong>the</strong>r Gulf floras.<br />

habitat & distribution: Recorded only once by Batanouny from a nor<strong>the</strong>rn farm.<br />

Photos: None.<br />

Malva parviflora L.<br />

Vernacular names: least mallow, cheeseweed, khubeizah<br />

Growth form: <strong>An</strong>nual herb. Fl. Feb–Apr.<br />

status: Native. Very common. Also recorded: Bahrain, Kuwait, E Saudi Arabia, UAE.<br />

habitat & distribution: Occasional to frequent in sandy soils after rains; is more common in and<br />

around inhabited and cultivated areas.<br />

قاول<br />

ةمتخ<br />

هزيبخ<br />

هزيبخ<br />

uses: Mandaville quotes earlier authors who reported that <strong>the</strong> leaves are eaten as a vegetable by<br />

Persians in Kuwait. Phillips notes that <strong>the</strong> leaves were used in <strong>the</strong> past to treat coughs and bladder<br />

complaints. Rizk & El-Ghazaly list a number <strong>of</strong> medicinal uses and state that <strong>the</strong> plant is poisonous<br />

to cattle and horses. Ghazanfar reports that <strong>the</strong> seeds and leaves are also used to treat fevers and<br />

ulcers.<br />

Photos: El Amin 16; Coll 557; Corn 117; Daoud 50, 51; Jong xix, 416; Phil 129; Pick 181; Shuaib 56; West<br />

100.<br />

MENISPERMACEAE<br />

Cocculus pendulus (J.R. & G. Forst.) Diels Plate 74<br />

قينخ<br />

Vernacular names: khuneeq<br />

Growth form: Climber. Fl. Mar–Apr.<br />

status: Native. Rare. Also recorded: UAE.<br />

habitat & distribution: Recorded from Al Karaanah and Al Kharrarah. Still at <strong>the</strong> latter location,<br />

and also present at <strong>Qatar</strong> University campus (MS). Usually grows over Ziziphus or Acacia trees.<br />

uses: Fruit edible (El Amin). Used in <strong>the</strong> treatment <strong>of</strong> fevers (Rizk & El-Ghazaly).<br />

Photos: El Amin 16; Bat 40; Coll 563; Ghaz 8, 9; Jong 421; West 55, 56.<br />

MOLLUGINACEAE<br />

Mollugo cerviana (L.) Ser.<br />

Vernacular names: carpetweed<br />

Growth form: <strong>An</strong>nual herb. Fl. Apr.<br />

status: Introduced. ?Rare. Not in o<strong>the</strong>r Gulf floras.<br />

habitat & distribution: Previously recorded by Batanouny from farmed rodat in N and C <strong>Qatar</strong>.<br />

uses: A number <strong>of</strong> medicinal properties are listed by Rizk & El-Ghazaly.<br />

Photos: Coll 565.<br />

MORACEAE<br />

Ficus carica L.<br />

Vernacular names: fig, teen<br />

نيت<br />

Growth form: Shrub or tree. Fl. Not known.<br />

status: Introduced. Also recorded: Bahrain, E Saudi Arabia, UAE.<br />

habitat & distribution: One <strong>of</strong> <strong>the</strong> more widely planted fig species. It may occasionally escape<br />

into <strong>the</strong> wild.<br />

uses: Grown for fruit and as an ornamental plant.<br />

Photos: Ghaz 16; West 31.<br />

Morus nigra L.<br />

MALVACEAE / MENISPERMACEAE / MOLLUGINACEAE / MORACEAE<br />

Vernacular names: black mulberry<br />

Growth form: Tree. Fl. Not known.<br />

status: Introduced. Also recorded: Bahrain, E Saudi Arabia.<br />

habitat & distribution: Present distribution in <strong>Qatar</strong> not known but widespread in <strong>the</strong> Gulf.<br />

uses: Ornamental planting.<br />

Photos: none.<br />

52 53


MyRSINACEAE / NEURADACEAE / OROBANCHACEAE<br />

MYRSINACEAE<br />

<strong>An</strong>agallis arvensis L.<br />

Vernacular names: ’ayn al qaT (cat’s-eye), ’uwaynah (little eye)<br />

ةنيوع ،طقلا نيع<br />

blue/scarlet pimpernel<br />

Growth form: <strong>An</strong>nual herb. Fl. Mar–Apr.<br />

status: Introduced. Common. Also recorded: Bahrain, Kuwait, E Saudi Arabia, UAE.<br />

habitat & distribution: Occasional in cultivated areas and rodat with trees, particularly in N and<br />

NW <strong>Qatar</strong>.<br />

uses: Ghazanfar reports <strong>the</strong> whole plant is used to treat skin rash and ulcers.<br />

notes: Formerly placed in <strong>the</strong> Primulaceae. The blue-flowered form, currently known as A. arvensis<br />

subsp. foemina Mill., appears to be <strong>the</strong> usual form in <strong>Qatar</strong> and o<strong>the</strong>r parts <strong>of</strong> <strong>the</strong> Gulf, but <strong>the</strong><br />

scarlet-flowered form has also been reported.<br />

Photos: Bat 75; Coll 621; Daoud 195; Ghaz 209-10; Jong 473; Mand 103; Phil 157, 158; Pick 35; Shuaib<br />

56; West 108.<br />

NEURADACEAE<br />

Neurada procumbens L. Plate 75<br />

Vernacular names: creeping thorn rose, sa’adan<br />

ناديعس<br />

Growth form: <strong>An</strong>nual herb. Fl. Mar–Apr.<br />

status: Native. Common. Also recorded: Bahrain, Kuwait, E Saudi Arabia, UAE.<br />

habitat & distribution: Occasional to frequent in shallow or deep sand; mainly S <strong>Qatar</strong>.<br />

uses: It is well known that <strong>the</strong> seed pods embed <strong>the</strong>mselves in <strong>the</strong> hooves <strong>of</strong> livestock and can cause<br />

lameness. El Amin states that <strong>the</strong> fruit is ‘edible by man and loved by many animals’ and Mandaville<br />

that <strong>the</strong> fruit is ‘not unpleasant… plucked well before <strong>the</strong>y begin to harden’.<br />

Photos: El Amin 17; Bat 40; Coll 573; Daoud 1; Jong 438; Mand 104; Phil 147; Shuaib 105; West 68.<br />

OROBANCHACEAE<br />

Cistanche tubulosa (Schrenk) Wight Plate 76<br />

Cistanche lutea Wight; Cistanche phelypaea (L.) Cout.; Cistanche tinctoria (Forssk.) G. Beck.<br />

ثوترط ,نونذ<br />

Vernacular names: desert hyacinth, dhanoon, Tartuth<br />

Growth form: Parasitic herb. Fl. Mar–Apr.<br />

status: Native. Common. Also recorded: Bahrain, Kuwait, E Saudi Arabia, UAE.<br />

habitat & distribution: Mainly in coastal areas, including coastal sands along beaches, where<br />

occasional or forming small clumps. Host plants include Arthrocnemum macrostachyum, Cornulaca<br />

spp, Limonium axillare, Suaeda vermiculata and Tetraena qatarensis.<br />

uses: ‘Said to be poisonous’ (El Amin). Rizk & El-Ghazaly describe its use as an aphrodisiac, and a<br />

tonic to cure impotence and diarrhoea.<br />

notes: C. phelypaea is here combined with C. tubulosa; but it is possible that <strong>the</strong>y are distinct but<br />

closely-related species and that both occur in <strong>Qatar</strong>.<br />

Photos: El Amin 17; Bat 93; Coll 586; Corn 40, 186; Daoud 22, 224, 225; Jong 444; Mand 189, 190; Phil<br />

54, 55; Pick 140; Shuaib 52; West 135.<br />

Orobanche ramosa L.<br />

Vernacular names: hemp broomrape, branched broomrape<br />

Growth form: Parasitic herb. Fl. Mar.<br />

status: Introduced. ?Rare. First published record: Abdel Bari (1997). Also recorded: Kuwait.<br />

habitat & distribution: A parasite on Solanum nigrum.<br />

Photos: Pick 36; Shuaib 53.<br />

OxALIDACEAE<br />

Oxalis corniculata L.<br />

Vernacular names: creeping sorrel, Hamd/muHummad<br />

Growth form: <strong>An</strong>nual herb. Fl. Apr–Jun.<br />

status: Introduced. Local. Also recorded: Kuwait, E Saudi Arabia, UAE.<br />

habitat & distribution: A weed <strong>of</strong> lawns and gardens in Doha.<br />

54 55<br />

دمحم/دمح<br />

uses: The plant has numerous medicinal uses, including <strong>the</strong> treatment <strong>of</strong> bruises, clots, fever, snake<br />

bite and infections (Rizk & El-Ghazaly).<br />

Photos: Coll 591; Jong 447; Pick 133; Shuaib 73.<br />

Oxalis debilis Kunth<br />

Oxalis corymbosa DC.<br />

Growth form: Perennial herb. Fl. Not known.<br />

status: Introduced. Rare. First published record: Abdel Bari (1997). Not in o<strong>the</strong>r Gulf floras.<br />

habitat & distribution: Recorded as a weed <strong>of</strong> gardens.<br />

notes: Plants recorded in <strong>Qatar</strong> are probably referable to var. corymbosa (DC.) Lourteig.<br />

Photos: None.<br />

Oxalis pes-caprae L.<br />

Oxalis cernua Thunb.<br />

Growth form: <strong>An</strong>nual herb. Fl. Mar–Apr.<br />

status: Introduced. ?Local. Also recorded: E Saudi Arabia.<br />

habitat & distribution: Cultivated areas, C and N <strong>Qatar</strong>. A weed <strong>of</strong> garden origin.<br />

uses: ‘Leaves eaten in Iran in stews but bitter in taste’ (El Amin).<br />

Photos: Bat 55.<br />

PLANTAGINACEAE<br />

Plantago amplexicaulis Cav.<br />

OROBANCHACEAE / OxALIDACEAE / PLANTAGINACEAE<br />

Vernacular names: rabl, rablat al mistaH<br />

Growth form: <strong>An</strong>nual herb. Fl. Feb–Apr.<br />

status: Native. Common. Also recorded: Kuwait, E Saudi Arabia, UAE.<br />

habitat & distribution: Occasional in silty and sandy depressions in C and N <strong>Qatar</strong>.<br />

uses: The plant has various medicinal properties.<br />

Photos: El Amin 94 L ; Bat 94; Coll 601; Daoud 197; Jong 452; Shuaib 103.<br />

حطسملا ةلبر ,لبر


PLANTAGINACEAE<br />

Plantago ciliata Desf. Plate 77<br />

Vernacular names: rabl, lisan al Hamal, quraiTah, wedaina<br />

ةنيدو ,ةطيرق ,لمحلا ناسل ,لبر<br />

Growth form: <strong>An</strong>nual herb. Fl. Feb–Apr.<br />

status: Native. Local. Also recorded: Bahrain, Kuwait, E Saudi Arabia, UAE.<br />

habitat & distribution: Only a single plant was found during a survey <strong>of</strong> western <strong>Qatar</strong> in 2007<br />

(JN and S. Aspinall). Batanouny recorded it as ‘fairly common especially in sandy habitats in S<br />

<strong>Qatar</strong>’.<br />

Photos: El Amin 17; Bat 94; Coll 602; Daoud 201; Jong 454; Shuaib 103; West 137.<br />

Plantago coronopus L.<br />

Vernacular names: buck’s-horn plantain, rabl, wedaina<br />

ةنيدو ,لبر<br />

Growth form: <strong>An</strong>nual herb. Fl. Mar–Apr.<br />

status: Native. Rare. Also recorded: Bahrain, Kuwait, E Saudi Arabia.<br />

habitat & distribution: Previously recorded from Dukhan; recorded twice recently in W <strong>Qatar</strong>.<br />

uses: The roots are used to treat haemorrhoids, malaria and fevers; <strong>the</strong> leaves are locally applied<br />

to wounds, burns, abscesses, bites and inflammations (Rizk & El-Ghazaly).<br />

notes: A specimen collected from western <strong>Qatar</strong> (JN, 2007) was determined by E. Clement as<br />

Plantago commutata Guss. (P. coronopus subsp. commutata (Guss.) Pilg.).<br />

Photos: Coll 602; Daoud 196; Phil 155; Shuaib 104.<br />

Plantago lanceolata L.<br />

Vernacular names: rabl<br />

Growth form: Perennial herb. Fl. Mar–Apr.<br />

status: Introduced. Local. Also recorded: Kuwait.<br />

habitat & distribution: Occasional in cultivated areas in C <strong>Qatar</strong>.<br />

uses: Medicinal uses are described by Rizk & El-Ghazaly.<br />

Photos: Coll 603; Shuaib 104.<br />

Plantago ovata Forssk.<br />

Vernacular names: rabl<br />

Growth form: <strong>An</strong>nual herb. Fl. Feb–Apr.<br />

status: Native. Local. Also recorded: Bahrain, Kuwait, E Saudi Arabia, UAE.<br />

habitat & distribution: Occasional in sandy habitats; S <strong>Qatar</strong>.<br />

uses: Seeds reportedly have been used by bedouin as a laxative (Mandaville). Rizk & El-Ghazaly<br />

report many medicinal uses and Ghazanfar that <strong>the</strong> seeds are used as an astringent, a diuretic and<br />

for treating venereal disease.<br />

Photos: El Amin 94 L ; Coll 604; Daoud 199; Jong 455; Mand 184; Shuaib 104; West 137.<br />

Plantago psyllium L.<br />

Vernacular names: rabl<br />

لبر<br />

Growth form: <strong>An</strong>nual herb. Fl. Apr.<br />

status: Native. Rare. Not in o<strong>the</strong>r Gulf floras.<br />

habitat & distribution: Recorded on <strong>the</strong> road from Doha to N <strong>Qatar</strong> near Umm Slal Mohammed<br />

by Batanouny.<br />

Photos: None.<br />

لبر<br />

لبر<br />

PLUMBAGINACEAE<br />

Limonium axillare (Forssk.) Kuntze Plate 79<br />

Statice axillaris Forssk.<br />

فطق<br />

Vernacular names: sea lavender, qaTaf<br />

Growth form: Small shrub. Fl. Mar–Apr.<br />

status: Native. Common. Also recorded: Bahrain, E Saudi Arabia, UAE.<br />

habitat & distribution: Frequent to dominant on beaches, sabkha edge and o<strong>the</strong>r saline habitats<br />

around <strong>the</strong> coasts.<br />

uses: Ghazanfar states <strong>the</strong> whole plant is used against diarrhoea and as an astringent.<br />

notes: The related species Limonium carnosum (Boiss.) Kuntze is recorded from Kuwait, Saudi<br />

Arabia, UAE and doubtfully Oman (Ghazanfar 2003), and is <strong>the</strong>refore quite likely to occur in <strong>Qatar</strong>.<br />

Photos: El Amin 17; Bat 76; Coll 605; Corn 94; Ghaz 93; Jong 457; Mand 61; Phil 20, 164; Pick 76; West 109.<br />

POLYGALACEAE<br />

Polygala erioptera DC.<br />

Growth form: <strong>An</strong>nual herb. Fl. Mar–Apr.<br />

status: Native. Common. Also recorded: E Saudi Arabia, UAE.<br />

habitat & distribution: Occasional in sandy areas and rodat in C <strong>Qatar</strong> including Dukhan area<br />

and Al Wakra.<br />

uses: Jongbloed et al. state that <strong>the</strong> generic name comes from <strong>the</strong> belief that ‘animals feeding on<br />

<strong>the</strong>se species would produce more milk’.<br />

Photos: El Amin 97 L ; Bat 65; Coll 607; Jong 459; West 96.<br />

Polygala irregularis Boiss.<br />

Growth form: <strong>An</strong>nual herb. Fl. Apr.<br />

status: Native. Common. Also recorded: UAE.<br />

habitat & distribution: Reported by earlier authors as common, especially in NE <strong>Qatar</strong> on sandy<br />

or gravelly soils.<br />

Photos: Coll 608; Jong 460.<br />

POLYGONACEAE<br />

PLUMBAGINACEAE / POLyGALACEAE / POLyGONACEAE<br />

Calligonum comosum L’Hér.<br />

Calligonum polygonoides subsp. comosum (L’Hér.) Soskov<br />

Vernacular names: abal, arta<br />

Growth form: Shrub. Fl. Mar–Apr.<br />

status: Native. Rare. Also recorded: Bahrain, Kuwait, E Saudi Arabia, UAE.<br />

habitat & distribution: Recorded by El Amin from a sand dune near Umm Bab.<br />

uses: ‘The fruits are edible and freshen <strong>the</strong> mouth with <strong>the</strong>ir tart taste’ (Cornes & Cornes). Twigs<br />

said to be pounded and added to milk as a flavouring or tonic (Mandaville). Cornes & Cornes also<br />

mention that it is said to be used in a balm for skin ailments. El Amin says <strong>the</strong> woody base was used<br />

for firewood. Also reported by early authors to be used by bedouin in <strong>the</strong> tanning <strong>of</strong> hides.<br />

Photos: El Amin 18, 99 L ; Coll 611; Corn 32, 90; Daoud 154, 155; Ghaz 86, 87, 88; Jong 462, 463; Phil 83, 84;<br />

Pick 185; Shuaib 100; West 32.<br />

56 57


POLyGONACEAE<br />

Emex spinosa (L.) Campd. Plate 78<br />

Rumex spinosus L.<br />

Vernacular names: prickly dock, Hanzab/Hambaz, Ders al ’ajooz<br />

زوجعلا سرض ,زابمح/بازنح<br />

Growth form: <strong>An</strong>nual herb. Fl. Mar–Apr.<br />

status: Native. Common. Also recorded: Bahrain, Kuwait, E Saudi Arabia, UAE.<br />

habitat & distribution: Occasional or frequent in sandy habitats.<br />

uses: Phillips and Mandaville note that <strong>the</strong> plant is edible. It is used as a remedy for stomach<br />

disorders (Rizk & El-Ghazaly).<br />

Photos: Bat 18; Coll 612; Corn 93; Daoud 156, 157; Jong 465; Phil 85; Shuaib 99; West 32.<br />

Polygonum argyrocoleum Steud. ex Kunze<br />

Vernacular names: ghardab<br />

بدرغ<br />

Growth form: <strong>An</strong>nual herb. Fl. Not known.<br />

status: ?Introduced. ?Local. Also recorded: E Saudi Arabia, UAE.<br />

habitat & distribution: A weed <strong>of</strong> moist ground in cultivated rodat.<br />

notes: The record <strong>of</strong> P. equisetiformis by Obeid (1975) reported by Batanouny, a Mediterranean<br />

species, is probably referable to this species (see Miller & Cope 1996, p. 129) as is Polygonum bellardii<br />

Sibth. & Sm. (P. bellardii All.) recorded by El Amin. Miller & Cope do not list P. bellardii for Arabia.<br />

Photos: Coll 613.<br />

Rumex cyprius Murb.<br />

Growth form: <strong>An</strong>nual herb. Fl. Mar–Apr.<br />

status: ?Introduced. Rare. Not in o<strong>the</strong>r Gulf floras.<br />

habitat & distribution: Recorded by Batanouny from silty depressions.<br />

uses: ‘It is grazed by animals and <strong>the</strong> leaves are eaten by man as salad or cooked in stews’ (El Amin).<br />

Photos: Coll 615.<br />

Rumex dentatus L.<br />

Vernacular names: khillah<br />

Growth form: <strong>An</strong>nual herb. Fl. Apr.<br />

status: ?Introduced. Rare. Also recorded: E Saudi Arabia, UAE.<br />

habitat & distribution: Previously recorded from damp, sandy soils and silty depressions in<br />

cultivated areas.<br />

uses: Rizk & El-Ghazaly mention that an infusion <strong>of</strong> <strong>the</strong> plant is used as a cooling agent against<br />

sunstroke and that <strong>the</strong> root is astringent and applied to skin disorders.<br />

Photos: Coll 615; Jong 467.<br />

Rumex vesicarius L. Plate 82<br />

Vernacular names: sorrel, bladderdock, hummayD<br />

Growth form: <strong>An</strong>nual herb. Fl. Mar–Apr.<br />

status: Native. Local. Also recorded: Kuwait, E Saudi Arabia, UAE.<br />

ةلخ<br />

ضيمح<br />

habitat & distribution: Occasional in rocky terrain in shallow deposits <strong>of</strong> sand or silt, but also<br />

recorded in cultivated areas.<br />

uses: The leaves can be eaten raw or cooked. Mandaville reports from an earlier source that it was<br />

sometimes added during preparation <strong>of</strong> iqt (dried milk shards) to increase <strong>the</strong> acidity. A long list <strong>of</strong><br />

medicinal properties are given by Rizk & El-Ghazaly and o<strong>the</strong>r authors. Ghazanfar states that <strong>the</strong><br />

leaves and seeds are used to treat scorpion stings.<br />

Photos: El Amin 18; Bat 18; Coll 616; Daoud 158, 159; Ghaz 85; Jong 470; Pick 79; Shuaib 100; West 34.<br />

PORTULACACEAE<br />

Portulaca oleracea L.<br />

Vernacular names: purslane, barbir, baqlah, rijlah<br />

ةلجر ,ةلقب ,ريبرب<br />

Growth form: <strong>An</strong>nual herb. Fl. Apr–Sep.<br />

status: Introduced. Common. Also recorded: Bahrain, Kuwait, E Saudi Arabia, UAE.<br />

habitat & distribution: Frequent in cultivated areas, including gardens.<br />

uses: A salad herb, widely cultivated and available in many <strong>of</strong> <strong>Qatar</strong>’s markets as well as grocery<br />

stores and supermarkets. Is also highly palatable to animals. Jongbloed et al. also note that it is used<br />

as a bactericide, and in <strong>the</strong> treatment <strong>of</strong> worms, among o<strong>the</strong>r uses. Phillips said it was eaten to help<br />

digestion and o<strong>the</strong>r authors also note its diuretic properties.<br />

Photos: El Amin 18; Bat 20; Coll 618; Corn 89, 272; Jong 471; Phil 159; Pick 135; Shuaib 73; West 36.<br />

RESEDACEAE<br />

Ochradenus aucheri Boiss.<br />

Ochradenus dewittii Abdallah<br />

Growth form: Shrub. Fl. Oct-May.<br />

status: Native. Rare. Also recorded: UAE.<br />

habitat & distribution: It is normally a species <strong>of</strong> rocky habitats. Batanouny collected this species<br />

near Al Wukair.<br />

Photos: Ghaz 202; Jong 476, 478; West 67.<br />

Ochradenus baccatus Delile<br />

POLyGONACEAE / PORTULACACEAE / RESEDACEAE<br />

Vernacular names: qurDi<br />

يضرق<br />

Growth form: Shrub. Fl. Mar–Apr.<br />

status: Native. Common. Also recorded: Bahrain, Kuwait, E Saudi Arabia, UAE.<br />

habitat & distribution: Occasional or frequent in sandy or gravelly depressions, growing amongst<br />

Acacia, Ziziphus and Lycium. It can be seen by <strong>the</strong> roadside <strong>of</strong> <strong>the</strong> Al Shamal highway from north <strong>of</strong><br />

Doha to Al Khor and from Al Rayyan to Rawdat Rashed (MS); strong populations also occur inside<br />

<strong>Qatar</strong> University and <strong>the</strong> surrounding areas.<br />

uses: Cornes & Cornes say that it is mainly eaten by goats which strip <strong>the</strong> plants down to a few<br />

centimetres. The twigs, leaves and flowers are applied to wounds and sores to kill maggots (Rizk &<br />

El-Ghazaly). Ghazanfar reports <strong>the</strong> fruit and leaves are used to treat stomach-ache.<br />

Photos: Bat 38; Coll 638; Corn 110; Daoud 12, 119, 120; Ghaz 196-7; Mand 98, 99; Phil 130; Shuaib 101.<br />

Oligomeris linifolia (Vahl) J.F. Macbr.<br />

Oligomeris subulata (Delile) Boiss.; Reseda subulata Delile<br />

Growth form: <strong>An</strong>nual or biennial herb. Fl. Mar–Apr.<br />

status: Native. Very common. Also recorded: Bahrain, Kuwait, E Saudi Arabia, UAE.<br />

habitat & distribution: Frequent to abundant on harder substrates; mainly C and S <strong>Qatar</strong>.<br />

uses: Grazed by animals, especially goats and sheep (El Amin).<br />

Photos: Bat 39; Coll 639; Corn 113; Daoud 122; Jong 479; Phil 131; Shuaib 101.<br />

58 59


RESEDACEAE / RHAMNACEAE<br />

Reseda arabica Boiss.<br />

Vernacular names: shawlah<br />

Growth form: <strong>An</strong>nual herb. Fl. Mar–Apr.<br />

status: Native. Rare. Also recorded: Kuwait, E Saudi Arabia, UAE.<br />

habitat & distribution: Recorded from C and S <strong>Qatar</strong>, including Umm Bab.<br />

Photos: Coll 639; Daoud 12, 126, 127; Mand 100; Shuaib 102.<br />

Reseda aucheri Boiss.<br />

Vernacular names: shawlah<br />

Growth form: <strong>An</strong>nual or perennial. Fl. Mar–Apr.<br />

status: Native. ?Rare. Also recorded: Kuwait, UAE.<br />

habitat & distribution: El Amin recorded it from gravelly depressions in C <strong>Qatar</strong>. The habitat in<br />

<strong>the</strong> UAE is gravel plains and mountains (Jongbloed).<br />

notes: The plant occurring in <strong>Qatar</strong> and o<strong>the</strong>r parts <strong>of</strong> <strong>the</strong> Gulf is R. aucheri var. bracteata (Boiss.)<br />

Abdallah & De Wit.<br />

Photos: Coll 639; Ghaz 203; Jong 480; Pick 186; West 67.<br />

Reseda muricata C. Presl.<br />

Vernacular names: shawlah<br />

Growth form: Perennial herb. Fl. Mar–May.<br />

status: Native. Common. Also recorded: Bahrain, Kuwait, E Saudi Arabia.<br />

habitat & distribution: Occasional in compacted silty sand, in gypsum desert, and in shallows<br />

soil in rocky areas.<br />

uses: The plant possesses antimicrobial activity (Rizk & El-Ghazaly).<br />

Photos: El Amin 106 L ; Bat 39; Coll 641; Daoud 13, 129, 130; Mand 101; Shuaib 102.<br />

RHAMNACEAE<br />

Ziziphus mauritiana Lam.<br />

Vernacular names: sidr, nabaq (fruit), kenar (fruit)<br />

Growth form: Tree. Fl. Mar–Apr.<br />

status: Introduced. Common. Not in o<strong>the</strong>r Gulf floras.<br />

ةلوش<br />

ةلوش<br />

ةلوش<br />

)ةرمث( رانك ,)ةرمث( قبن ,ردس<br />

habitat & distribution: Occasional to locally frequent where planted in farms, gardens and<br />

roadsides. Rarely naturalised.<br />

uses: Planted along roadsides as a shade tree. The fruits are edible.<br />

Photos: El Amin 18; Bat 68.<br />

Ziziphus nummularia (Burm. f.) Wight & Arn. Plate 83<br />

Vernacular names: sidr, nabaq (fruit), kenar (fruit), dom (fruit)<br />

Growth form: Shrub or tree. Fl. Mar–May.<br />

status: Native. Local. Also recorded: Bahrain, E Saudi Arabia.<br />

)ةرمث( رانك ,)ةرمث( قبن ,ردس<br />

habitat & distribution: Occurs in C and S <strong>Qatar</strong> in sandy depressions; <strong>of</strong>ten dominant or mixed<br />

with Acacia spp.<br />

uses: The fruit is edible, sweet and ‘also used to treat coughs’ (El Amin). The species can be used<br />

for firewood. El Amin also comments on its ability to trap sand and control erosion. Mandaville<br />

citing earlier o<strong>the</strong>rs reports that <strong>the</strong> sidr tree (including Z. spina-christi) was used to make <strong>the</strong> hookended<br />

camel stick known as <strong>the</strong> mush’ab and that some tribes had a tradition that sidr thickets were<br />

haunted by jinn (spirits), so were not cut for fuel. <strong>An</strong> infusion <strong>of</strong> <strong>the</strong> leaves is used to treat stomach<br />

complaints, sore throats and ease joint pains, whilst <strong>the</strong> leaves are also applied to treat scabies and<br />

boils (El Amin, Rizk & El-Ghazaly).<br />

Photos: El Amin 19; Bat 66 L , 67; Coll 645; Corn 158; Mand 136.<br />

Ziziphus spina-christi (L.) Desf.<br />

Vernacular names: Christ’s thorn, sidr, nabaq (fruit)<br />

)ةرمث( قبن ,ردس<br />

Growth form: Tree. Fl. Mar–Apr.<br />

status: Introduced. Local. Also recorded: UAE.<br />

habitat & distribution: Occasional to frequent in farms and roadside plantations. Rarely found<br />

naturalised.<br />

uses: A list <strong>of</strong> medicinal uses is given by Rizk & El-Ghazaly. Ghazanfar reports that <strong>the</strong> leaves, fruit<br />

and seeds are used to treat blisters, bruises, chest pains, dandruff, fractures and mouth and gum<br />

problems.<br />

Photos: Bat 68; Coll 645; Jong 481; Pick 247; West 97.<br />

RUBIACEAE<br />

Galium tricornutum Dandy<br />

Growth form: <strong>An</strong>nual herb. Fl. Mar–Apr.<br />

status: Introduced. Rare. Also recorded: Bahrain, Kuwait, E Saudi Arabia, UAE.<br />

habitat & distribution: Recorded from Al Magdah by Batanouny, growing over Ziziphus.<br />

Photos: Jong 488; Shuaib 57.<br />

RUTACEAE<br />

Haplophyllum tuberculatum (Forssk.) Juss.<br />

Vernacular names: musaykah (‘muskweed’), Tafr at-tais/sinan at-tais سيتلا نانس/سيتلا رفط ,ةكيسم<br />

(‘smell <strong>of</strong> <strong>the</strong> goat’)<br />

Growth form: Perennial herb. Fl. Mar–Apr.<br />

status: Native. Local. Also recorded: Bahrain, Kuwait, E Saudi Arabia, UAE.<br />

habitat & distribution: Occasional in S, W and C <strong>Qatar</strong> on sandy soils.<br />

uses: El Amin remarks that <strong>the</strong> plant is ‘not liked’ as grazing, and that <strong>the</strong> ‘bedouin use it to<br />

cure scorpion stings’. Ghazanfar mentions <strong>the</strong> leaves are used to ‘streng<strong>the</strong>n back muscles after<br />

childbirth, for chest pains, flatulence, stomach problems and as a sedative’.<br />

Photos: Bat 65; Coll 662; Daoud 71; Jong 493; Mand 137; Pick 138; Shuaib 58; West 95.<br />

SALVADORACEAE<br />

Salvadora persica L.<br />

RHAMNACEAE / RUBIACEAE / RUTACEAE / SALVADORACEAE<br />

Vernacular names: toothbrush tree, miswak/suwak, rak/’arak<br />

Growth form: Shrub. Fl. Mar–Jul.<br />

status: Introduced. ?Common. Also recorded: E Saudi Arabia, UAE.<br />

كرع/كر ,كاوس/كاوسم<br />

60 61


SALVADORACEAE / SAPINDACEAE / SCROPHULARIACEAE<br />

habitat & distribution: Distribution in <strong>Qatar</strong> uncertain. Widely planted and naturalised in <strong>the</strong><br />

Gulf and native to parts <strong>of</strong> Saudi Arabia, Yemen and <strong>the</strong> far east <strong>of</strong> UAE (Jongbloed). One longestablished<br />

clump occurs by an old well within Ras Laffan Industrial City (JN).<br />

uses: The twigs are traditionally chewed to clean <strong>the</strong> teeth and are a familiar sight in Arabic markets.<br />

Jongbloed et al. list many medicinal uses <strong>of</strong> <strong>the</strong> fresh and dried leaves, including <strong>the</strong> treatment <strong>of</strong><br />

sore joints and scorpion stings.<br />

Photos: Coll 665; Jong 495; Pick 219; West 97.<br />

SAPINDACEAE<br />

Dodonaea viscosa Jacq.<br />

Dodonaea angustifolia L. f.<br />

Growth form: Shrub or small tree. Fl. Jan–May.<br />

status: Introduced. Common. Also recorded: E Saudi Arabia, UAE.<br />

habitat & distribution: Mainly planted by roadsides. Native to parts <strong>of</strong> Saudi Arabia and <strong>the</strong><br />

Hajar Mountains in UAE (Jongbloed).<br />

uses: As an ornamental and for screening; <strong>of</strong>ten clipped into hedges. Ghazanfar reports that <strong>the</strong><br />

leaves are used for treating toothache; many o<strong>the</strong>r medicinal uses are listed by Rizk & El-Ghazaly.<br />

notes: Possibly two or more subspecies occur, including subsp. viscosa Jacq. and subsp. angustifolia<br />

(L. f.) J.G. West.<br />

Photos: Jong 496; Pick 81; West 96.<br />

SCROPHULARIACEAE<br />

Misopates orontium (L.) Raf.<br />

<strong>An</strong>tirrhinum orontium L.<br />

Vernacular names: weasel’s snout, saysam<br />

Growth form: <strong>An</strong>nual herb. Fl. Mar–Apr.<br />

status: Introduced. Rare. Also recorded: UAE.<br />

habitat & distribution: Recorded from cultivated fields in C <strong>Qatar</strong>.<br />

Photos: Coll 672; Jong 507; West 132.<br />

Scrophularia deserti Delile Plate 80<br />

بيدلا بنع ,هريفز ,ةنيفع ,ةتيز<br />

Vernacular names: desert figwort, zeita, ’afeena, zafairah, ’inab al Deeb<br />

Growth form: Perennial herb. Fl. Mar–Apr.<br />

status: Native. Common. Also recorded: Bahrain, Kuwait, E Saudi Arabia, UAE.<br />

habitat & distribution: Occasional; generally on harder substrates in shallow sand or sandy<br />

runnels, especially in S and C <strong>Qatar</strong>.<br />

Photos: El Amin 19; Bat 92; Coll 683; Corn 191; Daoud 218, 219; Jong 511; Phil 88; Shuaib 70; West 134.<br />

Veronica cymbalaria Bodard<br />

Growth form: <strong>An</strong>nual herb. Fl. Apr.<br />

status: Introduced. Rare. Not in o<strong>the</strong>r Gulf floras.<br />

habitat & distribution: A weed <strong>of</strong> gardens in Doha.<br />

Photos: Coll 695.<br />

مسيس<br />

SOLANACEAE<br />

Datura innoxia Mill.<br />

Growth form: <strong>An</strong>nual herb. Fl. Mar–Sep.<br />

status: Introduced. Rare. First published record: Abdel Bari (1997). Also recorded: Kuwait.<br />

habitat & distribution: Recorded from Doha.<br />

Photos: Coll 697; Shuaib 84.<br />

Lycium shawii Roem. & Schult. Plate 84<br />

جسوع<br />

Vernacular names: desert thorn, ’awsaj<br />

Growth form: Shrub. Fl. Jan–Apr.<br />

status: Native. Very common. Also recorded: Bahrain, Kuwait, E Saudi Arabia, UAE.<br />

habitat & distribution: Frequent to locally dominant on hamada, in sandy and silty depressions,<br />

runnels, wadis and on rocky slopes. Stunted specimens have been recorded in saline habitats. Often<br />

grows with stands <strong>of</strong> Acacia and Ziziphus trees.<br />

uses: Grazed by animals and <strong>the</strong> berries are eaten by man; Cornes & Cornes note that <strong>the</strong> leaves<br />

provide browsing for camels and gazelles. Mandaville gives a detailed account relating to <strong>the</strong> old<br />

bedouin superstition that <strong>the</strong> plant is <strong>the</strong> abode <strong>of</strong> jinn (spirits), so some tribes do not use it for<br />

firewood. He states that it does not in any case provide good fuel, being ‘ra<strong>the</strong>r thorny and difficult<br />

to break’. Medicinally, <strong>the</strong> stems are used as a diuretic, laxative and tonic (Ghazanfar).<br />

Photos: El Amin 20; Bat 90, 91 L ; Coll 701; Corn 192; Daoud 210, 211; Jong 515; Mand 159; Phil 149; Pick<br />

190; Shuaib 83; West 129.<br />

Physalis angulata L.<br />

Growth form: <strong>An</strong>nual herb. Fl. May.<br />

status: Introduced. Rare. Not in o<strong>the</strong>r Gulf floras.<br />

habitat & distribution: Recorded by Batanouny from Doha as a weed amongst a cultivated plants.<br />

Photos: None.<br />

Solanum elaeagnifolium Cav.<br />

Growth form: Perennial herb. Fl. Mar–May.<br />

status: Introduced. Rare. Not in o<strong>the</strong>r Gulf floras.<br />

habitat & distribution: A weed <strong>of</strong> irrigated, cultivated areas in N <strong>Qatar</strong>.<br />

Photos: Bat 92.<br />

Solanum villosum (L.) Mill.<br />

Solanum nigrum L. var humile (Bernh.) Asch.<br />

SOLANACEAE<br />

Vernacular names: hairy nightshade, ’inab al-deeb<br />

بيدلا بنع<br />

Growth form: <strong>An</strong>nual herb. Fl. Mar–May.<br />

status: Introduced. ?Local. Also recorded: Kuwait, E Saudi Arabia, UAE.<br />

habitat & distribution: A weed <strong>of</strong> disturbed and enriched habitats, including cultivated areas,<br />

sewage ponds and gardens.<br />

uses: Batanouny noted that <strong>the</strong> fruit is edible. Mandaville states that although sometimes listed as<br />

a poisonous plant, he has seen <strong>the</strong> ripe berries being eaten. The plant is used as a diuretic and a<br />

long list <strong>of</strong> additional medicinal uses is given by Rizk & El-Ghazaly.<br />

Photos: Coll 705; Jong 519; Pick 192; Shuaib 84; West 130.<br />

62 63


TAMARICACEAE / TILIACEAE<br />

TAMARICACEAE<br />

Tamarix aphylla (L.) Karst.<br />

Vernacular names: tamarisk, ’athl, Tarfah<br />

Growth form: Tree. Fl. Oct–Feb.<br />

status: Introduced. Common. Also recorded: UAE.<br />

ةفرط ,لثع<br />

habitat & distribution: A frequently planted tree, <strong>of</strong>ten used as screening or shelterbelt around<br />

farms.<br />

uses: Mandaville notes its economic importance in Saudi Arabia, listing various uses including<br />

traditional house construction, ornamental planting, afforestation and dune stabilization. A<br />

number <strong>of</strong> medicinal uses is given by Rizk & El-Ghazaly and Ghazanfar.<br />

Photos: Ghaz 139; Jong 523; Pick 85.<br />

Tamarix aucheriana (Decne.) Baum<br />

Vernacular names: tamarisk, ’athl, Tarfah<br />

Growth form: Shrub. Fl. Apr–May, Oct–Nov.<br />

status: Native. Rare. Also recorded: Kuwait, E Saudi Arabia, UAE.<br />

habitat & distribution: Recorded by El Amin from Jebel Dukhan.<br />

uses: Used as firewood and <strong>the</strong> bark provides tannins; used for shelter belts (El Amin).<br />

Photos: Coll 711; Daoud 44, 45; Jong 524; Shuaib 82.<br />

Tamarix passerinoides Delile<br />

ةفرط ,لثع<br />

Vernacular names: tamarisk, ’athl, Tarfah<br />

ةفرط ,لثع<br />

Growth form: Shrub. Fl. Apr–Aug.<br />

status: Native. Rare. Not in o<strong>the</strong>r Gulf floras.<br />

habitat & distribution: Previously recorded from saline soils in SW <strong>Qatar</strong>, including Jebel Dukhan.<br />

uses: Used as firewood (El Amin).<br />

Photos: Coll 713.<br />

Tamarix ramosissima Ledeb.<br />

Vernacular names: tamarisk, ’athl, Tarfah<br />

Growth form: Shrub. Fl. Mar–Dec.<br />

status: Native. ?Local. Also recorded: E Saudi Arabia.<br />

habitat & distribution: Recorded by Batanouny from ‘saline ground, mainly in S. <strong>Qatar</strong>’.<br />

Photos: None.<br />

TILIACEAE<br />

ةفرط ,لثع<br />

Corchorus depressus (L.) Stocks Plate 85<br />

ربلا ةيخلم<br />

Vernacular names: mulakhiyah al bar<br />

Growth form: A prostrate shrublet. Fl. Mar–Apr.<br />

status: Native. Common. Also recorded: Bahrain, E Saudi Arabia, UAE.<br />

habitat & distribution: Occasional to frequent on silty or sandy depressions with compacted soil.<br />

uses: Rizk & El-Ghazaly report that <strong>the</strong> plant is used for removing tumours and pain.<br />

Photos: El Amin 20; Bat 69; Coll 715; Corn 118; Ghaz 107; Jong 526; Mand 63; Phil 127; Pick 142; West 98.<br />

Corchorus olitorius L.<br />

Vernacular names: mulakhiyah<br />

Growth form: <strong>An</strong>nual herb. Fl. Feb–May.<br />

status: Introduced. ?Local. Also recorded: E Saudi Arabia, UAE.<br />

habitat & distribution: Occasional as a weed <strong>of</strong> crops.<br />

uses: The leaves are used as a vegetable to make <strong>the</strong> Arabic dish mulakhiyah. A number <strong>of</strong> medicinal<br />

uses are listed by Rizk & El-Ghazaly.<br />

Photos: Coll 716; Ghaz 108.<br />

Corchorus trilocularis L.<br />

Vernacular names: wild jute, mulakhiyah al bar<br />

ربلا ةيخلم<br />

Growth form: <strong>An</strong>nual herb. Fl. Feb–May.<br />

status: Introduced. Rare. Also recorded: UAE.<br />

habitat & distribution: Reported as rare in N <strong>Qatar</strong> by Rizk & El-Ghazaly. El Amin listed C.<br />

trilocularis in <strong>the</strong> Arabic index to his book, but it was omitted from <strong>the</strong> species accounts.<br />

uses: Cultivated as a vegetable. A number <strong>of</strong> medicinal uses are listed by Rizk & El-Ghazaly.<br />

Photos: Jong 527; West 98.<br />

URTICACEAE<br />

Forsskaolea tenacissima L.<br />

Vernacular names: lazzaq, tubbaq<br />

Growth form: <strong>An</strong>nual herb. Fl. Mar–Apr.<br />

status: ?Native. Rare. Also recorded: UAE.<br />

64 65<br />

ةيخلم<br />

قابط ,قازل<br />

habitat & distribution: Recorded by El Amin from sandy soils and gravelly depressions, but he<br />

does not mention a location.<br />

Photos: Coll 738; Jong xii, 538; Pick 221; West 30.<br />

Forsskaolea viridis Ehrenb.<br />

Growth form: <strong>An</strong>nual herb. Fl. Jan–Apr.<br />

status: Native. ?Rare. First published record: Obeid (1975), but needs confirmation; not listed for<br />

<strong>Qatar</strong> by Miller & Cope. Also recorded: UAE.<br />

habitat & distribution: The typical habitat in Saudi Arabia is wadi bottoms and rocky slopes<br />

(Miller & Cope).<br />

Photos: Coll 739; Jong 539.<br />

Parietaria alsinifolia Delile<br />

Growth form: <strong>An</strong>nual herb. Fl. Mar–Apr.<br />

status: ?Native. Rare. Also recorded: UAE.<br />

habitat & distribution: Previously recorded from rodat, including Al Magdah.<br />

Photos: Coll 739; Jong 540.<br />

Urtica urens L.<br />

Growth form: <strong>An</strong>nual herb. Fl. Mar–Apr.<br />

status: Introduced. Local. Not in o<strong>the</strong>r Gulf floras.<br />

TILIACEAE / URTICACEAE


URTICACEAE / VERBENACEAE / ZyGOPHyLLACEAE<br />

habitat & distribution: A weed <strong>of</strong> irrigated gardens.<br />

uses: Many medicinal uses such as to sooth wounds and ulcers and to treat diarrhoea (Rizk & El-<br />

Ghazaly).<br />

Photos: Coll 740.<br />

VERBENACEAE<br />

Phyla nodiflora (L.) Greene<br />

Lippia nodiflora (L.) Michx.<br />

Vernacular names: frogfruit<br />

Growth form: Perennial herb. Fl. May–Jun.<br />

status: Introduced. Rare. Also recorded: Bahrain, Kuwait, E Saudi Arabia, UAE.<br />

habitat & distribution: A weed <strong>of</strong> wet places.<br />

uses: Sometimes cultivated as ground cover in parts <strong>of</strong> <strong>the</strong> Gulf. A wide variety <strong>of</strong> medical uses are<br />

known (Rizk & El-Ghazaly).<br />

Photos: Bat 88; Coll 747; Corn 203; Jong 541; Phil 177; Pick 86; Shuaib 74; West 125.<br />

Stachytarphaeta angustifolia (Mill.) Vahl<br />

Growth form: <strong>An</strong>nual or perennial herb. Fl. Mar–Apr.<br />

status: ?Introduced. ?Local. Not in o<strong>the</strong>r Gulf floras.<br />

habitat & distribution: Recorded by El Amin from rodat and runnels in C <strong>Qatar</strong>, but not recorded<br />

by any o<strong>the</strong>r authors from Arabia.<br />

uses: El Amin mentions that it is used medicinally in West Africa to treat dysentery, heart troubles,<br />

eye and ear sores.<br />

Photos: None.<br />

ZYGOPHYLLACEAE<br />

Fagonia bruguieri DC.<br />

Vernacular names: DHuraymah<br />

ةميرظ<br />

Growth form: Biennial or perennial herb. Fl. Mar–May.<br />

status: Native. Common. Also recorded: Bahrain, Kuwait, E Saudi Arabia, UAE.<br />

habitat & distribution: Occasional in sandy and gravelly habitats.<br />

uses: Rizk & El-Ghazaly describe a number <strong>of</strong> medicinal uses.<br />

notes: Batanouny describes three varieties occurring in <strong>Qatar</strong>: var. bruguieri, var. laxa Boiss. and<br />

var. rechingeri Hadidi.<br />

Photos: El Amin 20; Coll 757; Daoud 55, 56; Jong 545; Mand 138; Shuaib 81; West 87.<br />

Fagonia glutinosa Delile<br />

Vernacular names: DHuraymah<br />

Growth form: Biennial or perennial herb. Fl. Mar–Apr.<br />

status: Native. Local. Also recorded: Kuwait, E Saudi Arabia.<br />

habitat & distribution: Occasional in sandy areas; probably widespread.<br />

Photos: El Amin 20; Bat 57; Coll 757; Daoud 54; Shuaib 81.<br />

ةميرظ<br />

Fagonia indica Burm. f. Plate 81<br />

Vernacular names: DHuraymah, shwaikah, shka’e<br />

يعاكش ,ةكيوش ,ةميرظ<br />

Growth form: Perennial herb. Fl. Mar–Apr.<br />

status: Native. Common. Also recorded: Bahrain, E Saudi Arabia, UAE.<br />

habitat & distribution: Occasional on harder substrates and disturbed areas around inhabited<br />

areas.<br />

uses: Ghazanfar reports that <strong>the</strong> leaves, stems and roots are used for treating abdominal colic,<br />

dyspepsia, fever and venereal disease.<br />

Photos: El Amin 21; Bat 57; Coll 758; Jong 546; Phil 173; Pick 87; West 87.<br />

Fagonia ovalifolia Hadidi Plate 86<br />

Vernacular names: DHuraymah, shwaikah, shka’e<br />

يعاكش ,ةكيوش ,ةميرظ<br />

Growth form: Perennial herb. Fl. Mar–Apr.<br />

status: Native. Common. Also recorded: Bahrain, E Saudi Arabia, UAE.<br />

habitat & distribution: Occasional to frequent in s<strong>of</strong>ter sand.<br />

notes: Subspecies ovalifolia and pakistanica Ghafoor both occur in <strong>Qatar</strong>; var. qatarensis Hadidi has<br />

also been described (IPNI), presumably from <strong>Qatar</strong>.<br />

Photos: El Amin 21; Bat 58 L , 59; Coll 758; Corn 153; Jong 547; West 88.<br />

Fagonia tenuifolia Steud. & Hochst. ex Boiss.<br />

Growth form: Perennial herb. Fl. Apr.<br />

status: Native. ?Local. Not in o<strong>the</strong>r Gulf floras.<br />

habitat & distribution: Recorded by El Amin from Al Sheehaniyah and Al Karaanah in sandy<br />

depressions and runnels.<br />

Photos: El Amin 21; Coll 759.<br />

Seetzenia lanata (Willd.) Bullock Plate 87<br />

Seetzenia orientalis Decne.<br />

Vernacular names: Habyan/Habein<br />

نيبح/نايبح<br />

Growth form: Perennial herb. Fl. Mar–Apr.<br />

status: Native. Local. Also recorded: Bahrain, Kuwait, E Saudi Arabia, UAE.<br />

habitat & distribution: Occasional in sandy habitats in C and S <strong>Qatar</strong>.<br />

Photos: El Amin 22, 118 L ; Bat 60; Coll 760; Corn 153; Daoud 53 L ; Jong 548; Mand 139; Pick 222; West 88.<br />

Tetraena qatarensis (Hadidi) Beier & Thulin Plate 88<br />

Zygophyllum qatarense Hadidi<br />

Vernacular names: harm<br />

Growth form: Shrub. Fl. Mar–Apr.<br />

ZyGOPHyLLACEAE<br />

status: Native. Very common. Endemic to <strong>the</strong> Arabian Peninsula. Also recorded: Bahrain, Kuwait,<br />

E Saudi Arabia, UAE.<br />

habitat & distribution: Frequent to abundant, or dominant in hamada, gravel plains and saline<br />

sandy habitats, including sabkha edge and beaches.<br />

uses: Not edible to humans and usually avoided by wild animals and livestock. Mandaville remarks<br />

that it provides ‘useful summer grazing, but considered unhealthy in excess’.<br />

66 67<br />

مره


ZyGOPHyLLACEAE<br />

notes: All Zygophyllum species in Arabia have recently been transferred to <strong>the</strong> genus Tetraena<br />

(Beier et al. 2003). Mandaville notes that this is a ra<strong>the</strong>r variable species that was <strong>of</strong>ten attributed<br />

incorrectly to Zygophyllum coccineum L. It is possible that o<strong>the</strong>r Gulf and Saudi Arabian species, such<br />

as T. mandavillei may also occur inside <strong>the</strong> borders <strong>of</strong> <strong>Qatar</strong>.<br />

Photos: El Amin 22, 120 L ; Bat 61; Coll 764, 766; Corn 145, 154; Daoud 58, 59, 60; Jong 290, 555; Mand 142;<br />

Phil 172, 175; Pick 196; Shuaib 82; West 90.<br />

Tetraena simplex (L.) Beier & Thulin Plate 89<br />

Zygophyllum simplex L.<br />

Vernacular names: harm/hureim, da’a<br />

Growth form: <strong>An</strong>nual herb. Fl. Mar–Apr.<br />

status: Native. Very common. Also recorded: Bahrain, E Saudi Arabia, UAE.<br />

اعد ,ميره/مره<br />

habitat & distribution: Occasional or frequent in stony habitats and compacted, <strong>of</strong>ten saline soils.<br />

uses: In Bahrain, <strong>the</strong> leaves are used as an antibiotic and laxative; infusion <strong>of</strong> leaves or <strong>the</strong> seeds<br />

is applied to <strong>the</strong> eyes in ophthalmia (Rizk & El-Ghazaly).<br />

Photos: El Amin 22, 120 L ; Bat 62; Coll 766; Corn 157; Jong 556; Phil 176; Pick 145; West 91.<br />

Tribulus arabicus H. Hosni<br />

Vernacular names: shershir<br />

Growth form: Perennial herb. Fl. Feb–Sep.<br />

status: Native. Rare. <strong>An</strong> Arabian endemic. Also recorded: E Saudi Arabia, UAE.<br />

habitat & distribution: Shallow, fine textured soil and pebbly habitats (Batanouny).<br />

uses: Mandaville states that it is a main camel grazing species in parts <strong>of</strong> <strong>the</strong> Rub’ Al Khali.<br />

Photos: Coll 761; Jong 549; Mand 143, 144.<br />

Tribulus cistoides L.<br />

Tribulus terrestris var. cistoides (L.) C. Moore & Betche<br />

Vernacular names: shershir<br />

Growth form: Perennial herb. Fl. Not known.<br />

status: Native. Rare. First published record: Abdel Bari (1997). Not in o<strong>the</strong>r Gulf floras.<br />

habitat & distribution: Recorded by Abdel Bari from Al Sheehaniyah race track.<br />

Photos: None.<br />

Tribulus pterophorus C. Presl<br />

Vernacular names: shershir<br />

Growth form: Perennial herb. Fl. Mar–Apr.<br />

status: Native. Rare. Not in o<strong>the</strong>r Gulf floras.<br />

habitat & distribution: Recorded only from Al Mashbiya (<strong>the</strong> area NE <strong>of</strong> Abu Samrah) by El Amin<br />

from an area with stony soils.<br />

Photos: El Amin 21, 118 L .<br />

Tribulus terrestris L.<br />

Vernacular names: shershir<br />

Growth form: <strong>An</strong>nual or biennial herb. Fl. Apr–Dec.<br />

status: Native. Local. Also recorded: Kuwait, E Saudi Arabia, UAE.<br />

رشرش<br />

رشرش<br />

رشرش<br />

رشرش<br />

habitat & distribution: Occasional in gravelly and sandy habitats, including disturbed areas<br />

around camps, etc.; S and C <strong>Qatar</strong>.<br />

uses: A wide variety <strong>of</strong> medicinal uses are described by Rizk & El-Ghazaly.<br />

Photos: El Amin 21, 118 L ; Bat 60; Coll 763; Daoud 61, 62; Jong 553; Pick 146; Shuaib 82; West 89.<br />

ANGIOSPERMAE: MONOCOTyLEDONEAE<br />

ARECACEAE<br />

Phoenix dactylifera L. Plate 90<br />

Vernacular names: date palm, nakhl<br />

لخن<br />

Growth form: Tree. Fl. Apr–May.<br />

status: Introduced. Very common. Also recorded: Bahrain, E Saudi Arabia, UAE.<br />

habitat & distribution: Planted all over <strong>Qatar</strong>. Naturalised on <strong>the</strong> coast at Umm Bab.<br />

uses: Cultivated in plantations and gardens, especially for <strong>the</strong> edible fruit (date); also along<br />

roadsides to provide greenery and shade. Various authors describe cultural uses, including making<br />

ropes, baskets, ro<strong>of</strong>ing and used in <strong>the</strong> construction <strong>of</strong> boats and houses. It can also be used as<br />

firewood. Jongbloed et al. note that it has medicinal uses.<br />

Photos: El Amin 27; Coll 593; Corn 253, 254; Jong 13; Phil 19; West 167.<br />

CYMODOCEACEAE<br />

Diplan<strong>the</strong>ra uninervis (Forssk.) F.N.Williams<br />

Halodule uninervis (Forssk.) Asch.<br />

Vernacular names: seagrass<br />

Growth form: Marine perennial herb. Fl. Not known.<br />

status: Native. Common. First published record: Abdel Bari (1997). Not in o<strong>the</strong>r Gulf floras,<br />

though marine flowering plants appear to have been overlooked by some authors.<br />

habitat & distribution: Intertidal areas around <strong>the</strong> coast; widespread (BB).<br />

uses: Diplan<strong>the</strong>ra spp. and Halophila spp. (p. 71) are an important component <strong>of</strong> <strong>the</strong> marine food<br />

chain, but fur<strong>the</strong>r study is needed on <strong>the</strong>m in <strong>Qatar</strong>.<br />

notes: Diplan<strong>the</strong>ra wrightii (Asch.) Asch has been reported from Bahrain by Cornes & Cornes and<br />

may also be found in <strong>Qatar</strong> (see Mandaville 1990, p. 331).<br />

Photos: None.<br />

CYPERACEAE<br />

ZyGOPHyLLACEAE / ARECACEAE / CyMODOCEACEAE / CyPERACEAE<br />

Bolboschoenus maritimus (L.) Palla<br />

Scirpus maritimus L.<br />

Growth form: Sedge. Fl. Mar–Jun.<br />

status: Native. Rare. Also recorded: E Saudi Arabia, UAE.<br />

habitat & distribution: Batanouny only recorded it in a permanently wet area at Ras Ushirij, at<br />

<strong>the</strong> edge <strong>of</strong> a patch <strong>of</strong> Typha domingensis.<br />

Photos: Coll 277; Jong 15; West 169.<br />

68 69


CyPERACEAE<br />

Cyperus conglomeratus Rottb. Plate 91<br />

Cyperus aucheri Jaub. & Spach<br />

Vernacular names: thanda, rasha, ’andeb, sa’ed, qassis<br />

سيسق ,دعس ,بدنع ,اشر ,ةدنث<br />

Growth form: Sedge. Fl. May–Jun.<br />

status: Native. Common. Also recorded: Bahrain, Kuwait, E Saudi Arabia, UAE.<br />

habitat & distribution: A plant characteristic <strong>of</strong> deeper sand, especially larger dunes, but in <strong>Qatar</strong><br />

also locally abundant around coastal areas on beach sand and on white sand inland. Occasional to<br />

locally frequent in o<strong>the</strong>r habitats, such as sandy wadis and runnels.<br />

uses: Grazed, but considered inferior as fodder. Jongbloed et al. note that <strong>the</strong> rhizomes are<br />

sometimes used as food by humans in times <strong>of</strong> hunger.<br />

notes: This is a very variable group comprising a number <strong>of</strong> different described varieties and<br />

forms, but fur<strong>the</strong>r study <strong>of</strong> plants in <strong>Qatar</strong> and o<strong>the</strong>r parts <strong>of</strong> <strong>the</strong> Gulf is still needed. For fur<strong>the</strong>r<br />

information see Batanouny and Mandaville.<br />

Photos: El Amin 23; Bat 124; Coll 279; Corn 233; Jong 18; Phil 167; Pick 261; Shuaib 51; West 168.<br />

Cyperus jeminicus Rottb<br />

Cyperus conglomeratus subsp. jeminicus (Rottb.) Lye<br />

Vernacular names: rukbah, sa’ed<br />

Growth form: Sedge. Fl. Apr–May.<br />

status: Native. Local. Also recorded E Saudi Arabia.<br />

دعس ,ةبكر<br />

habitat & distribution: Locally frequent in coastal saline sand, including Ras Laffan, Fuwairit<br />

and o<strong>the</strong>r areas in NE <strong>Qatar</strong> (MS).<br />

Notes: Not previously documented for <strong>Qatar</strong>.<br />

Photos: None.<br />

Cyperus laevigatus L.<br />

Juncellus laevigatus (L.) C.B. Baker<br />

Vernacular names: sa’ed<br />

دعس<br />

Growth form: Sedge. Fl. Mar–Apr.<br />

status: Native. Rare. Also recorded: Bahrain, E Saudi Arabia, UAE.<br />

habitat & distribution: A species <strong>of</strong> marshy places, <strong>of</strong>ten in slightly saline habitats. Recorded by<br />

Batanouny at Abu Samrah in SW <strong>Qatar</strong>.<br />

uses: Fodder for camels (El Amin).<br />

Photos: Bat 124; Coll 280; Corn 231; Jong 19.<br />

Cyperus rotundus L.<br />

Vernacular names: sa’ed<br />

دعس<br />

Growth form: Sedge. Fl. Jan–Dec.<br />

status: Native. Common. Also recorded: Bahrain, E Saudi Arabia, UAE.<br />

habitat & distribution: Frequent in marshy areas, such as gardens and cultivated fields. Rarely<br />

seen in desert areas.<br />

Photos: Coll 282; Corn 235; Jong 20; West 168.<br />

HYDROCHARITACEAE<br />

Halophila ovalis (R. Br.) Hook. f.<br />

Growth form: Marine perennial herb. Fl. Not known.<br />

status: Native. Common. Not previously documented for <strong>Qatar</strong>. Also recorded: Bahrain, E Saudi<br />

Arabia.<br />

habitat & distribution: Intertidal areas around <strong>the</strong> coast; widespread (BB).<br />

Notes: See comments under Diplan<strong>the</strong>ra spp.<br />

Photos: None.<br />

Halophila stipulacea (Forssk.) Asch.<br />

Growth form: Marine perennial herb. Fl. Apr.<br />

status: Native. Common. Also recorded: Bahrain, E Saudi Arabia.<br />

habitat & distribution: Intertidal areas around <strong>the</strong> coast; widespread (BB). Collected from Al<br />

Khor shore by El Amin.<br />

Photos: El Amin 143 L ; Bat 109; Corn 226.<br />

JUNCACEAE<br />

Juncus rigidus Desf.<br />

Juncus arabicus (Asch. & Buchenau) Adamson; Juncus maritimus Lam. var. arabicus Asch. & Buchenau<br />

ةدنت ,اشر ,لصأ<br />

Vernacular names: asal, rasha, tanda<br />

Growth form: Rush. Fl. Nov-Dec.<br />

status: Native. Local. Also recorded: Bahrain, E Saudi Arabia, UAE.<br />

habitat & distribution: Frequent in muddy saltmarshes and o<strong>the</strong>r damp, saline areas. Occurs at<br />

Abu Samrah, Al Dhakhira, Jebel Fuwairit (MS) and also previously recorded from Wadi El Ireig.<br />

uses: Eaten by camels. Used to make mats.<br />

Photos: El Amin 26; Coll 444; Corn 231; Jong 27; Phil 162; Pick 262.<br />

LILIACEAE<br />

HyDROCHARITACEAE / JUNCACEAE / LILIACEAE<br />

Asphodelus tenuifolius Cav.<br />

Asphodelus fistulosus L. var tenuifolius (Cav.) Baker<br />

Vernacular names: barwaq<br />

قورب<br />

Growth form: <strong>An</strong>nual herb. Fl. Feb–Apr.<br />

status: Native. Common. Also recorded: Bahrain, Kuwait, E Saudi Arabia, UAE.<br />

habitat & distribution: Occasional to locally frequent in shallow, <strong>of</strong>ten somewhat compact, sand;<br />

particularly in sandy or gravelly depressions.<br />

uses: Phillips quotes Dickson (1955) reporting that <strong>the</strong> seeds <strong>of</strong> <strong>the</strong> plant were used by <strong>the</strong> Bedu<br />

<strong>of</strong> Kuwait to make a kind <strong>of</strong> dried milk. ‘Ga<strong>the</strong>red for use as a po<strong>the</strong>rb’ (Cornes & Cornes). The<br />

seeds have diuretic properties and are applied externally to ulcers and inflamed parts (Rizk & El-<br />

Ghazaly).<br />

notes: A. fistulosus L. is best treated as a separate species; it has been reported from Egypt and<br />

nor<strong>the</strong>rn Saudi Arabia (Boulos).<br />

Photos: El Amin 26; Coll 78; Corn 228; Jong 31; Phil 124, 125; Pick 154; Shuaib 50; West 154.<br />

70 71


LILIACEAE / POACEAE<br />

Dipcadi erythraeum Webb & Ber<strong>the</strong>l. Plates 93, 94<br />

Dipcadi serotinum (L.) Medik.; Dipcadi unicolor (Stocks) Baker<br />

Vernacular names: miSelmow, busalamo, anSel, bessail al maa,<br />

ءاملا ليسب ,لصنأ ,وملسوب ,ومليصم<br />

brown lily<br />

Growth form: Perennial herb. Fl. Feb–Mar.<br />

status: Native. Common. Also recorded: Bahrain, Kuwait, E Saudi Arabia, UAE.<br />

habitat & distribution: Occasional to frequent in S and C <strong>Qatar</strong>, in sandy depressions and runnels.<br />

uses: The bulb is reputed to be edible, although Mandaville says it is bitter, whilst Jongbloed et al.<br />

state that <strong>the</strong> capsule is edible. Rizk & El-Ghazaly report that in Bahrain <strong>the</strong> leaves are used as a<br />

laxative and as an ointment for wounds.<br />

Photos: El Amin 27; Bat 109; Coll 431; Corn 35, 227; Jong 33; Mand 266; Phil 126; Shuaib 49; West 154.<br />

POACEAE<br />

Aeluropus lagopoides (L.) Trin. ex Thwaites Plate 99<br />

Vernacular names: ’ikrish<br />

شركع<br />

Growth form: Perennial grass. Fl. Mar–May.<br />

status: Native. Very common. Also recorded: Bahrain, E Saudi Arabia, UAE.<br />

habitat & distribution: Frequent to abundant in saline habitats, including sabkha edge, saltmarsh<br />

and <strong>the</strong> edges <strong>of</strong> permanent wetland areas. Sometimes dominates large areas <strong>of</strong> saline flat.<br />

uses: El Amin states this species to be good fodder.<br />

Photos: El Amin 126 L ; Bat 110; Coll 342 L ; Corn 242; Jong 37; Phil 107; West 156.<br />

Aristida abnormis Chiov.<br />

Growth form: <strong>An</strong>nual grass. Fl. Apr.<br />

status: Native. Local. Also recorded: UAE.<br />

habitat & distribution: Probably occasional throughout <strong>Qatar</strong> on shallow soils.<br />

uses: Used for fodder (El Amin).<br />

Photos: Coll 344 L ; Jong 38.<br />

Aristida adscensionis L.<br />

Vernacular names: samaa, SafSaf<br />

فصفص ,امس<br />

Growth form: <strong>An</strong>nual or perennial grass. Fl. Apr.<br />

status: Native. Rare. First published record: Cope (2007). Also recorded: E Saudi Arabia, UAE.<br />

habitat & distribution: Details <strong>of</strong> <strong>the</strong> Cope record(s) not known, but a specimen <strong>of</strong> this species<br />

was collected by JN from a sandy depression in SW <strong>Qatar</strong> in 2007.<br />

Photos: Coll 345 L ; Jong 39.<br />

Aristida mutabilis Trin. & Rupr.<br />

Aristida meccana Hochst. ex Trin. & Rupr.<br />

Growth form: <strong>An</strong>nual grass. Fl. Apr.<br />

status: Native. Rare. Also recorded: E Saudi Arabia, UAE.<br />

habitat & distribution: Previously recorded from stony and rocky ground in N <strong>Qatar</strong>.<br />

Photos: None.<br />

Arundo donax L.<br />

Vernacular names: giant reed, ghab, qalam<br />

Growth form: Perennial grass. Fl. Apr.<br />

status: Native. Rare. Also recorded: E Saudi Arabia, UAE.<br />

habitat & distribution: Recorded by Batanouny from Al Wabrah.<br />

uses: Used for fodder (El Amin).<br />

Photos: Coll 348 L ; Jong 40.<br />

Avena sterilis L.<br />

Vernacular names: shufan barri<br />

Growth form: <strong>An</strong>nual grass. Fl. Apr.<br />

status: Native. Common. Also recorded: E Saudi Arabia.<br />

habitat & distribution: Frequent in rodat and o<strong>the</strong>r irrigated places.<br />

uses: Used for fodder (El Amin).<br />

72 73<br />

ملق ,باغ<br />

يرب نافوش<br />

notes: Cope lists subspp. sterilis and ludoviciana (Durieu) M. Gillet & Magne for <strong>Qatar</strong>; <strong>the</strong> latter<br />

is <strong>of</strong>ten treated as a separate species.<br />

Photos: El Amin 23; Coll 350 L .<br />

Brachypodium distachyum (L.) P. Beauv.<br />

Trachynia distachya (L.) Link<br />

Growth form: <strong>An</strong>nual grass. Fl. Feb–May.<br />

status: Native. ?Local. Also recorded: Bahrain, UAE.<br />

habitat & distribution: Recorded by Batanouny as occasional in deep alluvial soil in rodat.<br />

Photos: Coll 354 L ; Jong 44.<br />

Bromus danthoniae Trin.<br />

Growth form: <strong>An</strong>nual grass. Fl. Apr.<br />

status: Native. Rare. Also recorded: E Saudi Arabia.<br />

habitat & distribution: Appears only to have been recorded by Batanouny (and possibly also by<br />

El Amin) from a single locality as Al Sadd, S <strong>of</strong> Doha.<br />

Photos: Coll 355 L .<br />

Bromus madritensis L.<br />

<strong>An</strong>isantha madritensis (L.) Nevski; Bromus madritensis subsp. haussknechtii (Boiss.) H. Scholz; Bromus<br />

haussknechtii Boiss.<br />

Vernacular names: brome grass, zeil abu al-Hossein<br />

Growth form: <strong>An</strong>nual grass. Fl. Feb–May.<br />

status: Native. Rare. Also recorded: E Saudi Arabia, UAE.<br />

نيسحلا وبأ ليذ<br />

habitat & distribution: Recorded by Batanouny from a stony area at Umm Slal Mohammed.<br />

Photos: Bat 110; Coll 356 L ; Jong 45.<br />

Bromus scoparius L.<br />

Growth form: <strong>An</strong>nual grass. Fl. Apr–May.<br />

status: Native. Rare. Also recorded: E Saudi Arabia.<br />

POACEAE


POACEAE<br />

habitat & distribution: Recorded by Batanouny from a stony area with accumulated fine soil at<br />

Umm Slal Mohammed.<br />

notes: Batanouny ascribed his plant to var. psilostachys Halácsy (not formally recognised by Cope).<br />

Photos: Coll 357 L .<br />

Cenchrus ciliaris L. Plate 92<br />

Vernacular names: foxtail grass, sabaT<br />

Growth form: Perennial grass. Fl. Apr.<br />

status: Native. Very common. Also recorded: Bahrain, E Saudi Arabia, UAE.<br />

habitat & distribution: Occasional to abundant or locally dominant in moderately compact sand<br />

or silt, especially in rocky areas and <strong>of</strong>ten around inhabited areas and farmland.<br />

uses: Described by Mandaville as ‘useful fodder grass’.<br />

Photos: Coll 360 L ; Jong 46; Mand 237; Phil 108; Pick 251; West 157.<br />

Cenchrus pennisetiformis Hoschst. & Steud.<br />

Growth form: <strong>An</strong>nual grass. Fl. Apr.<br />

status: Native. Rare. Also recorded: UAE.<br />

habitat & distribution: Recorded from cultivated rodat in C <strong>Qatar</strong> by Batanouny, but presumably<br />

also occurs in natural areas with sandy and silty soils.<br />

Photos: Bat 111; Coll 361 L ; Jong 47.<br />

Cenchrus setigerus Vahl<br />

Growth form: Perennial grass. Fl. Not known.<br />

status: Native. ?Rare. Also recorded: UAE.<br />

habitat & distribution: The habitat <strong>of</strong> this species is similar to o<strong>the</strong>r species <strong>of</strong> Cenchrus.<br />

notes: Tentatively listed for <strong>Qatar</strong> by Cope on <strong>the</strong> basis <strong>of</strong> <strong>the</strong> photograph included in Batanouny<br />

(Plate 111), mis-labelled as C. pennisetiformis.<br />

Photos: Bat 111; Jong 49.<br />

Centropodia forsskaolii (Vahl) Cope Plate 100<br />

As<strong>the</strong>na<strong>the</strong>rum forsskaolii (Vahl) Nevski<br />

Vernacular names: qasba’a, hajeen<br />

Growth form: Perennial grass. Fl. Apr–May.<br />

status: Native. Local. Also recorded: E Saudi Arabia, UAE.<br />

habitat & distribution: Frequent to abundant in sandy runnels in SW <strong>Qatar</strong>.<br />

uses: Grazed by camels (Mandaville) and used for fodder (El Amin).<br />

Photos: El Amin 126 L ; Coll 361 L ; Jong 50; West 156.<br />

Chloris barbata Sw.<br />

Vernacular names: finger grass, khazamzam<br />

Growth form: Perennial grass. Fl. Apr.<br />

status: Introduced. Common. Also recorded: Bahrain, UAE.<br />

habitat & distribution: Frequent in cultivated farms, gardens and o<strong>the</strong>r irrigated land.<br />

uses: Good fodder (El Amin).<br />

Photos: El Amin 23; Coll 362 L ; Phil 109; Pick 252.<br />

طبس<br />

نيجه ،ءابصق<br />

مزمزخ<br />

Chloris virgata Sw.<br />

Vernacular names: finger grass, khazamzam, sineim<br />

Growth form: <strong>An</strong>nual grass. Fl. Feb–Jul.<br />

status: Introduced. Common. Also recorded: E Saudi Arabia, UAE.<br />

74 75<br />

مينس ,مزمزخ<br />

habitat & distribution: Frequent in cultivated land and o<strong>the</strong>r irrigated areas, including gardens.<br />

Photos: Bat 112; Coll 363 L ; Jong 52; West 157.<br />

Chrysopogon gryllus (L.) Trin.<br />

Vernacular names: halta<br />

Growth form: Perennial grass. Fl. Apr–Jun.<br />

status: Native. Local. Not in o<strong>the</strong>r Gulf floras.<br />

habitat & distribution: Occasional to frequent in wadis and runnels, <strong>of</strong>ten within rocky areas.<br />

Photos: None.<br />

Chrysopogon plumulosus Hochst. Plates 95, 101<br />

Vernacular names: halta, gharaz<br />

Growth form: Perennial grass. Fl. Apr.<br />

status: Native. Common. Also recorded: E Saudi Arabia.<br />

اتله<br />

زارغ ,اتله<br />

habitat & distribution: Occasional to locally frequent in shallow sand in wadis and runnels; <strong>of</strong>ten<br />

within rocky areas.<br />

uses: Eaten by all animals (El Amin).<br />

notes: Recorded as C. aucheri (Boiss.) Stapf by Batanouny, but later apparently redetermined as C.<br />

plumulosus (see El Amin, p.128). C. aucheri could also occur in <strong>Qatar</strong>.<br />

Photos: Bat 123; Coll 363 L .<br />

Cutandia memphitica (Spreng.) Benth. Plate 96<br />

Vernacular names: Memphis grass<br />

Growth form: <strong>An</strong>nual grass. Fl. Apr.<br />

status: Native. Local. Also recorded: Bahrain, E Saudi Arabia, UAE.<br />

habitat & distribution: Occasional in sandy areas <strong>of</strong> S and W <strong>Qatar</strong>.<br />

uses: Used for fodder (El Amin).<br />

Photos: Coll 165 L ; Jong 54; Mand 251; Phil 110; West 158.<br />

Cymbopogon commutatus (Steud.) Stapf<br />

Cymbopogon parkeri Stapf<br />

Vernacular names: incense grass, Sakhbar<br />

Growth form: Perennial grass. Fl. Apr.<br />

status: Native. Common. Also recorded: Bahrain, E Saudi Arabia, UAE.<br />

POACEAE<br />

ربخص<br />

habitat & distribution: Occasional to locally abundant in silty or sandy depressions and runnels;<br />

mainly N and C <strong>Qatar</strong>. Large tussocks <strong>of</strong> this lemon-scented species dominate silt pans in parts <strong>of</strong><br />

Ras Laffan Industrial City where <strong>the</strong>re is no grazing.<br />

uses: Grazed, particularly by sheep. The leaves have a strong lemon scent when crushed. Mandaville<br />

mentions that <strong>the</strong>re is a history <strong>of</strong> medicinal use for this species in Arabia. Jongbloed et al. state that<br />

plants <strong>of</strong> <strong>the</strong> genus are cultivated for citronella oil for use as perfume and insect repellent.<br />

Photos: El Amin 126 L ; Bat 112, 113 L ; Coll 365 L ; Corn 238; Jong 55; Mand 235; West 158.


POACEAE<br />

Cynodon dactylon (L.) Pers.<br />

Vernacular names: Bermuda grass, star grass, dog’s-tooth grass, <strong>the</strong>el/thayyil,<br />

najeel, najm<br />

Growth form: Perennial grass. Fl. Most <strong>of</strong> year.<br />

status: ?Native. Very common. Also recorded: Bahrain, E Saudi Arabia, UAE.<br />

مجن ,ليجن ,ليث<br />

habitat & distribution: The main constituent <strong>of</strong> lawns in urban areas; occasional in natural<br />

habitats.<br />

uses: Of economic importance due to its widespread use in making lawns. Medicinal uses worldwide<br />

are described by Rizk & El-Ghazaly.<br />

Photos: El Amin 23; Coll 366 L ; Corn 246; Jong 57; Phil 112; West 159.<br />

Dactyloctenium aegyptium (L.) Willd.<br />

Vernacular names: najm<br />

Growth form: <strong>An</strong>nual grass. Fl. Apr–May.<br />

status: Native. Common. Also recorded: Bahrain, E Saudi Arabia, UAE.<br />

habitat & distribution: Occasional to frequent in rodat, irrigated lawns, date groves and o<strong>the</strong>r<br />

tree plantations.<br />

uses: El Amin notes that it ‘is used as fodder, but not <strong>of</strong> high value’. Various medicinal uses are<br />

reported.<br />

Photos: El Amin 24; Bat 114; Coll 367 L ; Corn 247; Jong 58; Mand 248; Phil 113; Pick 254; West 159.<br />

Dichanthium annulatum (Forssk.) Stapf<br />

Growth form: Perennial grass. Fl. Apr.<br />

status: Native. Local. Also recorded: Bahrain, E Saudi Arabia, UAE.<br />

habitat & distribution: Previously recorded as common in irrigation channels and o<strong>the</strong>r damp<br />

areas in cultivated rodat, but seen only once by <strong>the</strong> present authors.<br />

uses: Used for fodder (El Amin).<br />

Photos: Bat 115; Coll 369 L ; Corn 238; Jong 61.<br />

Dichanthium foveolatum (Delile) Roberty Plate 97<br />

Eremopogon foveolatus (Delile) Stapf<br />

Vernacular names: halta<br />

Growth form: Perennial grass. Fl. Apr.<br />

status: Native. Common. Also recorded: Bahrain, E Saudi Arabia, UAE.<br />

habitat & distribution: Occasional in compacted sandy and silty areas; <strong>of</strong>ten within rocky habitats.<br />

uses: Used for fodder (El Amin).<br />

Photos: El Amin 132 L ; Bat 111; Coll 369 L ; Corn 140; Jong 62.<br />

Digitaria sanguinalis (L.) Scop.<br />

Vernacular names: finger grass, crab grass<br />

Growth form: <strong>An</strong>nual grass. Fl. May.<br />

status: Introduced. Local. Also recorded: E Saudi Arabia, UAE.<br />

habitat & distribution: Occasional in damp areas, including <strong>the</strong> edges <strong>of</strong> cultivated fields.<br />

uses: Grazed, and used for fodder.<br />

Photos: Coll 371 L ; Jong 63.<br />

مجن<br />

اتله<br />

Echinochloa colona (L.) Link<br />

Vernacular names: jungle rice, millet rice, corn panic grass, moDeiD, Difra<br />

Growth form: <strong>An</strong>nual grass. Fl. Apr.<br />

status: Introduced. Common. Also recorded: Bahrain, E Saudi Arabia, UAE.<br />

habitat & distribution: Occasional in irrigated lawns and gardens.<br />

uses: Good fodder (El Amin).<br />

Photos: El Amin 132 L ; Bat 115; Coll 372 L ; Corn 250; Jong 64; West 160.<br />

Eragrostis barrelieri Daveau<br />

Vernacular names: love-grass<br />

Growth form: <strong>An</strong>nual grass. Fl. Apr.<br />

status: Native. Local. Also recorded: E Saudi Arabia, UAE.<br />

habitat & distribution: Occasional in sandy soils.<br />

Photos: Bat 117; Coll 378 L ; Jong 66; Mand 250.<br />

Eragrostis cilianensis (All.) Vignolo ex Janch.<br />

Vernacular names: stink-grass, love-grass, tearab<br />

Growth form: <strong>An</strong>nual grass. Fl. Apr.<br />

status: Native. Common. Also recorded: UAE.<br />

ةرفض ,ضيضم<br />

habitat & distribution: Occasional in sandy runnels, wadis and o<strong>the</strong>r sandy areas; also as a weed<br />

in irrigated gardens and rodat.<br />

uses: Probably a useful grazing species.<br />

Photos: Bat 117; Coll 378 L ; Jong 67.<br />

Eragrostis ciliaris (L.) R. Br.<br />

Growth form: <strong>An</strong>nual grass. Fl. Feb–Mar.<br />

status: Native. ?Local. Also recorded: UAE.<br />

habitat & distribution: Occasional in fine soils in depressions, <strong>of</strong>ten under trees.<br />

Photos: Bat 117; Coll 379 L .<br />

Eragrostis pilosa (L.) P. Beauv.<br />

Growth form: <strong>An</strong>nual grass. Fl. Apr–May.<br />

status: Native. Rare. Also recorded: UAE.<br />

habitat & distribution: Recorded by Batanouny from a depression at Al Magdah, growing under<br />

<strong>the</strong> shade <strong>of</strong> Ziziphus and Lycium bushes.<br />

Photos: Coll 381 L .<br />

Halopyrum mucronatum (L.) Stapf<br />

Growth form: Perennial grass. Fl. Apr.<br />

status: Native. Rare. Also recorded: E Saudi Arabia, UAE.<br />

habitat & distribution: Dominates an area <strong>of</strong> beach sand at Abu Samrah.<br />

uses: Used for fodder (El Amin). Much grazed by livestock, including horses along <strong>the</strong> coast<br />

(Mandaville).<br />

Photos: Coll 385 L ; Jong 69; West 161.<br />

76 77<br />

POACEAE<br />

باريت


POACEAE<br />

Henrardia pubescens (Bertol.) C.E. Hubb.<br />

Growth form: <strong>An</strong>nual grass. Fl. Apr.<br />

status: Native. Local. Not in o<strong>the</strong>r Gulf floras.<br />

habitat & distribution: Occasional in rodat in C <strong>Qatar</strong>; also recorded at Doha.<br />

notes: Cope notes that much <strong>of</strong> <strong>the</strong> material originally named as this species was in fact Parapholis<br />

incurva (L.) C.E. Hubb. However, he thinks that Henrardia pubescens is also likely to be correctly<br />

reported for <strong>Qatar</strong> based on published descriptions.<br />

Photos: None.<br />

Hordeum murinum L.<br />

Hordeum glaucum Steud.<br />

Growth form: <strong>An</strong>nual grass. Fl. Mar–Apr.<br />

status: Native. Rare. Also recorded: Bahrain, E Saudi Arabia.<br />

habitat & distribution: Batanouny recorded this taxon from Umm Slal Mohammed growing in<br />

small depressions on calcareous ridges. It may also occur as a weed species in cultivated areas, as<br />

it does elsewhere in <strong>the</strong> Gulf.<br />

uses: Mandaville remarks that it is grazed by livestock when young, but is less palatable when<br />

maturing due to <strong>the</strong> hard, sharp awns.<br />

notes: The form in <strong>the</strong> Gulf is currently recognised as Hordeum murinum subsp. glaucum (Steud.)<br />

Tzvelev.<br />

Photos: Bat 120; Coll 386 L ; Phil 114.<br />

Hordeum vulgare L.<br />

Vernacular name: barley, sha’ir<br />

Growth form: <strong>An</strong>nual grass. Fl. Apr.<br />

status: Introduced. Local. Also recorded: Bahrain, E Saudi Arabia.<br />

habitat & distribution: Occasional in cultivated fields.<br />

uses: <strong>An</strong> important crop and fodder plant. Rizk & El-Ghazaly describe <strong>the</strong> medicinal properties<br />

<strong>of</strong> this species.<br />

Photos: None.<br />

Hyparrhenia hirta (L.) Stapf<br />

Vernacular names: blue-stem grass<br />

Growth form: Perennial grass. Fl. Apr–May.<br />

status: Native. ?Local. Also recorded: Bahrain, E Saudi Arabia, UAE.<br />

habitat & distribution: Recorded by El Amin from C and W <strong>Qatar</strong>, including Dukhan.<br />

uses: Good fodder (El Amin).<br />

Photos: Coll 387 L ; Corn 140; Jong 70; Mand 236; Phil 115.<br />

Lasiurus scindicus Henrard<br />

Lasiurus hirsutus (Vahl) Boiss., comb. illeg.<br />

Plates 98, 102<br />

Vernacular names: daHaH<br />

حاحد<br />

Growth form: Perennial grass. Fl. Apr–May.<br />

status: Native. Very common. Also recorded: E Saudi Arabia, UAE.<br />

ريعش<br />

habitat & distribution: Frequent to abundant in shallow sandy areas, including runnels; mainly<br />

in C and S <strong>Qatar</strong>.<br />

Photos: Bat 119; Coll 388 L ; Jong 73; Pick 255.<br />

Leptochloa fusca (L.) Kunth<br />

Diplachne fusca (L.) P. Beauv. ex Roem. & Schult.<br />

Growth form: Perennial grass. Fl. May–Nov.<br />

status: ?Introduced. Rare. Also recorded: E Saudi Arabia, UAE.<br />

habitat & distribution: Recorded by Batanouny in a regularly irrigated lawn <strong>of</strong> Ramada Hotel,<br />

Doha. The species may also potentially occur in o<strong>the</strong>r wet habitats.<br />

Photos: None.<br />

Lolium rigidum Gaudin<br />

Vernacular names: rye grass, simbil, rabi’ya<br />

Growth form: <strong>An</strong>nual grass. Fl. Apr.<br />

status: ?Native. Local. Also recorded: E Saudi Arabia, UAE.<br />

78 79<br />

ةيعيبر ,لبمس<br />

habitat & distribution: Occasional in Doha gardens and lawns, and in damp cultivated fields.<br />

May be native in some areas, but widely naturalised.<br />

uses: Used for fodder (El Amin). Toxicity to sheep and cattle has been reported; ingestion <strong>of</strong> seedheads<br />

can cause neurological disorders and especially affects <strong>the</strong> bovine liver (Rizk & El-Ghazaly).<br />

Photos: El Amin 28, 132 L ; Coll 390 L ; Jong 74 L .<br />

Ochthochloa compressa (Forssk.) Hilu Plate 103<br />

Eleusine compressa (Forssk.) Asch. & Schweinf. ex C. Chr.<br />

ةرمح ,مينص<br />

Vernacular names: wire grass, Sineim, Hamrah<br />

Growth form: Perennial grass. Fl. Apr.<br />

status: Native. Common. Also recorded: Bahrain, E Saudi Arabia, UAE.<br />

habitat & distribution: Occasional to locally abundant in sandy depressions and runnels; mainly<br />

SW <strong>Qatar</strong>, including Al Karaanah.<br />

uses: Good fodder (El Amin).<br />

Photos: El Amin 132 L ; Bat 116; Coll 392 L ; Corn 241; Jong 76; West 160.<br />

Panicum turgidum Forssk. Plate 104<br />

مامثإ/مامث<br />

Vernacular names: thamam/ithmam<br />

Growth form: Perennial grass. Fl. Apr.<br />

status: Native. Very common. Also recorded: Bahrain, E Saudi Arabia, UAE.<br />

POACEAE<br />

habitat & distribution: Abundant on sand sheets, sandy depressions and o<strong>the</strong>r sandy habitats.<br />

Especially common in S and SW <strong>Qatar</strong>, where it is a major component <strong>of</strong> <strong>the</strong> natural perennial<br />

vegetation. It probably avoids saline sand.<br />

uses: Described as good fodder by El Amin. Mandaville said it was reported that bedouin used this<br />

as food during famine. Jongbloed et al. note that it was used as a wound dressing. The species is a<br />

natural sand binder and could have potential for stabilising sand dunes.<br />

Photos: El Amin 24, 136 L ; Bat 118; Coll 396 L ; Corn 240; Jong 79; Mand 239; Phil 106, 116; West 161.


POACEAE<br />

Parapholis incurva (L.) C.E. Hubb.<br />

Vernacular names: curved hard-grass<br />

Growth form: Grass. Fl. Not known.<br />

status: Native. ?Rare. First published record: Cope (2007). Also recorded: Bahrain, Kuwait, E<br />

Saudi Arabia.<br />

habitat & distribution: The typical habitats include seashores; but also found in irrigated lawns<br />

and gardens.<br />

notes: Cope maps a record for this species over Doha. He notes that much <strong>of</strong> <strong>the</strong> material originally<br />

named as Henrardia pubescens (Bertol.) C.E. Hubb. is this species, though he does accept that<br />

Henrardia also ‘probably’ occurs.<br />

Photos: None.<br />

Paspalidium desertorum (A. Rich.) Stapf<br />

Growth form: Perennial grass. Fl. Apr.<br />

status: Native. Rare. Not in o<strong>the</strong>r Gulf floras.<br />

habitat & distribution: Recorded by El Amin from irrigated soils at a large farm.<br />

Photos: Coll 397 L .<br />

Pennisetum divisum (J.F. Gmel.) Henrard Plate 105<br />

Pennisetum dichotomum (Forssk.) Delile<br />

Vernacular names: bristle grass, <strong>the</strong>imoom<br />

Growth form: Perennial grass. Fl. Apr–May.<br />

status: Native. Common. Also recorded: Bahrain, E Saudi Arabia, UAE.<br />

habitat & distribution: Frequent to abundant on sand sheets, drifting sand and small dunes,<br />

especially in S <strong>Qatar</strong>.<br />

uses: Batanouny remarks that it is grazed, but not preferred by animals. The species has potential<br />

to be used as a sand dune fixer (El Amin).<br />

Photos: El Amin 25, 136 L ; Bat 119; Coll 398 L ; Corn 142; Jong 80; Phil 117; West 162.<br />

Pennisetum setaceum (Forssk.) Chiov.<br />

Growth form: Perennial grass. Fl. Not known.<br />

status: Native. Rare. First published record: Cope (2007). Also recorded: UAE.<br />

habitat & distribution: A plant <strong>of</strong> rocky areas. The map in Cope shows a locality marked on <strong>the</strong><br />

west coast.<br />

Photos: Jong 81; Pick 256.<br />

Phalaris minor Retz.<br />

Vernacular names: small canary grass, sha’ir al-far<br />

Growth form: <strong>An</strong>nual grass. Fl. Apr.<br />

status: Introduced. Common. Also recorded: Bahrain, E Saudi Arabia, UAE.<br />

habitat & distribution: A weed species <strong>of</strong> cultivated areas.<br />

uses: Used for fodder (El Amin).<br />

Photos: El Amin 136 L ; Coll 402 L ; Jong 83.<br />

موميث<br />

رافلا ريعش<br />

Phalaris paradoxa L.<br />

Vernacular names: sha’ir al-far<br />

Growth form: <strong>An</strong>nual grass. Fl. Apr.<br />

status: Introduced. Common. Also recorded: E Saudi Arabia.<br />

habitat & distribution: Recorded from gardens and cultivated rodat.<br />

uses: Used for fodder (El Amin).<br />

Photos: Coll 402 L .<br />

Phragmites australis (Cav.) Trin. ex Steud.<br />

Phragmites communis Trin.<br />

Vernacular names: reed, qaSSab, ghab, booS<br />

Growth form: Perennial grass. Fl. Apr.<br />

status: Native. Common. Also recorded: Bahrain, Kuwait, E Saudi Arabia, UAE.<br />

80 81<br />

رافلا ريعش<br />

صوب ,باغ ,بصق<br />

habitat & distribution: Locally dominant as reedbed fringing waste water lagoons and o<strong>the</strong>r<br />

permanent wetlands, especially at Abu Nakhla and Abu Hamour lagoons; also at Ras Laffan,<br />

Mesaieed and Dukhan.<br />

uses: El Amin notes that it is used for fodder, and also for thatching and firewood. Rizk & El-Ghazaly<br />

list many medicinal uses, including in <strong>the</strong> treatment <strong>of</strong> leukaemia, cholera, arthritis, coughs,<br />

earaches, fevers, nausea and toothache. The rhizome is used for acute arthritis and jaundice.<br />

notes: All records for <strong>Qatar</strong> refer to subsp. australis, but subsp. altissimus (Benth.) D. Rivera & M.A.<br />

Carreras occurs in Saudi Arabia and UAE (Cope).<br />

Photos: El Amin 25; Bat 120; Coll 403 L ; Corn 245; Jong 84; Mand 243; Phil 118; West 162.<br />

Poa annua L.<br />

Vernacular names: annual meadow grass<br />

Growth form: <strong>An</strong>nual grass. Fl. Jan–Dec.<br />

status: Introduced. Rare. Also recorded: E Saudi Arabia, UAE.<br />

habitat & distribution: Reported by Batanouny as a rare weed <strong>of</strong> lawns.<br />

Photos: Coll 403 L ; Jong 85 L .<br />

Polypogon monspeliensis (L.) Desf. Plate 106<br />

طقلا ليذ<br />

Vernacular names: rabbit-foot grass, annual beard-grass, dhail al quT<br />

Growth form: <strong>An</strong>nual grass. Fl. Apr.<br />

status: Native. Common. Also recorded: Bahrain, E Saudi Arabia, UAE.<br />

habitat & distribution: Frequent to abundant in damp, sandy places; especially at <strong>the</strong> edges <strong>of</strong><br />

irrigation channels and freshwater lagoons. Tolerant <strong>of</strong> low levels <strong>of</strong> salinity.<br />

Photos: El Amin 25; Bat 120; Coll 406 L ; Corn 248; Jong 86; West 163.<br />

Rostraria pumila (Desf.) Tzvelev<br />

Koeleria pumila (Desf.) Domin; Lophochloa pumila (Desf.) Bor<br />

Growth form: <strong>An</strong>nual grass. Fl. Apr.<br />

status: Native. Local. Also recorded: E Saudi Arabia, UAE.<br />

POACEAE<br />

habitat & distribution: Sandy habitats; occasional in gardens; Dukhan township, 2007 (JN).<br />

uses: Mandaville and Jongbloed et al. note its value as a spring grazing grass.<br />

Photos: Coll 407 L ; Jong 88.


POACEAE<br />

Schismus arabicus Nees Plate 109<br />

Growth form: <strong>An</strong>nual grass. Fl. Mar–Apr.<br />

status: Native. ?Very common. Also recorded: Bahrain, Kuwait, E Saudi Arabia, UAE.<br />

habitat & distribution: Schismus species are frequent to abundant over <strong>Qatar</strong> in sandy soils. Cope<br />

describes <strong>the</strong> habitat <strong>of</strong> S. arabicus as ‘dry banks, wadi beds and sandy hollows’.<br />

uses: Widely known as an important grazing species across parts <strong>of</strong> Arabia, due to its abundance<br />

after rains.<br />

notes: Very closely related to S. barbatus; differences are given by Cope. Batanouny listed S. barbatus<br />

without mentioning S. arabicus, whilst El Amin listed as S. arabicus, with S. barbatus as a synonym.<br />

Both are confirmed for <strong>Qatar</strong> by Cope.<br />

Photos: Bat 120; Coll 408 L ; Corn 251.<br />

Schismus barbatus (L.) Thell.<br />

Growth form: <strong>An</strong>nual grass. Fl. Mar–Apr.<br />

status: Native. ?Common. Also recorded: Bahrain, UAE.<br />

habitat & distribution: Described as ‘sandy plains, <strong>of</strong>ten overlying limestone’ (Cope).<br />

notes: See comments under S. arabicus.<br />

Photos: Coll 409 L ; Corn 251; Jong 90; Mand 246.<br />

Setaria verticillata (L.) P. Beauv.<br />

Vernacular names: rough bristle-grass, luSeeq<br />

Growth form: <strong>An</strong>nual grass. Fl. May.<br />

status: Introduced. Local. Also recorded: Bahrain, E Saudi Arabia, UAE.<br />

habitat & distribution: A weed <strong>of</strong> cultivated land.<br />

Photos: El Amin 25; Coll 410 L ; Corn 249; Jong 91; Pick 258; West 163.<br />

Setaria viridis (L.) P. Beauv.<br />

Growth form: <strong>An</strong>nual grass. Fl. May.<br />

status: Introduced. Local. Also recorded: E Saudi Arabia, UAE.<br />

habitat & distribution: A weed <strong>of</strong> damp areas in farms and gardens.<br />

uses: The plant is applied externally for bruises; various medicinal uses <strong>of</strong> <strong>the</strong> seed are reported<br />

(Rizk & El-Ghazaly).<br />

Photos: Coll 411 L .<br />

Sorghum halepense (L.) Pers.<br />

Vernacular names: Johnson grass<br />

Growth form: Perennial grass. Fl. Apr.<br />

status: Introduced. Rare. Also recorded: E Saudi Arabia, UAE.<br />

habitat & distribution: Recorded from cultivated fields in C <strong>Qatar</strong>.<br />

uses: Used for fodder (El Amin). Cultivated as forage in many parts <strong>of</strong> <strong>the</strong> world (Mandaville).<br />

Photos: Coll 412 L ; Jong 93.<br />

Sorghum x drummondii (Nees ex Steud.) Millsp. & Chase<br />

Vernacular names: Sudan grass<br />

Growth form: <strong>An</strong>nual or short-lived perennial grass. Fl. Not known.<br />

قيصل<br />

status: Native. First published record: Cope (2007). Also recorded: E Saudi Arabia.<br />

habitat & distribution: May occur in farm areas as an escape from cultivation.<br />

uses: Cultivated for fodder.<br />

notes: Derived from Sorghum bicolor (L.) Moench. and its wild ancestor S. arundinaceum (Desv.)<br />

Stapf. (Cope).<br />

Photos: None.<br />

Sphenopus divaricatus (Gouan) Rchb.<br />

Growth form: <strong>An</strong>nual grass. Fl. Apr.<br />

status: Native. Rare. Also recorded: Bahrain, UAE.<br />

habitat & distribution: Previously recorded by El Amin from saline, sandy soils in S <strong>Qatar</strong>.<br />

Photos: Coll 412 L ; Jong 94 L .<br />

Sporobolus ioclados (Nees ex Trin.) Nees Plate 107<br />

Sporobolus arabicus Boiss.<br />

Vernacular names: Sakham, rashad, nejma<br />

Growth form: Perennial grass. Fl. Apr.<br />

status: Native. Common. Also recorded: Bahrain, E Saudi Arabia, UAE.<br />

ةمجن ,دشار ,ماخص<br />

habitat & distribution: Dominant in moderately saline sandy and silty areas, including sabkha<br />

edge and o<strong>the</strong>r coastal habitats.<br />

uses: Used for fodder (El Amin).<br />

Photos: El Amin 26, 141 L ; Bat 121; Coll 414 L ; Corn 242; Jong 95; Phil 119; West 164.<br />

Sporobolus spicatus (Vahl) Kunth<br />

Vernacular names: drop seed grass, Sakham<br />

Growth form: Perennial grass. Fl. Apr.<br />

status: Native. Common. Also recorded: Bahrain, E Saudi Arabia, UAE.<br />

habitat & distribution: Occasional to frequent in urban areas, particularly irrigated roadside<br />

plantations, lawns and gardens; tolerant <strong>of</strong> salinity.<br />

Photos: El Amin 136 L ; Coll 416 L ; Corn 251; Jong 96; Phil 120; West 164.<br />

Stipa capensis Thunb. Plates 98, 108<br />

Vernacular names: spear grass, fea<strong>the</strong>r grass, needle grass, Sama’a<br />

Growth form: <strong>An</strong>nual grass. Fl. Apr.<br />

status: Native. Very common. Also recorded: Bahrain, E Saudi Arabia, UAE.<br />

82 83<br />

POACEAE<br />

habitat & distribution: Frequent to abundant and <strong>of</strong>ten dominant after winter rains on compact,<br />

shallow sandy soils in rocky areas and hamada; also in shallow sandy depressions and runnels.<br />

uses: Batanouny remarks that it is grazed when young, but also that it can be injurious to animals,<br />

due to <strong>the</strong> sharp awns, when fruiting. Mandaville states that it is used as standing hay after fruits<br />

have fallen.<br />

Photos: El Amin 141 L ; Bat 122; Coll 417 L ; Corn 140; Jong 97; Mand 252; Phil 121.<br />

Stipagrostis ciliata (Desf.) De Winter<br />

Growth form: Perennial grass. Fl. Feb–Jul.<br />

status: Native. Rare. First published record: Cope (2007). Also recorded: E Saudi Arabia, UAE.<br />

ماخص<br />

ةعمص


POACEAE / TyPHACEAE<br />

habitat & distribution: Seen only once recently, in W <strong>Qatar</strong> (S. Aspinall & JN, 2007), growing in<br />

compact, silty soil.<br />

Photos: Coll 418 L ; Jong 98; Mand 241.<br />

Stipagrostis obtusa (Delile) Nees<br />

Vernacular names: nuSi/naSi<br />

Growth form: Perennial grass. Fl. Apr.<br />

status: Native. Common. Also recorded: E Saudi Arabia, UAE.<br />

habitat & distribution: Frequent in gravel plains, gravelly depressions and o<strong>the</strong>r areas with<br />

compact soil.<br />

uses: Used for fodder (El Amin).<br />

Photos: Bat 122; Coll 420 L .<br />

Stipagrostis plumosa (L.) Munro ex T. <strong>An</strong>derson Plate 110<br />

Vernacular names: nuSi/naSi<br />

يصن<br />

Growth form: Perennial grass. Fl. Apr.<br />

status: Native. Very common. Also recorded: Bahrain, E Saudi Arabia, UAE.<br />

habitat & distribution: Frequent in sandy habitats; possibly more so in C and S <strong>Qatar</strong>.<br />

uses: Noted by several authors to be good fodder for livestock. Mandaville and Batanouny mention<br />

it being ga<strong>the</strong>red in Saudi Arabia to supplement grazing, or sold as hay.<br />

Photos: El Amin 26; Bat 123; Coll 420 L ; Corn 237; Jong 99; Phil 122; West 165.<br />

Stipagrostis sokotrana (Vierh.) De Winter<br />

Vernacular names: nuSi/naSi<br />

يصن<br />

Growth form: Perennial grass. Fl. Not known.<br />

status: Native. ?Local. First published record: Cope (2007). Endemic to <strong>the</strong> Arabian Peninsula.<br />

Also recorded: Bahrain, UAE.<br />

habitat & distribution: Recorded at Ras Laffan (JN, 2004), but <strong>the</strong> specimen requires confirmation.<br />

Locally common here on compact, silty soils; may <strong>the</strong>refore be more widespread in N <strong>Qatar</strong>.<br />

Photos: Corn 237; Phil 123.<br />

TYPHACEAE<br />

Typha domingensis (Pers.) Poir. ex Steud.<br />

Vernacular names: cat’s-tail, reedmace, bardi, booS, deeS<br />

سيد ,صوب ,يدرب<br />

Growth form: Perennial herb. Fl. May–Aug.<br />

status: Native. Local. Also recorded: Bahrain, Kuwait, E Saudi Arabia, UAE.<br />

habitat & distribution: Recorded from a small number <strong>of</strong> sites by Batanouny. It grows at Abu<br />

Nakhla wastewater ponds south <strong>of</strong> Doha (MAS) and is likely to be more frequent now, due to <strong>the</strong><br />

increase in permanently wet areas associated with sewage treatment works and irrigation schemes.<br />

uses: Batanouny notes that this species is used for many purposes such as baskets, paper-making<br />

and cattle feed in o<strong>the</strong>r countries. Ghazanfar reports that <strong>the</strong> flowers are used as a coolant for burns.<br />

Photos: Bat 14; Coll 720; Corn 257; Jong 104; Pick 260; Shuaib 51; West 167.<br />

يصن<br />

References & Bibliography<br />

The list <strong>of</strong> references includes all those cited in <strong>the</strong> text and additional key publications relating to <strong>Qatar</strong>, o<strong>the</strong>r<br />

Gulf states or to <strong>the</strong> Arabian vegetation in general.<br />

Abbas, J.A. & El-Oqlah, A.A. (1992). Distribution and communities <strong>of</strong> halophytic plants in<br />

Bahrain. Journal <strong>of</strong> Arid Environments 22: 205–218.<br />

Abdel Bari E.M., Fahmy, G., Al Thani, N., Al Thani, R. & Abdel-Dayem, M. (2007). The Ghaf<br />

Tree, Prosopis cineraria in <strong>Qatar</strong>. <strong>Qatar</strong> University and National Council for Culture, Arts and<br />

Heritage, Doha.<br />

Abdel Bari, E.M. (1995). Ophioglossum polyphyllum A. Braun, A new record for <strong>Qatar</strong>. <strong>Qatar</strong><br />

University Science Journal 15(2): 369–372.<br />

Abdel Bari, E.M. (1997). Additions to <strong>the</strong> flora <strong>of</strong> <strong>Qatar</strong>. <strong>Qatar</strong> University Science Journal 17(2):<br />

303–312.<br />

Abdul Majid, S. (2008). Vegetation mapping and anthropogenic effects in Al-Reem Biosphere<br />

Reserve in <strong>Qatar</strong>. MSc <strong>the</strong>sis, University College London.<br />

Abulfatih, H.A., Abdel Bari, E.M., Alsubaey, A. & Ibrahim, Y.M. (2001). Vegetation <strong>of</strong> <strong>Qatar</strong>.<br />

Scientific and Applied Research Center (SARC), University <strong>of</strong> <strong>Qatar</strong>.<br />

Al-Madheed, M.A. (2004). Guide to <strong>the</strong> wild plants in <strong>the</strong> State <strong>of</strong> <strong>Qatar</strong>. [in Arabic]<br />

<strong>An</strong>giosperm Phylogeny Group (2003). <strong>An</strong> update <strong>of</strong> <strong>the</strong> <strong>An</strong>giosperm Phylogeny Group<br />

classification for <strong>the</strong> orders and families <strong>of</strong> flowering plants: APG II. Botanical Journal <strong>of</strong> <strong>the</strong><br />

Linnean Society 141: 399–436.<br />

<strong>An</strong>on (undated). Wild flora <strong>of</strong> Ras Laffan Industrial City. Information booklet published by<br />

Environment Section, Ras Laffan Industrial City.<br />

<strong>An</strong>on (undated). Mangrove Conservation Programme. Information booklet published by<br />

Environment Section, Ras Laffan Industrial City.<br />

Arab Organization for Agricultural Development (comp. El Amin, H.M.) (1983). Wild plants <strong>of</strong><br />

<strong>Qatar</strong>. Published by AOAD, Khartoum (printed Richmond, UK).<br />

Babikir, A.A.A. (1984). Vegetation and environment on <strong>the</strong> coastal sand dunes and playas <strong>of</strong><br />

Khor El-Odaid area, <strong>Qatar</strong>. Geo-Journal 9: 377–385.<br />

Babikir, A.A.A. (1986). The vegetation <strong>of</strong> natural depressions in <strong>Qatar</strong> in relation to climate and<br />

soil. Journal <strong>of</strong> Arid Environments 10: 165–173.<br />

Barth, H-J & Böer, B. (eds), (2002). Sabkha ecosystems, Volume I: The Arabian Peninsula and adjacent<br />

countries, Tasks for Vegetation Science 36. Kluwer Academic Publishers, The Ne<strong>the</strong>rlands.<br />

Batanouny, K.H. (1981). Ecology and <strong>Flora</strong> <strong>of</strong> <strong>Qatar</strong>. Alden Press, Oxford.<br />

Batanouny, K.H. (2001). Plants in <strong>the</strong> deserts <strong>of</strong> <strong>the</strong> Middle East. Springer-Verlag, Berlin.<br />

Batanouny, K.H. & Turki, A.A. (1983). Vegetation <strong>of</strong> South Western <strong>Qatar</strong>. Arab Gulf J. Scient. Res.<br />

1: 5–19.<br />

Beier, B.-A., Chase, M.W. & Thulin, M. (2003). Phylogenetic relationships and taxonomy <strong>of</strong><br />

subfamily Zygophylloideae (Zygophyllaceae) based on molecular and morphological data Plant<br />

Syst. Evol. 240: 11–39.<br />

Böer, B. (1999). Ecosystems, anthropogenic impacts and habitat management techniques in Abu<br />

Dhabi. Paderborner Geographische Studien 12 (University <strong>of</strong> Paderborn, Germany).<br />

84 85


Böer, B.B. & Norton, J.A. (1996a). Changes in <strong>the</strong> vegetation <strong>of</strong> <strong>the</strong> NARC Sweihan enclosure<br />

between 1993 and 1995. Internal Research Report, National Avian Research Centre, Abu Dhabi.<br />

Böer, B.B. & Norton, J.A. (1996b). Vegetation rehabilitation in an enclosure in Abu Dhabi.<br />

Fresenius Envir. Bull. 5: 79–84.<br />

Böer, B. & Al Hajiri, S. (2002). The coastal and sabkha flora <strong>of</strong> <strong>Qatar</strong>: <strong>An</strong> introduction. In: Barth,<br />

H-J & Böer, B. (eds), Sabkha ecosystems, Volume I: The Arabian Peninsula and adjacent countries, Tasks<br />

for Vegetation Science 36, pp. 63–70. Kluwer Academic Publishers, The Ne<strong>the</strong>rlands.<br />

Boulos, L. (1999–2005). <strong>Flora</strong> <strong>of</strong> Egypt, Vols 1–4. Al Hadara Publishing, Cairo.<br />

Boulos, L. & Al-Dosari, M. (1994). <strong>Checklist</strong> <strong>of</strong> <strong>the</strong> flora <strong>of</strong> Kuwait. Journal <strong>of</strong> <strong>the</strong> University <strong>of</strong><br />

Kuwait (Science) 21: 203–281.<br />

Chaudhary, S.A. (1989). Grasses <strong>of</strong> Saudi Arabia. Ministry <strong>of</strong> Agriculture and Water, Riyadh.<br />

Chaudhary, S.A. (1999). Vegetation <strong>of</strong> <strong>the</strong> Kingdom <strong>of</strong> Saudi Arabia. Ministry Of Agriculture, KSA.<br />

Collenette, S. (1985). <strong>An</strong> illustrated guide to <strong>the</strong> flowers <strong>of</strong> Saudi Arabia. Meteorological and<br />

Environmental Protection Administration Publication No. 1. Scorpion Publishing, London.<br />

Collenette, S. (1999). Wildflowers <strong>of</strong> Saudi Arabia. National Commission for Wildlife Conservation<br />

and Development & Sheila Collenette, Riyadh.<br />

Cope, T.A. (2007). <strong>Flora</strong> <strong>of</strong> <strong>the</strong> Arabian Peninsula and Socotra. Volume 5, Part 1. Edinburgh University<br />

Press.<br />

Cornes, M.D. & Cornes, C.D. (1989). The wild flowering plants <strong>of</strong> Bahrain; an illustrated guide.<br />

IMMEL Publishing, London.<br />

Daoud, H.S. & Al-Rawi, A. (1985). <strong>Flora</strong> <strong>of</strong> Kuwait. Volume 1: Dicotyledoneae. KPI Limited, London<br />

& University <strong>of</strong> Kuwait.<br />

Dickson (1938) and o<strong>the</strong>rs, etc. take refs from Mand. and mention in text.<br />

Dickson, H. (1951). To add. Referenced under Halocnemum strobilaceum.<br />

Dickson, V. (1955). The wild flowers <strong>of</strong> Kuwait and Bahrain. George Allen & Unwin, London.<br />

El Amin, H.M. (Arab Organization for Agricultural Development) (1983). Wild plants <strong>of</strong> <strong>Qatar</strong>.<br />

Published by AOAD, Khartoum (printed Richmond, UK).<br />

Ghazanfar, S.A. (1992). <strong>An</strong> annotated catalogue <strong>of</strong> <strong>the</strong> vascular plants <strong>of</strong> Oman and <strong>the</strong>ir vernacular<br />

names. Scripta Botanica Belgica 2, National Botanic Garden <strong>of</strong> Belgium, Meise, Belgium.<br />

Ghazanfar, S.A. (1994). Handbook <strong>of</strong> Arabian medicinal plants. Springer, New York.<br />

Ghazanfar, S.A. (2003). <strong>Flora</strong> <strong>of</strong> Oman, Volume 1, Piperaceae–Primulacaea. Scripta Botanica Belgica<br />

25. National National Botanic Garden <strong>of</strong> Belgium, Meise, Belgium.<br />

Ghazanfar, S.A. & Fisher, M., Eds (1998). Vegetation <strong>of</strong> <strong>the</strong> Arabian Peninsula. Geobotany 25. Kluwer<br />

Academic Publishers, Dordrecht, The Ne<strong>the</strong>rlands.<br />

Gillespie, F. (2006). Discovering <strong>Qatar</strong>. Creative Writing and Photography, Rimons, France.<br />

Jongbloed, M., Western, A.R. & Böer, B. (2000). <strong>An</strong>notated check-list for plants in <strong>the</strong> U.A.E. Zodiac<br />

Publishing, Dubai, UAE.<br />

Jongbloed, M.V.D., Feulner, G.R., Böer, B. & Western, A.R. (2003). The comprehensive guide to <strong>the</strong><br />

wildflowers <strong>of</strong> <strong>the</strong> United Arab Emirates. Environmental Research and Wildlife Development Agency,<br />

Abu Dhabi.<br />

Mandaville, J.P. (1990). <strong>Flora</strong> <strong>of</strong> Eastern Saudi Arabia. Kegan Paul International and NCWCD.<br />

Migahid, A.M. (1998). <strong>Flora</strong> <strong>of</strong> Saudi Arabia. Vols 1 & 2. Third Edition. University Libraries, King<br />

Saud University, Riyadh.<br />

Miller, A.G. & Cope, T.A. (1996). <strong>Flora</strong> <strong>of</strong> <strong>the</strong> Arabian Peninsula and Socotra. Volume 1. Edinburgh<br />

University Press.<br />

Obeid, M. (1975). <strong>Qatar</strong> – study <strong>of</strong> <strong>the</strong> natural vegetation. FAO AGO QAT/74/003.<br />

Phillips, D.C. (1988). Wild flowers <strong>of</strong> Bahrain. A field guide to herbs, shrubs and trees. Privately<br />

published, Bahrain.<br />

Pickering, H. & Patzelt, A. (2008). Field guide to <strong>the</strong> wild plants <strong>of</strong> Oman. Kew Publishing,<br />

Richmond, Surrey.<br />

Rizk, A.M. & El-Ghazaly, G.A. (1995). Medicinal and poisonous plants <strong>of</strong> <strong>Qatar</strong>, Scientific and<br />

Applied Research Centre, University <strong>of</strong> <strong>Qatar</strong>.<br />

Shuaib, L. (1995). Wild flowers <strong>of</strong> Kuwait. Stacey International, London.<br />

Tayab, M.R. (2002). Assessment <strong>of</strong> <strong>the</strong> distribution and abundance <strong>of</strong> flora and fauna along <strong>the</strong><br />

coastal areas <strong>of</strong> Ras Laffan Industrial City, <strong>Qatar</strong>. <strong>Qatar</strong> Petroleum Report.<br />

Western, A.R. (1989). The flora <strong>of</strong> <strong>the</strong> United Arab Emirates: an introduction. United Arab Emirates<br />

University.<br />

Zohary, M. (1973). Geobotanical foundations <strong>of</strong> <strong>the</strong> Middle East. Fischer Verlag.<br />

86 87


Internet Resources<br />

Aluka<br />

http://www.aluka.org<br />

‘Digital library <strong>of</strong> scholarly resources from and about Africa’. Resources for plants are extensive and<br />

include photographs <strong>of</strong> specimens from historical collections.<br />

ePIC (electronic Plant Information Centre) (Royal Botanic Gardens, Kew, UK)<br />

http://epic.kew.org/<br />

Provides a convenient way to search IPNI and various resources at Kew, including bibliographies,<br />

herbarium collections and seed database.<br />

Euro+Med PlantBase<br />

http://ww2.bgbm.org/EuroPlusMed/query.asp<br />

<strong>An</strong> on-line database and information system for <strong>the</strong> vascular plants <strong>of</strong> Europe and <strong>the</strong> Mediterranean<br />

region. At <strong>the</strong> time <strong>of</strong> going to press it covered three <strong>of</strong> <strong>the</strong> largest families: Asteraceae, Poaceae<br />

and Rosaceae, and 29 smaller families.<br />

Global Biodiversity Information Facility (GBIF)<br />

http://data.gbif.org/welcome.htm<br />

Provides mapped world distributions for species, based on herbarium records, though <strong>the</strong>se do not<br />

include any collections from <strong>the</strong> Arabian Peninsula.<br />

Global Compendium <strong>of</strong> Weeds (GCW)<br />

http://www.hear.org/gcw/<br />

Useful information on status and world distribution <strong>of</strong> weed species.<br />

Germplasm Resources Information Network (GRIN)<br />

http://www.ars-grin.gov/cgi-bin/npgs/html/index.pl?language=en<br />

Useful general information on taxonomy and world distribution <strong>of</strong> plants. Covers species occurring<br />

in <strong>the</strong> USA, but includes many species naturalised in <strong>Qatar</strong>.<br />

International Plant Names Index (IPNI)<br />

http://www.ipni.org/index.html<br />

Provides information on plant names and bibliographic details. A collaboration between The Royal<br />

Botanic Gardens, Kew, The Harvard University Herbaria, and <strong>the</strong> Australian National Herbarium.<br />

USDA Plants Database<br />

http://plants.usda.gov/index.html<br />

A searchable database <strong>of</strong> information about <strong>the</strong> vascular plants and lower plants <strong>of</strong> <strong>the</strong> USA and<br />

its territories.<br />

Wikipedia<br />

http://en.wikipedia.org/wiki<br />

Useful for general information on taxonomy, distribution and uses <strong>of</strong> many species.<br />

Index to Families and Species<br />

The alphabetical index covers scientific names <strong>of</strong> families and species listed in <strong>the</strong> main accounts and those<br />

referred to in <strong>the</strong> introductory pages, toge<strong>the</strong>r with additional species mentioned under Notes. Synonyms are<br />

shown in italics and plate numbers in bold.<br />

Aaronsohnia factorovskyi 6, 13<br />

Abutilon fruticosum 52<br />

Abutilon hirtum 52<br />

Abutilon pannosum 52<br />

Acacia arabica 38<br />

Acacia ehrenbergiana 38, 58<br />

Acacia flava 38<br />

Acacia nilotica 38<br />

Acacia raddiana 39<br />

Acacia spirocarpa 39<br />

Acacia tortilis 39, 59<br />

Acacia viii<br />

ACANTHACEAE 1<br />

Aeluropus lagopoides ix, 72, 99<br />

Aerva javanica 3<br />

Aerva persica 3<br />

Aerva tomentosa 3<br />

Agathophora alopecuroides 26<br />

Agathophora iraqensis 26<br />

Agriophyllum minus viii, 26, 43<br />

Agriophyllum montasiri 26<br />

AIZOACEAE 2<br />

Aizoon canariense 2, 6<br />

Aizoon hispanicum 2<br />

Alhagi camelorum 39<br />

Alhagi graecorum 39<br />

Alhagi maurorum 39<br />

Althaea ludwigii viii, 52<br />

AMARANTHACEAE 3<br />

Amaranthus chlorostachys 3<br />

Amaranthus gracilis 4<br />

Amaranthus graecizans 3<br />

Amaranthus hybridus 3<br />

Amaranthus viridis 3<br />

Ammi majus 4<br />

<strong>An</strong>abasis setifera 26, 40<br />

<strong>An</strong>agallis arvensis 54<br />

<strong>An</strong>astatica hierochuntica 17, 28, 29<br />

<strong>An</strong>chusa hispida 15, 24<br />

<strong>An</strong>drachne telephioides viii, 35, 56<br />

<strong>An</strong>ethum graveolens 4<br />

ANGIOSPERMAE 1<br />

<strong>An</strong>isantha madritensis 73<br />

<strong>An</strong>tirrhinum orontium 62<br />

APIACEAE 4<br />

APOCyNACEAE 5<br />

ARECACEAE 69<br />

Argyrolobium abyssinicum 39<br />

Argyrolobium arabicum 39<br />

Aristida abnormis 72<br />

Aristida adscensionis 72<br />

Aristida meccana 72<br />

Aristida mutabilis 72<br />

Arnebia decumbens 15<br />

Arnebia hispidissima 15, 22<br />

Artemisia herba-alba 6<br />

Artemisia inculta 6<br />

Artemisia sieberi 6<br />

Arthrocnemum glaucum 26<br />

Arthrocnemum macrostachyum ix, 26<br />

Arundo donax 73<br />

ASCLEPIADACEAE 5<br />

Asphodelus fistulosus 71<br />

Asphodelus tenuifolius 71<br />

Aster squamatus 14<br />

ASTERACEAE 6<br />

Asteriscus hierichunticus 11<br />

Asteriscus pygmaeus 11<br />

As<strong>the</strong>na<strong>the</strong>rum forsskaolii 74<br />

Astragalus annularis 39, 60<br />

Astragalus corrugatus 40<br />

Astragalus eremophilus viii, 40, 61<br />

Astragalus gyzensis 40<br />

Astragalus hamosus 40<br />

Astragalus hauarensis 40<br />

Astragalus schimperi 40<br />

Astragalus sieberi 41<br />

Astragalus tenuirugis 40<br />

Astragalus tribuloides 41<br />

Astragalus zubairensis 41<br />

Atractylis carduus 6, 14<br />

Atractylis flava 6<br />

88 89


Atriplex leucoclada 26, 41<br />

Avena sterilis 73<br />

Avicennia marina ix, 1, 4<br />

Avicennia <strong>of</strong>ficinalis 1<br />

Bassia eriophora 27<br />

Bassia muricata 27<br />

Beta vulgaris 27<br />

Bienertia cycloptera 27<br />

Blepharis ciliaris 2, 2<br />

Blepharis edulis 2<br />

Bolboschoenus maritimus 69<br />

BORAGINACEAE 15<br />

Brachypodium distachyum 73<br />

Brassica rapa 17<br />

Brassica tournefortii 17<br />

BRASSICACEAE 17<br />

Bromus danthoniae 73<br />

Bromus haussknechtii 73<br />

Bromus madritensis 73<br />

Bromus scoparius 73<br />

Bupleurum semicompositum 4<br />

Calendula arvensis 7<br />

Calendula tripterocarpa 7<br />

Calligonum comosum 57<br />

Calligonum polygonoides 57<br />

Calotropis procera 5, 8<br />

CAPPARACEAE x, 21<br />

Capparis spinosa 21<br />

Carduncellus eriocephalus 7<br />

Carrichtera annua 18<br />

Carthamnus eriocephalus 7<br />

CARyOPHyLLACEAE x, 22<br />

Cassia italica 46<br />

Cassia obovata 46<br />

Cassia occidentalis 47<br />

Cassia senna 46<br />

Casuarina cunninghamiana 25<br />

Casuarina equisetifolia 25<br />

CASUARINACEAE 25<br />

Cenchrus ciliaris 74, 92<br />

Cenchrus pennisetiformis 74<br />

Cenchrus setigerus 74<br />

Centaurea pseudosinaica 7<br />

Centaurea sinaica 7, 15<br />

Centaurium pulchellum 49<br />

Centropodia forsskaolii 74, 100<br />

Chamaesyce arabica 35<br />

Chamaesyce granulata 35<br />

Chamaesyce hirta 36<br />

Chamaesyce prostrata 36<br />

Chamaesyce serpens 36<br />

CHENOPODIACEAE 26<br />

Chenopodium aegyptiacum 31<br />

Chenopodium album 28<br />

Chenopodium murale 28<br />

Chloris barbata 74<br />

Chloris virgata 75<br />

Chrozophora obliqua 36<br />

Chrozophora oblongifolia 36<br />

Chrozophora tinctoria 36<br />

Chrozophora verbascifolia 36<br />

Chrysan<strong>the</strong>mum coronarium 8<br />

Chrysopogon gryllus 75<br />

Chrysopogon plumulosus 75, 95, 101<br />

Cichorium pumilum 7<br />

CISTACEAE 31<br />

Cistanche lutea 54<br />

Cistanche phelypaea 54<br />

Cistanche tinctoria 54<br />

Cistanche tubulosa 54, 76<br />

Citrullus colocynthis 34, 54<br />

CLEOMACEAE x<br />

Cleome africana 21<br />

Cleome amblyocarpa 21<br />

Cleome brachycarpa 21<br />

Cleome dolichostyla 22<br />

Cleome noeana 22<br />

Cleome scaposa 22<br />

Cocculus pendulus 53, 74<br />

COMBRETACEAE 32<br />

COMPOSITAE x<br />

Conocarpus lancifolius 32<br />

CONVOLVULACEAE 32<br />

Convolvulus arvensis 32<br />

Convolvulus buschiricus 32<br />

Convolvulus cephalopodus 32, 52<br />

Convolvulus deserti 33<br />

Convolvulus fatmensis 32<br />

Convolvulus glomeratus 33<br />

Convolvulus microphyllus 33<br />

Convolvulus pilosellifolius 33, 53<br />

Convolvulus prostratus 33<br />

Conyza bonariensis 8<br />

Conyza dioscoridis 11<br />

Conyza maritima 14<br />

Corchorus depressus viii, 64, 85<br />

Corchorus olitorius 65<br />

Corchorus trilocularis 65<br />

Cornulaca aucheri 28<br />

Cornulaca monacantha 28<br />

Cornulaca viii<br />

Coronopus didymus 19<br />

Cressa cretica 33<br />

CRUCIFERAE x<br />

Cryophytum nodiflorum 2<br />

Cucumis colocynthis 34<br />

Cucumis prophetarum 34<br />

CUCURBITACEAE 34<br />

Cullen plicata 41<br />

Cuscuta chinensis 33<br />

Cuscuta pedicellata 34<br />

Cuscuta pentagona 34<br />

Cutandia memphitica 75, 96<br />

Cymbopogon commutatus viii, 75<br />

Cymbopogon parkeri 75<br />

CyMODOCEACEAE 69<br />

Cynodon dactylon 76<br />

CyNOMORIACEAE 35<br />

Cynomorium coccineum 35, 55<br />

CyPERACEAE 69<br />

Cyperus aucheri 70<br />

Cyperus conglomeratus viii, 70, 70, 91<br />

Cyperus jeminicus 70<br />

Cyperus laevigatus 70<br />

Cyperus rotundus 70<br />

Dactyloctenium aegyptium 76<br />

Datura innoxia 63<br />

Dichanthium annulatum 76<br />

Dichanthium foveolatum 76, 97<br />

DICOTYLEDONEAE 1<br />

Digitaria sanguinalis 76<br />

Dipcadi erythraeum viii, 72, 93, 94<br />

Dipcadi serotinum 72<br />

Dipcadi unicolor 72<br />

Diplachne fusca 79<br />

Diplan<strong>the</strong>ra uninervis 69<br />

Diplan<strong>the</strong>ra wrightii 69<br />

Dipterygium glaucum 22, 33<br />

Dodonaea angustifolia 62<br />

Dodonaea viscosa 62<br />

Echinochloa colona 77<br />

Echiochilon jugatum 15, 23<br />

Echiochilon kotschyi 15<br />

Echium horridum 16<br />

Eclipta alba 7<br />

Eclipta prostrata 7<br />

Eleusine compressa 79<br />

Emex spinosa 58, 78<br />

Enicostema axillare 49<br />

Enicostema verticillatum 49<br />

Ephedra ciliata 1<br />

Ephedra foliata x, 1<br />

Ephedra peduncularis 1<br />

EPHEDRACEAE 1<br />

Eragrostis barrelieri 77<br />

Eragrostis cilianensis 77<br />

Eragrostis ciliaris 77<br />

Eragrostis pilosa 77<br />

Eremobium aegyptiacum 18, 30<br />

Eremobium lineare 18<br />

Eremopogon foveolatus 76<br />

Erigeron bonariensis 8<br />

Erodium bryoniifolium 50<br />

Erodium glaucophyllum ix, 49, 68<br />

Erodium laciniatum 50, 69<br />

Erodium oxyrrhynchum 50<br />

Erodium pulverulentum 50<br />

Eruca sativa 18<br />

Eruca vesicaria 18<br />

Erucaria crassifolia 18<br />

Erucaria hispanica 18<br />

Erucaria lineariloba 18<br />

Erythraea pulchella 49<br />

Euphorbia arabica 35<br />

Euphorbia cornuta 37<br />

Euphorbia dracunculoides 37<br />

Euphorbia geniculata 37<br />

Euphorbia granulata 35<br />

Euphorbia heterophylla 37<br />

Euphorbia hirta 36<br />

Euphorbia peplus 37<br />

Euphorbia prostrata 36<br />

Euphorbia retusa 37, 57<br />

Euphorbia serpens 36<br />

90 91


EUPHORBIACEAE 35<br />

FABACEAE 38<br />

Fagonia bruguieri 66<br />

Fagonia glutinosa 67<br />

Fagonia indica 66, 81<br />

Fagonia ovalifolia 67, 86<br />

Fagonia tenuifolia 67<br />

Farsetia arabica 19<br />

Farsetia hamiltonii 19<br />

Farsetia heliophila 19, 31<br />

Farsetia linearis 19<br />

Ficus carica 53<br />

Filago desertorum 8, 12<br />

Filago prolifera 8<br />

Filago spathulata 8<br />

Flaveria trinervia 8<br />

Foeniculum vulgare 4<br />

Forsskaolea tenacissima 65<br />

Forsskaolea viridis 65<br />

Francoeuria crispa 12<br />

Frankenia pulverulenta 49, 71<br />

FRANKENIACEAE 49<br />

Galium tricornutum 61<br />

Gastrocotyle hispida 15<br />

GENTIANACEAE 49<br />

GERANIACEAE 49<br />

Geranium molle 50<br />

Glebionis coronaria 8<br />

Glossonema edule 5<br />

Glossonema varians 5, 9, 10<br />

Gnaphalium luteo-album 9<br />

Gymnarrhena micrantha 8, 17<br />

GYMNOSPERMAE 1<br />

Halocnemum strobilaceum 28<br />

Halodule uninervis 69<br />

Halogeton alopecuroides 26<br />

Halopeplis perfoliata 29, 42<br />

Halophila ovalis 71<br />

Halophila stipulacea 71<br />

Halopyrum mucronatum 77<br />

Haloxylon ammodendron 29<br />

Haloxylon persicum 29<br />

Haloxylon salicornicum viii, 29, 44<br />

Hammada elegans 29<br />

Hammada salicornica 29<br />

Haplophyllum tuberculatum 61<br />

Helian<strong>the</strong>mum kahiricum ix, 31, 50<br />

Helian<strong>the</strong>mum lippii ix, 32, 51<br />

Heliotropium bacciferum 16, 25<br />

Heliotropium crispum 16<br />

Heliotropium kotschyi 16<br />

Heliotropium ramosissimum 16<br />

Heliotropium zeylanicum 16<br />

Henrardia pubescens 78<br />

Herniaria hemistemon 22, 34<br />

Herniaria hirsuta 22<br />

Hippocrepis areolata 41<br />

Hippocrepis bicontorta 41<br />

Hippocrepis constricta 41<br />

Hippocrepis multisiliquosa 42<br />

Hippocrepis unisiliquosa 42<br />

Hordeum glaucum 78<br />

Hordeum murinum 78<br />

Hordeum vulgare 78<br />

HyDROCHARITACEAE 71<br />

Hymenocarpos circinnatus 42<br />

Hyparrhenia hirta 78<br />

Ifloga spicata 9, 18<br />

Indig<strong>of</strong>era articulata 42<br />

Indig<strong>of</strong>era intricata 42, 62<br />

Indig<strong>of</strong>era oblongifolia 42<br />

JUNCACEAE 71<br />

Juncellus laevigatus 70<br />

Juncus arabicus 71<br />

Juncus maritimus 71<br />

Juncus rigidus 71<br />

Koeleria pumila 81<br />

Koelpinia linearis 9, 19<br />

LABIATAE x<br />

LAMIACEAE 50<br />

Laphangium luteoalbum 9<br />

Lappula spinocarpos 16<br />

Lasiurus hirsutus 78<br />

Lasiurus scindicus 78, 98, 102<br />

Lathyrus inconspicuus 42<br />

Launaea capitata 9<br />

Launaea cassiniana 10<br />

Launaea fallax 10<br />

Launaea glomerata 9<br />

Launaea goraeensis 9<br />

Launaea mucronata 10<br />

Launaea nudicaulis 10<br />

Launaea procumbens 10<br />

Lawsonia alba 51<br />

Lawsonia inermis 51<br />

LEGUMINOSAE x<br />

Lepidium aucheri 19<br />

Lepidium didymum 19<br />

Lepidium sativum 19<br />

Leptadenia pyrotechnica 6, 11<br />

Leptochloa fusca 79<br />

LILIACEAE 71<br />

Limonium axillare ix, 57, 79<br />

Limonium carnosum 57<br />

LINACEAE 51<br />

Linum strictum 51<br />

Lippia nodiflora 66<br />

Lithospermum callosum 16<br />

Lithospermum kotschyi 15<br />

Lolium rigidum 79<br />

Lophochloa pumila 81<br />

LORANTHACEAE 51<br />

Loranthus acaciae 51<br />

Lotononis dichotoma 43<br />

Lotononis platycarpa 43<br />

Lotus garcinii 43<br />

Lotus glinoides 43<br />

Lotus halophilus 43, 63<br />

Lotus platycarpa 43<br />

Lotus pusillus 43<br />

Lotus schimperi 43<br />

Lotus villosus 43<br />

Lycium shawii viii, ix, 63, 84<br />

LyTHRACEAE 51<br />

Malva nicaeensis 52<br />

Malva parviflora 52<br />

MALVACEAE 52<br />

Matricaria aurea 10<br />

Matricaria auriculata 14<br />

Matricaria chamomilla 10<br />

Matricaria recutita 10<br />

Matthiola longipetala 19<br />

Medicago aschersoniana 43<br />

Medicago laciniata 43<br />

Medicago polymorpha 44<br />

Medicago sativa 44<br />

Melilotus albus 44<br />

Melilotus indicus 44<br />

MENISPERMACEAE 53<br />

Mercurialis annua 37<br />

Mesembryan<strong>the</strong>mum<br />

cryptanthum 2, 5<br />

Mesembryan<strong>the</strong>mum forskahlii 2<br />

Mesembryan<strong>the</strong>mum<br />

nodiflorum 2, 7<br />

Misopates orontium 62<br />

MOLLUGINACEAE 53<br />

Mollugo cerviana 53<br />

Moltkiopsis ciliata 16, 27<br />

MONOCOTYLEDONEAE 69<br />

Monsonia heliotropioides 50<br />

Monsonia nivea 50, 72<br />

MORACEAE 53<br />

Morus nigra 53<br />

MyRSINACEAE 54<br />

Nerium mascatense 5<br />

Nerium oleander 5<br />

Neurada procumbens 54, 75<br />

NEURADACEAE 54<br />

Ochradenus aucheri 59<br />

Ochradenus baccatus 59<br />

Ochradenus dewittii 59<br />

Ochthochloa compressa 79, 103<br />

Odontospermum pygmaeum 11<br />

Ogastemma pusillum 17, 26<br />

Oligomeris linifolia 60<br />

Oligomeris subulata 60<br />

Ononis reclinata 45<br />

Ononis sicula 45<br />

OPHIOGLOSSACEAE 1<br />

Ophioglossum polyphyllum x, 1, 3<br />

Opophytum forskahlii 2<br />

OROBANCHACEAE 54<br />

Orobanche ramosa 55<br />

OxALIDACEAE 55<br />

Oxalis cernua 55<br />

Oxalis corniculata 55<br />

Oxalis corymbosa 55<br />

Oxalis debilis 55<br />

Oxalis pes-caprae 55<br />

Pallenis hierichuntica 11<br />

Panicum turgidum viii, 79, 104<br />

Parapholis incurva 80<br />

Parietaria alsinifolia 66<br />

92 93


Parkinsonia aculeata 45<br />

Paronychia arabica 23, 35<br />

Paspalidium desertorum 80<br />

Pennisetum dichotomum 80<br />

Pennisetum divisum 80, 105<br />

Pennisetum setaceum 80<br />

Pentanema divaricatum 11<br />

Phalaris minor 80<br />

Phalaris paradoxa 81<br />

Phoenix dactylifera 69, 90<br />

Phragmites australis ix, 81<br />

Phragmites communis 81<br />

Phyla nodiflora 66<br />

Phyllanthus amarus 38<br />

Phyllanthus niruri 38<br />

Physalis angulata 63<br />

Picris aspleniodes 11<br />

Picris cyanocarpa 11<br />

Picris radicata 11<br />

PLANTAGINACEAE 55<br />

Plantago amplexicaulis 55<br />

Plantago ciliata 56, 77<br />

Plantago commutate 56<br />

Plantago coronopus 56<br />

Plantago lanceolata 56<br />

Plantago ovata 56<br />

Plantago psyllium 57<br />

Plicosepalus acaciae 51<br />

Pluchea dioscoridis 11<br />

PLUMBAGINACEAE 57<br />

Poa annua 81<br />

POACEAE 72<br />

Polycarpaea repens ix, 23, 36<br />

Polycarpaea robbairea 23<br />

Polycarpaea spicata 23<br />

Polycarpon tetraphyllum 23<br />

Polygala erioptera 57<br />

Polygala irregularis 57<br />

POLyGALACEAE 57<br />

POLyGONACEAE 57<br />

Polygonum argyrocoleum 58<br />

Polygonum bellardii 58<br />

Polygonum equisetiformis 58<br />

Polypogon monspeliensis ix, 81, 106<br />

Portulaca oleracea 59<br />

PORTULACACEAE 59<br />

Prosopis cineraria vi, 45, 64<br />

Prosopis juliflora vi, viii, 45, 65<br />

Psoralea plicata 41<br />

PTERIDOPHYTA 1<br />

Pulicaria crispa 12<br />

Pulicaria gnaphalodes 12<br />

Pulicaria sicula 12<br />

Pulicaria undulata viii, 12, 16<br />

Pulicaria undulata 12<br />

Raphanus sativus 19<br />

Reichardia orientalis 12<br />

Reichardia tingitana 12, 20<br />

Reseda arabica 60<br />

Reseda aucheri 60<br />

Reseda muricata 60<br />

Reseda subulata 60<br />

RESEDACEAE 59<br />

RHAMNACEAE 60<br />

Rhanterium epapposum 12, 21<br />

Rhynchosia minima 46<br />

Ricinus communis 38<br />

Robbairea delileana 23<br />

Rostraria pumila 81<br />

RUBIACEAE 61<br />

Rumex cyprius 58<br />

Rumex dentatus 58<br />

Rumex spinosus 58<br />

Rumex vesicarius 59, 82<br />

RUTACEAE 61<br />

Salicornia europaea 29, 45<br />

Salicornia herbacea 29<br />

Salicornia maritima 29<br />

Salsola alopecuroides 26<br />

Salsola baryosma 30<br />

Salsola cyclophylla 30<br />

Salsola drummondii 30<br />

Salsola foetida 30<br />

Salsola imbricata 30, 46<br />

Salsola mandavillei 30<br />

Salsola schweinfurthii 30<br />

Salsola soda 29<br />

Salsola vermiculata 30<br />

Salsola villosa 30<br />

Salsola ix, 46<br />

Salvadora persica 61<br />

SALVADORACEAE 61<br />

Salvia aegyptiaca 50, 70<br />

SAPINDACEAE 62<br />

Savignya parviflora 20, 32<br />

Schanginia aegyptiaca 31<br />

Schanginia baccata 31<br />

Schimpera arabica 20<br />

Schimpera persica 20<br />

Schismus arabicus 82, 109<br />

Schismus barbatus 82<br />

Scirpus maritimus 69<br />

Sclerocephalus arabicus ix, 24, 38<br />

Scorpiurus muricatus 46<br />

Scrophularia deserti 62, 80<br />

SCROPHULARIACEAE 62<br />

Seetzenia lanata 67, 87<br />

Seetzenia orientalis 67<br />

Seidlitzia rosmarinus viii, 31, 47<br />

Senecio coronopifolius 13<br />

Senecio desfontainei 13<br />

Senecio glaucus 13<br />

Senecio vulgaris 13<br />

Senna alexandrina 46<br />

Senna italica 46, 66<br />

Senna occidentalis 47<br />

Sesbania aegyptiaca 47<br />

Sesbania sesban 47<br />

Setaria verticillata 82<br />

Setaria viridis 82<br />

Silene arabica 24<br />

Silene conica 24<br />

Silene villosa 24, 39<br />

Sinapis arvensis 20<br />

Sisymbrium erysimoides 20<br />

Sisymbrium irio 20<br />

Sisymbrium orientale 20<br />

SOLANACEAE 63<br />

Solanum elaeagnifolium 63<br />

Solanum nigrum 63<br />

Solanum villosum 63<br />

Sonchus asper 13<br />

Sonchus cassinianus 10<br />

Sonchus maritimus 13<br />

Sonchus oleraceus 14<br />

Sonchus tenerrimus 14<br />

Sorghum arundinaceum 83<br />

Sorghum bicolor 83<br />

Sorghum halepense 82<br />

Sorghum x drummondii 83<br />

Spergula fallax 24, 37<br />

Spergularia bocconei 24<br />

Spergularia diandra 25<br />

Sphenopus divaricatus 83<br />

Sporobolus arabicus 83<br />

Sporobolus ioclados 83, 107<br />

Sporobolus spicatus 83<br />

Stachytarphaeta angustifolia 66<br />

Statice axillaris 57<br />

Stellaria media 25<br />

Stellaria pallida 25<br />

Stipa capensis ix, 83, 98, 108<br />

Stipagrostis ciliata 83<br />

Stipagrostis obtusa 84<br />

Stipagrostis plumosa 84, 110<br />

Stipagrostis sokotrana 84<br />

Stipagrostis ix<br />

Suaeda aegyptiaca 31, 48<br />

Suaeda fruticosa 31<br />

Suaeda vermiculata 31, 49<br />

Symphyotrichum squamatum 14<br />

TAMARICACEAE 64<br />

Tamarix aphylla 64<br />

Tamarix aucheriana 64<br />

Tamarix passerinoides 64<br />

Tamarix ramosissima 64<br />

Tamarix ix<br />

Taverniera aegyptiaca 47<br />

Taverniera aegyptiaca 47<br />

Taverniera spartea 47<br />

Tetraena qatarensis viii, ix, 67, 88<br />

Tetraena simplex 68, 89<br />

Teucrium polium 51, 73<br />

TILIACEAE 64<br />

Torilis nodosa 5<br />

Trachynia distachya 73<br />

Traganum nudatum 31<br />

Trian<strong>the</strong>ma pentandra 3<br />

Tribulus arabicus 68<br />

Tribulus cistoides 68<br />

Tribulus pterophorus 68<br />

Tribulus terrestris 68<br />

Trifolium resupinatum 48<br />

Trigonella anguina 48<br />

94 95


Trigonella glabra 48<br />

Trigonella hamosa 48<br />

Trigonella monantha 48<br />

Trigonella stellata 48, 67<br />

Tripleurospermum auriculatum 14<br />

Typha domingensis 84<br />

TyPHACEAE 84<br />

UMBELLIFERAE x<br />

Urospermum picroides 14<br />

Urtica urens 66<br />

URTICACEAE 65<br />

Vaccaria hispanica 25<br />

Vaccaria pyramidata 25<br />

VERBENACEAE 66<br />

96<br />

Veronica cymbalaria 63<br />

Vicia monantha 49<br />

Vicoa pentanema 11<br />

xanthium spinosum 14<br />

Zaleya pentandra 3<br />

Zilla spinosa 21<br />

Ziziphus mauritiana 60<br />

Ziziphus nummularia viii, 61, 83<br />

Ziziphus spina-christi 61<br />

ZyGOPHyLLACEAE 66<br />

Zygophyllum coccineum 68<br />

Zygophyllum mandavillei 68<br />

Zygophyllum qatarense 67<br />

Zygophyllum simplex 68


About this Book<br />

The State <strong>of</strong> <strong>Qatar</strong>, situated in <strong>the</strong> Arabian Gulf, covers over 11,000 km 2 <strong>of</strong> land surface and has more<br />

than 600 km <strong>of</strong> coastline. It supports a wide variety <strong>of</strong> natural habitats including limestone desert, rocky<br />

escarpments, coastal saltmarshes, natural depressions filled with fertile sand and silt, gravel plains and sand<br />

dunes. Toge<strong>the</strong>r with various man-made habitats <strong>the</strong>se support a surprisingly rich and varied plant life. This<br />

checklist provides details <strong>of</strong> nearly 400 species <strong>of</strong> vascular plants that have been recorded in <strong>Qatar</strong>, including<br />

at least 270 that are native and about 125 that are introduced. English and Arabic vernacular names are given<br />

where known, toge<strong>the</strong>r with summary information on rarity status in <strong>Qatar</strong>, growth form, flowering period,<br />

habitat, distribution and uses.<br />

The checklist provides an up-to-date scientific reference for ecologists and o<strong>the</strong>rs carrying out research on<br />

<strong>Qatar</strong>’s habitats and vegetation. Two <strong>of</strong> <strong>the</strong> authors are local <strong>Qatar</strong>is, who have studied <strong>the</strong> flora as part <strong>of</strong> <strong>the</strong>ir<br />

academic careers. The checklist updates information published more than 25 years ago by previous botanists<br />

working in <strong>the</strong> country, notably Pr<strong>of</strong>essor Batanouny who published his Ecology and <strong>Flora</strong> <strong>of</strong> <strong>Qatar</strong> in 1981. The<br />

checklist will also be useful to local people and visitors interested in <strong>the</strong> flora <strong>of</strong> <strong>Qatar</strong>.<br />

In recent years many <strong>of</strong> <strong>Qatar</strong>’s natural habitats have come under threat from <strong>the</strong> inevitable increase in urban,<br />

industrial and tourism developments. Many non-native plants have been introduced with food, livestock and<br />

forestry imports and some are having detrimental effects on native species. Coastal ecosystems supporting<br />

mangroves, saltmarsh and seagrass beds are particularly under threat both in <strong>Qatar</strong> and o<strong>the</strong>r parts <strong>of</strong> <strong>the</strong><br />

Gulf. It is important, <strong>the</strong>refore, that both <strong>the</strong> native and naturalised flora are monitored to inform impact<br />

assessments, ecosystem management and conservation measures. In this regard <strong>the</strong> <strong>Qatar</strong> Foundation<br />

with support from UNESCO and Maersk Oil <strong>Qatar</strong> are setting up <strong>the</strong> Quranic Botanic Garden in <strong>Qatar</strong> to<br />

maintain a living collection <strong>of</strong> native plants for scientific and educational purposes. These organisations have<br />

generously supported <strong>the</strong> production <strong>of</strong> this checklist as an important element in biodiversity conservation<br />

and environmental education. It is hoped that <strong>the</strong> checklist will add to <strong>the</strong> developing knowledge-base <strong>of</strong><br />

plant taxonomy and vegetation ecology amongst <strong>Qatar</strong>i botanists and stimulate interest and understanding<br />

<strong>of</strong> <strong>the</strong> natural habitats and flora <strong>of</strong> <strong>Qatar</strong>.

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