Checklist of Beetles of the British Isles, 2012 edition

Checklist of 

Beetles of the British Isles 

2nd Edition 

Edited by A.G. Duff 

with an Introduction by D.A. Lott 

and a chapter on Fossil Beetles by P.I. Buckland & P.C. Buckland

Summary of changes in the 2012 edition 

This section details the main changes in the current edition of the checklist, compared 

to the previous edition (Duff, 2008). For further details of these changes, please refer 

to the endnotes. 

Family-group names: 

Nomenclature of family-group names follows Bouchard et al. (2011) 

Family PAELOBIIDAE is renamed to HYGROBIIDAE 

Family SCYDMAENIDAE becomes a subfamily of STAPHYLINIDAE 

Family BOLBOCERATIDAE becomes a subfamily of GEOTRUPIDAE 

Order of families in SCARABAEOIDEA changed 

Family ANOBIIDAE is renamed to PTINIDAE 

Family LANGURIIDAE deleted and included species moved to EROTYLIDAE 

Species added to the main list (47): 

Suphrodytes figuratus 

Somotrichus unifasciatus 

Hololepta plana 

Ochthebius nilssoni 

Catops borealis 

Brachygluta klimschi 

Mycetoporus ambiguus 

Mycetoporus reichei 

Acrotona convergens 

Gyrophaena transversalis 

Anotylus hammondi 

Bledius lohsei 

Carpelimus erichsoni 

Carpelimus zealandicus 

Carpelimus alutaceus 

Edaphus beszedesi 

Paederidus ruficollis 

Philonthus alpinus 

Quedius lyszkowskii 

Quedius lucidulus 

Gyrohypnus wagneri 

Agrilus cuprescens 

Agrilus cyanescens 

Trachys subglaber 

Species deleted from the main list (9): 

Hydaticus continentalis 

Abax parallelus 

Stenolophus comma 

Aglyptinus agathidioides 

Alevonota aurantiaca 

Bledius praetermissus 

Heterocerus hispidulus (duplication error) 

Protapion ryei 

Cathormiocerus maritimus 

1 

Isorhipis melasoides 

Gastrallus laevigatus 

Mesocoelopus collaris 

Mirosternomorphus heali 

Colotes punctatus 

Meligethes matronalis 

Meligethes symphyti 

Typhaea haagi 

Berginus tamarisci 

Semanotus russicus 

Poecilium lividum 

Leiopus linnei 

Bruchus brachialis 

Aphthona pallida 

Longitarsus symphyti 

Pseudoperapion 

brevirostre 

Bradybatus fallax 

Cotaster uncipes 

Allopentarthrum elumbe 

Conarthrus praeustus 

Otiorhynchus cribricollis 

Sitona gressorius 

Polygraphus grandiclava

Acknowledgements 

Contents 

I would like to thank everyone who has helped to create and maintain the Checklist of 

Beetles of the British Isles. During the early part of this project I was greatly assisted 

by draft checklist manuscripts written by Roger Booth and Eric Philp. For keeping track 

of new and deleted species, published updates to the British list were invaluable (Lott, 

1995, 1998; Mann, 2000b, 2002; Owen, 1993a, 1994c). Recent published checklists 

were also very helpful covering the Irish fauna (Anderson, Nash & O'Connor, 1997; 

Nash, Anderson & O'Connor, 1997), water beetles (Foster, 2004, 2005; Foster & 

Friday, 2011), Carabidae (Luff, 2007), Leiodidae (Cooter, 1996b), Staphylinidae part 

(Lott, 2009a; Lott & Anderson, 2011), Dermestidae (Peacock, 1993), Nitiduloidea 

(Cooter, 1995), Kateretidae and Nitidulidae (Kirk-Spriggs, 1996), Ciidae (Orledge & 

Booth, 2006; Orledge, 2009), Anthicidae (Telnov, 2010), Scraptiidae (Levey, 2009), 

seed and leaf beetles (Cox, 2007) and weevils (Morris, 2003, 2011). The late Derek 

Lott deserves a special mention, not only as the original section author for the very 

large family Staphylinidae, but also for writing his stimulating introduction to the 

project, and for his unstinting encouragement. With his encyclopaedic knowledge of 

the British fauna, and nomenclatural issues in particular, a great debt is owed to Roger 

Booth for undertaking to author many individual families and for answering numerous 

questions. All the family authors are to be warmly thanked: Keith Alexander 

(Cantharidae), Max Barclay (Tetratomidae and Tenebrionidae), Roger Booth 

(numerous families), Jon Cooter (Leiodidae), Mike Cox (Megalopodidae, 

Orsodacnidae and Chrysomelidae), Garth Foster (water beetles), Colin Johnson 

(Ptiliidae, Clambidae, Monotomidae, Cryptophagidae, Corylophidae and Latridiidae), 

Brian Levey (Buprestidae, Mordellidae and Scraptiidae), the late Derek Lott 

(Staphylinidae), Martin Luff (Carabidae), Darren Mann (Scarabaeoidea), Howard 

Mendel (Elateroidea), Mike Morris (Curculionoidea), Glenda Orledge (Ciidae) and 

Martin Rejzek (Cerambycidae). Many thanks also to Phil and Paul Buckland for their 

chapter on beetles known only as fossils from Quaternary sediments. 

Several coleopterists have reviewed all or part of the checklist. Tony (A.J.) Allen 

helped with literature, reviewed numerous drafts of the complete checklist and 

suggested several formatting improvements. Max Barclay reviewed the arrangement 

of Otiorhynchus. Martin Collier spotted several errors and suggested some format 

changes. Scotty Dodd spotted an important error in Altica. Peter Hammond is thanked 

for correcting several errors, for some formatting suggestions and for thoroughly 

reviewing the entire checklist. Mark Hill corrected some errors during the course of 

extracting data for the Biological Records Centre species dictionary. Peter Hodge 

assisted with literature and drew my attention to a change in Globicornis. Eric Philp 

spotted a duplication in Heteroceridae. Mark Telfer is to be thanked for his comments 

on various drafts of the checklist, and for maintaining the MapMate ® Coleoptera 

species dictionary. Finally, the Board of Governors of The Coleopterist journal are to 

be thanked for allowing the checklist to be hosted on the journal's website at 

www.coleopterist.org.uk. 

Andrew Duff 

Introduction ................................................................................................................ 3 

Extant or recently extinct native and naturalised species ...................................... 7 

Species found as fossils in Quaternary sediments ..............................................127 

Non-established introductions ...............................................................................131 

Bibliography ............................................................................................................136 

Index to family-group and genus-group names ...................................................157 

2

Introduction 

Scope 

Rationale 

D.A. Lott 

S 

3 

Introduction 

ince publication of the Coleoptera volume of the second edition of Kloet & Hincks' 

checklist of British insects (Pope, 1977), a large number of additions and deletions 

to the list have been published, as well as numerous nomenclatural changes. In 

addition, recent advances in our understanding of the phylogeny of Coleoptera have 

led to major modifications of higher classification in checklists and catalogues that 

have been published more recently in Europe and North America. In the absence of 

any widely accepted, modern checklist of the British fauna, British workers have been 

using binomial names taken from a variety of often conflicting sources. 

It is intended that this checklist should have a wider currency and be suitable for use in 

future conservation reviews, survey reports and taxonomic dictionaries for biological 

recording. It is, therefore, vital that the nomenclature used should be widely accepted 

by coleopterists working on the British fauna. It is also important that the checklist 

should be as useful as possible to those working at larger scales ranging from 

European through Palaearctic to the world fauna. It is envisaged that the checklist will 

be updated regularly to keep abreast of published changes and suggested 

amendments from correspondents. To this end, comments on the checklist are 

welcome and should be addressed to the Editor by e-mail to andrew.duff@virgin.net. 

The checklist aims to include all species that have been reliably recorded from the 

British Isles as possible residents. Exotic species which are only known from casual 

importation, and have never formed established populations, are listed in an appendix. 

The British Isles is here taken to include Great Britain and Ireland, including the Isle of 

Man but not the Channel Islands. 

The checklist aims to be comprehensive in its listing of subfamilies, genera and 

species. Tribes and subtribes are only listed when supported by a recent, authoritative 

review. Contemporary checklists vary in the extent to which they include subgenera 

and many subgeneric classifications are old and in need of review. Consequently, 

subgenera have only been listed when they enjoy current and widespread usage. 

It is considered that there are two main uses for a checklist: 

to provide current standard names for species, genera, families and any other 

levels that are in common usage 

to enable other commonly used names to be interpreted as current standard 

names. 

It is also recognised that checklists have several subsidiary functions including: 

an inventory of biodiversity in a particular region 

the ordering of names into a standard sequence. 

Three areas are considered to be outside this checklist's remit. Firstly, the exhaustive 

listing of old synonyms is more properly the job of a catalogue rather than a simple 

checklist. Secondly, new names and new approaches to classification should appear 

in comprehensive taxonomic reviews rather than checklists. Thirdly, the correction of

Introduction Checklist of Beetles of the British Isles, 2012 Edition 

errors is not the primary function of a checklist, although correctness will be an 

important factor in evaluating any recently proposed changes to classification or 

nomenclature; misspelt names which have become widely used, for example, are 

listed here in order to draw attention to the correct usage. 

As far as possible, therefore, this checklist uses a classification that already exists in 

the current literature, although the opportunity has been taken, through the 

cooperation of several correspondents, to include some changes in advance of 

publication, in order to ensure future currency. There is, of course, no universally 

agreed classification because of differences in approach between regions and 

between individual workers. Inevitably, choices between conflicting practices have had 

to be made. The main objective in making these choices has been to construct a list of 

binomial names, whose content and arrangement will be broadly acceptable, or at 

least comprehensible, for the next ten to twenty years. In order to meet this aspiration, 

it has been necessary to address four factors that work against the stability of 

checklists fossilised at one moment in time. They are: 

Additions and deletions of species 

These changes are essentially unpredictable and will be communicated in future 

editions as they arise. 

Nomenclatural changes 

Replacement names for British taxa are frequently published in various notes, 

taxonomic reviews and foreign checklists. These changes arise from new synonymies, 

discoveries of senior homonyms and corrections of misidentifications. Corrections of 

misidentifications can result in the transfer of a misapplied name from one species to 

another. For example, Fauvel (1862) misidentified Aleochara punctatella Motschulsky 

as A. obscurella Gravenhorst and renamed the true A.obscurella Gravenhorst as A. 

algarum Fauvel. This error persisted for over 100 years until its discovery (Lohse, 

1985b) led to the transfer of the name A. obscurella Gravenhorst back to the original 

species. Consequently the name A. obscurella Gravenhorst has been applied to two 

different species at different times. It is very easy to misinterpret such names in old 

publications. 

The older the published list, the more difficult becomes the task of interpreting names. 

This checklist aims to provide the means for interpreting names used since around 

1950 when Joy (1932) was the standard British identification guide. It must be 

recognised that the interpretation of names in older publications and manuscripts 

requires specialist knowledge that cannot be substituted by a simple checklist. Indeed 

the same can be said for any work dealing with taxonomically confused groups or 

using outdated reference sources. 

Sometimes nomenclatural changes are contentious and lead to inconsistencies 

between different checklists. Occasionally, they prove to be wrong and are followed by 

reversions to previous usage in subsequent editions of the checklist. Unfortunately, 

the adoption of inappropriate type species for genera, the unearthing of ancient 

misidentifications and homonymies, and the reinstitution of old, long forgotten names, 

have too often led to the overturning of long-established nomenclature. Such changes 

are a regrettable cause of instability in taxonomic nomenclature and they are best 

approached in the context of a comprehensive taxonomic review. It is the duty of a 

checklist compiler to be cautious in this regard. It is better to err on the side of 

established usage, rather than indulge in taxonomic sorties that cause unwarranted 

instability. 

It is to be hoped that recent changes in the ICZN code (ICZN, 1999) will prevent some 

of the worst excesses of past practice, by conserving widely used names under threat 

4

Format 

5 

Introduction 

from disused 18th- and 19th-Century names. However, some well-established names 

of British taxa are known to be incorrect, or at least in need of investigation. Some of 

these names may well be the subject of future applications to the ICZN for their 

conservation. It is not intended here to initiate any radical changes that have not been 

instituted elsewhere. 

Changes in the boundaries of higher taxa 

The large volume of work on higher classification in recent decades has led to major 

advances in phylogeny and revolutionised our understanding of the relationships of 

taxa from family level down to species level. For example, the family Staphylinidae has 

been expanded to include several new subfamilies that were formerly families in their 

own right, whereas the Curculionidae has been broken up into several smaller 

families. 

Some of these changes have arisen following cladistic analyses, which suggest that 

taxa are paraphyletic. Others appear to be based on more informal arguments. In 

either case, the ranking of a higher taxon, such as a genus, is largely based on 

opinion, because there is no universally accepted definition of a genus. Usually, the 

scale of morphological differences dictates the separation of genera, but numbers of 

species have also been used as an argument (see e.g. Seevers, 1978). It is therefore 

no surprise that the rank of a particular name as genus, subgenus or synonym has 

sometimes proved to be contentious and this has led to regional differences and to 

unstable binomial names. Inevitably, some judgment has to be made about which of 

two conflicting approaches is going to gain more currency, but, in general, this 

checklist follows widely accepted world and regional catalogues and subsequent 

authoritative taxonomic revisions. 

Changes in the sequence of taxa in the list 

Taxa are often ordered within checklists so that they are placed next to their closest 

relatives and so phylogenetic relationships provide the basis for a standard 

sequencing. Traditionally, basal groups precede more specialised taxa. Several 

existing checklists arrange taxa alphabetically rather than suggest phylogenetic 

relationships that are not backed up by a proper investigation. Pope (1977) ordered 

species alphabetically and ordered higher taxa so as to reflect relationships, or at least 

traditional ideas of relationships, and this general principle has been continued in this 

list except that junior synonyms are listed in ascending date order rather than 

alphabetically. It is expected that phylogenetic relationships will be of minor 

consequence to many checklist users. Nevertheless, advances in knowledge in this 

area have the capacity to disrupt the stability of any adopted standard arrangement of 

taxa. It should be noted that binomial names remain unaffected, whatever the 

arrangement of taxa. 

The sequence of taxa in this checklist generally follows arrangements in other 

checklists that will be familiar to a wide range of users. 

The checklist is divided into three sections in order to list separately: 

extant or recently extinct (i.e. within historical times) native and naturalised 

species, including natural immigrants/vagrants and established (either 

temporarily or permanently) introductions 

species that are known only from the Quaternary fossil record 

non-established introductions.

Introduction Checklist of Beetles of the British Isles, 2012 Edition 

Citation 

The checklists are followed by a bibliography, and then an index to genus-group 

names (names of genera and subgenera, including their synonyms) and family-group 

names (for all categories higher than genus). 

Within the main checklist, conventional Latin phrases are used to qualify certain 

names. Although written in a largely forgotten language, these phrases have the 

considerable merit of being very concise: 

ad until 

ante before 

auctt. authors (i.e. in entomological literature generally) 

auctt. Brit. British authors 

auctt. Europ. European authors 

nec nor of 

non but not of 

partes maj. for the most part 

partim partly 

sensu in the sense of 

sensu lato in the broad sense 

sensu stricto in the narrow sense 

Question marks are used to indicate uncertainty. A question mark before a name 

indicates unsure identification or placement as a synonym. A question mark before a 

citation indicates doubt over whether the author used the name in the sense shown. 

Endnotes are generally given for significant changes to the British list since Pope 

(1977), unless these have already been detailed in the family introduction. In addition, 

endnotes may draw attention to particular nomenclatural or taxonomic points. 

Relevant Opinions of the International Commission for Zoological Nomenclature are 

cited as "ICZN Op." in endnotes (see www.iczn.org/Opinions.htm). 

Citation should follow this example format: 

MANN, D.J. 2012. Scarabaeidae. In: Duff, A.G. (ed.) Checklist of Beetles of the British 

Isles. 2nd edition. Pemberley Books. 

6

7 

Extant or recently extinct native and naturalised species: contents 

Extant or recently extinct native and naturalised species 

Order COLEOPTERA Linnaeus, 1758 ..................................................................... 10 

Suborder MYXOPHAGA Crowson, 1955 ................................................................. 10 

1. Family SPHAERIUSIDAE Erichson, 1845 ................................................ 10 

Suborder ADEPHAGA Clairville, 1806 ..................................................................... 10 

2. Family GYRINIDAE Latreille, 1810 ............................................................ 10 

3. Family HALIPLIDAE Aubé, 1836 ............................................................... 10 

4. Family NOTERIDAE Thomson, C.G., 1860 .............................................. 10 

5. Family HYGROBIIDAE Régimbart, 1878 (1837) ...................................... 11 

6. Family DYTISCIDAE Leach, 1815 ............................................................ 11 

7. Family CARABIDAE Latreille, 1802 ........................................................... 13 

Suborder POLYPHAGA Emery, 1886 ...................................................................... 23 

Superfamily HYDROPHILOIDEA Latreille, 1802 ...................................................... 23 

8. Family HELOPHORIDAE Leach, 1815 ..................................................... 23 

9. Family GEORISSIDAE Laporte, 1840 ....................................................... 24 

10. Family HYDROCHIDAE Thomson, C.G., 1859 ...................................... 24 

11. Family SPERCHEIDAE Erichson, 1837 .................................................. 24 

12. Family HYDROPHILIDAE Latreille, 1802 ............................................... 24 

13. Family SPHAERITIDAE Shuckard, 1839 ................................................. 26 

14. Family HISTERIDAE Gyllenhal, 1808 ..................................................... 26 

Superfamily STAPHYLINOIDEA Latreille, 1802 ....................................................... 27 

15. Family HYDRAENIDAE Mulsant, 1844 .................................................... 27 

16. Family PTILIIDAE Erichson, 1845 ............................................................ 28 

17. Family LEIODIDAE Fleming, 1821 ........................................................... 31 

18. Family SILPHIDAE Latreille, 1806 ........................................................... 33 

19. Family STAPHYLINIDAE Latreille, 1802 ................................................. 33 

Superfamily SCARABAEOIDEA Latreille, 1802 ....................................................... 55 

20. Family GEOTRUPIDAE Latreille, 1802 ................................................... 55 

21. Family TROGIDAE MacLeay, 1819 ........................................................ 55 

22. Family LUCANIDAE Latreille, 1804 ........................................................ 55 

23. Family SCARABAEIDAE Latreille, 1802 ................................................. 55 

Superfamily SCIRTOIDEA Fleming, 1821 ................................................................ 58 

24. Family EUCINETIDAE Lacordaire, 1857 ................................................. 58 

25. Family CLAMBIDAE Fischer von Waldheim, 1821 .................................. 58 

26. Family SCIRTIDAE Fleming, 1821 ........................................................... 58 

Superfamily DASCILLOIDEA Guérin-Méneville, 1843 (1834) ................................. 59 

27. Family DASCILLIDAE Guérin-Méneville, 1843 (1834) ............................ 59 

Superfamily BUPRESTOIDEA Leach, 1815 ............................................................ 59 

28. Family BUPRESTIDAE Leach, 1815 ...................................................... 59 

Superfamily BYRRHOIDEA Latreille, 1804 .............................................................. 60 

29. Family BYRRHIDAE Latreille, 1804 ......................................................... 60 

30. Family ELMIDAE Curtis, 1830 ................................................................. 60 

31. Family DRYOPIDAE Billberg, 1820 (1817) .............................................. 60 

32. Family LIMNICHIDAE Erichson, 1846 ..................................................... 61 

33. Family HETEROCERIDAE MacLeay, 1825 ............................................. 61 

34. Family PSEPHENIDAE Lacordaire, 1854 ................................................ 61 

35. Family PTILODACTYLIDAE Laporte, 1836 ............................................. 61 

Superfamily ELATEROIDEA Leach, 1815 ............................................................... 61 

36. Family EUCNEMIDAE Eschscholtz, 1829 ............................................... 61 

37. Family THROSCIDAE Laporte, 1840 ....................................................... 61

Extant or recently extinct native and naturalised species: contents Checklist of Beetles of the British Isles, 2012 Edition 

38. Family ELATERIDAE Leach, 1815 .......................................................... 62 

39. Family DRILIDAE Blanchard, 1845 .......................................................... 64 

40. Family LYCIDAE Laporte, 1836 ............................................................... 64 

41. Family LAMPYRIDAE Rafinesque, 1815 ................................................. 64 

42. Family CANTHARIDAE Imhoff, 1856 (1815) ........................................... 64 

Superfamily DERODONTOIDEA LeConte, 1861 ..................................................... 65 

43. Family DERODONTIDAE LeConte, 1861 ................................................ 65 

Superfamily BOSTRICHOIDEA Latreille, 1802 ........................................................ 65 

44. Family DERMESTIDAE Latreille, 1804 ................................................... 65 

45. Family BOSTRICHIDAE Latreille, 1802 ................................................... 66 

46. Family PTINIDAE Latreille, 1802 ............................................................. 67 

Superfamily LYMEXYLOIDEA Fleming, 1821 .......................................................... 68 

47. Family LYMEXYLIDAE Fleming, 1821 ..................................................... 68 

Superfamily CLEROIDEA Latreille, 1802 ................................................................. 68 

48. Family PHLOIOPHILIDAE Kiesenwetter, 1863 ........................................ 68 

49. Family TROGOSSITIDAE Latreille, 1802 ................................................ 68 

50. Family CLERIDAE Latreille, 1802 ........................................................... 69 

51. Family DASYTIDAE Laporte, 1840 ......................................................... 69 

52. Family MALACHIIDAE Fleming, 1821 .................................................... 69 

Superfamily CUCUJOIDEA Latreille, 1802 .............................................................. 70 

53. Family SPHINDIDAE Jacquelin du Val, 1860 .......................................... 70 

54. Family KATERETIDAE Kirby, 1837 ........................................................ 70 

55. Family NITIDULIDAE Latreille, 1802 ........................................................ 71 

56. Family MONOTOMIDAE Laporte, 1840 .................................................. 72 

57. Family SILVANIDAE Kirby, 1837 ............................................................. 73 

58. Family CUCUJIDAE Latreille, 1802 ......................................................... 73 

59. Family LAEMOPHLOEIDAE Ganglbauer, 1899 ...................................... 73 

60. Family PHALACRIDAE Leach, 1815 ....................................................... 74 

61. Family CRYPTOPHAGIDAE Kirby, 1826 ................................................ 74 

62. Family EROTYLIDAE Latreille, 1802 ....................................................... 76 

63. Family BYTURIDAE Gistel, 1848 ............................................................. 76 

64. Family BIPHYLLIDAE LeConte, 1861 ...................................................... 77 

65. Family BOTHRIDERIDAE Erichson, 1845 ............................................... 77 

66. Family CERYLONIDAE Billberg, 1820 ..................................................... 77 

67. Family ALEXIIDAE Imhoff, 1856 .............................................................. 77 

68. Family ENDOMYCHIDAE Leach, 1815 .................................................. 77 

69. Family COCCINELLIDAE Latreille, 1807 ................................................. 77 

70. Family CORYLOPHIDAE LeConte, 1852 ............................................... 79 

71. Family LATRIDIIDAE Erichson, 1842 ..................................................... 79 

Superfamily TENEBRIONOIDEA Latreille, 1802 ...................................................... 81 

72. Family MYCETOPHAGIDAE Leach, 1815 ............................................... 81 

73. Family CIIDAE Leach, 1819 ..................................................................... 81 

74. Family TETRATOMIDAE Billberg, 1820 .................................................. 82 

75. Family MELANDRYIDAE Leach, 1815 .................................................... 82 

76. Family MORDELLIDAE Latreille, 1802 .................................................... 83 

77. Family RIPIPHORIDAE Gemminger, 1870 (1855) .................................. 83 

78. Family COLYDIIDAE Billberg, 1820 ......................................................... 83 

79. Family TENEBRIONIDAE Latreille, 1802 ................................................ 84 

80. Family OEDEMERIDAE Latreille, 1810 ................................................... 86 

81. Family MELOIDAE Gyllenhal, 1810 ......................................................... 86 

82. Family MYCTERIDAE Oken, 1843 .......................................................... 86 

83. Family PYTHIDAE Solier, 1834 ............................................................... 86 

8

9 

Extant or recently extinct native and naturalised species: contents 

84. Family PYROCHROIDAE Latreille, 1806 ................................................. 86 

85. Family SALPINGIDAE Leach, 1815 ......................................................... 86 

86. Family ANTHICIDAE Latreille, 1819 ....................................................... 87 

87. Family ADERIDAE Csiki, 1909 ................................................................ 87 

88. Family SCRAPTIIDAE Gistel, 1848 ......................................................... 87 

Superfamily CHRYSOMELOIDEA Latreille, 1802 .................................................... 88 

89. Family CERAMBYCIDAE Latreille, 1802 ................................................ 88 

90. Family MEGALOPODIDAE Latreille, 1802 .............................................. 90 

91. Family ORSODACNIDAE Thomson, C.G., 1859 ..................................... 91 

92. Family CHRYSOMELIDAE Latreille, 1802 ............................................... 91 

Superfamily CURCULIONOIDEA Latreille, 1802 .................................................... 96 

Division ORTHOCERI Schönherr, 1823 .................................................................. 96 

93. Family NEMONYCHIDAE Bedel, 1882 ................................................... 96 

94. Family ANTHRIBIDAE Billberg, 1820 ..................................................... 96 

95. Family RHYNCHITIDAE Gistel, 1848 ...................................................... 96 

96. Family ATTELABIDAE Billberg, 1820 ...................................................... 97 

97. Family APIONIDAE Schönherr, 1823 ..................................................... 97 

98. Family NANOPHYIDAE Gistel, 1848 ..................................................... 100 

99. Family DRYOPHTHORIDAE Schönherr, 1825 ...................................... 100 

100. Family ERIRHINIDAE Schönherr, 1825 .............................................. 100 

101. Family RAYMONDIONYMIDAE Reitter, 1913 .................................... 100 

Division GONATOCERI Schönherr, 1825 .............................................................. 100 

102. Family CURCULIONIDAE Latreille, 1802 ............................................ 100 

103. Family PLATYPODIDAE Shuckard, 1839 ........................................... 112 

Endnotes ...................................................................................................... 113

Family SPHAERIUSIDAE Erichson, 1845 Checklist of Beetles of the British Isles, 2012 Edition 

Order COLEOPTERA Linnaeus, 1758 

Suborder MYXOPHAGA Crowson, 1955 

Superfamily SPHAERIUSOIDEA Erichson, 1845 

1. Family SPHAERIUSIDAE Erichson, 1845 1 

MICROSPORIDAE Crotch, 1873 

Family author: G.N. Foster 

SPHAERIUS Waltl, 1838 2 

MICROSPORUS Kolenati, 1846 

acaroides Waltl, 1838 

obsidianus (Kolenati, 1846) 

Suborder ADEPHAGA Clairville, 1806 

2. Family GYRINIDAE Latreille, 1810 


Subfamily GYRININAE Latreille, 1810 3 

Tribe GYRININI Latreille, 1810 

GYRINUS Geoffroy, 1762 4 

Subgenus GYRINULUS Zaitzev, 1908 

minutus Fabricius, 1798 

Subgenus GYRINUS Geoffroy, 1762 

aeratus Stephens, 1835 

opacus sensu auctt. Brit. ad 1914 non Sahlberg, 

C.R., 1819 

thomsoni Zaitzev, 1907 

edwardsi Sharp, 1914 

caspius Ménétries, 1832 

elongatus Aubé, 1838 non Marsham, 1802 

distinctus Aubé, 1838 

colymbus sensu auctt. Brit. non Erichson, 1837 

marinus Gyllenhal, 1808 

natator (Linnaeus, 1758) 

mergus Ahrens, 1812 

opacus Sahlberg, C.R., 1819 

lecontei Omer-Cooper, 1930 non Fall, 1921 

blairi Omer-Cooper, 1931 

paykulli Ochs, 1927 

bicolor sensu Paykull, 1798 non Fabricius, 1787 

substriatus Stephens, 1828 

natator sensu auctt. Brit. partes maj. non (Linnaeus, 

1758) 

oblitus Sharp, 1914 

fowleri Omer-Cooper, 1930 

suffriani Scriba, 1855 

urinator Illiger, 1807 

Tribe ORECTOCHILINI Régimbart, 1882 

ORECTOCHILUS Dejean, 1833 

villosus (Müller, O.F., 1776) 

10 

3. Family HALIPLIDAE Aubé, 1836 


BRYCHIUS Thomson, C.G., 1859 

elevatus (Panzer, 1793) 

PELTODYTES Régimbart, 1879 

CNEMIDOTUS sensu Erichson, 1832 non Illiger, 

1802 

caesus (Duftschmid, 1805) 

impressus sensu (Panzer, 1794) non (Fabricius, 

1787) 

HALIPLUS Latreille, 1802 5 

Subgenus HALIPLIDIUS Guignot, 1928 

confinis Stephens, 1828 

pallens Fowler, 1887 

halberti Bullock, 1928 

obliquus (Fabricius, 1787) 

varius Nicolai, 1822 6 

Subgenus NEOHALIPLUS Netolitzky, 1911 

lineatocollis (Marsham, 1802) 

Subgenus HALIPLUS Latreille, 1802 

apicalis Thomson, C.G., 1868 

striatus Sharp, 1869 

fluviatilis Aubé, 1836 

furcatus Seidlitz, 1887 

heydeni Wehncke, 1875 

ruficollis sensu auctt. Brit. partim non (De Geer, 

1774) 

immaculatus Gerhardt, 1877 

lineolatus Mannerheim, 1844 

nomax Balfour-Browne, F., 1911 

browneanus Sharp, 1913 

ruficollis (De Geer, 1774) 

fulvicollis sensu Edwards, 1911 non Erichson, 1837 

sibiricus Motschulsky, 1860 

borealis Gerhardt, 1877 non LeConte, 1850 

wehnckei Gerhardt, 1877 7 

Subgenus LIAPHLUS Guignot, 1928 

flavicollis Sturm, 1834 

fulvus (Fabricius, 1801) 

laminatus (Schaller, 1783) 

cinereus Aubé, 1836 

mucronatus Stephens, 1828 

variegatus Sturm, 1834 

4. Family NOTERIDAE Thomson, C.G., 1860 8 


NOTERUS Clairville, 1806 

clavicornis (De Geer, 1774) 

capricornis sensu auctt. Brit. non (Herbst, 1784) 

sparsus (Marsham, 1802) 

crassicornis (Müller, O.F., 1776) 

capricornis (Herbst, 1784) 

clavicornis sensu auctt. Brit. partim non (De Geer, 

1774)

5. Family HYGROBIIDAE Régimbart, 1878 

(1837) 9 

PAELOBIIDAE Erichson, 1837 


HYGROBIA Latreille, 1804 

HYGRIOBIA Latreille, 1804 10 

PAELOBIUS Schönherr, 1808 

PELOBIUS Erichson, 1832 (misspelling) 

hermanni (Fabricius, 1775) 

herrmanni (Fabricius, 1775) 10 

tarda (Herbst, 1779) 

6. Family DYTISCIDAE Leach, 1815 11 


Subfamily AGABINAE Thomson, C.G., 1867 

AGABUS Leach, 1817 

Subgenus ACATODES Thomson, C.G., 1859 

arcticus (Paykull, 1798) 

congener (Thunberg, 1794) 

sturmii (Gyllenhal in Schönherr, 1808) 

sturmi auctt. (misspelling) 

Subgenus AGABUS Leach, 1817 

labiatus (Brahm, 1790) 

femoralis (Paykull, 1798) 

uliginosus (Linnaeus, 1761) 

dispar Bold, 1849 

undulatus (Schrank, 1776) 

abbreviatus (Fabricius, 1787) 

Subgenus GAURODYTES Thomson, C.G., 1859 

affinis (Paykull, 1798) 

biguttatus (Olivier, 1795) 12 

bipustulatus (Linnaeus, 1767) 

solieri Aubé, 1837 

brunneus (Fabricius, 1798) 

conspersus (Marsham, 1802) 

didymus (Olivier, 1795) 

guttatus (Paykull, 1798) 

melanarius Aubé, 1837 

nebulosus (Forster, 1771) 

paludosus (Fabricius, 1801) 

striolatus (Gyllenhal, 1808) 

unguicularis (Thomson, C.G., 1867) 

ILYBIUS Erichson, 1832 

aenescens Thomson, C.G., 1870 

angustior sensu auctt. non (Gyllenhal, 1808) 

ater (De Geer, 1774) 13 

chalconatus (Panzer, 1796) 14 

fenestratus (Fabricius, 1781) 

fuliginosus (Fabricius, 1792) 

guttiger (Gyllenhal, 1808) 

montanus (Stephens, 1828) 14 

melanocornis (Zimmermann, 1915) 

quadriguttatus (Lacordaire, 1835) 

obscurus (Marsham, 1802) non (Panzer, 1795) 

subaeneus Erichson, 1837 

14 15 

wasastjernae (Sahlberg, C.R., 1824) 

11 

Family HYGROBIIDAE Régimbart, 1878 (1837) 

PLATAMBUS Thomson, C.G., 1859 

maculatus (Linnaeus, 1758) 

pulchellus (Heer, 1839) 

immaculatus Donisthorpe, 1899 

Subfamily COLYMBETINAE Erichson, 1837 

COLYMBETES Clairville, 1806 

fuscus (Linnaeus, 1758) 

RHANTUS Dejean, 1833 

RANTUS Dejean, 1833 16 

Subgenus NARTUS Zaitzev, 1907 

grapii (Gyllenhal, 1808) 

Subgenus RHANTUS Dejean, 1833 

bistriatus (Bergsträsser, 1778) 17 

adspersus (Fabricius, 1801) non (Panzer, 1796) 

aberratus Gemminger & Harold, 1868 

exsoletus (Forster, 1771) 

frontalis (Marsham, 1802) 

notatus sensu (Fabricius, 1781) non (Bergsträsser, 

1778) 

suturalis (MacLeay, 1825) 

punctatus (Geoffroy, 1785) non (Scopoli, 1763) 

pulverosus (Stephens, 1828) 

suturellus (Harris, 1828) 

bistriatus sensu auctt. Brit. non (Bergsträsser, 1778) 

Subfamily COPELATINAE Branden, 1885 

LIOPTERUS Dejean, 1833 18 

COPELATUS sensu auctt. non Erichson, 1832 

haemorrhoidalis (Fabricius, 1787) 

ruficollis (Schaller, 1783) non (De Geer, 1774) 

schalleri (Gmelin in Linnaeus, 1790) 

agilis (Fabricius, 1792) 

oblongus (Illiger, 1801) non (Herbst, 1784) 

Subfamily DYTISCINAE Leach, 1815 

Tribe ACILIINI Thomson, C.G., 1867 

ACILIUS Leach, 1817 

canaliculatus (Nicolai, 1822) 

fasciatus sensu Fowler, 1887 non (De Geer, 1774) 

sulcatus (Linnaeus, 1758) 

scoticus Stephens, 1828 

GRAPHODERUS Dejean, 1833 

bilineatus (De Geer, 1774) 

cinereus (Linnaeus, 1758) 

zonatus (Hoppe, 1795) 

Tribe CYBISTRINI Sharp, 1880 19 

CYBISTER Curtis, 1827 

lateralimarginalis (De Geer, 1774)

Family DYTISCIDAE Leach, 1815 Checklist of Beetles of the British Isles, 2012 Edition 

Tribe DYTISCINI Leach, 1815 

DYTISCUS Linnaeus, 1758 

circumcinctus Ahrens, 1811 

circumflexus Fabricius, 1801 

dimidiatus Bergsträsser, 1778 

lapponicus Gyllenhal, 1808 

marginalis Linnaeus, 1758 

semisulcatus Müller, O.F., 1776 

punctulatus Fabricius, 1777 

Tribe HYDATICINI Sharp, 1880 

HYDATICUS Leach, 1817 20 

seminiger (De Geer, 1774) 

transversalis (Pontoppidan, 1763) 

Subfamily HYDROPORINAE Aubé, 1836 

Tribe BIDESSINI Sharp, 1880 

BIDESSUS Sharp, 1882 

minutissimus (Germar, 1824) 

unistriatus (Goeze, 1777) 

HYDROGLYPHUS Motschulsky, 1853 

GUIGNOTUS Houlbert, 1934 

geminus (Fabricius, 1792) 

pusillus (Fabricius, 1781) non (Müller, O.F., 1776) 

Tribe HYDROPORINI Aubé, 1836 

DERONECTES Sharp, 1882 

latus (Stephens, 1829) 

GRAPTODYTES Seidlitz, 1887 

bilineatus (Sturm, 1835) 

hopffgarteni (Schilsky, 1892) 

flavipes (Olivier, 1795) 

concinnus (Stephens, 1835) 

granularis (Linnaeus, 1767) 

pictus (Fabricius, 1787) 

HYDROPORUS Clairville, 1806 21 

STERNOPORUS Falkenström, 1930 

HYDROPORIDIUS Guignot, 1949 

angustatus Sturm, 1835 

discretus Fairmaire & Brisout de Barneville, 1859 22 

neuter Fairmaire & Laboulbène, 1854 

elongatulus Sturm, 1835 

erythrocephalus (Linnaeus, 1758) 

ferrugineus Stephens, 1829 

glabriusculus Aubé, 1838 

gyllenhalii Schiødte, 1841 

gyllenhali auctt. (misspelling) 

piceus sensu auctt. Europ. non Stephens, 1828 

incognitus Sharp, 1869 

longicornis Sharp, 1871 

parallelus Sharp, 1869 non Aubé, 1838 

longulus Mulsant & Rey, 1861 

celatus Clark, 1862 

marginatus (Duftschmid, 1805) 

melanarius Sturm, 1835 

12 

monticola Sharp, 1869 

memnonius Nicolai, 1822 

morio Aubé, 1838 

melanocephalus sensu (Gyllenhal, 1808) non 

(Marsham, 1802) 

necopinatus Fery, 1999 23 

ssp. roni Fery, 1999 

cantabricus sensu auctt. Brit. non Sharp, 1882 

neglectus Schaum, 1845 

nigrita (Fabricius, 1792) 

obscurus Sturm, 1835 

obsoletus Aubé, 1838 

palustris (Linnaeus, 1761) 

tinctus Clark, 1862 

planus (Fabricius, 1782) 

ater (Forster, 1771) 24 

pubescens (Gyllenhal, 1808) 

rufifrons (Müller, O.F., 1776) 

rufifrons (Duftschmid, 1805) non (Müller, O.F., 

1776) 

piceus Stephens, 1828 

duftschmidti Rye, 1872 

scalesianus Stephens, 1828 

striola (Gyllenhal, 1826) 

vittula Erichson, 1837 

tessellatus (Drapiez, 1819) 

lituratus sensu Fowler, 1887 non (Fabricius, 1781) 

tristis (Paykull, 1798) 

umbrosus (Gyllenhal, 1808) 

NEBRIOPORUS Régimbart, 1906 25 

POTAMODYTES Zimmermann, 1919 non 

Grouvelle, 1896 

POTAMONECTES Zimmermann, 1921 

Subgenus NEBRIOPORUS Régimbart, 1906 

assimilis (Paykull, 1798) 

depressus (Fabricius, 1775) 

elegans (Panzer, 1794) 

Subgenus ZIMMERMANNIUS Guignot, 1941 

canaliculatus (Lacordaire, 1835) 26 

OREODYTES Seidlitz, 1887 

alpinus (Paykull, 1798) 27 

davisii (Curtis, 1831) 

davisi auctt. (misspelling) 

borealis sensu auctt. non (Gyllenhal, 1826) 

sanmarkii (Sahlberg, C.R., 1826) 

sanmarki auctt. (misspelling) 

rivalis (Gyllenhal, 1827) 

septentrionalis (Gyllenhal, 1826) 

PORHYDRUS Guignot, 1945 

lineatus (Fabricius, 1775) 

SCARODYTES des Gozis, 1914 

halensis (Fabricius, 1787) 

STICTONECTES Brinck, 1943 

STICTONOTUS Zimmermann, 1930 non Foerster, 

1856 

lepidus (Olivier, 1795) 

STICTOTARSUS Zimmermann, 1919 

duodecimpustulatus (Fabricius, 1792)

BOREONECTES Angus, 2010 

multilineatus (Falkenström, 1922) 

griseostriatus sensu auctt. non (De Geer, 1774) 

SUPHRODYTES des Gozis, 1914 

dorsalis (Fabricius, 1787) 

figuratus (Gyllenhal, 1826) 28 

Tribe HYDROVATINI Sharp, 1880 

HYDROVATUS Motschulsky, 1853 

OXYNOPTILUS Schaum in Schaum & 

Kiesenwetter, 1868 

clypealis Sharp, 1876 

cuspidatus (Kunze, 1818) 29 

Tribe HYGROTINI Portevin, 1929 

HYGROTUS Stephens, 1828 

Subgenus COELAMBUS Thomson, C.G., 1860 

confluens (Fabricius, 1787) 

impressopunctatus (Schaller, 1783) 

nigrolineatus (Steven, 1808) 30 

lautus (Schaum, 1843) 

novemlineatus (Stephens, 1829) 

parallellogrammus (Ahrens, 1812) 

parallelogrammus auctt. (misspelling) 

Subgenus HYGROTUS Stephens, 1828 

decoratus (Gyllenhal, 1810) 

inaequalis (Fabricius, 1777) 

quinquelineatus (Zetterstedt, 1828) 

versicolor (Schaller, 1783) 

Tribe HYPHYDRINI Gistel, 1848 

HYPHYDRUS Illiger, 1802 

ovatus (Linnaeus, 1761) 

Tribe LACCORNINI Wolfe & Roughley, 1990 

LACCORNIS des Gozis, 1914 

AGAPORUS Zimmermann, 1919 

oblongus (Stephens, 1835) 

Subfamily LACCOPHILINAE Gistel, 1848 

LACCOPHILUS Leach, 1815 

hyalinus (De Geer, 1774) 

interruptus (Panzer, 1795) 

minutus (Linnaeus, 1758) 

variolosus (Herbst, 1784) 

obscurus (Panzer, 1795) 

poecilus Klug, 1834 

variegatus (Germar & Kaulfuss, 1816) non 

(Fourcroy, 1785) 

obsoletus sensu auctt. non Westhoff, 1881 

ponticus Sharp, 1882 

13 

Family CARABIDAE Latreille, 1802 

7. Family CARABIDAE Latreille, 1802 

Family author: M.L. Luff 

The currently used nomenclature on the British 

Carabidae is based on that in Lindroth (1974) and 

Pope (1977); see also Luff (2007). More recent 

lists that have updated the nomenclature of some 

or all of the British fauna include Lindroth (1985, 

Fennoscandia), Lohse & Lucht (1989, central 

Europe), Turin (1990, 2000, Netherlands), 

Kryzhanovskij et al. (1995, Russia), Hansen (1996, 

Denmark), Hůrka (1996, Czech & Slovak 

republics) and Anderson et al. (1997, 2000, 

Ireland). These have now all been superseded by 

the Palaearctic list edited by Löbl & Smetana 

(2003). There have also been specialised lists for 

the genus Carabus, sensu lato (Turin et al., 1993; 

Deuve, 1994; Casale & Březina, 2003). The most 

recent list of central European Carabidae (Müller- 

Motzfeld, 2004) does not include synonyms. The 

following list is based on that of Löbl & Smetana 

(2003), with only a few more recent amendments 

provided by Dr Roger Booth of the Natural History 

Museum, London. 

The limits of the family Carabidae have remained 

constant in regard to the British fauna, since the 

Cicindelidae were included (as subfamily 

Cicindelinae) by Lindroth (1974). On the larger 

scale of the European or world fauna, however, 

there have been changes that do not affect the 

more limited British list. In particular, Erwin & Sims 

(1984) and Erwin (1985) recognised the separate 

family Trachypachidae, but included the former 

families Paussidae and Rhysodidae within the 

Carabidae. Lawrence & Newton (1995) followed 

the first two of these changes, but retained the 

Rhysodidae as a separate family. 

There is also no general agreement as to the 

arrangement and status of supra-generic taxa 

within the Carabidae. In an introduction to the 

classification of Caraboidea, Ball et al. (1998) 

compare many of the various arrangements that 

have been advocated. They conclude that points 

of conflict far outnumber any generally held areas 

of agreement. The present list is based on the 

relatively simple system used by Müller-Motzfeld 

(2004) which divided the family into only four 

subfamilies, with the great majority of British 

species in the Carabinae. This is in contrast to the 

classification of Lawrence & Newton (1985), who 

had many subfamilies, and consider the broscines 

to be a tribe within the subfamily Trechinae. 

However all other authors either treat them as a 

separate higher taxon, e.g. family Broscidae 

alongside family Trechidae (Deuve, 1993), or as a 

subfamily separate from the Trechines e.g. 

subfamily Broscinae, whereas Trechitae are 

considered a supertribe within the subfamily 

Psydrinae (Erwin, 1985) or a separate subfamily 

Trechinae. Detailed studies of the broscines have 

since supported their separate status (Roig-

Family CARABIDAE Latreille, 1802 Checklist of Beetles of the British Isles, 2012 Edition 

Juňent, 1998). The present list treats the broscines 

as a tribe within Carabinae. 

A smaller confusion surrounds the placement of 

the Platynini (often including Sphodrini). 

Traditional British and European classifications 

have included these in the Pterostichini, e.g. 

Hansen (1996) for Denmark, Turin (2000) for the 

Netherlands. However Lawrence & Newton (1985) 

not only agreed with Erwin (1985) in considering 

the Platynini as a separate tribe, but placed them 

near the Perigonini, Zuphiini and other 

‘truncatipennes’ groups. The latter placement can 

be supported on larval characters (Arndt, 1998). 

However although retaining the Sphodrini and 

Platynini, I prefer to place them immediately after 

the Pterostichini, as in the traditional arrangement. 

The tribe Licinini is here considered to include 

Badistriini, following Kryzhanovskij et al. (1995). 

Subtribe names are not included in the list, 

although the Harpalini and Platynini, in particular, 

are often subdivided at this level. 

The treatment of the major genera, Carabus, 

Bembidion, Pterostichus, Amara, Harpalus and 

Agonum in the present list has tried to follow the 

divisions currently generally accepted by most 

European workers and as used in Löbl & Smetana 

(2003). This inevitably has resulted in some 

inconsistencies between these genera. The limits 

of Carabus are generally agreed, at least for the 

British fauna, so no 'new' generic names are used. 

The names and arrangement of subgenera of 

Carabus follow Casale & Březina (2003). 

Bembidion, although the largest genus of 

European carabids, is seldom split into separate 

genera. Only Ocys (two British species) and 

Table 1: Carabidae species added to the British list since Pope (1977). 

14 

Cillenus (a single species) are separated by 

Kryzhanovskij et al. (1995), Hůrka (1996) and 

Hansen (1996). I have used these genera, as well 

as treating Bracteon (two species) as a distinct 

genus, based at least partly on larval features 

(Luff, 1993). Most European taxonomic works 

(excepting Hansen, 1996) now consider Poecilus 

to be a separate genus from Pterostichus, and this 

is followed here. The remaining taxa within 

Pterostichus are considered as subgenera, 

although generic status for some or all may well be 

justified. Similarly, only Curtonotus is separated 

from Amara as a distinct genus, following 

Kryzhanovskij et al. (1995). I follow Telfer (2001b) 

in considering Ophonus as a distinct genus, but 

have not used the genus Pseudoophonus, which 

remains within Harpalus (see Kryzhanovskij et al., 

1995). Agonum is the only one of these 

traditionally large genera which is now generally 

split into a number of genera. However Europhilus 

remains as a subgenus of Agonum. 

Other than changes in synonymies given in the 

list, 12 species have been added since Pope 

(1977): the references for these are given in Table 

1, as well as in the endnotes in the main list. Two 

species have been deleted from Pope’s list as they 

are now considered to be non-established 

introductions: Abax parallelus and Stenolophus 

comma (= plagiatus), while some other names 

have been lost due to synonymy or homonymy. 

The list also includes species now considered 

extinct. Introduced species which have never 

formed established populations here are listed in 

the appendix "Non-established introductions" on 

page 131. 

Species Reason for addition Literature source 

Asaphidion curtum taxonomic split Speight, Martinez & Luff (1986) 

Asaphidion stierlini taxonomic split Speight, Martinez & Luff (1986) 

Bembidion caeruleum immigrant Telfer (2001a) 

Bembidion inustum ? rare & overlooked Levey & Pavett (1999) 

Calathus cinctus taxonomic split Anderson & Luff (1994) 

Agonum lugens ? rare & overlooked Anderson (1985) 

Harpalus griseus immigrant Owen (1996) 

Ophonus subsinuatus overlooked museum specimens Telfer (2001b) 

Acupalpus maculatus immigrant Telfer (2003) 

Somotrichus unifasciatus temporarily established introduction Denton (2007) 

Cymindis macularis ? immigrant Hammond (1982a) 

Microlestes minutulus immigrant Eversham & Collier (1997)

Subfamily CICINDELINAE Latreille, 1802 

CICINDELA Linnaeus, 1758 

campestris Linnaeus, 1758 

hybrida Linnaeus, 1758 

maritima Latreille & Dejean, 1822 

sylvatica Linnaeus, 1758 

silvatica auctt. (misspelling) 

CYLINDERA Westwood, 1831 

CICINDELA sensu auctt. partim non Linnaeus, 1758 

germanica (Linnaeus, 1758) 

Subfamily BRACHININAE Bonelli, 1810 

BRACHINUS Weber, 1801 

Subgenus BRACHINUS Weber, 1801 

crepitans (Linnaeus, 1758) 

Subgenus BRACHYNIDIUS Reitter, 1919 

sclopeta (Fabricius, 1792) 

Subfamily OMOPHRONINAE Bonelli, 1810 

OMOPHRON Latreille, 1802 

SCOLYTUS Fabricius, 1790 non Müller, O.F., 1764 

limbatum (Fabricius, 1777) 

Subfamily CARABINAE Latreille, 1802 

Tribe CARABINI Latreille, 1802 

CALOSOMA Weber, 1801 

inquisitor (Linnaeus, 1758) 

sycophanta (Linnaeus, 1758) 

CARABUS Linnaeus, 1758 31 

Subgenus LIMNOCARABUS Géhin, 1876 

clatratus Linnaeus, 1761 

clathratus auctt. (misspelling) 

Subgenus EUCARABUS Géhin, 1885 

arvensis Herbst, 1784 

ssp. sylvaticus Dejean, 1826 

silvaticus auctt. (misspelling) 

anglicanus Motschulsky, 1865/6 

Subgenus CARABUS Linnaeus, 1758 

granulatus Linnaeus, 1758 

ssp. granulatus Linnaeus, 1758 

ssp. hibernicus Lindroth, 1956 

Subgenus MORPHOCARABUS Géhin, 1885 

monilis Fabricius, 1792 

consitus sensu auctt. non Panzer, 1809 

insularis Born, 1908 

Subgenus ARCHICARABUS Seidlitz, 1887 

nemoralis Müller, O.F., 1764 

Subgenus TACHYPUS Weber, 1801 

auratus Linnaeus, 1761 

Subgenus HEMICARABUS Géhin, 1885 

nitens Linnaeus, 1758 

15 

Subgenus OREOCARABUS Géhin, 1885 

glabratus Paykull, 1790 

ssp. lapponicus Born, 1909 


Subgenus MESOCARABUS Thomson, C.G., 1875 

problematicus Herbst, 1786 

ssp. feroensis Lapouge, 1910 

ssp. harcyniae Sturm, 1815 

catenulatus sensu auctt. non Scopoli, 1763 

gallicus Géhin, 1885 

Subgenus CHAETOCARABUS Thomson, C.G., 1875 

intricatus Linnaeus, 1761 

Subgenus MEGODONTUS Solier, 1848 

violaceus Linnaeus, 1758 

ssp. purpurascens Fabricius, 1787 

exasperatus sensu auctt. non Duftschmid, 1812 

ssp. sollicitans Hartert, 1907 

britannicus Born, 1908 

browni Deuve, 1999 

Tribe CYCHRINI Perty, 1830 

CYCHRUS Fabricius, 1794 

caraboides (Linnaeus, 1758) 

ssp. rostratus (Linnaeus, 1761) 

Tribe NEBRIINI Laporte, 1834 

LEISTUS Frölich, 1799 

Subgenus POGONOPHORUS Latreille, 1802 

montanus Stephens, 1827 

ssp. rhaeticus Heer, 1837 

rufomarginatus (Duftschmid, 1812) 

spinibarbis (Fabricius, 1775) 

Subgenus LEISTOPHORUS Reitter, 1905 

fulvibarbis Dejean, 1826 

Subgenus LEISTUS Frölich, 1799 

ferrugineus (Linnaeus, 1758) 

rufescens sensu auctt. partim non (Fabricius, 1775) 

terminatus (Hellwig in Panzer, 1793) 

rufescens (Fabricius, 1775) non (Strøm, 1768) 

praeustus (Fabricius, 1792) 32 

NEBRIA Latreille, 1802 

Subgenus PARANEBRIA Jeannel, 1937 

livida (Linnaeus, 1758) 

lateralis (Fabricius, 1787) 

sabulosa (Fabricius, 1787) 

Subgenus NEBRIA Latreille, 1802 

brevicollis (Fabricius, 1792) 

salina Fairmaire & Laboulbène, 1854 

degenerata Schaufuss, 1862 

iberica Oliveira, 1876 

klinckowstroemi Mjöberg, 1915 

Subgenus BOREONEBRIA Jeannel, 1937 

nivalis (Paykull, 1790) 

rufescens (Strøm, 1768) 

gyllenhali (Schönherr, 1806) 

balbii Bonelli, 1810 

EURYNEBRIA Ganglbauer, 1891 

complanata (Linnaeus, 1767)


PELOPHILA Dejean, 1821 

borealis (Paykull, 1790) 

Tribe NOTIOPHILINI Motschulsky, 1850 

NOTIOPHILUS Duméril, 1806 

aesthuans Motschulsky, 1864 

aestuans auctt. (misspelling) 

pusillus Waterhouse, G.R., 1833 non (von Schreber, 

1759) 

aquaticus (Linnaeus, 1758) 

pusillus (von Schreber, 1759) 

strigifrons (Baudi, 1864) 

blacki Edwards, J., 1913 

biguttatus (Fabricius, 1779) 

germinyi Fauvel, 1863 

hypocrita sensu auctt. non Putzeys, 1866 

palustris (Duftschmid, 1812) 

quadripunctatus Dejean, 1826 

quadriguttatus Fowler, 1886 (error) 

rufipes Curtis, 1829 

substriatus Waterhouse, G.R., 1833 

Tribe ELAPHRINI Latreille, 1802 

BLETHISA Bonelli, 1810 

HELOBIUM Leach, 1815 

multipunctata (Linnaeus, 1758) 

ELAPHRUS Fabricius, 1775 

Subgenus ELAPHRUS Fabricius, 1775 

cupreus Duftschmid, 1812 


uliginosus Fabricius, 1792 

Subgenus TRICHELAPHRUS Semenov, 1926 

riparius (Linnaeus, 1758) 

Tribe LORICERINI Bonelli, 1810 

LORICERA Latreille, 1802 

LOROCERA auctt. (misspelling) 

pilicornis (Fabricius, 1775) 

coerulescens sensu auctt. non (Linnaeus, 1758) 

Tribe SCARITINI Bonelli, 1810 

CLIVINA Latreille, 1802 

collaris (Herbst, 1784) 

contracta (Geoffroy in Fourcroy, 1785) 

fossor (Linnaeus, 1758) 

DYSCHIRIUS Bonelli, 1810 

Subgenus DYSCHIRIUS Bonelli, 1810 

angustatus (Ahrens, 1830) 

obscurus (Gyllenhal, 1827) 

thoracicus (Rossi, 1790) 

arenosus Stephens, 1827 

Subgenus DYSCHIRIODES Jeannel, 1941 

aeneus (Dejean, 1825) 

extensus Putzeys, 1846 

elongatulus Dawson, 1856 

globosus (Herbst, 1784) 

16 

gibbus (Fabricius, 1792) 

impunctipennis Dawson, 1854 

luedersi Wagner, 1915 

aeneus sensu auctt. non (Dejean, 1825) 

?tristis Stephens, 1827 

unicolor sensu auctt. non Motschulsky, 1844 

nitidus (Dejean, 1825) 

politus (Dejean, 1825) 

salinus Schaum, 1843 

Tribe BROSCINI Hope, 1838 

BROSCUS Panzer, 1813 

cephalotes (Linnaeus, 1758) 

MISCODERA Eschscholtz, 1830 

arctica (Paykull, 1798) 

Tribe TRECHINI Bonelli, 1810 

PERILEPTUS Schaum, 1860 

BLEMUS sensu Laporte, 1840 non Dejean, 1821 

areolatus (Creutzer, 1799) 

AEPUS Leach, 1819 

AEPOPSIS Jeannel, 1922 

marinus (Strøm, 1783) 

fulvescens Samouelle, 1819 

robinii (Laboulbène, 1849) 

robini auctt. (misspelling) 

TRECHUS Clairville, 1806 

BLEMUS sensu Stephens, 1827 non Dejean, 1821 

Subgenus EPAPHIUS Leach, 1819 

rivularis (Gyllenhal, 1810) 

secalis (Paykull, 1790) 

Subgenus TRECHUS Clairville, 1806 

fulvus Dejean, 1831 

lapidosus (Dawson, 1849) 

obtusus Erichson, 1837 

quadristriatus (Schrank, 1781) 

minutus (Fabricius, 1781) 

rubens sensu Clairville, 1806 non (Fabricius, 1792) 

rubens (Fabricius, 1792) 

paludosus (Gyllenhal, 1810) 

subnotatus Dejean, 1831 

THALASSOPHILUS Wollaston, 1854 

TRECHUS sensu Fowler, 1886 partim non Clairville, 

1806 

longicornis (Sturm, 1825) 

BLEMUS Dejean, 1821 

LASIOTRECHUS Ganglbauer, 1892 

discus (Fabricius, 1792) 

TRECHOBLEMUS Ganglbauer, 1891 

micros (Herbst, 1784)

Tribe BEMBIDIINI Stephens, 1827 

TACHYS Dejean, 1821 

Subgenus PARATACHYS Casey, 1918 

EOTACHYS Jeannel, 1941 

bistriatus (Duftschmid, 1812) 

?pallidulus (Antoine, 1943) non (Ménétries, 1846) 33 

micros (Fischer von Waldheim, 1828) 

gregarius Chaudoir, 1846 

obtusiusculus (Jeannel, 1941) 

piceus Edmonds, 1934 non Dalla Torre, 1877 

edmondsi Moore, 1956 33 

Subgenus TACHYS Dejean, 1821 

scutellaris Stephens, 1828 

ELAPHROPUS Motschulsky, 1839 

TACHYS sensu auctt. partim non Dejean, 1821 

parvulus (Dejean, 1831) 

walkerianus (Sharp, 1913) 

ASAPHIDION des Gozis, 1886 

TACHYPUS sensu auctt. non Weber, 1801 

curtum (Heyden, 1870) 34 

flavipes sensu auctt. Brit. partim non (Linnaeus, 

1761) 

flavipes (Linnaeus, 1761) 

pallipes (Duftschmid, 1812) 

stierlini (Heyden, 1880) 35 

flavipes sensu auctt. Brit. partim non (Linnaeus, 

1761) 

OCYS Stephens, 1828 

BEMBIDION sensu auctt. partim non Latreille, 1802 

harpaloides (Audinet-Serville, 1821) 

rufescens (Guérin-Méneville, 1823) 

quinquestriatus (Gyllenhal, 1810) 

CILLENUS Leach, 1819 


lateralis Samouelle, 1819 

BRACTEON Bedel, 1879 


CHRYSOBRACTEON Netolitzky, 1914 

argenteolum (Ahrens, 1812) 

litorale (Olivier, 1790) 

littorale (Olivier, 1791) 

paludosum (Panzer, 1794) 

BEMBIDION Latreille, 1802 

BEMBIDIUM auctt. (misspelling) 

Subgenus NEJA Motschulsky, 1864 

nigricorne Gyllenhal, 1827 

Subgenus METALLINA Motschulsky, 1850 

lampros (Herbst, 1784) 

celere (Fabricius, 1792) 

properans (Stephens, 1828) 

velox Erichson, 1837 non (Linnaeus, 1761) 

coeruleotinctum Reitter, 1908 

cyaneotinctum Sharp, 1913 

caeruleipenne Saunders, 1936 

17 


Subgenus PRINCIDIUM Motschulsky, 1864 

punctulatum Drapiez, 1821 

Subgenus ACTEDIUM Motschulsky, 1864 

pallidipenne (Illiger, 1802) 

ruficolle (Illiger, 1801) non (Panzer, 1796) 

Subgenus TESTEDIUM Motschulsky, 1864 

bipunctatum (Linnaeus, 1761) 

Subgenus EUPETEDROMUS Netolitzky, 1911 

dentellum (Thunberg, 1787) 

flammulatum (Clairville, 1806) 

Subgenus NOTAPHUS Dejean, 1821 

obliquum Sturm, 1825 

semipunctatum (Donovan, 1806) 

adustum Schaum, 1860 

varium (Olivier, 1795) 36 

ustulatum sensu Sturm, 1825 non (Linnaeus, 1758) 

nebulosum (Stephens, 1828) 

Subgenus NOTAPHEMPHANES Netolitzky, 1920 

NOTHAPHEMPHANES auctt. (misspelling) 

ephippium (Marsham, 1802) 

Subgenus PLATAPHUS Motschulsky, 1864 

prasinum (Duftschmid, 1812) 

Subgenus TRICHOPLATAPHUS Netolitzky, 1914 

BLEPHAROPLATAPHUS Netolitzky, 1920 

virens Gyllenhal, 1827 

Subgenus BEMBIDIONETOLITZKYA Strand, E., 1929 

DANIELA Netolitzky, 1910 non Koch, 1891 

atrocaeruleum (Stephens, 1828) 

atrocoeruleum auctt. (misspelling) 

caeruleum Audinet-Serville, 1826 37 

coeruleum auctt. (misspelling) 

geniculatum Heer, 1837/8 

redtenbacheri Daniel, K., 1902 

tibiale (Duftschmid, 1812) 

Subgenus OCYDROMUS Clairville, 1806 

PERYPHUS Dejean, 1821 

bruxellense Wesmael, 1835 

rupestre sensu auctt. non (Linnaeus, 1767) 

bualei Jacquelin du Val, 1852 

andreae sensu auctt. non (Fabricius, 1787) 

cruciatum sensu auctt. non Dejean, 1831 

ssp. anglicanum Sharp, 1869 

ssp. polonicum Müller, J., 1930 

decorum (Zenker in Panzer, 1800) 

deletum Audinet-Serville, 1821 

nitidulum (Marsham, 1802) non (Schrank, 1781) 

dalmatinum var. latinum sensu MacKechnie-Jarvis, 

1932 non Netolitzky, 1911 

femoratum Sturm, 1825 

fluviatile Dejean, 1831 

lunatum (Duftschmid, 1812) 

maritimum (Stephens, 1835) 

concinnum sensu auctt. non (Stephens, 1828) 

dorsuarium Bedel, 1879 

monticola Sturm, 1825 

saxatile Gyllenhal, 1827 

vectense Fowler, 1886 

stephensii Crotch, 1866 

stephensi auctt. (misspelling) 

affine (Stephens, 1835) non Say, 1825 

testaceum (Duftschmid, 1812) 

tetracolum Say, 1825 

ustulatum sensu auctt. non (Linnaeus, 1758)


littorale sensu auctt. non (Olivier, 1791) 

Subgenus NEPHA Motschulsky, 1864 

illigeri Netolitzky, 1914 

quadriguttatum sensu auctt. Brit. non (Illiger, 1798) 

nec (Fabricius, 1775) 

tetragrammum sensu auctt. non Chaudoir, 1846 

genei sensu auctt. Brit. non Küster, 1847 

Subgenus SINECHOSTICTUS Motschulsky, 1864 

SYNECHOSTICTUS auctt. (misspelling) 

stomoides Dejean, 1831 

atroviolaceum sensu auctt. non Dufour, 1820 

Subgenus PSEUDOLIMNAEUM Kraatz, 1888 

inustum Jacquelin du Val, 1857 38 

Subgenus LYMNAEUM Stephens, 1828 

nigropiceum (Marsham, 1802) 

Subgenus SEMICAMPA Netolitzky, 1910 

gilvipes Sturm, 1825 

schuppelii Dejean, 1831 

schueppeli auctt. (misspelling) 

Subgenus DIPLOCAMPA Bedel, 1896 

assimile Gyllenhal, 1810 

clarkii (Dawson, 1849) 

clarki auctt. (misspelling) 

fumigatum (Duftschmid, 1812) 

Subgenus EMPHANES Motschulsky, 1850 

minimum (Fabricius, 1792) 39 

normannum Dejean, 1831 

Subgenus BEMBIDION Latreille, 1802 

LOPHA Dejean, 1821 non Bolten, 1798 

humerale Sturm, 1825 

quadrimaculatum (Linnaeus, 1761) 

quadriguttatum (Fabricius, 1775) 

quadripustulatum Audinet-Serville, 1821 

quadriguttatum sensu (Olivier, 1795) non (Fabricius, 

1775) 

antiquorum Crotch, 1871 

Subgenus TREPANEDORIS Netolitzky, 1918 

doris (Panzer, 1796) 

Subgenus TREPANES Motschulsky, 1864 

LEJA Dejean, 1821 32 

articulatum (Panzer, 1795) 

octomaculatum (Goeze, 1777) 

sturmii (Panzer, 1804) 

sturmi auctt. (misspelling) 

Subgenus PHYLA Motschulsky, 1844 

PHILA Motschulsky, 1846 40 

obtusum Audinet-Serville, 1821 

Subgenus PHILOCHTHUS Stephens, 1828 

PHILOCTHUS auctt. (misspelling) 

aeneum Germar, 1824 

biguttatum (Fabricius, 1779) 

guttula (Fabricius, 1792) 

iricolor Bedel, 1879 

riparium sensu Fowler, 1886 partim non (Olivier, 

1795) 

lunulatum (Geoffroy in Fourcroy, 1785) 

riparium (Olivier, 1795) 

mannerheimii Sahlberg, C.R., 1827 

mannerheimi auctt. (misspelling) 

haemorrhoum sensu auctt. non (Stephens, 1828) 

unicolor Chaudoir, 1850 

18 

Tribe POGONINI Laporte, 1834 

POGONUS Dejean, 1821 

chalceus (Marsham, 1802) 

littoralis (Duftschmid, 1812) 41 

litoralis auctt. (misspelling) 

luridipennis (Germar, 1822) 

Tribe PATROBINI Kirby, 1837 

PATROBUS Dejean, 1821 

assimilis Chaudoir, 1844 

clavipes Thomson, C.G., 1857 

atrorufus (Strøm, 1768) 

excavatus (Paykull, 1790) 

rufipes sensu (Duftschmid, 1812) non (Fabricius, 

1792) 

septentrionis Dejean, 1828 

Tribe PTEROSTICHINI Bonelli, 1810 

STOMIS Clairville, 1806 

pumicatus (Panzer, 1795) 

POECILUS Bonelli, 1810 

PTEROSTICHUS sensu auctt. partim non Bonelli, 

1810 

FERONIA Latreille, 1817 

cupreus (Linnaeus, 1758) 

affinis (Sturm, 1824) 

erythropus (Dejean, 1828) 

dinniki (Lutshnik, 1912) 

caesica Donisthorpe, 1931 

kugelanni (Panzer, 1797) 

dimidiatus (Olivier, 1795) non (Rossi, 1790) 

lepidus (Leske, 1785) 

versicolor (Sturm, 1824) 

caerulescens sensu auctt. non (Linnaeus, 1758) 

PTEROSTICHUS Bonelli, 1810 

Subgenus PTEROSTICHUS Bonelli, 1810 

cristatus (Dufour, 1820) 

ssp. parumpunctatus Germar, 1824 

Subgenus STEROPUS Stephens, 1828 

aethiops (Panzer, 1796) 42 

madidus (Fabricius, 1775) 

concinnus (Sturm, 1818) 

Subgenus PEDIUS Motschulsky, 1850 

longicollis (Duftschmid, 1812) 43 

inaequalis (Marsham, 1802) non (Panzer, 1795) 

ochraceus (Sturm, 1824) 

Subgenus LYPEROSOMUS Motschulsky, 1850 

aterrimus (Herbst, 1784) 

Subgenus ADELOSIA Stephens, 1835 

macer (Marsham, 1802) 

picimanus (Duftschmid, 1812) 

Subgenus PLATYSMA Bonelli, 1810 

niger (Schaller, 1783) 44 

scotus (Jeannel, 1942) 

Subgenus BOTHRIOPTERUS Chaudoir, 1838 

adstrictus Eschscholtz, 1823

orinomum (Stephens, 1828) 

vitreus (Dejean, 1828) 

oblongopunctatus (Fabricius, 1787) 

quadrifoveolatus Letzner, 1852 

angustatus (Duftchmid, 1812) non (Fabricius, 1787) 

Subgenus OMASEUS Stephens, 1828 

melanarius (Illiger, 1798) 

vulgaris sensu auctt. non (Linnaeus, 1758) 

Subgenus PSEUDOMASEUS Chaudoir, 1838 

anthracinus (Panzer, 1795) 

gracilis (Dejean, 1828) 

minor (Gyllenhal, 1827) 

nigrita (Paykull, 1790) 

rhaeticus Heer, 1837/8 45 

nigrita sensu auctt. partim non (Paykull, 1790) 

Subgenus LAGARUS Chaudoir, 1838 

vernalis (Panzer, 1795) 46 

crenatus (Duftschmid, 1812) non (Gmelin in 

Linnaeus, 1790) 

Subgenus ARGUTOR Stephens, 1828 

diligens (Sturm, 1824) 

strenuus sensu Dawson, 1854 non (Panzer, 1796) 

strenuus (Panzer, 1796) 

erythropus (Marsham, 1802) 

ABAX Bonelli, 1810 47 

parallelepipedus (Piller & Mitterpacher, 1783) 

parallelopipedus auctt. (misspelling) 

ater (Villers, 1790) 

striola (Fabricius, 1792) 

Tribe SPHODRINI Laporte, 1834 

PLATYDERUS Stephens, 1828 

PLATYDERES Stephens, 1827 32 

depressus (Audinet-Serville, 1821) 

ruficollis (Marsham, 1802) non (Fabricius, 1787) 

SYNUCHUS Gyllenhal, 1810 

TAPHRIA Latreille, 1819 

ODONTONYX Stephens, 1827 

vivalis (Illiger, 1798) 

nivalis (Panzer, 1797) non (Paykull, 1790) 

CALATHUS Bonelli, 1810 

Subgenus AMPHIGYNUS Haliday, 1841 

rotundicollis Dejean, 1828 

piceus sensu (Marsham, 1802) non (Linnaeus, 

1758) 

Subgenus CALATHUS Bonelli, 1810 

ambiguus (Paykull, 1790) 

fuscus (Fabricius, 1792) 

cinctus Motschulsky, 1850 48 

melanocephalus sensu auctt. partim non (Linnaeus, 

1758) 

mollis sensu auctt. partim non Marsham, 1802 

erythroderus Gemminger & Harold, 1868 

erratus (Sahlberg, C.R., 1827) 

fulvipes sensu (Gyllenhal, 1810) non (Fabricius, 

1792) 

flavipes sensu (Duftschmid, 1812) non (Geoffroy in 

Fourcroy, 1785) 

fuscipes (Goeze, 1777) 

19 


cisteloides (Panzer, 1793) 

melanocephalus (Linnaeus, 1758) 

ochropterus (Duftschmid, 1812) 

nubigena Haliday, 1838 

micropterus (Duftschmid, 1812) 

mollis (Marsham, 1802) 49 

ochropterus sensu auctt. non (Duftschmid, 1812) 

SPHODRUS Clairville, 1806 

leucophthalmus (Linnaeus, 1758) 

leucopthalmus Fowler, 1886 (misspelling) 


LAEMOSTENUS Bonelli, 1810 

LAEMOSTHENES Schaufuss, 1865 

Subgenus LAEMOSTENUS Bonelli, 1810 

complanatus (Dejean, 1828) 

Subgenus PRISTONYCHUS Dejean, 1828 

terricola (Herbst, 1784) 

subcyaneus (Illiger, 1801) 

Tribe PLATYNINI Bonelli, 1810 

OLISTHOPUS Dejean, 1828 

ODONTONYX sensu auctt. non Stephens, 1827 

rotundatus (Paykull, 1790) 50 

rotundicollis (Marsham, 1802) 

OXYPSELAPHUS Chaudoir, 1843 

AGONUM sensu auctt. partim non Bonelli, 1810 

ANCHUS LeConte, 1854 

obscurus (Herbst, 1784) 

?obscurus (Müller, O.F., 1776) 

oblongus (Fabricius, 1792) 

PARANCHUS Lindroth, 1974 


albipes (Fabricius, 1796) 

ruficornis (Goeze, 1777) non (De Geer, 1774) 

oblongus (Fabricius, 1792) non (Fabricius, 1792) 

pallipes (Fabricius, 1801) non (Fabricius, 1787) 

ANCHOMENUS Bonelli, 1810 


dorsalis (Pontoppidan, 1763) 

prasinus (Thunberg, 1784) 

PLATYNUS Bonelli, 1810 


assimilis (Paykull, 1790) 

?junceus (Scopoli, 1763) 

angusticollis (Fabricius, 1801) 

BATENUS Motschulsky, 1864 


livens (Gyllenhal, 1810) 

SERICODA Kirby, 1837 


quadripunctata (De Geer, 1774)


AGONUM Bonelli, 1810 

Subgenus EUROPHILUS Chaudoir, 1859 

fuliginosum (Panzer, 1809) 

gracile Sturm, 1824 

micans Nicolai, 1822 

piceum (Linnaeus, 1758) 

scitulum Dejean, 1828 

thoreyi Dejean, 1828 

pelidnum sensu (Paykull, 1792) non (Herbst, 1784) 

puellum Dejean, 1828 

Subgenus AGONUM Bonelli, 1810 

chalconotum Ménétries, 1832 

sahlbergii (Chaudoir, 1850) 

sahlbergi auctt. (misspelling) 

archangelicum Sahlberg, J., 1874 

emarginatum (Gyllenhal, 1827) 

afrum (Duftschmid, 1812) non (Thunberg, 1784) 

moestum sensu auctt. non (Duftschmid, 1812) nec 

(Gmelin in Linnaeus, 1790) 

ericeti (Panzer, 1809) 

gracilipes (Duftschmid, 1812) 

elongatum Dejean, 1828 

lugens (Duftschmid, 1812) 51 

marginatum (Linnaeus, 1758) 

muelleri (Herbst, 1784) 

parumpunctatum (Fabricius, 1792) 

chalybeum Sturm, 1824 

nigrum Dejean, 1828 

atratum sensu auctt. non (Duftschmid, 1812) 

dahli (Preudhomme de Borre, 1879) 

sexpunctatum (Linnaeus, 1758) 

versutum Sturm, 1824 

viduum (Panzer, 1796) 

moestum sensu Fowler & Donisthorpe, 1913 non 

(Duftschmid, 1812) 

Tribe ZABRINI Bonelli, 1810 

ZABRUS Clairville, 1806 

tenebrioides (Goeze, 1777) 


AMARA Bonelli, 1810 

Subgenus ZEZEA Csiki, 1929 

TRIAENA LeConte, 1847 non Hübner, 1818 

plebeja (Gyllenhal, 1810) 

plebeia auctt. (misspelling) 

strenua Zimmermann, 1832 

vectensis Dawson, 1849 

Subgenus AMARA Bonelli, 1810 

aenea (De Geer, 1774) 

trivialis (Gyllenhal, 1810) 

anthobia Villa & Villa, 1833 

communis (Panzer, 1797) 

vulgaris sensu Dawson, 1854 non (Linnaeus, 1758) 

convexior Stephens, 1828 

continua Thomson, C.G., 1873 

curta Dejean, 1828 

eurynota (Panzer, 1796) 

eyrinota auctt. (misspelling) 

acuminata (Paykull, 1798) 

famelica Zimmermann, 1832 

familiaris (Duftschmid, 1812) 

lucida (Duftschmid, 1812) 

lunicollis Schiødte, 1837 

20 

vulgaris sensu auctt. non (Linnaeus, 1758) 

montivaga Sturm, 1825 

nitida Sturm, 1825 

ovata (Fabricius, 1792) 52 

obsoleta sensu Dejean, 1828 non (Duftschmid, 

1812) 

adamantina Kolenati, 1845 

similata (Gyllenhal, 1810) 

obsoleta (Duftschmid, 1812) 

spreta Dejean, 1831 

tibialis (Paykull, 1798) 

Subgenus CELIA Zimmermann, 1832 

bifrons (Gyllenhal, 1810) 

livida sensu Schiødte, 1841 non (Fabricius, 1792) 

cursitans (Zimmermann, 1832) 

fuscicornis (Zimmermann, 1832) 

fusca Dejean, 1828 

complanata Dejean, 1828 

infima (Duftschmid, 1812) 53 

praetermissa (Sahlberg, C.R., 1827) 

rufocincta (Sahlberg, C.R., 1827) 

Subgenus PARACELIA Bedel, 1899 

quenseli (Schönherr, 1806) 

quenselii auctt. (misspelling) 

Subgenus BRADYTUS Stephens, 1827 

apricaria (Paykull, 1790) 

consularis (Duftschmid, 1812) 

fulva (Müller, O.F., 1776) 54 

Subgenus PERCOSIA Zimmermann, 1832 

equestris (Duftschmid, 1812) 

patricia (Duftschmid, 1812) 

CURTONOTUS Stephens, 1827 

CYRTONOTUS auctt. (misspelling) 

AMARA sensu auctt. partim non Bonelli, 1810 

alpinus (Paykull, 1790) 

aulicus (Panzer, 1796) 

spinipes sensu Schiødte, 1841 non (Linnaeus, 

1758) 

convexiusculus (Marsham, 1802) 

Tribe HARPALINI Bonelli, 1810 

HARPALUS Latreille, 1802 

Subgenus PSEUDOOPHONUS Motschulsky, 1844 

PSEUDOPHONUS auctt. (misspelling) 

PARDILEUS des Gozis, 1882 

calceatus (Duftschmid, 1812) 

griseus (Panzer, 1797) 55 

rufipes (De Geer, 1774) 

ruficornis (Fabricius, 1775) 

pubescens (Müller, O.F., 1776) 

Subgenus HARPALUS Latreille, 1802 

HAPLOHARPALUS Schauberger, 1926 

affinis (Schrank, 1781) 

aeneus (Fabricius, 1775) non (De Geer, 1774) 

anxius (Duftschmid, 1812) 

attenuatus Stephens, 1828 

consentaneus Dejean, 1829 

cupreus Dejean, 1829 

dimidiatus (Rossi, 1790) 

caspius sensu auctt. Brit. non (von Steven, 1806) 

froelichii Sturm, 1818 

froelichi auctt. (misspelling)

honestus (Duftschmid, 1812) 

ignavus (Duftschmid, 1812) 

laevipes Zetterstedt, 1828 

seriepunctatus sensu Gyllenhal, 1827 non Sturm, 

1818 

quadripunctatus Dejean, 1829 

montivagus Reitter, 1900 

latus (Linnaeus, 1758) 

erythrocephalus (Fabricius, 1787) 

metallescens Rye, 1874 

neglectus Audinet-Serville, 1821 

pumilus Sturm, 1818 

vernalis sensu (Duftschmid, 1812) non (Panzer, 

1796) 

picipennis sensu auctt. Brit. non Duftschmid, 1812 

funestus Audinet-Serville, 1821 

rubripes (Duftschmid, 1812) 

sobrinus Dejean, 1829 

rufipalpis Sturm, 1818 

rufitarsis (Duftschmid, 1812) non (Illiger, 1801) 

ignavus sensu auctt. Brit. non (Duftschmid, 1812) 

serripes (Quensel in Schönherr, 1806) 

servus (Duftschmid, 1812) 

smaragdinus (Duftschmid, 1812) 

discoideus Erichson, 1837 

tardus (Panzer, 1796) 

rufimanus (Marsham, 1802) 

luteicornis sensu auctt. Brit. non (Duftschmid, 1812) 

Subgenus CRYPTOPHONUS Brandmayr & Zetto 

Brandmayr, 1982 

melancholicus Dejean, 1829 

tenebrosus Dejean, 1829 

ssp. centralis Schauberger, 1929 

OPHONUS Dejean, 1821 

HARPALUS sensu auctt. partim non Latreille, 1802 

Subgenus OPHONUS Dejean, 1821 

ardosiacus (Lutshnik, 1922) 

obscurus sensu (Sturm, 1818) non (Fabricius, 1792) 

diffinis sensu Joy, 1932 non (Dejean, 1829) 

rotundicollis sensu auctt. Brit. non Kolenati, 1845 

azureus (Fabricius, 1775) 

subquadratus sensu auctt. Brit. non (Dejean, 1829) 

sabulicola (Panzer, 1796) 

stictus Stephens, 1828 

obscurus (Fabricius, 1792) non (Müller, O.F., 1776) 

monticola (Dejean, 1829) 

Subgenus METOPHONUS Bedel, 1895 

cordatus (Duftschmid, 1812) 56 

laticollis Mannerheim, 1825 

punctatulus (Duftschmid, 1812) non (Fabricius, 

1792) 

nitidulus sensu Stephens, 1828 non (Schrank, 

1781) 

melletii (Heer, 1837/8) 

melleti auctt. (misspelling) 

rectangulus Thomson, C.G., 1870 

championi Sharp, 1912 

rupicoloides Sharp, 1912 

brevicollis sensu Jeannel, 1942 ? (Audinet-Serville, 

1821) 

parallelus (Dejean, 1829) 

melleti sensu Jeannel, 1942 non (Heer, 1837/8) 

zigzag sensu auctt. Brit. non Costa, 1882 

puncticeps Stephens, 1828 

21 


rectangulus sensu Sharp, 1912 non Thomson, C.G., 

1870 

angusticollis Müller, J., 1921 

puncticollis (Paykull, 1798) 

rufibarbis (Fabricius, 1792) 

brevicollis sensu auctt. partim non (Audinet-Serville, 

1821) 

subpunctatus Stephens, 1828 

seladon (Schauberger, 1926) 

rupicola (Sturm, 1818) 

schaubergerianus (Puel, 1937) 

rufibarbis sensu auctt. non (Fabricius, 1792) 

brevicollis sensu auctt. partim non (Audinet-Serville, 

1821) 

subsinuatus Rey, 1886 57 

ANISODACTYLUS Dejean, 1829 

binotatus (Fabricius, 1787) 

spurcaticornis Dejean, 1829 

nemorivagus (Duftschmid, 1812) 

atricornis (Stephens, 1835) 

poeciloides (Stephens, 1828) 

pseudoaeneus sensu auctt. non Dejean, 1829 

DIACHROMUS Erichson, 1837 

germanus (Linnaeus, 1758) 

SCYBALICUS Schaum, 1862 

oblongiusculus (Dejean, 1829) 

DICHEIROTRICHUS Jacquelin du Val, 1855 

DICHIROTRICHUS auctt. (misspelling) 

gustavii Crotch, 1871 

gustavi auctt. (misspelling) 

pubescens (Paykull, 1790) non (Müller, O.F., 1776) 

obsoletus (Dejean, 1829) 

TRICHOCELLUS Ganglbauer, 1892 

BRADYCELLUS sensu Fowler, 1886 partim non 

Erichson, 1837 

cognatus (Gyllenhal, 1827) 

placidus (Gyllenhal, 1827) 

BRADYCELLUS Erichson, 1837 

caucasicus (Chaudoir, 1846) 

collaris (Paykull, 1798) non (Herbst, 1784) 

csikii Laczó, 1912 

distinctus (Dejean, 1829) 

harpalinus (Audinet-Serville, 1821) 

ruficollis (Stephens, 1828) 

similis (Dejean, 1829) 

sharpi Joy, 1912 

distinctus sensu Fowler, 1886 non (Dejean, 1829) 

verbasci (Duftschmid, 1812) 

STENOLOPHUS Dejean, 1821 58 

mixtus (Herbst, 1784) 

vespertinus (Panzer, 1796) 

ziegleri (Panzer, 1809) 

skrimshiranus Stephens, 1828 

teutonus (Schrank, 1781) 

vaporariorum sensu (Fabricius, 1775) non 

(Linnaeus, 1758) 

anglicus Schiødte, 1861-63


ACUPALPUS Latreille, 1829 

brunnipes (Sturm, 1825) 

brunneipes auctt. (misspelling) 

dubius Schilsky, 1888 

luridus sensu auctt. non Dejean, 1829 

luteatus sensu Joy, 1932 non (Duftschmid, 1812) 

elegans (Dejean, 1829) 

exiguus Dejean, 1829 

flavicollis (Sturm, 1825) 

luridus Dejean, 1829 

maculatus (Schaum, 1860) 59 

meridianus (Linnaeus, 1761) 

parvulus (Sturm, 1825) 

dorsalis (Fabricius, 1787) non (Pontoppidan, 1763) 

derelictus (Dawson, 1854) 

ANTHRACUS Motschulsky, 1850 

ACUPALPUS sensu auctt. partim non Latreille, 

1829 

consputus (Duftschmid, 1812) 

Tribe CHLAENIINI Brullé, 1834 

CHLAENIUS Bonelli, 1810 

CHLAENIELLUS Reitter, 1908 

nigricornis (Fabricius, 1787) 

melanocornis Dejean, 1826 

nitidulus (Schrank, 1781) 

schrankii (Duftschmid, 1812) 

tristis (Schaller, 1783) 

holosericeus (Fabricius, 1787) 

vestitus (Paykull, 1790) 

CALLISTUS Bonelli, 1810 

lunatus (Fabricius, 1775) 

Tribe OODINI LaFerté-Sénectère, 1851 

OODES Bonelli, 1810 

helopioides (Fabricius, 1792) 

Tribe LICININI Bonelli, 1810 

LICINUS Latreille, 1802 

depressus (Paykull, 1790) 

punctatulus (Fabricius, 1792) 

punctulatus Kloet & Hincks, 1945 (error) 

silphoides sensu (Fabricius, 1792) non (Rossi, 

1790) 

BADISTER Clairville, 1806 

Subgenus BADISTER Clairville, 1806 

bullatus (Schrank, 1798) 

bipustulatus (Fabricius, 1792) non (Fabricius, 1775) 

meridionalis Puel, 1925 

kineli Makolski, 1952 

unipustulatus Bonelli, 1813 

Subgenus TRIMORPHUS Stephens, 1828 

sodalis (Duftschmid, 1812) 

humeralis Bonelli, 1813 

Subgenus BAUDIA Ragusa, 1884 

collaris Motschulsky, 1844 

anomalus (Perris, 1866) 

22 

striatulus Hansen, 1944 

dilatatus Chaudoir, 1837 

peltatus (Panzer, 1797) 

Tribe PANAGAEINI Bonelli, 1810 

PANAGAEUS Latreille, 1802 

bipustulatus (Fabricius, 1775) 

quadripustulatus Sturm, 1815 

cruxmajor (Linnaeus, 1758) 

Tribe PERIGONINI Horn, 1881 

PERIGONA Laporte, 1835 

nigriceps (Dejean, 1831) 

Tribe MASOREINI Chaudoir, 1871 

MASOREUS Dejean, 1821 

wetterhallii (Gyllenhal, 1813) 

wetterhalli auctt. (misspelling) 

laticollis (Sturm, 1825) 

Tribe LEBIINI Bonelli, 1810 

LEBIA Latreille, 1802 

Subgenus LAMPRIAS Bonelli, 1810 

chlorocephala (Hoffmann, J., 1803) 

chrysocephala (Motschulsky, 1864) 

cyanocephala (Linnaeus, 1758) 

Subgenus LEBIA Latreille, 1802 

cruxminor (Linnaeus, 1758) 

marginata (Geoffroy in Fourcroy, 1785) 


scapularis (Geoffroy in Fourcroy, 1785) 

turcica (Fabricius, 1787) 

SOMOTRICHUS Seidlitz, 1887 

unifasciatus (Dejean, 1831) 60 

DEMETRIAS Bonelli, 1810 

Subgenus RISOPHILUS Leach, 1815 

AETOPHORUS Schmidt-Göbel, 1846 

imperialis (Germar, 1824) 

Subgenus DEMETRIAS Bonelli, 1810 

atricapillus (Linnaeus, 1758) 

monostigma Samouelle, 1819 

unipunctatus (Germar, 1824) 

CYMINDIS Latreille, 1806 

Subgenus CYMINDIS Latreille, 1806 

axillaris (Fabricius, 1794) 

Subgenus TARSOSTINUS Motschulsky, 1864 

macularis Mannerheim in Fischer von Waldheim, 

1824 61 

Subgenus TARULUS Bedel, 1906 

vaporariorum (Linnaeus, 1758) 

basalis Gyllenhal, 1810

PARADROMIUS Fowler, 1887 

DROMIUS sensu auctt. partim non Bonelli, 1810 

linearis (Olivier, 1795) 

longiceps (Dejean, 1826) 

DROMIUS Bonelli, 1810 

agilis (Fabricius, 1787) 62 

bimaculatus Dejean, 1825 

angustus Brullé, 1834 

meridionalis Dejean, 1825 

discus Puel, 1919 

quadrimaculatus (Linnaeus, 1758) 

CALODROMIUS Reitter, 1905 


spilotus (Illiger, 1798) 

quadrinotatus (Zenker in Panzer, 1800) non 

(Fabricius, 1798) 

PHILORHIZUS Hope, 1838 


DROMIOLUS Reitter, 1905 

melanocephalus (Dejean, 1825) 

notatus (Stephens, 1827) 

nigriventris (Thomson, C.G., 1857) 

quadrisignatus (Dejean, 1825) 

sigma (Rossi, 1790) 

vectensis (Rye, 1873) 

insignis sensu auctt. non (Lucas, 1846) 

MICROLESTES Schmidt-Göbel, 1846 

BLECHRUS Motschulsky, 1847 

maurus (Sturm, 1827) 

glabratus sensu auctt. non (Duftschmid, 1812) 

minutulus (Goeze, 1777) 63 

glabratus (Duftschmid, 1812) 

LIONYCHUS Wissmann, 1846 

quadrillum (Duftschmid, 1812) 

bipunctatus (Heer, 1838) 

unicolor Schilsky, 1888 

SYNTOMUS Hope, 1838 

METABLETUS Schmidt-Göbel, 1846 

foveatus (Geoffroy in Fourcroy, 1785) 

foveola (Gyllenhal, 1810) 

obscuroguttatus (Duftschmid, 1812) 

atratus (Dejean, 1825) 

truncatellus (Linnaeus, 1761) 

Tribe ODACANTHINI Laporte, 1834 

ODACANTHA Paykull, 1798 

COLLIURIS sensu auctt. non De Geer, 1774 

melanura (Linnaeus, 1767) 

Tribe DRYPTINI Bonelli, 1810 

DRYPTA Latreille, 1796 

dentata (Rossi, 1790) 

emarginata (Olivier, 1790) 

23 

Tribe ZUPHIINI Bedel, 1895 

Family HELOPHORIDAE Leach, 1815 

POLISTICHUS Bonelli, 1810 

POLYSTICHUS auctt. (misspelling) 

connexus (Geoffroy in Fourcroy, 1785) 

fasciolatus sensu auctt. non (Rossi, 1790) 

vittatus Brullé, 1834 

Suborder POLYPHAGA Emery, 1886 

Superfamily HYDROPHILOIDEA Latreille, 1802 

8. Family HELOPHORIDAE Leach, 1815 64 


HELOPHORUS Fabricius, 1775 

ELOPHORUS Fabricius, 1775 65 

Subgenus EMPLEURUS Hope, 1838 

nubilus Fabricius, 1776 

nubilis auctt. (misspelling) 

porculus Bedel, 1881 

rugosus sensu auctt. Brit. partim non Olivier, 1795 

rufipes (Bosc d'Antic, 1791) 

rugosus Olivier, 1795 

Subgenus CYPHELOPHORUS Kuwert, 1884 

tuberculatus Gyllenhal, 1808 

scaber LeConte, 1850 

Subgenus TRICHELOPHORUS Kuwert, 1886 

alternans Gené, 1836 

intermedius Mulsant, 1844 

Subgenus HELOPHORUS Fabricius, 1775 

aequalis Thomson, C.G., 1868 

aquaticus sensu auctt. Brit. ad 1965 partim non 

(Linnaeus, 1758) 66 

grandis Illiger, 1798 

aquaticus sensu auctt. Brit. partim, Strand, A., 1962 

non (Linnaeus, 1758) 

Subgenus ATRACTHELOPHORUS Kuwert, 1886 

arvernicus Mulsant, 1846 

brevipalpis Bedel, 1881 67 

creticus Kiesenwetter, 1858 

bulbipalpis Kuwert, 1890 

Subgenus RHOPALHELOPHORUS Kuwert, 1886 

dorsalis (Marsham, 1802) 

quadrisignatus Bach, 1851 

flavipes Fabricius, 1792 

viridicollis Stephens, 1829 

aeneipennis sensu auctt. non Thomson, C.G., 1854 

sphagnicola Hardy, 1871 

shetlandicus Kuwert, 1890 

phalleterus Sharp, 1916 

fulgidicollis Motschulsky, 1860 

mulsanti Cox, 1874 ex Rye, 1866 

flavipes var. mulsanti sensu auctt. Brit. partim 

exasperatus Sharp, 1916 

granularis (Linnaeus, 1761) 

brevicollis Thomson, C.G., 1868 

ytenensis Sharp, 1916 

griseus Herbst, 1793 

affinis (Marsham, 1802) non (Thunberg, 1794)

Family GEORISSIDAE Laporte, 1840 Checklist of Beetles of the British Isles, 2012 Edition 

semifulgens sensu d'Orchymont, 1937 non Rey, 

1885 

laticollis Thomson, C.G., 1854 

pallidulus Thomson, C.G., 1868 

longitarsis Wollaston, 1864 

erichsoni Bach, 1866 

semifulgens Rey, 1885 

diffinis Sharp, 1916 

minutus Fabricius, 1775 

nanus Sturm, 1836 

obscurus Mulsant, 1844 

aeneipennis Thomson, C.G., 1854 

walkeri Sharp, 1916 

strigifrons Thomson, C.G., 1868 

championi Sharp, 1915 

9. Family GEORISSIDAE Laporte, 1840 68 


GEORISSUS Latreille, 1809 

GEORYSSUS auctt. (misspelling) 

CATHAMMISTES Illiger, 1807 69 

crenulatus (Rossi, 1794) 

pygmaeus (Fabricius, 1798) 

10. Family HYDROCHIDAE Thomson, C.G., 

1859 70 


HYDROCHUS Leach, 1817 

angustatus Germar, 1824 

brevis (Herbst, 1793) 

crenatus (Fabricius, 1792) 

carinatus Germar, 1824 71 

elongatus (Schaller, 1783) 

ignicollis Motschulsky, 1860 

elongatus sensu auctt. Brit. partim non (Schaller, 

1783) 

megaphallus van Berge Henegouwen, 1988 72 

nitidicollis Mulsant, 1844 

interruptus sensu auctt. Brit. non von Heyden, 1870 

11. Family SPERCHEIDAE Erichson, 1837 73 


SPERCHEUS Kugelann in Illiger, 1798 

emarginatus (Schaller, 1783) 

12. Family HYDROPHILIDAE Latreille, 1802 74 


Subfamily HYDROPHILINAE Latreille, 1802 

ANACAENA Thomson, C.G., 1859 

bipustulata (Marsham, 1802) 

globulus (Paykull, 1798) 

limbata (Fabricius, 1792) 

24 

ovata (Reiche, 1861) 

lutescens (Stephens, 1829) 75 

limbata sensu auctt. partim non (Fabricius, 1792) 

nitida (Heer, 1841) 

PARACYMUS Thomson, C.G., 1867 

aeneus (Germar, 1824) 

scutellaris (Rosenhauer, 1856) 

aeneus sensu (Stephens, 1829) non (Germar, 

1824) 

nigroaeneus (Sahlberg, J., 1875) 

BEROSUS Leach, 1817 76 

Subgenus BEROSUS Leach, 1817 

affinis Brullé, 1835 

luridus (Linnaeus, 1761) 

signaticollis (Charpentier, 1825) 

Subgenus ENOPLURUS Hope, 1838 

fulvus Kuwert, 1888 

spinosus sensu auctt. partim non (von Steven in 

Schönherr, 1808) 

CHAETARTHRIA Stephens, 1833 

seminulum (Herbst, 1797) 

coccinelloides (Stephens, 1829) 

simillima Vorst & Cuppen, 2003 77 

seminulum sensu auctt. partim non (Herbst, 1797) 

CYMBIODYTA Bedel, 1881 

marginellus (Fabricius, 1792) 

marginella auctt. (misspelling) 

margipallens (Marsham, 1802) 

ovalis (Thomson, C.G., 1853) 

ENOCHRUS Thomson, C.G., 1859 

PHILYDRUS Solier, 1834 non Duftschmid, 1805 

PHILHYDRUS Brullé, 1835 non Brookes, 1828 

affinis (Thunberg, 1794) 

minutus sensu auctt., ?sensu (Fabricius, 1801) non 

(Fabricius, 1775, 1792) 

bicolor (Fabricius, 1792) 

maritimus (Thomson, C.G., 1853) 

coarctatus (Gredler, 1863) 

suturalis (Sharp, 1872) 

fuscipennis (Thomson, C.G., 1884) 78 

melanocephalus sensu auctt. Brit. partim non 

(Olivier, 1792) 

halophilus (Bedel, 1878) 79 



melanocephalus (Olivier, 1792) 

bicolor sensu (Fowler, 1887) non (Fabricius, 1792) 

nigritus (Sharp, 1872) 80 

affinis sensu auctt. partim non (Thunberg, 1794) 

isotae Hebauer, 1981 81 

ochropterus (Marsham, 1802) 

frontalis (Erichson, 1837) 

nigricans (Zetterstedt, 1838) 

quadripunctatus (Herbst, 1797) 



ytenensis (Sharp, 1915) 

testaceus (Fabricius, 1801)

HELOCHARES Mulsant, 1844 

lividus (Forster, 1771) 

griseus (Fabricius, 1787) 

obscurus (Müller, O.F., 1776) 82 

punctatus Sharp, 1869 

obscurus sensu auctt. partim non (Müller, O.F., 

1776) 

HYDROBIUS Leach, 1815 

fuscipes (Linnaeus, 1758) 

chalconotus Stephens, 1829 

subrotundus Stephens, 1829 

picicrus Thomson, C.G., 1884 

LIMNOXENUS Motschulsky, 1853 

HYDROBIUS sensu Fowler, 1887 partim non 

Leach, 1815 

niger (Gmelin in Linnaeus, 1790) 83 

oblongus (Herbst, 1797) 

HYDROCHARA Berthold, 1827 

HYDROPHILUS sensu Leach, 1817 non Geoffroy, 

1762 

HYDROCHARIS Hope, 1838 40 


HYDROPHILUS Geoffroy, 1762 

HYDROUS Linnaeus, 1775 

piceus (Linnaeus, 1758) 

LACCOBIUS Erichson, 1837 84 

atratus Rottenberg, 1874 

scutellaris sensu auctt. Brit. non Motschulsky, 1855 

regularis sensu auctt. Brit. non Rey, 1885 

bipunctatus (Fabricius, 1775) 

alutaceus Thomson, C.G., 1868 

colon (Stephens, 1829) 

biguttatus Gerhardt, 1877 

bipunctatus sensu Fowler, 1887 non (Fabricius, 

1775) 


simulatrix d'Orchymont, 1932 85 

simulator auctt. (misspelling) 

sculptus sensu Gentili & Chiesa, 1976 partim non 

d'Orchymont, 1936 

sinuatus Motschulsky, 1849 

oblongus Gorham, 1907 

striatulus (Fabricius, 1801) 

nigriceps Thomson, C.G., 1853 

sinuatus sensu auctt. Brit. ante 1907 non 

Motschulsky, 1849 

purpurascens Newbery, 1908 


atrocephalus sensu auctt. Brit. non Reitter, 1872 86 

Subfamily SPHAERIDIINAE Latreille, 1802 

COELOSTOMA Brullé, 1835 

CYCLONOTUM Erichson, 1837 

orbiculare (Fabricius, 1775) 

DACTYLOSTERNUM Wollaston, 1854 

abdominale (Fabricius, 1792) 87 

25 

Family HYDROPHILIDAE Latreille, 1802 

CERCYON Leach, 1817 88 

CERYCON Rey, 1886 

ERCYCON Rey, 1886 

PARALIOCERCYON Ganglbauer, 1904 

Subgenus DICYRTOCERCYON Ganglbauer, 1904 

ustulatus (Preyssler, 1790) 

haemorrhous (Gyllenhal, 1808) 

Subgenus PARACYCREON d’Orchymont, 1942 

laminatus Sharp, 1873 

Subgenus CERCYON Leach, 1817 

alpinus Vogt, 1969 89 

bifenestratus Küster, 1851 

convexiusculus Stephens, 1829 

lugubris sensu (Paykull, 1798) non (Olivier, 1790) 

intermixtus Sharp, 1918 

depressus Stephens, 1829 

granarius Erichson, 1837 


flavipes (Fabricius, 1792) 

aquatilis Donisthorpe, 1932 90 

impressus (Sturm, 1807) 

atomarius (Fabricius, 1775) non (Linnaeus, 1767) 

haemorrhoidalis sensu (Herbst, 1792) non 


lateralis (Marsham, 1802) 

littoralis (Gyllenhal, 1808) 

litoralis auctt. (misspelling) 

marinus Thomson, C.G., 1853 

aquaticus Laporte, 1840 non Stephens, 1829 

melanocephalus (Linnaeus, 1758) 

nigriceps (Marsham, 1802) 

atricapillus (Marsham, 1802) 91 

obsoletus (Gyllenhal, 1808) 

lugubris sensu (Olivier, 1790) non (Geoffroy in 


pygmaeus (Illiger, 1801) 

quisquilius (Linnaeus, 1761) 

flavus (Marsham, 1802) 

sternalis (Sharp, 1918) 

subsulcatus sensu auctt. non Rey, 1885 

pumilio (Sharp, 1918) 

terminatus (Marsham, 1802) 

tristis (Illiger, 1801) 

boletophagus (Marsham, 1802) 

minutus sensu auctt. Brit. non (Gyllenhal, 1808) 

unipunctatus (Linnaeus, 1758) 

Subgenus PARACERCYON Seidlitz, 1888 

analis (Paykull, 1798) 

flavipes sensu (Thunberg, 1794) non (Fabricius, 

1792) 

MEGASTERNUM Mulsant, 1844 

concinnum (Marsham, 1802) 

obscurum (Marsham, 1802) non (Fabricius, 1792) 

boletophagum sensu auctt. non (Marsham, 1802) 

minutum (Gyllenhal, 1808) 

CRYPTOPLEURUM Mulsant, 1844 

crenatum (Kugelann, 1794) 

minutum (Fabricius, 1775) 

atomarium sensu (Olivier, 1790) non (Linnaeus, 

1767) 

subtile Sharp, 1884

Family SPHAERITIDAE Shuckard, 1839 Checklist of Beetles of the British Isles, 2012 Edition 

SPHAERIDIUM Fabricius, 1775 

bipustulatum Fabricius, 1781 

?quadrimaculatum (Marsham, 1802) 

?semistriatum Laporte, 1840 

lunatum Fabricius, 1792 

scarabaeoides sensu auctt. Brit. partim non 


marginatum Fabricius, 1787 92 

bipustulatum sensu auctt. partim non Fabricius, 

1781 

scarabaeoides (Linnaeus, 1758) 

quadrimaculatum sensu auctt. Brit. partim non 

(Marsham, 1802) 

13. Family SPHAERITIDAE Shuckard, 1839 

Family author: R.G. Booth 

SPHAERITES Duftschmid, 1805 

glabratus (Fabricius, 1792) 

14. Family HISTERIDAE Gyllenhal, 1808 93 

Family author: A.G. Duff 

Subfamily ABRAEINAE MacLeay, 1819 

Tribe ABRAEINI MacLeay, 1819 

ABRAEUS Leach, 1817 

granulum Erichson, 1839 

perpusillus (Marsham, 1802) 

globosus (Hoffmann, J., 1803) 

Tribe PLEGADERINI Portevin, 1929 

PLEGADERUS Erichson, 1834 

dissectus Erichson, 1839 

vulneratus (Panzer, 1796) 

Tribe ACRITINI Wenzel, 1944 

ACRITUS LeConte, 1853 

Subgenus ACRITUS LeConte, 1853 

nigricornis (Hoffmann, J., 1803) 

minutus (Paykull, 1811) non (Herbst, 1792) 

Subgenus PYCNACRITUS Casey, 1916 

homoeopathicus Wollaston, 1857 

rhenanus Fuss, 1868 

AELETES Horn, 1873 

atomarius (Aubé, 1842) 

HALACRITUS Schmidt, J., 1893 

ACRITUS sensu Fowler, 1889 partim non LeConte, 

1853 

punctum (Aubé, 1843) 

26 

Tribe TERETRIINI Bickhardt, 1914 

TERETRIUS Erichson, 1834 

fabricii Mazur, 1972 

picipes sensu (Fabricius, 1792) non (Olivier, 1789) 

Subfamily SAPRININAE Blanchard, 1845 

SAPRINUS Erichson, 1834 

aeneus (Fabricius, 1775) 

immundus (Gyllenhal, 1827) 

planiusculus Motschulsky, 1849 

cuspidatus Ihssen, 1949 

semistriatus (Scriba, 1790) 

nitidulus (Fabricius, 1801) 

subnitescens Bickhardt, 1909 

virescens (Paykull, 1798) 

HYPOCACCUS Thomson, C.G., 1867 

SAPRINUS sensu Fowler, 1889 partim non 


Subgenus HYPOCACCUS Thomson, C.G., 1867 

crassipes (Erichson, 1834) 94 

metallicus (Herbst, 1792) 

rugiceps (Duftschmid, 1805) 

quadristriatus sensu (Hoffmann, J., 1803) non 

(Thunberg, 1794) 

rugifrons (Paykull, 1798) 

Subgenus BAECKMANNIOLUS Reichardt, 1926 

PACHYLOPUS sensu auctt. non Erichson, 1834 

dimidiatus (Illiger, 1807) 

ssp. maritimus (Stephens, 1830) 

GNATHONCUS Jacquelin du Val, 1857 

buyssoni Auzat, 1917 

communis (Marseul, 1862) 

schmidti Reitter, 1894 

nidicola Joy, 1907 

nannetensis (Marseul, 1862) 

rotundatus (Kugelann, 1792) 

nanus (Scriba, 1790) non (Piller & Mitterpacher, 

1783) 

punctulatus Thomson, C.G., 1862 

MYRMETES Marseul, 1862 

paykulli Kanaar, 1979 

piceus (Paykull, 1809) non (Marsham, 1802) 

Subfamily DENDROPHILINAE Reitter, 1909 

Tribe DENDROPHILINI Reitter, 1909 

DENDROPHILUS Leach, 1817 

punctatus (Herbst, 1792) 

pygmaeus (Linnaeus, 1758) 

xavieri Marseul, 1873 

KISSISTER Marseul, 1862 

CARCINOPS sensu Fowler, 1889 partim non 

Marseul, 1855 

minimus (Laporte, 1840)

Tribe PAROMALINI Reitter, 1909 

CARCINOPS Marseul, 1855 

pumilio (Erichson, 1834) 

quatuordecimstriatus (Stephens, 1835) 

PAROMALUS Erichson, 1834 

MICROLOMALUS Lewis, 1907 

flavicornis (Herbst, 1792) 

parallelepipedus (Herbst, 1792) 


Subfamily ONTHOPHILINAE MacLeay, 1819 

ONTHOPHILUS Leach, 1817 

punctatus (Müller, O.F., 1776) 

sulcatus (Geoffroy in Fourcroy, 1785) 

globulosus sensu Fowler, 1889 non (Olivier, 1789) 

striatus (Forster, 1771) 

Subfamily TRIBALINAE Bickhardt, 1914 

EPIERUS Erichson, 1834 

comptus Erichson, 1834 95 

Subfamily HISTERINAE Gyllenhal, 1808 

MARGARINOTUS Marseul, 1854 

HISTER sensu Fowler, 1889 partim non Linnaeus, 

1758 

Subgenus PTOMISTER Houlbert & Monnot, 1923 

brunneus (Fabricius, 1775) 

impressus (Fabricius, 1798) 

cadaverinus (Hoffmann, J., 1803) 

merdarius (Hoffmann, J., 1803) 

striola (Sahlberg, C.R., 1819) 

ssp. succicola (Thomson, C.G., 1862) 

Subgenus STENISTER Reichardt, 1926 

obscurus (Kugelann, 1792) 

stercorarius (Hoffmann, J., 1803) 

Subgenus PARALISTER Bickhardt, 1917 

neglectus (Germar, 1813) 

purpurascens (Herbst, 1792) 

ventralis (Marseul, 1854) 

carbonarius sensu auctt. Brit. non (Hoffmann, J., 

1803) 

Subgenus PROMETHISTER Kryzhanovskij, 1966 

GRAMMOSTETHUS sensu auctt. partim non Lewis, 

1906 

marginatus (Erichson, 1834) 

HISTER Linnaeus, 1758 

bissexstriatus Fabricius, 1801 

illigeri Duftschmid, 1805 

sinuatus sensu Illiger, 1798 non Fabricius, 1792 

quadrimaculatus Linnaeus, 1758 

gagates Illiger, 1807 

aethiops Heer, 1841 

quadrinotatus Scriba, 1790 

unicolor Linnaeus, 1758 

27 

Family HYDRAENIDAE Mulsant, 1844 

ATHOLUS Thomson, C.G., 1859 

HISTER sensu Fowler, 1889 partim non Linnaeus, 

1758 

PERANUS Lewis, 1906 

bimaculatus (Linnaeus, 1758) 

duodecimstriatus (Schrank, 1781) 

HOLOLEPTA Paykull, 1811 

plana (Sulzer, 1776) 96 

Subfamily HAETERIINAE Marseul, 1857 

HAETERIUS Dejean, 1833 

HETAERIUS Erichson, 1834 

ferrugineus (Olivier, 1789) 

Superfamily STAPHYLINOIDEA Latreille, 1802 

15. Family HYDRAENIDAE Mulsant, 1844 


Subfamily HYDRAENINAE Mulsant, 1844 

HYDRAENA Kugelann, 1794 

britteni Joy, 1907 

flavipes Sturm, 1836 

minutissima sensu Stephens, 1829 non (Weber & 

Mohr, 1804) 97 

?atricapilla (Waterhouse, G.R., 1833) 

gracilis Germar, 1824 

elongata Curtis, 1830 

nigrita Germar, 1824 

palustris Erichson, 1837 

pulchella Germar, 1824 

pygmaea Waterhouse, G.R., 1833 

riparia Kugelann, 1794 

rufipes Curtis, 1830 

angustata sensu auctt. Brit. non Sturm, 1836 

longior Rey, 1885 

testacea Curtis, 1830 

LIMNEBIUS Leach, 1815 

LIMNOBIUS auctt. (misspelling) 

aluta Bedel, 1881 

picinus sensu auctt. Brit. partim non (Marsham, 

1802) 

minutissimus sensu Balfour-Browne, J., 1938 non 

(Germar, 1824) 

crinifer Rey, 1885 98 

nitidus (Marsham, 1802) 

picinus (Marsham, 1802) 

papposus Mulsant, 1844 

truncatellus (Thunberg, 1794) 

Subfamily OCHTHEBIINAE Thomson, C.G., 1859 

ENICOCERUS Stephens, 1829 99 

HENICOCERUS Agassiz, 1846 

OCHTHEBIUS sensu auctt. partim non Leach, 1815 

exsculptus (Germar, 1824) 

viridiaeneus Stephens, 1829

Family PTILIIDAE Erichson, 1845 Checklist of Beetles of the British Isles, 2012 Edition 

gibsoni Curtis, 1830 

tristis Curtis, 1830 

sulcicollis Sturm, 1836 

lividipes Fairmaire & Laboulbène, 1854 

OCHTHEBIUS Leach, 1815 

Subgenus ASIOBATES Thomson, C.G., 1859 

auriculatus Rey, 1886 

bicolon Germar, 1824 

rufimarginatus Stephens, 1829 

dilatatus Stephens, 1829 

bicolon sensu Stephens, 1829 non Germar, 1824 

impressicollis Laporte, 1840 

kaninensis sensu Hellén, 1946 non Poppius, 1909 

Subgenus HOMALOCHTHEBIUS Kuwert, 1887 

aeneus Stephens, 1835 

lejolisii Mulsant & Rey, 1861 

lejolisi auctt. (misspelling) 

subinteger sensu auctt. non Mulsant & Rey, 1861 

minimus (Fabricius, 1792) 

pygmaeus sensu auctt. Brit. non (Fabricius, 1792) 

impressus (Marsham, 1802) 

Subgenus HYMENODES Mulsant, 1844 100 

nanus Stephens, 1829 


nilssoni Hebauer, 1986 101 

poweri Rye, 1869 

metallescens sensu Balfour-Browne, J., 1958 non 

Rosenhauer, 1847 

punctatus Stephens, 1829 

Subgenus OCHTHEBIUS Leach, 1815 102 

lenensis Poppius, 1907 

marinus (Paykull, 1798) 

pusillus Stephens, 1835 

margipallens (Latreille, 1807) non (Marsham, 1802) 

viridis sensu Fowler & Donisthorpe, 1913 non 

Peyron, 1858 

viridis Peyron, 1858 

margipallens sensu Fowler & Donisthorpe, 1913 

non Latreille, 1807 

ssp. viridis Peyron, 1858 

ssp. fallaciosus Ganglbauer, 1901 

AULACOCHTHEBIUS Kuwert, 1887 99 

OCHTHEBIUS sensu auctt. partim non Leach, 1815 

exaratus (Mulsant, 1844) 

28 

16. Family PTILIIDAE Erichson, 1845 

Family author: C. Johnson 

Higher classification, nomenclature and synonymy 

follow that in the Palaearctic catalogue (Johnson, 

2004), except that the listing below follows what 

seems to me to be a more natural system, instead 

of an alphabetical one. When thorough 

phylogenetic studies on the family have been 

carried out, however, we may expect further 

changes in higher classification. Regarding 

synonymy, only important foreign names are 

included below, but an attempt has been made to 

include all names in the British literature, 

especially misidentifications and misspellings. 

Subfamily NOSSIDIINAE Sörensson, in prep. 

NOSSIDIUM Erichson, 1845 

pilosellum (Marsham, 1802) 

brunneum (Marsham, 1802) 

nitidulum (Marsham, 1802) 

Subfamily PTILIINAE Erichson, 1845 

Tribe PTENIDIINI Flach, 1889 

PTENIDIUM Erichson, 1845 103 

ANISARTHRIA Stephens, 1830 

Subgenus GRESSNERIUM Ganglbauer, 1899 

gressneri Erichson, 1845 

Subgenus MATTHEWSIUM Flach, 1889 

laevigatum Erichson, 1845 

turgidum Thomson, C.G., 1855 

Subgenus WANKOWIZIUM Flach, 1889 

brenskei Flach, 1887 

intermedium Wankowicz, 1869 

vankoviezii Matthews, A., 1870 

wankowiezii auctt. (misspelling) 

Subgenus PTENIDIUM Erichson, 1845 

formicetorum Kraatz, 1851 

myrmecophilum (Motschulsky, 1845) non (Allibert, 

1844) 

kraatzii Matthews, A., 1872 

kraatzi Matthews, A., 1872 (misspelling) 

nigrifrons Britten, 1926 104 

fuscicorne Erichson, 1845 

picipes Matthews, A., 1860 

rugosum Britten, 1910 

longicorne Fuss, 1868 

brisoutii Matthews, A., 1872 

brisouti auctt. (misspelling) 

punctatum (Gyllenhal, 1827) 

pusillum (Gyllenhal, 1808) 103 

melas (Marsham, 1802) 

perpusillum (Marsham, 1802) 

nitidum (Stephens, 1830) 

punctulum (Stephens, 1830) 

apicale Erichson, 1845 

terminale Haldeman, 1848 

atamaroides Motschulsky, 1869 

atomaroides Motschulsky, 1869 (misspelling)

evanescens sensu Matthews, A., 1872 ? (Marsham, 

1802) 

Subgenus GILLMEISTERIUM Flach, 1889 

nitidum (Heer, 1841) 

minutissimum (Stephens, 1830) 

laevigatum sensu Matthews, A., 1872 non Erichson, 

1845 

Tribe PTILIINI Erichson, 1845 

EURYPTILIUM Matthews, A., 1872 

gillmeisteri Flach, 1889 105 

saxonicum (Gillmeister, 1845) 

marginatum sensu auctt. non Aubé, 1850 

PTILIOLA Haldeman, 1848 

NANOPTILIUM Flach, 1889 106 

brevicollis (Matthews, A., 1860) 

kunzei (Heer, 1841) 

nana (Stephens, 1830) 

trisulcata (Stephens, 1830) 

rugulosa (Allibert, 1844) 

PTILIOLUM Flach, 1888 

Subgenus PTILIOLUM Flach, 1888 

fuscum (Erichson, 1845) 

spencei sensu auctt. Brit. partim non (Allibert, 1844) 

flachi (Reitter, 1909) 

marginatum (Aubé, 1850) 

spencei sensu auctt. Brit. partim non (Allibert, 1844) 

lederi (Flach, 1888) 

sahlbergi (Flach, 1888) 

spencei (Allibert, 1844) 

angustatum (Erichson, 1845) 

oblongum (Gillmeister, 1845) 

foersteri (Matthews, A., 1872) 

Subgenus EUPTILIUM Flach, 1889 

caledonicum (Sharp, 1871) 

croaticum (Matthews, A., 1872) 

schwarzi (Flach, 1887) 

asperum (Britten, 1917) 

ACTIDIUM Matthews, A., 1868 

aterrimum (Motschulsky, 1845) 

concolor (Sharp, 1867) 

coarctatum (Haliday, 1855) 

OLIGELLA Motschulsky, 1869 

foveolata (Allibert, 1844) 

?minima (Herbst, 1793) 

excavata (Erichson, 1845) 

clandestina (Haliday, 1855) 

insignis (Matthews, A., 1861) 

intermedia Besuchet, 1971 

PTILIUM Gyllenhal, 1827 107 

affine Erichson, 1845 

incognitum Matthews, A., 1889 

caesum Erichson, 1845 108 

exaratum (Allibert, 1844) 

canaliculatum Erichson, 1845 

horioni Rosskothen, 1934 

exaratum sensu auctt. Brit. partim non Allibert, 1844 

29 

myrmecophilum (Allibert, 1844) 

inquilinum Erichson, 1845 

Family PTILIIDAE Erichson, 1845 

MILLIDIUM Motschulsky, 1855 109 

PTILIUM sensu auctt. partim non Gyllenhal, 1827 

minutissimum (Ljungh, 1804) 

trisulcatum (Aubé, 1833) 

MICROPTILIUM Matthews, A., 1872 110 

palustre Kuntzen, 1914 111 

pulchellum (Allibert, 1844) 

Tribe PTINELLINI Reitter, 1906 (1891) 

MICRIDIUM Motschulsky, 1869 112 

halidaii (Matthews, A., 1868) 

halidayi auctt. (misspelling) 

PTINELLA Motschulsky, 1844 

PLITIUM Besuchet, 1971 

aptera (Guérin-Méneville, 1839) 

tenella sensu Matthews, A., 1872 non Erichson, 

1845 

angustula sensu Matthews, A., 1872 non 

Gillmeister, 1845 

tenuis Csiki, 1911 

britannica Matthews, A., 1858 

brittannica auctt. (misspelling) 

cavelli (Broun, 1893) 

denticollis (Fairmaire, 1858) 

maria Matthews, A. 1862 

mariae auctt. (misspelling) 

errabunda Johnson, 1975 

limbata (Heer, 1841) 

testacea (Heer, 1841) 

proteus Matthews, A., 1862 

simsoni (Matthews, A., 1878) 

subvariolosa (Britten, 1932) 113 

flaviventris sensu (Donisthorpe, 1939) non 

(Motschulsky, 1869) 

taylorae Johnson, 1977 

PTERYX Matthews, A., 1858 

suturalis (Heer, 1841) 

mutabilis Matthews, A., 1858 

Subfamily ACROTRICHINAE Reitter, 1909 (1856) 

Tribe NEPHANINI Portevin, 1929 

NEPHANES Thomson, C.G., 1859 

TITAN Matthews, A., 1858 114 

ELACHYS Matthews, A., 1860 

titan (Newman, 1834) 

abbreviatella (Heer, 1841) 

curta (Allibert, 1844) 

SMICRUS Matthews, A., 1872 

MICRUS sensu Matthews, A., 1858 non 


filicornis (Fairmaire & Laboulbène, 1855)

Family PTILIIDAE Erichson, 1845 Checklist of Beetles of the British Isles, 2012 Edition 

BAEOCRARA Thomson, C.G., 1859 

BOEOCRARA auctt. (misspelling) 

variolosa (Mulsant & Rey, 1861) 

litoralis (Thomson, C.G., 1855) non (Motschulsky, 

1845) 

thomsoni (Sharp, 1866) 

Tribe ACROTRICHINI Reitter, 1909 (1856) 

ACTINOPTERYX Matthews, A., 1872 

fucicola (Allibert, 1844) 

mollis (Haliday, 1855) 

ACROTRICHIS Motschulsky, 1848 115 

ACRATRICHIS auctt. (misspelling) 

TRICHOPTERYX Kirby, 1826 non Hübner, 1825 

Subgenus ACROTRICHIS Motschulsky, 1848 

arnoldi Rosskothen, 1935 

atomaria (De Geer, 1774) 

minima (Marsham, 1802) non (Herbst, 1793) 

minutissima sensu (Marsham, 1802) ? (Linnaeus, 

1767) 

minuta (Stephens, 1830) 

convexa (Matthews, A., 1858) 

convexiuscula sensu (Matthews, A., 1872) non 

Motschulsky, 1851 116 

thoracica sensu (Matthews, A., 1872) non (Waltl, 

1838) 

seminitens (Matthews, A., 1877) 

brevicornis sensu (Fowler, 1884) non (Motschulsky, 

1868) 

brevipennis (Erichson, 1845) 

clavipes (Gillmeister, 1845) 

kirbii (Matthews, A., 1865) 116 

carbonaria (Matthews, A., 1873) 116 

cephalotes (Allibert, 1844) 117 

chevrolatii sensu auctt. non (Allibert, 1844) 

chevrolati auctt. (misspelling) 

pygmaea (Erichson, 1845) 

parallelogramma (Gillmeister, 1845) 

cognata (Matthews, A., 1877) 

platonoffi Renkonen, 1945 

danica Sundt, 1958 

atomaria sensu Joy, 1932 non (De Geer, 1774) 

dispar (Matthews, A., 1865) 

bovina sensu auctt. Brit. non (Motschulsky, 1845) 

fascicularis (Herbst, 1793) 

cantiana (Matthews, A., 1871) 116 

laetitiae (Matthews, A., 1873) 116 

laetitia auctt. (misspelling) 

henrici (Matthews, A., 1872) 

fratercula (Matthews, A., 1878) 

insularis (Mäklin, 1852) 

intermedia (Gillmeister, 1845) 118 

suffocata (Haliday, 1855) 116 

lata sensu (Matthews, A., 1865) non (Motschulsky, 

1845) 

josephi (Matthews, A., 1872) 119 

subcognata Johnson, 1975 

lucidula Rosskothen, 1935 

montandonii (Allibert, 1844) 120 

montandoni auctt. (misspelling) 116 

picicornis (Mannerheim, 1843) 

rivularis (Allibert, 1844) 

longicornis (Mannerheim, 1844) 

30 

similis (Gillmeister, 1845) 

jansoni (Matthews, A., 1866) 116 

angusta (Matthews, A., 1889) 116 

norvegica Strand, A., 1941 

parva Rosskothen, 1935 

silvatica sensu Sundt, 1958 partim non Rosskothen, 

1935 

pumila (Erichson, 1845) 

kirbii sensu (Matthews, A., 1865) partim 116 

fuscula (Matthews, A., 1871) 116 

guerinii sensu (Matthews, A., 1872) partim ? 

(Allibert, 1844) 116 

guerini auctt. (misspelling) 

longicornis sensu Matthews, A., 1872 non 

Mannerheim, 1844 

brevipennis sensu Joy, 1932 partim non (Erichson, 

1845) 

rosskotheni Sundt, 1971 

fraterna Johnson, 1975 

rugulosa Rosskothen, 1935 

volans sensu (Matthews, A., 1878) non 


sjobergi sensu Johnson, 1967 non Sundt, 1958 121 

sjoebergi auctt. (misspelling) 

sericans (Heer, 1841) 

picicornis sensu auctt. non (Mannerheim, 1843) 

depressa (Gillmeister, 1845) 

bovina (Motschulsky, 1845) 

brevis (Motschulsky, 1845) 116 

obscoena Wollaston, 1857 

obscaena auctt. (misspelling) 

ambigua (Matthews, A., 1865) 116 

chevrierii sensu (Matthews, A., 1866) non ? (Allibert, 

1844) 

waterhousii (Matthews, A., 1866) 116 

edithia (Matthews, A., 1871) 116 

longula (Matthews, A., 1871) 116 

guerinii sensu (Matthews, A., 1872) partim ? 

(Allibert, 1844) 116 

guerini auctt. (misspelling) 

poweri (Matthews, A., 1872) 

chevrolatii sensu Johnson, 2003 ? (Allibert, 1844) 

116 

silvatica Rosskothen, 1935 

sitkaensis (Motschulsky, 1845) 

fratercula sensu auctt. non (Matthews, A., 1878) 

strandi Sundt, 1958 

laetitiae sensu (Fowler, 1879) non (Matthews, A., 

1873) 116 

thoracica (Waltl, 1838) 

attenuata (Gillmeister, 1845) 116 

anthracina (Matthews, A., 1865) 

brevicornis Motschulsky, 1868 

Subgenus CTENOPTERYX Flach, 1889 

grandicollis (Mannerheim, 1844) 

sanctaehelenae Johnson, 1972 122

17. Family LEIODIDAE Fleming, 1821 

Family author: J. Cooter 

Subfamily LEIODINAE Fleming, 1821 123 

Tribe SOGDINI Lopatin, 1961 

SOGDA Lopatin, 1961 

HYDNOBIUS sensu auctt. partim non Schmidt, 

W.L.E., 1841 

TRICHOHYDNOBIUS Vogt, 1961 

suturalis (Zetterstedt, 1828) 

perrisii (Fairmaire, 1855) 

perrisi auctt. (misspelling) 

HYDNOBIUS Schmidt, W.L.E., 1841 

latifrons (Curtis, 1840) 

strigosus Schmidt, W.L.E., 1841 

punctatus (Sturm, 1807) 

punctatissimus (Stephens, 1829) 

spinipes (Gyllenhal, 1813) 

TRIARTHRON Märkel, 1840 

maerkelii Märkel, 1840 

maerkeli auctt. (misspelling) 

Tribe LEIODINI Fleming, 1821 

LEIODES Latreille, 1796 

LIODES Gemminger & Harold, 1868 (misspelling) 

OOSPHAERULA Ganglbauer, 1899 

TRICHOSPHAERULA Fleischer, 1904 

badia (Sturm, 1807) 

similata (Rye, 1870) 

calcarata (Erichson, 1845) 

polita (Marsham, 1802) non (Fuessly, 1775) 

?maritima Stephens, 1832 

nigrescens Fleischer, 1906 

ciliaris (Schmidt, W.L.E., 1841) 

cinnamomea (Panzer, 1793) 

lycoperdi Stephens, 1829 

stephensi Stephens, 1832 

ferruginea (Fabricius, 1787) 

?nigricollis Stephens, 1832 

ovalis (Schmidt, W.L.E., 1841) 

scita (Erichson, 1845) 

flavescens (Schmidt, W.L.E., 1841) 

stenocoryphe Joy, 1911 

furva (Erichson, 1845) 

gallica (Reitter, 1884) 

brunnea sensu auctt. Brit. non (Sturm, 1807) 

gyllenhalii Stephens, 1829 

brunnea (Gyllenhal, 1810) non (Sturm, 1807) 

parvula (Sahlberg, C.R., 1833) 

parvula sensu Fowler, 1889 partim 

litura Stephens, 1832 

punctulata sensu auctt. non Gyllenhal, 1810 

ornata (Fairmaire, 1855) 

longipes (Schmidt, W.L.E., 1841) 

curta (Fairmaire & Laboulbène, 1855) 

curta var. donisthorpei Fleischer, 1911 

lucens (Fairmaire, 1855) 

picea Illiger sensu Stephens, 1829 non (Panzer, 

1797) 

oblonga sensu auctt. non Erichson, 1845 

31 

Family LEIODIDAE Fleming, 1821 

lunicollis (Rye, 1872) 

macropus (Rye, 1873) 

curvipes sensu Reitter, 1884 non (Schmidt, W.L.E., 

1841) 

nigrita (Schmidt, W.L.E., 1841) 

scita sensu Reitter, 1884 non Erichson, 1845 

obesa (Schmidt, W.L.E., 1841) 

dubia sensu Joy, 1911 partim non (Kugelann, 1794) 

oblonga (Erichson, 1845) 

grandis (Fairmaire & Laboulbène, 1855) 

anglica (Rye, 1873) 

picea (Panzer, 1797) 

rufipennis (Paykull, 1798) 

thoracica Stephens, 1829 

maxillosa Stephens, 1829 

?testacea Stephens, 1829 

dyllwynii Stephens, 1832 

?suturalis Stephens, 1832 

clavicornis (Rye, 1875) 

davidiana (Joy, 1911) 

algirica sensu Donisthorpe, 1911 non (Rye, 1875) 

dubia sensu auctt. partim non (Kugelann, 1794) 

dubia var. subglobosa sensu Joy, 1911 non Reitter, 

1884 

dubia var. bicolor sensu Joy, 1911 non (Schmidt, 

W.L.E., 1841) 

rugosa Stephens, 1829 

arenaria Stephens, 1832 

silesiaca (Kraatz, 1852) 

strigipennis Daffner, 1983 

flavicornis sensu auctt. Brit. non (Brisout de 

Barneville, 1863) 124 

parvula sensu Fowler, 1889 partim non (Sahlberg, 

C.R., 1833) 

triepkii (Schmidt, W.L.E., 1841) 

triepkei auctt. (misspelling) 

pallens sensu auctt. Brit. non (Sturm, 1807) 125 

rubiginosa sensu auctt. Brit. non (Schmidt, W.L.E., 

1841) 

curvipes (Schmidt, W.L.E., 1841) 

rotundata sensu Donisthorpe, 1933 non (Erichson, 

1845) 

LIOCYRTUSA Daffner, 1982 

CYRTUSA sensu auctt. non Erichson, 1842 

minuta (Ahrens, 1812) 

vittata (Curtis, 1840) 

pauxilla (Schmidt, W.L.E., 1841) 

Tribe PSEUDOLIODINI Portevin, 1926 

AGARICOPHAGUS Schmidt, W.L.E., 1841 

cephalotes Schmidt, W.L.E., 1841 

conformis Erichson, 1845 

COLENIS Erichson, 1845 

immunda (Sturm, 1807) 

dentipes (Gyllenhal, 1810) 

aciculata (Stephens, 1829) 

Tribe AGATHIDIINI Westwood, 1838 

ANISOTOMA Panzer, 1797 

LIODES Erichson, 1845 non von Heyden, 1826 

castanea (Herbst, 1792) 

glabra (Fabricius, 1792)

Family LEIODIDAE Fleming, 1821 Checklist of Beetles of the British Isles, 2012 Edition 

axillaris sensu Stephens, 1829 non Gyllenhal, 1810 

humeralis (Fabricius, 1792) 

armata Stephens, 1829 

orbicularis (Herbst, 1792) 

seminulum (Kugelann, 1794) 

AMPHICYLLIS Erichson, 1845 

globus (Fabricius, 1792) 

ruficollis (Olivier, 1792) 

ferrugineus (Sturm, 1807) 

AGATHIDIUM Panzer, 1797 

Subgenus CYPHOCEBLE Thomson, C.G., 1859 

arcticum Thomson, C.G., 1862 

rhinoceros Sharp, 1866 

nigrinum Sturm, 1807 

staphylaeum (Gyllenhal, 1810) 

ferrugineum sensu Stephens, 1829 non Sturm, 

1807 

seminulum sensu Stephens, 1829 partim non 


Subgenus NEOCEBLE des Gozis, 1886 

confusum Brisout de Barneville, 1863 

piceum Thomson, C.G., 1862 non Erichson, 1845 

clypeatum Sharp, 1866 

convexum Sharp, 1866 

piceum Erichson, 1845 non Melsheimer, 1844 

globosum Mulsant & Rey, 1861 non (Kugelann, 

1794) 

marginatum Sturm, 1807 

seminulum sensu Stephens, 1829 partim non 


mandibulare sensu Stephens, 1829 partim non 

Sturm, 1807 

pumilum Hardy, 1852 

nigripenne (Fabricius, 1792) 


Sturm, 1807 

rotundatum (Gyllenhal, 1827) 

lycogalae Hardy, 1852 

sphaerulum Reitter, 1898 

reitteri Ganglbauer, 1899 

varians Beck, 1817 


Sturm, 1807 

Subgenus AGATHIDIUM Panzer, 1797 

atrum (Paykull, 1798) 

atrum sensu Stephens, 1829 partim 

globus sensu Stephens, 1829 partim non (Paykull, 

1798) 

rufipes Stephens, 1832 

laevigatum Erichson, 1845 

atrum sensu Stephens, 1829 partim 

orbiculare sensu Stephens, 1829 non (Herbst, 

1792) 

nigrinum sensu Stephens, 1829 non Sturm, 1807 

affine sensu Stephens, 1829 

pisanum Brisout de Barneville, 1872 

badium sensu auctt. Brit. non Erichson, 1845 126 

seminulum (Linnaeus, 1758) 

32 

Subfamily COLONINAE Horn, 1880 (1859) 

COLON Herbst, 1797 

KOLON Herbst, 1797 127 

Subgenus EURYCOLON Ganglbauer, 1899 

latum Kraatz, 1850 

rufescens Kraatz, 1850 

Subgenus MYLOECHUS Latreille, 1807 

angulare Erichson, 1837 

appendiculatum (Sahlberg, C.R., 1822) 

calcaratum sensu auctt. Brit. non Erichson, 1837 

denticulatum Kraatz, 1850 

microps Czwalina, 1881 

brunneum (Latreille, 1807) 

dentipes (Sahlberg, C.R., 1822) 

zebei Kraatz, 1854 

barnevillei sensu auctt. Brit. non Kraatz, 1850 

Subgenus COLON Herbst, 1797 

serripes (Sahlberg, C.R., 1822) 

fusculum Erichson, 1837 

puncticolle sensu auctt. Brit. partim non Kraatz, 

1850 

viennense Herbst, 1797 

puncticolle sensu auctt. Brit. partim non Kraatz, 

1850 

Subfamily CHOLEVINAE Kirby, 1837 

Tribe ANEMADINI Hatch, 1928 

NEMADUS Thomson, C.G., 1867 

colonoides (Kraatz, 1851) 

Tribe CHOLEVINI Kirby, 1837 

NARGUS Thomson, C.G., 1867 

Subgenus NARGUS Thomson, C.G., 1867 

velox (Spence, 1813) 

Subgenus DEMOCHRUS Thomson, C.G., 1867 

anisotomoides (Spence, 1813) 

wilkinii (Spence, 1813) 

wilkini auctt. (misspelling) 

CHOLEVA Latreille, 1796 

Subgenus CHOLEVOPSIS Jeannel, 1922 

spadicea (Sturm, 1839) 

Subgenus CHOLEVA Latreille, 1796 

agilis (Illiger, 1798) 

angustata (Fabricius, 1781) 

cisteloides sensu auctt. Brit. partim non (Frölich, 

1799) 

cisteloides (Frölich, 1799) 

elongata (Paykull, 1798) 128 

fagniezi Jeannel, 1922 

angustata sensu auctt. Brit. partim non (Fabricius, 

1781) 

glauca Britten, 1918 

jeanneli Britten, 1922 

sturmi sensu auctt. Brit. non Brisout de Barneville, 

1863 

lederiana Reitter, 1902 129

agilis sensu auctt. Brit. ?partim non (Illiger, 1798) 

septentrionis Jeannel, 1923 

oblonga Latreille, 1807 

intermedia (Kraatz, 1852) 

SCIODREPOIDES Hatch, 1933 

SCIODREPA sensu auctt. non Thomson, C.G., 

1859 

fumatus (Spence, 1813) 

watsoni (Spence, 1813) 

CATOPS Paykull, 1798 

borealis Krogerus, 1931 130 

chrysomeloides (Panzer, 1798) 

coracinus Kellner, 1846 

fuliginosus Erichson, 1837 

fuscus (Panzer, 1794) 

grandicollis Erichson, 1837 

kirbii (Spence, 1813) 

kirbyi auctt. (misspelling) 

longulus Kellner, 1846 

morio (Fabricius, 1787) 

nigricans (Spence, 1813) 

nigriclavis Gerhardt, 1900 

nigrita Erichson, 1837 

tristis (Panzer, 1793) 131 

leachi (Spence, 1813) 

montivagus Heer, 1841 

CATOPIDIUS Jeannel, 1922 

depressus (Murray, 1856) 

PARABATHYSCIA Jeannel, 1908 

BATHYSCIA sensu auctt. partim non Schiødte, 

1849 

wollastoni (Janson, E.W., 1857) 

Tribe PTOMAPHAGINI Jeannel, 1911 

PTOMAPHAGUS Illiger, 1798 

medius Rey, 1889 

sericeus sensu auctt. Brit. partim non (Fabricius, 

1792) 

sericatus sensu auctt. Brit. non Chaudoir, 1845 

subvillosus (Goeze, 1777) 

sericeus (Fabricius, 1792) 

varicornis (Rosenhauer, 1847) 

variicornis auctt. (misspelling) 

Subfamily PLATYPSYLLINAE Ritsema, 1869 

LEPTINUS Müller, P.W.J., 1817 

testaceus Müller, P.W.J., 1817 

18. Family SILPHIDAE Latreille, 1806 


Subfamily SILPHINAE Latreille, 1806 

NECRODES Leach, 1815 

littoralis (Linnaeus, 1758) 

33 

THANATOPHILUS Leach, 1815 

dispar (Herbst, 1793) 

rugosus (Linnaeus, 1758) 

sinuatus (Fabricius, 1775) 

Family SILPHIDAE Latreille, 1806 

OICEOPTOMA Leach, 1815 

OECEOPTOMA auctt. (misspelling) 

THANATOPHILUS sensu Fowler, 1888 partim non 

Leach, 1815 

thoracicum (Linnaeus, 1758) 

ACLYPEA Reitter, 1884 

BLITOPHAGA Reitter, 1884 

opaca (Linnaeus, 1758) 

undata (Müller, O.F., 1776) 

reticulata (Fabricius, 1787) non (Linnaeus, 1767) 

DENDROXENA Motschulsky, 1858 

XYLODREPA Thomson, C.G., 1859 

quadrimaculata (Scopoli, 1772) 

quadripunctata sensu auctt. non (Linnaeus, 1758) 

SILPHA Linnaeus, 1758 

PHOSPHUGA Leach, 1817 

ABLATTARIA Reitter, 1884 

atrata Linnaeus, 1758 

paedemontana Fabricius, 1775 

brunnea Herbst, 1793 

ssp. atrata Linnaeus, 1758 

ssp. subrotundata (Leach, 1817) 

carinata Herbst, 1783 

griesbachiana Stephens, 1830 

laevigata Fabricius, 1775 

obscura Linnaeus, 1758 

tristis Illiger, 1798 

tyrolensis Laicharting, 1781 

nigrita Creutzer, 1799 

Subfamily NICROPHORINAE Kirby, 1837 

NICROPHORUS Fabricius, 1775 

NECROPHORUS auctt. (misspelling) 

germanicus (Linnaeus, 1758) 

humator (Gleditsch, 1767) 

interruptus Stephens, 1830 

fossor Erichson, 1837 

investigator Zetterstedt, 1824 

ruspator Erichson, 1837 

vespillo (Linnaeus, 1758) 

vespilloides Herbst, 1783 

mortuorum Fabricius, 1792 

vestigator Herschel, 1807 

19. Family STAPHYLINIDAE Latreille, 1802 

Family author: A,G. Duff 

Subfamily OMALIINAE MacLeay, 1825 132 

Tribe ANTHOPHAGINI Thomson, C.G., 1859 

ACIDOTA Stephens, 1829 

crenata (Fabricius, 1792)

Family STAPHYLINIDAE Latreille, 1802 Checklist of Beetles of the British Isles, 2012 Edition 

cruentata Mannerheim, 1830 

ANTHOBIUM Leach, 1819 133 

LATHRIMAEUM Erichson, 1839 

atrocephalum (Gyllenhal, 1827) 

unicolor (Marsham, 1802) 

ANTHOPHAGUS Gravenhorst, 1802 

Subgenus ANTHOPHAGUS Gravenhorst, 1802 


Subgenus PHAGANTHUS Mulsant & Rey, 1880 


ARPEDIUM Erichson, 1939 

EUCNECOSUM Reitter, 1909 134 

brachypterum (Gravenhorst, 1802) 

DELIPHRUM Erichson, 1839 

tectum (Paykull, 1789) 

GEODROMICUS Redtenbacher, 1857 135 

PSEPHIDONUS Gistel, 1856 

longipes (Mannerheim, 1830) 

globulicollis sensu auctt. Brit. non (Mannerheim, 

1830) 

nigrita (Müller, P.W.J., 1821) 

plagiatus sensu auctt. non (Fabricius, 1798) 

LESTEVA Latreille, 1796 136 

hanseni Lohse, 1953 

fontinalis sensu auctt. non Kiesenwetter, 1850 

longoelytrata (Goeze, 1777) 

monticola Kiesenwetter, 1847 

pubescens Mannerheim, 1830 

luctuosa sensu auctt. Brit. non Fauvel, 1871 

punctata Erichson, 1839 137 

villosus (Waltl, 1838) 

sicula Erichson, 1840 

ssp. heeri Fauvel, 1871 138 

OLOPHRUM Erichson, 1839 

assimile (Paykull, 1800) 

consimile (Gyllenhal, 1810) 

fuscum (Gravenhorst, 1806) 

nicholsoni Donisthorpe, 1910 

piceum (Gyllenhal, 1810) 

OROCHARES Kraatz, 1857 

angustatus (Erichson, 1840) 

PHILORINUM Kraatz, 1858 

sordidum (Stephens, 1834) 

PHYLLODREPOIDEA Ganglbauer, 1895 

crenata Ganglbauer, 1895 

Tribe CORYPHIINI Jakobson, 1908 

CORYPHIUM Stephens, 1834 

angusticolle Stephens, 1834 

34 

EUDECTUS Redtenbacher, 1857 

whitei Sharp, 1871 

giraudi sensu auctt. Brit. non Redtenbacher, 1857 

Tribe EUSPHALERINI Hatch, 1957 

EUSPHALERUM Kraatz, 1858 

ANTHOBIUM sensu auctt. non Leach, 1819 

luteum (Marsham, 1802) 

ophthalmicum (Paykull, 1800) non (Scopoli, 1763) 

minutum (Fabricius, 1792) 

primulae (Stephens, 1834) 

sorbi (Gyllenhal, 1810) 137 

testaceum (Gravenhorst, 1806) 

sorbicola (Kangas, 1941) 

lapponicum sensu auctt. non (Mannerheim, 1830) 

torquatum (Marsham, 1802) 

Tribe HADROGNATHINI Portevin, 1929 

HADROGNATHUS Schaum, 1852 

longipalpis (Mulsant & Rey, 1851) 139 

longipalpus auctt. (misspelling) 

Tribe OMALIINI MacLeay, 1825 

ACROLOCHA Thomson, C.G., 1858 140 

ELONIUM Leach, 1819 

minuta (Olivier, 1795) 

striata (Gravenhorst, 1802) 141 

sulcula (Stephens, 1834) 

striata sensu auctt. Brit. non (Gravenhorst, 1802) 

ACRULIA Thomson, C.G., 1858 

inflata (Gyllenhal, 1813) 

DROPEPHYLLA Mulsant & Rey, 1880 142 

devillei Bernhauer, 1902 

grandiloqua Luze, 1910 143 

propinqua Bernhauer, 1943 144 

gracilicornis (Fairmaire & Laboulbène, 1856) 

heerii (Heer, 1841) 145 

ioptera (Stephens, 1832) 

linearis sensu auctt. Brit. non (Zetterstedt, 1828) 

koltzei Jászay & Hlavac, 2006 146 

vilis (Erichson, 1840) 

HAPALARAEA Thomson, C.G., 1858 

pygmaea (Paykull, 1800) 147 

HYPOPYCNA Mulsant & Rey, 1880 

HAPALARAEA sensu auctt. partim non Thomson, 

C.G., 1858 

rufula (Erichson, 1840) 

MICRALYMMA Westwood, 1838 

marinum (Strøm, 1783) 

OMALIUM Gravenhorst, 1802 

allardi Fairmaire & Brisout de Barneville, 1859 

caesum Gravenhorst, 1806 

excavatum Stephens, 1834

exiguum Gyllenhal, 1810 

italicum Bernhauer, 1902 

tricolor Mulsant & Rey, 1880 non Wollaston, 1865 

laeviusculum Gyllenhal, 1827 

laticolle Kraatz, 1858 

brevicolle sensu auctt. Brit. non Thomson, C.G., 

1884 

foraminosum sensu auctt. Brit. non Mäklin, 1852 

oxyacanthae Gravenhorst, 1806 

riparium Thomson, C.G., 1857 

rivulare (Paykull, 1789) 137 

cursor (Müller, O.F., 1776) 

rugatum Mulsant & Rey, 1880 

rugulipenne Rye, 1864 

septentrionis Thomson, C.G., 1856 

PARAPHLOEOSTIBA Steel, 1960 

gayndahensis (MacLeay, 1873) 148 

PHLOEONOMUS Heer, 1839 

punctipennis Thomson, C.G., 1867 

pusillus (Gravenhorst, 1806) 

PHLOEOSTIBA Thomson, C.G., 1858 149 

PHLOEONOMUS sensu auctt. partim non Heer, 

1839 

lapponica (Zetterstedt, 1838) 

plana (Paykull, 1792) 137 


PHYLLODREPA Thomson, C.G., 1859 150 

HAPALARAEA sensu auctt. partim non Thomson, 

C.G., 1858 

floralis (Paykull, 1789) 

nigra (Gravenhorst, 1806) 

puberula Bernhauer, 1903 

salicis (Gyllenhal, 1810) 

XYLODROMUS Heer, 1839 

concinnus (Marsham, 1802) 151 

brunnipennis sensu auctt. non (Stephens, 1834) 152 

depressus (Gravenhorst, 1802) 

testaceus (Erichson, 1840) 137 

pygmaeus (Gravenhorst, 1806) 

XYLOSTIBA Ganglbauer, 1895 153 

PHLOEONOMUS sensu auctt. partim non Heer, 

1839 

bosnica (Bernhauer, 1902) 154 

monilicornis sensu auctt. Brit. partim non (Gyllenhal, 

1810) 

monilicornis (Gyllenhal, 1810) 

Subfamily PROTEININAE Erichson, 1839 

MEGARTHRUS Stephens, 1829 

bellevoyei (Saulcy, 1862) 

affinis Miller, 1853 non Stephens, 1834 155 

denticollis (Beck, 1817) 

depressus (Paykull, 1789) 156 

sinuatocollis (Lacordaire, 1835) 157 

hemipterus (Illiger, 1794) 

prosseni Schatzmayr, 1904 

depressus sensu auctt. non (Paykull, 1789) 

35 

METOPSIA Wollaston, 1854 

clypeata (Müller, P.W.J., 1821) 

gallica (Koch, 1938) 

retusa (Stephens, 1834) 158 

Family STAPHYLINIDAE Latreille, 1802 

PROTEINUS Latreille, 1796 

atomarius Erichson, 1840 159 

brachypterus (Fabricius, 1792) 

crenulatus Pandellé, 1867 

limbatus sensu auctt. non Mäklin, 1852 

laevigatus Hochhuth, 1871 

macropterus sensu auctt. non (Gravenhorst, 

1806) 160 

serrifer Muona, 1977 

ovalis Stephens, 1834 

Subfamily MICROPEPLINAE Leach, 1815 

MICROPEPLUS Latreille, 1809 

ARRHENOPEPLUS Koch, 1937 161 

caelatus Erichson, 1839 

fulvus Erichson, 1840 

porcatus (Paykull, 1789) 

staphylinoides (Marsham, 1802) 

tesserula Curtis, 1828 

Subfamily PSELAPHINAE Latreille, 1802 162 

Supertribe BATRISITAE Reitter, 1882 

Tribe BATRISINI Reitter, 1882 

BATRISODES Reitter, 1882 

adnexus (Hampe, 1863) 

buqueti sensu auctt. Brit. non (Aubé, 1833) 

delaporti (Aubé, 1833) 

laporti auctt. (misspelling) 

venustus (Reichenbach, 1816) 

Supertribe CLAVIGERITAE Leach, 1815 

Tribe CLAVIGERINI Leach, 1815 

CLAVIGER Preyssler, 1790 

longicornis Müller, P.W.J., 1818 

testaceus Preyssler, 1790 

Supertribe EUPLECTITAE Streubel, 1839 

Tribe EUPLECTINI Streubel, 1839 

EUPLECTUS Leach, 1817 163 

EUPLECTOIDES Jeannel, 1954 

bescidicus Reitter, 1881 

bohemicus Machulka, 1935 

brunneus sensu Pearce, 1957 non (Grimmer, 1841) 

bonvouloiri Reitter, 1881 164 

ssp. rosae Raffray, 1910 165 

brunneus (Grimmer, 1841) 166 

decipiens Raffray, 1910 

duponti sensu Pearce, 1974 non Aubé, 1833 

duponti Aubé, 1833 

aubeanus sensu Pearce, 1974 non Reitter, 1881


infirmus Raffray, 1910 

boeticus Jeannel, 1950 

karstenii (Reichenbach, 1816) 

kirbii Denny, 1825 

kirbyi auctt. (misspelling) 

nanus sensu auctt. partim non (Reichenbach, 1816) 

mutator Fauvel, 1895 

fauveli Guillebeau, 1888 167 

falsus Bedel, 1906 

tomlini Joy, 1906 

nanus (Reichenbach, 1816) 

reichenbachi Leach, 1817 

carolae Allen, 1940 

piceus Motschulsky, 1835 

pearcei (Jeannel, 1954) 

punctatus Mulsant, 1861 168 

sanguineus Denny, 1825 

signatus (Reichenbach, 1816) 

tholini Guillebeau, 1888 169 

punctatus sensu auctt. partim non Mulsant, 1861 

PLECTOPHLOEUS Reitter, 1891 

erichsoni (Aubé, 1844) 170 

ssp. occidentalis Besuchet, 1969 

nitidus (Fairmaire, 1857) 

Tribe TRICHONYCHINI Reitter, 1882 

Subtribe BIBLOPORINA Park, 1951 

BIBLOPORUS Thomson, C.G., 1859 

bicolor (Denny, 1825) 

minutus Raffray, 1914 

bicolor sensu auctt. Brit. partim non (Denny, 1825) 

hoeglundi Palm, 1948 

sulcatus Jeannel, 1950 

Subtribe PANAPHANTINA Jeannel, 1950 

BIBLOPLECTUS Reitter, 1882 171 

ambiguus (Reichenbach, 1816) 

championi Jeannel, 1950 

delhermi Guillebeau, 1888 

pusillus sensu Jeannel, 1950 non (Denny, 1825) 

intermedius Pearce, 1951 

minutissimus (Aubé, 1833) 

pusillus (Denny, 1825) 

academicus Pearce, 1951 

therondi Besuchet, 1953 

spinosus Raffray, 1914 

ambiguus sensu Pearce, 1951 non (Reichenbach, 

1816) 

pusillus sensu Pearce, 1951 non (Denny, 1825) 

tenebrosus (Reitter, 1880) 

margaretae Sharp, 1916 

Subtribe TRICHONYCHINA Reitter, 1882 

AMAURONYX Reitter, 1882 

maerkelii (Aubé, 1844) 

TRICHONYX Chaudoir, 1845 

sulcicollis (Reichenbach, 1816) 

36 

Subtribe TRIMIINA Bowman, 1934 

TRIMIUM Aubé, 1833 

brevicorne (Reichenbach, 1816) 

Supertribe GONIACERITAE Reitter, 1882 (1872) 

Tribe BRACHYGLUTINI Raffray, 1904 

BRACHYGLUTA Thomson, C.G., 1859 

fossulata (Reichenbach, 1816) 

haematica (Reichenbach, 1816) 

helferi (Schmidt-Göbel, 1836) 

klimschi Holdhaus, 1902 172 

pandellei (Saulcy, 1876) 

cotus (Saulcy, 1876) 

simplicior Raffray, 1904 173 

haematica sensu auctt. partim non (Reichenbach, 

1816) 

sinuata (Aubé, 1833) 173 

haematica sensu auctt. partim non (Reichenbach, 

1816) 

waterhousei (Rye, 1869) 174 

depressa sensu auctt. non (Aubé, 1833) 

simplex (Waterhouse, G.R., 1862) non 


FAGNIEZIA Jeannel, 1950 175 

TRISSEMUS sensu auctt. partim non Jeannel, 1949 

impressa (Panzer, 1803) 

REICHENBACHIA Leach, 1826 

juncorum (Leach, 1817) 

RYBAXIS Saulcy, 1876 

longicornis (Leach, 1817) 

laminata (Motschulsky, 1836) 176 

Tribe BYTHININI Raffray, 1890 

BRYAXIS Kugelann, 1794 

bulbifer (Reichenbach, 1816) 

curtisii (Leach, 1817) 

puncticollis (Denny, 1825) 

validus (Aubé, 1844) 

BYTHINUS Leach, 1817 

burrellii Denny, 1825 

macropalpus Aubé, 1833 

TYCHOBYTHINUS Ganglbauer, 1896 

glabratus (Rye, 1870) 

ludyi (Reitter, 1882) 

Tribe TYCHINI Raffray, 1904 

TYCHUS Leach, 1817 

niger (Paykull, 1800) 

striola Guillebeau, 1888

Supertribe PSELAPHITAE Latreille, 1802 

Tribe PSELAPHINI Latreille, 1802 

PSELAPHAULAX Reitter, 1909 

dresdensis (Herbst, 1792) 

PSELAPHUS Herbst, 1792 

heisei Herbst, 1792 

Subfamily PHLOEOCHARINAE Erichson, 1839 

PHLOEOCHARIS Mannerheim, 1830 

subtilissima Mannerheim, 1830 

Subfamily TACHYPORINAE MacLeay, 1825 

Tribe MYCETOPORINI Thomson, C.G., 1859 

BOLITOBIUS Leach, 1819 

BRYOCHARIS Lacordaire, 1835 

castaneus (Stephens, 1832) 

analis sensu auctt. non (Fabricius, 1787) 177 

cingulatus (Mannerheim, 1830) 

BRYOPHACIS Reitter, 1909 178 

BRYOPORUS sensu auctt. partim non Kraatz, 1857 

crassicornis (Mäklin, 1847) 

maklini (Sahlberg, J., 1871) 

rugipennis sensu auctt. Brit. non (Pandellé, 

1869) 179 

BRYOPORUS Kraatz, 1857 

cernuus (Gravenhorst, 1806) 

merdarius (Olivier, 1795) non (Fabricius, 1775) 

ISCHNOSOMA Stephens, 1829 180 

MYCETOPORUS sensu auctt. partim non 


longicorne (Mäklin, 1847) 

splendidum (Gravenhorst, 1806) 

LORDITHON Thomson, C.G., 1858 

BOBITOBUS Tottenham, 1939 

BOLITOBIUS sensu auctt. non Leach, 1819 

exoletus (Erichson, 1839) 

lunulatus (Linnaeus, 1761) 

thoracicus (Fabricius, 1777) 

trinotatus (Erichson, 1839) 

MYCETOPORUS Mannerheim, 1830 

ambiguus Luze, 1901 181 

clavicornis sensu auctt. Brit. partim non (Stephens, 

1832) 

angularis Mulsant & Rey, 1853 

baudueri Mulsant & Rey, 1875 

hellieseni Strand, A., 1950 

bimaculatus Lacordaire, 1835 

clavicornis (Stephens, 1832) 

forticornis sensu auctt. Brit. non Fauvel, 1875 

despectus Strand, A., 1969 

lepidus sensu auctt. partim non (Gravenhorst, 1802) 

37 


longulus sensu auctt. partim non Mannerheim, 1830 

erichsonanus Fagel, 1965 

baudueri sensu auctt. partim non Mulsant & Rey, 

1875 

lepidus (Gravenhorst, 1806) 

brunneus sensu auctt. partim non (Paykull, 1789) 

longulus Mannerheim, 1830 

brunneus sensu auctt. partim non (Paykull, 1789) 

monticola Fowler, 1888 

nigricollis Stephens, 1835 

splendens (Marsham, 1802) non (Fabricius, 1792) 

piceolus Rey, 1883 

baudueri sensu auctt. partim non Mulsant & Rey, 

1875 

punctus (Gravenhorst, 1806) 

reichei (Pandellé, 1869) 181 

clavicornis sensu auctt. Brit. partim non (Stephens, 

1832) 

rufescens (Stephens, 1832) 

lucidus Erichson, 1839 

PARABOLITOBIUS Li, Zhao & Sakai, 2000 

BOLITOBIUS sensu auctt. partim non Leach, 1819 

inclinans (Gravenhorst, 1806) 182 

Tribe TACHYPORINI MacLeay, 1825 

CILEA Jacquelin du Val, 1856 

LEUCOPARYPHUS Kraatz, 1857 

silphoides (Linnaeus, 1767) 

LAMPRINODES Luze, 1901 

saginatus (Gravenhorst, 1806) 

SEPEDOPHILUS Gistel, 1856 

CONOSOMA sensu auctt. non Kraatz, 1857 

CONOSOMUS sensu auctt. non Motschulsky, 1857 

bipunctatus (Gravenhorst, 1802) 

constans (Fowler, 1888) 

testaceus sensu auctt. partim non (Fabricius, 1792) 

pubescens sensu auctt. partim non (Gravenhorst, 

1802) 

immaculatus (Stephens, 1832) 

littoreus (Linnaeus, 1758) 

lusitanicus Hammond, 1973 



1802) 

marshami (Stephens, 1832) 



1802) 

nigripennis (Stephens, 1832) 

lividus (Erichson, 1839) 

pedicularius (Gravenhorst, 1802) 

testaceus (Fabricius, 1792) 183 

pubescens (Gravenhorst, 1802) 

TACHINUS Gravenhorst, 1802 184 

DRYMOPORUS Thomson, C.G., 1859 185 


corticinus Gravenhorst, 1802 

collaris Gravenhorst, 1802 

elongatus Gyllenhal, 1810 

flavolimbatus Pandellé, 1869


humeralis Gravenhorst, 1802 

laticollis Gravenhorst, 1802 

lignorum (Linnaeus, 1758) 


marginellus (Fabricius, 1781) 

pallipes (Gravenhorst, 1806) 

benicki Ullrich, 1975 186 

proximus Kraatz, 1855 

rufipennis Gyllenhal, 1810 

rufipes (Linnaeus, 1758) 187 

signatus Gravenhorst, 1802 

scapularis Stephens, 1832 

subterraneus (Linnaeus, 1758) 

TACHYPORUS Gravenhorst, 1802 

atriceps Stephens, 1832 

chrysomelinus (Linnaeus, 1758) 

fasciatus Nicholson, 1911 

dispar (Paykull, 1789) 188 

chrysomelinus sensu auctt. partim non (Linnaeus, 

1758) 

formosus Matthews, A.H., 1838 

hypnorum (Fabricius, 1775) 

nitidulus (Fabricius, 1781) 189 

obtusus (Linnaeus, 1767) 

nitidicollis Stephens, 1832 

pallidus Sharp, 1871 

scutellaris Rye, 1871 non Lacordaire, 1835 

pusillus Gravenhorst, 1806 

quadriscopulatus Pandellé, 1869 

scitulus Erichson, 1839 

macropterus sensu auctt. non Stephens, 1832 

solutus Erichson, 1839 

tersus Erichson, 1839 

transversalis Gravenhorst, 1806 

Subfamily TRICHOPHYINAE Thomson, C.G., 

1858 

TRICHOPHYA Mannerheim, 1830 

pilicornis (Gyllenhal, 1810) 

Subfamily HABROCERINAE Mulsant & Rey, 1876 

HABROCERUS Erichson, 1839 

capillaricornis (Gravenhorst, 1806) 

Subfamily ALEOCHARINAE Fleming, 1821 190 

Tribe ACTOCHARINI Bernhauer & Schubert, 1911 191 

ACTOCHARIS Sharp, 1870 

readingii Sharp, 1870 

marina Fauvel, 1871 

Tribe ALEOCHARINI Fleming, 1821 

ALEOCHARA Gravenhorst, 1802 192 

Subgenus ALEOCHARA Gravenhorst, 1802 

brevipennis Gravenhorst, 1806 

curtula (Goeze, 1777) 

lata Gravenhorst, 1802 

38 

Subgenus BARYODMA Thomson, C.G., 1858 

intricata Mannerheim, 1830 

Subgenus CERANOTA Stephens, 1839 

ruficornis Gravenhorst, 1802 

Subgenus COPROCHARA Mulsant & Rey, 1874 

bilineata Gyllenhal, 1810 

binotata Kraatz, 1858 193 

verna sensu auctt. non Say, 1836 194 

pauxilla sensu auctt. non Mulsant & Rey, 1874 

bipustulata (Linnaeus, 1761) 

verna Say, 1836 195 

pauxilla Mulsant & Rey, 1874 196 

Subgenus EMPLENOTA Casey, 1884 

obscurella Gravenhorst, 1806 197 

algarum Fauvel, 1862 

phycophila Allen, 1937 198 

Subgenus POLYSTOMOTA Casey, 1906 199 

POLYSTOMA Stephens, 1835 non Zeder, 1800 

grisea Kraatz, 1856 

punctatella Motschulsky, 1858 

obscurella sensu auctt. non Gravenhorst, 1806 

Subgenus RHEOCHARA Mulsant & Rey, 1875 

spadicea (Erichson, 1837) 

Subgenus XENOCHARA Mulsant & Rey, 1874 

cuniculorum Kraatz, 1858 

discipennis Mulsant & Rey, 1853 

fumata Gravenhorst, 1802 

funebris Wollaston, 1864 

diversa sensu auctt. partim non (Sahlberg, J., 1876) 

albovillosa Bernhauer, 1901 200 

inconspicua Aubé, 1850 

kamila Likovský, 1984 201 

diversa (Sahlberg, J., 1876) non Mulsant & Rey, 

1853 

lanuginosa Gravenhorst, 1802 

lygaea Kraatz, 1862 

maculata Brisout de Barneville, 1863 

moerens Gyllenhal, 1827 

moesta Gravenhorst, 1802 

crassiuscula Sahlberg, C.R., 1831 

sanguinea (Linnaeus, 1758) 

sparsa Heer, 1839 

stichai Likovský, 1965 

tristis Gravenhorst, 1806 

villosa Mannerheim, 1830 

TINOTUS Sharp, 1883 202 

morion (Gravenhorst, 1802) 

Tribe ATHETINI Casey, 1910 

Subtribe ATHETINA Casey, 1910 203 

ACROTONA Thomson, C.G., 1859 204 

ATHETA sensu auctt. partim non Thomson, C.G., 

1858 

COLPODOTA Mulsant & Rey, 1873 

aterrima (Gravenhorst, 1802) 

benicki (Allen, 1940) 

pusilla (Brundin, 1952) 205 

convergens (Strand, 1958) 206

obfuscata sensu Brundin (1952) non (Gravenhorst, 

1802) 

exigua (Erichson, 1837) 

muscorum (Brisout de Barneville, 1860) 

obfuscata (Gravenhorst, 1802) 

parens (Mulsant & Rey, 1852) 

parvula (Mannerheim, 1830) 

parva sensu auctt. Brit. non (Sahlberg, C.R., 1831) 

pseudotenera (Cameron, 1933) 207 

rassii Muona, 1993 

pygmaea (Gravenhorst, 1802) 

sylvicola (Kraatz, 1856) 

silvicola auctt. (misspelling) 

planipennis (Thomson, C.G., 1855) 32 

troglodytes (Motschulsky, 1858) 

consanguinea (Eppelsheim, 1875) 208 

ADOTA Casey, 1909 


1858 

PHYCONOMA Easton, 1971 209 

maritima (Mannerheim, 1843) 

immigrans (Easton, 1971) 210 

ALAOBIA Thomson, C.G., 1858 211 

MYCOTA Mulsant & Rey, 1874 

ATHETA 'LOHSE GROUP II' sensu auctt. partim 

gagatina (Baudi, 1848) 

hybrida (Sharp, 1869) 

linderi (Brisout de Barneville, 1863) 212 

pallidicornis (Thomson, C.G., 1856) 

humeralis (Kraatz, 1856) 

scapularis (Sahlberg, C.R., 1831) 

sodalis (Erichson, 1837) 

subglabra (Sharp, 1869) 213 

sparreschneideri sensu auctt. Brit. non (Munster, 

1923) 

nondescripta (Ashe, 1957) 

taxiceroides Munster, 1932 214 

trinotata (Kraatz, 1856) 

ALEVONOTA Thomson, C.G., 1858 215 

ALEUONOTA Thomson, C.G., 1861 

egregia (Rye, 1876) 

gracilenta (Erichson, 1839) 

rufotestacea (Kraatz, 1856) 

atricapilla (Mulsant & Rey, 1852) 

aurantiaca Fauvel, 1895 216 

ALIANTA Thomson, C.G., 1858 

incana (Erichson, 1837) 

ALOCONOTA Thomson, C.G., 1858 


1858 

GLOSSOLA Fowler, 1887 

Subgenus ALOCONOTA Thomson, C.G., 1858 

cambrica (Wollaston, 1855) 

currax (Kraatz, 1856) 

eichhoffi (Scriba, 1867) 

aegyptiaca sensu (Ganglbauer, 1895) non 


gregaria (Erichson, 1839) 

insecta (Thomson, C.G., 1856) 

mihoki Bernhauer, 1913 217 

39 


planifrons (Waterhouse, G.R., 1864) 

debilicornis sensu auctt. Brit. non Erichson, 1839 218 

subgrandis Brundin, 1954 219 

sulcifrons (Stephens, 1832) 

Subgenus DISOPORA Thomson, C.G., 1859 220 

coulsoni (Last, 1952) 

longicollis sensu auctt. Brit. non (Mulsant & Rey, 

1852) 

languida (Erichson, 1837) 221 

walshi (Williams, 1930) 

longicollis (Mulsant & Rey, 1852) 222 

languida sensu auctt. Brit. non (Erichson, 1837) 

AMIDOBIA Thomson, C.G., 1858 


1858 

talpa (Heer, 1841) 

AMISCHA Thomson, C.G., 1858 

analis (Gravenhorst, 1802) 

bifoveolata (Mannerheim, 1830) 

cavifrons (Sharp, 1869) 223 

decipiens (Sharp, 1869) 

forcipata Mulsant & Rey, 1873 

nigrofusca (Stephens, 1829) 

soror (Kraatz, 1856) 223 

simillima (Sharp, 1869) 224 

ANOPLETA Mulsant & Rey, 1874 225 


1858 

corvina (Thomson, C.G., 1856) 

ellimani (Bernhauer, 1914) 226 

kochi (Roubal, 1937) 227 

verulamii Allen, 1994 228 

puberula (Sharp, 1869) 

luctuosa sensu auctt. non (Mulsant & Rey, 1853) 229 

picicornis (Mulsant & Rey, 1873) 

soedermani (Bernhauer, 1931) 230 

liliputana sensu auctt. Brit. non (Brisout de 

Barneville, 1860) 

ATHETA Thomson, C.G., 1858 

MEGISTA Mulsant & Rey, 1874 

?TETROPLA Mulsant & Rey, 1874 231 

aeneicollis (Sharp, 1869) 

pertyi sensu auctt. non (Heer, 1839) 

aquatica (Thomson, C.G., 1852) 

aquatilis (Thomson, C.G., 1867) 

brunneipennis (Thomson, C.G., 1852) 

valida (Kraatz, 1856) 

castanoptera (Mannerheim, 1830) 

ebenina (Mulsant & Rey, 1874) 

graminicola (Gravenhorst, 1806) 

heymesi Hubenthal, 1913 232 

hypnorum (Kiesenwetter, 1850) 

incognita (Sharp, 1869) 

laevicauda Sahlberg, J., 1876 233 

triangulum (Kraatz, 1856) 

xanthopus (Thomson, C.G., 1856) 

'species provisionally assigned to Atheta' 234 

autumnalis (Erichson, 1839) 235 

basicornis (Mulsant & Rey, 1852) 

boletophila (Thomson, C.G., 1856)


britanniae (Bernhauer & Scheerpeltz, 1926) 

repanda sensu auctt. non (Mulsant & Rey, 1874) 236 

reperta (Sharp, 1913) non Casey, 1910 

crassicornis (Fabricius, 1792) 

inoptata (Sharp, 1913) 

diversa (Sharp, 1869) 

divisa (Märkel, 1844) 

euryptera (Stephens, 1832) 

fungicola (Thomson, C.G., 1852) 

nitidicollis (Fairmaire & Laboulbène, 1856) 

harwoodi (Williams, 1930) 

intermedia (Thomson, C.G., 1852) 237 

liturata (Stephens, 1832) 

nidicola (Johansen, 1914) 

nigritula (Gravenhorst, 1802) 238 

oblita (Erichson, 1839) 

paracrassicornis Brundin, 1954 

pilicornis (Thomson, C.G., 1852) 

procera (Kraatz, 1856) 

ravilla (Erichson, 1839) 

angusticollis (Thomson, C.G., 1856) 

strandiella (Brundin, 1954) 

vaga (Heer, 1839) 

nigricornis (Thomson, C.G., 1852) non (Stephens, 

1832) 239 

BADURA Mulsant & Rey, 1873 240 


1858 

DIMETROTA sensu auctt. partim non Mulsant & 

Rey, 1873 

macrocera (Thomson, C.G., 1856) 

puncticollis (Benick, 1938) 

BESSOBIA Thomson, C.G., 1858 241 


1858 

excellens (Kraatz, 1856) 

fungivora (Thomson, C.G., 1867) 

monticola (Thomson, C.G., 1852) 

occulta (Erichson, 1837) 

BOREOPHILIA Benick, 1973 


1858 

eremita (Rye, 1866) 242 

islandica sensu auctt. Brit. non (Kraatz, 1857) 

hercynica sensu auctt. Brit. non (Renkonen, 1936) 

BRUNDINIA Tottenham, 1949 


1858 

METAXYA Mulsant & Rey, 1873 non Walker, 1856 

ACTOPHYLLA sensu auctt. partim non Bernhauer, 

1909 

marina (Mulsant & Rey, 1853) 243 

granosa (Hochhuth, 1849) 

meridionalis (Mulsant & Rey, 1853) 

CADAVEROTA Yosii & Sawada, 1976 244 

ATHETA 'LOHSE GROUP II' auctt. partim 

cadaverina (Brisout de Barneville, 1860) 

hansseni (Strand, 1943) 245 

40 

CALLICERUS Gravenhorst, 1802 

SEMIRIS Heer, 1839 246 

obscurus Gravenhorst, 1802 

rigidicornis (Erichson, 1839) 

CERITAXA Mulsant & Rey, 1873 247 


1858 

dilaticornis (Kraatz, 1856) 

spissata (Mulsant & Rey, 1873) 248 

pervagata (Benick, 1974) 

dilaticornis sensu auctt. non (Kraatz, 1856) 

testaceipes (Heer, 1839) 

brevicollis (Baudi, 1848) 

CHAETIDA Mulsant & Rey, 1874 249 


1858 

longicornis (Gravenhorst, 1802) 

COPROTHASSA Thomson, C.G., 1859 250 


1858 

melanaria (Mannerheim, 1830) 

testudinea (Erichson, 1839) 

DADOBIA Thomson, C.G., 1858 

immersa (Erichson, 1837) 

DALOTIA Casey, 1910 

ATHETA 'LOHSE GROUP I' auctt. partim 

coriaria (Kraatz, 1856) 251 

DATOMICRA Mulsant & Rey, 1874 252 


1858 

DATOSTIBA Sawada, 1976 

canescens (Sharp, 1869) 

celata (Erichson, 1837) 253 

arenicola (Thomson, C.G., 1868) 

germana (Sharp, 1869) 

dadopora (Thomson, C.G., 1867) 

celata sensu auctt. non (Erichson, 1837) 

nigra (Kraatz, 1856) 

zosterae sensu auctt. non (Thomson, C.G., 1856) 

sordidula (Erichson, 1837) 254 

zosterae (Thomson, C.G., 1856) 255 

hodierna (Sharp, 1869) 

olorophila Keys, 1933 

DILACRA Thomson, C.G., 1858 256 


1858 

DRALICA Mulsant & Rey, 1874 

luteipes (Erichson, 1837) 

vilis (Erichson, 1837) 257 

DIMETROTA Mulsant & Rey, 1873 258 


1858 

aeneipennis (Thomson, C.G., 1856) 

picipennis sensu auctt. non (Mannerheim, 1843) 259 

atramentaria (Gyllenhal, 1810)

cauta (Erichson, 1837) 

parvula sensu auctt. partim non (Mannerheim, 

1830) 

cinnamoptera (Thomson, C.G., 1856) 

ischnocera (Thomson, C.G., 1870) 

laevana (Mulsant & Rey, 1852) 

clintoni (Kevan, 1969) 260 

marcida (Erichson, 1837) 

nigripes (Thomson, C.G., 1856) 

setigera (Sharp, 1869) 

DINARAEA Thomson, C.G., 1858 


1858 

aequata (Erichson, 1837) 

angustula (Gyllenhal, 1810) 

linearis (Gravenhorst, 1802) 

DOCHMONOTA Thomson, C.G., 1859 


1858 

clancula (Erichson, 1837) 

ENALODROMA Thomson, C.G., 1859 


1858 

PTYCHANDRA Ganglbauer, 1895 non Felder, 1861 

hepatica (Erichson, 1839) 

GEOSTIBA Thomson, C.G., 1858 261 

SIPALIA sensu auctt. non Mulsant & Rey, 1853 

EVANYSTES Gistel, 1856 

circellaris (Gravenhorst, 1806) 

HALOBRECTA Thomson, C.G., 1858 

algae (Hardy, 1851) 262 

puncticeps (Thomson, C.G., 1852) 

algophila (Fenyes, 1909) 263 

flavipes Thomson, C.G., 1861 

princeps (Sharp, 1869) 264 

HYDROSMECTA Thomson, C.G., 1858 


1858 

HYDROSMECTINA Ganglbauer, 1895 265 

delicatissima (Bernhauer, 1908) 266 

delicatula (Sharp, 1869) 

eximia (Sharp, 1869) 

fragilis (Kraatz, 1854) 

longula (Heer, 1839) 

thinobioides (Kraatz, 1854) 

subtilissima (Kraatz, 1854) 

septentrionum (Benick, 1969) 267 

LIOGLUTA Thomson, C.G., 1858 

HYPNOTA Mulsant & Rey, 1874 

alpestris (Heer, 1839) 

nitidiuscula (Sharp, 1869) 268 

nitidula (Kraatz, 1856) 269 

granigera (Kiesenwetter, 1850) 270 

longiuscula (Gravenhorst, 1802) 

vicina (Stephens, 1832) 

microptera Thomson, C.G., 1867 

oblonga (Erichson, 1839) non (Lacordaire, 1835) 

41 

oblongiuscula Sharp, 1869 271 

pagana (Erichson, 1839) 


LYPROCORRHE Thomson, C.G., 1859 

NOTOTHECTA sensu auctt. partim non Thomson, 

C.G., 1858 

anceps (Erichson, 1837) 

MICRODOTA Mulsant & Rey, 1873 272 


1858 

aegra (Heer, 1841) 

amicula (Stephens, 1832) 

atomaria (Kraatz, 1856) 273 

atricolor (Sharp, 1869) 

mortuorum sensu auctt. Brit. non (Thomson, C.G., 

1867) 

benickiella (Brundin, 1948) 

validiuscula sensu auctt. non (Kraatz, 1856) 

boreella (Brundin, 1948) 

excelsa (Bernhauer, 1911) 

glabricula (Thomson, C.G., 1867) 

atomaria sensu (Sharp, 1869) non (Kraatz, 1856) 

minuscula sensu auctt. non (Brisout de Barneville, 

1860) 

indubia (Sharp, 1869) 

inquinula (Gravenhorst, 1802) 

liliputana (Brisout de Barneville, 1860) 274 

alpina (Benick, 1940) 

minuscula (Brisout de Barneville, 1860) 

atomaria sensu auctt. non (Kraatz, 1856) 

perexigua (Sharp, 1869) 

palleola (Erichson, 1837) 

spatuloides (Benick, 1939) 

spatula sensu auctt. Brit. non (Fauvel, 1876) 

subtilis (Scriba, 1866) 

MOCYTA Mulsant & Rey, 1874 275 


1858 

amplicollis (Mulsant & Rey, 1873) 

clientula (Erichson, 1839) 276 

fungi (Gravenhorst, 1806) 

fussi (Bernhauer, 1908) 

orbata (Erichson, 1837) 

orphana (Erichson, 1837) 

MYCETOTA Ádám, 1987 277 


fimorum (Brisout de Barneville, 1860) 278 

laticollis (Stephens, 1832) 

NEHEMITROPIA Lohse, 1971 

lividipennis (Mannerheim, 1830) 

sordida (Marsham, 1802) non (Gravenhorst, 

1802) 279 

NEOHILARA Lohse, 1971 


1858 

subterranea (Mulsant & Rey, 1853)


NOTOTHECTA Thomson, C.G., 1858 


1858 

confusa (Märkel, 1844) 

flavipes (Gravenhorst, 1806) 

OREOSTIBA Ganglbauer, 1895 280 


tibialis (Heer, 1839) 

OUSIPALIA des Gozis, 1886 


1858 

caesula (Erichson, 1839) 

PACHNIDA Mulsant & Rey, 1874 

ALIANTA sensu auctt. partim non Thomson, C.G., 

1858 

nigella (Erichson, 1837) 

PACHYATHETA Munster, 1925 281 


1858 

cribrata (Kraatz, 1856) 

mortuorum (Thomson, C.G., 1867) 282 

nesslingi sensu auctt. Brit. non (Bernhauer, 1928) 

PARAMEOTICA Ganglbauer, 1895 283 


1858 

DILACRA sensu auctt. partim non Thomson, C.G., 

1858 

difficilis (Brisout de Barneville, 1860) 

laticeps sensu auctt. Brit. non (Thomson, C.G., 

1856) 

rigua (Williams, 1929) 

PARANOPLETA Brundin in Hansen, 1954 

inhabilis (Kraatz, 1856) 

PHILHYGRA Mulsant & Rey, 1873 284 


1858 

HYGROECIA Mulsant & Rey, 1873 

PELURGA Mulsant & Rey, 1874 

arctica (Thomson, C.G., 1856) 

clavipes (Sharp, 1869) 

britteni Joy, 1913 

debilis (Erichson, 1837) 

deformis (Kraatz, 1856) 

complanata sensu auctt. non (Mannerheim, 1830) 

elongatula (Gravenhorst, 1802) 

balcanica (Brundin, 1944) non (Bernhauer, 1936) 

fallaciosa (Sharp, 1869) 

gyllenhalii (Thomson, C.G., 1856) 

hygrobia (Thomson, C.G., 1856) 285 

magniceps (Sahlberg, J., 1876) 

hygrotopora (Kraatz, 1856) 

luridipennis (Mannerheim, 1830) 

malleus (Joy, 1913) 

hygrobia sensu auctt. non (Thomson, C.G., 1856) 

melanocera (Thomson, C.G., 1856) 

?vaga (Heer, 1839) 

obtusangula (Joy, 1913) 

palustris (Kiesenwetter, 1844) 

42 

parca (Mulsant & Rey, 1873) 

nannion (Joy, 1931) 286 

scotica (Elliman, 1909) 

caucasica (Brundin, 1944) 

terminalis (Gravenhorst, 1806) 

volans (Scriba, 1859) 

halophila (Thomson, C.G., 1861) 

tomlini (Joy, 1913) 

PLATARAEA Thomson, C.G., 1858 


1858 

brunnea (Fabricius, 1798) 

nigriceps (Marsham, 1802) 

PYCNOTA Mulsant & Rey, 1874 

paradoxa (Mulsant & Rey, 1861) 

nidorum (Thomson, C.G., 1868) 

RHAGOCNEME Munster, 1923 287 


1858 

subsinuata (Erichson, 1839) 

SCHISTOGLOSSA Kraatz, 1856 

aubei (Brisout de Barneville, 1860) 

bergvalli Palm, 1968 288 

benicki Lohse, 1981 289 

curtipennis (Sharp, 1869) 

gemina (Erichson, 1837) 

viduata (Erichson, 1837) 

THINOBAENA Thomson, C.G., 1859 249 


1858 

vestita (Gravenhorst, 1806) 

TRAUMOECIA Mulsant & Rey, 1874 290 


picipes (Thomson, C.G., 1856) 

?complana (Mannerheim, 1830) 

TRICHIUSA Casey, 1893 

immigrata Lohse, 1984 291 

XENOTA Mulsant & Rey, 1874 292 


myrmecobia (Kraatz, 1856) 

Subtribe THAMIARAEINA Fenyes, 1921 

THAMIARAEA Thomson, C.G., 1858 

cinnamomea (Gravenhorst, 1802) 

hospita (Märkel, 1844) 

Tribe AUTALIINI Thomson, C.G., 1859 

AUTALIA Leach, 1819 

impressa (Olivier, 1795) 

brevicornis Blair, 1944 

longicornis Scheerpeltz, 1947 

impressa sensu auctt. partim non (Olivier, 1795) 

puncticollis Sharp, 1864

ivularis (Gravenhorst, 1802) 

Tribe DEINOPSINI Sharp, 1883 

DEINOPSIS Matthews, A.H., 1838 

DINOPSIS Agassiz, 1846 40 

erosa (Stephens, 1832) 

Tribe DIGLOTTINI Jakobson, 1909 

DIGLOTTA Champion, G.C., 1899 

DIGLOSSA Haliday, 1837 non Wagler, 1832 

mersa (Haliday, 1837) 

submarina (Fairmaire & Laboulbène, 1856) 293 

sinuaticollis (Mulsant & Rey, 1871) 294 

mersa sensu Lohse, 1985 non (Haliday, 1837) 

subsinuata Lohse, 1985 

Tribe FALAGRIINI Mulsant & Rey, 1873 

BOHEMIELLINA Machulka, 1941 

flavipennis (Cameron, 1920) 

paradoxa Machulka, 1941 

BORBOROPORA Kraatz in Kraatz & Fuss, 1862 

kraatzii Fuss in Kraatz & Fuss, 1862 

CORDALIA Jacobs, 1925 

FALAGRIA sensu auctt. partim non Leach, 1819 

obscura (Gravenhorst, 1802) 

FALAGRIA Leach, 1819 

caesa Erichson, 1837 

sulcata (Paykull, 1789) non (Müller, O.F., 1776) 

sulcatula (Gravenhorst, 1806) 

FALAGRIOMA Casey, 1906 


thoracica (Stephens, 1832) 

MYRMECOCEPHALUS MacLeay, 1873 


concinnus (Erichson, 1839) 

MYRMECOPORA Saulcy, 1865 

Subgenus XENUSA Mulsant & Rey, 1875 

brevipes Butler, 1909 295 

oweni Assing, 1997 296 

sulcata (Kiesenwetter, 1850) 

simillima (Wollaston, 1864) 297 

uvida (Erichson, 1840) 

Tribe GYMNUSINI Heer, 1839 

GYMNUSA Gravenhorst, 1806 

brevicollis (Paykull, 1800) 

variegata Kiesenwetter, 1845 

43 

Tribe HOMALOTINI Heer, 1839 


Subtribe BOLITOCHARINA Thomson, C.G., 1859 

BOLITOCHARA Mannerheim, 1830 

bella Märkel, 1844 

lucida (Gravenhorst, 1802) 

mulsanti Sharp, 1875 

obliqua Erichson, 1837 

pulchra (Gravenhorst, 1806) 

lunulata sensu (Paykull, 1789) non (Linnaeus, 1761) 

EURYUSA Erichson, 1837 

optabilis Heer, 1839 

sinuata Erichson, 1837 

HETEROTA Mulsant & Rey, 1874 

plumbea (Waterhouse, G.R., 1858) 

LEPTUSA Kraatz, 1856 298 

PACHYGLUTA Thomson, C.G., 1858 

fumida (Erichson, 1839) 

haemorrhoidalis sensu Ganglbauer, 1895 non 

(Heer, 1839) 

norvegica Strand, A., 1941 

pulchella (Mannerheim, 1830) 

angusta (Aubé, 1850) 

ruficollis (Erichson, 1839) 

PSEUDOMICRODOTA Machulka, 1935 

paganettii (Bernhauer, 1909) 299 

jelineki (Krása, 1914) 300 

PSEUDOPASILIA Ganglbauer, 1895 

testacea (Brisout de Barneville, 1863) 301 

RHOPALOCERINA Reitter, 1909 

clavigera (Scriba, 1859) 

TACHYUSIDA Mulsant & Rey, 1871 

gracilis (Erichson, 1837) 

THECTUROTA Casey, 1893 

CRYPTUSA sensu auctt. non Mulsant & Rey, 1873 

PRAGENSIELLA Machulka, 1941 

marchii (Dodero, 1922) 

capitalis sensu auctt. non (Mulsant & Rey, 1873) 

williamsi (Bernhauer, 1936) 302 

Subtribe GYROPHAENINA Kraatz, 1856 

AGARICOCHARA Kraatz, 1856 

GYROPHAENA sensu auctt. partim non 


latissima (Stephens, 1832) 

laevicollis (Kraatz, 1854) 

BRACHIDA Mulsant & Rey, 1871 

exigua (Heer, 1839) 

ENCEPHALUS Stephens, 1832 

complicans Stephens, 1832


GYROPHAENA Mannerheim, 1830 

affinis Mannerheim, 1830 

bihamata Thomson, C.G., 1867 

congrua Erichson, 1837 

fasciata sensu auctt. Brit. non (Marsham, 1802) 

fasciata (Marsham, 1802) 303 

laevipennis Kraatz, 1857 

gentilis Erichson, 1839 

hanseni Strand, A., 1946 

joyi Wendeler, 1924 

convexicollis Joy, 1912 non Sharp, 1883 

joyioides Wüsthoff, 1937 

lucidula Erichson, 1837 

manca Erichson, 1839 

angustata (Stephens, 1832) 304 

minima Erichson, 1837 

munsteri Strand, A., 1935 

nana (Paykull, 1800) 

poweri Crotch, 1867 

pseudonana Strand, A., 1939 

pulchella Heer, 1839 

rousi Dvorák, 1966 305 

strictula Erichson, 1839 

transversalis Strand, 1939 306 

williamsi Strand, A., 1935 

Subtribe HOMALOTINA Heer, 1839 

ANOMOGNATHUS Solier, 1849 

THECTURA Thomson, C.G., 1859 

cuspidatus (Erichson, 1839) 

CYPHEA Fauvel, 1863 

curtula (Erichson, 1837) 307 

HOMALOTA Mannerheim, 1830 

plana (Gyllenhal, 1810) 

Subtribe SILUSINA Fenyes, 1918 

SILUSA Erichson, 1837 

rubiginosa Erichson, 1837 

Tribe HYGRONOMINI Thomson, C.G., 1859 

HYGRONOMA Erichson, 1837 

dimidiata (Gravenhorst, 1806) 

Tribe HYPOCYPHTINI Laporte, 1835 

CYPHA Leach, 1819 308 

HYPOCYPHTUS Gyllenhal, 1827 

HYPOCYPTUS Erichson, 1839 (misspelling) 

apicalis (Brisout de Barneville, 1863) 309 

nitida (Palm, 1935) 310 

aprilis (Rey, 1882) 

imitator (Luze, 1902) 310 

discoidea (Erichson, 1839) 

laeviuscula (Mannerheim, 1830) 

longicornis (Paykull, 1800) 

pulicaria (Erichson, 1839) 

punctum (Motschulsky, 1857) 

seminulum (Erichson, 1839) 

seminula auctt. (misspelling) 

44 

hanseni (Palm, 1949) 310 

tarsalis (Luze, 1902) 311 

HOLOBUS Solier, 1849 

OLIGOTA sensu auctt. partim non Mannerheim, 

1830 

flavicornis (Lacordaire, 1835) 

OLIGOTA Mannerheim, 1830 

apicata (Erichson, 1837) 312 

granaria Erichson, 1837 

inflata (Mannerheim, 1830) 

parva Kraatz, 1862 

pygmaea Kraatz, 1858 non Solier, 1849 

picipes (Stephens, 1832) 

atomaria sensu auctt. Brit. non Erichson, 1837 

pumilio Kiesenwetter, 1858 

pusillima sensu auctt. Brit. partim non (Gravenhorst, 

1806) 

punctulata Heer, 1839 

ruficornis Sharp, 1870 

pusillima (Gravenhorst, 1806) 


Tribe LOMECHUSINI Fleming, 1821 

Subtribe LOMECHUSINA Fleming, 1821 

LOMECHUSA Gravenhorst, 1806 

ATEMELES Dillwyn, 1829 

emarginata (Paykull, 1789) 

paradoxa Gravenhorst, 1806 

LOMECHUSOIDES Tottenham, 1939 

LOMECHUSA sensu auctt. non Gravenhorst, 1806 

strumosus (Fabricius, 1775) 

Subtribe MYRMEDONIINA Thomson, C.G., 1867 

DRUSILLA Leach, 1819 

ASTILBUS Dillwyn, 1829 

canaliculata (Fabricius, 1787) 

MYRMOECIA Mulsant & Rey, 1874 

ZYRAS sensu auctt. partim non Stephens, 1835 

plicata (Erichson, 1837) 

PELLA Stephens, 1835 313 

ZYRAS sensu auctt. partim non Stephens, 1835 

MYRMEDONIA sensu auctt. non Erichson, 1837 

cognata (Märkel, 1842) 

funesta (Gravenhorst, 1806) 

humeralis (Gravenhorst, 1802) 

laticollis (Märkel, 1845) 

limbata (Paykull, 1789) 

lugens (Gravenhorst, 1802) 

ZYRAS Stephens, 1835 

collaris (Märkel, 1842) 

haworthi Stephens, 1835

Tribe MYLLAENINI Ganglbauer, 1895 

MYLLAENA Erichson, 1837 

brevicornis (Matthews, A.H., 1838) 

dubia (Gravenhorst, 1806) 

elongata (Matthews, A.H., 1838) 

fowleri Matthews, A., 1883 

gracilicornis Fairmaire & Brisout de Barneville, 1859 

gracilis (Matthews, A.H., 1838) 

infuscata Kraatz, 1853 

intermedia Erichson, 1837 

kraatzi Sharp, 1871 

masoni Matthews, A., 1883 

minuta (Gravenhorst, 1806) 

Tribe OXYPODINI Thomson, C.G., 1859 

Subtribe DINARDINA Mulsant & Rey, 1873 

DINARDA Leach, 1819 

dentata (Gravenhorst, 1806) 

hagensi Wasmann, 1889 

maerkeli Kiesenwetter, 1843 

pygmaea Wasmann, 1894 

HOMOEUSA Kraatz, 1856 

acuminata (Märkel, 1842) 

Subtribe MEOTICINA Seevers, 1978 

MEOTICA Mulsant & Rey, 1873 314 

anglica Benick in Muona, 1991 

indocilis sensu auctt. Brit. non (Heer, 1839) 

pallens sensu auctt. Brit. non (Redtenbacher, 1849) 

exilis (Knoch in Gravenhorst, 1806) 

exiliformis Joy, 1915 

exillima Sharp, 1915 

filiformis (Motschulsky, 1860) 315 

apicalis Benick, 1954 

capitalis sensu auctt. ?non Mulsant & Rey, 1873 316 

exilis sensu auctt. Brit. partim non (Knoch in 

Gravenhorst, 1806) 

pallens (Redtenbacher, 1849) 317 

lohsei Benick, 1953 

Subtribe OXYPODINA Thomson, C.G., 1859 

AMAROCHARA Thomson, C.G., 1858 

bonnairei (Fauvel, 1865) 

forticornis (Lacordaire, 1835) 

umbrosa (Erichson, 1837) 

CALODERA Mannerheim, 1830 318 

ITYOCARA Thomson, C.G., 1867 319 

aethiops (Gravenhorst, 1802) 

nigrita Mannerheim, 1830 

protensa Mannerheim, 1830 

riparia Erichson, 1837 

rubens (Erichson, 1837) 

rufescens Kraatz, 1856 

uliginosa Erichson, 1837 

45 


COUSYA Mulsant & Rey, 1875 320 

OCYUSA sensu auctt. Brit. partim non Kraatz, 1856 

CHILOMORPHA Krása, 1914 

defecta Mulsant & Rey, 1875 

longitarsis (Thomson, C.G., 1867) 

hibernica (Rye, 1876) 

laticollis sensu auctt. Brit. non Thomson, C.G., 1870 

nigrata (Fairmaire & Laboulbène, 1856) 

nitidiventris (Fagel, 1958) 321 

CRATARAEA Thomson, C.G., 1858 

suturalis (Mannerheim, 1830) 

DEXIOGYIA Thomson, C.G., 1858 

DEXIOGYA auctt. (misspelling) 

corticina (Erichson, 1837) 

HAPLOGLOSSA Kraatz, 1856 

MICROGLOTTA Kraatz, 1862 

MICROGLOSSA Stein, 1868 (misspelling) 

gentilis (Märkel, 1844) 

marginalis (Gravenhorst, 1806) 

obscura Joy, 1912 

nidicola (Fairmaire, 1852) 

picipennis (Gyllenhal, 1827) 

villosula (Stephens, 1832) 

pulla sensu auctt. non (Gravenhorst, 1802) 322 

puncticollis (Stephens, 1832) 

HYGROPORA Kraatz, 1856 

cunctans (Erichson, 1837) 

ILYOBATES Kraatz, 1856 

bennetti Donisthorpe, 1914 

nigricollis sensu auctt. Brit. partim non (Paykull, 

1800) 

subopacus Palm, 1935 323 

nigricollis (Paykull, 1800) 

propinquus (Aubé, 1850) 

ISCHNOGLOSSA Kraatz, 1856 

obscura Wunderle, 1990 324 

prolixa (Gravenhorst, 1802) 

turcica Wunderle, 1992 325 

MNIUSA Mulsant & Rey, 1875 326 

incrassata (Mulsant & Rey, 1852) 

OCALEA Erichson, 1837 

badia Erichson, 1837 

latipennis Sharp, 1870 

picata (Stephens, 1832) 

rivularis Miller, L., 1851 

OCYUSA Kraatz, 1856 

DEUBELIA Bernhauer, 1899 327 

maura (Erichson, 1837) 

picina (Aubé, 1850) 

OXYPODA Mannerheim, 1830 

acuminata (Stephens, 1832) 

lividipennis sensu auctt. non Mannerheim, 1830 328 

alternans (Gravenhorst, 1802)


annularis Mannerheim, 1830 

brachyptera (Stephens, 1832) 

brevicornis (Stephens, 1832) 

umbrata sensu auctt. non (Gravenhorst, 1802) 

carbonaria (Heer, 1841) 

sericea Heer, 1839 non Lacordaire, 1835 

elongatula Aubé, 1850 

exoleta Erichson, 1839 

ferruginea Erichson, 1839 

flavicornis Kraatz, 1856 

amoena Fairmaire & Laboulbène, 1856 329 

formiceticola Märkel, 1841 

haemorrhoa (Mannerheim, 1830) 

induta Mulsant & Rey, 1861 

islandica Kraatz, 1857 

lentula Erichson, 1837 

longipes Mulsant & Rey, 1861 

lurida Wollaston, 1857 

exoleta sensu auctt. non Erichson, 1839 

perplexa sensu auctt. partim non Mulsant & Rey, 

1860 

mutata Sharp, 1871 330 

riparia Fairmaire, 1859 non Thomson, C.G., 1855 

rugulosa sensu auctt. non Kraatz, 1856 

nigricornis Motschulsky, 1860 

opaca sensu auctt. Brit. partim non (Gravenhorst, 

1802) 

nigrocincta Mulsant & Rey, 1875 

opaca (Gravenhorst, 1802) 

praecox Erichson, 1839 331 

procerula Mannerheim, 1830 

elongatula sensu auctt. Brit. partim non Aubé, 1850 

recondita Kraatz, 1856 

soror Thomson, C.G., 1855 

spectabilis Märkel, 1844 

tarda Sharp, 1871 332 

brachyptera sensu auctt. partim ? non (Stephens, 

1832) 

tirolensis Gredler, 1863 

rupicola Rye, 1866 

vittata Märkel, 1842 

PHLOEOPORA Erichson, 1837 

PHLOEODROMA Kraatz, 1856 333 

concolor Kraatz, 1856 

corticalis (Gravenhorst, 1802) 

angustiformis Baudi, 1869 

nitidiventris Fauvel, 1904 334 

scribae Eppelsheim, 1884 

bernhaueri Lohse, 1984 335 

corticalis sensu auctt. Brit. partim non (Gravenhorst, 

1802) 

teres sensu auctt. Brit. ?partim non (Gravenhorst, 

1802) 336 

testacea (Mannerheim, 1830) 

STICHOGLOSSA Fairmaire & Laboulbène, 1856 

semirufa (Erichson, 1839) 

TETRALAUCOPORA Bernhauer, 1928 337 

CHILOPORA Kraatz, 1856 non Haime, 1854 

PAROCYUSA sensu auctt. non Bernhauer, 1902 

CHILOPORATA Strand, E., 1935 

longitarsis (Erichson, 1837) 

rubicunda (Erichson, 1837) 

46 

THIASOPHILA Kraatz, 1856 

THYASOPHILA Fairmaire & Laboulbène, 1856 

angulata (Erichson, 1837) 

inquilina (Märkel, 1842) 

Subtribe TACHYUSINA Thomson, C.G., 1859 

BRACHYUSA Mulsant & Rey, 1874 

concolor (Erichson, 1839) 

DACRILA Mulsant & Rey, 1874 338 

fallax (Kraatz, 1856) 

pruinosa (Kraatz, 1856) 

DASYGNYPETA Lohse, 1974 339 

GNYPETA sensu auctt. partim non Thomson, C.G., 

1858 

velata (Erichson, 1837) 

GNYPETA Thomson, C.G., 1858 

caerulea (Sahlberg, C.R., 1831) 

carbonaria (Mannerheim, 1830) 

labilis (Erichson, 1839) 

ripicola (Kiesenwetter, 1844) 

rubrior Tottenham, 1939 

ISCHNOPODA Stephens, 1835 340 

leucopus (Marsham, 1802) 

scitula (Erichson, 1837) 341 

umbratica (Erichson, 1837) 

TACHYUSA Erichson, 1837 342 

concinna Heer, 1839 343 

coarctata sensu auctt. Brit. non Erichson, 1837 

constricta Erichson, 1837 

objecta (Mulsant & Rey, 1870) 344 

THINONOMA Thomson, C.G., 1859 340 

atra (Gravenhorst, 1806) 

Tribe PHYTOSINI Thomson, C.G., 1867 

ARENA Fauvel, 1862 

tabida (Kiesenwetter, 1850) 

octavii Fauvel, 1862 

PHYTOSUS Curtis, 1838 

ACTOSUS Mulsant & Rey, 1871 

balticus Kraatz, 1859 

nigriventris (Chevrolat, 1843) 

spinifer Curtis, 1838 

Tribe PLACUSINI Mulsant & Rey, 1871 

PLACUSA Erichson, 1837 

complanata Erichson, 1839 345 

depressa Mäklin, 1845 

complanata sensu auctt. Brit. partim non Erichson, 

1839 

?humilis Erichson, 1839 

pumilio (Gravenhorst, 1802) 

tachyporoides (Waltl, 1838)

denticulata Sharp, 1870 

Subfamily SCAPHIDIINAE Latreille, 1806 346 

Tribe SCAPHIDIINI Latreille, 1806 

SCAPHIDIUM Olivier, 1790 

quadrimaculatum Olivier, 1790 

Tribe SCAPHIINI Achard, 1924 

SCAPHIUM Kirby, 1837 

immaculatum (Olivier, 1790) 

Tribe SCAPHISOMATINI Casey, 1893 

SCAPHISOMA Leach, 1815 

agaricinum (Linnaeus, 1758) 

assimile Erichson, 1845 

boleti (Panzer, 1793) 

Subfamily PIESTINAE Erichson, 1839 

SIAGONIUM Kirby, 1815 

quadricorne Kirby, 1815 

Subfamily OXYTELINAE Fleming, 1821 347 

Tribe COPROPHILINI Heer, 1839 

COPROPHILUS Latreille, 1829 348 

ELONIUM sensu auctt. non Leach, 1819 

striatulus (Fabricius, 1792) 

Tribe EUPHANIINI Reitter, 1909 

DELEASTER Erichson, 1839 

dichrous (Gravenhorst, 1802) 

SYNTOMIUM Curtis, 1828 

aeneum (Müller, P.W.J., 1821) 

Tribe OXYTELINI Fleming, 1821 

ANOTYLUS Thomson, C.G., 1859 

clypeonitens (Pandellé, 1867) 

speculifrons sensu auctt. Brit. non (Kraatz, 1857) 

complanatus (Erichson, 1839) 

fairmairei (Pandellé, 1867) 

hamatus (Fairmaire & Laboulbène, 1856) 

hammondi Schülke, 2009 349 

complanatus sensu auctt. partim non (Erichson, 

1839) 

insecatus (Gravenhorst, 1806) 

inustus (Gravenhorst, 1806) 

maritimus Thomson, C.G., 1861 

perrisi (Fauvel, 1862) 

mutator (Lohse, 1963) 

sculpturatus sensu auctt. partim non (Gravenhorst, 

1806) 

nitidulus (Gravenhorst, 1802) 

47 

rugosus (Fabricius, 1775) 137 

striatus (Strøm, 1768) 

saulcyi (Pandellé, 1867) 

sculpturatus (Gravenhorst, 1806) 

tetracarinatus (Block, 1799) 

OXYTELUS Gravenhorst, 1802 

fulvipes Erichson, 1839 

laqueatus (Marsham, 1802) 

migrator Fauvel, 1904 350 

piceus (Linnaeus, 1767) 

sculptus Gravenhorst, 1806 

PLATYSTETHUS Mannerheim, 1830 


Subgenus CRAETOPYCRUS Tottenham, 1939 

alutaceus Thomson, C.G., 1861 

capito Heer, 1839 

cornutus (Gravenhorst, 1802) 

degener Mulsant & Rey, 1878 

cornutus sensu auctt. Brit. partim non Gravenhorst, 

1802 

nitens (Sahlberg, C.R., 1832) 

nodifrons Mannerheim, 1830 

Subgenus PLATYSTETHUS Mannerheim, 1830 

arenarius (Fourcroy, 1785) 

Tribe THINOBIINI Sharp, 1887 

APLODERUS Stephens, 1833 

HAPLODERUS Agassiz, 1846 

caelatus (Gravenhorst, 1802) 

BLEDIUS Leach, 1819 351 

Subgenus ASTYCOPS Thomson, C.G., 1859 

subterraneus Erichson, 1839 

Subgenus BARGUS Schiødte, 1866 352 

annae Sharp, 1911 353 

arcticus Sahlberg, J., 1890 

denticollis sensu auctt. Brit. non Fauvel, 1872 

defensus Fauvel, 1872 

erraticus Erichson, 1839 

filipes Sharp, 1911 354 

fuscipes Rye, 1865 

longulus Erichson, 1839 

opacus (Block, 1799) 

pallipes (Gravenhorst, 1806) 355 

annae sensu Pope, 1977 partim ?non Sharp, 1911 

larseni Hansen, 1940 

terebrans (Schiødte, 1866) 

Subgenus BLEDIUS Leach, 1819 

frisius Lohse, 1978 356 

spectabilis sensu auctt. partim non Kraatz, 1857 

spectabilis Kraatz, 1857 

germanicus Wagner, 1935 non (Gravenhorst, 1806) 

356 357 

limicola Tottenham, 1940 

tricornis (Herbst, 1784) 

unicornis (Germar, 1825) 

Subgenus DICARENUS Gistel, 1834 

fergussoni Joy, 1912 

arenarius (Paykull, 1800) non (Fourcroy, 1785) 

arenoides Tottenham, 1939 

subniger Schneider, O., 1900


Subgenus ELBIDUS Mulsant & Rey, 1878 

bicornis (Germar, 1822) 358 

dama Motschulsky, 1857 

ssp. jutlandensis Herman, 1986 359 

atlanticus Lohse, 1978 non Koch, 1938 

diota Schiødte, 1866 

Subgenus EUCERATOBLEDIUS Znojko, 1929 

furcatus (Olivier, 1811) 

Subgenus HESPEROPHILUS Stephens, 1829 

atricapillus (Germar, 1825) 360 

praetermissus Williams, 1929 361 

crassicollis Lacordaire, 1835 

dissimilis Erichson, 1840 

femoralis (Gyllenhal, 1827) 

gallicus (Gravenhorst, 1806) 

fracticornis (Paykull, 1790) non (Müller, O.F., 1776) 

laetior Mulsant & Rey, 1878 

sharpi Fowler, 1913 

lohsei Schülke, 2011 362 

atricapillus sensu auctt. partim non (Germar, 1825) 

praetermissus sensu auctt. Brit. partim non 

Williams, 1929 

occidentalis Bondroit, 1907 

crassicollis sensu auctt. Brit. non Lacordaire, 1835 

CARPELIMUS Leach, 1819 

Subgenus CARPELIMUS Leach, 1819 

fuliginosus (Gravenhorst, 1802) 

lindrothi Palm, 1942 363 

obesus (Kiesenwetter, 1844) 

pusillus (Gravenhorst, 1802) 

lasti (Scheerpeltz, 1946) 

Subgenus MYOPINUS Scheerpeltz, 1937 

elongatulus (Erichson, 1839) 

Subgenus PARATROGOPHLOEUS Hatch, 1957 

bilineatus Stephens, 1834 

erichsoni (Sharp, 1871) 364 

bilineatus sensu auctt. partim non Stephens, 1834 

rivularis (Motschulsky, 1860) 137 

obscurus Stephens, 1834 

similis Smetana, 1967 365 

Subgenus TROGINUS Mulsant & Rey, 1878 

despectus (Baudi, 1869) 366 

incongruus Steel, 1969 367 

zealandicus sensu auctt. Brit. partim non (Sharp, 

1900) 

schneideri (Ganglbauer, 1895) 

hemerinus (Joy, 1913) 

zealandicus (Sharp, 1900) 364 

Subgenus TROGOPHLOEUS Mannerheim, 1830 

alutaceus (Fauvel, 1898) 368 

corticinus (Gravenhorst, 1806) 

dispersepunctatus (Scheerpeltz, 1947) 

foveolatus (Sahlberg, C.R., 1832) 

gracilis (Mannerheim, 1830) 

halophilus (Kiesenwetter, 1844) 

impressus (Lacordaire, 1835) 

manchuricus (Bernhauer, 1938) 

ssp. subtilicornis (Roubal, 1946) 369 

corticinus sensu auctt. Brit. partim non 

(Gravenhorst, 1806) 

strandi (Scheerpeltz, 1950) 

subtilis (Erichson, 1839) 

48 

MANDA Blackwelder, 1952 

ACROGNATHUS Erichson, 1839 non Agassiz, 

1836 

mandibularis (Gyllenhal, 1827) 

OCHTHEPHILUS Mulsant & Rey, 1856 370 

ANCYROPHORUS Kraatz, 1858 

andalusiacus (Fagel, 1957) 

omalinus sensu auctt. Brit. partim non (Erichson, 

1840) 

angustior (Bernhauer, 1943) 

omalinus sensu auctt. Brit. partim non (Erichson, 

1840) 

venustulus sensu auctt. Brit. non (Rosenhauer, 

1856) 

aureus (Fauvel, 1871) 

omalinus (Erichson, 1840) 

PLANEUSTOMUS Jacquelin du Val, 1857 

flavicollis Fauvel, 1871 

palpalis (Erichson, 1839) 

TEROPALPUS Solier, 1849 

CARPELIMUS sensu auctt. partim non Leach, 1819 

unicolor (Sharp, 1900) 

THINOBIUS Kiesenwetter, 1844 

bicolor Joy, 1911 371 

brunneipennis sensu auctt. Brit. non Kraatz, 1857 

linearis sensu auctt. Brit. non Kraatz, 1857 

brevipennis Kiesenwetter, 1850 

ciliatus Kiesenwetter, 1844 

longipennis sensu auctt. partim non (Heer, 1841) 

praetor Smetana, 1959 372 

crinifer Smetana, 1959 373 

longipennis sensu auctt. partim non (Heer, 1841) 

strandi Smetana, 1960 374 

longipennis (Heer, 1841) 375 

major Kraatz, 1857 

angusticeps sensu auctt. non Fauvel, 1889 

macroceros Joy, 1913 

macrocerus auctt. (misspelling) 

newberyi Scheerpeltz, 1925 

pallidus Newbery, 1909 non Casey, 1889 

THINODROMUS Kraatz, 1857 

AMISAMMUS des Gozis, 1886 

CARPELIMUS sensu auctt. partim non Leach, 1819 

arcuatus (Stephens, 1834) 

Subfamily OXYPORINAE Fleming, 1821 

OXYPORUS Fabricius, 1775 

rufus (Linnaeus, 1758) 

Subfamily SCYDMAENINAE Leach, 1815 376 

Tribe EUTHEIINI Casey, 1897 

EUTHEIA Stephens, 1830 

EUTHIA auctt. (misspelling) 

formicetorum Reitter, 1881 

linearis Mulsant, 1861

clavata Reitter, 1881 

plicata (Gyllenhal, 1813) 

schaumii Kiesenwetter, 1858 

schaumi auctt. (misspelling) 

scydmaenoides Stephens, 1830 

EUTHICONUS Reitter, 1881/2 

conicicollis (Fairmaire & Laboulbène, 1855) 377 

Tribe CEPHENNIINI Reitter, 1882 

CEPHENNIUM Müller, P.W.J. & Kunze, 1822 

gallicum Ganglbauer, 1899 

thoracicum sensu auctt. Brit. non Müller, P.W.J. & 

Kunze, 1822 

edmondsi Donisthorpe, 1931 

pallidum Edmonds, 1931 

Tribe CYRTOSCYDMINI Schaufuss, 1889 

EUCONNUS Thomson, C.G., 1859 

Subgenus CLADOCONNUS Reitter, 1909 

denticornis (Müller, P.W.J. & Kunze, 1822) 

Subgenus EUCONNUS Thomson, C.G., 1859 

fimetarius (Chaudoir, 1845) 

hirticollis (Illiger, 1798) 

rutilipennis (Müller, P.W.J. & Kunze, 1822) 

Subgenus NAPOCHUS Thomson, C.G., 1859 

duboisi Méquignon, 1929 

murielae Last, 1945 

pragensis (Machulka, 1923) 

claviger sensu auctt. Brit. non (Müller, P.W.J. & 

Kunze, 1822) 

Subgenus NEONAPOCHUS Machulka, 1929 

maeklinii (Mannerheim, 1844) 

maeklini auctt. (misspelling) 

MICROSCYDMUS Saulcy & Croissandeau, 1893 

EUCONNUS sensu auctt. partim non Thomson, 

C.G., 1859 

minimus (Chaudoir, 1845) 

nanus sensu auctt. Brit. partim non (Schaum, 1844) 

nanus (Schaum, 1844) 

NEURAPHES Thomson, C.G., 1859 378 

Subgenus NEURAPHES Thomson, C.G., 1859 

angulatus (Müller, P.W.J. & Kunze, 1822) 

carinatus (Mulsant & Rey, 1861) 

elongatulus (Müller, P.W.J. & Kunze, 1822) 

talparum Lokay, 1921 

rubicundus sensu auctt. Brit. non (Schaum, 1841) 

carinatus sensu auctt. Brit. partim non (Mulsant, 

1861) 

Subgenus PARARAPHES Reitter, 1891 

plicicollis Reitter, 1879 

planifrons Blatch, 1890 

praeteritus Rye, 1872 

longicollis sensu auctt. Brit. non (Motschulsky, 

1845) 

49 


SCYDMORAPHES Reitter, 1891 

NEURAPHES sensu auctt. partim non Thomson, 

C.G., 1859 

helvolus (Schaum, 1844) 

minutus sensu auctt. Brit. non (Chaudoir, 1845) 

nigrescens (Reitter, 1881) 

sparshalli (Denny, 1825) 

STENICHNUS Thomson, C.G., 1859 

CYRTOSCYDMUS Motschulsky, 1870 379 

SCYDMAENUS sensu Fowler, 1888 non Latreille, 

1802 

bicolor (Denny, 1825) 

exilis (Erichson, 1837) 

collaris (Müller, P.W.J. & Kunze, 1822) 

godarti (Latreille, 1806) 

poweri (Fowler, 1884) 

barnevillei (Reitter, 1884) 

harwoodianus Williams, 1927 

pusillus (Müller, P.W.J. & Kunze, 1822) 

scutellaris sensu auctt. Brit. partim non (Müller, 

P.W.J. & Kunze, 1822) 

stotti Donisthorpe, 1932 

scutellaris (Müller, P.W.J. & Kunze, 1822) 

pusillus sensu auctt. Brit. partim non (Müller, P.W.J. 

& Kunze, 1822) 

Tribe SCYDMAENINI Leach, 1815 

SCYDMAENUS Latreille, 1802 

EUMICRUS Laporte, 1840 

Subgenus CHOLERUS Thomson, C.G., 1859 

rufus Müller, P.W.J. & Kunze, 1822 

Subgenus SCYDMAENUS Latreille, 1802 

tarsatus Müller, P.W.J. & Kunze, 1822 

Subfamily STENINAE MacLeay, 1825 

DIANOUS Leach, 1819 

coerulescens (Gyllenhal, 1810) 

STENUS Latreille, 1796 380 

Subgenus HEMISTENUS Motschulsky, 1860 381 

PARASTENUS Heyden, 1905 382 

aceris Stephens, 1833 

fuscicornis Erichson, 1840 

geniculatus Gravenhorst, 1806 

glacialis Heer, 1839 

impressus Germar, 1824 

ludyi Fauvel, 1855 

coarcticollis sensu auctt. non Eppelsheim, 1890 

ochropus Kiesenwetter, 1858 

erichsoni Rye, 1864 

ossium Stephens, 1833 

pallipes Gravenhorst, 1802 

palustris Erichson, 1839 

subaeneus Erichson, 1840 

Subgenus HYPOSTENUS Rey, 1884 

cicindeloides (Schaller, 1783) 

fornicatus Stephens, 1833 

fulvicornis Stephens, 1833 

kiesenwetteri Rosenhauer, 1856 

latifrons Erichson, 1839


oscillator Rye, 1870 

similis (Herbst, 1784) 

solutus Erichson, 1840 

tarsalis Ljungh, 1810 

Subgenus METATESNUS Ádám, 2001 

METASTENUS Ádám, 1987 non Walker, 1834 383 

HEMISTENUS sensu auctt. non Motschulsky, 1860 

bifoveolatus Gyllenhal, 1827 

binotatus Ljungh, 1804 

brevipennis Thomson, C.G., 1851 384 

foveicollis Kraatz, 1857 

picipes sensu auctt. partim ?non Stephens, 1833 

butrintensis Smetana, 1959 

pallitarsis sensu auctt. partim non Stephens, 1833 

canescens Rosenhauer, 1856 

flavipes Stephens, 1833 

nitidiusculus Stephens, 1833 

niveus Fauvel, 1865 

pallitarsis Stephens, 1833 

picipennis Erichson, 1840 

picipes Stephens, 1833 

pubescens Stephens, 1833 

umbratilis Casey, 1884 

pseudopubescens Strand, A., 1940 

pubescens sensu auctt. Brit. partim non Stephens, 

1833 

Subgenus STENUS Latreille, 1796 

NESTUS Rey, 1884 

argus Gravenhorst, 1806 

asphaltinus Erichson, 1840 

assequens Rey, 1844 

simillimus Benick, 1949 

ater Mannerheim, 1830 

atratulus Erichson, 1839 

biguttatus (Linnaeus, 1758) 

bimaculatus Gyllenhal, 1810 

boops Ljungh, 1810 385 

buphthalmus sensu Gravenhorst, 1802 non 

(Schrank, 1776) 

calcaratus Scriba, 1864 386 

canaliculatus Gyllenhal, 1827 

carbonarius Gyllenhal, 1827 

circularis Gravenhorst, 1802 

clavicornis (Scopoli, 1763) 

comma LeConte, 1863 

bipunctatus Erichson, 1839 non Ljungh, 1804 

contumax Assing, 1994 387 

europaeus Puthz, 1966 

cautus sensu auctt. non Erichson, 1839 

vafellus sensu auctt. Brit. non Erichson, 1839 

fossulatus Erichson, 1840 

fuscipes Gravenhorst, 1802 

glabellus Thomson, C.G., 1870 388 

guttula Müller, P.W.J., 1821 

guynemeri Jacquelin du Val, 1850 

incanus Erichson, 1839 

incrassatus Erichson, 1839 

juno (Paykull, 1789) 

longitarsis Thomson, C.G., 1851 

lustrator Erichson, 1839 

melanarius Stephens, 1833 

melanopus (Marsham, 1802) 

morio Gravenhorst, 1806 

nanus Stephens, 1833 

declaratus Erichson, 1839 

nitens Stephens, 1833 

palposus Zetterstedt, 1838 

50 

proditor Erichson, 1839 

providus Erichson, 1839 

rogeri Kraatz, 1857 389 

pusillus Stephens, 1833 

exiguus Erichson, 1840 390 

subdepressus Mulsant & Rey, 1861 

Subgenus TESNUS Rey, 1884 391 

brunnipes Stephens, 1833 

crassus Stephens, 1833 

formicetorum Mannerheim, 1843 

nigritulus Gyllenhal, 1827 

opticus Gravenhorst, 1806 

Subfamily EUAESTHETINAE Thomson, C.G., 

1859 

EDAPHUS Motschulsky, 1857 

beszedesi Reitter, 1914 392 

EUAESTHETUS Gravenhorst, 1806 

bipunctatus (Ljungh, 1804) 

laeviusculus Mannerheim, 1844 

ruficapillus Lacordaire, 1835 

Subfamily PSEUDOPSINAE Ganglbauer, 1895 

PSEUDOPSIS Newman, 1834 

sulcata Newman, 1834 

Subfamily PAEDERINAE Fleming, 1821 393 

Tribe PAEDERINI Fleming, 1821 

Subtribe ASTENINA Hatch, 1957 

ASTENUS Dejean, 1833 

ASTENOGNATHUS Reitter, 1909 394 

immaculatus Stephens, 1833 

lyonessius (Joy, 1908) 

angustatus sensu auctt. non (Paykull, 1789) 

longelytratus Palm, 1936 

brevelytratus Coiffait, 1960 

procerus (Gravenhorst, 1806) 

filiformis (Latreille, 1806) non (Fabricius, 1792) 

pulchellus (Heer, 1839) 

melanurus sensu Townsend, 1948 non Küster, 1853 

serpentinus (Motschulsky, 1858) 

procerus sensu auctt. Brit. partim non (Gravenhorst, 

1806) 

subditus (Mulsant & Rey, 1878) 395 

Subtribe CRYPTOBIINA Casey, 1905 

OCHTHEPHILUM Stephens, 1829 

CRYPTOBIUM Mannerheim, 1830 

collare (Reitter, 1884) 396 

fracticorne (Paykull, 1800) 397 

brevipenne sensu auctt. non Mulsant & Rey, 1861 

jacquelini (Boieldieu, 1859)

Subtribe LATHROBIINA Laporte, 1835 

ACHENIUM Leach, 1819 

depressum (Gravenhorst, 1802) 

humile (Nicolai, 1822) 

LATHROBIUM Gravenhorst, 1802 398 

Subgenus LATHROBIUM Gravenhorst, 1802 

brunnipes (Fabricius, 1792) 

dilutum Erichson, 1839 

elongatum (Linnaeus, 1767) 

fovulum Stephens, 1833 

britannicum (Coiffait, 1972) 

fulvipenne (Gravenhorst, 1806) 399 

geminum Kraatz, 1857 400 

boreale Hochhuth, 1851 

volgense Hochhuth, 1851 

impressum Heer, 1841 

filiforme Gravenhorst, 1806 non (Fabricius, 1792) 

longulum Gravenhorst, 1802 

longipenne Fairmaire & Laboulbène, 1856 

pallidipenne Hochhuth, 1851 

ripicola Czwalina, 1888 401 

pallidum Nordmann, 1837 

rufipenne Gyllenhal, 1813 

LOBRATHIUM Mulsant & Rey, 1878 402 

LATHROBIUM sensu auctt. partim non 

Gravenhorst, 1802 

multipunctum (Gravenhorst, 1802) 

PLATYDOMENE Ganglbauer, 1895 403 

angusticollis (Lacordaire, 1835) 

TETARTOPEUS Czwalina, 1888 403 

angustatus (Lacordaire, 1835) 

quadratus (Paykull, 1789) 

rufonitidus (Reitter, 1909) 

fennicus (Renkonen, 1938) 404 

terminatus (Gravenhorst, 1802) 

zetterstedti (Rye, 1872) 

punctatus (Zetterstedt, 1828) non (Fourcroy, 1785) 

foveatus sensu auctt. non (Stephens, 1839) 

Subtribe MEDONINA Casey, 1905 

HYPOMEDON Mulsant & Rey, 1878 405 

CHLOECHARIS Lynch Arribálzaga, 1884 

debilicornis (Wollaston, 1857) 406 

LITHOCHARIS Dejean, 1833 

nigriceps Kraatz, 1859 

ochracea (Gravenhorst, 1802) 407 

MEDON Stephens, 1833 408 

apicalis (Kraatz, 1857) 

brunneus (Erichson, 1839) 

castaneus (Gravenhorst, 1802) 

dilutus (Erichson, 1839) 

fusculus (Mannerheim, 1830) 

piceus (Kraatz, 1858) 

pocofer (Peyron, 1857) 

ripicola (Kraatz, 1854) 

51 


PSEUDOMEDON Mulsant & Rey, 1877 409 

LITHOCHARIS sensu auctt. partim non Dejean, 

1833 

obscurellus (Erichson, 1840) 

obsoletus (Nordmann, 1837) 

SUNIUS Stephens, 1829 

HYPOMEDON sensu auctt. non Mulsant & Rey, 

1878 

bicolor (Olivier, 1795) 

melanocephalus (Fabricius, 1792) 

propinquus (Brisout de Barneville, 1867) 

Subtribe PAEDERINA Fleming, 1821 

PAEDERIDUS Mulsant & Rey, 1878 410 

PAEDERUS sensu auctt. partim non Fabricius, 

1775 

rubrothoracicus (Goeze, 1777) 

ruficollis (Fabricius, 1781) 392 

PAEDERUS Fabricius, 1775 

caligatus Erichson, 1840 

fuscipes Curtis, 1826 

littoralis Gravenhorst, 1802 

riparius (Linnaeus, 1758) 

Subtribe SCOPAEINA Mulsant & Rey, 1878 

SCOPAEUS Erichson, 1840 411 

gracilis (Sperk, 1835) 

laevigatus (Gyllenhal, 1827) 

gracilis sensu auctt. Brit. partim non (Sperk, 1835) 

minutus Erichson, 1840 

sulcicollis sensu Fowler, 1888 non (Stephens, 1833) 

gracilipes Edmonds, 1933 

ryei Wollaston, 1872 

minimus sensu auctt. Brit. non Erichson, 1839 412 

rubidus sensu auctt. Brit. non Mulsant & Rey, 1855 

sulcicollis (Stephens, 1833) 

minutus sensu auctt. partim non Erichson, 1840 

cognatus Mulsant & Rey, 1855 

abbreviatus sensu Edmonds, 1931 non Mulsant & 

Rey, 1855 

pusillus sensu Tottenham, 1949 non Kiesenwetter, 

1843 

Subtribe STILICINA Casey, 1905 

RUGILUS Leach, 1819 

STILICUS Berthold, 1827 

angustatus (Geoffroy, 1785) 

fragilis (Gravenhorst, 1806) 413 

scutellatus (Motschulsky, 1858) 

erichsonii (Fauvel, 1867) 

geniculatus (Erichson, 1839) 

orbiculatus (Paykull, 1789) 

rufipes Germar, 1836 

similis (Erichson, 1839) 

subtilis (Erichson, 1840)


Subfamily STAPHYLININAE Latreille, 1802 

Tribe OTHIINI Thomson, C.G., 1859 

ATRECUS Jacquelin du Val, 1856 

BAPTOLINUS Kraatz, 1857 


OTHIUS Stephens, 1829 

angustus Stephens, 1833 

melanocephalus (Gravenhorst, 1806) non (Geoffroy, 

1785) 

laeviusculus Stephens, 1833 

lapidicola Märkel & Kiesenwetter, 1848 414 

punctulatus (Goeze, 1777) 

subuliformis Stephens, 1833 

myrmecophilus Kiesenwetter, 1843 415 

Tribe STAPHYLININI Latreille, 1802 

Subtribe PHILONTHINA Kirby, 1837 

BISNIUS Stephens, 1829 416 

PHILONTHUS sensu auctt. partim non Stephens, 

1829 

cephalotes (Gravenhorst, 1802) 417 

fimetarius (Gravenhorst, 1802) 

rigidicornis (Gravenhorst, 1802) 

nigriventris (Thomson, C.G., 1867) 

parcus (Sharp, 1874) 

felix (Last, 1961) 

pseudoparcus (Brunne, 1976) 418 

puella (Nordmann, 1837) 

scoticus (Joy & Tomlin, 1913) 419 

sordidus (Gravenhorst, 1802) 

pachycephalus (Nordmann, 1837) 

subuliformis (Gravenhorst, 1802) 

fuscus (Gravenhorst, 1802) non (Gmelin in 


CAFIUS Stephens, 1829 

cicatricosus (Erichson, 1840) 

fucicola Curtis, 1830 

xantholoma (Gravenhorst, 1806) 

ERICHSONIUS Fauvel, 1874 

ACTOBIUS Fauvel, 1875 

cinerascens (Gravenhorst, 1802) 

signaticornis (Mulsant & Rey, 1853) 

ytenensis (Sharp, 1913) 

GABRIUS Stephens, 1829 420 

appendiculatus Sharp, 1910 

subnigritulus sensu auctt. non Joy, 1913 421 

astutoides (Strand, A., 1946) 

astutus sensu auctt. partim non Erichson, 1840 

bishopi Sharp, 1910 

breviventer (Sperk, 1835) 

coxalus (Hochhuth, 1871) 

pennatus Sharp, 1910 422 

exiguus (Nordmann, 1837) 

keysianus Sharp, 1910 

nigritulus (Gravenhorst, 1802) 

osseticus (Kolenati, 1846) 

52 

vernalis (Gravenhorst, 1806) non (Müller, O.F., 

1776) 

piliger Mulsant & Rey, 1876 

stipes Sharp, 1910 

splendidulus (Gravenhorst, 1802) 

trossulus (Nordmann, 1837) 

velox Sharp, 1910 

primigenius Joy, 1913 

GABRONTHUS Tottenham, 1955 

thermarum (Aubé, 1850) 

NEOBISNIUS Ganglbauer, 1895 

lathrobioides (Baudi, 1848) 

cerrutii Gridelli, 1943 423 

rubripennis Gridelli, 1943 

procerulus (Gravenhorst, 1806) 

prolixus (Erichson, 1840) 

villosulus (Stephens, 1833) 

PHILONTHUS Stephens, 1829 424 

SPATULONTHUS Tottenham, 1955 425 

PARAGABRIUS Coiffait, 1963 425 

addendus Sharp, 1867 

albipes (Gravenhorst, 1802) 

alpinus sensu auctt. Brit. partim non Eppelsheim, 

1875 426 

alpinus Eppelsheim, 1875 392 

atratus (Gravenhorst, 1802) 

carbonarius (Gravenhorst, 1802) 427 

varius (Gyllenhal, 1810) 

cognatus Stephens, 1832 

fuscipennis (Mannerheim, 1830) non (Block, 1799) 

concinnus (Gravenhorst, 1802) 

ochropus (Gravenhorst, 1802) 

confinis Strand, A., 1941 

coprophilus Jarrige, 1949 

corruscus (Gravenhorst, 1802) 

corvinus Erichson, 1839 

cruentatus (Gmelin in Linnaeus, 1790) 

debilis (Gravenhorst, 1802) 

decorus (Gravenhorst, 1802) 

dimidiatipennis Erichson, 1840 

discoideus (Gravenhorst, 1802) 

ebeninus (Gravenhorst, 1802) 

fumarius (Gravenhorst, 1806) 

furcifer Renkonen, 1937 

intermedius (Lacordaire, 1835) 

jurgans Tottenham, 1937 

varians sensu auctt. partim non (Paykull, 1789) 

laminatus (Creutzer, 1799) 

lepidus (Gravenhorst, 1802) 

longicornis Stephens, 1832 

mannerheimi Fauvel, 1869 

marginatus (Müller, O.F., 1764) 

micans (Gravenhorst, 1802) 

micantoides Benick & Lohse, 1956 

micans sensu auctt. partim non (Gravenhorst, 1802) 

nigrita (Gravenhorst, 1806) 

nitidicollis (Lacordaire, 1835) 

bimaculatus (Gravenhorst, 1802) non (Schrank, 

1798) 

parvicornis (Gravenhorst, 1802) 

agilis (Gravenhorst, 1806) 428 

politus (Linnaeus, 1758) 

aeneus (Rossi, 1790) non (De Geer, 1774)

temporalis sensu Crowson, 1970 non Mulsant & 

Rey, 1853 

punctus (Gravenhorst, 1802) 

quisquiliarius (Gyllenhal, 1810) 

rectangulus Sharp, 1874 

rotundicollis (Ménétries, 1832) 

rubripennis Stephens, 1832 

fulvipes sensu auctt. Brit. non (Fabricius, 1792) 

varipes Mulsant & Rey, 1861 429 

rufipes (Stephens, 1832) 430 

immundus sensu auctt. non (Gravenhorst, 1806) 

sanguinolentus (Gravenhorst, 1802) 

spinipes Sharp, 1874 431 

splendens (Fabricius, 1792) 137 

niger (Müller, O.F., 1764) 

succicola Thomson, C.G., 1860 

chalceus sensu Ganglbauer, 1895 non Stephens, 

1832 

tenuicornis Mulsant & Rey, 1853 

carbonarius sensu (Gyllenhal, 1810) non 


umbratilis (Gravenhorst, 1802) 

varians (Paykull, 1789) 432 

ventralis (Gravenhorst, 1802) 

RABIGUS Mulsant & Rey, 1876 433 

PHILONTHUS sensu auctt. partim non Stephens, 

1829 

pullus (Nordmann, 1837) 

REMUS Holme, 1837 434 

CAFIUS sensu auctt. partim non Stephens, 1829 

sericeus Holme, 1837 

Subtribe QUEDIINA Kraatz, 1857 

ACYLOPHORUS Nordmann, 1837 

glaberrimus (Herbst, 1784) 

ASTRAPAEUS Gravenhorst, 1802 

ulmi (Rossi, 1790) 435 

EURYPORUS Erichson, 1839 

picipes (Paykull, 1800) 

HETEROTHOPS Stephens, 1829 

binotatus (Gravenhorst, 1802) 436 

dissimilis (Gravenhorst, 1802) 

minutus Wollaston, 1860 437 

dissimilis sensu auctt. partim non (Gravenhorst, 

1802) 

niger Kraatz, 1868 438 

praevius Erichson, 1839 

QUEDIUS Stephens, 1829 

Subgenus DISTICHALIUS Casey, 1915 

cinctus (Paykull, 1790) 

Subgenus MICROSAURUS Dejean, 1833 439 

aetolicus Kraatz, 1858 

brevicornis (Thomson, C.G., 1860) 

brevis Erichson, 1840 

cruentus (Olivier, 1795) 

fulgidus (Fabricius, 1792) 137 

53 


?rufitarsis (Marsham, 1802) 

assimilis (Nordmann, 1837) 

invreae Gridelli, 1924 

ochripennis sensu auctt. Brit. non Ménétries, 1832 

assecla sensu auctt. non Mulsant & Rey, 1876 

lateralis (Gravenhorst, 1802) 

longicornis Kraatz, 1857 

lyszkowskii Lott, 2010 440 

maurus (Sahlberg, C.R., 1830) 

mesomelinus (Marsham, 1802) 

microps Gravenhorst, J.L.C., 1847 

infuscatus sensu Donisthorpe, 1928 non Erichson, 

1840 

nigrocaeruleus Fauvel, 1874 

puncticollis (Thomson, C.G., 1867) 

othiniensis Johansen, 1907 

scitus (Gravenhorst, 1806) 137 

analis (Fabricius, 1787) 

truncicola Fairmaire & Laboulbène, 1856 

ventralis (Aragona, 1830) non (Gravenhorst, 1802) 

xanthopus Erichson, 1839 

Subgenus QUEDIONUCHUS Sharp, 1884 

plagiatus Mannerheim, 1843 137 

glaber (Müller, O.F., 1764) 

flavopterus (Geoffroy, 1785) 

laevigatus (Gyllenhal, 1810) non (Marsham, 1802) 

Subgenus QUEDIUS Stephens, 1829 

balticus Korge, 1960 

molochinus sensu Last, 1952 non (Gravenhorst, 

1806) 

curtipennis Bernhauer, 1908 

fuliginosus sensu Britten, 1944 non (Gravenhorst, 

1802) 

fuliginosus (Gravenhorst, 1802) 

subfuliginosus Britten, 1944 

levicollis (Brullé, 1832) 

tristis sensu auctt. 441 

molochinus (Gravenhorst, 1806) 

simplicifrons Fairmaire, 1861 442 

hispanicus Bernhauer, 1898 

pallipes sensu Gridelli, 1924 partim non Lucas, 

1849 

secundus Last, 1952 

Subgenus RAPHIRUS Stephens, 1829 

auricomus Kiesenwetter, 1850 

boopoides Munster, 1923 

picipennis sensu auctt. Brit. non Heer, 1839 

scribae sensu auctt. Brit. non Ganglbauer, 1895 

boops (Gravenhorst, 1802) 

asturicus sensu auctt. non Bernhauer, 1918 

arestor Tottenham, 1948 

crius Tottenham, 1948 

fulvicollis (Stephens, 1833) 

fumatus (Stephens, 1833) 

humeralis Stephens, 1832 

obliteratus Erichson, 1840 

suturalis sensu auctt. Brit. partim non Kiesenwetter, 

1845 

lucidulus Erichson, 1839 392 

maurorufus (Gravenhorst, 1806) 

nemoralis Baudi, 1848 

humeralis sensu auctt. Brit. partim non Stephens, 

1832 

suturalis sensu auctt. Brit. partim non Kiesenwetter, 

1845 

nigriceps Kraatz, 1857


nitipennis (Stephens, 1833) 

attenuatus sensu (Gyllenhal, 1810) non 


hyperboreus sensu auctt. Brit. non Erichson, 1840 

persimilis Mulsant & Rey, 1876 

boops sensu auctt. Brit. partim non (Gravenhorst, 

1802) 

aridulus Jansson, 1939 443 

corion Tottenham, 1948 444 

mallius Tottenham, 1948 

picipes (Mannerheim, 1830) 

plancus Erichson, 1840 

riparius Kellner, 1843 

schatzmayri Gridelli, 1922 

scintillans (Gravenhorst, 1806) 

semiaeneus (Stephens, 1833) 

semiobscurus (Marsham, 1802) 

rufipes (Gravenhorst, 1802) non (Linnaeus, 1758) 

umbrinus Erichson, 1839 

pseudoumbrinus Lohse, 1961 445 

VELLEIUS Leach, 1819 

dilatatus (Fabricius, 1787) 

Subtribe STAPHYLININA Latreille, 1802 

CREOPHILUS Leach, 1819 

maxillosus (Linnaeus, 1758) 

DINOTHENARUS Thomson, C.G., 1858 

STAPHYLINUS sensu auctt. partim non Linnaeus, 

1758 

TRICHODERMA sensu auctt. non Stephens, 1835 

PLATYDRACUS sensu auctt. partim non Thomson, 

C.G., 1858 

PARABEMUS Reitter, 1909 

pubescens (De Geer, 1774) 446 

EMUS Leach, 1819 

hirtus (Linnaeus, 1758) 

OCYPUS Leach, 1819 

GOERIUS Westwood, 1827 


1758 447 

Subgenus MATIDUS Motschulsky, 1860 448 

brunnipes (Fabricius, 1781) 

nitens (Schrank, 1781) 

nero (Faldermann, 1835) 449 

similis sensu (Fabricius, 1792) non (Herbst, 1784) 

Subgenus OCYPUS Leach, 1819 

olens (Müller, O.F., 1764) 

ophthalmicus (Scopoli, 1763) 

Subgenus PSEUDOCYPUS Mulsant & Rey, 1876 450 

aeneocephalus (De Geer, 1774) 

cupreus sensu auctt. Brit. partim non (Rossi, 1790) 

fortunatarum (Wollaston, 1871) 

cupreus sensu auctt. Brit. partim non (Rossi, 1790) 

fuscatus (Gravenhorst, 1802) 

ONTHOLESTES Ganglbauer, 1895 

murinus (Linnaeus, 1758) 

tessellatus (Geoffroy, 1785) 

54 

PLATYDRACUS Thomson, C.G., 1858 


1758 

fulvipes (Scopoli, 1763) 

latebricola (Gravenhorst, 1806) 

stercorarius (Olivier, 1795) 

STAPHYLINUS Linnaeus, 1758 

caesareus Cederhjelm, 1798 

dimidiaticornis Gemminger, 1851 

caesareus sensu auctt. Brit. partim non 

(Cederhjelm, 1798) 

parumtomentosus Stein, 1903 

erythropterus Linnaeus, 1758 

TASGIUS Stephens, 1829 451 


1758 452 

OCYPUS sensu auctt. partim non Leach, 1819 

ALAPSODUS Tottenham, 1939 

Subgenus RAYACHEILA Motschulsky, 1845 453 

globulifer (Geoffroy, 1785) 

siculus Stierlin, 1864 non (Aubé, 1842) 

evitendus Tottenham, 1945 

melanarius (Heer, 1839) 

globulifer sensu auctt. non (Geoffroy, 1785) 

morsitans (Rossi, 1790) 

compressus (Marsham, 1802) 454 

winkleri Bernhauer, 1906 

Subgenus TASGIUS Stephens, 1829 

ater (Gravenhorst, 1802) 

pedator (Gravenhorst, 1802) 

Tribe XANTHOLININI Erichson, 1839 

GAUROPTERUS Thomson, C.G., 1860 

fulgidus (Fabricius, 1787) 

GYROHYPNUS Leach, 1819 

angustatus Stephens, 1833 

scoticus (Joy, 1913) 455 

atratus (Heer, 1839) 

fracticornis (Müller, O.F., 1776) 

punctulatus sensu auctt. partim non (Paykull, 1789) 

punctulatus (Paykull, 1789) 456 

liebei sensu auctt. non (Scheerpeltz, 1926) 

wagneri (Scheerpeltz, 1926) 392 

HYPNOGYRA Casey, 1906 457 

XANTHOLINUS sensu auctt. partim non Dejean, 

1821 

PHALACROLINUS Coiffait, 1972 

angularis Ganglbauer, 1895 

glabra (Gravenhorst, 1806) non (Müller, O.F., 1776) 

LEPTACINUS Erichson, 1839 

batychrus (Gyllenhal, 1827) 

formicetorum Märkel, 1841 

intermedius Donisthorpe, 1936 

sulcifrons sensu auctt. non (Stephens, 1833) 

pusillus (Stephens, 1833) 

linearis (Gravenhorst, 1802) non (Olivier, 1795)

MEGALINUS Mulsant & Rey, 1877 458 

XANTHOLINUS sensu auctt. partim non Dejean, 

1821 

glabratus (Gravenhorst, 1802) 

NUDOBIUS Thomson, C.G., 1860 

lentus (Gravenhorst, 1806) 

PHACOPHALLUS Coiffait, 1956 

parumpunctatus (Gyllenhal, 1827) 

pallidipennis (Motschulsky, 1858) 459 

tricolor (Kraatz, 1859) 

XANTHOLINUS Dejean, 1821 

Subgenus PURROLINUS Coiffait, 1956 

elegans (Olivier, 1795) 460 

tricolor sensu auctt. partim non (Fabricius, 1787) 

meridionalis (Lacordaire, 1835) 

semirufus sensu auctt. non Reitter, 1901 

jarrigei Coiffait, 1956 

tricolor (Fabricius, 1787) 461 

meyeri Drugmand, 1994 

Subgenus TYPHLOLINUS Reitter, 1908 

laevigatus Jacobsen, 1849 

distans sensu auctt. Brit. non Mulsant & Rey, 1853 

cribripennis Fauvel, 1873 

Subgenus XANTHOLINUS Dejean, 1821 

gallicus Coiffait, 1956 

linearis (Olivier, 1795) 

longiventris Heer, 1839 

Superfamily SCARABAEOIDEA Latreille, 1802 

20. Family GEOTRUPIDAE Latreille, 1802 462 

Family author: D.J. Mann 

Subfamily BOLBOCERATINAE Mulsant, 1842 

ODONTEUS Samouelle, 1819 

ODONTAEUS Dejean, 1821 

armiger (Scopoli, 1772) 

mobilicornis (Fabricius, 1775) 

testaceus (Fabricius, 1775) 

Subfamily GEOTRUPINAE Latreille, 1802 

TYPHAEUS Leach, 1815 

TYPHOEUS auctt. (misspelling) 

MINOTAURUS Mulsant, 1855 

typhoeus (Linnaeus, 1758) 

pumilus (Marsham, 1802) 

vulgaris (Leach, 1815) 

thyphoeus (Leach, 1815) 

ANOPLOTRUPES Jekel, 1866 

GEOTRUPES sensu auctt. partim non Latreille, 

1796 

stercorosus (Scriba, 1791) 

sylvaticus (Panzer, 1798) 

55 

GEOTRUPES Latreille, 1797 

mutator (Marsham, 1802) 

spiniger (Marsham, 1802) 463 

foveatus (Marsham, 1802) 464 

puncticollis Malinowsky, 1811 

sublaevigatus Stephens, 1830 

stercorarius (Linnaeus, 1758) 

punctatostriatus Stephens, 1830 

Family GEOTRUPIDAE Latreille, 1802 

TRYPOCOPRIS Motschulsky, 1860 

GEOTRUPES sensu auctt. partim non Latreille, 

1796 

pyrenaeus (Charpentier, 1825) 

vernalis (Linnaeus, 1758) 

21. Family TROGIDAE MacLeay, 1819 465 


TROX Fabricius, 1775 

perlatus (Goeze, 1777) 

hispidus sensu auctt. Brit. non (Pontoppidan, 1763) 

sabulosus (Linnaeus, 1758) 

scaber (Linnaeus, 1767) 

arenosus (Gmelin in Linnaeus, 1790) 

arenarius (Paykull, 1798) 

trisulcatus Curtis, 1845 

22. Family LUCANIDAE Latreille, 1804 466 


Subfamily SYNDESINAE MacLeay, 1819 

SINODENDRON Hellwig, 1792 

cylindricum (Linnaeus, 1758) 

Subfamily LUCANINAE Latreille, 1804 

DORCUS MacLeay, 1819 

parallelipipedus (Linnaeus, 1758) 


LUCANUS Scopoli, 1763 

cervus (Linnaeus, 1758) 

inermis (Marsham, 1802) 

PLATYCERUS Geoffroy, 1762 


23. Family SCARABAEIDAE Latreille, 1802 467 


Subfamily AEGIALIINAE Laporte, 1840 

AEGIALIA Latreille, 1807 

Subgenus AEGIALIA Latreille, 1807 

arenaria (Fabricius, 1787) 

globosa (Kugelann, 1794)

Family SCARABAEIDAE Latreille, 1802 Checklist of Beetles of the British Isles, 2012 Edition 

Subgenus PSAMMOPORUS Thomson, C.G., 1859 

insularis Pittino, 2006 468 

sabuleti sensu auctt. Brit. non (Panzer, 1797) 

Subgenus RHYSOTHORAX Bedel, 1911 

rufa (Fabricius, 1792) 469 

rufina Silfverberg, 1977 

Subfamily APHODIINAE Leach, 1815 470 

Tribe APHODIINI Leach, 1815 

APHODIUS Hellwig, 1798 

471 472 473 

Subgenus ACROSSUS Mulsant, 1842 

depressus (Kugelann, 1792) 

luridus (Fabricius, 1775) 

gagates (Geoffroy, 1764) 

variegatus (Herbst, 1783) 

nigrosulcatus (Marsham, 1802) 

rufipes (Linnaeus, 1758) 

muticus (Stephens, 1830) 

Subgenus AGOLIINUS Schmidt, 1913 

lapponum Gyllenhal, 1808 

axillaris Stephens, 1839 

subalpinus Hardy, 1847 

nemoralis Erichson, 1848 

Subgenus AGRILINUS Mulsant & Rey, 1870 

ater (De Geer, 1774) 

terrestris (Fabricius, 1775) 

obscurus (Marsham, 1802) non (Fabricius, 1792) 

terrenus Stephens, 1830 

constans Duftschmid, 1805 

nitidus Stephens, 1830 

rufus (Moll, 1782) 474 

scybalarius (Fabricius, 1781) 

rufescens (Fabricius, 1801) 

castaneus (Marsham, 1802) 


ochraceus Stephens, 1830 

sordidus (Fabricius, 1775) 

Subgenus AMMOECIUS Mulsant, 1842 

brevis Erichson, 1848 

Subgenus APHODIUS Hellwig, 1798 

fimetarius (Linnaeus, 1758) 

foetens (Fabricius, 1787) 

scrutator sensu (Marsham, 1802) non (Herbst, 

1789) 

aestivalis Stephens, 1839 

foetidus (Herbst, 1783) 475 

scybalarius sensu auctt. non (Fabricius, 1781) 

coprinus (Marsham, 1802) 

pedellus (De Geer, 1774) 476 

fimetarius sensu auctt. partim non (Linnaeus, 1758) 

Subgenus BODILUS Mulsant & Rey, 1870 

ictericus (Laicharting, 1781) 


Subgenus CALAMOSTERNUS Motschulsky, 1860 

granarius (Linnaeus, 1767) 

emarginatus Stephens, 1830 

haemorrhous Stephens, 1830 

lucens Stephens, 1830 

melanopus Stephens, 1830 

56 

Subgenus CHILOTHORAX Motschulsky, 1860 

conspurcatus (Linnaeus, 1758) 

distinctus (Müller, O.F., 1776) 

inquinatus (Herbst, 1783) 

attaminatus (Marsham, 1802) 

foedatus (Marsham, 1802) 

paykulli Bedel, 1907 

inquinatus sensu (Olivier, 1789) non (Herbst, 1783) 

tessulatus (Paykull, 1798) non (Laicharting, 1781) 

Subgenus COLOBOPTERUS Mulsant, 1842 

erraticus (Linnaeus, 1758) 

Subgenus ESYMUS Mulsant & Rey, 1870 

merdarius (Fabricius, 1775) 

pusillus (Herbst, 1789) 

phaeopterus Stephens, 1830 

Subgenus EUORODALUS Dellacasa, G., 1983 

coenosus (Panzer, 1798) 

tristis Panzer, 1833 

Subgenus EUPLEURUS Mulsant, 1842 

subterraneus (Linnaeus, 1758) 

Subgenus LABARRUS Mulsant & Rey, 1870 

lividus (Olivier, 1789) 

bilituratus (Marsham, 1802) 

Subgenus LIMARUS Mulsant & Rey, 1870 

zenkeri Germar, 1813 

Subgenus LIOTHORAX Motschulsky, 1860 

niger (Illiger, 1798) 477 

niger (Panzer, 1797) 

plagiatus (Linnaeus, 1767) 

elongatus Stephens, 1830 

Subgenus MELINOPTERUS Mulsant, 1842 

consputus Creutzer, 1799 

prodromus (Brahm, 1790) 

punctatosulcatus Sturm, 1805 478 

sabulicola Thomson, C.G., 1868 

sphacelatus (Panzer, 1798) 

marginellus Stephens, 1830 

punctatosulcatus sensu auctt. Brit. non Sturm, 1805 

Subgenus NIMBUS Mulsant & Rey, 1870 

contaminatus (Herbst, 1783) 

ciliaris (Marsham, 1802) 

obliteratus Sturm, 1823 

Subgenus OTOPHORUS Mulsant, 1842 

haemorrhoidalis (Linnaeus, 1758) 

sanguinolentus (Marsham, 1802) 

Subgenus PHALACRONOTHUS Motschulsky, 1860 


Subgenus PLAGIOGONUS Mulsant, 1842 

arenarius (Olivier, 1789) 479 

putridus (Geoffroy in Fourcroy, 1785) 480 

rhododactylus (Marsham, 1802) 

Subgenus PLANOLINUS Mulsant & Rey, 1870 

borealis Gyllenhal, 1827 

putridus sensu (Sturm, 1805) non (Geoffroy in 


fasciatus (Olivier, 1789) 481 

putridus sensu (Herbst, 1789) non (Geoffroy in 


foetidus (Fabricius, 1792) non (Herbst, 1783) 

uliginosus Hardy, 1847 

tenellus sensu auctt. non Say, 1823

Subgenus SIGORUS Mulsant & Rey, 1870 

porcus (Fabricius, 1792) 

turpis (Marsham, 1802) 

ruficrus (Marsham, 1802) 

Subgenus TEUCHESTES Mulsant, 1842 

fossor (Linnaeus, 1758) 

Subgenus VOLINUS Mulsant & Rey, 1870 

sticticus (Panzer, 1798) 482 

equestris (Panzer, 1798) 

EUHEPTAULACUS Dellacasa, G., 1983 

HEPTAULACUS sensu auctt. non Mulsant, 1842 

sus (Herbst, 1783) 

villosus (Gyllenhal in Schönherr, 1806) 

HEPTAULACUS Mulsant, 1842 

testudinarius (Fabricius, 1775) 

OXYOMUS Dejean, 1833 

sylvestris (Scopoli, 1763) 

silvestris auctt. (misspelling) 

porcatus (Fabricius, 1775) 

platycephalus (Marsham, 1802) 

Tribe EUPARIINI Schmidt, 1910 

SAPROSITES Redtenbacher, 1858 

mendax (Blackburn, 1892) 

parallelus sensu Champion, 1921 non Harold, 1867 

natalensis (Peringuey, 1901) 483 

Tribe PSAMMODIINI Mulsant, 1842 

Subtribe PSAMMODIINA Mulsant, 1842 

BRINDALUS Landin, 1960 

porcicollis (Illiger, 1803) 484 

DIASTICTUS Mulsant, 1842 

vulneratus (Sturm, 1805) 

PSAMMODIUS Fallén, 1807 

PSAMMOBIUS Heer, 1841 

asper (Fabricius, 1775) 

sulcicollis (Illiger, 1802) 

TESARIUS Rakovic, 1981 

PSAMMODIUS sensu auctt. partim non Fallén, 

1807 

caelatus (LeConte, 1857) 

mcclayi (Cartwright, 1955) 485 

Subtribe RHYSSEMINA Pittino & Mariani, 1986 

PLEUROPHORUS Mulsant, 1842 

PSAMMOBIUS sensu auctt. partim non Heer, 1841 

caesus (Creutzer in Panzer, 1796) 

RHYSSEMUS Mulsant, 1842 


57 

Family SCARABAEIDAE Latreille, 1802 

Subfamily SCARABAEINAE Latreille, 1802 

Tribe COPRINI Leach, 1815 

COPRIS Geoffroy, 1762 

lunaris (Linnaeus, 1758) 

Tribe ONTHOPHAGINI Burmeister, 1846 

ONTHOPHAGUS Latreille, 1802 

486 487 

Subgenus ONTHOPHAGUS Latreille, 1802 

taurus (Schreber, 1759) 

Subgenus PALAEONTHOPHAGUS Zunino, 1979 

coenobita (Herbst, 1783) 

fracticornis (Preyssler, 1790) 488 

similis sensu auctt. non (Scriba, 1790) 

joannae Goljan, 1953 

ovatus sensu auctt. partim non (Linnaeus, 1767) 

medius (Kugelann, 1792) 489 

vacca sensu auctt. partim non (Linnaeus, 1767) 

nuchicornis (Linnaeus, 1758) 

verticicornis (Fabricius, 1775) 

xiphias (Fabricius, 1792) 

dillwynii (Stephens, 1830) 

similis (Scriba, 1790) 

fracticornis sensu auctt. Brit. non (Preyssler, 1790) 

verticicornis (Laicharting, 1781) 490 

nutans (Fabricius, 1787) 

Subfamily MELOLONTHINAE Leach, 1819 

Tribe HOPLIINI Latreille, 1829 

HOPLIA Illiger, 1803 

Subgenus DECAMERA Mulsant, 1842 

philanthus (Füessly, 1775) 

farinosa sensu auctt. non (Linnaeus, 1761) 491 

Tribe MELOLONTHINI Leach, 1819 

Subtribe MELOLONTHINA Leach, 1819 

MELOLONTHA Fabricius, 1775 492 

hippocastani Fabricius, 1801 

melolontha (Linnaeus, 1758) 

vulgaris Fabricius, 1775 

POLYPHYLLA Harris, 1842 

fullo (Linnaeus, 1758) 

Subtribe RHIZOTROGINA Burmeister, 1855 

AMPHIMALLON Latreille, 1825 

AMPHIMALLUS Mulsant, 1842 (misspelling) 

RHIZOTROGUS sensu auctt. partim non Berthold, 

1827 

fallenii (Gyllenhal, 1817) 493 

ochraceum sensu auctt. partim non (Knoch, 1801) 

solstitiale (Linnaeus, 1758)

Family EUCINETIDAE Lacordaire, 1857 Checklist of Beetles of the British Isles, 2012 Edition 

Tribe SERICINI Kirby, 1837 

OMALOPLIA Schönherr, 1817 

HOMALOPLIA Erichson, 1847 

ruricola (Fabricius, 1775) 

varius (Marsham, 1802) 

SERICA MacLeay, 1819 

brunnea (Linnaeus, 1758) 

Subfamily RUTELINAE MacLeay, 1819 

Tribe ANOMALINI Streubel, 1839 

ANOMALA Samouelle, 1819 

EUCHLORA MacLeay, 1819 

dubia (Scopoli, 1763) 

aenea (De Geer, 1774) 

frischii (Fabricius, 1775) 

PHYLLOPERTHA Stephens, 1830 

horticola (Linnaeus, 1758) 

arvicola sensu (Marsham, 1802) non (Fabricius, 

1775) 

Subfamily CETONIINAE Leach, 1815 494 

Tribe CETONIINI Leach, 1815 

CETONIA Fabricius, 1775 

aurata (Linnaeus, 1761) 

PROTAETIA Burmeister, 1842 

CETONIA sensu auctt. partim non Fabricius, 1775 

Subgenus POTOSIA Mulsant & Rey, 1870 

metallica (Herbst, 1782) 

cuprea sensu auctt. Brit. non (Fabricius, 1775) 

floricola (Herbst, 1790) 

Tribe TRICHIINI Fleming, 1821 

GNORIMUS Lepeletier & Audinet-Serville, 1828 495 

ALEUROSTICTUS Kirby, 1827 

nobilis (Linnaeus, 1758) 

variabilis (Linnaeus, 1758) 

octopunctatus (Fabricius, 1775) 

TRICHIUS Fabricius, 1775 

fasciatus (Linnaeus, 1758) 

gallicus Dejean, 1821 

rosaceus (Voët, 1769) 496 

zonatus sensu auctt. Brit. non Germar, 1831 

abdominalis sensu auctt. Brit. non (Ménétries, 1832) 

58 

Superfamily SCIRTOIDEA Fleming, 1821 

24. Family EUCINETIDAE Lacordaire, 1857 


EUCINETUS Germar, 1818 

meridionalis (Laporte, 1836) 

25. Family CLAMBIDAE Fischer von Waldheim, 

1821 


Subfamily CALYPTOMERINAE Crowson, 1955 

CALYPTOMERUS Redtenbacher, 1849 

CLAMBUS sensu auctt. Brit. partim non Fischer von 

Waldheim, 1821 

COMAZUS Fairmaire & Laboulbène, 1854 

dubius (Marsham, 1802) 

enshamensis (Stephens, 1829) 

Subfamily CLAMBINAE Fischer von Waldheim, 

1821 

CLAMBUS Fischer von Waldheim, 1821 

armadillo (De Geer, 1774) 

armadillus auctt. (misspelling) 

convexus (Marsham, 1802) 

atomarius (Sturm, 1807) 

nanus (Stephens, 1829) 

evae Endrödy-Younga, 1960 

minutus sensu auctt. Brit. partim non (Sturm, 1807) 

gibbulus (LeConte, 1850) 

punctulum sensu auctt. Brit. partim non (Beck, 

1817) 

radula Endrödy-Younga, 1960 

nigrellus Reitter, 1914 


nigriclavis Stephens, 1832 

minutus sensu (Stephens, 1829) non (Sturm, 1807) 

pallidulus Reitter, 1911 


pubescens Redtenbacher, 1849 

punctulum (Beck, 1817) 

punctulus auctt. (misspelling) 


borealis Strand, A., 1946 

simsoni Blackburn, 1902 497 

26. Family SCIRTIDAE Fleming, 1821 


ELODES Latreille, 1796 

HELODES sensu auctt. non Paykull, 1799 

elongata Tournier, 1868 498 

minuta sensu auctt. partim non (Linnaeus, 1767) 

koelleri Klausnitzer, 1971 

minuta (Linnaeus, 1767) 

pseudominuta Klausnitzer, 1971 498


tricuspis Nyholm, 1985 498 


elongata sensu auctt. partim non Tournier, 1868 

ODELES Klausnitzer, 2004 

marginata (Fabricius, 1798) 499 

MICROCARA Thomson, C.G., 1859 

testacea (Linnaeus, 1767) 

livida (Fabricius, 1792) non (Geoffroy in Fourcroy, 

1785) 

bohemani sensu auctt. Brit. non (Mannerheim, 

1844) 

CYPHON Paykull, 1799 

coarctatus Paykull, 1799 

paykulli Guérin-Méneville, 1843 

nitidulus Thomson, C.G., 1855 

hilaris Nyholm, 1944 

ochraceus sensu auctt. partim non Stephens, 1830 

kongsbergensis Munster, 1924 500 

laevipennis Tournier, 1868 

phragmiteticola Nyholm, 1955 501 

variabilis sensu auctt. Brit. partim non (Thunberg, 

1787) 

ochraceus Stephens, 1830 

pallidulus Boheman, 1849 

padi (Linnaeus, 1758) 

palustris Thomson, C.G., 1855 

coarctatus sensu auctt. non Paykull, 1799 

pubescens (Fabricius, 1792) 

variabilis sensu auctt. Brit. partim non (Thunberg, 

1787) 

punctipennis Sharp, 1873 

variabilis (Thunberg, 1787) 

PRIONOCYPHON Redtenbacher, 1858 

serricornis (Müller, P.W.J., 1821) 

HYDROCYPHON Redtenbacher, 1858 

deflexicollis (Müller, P.W.J., 1821) 

SCIRTES Illiger, 1807 

hemisphaericus (Linnaeus, 1758) 

orbicularis (Panzer, 1793) 

Superfamily DASCILLOIDEA Guérin-Méneville, 

1843 (1834) 

27. Family DASCILLIDAE Guérin-Méneville, 

1843 (1834) 


DASCILLUS Latreille, 1796 

cervinus (Linnaeus, 1758) 

59 

Family DASCILLIDAE Guérin-Méneville, 1843 (1834) 

Superfamily BUPRESTOIDEA Leach, 1815 

28. Family BUPRESTIDAE Leach, 1815 502 

Family author: B. Levey 

Subfamily BUPRESTINAE Leach, 1815 

Tribe ANTHAXIINI Gory & Laporte, 1839 

ANTHAXIA Eschscholtz, 1829 

Subgenus ANTHAXIA Eschscholtz, 1829 

nitidula (Linnaeus, 1758) 

Subgenus MELANTHAXIA Rikhter, 1949 

quadripunctata (Linnaeus, 1758) 503 

Tribe MELANOPHILINI Bedel, 1921 

MELANOPHILA Eschscholtz, 1829 

acuminata (De Geer, 1774) 

Subfamily AGRILINAE Laporte, 1835 

Tribe AGRILINI Laporte, 1835 

AGRILUS Curtis, 1825 

Subgenus AGRILUS Curtis, 1825 

viridis (Linnaeus, 1758) 

littlei Curtis, 1840 504 

Subgenus ANAMBUS Thomson, C.G., 1864 

angustulus (Illiger, 1803) 505 

pavidus (Fabricius, 1792) 

viridis sensu Stephens, 1830 partim non (Linnaeus, 

1758) 

biguttatus (Fabricius, 1777) 506 

pannonicus (Piller & Mitterpacher, 1783) 

cuprescens (Ménétries, 1832) 507 

cyanescens Ratzeburg, 1837 507 

laticornis (Illiger, 1803) 

viridis sensu Stephens, 1830 partim non (Linnaeus, 

1758) 

sinuatus (Olivier, 1790) 

sulcicollis Lacordaire, 1835 508 

Tribe APHANISTICINI Jacquelin du Val, 1859 

APHANISTICUS Latreille, 1829 

emarginatus (Olivier, 1790) 

pusillus (Olivier, 1790) 

Tribe TRACHEINI Laporte, 1835 

TRACHYS Fabricius, 1801 


scrobiculatus Kiesenwetter, 1857 

pumilus sensu auctt. non (Illiger, 1803) 

subglaber Rey, 1891 509 

troglodytes Gyllenhal in Schönherr, 1817

Family BYRRHIDAE Latreille, 1804 Checklist of Beetles of the British Isles, 2012 Edition 

Superfamily BYRRHOIDEA Latreille, 1804 

29. Family BYRRHIDAE Latreille, 1804 


Subfamily BYRRHINAE Latreille, 1804 

SIMPLOCARIA Stephens, 1829 

maculosa Erichson, 1847 

semistriata (Fabricius, 1794) 510 

picipes (Olivier, 1790) 

MORYCHUS Erichson, 1846 


CYTILUS Erichson, 1846 

sericeus (Forster, 1771) 

varius (Fabricius, 1775) 

BYRRHUS Linnaeus, 1767 511 

arietinus Steffahny, 1842 

fasciatus sensu auctt. Brit. partim non (Forster, 

1770) 

fasciatus (Forster, 1770) 

pilula (Linnaeus, 1758) 

dennii Curtis, 1826 

pustulatus (Forster, 1770) 

niger (Forster, 1770) 

dorsalis Fabricius, 1787 

PORCINOLUS Mulsant & Rey, 1869 

BYRRHUS sensu Fowler, 1889 partim non 

Linnaeus, 1767 

murinus (Fabricius, 1794) 

rubidus (Kugelann, 1792) 512 

undulatus (Kugelann, 1792) 512 

Subfamily SYNCALYPTINAE Mulsant & Rey, 1869 

CHAETOPHORA Kirby & Spence, 1823 

CHAETOPHORUS Kirby & Spence, 1828 

SYNCALYPTA Dillwyn, 1829 

spinosa (Rossi, 1794) 

CURIMOPSIS Ganglbauer, 1902 

SYNCALYPTA sensu auctt. partim non Dillwyn, 

1829 

maritima (Marsham, 1802) 

striatopunctata (Steffahny, 1842) 

hirsuta (Sharp, 1871) 

nigrita (Palm, 1934) 513 

setigera (Illiger, 1798) 

30. Family ELMIDAE Curtis, 1830 


ELMIS Latreille, 1802 

HELMIS Bedel, 1878 (misspelling) 

aenea (Müller, P.W.J., 1806) 

60 

megerlei sensu auctt. Brit. non (Duftschmid, 1805) 

maugei sensu auctt. Brit. non Bedel, 1878 

ESOLUS Mulsant & Rey, 1872 

ELMIS sensu Fowler, 1889 partim non Latreille, 

1802 

parallelepipedus (Müller, P.W.J., 1806) 


LIMNIUS Illiger, 1802 


1802 

LATELMIS Reitter, 1883 

volckmari (Panzer, 1793) 

volkmari auctt. (misspelling) 

MACRONYCHUS Müller, P.W.J., 1806 

quadrituberculatus Müller, P.W.J., 1806 

NORMANDIA Pic, 1900 

RIOLUS sensu Fowler, 1889 partim non Mulsant & 

Rey, 1872 

APTYKTOPHALLUS Steffan, 1958 

nitens (Müller, P.W.J., 1817) 

sodalis sensu auctt. Brit. non (Erichson, 1847) 

OULIMNIUS des Gozis, 1886 

LIMNIUS sensu auctt. Brit. non Illiger, 1802 

major (Rey, 1889) 514 

falcifer Berthélemy, 1962 

rivularis (Rosenhauer, 1856) 

variabilis sensu auctt. Brit. non (Stephens, 1828) 

troglodytes (Gyllenhal, 1827) 

tuberculatus (Müller, P.W.J., 1806) 

variabilis (Stephens, 1828) 

RIOLUS Mulsant & Rey, 1872 


1802 

cupreus (Müller, P.W.J., 1806) 

subviolaceus (Müller, P.W.J., 1817) 

STENELMIS Dufour, 1835 

canaliculata (Gyllenhal, 1808) 

31. Family DRYOPIDAE Billberg, 1820 (1817) 


POMATINUS Sturm, 1853 

HELICHUS sensu auctt. Europ. non Erichson, 

1847 515 

substriatus (Müller, P.W.J., 1806) 

DRYOPS Olivier, 1791 

Subgenus DRYOPS Olivier, 1791 

PARNUS Fabricius, 1792 

anglicanus Edwards, J., 1909 

auriculatus (Fourcroy, 1785) 

prolifericornis (Fabricius, 1792) 

ernesti des Gozis, 1886 

auriculatus sens (Fowler, 1889) non (Fourcroy, 

1785)

griseus (Erichson, 1847) 

luridus (Erichson, 1847) 

prolifericornis sensu auctt. Brit. non (Fabricius, 

1792) 

similaris Bollow, 1936 516 

griseus sensu auctt. partim non (Erichson, 1847) 

striatellus (Fairmaire & Brisout de Barneville, 1859) 

algiricus sensu auctt. Brit. non Lucas, 1846 

Subgenus YRDOPS Steffan, 1961 

nitidulus (Heer, 1841) 

32. Family LIMNICHIDAE Erichson, 1846 


LIMNICHUS Dejean, 1821 

pygmaeus (Sturm, 1807) 

33. Family HETEROCERIDAE MacLeay, 1825 


AUGYLES Schiødte, 1866 517 

HETEROCERUS sensu auctt. partim non Fabricius, 

1792 

hispidulus (Kiesenwetter, 1843) 

maritimus (Guérin-Méneville, 1844) 

sericans sensu auctt. Brit. non (Kiesenwetter, 1843) 

britannicus (Kuwert, 1890) 

HETEROCERUS Fabricius, 1792 

fenestratus (Thunberg, 1784) 

laevigatus Panzer, 1794 

pulchellus Kiesenwetter, 1843 

flexuosus Stephens, 1828 

femoralis Krynicky, 1832 

fossor Kiesenwetter, 1843 

rectus Waterhouse, G.R., 1859 

fusculus Kiesenwetter, 1843 

marginatus (Fabricius, 1787) 

obsoletus Curtis, 1828 

34. Family PSEPHENIDAE Lacordaire, 1854 


Subfamily EUBRIINAE Lacordaire, 1857 

EUBRIA Germar, 1818 

palustris Germar, 1818 

35. Family PTILODACTYLIDAE Laporte, 1836 


PTILODACTYLA Illiger, 1807 

exotica Chapin, 1927 518 

61 

Family LIMNICHIDAE Erichson, 1846 

Superfamily ELATEROIDEA Leach, 1815 

36. Family EUCNEMIDAE Eschscholtz, 1829 

Family author: H. Mendel 

Subfamily MELASINAE Fleming, 1821 

Tribe MELASINI Fleming, 1821 

MELASIS Olivier, 1790 

buprestoides (Linnaeus, 1761) 

ISORHIPIS Lacordaire, 1835 

melasoides (Laporte, 1835) 519 

Tribe EPIPHANINI Muona, 1993 

HYLIS des Gozis, 1886 

HYPOCAELUS sensu Guérin-Méneville, 1843 non 

Dejean, 1833 

HYPOCOELUS auctt. (misspelling) 

cariniceps (Reitter, 1902) 

olexai (Palm, 1955) 

procerulus sensu auctt. non (Mannerheim, 1823) 

EPIPHANIS Eschscholtz, 1829 

cornutus Eschscholtz, 1829 

Tribe DIRHAGINI Reitter, 1911 

MICRORHAGUS Dejean, 1833 

DIRHAGUS Latreille, 1834 

pygmaeus (Fabricius, 1792) 

Subfamily EUCNEMINAE Eschscholtz, 1829 

EUCNEMIS Ahrens, 1812 

capucina Ahrens, 1812 

37. Family THROSCIDAE Laporte, 1840 


AULONOTHROSCUS Horn, 1890 

TRIXAGUS sensu auctt. partim non Kugelann, 1794 

brevicollis (de Bonvouloir, 1859) 

TRIXAGUS Kugelann, 1794 

THROSCUS Latreille, 1796 

carinifrons (de Bonvouloir, 1859) 

seriatus Blair, 1942 

dermestoides (Linnaeus, 1767) 

gracilis Wollaston, 1854 

elateroides sensu auctt. Brit. non (Heer, 1841) 

obtusus (Curtis, 1827)

Family ELATERIDAE Leach, 1815 Checklist of Beetles of the British Isles, 2012 Edition 

38. Family ELATERIDAE Leach, 1815 


Subfamily AGRYPNINAE Candèze, 1857 

AGRYPNUS Eschscholtz, 1829 

ADELOCERA sensu auctt. partim non Latreille, 

1829 

LACON sensu auctt. non Laporte, 1836 

BRACHYLACON Motschulsky, 1858 

murinus (Linnaeus, 1758) 

LACON Laporte, 1836 

ADELOCERA sensu auctt. partim non Latreille, 

1829 

querceus (Herbst, 1784) 

varius (Olivier, 1790) 

Subfamily HYPNOIDINAE Schwarz, 1906 (1860) 

HYPNOIDUS Dillwyn, 1829 

CRYPTOHYPNUS Eschscholtz, 1830 

riparius (Fabricius, 1792) 

Subfamily DENTICOLLINAE Stein & Weise, 1877 

(1848) 

Tribe CTENICERINI Fleutiaux, 1936 

ACTENICERUS Kiesenwetter, 1858 

CORYMBITES sensu Fowler, 1889 partim non 

Latreille, 1834 

sjaelandicus (Müller, O.F., 1764) 

tessellatus sensu (Fabricius, 1775) non (Linnaeus, 

1758) 

ANOSTIRUS Thomson, C.G., 1859 

CALOSTIRUS Thomson, C.G., 1864 



castaneus (Linnaeus, 1758) 

CTENICERA Latreille, 1829 

LUDIUS sensu Eschscholtz, 1829 non Berthold, 

1827 

CTENICERUS Stephens, 1830 (misspelling) 

CORYMBITES Latreille, 1834 

cuprea (Fabricius, 1775) 

aeruginosa sensu (Fabricius, 1798) non (Olivier, 

1790) 

pectinicornis (Linnaeus, 1758) 

CALAMBUS Thomson, C.G., 1859 

SELATOSOMUS sensu auctt. partim non Stephens, 

1830 


APLOTARSUS Stephens, 1830 


1830 

angustulus (Kiesenwetter, 1858) 

incanus (Gyllenhal, 1827) 

62 

quercus sensu (Gyllenhal, 1808) non (Olivier, 1790) 

ochropterus (Stephens, 1830) 

PARAPHOTISTUS Kishii, 1966 


1830 

impressus (Fabricius, 1792) 

nigricornis (Panzer, 1799) 

metallicus (Paykull, 1800) 

PROSTERNON Latreille, 1834 

CORYMBITES sensu Fowler, 1889 non Latreille, 

1834 

tessellatum (Linnaeus, 1758) 

holosericeum (Olivier, 1790) 

SELATOSOMUS Stephens, 1830 



aeneus (Linnaeus, 1758) 

cruciatus (Linnaeus, 1758) 

melancholicus (Fabricius, 1798) 520 

Tribe DENTICOLLINI Stein & Weise, 1877 (1848) 

CIDNOPUS Thomson, C.G., 1859 

LIMONIUS sensu auctt. partim non Eschscholtz, 

1829 

aeruginosus (Olivier, 1790) 

cylindricus (Rossi, 1792) 

KIBUNEA Kishii, 1966 

LIMONIUS sensu auctt. partim non Eschscholtz, 

1829 

CIDNOPUS sensu auctt. partim non Thomson, 

C.G., 1859 


LIMONISCUS Reitter, 1905 

violaceus (Müller, P.W.J., 1821) 

DENTICOLLIS Piller & Mitterpacher, 1783 

CAMPYLUS Fischer von Waldheim, 1823/4 

linearis (Linnaeus, 1758) 

ATHOUS Eschscholtz, 1829 

Subgenus ATHOUS Eschscholtz, 1829 


obscurus sensu (Paykull, 1800) non (Linnaeus, 

1758) 

vittatus (Fabricius, 1792) 521 

advena (Scopoli, 1763) 32 

Subgenus EXANATHROTUS Méquignon, 1930 

subfuscus (Müller, O.F., 1764) 

Subgenus ORTHATHOUS Reitter, 1905 

bicolor (Goeze, 1777) 

longicollis (Olivier, 1790) 

campyloides Newman, 1833 

difformis Lacordaire, 1835 

cavus sensu Waterhouse, G.R., 1858 non (Germar, 

1817)

DIACANTHOUS Reitter, 1905 

HARMINIUS sensu auctt. non Fairmaire, 1852 

undulatus (De Geer, 1774) 

STENAGOSTUS Thomson, C.G., 1859 

ATHOUS sensu auctt. Brit. partim non Eschscholtz, 

1829 

rhombeus (Olivier, 1790) 

villosus sensu auctt. non (Fourcroy, 1785) 

HEMICREPIDIUS Germar, 1839 

ATHOUS sensu auctt. Brit. partim non Eschscholtz, 

1829 

hirtus (Herbst, 1784) 

niger sensu auctt. Brit. non (Linnaeus, 1758) 

Subfamily ELATERINAE Leach, 1815 

Tribe ADRASTINI Candèze, 1863 

ADRASTUS Eschscholtz, 1829 

pallens (Fabricius, 1792) 

nitidulus (Marsham, 1802) non (Fabricius, 1787) 

limbatus sensu (Stephens, 1830) non (Fabricius, 

1777) 

rachifer (Fourcroy, 1785) 

minutus sensu (Olivier, 1790) non (Linnaeus, 1758) 

pusillus (Fabricius, 1801) 

SYNAPTUS Eschscholtz, 1829 

filiformis (Fabricius, 1781) 

Tribe AGRIOTINI Laporte, 1840 

AGRIOTES Eschscholtz, 1829 

acuminatus (Stephens, 1830) 

sobrinus Kiesenwetter, 1858 

lineatus (Linnaeus, 1767) 

obscurus (Linnaeus, 1758) 

pallidulus (Illiger, 1807) 

sordidus (Illiger, 1807) 

sputator (Linnaeus, 1758) 

DALOPIUS Eschscholtz, 1829 

DOLOPIUS auctt. (misspelling) 

marginatus (Linnaeus, 1758) 

Tribe AMPEDINI Gistel, 1848 

AMPEDUS Dejean, 1833 

ELATER sensu auctt. non Linnaeus, 1758 

balteatus (Linnaeus, 1758) 

cardinalis (Schiødte, 1865) 

praeustus sensu auctt. non (Fabricius, 1792) 

coccinatus (Rye, 1867) 

cinnabarinus (Eschscholtz, 1829) 

lythropterus Germar, 1844 

elongantulus (Fabricius, 1787) 

elongatulus auctt. (misspelling) 

nigerrimus (Lacordaire, 1835) 

aethiops sensu auctt. Brit. non (Lacordaire, 1835) 

nigrinus (Herbst, 1784) 

pomonae (Stephens, 1830) 

pomorum (Herbst, 1784) 

63 

Family ELATERIDAE Leach, 1815 

ferrugatus (Lacordaire, 1835) 

quercicola (du Buysson, 1887) 

pomonae sensu auctt. non (Stephens, 1830) 

miniatus (Gorham, 1892) 

rufipennis (Stephens, 1830) 

sanguineus (Linnaeus, 1758) 

sanguinolentus (Schrank, 1776) 

tristis (Linnaeus, 1758) 

BRACHYGONUS du Buysson, 1912 

AMPEDUS sensu auctt. partim non Dejean, 1833 

ruficeps (Mulsant & Guillebeau, 1855) 

pallidulus (Reitter, 1911) 

ISCHNODES Germar, 1844 

sanguinicollis (Panzer, 1793) 

Tribe MEGAPENTHINI Gurjeva, 1973 

MEGAPENTHES Kiesenwetter, 1858 

lugens (Redtenbacher, 1842) 

PROCRAERUS Reitter, 1905 

MEGAPENTHES sensu Fowler, 1889 partim non 

Kiesenwetter, 1858 

tibialis (Lacordaire, 1835) 

Tribe ELATERINI Leach, 1815 

ELATER Linnaeus, 1758 

LUDIUS Berthold, 1827 

ferrugineus Linnaeus, 1758 

SERICUS Eschscholtz, 1829 

SERICOSOMUS Dejean, 1833 

brunneus (Linnaeus, 1758) 

Tribe incertae sedis 

PANSPAEUS Sharp, 1877 

PANSPOEUS auctt. (misspelling) 

guttatus Sharp, 1877 522 

Subfamily MELANOTINAE Candèze, 1859 (1848) 

MELANOTUS Eschscholtz, 1829 

castanipes (Paykull, 1800) 523 

villosus sensu auctt. partim non (Geoffory in 


punctolineatus (Pelerin, 1829) 

niger sensu (Fabricius, 1792) non (Linnaeus, 1758) 

aterrimus sensu (Stephens, 1830) non (Linnaeus, 

1761) 

villosus (Geoffroy in Fourcroy, 1785) 

rufipes (Herbst, 1784) non (De Geer, 1774) 

erythropus (Gmelin in Linnaeus, 1790) 

rugosus (Marsham, 1802)

Family DRILIDAE Blanchard, 1845 Checklist of Beetles of the British Isles, 2012 Edition 

Subfamily NEGASTRIINAE Nakane & Kishii, 1956 

FLEUTIAUXELLUS Méquignon, 1930 

HYPNOIDUS sensu auctt. Brit. partim non Dillwyn, 

1829 

maritimus (Curtis, 1840) 

NEGASTRIUS Thomson, C.G., 1859 


1829 

arenicola (Boheman, 1852) 524 

pulchellus (Linnaeus, 1761) 

sabulicola (Boheman, 1851) 

OEDOSTETHUS LeConte, 1853 


1829 

FLEUTIAUXELLUS sensu auctt. partim non 

Méquignon, 1930 

quadripustulatus (Fabricius, 1792) 

ZOROCHROS Thomson, C.G., 1859 


1829 

meridionalis (Laporte, 1840) 

minimus (Lacordaire, 1835) 

dermestoides (Herbst, 1806) non (Linnaeus, 1767) 

quadriguttatus sensu auctt. Brit. non (Laporte, 1840) 

tetragraphus sensu auctt. Brit. non (Germar, 1844) 

flavipes sensu auctt. Brit. non (Aubé, 1850) 

Subfamily CARDIOPHORINAE Candèze, 1859 

CARDIOPHORUS Eschscholtz, 1829 

asellus Erichson, 1840 

gramineus (Scopoli, 1763) 

thoracicus (Fabricius, 1775) 

ruficollis (Linnaeus, 1758) 

vestigialis Erichson, 1840 

erichsoni du Buysson, 1901 

DICRONYCHUS Brullé, 1832 

CARDIOPHORUS sensu auctt. partim non 

Eschscholtz, 1829 

equisetioides Lohse, 1976 

equiseti sensu auctt. Brit. non (Herbst, 1784) 

39. Family DRILIDAE Blanchard, 1845 


DRILUS Olivier, 1790 

flavescens (Fourcroy, 1785) 

64 

40. Family LYCIDAE Laporte, 1836 


Subfamily EROTINAE LeConte, 1881 525 

Tribe EROTINI LeConte, 1881 

PLATYCIS Thomson, C.G., 1859 

minutus (Fabricius, 1787) 

EROTIDES Waterhouse, C.O., 1879 

cosnardi (Chevrolat, 1829) 

Tribe DICTYOPTERINI Kleine, 1928 

PYROPTERUS Mulsant, 1838 

nigroruber (De Geer, 1774) 


DICTYOPTERA Latreille, 1829 

EROS Newman, 1838 

aurora (Herbst, 1784) 

41. Family LAMPYRIDAE Rafinesque, 1815 


LAMPYRIS Geoffroy, 1762 

noctiluca (Linnaeus, 1758) 

LAMPROHIZA Motschulsky, 1853 

splendidula (Linnaeus, 1767) 526 

PHOSPHAENUS Laporte, 1833 

hemipterus (Goeze, 1777) 

42. Family CANTHARIDAE Imhoff, 1856 (1815) 

Family author: K.N.A. Alexander 

Subfamily CANTHARINAE Imhoff, 1856 (1815) 

Tribe PODABRINI Gistel, 1856 

PODABRUS Westwood, 1838 


Tribe CANTHARINI Imhoff, 1856 (1815) 

ANCISTRONYCHA Märkel in Kiesenwetter, 1852 

abdominalis (Fabricius, 1798) 

cyanea (Curtis, 1828) 

CANTHARIS Linnaeus, 1758 

TELEPHORUS Schaeffer, 1766 

cryptica Ashe, 1947 

pallida sensu auctt. partim non Goeze, 1777

icolor sensu (Fowler, 1889) partim non Fabricius, 

1798 

decipiens Baudi, 1871 

haemorrhoidalis sensu auctt. Brit. non Fabricius, 

1792 

clypeata sensu (Fowler, 1889) non Illiger, 1798 

figurata Mannerheim, 1843 

scotica (Sharp, 1866) 

cruachana Chitty, 1893 

fusca Linnaeus, 1758 

lateralis Linnaeus, 1758 

oralis Germar, 1824 

livida Linnaeus, 1758 

rufipes Herbst, 1784 

dispar Fabricius, 1792 

nigra (De Geer, 1774) 

fulvicollis Fabricius, 1792 non Scopoli, 1763 

flavilabris Fallén, 1807 

nigricans (Müller, O.F., 1776) 

discoidea (Stephens, 1830) 

obscura Linnaeus, 1758 

pallida Goeze, 1777 

bicolor Panzer, 1797 non Linnaeus, 1763 

bicolor sensu (Fowler, 1889) partim non Fabricius, 

1798 

paludosa Fallén, 1807 

pellucida Fabricius, 1792 

rufa Linnaeus, 1758 

lituratus Fallén, 1807 

darwiniana (Sharp, 1866) 

rustica Fallén, 1807 

thoracica (Olivier, 1790) 

bicolor Herbst, 1784 non Linnaeus, 1763 

?caeruleocephala Thunberg, 1784 

RHAGONYCHA Eschscholtz, 1830 

elongata (Fallén, 1807) 

fulva (Scopoli, 1763) 

lignosa (Müller, O.F., 1764) 

pallida (Fabricius, 1787) non (Goeze, 1777) 

limbata Thomson, C.G., 1864 

femoralis sensu auctt. Brit. non (Brullé, 1832) 

lutea (Müller, O.F., 1764) 

fuscicornis (Olivier, 1790) 

testacea (Linnaeus, 1758) 

translucida (Krynicki, 1832) 

pilosa sensu (Stephens, 1830) non (Paykull, 1798) 

unicolor sensu Fowler, 1889 ?non (Curtis, 1840) 

Subfamily SILINAE Mulsant, 1862 

SILIS Charpentier, 1825 

CRUDOSILIS Kazantsev, 1995 

ruficollis (Fabricius, 1775) 

Subfamily MALTHININAE Kiesenwetter, 1852 

Tribe MALTHININI Kiesenwetter, 1852 

MALTHINUS Latreille, 1806 

balteatus Suffrian, 1851 

flaveolus (Herbst, 1786) 

punctatus sensu Fowler, 1889 ?non (Fourcroy, 

1785) 

frontalis (Marsham, 1802) 

seriepunctatus Kiesenwetter, 1852 

65 

Family DERODONTIDAE LeConte, 1861 

fasciatus sensu auctt. non (Olivier, 1790) 

Tribe MALTHODINI Böving & Craighead, 1931 

MALTHODES Kiesenwetter, 1852 

crassicornis (Mäklin, 1846) 

nigellus sensu Fowler 1890 non Kiesenwetter, 1852 

brevicollis sensu auctt. Brit. non (Paykull, 1798) 527 

dispar (Germar, 1824) 

fibulatus Kiesenwetter, 1852 

flavoguttatus Kiesenwetter, 1852 

fuscus (Waltl, 1838) 

pellucidus Kiesenwetter, 1852 

guttifer Kiesenwetter, 1852 

lobatus Kiesenwetter, 1852 528 

marginatus (Latreille, 1806) 

maurus (Laporte, 1840) 

misellus sensu auctt. non Kiesenwetter, 1852 

minimus (Linnaeus, 1758) 

marginicollis Schilsky, 1892 

mysticus Kiesenwetter, 1852 

pumilus (Brébisson, 1835) 

brevicollis sensu Kiesenwetter, 1852 non (Paykull, 

1798) 

atomus Thomson, C.G., 1864 

Superfamily DERODONTOIDEA LeConte, 1861 

43. Family DERODONTIDAE LeConte, 1861 


Subfamily LARICOBIINAE Mulsant & Rey, 1864 

LARICOBIUS Rosenhauer, 1846 

erichsonii Rosenhauer, 1846 529 

erichsoni auctt. (misspelling) 

Superfamily BOSTRICHOIDEA Latreille, 1802 

44. Family DERMESTIDAE Latreille, 1804 530 


Subfamily THYLODRIINAE Semenov, 1909 

THYLODRIAS Motschulsky, 1839 

contractus Motschulsky, 1839 

Subfamily THORICTINAE Agassiz, 1846 

THORICTODES Reitter, 1875 

THAUMAPHRASTUS Blaisdell, 1927 

heydeni Reitter, 1875 531 

karanisensis (Blaisdell, 1927)

Family BOSTRICHIDAE Latreille, 1802 Checklist of Beetles of the British Isles, 2012 Edition 

Subfamily DERMESTINAE Latreille, 1804 

DERMESTES Linnaeus, 1758 

Subgenus DERMESTINUS Zhantiev, 1967 

carnivorus Fabricius, 1775 

carniforus Fabricius, 1775 

frischii Kugelann, 1792 

maculatus De Geer, 1774 

vulpinus Fabricius, 1781 

murinus Linnaeus, 1758 

undulatus Brahm, 1790 

Subgenus DERMESTES Linnaeus, 1758 

ater De Geer, 1774 

cadaverinus Fabricius, 1775 

haemorrhoidalis Küster, 1852 

peruvianus sensu auctt. Brit. partim non Laporte, 

1840 

lardarius Linnaeus, 1758 

leechi Kalik, 1952 

peruvianus Laporte, 1840 

oblongus sensu auctt. Brit. non Solier in Gay, 1849 

Subfamily ATTAGENINAE Laporte, 1840 

ATTAGENUS Latreille, 1802 

brunneus Faldermann, 1835 

elongatulus Casey, 1900 

cyphonoides Reitter, 1881 

alfierii Pic, 1910 

fasciatus (Thunberg, 1795) 

gloriosae (Fabricius, 1798) 

pellio (Linnaeus, 1758) 

smirnovi Zhantiev, 1973 532 

trifasciatus (Fabricius, 1787) 

unicolor (Brahm, 1791) 

piceus (Olivier, 1790) non (Thunberg, 1781) 

megatoma (Fabricius, 1798) 

Subfamily MEGATOMINAE Leach, 1815 

TROGODERMA Dejean, 1821 

angustum (Solier, 1849) 533 

glabrum (Herbst, 1783) 

boron Beal, 1954 

granarium Everts, 1898 

khapra Arrow, 1917 

inclusum LeConte, 1854 

versicolor sensu auctt. Brit. non (Creutzer, 1799) 

variabile Ballion, 1878 534 

parabile Beal, 1954 

REESA Beal, 1967 

vespulae (Milliron, 1939) 535 

GLOBICORNIS Latreille, 1829 

rufitarsis (Creutzer in Panzer, 1796) 

nigripes (Fabricius, 1792) non (Olivier, 1790) 

plantaris (Curtis, 1838) 

MEGATOMA Herbst, 1792 

undata (Linnaeus, 1758) 

66 

ANTHRENOCERUS Arrow, 1915 

australis (Hope, 1843) 

ORPHINUS Motschulsky, 1858 

fulvipes (Guérin-Méneville, 1838) 

brevicornis (Sharp, 1885) 

CTESIAS Stephens, 1830 

TIRESIAS Stephens, 1835 

serra (Fabricius, 1792) 

Subfamily ANTHRENINAE Gistel, 1848 

ANTHRENUS Geoffroy, 1762 

Subgenus ANTHRENUS Müller, O.F., 1764 

flavipes LeConte, 1854 

vorax Waterhouse, C.O. 1883 

pimpinellae Fabricius, 1775 

scrophulariae (Linnaeus, 1758) 

Subgenus ANTHRENODES Chobaut, 1898 

sarnicus Mroczkowski, 1963 

Subgenus ANTHRENOPS Reitter, 1881 

coloratus Reitter, 1881 536 

Subgenus FLORILINUS Mulsant & Rey, 1868 

museorum (Linnaeus, 1761) 

olgae Kalik, 1946 537 

Subgenus HELOCERUS Mulsant & Rey, 1868 

fuscus Olivier, 1789 

claviger Erichson, 1846 

Subgenus NATHRENUS Casey, 1900 

verbasci (Linnaeus, 1767) 


Subfamily TRINODINAE Casey, 1900 

TRINODES Dejean, 1821 

hirtus (Fabricius, 1781) 

45. Family BOSTRICHIDAE Latreille, 1802 


Subfamily BOSTRICHINAE Latreille, 1802 

BOSTRICHUS Geoffroy, 1762 538 

BOSTRYCHUS auctt. (misspelling) 

capucinus (Linnaeus, 1758) 

Subfamily DINODERINAE Thomson, C.G., 1863 

RHYZOPERTHA Stephens, 1830 

RHIZOPERTHA Guérin-Méneville, 1845 

(misspelling) 

dominica (Fabricius, 1792) 

pusilla (Fabricius, 1798) 

STEPHANOPACHYS Waterhouse, C.O., 1888 

substriatus (Paykull, 1800)

Subfamily LYCTINAE Billberg, 1820 

LYCTUS Fabricius, 1792 

XYLOTROGUS Stephens, 1830 

brunneus (Stephens, 1830) 

linearis (Goeze, 1777) 

fuscus (Linnaeus, 1767) 

canaliculatus Fabricius, 1792 

46. Family PTINIDAE Latreille, 1802 


Subfamily EUCRADINAE LeConte, 1861 

HEDOBIA Dejean, 1821 

Subgenus PTINOMORPHUS Mulsant & Rey, 1868 

imperialis (Linnaeus, 1767) 

Subfamily PTININAE Latreille, 1802 

Tribe GIBBIINI Jacquelin du Val, 1860 

GIBBIUM Scopoli, 1777 

SCOTIAS de Czenpinski, 1778 

aequinoctiale Boieldieu, 1854 539 

psylliodes sensu auctt. partim non (de Czenpinski, 

1778) 

psylloides (de Czenpinski, 1778) 

scotias (Fabricius, 1781) 

STETHOMEZIUM Hinton, 1943 

squamosum Hinton, 1943 

MEZIUM Samouelle, 1819 

affine Boieldieu, 1856 

sulcatum sensu (Marsham, 1802) non (Fabricius, 

1781) 

Tribe PTININI Latreille, 1802 

TRIGONOGENIUS Solier, 1849 

globulus Solier, 1849 

SPHAERICUS Wollaston, 1854 

gibboides (Boieldieu, 1854) 

NIPTUS Boieldieu, 1856 

hololeucus (Faldermann, 1835) 

TIPNUS Thomson, C.G., 1859 

NIPTUS sensu Fowler, 1890 partim non Boieldieu, 

1856 

unicolor (Piller & Mitterpacher, 1783) 


PSEUDEUROSTUS von Heyden, L., 1906 

EUROSTUS Mulsant & Rey, 1868 non Dallas, 1851 

hilleri (Reitter, 1877) 

67 

Family PTINIDAE Latreille, 1802 

PTINUS Linnaeus, 1767 540 

BRUCHUS Geoffroy, 1762 541 

clavipes Panzer, 1792 

latro sensu auctt. partim non Fabricius, 1775 

testaceus Olivier, 1790 non Thunberg, 1784 

brunneus Duftschmid, 1825 non Gmelin in 


hirtellus Sturm, 1837 

female form mobilis Moore, 1957 

dubius Sturm, 1837 542 

exulans Erichson, 1842 

fur (Linnaeus, 1758) 

latro Fabricius, 1775 

latefasciatus Gorham, 1883 

lichenum Marsham, 1802 

palliatus Perris, 1847 

germanus sensu Fabricius, 1781 non Goeze, 1777 

?Linnaeus, 1767 

pusillus Sturm, 1837 

raptor Sturm, 1837 

sexpunctatus Panzer, 1792 

subpilosus Sturm, 1837 

tectus Boieldieu, 1856 

villiger Reitter, 1884 

Subfamily DRYOPHILINAE Gistel, 1848 

GRYNOBIUS Thomson, C.G., 1859 

PRIOBIUM sensu auctt. non Motschulsky, 1845 


tricolor (Olivier, 1790) 

castaneus (Fabricius, 1792) 

excavatus (Kugelann, 1792) 

eichhoffi (Seidlitz, 1889) 

kiesenwetteri (Edwards, 1921) 

DRYOPHILUS Chevrolat, 1832 

anobioides Chevrolat, 1832 

pusillus (Gyllenhal, 1808) 

Subfamily ERNOBIINAE Pic, 1912 

OCHINA Dejean, 1821 

ptinoides (Marsham, 1802) 

hederae (Müller, P.W.J., 1821) 

XESTOBIUM Motschulsky, 1845 

rufovillosum (De Geer, 1774) 

tessellatum (de Villers, C.J., 1789) 

ERNOBIUS Thomson, C.G., 1859 

abietis (Fabricius, 1792) 543 

angusticollis (Ratzeburg, 1837) 

parvicollis (Mulsant & Rey, 1863) 

gigas (Mulsant & Rey, 1863) 

reflexus sensu auctt. non (Mulsant & Rey, 1863) 

subelongatus Pic, 1914 

mulsantianus Sharp, 1916 

mollis (Linnaeus, 1758) 

reversus Sharp, 1916 

nigrinus (Sturm, 1837) 

pini (Sturm, 1837) 

oblitus Sharp, 1916

Family LYMEXYLIDAE Fleming, 1821 Checklist of Beetles of the British Isles, 2012 Edition 

Subfamily ANOBIINAE Fleming, 1821 

STEGOBIUM Motschulsky, 1860 

SITODREPA Thomson, C.G., 1863 

paniceum (Linnaeus, 1758) 

GASTRALLUS Jacquelin du Val, 1860 

immarginatus (Müller, P.W.J., 1821) 

laevigatus sensu Donisthorpe, 1936 non (Olivier, 

1790) 

laevigatus (Olivier, 1790) 544 

ANOBIUM Fabricius, 1775 

HADROBREGMUS sensu auctt. Europ. partim non 

Thomson, C.G., 1859 

HEMICOELUS LeConte, 1861 

fulvicorne (Sturm, 1837) 

inexspectatum Lohse, 1954 

nitidum Fabricius, 1792 545 

punctatum (De Geer, 1774) 

domesticum (Geoffroy in Fourcroy, 1785) 

striatum sensu Olivier, 1790 non Fabricius, 1787 

HADROBREGMUS Thomson, C.G., 1859 

ANOBIUM sensu auctt. partim non Fabricius, 1775 

COELOSTETHUS LeConte, 1861 

DENDROBIUM Mulsant & Rey, 1864 

denticollis (Creutzer in Panzer, 1796) 

PRIOBIUM Motschulsky, 1845 

carpini (Herbst, 1793) 546 

Subfamily PTILININAE Shuckard, 1839 

PTILINUS Geoffroy, 1762 547 


Subfamily XYLETININAE Gistel, 1848 

XYLETINUS Latreille, 1809 

longitarsis Jansson, 1942 

ater sensu auctt. Brit. partim non (Creutzer in 

Panzer, 1796) 

LASIODERMA Stephens, 1835 

serricorne (Fabricius, 1792) 

Subfamily DORCATOMINAE Thomson, C.G., 

1859 

MESOCOELOPUS Jacquelin du Val, 1860 

collaris Mulsant & Rey, 1864 548 

DORCATOMA Herbst, 1792 

DORKATOMA Herbst, 1792 549 

ambjoerni Baranowski, 1985 550 

chrysomelina Sturm, 1837 

dresdensis Herbst, 1792 

flavicornis (Fabricius, 1792) 

substriata Hummel, 1829 

serra (Panzer, 1795) non Fabricius, 1792 

68 

punctulata sensu Sharp, 1914 non Mulsant & Rey, 

1864 

CAENOCARA Thomson, C.G., 1859 


subglobosa sensu Donisthorpe, 1918 non Mulsant & 

Rey, 1864 

bovistae (Hoffmann, J., 1803) 

ANITYS Thomson, C.G., 1863 

rubens (Hoffmann, J., 1803) 

MIROSTERNOMORPHUS Español, 1977 

heali Bercedo & Arnáiz, 2010 551 

Superfamily LYMEXYLOIDEA Fleming, 1821 

47. Family LYMEXYLIDAE Fleming, 1821 


Subfamily HYLECOETINAE Germar, 1818 

HYLECOETUS Latreille, 1806 

dermestoides (Linnaeus, 1761) 

Subfamily LYMEXYLINAE Fleming, 1821 

LYMEXYLON Fabricius, 1775 

navale (Linnaeus, 1758) 

Superfamily CLEROIDEA Latreille, 1802 

48. Family PHLOIOPHILIDAE Kiesenwetter, 

1863 


PHLOIOPHILUS Stephens, 1830 

PHLOEOPHILUS auctt. (misspelling) 

edwardsii Stephens, 1830 

edwardsi auctt. (misspelling) 

49. Family TROGOSSITIDAE Latreille, 1802 


Subfamily PELTINAE Latreille, 1806 

OSTOMA Laicharting, 1781 

ferrugineum (Linnaeus, 1758) 

THYMALUS Latreille, 1802 

limbatus (Fabricius, 1787)

Subfamily LOPHOCATERINAE Crowson, 1964 

LOPHOCATERES Olliff, 1883 

pusillus (Klug, 1832) 552 

Subfamily TROGOSSITINAE Latreille, 1802 

TENEBROIDES Piller & Mitterpacher, 1783 

TENEBRIOIDES auctt. (misspelling) 

mauritanicus (Linnaeus, 1758) 

NEMOZOMA Latreille, 1804 

NEMOSOMA Latreille, 1809 40 

elongatum (Linnaeus, 1761) 

50. Family CLERIDAE Latreille, 1802 553 


Subfamily THANEROCLERINAE Chapin, 1924 

THANEROCLERUS Lefebvre, 1838 

buqueti (Lefebvre, 1835) 554 

Subfamily TILLINAE Fischer von Waldheim, 1813 

TILLUS Olivier, 1790 

elongatus (Linnaeus, 1758) 

TILLOIDEA Laporte, 1833 

unifasciata (Fabricius, 1787) 

Subfamily CLERINAE Latreille, 1802 

OPILO Latreille, 1802 

mollis (Linnaeus, 1758) 

THANASIMUS Latreille, 1806 

femoralis (Zetterstedt, 1838) 

rufipes (Brahm, 1797) non (De Geer, 1775) 

formicarius (Linnaeus, 1758) 

TRICHODES Herbst, 1792 

alvearius (Fabricius, 1792) 

alveolarius Latreille, 1804 

interruptus Kraatz, 1894 

apiarius (Linnaeus, 1758) 

Subfamily TARSOSTENINAE Jacquelin du Val, 

1860 

PARATILLUS Gorham, 1878 

carus (Newman, 1840) 

TARSOSTENUS Spinola, 1844 

univittatus (Rossi, 1792) 

69 

Family CLERIDAE Latreille, 1802 

Subfamily KORYNETINAE Laporte, 1836 

KORYNETES Herbst, 1792 

CORYNETES Paykull, 1798 40 

caeruleus (De Geer, 1775) 

cyanellus Dejean, 1837 

NECROBIA Olivier, 1795 

ruficollis (Fabricius, 1775) 

rufipes (De Geer, 1775) 

violacea (Linnaeus, 1758) 

51. Family DASYTIDAE Laporte, 1840 555 


Subfamily RHADALINAE LeConte, 1861 

APLOCNEMUS Stephens, 1830 

HAPLOCNEMUS auctt. (misspelling) 

impressus (Marsham, 1802) 

pini (Redtenbacher, 1849) 


Subfamily DASYTINAE Laporte, 1840 

DASYTES Paykull, 1799 


serricornis Stephens, 1830 

aerosus Kiesenwetter, 1867 

cyaneus (Fabricius, 1775) 556 

caeruleus (De Geer, 1774) non (Linnaeus, 1758) 

niger (Linnaeus, 1761) 

floralis sensu (Stephens, 1830) non (Olivier, 1790) 

plumbeus (Müller, O.F., 1776) 


tibialis Zetterstedt, 1828 

flavipes sensu Joy, 1932 non (Olivier, 1790) 

oculatus sensu auctt. Brit. non Kiesenwetter, 1867 

virens (Marsham, 1802) 

flavipes sensu (Olivier, 1790) non (Fabricius, 1777) 

flavipes sensu Fowler, 1889 non (Fabricius, 1777) 

puncticollis Reitter, 1888 non Germain, 1855 

plumbeus sensu Joy, 1932 non (Müller, O.F., 1776) 

PSILOTHRIX Küster, 1850 

viridicoeruleus (Geoffroy in Fourcroy, 1785) 

cyaneus (Olivier, 1790) 

nobilis (Illiger, 1798) 

caeruleus sensu (Stephens, 1830) non (Fabricius, 

1775) 

viridis sensu (Stephens, 1830) non (Rossi, 1792) 

DOLICHOSOMA Stephens, 1830 

lineare (Rossi, 1794) 

52. Family MALACHIIDAE Fleming, 1821 557 


COLOTES Erichson, 1840 

punctatus (Erichson, 1840) 558

Family SPHINDIDAE Jacquelin du Val, 1860 Checklist of Beetles of the British Isles, 2012 Edition 

EBAEUS Erichson, 1840 

pedicularius (Linnaeus, 1758) 

productus sensu (Stephens, 1830) non (Olivier, 

1790) 

HYPEBAEUS Kiesenwetter, 1863 

EBAEUS sensu auctt. Brit. partim non Erichson, 

1840 


abietinus sensu auctt. Brit. non Abeille de Perrin, 

1869 

AXINOTARSUS Motschulsky, 1853 

marginalis (Laporte, 1840) 

pulicarius (Fabricius, 1777) 

ruficollis (Olivier, 1790) 

TROGLOPS Erichson, 1840 

cephalotes (Olivier, 1790) 559 

CORDYLEPHERUS Evers, 1985 

MALACHIUS sensu auctt. partim non Fabricius, 

1775 

viridis (Fabricius, 1787) 

MALACHIUS Fabricius, 1775 

aeneus (Linnaeus, 1758) 


immaculicollis Rey, 1867 

lusitanicus var. australis sensu Donisthorpe, 1931 

non Rey, 1867 

CLANOPTILUS Motschulsky, 1854 

MALACHIUS sensu auctt. partim non Fabricius, 

1775 

barnevillei (Puton, 1865) 

marginellus (Olivier, 1790) 

elegans sensu (Donisthorpe, 1931) non (Geoffroy in 


pseudosardous (Reclairie & van der Wiel, 1932) 

angustimarginalis (Donisthorpe, 1933) 

strangulatus (Abeille de Perrin, 1885) 

vulneratus Abeille de Perrin, 1891 

SPHINGINUS Mulsant & Rey, 1867 

lobatus (Olivier, 1790) 560 

CERAPHELES Mulsant & Rey, 1867 

DIAPHONUS Mulsant & Rey, 1867 

ANTHOCOMUS sensu Fowler, 1889 partim non 


terminatus (Ménétries, 1832) 

ANTHOCOMUS Erichson, 1840 

fasciatus (Linnaeus, 1758) 

rufus (Herbst, 1784) 

coccineus (Schaller, 1783) non (Linnaeus, 1761) 

70 

Superfamily CUCUJOIDEA Latreille, 1802 

53. Family SPHINDIDAE Jacquelin du Val, 1860 


SPHINDUS Dejean, 1821 

dubius (Gyllenhal, 1808) 

ASPIDIPHORUS Dejean, 1821 

ARPIDIPHORUS Dejean, 1821 561 

orbiculatus (Gyllenhal, 1808) 

54. Family KATERETIDAE Kirby, 1837 562 


KATERETES Herbst, 1793 

CATERETES auctt. (misspelling) 

CERCUS Latreille, 1796 

ANISOCERA Stephens, 1832 

ANOMAEOCERA Shuckard, 1840 


similis (Marsham, 1802) 


spiraeae (Stephens, 1830) 

pusillus (Thunberg, 1794) 

bipustulatus (Paykull, 1798) non (Thunberg, 

1781) 563 

punctulatus (Marsham, 1802) 

rufilabris (Latreille, 1807) 

caricis Stephens, 1830 

junci Stephens, 1830 

BRACHYPTERUS Kugelann, 1794 

glaber (Newman, 1834) 

pubescens Erichson, 1843 

urticae (Fabricius, 1792) 

erythropus (Marsham, 1802) 

BRACHYPTEROLUS Grouvelle, 1913 

HETEROSTOMUS Jacquelin du Val, 1858 non 

Bigot, 1857 

BRACHYPTERUS sensu Fowler, 1889 partim non 

Kugelann, 1794 

antirrhini (Murray, 1864) 

villiger (Reitter, 1885) 564 

linariae (Stephens, 1830) 

cornelii Spornraft, 1966 

pulicarius (Linnaeus, 1758) 

gravidus (Illiger, 1798) 

vestitus (Kiesenwetter, 1850)

55. Family NITIDULIDAE Latreille, 1802 


Subfamily CARPOPHILINAE Erichson, 1842 

Tribe CARPOPHILINI Erichson, 1842 

UROPHORUS Murray, 1864 

CARPOPHILUS sensu auctt. partim non Stephens, 

1829 


CARPOPHILUS Stephens, 1829 

dimidiatus (Fabricius, 1792) 

flavipes Murray, 1864 

hemipterus (Linnaeus, 1758) 

ligneus Murray, 1864 

maculatus Murray, 1864 

marginellus Motschulsky, 1858 

mutilatus Erichson, 1843 

nepos Murray, 1864 

freemani Dobson, 1956 

obsoletus Erichson, 1843 

sexpustulatus (Fabricius, 1792) 

bimaculatus (Marsham, 1802) 

truncatus Murray, 1864 565 

pilosellus sensu auctt. non Motschulsky, 1858 

halli Dobson, 1954 

Tribe EPURAEINI Kirejtshuk, 1986 

EPURAEA Erichson, 1843 

DADOPORA Thomson, C.G., 1859 

DADOPERA Joy, 1932 (misspelling) 

Subgenus EPURAEA Erichson, 1843 

aestiva (Linnaeus, 1758) 

depressa sensu (Illiger, 1798) non (Linnaeus, 1758) 

angustula Sturm, 1844 

biguttata (Thunberg, 1784) 

unicolor (Olivier, 1790) 566 

obsoleta sensu Fowler, 1889 partim non (Fabricius, 

1792) 

binotata Reitter, 1872 

nana Reitter, 1873 

distincta (Grimmer, 1841) 

fuscicollis (Stephens, 1835) 

diffusa Brisout de Barneville, 1863 

guttata (Olivier, 1811) 

decemguttata sensu (Fabricius, 1792) non (Olivier, 

1790) 

longula Erichson, 1845 

marseuli Reitter, 1872 

pusilla (Illiger, 1798) non (Thunberg, 1794) 567 

melina Erichson, 1843 

neglecta (Heer, 1841) 

pallescens (Stephens, 1835) 

aestiva sensu (Illiger, 1798) non (Linnaeus, 1758) 

florea Erichson, 1845 567 

rufomarginata (Stephens, 1830) 

parvula Sturm, 1844 

silacea (Herbst, 1783) 

deleta Sturm, 1844 567 

terminalis (Mannerheim, 1843) 

immunda Sturm, 1844 

71 

Family NITIDULIDAE Latreille, 1802 

adumbrata Mannerheim, 1852 567 

thoracica Tournier, 1872 

oblonga sensu auctt. Brit. non (Herbst, 1793) 

variegata (Herbst, 1793) 

Subgenus EPURAEANELLA Crotch, 1874 

OMOSIPHORA Reitter, 1875 

limbata (Fabricius, 1787) 

Subgenus MICRURIA Reitter, 1875 

MICRURULA Reitter, 1884 

melanocephala (Marsham, 1802) 

affinis (Stephens, 1830) 

truncata (Stephens, 1830) non (Fabricius, 1792) 

Subfamily MELIGETHINAE Thomson, C.G., 1859 

PRIA Stephens, 1829 

LARIA sensu auctt. partim non Scopoli, 1763 

dulcamarae (Scopoli, 1763) 

truncatella (Marsham, 1802) 

MELIGETHES Stephens, 1829 568 


caeruleus (Marsham, 1802) 

latipes (Marsham, 1802) 

nigrinus (Marsham, 1802) 

nigricornis Stephens, 1830 

urticae Stephens, 1830 

atramentarius Förster, 1849 

symphyti sensu auctt. Brit. partim non (Heer, 1841) 

atratus (Olivier, 1790) 

rufipes sensu (Marsham, 1802) non (Linnaeus, 

1767) 

bidens Brisout de Barneville, 1863 

bidentatus sensu Joy, 1932 non Brisout de 

Barneville, 1863 

bidentatus Brisout de Barneville, 1863 

gresseri Bach, 1875 

corsicus Deville, 1908 

brevis Sturm, 1845 

pictus Rye, 1871 

brunnicornis Sturm, 1845 

carinulatus Förster, 1849 

erythropus sensu auctt. non (Marsham, 1802) 569 

coracinus Sturm, 1845 

corvinus Erichson, 1845 

difficilis (Heer, 1841) 

erichsonii Brisout de Barneville, 1863 

erichsoni auctt. (misspelling) 

exilis Sturm, 1845 

flavimanus Stephens, 1830 

lumbaris Sturm, 1845 

fulvipes Brisout de Barneville, 1863 

rubripes Mulsant & Rey, 1863 

gagathinus Erichson, 1845 570 

gagatinus auctt. (misspelling) 

lugubris sensu auctt. partim non Sturm, 1845 

haemorrhoidalis Förster, 1849 571 

incanus Sturm, 1845 

kunzei Erichson, 1845 

lugubris Sturm, 1845 

matronalis Audisio & Spornraft, 1990 572 

morosus Erichson, 1845 

memnonius Erichson, 1845 

nanus Erichson, 1845 

nigrescens Stephens, 1830

Family MONOTOMIDAE Laporte, 1840 Checklist of Beetles of the British Isles, 2012 Edition 

xanthoceros Stephens, 1830 

picipes Sturm, 1845 

obscurus Erichson, 1845 

palmatus Erichson, 1845 

ochropus Sturm, 1845 

ovatus Sturm, 1845 

pedicularius (Gyllenhal, 1808) 573 

viduatus (Heer, 1841) 574 

aestimabilis Reitter, 1872 

persicus (Faldermann, 1835) 

pedicularius sensu auctt. ante 1993 non (Gyllenhal, 

1808) 574 

tenebrosus Förster, 1849 

planiusculus (Heer, 1841) 

murinus Erichson, 1845 

rotundicollis Brisout de Barneville, 1863 

ruficornis (Marsham, 1802) 

subrugosus sensu Stephens, 1830 partim non 

(Gyllenhal, 1808) 

flavipes Sturm, 1845 569 

serripes (Gyllenhal, 1827) 

solidus (Kugelann, 1794) 

subrugosus (Gyllenhal, 1808) 575 

symphyti (Heer, 1841) 576 

umbrosus Sturm, 1845 

viridescens (Fabricius, 1787) 

Subfamily NITIDULINAE Latreille, 1802 

NITIDULA Fabricius, 1775 

bipunctata (Linnaeus, 1758) 

bipustulata (Linnaeus, 1761) 

carnaria (Schaller, 1783) 

quadripustulata Fabricius, 1792 

variata Stephens, 1830 

flavomaculata Rossi, 1790 

flexuosa Olivier, 1790 


OMOSITA Erichson, 1843 

colon (Linnaeus, 1758) 

depressa (Linnaeus, 1758) 

discoidea (Fabricius, 1775) 

SORONIA Erichson, 1843 

grisea (Linnaeus, 1758) 

punctatissima (Illiger, 1794) 

AMPHOTIS Erichson, 1843 

marginata (Fabricius, 1781) 

CYCHRAMUS Kugelann, 1794 

luteus (Fabricius, 1787) 

fungicola Heer, 1841 

POCADIUS Erichson, 1843 

adustus Reitter, 1888 

lanuginosus Franz, 1969 567 

ferrugineus (Fabricius, 1775) 

THALYCRA Erichson, 1843 

fervida (Olivier, 1790) 

sericea (Sturm, 1839) 

72 

Subfamily CRYPTARCHINAE Thomson, C.G., 

1859 

CRYPTARCHA Shuckard, 1839 

strigata (Fabricius, 1787) 

undata sensu (Marsham, 1802) non (Olivier, 1790) 

undata (Olivier, 1790) 

imperialis (Fabricius, 1792) 

nebulosa (Marsham, 1802) 

GLISCHROCHILUS Reitter, 1873 

IPS Fabricius, 1777 non De Geer, 1775 

Subgenus GLISCHROCHILUS Reitter, 1873 

quadripunctatus (Linnaeus, 1758) 

quadripustulatus (Linnaeus, 1761) 

Subgenus LIBRODOR Reitter, 1884 

hortensis (Geoffroy in Fourcroy, 1785) 

quadripunctatus sensu (Olivier, 1790) non 


olivieri Bedel, 1891 

quadriguttatus (Fabricius, 1777) 

PITYOPHAGUS Shuckard, 1839 

ferrugineus (Linnaeus, 1761) 

Subfamily CYBOCEPHALINAE Jacquelin du Val, 

1858 

CYBOCEPHALUS Erichson, 1844 

fodori Endrödy-Younga, 1965 577 

56. Family MONOTOMIDAE Laporte, 1840 578 


Subfamily RHIZOPHAGINAE Redtenbacher, 1845 

RHIZOPHAGUS Herbst, 1793 

RYZOPHAGUS Herbst, 1793 579 

RHYZOPHAGUS Gyllenhal, 1813 (misspelling) 

Subgenus RHIZOPHAGUS Herbst, 1793 


longicollis (Gyllenhal, 1827) 

gyllenhalii Thomson, C.G., 1885 

dispar (Paykull, 1800) 

punctulatus Guillebeau, 1897 

fenestralis (Linnaeus, 1758) 

parvulus (Paykull, 1800) 

ferrugineus (Paykull, 1800) 

minor Méquignon, 1909 


oblongicollis Blatch & Horner, 1892 

punctulatus sensu auctt. Brit. non Guillebeau, 1897 

simplex sensu auctt. non Reitter, 1884 

parallelocollis Gyllenhal, 1827 

terebrans sensu Stephens, 1829 non (Olivier, 1790) 

perforatus Erichson, 1845 

picipes (Olivier, 1790) 

politus (Hellwig, 1792) 

Subgenus ANOMOPHAGUS Reitter, 1907 

cribratus Gyllenhal, 1827

Subgenus EURHIZOPHAGUS Méquignon, 1909 

depressus (Fabricius, 1792) 

grandis Gyllenhal, 1827 580 

CYANOSTOLUS Ganglbauer, 1899 

aeneus (Richter, 1820) 

caeruleipennis (Sahlberg, C.R., 1837) 

caeruleus (Waltl, 1839) 

cyaneipennis (Hardy, 1847) 

Subfamily MONOTOMINAE Laporte, 1840 

MONOTOMA Herbst, 1793 

Subgenus GYROCECIS Thomson, C.G., 1859 

angusticollis (Gyllenhal, 1827) 

formicetorum Thomson, C.G., 1863 

conicicollis Guérin-Méneville, 1837 

angusticollis sensu auctt. non (Gyllenhal, 1827) 

quadrifoveolata Aubé, 1838 

subquadrifoveolata sensu Fowler, 1889 non 

Waterhouse, G.R., 1858 

Subgenus MONOTOMA Herbst, 1793 

bicolor Villa & Villa, 1835 

brevicollis Aubé, 1838 

quadridentata Thomson, C.G., 1870 

longicollis (Gyllenhal, 1827) 

gracilis Curtis, 1830 

angustata Stephens, 1830 

flavipes Kunze, 1839 

picipes Herbst, 1793 

contracta Marsham, 1802 

pallida Stephens, 1830 

quadricollis Aubé, 1838 581 

bicolor sensu auctt. partim non Villa & Villa, 1835 

spinicollis Aubé, 1838 

spinigera sensu auctt. non Chaudoir, 1845 

testacea Motschulsky, 1845 

quadriimpressa Motschulsky, 1845 

rufa Redtenbacher, 1849 

subquadrifoveolata Waterhouse, G.R., 1858 

57. Family SILVANIDAE Kirby, 1837 


Subfamily BRONTINAE Blanchard, 1845 

ULEIOTA Latreille, 1796 

BRONTES Fabricius, 1801 

planatus (Linnaeus, 1761) 

pallens (Fabricius, 1792) 

DENDROPHAGUS Schönherr, 1809 

crenatus (Paykull, 1799) 

CRYPTAMORPHA Wollaston, 1854 

desjardinsii (Guérin-Méneville, 1844) 

desjardinsi auctt. (misspelling) 

PSAMMOECUS Latreille, 1829 

PSAMMOECHUS Boudier, 1834 (misspelling) 

bipunctatus (Fabricius, 1792) 

73 

Subfamily SILVANINAE Kirby, 1837 

CATHARTUS Reiche, 1854 

quadricollis (Guérin-Méneville, 1844) 

NAUSIBIUS Lentz, 1857 

clavicornis (Kugelann, 1794) 

dentatus (Marsham, 1802) 

Family SILVANIDAE Kirby, 1837 

AHASVERUS des Gozis, 1881 

CATHARTUS sensu Fowler, 1889 partim non 

Reiche, 1854 

advena (Waltl, 1834) 

brunneus (Fabricius, 1792) 582 

ORYZAEPHILUS Ganglbauer, 1899 

SILVANUS sensu Fowler, 1889 partim non Latreille, 

1804 

mercator (Fauvel, 1889) 

surinamensis (Linnaeus, 1758) 

SILVANUS Latreille, 1804 

LEPTUS Duftschmid, 1825 non Latreille, 1796 

bidentatus (Fabricius, 1792) 

unidentatus (Olivier, 1790) 

SILVANOPRUS Reitter, 1911 

SILVANUS sensu Fowler, 1889 partim non Latreille, 

1804 

fagi (Guérin-Méneville, 1844) 

similis (Erichson, 1846) 

58. Family CUCUJIDAE Latreille, 1802 


PEDIACUS Shuckard, 1839 

depressus (Herbst, 1797) 

dermestoides (Fabricius, 1792) 

59. Family LAEMOPHLOEIDAE Ganglbauer, 

1899 


LAEMOPHLOEUS Dejean, 1835 

monilis (Fabricius, 1787) 

CRYPTOLESTES Ganglbauer, 1899 

LEPTUS Thomson, C.G., 1859 non Latreille, 1796 

capensis (Waltl, 1834) 

duplicatus (Waltl, 1839) 

ferrugineus (Stephens, 1831) 

pusilloides (Steel & Howe, 1952) 

pusillus (Schönherr, 1817) 

minutus (Olivier, 1791) non (Geoffroy in Fourcroy, 

1785) 

spartii (Curtis, 1834) 

ater (Olivier, 1795) non (Geoffroy in Fourcroy, 1785) 

turcicus (Grouvelle, 1876)

Family PHALACRIDAE Leach, 1815 Checklist of Beetles of the British Isles, 2012 Edition 

NOTOLAEMUS Lefkovitch, 1959 

LAEMOPHLOEUS sensu Fowler, 1889 partim non 

Dejean, 1835 

unifasciatus (Latreille, 1804) 

bimaculatus (Paykull, 1801) non (Olivier, 1791) 

fasciatus (Stephens, 1839) 

LEPTOPHLOEUS Casey, 1916 

LAEMOPHLOEUS sensu Fowler, 1889 partim non 

Dejean, 1835 

clematidis (Erichson, 1846) 

janeti (Grouvelle, 1899) 

60. Family PHALACRIDAE Leach, 1815 


PHALACRUS Paykull, 1800 

caricis Sturm, 1807 

nigrinus sensu auctt. Brit. non (Marsham, 1802) 

millefolii sensu Gyllenhal, 1813 non Paykull, 1800 

championi Guillebeau, 1892 

brunnipes sensu Rye, 1872 non Brisout de 

Barneville, 1863 

corruscus (Panzer, 1796) 

coruscus auctt. (misspelling) 

fimetarius sensu (Paykull, 1798) non (Fabricius, 

1775) 

fimetarius (Fabricius, 1775) 

brisouti Rye, 1872 

hybridus Flach, 1888 

substriatus Gyllenhal, 1813 

OLIBRUS Erichson, 1845 



particeps sensu auctt. Brit. non Mulsant, 1861 

corticalis (Panzer, 1796) 

flavicornis (Sturm, 1807) 

helveticus Rye, 1876 

liquidus Erichson, 1845 

bicolor sensu auctt. Brit. non (Fabricius, 1792) 

millefolii (Paykull, 1800) 

pygmaeus (Sturm, 1807) 

STILBUS Seidlitz, 1872 

atomarius (Linnaeus, 1767) 

oblongus (Erichson, 1845) 

testaceus (Panzer, 1796) 

61. Family CRYPTOPHAGIDAE Kirby, 1826 583 


Subfamily CRYPTOPHAGINAE Kirby, 1826 

Tribe CRYPTOPHAGINI Kirby, 1826 

TELMATOPHILUS Heer, 1841 

brevicollis Aubé, 1862 

caricis (Olivier, 1790) 

obscurus (Fabricius, 1792) 

schonherrii (Gyllenhal, 1808) 

74 

schoenherri auctt. (misspelling) 

sparganii (Ahrens, 1812) 

typhae (Fallén, 1802) 

pumilus Reitter, 1875 

PARAMECOSOMA Curtis, 1833 

melanocephalum (Herbst, 1793) 

bicolor Curtis, 1833 

infuscatum Halbert, 1910 

HENOTICUS Thomson, C.G., 1868 

californicus (Mannerheim, 1843) 

germanicus Reitter, 1906 

serratus (Gyllenhal, 1808) 

CRYPTOPHAGUS Herbst, 1792 

KRYPTOPHAGUS Herbst, 1792 584 

MNIONOMUS Wollaston, 1864 

acutangulus Gyllenhal, 1827 

waterhousei Rye, 1866 

badius Sturm, 1845 

cellaris (Scopoli, 1763) 

crenatus (Herbst, 1792) 

confusus Bruce, 1934 585 

corticinus Thomson, C.G., 1863 

dentatus (Herbst, 1793) 

fumatus (Marsham, 1802) 

niger sensu auctt. non Brisout de Barneville, 1863 

denticulatus Heer, 1841 

pilosus Gyllenhal, 1827 non Herbst, 1792 

pallidus sensu auctt. Brit. non Sturm, 1845 

pseudodentatus Bruce, 1934 

dilutus Reitter, 1874 

hexagonalis sensu auctt. ante 2006 non Tournier, 

1872 

subvittatus sensu auctt. Brit. non Reitter, 1888 586 

distinguendus Sturm, 1845 

umbratus Erichson, 1846 

stramenti Donisthorpe, 1935 

falcozi Roubal, 1927 

westi Bruce, 1943 

fallax Balfour-Browne, 1953 

fumatus sensu auctt. non (Marsham, 1802) 

spadiceus sensu auctt. non Falcoz, 1925 

fuscicornis Sturm, 1845 

insulicola Roubal, 1919 

vagus Bruce, 1938 

acuminatus Coombs & Woodroffe, 1955 

intermedius Bruce, 1934 

labilis Erichson, 1846 


validus Kraatz, 1856 

laticollis Lucas, 1846 

affinis Sturm, 1845 non Sahlberg, R.F., 1834 

lycoperdi (Scopoli, 1763) 

rufus (Marsham, 1802) 

bituberculatus Stephens, 1830 

marshami Stephens, 1830 

micaceus Rey, 1889 

lovendali Ganglbauer, 1899 

loevendali auctt. (misspelling) 

obsoletus Reitter, 1879 

parallelus Brisout de Barneville, 1863 

angustus sensu auctt. ante 2006 non Ganglbauer, 

1899 

cylindrus sensu auctt. ? Kiesenwetter, 1858 

populi Paykull, 1800

grandis Kraatz, 1856 

pubescens Sturm, 1845 

puncticollis Lucas, 1846 

rotundatus Coombs & Woodroffe, 1955 

punctipennis Brisout de Barneville, 1863 

pilosus sensu auctt. ad 2007 non Gyllenhal, 1827 

badius sensu auctt. Brit. partim non Sturm, 1845 

reflexus Rey, 1889 

pallidus sensu auctt. ad 2007 non Sturm, 1845 

fowleri Joy, 1910 

ruficornis Stephens, 1830 

nigritulus Reitter, 1888 

saginatus Sturm, 1845 

scanicus (Linnaeus, 1758) 

patruelis Sturm, 1845 

niger Brisout de Barneville, 1863 

schmidtii Sturm, 1845 

schmidti auctt. (misspelling) 

scutellatus Newman, 1834 

crenatus sensu auctt. Brit. non Herbst, 1792 

bicolor Sturm, 1845 

setulosus Sturm, 1845 

simplex Miller, 1858 

subdepressus Gyllenhal, 1827 

subfumatus Kraatz, 1856 

uncinatus Stephens, 1830 

badius sensu auctt. Brit. partim non Sturm, 1845 

hirtulus Kraatz, 1858 

immixtus Rey, 1889 

postpositus Sahlberg, J., 1903 

MICRAMBE Thomson, C.G., 1863 

CRYPTOPHAGUS sensu auctt. partim non Herbst, 

1792 

MICRAMBINUS Reitter, 1906 586 

abietis (Paykull, 1798) 

bimaculata (Panzer, 1798) 

pilosula (Erichson, 1846) 

?villosa (Heer, 1841) non (Beck, 1817) nec auctt. 

lindbergorum (Bruce, 1934) 

ulicis (Stephens, 1830) 

vini (Panzer, 1797) non (Laicharting, 1781) 

?obcordata (Marsham, 1802) 

woodroffei Johnson, 2007 

?ocularis (Reitter, 1872) 

villosa sensu auctt. ante 2007 non (Heer, 1841) nec 

(Beck, 1817) 

ANTHEROPHAGUS Dejean, 1821 

pallens (Linnaeus, 1758) 


silaceus (Herbst, 1792) 

canescens Grouvelle, 1916 

similis Curtis, 1835 

pallens sensu auctt. ad 2007 non (Linnaeus, 1758) 

Tribe CAENOSCELINI Casey, 1900 

CAENOSCELIS Thomson, C.G., 1863 

COENOSCELIS auctt. Brit. (misspelling) 

ferruginea (Sahlberg, C.R., 1820) 

pallida (Wollaston, 1847) 

sibirica Reitter, 1889 587 

subdeplanata Brisout de Barneville, 1882 

75 

Family CRYPTOPHAGIDAE Kirby, 1826 

Subfamily ATOMARIINAE LeConte, 1861 

Tribe HYPOCOPRINI Reitter, 1879 

HYPOCOPRUS Motschulsky, 1839 

latridioides Motschulsky, 1839 

lathridioides auctt. (misspelling) 

quadricollis Reitter, 1878 

Tribe ATOMARIINI LeConte, 1861 

ATOMARIA Stephens, 1829 

Subgenus ATOMARIA Stephens, 1829 

badia Erichson, 1846 

sahlbergi Sjöberg, 1947 

bella Reitter, 1875 

diluta Erichson, 1846 

fimetarii (Fabricius, 1792) 

fimetarii (Herbst, 1793) 

fimetarius auctt. (misspelling) 

impressa Erichson, 1846 

linearis Stephens, 1830 

lohsei Johnson & Strand, A., 1968 

longicornis Thomson, C.G., 1863 

procerula sensu auctt. non Erichson, 1846 

nigrirostris Stephens 1830 

umbrina sensu Erichson, 1846 non (Gyllenhal, 

1827) 

fuscicollis Mannerheim, 1852 588 

plicicollis Mäklin, 1863 

nigriventris Stephens, 1830 

nana Erichson, 1846 

pulchra Erichson, 1846 

barani Brisout de Barneville, 1863 

puncticollis Thomson, C.G., 1868 

nigriventris sensu auctt. Brit. partim non Stephens, 

1830 

punctithorax Reitter, 1888 

nigriventris sensu auctt. Brit. partim non Stephens, 

1830 

consanguinea Johnson, 1976 

strandi Johnson, 1967 

alpina sensu auctt. Brit. non Heer, 1841 

elongatula sensu auctt. Brit. non Erichson, 1846 

umbrina (Gyllenhal, 1827) 

fumata sensu auctt. Brit. non Erichson, 1846 

vespertina Mäklin, 1853 

affinis sensu auctt. Brit. non (Sahlberg, R.F., 1834) 

badia sensu auctt. Brit. non Erichson, 1846 

prolixa sensu auctt. non Erichson, 1846 

pulchra sensu auctt. ad 2006 non Erichson, 1846 

wollastoni Sharp, 1867 

Subgenus ANCHICERA Thomson, C.G., 1863 

apicalis Erichson, 1846 

atra (Herbst, 1793) 

atricapilla Stephens, 1830 

nigriceps Erichson, 1846 

basalis Erichson, 1846 

nitidula sensu auctt. non Heer, 1841 nec (Marsham, 

1802) 

clavigera Ganglbauer, 1899 

fuscata (Schönherr, 1808) 

castanea Stephens, 1830 


fuscipes (Gyllenhal, 1808)

Family EROTYLIDAE Latreille, 1802 Checklist of Beetles of the British Isles, 2012 Edition 

carbonaria Stephens, 1830 

concolor Märkel, 1844 

gutta Newman, 1834 

hislopi Wollaston, 1857 

gibbula sensu auctt. Brit. non Erichson, 1846 

lewisi Reitter, 1877 

mesomela (Herbst, 1792) 

mesomelaena auctt. (misspelling) 

dimidiata (Marsham, 1802) 

phaeorrhaea (Marsham, 1802) 

morio Kolenati, 1846 

munda Erichson, 1846 

nigripennis (Kugelann, 1794) 

nitidula (Marsham, 1802) 

testacea sensu auctt. Brit. non Stephens, 1830 

analis sensu auctt. Brit. non Erichson, 1846 

borealis Sjöberg, 1947 

ornata Heer, 1841 

contaminata Erichson, 1846 

peltata Kraatz, 1853 

nitidula Heer, 1841 non (Marsham, 1802) 

pseudatra Reitter, 1888 

reitteri Løvendal, 1893 

pusilla (Paykull, 1798) 

phaeogaster (Marsham, 1802) 

evanescens sensu auctt. Brit. non (Marsham, 1802) 

basella Stephens, 1830 

fulvicollis Stephens, 1830 

thoracica Stephens, 1830 

castanea sensu auctt. Brit. non Stephens, 1830 

rhenonum Kraatz, 1853 

rhenana auctt. (misspelling) 

elevata Allen, 1938 

godarti sensu Sjöberg, 1947 non Guillebeau, 1885 

rubella Heer, 1841 

bicolor sensu auctt. Brit. non Erichson, 1846 

berolinensis Kraatz, 1853 588 

rubida Reitter, 1875 

ornata sensu auctt. Brit. non Heer, 1841 

cognata sensu auctt. Europ. non Erichson, 1846 

gibbula sensu Allen, 1968 non Erichson, 1846 

versicolor sensu auctt. Brit. non Erichson, 1846 

viennensis Reitter, 1875 

rubricollis Brisout de Barneville, 1863 

divisa Rye, 1876 

scutellaris Motschulsky, 1849 

testacea Stephens, 1830 

ruficornis (Marsham, 1802) non (Gmelin in 

Linnaeus, 1790) 588 

dorsalis Stephens, 1830 

terminata (Comolli, 1837) 

turgida Erichson, 1846 589 

zetterstedti (Zetterstedt, 1838) 

OOTYPUS Ganglbauer, 1899 

EPHISTEMUS sensu Fowler, 1889 partim non 

Stephens, 1829 

globosus (Waltl, 1838) 

nigriclavis sensu auctt. Brit. non (Stephens, 1829) 

palustris (Wollaston, 1846) 

EPHISTEMUS Stephens, 1829 

globulus (Paykull, 1798) 

gyrinoides (Marsham, 1802) 

nitens (Marsham, 1802) 

piceorhaeus (Marsham, 1802) 

pulchellus (Marsham, 1802) 

76 

dimidiatus (Sturm, 1807) 

confinis Stephens, 1829 

histeroides Stephens, 1835 


ovulum (Erichson, 1846) 

dubius Fowler, 1889 

reitteri Casey, 1900 590 

62. Family EROTYLIDAE Latreille, 1802 


Subfamily CRYPTOPHILINAE Casey, 1900 591 

CRYPTOPHILUS Reitter, 1874 

integer (Heer, 1841) 592 

Subfamily EROTYLINAE Latreille, 1802 

Tribe DACNINI Gistel, 1848 

DACNE Latreille, 1796 

bipustulata (Thunberg, 1781) 


rufifrons (Fabricius, 1775) 

fowleri Joy, 1905 

Tribe TRITOMINI Curtis, 1834 

TRIPLAX Herbst, 1793 

aenea (Schaller, 1783) 

bicolor (Marsham, 1802) 

lacordairii Crotch, 1870 

lacordairei auctt. (misspelling) 

russica (Linnaeus, 1758) 

scutellaris Charpentier, 1825 

bicolor sensu Gyllenhal, 1808 non (Marsham, 1802) 

gyllenhalii Crotch, 1870 

TRITOMA Fabricius, 1775 593 

CYRTOTRIPLAX Crotch, 1873 

bipustulata Fabricius, 1775 

humeralis (Marsham, 1802) 

63. Family BYTURIDAE Gistel, 1848 


BYTURUS Latreille, 1796 

ochraceus (Scriba, 1790) 

aestivus sensu auctt. non (Linnaeus, 1758) 

fumatus sensu auctt. partim non (Fabricius, 1775) 

tomentosus (De Geer, 1774) 

flavescens (Marsham, 1802) 

urbanus (Lindemann, 1865)

64. Family BIPHYLLIDAE LeConte, 1861 


BIPHYLLUS Dejean, 1821 

DIPHYLLUS Berthold, 1827 


DIPLOCOELUS Guérin-Méneville, 1844 

fagi (Chevrolat in Guérin-Méneville, 1837) 

65. Family BOTHRIDERIDAE Erichson, 1845 


Subfamily TEREDINAE Seidlitz, 1888 

TEREDUS Dejean, 1835 

cylindricus (Olivier, 1790) 

nitidus (Fabricius, 1792) 

OXYLAEMUS Erichson, 1845 

cylindricus (Creutzer in Panzer, 1796) 

variolosus (Dufour, 1843) 

Subfamily ANOMMATINAE Ganglbauer, 1899 

ANOMMATUS Wesmael, 1835 

diecki Reitter, 1875 594 

duodecimstriatus (Müller, P.W.J., 1821) 

obsoletus (Stephens, 1830) 

66. Family CERYLONIDAE Billberg, 1820 


Subfamily MURMIDIINAE Jacquelin du Val, 1858 

MURMIDIUS Leach, 1821 

ovalis (Beck, 1817) 

segregatus Waterhouse, C.O., 1876 

Subfamily CERYLONINAE Billberg, 1820 

CERYLON Latreille, 1802 

fagi Brisout de Barneville, 1867 

ferrugineum Stephens, 1830 

primroseae Donisthorpe, 1942 

histeroides (Fabricius, 1792) 

67. Family ALEXIIDAE Imhoff, 1856 


SPHAEROSOMA Stephens, 1832 

ALEXIA Stephens, 1833 

pilosum (Panzer, 1793) 

piliferum sensu auctt. Brit. non (Müller, P.W.J., 

1821) 

quercus Stephens, 1832 

77 

Family BIPHYLLIDAE LeConte, 1861 

68. Family ENDOMYCHIDAE Leach, 1815 595 


Subfamily ENDOMYCHINAE Leach, 1815 

ENDOMYCHUS Panzer, 1795 

coccineus (Linnaeus, 1758) 

Subfamily ANAMORPHINAE Strohecker, 1953 

SYMBIOTES Redtenbacher, 1849 

latus Redtenbacher, 1849 

Subfamily MEROPHYSIINAE Seidlitz, 1872 

HOLOPARAMECUS Curtis, 1833 

caularum (Aubé, 1843) 

depressus Curtis, 1833 

kunzei (Aubé, 1843) 

singularis (Beck, 1817) 

Subfamily LYCOPERDININAE Bromhead, 1838 

LYCOPERDINA Latreille, 1807 

bovistae (Fabricius, 1792) 

succincta (Linnaeus, 1767) 

Subfamily MYCETAEINAE Jacquelin du Val, 1857 

MYCETAEA Stephens, 1829 

subterranea (Fabricius, 1801) 

fumata sensu (Marsham, 1802) non (Linnaeus, 

1767) 

hirta (Marsham, 1802) non (Herbst, 1783) 

villosa (Beck, 1817) 

69. Family COCCINELLIDAE Latreille, 1807 


Subfamily COCCIDULINAE Mulsant, 1846 

COCCIDULA Kugelann in Illiger, 1798 

rufa (Herbst, 1783) 

scutellata (Herbst, 1783) 

RHYZOBIUS Stephens, 1829 

RHIZOBIUS sensu auctt. non Burmeister, 1835 

RHIZOBIELLUS Oke, 1951 

chrysomeloides (Herbst, 1792) 596 

litura (Fabricius, 1787) 

lophanthae (Blaisdell, 1892) 597 

RODOLIA Mulsant, 1850 

cardinalis (Mulsant, 1850) 598

Family COCCINELLIDAE Latreille, 1807 Checklist of Beetles of the British Isles, 2012 Edition 

STETHORUS Weise, 1885 

SCYMNUS sensu Fowler, 1889 partim non 


punctillum (Weise, 1891) 

ater sensu auctt. Brit. partim non (Kugelann, 1794) 

minimus (Rossi, 1794) non (Müller, O.F., 1776) 

CLITOSTETHUS Weise, 1885 



arcuatus (Rossi, 1794) 599 

SCYMNUS Kugelann, 1794 

Subgenus SCYMNUS Kugelann, 1794 

femoralis (Gyllenhal, 1827) 

rubromaculatus sensu auctt. Brit. partim non 

(Goeze, 1777) 

pygmaeus sensu auctt. Brit. partim non (Geoffroy in 


frontalis (Fabricius, 1787) 

interruptus (Goeze, 1777) 600 

jakowlewi Weise, 1892 601 

nigrinus Kugelann, 1794 

rubromaculatus (Goeze, 1777) 602 

schmidti Fürsch, 1958 

pygmaeus sensu auctt. Brit. partim non (Geoffroy in 


frontalis var. immaculatus sensu auctt. Brit. partim 

non Suffrian, 1843 

mimulus Capra & Fürsch, 1967 

Subgenus PULLUS Mulsant, 1846 

auritus Thunberg, 1795 

capitatus (Fabricius, 1798) 

suturalis Thunberg, 1795 

Subgenus NEOPULLUS Sasaji, 1971 

haemorrhoidalis Herbst, 1797 

limbatus Stephens, 1832 

testaceus sensu auctt. Europ. ante 1967 non 


NEPHUS Mulsant, 1846 



bisignatus (Boheman, 1850) 

bohemani (Stenius, 1952) 

quadrimaculatus (Herbst, 1783) 

pulchellus (Herbst, 1797) 

redtenbacheri (Mulsant, 1846) 

testaceus sensu auctt. Brit. non (Motschulsky, 1837) 

mulsanti (Waterhouse, G.R., 1862) 

lividus (Bold, 1872) 

limonii (Donisthorpe, 1903) 603 

CRYPTOLAEMUS Mulsant, 1853 

montrouzieri Mulsant, 1853 604 

HYPERASPIS Dejean, 1836 

pseudopustulata Mulsant, 1853 

reppensis sensu auctt. Brit. non (Herbst, 1783) 

78 

Subfamily CHILOCORINAE Mulsant, 1846 

PLATYNASPIS Redtenbacher, 1843 

luteorubra (Goeze, 1777) 

CHILOCORUS Leach, 1815 


renipustulatus (Scriba, 1791) 

similis (Rossi, 1790) non (Thunberg, 1781) 

EXOCHOMUS Redtenbacher, 1843 


Subfamily COCCINELLINAE Latreille, 1807 

COCCINULA Dobzhansky, 1925 

quattuordecimpustulata (Linnaeus, 1758) 605 

ANISOSTICTA Dejean, 1836 

novemdecimpunctata (Linnaeus, 1758) 

TYTTHASPIS Crotch, 1874 

MICRASPIS Redtenbacher, 1843 non Dejean, 1836 

sedecimpunctata (Linnaeus, 1761) 

sexdecimpunctata auctt. (misspelling) 

MYZIA Mulsant, 1846 

MYSIA Mulsant, 1846 non Lamarck, 1818 

NEOMYSIA Casey, 1899 

PARAMYSIA Reitter, 1911 

oblongoguttata (Linnaeus, 1758) 

MYRRHA Mulsant, 1846 

HALYZIA sensu Fowler, 1889 partim non Mulsant, 

1846 

octodecimguttata (Linnaeus, 1758) 

PROPYLEA Mulsant, 1846 

PROPYLAEA auctt. (misspelling) 


1846 

quattuordecimpunctata (Linnaeus, 1758) 

quatuordecimpunctata auctt. (misspelling) 

conglobata sensu auctt. Brit. non (Linnaeus, 1758) 

CALVIA Mulsant, 1846 


1846 

quattuordecimguttata (Linnaeus, 1758) 

quatuordecimguttata auctt. (misspelling) 

VIBIDIA Mulsant, 1846 


1846 

duodecimguttata (Poda, 1761) 

HALYZIA Mulsant, 1846 

sedecimguttata (Linnaeus, 1758) 

sexdecimguttata auctt. (misspelling)

PSYLLOBORA Dejean, 1836 

THEA Mulsant, 1846 


1846 

vigintiduopunctata (Linnaeus, 1758) 

ANATIS Mulsant, 1846 

ocellata (Linnaeus, 1758) 

APHIDECTA Weise, 1899 

APHIDEITA Weise, 1893 32 

ADALIA sensu Fowler, 1889 partim non Mulsant, 

1846 

obliterata (Linnaeus, 1758) 

HIPPODAMIA Dejean, 1836 

Subgenus HIPPODAMIA Dejean, 1836 

tredecimpunctata (Linnaeus, 1758) 

Subgenus ADONIA Mulsant, 1846 

variegata (Goeze, 1777) 

COCCINELLA Linnaeus, 1758 

hieroglyphica Linnaeus, 1758 

magnifica Redtenbacher, 1843 

divaricata sensu auctt. non Olivier, 1808 

distincta Faldermann, 1837 non Thunberg, 1781 

quinquepunctata Linnaeus, 1758 

septempunctata Linnaeus, 1758 

undecimpunctata Linnaeus, 1758 

ADALIA Mulsant, 1846 

bipunctata (Linnaeus, 1758) 

decempunctata (Linnaeus, 1758) 

bothnica sensu Fowler, 1889 non (Paykull, 1799) 

biabilis Marriner, 1926 

HARMONIA Mulsant, 1846 

axyridis (Pallas, 1773) 606 

quadripunctata (Pontoppidan, 1763) 

Subfamily EPILACHNINAE Mulsant, 1846 

HENOSEPILACHNA Li, 1961 

EPILACHNA sensu auctt. partim non Dejean, 1836 

argus (Geoffory in Fourcroy, 1762) 607 

SUBCOCCINELLA Guérin-Méneville, 1844 

vigintiquattuorpunctata (Linnaeus, 1758) 

vigintiquatuorpunctata auctt. (misspelling) 

70. Family CORYLOPHIDAE LeConte, 1852 608 


Subfamily CORYLOPHINAE LeConte, 1852 

SERICODERUS Stephens, 1829 

brevicornis Matthews, A., 1890 609 

lateralis (Gyllenhal, 1827) 

thoracicus Stephens, 1829 

79 

Family CORYLOPHIDAE LeConte, 1852 

CORYLOPHUS Stephens, 1833 

cassidoides (Marsham, 1802) 

cassidioides auctt. (misspelling) 

sublaevipennis Jacquelin du Val, 1859 

Subfamily ORTHOPERINAE Jacquelin du Val, 

1857 

ORTHOPERUS Stephens, 1829 

aequalis Sharp, 1885 

nitidulus Allen, 1942 

atomarius (Heer, 1841) 

atomus (Gyllenhal, 1808) 

?punctum (Marsham, 1802) 

brunnipes sensu auctt. partim non (Gyllenhal, 1808) 

picea Stephens, 1829 

corticalis sensu Matthews, A., 1885 non 

(Redtenbacher, 1849) 

brunnipes (Gyllenhal, 1808) 

nigriclavis (Stephens, 1829) 

kluki Wankowicz, 1865 

corticalis (Redtenbacher, 1849) 610 

brunnipes sensu auctt. partim non (Gyllenhal, 1808) 

mundus sensu auctt. non Matthews, A., 1885 

punctatulus Matthews, A., 1885 

improvisus Bruce, 1948 

nigrescens Stephens, 1829 

?picatus (Marsham, 1802) 

atomos Stephens, 1829 

truncatus Stephens, 1829 

pilosiusculus sensu auctt. Brit. non Jacquelin du 

Val, 1859 

coriaceus Mulsant & Rey, 1861 

mundus Matthews, A., 1885 611 

Subfamily RYPOBIINAE Paulian, 1950 

RYPOBIUS LeConte, 1852 

RHYPOBIUS auctt. (misspelling) 

praetermissus Bowestead, 1999 

ruficollis sensu auctt. non (Jacquelin du Val, 1854) 

71. Family LATRIDIIDAE Erichson, 1842 612 


Subfamily LATRIDIINAE Erichson, 1842 

LITHOSTYGNUS Broun, 1886 

METOPHTHALMUS sensu auctt. non Motschulsky, 

1850 

serripennis Broun, 1914 

STEPHOSTETHUS LeConte, 1878 

LATHRIDIUS sensu auctt. Brit. partim non Illiger, 

1801 

alternans (Mannerheim, 1844) 613 

angusticollis (Gyllenhal, 1827) 

angulatus (Mannerheim, 1844) 

undulatus (Motschulsky, 1866) 

kokujevi (Semenov, 1898) 

kokujewi auctt. (misspelling) 

caucasicus (Mannerheim, 1844) 614 

sinuatocollis sensu auctt. non (Faldermann, 1837)

Family LATRIDIIDAE Erichson, 1842 Checklist of Beetles of the British Isles, 2012 Edition 

campicola sensu auctt. non (Gerhardt, 1911) 

lardarius (De Geer, 1775) 

dilaticollis (Motschulsky, 1866) 

pini (Motschulsky, 1866) 

CARTODERE Thomson, C.G., 1859 

LATHRIDIUS sensu auctt. partim non Illiger, 1801 

CONINOMUS Thomson, C.G., 1863 

Subgenus CARTODERE Thomson, C.G., 1859 

constricta (Gyllenhal, 1827) 

carinata (Gyllenhal, 1827) 

Subgenus ARIDIUS Motschulsky, 1866 

LATHRIDIUS sensu auctt. partim non Illiger, 1801 

CONINOMUS sensu auctt. Brit. partim non 


bifasciata (Reitter, 1877) 

nodifer (Westwood, 1839) 

satelles (Blackburn, 1888) 

australica sensu auctt. non (Belon, 1887) 

norvegica (Strand, 1940) 

LATRIDIUS Herbst, 1793 

LATHRIDIUS Illiger, 1801 40 

CONITHASSA Thomson, C.G., 1859 

ENICMUS sensu auctt. Brit. partim non Thomson, 

C.G., 1859 

assimilis Mannerheim, 1844 

minutus sensu auctt. partim non (Linnaeus, 1767) 

pseudominutus (Strand, A., 1958) 

consimilis Mannerheim, 1844 


porcatus Herbst, 1793 

minutus sensu auctt. partim non (Linnaeus, 1767) 

?pulla Marsham, 1802 

anthracinus Mannerheim, 1844 

minutissimus Motschulsky, 1858 

THES Semenov, 1910 

LAR Semenov, 1904 non Gosse, 1857 

LATHRIDIUS sensu auctt. Brit. partim non Illiger, 

1801 

bergrothi (Reitter, 1880) 

ENICMUS Thomson, C.G., 1859 

brevicornis (Mannerheim, 1844) 

carbonarius (Mannerheim, 1844) 

fungicola Thomson, C.G., 1868 

histrio Joy & Tomlin, 1910 

transversus sensu auctt. Brit. partim non (Olivier, 

1790) 

rugosus (Herbst, 1793) 

testaceus (Stephens, 1830) 

cordaticollis (Aubé, 1850) 

transversus (Olivier, 1790) 

DIENERELLA Reitter, 1911 

CARTODERE sensu Thomson, C.G., 1863 non 


MICROGRAMME Walkley, 1948 

Subgenus CARTODEREMA Reitter, 1911 

clathrata (Mannerheim, 1844) 615 

elongata sensu auctt. Brit. partes maj. non (Curtis, 

1830) 

separanda sensu auctt. non (Reitter, 1887) 

80 

ruficollis (Marsham, 1802) 

collaris (Mannerheim, 1844) 

exilis (Mannerheim, 1844) 

nanula (Mannerheim, 1844) 

vincenti Johnson, 2007 

elongata (Curtis, 1830) non (Gyllenhal, 1827) 

Subgenus DIENERELLA Reitter, 1911 

argus (Reitter, 1884) 

elegans (Aubé, 1850) 

depilis (Belon, 1895) 

pilifera sensu auctt. Brit. partim non (Reitter, 1875) 

filiformis (Gyllenhal, 1827) 

filum (Aubé, 1850) 

schueppeli (Reitter, 1880) 

ADISTEMIA Fall, 1899 

watsoni (Wollaston, 1871) 

Subfamily CORTICARIINAE Curtis, 1829 

CORTICARIA Marsham, 1802 

alleni Johnson, 1974 

longicollis sensu auctt. Brit. partim non (Zetterstedt, 

1838) 

obscura sensu auctt. Brit. non Brisout de Barneville, 

1863 

depressa sensu auctt. Brit. non Thomson, C.G., 

1871 

corsica sensu auctt. Brit. non Brisout de Barneville, 

1878 

crenulata (Gyllenhal, 1827) 

dubia Dajoz, 1970 

eppelsheimi sensu auctt. Brit. non Reitter, 1875 

epelsheimi auctt. Brit. (misspelling) 

elongata (Gyllenhal, 1827) 

fagi Wollaston, 1854 

pietschi Ganglbauer, 1899 

aequidentata Allen, 1937 

ferruginea Marsham, 1802 

rufula Zetterstedt, 1838 

fenestralis sensu Reitter, 1875 non (Linnaeus, 

1758) 

fulva (Comolli, 1837) 

hirtella Thomson, C.G., 1863 

flavescens Thomson, C.G., 1871 

impressa (Olivier, 1790) 

denticulata (Gyllenhal, 1827) 

inconspicua Wollaston, 1860 

crenicollis sensu auctt. Brit. non Mannerheim, 1844 

saginata sensu auctt. Brit. non Mannerheim, 

1844 616 

longicollis (Zetterstedt, 1838) 

longicornis (Herbst, 1783) 617 

abietorum Motschulsky, 1867 

abietum auctt. (misspelling) 

depressa Thomson, C.G., 1871 

polypori Sahlberg, J., 1900 618 

alleni sensu auctt. Brit. partim non Johnson, 1974 

punctulata Marsham, 1802 

pubescens sensu auctt. Brit. non (Gyllenhal, 1827) 

rubripes Mannerheim, 1844 

linearis (Paykull, 1798) non (Thunberg, 1784) 

serrata (Paykull, 1798) 

laticollis Mannerheim, 1844 

umbilicata (Beck, 1817) 

cylindrica Mannerheim, 1844

orealis Wollaston, 1855 

CORTICARINA Reitter, 1880 

MELANOPHTHALMA sensu Fowler, 1889 partim 

non Motschulsky, 1866 

curta (Wollaston, 1854) 

fulvipes sensu auctt. non (Comolli, 1837) 

fuscipennis sensu auctt. Brit. non (Mannerheim, 

1844) 

meridionalis (Reitter, 1875) 

lambiana (Sharp, 1910) 

latipennis (Sahlberg, J., 1871) 

fowleriana (Sharp, 1910) 

minuta (Fabricius, 1792) 

fuscula (Gyllenhal, 1827) 

similata (Gyllenhal, 1827) 

?fulvipes (Comolli, 1837) non auctt. 

truncatella (Mannerheim, 1844) 

CORTINICARA Johnson, 1975 

CORTICARINA sensu auctt. partim non Reitter, 

1880 

MELANOPHTHALMA sensu Fowler, 1889 partim 

non Motschulsky, 1866 

gibbosa (Herbst, 1793) 

pallida (Marsham, 1802) 

impressa sensu (Marsham, 1802) non (Olivier, 

1790) 

MELANOPHTHALMA Motschulsky, 1866 

distinguenda (Comolli, 1837) 

angulata sensu auctt. Brit. non (Wollaston, 1864) 

suturalis (Mannerheim, 1844) 619 

transversalis sensu auctt. partim non (Gyllenhal, 

1827) 

transversalis (Gyllenhal, 1827) 

curticollis (Mannerheim, 1844) 

wollastoni (Waterhouse, G.R., 1859) 

MIGNEAUXIA Jacquelin du Val, 1859 

lederi Reitter, 1875 620 

?inflata (Rosenhauer, 1856) non auctt. 

orientalis Reitter, 1877 

Superfamily TENEBRIONOIDEA Latreille, 1802 

72. Family MYCETOPHAGIDAE Leach, 1815 


PSEUDOTRIPHYLLUS Reitter, 1880 

TRIPHYLLUS sensu Fowler, 1889 partim non 

Dejean, 1821 

suturalis (Fabricius, 1801) 

humeralis (Stephens, 1830) 

TRIPHYLLUS Dejean, 1821 

bicolor (Fabricius, 1777) 

punctatus (Hellwig, 1792) 

pilosus (Herbst, 1792) 

humeralis (Marsham, 1802) 

immaculatus Roubal, 1936 

81 

Family MYCETOPHAGIDAE Leach, 1815 

LITARGUS Erichson, 1846 

balteatus LeConte, 1856 

coloratus sensu auctt. Brit. non Rosenhauer, 1856 

connexus (Geoffroy in Fourcroy, 1785) 

bifasciatus (Fabricius, 1787) 

MYCETOPHAGUS Hellwig, 1792 

TRITOMA Geoffory, 1762 621 

atomarius (Fabricius, 1787) 

fulvicollis Fabricius, 1792 

multipunctatus Fabricius, 1792 

piceus (Fabricius, 1777) 

populi Fabricius, 1798 

quadriguttatus Müller, P.W.J., 1821 


TYPHAEA Stephens, 1829 

haagi Reitter, 1874 622 

decipiens Lohse, 1989 

stercorea (Linnaeus, 1758) 

fumata (Linnaeus, 1767) 

EULAGIUS Motschulsky, 1845 

ATRITOMUS Reitter, 1877 

filicornis (Reitter, 1887) 623 

BERGINUS Erichson, 1846 

tamarisci Wollaston, 1854 624 

73. Family CIIDAE Leach, 1819 

Family authors: G.M. Orledge & R.G. Booth 

Subfamily CIINAE Leach, 1819 

Tribe OROPHIINI Thomson, C.G., 1863 

OCTOTEMNUS Mellié, 1847 

glabriculus (Gyllenhal, 1827) 

ROPALODONTUS Mellié, 1847 

RHOPALODONTUS Dohrn in Strübing, 1851 40 

perforatus (Gyllenhal, 1813) 

Tribe CIINI Leach, 1819 

SULCACIS Dury, 1917 

ENTYPUS Redtenbacher, 1849 non Dahlbom, 1843 

ENNEARTHRON sensu Fowler, 1889 non Mellié, 

1847 

nitidus (Fabricius, 1792) 

affinis (Gyllenhal, 1827) 

STRIGOCIS Dury, 1917 

RHOPALODONTUS sensu Fowler, 1889 non Dohrn 

in Strübing, 1851 

SULCACIS sensu auctt. partim. non Dury, 1917 

bicornis (Mellié, 1849) 

fronticornis sensu auctt. Brit. non (Panzer, 1809)

Family TETRATOMIDAE Billberg, 1820 Checklist of Beetles of the British Isles, 2012 Edition 

ORTHOCIS Casey, 1898 

CIS sensu auctt. partim non Latreille, 1796 

MELLIEICIS Lohse, 1964 

alni (Gyllenhal, 1813) 

coluber (Abeille de Perrin, 1874) 

latifrons (Pool, 1917) 

CIS Latreille, 1796 

bidentatus (Olivier, 1790) 

inermis (Marsham, 1802) 

bilamellatus Wood, 1884 

boleti (Scopoli, 1763) 

castaneus (Herbst, 1793) 

nitidus sensu auctt. non (Fabricius, 1792) 

dentatus Mellié, 1849 

fagi Waltl, 1839 

fuscatus Mellié, 1849 

castaneus sensu Joy, 1932 non (Herbst, 1793) nec 

Mellié, 1849 

quadridentulus sensu Donisthorpe, 1933 non Perris, 

1874 

festivus (Panzer, 1793) 

jacquemartii Mellié, 1849 

jacquemarti auctt. (misspelling) 

lineatocribratus Mellié, 1849 

micans (Fabricius, 1792) 

hispidus (Paykull, 1798) 

savilli Donisthorpe, 1936 

punctulatus Gyllenhal, 1827 

pygmaeus (Marsham, 1802) 

nigricornis (Marsham, 1802) 


oblongus Mellié, 1849 

submicans Abeille de Perrin, 1874 

micans sensu auctt. non (Fabricius, 1792) 

vestitus Mellié, 1849 

pygmaeus sensu Nyholm, 1953 non (Marsham, 

1802) 

villosulus (Marsham, 1802) 

pyrrhocephalus (Marsham, 1802) 

flavus Stephens, 1830 

setiger Mellié, 1849 

ENNEARTHRON Mellié, 1847 

cornutum (Gyllenhal, 1827) 

concinnum (Marsham, 1802) 32 

74. Family TETRATOMIDAE Billberg, 1820 

Family author: M.V.L. Barclay 

Subfamily HALLOMENINAE Gistel, 1848 625 

HALLOMENUS Panzer, 1793 

binotatus (Quensel, 1790) 626 

humeralis Panzer, 1793 

Subfamily TETRATOMINAE Billberg, 1820 

TETRATOMA Fabricius, 1790 

ancora Fabricius, 1790 

desmarestii Latreille, 1807 627 

desmaresti auctt. (misspelling) 

fungorum Fabricius, 1790 

82 

75. Family MELANDRYIDAE Leach, 1815 


Subfamily MELANDRYINAE Leach, 1815 

ORCHESIA Latreille, 1807 

CLINOCRARA Thomson, C.G., 1859 

micans (Panzer, 1793) 

minor Walker, 1836 

tetratoma Thomson, C.G., 1864 

undulata Kraatz, 1853 

ANISOXYA Mulsant, 1856 

fuscula (Illiger, 1798) 

ABDERA Stephens, 1832 

CARIDA Mulsant, 1856 


biflexuosa (Curtis, 1829) 

bifasciata sensu auctt. Brit. non (Marsham, 1802) 

flexuosa (Paykull, 1799) 

quadrifasciata (Curtis, 1829) 

?bifasciata (Marsham, 1802) 

triguttata (Gyllenhal, 1810) 

PHLOIOTRYA Stephens, 1832 

PHLOEOTRYA auctt. (misspelling) 

vaudoueri Mulsant, 1856 

rufipes sensu auctt. Brit. non (Gyllenhal, 1810) 

XYLITA Paykull, 1798 

laevigata (Hellenius, 1786) 

buprestoides (Fabricius, 1792) 

HYPULUS Paykull, 1798 

quercinus (Quensel, 1790) 628 

ZILORA Mulsant, 1856 

ferruginea (Paykull, 1798) 

MELANDRYA Fabricius, 1801 

barbata (Fabricius, 1787) 

dubia sensu auctt. Brit. non (Schaller, 1783) 


Subfamily OSPHYINAE Mulsant, 1856 (1839) 

CONOPALPUS Gyllenhal, 1810 

testaceus (Olivier, 1790) 

OSPHYA Illiger, 1807 

bipunctata (Fabricius, 1775)

76. Family MORDELLIDAE Latreille, 1802 


The higher classification used in this checklist 

follows Franciscolo (1957), the most recent 

authoritative treatment of the the family as a 

whole. The systematics of the British species of 

Mordellistena have not been fully resolved and it is 

likely that further species will be found here and 

some species currently on the British list may 

prove to be based on misidentifications. I have 

listed some currently accepted synonyms that 

have not appeared in the most recent British 

checklists, where these names were based on 

British material or used in past British works. 

Subfamily MORDELLINAE Latreille, 1802 

Tribe MORDELLINI Latreille, 1802 

TOMOXIA Costa, A., 1853 

bucephala Costa, A., 1853 

biguttata (Gyllenhal, 1827) non (Rossi, 1794) 

fasciata sensu (Paykull, 1800) non (Fabricius, 1775) 

MORDELLA Linnaeus, 1758 

holomelaena Apfelbeck, 1914 

aculeata sensu auctt. Brit. partim non Linnaeus, 

1758 629 

leucaspis Küster, 1849 630 

aculeata sensu auctt. Brit. partim non Linnaeus, 

1758 

VARIIMORDA Méquignon, 1946 

MORDELLA sensu auctt. partim non Linnaeus, 

1758 

villosa (Schrank, 1781) 

fasciata (Fabricius, 1775) non (Forster, 1771) 

Tribe MORDELLISTENINI Ermisch, 1941 

MORDELLISTENA Costa, A., 1854 

Subgenus MORDELLISTENA Costa, A., 1854 

brevicauda (Boheman, 1849) 

humeralis (Linnaeus, 1758) 

flavescens (Marsham, 1802) 

axillaris (Gyllenhal, 1810) 

neuwaldeggiana (Panzer, 1796) 

brunnea (Fabricius, 1801) 

ferruginea (Marsham, 1802) 

parvula (Gyllenhal, 1827) 

inaequalis sensu auctt. Brit. non Mulsant, 1856 

pseudoparvula Ermisch, 1956 631 

parvula sensu auctt. Brit. partim non (Gyllenhal, 

1827) 

parvuloides Ermisch, 1956 632 

eludens Allen, 1999 633 

pseudopumila Ermisch, 1962 634 

pumila sensu auctt. Brit. partim non (Gyllenhal, 

1810) 

pumila (Gyllenhal, 1810) 

pygmaeola Ermisch, 1956 635 

pygmeola auctt. (misspelling) 

83 

Family MORDELLIDAE Latreille, 1802 

pumila sensu auctt. Brit. partim non (Gyllenhal, 

1810) 

secreta Horák, 1983 636 

variegata (Fabricius, 1798) 

humeralis sensu auctt. Brit. partim non (Linnaeus, 

1758) 

bicolor (Marsham, 1802) 

Subgenus MORDELLINA Schilsky, 1908 

PSEUDOMORDELLINA Ermisch, 1951 

acuticollis Schilsky, 1895 637 


1827) 

imitatrix Allen, 1995 638 

nanuloides Ermisch, 1967 639 


1827) 

MORDELLOCHROA Emery, 1876 

abdominalis (Fabricius, 1775) 

ventralis (Fabricius, 1792) 

nigra (Marsham, 1802) 

77. Family RIPIPHORIDAE Gemminger, 1870 

(1855) 

RHIPIPHORIDAE auctt. (misspelling) 


METOECUS Dejean, 1834 

paradoxus (Linnaeus, 1761) 

78. Family COLYDIIDAE Billberg, 1820 


Subfamily PYCNOMERINAE Erichson, 1845 

PYCNOMERUS Erichson, 1842 

fuliginosus Erichson, 1842 

Subfamily COLYDIINAE Billberg, 1820 

Tribe ORTHOCERINI Blanchard, 1845 

ORTHOCERUS Latreille, 1796 

SARROTRIUM Illiger, 1798 

clavicornis (Linnaeus, 1758) 

muticus (Linnaeus, 1767) 

Tribe SYNCHITINI Erichson, 1845 

SYNCHITA Hellwig, 1792 


juglandis (Hellwig, 1792) 

separanda (Reitter, 1882) 

humeralis sensu auctt. Brit. partes maj. non 


angularis Abeille de Perrin, 1901

Family TENEBRIONIDAE Latreille, 1802 Checklist of Beetles of the British Isles, 2012 Edition 

CICONES Curtis, 1827 

undatus Guérin-Méneville, 1829 640 

variegatus (Hellwig, 1792) 

BITOMA Herbst, 1793 

MONOTOMA Panzer, 1792 32 

DITOMA Illiger, 1807 

crenata (Fabricius, 1775) 

ENDOPHLOEUS Dejean, 1834 

markovichianus (Piller & Mitterpacher, 1783) 

spinosulus (Latreille, 1807) 

LANGELANDIA Aubé, 1843 

anophthalma Aubé, 1843 

Tribe COLYDIINI Billberg, 1820 

COLYDIUM Fabricius, 1792 

elongatum (Fabricius, 1787) 

AULONIUM Erichson, 1845 

ruficorne (Olivier, 1790) 

trisulcus (Geoffroy in Fourcroy, 1785) 

trisulcatum (misspelling) 

trisulcum (misspelling) 

79. Family TENEBRIONIDAE Latreille, 1802 

Family author: M.V.L. Barclay 

Subfamily LAGRIINAE Latreille, 1825 (1820) 

LAGRIA Fabricius, 1775 

atripes Mulsant & Guillebeau, 1855 

hirta (Linnaeus, 1758) 

Subfamily TENEBRIONINAE Latreille, 1802 

Tribe BOLITOPHAGINI Kirby, 1837 

BOLITOPHAGUS Illiger, 1798 

BOLETOPHAGUS auctt. (misspelling) 

reticulatus (Linnaeus, 1767) 

ELEDONA Latreille, 1796 

HELEDONA Fowler, 1890 (misspelling) 

agricola (Herbst, 1783) 

agaricola Fowler, 1890 (misspelling) 

Tribe TENEBRIONINI Latreille, 1802 

TENEBRIO Linnaeus, 1758 

molitor Linnaeus, 1758 

laticollis Stephens, 1832 

obscurus Fabricius, 1792 

84 

Tribe ALPHITOBIINI Reitter, 1917 

ALPHITOBIUS Stephens, 1829 

diaperinus (Panzer, 1796) 

laevigatus (Fabricius, 1781) 

Tribe TRIBOLIINI Gistel, 1848 

TRIBOLIUM MacLeay, 1825 

castaneum (Herbst, 1797) 

navale (Fabricius, 1775) 641 

confusum Jacquelin du Val, 1863 

destructor Uyttenboogaart, 1934 

LATHETICUS Waterhouse, C.O., 1880 

oryzae Waterhouse, C.O., 1880 

Tribe PALORINI Matthews, 2003 

PALORUS Mulsant, 1854 

CAENOCORSE Thomson, C.G., 1859 

ratzeburgii (Wissmann, 1848) 

melinus sensu Fowler, 1890 non (Herbst, 1783) 

subdepressus (Wollaston, 1864) 

Tribe ULOMINI Blanchard, 1845 

ULOMA Dejean, 1821 

culinaris (Linnaeus, 1758) 

Tribe PEDININI Eschscholtz, 1829 

PHYLAN Dejean, 1821 

HELIOPATHES sensu Fowler, 1890 non Dejean, 

1834 


Tribe MELANIMONINI Seidlitz, 1894 (1854) 

MELANIMON von Steven, 1829 

tibialis (Fabricius, 1781) 

Tribe OPATRINI Brullé, 1832 

OPATRUM Fabricius, 1775 

sabulosum (Linnaeus, 1760) 

Tribe HELOPINI Latreille, 1802 

HELOPS Fabricius, 1775 

caeruleus (Linnaeus, 1758) 

coeruleus Fowler, 1890 (misspelling) 

NALASSUS Mulsant, 1854 

CYLINDRINOTUS sensu auctt. partim non 

Faldermann, 1837 

CYLINDRONOTUS auctt. partim (misspelling) 

HELOPS sensu Fowler, 1890 partim non Fabricius, 

1775 

laevioctostriatus (Goeze, 1777) 

striatus (Geoffroy in Fourcroy, 1785)

XANTHOMUS Mulsant, 1854 

CYLINDRINOTUS sensu auctt. partim non 

Faldermann, 1837 

CYLINDRONOTUS auctt. partim (misspelling) 

HELOPS sensu Fowler, 1890 partim non Fabricius, 

1775 

pallidus (Curtis, 1830) 

Tribe BLAPTINI Leach, 1815 

BLAPS Fabricius, 1775 

lethifera Marsham, 1802 

similis Latreille, 1804 

mortisaga (Linnaeus, 1758) 

mucronata Latreille, 1804 

Subfamily DIAPERINAE Latreille, 1802 

Tribe CRYPTICINI Brullé, 1832 

CRYPTICUS Latreille, 1817 

quisquilius (Linnaeus, 1760) 

Tribe PHALERIINI Blanchard, 1845 

PHALERIA Latreille, 1802 

cadaverina (Fabricius, 1792) 

Tribe MYRMECHIXENINI Jacquelin du Val, 1858 

MYRMECHIXENUS Chevrolat, 1835 

MYRMECOXENUS Märkel, 1844 

subterraneus Chevrolat, 1835 

vaporariorum Guérin-Méneville, 1843 

Tribe HYPOPHLAEINI Billberg, 1820 

CORTICEUS Piller & Mitterpacher, 1783 

HYPOPHLAEUS Fabricius, 1790 

HYPOPHLOEUS auctt. (misspelling) 


fraxini (Kugelann, 1794) 

linearis (Fabricius, 1790) 

unicolor Piller & Mitterpacher, 1783 

castaneus (Fabricius, 1790) 

Tribe SCAPHIDEMINI Reitter, 1922 

SCAPHIDEMA Redtenbacher, 1849 

metallicum (Fabricius, 1792) 

Tribe DIAPERINI Latreille, 1802 

ALPHITOPHAGUS Stephens, 1832 

bifasciatus (Say, 1824) 

quadripustulatus Stephens, 1832 

unifasciatus Donisthorpe, 1925 

GNATOCERUS Thunberg, 1814 

GNATHOCERUS Agassiz, 1846 40 

cornutus (Fabricius, 1798) 

maxillosus (Fabricius, 1801) 

85 

PENTAPHYLLUS Dejean, 1821 

testaceus (Hellwig, 1792) 642 

Family TENEBRIONIDAE Latreille, 1802 

PLATYDEMA Laporte & Brullé, 1831 

violaceum (Fabricius, 1790) 

dytiscoides sensu Fowler, 1890 non (Rossi, 1790) 

DIAPERIS Geoffroy, 1762 

boleti (Linnaeus, 1758) 

Subfamily ALLECULINAE Laporte, 1840 

Tribe ALLECULINI Laporte, 1840 

PRIONYCHUS Solier, 1835 

ERYX Stephens, 1832 non Daudin, 1803 

ater (Fabricius, 1775) 

melanarius (Germar, 1813) 

fairmairei sensu auctt. Brit. non Reiche, 1861 

GONODERA Mulsant, 1856 

CISTELA sensu auctt. partim non Forster, 1771 

luperus (Herbst, 1783) 

PSEUDOCISTELA Crotch, 1873 

CISTELA Fabricius, 1775 non Geoffroy, 1762 643 

ceramboides (Linnaeus, 1758) 

ISOMIRA Mulsant, 1856 

CISTELA sensu Fowler, 1890 partim non Fabricius, 

1775 

murina (Linnaeus, 1758) 

MYCETOCHARA Berthold, 1827 

MYCETOCHARES Latreille, 1829 40 


bipustulata (Illiger, 1794) 

Tribe CTENIOPODINI Solier, 1835 

CTENIOPUS Solier, 1835 

sulphureus (Linnaeus, 1758) 

flavus (Scopoli, 1763) 

OMOPHLUS Dejean, 1834 

pubescens (Linnaeus, 1758) 

betulae (Herbst, 1783) 

rufitarsis (Leske, 1785) 

armeriae (Curtis, 1836)

Family OEDEMERIDAE Latreille, 1810 Checklist of Beetles of the British Isles, 2012 Edition 

80. Family OEDEMERIDAE Latreille, 1810 


Subfamily OEDEMERINAE Latreille, 1810 

Tribe NACERDINI Mulsant, 1858 

NACERDES Dejean, 1834 

NACERDA Stephens, 1839 (misspelling) 

ANONCODES Redtenbacher, 1845 


Tribe DITYLINI Mulsant, 1858 

CHRYSANTHIA Schmidt, W., 1846 

CHRYSARTHRIA Schmidt, W., 1844 32 

nigricornis Westhoff, 1881 

viridis sensu Schmidt, W., 1846 non (De Geer, 

1775) 

Tribe ASCLERINI Gistel, 1848 

ISCHNOMERA Stephens, 1832 

ASCLERA Dejean, 1834 

caerulea (Linnaeus, 1758) 

coerulea Fowler, 1890 (misspelling) 

cinerascens (Pandellé, 1867) 644 

cyanea (Fabricius, 1792) 645 

caerulea sensu auctt. partim non (Linnaeus, 1758) 

graeca (Dahlgren, 1976) 

sanguinicollis (Fabricius, 1787) 

Tribe OEDEMERINI Latreille, 1810 

OEDEMERA Olivier, 1789 

Subgenus ONCOMERA Stephens, 1832 

femoralis (Olivier, 1803) 

femorata (Fabricius, 1792) non (Scopoli, 1763) 

Subgenus OEDEMERA Olivier, 1789 

lurida (Marsham, 1802) 

nobilis (Scopoli, 1763) 

virescens (Linnaeus, 1767) 

81. Family MELOIDAE Gyllenhal, 1810 


Subfamily MELOINAE Gyllenhal, 1810 

LYTTA Fabricius, 1775 

CANTHARIS sensu auctt. non Linnaeus, 1758 

vesicatoria (Linnaeus, 1758) 

MELOE Linnaeus, 1758 

autumnalis Olivier, 1792 

brevicollis Panzer, 1793 

cicatricosus Leach, 1813 

mediterraneus Müller, J., 1925 646 

rugosus sensu auctt. partim non Marsham, 1802 

proscarabaeus Linnaeus, 1758 

86 

rugosus Marsham, 1802 647 

variegatus Donovan, 1793 

violaceus Marsham, 1802 

Subfamily NEMOGNATHINAE Laporte, 1840 

SITARIS Latreille, 1802 

APALUS sensu auctt. Brit. non Fabricius, 1775 

muralis (Forster, 1771) 

82. Family MYCTERIDAE Oken, 1843 


MYCTERUS Clairville, 1798 

curculioides (Fabricius, 1781) 

curculionoides auctt. (misspelling) 

83. Family PYTHIDAE Solier, 1834 


PYTHO Latreille, 1796 

depressus (Linnaeus, 1767) 

84. Family PYROCHROIDAE Latreille, 1806 


PYROCHROA Geoffroy, 1762 648 

coccinea (Linnaeus, 1761) 

serraticornis (Scopoli, 1763) 

SCHIZOTUS Newman, 1838 

PYROCHROA sensu Fowler, 1890 partim non 

Geoffroy, 1762 

PYROCHROELLA Reitter, 1911 


85. Family SALPINGIDAE Leach, 1815 


Subfamily AGLENINAE Horn, 1878 

AGLENUS Erichson, 1845 

brunneus (Gyllenhal, 1813) 

Subfamily SALPINGINAE Leach, 1815 

LISSODEMA Curtis, 1833 

cursor (Gyllenhal, 1813) 

kirkae Donisthorpe, 1925 

denticolle (Gyllenhal, 1813) 

quadripustulatum (Marsham, 1802) non (Fabricius, 

1775)

RABOCERUS Mulsant, 1859 

SALPINGUS sensu Fowler, 1890 partim non Illiger, 

1801 

foveolatus (Ljungh, 1824) 

mutilatus sensu Champion, 1886 non (Beck, 1817) 

bishopi Sharp, 1909 

gabrieli Gerhardt, 1901 

foveolatus sensu auctt. Brit. non (Ljungh, 1824) 

SPHAERIESTES Stephens, 1829 

SALPINGUS sensu auctt. non Illiger, 1801 

SALPINGELLUS Reitter, 1911 

ater (Paykull, 1798) 649 

picea Germar, 1825 

stockmanni (Biström, 1977) 

castaneus (Panzer, 1796) 

reyi (Abeille de Perrin, 1874) 

ater sensu Stephens, 1831 non (Paykull, 1798) 

aeratus sensu auctt. Brit. non Mulsant, 1859 

VINCENZELLUS Reitter, 1911 

RHINOSIMUS sensu Fowler, 1890 partim non 


ruficollis (Panzer, 1794) 

viridipennis (Latreille, 1804) 

SALPINGUS Illiger, 1801 

RHINOSIMUS Latreille, 1802 

planirostris (Fabricius, 1787) 650 

ruficollis (Linnaeus, 1761) 

86. Family ANTHICIDAE Latreille, 1819 651 


Subfamily ANTHICINAE Latreille, 1819 

ANTHICUS Paykull, 1798 

angustatus Curtis, 1838 

antherinus (Linnaeus, 1760) 

bimaculatus (Illiger, 1801) 

flavipes (Panzer, 1797) 

scoticus Rye, 1872 652 

tristis Schmidt, 1842 

ater sensu Stephens, 1832 non (Panzer, 1796) 

ssp. schaumii Wollaston, 1857 

CORDICOLLIS Marseul, 1879 

CORDICOMUS Pic, 1894 

ANTHICUS sensu auctt. partim non Paykull, 1798 

instabilis (Schmidt, 1842) 

tibialis (Curtis, 1838) non (Waltl, 1835) 

CYCLODINUS Mulsant & Rey, 1866 


constrictus (Curtis, 1838) 653 

humilis sensu auctt. Brit. non (Germar, 1824) 

albionis (Krekich-Strassoldo, 1919) 

larvipennis sensu (Krekich-Strassoldo, 1919) non 

(Marseul, 1879) 

salinus (Crotch, 1867) 

crotchi (Pic, 1893) 

87 

Family ANTHICIDAE Latreille, 1819 

OMONADUS Mulsant & Rey, 1866 


bifasciatus (Rossi, 1792) 

floralis (Linnaeus, 1758) 

formicarius (Goeze, 1777) 

quisquilius (Thomson, C.G., 1864) 

STRICTICOLLIS Marseul, 1879 

STRICTICOMUS Pic, 1894 


tobias (Marseul, 1879) 

Subfamily NOTOXINAE Stephens, 1829 

NOTOXUS Geoffroy, 1762 654 

monoceros (Linnaeus, 1760) 

87. Family ADERIDAE Csiki, 1909 


ADERUS Stephens, 1829 

XYLOPHILUS Latreille, 1825 non Mannerheim, 

1823 

HYLOPHILUS Berthold, 1827 non Temminck, 1822 

populneus (Creutzer in Panzer, 1796) 

boleti (Marsham, 1802) 

EUGLENES Westwood, 1830 

ADERUS sensu auctt. partim non Stephens, 1829 

oculatus (Paykull, 1798) 

pygmaeus sensu auctt. Brit. non (De Geer, 1774) 

VANONUS Casey, 1895 

ADERUS sensu auctt. partim non Stephens, 1829 

brevicornis (Perris, 1869) 

neglectus sensu auctt. Brit. partim non (Jacquelin 

du Val, 1862) 

88. Family SCRAPTIIDAE Gistel, 1848 


The higher classification used in this checklist 

follows Franciscolo (1972), the most recent 

authoritative treatment of the family as a whole. I 

have listed some currently accepted synonyms 

that have not appeared in the most recent British 

checklists, where these names were based on 

British material or used in past British works. I 

have not included those infrasubspecific names 

which are not available under article 45 of the 

Fourth Edition of the International Code of 

Zoological Nomenclature. 

Subfamily SCRAPTIINAE Gistel, 1848 

SCRAPTIA Latreille, 1807 

dubia (Olivier, 1790) 

fuscula Müller, P.W.J., 1821 

testacea Allen, 1940

Family CERAMBYCIDAE Latreille, 1802 Checklist of Beetles of the British Isles, 2012 Edition 

fuscula sensu auctt. Brit. partim non Müller, P.W.J., 

1821 

Subfamily ANASPIDINAE Mulsant, 1856 

ANASPIS Geoffroy, 1762 

Subgenus ANASPIS Geoffroy, 1762 

bohemica Schilsky, 1898 

hudsoni sensu Buck, 1954 partim non Donisthorpe, 

1909 

fasciata (Forster, 1771) 

humeralis sensu auctt. non (Fabricius, 1775) 

biguttata (Rossi, 1794) 

geoffroyi Müller, P.W.J., 1821 

quadrinotata Stephens, 1832 

quadripustulata Stephens, 1832 

scapularis Stephens, 1832 

subfasciata Stephens, 1832 

frontalis (Linnaeus, 1758) 

lateralis (Fabricius, 1792) 

garneysi Fowler, 1889 

lurida Stephens, 1832 

testacea (Marsham, 1802) non (Fabricius, 1787) 

subtestacea Stephens, 1832 

maculata (Geoffroy in Fourcroy, 1785) 

melanopa (Forster, 1771) 655 

nigricollis (Marsham, 1802) 

obscura (Marsham, 1802) 

pallida (Marsham, 1802) 

florenceae Donisthorpe, 1928 

pulicaria Costa, A., 1854 

lateralis sensu Stephens, 1832 non (Fabricius, 

1792) 

forcipata Mulsant, 1856 

regimbarti Schilsky, 1895 

ruficollis sensu auctt. Brit. non (Fabricius, 1792) 

fraudulenta Joy, 1912 

thoracica (Linnaeus, 1758) 656 

septentrionalis Champion, 1891 

latipalpis Schilsky, 1895 

schilskyana Csiki, 1915 657 

marginicollis Lindberg, 1925 

Subgenus NASSIPA Emery, 1876 658 

costai Emery, 1876 

costae auctt. (misspelling) 

fuscescens Stephens, 1832 659 

flava sensu auctt. non (Linnaeus, 1758) 

thoracica sensu Fowler, 1889 non (Linnaeus, 1758) 

rufilabris (Gyllenhal, 1827) 

hudsoni Donisthorpe, 1909 

Superfamily CHRYSOMELOIDEA Latreille, 

1802 

89. Family CERAMBYCIDAE Latreille, 1802 660 

Family author: M. Rejzek 

Subfamily PRIONINAE Latreille, 1802 

PRIONUS Geoffroy, 1762 

coriarius (Linnaeus, 1758) 

88 

Subfamily LEPTURINAE Latreille, 1802 

Tribe RHAGIINI Kirby, 1837 

RHAGIUM Fabricius, 1775 

Subgenus RHAGIUM Fabricius, 1775 

inquisitor (Linnaeus, 1758) 

indagator Fabricius, 1787 

Subgenus HAGRIUM Villiers, 1978 

bifasciatum Fabricius, 1775 

Subgenus MEGARHAGIUM Reitter, 1913 

mordax (De Geer, 1775) 

inquisitor sensu auctt. Brit. partim non (Linnaeus, 

1758) 

STENOCORUS Geoffroy, 1762 

STENOCHORUS auctt. (misspelling) 

TOXOTUS Dejean, 1821 

meridianus (Linnaeus, 1758) 

DINOPTERA Mulsant, 1863 

PACHYTA sensu auctt. partim non Dejean, 1821 

ACMAEOPS sensu auctt. non LeConte, 1850 

collaris (Linnaeus, 1758) 

GRAMMOPTERA Audinet-Serville, 1835 

abdominalis (Stephens, 1831) 

variegata (Germar, 1824) non (Fabricius, 1775) 

analis (Herrich-Schäffer in Panzer, 1832) 

ruficornis (Fabricius, 1781) 

holomelina Pool, 1905 661 

ustulata (Schaller, 1783) 

praeusta (Fabricius, 1787) 

Tribe LEPTURINI Latreille, 1802 

PEDOSTRANGALIA Sokolov, 1896 

STRANGALIA sensu auctt. partim non Audinet- 

Serville, 1835 

revestita (Linnaeus, 1767) 

LEPTUROBOSCA Reitter, 1913 

LEPTURA sensu auctt. partim non Linnaeus, 1758 

virens (Linnaeus, 1758) 

LEPTURA Linnaeus, 1758 



aurulenta Fabricius, 1792 

quadrifasciata Linnaeus, 1758 

ANASTRANGALIA Casey, 1924 


sanguinolenta (Linnaeus, 1761) 

STICTOLEPTURA Casey, 1924 


rubra (Linnaeus, 1758) 

scutellata (Fabricius, 1781)

PARACORYMBIA Miroshnikov, 1998 


fulva (De Geer, 1775) 

ANOPLODERA Mulsant, 1839 


sexguttata (Fabricius, 1775) 

JUDOLIA Mulsant, 1863 

PACHYTA sensu Fowler, 1890 partim non Dejean, 

1821 

sexmaculata (Linnaeus, 1758) 

PACHYTODES Pic, 1891 

PACHYTA sensu auctt. partim non Dejean, 1821 

JUDOLIA sensu auctt. partim non Mulsant, 1863 

cerambyciformis (Schrank, 1781) 

ALOSTERNA Mulsant, 1863 

GRAMMOPTERA sensu Fowler, 1890 partim non 

Audinet-Serville, 1835 

tabacicolor (De Geer, 1775) 

PSEUDOVADONIA Lobanov, Danilevsky & Murzin, 

1981 


livida (Fabricius, 1777) 

STRANGALIA Audinet-Serville, 1835 

attenuata (Linnaeus, 1758) 

RUTPELA Nakane & Ohbayashi, 1959 



maculata (Poda, 1761) 

armata (Herbst, 1784) 

STENURELLA Villiers, 1974 




nigra (Linnaeus, 1758) 

Subfamily SPONDYLIDINAE Audinet-Serville, 

1832 

Tribe ASEMINI Thomson, J., 1861 

ASEMUM Eschscholtz, 1830 

striatum (Linnaeus, 1758) 

TETROPIUM Kirby, 1837 

ISARTHRON Dejean, 1835 662 

CRIOMORPHUS Mulsant, 1839 662 

castaneum (Linnaeus, 1758) 

gabrieli Weise, 1905 

fuscum sensu auctt. Brit. non (Fabricius, 1787) 

crawshayi Sharp, 1905 

89 

Family CERAMBYCIDAE Latreille, 1802 

ARHOPALUS Audinet-Serville, 1834 

CRIOCEPHALUS Dejean, 1835 

CRIOCEPHALUM Mulsant, 1839 

ferus (Mulsant, 1839) 

tristis sensu auctt. non (Fabricius, 1787) 

polonicus (Motschulsky, 1845) 

rusticus (Linnaeus, 1758) 

Subfamily CERAMBYCINAE Latreille, 1802 

Tribe CALLIDIOPINI Lacordaire, 1868 

TRINOPHYLUM Bates, 1878 

TRINOPHYLLUM auctt. (misspelling) 

cribratum Bates, 1878 

Tribe CERAMBYCINI Latreille, 1802 

CERAMBYX Linnaeus, 1758 

cerdo Linnaeus, 1758 

heros sensu Stephens, 1831 non Scopoli, 1763 

scopolii Fuessly, 1775 

cerdo sensu Scopoli, 1763 non Linnaeus, 1758 

Tribe GRACILIINI Mulsant, 1839 

GRACILIA Audinet-Serville, 1834 

minuta (Fabricius, 1781) 

Tribe OBRIINI Mulsant, 1839 

OBRIUM Dejean, 1821 

brunneum (Fabricius, 1792) 

cantharinum (Linnaeus, 1767) 

Tribe PSEBIINI Lacordaire, 1868 

NATHRIUS Brèthes, 1916 

LEPTIDEA Mulsant, 1839 non Billberg, 1820 

LEPTIDEELLA Strand, E., 1936 

brevipennis (Mulsant, 1839) 

Tribe MOLORCHINI Gistel, 1848 

MOLORCHUS Fabricius, 1792 

CAENOPTERA Thomson, C.G., 1859 

minor (Linnaeus, 1758) 

GLAPHYRA Newman, 1840 

MOLORCHUS sensu Fowler, 1890 partim non 

Fabricius, 1792 

CAENOPTERA sensu Joy, 1932 partim non 


umbellatarum (von Schreber, 1759) 

Tribe CALLICHROMATINI Swainson, 1840 

AROMIA Audinet-Serville, 1833 

moschata (Linnaeus, 1758)

Family MEGALOPODIDAE Latreille, 1802 Checklist of Beetles of the British Isles, 2012 Edition 

Tribe CALLIDIINI Kirby, 1837 

HYLOTRUPES Audinet-Serville, 1834 

bajulus (Linnaeus, 1758) 

SEMANOTUS Mulsant, 1839 

russicus (Fabricius, 1776) 663 

CALLIDIUM Fabricius, 1775 

violaceum (Linnaeus, 1758) 

PYRRHIDIUM Fairmaire, 1864 

CALLIDIUM sensu Fowler, 1890 partim non 


sanguineum (Linnaeus, 1758) 

PHYMATODES Mulsant, 1839 664 



testaceus (Linnaeus, 1758) 

variabilis (Linnaeus, 1761) 

POECILIUM Fairmaire, 1864 



alni (Linnaeus, 1767) 

lividum (Rossi, 1794) 665 

Tribe CLYTINI Mulsant, 1839 

CLYTUS Laicharting, 1784 

arietis (Linnaeus, 1758) 

medioniger Allen, 1959 

PLAGIONOTUS Mulsant, 1842 

CLYTUS sensu Fowler, 1890 partim non 

Laicharting, 1784 

arcuatus (Linnaeus, 1758) 

Tribe ANAGLYPTINI Lacordaire, 1868 

ANAGLYPTUS Mulsant, 1839 

ANACLYPTUS Mulsant, 1839 

CLYTUS sensu Fowler, 1890 partim non 

Laicharting, 1784 

mysticus (Linnaeus, 1758) 

Subfamily LAMIINAE Latreille, 1825 

Tribe MESOSINI Mulsant, 1839 

MESOSA Latreille, 1829 

nebulosa (Fabricius, 1781) 

nubila (Gmelin in Linnaeus, 1790) 

Tribe AGAPANTHIINI Mulsant, 1839 

AGAPANTHIA Audinet-Serville, 1835 

villosoviridescens (De Geer, 1775) 

lineatocollis (Donovan, 1797) 

90 

Tribe LAMIINI Latreille, 1825 

LAMIA Fabricius, 1775 

textor (Linnaeus, 1758) 

Tribe POGONOCHERINI Mulsant, 1839 

POGONOCHERUS Dejean, 1821 

POGONOCHAERUS auctt. (misspelling) 

fasciculatus (De Geer, 1775) 

hispidulus (Piller & Mitterpacher, 1783) 

bidentatus Thomson, C.G., 1866 

hispidus (Linnaeus, 1758) 

dentatus (Geoffroy in Fourcroy, 1785) 

Tribe ACANTHOCININI Blanchard, 1845 

ACANTHOCINUS Dejean, 1821 

aedilis (Linnaeus, 1758) 

LEIOPUS Audinet-Serville, 1835 

linnei Wallin, Nylander & Kvamme, 2009 666 

nebulosus sensu auctt. partim non (Linnaeus, 1758) 

nebulosus (Linnaeus, 1758) 

Tribe SAPERDINI Mulsant, 1839 

SAPERDA Fabricius, 1775 

carcharias (Linnaeus, 1758) 

populnea (Linnaeus, 1758) 

scalaris (Linnaeus, 1758) 

STENOSTOLA Dejean, 1835 

dubia (Laicharting, 1784) 

ferrea sensu auctt. Brit. non (Schrank, 1776) 

Tribe PHYTOECIINI Mulsant, 1839 

PHYTOECIA Dejean, 1835 

cylindrica (Linnaeus, 1758) 

OBEREA Dejean, 1835 

oculata (Linnaeus, 1758) 

Tribe TETROPINI Portevin, 1927 

TETROPS Stephens, 1829 

praeustus (Linnaeus, 1758) 

starkii Chevrolat, 1859 667 

90. Family MEGALOPODIDAE Latreille, 1802 

Family author: M.L. Cox 

Subfamily ZEUGOPHORINAE Böving & 

Craighead, 1931 

ZEUGOPHORA Kunze, 1818 

flavicollis (Marsham, 1802) 

subspinosa (Fabricius, 1781)

turneri Power, 1863 

91. Family ORSODACNIDAE Thomson, C.G., 

1859 


ORSODACNE Latreille, 1802 

ORSODACNA Latreille, 1804 

cerasi (Linnaeus, 1758) 

humeralis Latreille, 1804 

lineola (Panzer, 1795) non (Fabricius, 1781) 668 

92. Family CHRYSOMELIDAE Latreille, 1802 


Subfamily BRUCHINAE Latreille, 1802 

BRUCHUS Linnaeus, 1767 669 

MYLABRIS Geoffroy, 1762 670 

LARIA sensu auctt. non Scopoli, 1763 

atomarius (Linnaeus, 1761) 

fahraei Gyllenhal, 1839 

viciae sensu Fowler, 1890 non Olivier, 1795 

brachialis Fåhraeus, 1839 671 

ervi Frölich, 1799 672 

loti Paykull, 1800 

pisorum (Linnaeus, 1758) 

pisi Linnaeus, 1767 

rufimanus Boheman, 1833 

affinis sensu auctt. Brit. non Frölich, 1799 

velutinus Mulsant & Rey, 1858 

rufipes Herbst, 1783 

luteicornis sensu auctt. Brit. non Illiger, 1794 

BRUCHIDIUS Schilsky, 1905 

BRUCHUS sensu Fowler, 1890 partim non 


cisti (Fabricius, 1775) 

unicolor (Olivier, 1795) 

canus (Germar, 1824) 

debilis (Gyllenhal, 1833) 

cisti "(Paykull, 1800)" sensu Joy, 1932 

incarnatus (Boheman, 1833) 673 

olivaceus (Germar, 1824) 674 

canus sensu auctt. Brit. non (Germar, 1824) 

unicolor sensu Schilsky, 1905 non (Olivier, 1795) 

varius (Olivier, 1795) 675 

villosus (Fabricius, 1792) 

cisti sensu (Paykull, 1800) non (Fabricius, 1775) 

ater (Marsham, 1802) non (Scriba, 1790) 

fasciatus sensu auctt. non (Olivier, 1795) 

ACANTHOSCELIDES Schilsky in Küster & Kraatz, 

1905 

obtectus (Say, 1831) 

obsoletus sensu auctt. non Say, 1831 

CALLOSOBRUCHUS Pic, 1902 

chinensis (Linnaeus, 1758) 


maculatus (Fabricius, 1775) 

91 

Family ORSODACNIDAE Thomson, C.G., 1859 

quadrimaculatus (Fabricius, 1792) 

Subfamily DONACIINAE Kirby, 1837 

MACROPLEA Samouelle, 1819 

HAEMONIA Dejean, 1821 

appendiculata (Panzer, 1794) 

equiseta (Fabricius, 1798) 

mutica (Fabricius, 1792) 

curtisi (Lacordaire, 1845) 

DONACIA Fabricius, 1775 

aquatica (Linnaeus, 1758) 

dentipes Fabricius, 1792 

bicolora Zschach, 1788 

bicolor auctt. (misspelling) 

sagittariae Fabricius, 1792 

cinerea Herbst, 1784 

clavipes Fabricius, 1792 

menyanthis Fabricius, 1801 

menyanthidis Gyllenhal, 1813 

crassipes Fabricius, 1775 

dentata Hoppe, 1795 

impressa Paykull, 1799 

marginata Hoppe, 1795 

limbata Panzer, 1796 

lemnae Fabricius, 1801 

obscura Gyllenhal, 1813 

semicuprea Panzer, 1796 

simplex Fabricius, 1775 

linearis Hoppe, 1795 

sparganii Ahrens, 1810 

thalassina Germar, 1811 

versicolorea (Brahm, 1791) 

vulgaris Zschach, 1788 

typhae Ahrens, 1810 

PLATEUMARIS Thomson, C.G., 1859 

DONACIA sensu Fowler, 1890 partim non Fabricius, 

1775 

bracata (Scopoli, 1772) 

braccata auctt. (misspelling) 

discolor (Panzer, 1795) 

rustica (Kunze, 1818) 

affinis (Kunze, 1818) 

sericea (Linnaeus, 1758) 

Subfamily CRIOCERINAE Latreille, 1804 

LEMA Fabricius, 1798 

cyanella (Linnaeus, 1758) 

puncticollis Curtis, 1830 

OULEMA des Gozis, 1886 

erichsoni (Suffrian, 1841) 

melanopus (Linnaeus, 1758) 676 

melanopa auctt. 

obscura (Stephens, 1831) 

cyanella sensu auctt. partim non (Linnaeus, 1758) 

gallaeciana (Heyden, 1870) 

lichenis sensu (Weise, 1882) non (Voet, 1806) 

rufocyanea (Suffrian, 1847) 677 

melanopus sensu auctt. partim non (Linnaeus, 

1758) 

duftschmidi (Redtenbacher, 1874)

Family CHRYSOMELIDAE Latreille, 1802 Checklist of Beetles of the British Isles, 2012 Edition 

septentrionis (Weise, 1880) 

CRIOCERIS Geoffroy, 1762 

asparagi (Linnaeus, 1758) 

LILIOCERIS Reitter, 1912 

CRIOCERIS sensu Fowler, 1890 partim non Müller, 

O.F., 1764 

lilii (Scopoli, 1763) 

Subfamily CRYPTOCEPHALINAE Gyllenhal, 1813 

Tribe CLYTRINI Kirby, 1837 

LABIDOSTOMIS Dejean, 1836 

tridentata (Linnaeus, 1758) 

CLYTRA Laicharting, 1781 

CLYTHRA Fabricius, 1798 (misspelling) 

laeviuscula Ratzeburg, 1837 

quadripunctata (Linnaeus, 1758) 

SMARAGDINA Dejean, 1836 

CYANIRIS Dejean, 1836 non Dalman, 1816 

GYNANDROPHTALMA Lacordaire, 1848 

GYNANDROPHTHALMA Jacquelin du Val, 1865 

affinis (Illiger, 1794) 

Tribe CRYPTOCEPHALINI Gyllenhal, 1813 

CRYPTOCEPHALUS Geoffroy, 1762 

aureolus Suffrian, 1847 

biguttatus (Scopoli, 1763) 

bilineatus (Linnaeus, 1767) 

bipunctatus (Linnaeus, 1758) 

coryli (Linnaeus, 1758) 

decemmaculatus (Linnaeus, 1758) 

exiguus Schneider, D.H., 1792 

frontalis Marsham, 1802 

fulvus (Goeze, 1777) 

hypochaeridis (Linnaeus, 1758) 

cristula Dufour, 1843 

labiatus (Linnaeus, 1761) 

moraei (Linnaeus, 1758) 

nitidulus Fabricius, 1787 

ochrostoma Harold, 1872 

parvulus Müller, O.F., 1776 

nigrocoeruleus (Goeze, 1777) 

primarius Harold, 1872 

punctiger Paykull, 1799 

pusillus Fabricius, 1777 

querceti Suffrian, 1848 

sexpunctatus (Linnaeus, 1758) 

violaceus Laicharting, 1781 

Subfamily LAMPROSOMATINAE Lacordaire, 1848 

OOMORPHUS Curtis, 1831 

LAMPROSOMA sensu auctt. non Kirby, 1818 678 

concolor (Sturm, 1807) 

92 

Subfamily EUMOLPINAE Hope, 1840 

BROMIUS Dejean, 1836 

ADOXUS Kirby, 1837 

obscurus (Linnaeus, 1758) 679 

Subfamily CHRYSOMELINAE Latreille, 1802 

TIMARCHA Samouelle, 1819 

goettingensis (Linnaeus, 1758) 

coriaria (Laicharting, 1781) 

tenebricosa (Fabricius, 1775) 

CHRYSOLINA Motschulsky, 1860 680 

CHRYSOMELA sensu auctt. non Linnaeus, 1758 

americana (Linnaeus, 1758) 681 

banksii (Fabricius, 1775) 682 

banksi auctt. (misspelling) 

brunsvicensis (Gravenhorst, 1807) 

didymata sensu auctt. Brit. non (Scriba, 1791) 

cerealis (Linnaeus, 1767) 

coerulans (Scriba, 1791) 683 

caerulans auctt. (misspelling) 

fastuosa (Scopoli, 1763) 

graminis (Linnaeus, 1758) 

haemoptera (Linnaeus, 1758) 

herbacea (Duftschmid, 1825) 

menthastri (Suffrian, 1851) 

menthrasti (Fowler, 1890) (misspelling) 

hyperici (Forster, 1771) 

latecincta (Demaison, 1896) 

sanguinolenta sensu auctt. Brit. partim non 


crassicornis (Helliesin, 1911) non (Fabricius, 1775) 

hellieseni Silfverberg, 1977 684 

ssp. intermedia (Franz, 1938) 685 

marginata (Linnaeus, 1758) 

oricalcia (Müller, O.F., 1776) 

orichalcia auctt. (misspelling) 

hobsoni (Stephens, 1831) 

polita (Linnaeus, 1758) 

sanguinolenta (Linnaeus, 1758) 

marginalis (Duftschmid, 1825) 

staphylaea (Linnaeus, 1758) 

staphylea (Fowler, 1890) (misspelling) 

sturmi (Westhoff, 1882) 

goettingensis sensu (Linnaeus, 1761) non 


violacea Weise, 1916 non (Müller, O.F., 1776) 

varians (Schaller, 1783) 

GASTROPHYSA Dejean, 1836 

GASTROIDEA Hope, 1840 

polygoni (Linnaeus, 1758) 

viridula (De Geer, 1775) 

PHAEDON Latreille, 1829 

armoraciae (Linnaeus, 1758) 

cochleariae (Fabricius, 1792) 

regnianum Tottenham, 1941 

concinnus Stephens, 1831 

tumidulus (Germar, 1824)

HYDROTHASSA Thomson, C.G., 1859 

glabra (Herbst, 1783) 

aucta (Fabricius, 1787) 

hannoveriana (Fabricius, 1775) 

hannoverana Fowler, 1890 (misspelling) 

marginella (Linnaeus, 1758) 

PRASOCURIS Latreille, 1802 

HELODES Paykull, 1799 non Latreille, 1796 

junci (Brahm, 1790) 

phellandrii (Linnaeus, 1758) 

PLAGIODERA Dejean, 1836 

versicolora (Laicharting, 1781) 

armoraciae sensu (Fabricius, 1775) non (Linnaeus, 

1758) 

CHRYSOMELA Linnaeus, 1758 

MELASOMA Stephens, 1831 

aenea Linnaeus, 1758 

populi Linnaeus, 1758 

tremula Fabricius, 1787 

tremulae auctt. (misspelling) 

longicollis (Suffrian, 1851) 

GONIOCTENA Dejean, 1836 

PHYTODECTA Kirby, 1837 

decemnotata (Marsham, 1802) 

rufipes (De Geer, 1775) non (Linnaeus, 1758) 

olivacea (Forster, 1771) 

pallida (Linnaeus, 1758) 

viminalis (Linnaeus, 1758) 

PHRATORA Dejean, 1836 

PHYLLODECTA Kirby, 1837 

laticollis Suffrian, 1851 

cavifrons Thomson, C.G., 1866 

polaris Schneider, J.S., 1886 

vitellinae (Linnaeus, 1758) 

vulgatissima (Linnaeus, 1758) 

Subfamily GALERUCINAE Latreille, 1802 

Tribe GALERUCINI Latreille, 1802 

GALERUCELLA Crotch, 1873 

calmariensis (Linnaeus, 1767) 

lineola (Fabricius, 1781) 

nymphaeae (Linnaeus, 1758) 

pusilla (Duftschmid, 1825) 

sagittariae (Gyllenhal, 1813) 

fergussoni Fowler, 1910 

grisescens sensu auctt. Brit. non (Joannis, 1866) 686 

tenella (Linnaeus, 1761) 

PYRRHALTA Joannis, 1865 

GALERUCELLA sensu Fowler, 1890 partim non 

Crotch, 1873 

viburni (Paykull, 1799) 

XANTHOGALERUCA Laboissière, 1934 

luteola (Müller, O.F., 1766) 687 

93 

Family CHRYSOMELIDAE Latreille, 1802 

GALERUCA Geoffroy, 1762 

ADIMONIA Laicharting, 1781 

GALLERUCA Fabricius, 1792 (misspelling) 

laticollis (Sahlberg, C.R., 1838) 

interrupta sensu auctt. non Illiger, 1802 

circumdata Duftschmid, 1825 

tanaceti (Linnaeus, 1758) 688 

LOCHMAEA Weise, 1883 

LOCHMAEATA Strand, E., 1935 

caprea (Linnaeus, 1758) 

capreae auctt. (misspelling) 

crataegi (Forster, 1771) 

suturalis (Thomson, C.G., 1866) 

DIABROTICA Dejean, 1836 

virgifera LeConte, 1858 689 

PHYLLOBROTICA Dejean, 1836 

quadrimaculata (Linnaeus, 1758) 

LUPERUS Geoffroy, 1762 


longicornis (Fabricius, 1781) 

rufipes sensu auctt. Brit. ? (Scopoli, 1763) 

CALOMICRUS Dillwyn, 1829 

LUPERUS sensu Fowler, 1890 partim non Müller, 

O.F., 1764 

circumfusus (Marsham, 1802) 

circumfuscus auctt. (misspelling) 

nigrofasciatus sensu Weise, 1886 non (Goeze, 

1777) 

AGELASTICA Dejean, 1836 


SERMYLASSA Reitter, 1912 

SERMYLA Chapuis, 1875 non Adams, 1854 

halensis (Linnaeus, 1767) 

Tribe ALTICINI Newman, 1834 

LUPEROMORPHA Weise, 1887 

xanthodera (Fairmaire, 1888) 690 

PHYLLOTRETA Dejean, 1836 

atra (Fabricius, 1775) 

consobrina (Curtis, 1837) 

hintoni Donisthorpe, 1944 

cruciferae (Goeze, 1777) 

diademata Foudras, 1860 

exclamationis (Thunberg, 1784) 

flexuosa (Illiger, 1794) 

sinuata (Stephens, 1831) 

nemorum (Linnaeus, 1758) 

nigripes (Fabricius, 1775) 

nodicornis (Marsham, 1802) 

ochripes (Curtis, 1837) 

punctulata (Marsham, 1802) 

aerea Allard, 1859 

striolata (Fabricius, 1803) 

vittata sensu auctt. non (Fabricius, 1801) 

tetrastigma (Comolli, 1837)

Family CHRYSOMELIDAE Latreille, 1802 Checklist of Beetles of the British Isles, 2012 Edition 

undulata Kutschera, 1860 

vittula (Redtenbacher, 1849) 

APHTHONA Dejean, 1836 

?atratula Allard, 1859 

atrovirens sensu auctt. Brit. non Förster, 1849 691 

atrocaerulea (Stephens, 1829) 

cyanella (Redtenbacher, 1849) 

puncticollis Allard, 1866 

euphorbiae (Schrank, 1781) 

virescens Foudras, 1860 

aeneomicans sensu auctt. non Allard, 1875 

herbigrada (Curtis, 1837) 

lutescens (Gyllenhal, 1808) 

melancholica Weise, 1888 

venustula sensu auctt. Brit. non Kutschera, 1861 

nigriceps (Redtenbacher, 1842) 

pallida sensu auctt. Brit. partim non Bach, 1856 

nonstriata (Goeze, 1777) 

coerulea (Fourcroy, 1785) 

pallida (Bach, 1856) 692 

nigriceps sensu auctt. Brit. partim non 

(Redtenbacher, 1842) 

LONGITARSUS Berthold, 1827 

absynthii Kutschera, 1862 

absinthii auctt. (misspelling) 

aeneicollis (Faldermann, 1837) 

suturalis (Marsham, 1802) non (Fabricius, 1775) 

nigricollis (Foudras, 1860) 

aeruginosus (Foudras, 1860) 

agilis (Rye, 1868) 

anchusae (Paykull, 1799) 

atricillus (Linnaeus, 1761) 

ballotae (Marsham, 1802) 

cerinus sensu auctt. Brit. partim non (Foudras, 

1860) 

brunneus (Duftschmid, 1825) 

castaneus sensu auctt. Brit. non (Duftschmid, 1825) 

curtus (Allard, 1860) 

dorsalis (Fabricius, 1781) 693 

exoletus (Linnaeus, 1758) 

femoralis (Marsham, 1802) 

ferrugineus (Foudras, 1860) 

waterhousei Kutschera, 1864 

flavicornis (Stephens, 1831) 

tabidus sensu auctt. partim non (Fabricius, 1775) 

jacobaeae sensu auctt. partim non (Waterhouse, 

G.R., 1858) 

rufescens Fowler, 1890 

fowleri Allen, 1967 

abdominalis sensu Fowler, 1890 non (Duftschmid, 

1825) 

ganglbaueri Heikertinger, 1912 

piciceps sensu auctt. non (Stephens, 1831) 

senecionis Brisout de Barneville, 1873 non 


gracilis Kutschera, 1864 

poweri (Allard, 1866) 

holsaticus (Linnaeus, 1758) 

jacobaeae (Waterhouse, G.R., 1858) 

tabidus sensu auctt. partim non (Fabricius, 1775) 

kutscherae (Rye, 1872) 

atriceps Kutschera, 1863 non (Stephens, 1831) 

longiseta Weise, 1889 

clarus Allen, 1967 694 

luridus (Scopoli, 1763) 

94 

brunneus sensu auctt. non (Duftschmid, 1825) 

castaneus (Duftschmid, 1825) 

fusculus sensu auctt. Brit. non Kutschera, 1864 

lycopi (Foudras, 1860) 

abdominalis sensu (Allard, 1860) non (Duftschmid, 

1825) 

melanocephalus (De Geer, 1775) 

piciceps (Stephens, 1831) 

membranaceus (Foudras, 1860) 

cerinus sensu auctt. Brit. partim non (Foudras, 

1860) 

teucrii (Allard, 1860) 

nasturtii (Fabricius, 1792) 

nigerrimus (Gyllenhal, 1827) 

nigrofasciatus (Goeze, 1777) 

patruelis (Allard, 1866) 

distinguendus (Rye, 1872) 

obliteratoides Gruev, 1973 695 

obliteratus (Rosenhauer, 1847) 

pulex sensu Fowler, 1890 non (Schrank, 1781) 

ochroleucus (Marsham, 1802) 

parvulus (Paykull, 1799) 

pumilus (Illiger, 1807) 

ater sensu Fowler, 1890 non (Fabricius, 1775) 

pellucidus (Foudras, 1860) 

medicaginis (Allard, 1860) 

plantagomaritimus Dollman, 1912 

pratensis (Panzer, 1794) 

pusillus (Gyllenhal, 1813) 

collaris (Stephens, 1831) 

bearei Kevan, 1967 696 

quadriguttatus (Pontoppidan, 1763) 

reichei (Allard, 1860) 

fusculus Kutschera, 1864 

rubiginosus (Foudras, 1860) 

flavicornis sensu (Allard, 1860) non (Stephens, 

1831) 

rutilus (Illiger, 1807) 

succineus (Foudras, 1860) 

laevis sensu (Allard, 1860) non (Duftschmid, 1825) 

suturellus (Duftschmid, 1825) 

thoracicus (Stephens, 1831) 

symphyti Heikertinger, 1912 697 

tabidus (Fabricius, 1775) 

ALTICA Geoffroy, 1762 698 

HALTICA Illiger, 1801 40 

brevicollis Foudras, 1860 

coryli (Allard, 1860) 

carinthiaca Weise, 1888 699 

helianthemi (Allard, 1859) 700 

pusilla (Duftschmid, 1825) non (Müller, O.F., 1776) 

longicollis (Allard, 1860) 701 

ericeti sensu auctt. Brit. non (Allard, 1859) 

britteni Sharp, 1914 702 

sandini Kemner, 1919 

lythri Aubé, 1843 

tamaricis sensu auctt. Brit. partim non Schrank, 

1785 

oleracea (Linnaeus, 1758) 

ampelophaga sensu auctt. Brit. non Guérin- 

Méneville, 1858 


palustris Weise, 1888 

HERMAEOPHAGA Foudras, 1860 

mercurialis (Fabricius, 1792)

BATOPHILA Foudras, 1860 

aerata (Marsham, 1802) 

rubi (Paykull, 1799) 

LYTHRARIA Bedel, 1897 

OCHROSIS sensu Fowler, 1890 non Foudras, 1860 

salicariae (Paykull, 1800) 

OCHROSIS Foudras, 1860 

CREPIDODERA sensu Fowler, 1890 partim non 

Dejean, 1836 

ventralis (Illiger, 1807) 

NEOCREPIDODERA Heikertinger, 1911 

CREPIDODERA sensu auctt. non Dejean, 1836 

ASIORESTIA Jacobson, 1925 

ferruginea (Scopoli, 1763) 

impressa (Fabricius, 1801) 

transversa (Marsham, 1802) 

DEROCREPIS Weise, 1886 

CREPIDODERA sensu Fowler, 1890 partim non 

Dejean, 1836 


HIPPURIPHILA Foudras, 1860 

modeeri (Linnaeus, 1761) 

CREPIDODERA Dejean, 1836 

CHALCOIDES Foudras, 1860 

aurata (Marsham, 1802) 703 

aurea (Fourcroy, 1785) 

helxines sensu auctt. non (Linnaeus, 1758) 

fulvicornis (Fabricius, 1792) 

smaragdina Foudras, 1860 

nitidula (Linnaeus, 1758) 

plutus (Latreille, 1804) 

chloris Foudras, 1860 

EPITRIX Foudras, 1860 

EPITHRIX Foudras, 1860 

atropae Foudras, 1860 

pubescens (Koch, J.D.W., 1803) 

PODAGRICA Dejean, 1836 

fuscicornis (Linnaeus, 1767) 

fuscipes (Fabricius, 1775) 

MANTURA Stephens, 1831 

chrysanthemi (Koch, J.D.W., 1803) 

matthewsii (Curtis, 1833) 

obtusata (Gyllenhal, 1813) 

rustica (Linnaeus, 1767) 

CHAETOCNEMA Stephens, 1831 704 

PLECTROSCELIS Dejean, 1836 

aerosa (Letzner, 1847) 

arida Foudras, 1860 

aridula sensu auctt. Brit. partim non (Gyllenhal, 

1827) 

concinna (Marsham, 1802) 

confusa (Boheman, 1851) 

hortensis (Fourcroy, 1785) 

picipes Stephens, 1831 705 

95 


concinna sensu auctt. partim non (Marsham, 1802) 

laevicollis (Thomson, C.G., 1866) 706 

heikertingeri Lubischev, 1963 706 

sahlbergii (Gyllenhal, 1827) 

subcoerulea (Kutschera, 1864) 

SPHAERODERMA Stephens, 1831 

rubidum (Graëlls, 1858) 

testaceum sensu auctt. partim non Fabricius, 1775 

testaceum (Fabricius, 1775) 

cardui (Gyllenhal, 1813) 

APTEROPEDA Dejean, 1836 

globosa (Illiger, 1794) 

orbiculata (Marsham, 1802) 

splendida Allard, 1860 

MNIOPHILA Stephens, 1831 

muscorum (Koch, J.D.W., 1803) 

DIBOLIA Latreille, 1829 

cynoglossi (Koch, J.D.W., 1803) 

PSYLLIODES Latreille, 1829 


attenuata (Koch, J.D.W., 1803) 

chalcomera (Illiger, 1807) 

chrysocephala (Linnaeus, 1758) 

anglica (Fabricius, 1775) 

luridipennis sensu auctt. partim non Kutschera, 

1864 

cucullata (Illiger, 1807) 707 

cuprea (Koch, J.D.W., 1803) 

instabilis sensu auctt. Brit. non Foudras, 1860 

dulcamarae (Koch, J.D.W., 1803) 

hyoscyami (Linnaeus, 1758) 

laticollis Kutschera, 1860 

weberi Lohse, 1955 708 

luridipennis Kutschera, 1864 

hospes sensu auctt. non Wollaston, 1854 

luteola (Müller, O.F., 1776) 

marcida (Illiger, 1807) 

napi (Fabricius, 1792) 

picina (Marsham, 1802) 

sophiae Heikertinger, 1914 

cyanoptera sensu auctt. non (Illiger, 1807) 

Subfamily CASSIDINAE Gyllenhal, 1813 

PILEMOSTOMA Desbrochers, 1891 

CASSIDA sensu Fowler, 1890 partim non Linnaeus, 

1758 

fastuosa (Schaller, 1783) 

HYPOCASSIDA Weise, 1893 

subferruginea (Schrank, 1776) 

CASSIDA Linnaeus, 1758 

denticollis Suffrian, 1844 

chloris sensu auctt. Brit. non Suffrian, 1844 

flaveola Thunberg, 1794 

hemisphaerica Herbst, 1799 

murraea Linnaeus, 1767 

maculata Linnaeus, 1767

Family NEMONYCHIDAE Bedel, 1882 Checklist of Beetles of the British Isles, 2012 Edition 

nebulosa Linnaeus, 1758 

nobilis Linnaeus, 1758 

prasina Illiger, 1798 

sanguinolenta sensu auctt. Brit. non Müller, O.F., 

1776 

chloris Suffrian, 1844 

rubiginosa Müller, O.F., 1776 

viridis sensu Scopoli, 1763 non Linnaeus, 1758 

sanguinosa Suffrian, 1844 

vibex Linnaeus, 1767 

viridis Linnaeus, 1758 

equestris Fabricius, 1787 

vittata de Villers, 1789 

Superfamily CURCULIONOIDEA Latreille, 

1802 709 

Division ORTHOCERI Schönherr, 1823 710 

93. Family NEMONYCHIDAE Bedel, 1882 711 

Family author: M.G. Morris 

Subfamily CIMBERIDINAE des Gozis, 1882 712 

CIMBERIS des Gozis, 1881 

RHINOMACER Fabricius, 1781 713 

attelaboides (Fabricius, 1787) 

94. Family ANTHRIBIDAE Billberg, 1820 714 


Subfamily ANTHRIBINAE Billberg, 1820 

Tribe ANTHRIBINI Billberg, 1820 

ANTHRIBUS Geoffroy, 1762 715 

BRACHYTARSUS Schönherr, 1823 

fasciatus Forster, 1770 

nebulosus Forster, 1770 

variegatus Fourcroy, 1785 


Tribe PLATYRHININI Bedel, 1882 

PLATYRHINUS Clairville, 1798 

PLATYRRHINUS Fowler, 1891 (misspelling) 

resinosus (Scopoli, 1763) 

latirostris (Fabricius, 1775) 

Tribe PLATYSTOMINI Pierce, 1916 

PLATYSTOMOS Schneider, D.H., 1791 715 

PLATYSTOMUS auctt. (misspelling) 

ANTHRIBUS sensu auctt. non Geoffroy, 1762 

MACROCEPHALUS Olivier, 1789 non Swederus, 

1787 

albinus (Linnaeus, 1758) 

96 

Tribe DISCOPTENINI Lacordaire, 1865 

PSEUDEUPARIUS Jordan, 1914 716 

TROPIDERES sensu auctt. non Schönherr, 1823 

sepicola (Fabricius, 1792) 

Tribe ZYGAENODINI Lacordaire, 1865 

DISSOLEUCAS Jordan, 1925 717 

niveirostris (Fabricius, 1798) 

Subfamily CHORAGINAE Kirby, 1819 

Tribe CHORAGINI Kirby, 1819 

CHORAGUS Kirby, 1819 

sheppardi Kirby, 1819 

Tribe ARAECERINI Lacordaire, 1865 

ARAECERUS Schönherr, 1823 

ARAEOCERUS auctt. (misspelling) 

fasciculatus (De Geer, 1775) 

Subfamily URODONTINAE Thomson, C.G., 

1859 718 

BRUCHELA Dejean, 1821 

BRUCHELLA auctt. (misspelling) 

URODON Schönherr, 1823 

rufipes (Olivier, 1790) 719 

95. Family RHYNCHITIDAE Gistel, 1848 


Subfamily RHYNCHITINAE Gistel, 1848 

Tribe RHYNCHITINI Gistel, 1848 

INVOLVULUS Schrank, 1798 715 

caeruleus (De Geer, 1775) 

cupreus (Linnaeus, 1758) 

LASIORHYNCHITES Jekel, 1860 720 

RHYNCHITES sensu auctt. partim non Schneider, 

D.H., 1791 

cavifrons (Gyllenhal, 1833) 

pubescens sensu auctt. non (Fabricius, 1775) 

olivaceus (Gyllenhal, 1833) 

ophthalmicus sensu auctt. Brit. non (Stephens, 

1831) 

sericeus sensu (Fowler, 1890) non (Herbst, 1797)

NEOCOENORRHINUS Voss, 1951 

RHYNCHITES sensu auctt. partim non Schneider, 

D.H., 1791 

CAENORHINUS sensu auctt. non Thomson, C.G., 

1859 721 

COENORRHINUS Voss, 1932 

germanicus (Herbst, 1797) 

minutus sensu (Thomson, C.G., 1866) non (Herbst, 

1797) 

interpunctatus (Stephens, 1831) 

minutus (Herbst, 1797) 

aeneovirens (Marsham, 1802) 

pauxillus (Germar, 1824) 

RHYNCHITES Schneider, D.H., 1791 

auratus (Scopoli, 1763) 

bacchus (Linnaeus, 1758) 

TATIANAERHYNCHITES Legalov, 2002 

NEOCOENORRHINUS sensu auctt. partim non 

Voss, 1951 

aequatus (Linnaeus, 1767) 

TEMNOCERUS Thunberg, 1815 715 

PSELAPHORHYNCHITES Schilsky, 1903 

coeruleus (Fabricius, 1798) 

tomentosus (Gyllenhal, 1839) 

uncinatus (Thomson, C.G., 1865) 

longiceps (Thomson, C.G., 1888) 

harwoodi (Joy, 1911) 

nanus (Paykull, 1792) 

Tribe BYCTISCINI Voss, 1923 

BYCTISCUS Thomson, C.G., 1859 

betulae (Linnaeus, 1758) 

betuleti (Fabricius, 1792) 

populi (Linnaeus, 1758) 

Tribe DEPORAINI Voss, 1929 

CAENORHINUS Thomson, C.G., 1859 722 

mannerheimii (Hummel, 1823) 

mannerheimi auctt. (misspelling) 

megacephalus (Germar, 1824) 

DEPORAUS Samouelle, 1819 

betulae (Linnaeus, 1758) 

96. Family ATTELABIDAE Billberg, 1820 


Subfamily ATTELABINAE Billberg, 1820 

ATTELABUS Linnaeus, 1758 

nitens (Scopoli, 1763) 

97 

curculionoides Linnaeus, 1763 

Family ATTELABIDAE Billberg, 1820 

Subfamily APODERINAE Jekel, 1860 

APODERUS Olivier, 1807 

coryli (Linnaeus, 1758) 

97. Family APIONIDAE Schönherr, 1823 723 


Subfamily APIONINAE Schönherr, 1823 

Tribe APIONINI Schönherr, 1823 

APION Herbst, 1797 

ERYTHRAPION Schilsky, 1906 

cruentatum Walton, 1844 

sanguineum sensu Stephens, 1839 non (De Geer, 

1775) 

desideratum Sharp, 1918 

frumentarium (Linnaeus, 1758) 

miniatum Germar, 1833 724 

haematodes Kirby, 1808 

frumentarium sensu (Paykull, 1792) non (Linnaeus, 

1758) 

brachypterum Sharp, 1918 

fraudator Sharp, 1918 

rubens Stephens, 1839 

rubiginosum Grill, 1893 

sanguineum (De Geer, 1775) 

Tribe APLEMONINI Kissinger, 1968 

AIZOBIUS Alonso-Zarazaga, 1991 

APION sensu auctt. partim non Herbst, 1797 

sedi (Germar, 1818) 

HELIANTHEMAPION Wagner, 1930 


aciculare (Germar, 1817) 725 

PERAPION Wagner, 1907 


Subgenus PERAPION Wagner, 1907 

affine (Kirby, 1808) 

curtirostre (Germar, 1817) 

brevirostre sensu (Kirby, 1808) non (Herbst, 1797) 

humile (Germar, 1817) 

hydrolapathi (Marsham, 1802) 

marchicum (Herbst, 1797) 

violaceum (Kirby, 1808) 

Subgenus EROOSAPION Ehret, 1994 

lemoroi (Brisout de Barneville, 1880) 

PSEUDAPLEMONUS Wagner, 1930 


limonii (Kirby, 1808) 

PSEUDOPERAPION Wagner, 1930 

brevirostre (Herbst, 1797) 726

Family APIONIDAE Schönherr, 1823 Checklist of Beetles of the British Isles, 2012 Edition 

Tribe ASPIDAPIINI Alonso-Zarazaga, 1990 

ASPIDAPION Schilsky, 1901 


Subgenus ASPIDAPION Schilsky, 1901 

radiolus (Marsham, 1802) 

soror (Rey, 1895) 727 

foveatoscutellatum (Wagner, 1906) 

Subgenus KOESTLINIA Alonso-Zarazaga, 1990 

aeneum (Fabricius, 1775) 

Tribe CERATAPIINI Alonso-Zarazaga, 1990 

ACENTROTYPUS Alonso-Zarazaga, 1990 


brunnipes (Boheman, 1839) 

laevigatum (Kirby, 1808) non (Paykull, 1792) 

CERATAPION Schilsky, 1901 728 


Subgenus CERATAPION Schilsky, 1901 

armatum (Gerstaecker, 1854) 

carduorum (Kirby, 1808) 

dentirostre sensu auctt. Brit. non (Gerstaecker, 

1854) 

lacertense (Tottenham, 1941) 

gibbirostre (Gyllenhal, 1813) 

carduorum sensu auctt. non (Kirby, 1808) 

Subgenus ACANEPHODUS Alonso-Zarazaga, 1990 

onopordi (Kirby, 1808) 

DIPLAPION Reitter, 1916 


confluens (Kirby, 1808) 

stolidum (Germar, 1817) 

OMPHALAPION Schilsky, 1901 


beuthini (Hoffmann, Anton, 1874) 

dispar sensu auctt. Brit. non (Germar, 1817) 729 

hookerorum (Kirby, 1808) 

hookeri auctt. (misspelling) 730 

laevigatum (Paykull, 1792) 731 

sorbi (Fabricius, 1792) 

Tribe EXAPIINI Alonso-Zarazaga, 1990 

EXAPION Bedel, 1887 


ULAPION Ehret, 1997 

difficile (Herbst, 1797) 

kiesenwetteri (Desbrochers, 1870) 

fuscirostre (Fabricius, 1775) 

genistae (Kirby, 1811) 

ulicis (Forster, 1771) 

Tribe IXAPIINI Alonso-Zarazaga, 1990 

IXAPION Roudier & Tempère, 1973 


variegatum (Wencker, 1864) 732 

98 

Tribe KALCAPIINI Alonso-Zarazaga, 1990 

KALCAPION Schilsky, 1906 


pallipes (Kirby, 1808) 

semivittatum (Gyllenhal, 1833) 

MELANAPION Wagner, 1930 


minimum (Herbst, 1797) 

SQUAMAPION Bokor, 1923 


THYMAPION Deville, 1924 

atomarium (Kirby, 1808) 

minutissimum sensu auctt. Brit. non (Rosenhauer, 

1856) 

serpyllicola sensu auctt. Brit. non (Wencker, 1864) 

cineraceum (Wencker, 1864) 

millum sensu (Bach, 1854) non (Gyllenhal, 1833) 

annulipes sensu auctt. non (Wencker, 1864) 

flavimanum (Gyllenhal, 1833) 

vicinum (Kirby, 1808) 

TAENIAPION Schilsky, 1906 


urticarium (Herbst, 1784) 

Tribe MALVAPIINI Alonso-Zarazaga, 1990 

MALVAPION Hoffmann, Adolfe, 1958 


malvae (Fabricius, 1775) 

PSEUDAPION Schilsky, 1906 


rufirostre (Fabricius, 1775) 

RHOPALAPION Schilsky, 1906 


longirostre (Olivier, 1807) 733 

Tribe OXYSTOMATINI Alonso-Zarazaga, 1990 

Subtribe OXYSTOMATINA Alonso-Zarazaga, 1990 

CYANAPION Bokor, 1923 


Subgenus CYANAPION Bokor, 1923 

spencii (Kirby, 1808) 

Subgenus BOTHRYORRHYNCHAPION Bokor, 1923 

afer (Gyllenhal, 1833) 734 

afrum auctt. (misspelling) 

unicolor sensu auctt. non (Kirby, 1808) 

platalea sensu auctt. Brit. non (Germar, 1817) 

gyllenhalii (Kirby, 1808) 

gyllenhali auctt. (misspelling) 

unicolor (Kirby, 1808)

EUTRICHAPION Reitter, 1916 


Subgenus EUTRICHAPION Reitter, 1916 

ervi (Kirby, 1808) 

viciae (Paykull, 1800) 

Subgenus CNEMAPION Bokor, 1923 

vorax (Herbst, 1797) 

Subgenus PSILOCALYMMA Alonso-Zarazaga, 1990 

punctigerum (Paykull, 1792) 

HEMITRICHAPION Voss, 1959 


Subgenus TINOCYBA Alonso-Zarazaga, 1990 

reflexum (Gyllenhal, 1833) 

livescerum (Gyllenhal, 1833) 

waltoni (Stephens, 1839) 

HOLOTRICHAPION Györffy, 1956 


Subgenus HOLOTRICHAPION Györffy, 1956 

ononis (Kirby, 1808) 

Subgenus APIOPS Alonso-Zarazaga, 1990 

pisi (Fabricius, 1801) 

Subgenus LEGARICAPION Ehret, 1990 

aethiops (Herbst, 1797) 

OXYSTOMA Duméril, 1806 


cerdo (Gerstaecker, 1854) 

craccae (Linnaeus, 1767) 

pomonae (Fabricius, 1798) 

opeticum sensu auctt. Brit. non (Bach, 1854) 

subulatum (Kirby, 1808) 

PIRAPION Reitter, 1916 


immune (Kirby, 1808) 

Subtribe CATAPIINA Alonso-Zarazaga, 1990 

CATAPION Schilsky, 1906 


curtisii (Stephens, 1831) 735 

curtisi auctt. (misspelling) 

curtulum Desbrochers, 1870 

pubescens (Kirby, 1811) 

seniculus (Kirby, 1808) 

Subtribe SYNAPIINA Alonso-Zarazaga, 1990 

ISCHNOPTERAPION Bokor, 1923 


Subgenus ISCHNOPTERAPION Bokor, 1923 

loti (Kirby, 1808) 

modestum (Germar, 1817) 

sicardi (Desbrochers, 1893) 

Subgenus CHLORAPION Györffy, 1956 

virens (Herbst, 1797) 

99 

Family APIONIDAE Schönherr, 1823 

PROTOPIRAPION Alonso-Zarazaga, 1990 


atratulum (Germar, 1817) 

striatum (Marsham, 1802) non (Müller, O.F., 1776) 

STENOPTERAPION Bokor, 1923 


Subgenus STENOPTERAPION Bokor, 1923 

intermedium (Eppelsheim, 1875) 736 

meliloti (Kirby, 1808) 

tenue (Kirby, 1808) 

Subgenus COBOSIOTHERIUM Alonso-Zarazaga, 1990 

scutellare (Kirby, 1811) 

SYNAPION Schilsky, 1902 


ebeninum (Kirby, 1808) 

Subtribe TRICHAPIINA Alonso-Zarazaga, 1990 

BETULAPION Ehret, 1994 


TRICHAPION sensu Wagner, 1912 partim 

simile (Kirby, 1811) 

Tribe PIEZOTRACHELINI Voss, 1959 

PROTAPION Schilsky, 1908 


apricans (Herbst, 1797) 

assimile (Kirby, 1808) 

bohemani sensu (Bedel, 1885) non (Thomson, 

C.G., 1865) 

ssp. assimile (Kirby, 1808) 

ssp. ryei (Blackburn, 1874) 737 

difforme (Germar, 1818) 

dissimile (Germar, 1817) 

filirostre (Kirby, 1808) 

cantianum (Wagner, 1906) 

fulvipes (Geoffroy in Fourcroy, 1785) 

flavipes sensu (Paykull, 1792) non (Fabricius, 1775) 

dichroum (Bedel, 1886) 

laevicolle (Kirby, 1811) 

nigritarse (Kirby, 1808) 

ononidis (Gyllenhal, 1827) 

ononicola (Bach, 1854) 

bohemani (Thomson, C.G., 1865) 

schoenherri (Boheman, 1839) 

trifolii (Linnaeus, 1768) 

aestivum (Germar, 1817) 

varipes (Germar, 1817) 

PSEUDOPROTAPION Ehret, 1990 


astragali (Paykull, 1800)

Family NANOPHYIDAE Gistel, 1848 Checklist of Beetles of the British Isles, 2012 Edition 

98. Family NANOPHYIDAE Gistel, 1848 


DIECKMANNIELLUS Alonso-Zarazaga, 1989 

NANOPHYES sensu auctt. partim non Schönherr, 

1838 

gracilis (Redtenbacher, 1849) 

NANOPHYES Schönherr, 1838 

NANODES Schönherr, 1825 738 

marmoratus (Goeze, 1777) 

lythri (Fabricius, 1787) 

99. Family DRYOPHTHORIDAE Schönherr, 

1825 


Subfamily DRYOPHTHORINAE Schönherr, 1825 

DRYOPHTHORUS Germar, 1824 739 

corticalis (Paykull, 1792) 

Subfamily RHYNCHOPHORINAE Schönherr, 

1833 

Tribe LITOSOMINI Lacordaire, 1865 

SITOPHILUS Schönherr, 1838 

CALANDRA Schellenberg, 1798 740 

CALENDRA Clairville, 1798 741 

granarius (Linnaeus, 1758) 

oryzae (Linnaeus, 1763) 

zeamais Motschulsky, 1855 

100. Family ERIRHINIDAE Schönherr, 1825 742 


Subfamily ERIRHININAE Schönherr, 1825 

Tribe ERIRHININI Schönherr, 1825 

GRYPUS Germar, 1817 

GRYPIDIUS Schönherr, 1826 

equiseti (Fabricius, 1775) 

NOTARIS Germar, 1817 

ERIRHINUS Schönherr, 1825 743 

acridulus (Linnaeus, 1758) 

aethiops (Fabricius, 1792) 

scirpi (Fabricius, 1792) 744 

rotundicollis (Motschulsky, 1860) 

PROCAS Stephens, 1831 745 

granulicollis Walton, 1848 

picipes (Marsham, 1802) 

armillatus sensu auctt. non (Fabricius, 1801) 

100 

THRYOGENES Bedel, 1884 

ERIRHINUS sensu auctt. non Schönherr, 1825 

festucae (Herbst, 1795) 

fiorii Zumpt, 1928 746 

atrirostris Lohse, 1991 

nereis (Paykull, 1800) 

scirrhosus (Gyllenhal, 1836) 

TOURNOTARIS Alonso-Zarazaga & Lyal, 1999 

NOTARIS sensu auctt. partim non Germar, 1817 

bimaculata (Fabricius, 1787) 

Tribe STENOPELMINI LeConte, 1876 

STENOPELMUS Schönherr, 1836 

rufinasus Gyllenhal, 1836 

Tribe TANYSPHYRINI Gistel, 1848 

TANYSPHYRUS Germar, 1817 

lemnae (Paykull, 1792) 

101. Family RAYMONDIONYMIDAE Reitter, 

1913 747 


FERRERIA Alonso-Zarazaga & Lyal, 1999 

RAYMONDIA Aubé, 1861 non Frauenfeld, 1856 

RAYMONDIONYMUS Ganglbauer, 1906 non 

Wollaston, 1873 

marqueti (Aubé, 1863) 748 

Division GONATOCERI Schönherr, 1825 

102. Family CURCULIONIDAE Latreille, 1802 


Subfamily CURCULIONINAE Latreille, 1802 

Tribe CURCULIONINI Latreille, 1802 

ARCHARIUS Gistel, 1848 

CURCULIO sensu auctt. partim non Linnaeus, 1758 

BALANOBIUS Jekel, 1861 

pyrrhoceras (Marsham, 1802) 

salicivorus (Paykull, 1792) 

CURCULIO Linnaeus, 1758 

BALANINUS Germar, 1817 

betulae (Stephens, 1831) 

cerasorum sensu (Paykull, 1792) non Fabricius, 

1775 

glandium Marsham, 1802 

turbatus (Gyllenhal, 1836) 

nucum Linnaeus, 1758 

rubidus (Gyllenhal, 1836) 

venosus (Gravenhorst, 1807) 


cerasorum (Fabricius, 1775)

Tribe ACALYPTINI Thomson, C.G., 1859 

ACALYPTUS Schönherr, 1833 

carpini (Fabricius, 1792) 

Tribe ANTHONOMINI Thomson, C.G., 1859 

ANTHONOMUS Germar, 1817 749 

Subgenus ANTHONOMUS Germar, 1817 

bituberculatus Thomson, C.G., 1868 

fasciatus (Marsham, 1802) non (Scopoli, 1763) 

rosinae des Gozis, 1882 

brunnipennis (Curtis, 1840) 

comari Crotch, 1869 

chevrolati Desbrochers, 1868 

conspersus Desbrochers, 1868 

humeralis (Panzer, 1795) 

incurvus (Panzer, 1795) 


ulmi sensu auctt. partim non (De Geer, 1775) 

piri Kollar, 1837 

cinctus Redtenbacher, 1858 

pomorum (Linnaeus, 1758) 

rubi (Herbst, 1795) 

rufus Gyllenhal, 1836 

ulmi (De Geer, 1775) 

inversus Bedel, 1884 

Subgenus ANTHOMORPHUS Wiese, 1883 

PARANTHONOMUS Dietz, 1891 

varians (Paykull, 1792) 

phyllocola (Herbst, 1795) 

Subgenus FURCIPUS Desbrochers, 1868 750 

rectirostris (Linnaeus, 1758) 751 

BRACHONYX Schönherr, 1825 

pineti (Paykull, 1792) 

BRADYBATUS Germar, 1824 

fallax Gerstaecker, 1860 752 

Tribe CIONINI Schönherr, 1825 

CIONUS Clairville, 1798 

alauda (Herbst, 1784) 

blattariae (Fourcroy, 1785) 

hortulanus (Fourcroy, 1785) 

longicollis Brisout de Barneville, 1863 

nigritarsis Reitter, 1904 

thapsus sensu auctt. Brit. non (Fabricius, 1792) 

scrophulariae (Linnaeus, 1758) 

woodi Donisthorpe, 1921 

tuberculosus (Scopoli, 1763) 

verbasci (Fabricius, 1787) 

CLEOPUS Dejean, 1821 

CIONUS sensu Fowler, 1891 partim non Clairville, 

1798 

pulchellus (Herbst, 1795) 

101 

Family CURCULIONIDAE Latreille, 1802 

Tribe ELLESCINI Thomson, C.G., 1859 

Subtribe ELLESCINA Thomson, C.G., 1859 

ELLESCUS Dejean, 1821 753 

ELLESCHUS auctt. (misspelling) 

bipunctatus (Linnaeus, 1758) 

Subtribe DORYTOMINA Bedel, 1886 

DORYTOMUS Germar, 1817 754 

ETEOPHILUS Bedel, 1886 

affinis (Paykull, 1800) 755 

edoughensis Desbrochers, 1875 

dejeani Faust, 1882 

costirostris (Gyllenhal, 1836) non Sahlberg, C.R., 

1835 

filirostris (Gyllenhal, 1836) 

hirtipennis Bedel, 1884 

ictor (Herbst, 1795) 

validirostris (Gyllenhal, 1836) 756 

longimanus (Forster, 1771) 

vorax (Fabricius, 1792) 

majalis (Paykull, 1792) 

melanophthalmus (Paykull, 1792) 

agnathus (Boheman, 1843) 

rufatus (Bedel, 1888) 

pectoralis sensu Gyllenhal, 1843 non (Panzer, 

1795) 

rufulus Bedel, 1884 non Mannerheim, 1853 

salicinus (Gyllenhal, 1827) 

salicis Walton, 1851 

taeniatus (Fabricius, 1781) 

maculatus (Marsham, 1802) 

silbermanni Wencker, 1866 

tortrix (Linnaeus, 1761) 

tremulae (Fabricius, 1787) 

Tribe MECININI Gistel, 1848 757 

CLEOPOMIARUS Pierce, 1919 

MIARUS sensu auctt. partim non Schönherr, 1826 

graminis (Gyllenhal, 1813) 

degorsi sensu auctt. ?non Abeille de Perrin, 

1906 758 

micros (Germar, 1821) 

plantarum (Germar, 1824) 

GYMNETRON Schönherr, 1825 759 

GYMNAETRON auctt. (misspelling) 

beccabungae (Linnaeus, 1761) 

squamicolle Reitter, 1907 

melanarium (Germar, 1821) 

rostellum (Herbst, 1795) 

veronicae (Germar, 1821) 

nigrum Hardy & Bold, 1852 

beccabungae sensu auctt. Brit. ante 1931 non 


villosulum Gyllenhal, 1838 

MECINUS Germar, 1821 

circulatus (Marsham, 1802) 

collaris Germar, 1821 

janthinus Germar, 1821

Family CURCULIONIDAE Latreille, 1802 Checklist of Beetles of the British Isles, 2012 Edition 

labilis (Herbst, 1795) 760 

pascuorum (Gyllenhal, 1813) 760 

marshalli Donisthorpe, 1943 

melas sensu Donisthorpe, 1948 non Boheman, 

1838 

pyraster (Herbst, 1795) 

MIARUS Schönherr, 1826 

campanulae (Linnaeus, 1767) 

RHINUSA Stephens, 1829 

GYMNETRON sensu auctt. partim non Schönherr, 

1825 

antirrhini (Paykull, 1800) 

collina (Gyllenhal, 1813) 

linariae (Panzer, 1795) 

Tribe RHAMPHINI Rafinesque, 1815 761 

ISOCHNUS Thomson, C.G., 1859 

RHYNCHAENUS sensu auctt. partim non Clairville, 

1798 

foliorum (Müller, O.F., 1764) 

saliceti (Paykull, 1792) 

sequensi (Stierlin, 1894) 762 

populi (Fabricius, 1792) non (Linnaeus, 1758) 

populicola (Silfverberg, 1977) 

ORCHESTES Illiger, 1798 


1798 

Subgenus ORCHESTES Illiger, 1798 


saltator (Fourcroy, 1785) 

ferrugineus (Marsham, 1802) 

calceatus (Germar, 1821) 763 

scutellaris (Fabricius, 1801) 32 

iota (Fabricius, 1787) 

pilosus (Fabricius, 1781) 

ilicis (Fabricius, 1787) 

sparsus sensu auctt. Brit. partim non (Fåhraeus, 

1843) 

quercus (Linnaeus, 1758) 

rusci (Herbst, 1795) 

signifer (Creutzer, 1799) 

avellanae (Donovan, 1797) non (Paykull, 1792) 764 

erythropus sensu auctt. Brit. non (Germar, 1821) 765 

sparsus sensu auctt. Brit. partim non (Fåhraeus, 

1843) 

testaceus (Müller, O.F., 1776) 

alni sensu (Hoffmann, 1958) non (Linnaeus, 1758) 

Subgenus SALIUS Schrank, 1798 

EUTHORON Thomson, C.G., 1859 

fagi (Linnaeus, 1758) 

PSEUDORCHESTES Bedel, 1894 


1798 

pratensis (Germar, 1821) 

RHAMPHUS Clairville, 1798 

RAMPHUS auctt. (misspelling) 

oxyacanthae (Marsham, 1802) 

pulicarius (Herbst, 1795) 

102 

flavicornis Clairville, 1798 

subaeneus Illiger, 1807 766 

RHYNCHAENUS Clairville, 1798 

lonicerae (Herbst, 1795) 767 

xylostei Clairville, 1798 

TACHYERGES Schönherr, 1825 


1798 

decoratus (Germar, 1821) 

pseudostigma (Tempère, 1982) 768 

salicis (Linnaeus, 1758) 

stigma (Germar, 1821) 

Tribe SMICRONYCHINI Seidlitz, 1891 

SMICRONYX Schönherr, 1843 

coecus (Reich, 1797) 

jungermanniae (Reich, 1797) 

reichi (Gyllenhal, 1836) 

seriepilosus Tournier, 1874 

Tribe STOREINI Lacordaire, 1863 

PACHYTYCHIUS Jekel, 1861 

haematocephalus (Gyllenhal, 1836) 

Tribe STYPHLINI Jekel, 1861 

ORTHOCHAETES Germar, 1824 

insignis (Aubé, 1863) 

setiger (Beck, 1817) 

PSEUDOSTYPHLUS Tournier, 1874 

pillumus (Gyllenhal, 1836) 

pilumnus auctt. (misspelling) 

Tribe TYCHIINI Gistel, 1848 

SIBINIA Germar, 1817 769 

arenariae Stephens, 1831 

primita (Herbst, 1795) 770 

signata (Gyllenhal, 1813) ? Panzer, 1795 

plantaginis sensu auctt. Brit. non Eppelsheim, 1875 

pyrrhodactyla (Marsham, 1802) 

potentillae Germar, 1824 771 

lloydi (Donisthorpe, 1929) 772 

sodalis Germar, 1824 

TYCHIUS Germar, 1817 

MICCOTROGUS Schönherr, 1825 773 

breviusculus Desbrochers, 1873 774 

haematopus sensu auctt. non Gyllenhal, 1836 

crassirostris Kirsch, 1871 775 

junceus (Reich, 1797) 

flavicollis Stephens, 1831 

haematopus Gyllenhal, 1836 

lineatulus Stephens, 1831 

meliloti Stephens, 1831 

parallelus (Panzer, 1794) 

venustus sensu auctt. non (Fabricius, 1781)

picirostris (Fabricius, 1787) 

cinerascens (Marsham, 1802) 

posticus (Gyllenhal, 1836) 

polylineatus (Germar, 1824) 

pusillus Germar, 1842 

pygmaeus Brisout de Barneville, H., 1860 

quinquepunctatus (Linnaeus, 1758) 

schneideri (Herbst, 1795) 

squamulatus Gyllenhal, 1836 

flavicollis sensu auctt. non Stephens, 1831 776 

stephensi Gyllenhal, 1836 

tomentosus (Herbst, 1795) non (Olivier, 1790) 

tibialis Boheman, 1843 

Subfamily BAGOINAE Thomson, C.G., 1859 

BAGOUS Germar, 1817 777 

Subgenus BAGOUS Germar, 1817 

PROBAGOUS Sharp, 1917 

argillaceus Gyllenhal, 1836 

binodulus (Herbst, 1795) 

brevis Gyllenhal, 1836 

collignensis (Herbst, 1797) 

frit sensu auctt. partim non (Herbst, 1795) 

claudicans Boheman, 1845 

czwalinae Seidlitz, 1891 

czwalinai auctt. (misspelling) 

heasleri Newbery, 1902 

diglyptus Boheman, 1845 

frit (Herbst, 1795) 

limosus (Gyllenhal, 1827) 

longitarsis Thomson, C.G., 1868 

arduus Sharp, 1917 778 

tomlini Sharp, 1917 

lutulosus (Gyllenhal, 1827) 

nodulosus Gyllenhal, 1836 

subcarinatus Gyllenhal, 1836 

frit sensu Newbery, 1902 non (Herbst, 1795) 

tempestivus (Herbst, 1795) 

cnemerythrus (Marsham, 1802) 

angustulus Thomson, C.G., 1870 

Subgenus ABAGOUS Sharp, 1917 

glabrirostris (Herbst, 1795) 

lutosus (Gyllenhal, 1813) 

lutulentus (Gyllenhal, 1813) 

binotatus Stephens, 1831 

nigritarsis Thomson, C.G., 1865 

puncticollis Boheman, 1854 

collignensis sensu auctt. partim non (Herbst, 1797) 

robustus Brisout de Barneville, H., 1863 

rudis Sharp, 1917 779 

Subgenus CYPRUS Schönherr, 1825 

tubulus Caldara & O'Brien, 1994 

cylindrus (Paykull, 1800) non (Fabricius, 1781) 

attenuatus (Ahrens, 1812) non (Fabricius, 1801) 

angustus Silfverberg, 1977 non (Tanner, 1954) 780 

Subgenus EPHIMEROPUS Hochhuth, 1847 

petro (Herbst, 1795) 

aubei (Cussac, 1851) 

Subgenus HYDRONOMUS Schönherr, 1825 

alismatis (Marsham, 1802) 

103 


Subfamily BARIDINAE Schönherr, 1836 781 

Tribe BARIDINI Schönherr, 1836 

AULACOBARIS Desbrochers, 1892 

BARIS sensu auctt. partim non Germar, 1817 

lepidii (Germar, 1824) 

picicornis (Marsham, 1802) 

BARIS Germar, 1817 782 

analis (Olivier, 1790) 

COSMOBARIS Casey, 1920 


scolopacea (Germar, 1818) 

MELANOBARIS Alonso-Zarazaga & Lyal, 1999 


laticollis (Marsham, 1802) 

Tribe APOSTASIMERINI Schönherr, 1844 

LIMNOBARIS Bedel, 1885 783 

dolorosa (Goeze, 1777) 

pilistriata (Stephens, 1831) 

t-album (Linnaeus, 1758) 

ssp. atriplicis (Fabricius, 1777) 

Subfamily CEUTORHYNCHINAE Gistel, 1848 784 

Tribe CEUTORHYNCHINI Gistel, 1848 

AMALORRHYNCHUS Reitter, 1913 

CEUTHORHYNCHIDIUS sensu Fowler, 1891 

partim non Jacquelin du Val, [1855] 

melanarius (Stephens, 1831) 

CALOSIRUS Thomson, C.G., 1859 

CEUTORHYNCHUS sensu auctt. partim non 

Germar, 1824 

terminatus (Herbst, 1795) 

CEUTORHYNCHUS Germar, 1824 785 

CEUTHORHYNCHUS Schönherr, 1837 40 

CEUTHORRHYNCHUS Fowler, 1891 (misspelling) 

CEUTHORHYNCHIDIUS sensu Fowler, 1891 

partim non Jacquelin du Val, [1855] 

alliariae Brisout de Barneville, H., 1860 

roberti sensu auctt. Brit. non Gyllenhal, 1837 

assimilis (Paykull, 1792) 786 

pleurostigma (Marsham, 1802) 787 

atomus Boheman, 1845 

setosus Boheman, 1845 

cakilis (Hansen, 1917) 788 

chalybaeus Germar, 1824 

timidus Weise, 1883 

moguntiacus Schultze, 1895 

cochleariae (Gyllenhal, 1813) 

constrictus (Marsham, 1802) 

contractus (Marsham, 1802) 789 

minutus (Reich, 1797) non (Drury, 1773) 

erysimi (Fabricius, 1787)


hepaticus Gyllenhal, 1837 

hirtulus Germar, 1824 

insularis Dieckmann, 1971 

contractus var. pallipes Crotch, 1865 

obstrictus (Marsham, 1802) 

assimilis sensu auctt. non (Paykull, 1792) 787 

pallidactylus (Marsham, 1802) 

quadridens (Panzer, 1795) non (Fabricius, 1775) 

parvulus Brisout de Barneville, Ch., 1869 

pectoralis Weise, 1895 

chalybeus sensu auctt. Brit. ? Germar, 1824 

pervicax Weise, 1883 

suturellus sensu auctt. non Gyllenhal, 1837 

picitarsis Gyllenhal, 1837 

pulvinatus Gyllenhal, 1837 

pumilio (Gyllenhal, 1827) 

posthumus sensu auctt. Brit. non Germar, 1824 

pyrrhorhynchus (Marsham, 1802) 

querceti (Gyllenhal, 1813) 

rapae Gyllenhal, 1837 

resedae (Marsham, 1802) 

sulcicollis (Paykull, 1800) 

cyanipennis Germar, 1824 

syrites Germar, 1824 

thomsoni Kolbe, 1900 

moguntiacus sensu auctt. non Schultze, 1895 

turbatus Schultze, 1903 

typhae (Herbst, 1795) 

floralis (Paykull, 1792) non (Olivier, 1790) 790 

palustris (Edmonds, T., 1930) 

unguicularis Thomson, C.G., 1871 

curvirostris Schultze, 1898 

COELIODES Schönherr, 1837 

rana (Fabricius, 1787) 

quercus sensu (Fabricius, 1787) non (Linnaeus, 

1758) 

dryados (Gmelin in Linnaeus, 1790) 790 

ruber (Marsham, 1802) 

transversealbofasciatus (Goeze, 1777) 

cinctus (Fourcroy, 1785) non (Drury, 1772) 

erythroleucos (Gmelin in Linnaeus, 1790) 790 

erythroleucus Fowler, 1891 (misspelling) 

COELIODINUS Dieckmann, 1972 791 

COELIODES sensu auctt. partim non Schönherr, 

1837 

nigritarsis (Hartmann, 1895) 

rubicundus (Herbst, 1795) 

DATONYCHUS Wagner, 1944 


Germar, 1824 

angulosus (Boheman, 1845) 

arquatus (Herbst, 1795) 792 

arquata (Herbst, 1795) (incorrect original spelling) 

melanostictus (Marsham, 1802) 

urticae (Boheman, 1845) 

DRUPENATUS Reitter, 1913 

DRUSENATUS Reitter, 1916 

POOPHAGUS sensu Fowler, 1891 partim non 

Schönherr, 1837 

nasturtii (Germar, 1824) 

104 

ETHELCUS Reitter, 1916 


Germar, 1824 

verrucatus (Gyllenhal, 1837) 

GLOCIANUS Reitter, 1916 


Germar, 1824 

distinctus (Brisout de Barneville, Ch., 1870) 

marginatus (Paykull, 1792) non (Fabricius, 1775) 

simillimus Edwards, J., 1911 

moelleri (Thomson, C.G., 1868) 

pilosellus (Gyllenhal, 1837) 

punctiger (Gyllenhal, 1837) 

HADROPLONTUS Thomson, C.G., 1859 


Germar, 1824 

litura (Fabricius, 1775) 

trimaculatus (Fabricius, 1775) 

MICRELUS Thomson, C.G., 1859 

CEUTHORRHYNCHUS sensu Fowler, 1891 partim 

non Germar, 1824 

ericae (Gyllenhal, 1813) 

MICROPLONTUS Wagner, 1944 


Germar, 1824 

campestris (Gyllenhal, 1837) 

chrysanthemi sensu auctt. Brit. non auctt. Europ. 

nec (Germar, 1824) 

rugulosus (Herbst, 1795) 

cinereus (Marsham, 1802) 

melanostigma (Marsham, 1802) 

uniguttatus (Marsham, 1802) 

triangulum (Boheman, 1845) 

MOGULONES Reitter, 1916 


Germar, 1824 

BORAGINOBIUS Wagner, 1944 

asperifoliarum (Gyllenhal, 1813) 

euphorbiae (Brisout de Barneville, Ch., 1866) 

geographicus (Goeze, 1777) 

NEDYUS Schönherr, 1825 

CIDNORHINUS Thomson, C.G., 1859 

CIDNORRHINUS auctt. (misspelling) 

COELIODES sensu Fowler, 1891 partim non 



urticae (Marsham, 1802) 

PARETHELCUS Dieckmann, 1972 


Germar, 1824 

pollinarius (Forster, 1771) 

POOPHAGUS Schönherr, 1837 

sisymbrii (Fabricius, 1777)

SIROCALODES Voss, 1958 


Germar, 1824 

depressicollis (Gyllenhal, 1813) 

nigrinus (Marsham, 1802) non (Herbst, 1795) 

mixtus (Mulsant & Rey, 1858) 

quercicola (Paykull, 1792) 

STENOCARUS Thomson, C.G., 1859 



ruficornis (Stephens, 1831) 

cardui sensu auctt. Brit. non (Herbst, 1784) 

fuliginosus (Marsham, 1802) non (Gmelin in 


umbrinus (Gyllenhal, 1837) 793 

TAPEINOTUS Schönherr, 1826 

TAPINOTUS Schönherr, 1826 794 

sellatus (Fabricius, 1794) 

THAMIOCOLUS Thomson, C.G., 1859 


Germar, 1824 

viduatus (Gyllenhal, 1813) 

TRICHOSIROCALUS Colonnelli, 1979 

CEUTHORHYNCHIDIUS sensu auctt. non 

Jacquelin du Val, [1855] 

barnevillei (Grenier, 1866) 

chevrolati (Crotch, 1865) 32 

dawsoni (Brisout de Barneville, Ch., 1869) 

horridus (Panzer, 1801) 

rufulus (Dufour, 1851) 

thalhammeri (Schultze, 1906) 795 

troglodytes (Fabricius, 1787) 

ZACLADUS Reitter, 1913 

ALLODACTYLUS Weise, 1883 non Lataste & 

Rochebrune, 1876 



exiguus (Olivier, 1807) 

geranii (Paykull, 1800) 

affinis (Paykull, 1792) non (Schrank, 1781) 

Tribe AMALINI Wagner, 1936 

AMALUS Schönherr, 1825 

scortillum (Herbst, 1795) 

haemorrhous (Herbst, 1795) non (Gmelin in 


Tribe MONONYCHINI LeConte, 1876 

MONONYCHUS Germar, 1824 

punctumalbum (Herbst, 1784) 

pseudacori (Fabricius, 1792) non (Rossi, 1790) 

Tribe PHYTOBIINI Gistel, 1848 

EUBRYCHIUS Thomson, C.G., 1859 

velutus (Beck, 1817) 

105 


velatus (Germar, 1818) (misspelling) 

NEOPHYTOBIUS Wagner, 1936 

PHYTOBIUS sensu auctt. partim non Schönherr, 

1833 

muricatus (Brisout de Barneville, Ch., 1867) 

quadrinodosus (Gyllenhal, 1813) 

denticollis (Gyllenhal, 1837) 

PELENOMUS Thomson, C.G., 1859 

PHYTOBIUS sensu Dejean, 1835 non Schönherr, 

1833 

RHINONCUS sensu Fowler, 1891 partim non 


canaliculatus (Fåhraeus, 1843) 

comari (Herbst, 1795) 796 

olssoni (Israelson, 1972) 797 

quadricorniger (Colonnelli, 1986) 

quadricornis (Gyllenhal, 1813) non (Paykull, 1792) 

quadrituberculatus (Fabricius, 1787) 

waltoni (Boheman, 1843) 

zumpti (Wagner, 1939) 798 

PHYTOBIUS Schönherr, 1833 

LITODACTYLUS Redtenbacher, 1849 

leucogaster (Marsham, 1802) 

RHINONCUS Schönherr, 1825 

AMALORHINONCUS Wagner, 1936 

albicinctus Gyllenhal, 1837 

bruchoides (Herbst, 1784) 

castor (Fabricius, 1792) 

inconspectus (Herbst, 1795) 

gramineus (Fabricius, 1792) non (Gmelin in 


pericarpius (Linnaeus, 1758) 

perpendicularis (Reich, 1797) 

Tribe SCLEROPTERINI Schultze, 1902 

RUTIDOSOMA Stephens, 1831 

RHYTIDOSOMA auctt. (misspelling) 

RHYTIDOSOMUS Schönherr, 1837 40 

globulus (Herbst, 1795) 

Subfamily COSSONINAE Schönherr, 1825 

Tribe COSSONINI Schönherr, 1825 

COSSONUS Clairville, 1798 

linearis (Fabricius, 1775) 

planatus Bedel, 1881 

parallelepipedus (Herbst, 1795) 

ferrugineus sensu Fowler, 1891 ? Clairville, 1798 

RHOPALOMESITES Wollaston, 1873 

MESITES sensu auctt. non Schönherr, 1838 

tardyi (Curtis, 1825) 

tardii auctt. (misspelling)


Tribe DRYOTRIBINI LeConte, 1876 

COTASTER Motschulsky, 1851 

uncipes (Boheman, 1838) 799 

Tribe ONYCHOLIPINI Wollaston, 1873 

PSELACTUS Broun, 1886 

CODIOSOMA Bedel, 1885 non Kirby, W.F., 1874 

PHLOEOPHAGIA Aurivillius, 1924 

spadix (Herbst, 1795) 

PSEUDOPHLOEOPHAGUS Wollaston, 1873 

CAULOTRUPIS sensu auctt. non Wollaston, 1854 

CAULOTRYPIS Fowler, 1891 (misspelling) 

aeneopiceus (Boheman, 1845) 

STEREOCORYNES Wollaston, 1873 

RHYNCOLUS sensu auctt. Brit. partim non Germar, 

1817 

truncorum (Germar, 1824) 

Tribe PENTARTHRINI Lacordaire, 1865 

ALLOPENTARTHRUM Kuschel in Wibmer & O’Brien, 

1986 

elumbe (Boheman, 1838) 800 

EUOPHRYUM Broun, 1909 

confine (Broun, 1881) 

rufum (Broun, 1880) 

PENTARTHRUM Wollaston, 1854 

huttoni Wollaston, 1854 

Tribe PROECINI Voss, 1956 

CONARTHRUS Wollaston, 1873 

praeustus (Boheman in Schönherr, 1838) 801 

Tribe RHYNCOLINI Gistel, 1848 

Subtribe RHYNCHOLINA Gistel, 1848 

MACRORHYNCOLUS Wollaston, 1873 

littoralis (Broun, 1880) 802 

RHYNCOLUS Germar, 1817 

EREMOTES Wollaston, 1861 

ater (Linnaeus, 1758) 803 

chloropus sensu auctt. non (Linnaeus, 1758) 

Subtribe PHLOEOPHAGINA Voss, 1955 

PHLOEOPHAGUS Schönherr, 1838 804 

RHYNCOLUS sensu auctt. Brit. partim non Germar, 

1817 

lignarius (Marsham, 1802) 

106 

Subfamily CRYPTORHYNCHINAE Schönherr, 

1825 

Tribe CRYPTORHYNCHINI Schönherr, 1825 

Subtribe CRYPTORHYNCHINA Schönherr, 1825 

CRYPTORHYNCHUS Illiger, 1807 

CRYPTORHYNCHIDIUS Pierce, 1919 non 

Champion, 1914 

lapathi (Linnaeus, 1758) 

Subtribe TYLODINA Lacordaire, 1865 

ACALLES Schönherr, 1825 805 

misellus Boheman, 1844 

?parvulus Boheman, 1837 

turbatus sensu auctt. non Boheman, 1844 

ptinoides (Marsham, 1802) 

KYKLIOACALLES Stüben, 1999 

ACALLES sensu auctt. partim non Schönherr, 1825 

roboris (Curtis, 1835) 

Subfamily CYCLOMINAE Schönherr, 1826 

Tribe RHYTHIRRININI Lacordaire, 1863 

Subtribe GRONOPINA Bedel, 1884 

GRONOPS Schönherr, 1823 

inaequalis Boheman, 1842 806 


Subfamily ENTIMINAE Schönherr, 1823 

Tribe ALOPHINI LeConte, 1874 

GRAPTUS Schönherr, 1823 

ALOPHUS Schönherr, 1826 

triguttatus (Fabricius, 1775) 

Tribe BRACHYDERINI Schönherr, 1826 

BRACHYDERES Schönherr, 1826 

incanus (Linnaeus, 1758) 807 

lusitanicus (Fabricius, 1781) 808 

NELIOCARUS Thomson, C.G., 1859 

STROPHOSOMUS sensu auctt. partim non 


faber (Herbst, 1784) 

nebulosus (Stephens, 1831) 

retusus (Marsham, 1802) non (Fabricius, 1781) 

sus (Stephens, 1831) 

lateralis (Paykull, 1792) non (Panzer, 1789) 

STROPHOSOMA Billberg, 1820 

STROPHOSOMUS Schönherr, 1823 

capitatum (De Geer, 1775) 

obesum (Marsham, 1802) non (Fabricius, 1775)

subrotundum (Marsham, 1802) 


fulvicorne (Walton, 1846) 

curvipes Thomson, C.G., 1868 809 

melanogrammum (Forster, 1771) 

coryli (Fabricius, 1775) 

Tribe CNEORHININI Lacordaire, 1863 

ATTACTAGENUS Tournier, 1876 

ATACTOGENUS Fowler, 1891 (misspelling) 

CNEORHINUS sensu auctt. non Schönherr, 1823 

CNEORRHINUS auctt. (misspelling) 

plumbeus (Marsham, 1802) 

exaratus (Marsham, 1802) non (Gmelin in Linnaeus, 

1790) 

PHILOPEDON Schönherr, 1826 810 

plagiatum (Schaller, 1783) 

geminatus (Fabricius, 1787) 

Tribe GEONEMINI Gistel, 1848 

BARYNOTUS Germar, 1817 

moerens (Fabricius, 1792) 

elevatus (Marsham, 1802) 

obscurus (Fabricius, 1775) 

squamosus Germar, 1824 

schoenherri (Zetterstedt, 1838) non Dalman, 1823 

Tribe OMIINI Shuckard, 1839 

OMIAMIMA Silfverberg, 1977 811 

OMIAS sensu auctt. non Germar, 1817 

mollina (Boheman, 1834) 

Tribe OTIORHYNCHINI Schönherr, 1826 

OTIORHYNCHUS Germar, 1824 812 

OTIORRHYNCHUS auctt. (misspelling) 

BRACHYRHINUS Latreille, 1802 813 

arcticus (Fabricius, O., 1780) 

blandus Gyllenhal, 1834 

armadillo (Rossi, 1792) 814 

atroapterus (De Geer, 1775) 

aurifer Boheman, 1843 815 

auropunctatus Gyllenhal, 1834 

clavipes (Bonsdorff, 1785) 

tenebricosus sensu (Olivier, 1790) non (Herbst, 

1784) 

fuscipes (Olivier, 1807) 

lugdunensis Boheman, 1843 

crataegi Germar, 1824 816 

cribricollis Gyllenhal, 1834 817 

desertus Rosenhauer, 1847 

muscorum Brisout de Barneville, H., 1863 

dieckmanni Magnano, 1979 818 

setosulus sensu auctt. Brit. non Stierlin, 1861 819 

ligneus (Olivier, 1807) 

ligustici (Linnaeus, 1758) 

morio (Fabricius, 1781) 

ebeninus Gyllenhal, 1834 

nodosus (Müller, O.F., 1764) 

dubius (Strøm, 1783) 

107 


maurus (Gyllenhal, 1813) non (Marsham, 1802) 

ovatus (Linnaeus, 1758) 

porcatus (Herbst, 1795) 

raucus (Fabricius, 1777) 

rugifrons (Gyllenhal, 1813) 

dillwyni Stephens, 1831 

rugosostriatus (Goeze, 1777) 

scabrosus (Marsham, 1802) 

salicicola Heyden, 1908 820 

scaber (Linnaeus, 1758) 

septentrionis (Herbst, 1795) 

singularis (Linnaeus, 1767) 

picipes (Fabricius, 1777) 

sulcatus (Fabricius, 1775) 

uncinatus Germar, 1824 

Tribe PERITELINI Lacordaire, 1863 

PERITELUS Germar, 1824 

sphaeroides Germar, 1824 

griseus sensu (Olivier, 1807) non (Fabricius, 1775) 

Tribe PHYLLOBIINI Schönherr, 1826 

PHYLLOBIUS Germar, 1824 

Subgenus PHYLLOBIUS Germar, 1824 

pyri (Linnaeus, 1758) 

vespertinus (Fabricius, 1792) 821 

artemisiae Desbrochers, 1873 

Subgenus DIELETUS Reitter, 1916 

argentatus (Linnaeus, 1758) 

Subgenus METAPHYLLOBIUS Smirnov, 1913 

glaucus (Scopoli, 1763) 

calcaratus (Fabricius, 1792) 

pomaceus Gyllenhal, 1834 

urticae (De Geer, 1775) non (Scopoli, 1763) 

alneti (Fabricius, 1792) non (Schrank, 1781) 

Subgenus NEMOICUS Dillwyn, 1829 

oblongus (Linnaeus, 1758) 

Subgenus PARNEMOICUS Schilsky, 1911 

roboretanus Gredler, 1882 

virideaeris sensu auctt. Brit. non (Laicharting, 1781) 

parvulus sensu (Olivier, 1807) non (Fabricius, 1792) 

viridicollis (Fabricius, 1801) 

pomonae (Olivier, 1807) 

Subgenus PTERYGORRHYNCHUS Pesarini, 1969 

maculicornis Germar, 1824 

Subgenus SUBPHYLLOBIUS Schilsky, 1911 

virideaeris (Laicharting, 1781) 

viridiaeris auctt. (misspelling) 

Tribe POLYDRUSINI Schönherr, 1823 

LIOPHLOEUS Germar, 1817 

tessulatus (Müller, O.F., 1776) 

nubilus (Fabricius, 1777) 

maurus (Marsham, 1802) 

PACHYRHINUS Schönherr, 1823 

lethierryi (Desbrochers, 1875) 822 

mustela (Herbst, 1797) 823


POLYDRUSUS Germar, 1817 824 

POLYDROSOS auctt. (misspelling) 

Subgenus POLYDRUSUS Germar, 1817 

tereticollis (De Geer, 1775) 

undatus (Fabricius, 1781) 

Subgenus CHRYSOPHIS des Gozis, 1882 

THOMSONEONYMUS Desbrochers, 1902 

formosus (Mayer, 1779) 825 

sericeus (Schaller, 1783) non (Goeze, 1777) 

splendidus (Herbst, 1784) 

Subgenus EUDIPNUS Thomson, C.G., 1859 

mollis (Strøm, 1768) 

micans (Fabricius, 1792) 

Subgenus EURODRUSUS Korotyaev & Meleshko, 1997 

confluens Stephens, 1831 

Subgenus EUSTOLUS Thomson, C.G., 1859 

TYLODRUSUS Stierlin, 1884 

flavipes (De Geer, 1775) 

pterygomalis Boheman, 1840 

Subgenus METALLITES Germar, 1824 

marginatus Stephens, 1831 

Subgenus NEOEUSTOLUS Alonso-Zarazaga & Lyal, 

1999 

EUSTOLUS sensu auctt. non Thomson, C.G., 1859 

cervinus (Linnaeus, 1758) 

pilosus Gredler, 1866 

pulchellus Stephens, 1831 

chrysomela sensu auctt. Brit. non (Olivier, 1807) 826 

salsicola Fairmaire, 1852 

insquamosus (Everts, 1921) 

Tribe SCIAPHILINI Sharp, 1891 827 

BARYPEITHES Jacquelin du Val, 1855 

BARYPITHES Gemminger & Harold, 1871 40 

Subgenus BARYPEITHES Jacquelin du Val, 1855 

sulcifrons (Boheman, 1843) 

Subgenus EXOMIAS Bedel, 1883 

araneiformis (Schrank, 1781) 

curvimanus (Jacquelin du Val, 1855) 

pellucidus (Boheman, 1834) 

duplicatus Keys, 1911 

pyrenaeus Seidlitz, 1868 

BRACHYSOMUS Schönherr, 1823 

echinatus (Bonsdorff, 1785) 

hirtus (Boheman, 1845) 

SCIAPHILUS Schönherr, 1823 

asperatus (Bonsdorff, 1785) 

muricatus (Fabricius, 1792) non (Drury, 1773) 

Tribe SITONINI Gistel, 1848 828 

ANDRION Velázquez, 2007 

SITONA sensu auctt. partim non Germar, 1817 

regensteinense (Herbst, 1797) 

108 

CHARAGMUS Schönherr, 1826 


griseus (Fabricius, 1775) 

COELOSITONA González, 1971 


cambricus (Stephens, 1831) 

cinerascens (Fåhraeus, 1840) 829 

puberulus (Reitter, 1903) 

cambricus sensu auctt. Brit. partim non (Stephens, 

1831) 

brevirostris (Solari, 1948) 

SITONA Germar, 1817 

ambiguus Gyllenhal, 1834 

brevicollis sensu auctt. partim non Gyllenhal, 1834 

lineellus sensu Lindberg, 1933, auctt. Brit. post 

1930 non (Bonsdorff, 1785) 

cylindricollis (Fåhraeus, 1840) 

meliloti Walton, 1846 

gemellatus Gyllenhal, 1834 

gressorius (Fabricius, 1792) 830 

hispidulus (Fabricius, 1777) 

humeralis Stephens, 1831 

lepidus Gyllenhal, 1834 

flavescens (Marsham, 1802) non (Fabricius, 1787) 

lineatus (Linnaeus, 1758) 

lineellus (Bonsdorff, 1785) 

decipiens Lindberg, 1933 

macularius (Marsham, 1802) 

crinitus (Herbst, 1795) non (Gmelin in Linnaeus, 

1790) 

ononidis Sharp, 1866 

puncticollis Stephens, 1831 

striatellus Gyllenhal, 1834 

tibialis (Herbst, 1795) non (Sparrman, 1785) 

brevicollis Gyllenhal, 1834 partim 

sulcifrons (Thunberg, 1798) 

suturalis Stephens, 1831 

waterhousei Walton, 1846 

Tribe TANYMECINI Lacordaire, 1863 

TANYMECUS Germar, 1817 

palliatus (Fabricius, 1787) 

Tribe TRACHYPHLOEINI Gistel, 1848 831 

CAENOPSIS Bach, 1854 

fissirostris (Walton, 1847) 

waltoni (Boheman, 1843) 

CATHORMIOCERUS Schönherr, [1842] 

aristatus (Gyllenhal, 1827) 

attaphilus Brisout de Barneville, Ch., 1880 

myrmecophilus (Seidlitz, 1868) 

britannicus Blair, 1934 832 

socius Boheman, 1843 

maritimus Rye, 1874 

spinosus (Goeze, 1777) 

asperatus (Boheman, 1843) 

squamulatus sensu (Olivier, 1807) non (Herbst, 

1795) 

olivieri (Bedel, 1883) 

alleni (Donisthorpe, 1948)

ROMUALDIUS Borovec, 2009 

angustisetulus (Hansen, 1915) 833 

bifoveolatus sensu auctt. Brit. partim non (Beck, 

1817) 

bifoveolatus (Beck, 1817) 

scaber sensu auctt. non (Linnaeus, 1758) 

TRACHYPHLOEUS Germar, 1817 

TRACHYPHLAEUS Stephens, 1829 (misspelling) 

alternans Gyllenhal, 1834 

digitalis (Gyllenhal, 1827) 

rectus Thomson, C.G., 1865 

laticollis sensu auctt. Brit. non Boheman, 1843 834 

scabricul (Linnaeus, 1771) 

scabriculus auctt. (misspelling) 

scaber sensu Boheman, 1843 non (Linnaeus, 1758) 

spinimanus Germar, 1824 

Tribe TROPIPHORINI Marseul, 1863 

TROPIPHORUS Schönherr, 1842 

elevatus (Herbst, 1795) 

carinatus sensu (Müller, O.F., 1776) non (Linnaeus, 

1767) 

obtusus (Bonsdorff, 1785) 

terricola (Newman, 1838) 

tomentosus (Marsham, 1802) non (Olivier, 1790) 

Subfamily HYPERINAE Marseul, 1863 (1848) 

HYPERA Germar, 1817 

PHYTONOMUS Schönherr, 1823 835 

Subgenus HYPERA Germar, 1817 

arator (Linnaeus, 1758) 

polygoni (Linnaeus, 1761) 

fuscocinerea (Marsham, 1802) 

murina (Fabricius, 1792) non (Müller, O.F., 1764) 

nigrirostris (Fabricius, 1775) 

ononidis Chevrolat, 1863 

plantaginis (De Geer, 1775) 

postica (Gyllenhal, 1813) 

variabilis (Herbst, 1795) non (Fabricius, 1777) 

suspiciosa (Herbst, 1795) 

pedestris (Paykull, 1792) non (Poda, 1761) 

miles (Paykull, 1792) 32 

venusta (Fabricius, 1781) 

trilineata (Marsham, 1802) 

Subgenus ANTIDONUS Bedel, 1886 

dauci (Olivier, 1807) 

fasciculata (Herbst, 1795) non (De Geer, 1775) 

zoilus (Scopoli, 1763) 

punctata (Fabricius, 1775) non (Scopoli, 1763) 

?austriaca (Schrank, 1781) 

Subgenus BOREOHYPERA Korotyaev, 1999 

diversipunctata (Schrank, 1798) 

elongata (Paykull, 1792) non (Fabricius, 1775) 

Subgenus DAPALINUS Capiomont, 1868 

meles (Fabricius, 1792) 

Subgenus ERIRINOMORPHUS Capiomont, 1868 

arundinis (Paykull, 1792) 

pollux (Fabricius, 1801) 

adspersa (Fabricius, 1792) non (Fabricius, 1775) 

alternans (Stephens, 1831) 

109 

rumicis (Linnaeus, 1758) 


Subgenus TIGRINELLUS Capiomont, 1868 

pastinaceae (Rossi, 1790) 

tigrina Boheman, 1834 

LIMOBIUS Schönherr, 1843 

borealis (Paykull, 1792) 

dissimilis (Herbst, 1795) 

mixtus (Boheman, 1834) 

Subfamily LIXINAE Schönherr, 1823 

Tribe LIXINI Schönherr, 1823 

LARINUS Germar, 1824 


carlinae (Olivier, 1807) 

LIXUS Fabricius, 1801 836 

Subgenus LIXUS Fabricius, 1801 

paraplecticus (Linnaeus, 1758) 

productus Stephens, 1831 

Subgenus DILIXELLUS Reitter, 1916 

angustatus (Fabricius, 1775) 

algirus sensu auctt. non (Linnaeus, 1758) 

vilis (Rossi, 1790) 


lateralis Stephens, 1831 

Subgenus EULIXUS Reitter, 1916 

iridis Olivier, 1807 

paraplecticus sensu auctt. Brit. partim non 


turbatus Gyllenhal, 1824 

gemellatus Gyllenhal in Schönherr, 1836 

scabricollis Boheman, 1843 837 

Tribe CLEONINI Schönherr, 1826 

BOTHYNODERES Schönherr, 1823 

CHROMODERUS Motschulsky, 1860 

affinis (Schrank, 1781) 

fasciatus (Müller, O.F., 1776) non (Scopoli, 1763) 

albidus (Fabricius, 1787) 

CLEONIS Dejean, 1821 

CLEONUS Schönherr, 1826 

pigra (Scopoli, 1763) 

sulcirostris (Linnaeus, 1767) 

CONIOCLEONUS Motschulsky, 1860 

CLEONUS sensu Fowler, 1891 partim non 


hollbergii (Fåhraeus, 1842) 

hollbergi auctt. (misspelling) 

glaucus (Fabricius, 1787) non (Scopoli, 1763) 

nebulosus (Linnaeus, 1758) 

Tribe RHINOCYLLINI Lacordaire, 1863 

RHINOCYLLUS Germar, 1817 

conicus (Frölich, 1792)


latirostris (Latreille, 1804) 

Subfamily MESOPTILIINAE Lacordaire, 1863 

Tribe MAGDALIDINI Pascoe, 1870 

MAGDALIS Germar, 1817 

Subgenus MAGDALIS Germar, 1817 

duplicata Germar, 1818 

memnonia (Gyllenhal in Faldermann, 1837) 

phlegmatica (Herbst, 1797) 

Subgenus EDO Germar, 1819 

ruficornis (Linnaeus, 1758) 

pruni (Linnaeus, 1767) non (Scopoli, 1763) 

Subgenus ODONTOMAGDALIS Barrios, 1984 

MAGDALINUS sensu auctt. non Germar, 1843 

armigera (Fourcroy, 1785) 

carbonaria (Linnaeus, 1758) 

Subgenus PANUS Schönherr, 1826 

barbicornis (Latreille, 1804) 

Subgenus PORROTHUS Dejean, 1821 

NEOPANUS Reitter, 1916 

cerasi (Linnaeus, 1758) 

Subfamily MOLYTINAE Schönherr, 1823 

Tribe MOLYTINI Schönherr, 1823 

Subtribe MOLYTINA Schönherr, 1823 

LIPARUS Olivier, 1807 

MOLYTES Schönherr, 1823 

coronatus (Goeze, 1777) 

anglicanus (Marsham, 1802) 


Subtribe LEIOSOMATINA Reitter, 1913 

LEIOSOMA Stephens, 1829 

LEIOSOMUS Schönherr, 1842 40 

LIOSOMA Agassiz, 1847 non Brandt, 1835 40 

deflexum (Panzer, 1795) 

ovatulum (Clairville, 1798) 

oblongulum Boheman, 1842 

troglodytes (Rye, 1873) 838 

pyrenaeum sensu auctt. Brit. non Brisout de 

Barneville, Ch., 1866 

Subtribe PLINTHINA Lacordaire, 1863 

MITOPLINTHUS Reitter, 1897 

PLINTHUS Westwood, (1838) non Germar, 1817 

EPIPOLAEUS Weise, 1907 

caliginosus (Fabricius, 1775) 

Subtribe TYPODERINA Voss, 1965 

ANCHONIDIUM Bedel, 1884 

unguiculare (Aubé, 1850) 

110 

Tribe ANOPLINI Bedel, 1883 

ANOPLUS Germar, 1820 

plantaris (Naezen, 1794) 

roboris Suffrian, 1840 

Tribe HYLOBIINI Kirby, 1837 

HYLOBIUS Germar, 1817 

CURCULIO sensu Bedel, 1888 non Linnaeus 1758 

Subgenus CALLIRUS Dejean, 1821 

abietis (Linnaeus, 1758) 

transversovittatus (Goeze, 1777) 

fatuus (Rossi, 1790) 

Tribe LEPYRINI Kirby, 1837 

LEPYRUS Germar, 1817 

capucinus (Schaller, 1783) 

binotatus (Fabricius, 1792) 

Tribe PHRYNIXINI Kuschel, 1964 

SYAGRIUS Pascoe, 1875 

intrudens Waterhouse, C.O., 1903 

Tribe PISSODINI Gistel, 1848 

PISSODES Germar, 1817 

castaneus (De Geer, 1775) 

notatus (Fabricius, 1787) non (Bonsdorff, 1785) 

pini (Linnaeus, 1758) 

validirostris (Sahlberg, C.R., 1834) 

Tribe TRACHODINI Gistel, 1848 

TRACHODES Germar, 1824 

hispidus (Linnaeus, 1758) 

Subfamily OROBITIDINAE Thomson, C.G., 1859 

OROBITIS Germar, 1817 

cyaneus (Linnaeus, 1758) 

Subfamily SCOLYTINAE Latreille, 1804 839 

Tribe SCOLYTINI Latreille, 1804 840 

Subtribe SCOLYTINA Latreille, 1804 

SCOLYTUS Geoffroy, 1762 

ECCOPTOGASTER Illiger, 1798 

intricatus (Ratzeburg, 1837) 

laevis Chapuis, 1873 841 

mali (Bechstein & Scharfenberg, 1805) 

pruni (Ratzeburg, 1837) 

multistriatus (Marsham, 1802) 

pygmaeus (Fabricius, 1787) 842 

ratzeburgi Janson, 1856 

destructor sensu Ratzeburg, 1837 non Olivier, 1795

ugulosus (Müller, P.W.J., 1818) 

scolytus (Fabricius, 1775) 

destructor Olivier, 1795 

Subtribe CORTHYLINA LeConte, 1876 

PITYOPHTHORUS Eichhoff, 1864 

lichtensteinii (Ratzeburg, 1837) 843 

lichtensteini auctt. (misspelling) 

scoticus Blandford, 1891 

pubescens (Marsham, 1802) 

ramulorum (Perris, 1856) 

Subtribe CRYPHALINA Lindemann, 1877 

CRYPHALUS Erichson, 1836 

asperatus (Gyllenhal, 1813) 

abietis (Ratzeburg, 1837) 

ERNOPORICUS Berger, 1917 

ERNOPORUS sensu auctt. partim non Thomson, 

C.G., 1859 

caucasicus (Lindemann, 1876) 

fagi (Fabricius, 1798) 

ERNOPORUS Thomson, C.G., 1859 

tiliae (Panzer, 1793) 

TRYPOPHLOEUS Fairmaire, 1864 

CRYPHALUS sensu Fowler, 1891 partim non 


binodulus (Ratzeburg, 1837) 

asperatus sensu auctt. partim non (Gyllenhal, 1813) 

Subtribe CRYPTURGINA LeConte, 1876 

CRYPTURGUS Erichson, 1836 

subcribrosus Eggers, 1933 844 

Subtribe DRYOCOETINA Lindemann, 1877 

DRYOCOETES Eichhoff, 1864 

DRYOCAETES auctt. (misspelling) 

DRYOCOETINUS Balachowsky, 1949 

alni (Georg, 1856) 

autographus (Ratzeburg, 1837) 


LYMANTOR Løvendal, 1889 

DRYOCOETES sensu Fowler, 1891 partim non 

Eichhoff, 1864 

coryli (Perris, 1855) 

TAPHRORYCHUS Eichhoff, 1878 

bicolor (Herbst, 1794) 

villifrons (Dufour, 1843) 845 

XYLOCLEPTES Ferrari, 1867 

bispinus (Duftschmid, 1825) 

111 

Subtribe IPINA Bedel, 1888 


IPS De Geer, 1775 

TOMICUS sensu Fowler, 1891 partim non Latreille, 

1802 

acuminatus (Gyllenhal, 1827) 

cembrae (Heer, 1836) 

sexdentatus (Boerner, 1766) 

typographus (Linnaeus, 1758) 

ORTHOTOMICUS Ferrari, 1867 

ONTHOTOMICUS Ferrari, 1867 

TOMICUS sensu Fowler, 1891 partim non Latreille, 

1802 

erosus (Wollaston, 1857) 

laricis (Fabricius, 1792) 

suturalis (Gyllenhal, 1827) 

PITYOGENES Bedel, 1888 

bidentatus (Herbst, 1784) 

ater (Fabricius, 1792) 

chalcographus (Linnaeus, 1760) 

quadridens (Hartig, 1834) 

trepanatus (Nördlinger, 1848) 

Subtribe XYLEBORINA LeConte, 1876 

XYLEBORINUS Reitter, 1913 

XYLEBORUS sensu auctt. partim non Eichhoff, 

1864 

saxesenii (Ratzeburg, 1837) 

saxeseni auctt. (misspelling) 

xylographus sensu auctt. Brit. non (Say, 1826) 

XYLEBORUS Eichhoff, 1864 

ANISANDRUS Ferrari, 1867 

dispar (Fabricius, 1792) 

dryographus (Ratzeburg, 1837) 

villifrons sensu Donisthorpe, 1924 non (Dufour, 

1843) 

sampsoni Donisthorpe, 1940 

donisthorpei Schedl, 1951 

monographus (Fabricius, 1792) 846 

Subtribe XYLOTERINA LeConte, 1876 

TRYPODENDRON Stephens, 1830 

XYLOTERUS Erichson, 1836 

domesticum (Linnaeus, 1758) 

lineatum (Olivier, 1795) 

signatum (Fabricius, 1792) 

Tribe HYLESININI Erichson, 1836 

Subtribe HYLESININA Erichson, 1836 

HYLESINUS Fabricius, 1801 

LEPERISINUS Reitter, 1913 


toranio (Danthoine in Bernard, 1788) 

oleiperda (Fabricius, 1792) 847 

bicolor (Brullé, 1832) 


fraxini sensu auctt. Brit. non (Panzer, 1799) 848

Family PLATYPODIDAE Shuckard, 1839 Checklist of Beetles of the British Isles, 2012 Edition 

wachtli Reitter, 1887 849 

ssp. orni Fuchs, 1906 850 

HYLASTINUS Bedel, 1888 

obscurus (Marsham, 1802) 

KISSOPHAGUS Chapuis, 1869 

CISSOPHAGUS auctt. (misspelling) 

vicinus (Comolli, 1837) 

hederae (Schmitt, 1843) 

PTELEOBIUS Bedel, 1888 

ACRANTUS sensu auctt. Brit. non Broun, 1881 

HYLESINUS sensu Fowler, 1891 partim non 


vittatus (Fabricius, 1792) 

Subtribe HYLASTINA LeConte, 1876 

HYLASTES Erichson, 1836 

angustatus (Herbst, 1794) 

ater (Paykull, 1800) 851 

pinicola Bedel, 1888 852 

attenuatus Erichson, 1836 

brunneus Erichson, 1836 

cunicularius Erichson, 1836 

opacus Erichson, 1836 

HYLURGOPS LeConte, 1876 

HYLASTES sensu Fowler, 1891 partim non 


palliatus (Gyllenhal, 1813) 

Subtribe HYLURGINA Gistel, 1848 

DENDROCTONUS Erichson, 1836 

micans (Kugelann, 1794) 853 

TOMICUS Latreille, 1802 

BLASTOPHAGUS Eichhoff, 1864 

MYELOPHILUS Eichhoff, 1878 

minor (Hartig, 1834) 

piniperda (Linnaeus, 1758) 

XYLECHINUS Chapuis, 1869 

pilosus (Ratzeburg, 1837) 

Subtribe PHLOEOTRIBINA Chapuis, 1869 

PHLOEOSINUS Chapuis, 1869 

aubei (Perris, 1855) 854 

bicolor sensu auctt. non (Brullé, 1832) 855 

thujae (Perris, 1855) 

PHLOEOTRIBUS Latreille, 1797 

PHLOEOPHTHORUS Wollaston, 1854 


Subtribe POLYGRAPHINA Chapuis, 1869 

POLYGRAPHUS Erichson, 1836 

grandiclava (Thomson, C.G., 1886) 856 

112 

poligraphus (Linnaeus, 1758) 

pubescens (Fabricius, 1792) 

103. Family PLATYPODIDAE Shuckard, 1839 857 


PLATYPUS Herbst, 1793 

cylindrus (Fabricius, 1792) 

Families: 103 

Genera: 1265 

Species: 4072

Endnotes 

1 

Emended from original Sphaeriidae to remove 

homonymy with Sphaeriidae Deshayes, 1854 (1820) 

(Mollusca, Bivalvia), ICZN Op. 1957. 

2 

Name conserved, ICZN Op. 1957. 

3 

Aulonogyrus striatus (Fabricius), listed in Pope 

(1977), has been deleted from the British List. 

4 


5 

Subgenera of Haliplus follow Holmen (1987). 

6 

Added (reinstated?) by Parry (1982a) and more fully 

by Parry (1983). 

7 

Synonymized by Lundmark, Drotz & Nilsson (2001). 

8 

This checklist is based on Nilsson (2003). 

9 

Use of the younger family name Hygrobiidae is in 

accordance with ICZN Article 40.2 and follows 

Bouchard et al. (2011). 

10 

Name rejected, ICZN Op. 280. 

11 

This checklist is based on Nilsson (2003[2006]). 

12 

Junior primary homonym of biguttatus Gmelin in 

Linnaeus, 1789. 

13 Name conserved, ICZN Op. 1556. 

14 Transferred from Agabus to Ilybius by Nilsson 

(2000). 

15 Added by Owen et al. (1992a) and more fully by 

Owen et al. (1992b) [as Agabus wasastjernae]. 

16 Name rejected, ICZN Op. 289. 

17 Name established by Nilsson & Holmen (1995). 

18 Separated from Copelatus s.str. by Balke, Ribera & 

Vogler (2004). 

19 Justified emendation of the original “Cybisterini”. 

20 Hydaticus continentalis Balfour-Browne, J., has been 

deleted from the British List (Foster, 2012). 

21 Hydroporus foveolatus Heer, listed in Pope (1977), 

has been deleted from the British List. 


23 

Renamed by Fery (1999). 

24 


25 

N. canariensis (Bedel), listed in Pope (1977) [in 

Potamonectes], has been deleted from the British 

List. 

26 

Added by Carr (1999) and more fully by Carr (2000). 

27 

Added by Foster & Spirit (1986). 

28 

Added by Foster & Friday (2011). 

29 

Added by Drake (2006). 

30 

Added by Carr (1984b). 

31 

C. (Autocarabus) cancellatus Illiger, listed by Pope 

(1977), is here treated as a non-established 

introduction. 

32 

Forgotten senior synonym. 

33 

Synonymized by Coulon (2004). See also Hammond 

(2002). 

34 

Added by Speight, Martinez & Luff (1986). 

35 

Added by Focarile (1964); see also Speight, 

Martinez & Luff (1986). 

36 

Junior primary homonym of varium Müller, O.F., 

1776. 

37 

Added by Telfer (2001a). 

38 

Added by Levey & Pavett (1999). 

39 

Junior primary homonym of minimum Müller, O.F. 

1776. 

40 

Unjustified emendation. 

113 

Endnotes 

41 

Junior primary homonym of littoralis Olivier, 1795. 

42 

Junior primary homonym of aethiops Herbst, 1784. 

43 

Junior primary homonym of longicollis Lichtenstein, 

1796. 

44 

Junior primary homonym of niger Linnaeus, 1764. 

45 

Added by J.A. Owen in Jones (1989) and more fully 

by Luff (1990). 

46 Junior primary homonym of vernalis Müller, O.F., 

1776. 

47 Abax parallelus (Duftschmid) has been moved to the 

list of Non-established introductions. 

48 Added by Anderson & Luff (1994). 

49 Junior primary homonym of mollis Strøm, 1768. 

ICZN Op. 1723 rejects Strøm's name for Principle of 

Priority but not for Principle of Homonymy. 

50 Junior primary homonym of rotundatus Gmelin in 


51 Added by Anderson (1985). 

52 Junior primary homonym of ovata Paykull, 1790. 

53 Junior primary homonym of infima Gravenhorst, 

1807. 

54 Junior primary homonym of fulva De Geer, 1774. 

55 Confirmed as British by Owen (1996). 

56 Junior primary homonym of cordatus Scopoli, 1763. 

57 

Added by Telfer (2001b). 

58 

Stenolophus comma (Fabricius) has been moved to 

the list of Non-established introductions. 

59 

Added by Telfer (2003). 

60 

British records reviewed by Denton (2007), who 

showed that the species has been temporarily 

established in Britain in the past. 

61 

Added by Hammond (1982a). 

62 

Junior primary homonym of agilis Müller, O.F., 1776. 

63 

Added by Eversham & Collier (1997). 

64 

Hansen (1991) elevated the Helophorinae 

(Hydrophilidae) to the level of a family within the 

Hydrophiloidea. Classification follows Angus (1999). 

65 


66 

Synonymized by Angus (1982). 

67 


68 

Hansen (1991) elevated the Georissinae 


Hydrophiloidea. 

69 


70 

Hansen (1991) elevated the Hydrochinae 



71 

Synonymized by Hansen (1999). 

72 

Added by van Berge Henegouwen (1988). 

73 

Hansen (1991) elevated the Spercheinae 



74 

Subfamilies and sequence of genera follows Löbl & 

Smetana (2004), after Foster (2005). 

75 

Status revised by van Berge Henegouwen (1986). 

76 


77 

Added by Foster (2005). See also Levey (2005). 

78 

Added by Sharp (1914) but since synonymized by 

authors including Pope (1977); status revised by 

Hansen (1987). 

79 

Added by Newbery (1914) but since synonymized by 

authors including Pope (1977); status revised by 

Hansen (1987).

Endnotes Checklist of Beetles of the British Isles, 2012 Edition 

80 

Added by Foster (1984) [as Enochrus isotae]. 

81 

Synonymized by Schödl (1997). 

82 

Added by Hansen (1982). 

83 

Junior primary homonym of niger Fabricius, 1775. 

84 

British records of L. obscuratus Rottenberg require 

confirmation. See Gentili & Chiesa (1975) for a 

putative record. 

85 

Added by Gentili & Chiesa (1975) [as “L. simulator”]. 

86 

Synonymized by Gentili & Ribera (1998). 

87 

Added by Allen (2004). 

88 

Arrangement of subgenera follows Hansen (1987). 

89 Added by Owen & Mendel (1990). 

90 Synonymized by Allen (1969). 

91 Synonymized by Hansen (1999). 

92 Status revised by van Berge Henegouwen (1989). 

93 Nomenclature and arrangement of this family follow 

Mazur (1984). 

94 Added by Lackner (2005). 

95 Added by Nash (1982). 

96 Added by Allen & Hance (2009). 

97 Synonymized by Hansen (1998). 

98 Added by Carr (1984a). 

99 Raised from level of subgenus of Ochthebius by 


100 Three further species are in need of confirmation as 

British. Two specimens of O. (Hymenodes) difficilis 

Mulsant, labelled from “Britannia” (M. von Pfaundler 

coll.), are in the Zoologische Staatssammlung, 

Munich (Hebauer, 1985); one male specimen of O. 

(Hymenodes) pilosus Waltl, labelled from “Anglia”, is 

in the Deutsches Entomologisches Institut, 

Eberswalde (Jäch, 1989); and a specimen of O. 

(Hymenodes) quadrifoveolatus Wollaston, labelled 

from “Anglia” (leg. Jekel), is in the Universitets 

Zoologiska Museum, Helsingfors (Jäch, 1989). In the 

absence of specimens with more accurate collection 

data the evidence that these three species have 

occurred in Britain must remain inconclusive. 

101 Added by Foster (2008) and more fully by 

O'Callaghan et al. (2009). 

102 For a review of the O. viridis species group, see 

Foster (2009). 


104 Synonymized by Johnson (2001a). 

105 Added by Johnson (2001b). 

106 Synonymized by Biström & Silfverberg (1979); see 

also Johnson (1987a). 

107 Not Erichson, 1845; name conserved, ICZN Op. 

1277. 

108 

Reinstated by Johnson (1987a). 

109 

Now a separate genus (Dybas & Dybas, 1990). 

110 

Transferred from Ptinellini to Ptiliini by Darby (2012). 

111 

Added by Johnson (1987a). 

112 

Transferred from Ptiliini to Ptinellini by Darby (2012). 

113 

Synonymized by Johnson (1982b); see also Johnson 

(1987a). 


115 Checklist is based on Johnson (1987b). This paper 

showed that A. championi and A. sarae, listed as 

'species incertae sedis' in Pope (1977), are foreign 

species which have no valid place on the British 

Isles List. 

116 Listed as a 'species incertae sedis' in Pope (1977); 

synonymized by Johnson (1987b). 

114 

117 

See Johnson (2003). 

118 

thomsoni Erichson is an invalid varietal name lacking 

description, so has no standing in nomenclature 

(Johnson, 1987b). 

119 

Added by Johnson (1992b). 

120 

See Sörensson (2003). 

121 

Added by Johnson (1967), but shown to be a 

misidentification of A. rugulosa by Johnson (1975a). 

122 

Added by Johnson (1987a). 

123 

The checklist for this subfamily is based on the 

checklist in Cooter (1996b). Aglyptinus agathidioides 

Blair has been moved to the list of Non-established 

introductions. 

124 

Reidentified by Cooter (1992). 

125 

See Cooter (1996a). 

126 

Reidentified by Cooter (2004). 

127 


128 

Added by Collier (1986). 

129 

Added by Schilthuizen (1990) [as Choleva 

septentrionis septentrionis] and confirmed by 

Růžička & Vávra (2003) [as C. lederiana]. 

130 

Added by Schilthuizen (2010). 

131 

Junior primary homonym of tristis Rossi, 1790. 

132 

Allocation of genera to tribes follows Herman 

(2001b). 

133 

Zanetti (1987) used the name Lathrimaeum for this 

genus, because he considered that the type species 

of Anthobium is a species of Eusphalerum. However, 

current usage of Anthobium has been conserved by 

redesignation of the type species (Herman, 2001a). 

134 

Synonymized by Assing & Schülke (2012). 

135 


136 

Current usage of this name has been conserved by 

redesignation of its type species (Herman, 2001a). 

137 


138 

Regarded as a subspecies by Zanetti (1987) and 

Smetana (2004a), but merely as a synonym by 


139 

Added by Lott (1989). 

140 


141 

Tottenham (1954) used this name for A. minuta, 

which had been recently found for the first time in 

Britain (Steel, 1957), although Joy (1932) used the 

name for the more widespread A. sulcula. 

142 

Raised to generic rank by Steel (1970), though not 

followed by most subsequent authors until generic 

status reaffirmed by Thayer (2003). 

143 

Synonymy appears in Zanetti (1987). 

144 

Synonymized in Hammond (1996). 

145 

Not listed by Smetana (2004a), but specific status 

and presence in Britain confirmed by Jászay & 

Hlavac (2006). 

146 

Added by Jászay & Hlavac (2006). 

147 


148 

Added by Hammond (2007b). 

149 

Raised to generic rank by Steel (1970) and most 

contemporary authors, though still combined with 

Phloeonomus by Poole & Gentili (1996). 

150 

Raised to generic rank by Steel (1970) and most 

contemporary authors, though still combined with 

Hapalaraea by Poole & Gentili (1996). 

151 


152 

The type series of brunnipennis is conspecific with 

depressus (Hammond, pers. comm.), so this name

cannot be used to replace concinnus as suggested 

by Hansen (1996). 

153 

Raised to generic rank by Steel (1970). 

154 

Added by Allen & Booth (2005). 

155 

Junior homonym replaced in Cuccodoro & Löbl 

(1997). 

156 

This name was previously used as a 

misidentification of what is now known as M. 

prosseni (Cuccodoro & Löbl, 1997). Consequently, 

great care needs to be taken in interpreting the 

name depressus in the literature and it is 

recommended that when using this name in future, it 

should be quoted as 'depressus (=sinuatocollis)'. 

157 

Synonymized by Cuccodoro & Löbl (1997). 

158 

Synonymized by Zerche (1998). 

159 

Name protected by Herman (2001a). 

160 

First replaced with the name serrifer (Muona, 1977), 

which was later synonymized by Gusarov (1992). 

161 

Synonymized by Assing & Schülke (2012). 

162 

Formerly regarded as a separate family, the 

Pselaphinae were placed within the Staphylinidae by 

Newton & Thayer (1995). Supertribes, tribes and 

subtribes follow Löbl & Besuchet (2004). 

163 

The species in this genus have been confused in the 

British literature up until very recently. Names in old 

lists should be interpreted with caution. 

164 

Added by Bowestead & Eccles (1987). 

165 

Subspecific status reaffirmed by Besuchet (1999). 

166 

This name has been frequently misapplied to E. 

bescidicus by British authors including Pearce 

(1957). See also Allen (1994b). 

167 

Junior homonym replaced in Löbl & Besuchet 

(2004). 

168 

Until recently the name, E. punctatus, was applied to 

two British species formerly considered to be of 

subspecific status. 

169 

Raised from subspecific rank by Besuchet (1989). 

170 

Added by Johnson & Eccles (1983). 

171 

The species in this genus have been confused in the 

British literature in the past. Names in old lists should 

be interpreted with caution. 

172 

Added by Johnson (2012c). 

173 

Formerly considered to be a subspecies of 

haematica (Reichenbach), but raised to specific rank 

by Besuchet (1999) and recorded from Britain by 

Sabella et al. (2004). 

174 

Löbl (1998) considered this name to be a junior 

synonym of hipponensis (Saulcy) because he 

mistakenly believed the date of publication to be 

1882. In fact Rye (1869) had published the name 

earlier to replace simplex (Waterhouse) which was 

preoccupied. 

175 

Usage follows Löbl & Besuchet (2004). 

176 

Considered to be an intraspecific male form of 

longicornis by Besuchet (1989), but regarded as a 

separate species by Odegaard (2001). 

177 

Lectotype designated by Schülke (1999), 

establishing synonymy with Quedius scitus 

(Gravenhorst, 1806). All uses of the name Bolitobius 

analis (Fabricius) are therefore misidentifications. 

178 

Raised to generic rank by Campbell (1993). 

179 

Synonymized by Schülke (2012). 

180 

Formerly treated as a subgenus of Mycetoporus, but 

raised to full generic rank by Campbell (1991). 

181 

Added by Schülke (2011b). 

115 

182 Transferred from Bolitobius by Assing & Schülke 

Endnotes 

(2001). 

183 Prior to Hammond (1973), this name was also used 

for three other closely related species, which were 

then unrecognised. 

184 Species groups have been recognised by Ullrich 

(1975). 

185 Formerly used as a subgenus, but synonymized by 

Campbell (1973). 

186 Synonymy appears in Lohse (1989). The Scottish 

paratype is also conspecific with pallipes (Hammond, 

pers. comm.). 

187 Usage and synonymy reinstated by Schülke (2005). 

188 Added by Booth (1988). 

189 The subgenus Palporus was created for this species 

by Campbell (1979). 

190 The list of tribes follows Newton & Thayer (1992) 

and is the same as that used by Smetana (2004d). 

Subtribes used by Smetana (2004d) are also 

included. 

191 This tribe is included within the Homalotini by Ashe 

(1998). 

192 Most subgenera are assigned to full generic rank by 

Seevers (1978), but this practice has not been 

followed by recent authors (e.g. Maus et al., 2000). 

Subgenera and species names follow Welch (1997), 

except where indicated. 

193 Added by Welch (1969) [as Aleochara verna]. This 

species was misidentified as Aleochara pauxilla by 

Lohse (1974). 

194 Reidentified by Welch (1990b). 

195 Added by Owen (1990c) [as Aleochara pauxilla]. 

196 

Synonymized by Maus (2001). 

197 

This name has been previously misapplied to A. 

punctatella (Lohse, 1985b). 

198 

Only known from the unique type specimen taken in 

Dorset in 1937; listed as a synonym of A. obscurella 

by Smetana (2004d), but regarded as a good 

species by Welch (1997). 

199 

Combined with Emplenota in Welch (1997), but its 

distinctness was emphasised by Maus et al. (2000). 

200 

Synonymized by Welch (1997). 

201 

Replacement due to Likovský (1984). 

202 

Previously placed in the tribe Hoplandriini, but 

transferred to the Aleocharini by Hanley (2002). 

203 

The generic limits in this subtribe are in need of 

review and are likely to be fluid for the foreseeable 

future (see also notes in family introduction). There 

are inconsistencies between the checklists published 

in Central Europe and Scandinavia (see e.g. Lohse 

1985a, 1987; Muona 1987). The list of genera in the 

Palaearctic catalogue (Smetana 2004d) largely 

follows central European usage while incorporating 

as subgenera several generic names recognised in 

Scandinavian checklists. While this approach 

outlines divisions in the large amorphous genus, 

Atheta, it leaves several fairly distinctive groups 

within Atheta unrecognised in binomial names. 

Consequently, this checklist continues the expedient 

approach of splitting Atheta into manageable groups 

and recognising them as genera. A note is given 

wherever appropriate to explain how taxa are treated 

differently by Smetana (2004d). 

204 

Species groups have been identified by Brundin 

(1952).


205 Synonymized by Allen (1994a). 

206 Added by Lott (2009b). 

207 Added by A.C. Galsworthy & R.G. Booth in Hodge 

(2005). 

208 Synonymized by Nikitsky et al. (1998). 

209 Synonymized by Lohse (1988). 

210 Synonymized by Gusarov (2003). 

211 Listed by Smetana (2004d) as a subgenus of Atheta, 

but widely ranked as a genus by previous authors. 

Several species listed in Lohse's Atheta 

'Mischgruppe II' are included here. 

212 Added by P.J. Hodge in Hodge (2003) [as Atheta 

linderi]. 

213 Listed by Smetana (2004d) in Oreostiba. 

214 Listed by Smetana (2004d) in Traumoecia. 

215 British records of Alevonota spp. require review, 

following publication of Assing & Wunderle (2008). 

216 Synonymized by Assing & Wunderle (2008). 

217 Added by Last (1980). 

218 Reidentified by Lott (1993b). 

219 Added by Hammond (1981). 

220 Raised to generic rank by Muona (1979a) and 

Köhler & Klausnitzer (1998), though not by Assing & 

Schülke (2007). 

221 Before Last (1952), this name was misapplied to A. 

longicollis. 

222 Before Last (1952), this name was misapplied to A. 

coulsoni. 

223 Synonymized by Muona (1990). 

224 Synonymized by Williams (1979a). 


but ranked as a genus by Seevers (1978) and 

Silfverberg (2004). 

226 Synonymized by Benick (1970); formerly placed in 

Microdota. 

227 Added by Allen (1994a) [as Atheta verulamii]. 

228 

Synonymized by Vogel (2004). 

229 

Kevan (1963) placed this species in Atheta 

subgenus Amidobia. Muona (1979b) found that the 

species had been misinterpreted by Brundin (1948), 

who, according to Allen (1994a) and Assing et al. 

(1998), confused it with A. puberula, a species 

assigned to Anopleta by Benick (1970). 

230 

Formerly placed in Microdota but transferred to 

Anopleta by Benick (1970). 

231 

Used as a subgenus of Atheta by Likovský in Boháĉ 

(1993) for A. liturata and A. nigritula and retained by 

Smetana (2004d). 

232 

Added by Lyszkowski (1992). 

233 

Listed by Pope (1977) but first formally recorded by 

Allen (1994a). 

234 

The list below contains species assigned to Atheta s. 

str. by Smetana (2004d) that should be treated 

provisionally as belonging to the genus Atheta. The 

list is made up of various species that were placed in 

Lohse's 'Mischgruppen I and III/IV' in Atheta sensu 

lato (Freude et al., 1974). It contains a number of 

species groups that are not necessarily closely 

related, and which await a proper revision of generic 

relationships within the Athetini (see discussion by 

Gusarov (2003)). 

235 

Listed by Pope (1977) but first formally recorded by 

Allen (1994a). 

236 

Replacement due to Muona (1979b). 

116 

237 Included in Lohse's 'Mischgruppe I’, but assigned to 

Dimetrota by Brundin (1963) and Smetana (2004d). 

238 Placed in the genus Plataraea by Muona (1979a), 

but not by Lohse (1985a). 

239 Nomenclature as in Smetana (2004d). 


and not ranked as a genus by recent authors. 

Treated as separate from Dimetrota by Muona 

(1979a) and Baranowski (1982), though not by 

Lohse (1989). Synonymized with Atheta rather than 

Dimetrota by Poole & Gentili (1996). 


but ranked as a genus by Seevers (1978). 

242 Regarded as a good species by Lohse (1989). 

243 Formerly included in Atheta subgenus Actophylla; 

name protected by Vogel (2004). 

244 Listed as a synonym of Dimetrota by Smetana 

(2004d). 

245 Added by Owen (1983) [as Atheta hansseni]. 

246 Synonymized by Assing (2001). 



248 Synonymy by Benick (1974) confirmed by Muona 

(1979b). 




and not ranked as a genus by recent authors, but 

regarded as a subgenus of Acrotona by Seevers 

(1978). 

251 Generic assignment established by Gusarov (2003) 

following discussion by Muona (1979b). 



253 In Joy (1932) and Strand & Vik (1964), this name 

was misapplied to D. dadopora. 

254 Listed on its own in Atheta subgenus Datostiba by 

Likovský in Boháč (1993). 

255 In Joy (1932), this name was misapplied to D. nigra. 


but ranked as a genus by Seevers (1978) and 

Silfverberg (2004). 

257 This is the type species of Dralica, which was 

retained as a separate subgenus of Atheta by 

Likovský in Boháč (1993) and Vogel (2004). 


but ranked as a genus by Seevers (1978) and Lohse 

et al. (1990). The traditional concept of Dimetrota 

has been threatened by Blackwelder’s designation of 

Homalota tristicula (=Cadaverota cadaverina) as the 

type species (Muona, 1979b), but Lohse & Smetana 

(1985) argued that Fenyes’ previous designation of 

D. marcida is valid. Smetana (2004d) includes both 

Cadaverota cadaverina and traditional Dimetrota 

species within the genus. 

259 The true identity of this species was established by 

Lohse & Smetana (1985). 

260 Synonymized by Lott (2002). 


262 Listed incorrectly as a junior synonym of H. 

puncticeps by Smetana (2004d). 

263 Listed as British by Hammond (2000a) on the basis 

of an as yet unpublished revision of the British 

species of Halobrecta. Diagnostic characters are

given by Gusarov (2004). Listed incorrectly as a 

synonym of H. flavipes by Smetana (2004d). 

264 Not listed by Smetana (2004d). 

265 Synonymized by Assing & Schülke (2001). 

266 Added by Allen & Eccles (1988) [as Hydrosmectina 

delicatissima]. 

267 Allen (1992a), following Lohse (1988), replaced H. 

subtilissima in the British list with this name, but it is 

synonymized in Smetana (2004d). 

268 Synonymy appears in Muona (1979a) and is 

followed by subsequent checklists. 

269 Listed as a subspecies by Muona (1979a) but Assing 

& Schülke (2007) argued that it should be regarded 

as a synonym. 

270 Placed in Atheta by Poole & Gentili (1996), though 

assigned to Liogluta by all recent European authors 

and Seevers (1978). 

271 Synonymy confirmed by Assing & Schülke (1999). 


but ranked as a genus by Seevers (1978). Species 

groups have been identified by Brundin (1948). 

273 Added (reinstated) by Allen (1994a). In Joy (1932) 

this name was misapplied to M. glabricula. 

274 In Joy (1932) this name was misapplied to Anopleta 

soedermani. 

275 Listed as a synonym of Acrotona by Smetana 

(2004d). Carter & Owen (1987) listed M. gilvicollis 

Scheerpeltz as a British species, although there are 

no published British records for this species, whose 

taxonomic status in any case requires further 

scrutiny, preferably as part of a wider study of the M. 

fungi species group.(Assing & Schülke, 2001). 

276 Hodge & Jones (1995) reported that some British 

records of M. clientula were referable to M. 

negligens (Mulsant & Rey), but provided no 

supporting evidence. 



278 Pace (2006) placed this species in Atheta subgenus 

Oxypodera Bernhauer, 1915. 

279 Replaced by Zerche (1991). 


but ranked as a genus by Brundin (1940) and 

Likovský in Boháĉ (1993). 


but ranked as a genus by Silfverberg (2004). 

282 In Joy (1932) this name was misapplied to Microdota 

atricolor. 




even though the promotion of this taxon to full 

generic status was proposed by Muona (1995) and it 

was used as a genus by Seevers (1978) and Lohse 

et al. (1990). Species groups have been recognised 

by Brundin (1943). 

285 In Strand & Vik (1964), this name was misapplied to 

P. malleus. 

286 Synonymized by Muona (1995). 



288 Added by Owen (1990b) [as Schistoglossa benicki]. 

289 Synonymized by Mahler & Vagtholm-Jensen (2002). 



117 

Endnotes 

291 Added by Heal (1993). 


but ranked as a genus by Seevers (1978). The 

American concept of this genus is much wider than 

that used here. It includes a number of species 

whose relationships are unclear and which in Europe 

would be placed in Atheta sensu lato. 

293 Synonymized by Good (1998). 

294 Correct name established by Good (1998). 

295 Listed as Myrmecoporus sp. by Hammond (2000a); 

see note below. 

296 

Added by Assing (1997a), but considered to be the 

true brevipes by Hammond (2000a) and listed as 

such in his list of British coastal Staphylinidae. 

297 

Synonymized by Assing (1999c). 

298 

Usage conserved by designation of Leptusa 

pulchella as type species (ICZN Op. 2113). 

299 

Added by Owen (1991) [as Pseudomicrodota 

jelineki]. 

300 

Synonymized by Vogel (2004). 

301 

Placed in Hygroecia (= Philhygra) by Joy (1932). 

302 

Taxonomic status doubtful (Hammond, 1996); only 

known from the unique type specimen taken in 

Hertfordshire in 1930. 

303 

In Joy (1932) this name is misapplied to Gyrophaena 

congrua. 

304 A junior homonym according to Nikitsky et al. (1998). 

305 

Added by Welch (2000). 

306 

Added by R.G. Booth and A.C. Galsworthy in Booth 

(2008). 

307 


308 

British records of Cypha ovulum (Heer) require 

confirmation. See Hyman (1994). 

309 

This name was used for Cypha pulicaria by Joy 

(1932). 

310 

Synonymized by Dauphin (2003, 2004). 

311 


312 

Placed in the genus Oligota by Williams (1979c), 

although this has been overlooked by subsequent 

authors who allocate it to the genus Holobus. 

313 

Treated as a genus by Maruyama (2006) following 

Kistner (1971). 

314 Species groups have been identified by Muona 

(1991). 

315 Usage follows Nikitsky et al. (1998). 

316 Muona (1979b) considered this name to be a nomen 

dubium. 

317 Added by Owen (1982) [as Meotica lohsei]. Before 

1976, this name was misapplied to Meotica anglica 

by British coleopterists. 

318 Species groups have been identified by Assing 

(1996a). 


320 Raised to generic rank by Assing & Schülke (2001). 

321 Added by Williams (1979b) [as Ocyusa nitidiventris], 

but central European and British records are referred 

to Cousya nigrata by Assing & Schülke (2007). 

322 See discussion by Smetana (2004d). 

323 

Synonymized by Assing (1999b). 

324 

Added by Lott (1993a). 

325 

Added by Owen (1994b) and more fully by Owen 

(1994d). 

326 

Listed as a synonym of Ocyusa by Poole & Gentili 

(1996), even though its generic status has long been


established in Europe and even though Seevers 

(1978) suggested that it was distinct. 

327 

Synonymized by Assing & Schülke (2001). 

328 

This name is now used for a species in the Athetine 

genus Nehemitropia (Zerche, 1991). 

329 

This synonymy appears in an update of the Central 

European list (Lohse, 1989) and is followed in other 

checklists. 

330 

The use of this name follows Zerche (1994). Not 

recorded since before 1970 (Hyman, 1994). 

331 

Added by Williams (1990) from a single female found 

at Dungeness, Kent. 

332 

Regarded as a long-winged form of Oxypoda 

brachyptera by Pope (1977), but raised to specific 

rank in most European works. 

333 

Synonymized by Lohse (1984). 

334 

Added by R.G. Booth in Hodge (1999). 

335 Synonymized by Dauphin (2001). 

336 Reidentified by Whitehead (1994). 

337 Nomenclatural change due to Assing & Schülke 

(2007). 

338 Listed by Smetana (2004d) under the tribe Athetini, 

but transferred to the Oxypodini by Lohse (1989) and 


339 The name as first used by Scheerpeltz (1968) is 

unavailable, and the correct authorship was 

established by Assing & Schülke (2001). 

340 Raised to generic rank in Assing et al. (1998). 

341 Included in Tachyusa by Assing et al. (1998) but 

assigned to Ischnopoda by Paśnik (2006b). 

342 The type species of Tachyusa and Ischnopoda were 

set by ICZN Op. 600. 

343 Status revised by Paśnik (2006a). British records of 

Tachyusa coarctata (Erichson) apparently all relate 

to T. concinna Heer (R.G. Booth, pers. comm.). 

344 Added by R.G. Booth and P.J. Hodge in Hodge 

(2003). British records summarised by Telfer 

(2007b). 

345 Added by R.G. Booth in Hodge (1999). 

346 Formerly regarded as a separate family, but 

incorporated into the Staphylinidae as a subfamily on 

grounds of larval morphology by Kasule (1966). This 

approach was supported by Lawrence & Newton 

(1982) and adopted by most subsequent authors 

except for Hansen (1997). Tribal classification and 

arrangement follow Löbl (1997). 

347 Tribal classification follows Herman (2001b). Hansen 

(1996) gives an alternative classification. Gildenkov 

(2003) erected a new tribe, Mandini, which includes 

the genera Manda and Planeustomus. 


349 Described as new to science by Schülke (2009). For 

further British records see Lott (2011). 

350 Added by Owen (1997b). 

351 Herman (1986) used species groups in his world 

revision rather than existing subgenera. However, 

subgenera are widely referred to in the literature and 

are included here as listed by Smetana (2004e). 

352 Status follows Assing & Schülke (2012). 

353 The distinctness of this species from B. pallipes is 

not widely recognised outside Britain and is in need 

of review. Several authors including Anderson et al. 

(1997) have followed Pope (1977) in using this name 

in a wider sense to include B. pallipes s. str. when 

referring to the British fauna, as did Herman (1986), 

118 

while continuing to use the name pallipes in listing 

the continental fauna. 

354 

This name was misapplied in part by Horion (1963) 

to B. pallipes. 

355 

All British records were referred to B. annae in Pope 

(1977), but B. pallipes was listed as a separate 

species by Herman (1986) and this name is now 

applied to one of the more frequently encountered 

species of the subgenus in Western Europe. 

356 

Status revised by Schülke (2010). 

357 

Name replacement due to Herman (1986). 

358 


359 

British populations were referred to this subspecies 

by Hammond (2000a). 

360 


361 

Synonymized by Schülke (2011a). 

362 

Described as new to science by Schülke (2011a). 

363 

Added by Owen (1979). 

364 

Added by Lott (2009a). 

365 

Recognised for some time (P.M. Hammond, 

unpublished) but first formally listed in Boyce (1991) 

and Fowles & Boyce (1992). 

366 

Previous British records based on misidentifications, 

but listed by Hammond (2000a) as British on the 

basis of a single museum specimen collected at 

Gretna in 1934. The species was not keyed in the 

monograph by Lott (2009a). 

367 

Status revised by Lott (2009a). 

368 

Lott (2009a) briefly mentions a single male found in 

Leicestershire in 1990, although the species was not 

included in the keys because there is no evidence of 

an established breeding population in Britain. C. 

alutaceus is included in the British list here on the 

basis that the species is putatively a natural vagrant 

(cf. Oxypoda praecox). 

369 

Relegated to a subspecies by Gildenkov (2001). 

370 

Species list based on Lott (2008). 

371 

A reassessment of the identity of this taxon is 

needed following work on the types of related 

species by Makranczy & Schülke (2001). 

372 

Synonymized by Schülke (1998). 

373 


374 

Regarded as a subspecies by Muona (1979a), but 

merely treated as a synonym by Schülke in Assing & 

Schülke (1999). 

375 

Added by Lott (1993c). 

376 

Position as a subfamily within the Staphylinidae, and 

as a sister group to Steninae + Euaesthetinae, 

follows Grebennikov & Newton (2009). Higher 

classification for the Scydmaeninae follows Newton 

& Franz (1998). This work uses an alphabetic listing 

of genera within family groups, and subgenera within 

genera. 

377 

Added by R.G. Booth in Booth (2001). 

378 

Originally Nevraphes, treated as an incorrect original 

spelling by Newton & Franz (1998). 

379 

Treated as a valid subgenus by Newton & Franz 

(1998), but according to Franz & Besuchet (1971) 

Cyrtoscydmus includes exilis (Erichson) which is the 

type species of Stenichnus s.str., so Cyrtoscydmus 

has been synonymized until this problem can be 

resolved. 

380 

Puthz (1967) discussed the lack of compatibility 

between the subgenera listed here and a natural 

phylogenetic classification. However, these

subgenera are well established and continue to be 

widely used as a matter of convenience to break up 

this species-rich genus into manageable groups. 

381 This name was formerly applied to the subgenus 

Metastenus. 

382 Synonymized by Ádám (1987) and followed by 

Herman (2001b). 

383 Originally proposed as a generic name (Ádám, 

1987), but downgraded by Puthz (1999). Species 

groups have been recognised by Puthz (1972) [as 

Hemistenus]. 

384 Considered to be a short-winged form of picipes by 

Puthz (1972) and listed as a subspecies by Smetana 

(2004f), but Kevan & Allen (1961) argued that it was 

distinct. 

385 Junior primary homonym of boops Gravenhorst, 

1806. 

386 

Added by Shirt (1987). 

387 

Added by R.G. Booth in Hodge (2003). 

388 

Added by Lott & Foster (1990). 

389 

Synonymized by Puthz (1974). 

390 

Synonymized by Puthz (1993). 

391 

Species groups have been recognised by Puthz 

(1968). 

392 

Added by Lott & Anderson (2011). 

393 

Tribes and subtribes are arranged according to Lott 

& Anderson (2011). 



396 Added by Lott et al. (2007). 

397 Marked as a junior primary homonym in Pope 

(1977), but without further details. 

398 The subgenera listed here have been accorded 

generic rank by Coiffait (1982a), Assing & Schülke 

(2001, 2007), Herman (2003) and Smetana (2004g), 

but are retained as subgenera here pending a world 

revision of paederine genera. 

399 Junior primary homonym of fulvipenne Turton in 




402 Raised to generic rank by Coiffait (1982a). 

403 Raised to generic rank by Assing in Assing & 

Schülke (2012). 


405 Duff (1995) and Herman (2003) have pointed out 

that Hypomedon and Chloecharis share the same 

type species, because Blackwelder (1939) 

designated Lithocharis debilicornis as the type 

species of Hypomedon. Most authors follow Bordoni 

(1975a) in separating debilicornis from species now 

placed in Sunius, so Hypomedon should be removed 

from synonymy with Sunius. 

406 Added by Drane (1994) [as Chloecharis debilicornis]. 

407 Drugmand (1994) regarded ochracea as a junior 

synonym of tricolor (Fabricius), although the latter 

name is traditionally applied to a species of 

Xantholinus. 

408 Species groups have been recognised by Coiffait 

(1984). 

409 Treated as a genus by Coiffait (1984), but still 

combined with Lithocharis by Poole & Gentili (1996). 

119 

Endnotes 

410 

Raised to generic rank by Coiffait (1982a) and Frank 

(1988), but still combined with Paederus by Poole & 

Gentili (1996). 

411 

Species groups have been allocated to subgenera 

by Coiffait (1984). 

412 

Reidentified by Allen (1995b). 

413 

Junior primary homonym of fragilis Latreille, 1804. 

Synonymized by Herman (2003). 

414 

Added by Skidmore (1988). 

415 

Synonymized by Assing (1997b). 

416 

Genus re-established by Smetana (1995), who 

identified species groups. 

417 


418 Added by Owen (1993b). 

419 Transferred from Gabrius by Smetana (1995). 


(1974) and amended by Smetana (1995). 

421 According to Silfverberg (1991) and Herman 

(2001a), subnigritulus Reitter, 1909 is unavailable 

and subnigritulus Joy is a different species. 

422 Identified as a junior synonym of coxalus by Gusarov 

(1991), which in turn was synonymized by 

Schillhammer (1997). 

423 Synonymized by Bordoni (1975b). 


(1974) and amended by Smetana (1995). 

425 Synonymized by Smetana (1995). 

426 Allen (1970) drew attention to an old British record of 

P. albipes var. alpinus. However, Allen (op. cit.) 

never examined the specimen in question and P. 

alpinus was not listed as British by Pope (1977). 

Regarded as British by Hyman (1994) and Hodge & 

Jones (1995) on the basis of reports of a more 

recent unpublished record from Kent. However, it 

cannot be included in a British Isles checklist on 

currently published evidence. 

427 The name, P. carbonarius, has also been widely 

applied to P. tenuicornis Mulsant & Rey until very 

recently. Therefore, great care is needed when 

interpreting the use of this name. 

428 Synonymized in Coiffait (1974). 

429 Introduced to the British list as a distinct species by 

Smetana (1966), but considered to be a variety by 

Coiffait (1972). 

430 The familiar use of the name immundus for this 

species is based on an early misidentification by 

Gyllenhal (Smetana & Herman, 1999). 

431 Added by Allen & Owen (1997). 

432 Lott & Anderson (2011) consider that some 

specimens standing as varians may be referable to 

P. pseudovarians Strand, A., 1941. 

433 Raised to generic rank by Coiffait (1974). 

434 Status as a separate genus questioned by Smetana 

(1995) and included in Cafius by Hammond (2000a), 

but retained by Herman (2001b) and Smetana 

(2004h). 

435 Added by P.J. Hodge in Booth (2006) and more fully 

by Hance (2007). 

436 Junior primary homonym of binotatus Thunberg, 

1796. 

437 Added by J.A. Owen in Jones (1989). 

438 Whitehead (2002), Smetana (2004h) and 

Solodovnikov in Assing & Schülke (2012) followed 

Israelson (1979) in regarding this taxon as a 

subspecies or junior synonym of H. praevius.


439 

Usage conserved by designation of Q. obscuripennis 

as type species (ICZN Op. 2115). 

440 

Described as new to science by Lott (2010). 

441 

Herman (2001b) has shown that Gravenhorst's 

(1802) usage of this name is a nomen nudum. 

442 

Usage of simplicifrons follows Pope (1977) and 

Herman (2001b), although Bordoni (1976) regarded 

hispanicus as the correct name for the species that 

occurs in Britain. 

443 

Synonymized by Assing (1999a). 

444 

Listed as a separate species by Smetana (2004h). 

445 

Appears as a synonym in Pope (1977) following 

Korge (1963). Synonymy reaffirmed by Assing 

(1999a). 

446 

Placed in the genus Platydracus Thomson by Pope 

(1977), though not by other authors. 

447 

Pope (1977) listed all Ocypus species under 

Staphylinus, unlike other contemporary authors. 

448 

Status follows Smetana & Davies (2000). 

449 

Synonymized by Smetana & Davies (2000). 

450 

Raised to generic rank by Coiffait (1974) but retained 

as subgenus by Smetana & Davies (2000). 

451 

Raised to generic rank by Coiffait (1974) and by 

Smetana & Davies (2000). 

452 

Pope (1977) listed all Tasgius species under 

Staphylinus, unlike other contemporary authors. 

453 

Raised to generic rank by Coiffait (1974) as 

Alapsodus Tottenham, but retained as subgenus by 

Smetana & Davies (2000). 

454 

Junior primary homonym of compressus Fourcroy, 

1785. Synonymized by Steel (1948). 

455 

Regarded as a synonym by Pope (1977) and others 

until re-established as a good species by Lohse 

(1989). Again synonymized by Assing (1996b). 

456 


457 

Raised to generic rank by Coiffait (1972) as 

Phalacrolinus, which was later synonymized by 

Smetana (1982). 

458 

Raised to generic rank by Coiffait (1972). 

459 

Added by Hammond (1982b) [as Phacophallus 

tricolor]. The use of pallidipennis as the name for this 

species follows Bordoni (2002) and subsequent 

authors. 

460 

Replacement due to Drugmand (1994). 

461 

Drugmand’s (1994) interpretation of the identity of 

Fabricius' Staphylinus tricolor, as the species 

traditionally known as Lithocharis ochraceus, was 

not accepted by Herman (2001a). 

462 

The generic placement of the species follows that of 

Zunino (1984) and Löbl & Smetana (2006). The use 

of Anoplotrupes and Trypocopris is accepted 

throughout the European literature (e.g. Baraud, 

1992; Král, 1993; Krell, 1998; Martín-Piera & López- 

Colón, 2000) and the Palaearctic Catalogue (Löbl & 

Smetana, 2006). 

463 

Junior primary homonym of spiniger Gmelin in 

Linnaeus, 1790. The name puncticollis (Malinowsky, 

1811) has been applied to this species (e.g. Martín- 

Piera & López-Colón, 2000). 

464 

The synonmy of Scarabaeus fovetaus Marsham, 

1802 under G. stercorarius (Linnaeus, 1758) has 

been found to be incorrect and it should be placed 

under G. spiniger (Marsham, 1802) (D.J. Mann, 

unpublished). 

120 

465 The nomenclature of the Trogidae follows that of 

Löbl & Smetana (2006). 

466 The nomenclature of the Lucanidae follows that of 


467 The nomenclature follows that of Löbl & Smetana 

(2006). 

468 

The taxonomy of the European members of 

Psammoporus was reviewed by Pittino (2006). P. 

insularis is at present only known from Great Britain. 

469 


470 

The placement of the species follows Dellacasa, 

Bordat & Dellacasa (2001) and Löbl & Smetana 

(2006). 

471 

Authorship of Aphodius follows the recommendation 

of Alonso-Zarazaga & Krell (2011). 

472 

Dellacasa, Bordat & Dellacasa (2001) elevate all of 

the current subgenera under Aphodius to generic 

level. The subgenera under Aphodius are placed in 

alphabetical order, as the taxonomic placement of 

most subgenera is uncertain. 

473 

A. (Coprimorphus) scrutator (Herbst, 1789) is listed 

in Löbl & Smetana (2006) as occurring in Britain, but 

this is believed to be in error. 

474 


475 


476 

Added by Wilson (2001). A species currently 

confused under A. fimetarius (Linnaeus, 1758). 

Dellacasa & Dellacasa (2007) still consider pedellus 

to be synonymous with A. fimetarius (Linnaeus). The 

status of pedellus as a valid species is considered 

valid by Löbl & Smetana (2006) and here. 

477 


478 

Added by D.J. Mann in Hodge (2000). A species 

currently confused under A. sphacelatus (Panzer, 

1798). 

479 


480 

Name suppressed, ICZN Op. 2150. 

481 


482 

The name sticticus (Panzer) has long been used in 

the continental literature (e.g. Dellacasa, 1988; 

Baraud, 1992) and is followed here. 

483 

Added by Angus et al. (2003). 

484 

The generic placement of this species is discussed 

by Mann & Booth (2000). 

485 

Added by D.J. Mann in Hodge (1998). 

486 

The nomenclature follows Zunino (1979) and Löbl & 


487 

O. (Palaeonthophagus) ovatus (Linnaeus, 1758) is 

listed by Löbl & Smetana (2006) as occurring in 'GB', 

but this is believed to be in error as no specimens 

have been found in collections. 

488 

Confirmed as British by Johnson (1991). 

489 

Status revised by Rössner, Schönfeld & Ahrens 

(2010). The true O. vacca is not currently known 

from the British Isles. 

490 

Junior primary homonym of verticicornis Fabricius, 

1775, but see Branco (2002). 

491 

The name farinosa (Linnaeus) is used in the 

continental literature (e.g. Baraud, 1992). However, 

the use of the name philanthus (Füessly) was reestablished 

by Krell (1991). 

492 


493 

Re-identification due to Krell & Rößner (2009).

494 

The nomenclature of the Cetoniinae follows Krajčík 

(1998). 

495 


496 

Name rejected because it was published in a work 

that was not consistently binominal (Krell, 2012). 

497 

Added by Johnson (1997). 

498 


499 

Members of the marginata-group of Elodes were 

transferred to Odeles by Klausnitzer (2004). 

500 

Added by Skidmore (1985b). 

501 

Synonymized by Klausnitzer (1998). 

502 

The higher classification of the Buprestidae is 

currently undergoing a period of change. The most 

recent summary of the higher classification is 

Bellamy (2003). 

503 

Added by Cooter (1992b). 

504 

This enigmatic species was described from 

specimens collected in Dumfriesshire. It is given as a 

synonym of viridis Linnaeus in G.R. Waterhouse's 

1858 Catalogue of British Coleoptera. It is not certain 

if Waterhouse was the originator of this synonymy, 

or if he was following an earlier opinion. The true 

identity of littlei will only be resolved if the type 

material is found. 

505 


506 

Junior primary homonym of biguttata Scopoli, 1763, 

but this name is used by all European authors. 

507 

Added by P.J. Hodge in Booth (2009) and more fully 

by Hodge (2010). 

508 

Added by James (1994). 

509 

Added by Levey (2012). 

510 


511 


512 


513 

Added by Johnson (1978). 

514 

Added by Parry (1980). 

515 

Synonymized by Nelson (1990). 

516 


517 

Split from Heterocerus Fabricius, 1792, by Pacheco 

(1964). 

518 

Added by Mann (2006). 

519 

Added by Mendel, Jeffery & Pledger (2011). 

520 

Added by Speight (1986). 

521 

Junior primary homonym of vittatus Gmelin in 


522 Added by Owen et al. (1985) [as Panspoeus 

guttatus]. 

523 Status revised by Mendel (2004). 

524 Added by Mendel (2002). 

525 Arrangement of the subfamily follows Kazantsev 

(2004). 

526 Added by Allen (1989). 

527 Synonymized by Alexander (2003). 

528 Added by Barclay & Kopetz (2003). 

529 Added by Hammond & Barham (1982) [as Laricobius 

erichsoni]. 

530 The checklist for this family is taken from Peacock 

(1993), slightly simplified. 

531 Added by Peacock (1993). 

532 Added by Peacock (1979). 

533 Added by Shaw (1999). 

534 Added by Thompson (1978). 

535 Added by Adams (1978). 

121 

536 Added by Hinton (1945) but not listed in Pope 

Endnotes 

(1977). 

537 Added by Adams (1988). 


539 Justified emendation of the original oequinoctiale. 

Added by Bellès & Halstead (1985). 

540 British records of P. pilosus Müller, P.W.J. require 

confirmation. 


542 Added by Hodge & Parry (1981). 


544 Added by Mendel & Hatton (2012). 

545 Added by Mendel (1982) [as Hemicoelus nitidus]. 

546 Added by M.V.L. Barclay in Booth (2002a) and more 

fully by Barclay (2005). 


548 Added by M.V.L. Barclay in Booth (2009). 

549 Rejected spelling, ICZN Op. 1810. 

550 Added by Mendel & Owen (1991). 

551 Described as new to science by Bercedo & Arnáiz 

(2010). 

552 Long recognised as a stored-product pest (e.g. 

Mound, 1989) but not included in Pope (1977). 

553 Arrangement and nomenclature of the Cleridae 

generally follow Gerstmeier (1998).The 

Thaneroclerinae were raised to family rank by 

Kolibáč (1992), followed by Gerstmeier (1998), but 

this checklist retains it as a subfamily of Cleridae 

following Lawrence & Newton (1995). 

554 Justified emendation of the original buquet. 

555 Separation of Dasytidae from Melyridae follows Löbl 

& Smetana (2007). 

556 Added by Johnson (1975a) [as Dasytes caeruleus]. 

557 Separation of Malachiidae from Melyridae follows 


558 Added by Welch (2008). 

559 Added by Key (1983). 

560 Added by Allen (1984b). 



563 Synonymized by Biström (1977). 

564 Synonymized by Audisio (1993). 

565 Added by P.M. Hammond & R.G. Booth in Booth 

(2008). 

566 Synonymized by Kurochkin & Kirejtshuk (2006). 

567 Synonymized by Audisio (1993). 

568 Species groups are recognised by Kirk-Spriggs 

(1996). 

569 Synonymized by Bacchus & Kirk-Spriggs (1991). 

570 Cited incorrectly as a synonym of M. lugubris in 

Pope (1977); reinstated by Kirk-Spriggs (1988). 

571 Added by Parry (1990); see also Kirk-Spriggs 

(1992). 

572 Added by R.G. Booth in Hodge (in prep.). 

573 New sense from 1993 onwards, cf. persicus 

(Faldermann). 

574 Synonymized by Silfverberg (1979) . 

575 Omitted from Pope (1977) in error; reinstated by 

Booth (1986). 

576 Added by James (2011). 

577 Added by Prance (2001).


578 Nomenclature for this family largely follows Löbl & 

Smetana (2007) except that the treatment of 

Cyanostolus follows Peacock (1977). 


580 Added by King & Fielding (1989). 

581 Added by R.G. Booth in Booth (2001). 


583 Nomenclature for this family follows Löbl & Smetana 

(2007). 


585 Added by Johnson (1988). 

586 Synonymized by Johnson (2012a). 


588 Synonymized by Johnson (1992a). 

589 Added by B. Levey & P.M. Pavett in Hodge (1999). 

See also Johnson (2002). 

590 Added by R.G. Booth & A.C. Galsworthy in Booth 

(2008), 

591 Arrangement of subfamilies follows Leschen (2003). 

592 

Added by Hammond (2007c). 

593 


594 

Added by Eccles & Bowestead (1987). 

595 

Names and arrangement of subfamilies follows 

Tomaszewska (2000). 

596 

Added by Hawkins (2001). 

597 

Added by D.A. Coleman per R.G. Booth in Hodge 

(2000). 

598 

Added by R.G. Booth & A. Salisbury in Booth (2004). 

599 

Junior primary homonym of arcuatus Fabricius, 

1787. 

600 

Added by Denton (2004). 

601 

Added by R.M. Lyszkowski in Hodge & Jones 

(1995). 

602 

Added by D.S. Hackett in Booth (2002a). 

603 Possibly a valid species. 

604 Added by Constantine & Majerus (1994). 

605 Added by Majerus et al. (1997). See also Muggleton 

(1999). 

606 Added by Majerus & Roy (2005). 

607 Added by I.S. Menzies in Hodge (1999) and more 

fully by Menzies & Spooner (2000). The generic 

placement of argus is probably not correct. 

608 This checklist is based upon that in Bowestead 

(1999). 


(2007). For identification characters see Bowestead 

(2002). 

610 Status revised by Bowestead (1999). Name 

resurrected by Bowestead for the species long 

known as Orthoperus mundus. 

611 Synonymized by Bowestead (1999). 

612 Nomenclature for this family follows Löbl & Smetana 

(2007). 

613 Added by Levey (1997). 


615 Reidentified by Krell et al. (2005). 

616 See Johnson (2012b). 

617 Added by Lyszkowski, Owen & Taylor (1992) [as 

Corticaria abietorum]. 




(2007) [as Migneauxia orientalis]. 

122 

621 


622 

Added by Dorning, Halstead & Hammond (2012). 

623 

Added by Harrison (1996). 

624 

Added by Allen & Booth (2008). 

625 

Hallomeninae was transferred from Melandryidae to 

Tetratomidae by Nikitsky (1998). 

626 

Junior primary homonym of binotatus Gmelin in 


627 Justified emendation of the original misspelling 

Desmaretsii. 

628 Junior primary homonym of quercinus Gmelin in 


629 Reidentified by Batten (1986). 

630 Added by Batten (1986). 

631 Added by Allen (1986) [as Mordellistena 

parvuloides]; see also Owen (1999). 

632 Synonymized by Horák (1996). 

633 Described as new to science by Allen (1999). 

Synonymized by Levey (2002). 

634 Added by Batten (1986). The status of M. 

pseudopumila needs further study. 

635 Added by Cooter (1991) [as Mordellistena 

pygmeola]. The status of M. pygmaeola needs 

further study. 

636 Added by Levey (1999). 


638 Described as new to science by Allen (1995c). 

Synonymized by Levey (2002). 


640 Added by Mendel & Owen (1987) [as Cicones 

undata]. 


642 Confirmed as a British species by Hammond 

(2007a). 

643 Geoffroy's name is suppressed for purposes of the 

Principle of Priority but not for those of the Principle 

of Homonymy (ICZN Op. 1754). 

644 Added by Skidmore & Hunter (1980). 

645 Added by Allen (1988b). 

646 Added by Whitehead (1992). 

647 Junior primary homonym of rugosus Thunberg, 

1781. 


649 Junior primary homonym of ater De Geer, 1774. 

650 Junior primary homonym of planirostris Piller & 

Mitterpacher, 1783. 

651 Genera of Anthicidae follow Chandler, Nardi & 

Telnov (2004). See also Telnov (2010). 

652 Synonymized by Chandler et al. (2004). 

653 Junior primary homonym of constrictus Say, 1826. 


655 An application is being prepared to give maculata 

(Geoffroy in Fourcroy, 1785) precedence over 

melanopa Forster, 1771. 

656 Levey (2002) suggested that A. thoracica and A. 

septentrionalis belong to a single variable species. 

657 Levey (1996) synonymized A. schilskyana with 

septentrionalis. 

658 Levey (2003) revised the European species of this 

subgenus. 

659 Csiki (1915) gives fuscescens as a junior synonym 

of A. thoracica (Linnaeus, 1758), however all the 

specimens standing over this name in the Stephens

collection in the Natural History Museum, London 

are costai Emery. For reasons of stability it is not 

proposed to invoke the principle of priority and it is 

likely that the prevailing usage of the name costai 

would be validated under article 23.9 of the Fourth 

Edition of the International Code of Zoological 

Nomenclature. 

660 

This checklist is based on that of Rejzek (2004), 

which contains details of the higher classification 

used. 

661 

Synonymized by Uhthoff-Kaufmann (1989). 

662 


663 

Added by Mendel & Barclay (2008). 

664 


665 

Established introduction in the Reading, Berkshire, 

area in the late 19th and early 20th Centuries 

(Barclay, 2003b). 

666 

Described as new to science by Wallin, Nylander & 

Kvamme (2009). 

667 

Added by Harrison (1992a, 1992b). 

668 

Synonymized by Petitpierre (2000). 

669 


670 


671 

Added by Hammond & Harvey (2011). 

672 

Added by D. Hance & P.J. Hodge in Cox (2001). 

673 

Added by Cox (2001). 

674 

Added (reinstated) by Aldridge & Pope (1986). 

675 Added by Hodge (1997). 

676 Oulema melanopus sensu stricto added by Cox 

(1995a). 

677 Added by Booth (1994b) [as Oulema duftschmidi]. 

678 Synonymized by Monrós (1956), who showed that 

the beetle known to European coleopterists as 

Lamprosoma concolor Sturm is not congeneric with 

L. bicolor Kirby, the type species of Lamprosoma 

Kirby, and that the valid generic name for the 

European species is Oomorphus Curtis, 1831. 

679 Added (reinstated?) by Kendall (1981). 

680 Subgenera are recognised by European authors. 

Species-level nomenclature for this genus follows 

Bieńkowski (2001). 

681 First recorded indoors by Johnson (1963), then as a 

possibly established introduction by Halstead (1996). 

682 Justified emendation of the original misspelling 

bankii. 

683 A breeding population was documented by 

Salisbury, Malumphy & Halstead (2012). 

684 See also Cox (2000). 

685 Status revised by Cox (2000). 

686 Synonymized by Shute in Cox (2000). 

687 Added as an accidental introduction by Smith (1990), 

then recorded as probably wild by Buckland & 

Skidmore (1999). 

688 Cox (2007) describes morphological variation within 

British and Irish G. tanaceti and discusses the 

possibility that another Galeruca species may be 

present. 

689 Added by Ostojá-Starzewski (2005). 

690 Added by Johnson & Booth (2004). 

691 See Cox (2000) for a discussion of the identity of A. 

atrovirens of British authors. 

692 Added by Sinclair & Hutchins (2009). 

693 Junior primary homonym of dorsalis Fabricius, 1777. 

123 

Endnotes 

694 

Synonymized by Döberl (1987); see also Allen 

(1993). 

695 

Added by Booth (1994a). 

696 

Synonymized by Booth (1998). 

697 


698 


699 

Added by P.M. Hammond in Cox (2000). 

700 

Replacement name due to Warchałowski (1995). 

701 

Synonymy follows Cox (2007). 

702 

Synonymized by Arnold (1990). 

703 

Junior primary homonym of aurata Fabricius, 1775. 

704 

C. aridula (Gyllenhal) and C. conducta (Motschulsky) 

have been deleted from the list, as all records are 

considered to be due to misidentification or 

mislabelling. See Cox (2000). 

705 

Added by Booth & Owen (1997). 

706 

Synonymized by Booth & Owen (1997). 

707 Added by Cox (1995b). 

708 Synonymized by Döberl (1995). 

709 The checklists for the families of Curculionoidea 

follow Morris (2003), with more recent amendments 

referred to in relevant endnotes. 

710 Following Thompson (1992) and others. 

711 See Alonso-Zaraga & Lyal (1999) for a discussion of 

this family and the nomenclature of its constituents. 

712 Subfamily Nemonychinae contains only Nemonyx 

(not British). 

713 Rhinomacer, and family-group names based 

thereon, have been effectively suppressed (ICZN 

Op. 1754). 

714 Given precedence over Choragidae (ICZN Op. 

1756). Nomenclature and arrangement follow M. 

Trýzna & B.D. Valentine in Löbl & Smetana (2011). 

715 See Alonso-Zaraga & Lyal (1999). 

716 Generic assignment per Löbl & Smetana (2011). 

717 Placed as a subgenus of Tropideres in Pope (1977). 

718 Given family rank by Crowson (1984) and so treated 

by Morris (1990, 1995). Though currently regarded 

as a subfamily of Anthribidae, as here, this may be 

controversial. 

719 (Re)discovered in Britain by Hyman (1987b) [as 

Bruchella rufipes]; see also Morris (1990). 

720 British records of L. sericeus (Herbst) require 

confirmation; see Morris & Johnson (2005). 

721 See Morris (1990). 

722 Generic status follows Löbl & Smetana (2011). 

723 Not now regarded as a subfamily of Brentidae (cf. 

Gønget, 1997). Nomenclature and arrangement 

follow M.A. Alonso-Zarazaga in Löbl & Smetana 

(2011). 

724 Synonymized by Thompson & Alonso-Zarazaga 

(1988). 

725 Added by Fowles & Morris (1994). 

726 Added by P.J. Hodge in Booth (2009) and more fully 

in Hodge (2011). 

727 Added by Morris & Péricart (1988). 

728 Taxonomy of Ceratapion follows Alonso-Zarazaga 

(1991) and Wanat (1995); see also Morris (1993a). 

729 Synonymized by Wanat (1995); see also Morris & 

Booth (1997). 

730 Incorrect original spelling, established by Wanat 

(1995); see also Morris & Booth (1997). 

731 Junior primary homonym of laevigatum Fabricius, 

1792. Species-group name unchanged from Pope


(1977), restored from Morris (1990); see also Morris 

(1993a). 

732 Added by Foster, Morris & Whitehead (2001). 

733 Added by Jones (2006). 

734 Taxonomy of group clarified by Dieckmann (1976). 

735 Wrongly placed in Apion (Eutrichapion) in Pope 

(1977). 

736 Added by Parry (1982b) [as Apion intermedium]. 

737 Status as a good species doubtful, but to be 

investigated. 


739 Included in Cossoninae by Pope (1977). 



742 Nomenclature and arrangement follow R. Caldara in 


743 This synonymy (Alonso-Zarazaga & Lyal, 1999) was 

overlooked by Morris (2002), who incorrectly 

separated aethiops from the other species. It is 

bimaculatus which should be placed in a separate 

genus, following recent research which is continuing. 

For a radically different treatment of Notaris sensu 

lato see for example Abbazzi & Osella (1992). 

744 Junior primary homonym of scirpi Rossi, 1790. 

745 The taxonomy and nomenclature of the species in 

this genus were revised by Thompson (2006). 

746 Added by Booth (1993). The synonymy was 

established by Booth (2002b). 

747 Treated as a subfamily of Curculionidae (Gonatoceri) 

by many authors, including Osella (1977) and 

Lawrence & Newton (1995). 

748 Added as a casual introduction by Williams (1968), 

then recorded as an established introduction by 

Thompson (1995) and Owen (1995) [as 

Raymondionymus marqueti]. See also Owen 

(1997a). 

749 A. britannus Desbrochers is here treated as a non- 

established introduction. 

750 Often treated as a full genus. 

751 Added by Read (1981) [as Furcipus rectirostris]. 

752 Added by R.G. Booth in Hodge (in prep.). 

753 British records of E. scanicus (Paykull) require 

confirmation. 

754 Subgenera, used by Alonso-Zarazaga & Lyal (1999) 

and widely in the Continental literature, are not 

employed here because both Dieckmann (1986) (the 

German fauna) and O'Brien (1970) (the Nearctic 

species) found them to be unsustainable. They have 

not been used to any extent in the British literature. 

755 Junior primary homonym of affinis (Schrank, 1781). 

Some Continental literature already uses 

edoughensis for this species. 

756 Synonymized by Dieckmann (1979); see also Morris 

& Booth (1997). 

757 The treatment of this tribe follows the phylogenetic 

analysis of Caldara (2001). 

758 Owen (1993a) is followed in believing that this taxon, 

at least in Britain, is to be referred to graminis. This 

was the opinion of the late Dr L. Dieckmann, who 

examined some British examples of putative degorsi 

(M.G. Morris in litt.); see also Owen (1988). It is 

uncertain whether a good species, degorsi, is part of 

the European fauna, but it seems doubtful. 

759 G. lloydi Donisthorpe is now treated as a synonym of 

Sibinia pyrrhodactyla (Marsham). 

124 

760 

Transferred from Gymnetron by Caldara (2001). 

761 

Treatment of Rhynchaenus and Orchestes as 

different genera follows the detailed study by Kojima 

& Morimoto (1996). 

762 

Added (reinstated?) by Parry (1981) [as 

Rhynchaenus populi]. 

763 

Added by Allen (1988a) [as Rhynchaenus 

calceatus]. See also Mendel (1994), Morris & Booth 

(1997), and Morris & Owen (1997). 

764 

Synonymized by Morris & Booth (1997). 

765 

See the conclusions of Thompson (1994). 

766 

Added by Hammond (2000b). 

767 

Junior primary homonym of lonicerae de 

Razoumovsky, 1789. 

768 

Added by Tempère (1982) [as Rhynchaenus 

pseudostigma]. See also Morris (1987, 1993b). 

769 

S. pellucens (Scopoli) has been omitted from the 

current list. Its history as a 'British' species is in 

Fowler (1891), but its status in Britain is thought to 

be dubious. See Morris & Johnson (2005) for details. 

770 

Correctly ‘primitus’, but treated as adjectival in form 

by later revisers and in recent European checklists; 

see Morris & Booth (1997). 

771 

Synonymized by Caldara (1990a); see also Morris & 

Booth (1997). 

772 

Synonymized by Morris (1989). 

773 

Not regarded as a valid genus or subgenus by 

Caldara (1990b). 

774 Added by R.G. Booth in Hodge (1999). 

775 Rediscovered as a resident species by Morris 

(2004a, 2004b). 

776 Synonymized by Caldara (1983, 1990a); see also 

Allen (1984a) and Morris & Booth (1997). 

777 The subgenera of Bagous used here have no 

phylogenetic significance and are unsustainable on a 

world basis (Caldara & O'Brien, 1998; Alonso- 

Zarazaga & Lyal, 1999); they are retained here 

because of their familiarity and use in the 

identification of species (Dieckmann, 1964; Morris, 

2002). 

778 Synonymized by Caldara & O'Brien (1998). 

779 Synonymized by Allen (1992b); see also Morris & 

Booth (1997). 

780 Synonymized by Caldara & O’Brien (1994); see also 

Morris & Booth (1997). 

781 Nomenclature and arrangement follow J. Prena in 


782 B. chlorizans Germar is noted by Fowler (1891) but 

has not been confirmed as British and is omitted 

from the current list. See Morris & Johnson (2005) 

for details. 

783 Taxonomy and nomenclature of the European taxa 

clarified by Dieckmann (1991) and briefly discussed 

by Morris & Booth (1997). 

784 The checklist for this subfamily follows Morris 

(1991a) with the necessary changes for an 

alphabetical arrangement and incorporating 

important nomenclatural amendments by Colonnelli 

(1998, 2004). 

785 Subgenera of Ceutorhynchus are not used by 

Alonso-Zarazaga & Lyal (1999), probably do not 

express phylogenetic relationships, and are mostly 

unfamiliar to British workers. Consequently, they are 

not included here.

786 Junior primary homonym of assimilis Fabricius, 

1775. 

787 Synonymized by Colonnelli (1998). See also Morris 

& Booth (1997). 

788 Added to the British list by Owen (1990a), and to the 

Irish list by Morris (1992). 

789 Junior primary homonym of contractus Fourcroy, 

1785. 

790 Synonymized by Colonnelli (1998). 

791 Treated as a full genus in Alonso-Zarazaga & Lyal 

(1999). 

792 Rediscovered as a resident species by Fowles & 

Morris (1999). 

793 Synonymized by Colonnelli (1990). 

794 Rejected alternative original spelling. 

795 First recognised in Britain by J. Parry. First published 

record appears in Atty (1983). See Owen (1994d). 

796 Retained over commari (Panzer, 1795) (Colonnelli, 

1998); Alonso-Zarazaga & Lyal (2002) have shown 

that Colonnelli's interpretation is based on an error of 

dating. 

797 Added by Johnson (1982a) [as Phytobius olssoni]. 

798 Added by Morris (1991b) [as Phytobius zumpti]. 

799 Added by Hammond (2008). 

800 Added by Turner (2011) [as Pentarthrum elumbe]. 

801 Added by Allen & Turner (2012). 

802 Added by Welch (1990a). 

803 Name conserved, ICZN Op. 1655; see also Morris & 

Booth (1997). 

804 British records of P. gracilis (Rosenhauer) require 

confirmation. 

805 

Revised by Stüben et al. (2003). 

806 

Added by Clemons (1983). 

807 

Added by Denton (2005). 

808 

Added by M.V.L. Barclay in Hodge (2007). 

809 

Synonymized by Dieckmann (1980). 

810 

The genus Philopedon must be treated as neuter; 

see Morris & Booth (1997). 

811 

Replacement name; see Morris (1997). 

812 

British records of O. coecus Germar, 1824 (= niger 

(Fabricius, 1775) non (Drury, 1773)) require 

confirmation. 

813 


814 

Added by M.V.L. Barclay in Hodge (2000) and more 

fully by Barclay (2003a). 

815 

Added by Hyman (1987a). 

816 

The species was first detected by MAFF in the mid- 

1980s but was formally added by Morris (1997). 

817 


818 

Re-identification due to Barclay (2009). 

819 

Added by N.F. Heal; see Morris (1997). 

820 

Added by S.G. Cole in Booth (2002a) and more fully 

by Barclay (2003a). 

821 Generally considered to be a good species distinct 

from P. pyri but not accepted as such by Pesarini 

(1981). 

822 Added by Plant, Morris & Heal (2006). 

823 Added by Denton (2005). 

824 P. prasinus (Olivier), included by Pope (1977), is 

omitted as being very doubtfuly British (Morris, 

1997). See also Morris & Johnson (2005). 

825 cf. Morris (1997). 

125 

Endnotes 

826 Synonymized by Roudier (1963), who established 

that the true Polydrusus chrysomela is confined to 

Portugal; see also Morris (1996). 

827 Eusomus ovulum (Germar) is omitted; see Morris 

(1997) and Morris & Johnson (2005). 

828 Classification of the Sitonini follows Velázquez et al. 

(2007). 

829 Recently confirmed as an established and probably 

native species (P.M. Hammond, pers. comm.). 

830 Added by Cunningham (in prep.). 

831 Arrangement of this tribe follows Morris (2011). 

832 Detailed work by Piper et al. (2001) has failed to 

confirm this taxon as a good species. 

833 Shown to be distinct from T. bifoveolatus by Jermiin 

et al. (1991). Added to the British list by Harrison 

(1993). 

834 Synonymized by Borovec (1991); see also Morris & 

Booth (1997). 

835 Unjustified replacement name. 

836 The single British record of L. (Lixochelus) filiformis 

(Fabricius) [listed as L. elongatus (Goeze) in Pope 

(1977)] is of dubious provenance and the species 

has been omitted from the current list. See Mann, 

Hancock & Morris (2005) and Morris & Johnson 

(2005) for details. 


838 Tempère (1979) established that L. troglodytes is a 

good species and not a subspecies of L. pyrenaeum; 

see also Morris & Booth (1997). 

839 The genera in this group are not included in Alonso- 

Zarazaga & Lyal (1999), but it is regarded by them 

as a subfamily of Curculionidae. Nomenclature and 

arrangement follow M. Knížek in Löbl & Smetana 

(2011). 

840 Only two tribes of Scolytinae are recognised here (cf. 

Silfverberg, 1992). Tribes and subtribes follow Wood 

(1982), with adjustments for that author's view that 

Scolytinae have family (not subfamily) rank. 

841 Added by Atkins, O’Callaghan & Kirby (1981). 


843 Confirmed as a British breeding species by 

Lyszkowski (1993). See also Owen (1994a), who 

showed that some earlier records were inaccurate. 

844 Added by Winter (1990); see also Owen (1993a). 


846 Added by Telfer (2007a). 

847 Synonymized by Wood & Bright (1992). 

848 Synonymy follows Pope (1977) and Owen (1993a), 

but see Silfverberg (1992). 

849 Status of Hylesinus orni Fuchs, 1906 follows Löbl & 


850 Synonymized with Leperisinus varius in error by 

Pope (1977). 

851 Junior primary homonym of ater Rossi, 1792. 

Possibly a misidentification of ater Fabricius, 1792. 

852 Replacement name for ater Paykull, 1800 non Rossi, 

1792. 

853 Added by Cooter (1982, 1983). See also King & 

Fielding (1989). 

854 Added as a casual introduction by Winter (1991) [as 

P. bicolor], then recorded as apparently established 

by Winter (1998). 

855 Synonymy follows Schedl (1981). 

856 Added by Chuter (2010).


857 Genera in this group are not included in Alonso- 

Zarazaga & Lyal (1999), who regard it as a family 

separate from Curculionidae. The treatment in the 

current list follows Pope (1977). 

126

Species found as fossils in Quaternary sediments 

P.I. Buckland & P.C. Buckland 

F 

127 


or over 40 years the study of fossil insects has provided a significant contribution 

to our understanding of environmental and climate change. Beetles, by way of 

their frequency as fossils in Quaternary terrestrial and freshwater sediments, relative 

ease of identification as fossils, niche specificity and apparent morphological and 

physiological stability over the Quaternary and beyond 1 , have been the most studied 

group. Their study has been particularly instrumental in elucidating climate change 

over the past 250,000 years and in defining the rapidity of climatic transitions during 

the last glaciation and into the present interglacial, the Holocene. Although a relatively 

small group, palaeoentomologists continue to provide an important contribution to 

Quaternary science and archaeology, as well as to the broader issues of biodiversity, 

landscape change and human impact. What follows is a list of those species which 

have been found in Quaternary sediments in the British Isles, but which have not been 

recorded as native in historic times. This includes both species that have Holocene 

records and are now extinct in Britain and species that have only pre-Holocene 

records, and thus no Red Data Book status. The status of a number of species is 

inevitably uncertain with some previously known only as fossils being discovered 

either as residents or introductions 2 , and others having doubtfully extant breeding 

populations. 

All data are extracted from the Bugs Coleopteran Ecology Package (BugsCEP), a 

database of Coleopteran ecology, distribution and fossil records, which is available for 

free download from http://www.bugscep.com. BugsCEP is currently being developed 

and extended by Phil Buckland (Umeå University, Sweden, e-mail phil.buckland 

@arke.umu.se) and Paul Buckland (Sheffield, UK, e-mail paul.buckland 

@bugscep.com), and provides a variety of tools to aid the entomologist and 

palaeoentomologist in teaching and research. These include, among other things, 

complex habitat queries, graphical ecology summaries, and palaeotemperature 

reconstruction. Please see the website for details, and also access to a full 

bibliography (Qbib) of relevant papers. A mapping interface, and the ability to crossquery 

with other proxy data such as plant macrofossils and geochemistry, is being 

developed as part of the Strategic Environmental Archaeology Database (SEAD, 

http://www.sead.se). 

Key 

Q = (prior to Marine Isotope Stage 5e / >~130kaBP). 

I = last interglacial (=Ipswichian / Eemian / Marine Isotope Stage 5e / ~130-115kaBP) 

G = last glacial (=Devensian / Weichselian / Marine Isotope Stages 5d-2 / ~115- 

15kaBP) 

LG = late glacial (= Pollen Zones I-III / ~13.5-10kaBP) 

H = Holocene 

* = species with historical records but probably only as non-established introductions 

Note: records described as G or LG do not necessarily equate with cold (stadial) 

phases and may indicate conditions at least as warm as the present day, but in an 

open (=interstadial) landscape. 

When publishing data from this list, or the BugsCEP computer package, please use 

the full citation given for Buckland & Buckland (2006) in "Bibliography" on page 136. 

Further information on using BugsCEP and examples can be found in Buckland (2007) 

and on the project website. 

1 Most recently on this see Fikáćek, Prokin & Angus (2011). 

2 See for example Mendel, Jeffrey & Pledger (2011).

Species found as fossils in Quaternary sediments Checklist of Beetles of the British Isles, 2012 Edition 

Family GYRINIDAE Latreille, 1810 

Gyrinus colymbus Erichson H 

Family DYTISCIDAE Leach, 1815 

Agabus serricornis (Paykull) G 

Bidessus grossepunctatus Vorbringer H 

Colymbetes dolabratus (Paykull) Q;G;LG 

Colymbetes paykulli Erichson G;LG 

Colymbetes striatus (Linnaeus) LG;H 

Hydroporus lapponum (Gyllenhal) LG 

Hydroporus nigellus Mannerheim LG 

Hydroporus notabilis LeConte LG 

Hygrotus (Coelambus) unguicularis Crotch LG 

Ilybius angustior (Gyllenhal) LG 

Ilybius vittiger (Gyllenhal) G 

Family RHYSODIDAE Laporte, 1840 

Rhysodes sulcatus (Fabricius) Q;I;H 


*Abax parallelus (Duftschmid) Q 

Agonum consimile (Gyllenhal) G;LG 

Agonum exaratum (Mannerheim) Q;G 

Amara erratica (Duftschmid) G 

Amara interstitialis Dejean G 

Amara municipalis (Duftschmid) Q;G 

Amara torrida (Panzer) G;LG 

Asaphidion cyanicorne (Pandellé) LG 

*Bembidion callosum Küster LG 

Bembidion dauricum (Motschulsky) G;LG 

Bembidion difficile (Motschulsky) Q;G;LG 

Bembidion elongatum Dejean I;Q 

Bembidion fellmanni (Mannerheim) Q;G;LG 

Bembidion grisvardi Dewailly / ibericum Piochard de la 

Brulerie LG 

Bembidion hastii Sahlberg, C.R. Q;G;LG 

Bembidion hyperboraeum Munster G;LG 

Bembidion lapponicum Zetterstedt G;LG 

Bembidion mckinleyi Fall G;LG 

Bembidion obscurellum (Motschulsky) G;LG 

Bembidion petrosum Gebler G;LG 

Bembidion transparens (Gebler) Q;G;LG 

Bembidion velox (Linnaeus) Q;LG 

*Carabus (Autocarabus) cancellatus Illiger G 

Carabus (Carabus) menetriesi Hummel G 

Carabus (Homoeocarabus) maeander Fischer von 

Waldheim G 

Carabus (Oreocarabus) hortensis Linnaeus G 

*Carabus (Tomocarabus) convexus Fabricius LG 

Chlaenius costulatus Motschulsky Q;G;LG 

Chlaenius festivus (Panzer) I 

Chlaenius quadrisulcatus (Paykull) I 

Chlaenius sulcicollis (Paykull) Q;I;H 

Chlaenius variegatus (Geoffroy in Fourcroy) I 

Cymindis angularis Gyllenhal G;LG 

Cymindis humeralis (Geoffroy in Fourcroy) LG 

Diacheila arctica (Gyllenhal) G;LG 

Diacheila polita (Faldermann) Q;G;LG 

Dyschirius nigricornis Motschulsky 

= septentrionis Munster Q;G;LG 

Oodes gracilis Villa & Villa Q;I;H 

Pterostichus blandulus Miller G;LG 

Pterostichus brevicornis (Kirby) Q 

Pterostichus kokeilii Miller G 

Pterostichus magus Mannerheim LG 

128 

Pterostichus middendorffi (Sahlberg, J.R.) Q 

Syntomus parallelus (Ball) LG 

Trechus amplicollis Fairmaire Q 

Family HYDROCHIDAE Thomson, C.G., 1859 

Hydrochus flavipennis Küster G 

Family HELOPHORIDAE Leach, 1815 

Helophorus aquaticus (Linnaeus) s. str. G 

Helophorus arcticus Brown Q 

Helophorus aspericollis Angus G 

Helophorus discrepans Rey G 

Helophorus glacialis Villa G;LG 

Helophorus lapponicus Thomson, C.G. G;LG 

Helophorus mongoliensis Angus G 

Helophorus oblongus LeConte Q;LG 

Helophorus obscurellus Poppius Q;G;LG 

Helophorus orientalis Motschulsky G 

Helophorus pallidus Gebler G 

Helophorus praenanus (Łomnicki) G;LG 

Helophorus sibiricus (Motschulsky) G;LG 

Helophorus splendidus Sahlberg, J. G;LG 


Hydrobius arcticus Kuwert G 

Laccobius decorus (Gyllenhal) G 

Family HISTERIDAE Gyllenhal, 1808 

Chetabraeus globulus (Creutzer) Q;G 

Hister funestus Erichson G 

Margarinotus terricola Germar LG 

Family HYDRAENIDAE Mulsant, 1844 

Hydraena coopei Angus Q 

Hydraena cf. latebricola Jäch G 

Ochthebius foveolatus Germar LG 

Ochthebius kaninensis Poppius G 

Ochthebius pedicularius Kuwert LG 

Family SILPHIDAE Latreille, 1806 

Ptomascopus zhangla Háva, Schneider & Růžička G 

Thanatophilus lapponicus (Herbst) LG 


Acidota quadrata (Zetterstedt) Q;G;LG;H 

Anotylus gibbulus (Eppelsheim) Q;G 

Anotylus mendus Herman Q 

Anotylus tetratoma Czwalina Q 

Anthophagus bicornis (Block) LG 

Aploderus caesus (Erichson) Q;G 

Arpedium quadrum (Gravenhorst) Q;LG;H 

Batrisus formicarius Aubé H 

Bledius cribricollis Heer, 1839 

Bledius littoralis Heer Q;G 

Boreaphilus henningianus Sahlberg, C.R. Q;G;LG 

Eucnecosum cf. tenue LeConte LG 

Eudectus giraudi Redtenbacher LG 

Geodromicus kunzei Heer Q;LG 

Holoboreaphilus nordenskioldi (Mäklin) Q;G;LG 

Leptobium gracile (Gravenhorst) Q 

Micropeplus hoogendorni Matthews 

=dokuchaevi (Ryabukhin) Q 

Ocypus picipennis (Fabricius) LG 

Olophrum boreale (Paykull) Q;G;LG

Olophrum rotundicolle (Sahlberg, C.R.) G;LG 

Philonthus linkei Solsky G 

Pycnoglypta lurida (Gyllenhal) Q;G;LG 

Stenus plicipennis (Casey) LG 

Tachinus arcticus (Motschulsky) G 

Tachinus caelatus Ullrich Q;G 

Tachinus jacuticus Poppius Q;G 

Tachinus marginatus Gyllenhal G 


Aphodius (Acrossus) carpetanus Gräells Q;I 

Aphodius (Alocoderus) holdereri Reitter G 

Aphodius (Amidorus) obscurus (Fabricius) LG 

Aphodius (Calaphodius) bonvouloiri Harold G 

Aphodius (Chilothorax) jacobsoni Koshantschikov G 

Aphodius (Esymus) quadriguttatus (Herbst) H 

Aphodius (Neagolius) montivagus Erichson LG 

*Aphodius (Nialus) varians Duftschmid H 

Aphodius (Phaeaphodius) costalis Gebler G 

Aphodius (Teuchestes) brachysomus Solsky G 

Ataenius horticola Harold Q 

Caccobius histeroides (Ménétries) I 

Caccobius schreberi (Linnaeus) Q;I;H 

Drepanocerus sp. I 

Euoniticellus fulvus (Goeze) I 

Onthophagus furcatus (Fabricius) I 

Onthophagus gibbulus (Pallas) I;G 

Onthophagus massai Baraud I 

Onthophagus opacicollis Reitter I 

Osmoderma eremita (Scopoli) Q 

Rhyssemus algiricus Lucas Q 

Triodonta sp. Q 

Valgus hemipterus (Linnaeus) Q;I;H 

Family BUPRESTIDAE Leach, 1815 

Buprestis rustica Linnaeus H 

Family BYRRHIDAE Latreille, 1804 

Curimopsis cyclolepidia (Munster) G;LG 

Simplocaria metallica (Sturm) Q;G;LG 

Family ELMIDAE Curtis, 1830 

Dupophilus brevis Mulsant & Rey Q 

Esolus pygmaeus (Müller, P.W.J.) Q;H 

Limnius opacus Müller, P.W.J. Q 

Stenelmis consobrina Dufour I;H 

Family LIMNICHIDAE Erichson, 1846 

Pelochares versicolor (Waltl) Q;I 

Family HETEROCERIDAE MacLeay, 1825 

Heterocerus intermedius Kiesenwetter G;LG 

Family EUCNEMIDAE Eschscholtz, 1829 

Dromaeolus barnabita (Villa) H 


Hypnoidus rivularius (Gyllenhal) G;LG 

Idolus picipennis (Bach) Q 

Porthmidius austriacus (Schrank) H 

Family DERMESTIDAE Latreille, 1804 

Dermestes laniarius Illiger I;H 

129 



Stagetus borealis Israelson H 

Family TROGOSSITIDAE Latreille, 1802 

Peltis grossa (Linnaeus) Q;H 

Temnochila caerulea (Olivier) H 

Tenebroides fuscus (Goeze) H 


Opetiopalpus scutellaris (Panzer) Q;I;G 

Family KATERETIDAE Kirby, 1837 

Heterhelus scutellaris (Heer) Q 


Airaphilus elongatus (Gyllenhal) Q;LG;H 

Family LAEMOPHLOEIDAE Ganglbauer, 1899 

Cryptolestes corticinus (Erichson) H 

Notolaemus castaneus (Erichson) H 

Family EROTYLIDAE Latreille, 1802 

Leucohimatium sp. H 

Family BOTHRIDERIDAE Erichson, 1845 

Bothrideres bipunctatus (Gmelin) I;H 

Family ENDOMYCHIDAE Leach, 1815 

Mycetina cruciata (Schaller) H 

Family COCCINELLIDAE Latreille, 1807 

Anisosticta strigata (Thunberg) G 

Ceratomegilla ulkei Crotch G;LG 

Hippodamia arctica (Schneider, D.H.) Q;G;LG 

Hippodamia baratovskii Semenov & Dobzhansky G 

Hippodamia septemmaculata (De Geer) Q 

Nephus bipunctatus (Kugelann) G;LG 

Family CIIDAE Leach, 1819 

Ropalodontus baudueri Abeille de Perrin H 

Family COLYDIIDAE Billberg, 1820 

Pycnomerus terebrans (Olivier) Q;I;H 


Diaclina fagi (Panzer) Q 

Family PROSTOMIDAE Thomson, C.G., 1859 

Prostomis mandibularis (Fabricius) H 


Anthicus ater (Panzer) G;LG 

Cordicomus gracilis (Panzer) H 

Cordicomus sellatus (Panzer) I 


Chaetocnema obesa (Boieldieu) G 

Chrysolina limbata (Fabricius) Q;G 

Chrysomela collaris Linnaeus G;LG 

Chrysomela septentrionalis (Ménétries) G;LG 

Donacia polita Kunze Q 

Entomoscelis adonidis (Pallas) G;LG

Species found as fossils in Quaternary sediments Checklist of Beetles of the British Isles, 2012 Edition 

Neocrepidodera interpunctata (Motschulsky) G 

Pachnephorus tesselatus (Duftschmid) Q 

Family RHYNCHITIDAE Gistel, 1848 

Rhynchites pubescens (Fabricius) G 


Aphanommata filum (Mulsant & Rey) I 

Aulacobaris violaceomicans (Solari) LG 

Cathormiocerus curviscapus Seidlitz G 

Cathormiocerus validiscapus Rouget Q;G;H 

Cyphocleonus trisulcatus (Herbst) H 

Hypera obovata (Csiki) G;LG 

Isochnus flagellum (Ericson) LG 

Otiorhynchus fuscipes (Olivier) G;LG 

Otiorhynchus mandibularis Redtenbacher G 

130 

Otiorhynchus politus Gyllenhal G 

Otiorhynchus proximus Stierlin I 

Phthorophloeus spinulosus Rey Q 

Pissodes gyllenhali (Sahlberg, C.R.) H 

Pseudocleonus cinereus (Schrank) G 

Rhyncolus elongatus (Gyllenhal) Q;G;H 

Rhyncolus punctulatus Boheman Q;H 

Rhyncolus sculpturatus Waltl I;H 

Rhyncolus strangulatus Perris Q;H 

Scolytus carpini (Ratzeburg) I 

Scolytus koenigi Schevyrev Q;I 

Stenoscelis submuricatus (Schönherr) Q;I 

Stomodes gyrosicollis (Boheman) Q 

Tachyerges rufitarsis (Germar) Q 

Family PLATYPODIDAE Shuckard, 1839 

Platypus oxyurus Dufour Q

Non-established introductions 

A. G. Duff 

A 

Family DYTISCIDAE Leach, 1815 

Megadytes costalis (Fabricius) 

131 


ll published records of the following species appear to have been of non-breeding 

introductions, or at least no definite evidence has yet been forthcoming that the 

specimens found are natural vagrants or represent established populations, even if 

they have occurred apparently ‘in the wild’. Non-established introductions are not 

considered to be an integral part of our fauna and so have been omitted from the main 

checklist. This checklist has mainly been compiled from a limited range of sources 

(Aitken, 1975; Coquempot, 2006; Hincks, 1953; Hinton, 1945; Hodge & Jones, 1995; 

Ostojá-Starzewski & Hammon, 2008; Winter, 1991) and is likely to be incomplete. 

Note that the taxonomy and nomenclature of some species may have changed since 

the original record(s). 


Abax parallelus (Duftschmid) 

Anchista binotata (Dejean) 

Bembidion (Nepha) callosum Küster 

Carabus (Autocarabus) cancellatus Illiger 

Carabus (Tomocarabus) convexus Fabricius 

Casnoidea indica (Thunberg) 

Celaenephes parallelus Schmidt-Göbel 

Coleolissus iris Andrewes 

Colliuris suturalis Péringuey 

Cymindis setifensis Lucas 

Elaphropus quadrisignatus (Duftschmid) 

Mochtherus tetraspilotus (MacLeay) 

Plochionus pallens (Fabricius) 

Porotachys bisulcatus (Nicolai) 

Stenolophus comma (Fabricius) 

= plagiatus Gorham 

Syntomus fuscomaculatus (Motschulsky) 

Tachyta guineensis Alluaud 


Cercyon variegatus Sharp 

Family HISTERIDAE Gyllenhal, 1808 

Carcinops troglodytes (Paykull) 

Macrolister major (Linnaeus) 

Platysoma elongatum (Thunberg) 

= oblongum (Fabricius) 

Saprinus chalcites (Illiger) 

Teretrius punctulatus Fåhraeus 

Family LEIODIDAE Fleming, 1821 

Aglyptinus agathidioides Blair 


Coenonica puncticollis (Kraatz) 

Oligota chrysopyga Kraatz 

Oligota pseudoparva Williams 

Reichenbachia coronatus Westwood 

Family TROGIDAE MacLeay, 1819 

Trox suberosus (Fabricius) 


Anisoplia agricola (Poda) 

Aphodius (Nialus) varians Duftschmid 

Aphodius (Subrinus) sturmi Harold 

Aphodius (Trichonotulus) scrofa (Fabricius) 

Archophileurus santafeanus Arrow 

Bothynus dasypleurus (Germar) 

Catharsius molossus (Linnaeus) 

Catharsius satyrus Kolbe 

Heteronychus licas (Klug) 

Megasoma gygas Herbst 

Onitis philemon Fabricius 

Onthophagus flavocinctus Klug 

Onthophagus furcatus (Fabricius) 

Oxythyrea funesta (Poda) 

Phyllotocus rufipennis (Boisduval) 

Smaragdesthes africana (Drury) 

Strategus simson (Linnaeus) 

Trochalus fraterculus Kolbe 

Valgus hemipterus (Linnaeus) 

Family BUPRESTIDAE Leach, 1815 

Anthaxia dimidiata Thunberg 

Buprestis aurulenta Linnaeus 

Buprestis novemmaculata Linnaeus 

Buprestis rusticorum Kirby 

Buprestis splendens Fabricius 

Chrysobothris chrysostigma (Linnaeus) 

Chrysobothris femorata (Olivier) 

Eurythrea austrica Linnaeus 

Stenocera sternicornis (Linnaeus) 


Drasterius bimaculatus (Rossi) 

Heteroderes rufangulus (Gyllenhal) 

Family DERMESTIDAE Latreille, 1804 

Dermestes laniarius Illiger 

Phradonoma cercyonoides (Reitter) 

Phradonoma nobile (Reitter) 

Thorictodes dartevellei John 

Thorictus grandicollis Germar 

Thorictus indicus Grouvelle 

Trogoderma anthrenoides (Sharp) 

Trogoderma sternale Jayne

Non-established introductions Checklist of Beetles of the British Isles, 2012 Edition 

Family BOSTRICHIDAE Latreille, 1802 

Amphicerus cornutus (Pallas) 

Apate monachus (Fabricius) 

Bostrychoplites cornutus (Olivier) 

Bostrychus jesuita Fabricius 

Dinoderus bifoveolatus (Wollaston) 

Dinoderus brevis Horn 

Dinoderus distinctus Lesne 

Dinoderus minutus (Fabricius) 

Dinoderus ocellaris Stephens 

Heterobostrychus aequalis (Waterhouse) 

Heterobostrychus brunneus (Murray) 

Heterobostrychus hamatipennis (Lesne) 

Lyctoxylon dentata Pascoe 

Lyctoxylon japonicum Reitter 

Lyctus africanus Lesne 

Lyctus cavicollis LeConte 

Lyctus discedens Blackburn 

Lyctus planicollis LeConte 

Lyctus simplex Reitter 

Lyctus sinensis Lesne 

Micrapate xyloperthoides (Jacquelin du Val) 

Minthea rugicollis Walker 

Octodesmus parvulus Lesne 

Phonopate stridula Lesne 

Scobicia declivis LeConte 

Scobicia pustulata (Fabricius) 

Sinoxylon anale Lesne 

Sinoxylon conigerum Gerstaecker 

Sinoxylon crassum Lesne 

Sinoxylon doliolum Lesne 

Sinoxylon pugnax Lesne 

Sinoxylon ruficorne Fåhraeus 

Tetrapriocera longicornis (Olivier) 

Tristaria grouvellei Reitter 

Trogoxylon impressum (Comolli) 

Trogoxylon parallelopipedum (Melsheimer) 

Xylion adusta Fåhraeus 

Xylion collaris (Erichson) 

Xylion cylindricus (MacLeay) 

Xylion securifer Lesne 

Xylobiops basilaris (Say) 

Xylodeleis obsipa (Germar) 

Xyloperthella crinitarsis (Imhoff) 

Xyloperthella picea (Olivier) 

Xylothrips flavipes (Illiger) 


Mezium americanum Laporte 

Nicobium castaneum (Olivier) 

Ptinus incisicollis Pic 

Ptinus japonicus Reitter 

Stegatus palliatus (Chevrolat) 

Tricorynus herbarium (Gorham) 

Trypopitys carpini Herbst 

Xyletinus ater (Creutzer) 

Family TROGOSSITIDAE Latreille, 1802 

Tenebroides oblongus Sharp 


Cylidroctenus chalybaeum (Westwood) 

Cylidrus fasciatus Laporte 

Cymatodera cylindricollis Chevrolat 

Exkorynetes analis (Klug) 

Opilo domesticus Sturm 

Tillus notatus Klug 

132 

Family MELYRIDAE Leach, 1815 

Melyris oblonga Fabricius 

Family MALACHIIDAE Fleming, 1821 

Anthocomus bipunctatus Harrer 

Ebaeus bicaudatus Thomson 

Family NITIDULIDAE Latreille, 1802 

Brachypeplus depressus Erichson 

Brachypeplus niger Murray 

Brachypeplus pilosellus (Murray) 

Brachypeplus rubidus Murray 

Carpophilus fumatus Boheman 

Carpophilus lugubris Murray 

Colopterus morio Erichson 

Colopterus truncatus Randall 

Glischrochilus fasciatus (Olivier) 

Glischrochilus vittatus (Say) 

Lasiodactylus tibialis Boheman 

Stelidota ferruginea Reitter 

Stelidota strigosa (Gyllenhal) 


Cathartosilvanus vulgaris (Grouvelle) 

Monanus concinnulus (Walker) 

Oryzaephilus acuminatus Halstead 

Oryzaephilus gibbosus Aitken 

Parasilvanus fairmairei (Grouvelle) 

Parasilvanus ocellatus (Grouvelle) 

Protosilvanus lateritius (Reitter) 

Silvanoprus cephalotes (Reitter) 

Silvanoprus scuticollis (Walker) 

Silvanus castaneus MacLeay 

Silvanus difficilis Halstead 

Silvanus inarmatus Wollaston 

Silvanus lateritius (Broun) 

Silvanus lewisi Reitter 

Silvanus productus Halstead 

Silvanus proximus Grouvelle 

Family PASSANDRIDAE Blanchard, 1845 

Laemotmetus rhizophagoides (Walker) 

Family LAEMOPHLOEIDAE Ganglbauer, 1899 

Placonotus politissimus (Wollaston) 

Family CRYPTOPHAGIDAE Kirby, 1826 

Micrambe punctata Grouvelle 

= aubrooki Donisthorpe 

Salebius tarsalis Casey 

Family EROTYLIDAE Latreille, 1802 

Leucohimatium arundinaceum (Forskål) 

Pharaxonotha kirschi Reitter 

Family ENDOMYCHIDAE Leach, 1815 

Trochoideus desjardinsi Guérin-Méneville 

Family COCCINELLIDAE Latreille, 1807 

Calvia decemguttata (Linnaeus) 

Chilocorus distigma Klug 

?Cynegetis impunctata (Linnaeus) 3 

Eriopis connexa Germar 

3 Added by Roy et al. (2011) but the identification has 

since been challenged.

Exochomus nigromaculatus (Goeze) 

Hippodamia convergens (Guérin-Méneville) 

Hyperaspis festiva Mulsant 

Procula douei Mulsant 

Scymnus impexus Mulsant 

Family CORYLOPHIDAE LeConte, 1852 

Arthrolips croceus Matthews 

Family LATRIDIIDAE Erichson, 1842 

Cartodere sp. cf. constricta (Gyllenhal) 

Dienerella costulata (Reitter) 

Dienerella lurida Rücker 

Family CIIDAE Leach, 1819 

Cis sp. cf. delagoensis Reitter 

Family MELANDRYIDAE Leach, 1815 

Serropalpus barbatus (Schaller) 

Family COLYDIIDAE Billberg, 1820 

Aprostoma planifrons Westwood 

Bitoma siccana (Pascoe) 

Colobicus parilis Pascoe 

Colobicus specialis Grouvelle 

Microprius confusus Grouvelle 

Microprius raffragi Grouvelle 

Tyrtaeus dobsoni Hinton 


Ammophorus peruvianus Guérin-Méneville 

Ammophorus rubripes Solier 

Anemia sardoa Géné 

Anthrenopsis scriptipennis (Fairmaire) 

Blaps binominata Escalera 

Blapstinus punctulatus Solier 

Coelopalorus carinatus (Blair) 

Coelopalorus foveicollis (Blair) 

Cossyphus depressus (Fabricius) 

Cynaeus angustus (LeConte) 

Gonocephalum bilineatum (Walker) 

Gonocephalum prolixum (Erichson) 

Gonocephalum rusticum (Olivier) 

Gonocephalum simplex (Fabricius) 

Helenophorus collaris Linnaeus 

Himatismus villosus (Haag-Rutenberg) 

Leiochrodes varicolor Westwood 

Lyphia depressa Hinton 

Lyphia orientalis Blair 

Lyprops curticollis Fairmaire 

Meneristes australis (Blessinger) 

Mesomorphus villiger (Blanchard) 

Mesostena puncticollis Solier 

Mesostenopa arabica Gestro 

Mesostenopa rathjensi Gebien 

Morica planata (Fabricius) 

Ocnera hispida (Forskål) 

Opatroides judaicus Baudi 

Opatroides punctulatus Brullé 

Opatrum verrucosum Germar 

Pachychila salzmanni Solier 

Palorinus humeralis (Gebien) 

Palorus beesoni Blair 

Palorus cerylonoides (Pascoe) 

Palorus ficicola (Wollaston) 

Palorus genalis Blair 

Pimelia rugosa (Fabricius) 

133 

Platydema convexifrons Chevrolat 

Polycoelogastridion tenuipes Kaszab 

Rhytinota laticollis Schaufuss 

Rhytinota praelonga Reiche 

Scotobius crispatus Germar 

Sitophagus hololeptoides (Laporte) 

Tenebrio guineensis Imhoff 

Tentyria cypria Kraatz 

Tribolium anaphe Hinton 

Tribolium audax Halstead 

Tribolium madens (Charpentier) 

Family MELOIDAE Gyllenhal, 1810 

Meloe decorus (Brandt & Erichson) 


Anthicus crinitus Laferté-Sénectère 

Anthicus hesperi King 

Anthicus troilus Hinton 

Floydwernerius australis (King) 

Formicomus pedestris Rossi 

Notoxus monodon (Fabricius) 

Notoxus numidicus (Lucas) 


Family CERAMBYCIDAE Latreille, 1802 

Acanthocinus griseus (Fabricius) 

Ambona electus Gahan 

Ancylonotus tribulus (Fabricius) 

Anoplophora chinensis (Forster) 

Anoplophora glabripennis (Motschulsky) 

Aphoplistis pilosellus (Chevrolat) 

Apriona germari (Hope) 

Arhopalus productus LeConte 

Asemum moestum Haldeman 

Batocera rufomaculata (De Geer) 

Chlorophorus annularis (Fabricius) 

Chlorophorus annulatus Hope 

Chlorophorus figuratus (Scopoli) 

Chlorophorus pilosus (Forster) 

Chlorophorus varius (Müller, O.F.) 

Chlorophorus viticis Gressitt & Rondonare 

Clytus rhamni Germar 

Compsocerus euqestris (Guérin-Méneville) 

Coptocerus rubripes Boisduval 

Coptops aedificator (Fabricius) 

Cordylomera spinicornis (Fabricius) 

Cordylomera suturalis Chevrolat 

Cyllene caryae Gahan 

Cyrtophorus verrucosus (Olivier) 

Deilus fugax (Olivier) 

Diaxenes dendrobii Gahan 

Eburia quadrigeminata (Say) 

Elaphidion nanum (Fabricius) 

Enaphalodes rufulus (Haldeman) 

Ergates faber (Linnaeus) 

Ergates spiculatus LeConte 

Euderces pini (Olivier) 

Geloneatha hirta (Fairmaire) 

Hesperophanes gayi Plavilstshikov 

Leptura obliterata (Haldeman) 

Lagocheirus undulatus (Voet) 

Mallodon downesi (Hope) 

Megacyllene guttata (Chevrolat) 

Monochamus galloprovincialis (Olivier) ssp. pistor 

(Germar) 

Monochamus rosenmuelleri (Cederhjelm) 

Monochamus sartor (Fabricius) 

Monochamus scutellatus (Say)

Non-established introductions Checklist of Beetles of the British Isles, 2012 Edition 

Monochamus sutor (Linnaeus) 

Monochamus titillator (Fabricius) 

Morimus asper (Sulzer) 

Neoclytus acuminatus (Fabricius) 

Neoclytus caprea (Say) 

Neoclytus scutellaris (Olivier) 

Oemona hirta (Fabricius) 

Oxypleurus nodieri Mulsant 

Pachydissus hector Kolbe 

Paramoeocerus barbicornis (Fabricius) 

Phoracantha recurva Newman 

Phoracantha semipunctata (Fabricius) 

Phryneta leprosa (Fabricius) 

Plagionotus detritus (Linnaeus) 

Plocaederus basalis Gahan 

Plocaederus viridipennis (Hope) 

Rhagiomorpha lepturoides (Boisduval) 

Romaleum rufulum Haldeman 

Rosalia alpina (Linnaeus) 

Smodicum cucujiforme (Say) 

Sophronica cinerascens Breuning 

Stenygrinum quadrinotatum Bates 

Stromatium barbatum (Fabricius) 

Tetropium cinnamopterum Kirby, W. 

Tetropium fuscum (Fabricius) 

Trichoferus campestris (Faldermann) 

Trichoferus griseus (Fabricius) 

Xylotrechus colonus (Fabricius) 


Altica carduorum (Guérin-Méneville) 

Bruchidius albopubens (Pic) 

Bruchidius atrolineatus (Pic) 

Bruchidius lividimanus (Gyllenhal) 

Bruchidius pauper (Boheman) 

Bruchidius quinqueguttatus (Olivier) 

Bruchus dentipes (Baudi) 

Bruchus emarginatus Allard 

Bruchus lentis Frölich 

Bruchus natalensis (Pic) 

Bruchus signaticornis Gyllenhal 

Bruchus tristis Boheman 

Callosobruchus analis (Fabricius) 

Callosobruchus phaseoli (Gyllenhal) 

Callosobruchus rhodesianus (Pic) 

Calopepla leayana Latreille 

Caryedon longipennis Pic 

Caryedon serratus (Olivier) 

Caryedon sudanensis Southgate 

Caryopemon cruciger Stephens 

Cerotoma trifurcata (Forster) 

Charidotis circulus Boheman 

Chelymorpha variabilis Boheman 

Chrysolina viridana (Küster) 

Chrysomela cuprea Baly 

Chrysomela superba Thunberg 

Coptocycla laqueata Spaeth 

Crioceris duodecimpunctata (Linnaeus) 

Deloyala cruciata (Linnaeus) 

Euspermophagus sericeus (Geoffroy in Fourcroy) 

Hilarocassis albida (Germar) 

Hispa atra Linnaeus 

Leptinotarsa decemlineata (Say) 

Mesomphalia sextillata Boheman 

Pachymerus cardo (Fåhraeus) 

Pachymerus nigriventris Pic 

Pachynephorus cylindricus Lucas 

Plagiometriona tenella (Klug) 

134 

Poecilaspis angulata (Germar) 

Pseudomesophila indigacea (Boheman) 

Smaragdina salicina (Scopoli) 

Specularius erythraeus (Pic) 

Specularius impressithorax (Pic) 

Spermophagus elizabethae Pic 

Spermophagus natalensis Fåhraeus 

Stolas cyanea (Linnaeus) 

Stolas impluviata (Boheman) 

Stolas lacordairei var. subrugosa (Boheman) 

Stolas redtenbacheri imperialis (Boheman) 

Zabrotes subfasciatus (Boheman) 

Family ANTHRIBIDAE Billberg, 1820 

Anaemerus fuscus (Olivier) 

Trigonorhinus secundus (Wolfrum) 

Family APIONIDAE Schönherr, 1823 

Piezotrachelus sp. cf. gerstaeckeri (Faust) 

Family DRYOPHTHORIDAE Schönherr, 1825 

Cosmopolites sordidus (Germar) 

Diocalandra impressicollis (Quedenfeldt) 

Sipalinus gigas (Fabricius) 

Sitophilus linearis (Herbst) 

Trochorhopalus strangulatus (Gyllenhal) 


Achopera alternata Lea 

Anthonomus britannus Desbrochers 

= pubescens sensu auctt. non (Fabricius) 

Antliarhinus verdcourti Marshall, G. 

Balanogastris kolae (Desbrochers) 

Baris janthina (Boheman) 

Blosyrus asellus (Olivier) 

Brachycerus albidentatus Gyllenhal 

Brachytemnus porcatus (Germar) 

Caulophilus oryzae (Gyllenhal) 

Coccotrypes congonus Eggers 

Coccotrypes dactyliperda (Fabricius) 

Conarthrus cylindricus Wollaston 

Conorhynchus brevirostris (Gyllenhal) 

Conotrachelus histrio Boheman 

Cryphaloides donisthorpei Formanek 

Cryphalus buscki Hopkins 

Cryphalus pallidus Eichhoff 

Cryphalus piceae (Ratzeburg) 

Crypturgus borealis Swaine 

Crypturgus cinereus (Herbst) 

Crypturgus pusillus (Gyllenhal) 

Cycloderes espanoli Roudier 

Dendroctonus brevicornis LeConte 

Dendroctonus pseudotsugae Hopkins 

Dendroctonus rufipennis (Kirby) 

Dryocoetes affaber (Mannerheim) 

Episus cyathiformis Gyllenhal 

Gnathotrichus sulcatus (LeConte) 

Hylastes ruber Swaine 

Hylurgus ligniperda (Fabricius) 

Hylurgops porosus (LeConte) 

Hylurgops rugipennis (Mannerheim) 

Hyperodes bonariensis Kuschel 

Hypothenemus eruditus Westwood 

Hypothenemus hampei (Ferrari) 

Hypothenemus moschatae (Schauffer) 

Ips amitinus (Eichhoff) 

Ips curvidens Germar

Ips duplicatus (Sahlberg) 

Ips latidens (LeConte) 

Ips pini (Say) 

Larinus turbinatus Gyllenhal 

Leurostenus filum Marshall 

Listroderes aequivoca Kuschel 

Listroderes costirostris ssp. obliquus (Klug) 

Lixus ochraceus Boheman 

Lixus terminalis LeConte 

Mesites pallidipennis Boheman 

Orchidophilus aterrimus (Waterhouse) 

Orthorhinus cylindirostris (Fabricius) 

Orthotomicus caelatus (Eichhoff) 

Orthotomicus proximus (Eichhoff) 

Otiorhynchus corruptor (Host) 

Otiorhynchus parvicollis Gyllenhal 

Phaenomerus strigicollis Faust 

Phlyctinus callosus Boheman 

Pityogenes plagiatus (LeConte) 

Poecilips myristicae (Roepke) 

Polydrusus (Metallites) impar des Gozis 

Polygraphus rufipennis (Kirby) 

Polygraphus subopacus (Thomson) 

Pseudohylesinus tsugae Swaine 

Pseudostenotrupis marshalli Zimmermann 

Pseudostoreus placitus Lea 

= harrisoni (Pool) 

Rhyncolus elongatus (Gyllenhal) 

Rhytideres plicatus (Olivier) 

135 

Scierus annectens LeConte 

Sclerorhinus bubalus (Olivier) 

Scolytus dimidiatus Chapuis 

Scolytus laricis Blackman 

Scolytus unispinosus LeConte 

Sipalinus gigas (Fabricius) 

Sipalinus squalidus Kolbe 

Stephanoderes cassiae Eichhoff 

Stephanoderes coffeae Hagedorn 

Sternochetus mangiferae (Fabricius) 

Trypodendron retusum (LeConte) 

Trypophloeus granulatus (Ratzeburg) 

Tychius pumilus Brisout 

Xyleborus eurygraphus (Ratzeburg) 

Xyleborus ferrugineus (Fabricius) 

Xyleborus mascarensis Eichhoff 

Xylosandrus morigerus (Blandford) 

Zeugenia rosacea Heller 


Family PLATYPODIDAE Shuckard, 1839 

Doliopygus bidentatus ssp. kenyaensis Schedl 

Doliopygus crinitus (Chapuis) 

Doliopygus serratus (Strohmeyer) 

Platypus curtus Chapuis 

Platypus hintzi Schaufuss 

Platypus linearis Stephens 

Platypus parallelus (Fabricius) 

Platypus penetralis Sampson

Bibliography Checklist of Beetles of the British Isles, 2012 Edition 

Bibliography 

ABBAZZI, P. & OSELLA, G. 1992. Elenco sistematicofaunistico 

degli Anthribidae, Rhinomaceridae, 

Attelabidae, Apionidae, Brentidae, Curculionidae Italiani 

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Index to family-group and genus-group names 

A 

Abagous, 103 

Abax, 19, 128 

Abdera, 82 

Ablattaria, 33 

ABRAEINAE, 26 

ABRAEINI, 26 

Abraeus, 26 

Acalles, 106 

ACALYPTINI, 101 

Acalyptus, 101 

Acanephodus, 98 

ACANTHOCININI, 90 

Acanthocinus, 90, 133 

Acanthoscelides, 91 

Acatodes, 11 

Acentrotypus, 98 

Achenium, 51 

Achopera, 134 

Acidota, 33, 128 

ACILIINI, 11 

Acilius, 11 

Aclypea, 33 

Acmaeops, 88 

Acrantus, 112 

Acratrichis, 30 

ACRITINI, 26 

Acritus, 26 

Acrognathus, 48 

Acrolocha, 34 

Acrossus, 56, 129 

Acrotona, 38 

ACROTRICHINAE, 29 

ACROTRICHINI, 30 

Acrotrichis, 30 

Acrulia, 34 

Actedium, 17 

Actenicerus, 62 

Actidium, 29 

Actinopteryx, 30 

Actobius, 52 

ACTOCHARINI, 38 

Actocharis, 38 

Actophylla, 40 

Actosus, 46 

Acupalpus, 22 

Acylophorus, 53 

Adalia, 79 

Adelocera, 62 

Adelosia, 18 

ADEPHAGA, 10 

ADERIDAE, 87 

Aderus, 87 

Adimonia, 93 

Adistemia, 80 

Adonia, 79 

Adota, 39 

Adoxus, 92 

ADRASTINI, 63 

Adrastus, 63 

Aegialia, 55 

AEGIALIINAE, 55 

Aeletes, 26 

Aepopsis, 16 

Aepus, 16 

Aetophorus, 22 

AGABINAE, 11 

Agabus, 11, 128 

Agapanthia, 90 

AGAPANTHIINI, 90 

Agaporus, 13 

Agaricochara, 43 

Agaricophagus, 31 

AGATHIDIINI, 31 

Agathidium, 32 

Agelastica, 93 

AGLENINAE, 86 

Aglenus, 86 

Aglyptinus, 131 

Agoliinus, 56 

Agonum, 20, 128 

AGRILINAE, 59 

AGRILINI, 59 

Agrilinus, 56 

Agrilus, 59 

Agriotes, 63 

AGRIOTINI, 63 

AGRYPNINAE, 62 

Agrypnus, 62 

Ahasverus, 73 

Airaphilus, 129 

Aizobius, 97 

Alaobia, 39 

Alapsodus, 54 

Aleochara, 38 

ALEOCHARINAE, 38 

ALEOCHARINI, 38 

Aleuonota, 39 

Aleurostictus, 58 

Alevonota, 39 

Alexia, 77 

ALEXIIDAE, 77 

Alianta, 39 

ALLECULINAE, 85 

ALLECULINI, 85 

Allodactylus, 105 

Allopentarthrum, 106 

Alocoderus, 129 

Aloconota, 39 

ALOPHINI, 106 

Alophus, 106 

Alosterna, 89 

ALPHITOBIINI, 84 

Alphitobius, 84 

Alphitophagus, 85 

Altica, 94, 134 

ALTICINI, 93 

AMALINI, 105 

Amalorhinoncus, 105 

Amalorrhynchus, 103 

Amalus, 105 

Amara, 20, 128 

Amarochara, 45 

Amauronyx, 36 

Ambona, 133 

Amidobia, 39 

Amidorus, 129 

Amisammus, 48 

Amischa, 39 

Ammoecius, 56 

157 


Ammophorus, 133 

AMPEDINI, 63 

Ampedus, 63 

Amphicerus, 132 

Amphicyllis, 32 

Amphigynus, 19 

Amphimallon, 57 

Amphimallus, 57 

Amphotis, 72 

Anacaena, 24 

Anaclyptus, 90 

Anaemerus, 134 

ANAGLYPTINI, 90 

Anaglyptus, 90 

Anambus, 59 

ANAMORPHINAE, 77 

ANASPIDINAE, 88 

Anaspis, 88 

Anastrangalia, 88 

Anatis, 79 

Anchicera, 75 

Anchista, 131 

Anchomenus, 19 

Anchonidium, 110 

Anchus, 19 

Ancistronycha, 64 

Ancylonotus, 133 

Ancyrophorus, 48 

Andrion, 108 

ANEMADINI, 32 

Anemia, 133 

Anisandrus, 111 

Anisarthria, 28 

Anisocera, 70 

Anisodactylus, 21 

Anisoplia, 131 

Anisosticta, 78, 129 

Anisotoma, 31 

Anisoxya, 82 

Anitys, 68 

ANOBIINAE, 68 

Anobium, 68 

Anomaeocera, 70 

Anomala, 58 

ANOMALINI, 58 

ANOMMATINAE, 77 

Anommatus, 77 

Anomognathus, 44 

Anomophagus, 72 

Anoncodes, 86 

Anopleta, 39 

ANOPLINI, 110 

Anoplodera, 89 

Anoplophora, 133 

Anoplotrupes, 55 

Anoplus, 110 

Anostirus, 62 

Anotylus, 47, 128 

Anthaxia, 59, 131 

ANTHAXIINI, 59 

Antherophagus, 75 

ANTHICIDAE, 87, 129, 133 

ANTHICINAE, 87 

Anthicus, 87, 129, 133 

Anthobium, 34 

Anthocomus, 70, 132

Index to family-group and genus-group names Checklist of Beetles of the British Isles, 2012 Edition 

Anthomorphus, 101 

ANTHONOMINI, 101 

Anthonomus, 101, 134 

ANTHOPHAGINI, 33 

Anthophagus, 34, 128 

Anthracus, 22 

ANTHRENINAE, 66 

Anthrenocerus, 66 

Anthrenodes, 66 

Anthrenops, 66 

Anthrenopsis, 133 

Anthrenus, 66 

ANTHRIBIDAE, 96, 134 

ANTHRIBINAE, 96 

ANTHRIBINI, 96 

Anthribus, 96 

Antidonus, 109 

Antliarhinus, 134 

Apalus, 86 

Apate, 132 

APHANISTICINI, 59 

Aphanisticus, 59 

Aphanommata, 130 

Aphidecta, 79 

Aphideita, 79 

APHODIINAE, 56 

APHODIINI, 56 

Aphodius, 56, 129, 131 

Aphoplistis, 133 

Aphthona, 94 

Apion, 97 

APIONIDAE, 97, 134 

APIONINAE, 97 

APIONINI, 97 

Apiops, 99 

APLEMONINI, 97 

Aplocnemus, 69 

Aploderus, 47, 128 

Aplotarsus, 62 

APODERINAE, 97 

Apoderus, 97 

APOSTASIMERINI, 103 

Apriona, 133 

Aprostoma, 133 

Apteropeda, 95 

Aptyktophallus, 60 

ARAECERINI, 96 

Araecerus, 96 

Araeocerus, 96 

Archarius, 100 

Archicarabus, 15 

Archophileurus, 131 

Arena, 46 

Argutor, 19 

Arhopalus, 89, 133 

Aridius, 80 

Aromia, 89 

Arpedium, 34, 128 

Arpidiphorus, 70 

Arrhenopeplus, 35 

Arthrolips, 133 

Asaphidion, 17, 128 

Asclera, 86 

ASCLERINI, 86 

ASEMINI, 89 

Asemum, 89, 133 

Asiobates, 28 

Asiorestia, 95 

ASPIDAPIINI, 98 

Aspidapion, 98 

Aspidiphorus, 70 

ASTENINA, 50 

Astenognathus, 50 

Astenus, 50 

Astilbus, 44 

Astrapaeus, 53 

Astycops, 47 

Atactogenus, 107 

Ataenius, 129 

Atemeles, 44 

Atheta, 39 

ATHETINA, 38 

ATHETINI, 38 

Atholus, 27 

Athous, 62 

Atomaria, 75 

ATOMARIINAE, 75 

ATOMARIINI, 75 

Atracthelophorus, 23 

Atrecus, 52 

Atritomus, 81 

Attactagenus, 107 

ATTAGENINAE, 66 

Attagenus, 66 

ATTELABIDAE, 97 

ATTELABINAE, 97 

Attelabus, 97 

Augyles, 61 

Aulacobaris, 103, 130 

Aulacochthebius, 28 

Aulonium, 84 

Aulonothroscus, 61 

Autalia, 42 

AUTALIINI, 42 

Autocarabus, 128, 131 

Axinotarsus, 70 

B 

Badister, 22 

Badura, 40 

Baeckmanniolus, 26 

Baeocrara, 30 

BAGOINAE, 103 

Bagous, 103 

Balaninus, 100 

Balanobius, 100 

Balanogastris, 134 

Baptolinus, 52 

Bargus, 47 

BARIDINAE, 103 

BARIDINI, 103 

Baris, 103, 134 

Barynotus, 107 

Baryodma, 38 

Barypeithes, 108 

Barypithes, 108 

Batenus, 19 

Bathyscia, 33 

Batocera, 133 

Batophila, 95 

BATRISINI, 35 

BATRISITAE, 35 

Batrisodes, 35 

Batrisus, 128 

Baudia, 22 

BEMBIDIINI, 17 

Bembidion, 17, 128, 131 

158 

Bembidionetolitzkya, 17 

Bembidium, 17 

Berginus, 81 

Berosus, 24 

Bessobia, 40 

Betulapion, 99 

Bibloplectus, 36 

BIBLOPORINA, 36 

Bibloporus, 36 

BIDESSINI, 12 

Bidessus, 12, 128 

BIPHYLLIDAE, 77 

Biphyllus, 77 

Bisnius, 52 

Bitoma, 84, 133 

Blaps, 85, 133 

Blapstinus, 133 

BLAPTINI, 85 

Blastophagus, 112 

Blechrus, 23 

Bledius, 47, 128 

Blemus, 16 

Blepharoplataphus, 17 

Blethisa, 16 

Blitophaga, 33 

Blosyrus, 134 

Bobitobus, 37 

Bodilus, 56 

Boeocrara, 30 

Bohemiellina, 43 

BOLBOCERATINAE, 55 

Boletophagus, 84 

Bolitobius, 37 

Bolitochara, 43 

BOLITOCHARINA, 43 

BOLITOPHAGINI, 84 

Bolitophagus, 84 

Boraginobius, 104 

Borboropora, 43 

Boreaphilus, 128 

Boreohypera, 109 

Boreonebria, 15 

Boreonectes, 13 

Boreophilia, 40 

BOSTRICHIDAE, 66, 132 

BOSTRICHINAE, 66 

BOSTRICHOIDEA, 65 

Bostrichus, 66 

Bostrychoplites, 132 

Bostrychus, 66, 132 

Bothrideres, 129 

BOTHRIDERIDAE, 77, 129 

Bothriopterus, 18 

Bothryorrhynchapion, 98 

Bothynoderes, 109 

Bothynus, 131 

Brachida, 43 

BRACHININAE, 15 

Brachinus, 15 

Brachonyx, 101 

Brachycerus, 134 

Brachyderes, 106 

BRACHYDERINI, 106 

Brachygluta, 36 

BRACHYGLUTINI, 36 

Brachygonus, 63 

Brachylacon, 62 

Brachynidius, 15 

Brachypeplus, 132

Brachypterolus, 70 

Brachypterus, 70 

Brachyrhinus, 107 

Brachysomus, 108 

Brachytarsus, 96 

Brachytemnus, 134 

Brachyusa, 46 

Bracteon, 17 

Bradybatus, 101 

Bradycellus, 21 

Bradytus, 20 

Brindalus, 57 

Bromius, 92 

Brontes, 73 

BRONTINAE, 73 

BROSCINI, 16 

Broscus, 16 

Bruchela, 96 

Bruchella, 96 

Bruchidius, 91, 134 

BRUCHINAE, 91 

Bruchus, 67, 91, 134 

Brundinia, 40 

Bryaxis, 36 

Brychius, 10 

Bryocharis, 37 

Bryophacis, 37 

Bryoporus, 37 

BUPRESTIDAE, 59, 129, 131 

BUPRESTINAE, 59 

Buprestis, 129, 131 

BUPRESTOIDEA, 59 

BYCTISCINI, 97 

Byctiscus, 97 

BYRRHIDAE, 60, 129 

BYRRHINAE, 60 

BYRRHOIDEA, 60 

Byrrhus, 60 

BYTHININI, 36 

Bythinus, 36 

BYTURIDAE, 76 

Byturus, 76 

C 

Caccobius, 129 

Cadaverota, 40 

Caenocara, 68 

Caenocorse, 84 

Caenopsis, 108 

Caenoptera, 89 

Caenorhinus, 97 

CAENOSCELINI, 75 

Caenoscelis, 75 

Cafius, 52, 53 

Calambus, 62 

Calamosternus, 56 

Calandra, 100 

Calaphodius, 129 

Calathus, 19 

Calendra, 100 

Callicerus, 40 

CALLICHROMATINI, 89 

CALLIDIINI, 90 

CALLIDIOPINI, 89 

Callidium, 90 

Callirus, 110 

Callistus, 22 

Callosobruchus, 91, 134 

Calodera, 45 

Calodromius, 23 

Calomicrus, 93 

Calopepla, 134 

Calosirus, 103 

Calosoma, 15 

Calostirus, 62 

Calvia, 78, 132 

CALYPTOMERINAE, 58 

Calyptomerus, 58 

Campylus, 62 

CANTHARIDAE, 64 

CANTHARINAE, 64 

CANTHARINI, 64 

Cantharis, 64, 86 

CARABIDAE, 13, 128, 131 

CARABINAE, 15 

CARABINI, 15 

Carabus, 15, 128, 131 

Carcinops, 27, 131 

CARDIOPHORINAE, 64 

Cardiophorus, 64 

Carida, 82 

Carpelimus, 48 

CARPOPHILINAE, 71 

CARPOPHILINI, 71 

Carpophilus, 71, 132 

Cartodere, 80, 133 

Cartoderema, 80 

Caryedon, 134 

Caryopemon, 134 

Casnoidea, 131 

Cassida, 95 

CASSIDINAE, 95 

CATAPIINA, 99 

Catapion, 99 

Cateretes, 70 

Cathammistes, 24 

Catharsius, 131 

Cathartosilvanus, 132 

Cathartus, 73 

Cathormiocerus, 108, 130 

Catopidius, 33 

Catops, 33 

Caulophilus, 134 

Caulotrupis, 106 

Caulotrypis, 106 

Celaenephes, 131 

Celia, 20 

CEPHENNIINI, 49 

Cephennium, 49 

CERAMBYCIDAE, 88, 133 

CERAMBYCINAE, 89 

CERAMBYCINI, 89 

Cerambyx, 89 

Ceranota, 38 

Cerapheles, 70 

CERATAPIINI, 98 

Ceratapion, 98 

Ceratomegilla, 129 

Cercus, 70 

Cercyon, 25, 131 

Ceritaxa, 40 

Cerotoma, 134 

Cerycon, 25 

Cerylon, 77 

CERYLONIDAE, 77 

CERYLONINAE, 77 

Cetonia, 58 

159 


CETONIINAE, 58 

CETONIINI, 58 

Ceuthorhynchidius, 105 

Ceuthorhynchus, 103 

Ceuthorrhynchus, 103 

CEUTORHYNCHINAE, 103 

CEUTORHYNCHINI, 103 

Ceutorhynchus, 103 

Chaetarthria, 24 

Chaetida, 40 

Chaetocarabus, 15 

Chaetocnema, 95, 129 

Chaetophora, 60 

Chaetophorus, 60 

Chalcoides, 95 

Charagmus, 108 

Charidotis, 134 

Chelymorpha, 134 

Chetabraeus, 128 

CHILOCORINAE, 78 

Chilocorus, 78, 132 

Chilomorpha, 45 

Chilopora, 46 

Chiloporata, 46 

Chilothorax, 56, 129 

Chlaeniellus, 22 

CHLAENIINI, 22 

Chlaenius, 22, 128 

Chloecharis, 51 

Chlorapion, 99 

Chlorophorus, 133 

Cholerus, 49 

Choleva, 32 

CHOLEVINAE, 32 

CHOLEVINI, 32 

Cholevopsis, 32 

CHORAGINAE, 96 

CHORAGINI, 96 

Choragus, 96 

Chromoderus, 109 

Chrysanthia, 86 

Chrysarthia, 86 

Chrysobothris, 131 

Chrysobracteon, 17 

Chrysolina, 92, 129, 134 

Chrysomela, 93, 129, 134 

CHRYSOMELIDAE, 91, 129, 134 

CHRYSOMELINAE, 92 

CHRYSOMELOIDEA, 88 

Chrysophis, 108 

Cicindela, 15 

CICINDELINAE, 15 

Cicones, 84 

Cidnopus, 62 

Cidnorhinus, 104 

Cidnorrhinus, 104 

CIIDAE, 81, 129, 133 

CIINAE, 81 

CIINI, 81 

Cilea, 37 

Cillenus, 17 

CIMBERIDINAE, 96 

Cimberis, 96 

CIONINI, 101 

Cionus, 101 

Cis, 82, 133 

Cissophagus, 112 

Cistela, 85 

Cladoconnus, 49


CLAMBIDAE, 58 

CLAMBINAE, 58 

Clambus, 58 

Clanoptilus, 70 

Claviger, 35 

CLAVIGERINI, 35 

CLAVIGERITAE, 35 

CLEONINI, 109 

Cleonis, 109 

Cleonus, 109 

Cleopomiarus, 101 

Cleopus, 101 

CLERIDAE, 69, 129, 132 

CLERINAE, 69 

CLEROIDEA, 68 

Clinocrara, 82 

Clitostethus, 78 

Clivina, 16 

Clythra, 92 

CLYTINI, 90 

Clytra, 92 

CLYTRINI, 92 

Clytus, 90, 133 

Cnemapion, 99 

Cnemidotus, 10 

CNEORHININI, 107 

Cneorhinus, 107 

Cneorrhinus, 107 

Cobosiotherium, 99 

Coccidula, 77 

COCCIDULINAE, 77 

Coccinella, 79 

COCCINELLIDAE, 77, 129, 132 

COCCINELLINAE, 78 

Coccinula, 78 

Coccotrypes, 134 

Codiosoma, 106 

Coelambus, 13, 128 

Coeliodes, 104 

Coeliodinus, 104 

Coelopalorus, 133 

Coelositona, 108 

Coelostethus, 68 

Coelostoma, 25 

Coenonica, 131 

Coenorrhinus, 97 

Coenoscelis, 75 

Colenis, 31 

Coleolissus, 131 

COLEOPTERA, 10 

Colliuris, 23, 131 

Colobicus, 133 

Colobopterus, 56 

Colon, 32 

COLONINAE, 32 

Colopterus, 132 

Colotes, 69 

Colpodota, 38 

COLYDIIDAE, 83, 129, 133 

COLYDIINAE, 83 

COLYDIINI, 84 

Colydium, 84 

Colymbetes, 11, 128 

COLYMBETINAE, 11 

Comazus, 58 

Compsocerus, 133 

Conarthrus, 106, 134 

Coninomus, 80 

Coniocleonus, 109 

Conithassa, 80 

Conopalpus, 82 

Conorhynchus, 134 

Conosoma, 37 

Conosomus, 37 

Conotrachelus, 134 

COPELATINAE, 11 

Copelatus, 11 

COPRINI, 57 

Copris, 57 

Coprochara, 38 

COPROPHILINI, 47 

Coprophilus, 47 

Coprothassa, 40 

Coptocerus, 133 

Coptocycla, 134 

Coptops, 133 

Cordalia, 43 

Cordicollis, 87 

Cordicomus, 87, 129 

Cordylepherus, 70 

Cordylomera, 133 

CORTHYLINA, 111 

Corticaria, 80 

CORTICARIINAE, 80 

Corticarina, 81 

Corticeus, 85 

Cortinicara, 81 

CORYLOPHIDAE, 79, 133 

CORYLOPHINAE, 79 

Corylophus, 79 

Corymbites, 62 

Corynetes, 69 

CORYPHIINI, 34 

Coryphium, 34 

Cosmobaris, 103 

Cosmopolites, 134 

COSSONINAE, 105 

COSSONINI, 105 

Cossonus, 105 

Cossyphus, 133 

COTASTER, 106 

Cousya, 45 

Craetopycrus, 47 

Crataraea, 45 

Creophilus, 54 

Crepidodera, 95 

Criocephalum, 89 

Criocephalus, 89 

CRIOCERINAE, 91 

Crioceris, 92, 134 

Criomorphus, 89 

Crudosilis, 65 

CRYPHALINA, 111 

Cryphaloides, 134 

Cryphalus, 111, 134 

Cryptamorpha, 73 

Cryptarcha, 72 

CRYPTARCHINAE, 72 

CRYPTICINI, 85 

Crypticus, 85 

CRYPTOBIINA, 50 

Cryptobium, 50 

CRYPTOCEPHALINAE, 92 

CRYPTOCEPHALINI, 92 

Cryptocephalus, 92 

Cryptohypnus, 62 

Cryptolaemus, 78 

Cryptolestes, 73, 129 

160 

CRYPTOPHAGIDAE, 74, 132 

CRYPTOPHAGINAE, 74 

CRYPTOPHAGINI, 74 

Cryptophagus, 74 

CRYPTOPHILINAE, 76 

Cryptophilus, 76 

Cryptophonus, 21 

Cryptopleurum, 25 

Cryptorhynchidius, 106 

CRYPTORHYNCHINA, 106 

CRYPTORHYNCHINAE, 106 

CRYPTORHYNCHINI, 106 

Cryptorhynchus, 106 

CRYPTURGINA, 111 

Crypturgus, 111, 134 

Cryptusa, 43 

Ctenicera, 62 

CTENICERINI, 62 

Ctenicerus, 62 

CTENIOPODINI, 85 

Cteniopus, 85 

Ctenopteryx, 30 

Ctesias, 66 

CUCUJIDAE, 73 

CUCUJOIDEA, 70 

Curculio, 100 

CURCULIONIDAE, 100, 130, 134 

CURCULIONINAE, 100 

CURCULIONINI, 100 

CURCULIONOIDEA, 96 

Curimopsis, 60, 129 

Curtonotus, 20 

Cyanapion, 98 

Cyaniris, 92 

Cyanostolus, 73 

Cybister, 11 

CYBISTRINI, 11 

CYBOCEPHALINAE, 72 

Cybocephalus, 72 

Cychramus, 72 

CYCHRINI, 15 

Cychrus, 15 

Cycloderes, 134 

Cyclodinus, 87 

CYCLOMINAE, 106 

Cyclonotum, 25 

Cylidroctenus, 132 

Cylidrus, 132 

Cylindera, 15 

Cylindrinotus, 84 

Cylindronotus, 84 

Cyllene, 133 

Cymatodera, 132 

Cymbiodyta, 24 

Cymindis, 22, 128, 131 

Cynaeus, 133 

Cynegetis, 132 

Cypha, 44 

Cyphea, 44 

Cyphelophorus, 23 

Cyphoceble, 32 

Cyphocleonus, 130 

Cyphon, 59 

Cyprus, 103 

Cyrtonotus, 20 

Cyrtophorus, 133 

CYRTOSCYDMINI, 49 

Cyrtoscydmus, 49 

Cyrtotriplax, 76

Cyrtusa, 31 

Cytilus, 60 

D 

Dacne, 76 

DACNINI, 76 

Dacrila, 46 

Dactylosternum, 25 

Dadobia, 40 

Dadopera, 71 

Dadopora, 71 

Dalopius, 63 

Dalotia, 40 

Daniela, 17 

Dapalinus, 109 

DASCILLIDAE, 59 

DASCILLOIDEA, 59 

Dascillus, 59 

Dasygnypeta, 46 

Dasytes, 69 

DASYTIDAE, 69 

DASYTINAE, 69 

Datomicra, 40 

Datonychus, 104 

Datostiba, 40 

Decamera, 57 

Deilus, 133 

DEINOPSINI, 43 

Deinopsis, 43 

Deleaster, 47 

Deliphrum, 34 

Deloyala, 134 

Demetrias, 22 

Demochrus, 32 

Dendrobium, 68 

Dendroctonus, 112, 134 

Dendrophagus, 73 

DENDROPHILINAE, 26 

DENDROPHILINI, 26 

Dendrophilus, 26 

Dendroxena, 33 

DENTICOLLINAE, 62 

DENTICOLLINI, 62 

Denticollis, 62 

DEPORAINI, 97 

Deporaus, 97 

Dermestes, 66, 129, 131 

DERMESTIDAE, 65, 129, 131 

DERMESTINAE, 66 

Dermestinus, 66 

Derocrepis, 95 

DERODONTIDAE, 65 

DERODONTOIDEA, 65 

Deronectes, 12 

Deubelia, 45 

Dexiogya, 45 

Dexiogyia, 45 

Diabrotica, 93 

Diacanthous, 63 

Diacheila, 128 

Diachromus, 21 

Diaclina, 129 

Dianous, 49 

DIAPERINAE, 85 

DIAPERINI, 85 

Diaperis, 85 

Diaphonus, 70 

Diastictus, 57 

Diaxenes, 133 

Dibolia, 95 

Dicarenus, 47 

Dicheirotrichus, 21 

Dichirotrichus, 21 

Dicronychus, 64 

Dictyoptera, 64 

DICTYOPTERINI, 64 

Dicyrtocercyon, 25 

Dieckmanniellus, 100 

Dieletus, 107 

Dienerella, 80, 133 

Diglossa, 43 

Diglotta, 43 

DIGLOTTINI, 43 

Dilacra, 40 

Dilixellus, 109 

Dimetrota, 40 

Dinaraea, 41 

Dinarda, 45 

DINARDINA, 45 

DINODERINAE, 66 

Dinoderus, 132 

Dinopsis, 43 

Dinoptera, 88 

Dinothenarus, 54 

Diocalandra, 134 

Diphyllus, 77 

Diplapion, 98 

Diplocampa, 18 

Diplocoelus, 77 

DIRHAGINI, 61 

Dirhagus, 61 

DISCOPTENINI, 96 

Disopora, 39 

Dissoleucas, 96 

Distichalius, 53 

Ditoma, 84 

DITYLINI, 86 

Dochmonota, 41 

Dolichosoma, 69 

Doliopygus, 135 

Dolopius, 63 

Donacia, 91, 129 

DONACIINAE, 91 

Dorcatoma, 68 

DORCATOMINAE, 68 

Dorcus, 55 

Dorkatoma, 68 

DORYTOMINA, 101 

Dorytomus, 101 

Dralica, 40 

Drasterius, 131 

Drepanocerus, 129 

DRILIDAE, 64 

Drilus, 64 

Dromaeolus, 129 

Dromiolus, 23 

Dromius, 23 

Dropephylla, 34 

Drupenatus, 104 

Drusenatus, 104 

Drusilla, 44 

Drymoporus, 37 

Dryocaetes, 111 

Dryocoetes, 111, 134 

DRYOCOETINA, 111 

Dryocoetinus, 111 

DRYOPHILINAE, 67 

161 


Dryophilus, 67 

DRYOPHTHORIDAE, 100, 134 

DRYOPHTHORINAE, 100 

Dryophthorus, 100 

DRYOPIDAE, 60 

Dryops, 60 

DRYOTRIBINI, 106 

Drypta, 23 

DRYPTINI, 23 

Dupophilus, 129 

Dyschiriodes, 16 

Dyschirius, 16, 128 

DYTISCIDAE, 11, 128, 131 

DYTISCINAE, 11 

DYTISCINI, 12 

Dytiscus, 12 

E 

Ebaeus, 70, 132 

Eburia, 133 

Eccoptogaster, 110 

Edaphus, 50 

Edo, 110 

Elachys, 29 

Elaphidion, 133 

ELAPHRINI, 16 

Elaphropus, 17, 131 

Elaphrus, 16 

Elater, 63 

ELATERIDAE, 62, 129, 131 

ELATERINAE, 63 

ELATERINI, 63 

ELATEROIDEA, 61 

Elbidus, 48 

Eledona, 84 

Elleschus, 101 

ELLESCINA, 101 

ELLESCINI, 101 

Ellescus, 101 

ELMIDAE, 60, 129 

Elmis, 60 

Elodes, 58 

Elonium, 34, 47 

Elophorus, 23 

Emphanes, 18 

Emplenota, 38 

Empleurus, 23 

Emus, 54 

Enalodroma, 41 

Enaphalodes, 133 

Encephalus, 43 

ENDOMYCHIDAE, 77, 129, 132 

ENDOMYCHINAE, 77 

Endomychus, 77 

Endophloeus, 84 

Enicmus, 80 

Enicocerus, 27 

Ennearthron, 82 

Enochrus, 24 

Enoplurus, 24 

ENTIMINAE, 106 

Entomoscelis, 129 

Entypus, 81 

Eotachys, 17 

Epaphius, 16 

Ephimeropus, 103 

Ephistemus, 76 

Epierus, 27


Epilachna, 79 

EPILACHNINAE, 79 

EPIPHANINI, 61 

Epiphanis, 61 

Epipolaeus, 110 

Episus, 134 

Epithrix, 95 

Epitrix, 95 

Epuraea, 71 

Epuraeanella, 71 

EPURAEINI, 71 

Ercycon, 25 

Eremotes, 106 

Ergates, 133 

Erichsonius, 52 

Eriopis, 132 

ERIRHINIDAE, 100 

ERIRHININAE, 100 

ERIRHININI, 100 

Erirhinus, 100 

Eririnomorphus, 109 

ERNOBIINAE, 67 

Ernobius, 67 

Ernoporicus, 111 

Ernoporus, 111 

Eroosapion, 97 

Eros, 64 

Erotides, 64 

EROTINAE, 64 

EROTINI, 64 

EROTYLIDAE, 76, 129 

EROTYLINAE, 76 

Erythrapion, 97 

Eryx, 85 

Esolus, 60, 129 

Esymus, 56, 129 

Eteophilus, 101 

Ethelcus, 104 

EUAESTHETINAE, 50 

Euaesthetus, 50 

Eubria, 61 

EUBRIINAE, 61 

Eubrychius, 105 

Eucarabus, 15 

Euceratobledius, 48 

Euchlora, 58 

EUCINETIDAE, 58 

Eucinetus, 58 

Eucnecosum, 34, 128 

EUCNEMIDAE, 61, 129 

EUCNEMINAE, 61 

Eucnemis, 61 

Euconnus, 49 

EUCRADINAE, 67 

Eudectus, 34, 128 

Euderces, 133 

Eudipnus, 108 

Euglenes, 87 

Euheptaulacus, 57 

Eulagius, 81 

Eulixus, 109 

Eumicrus, 49 

EUMOLPINAE, 92 

Euoniticellus, 129 

Euophryum, 106 

Euorodalus, 56 

EUPARIINI, 57 

Eupetedromus, 17 

EUPHANIINI, 47 

EUPLECTINI, 35 

EUPLECTITAE, 35 

Euplectoides, 35 

Euplectus, 35 

Eupleurus, 56 

Euptilium, 29 

Eurhizophagus, 73 

Eurodrusus, 108 

Europhilus, 20 

Eurostus, 67 

Eurycolon, 32 

Eurynebria, 15 

Euryporus, 53 

Euryptilium, 29 

Eurythrea, 131 

Euryusa, 43 

Euspermophagus, 134 

EUSPHALERINI, 34 

Eusphalerum, 34 

Eustolus, 108 

Eutheia, 48 

EUTHEIINI, 48 

Euthia, 48 

Euthiconus, 49 

Euthoron, 102 

Eutrichapion, 99 

Evanystes, 41 

Exanathrotus, 62 

EXAPIINI, 98 

Exapion, 98 

Exkorynetes, 132 

Exochomus, 78, 133 

Exomias, 108 

F 

Fagniezia, 36 

Falagria, 43 

FALAGRIINI, 43 

Falagrioma, 43 

Feronia, 18 

Ferreria, 100 

Fleutiauxellus, 64 

Florilinus, 66 

Floydwernerius, 133 

Formicomus, 133 

Furcipus, 101 

G 

Gabrius, 52 

Gabronthus, 52 

Galeruca, 93 

Galerucella, 93 

GALERUCINAE, 93 

GALERUCINI, 93 

Galleruca, 93 

Gastrallus, 68 

Gastroidea, 92 

Gastrophysa, 92 

Gaurodytes, 11 

Gauropterus, 54 

Geloneatha, 133 

Geodromicus, 34, 128 

GEONEMINI, 107 

GEORISSIDAE, 24 

Georissus, 24 

Georyssus, 24 

Geostiba, 41 

162 

Geotrupes, 55 

GEOTRUPIDAE, 55 

GEOTRUPINAE, 55 

GIBBIINI, 67 

Gibbium, 67 

Gillmeisterium, 29 

Glaphyra, 89 

Glischrochilus, 72, 132 

Globicornis, 66 

Glocianus, 104 

Glossola, 39 

Gnathocerus, 85 

Gnathoncus, 26 

Gnathotrichus, 134 

Gnatocerus, 85 

Gnorimus, 58 

Gnypeta, 46 

Goerius, 54 

GONATOCERI, 100 

GONIACERITAE, 36 

Gonioctena, 93 

Gonocephalum, 133 

Gonodera, 85 

Gracilia, 89 

GRACILIINI, 89 

Grammoptera, 88 

Grammostethus, 27 

Graphoderus, 11 

Graptodytes, 12 

Graptus, 106 

Gressnerium, 28 

GRONOPINA, 106 

Gronops, 106 

Grynobius, 67 

Grypidius, 100 

Grypus, 100 

Guignotus, 12 

Gymnaetron, 101 

Gymnetron, 101 

Gymnusa, 43 

GYMNUSINI, 43 

Gynandrophtalma, 92 

Gynandrophthalma, 92 

GYRINIDAE, 10, 128 

GYRININAE, 10 

GYRININI, 10 

Gyrinulus, 10 

Gyrinus, 10, 128 

Gyrocecis, 73 

Gyrohypnus, 54 

Gyrophaena, 44 

GYROPHAENINA, 43 

H 

HABROCERINAE, 38 

Habrocerus, 38 

Hadrobregmus, 68 

HADROGNATHINI, 34 

Hadrognathus, 34 

Hadroplontus, 104 

Haemonia, 91 

HAETERIINAE, 27 

Haeterius, 27 

Hagrium, 88 

Halacritus, 26 

HALIPLIDAE, 10 

Haliplidius, 10 

Haliplus, 10

HALLOMENINAE, 82 

Hallomenus, 82 

Halobrecta, 41 

Haltica, 94 

Halyzia, 78 

Hapalaraea, 34 

Haplocnemus, 69 

Haploderus, 47 

Haploglossa, 45 

Haploharpalus, 20 

Harminius, 63 

Harmonia, 79 

HARPALINI, 20 

Harpalus, 20 

Hedobia, 67 

Heledona, 84 

Helenophorus, 133 

Helianthemapion, 97 

Helichus, 60 

Heliopathes, 84 

Helmis, 60 

Helobium, 16 

Helocerus, 66 

Helochares, 25 

Helodes, 58, 93 

HELOPHORIDAE, 23, 128 

Helophorus, 23, 128 

HELOPINI, 84 

Helops, 84 

Hemicarabus, 15 

Hemicoelus, 68 

Hemicrepidius, 63 

Hemistenus, 49, 50 

Hemitrichapion, 99 

Henicocerus, 27 

Henosepilachna, 79 

Henoticus, 74 

Heptaulacus, 57 

Hermaeophaga, 94 

Hesperophanes, 133 

Hesperophilus, 48 

Hetaerius, 27 

Heterhelus, 129 

Heterobostrychus, 132 

HETEROCERIDAE, 61, 129 

Heterocerus, 61, 129 

Heteroderes, 131 

Heteronychus, 131 

Heterostomus, 70 

Heterota, 43 

Heterothops, 53 

Hilarocassis, 134 

Himatismus, 133 

Hippodamia, 79, 129, 133 

Hippuriphila, 95 

Hispa, 134 

Hister, 27, 128 

HISTERIDAE, 26, 128, 131 

HISTERINAE, 27 

Holoboreaphilus, 128 

Holobus, 44 

Hololepta, 27 

Holoparamecus, 77 

Holotrichapion, 99 

Homalochthebius, 28 

Homaloplia, 58 

Homalota, 44 

HOMALOTINA, 44 

HOMALOTINI, 43 

Homoeocarabus, 128 

Homoeusa, 45 

Hoplia, 57 

HOPLIINI, 57 

HYDATICINI, 12 

Hydaticus, 12 

Hydnobius, 31 

Hydraena, 27, 128 

HYDRAENIDAE, 27, 128 

HYDRAENINAE, 27 

Hydrobius, 25, 128 

Hydrochara, 25 

Hydrocharis, 25 

HYDROCHIDAE, 24, 128 

Hydrochus, 24, 128 

Hydrocyphon, 59 

Hydroglyphus, 12 

Hydronomus, 103 

HYDROPHILIDAE, 24, 128, 131 

HYDROPHILINAE, 24 

HYDROPHILOIDEA, 23 

Hydrophilus, 25 

Hydroporidius, 12 

HYDROPORINAE, 12 

HYDROPORINI, 12 

Hydroporus, 12, 128 

Hydrosmecta, 41 

Hydrosmectina, 41 

Hydrothassa, 93 

Hydrous, 25 

HYDROVATINI, 13 

Hydrovatus, 13 

Hygriobia, 11 

Hygrobia, 11 

HYGROBIIDAE, 11 

Hygroecia, 42 

Hygronoma, 44 

HYGRONOMINI, 44 

Hygropora, 45 

HYGROTINI, 13 

Hygrotus, 13, 128 

Hylastes, 112, 134 

HYLASTINA, 112 

Hylastinus, 112 

HYLECOETINAE, 68 

Hylecoetus, 68 

HYLESININA, 111 

HYLESININI, 111 

Hylesinus, 111 

Hylis, 61 

HYLOBIINI, 110 

Hylobius, 110 

Hylophilus, 87 

Hylotrupes, 90 

HYLURGINA, 112 

Hylurgops, 112, 134 

Hylurgus, 134 

Hymenodes, 28 

Hypebaeus, 70 

Hypera, 109, 130 

Hyperaspis, 78, 133 

HYPERINAE, 109 

Hyperodes, 134 

HYPHYDRINI, 13 

Hyphydrus, 13 

Hypnogyra, 54 

HYPNOIDINAE, 62 

Hypnoidus, 62, 64, 129 

Hypnota, 41 

163 


Hypocaccus, 26 

Hypocaelus, 61 

Hypocassida, 95 

Hypocoelus, 61 

HYPOCOPRINI, 75 

Hypocoprus, 75 

HYPOCYPHTINI, 44 

Hypocyphtus, 44 

Hypocyptus, 44 

Hypomedon, 51 

HYPOPHLAEINI, 85 

Hypophlaeus, 85 

Hypophloeus, 85 

Hypopycna, 34 

Hypostenus, 49 

Hypothenemus, 134 

Hypulus, 82 

I 

Idolus, 129 

Ilybius, 11, 128 

Ilyobates, 45 

Involvulus, 96 

IPINA, 111 

Ips, 72, 111, 134 

Isarthron, 89 

Ischnodes, 63 

Ischnoglossa, 45 

Ischnomera, 86 

Ischnopoda, 46 

Ischnopterapion, 99 

Ischnosoma, 37 

Isochnus, 102, 130 

Isomira, 85 

Isorhipis, 61 

Ityocara, 45 

IXAPIINI, 98 

Ixapion, 98 

J 

Judolia, 89 

K 

KALCAPIINI, 98 

Kalcapion, 98 

Kateretes, 70 

KATERETIDAE, 70, 129 

Kibunea, 62 

Kissister, 26 

Kissophagus, 112 

Koestlinia, 98 

Kolon, 32 

Korynetes, 69 

KORYNETINAE, 69 

Kryptophagus, 74 

Kyklioacalles, 106 

L 

Labarrus, 56 

Labidostomis, 92 

Laccobius, 25, 128 

LACCOPHILINAE, 13 

Laccophilus, 13 

LACCORNINI, 13


Laccornis, 13 

Lacon, 62 

LAEMOPHLOEIDAE, 73, 129, 132 

Laemophloeus, 73 

Laemostenus, 19 

Laemosthenes, 19 

Laemotmetus, 132 

Lagarus, 19 

Lagocheirus, 133 

Lagria, 84 

LAGRIINAE, 84 

Lamia, 90 

LAMIINAE, 90 

LAMIINI, 90 

Lamprias, 22 

Lamprinodes, 37 

Lamprohiza, 64 

Lamprosoma, 92 

LAMPROSOMATINAE, 92 

LAMPYRIDAE, 64 

Lampyris, 64 

Langelandia, 84 

LANGURIIDAE, 132 

Lar, 80 

Laria, 71, 91 

LARICOBIINAE, 65 

Laricobius, 65 

Larinus, 109, 135 

Lasiodactylus, 132 

Lasioderma, 68 

Lasiorhynchites, 96 

Lasiotrechus, 16 

Latelmis, 60 

Latheticus, 84 

Lathridius, 80 

Lathrimaeum, 34 

LATHROBIINA, 51 

Lathrobium, 51 

LATRIDIIDAE, 79, 133 

LATRIDIINAE, 79 

Latridius, 80 

Lebia, 22 

LEBIINI, 22 

Legaricapion, 99 

Leiochrodes, 133 

Leiodes, 31 

LEIODIDAE, 31, 131 

LEIODINAE, 31 

LEIODINI, 31 

Leiopus, 90 

Leiosoma, 110 

LEIOSOMATINA, 110 

Leiosomus, 110 

Leistophorus, 15 

Leistus, 15 

Leja, 18 

Lema, 91 

Leperisinus, 111 

Leptacinus, 54 

Leptidea, 89 

Leptideella, 89 

Leptinotarsa, 134 

Leptinus, 33 

Leptobium, 128 

Leptophloeus, 74 

Leptura, 88, 133 

LEPTURINAE, 88 

LEPTURINI, 88 

Lepturobosca, 88 

Leptus, 73 

Leptusa, 43 

LEPYRINI, 110 

Lepyrus, 110 

Lesteva, 34 

Leucohimatium, 129, 132 

Leucoparyphus, 37 

Leurostenus, 135 

Liaphlus, 10 

Librodor, 72 

LICININI, 22 

Licinus, 22 

Limarus, 56 

Limnebius, 27 

LIMNICHIDAE, 61, 129 

Limnichus, 61 

Limnius, 60, 129 

Limnobaris, 103 

Limnobius, 27 

Limnocarabus, 15 

Limnoxenus, 25 

Limobius, 109 

Limoniscus, 62 

Limonius, 62 

Liocyrtusa, 31 

Liodes, 31 

Liogluta, 41 

Lionychus, 23 

Liophloeus, 107 

Liopterus, 11 

Liosoma, 110 

Liothorax, 56 

Liparus, 110 

Lissodema, 86 

Listroderes, 135 

Litargus, 81 

Lithocharis, 51 

Lithostygnus, 79 

Litodactylus, 105 

LITOSOMINI, 100 

LIXINAE, 109 

LIXINI, 109 

Lixus, 109, 135 

Llilioceris, 92 

Lobrathium, 51 

Lochmaea, 93 

Lochmaeata, 93 

Lomechusa, 44 

LOMECHUSINA, 44 

LOMECHUSINI, 44 

Lomechusoides, 44 

Longitarsus, 94 

Lopha, 18 

Lophocateres, 69 

LOPHOCATERINAE, 69 

Lordithon, 37 

Loricera, 16 

LORICERINI, 16 

Lorocera, 16 

LUCANIDAE, 55 

LUCANINAE, 55 

Lucanus, 55 

Luperomorpha, 93 

Luperus, 93 

LYCIDAE, 64 

Lycoperdina, 77 

LYCOPERDININAE, 77 

LYCTINAE, 67 

Lyctoxylon, 132 

164 

Lyctus, 67, 132 

Lymantor, 111 

LYMEXYLIDAE, 68 

LYMEXYLINAE, 68 

LYMEXYLOIDEA, 68 

Lymexylon, 68 

Lymnaeum, 18 

Lyperosomus, 18 

Lyphia, 133 

Lyprocorrhe, 41 

Lyprops, 133 

Lythraria, 95 

Lytta, 86 

M 

Macrocephalus, 96 

Macrolister, 131 

Macronychus, 60 

Macroplea, 91 

Macrorhyncolus, 106 

MAGDALIDINI, 110 

Magdalinus, 110 

Magdalis, 110 

MALACHIIDAE, 69, 132 

Malachius, 70 

Mallodon, 133 

MALTHININAE, 65 

MALTHININI, 65 

Malthinus, 65 

Malthodes, 65 

MALTHODINI, 65 

MALVAPIINI, 98 

Malvapion, 98 

Manda, 48 

Mantura, 95 

Margarinotus, 27, 128 

MASOREINI, 22 

Masoreus, 22 

Matidus, 54 

Matthewsium, 28 

MECININI, 101 

Mecinus, 101 

Medon, 51 

MEDONINA, 51 

Megacyllene, 133 

Megadytes, 131 

Megalinus, 55 

MEGALOPODIDAE, 90 

Megapenthes, 63 

MEGAPENTHINI, 63 

Megarhagium, 88 

Megarthrus, 35 

Megasoma, 131 

Megasternum, 25 

Megatoma, 66 

MEGATOMINAE, 66 

Megista, 39 

Megodontus, 15 

Melanapion, 98 

Melandrya, 82 

MELANDRYIDAE, 82, 133 

MELANDRYINAE, 82 

Melanimon, 84 

MELANIMONINI, 84 

Melanobaris, 103 

Melanophila, 59 

MELANOPHILINI, 59 

Melanophthalma, 81

MELANOTINAE, 63 

Melanotus, 63 

Melanthaxia, 59 

MELASINAE, 61 

MELASINI, 61 

Melasis, 61 

Melasoma, 93 

Meligethes, 71 

MELIGETHINAE, 71 

Melinopterus, 56 

Mellieicis, 82 

Meloe, 86, 133 

MELOIDAE, 86, 133 

MELOINAE, 86 

Melolontha, 57 

MELOLONTHINA, 57 

MELOLONTHINAE, 57 

MELOLONTHINI, 57 

MELYRIDAE, 132 

Melyris, 132 

Meneristes, 133 

Meotica, 45 

MEOTICINA, 45 

MEROPHYSIINAE, 77 

Mesites, 105, 135 

Mesocarabus, 15 

Mesocoelopus, 68 

Mesomorphus, 133 

Mesomphalia, 134 

MESOPTILIINAE, 110 

Mesosa, 90 

MESOSINI, 90 

Mesostena, 133 

Mesostenopa, 133 

Metabletus, 23 

Metallina, 17 

Metallites, 108 

Metaphyllobius, 107 

Metastenus, 50 

Metatesnus, 50 

Metaxya, 40 

Metoecus, 83 

Metophonus, 21 

Metophthalmus, 79 

Metopsia, 35 

Mezium, 67, 132 

Miarus, 102 

Miccotrogus, 102 

Micralymma, 34 

Micrambe, 75, 132 

Micrambinus, 75 

Micrapate, 132 

Micraspis, 78 

Micrelus, 104 

Micridium, 29 

Microcara, 59 

Microdota, 41 

Microglossa, 45 

Microglotta, 45 

Microgramme, 80 

Microlestes, 23 

Microlomalus, 27 

MICROPEPLINAE, 35 

Micropeplus, 35, 128 

Microplontus, 104 

Microprius, 133 

Microptilium, 29 

Microrhagus, 61 

Microsaurus, 53 

Microscydmus, 49 

MICROSPORIDAE, 10 

Microsporus, 10 

Micruria, 71 

Micrurula, 71 

Micrus, 29 

Migneauxia, 81 

Millidium, 29 

Minotaurus, 55 

Minthea, 132 

Mirosternomorphus, 68 

Miscodera, 16 

Mitoplinthus, 110 

Mnionomus, 74 

Mniophila, 95 

Mniusa, 45 

Mochtherus, 131 

Mocyta, 41 

Mogulones, 104 

MOLORCHINI, 89 

Molorchus, 89 

Molytes, 110 

MOLYTINA, 110 

MOLYTINAE, 110 

MOLYTINI, 110 

Monanus, 132 

Monochamus, 133 

MONONYCHINI, 105 

Mononychus, 105 

Monotoma, 73, 84 

MONOTOMIDAE, 72 

MONOTOMINAE, 73 

Mordella, 83 

MORDELLIDAE, 83 

Mordellina, 83 

MORDELLINAE, 83 

MORDELLINI, 83 

Mordellistena, 83 

MORDELLISTENINI, 83 

Mordellochroa, 83 

Morica, 133 

Morimus, 134 

Morphocarabus, 15 

Morychus, 60 

MURMIDIINAE, 77 

Murmidius, 77 

Mycetaea, 77 

MYCETAEINAE, 77 

Mycetina, 129 

Mycetochara, 85 

Mycetochares, 85 

MYCETOPHAGIDAE, 81 

Mycetophagus, 81 

MYCETOPORINI, 37 

Mycetoporus, 37 

Mycetota, 41 

Mycota, 39 

MYCTERIDAE, 86 

Mycterus, 86 

Myelophilus, 112 

Mylabris, 91 

Myllaena, 45 

MYLLAENINI, 45 

Myloechus, 32 

Myopinus, 48 

MYRMECHIXENINI, 85 

Myrmechixenus, 85 

Myrmecocephalus, 43 

Myrmecopora, 43 

165 


Myrmecoxenus, 85 

Myrmedonia, 44 

MYRMEDONIINA, 44 

Myrmetes, 26 

Myrmoecia, 44 

Myrrha, 78 

Mysia, 78 

MYXOPHAGA, 10 

Myzia, 78 

N 

Nacerda, 86 

Nacerdes, 86 

NACERDINI, 86 

Nalassus, 84 

Nanodes, 100 

Nanophyes, 100 

NANOPHYIDAE, 100 

Nanoptilium, 29 

Napochus, 49 

Nargus, 32 

Nartus, 11 

Nassipa, 88 

Nathrenus, 66 

Nathrius, 89 

Nausibius, 73 

Neagolius, 129 

Nebria, 15 

NEBRIINI, 15 

Nebrioporus, 12 

Necrobia, 69 

Necrodes, 33 

Necrophorus, 33 

Nedyus, 104 

NEGASTRIINAE, 64 

Negastrius, 64 

Nehemitropia, 41 

Neja, 17 

Neliocarus, 106 

Nemadus, 32 

NEMOGNATHINAE, 86 

Nemoicus, 107 

NEMONYCHIDAE, 96 

Nemosoma, 69 

Nemozoma, 69 

Neobisnius, 52 

Neoceble, 32 

Neoclytus, 134 

Neocoenorrhinus, 97 

Neocrepidodera, 95, 130 

Neoeustolus, 108 

Neohaliplus, 10 

Neohilara, 41 

Neomysia, 78 

Neonapochus, 49 

Neopanus, 110 

Neophytobius, 105 

Neopullus, 78 

Nepha, 18, 131 

Nephanes, 29 

NEPHANINI, 29 

Nephus, 78, 129 

Nestus, 50 

Neuraphes, 49 

Nialus, 129, 131 

Nicobium, 132 

NICROPHORINAE, 33 

Nicrophorus, 33


Nimbus, 56 

Niptus, 67 

Nitidula, 72 

NITIDULIDAE, 71, 132 

NITIDULINAE, 72 

Normandia, 60 

NOSSIDIINAE, 28 

Nossidium, 28 

Notaphemphanes, 17 

Notaphus, 17 

Notaris, 100 

NOTERIDAE, 10 

Noterus, 10 

Nothaphemphanes, 17 

NOTIOPHILINI, 16 

Notiophilus, 16 

Notolaemus, 74, 129 

Notothecta, 42 

NOTOXINAE, 87 

Notoxus, 87, 133 

Nudobius, 55 

O 

Oberea, 90 

OBRIINI, 89 

Obrium, 89 

Ocalea, 45 

Ochina, 67 

Ochrosis, 95 

OCHTHEBIINAE, 27 

Ochthebius, 28, 128 

Ochthephilum, 50 

Ochthephilus, 48 

Ocnera, 133 

Octodesmus, 132 

Octotemnus, 81 

Ocydromus, 17 

Ocypus, 54, 128 

Ocys, 17 

Ocyusa, 45 

Odacantha, 23 

ODACANTHINI, 23 

Odeles, 59 

Odontaeus, 55 

Odonteus, 55 

Odontomagdalis, 110 

Odontonyx, 19 

Oeceoptoma, 33 

Oedemera, 86 

OEDEMERIDAE, 86 

OEDEMERINAE, 86 

OEDEMERINI, 86 

Oedostethus, 64 

Oemona, 134 

Oiceoptoma, 33 

Olibrus, 74 

Oligella, 29 

Oligota, 44, 131 

Olisthopus, 19 

Olophrum, 34, 128 

OMALIINAE, 33 

OMALIINI, 34 

Omalium, 34 

Omaloplia, 58 

Omaseus, 19 

Omiamima, 107 

Omias, 107 

OMIINI, 107 

Omonadus, 87 

Omophlus, 85 

Omophron, 15 

OMOPHRONINAE, 15 

Omosiphora, 71 

Omosita, 72 

Omphalapion, 98 

Oncomera, 86 

Onitis, 131 

Ontholestes, 54 

ONTHOPHAGINI, 57 

Onthophagus, 57, 129, 131 

ONTHOPHILINAE, 27 

Onthophilus, 27 

Onthotomicus, 111 

ONYCHOLIPINI, 106 

Oodes, 22, 128 

OODINI, 22 

Oomorphus, 92 

Oosphaerula, 31 

Ootypus, 76 

OPATRINI, 84 

Opatroides, 133 

Opatrum, 84, 133 

Opetiopalpus, 129 

Ophonus, 21 

Opilo, 69, 132 

Orchesia, 82 

Orchestes, 102 

Orchidophilus, 135 

ORECTOCHILINI, 10 

Orectochilus, 10 

Oreocarabus, 15, 128 

Oreodytes, 12 

Oreostiba, 42 

OROBITIDINAE, 110 

Orobitis, 110 

Orochares, 34 

OROPHIINI, 81 

Orphinus, 66 

Orsodacna, 91 

Orsodacne, 91 

ORSODACNIDAE, 91 

Orthathous, 62 

ORTHOCERI, 96 

ORTHOCERINI, 83 

Orthocerus, 83 

Orthochaetes, 102 

Orthocis, 82 

ORTHOPERINAE, 79 

Orthoperus, 79 

Orthorhinus, 135 

Orthotomicus, 111, 135 

Oryzaephilus, 73, 132 

Osphya, 82 

OSPHYINAE, 82 

Ostoma, 68 

OTHIINI, 52 

Othius, 52 

OTIORHYNCHINI, 107 

Otiorhynchus, 107, 130, 135 

Otiorrhynchus, 107 

Otophorus, 56 

Oulema, 91 

Oulimnius, 60 

Ousipalia, 42 

Oxylaemus, 77 

Oxynoptilus, 13 

Oxyomus, 57 

166 

Oxypleurus, 134 

Oxypoda, 45 

OXYPODINA, 45 

OXYPODINI, 45 

OXYPORINAE, 48 

Oxyporus, 48 

Oxypselaphus, 19 

Oxystoma, 99 

OXYSTOMATINA, 98 

OXYSTOMATINI, 98 

OXYTELINAE, 47 

OXYTELINI, 47 

Oxytelus, 47 

Oxythyrea, 131 

P 

Pachnephorus, 130 

Pachnida, 42 

Pachyatheta, 42 

Pachychila, 133 

Pachydissus, 134 

Pachygluta, 43 

Pachylopus, 26 

Pachymerus, 134 

Pachynephorus, 134 

Pachyrhinus, 107 

Pachyta, 88, 89 

Pachytodes, 89 

Pachytychius, 102 

Paederidus, 51 

PAEDERINA, 51 

PAEDERINAE, 50 

PAEDERINI, 50 

Paederus, 51 

PAELOBIIDAE, 11 

Paelobius, 11 

Palaeonthophagus, 57 

PALORINI, 84 

Palorinus, 133 

Palorus, 84, 133 

PANAGAEINI, 22 

Panagaeus, 22 

PANAPHANTINA, 36 

Panspaeus, 63 

Panspoeus, 63 

Panus, 110 

Parabathyscia, 33 

Parabemus, 54 

Parabolitobius, 37 

Paracelia, 20 

Paracercyon, 25 

Paracorymbia, 89 

Paracycreon, 25 

Paracymus, 24 

Paradromius, 23 

Paragabrius, 52 

Paraliocercyon, 25 

Paralister, 27 

Paramecosoma, 74 

Parameotica, 42 

Paramoeocerus, 134 

Paramysia, 78 

Paranchus, 19 

Paranebria, 15 

Paranopleta, 42 

Paranthonomus, 101 

Paraphloeostiba, 35 

Paraphotistus, 62

Pararaphes, 49 

Parasilvanus, 132 

Parastenus, 49 

Paratachys, 17 

Paratillus, 69 

Paratrogophloeus, 48 

Pardileus, 20 

Parethelcus, 104 

Parnemoicus, 107 

Parnus, 60 

Parocyusa, 46 

PAROMALINI, 27 

Paromalus, 27 

PASSANDRIDAE, 132 

PATROBINI, 18 

Patrobus, 18 

Pediacus, 73 

PEDININI, 84 

Pedius, 18 

Pedostrangalia, 88 

Pelenomus, 105 

Pella, 44 

Pelobius, 11 

Pelochares, 129 

Pelophila, 16 

PELTINAE, 68 

Peltis, 129 

Peltodytes, 10 

Pelurga, 42 

Pentaphyllus, 85 

PENTARTHRINI, 106 

Pentarthrum, 106 

Peranus, 27 

Perapion, 97 

Percosia, 20 

Perigona, 22 

PERIGONINI, 22 

Perileptus, 16 

PERITELINI, 107 

Peritelus, 107 

Peryphus, 17 

Phacophallus, 55 

Phaeaphodius, 129 

Phaedon, 92 

Phaenomerus, 135 

Phaganthus, 34 

PHALACRIDAE, 74 

Phalacrolinus, 54 

Phalacronothus, 56 

Phalacrus, 74 

Phaleria, 85 

PHALERIINI, 85 

Pharaxonotha, 132 

Phila, 18 

Philhydrus, 24 

Philhygra, 42 

Philochthus, 18 

Philocthus, 18 

PHILONTHINA, 52 

Philonthus, 52, 129 

Philopedon, 107 

Philorhizus, 23 

Philorinum, 34 

Philydrus, 24 

PHLOEOCHARINAE, 37 

Phloeocharis, 37 

Phloeodroma, 46 

Phloeonomus, 35 

Phloeophagia, 106 

PHLOEOPHAGINA, 106 

Phloeophagus, 106 

Phloeophilus, 68 

Phloeophthorus, 112 

Phloeopora, 46 

Phloeosinus, 112 

Phloeostiba, 35 

PHLOEOTRIBINA, 112 

Phloeotribus, 112 

Phloeotrya, 82 

PHLOIOPHILIDAE, 68 

Phloiophilus, 68 

Phloiotrya, 82 

Phlyctinus, 135 

Phonopate, 132 

Phoracantha, 134 

Phosphaenus, 64 

Phosphuga, 33 

Phradonoma, 131 

Phratora, 93 

Phryneta, 134 

PHRYNIXINI, 110 

Phthorophloeus, 130 

Phyconoma, 39 

Phyla, 18 

Phylan, 84 

PHYLLOBIINI, 107 

Phyllobius, 107 

Phyllobrotica, 93 

Phyllodecta, 93 

Phyllodrepa, 35 

Phyllodrepoidea, 34 

Phyllopertha, 58 

Phyllotocus, 131 

Phyllotreta, 93 

Phymatodes, 90 

PHYTOBIINI, 105 

Phytobius, 105 

Phytodecta, 93 

Phytoecia, 90 

PHYTOECIINI, 90 

Phytonomus, 109 

PHYTOSINI, 46 

Phytosus, 46 

PIESTINAE, 47 

PIEZOTRACHELINI, 99 

Piezotrachelus, 134 

Pilemostoma, 95 

Pimelia, 133 

Pirapion, 99 

Pissodes, 110, 130 

PISSODINI, 110 

Pityogenes, 111, 135 

Pityophagus, 72 

Pityophthorus, 111 

Placonotus, 132 

Placusa, 46 

PLACUSINI, 46 

Plagiodera, 93 

Plagiogonus, 56 

Plagiometriona, 134 

Plagionotus, 90, 134 

Planeustomus, 48 

Planolinus, 56 

Platambus, 11 

Plataphus, 17 

Plataraea, 42 

Plateumaris, 91 

Platycerus, 55 

167 


Platycis, 64 

Platydema, 85, 133 

Platyderes, 19 

Platyderus, 19 

Platydomene, 51 

Platydracus, 54 

Platynaspis, 78 

PLATYNINI, 19 

Platynus, 19 

PLATYPODIDAE, 112, 130, 135 

PLATYPSYLLINAE, 33 

Platypus, 112, 130, 135 

PLATYRHININI, 96 

Platyrhinus, 96 

Platyrrhinus, 96 

Platysma, 18 

Platysoma, 131 

Platystethus, 47 

PLATYSTOMINI, 96 

Platystomos, 96 

Platystomus, 96 

Plectophloeus, 36 

Plectroscelis, 95 

PLEGADERINI, 26 

Plegaderus, 26 

Pleurophorus, 57 

PLINTHINA, 110 

Plinthus, 110 

Plitium, 29 

Plocaederus, 134 

Plochionus, 131 

Pocadius, 72 

PODABRINI, 64 

Podabrus, 64 

Podagrica, 95 

Poecilaspis, 134 

Poecilips, 135 

Poecilium, 90 

Poecilus, 18 

POGONINI, 18 

Pogonochaerus, 90 

POGONOCHERINI, 90 

Pogonocherus, 90 

Pogonophorus, 15 

Pogonus, 18 

Polistichus, 23 

Polycoelogastridion, 133 

Polydrosus, 108 

POLYDRUSINI, 107 

Polydrusus, 108, 135 

POLYGRAPHINA, 112 

Polygraphus, 112, 135 

POLYPHAGA, 23 

Polyphylla, 57 

Polystichus, 23 

Polystoma, 38 

Polystomota, 38 

Pomatinus, 60 

Poophagus, 104 

Porcinolus, 60 

Porhydrus, 12 

Porotachys, 131 

Porrothus, 110 

Porthmidius, 129 

Potamodytes, 12 

Potamonectes, 12 

Potosia, 58 

Pragensiella, 43 

Prasocuris, 93


Pria, 71 

Princidium, 17 

Priobium, 67, 68 

PRIONINAE, 88 

Prionocyphon, 59 

Prionus, 88 

Prionychus, 85 

Pristonychus, 19 

Probagous, 103 

Procas, 100 

Procraerus, 63 

Procula, 133 

PROECINI, 106 

Promethister, 27 

Propylaea, 78 

Propylea, 78 

Prosternon, 62 

PROSTOMIDAE, 129 

Prostomis, 129 

Protaetia, 58 

Protapion, 99 

PROTEININAE, 35 

Proteinus, 35 

Protopirapion, 99 

Protosilvanus, 132 

Psammobius, 57 

PSAMMODIINA, 57 

PSAMMODIINI, 57 

Psammodius, 57 

Psammoechus, 73 

Psammoecus, 73 

Psammoporus, 56 

PSEBIINI, 89 

Pselactus, 106 

Pselaphaulax, 37 

PSELAPHINAE, 35 

PSELAPHINI, 37 

PSELAPHITAE, 37 

Pselaphorhynchites, 97 

Pselaphus, 37 

PSEPHENIDAE, 61 

Psephidonus, 34 

Pseudapion, 98 

Pseudaplemonus, 97 

Pseudeuparius, 96 

Pseudeurostus, 67 

Pseudocistela, 85 

Pseudocleonus, 130 

Pseudocypus, 54 

Pseudohylesinus, 135 

Pseudolimnaeum, 18 

PSEUDOLIODINI, 31 

Pseudomaseus, 19 

Pseudomedon, 51 

Pseudomesophila, 134 

Pseudomicrodota, 43 

Pseudomordellina, 83 

Pseudoophonus, 20 

Pseudopasilia, 43 

Pseudoperapion, 97 

Pseudophloeophagus, 106 

Pseudophonus, 20 

Pseudoprotapion, 99 

PSEUDOPSINAE, 50 

Pseudopsis, 50 

Pseudorchestes, 102 

Pseudostenotrupis, 135 

Pseudostoreus, 135 

Pseudostyphlus, 102 

Pseudotriphyllus, 81 

Pseudovadonia, 89 

Psilocalymma, 99 

Psilothrix, 69 

Psylliodes, 95 

Psyllobora, 79 

Pteleobius, 112 

PTENIDIINI, 28 

Ptenidium, 28 

PTEROSTICHINI, 18 

Pterostichus, 18, 128 

Pterygorrhynchus, 107 

Pteryx, 29 

PTILIIDAE, 28 

PTILIINAE, 28 

PTILIINI, 29 

PTILININAE, 68 

Ptilinus, 68 

Ptiliola, 29 

Ptiliolum, 29 

Ptilium, 29 

Ptilodactyla, 61 

PTILODACTYLIDAE, 61 

Ptinella, 29 

PTINELLINI, 29 

PTINIDAE, 67, 129, 132 

PTININAE, 67 

PTININI, 67 

Ptinomorphus, 67 

Ptinus, 67, 132 

PTOMAPHAGINI, 33 

Ptomaphagus, 33 

Ptomascopus, 128 

Ptomister, 27 

Ptychandra, 41 

Pullus, 78 

Purrolinus, 55 

Pycnacritus, 26 

Pycnoglypta, 129 

PYCNOMERINAE, 83 

Pycnomerus, 83, 129 

Pycnota, 42 

Pyrochroa, 86 

Pyrochroella, 86 

PYROCHROIDAE, 86 

Pyropterus, 64 

Pyrrhalta, 93 

Pyrrhidium, 90 

PYTHIDAE, 86 

Pytho, 86 

Q 

QUEDIINA, 53 

Quedionuchus, 53 

Quedius, 53 

R 

Rabigus, 53 

Rabocerus, 87 

Ramphus, 102 

Rantus, 11 

Raphirus, 53 

Rayacheila, 54 

Raymondia, 100 

RAYMONDIONYMIDAE, 100 

Raymondionymus, 100 

Reesa, 66 

168 

Reichenbachia, 36, 131 

Remus, 53 

RHADALINAE, 69 

RHAGIINI, 88 

Rhagiomorpha, 134 

Rhagium, 88 

Rhagocneme, 42 

Rhagonycha, 65 

RHAMPHINI, 102 

Rhamphus, 102 

Rhantus, 11 

Rheochara, 38 

RHINOCYLLINI, 109 

Rhinocyllus, 109 

Rhinomacer, 96 

Rhinoncus, 105 

Rhinosimus, 87 

Rhinusa, 102 

RHIPIPHORIDAE, 83 

Rhizobiellus, 77 

Rhizobius, 77 

Rhizopertha, 66 

RHIZOPHAGINAE, 72 

Rhizophagus, 72 

RHIZOTROGINA, 57 

Rhizotrogus, 57 

Rhopalapion, 98 

Rhopalhelophorus, 23 

Rhopalocerina, 43 

Rhopalodontus, 81 

Rhopalomesites, 105 

Rhynchaenus, 102 

Rhynchites, 97, 130 

RHYNCHITIDAE, 96, 130 

RHYNCHITINAE, 96 

RHYNCHITINI, 96 

RHYNCHOPHORINAE, 100 

RHYNCOLINA, 106 

RHYNCOLINI, 106 

Rhyncolus, 106, 130, 135 

Rhypobius, 79 

Rhysodes, 128 

RHYSODIDAE, 128 

Rhysothorax, 56 

RHYSSEMINA, 57 

Rhyssemus, 57, 129 

RHYTHIRRININI, 106 

Rhytideres, 135 

Rhytidosoma, 105 

Rhytidosomus, 105 

Rhytinota, 133 

Rhyzobius, 77 

Rhyzopertha, 66 

Rhyzophagus, 72 

Riolus, 60 

RIPIPHORIDAE, 83 

Risophilus, 22 

Rodolia, 77 

Romaleum, 134 

Romualdius, 109 

Ropalodontus, 81, 129 

Rosalia, 134 

Rugilus, 51 

RUTELINAE, 58 

Rutidosoma, 105 

Rutpela, 89 

Rybaxis, 36 

RYPOBIINAE, 79 

Rypobius, 79

Ryzophagus, 72 

S 

Salebius, 132 

Salius, 102 

Salpingellus, 87 

SALPINGIDAE, 86 

SALPINGINAE, 86 

Salpingus, 87 

Saperda, 90 

SAPERDINI, 90 

SAPRININAE, 26 

Saprinus, 26, 131 

Saprosites, 57 

Sarrotrium, 83 

Scaphidema, 85 

SCAPHIDEMINI, 85 

SCAPHIDIINAE, 47 

SCAPHIDIINI, 47 

Scaphidium, 47 

SCAPHIINI, 47 

Scaphisoma, 47 

SCAPHISOMATINI, 47 

Scaphium, 47 

SCARABAEIDAE, 55, 129, 131 

SCARABAEINAE, 57 

SCARABAEOIDEA, 55 

SCARITINI, 16 

Scarodytes, 12 

Schistoglossa, 42 

Schizotus, 86 

SCIAPHILINI, 108 

Sciaphilus, 108 

Scierus, 135 

Sciodrepa, 33 

Sciodrepoides, 33 

Scirtes, 59 

SCIRTIDAE, 58 

SCIRTOIDEA, 58 

SCLEROPTERINI, 105 

Sclerorhinus, 135 

Scobicia, 132 

SCOLYTINA, 110 

SCOLYTINAE, 110 

SCOLYTINI, 110 

Scolytus, 15, 110, 130, 135 

SCOPAEINA, 51 

Scopaeus, 51 

Scotias, 67 

Scotobius, 133 

Scraptia, 87 

SCRAPTIIDAE, 87 

SCRAPTIINAE, 87 

Scybalicus, 21 

SCYDMAENINAE, 48 

SCYDMAENINI, 49 

Scydmaenus, 49 

Scydmoraphes, 49 

Scymnus, 78, 133 

Selatosomus, 62 

Semanotus, 90 

Semicampa, 18 

Semiris, 40 

Sepedophilus, 37 

Serica, 58 

SERICINI, 58 

Sericoda, 19 

Sericoderus, 79 

Sericosomus, 63 

Sericus, 63 

Sermyla, 93 

Sermylassa, 93 

Serropalpus, 133 

Siagonium, 47 

Sibinia, 102 

Sigorus, 57 

SILINAE, 65 

Silis, 65 

Silpha, 33 

SILPHIDAE, 33, 128 

SILPHINAE, 33 

Silusa, 44 

SILUSINA, 44 

SILVANIDAE, 73, 129, 132 

SILVANINAE, 73 

Silvanoprus, 73, 132 

Silvanus, 73, 132 

Simplocaria, 60, 129 

Sinechostictus, 18 

Sinodendron, 55 

Sinoxylon, 132 

Sipalia, 41 

Sipalinus, 134, 135 

Sirocalodes, 105 

Sitaris, 86 

Sitodrepa, 68 

Sitona, 108 

SITONINI, 108 

Sitophagus, 133 

Sitophilus, 100, 134 

Smaragdesthes, 131 

Smaragdina, 92, 134 

SMICRONYCHINI, 102 

Smicronyx, 102 

Smicrus, 29 

Smodicum, 134 

Sogda, 31 

SOGDINI, 31 

Somotrichus, 22 

Sophronica, 134 

Soronia, 72 

Spatulonthus, 52 

Specularius, 134 

SPERCHEIDAE, 24 

Spercheus, 24 

Spermophagus, 134 

Sphaericus, 67 

SPHAERIDIINAE, 25 

Sphaeridium, 26 

Sphaeriestes, 87 

Sphaerites, 26 

SPHAERITIDAE, 26 

Sphaerius, 10 

SPHAERIUSIDAE, 10 

SPHAERIUSOIDEA, 10 

Sphaeroderma, 95 

Sphaerosoma, 77 

SPHINDIDAE, 70 

Sphindus, 70 

Sphinginus, 70 

SPHODRINI, 19 

Sphodrus, 19 

SPONDYLIDINAE, 89 

Squamapion, 98 

Stagetus, 129 

STAPHYLINIDAE, 33, 128, 131 

STAPHYLININA, 54 

169 


STAPHYLININAE, 52 

STAPHYLININI, 52 

STAPHYLINOIDEA, 27 

Staphylinus, 54 

Stegatus, 132 

Stegobium, 68 

Stelidota, 132 

Stenagostus, 63 

Stenelmis, 60, 129 

Stenichnus, 49 

STENINAE, 49 

Stenister, 27 

Stenocarus, 105 

Stenocera, 131 

Stenochorus, 88 

Stenocorus, 88 

Stenolophus, 21, 131 

STENOPELMINI, 100 

Stenopelmus, 100 

Stenopterapion, 99 

Stenoscelis, 130 

Stenostola, 90 

Stenurella, 89 

Stenus, 49, 129 

Stenygrinum, 134 

Stephanoderes, 135 

Stephanopachys, 66 

Stephostethus, 79 

Stereocorynes, 106 

Sternochetus, 135 

Sternoporus, 12 

Steropus, 18 

Stethomezium, 67 

Stethorus, 78 

Stichoglossa, 46 

Stictoleptura, 88 

Stictonectes, 12 

Stictonotus, 12 

Stictotarsus, 12 

Stilbus, 74 

STILICINA, 51 

Stilicus, 51 

Stolas, 134 

Stomis, 18 

Stomodes, 130 

STOREINI, 102 

Strangalia, 89 

Strategus, 131 

Stricticollis, 87 

Stricticomus, 87 

Strigocis, 81 

Stromatium, 134 

Strophosoma, 106 

Strophosomus, 106 

STYPHLINI, 102 

Subcoccinella, 79 

Subphyllobius, 107 

Subrinus, 131 

Sulcacis, 81 

Sunius, 51 

Suphrodytes, 13 

Syagrius, 110 

Symbiotes, 77 

SYNAPIINA, 99 

Synapion, 99 

Synaptus, 63 

Syncalypta, 60 

SYNCALYPTINAE, 60 

Synchita, 83


SYNCHITINI, 83 

SYNDESINAE, 55 

Synechostictus, 18 

Syntomium, 47 

Syntomus, 23, 128, 131 

Synuchus, 19 

T 

Tachinus, 37, 129 

Tachyerges, 102, 130 

TACHYPORINAE, 37 

TACHYPORINI, 37 

Tachyporus, 38 

Tachypus, 15, 17 

Tachys, 17 

Tachyta, 131 

Tachyusa, 46 

Tachyusida, 43 

TACHYUSINA, 46 

Taeniapion, 98 

TANYMECINI, 108 

Tanymecus, 108 

TANYSPHYRINI, 100 

Tanysphyrus, 100 

Tapeinotus, 105 

Taphria, 19 

Taphrorychus, 111 

Tapinotus, 105 

TARSOSTENINAE, 69 

Tarsostenus, 69 

Tarsostinus, 22 

Tarulus, 22 

Tasgius, 54 

Tatianaerhynchites, 97 

Telephorus, 64 

Telmatophilus, 74 

Temnocerus, 97 

Temnochila, 129 

Tenebrio, 84, 133 

Tenebrioides, 69 

TENEBRIONIDAE, 84, 129, 133 

TENEBRIONINAE, 84 

TENEBRIONINI, 84 

TENEBRIONOIDEA, 81 

Tenebroides, 69, 129, 132 

Tentyria, 133 

TEREDINAE, 77 

Teredus, 77 

TERETRIINI, 26 

Teretrius, 26, 131 

Teropalpus, 48 

Tesarius, 57 

Tesnus, 50 

Testedium, 17 

Tetartopeus, 51 

Tetralaucopora, 46 

Tetrapriocera, 132 

Tetratoma, 82 

TETRATOMIDAE, 82 

TETRATOMINAE, 82 

TETROPINI, 90 

Tetropium, 89, 134 

Tetropla, 39 

Tetrops, 90 

Teuchestes, 57, 129 

Thalassophilus, 16 

Thalycra, 72 

Thamiaraea, 42 

THAMIARAEINA, 42 

Thamiocolus, 105 

Thanasimus, 69 

Thanatophilus, 33, 128 

THANEROCLERINAE, 69 

Thaneroclerus, 69 

Thaumaphrastus, 65 

Thea, 79 

Thectura, 44 

Thecturota, 43 

Thes, 80 

Thiasophila, 46 

Thinobaena, 42 

THINOBIINI, 47 

Thinobius, 48 

Thinodromus, 48 

Thinonoma, 46 

Thomsoneonymus, 108 

THORICTINAE, 65 

Thorictodes, 65, 131 

Thorictus, 131 

THROSCIDAE, 61 

Throscus, 61 

Thryogenes, 100 

Thyasophila, 46 

Thylodrias, 65 

THYLODRIINAE, 65 

Thymalus, 68 

Thymapion, 98 

Tigrinellus, 109 

TILLINAE, 69 

Tilloidea, 69 

Tillus, 69, 132 

Timarcha, 92 

Tinocyba, 99 

Tinotus, 38 

Tipnus, 67 

Tiresias, 66 

Titan, 29 

Tomicus, 111, 112 

Tomocarabus, 128, 131 

Tomoxia, 83 

Tournotaris, 100 

Toxotus, 88 

TRACHEINI, 59 

Trachodes, 110 

TRACHODINI, 110 

Trachyphlaeus, 109 

TRACHYPHLOEINI, 108 

Trachyphloeus, 109 

Trachys, 59 

Traumoecia, 42 

TRECHINI, 16 

Trechoblemus, 16 

Trechus, 16, 128 

Trepanedoris, 18 

Trepanes, 18 

Triaena, 20 

Triarthron, 31 

TRIBALINAE, 27 

TRIBOLIINI, 84 

Tribolium, 84, 133 

TRICHAPIINA, 99 

Trichapion, 99 

Trichelaphrus, 16 

Trichelophorus, 23 

TRICHIINI, 58 

Trichius, 58 

Trichiusa, 42 

170 

Trichocellus, 21 

Trichoderma, 54 

Trichodes, 69 

Trichoferus, 134 

Trichohydnobius, 31 

Trichonotulus, 131 

TRICHONYCHINA, 36 

TRICHONYCHINI, 36 

Trichonyx, 36 

Trichophya, 38 

TRICHOPHYINAE, 38 

Trichoplataphus, 17 

Trichopteryx, 30 

Trichosirocalus, 105 

Trichosphaerula, 31 

Tricorynus, 132 

Trigonogenius, 67 

trigonorhinus, 134 

TRIMIINA, 36 

Trimium, 36 

Trimorphus, 22 

Trinodes, 66 

TRINODINAE, 66 

Trinophyllum, 89 

Trinophylum, 89 

Triodonta, 129 

Triphyllus, 81 

Triplax, 76 

Trissemus, 36 

Tristaria, 132 

Tritoma, 76, 81 

TRITOMINI, 76 

Trixagus, 61 

Trochalus, 131 

Trochoideus, 132 

Trochorhopalus, 134 

TROGIDAE, 55, 131 

Troginus, 48 

Troglops, 70 

Trogoderma, 66, 131 

Trogophloeus, 48 

TROGOSSITIDAE, 68, 129, 132 

TROGOSSITINAE, 69 

Trogoxylon, 132 

Tropideres, 96 

TROPIPHORINI, 109 

Tropiphorus, 109 

Trox, 55, 131 

Trypocopris, 55 

Trypodendron, 111, 135 

Trypophloeus, 111, 135 

Trypopitys, 132 

TYCHIINI, 102 

TYCHINI, 36 

Tychius, 102, 135 

Tychobythinus, 36 

Tychus, 36 

TYLODINA, 106 

Tylodrusus, 108 

Typhaea, 81 

Typhaeus, 55 

Typhlolinus, 55 

Typhoeus, 55 

TYPODERINA, 110 

Tyrtaeus, 133 

Tytthaspis, 78

U 

Ulapion, 98 

Uleiota, 73 

Uloma, 84 

ULOMINI, 84 

Urodon, 96 

URODONTINAE, 96 

Urophorus, 71 

V 

Valgus, 129, 131 

Vanonus, 87 

Variimorda, 83 

Velleius, 54 

Vibidia, 78 

Vincenzellus, 87 

Volinus, 57 

W 

Wankowizium, 28 

X 

Xanthogaleruca, 93 

XANTHOLININI, 54 

Xantholinus, 55 

Xanthomus, 85 

Xenochara, 38 

Xenota, 42 

Xenusa, 43 

Xestobium, 67 

XYLEBORINA, 111 

Xyleborinus, 111 

Xyleborus, 111, 135 

Xylechinus, 112 

XYLETININAE, 68 

Xyletinus, 68, 132 

Xylion, 132 

Xylita, 82 

Xylobiops, 132 

Xylocleptes, 111 

Xylodeleis, 132 

Xylodrepa, 33 

Xylodromus, 35 

Xyloperthella, 132 

Xylophilus, 87 

Xylosandrus, 135 

Xylostiba, 35 

XYLOTERINA, 111 

Xyloterus, 111 

Xylothrips, 132 

171 


Xylotrechus, 134 

Xylotrogus, 67 

Y 

Yrdops, 61 

Z 

ZABRINI, 20 

Zabrotes, 134 

Zabrus, 20 

Zacladus, 105 

Zeugenia, 135 

Zeugophora, 90 

ZEUGOPHORINAE, 90 

Zezea, 20 

Zilora, 82 

Zimioma, 129 

Zimmermannius, 12 

Zorochros, 64 

ZUPHIINI, 23 

ZYGAENODINI, 96 

Zyras, 44

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