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49-77 - SABONET
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20 mm<br />
Acanthopale confertiflora<br />
Del. Sandie Burrows<br />
Blepharis grandis<br />
Del. Sandie Burrows<br />
20 mm<br />
Acanthus ueleensis<br />
Del. Sandie Burrows<br />
Brillantaisia kirungae<br />
Del. Sandie Burrows<br />
20 mm<br />
20 mm<br />
Plants of the Nyika Plateau<br />
<strong>49</strong>
Dicliptera maculata<br />
Del. Sandie Burrows<br />
20 mm<br />
Hypoestes aristata<br />
Del. Sandie Burrows<br />
50 Plants of the Nyika Plateau<br />
20 mm<br />
Dyschoriste verticillaris<br />
Del. Sandie Burrows<br />
20 mm<br />
Isoglossa strigulosa<br />
Del. Sandie Burrows<br />
20 mm
& Synge 214 (K, MAL).<br />
Zambia. Nyika Plateau, Chowo Forest, cf. Dowsett-Lemaire 96 (K?).<br />
Isoglossa substrobilina C.B.Clarke or aff.<br />
Trailing herb with ascending stems and ovate-elliptic tapering leaves, flowers<br />
glandular-hairy in terminal and lateral uninterrupted spike-like racemes,<br />
bracts spoon-shaped or obovate. It is not known whether Dowsett-Lemaire’s<br />
plant has been described as a separate taxon since the publication of her<br />
report and, as such, no detail was available on how it differs from the<br />
above description. Known from the Nyika Plateau and the Misuku Hills,<br />
in montane forest undergrowth.<br />
Source. Dowsett-Lemaire (1985).<br />
Zambia. Nyika Plateau, Chowo and Kasoma Forests, cf. Dowsett-Lemaire<br />
260 (K?).<br />
Isoglossa strigulosa C.B.Clarke �<br />
Ascending herb, 1–1.5 m tall; corolla expanded, white with dark red-purple<br />
marks; in montane forest undergrowth. Known only from the Nyika<br />
Plateau and the Misuku Hills.<br />
Source. Brummitt (1973); PRE.<br />
Zambia. Nyika Plateau, Chowo Forest, 2,160 m, Pawek 10027 (MO, PRE,<br />
UC, K, SRGH, MAL).<br />
Malawi. Nyika National Park, Kawozya, forest E. of summit, 1972, Brummitt<br />
& Synge 215 (K, MAL, SRGH).<br />
Justicia linearispica C.B.Clarke<br />
Herbaceous perennial herb from a woody root; leaves sessile, 25 mm x<br />
6 mm, glabrescent; floral leaves scarcely 6 mm long, narrowly oblong or<br />
linear, sepals 4, flowers ca 8 mm long. Eastern and southern tropical Africa.<br />
Source. UZL; PRE.<br />
Malawi. Nyika National Park, without locality, 1958, Robson 193 (PRE);<br />
without locality, 2000, Mwanyambo 518 (MAL).<br />
No country given. 1967, Richards 22451 (UZL).<br />
Justicia mollugo C.B.Clarke<br />
Minute scabrous hispid annual herb; leaves oblong, up to 19 mm long;<br />
flowers axillary, sessile, clustered, corolla scarcely 5 mm long; in ephemeral<br />
habitats. Eastern tropical Africa.<br />
Malawi. Nyika National Park, Domwe Viewpoint parking area, 2000, Winter<br />
4103 (MAL); Fingira Rock, 2000, Winter 4233 (PRE).<br />
Justicia nuttii C.B.Clarke<br />
Justicia goetzei Lindau; Justicia schliebenii Mildbr.<br />
Small, pubescent herbaceous perennial with clustered stems up to 260 mm<br />
tall; leaves 20–37 mm x ca 15 mm; flowers ca 12 mm long, white with a<br />
few red markings on midlobe of lower lip; common in frequently burnt<br />
high-altitude short grassland. From the Iringa region in S. Tanzania to Dedza<br />
in central Malawi.<br />
Malawi. Nyika National Park, hill E.S.E. of Dembo Bridge, 2000, Winter<br />
4046 (MAL, PRE, SRGH, UZL); Mpopoti, 2000, Mwanyambo 553 (MAL);<br />
2000, Thera 3056 (MAL).<br />
Justicia phyllostachys C.B.Clarke<br />
Slender hairy perennial herb up to 0.6 m tall; leaves up to 100 x 41 mm,<br />
petiole up to 37 mm long; flowers in terminal spikes, floral leaves ca 16 mm<br />
long, ovate or elliptic, sepals 5, flowers 10–19 mm long, varying from mauvepink<br />
to red purple with purple nectar guides or with white markings on<br />
midlobe of lower lip; in scrub or forest margins. D.R.C., Tanzania, Angola,<br />
Zambia, Malawi, Mozambique, and Zimbabwe.<br />
Malawi. Nyika Plateau, 1896, Whyte 118 (K, holo.); North Rukuru Valley,<br />
2000, Winter 4117 (MAL).<br />
Zambia. Nyika Plateau, near Rest House on Chisanga Falls Road, 2000,<br />
Winter 4014 (UZL).<br />
Justicia striata (Klotzsch) Bullock �<br />
Adhatoda striata Klotsch; Justicia melampyrum S.Moore<br />
Very variable, slender herb of open forest and woodland, supported by<br />
surrounding shrubs and grasses; leaves mostly elliptic-oblong, short-petioled,<br />
38–52 mm long, but lower leaves can be broadly ovate with longer petioles;<br />
flowers axillary, floral leaves usually round or ovate, flowers usually<br />
white, also pale mauve or magenta, 8.5–10(–13) mm long. Widespread<br />
throughout tropical Africa. This name may have been misapplied to J.<br />
phyllostachys C.B.Clarke (above).<br />
Malawi. Nyika National Park, Vitinthiza Hill slope, 2000, Mwanyambo<br />
575 (MAL).<br />
Lepidagathis sparsiceps C.B.Clarke �<br />
Procumbent perennial herb with linear leaves; flowers all at base of stems,<br />
heads woolly; on sparsely vegetated, gravelly slopes. Eastern Africa.<br />
Source. Brummitt (1973).<br />
Malawi. Nyika National Park, S. slopes of Kawozya, 1972, Brummitt &<br />
Synge 173 (K, MAL, SRGH); N. Rukuru Valley, 2000, Winter 4110 (MAL).<br />
Mellera submutica C.B.Clarke �<br />
Onus submuticus (C.B.Clarke) Gilli; Pseudobarleria lindaui Dewevre<br />
Perennial similar to Mimulopsis, 1 m tall; leaves ovate, 75 x 50 mm, cordate,<br />
margin toothed; inflorescence pyramidal, glandular, flowers 25 mm<br />
long, lilac-blue with orange throat, tube gradually dilated upwards, mouth<br />
very oblique, 2-lipped, anticous lobe twice as broad as the others. Eastern<br />
Africa.<br />
Source. Brummitt (1973).<br />
Malawi. Nyika National Park, S. slopes of Kawozya, 1972, Brummitt &<br />
Synge 186 (K, MAL, SRGH).<br />
Metarungia pubinervia (T.Anders.) Baden �<br />
Anisotes pubinervis (T.Anders.) Heine; Macrorungia pubinervia<br />
(T.Anders.) C.B.Clarke<br />
Shrub or rarely small tree similar to Anisotes nyassae, 2–5 m high, sometimes<br />
with aerial roots; leaves short-stalked, to 200 x 80 mm; spikes axillary,<br />
one-sided, 60 mm long, hanging, often borne on bare branches below<br />
the leaves; bracts pinkish, ovate to elliptic, flat, 10–14(19) mm long, with a<br />
hyaline margin; flowers red or maroon, rarely white, 30 mm long; lower<br />
lip lobed at tip only; in undergrowth of drier forests. Ethiopia to South<br />
Africa. Baden’s (1981b) assertion (based on a single character) that<br />
Macrorungia was not closely related to Anisotes is unconvincing.<br />
Source. Dowsett-Lemaire (1985).<br />
Malawi. Nyika National Park, Kasyaula Forest, cf. Dowsett-Lemaire 188<br />
(K?).<br />
Zambia. Nyika Plateau, Kasoma Forest, Dowsett-Lemaire 259 (K?).<br />
Mimulopsis solmsii Schweinf. �<br />
Mimulopsis violacea Lindau; Mimulopsis sesamoides S.Moore<br />
Ascending, scrambling shrub up to 2.5 m tall, often covered with long reddish<br />
hairs; leaves ovate, to 150 mm, often doubly round- or sharp-toothed;<br />
inflorescence glandular, flowers trumpet-shaped subequally 5-lobed, 30–<br />
35 mm long, white to pale blue or lilac with orange markings, or yellowish<br />
with purple guide lines within. Widespread in tropical Africa.<br />
Source. Brummitt (1973); Dowsett-Lemaire (1985).<br />
Malawi. Nyika National Park, Kawozya summit, 1972, Brummitt & Synge<br />
200 (K, MAL).<br />
Zambia. Nyika Plateau, Manyenjere and Chowo Forests, cf. Dowsett-Lemaire<br />
261 (K?).<br />
Monechma varians C.B.Clarke �<br />
Stems simple, 300 mm from a woody rootstock; flowers white, similar to<br />
those of Justicia; seeds solitary in each cell, and smooth (2 per cell and<br />
rough in Justicia). Endemic to the Nyika Plateau.<br />
Source. Brummitt (1973).<br />
Malawi. Nyika National Park, Sawi Valley, 1972, Synge 437 (K, MAL).<br />
Peristrophe aculeata (C.B.Clarke) R.K.Brummitt �<br />
Dicliptera aculeata C.B.Clarke<br />
Similar to P. paniculata, but perennial, and with a 17–27 mm long corolla;<br />
in woodland. Northern Malawi. This record casts further doubt onto the<br />
occurrence of P. paniculata on the Nyika Plateau. The placental type is the<br />
only consistent character for distinguishing Peristrophe from Dicliptera<br />
(Balkwill 1996a, 1996b).<br />
Plants of the Nyika Plateau<br />
51<br />
ACANTHACEAE
Justicia striata<br />
Del. Sandie Burrows<br />
10 mm<br />
20 mm<br />
52 Plants of the Nyika Plateau<br />
20 mm<br />
20 mm<br />
Lepidagathis sparsiceps<br />
Del. Sandie Burrows<br />
Mellera submutica<br />
Del. Sandie Burrows
Mimulopsis solmsii<br />
Del. Sandie Burrows<br />
20 mm<br />
20 mm<br />
Monechma varians<br />
Del. Sandie Burrows<br />
Metarungia pubinervia<br />
Del. Sandie Burrows<br />
20 mm<br />
Peristrophe aculeata<br />
Del. Sandie Burrows<br />
20 mm<br />
Plants of the Nyika Plateau<br />
53
ACANTHACEAE<br />
Source. Brummitt (1991).<br />
Malawi. Nyika Plateau, 1,220–1,830 m, 1896, Whyte s.n. (K, lecto.), 1,830–<br />
2,135 m, Whyte s.n. (K).<br />
Peristrophe paniculata (Forssk.) Brummitt<br />
Dianthera bicalyculata Retz.; Justicia bicalyculata (Retz.) Vahl;<br />
?Peristrophe dewewrei De Wild. & Durand<br />
Usually unbranched erect annual herb to 2 m, stems 6-ridged; leaves shortlystalked,<br />
60–110 x 30–40 mm; flowers magenta, 10 mm long, in a large<br />
branched panicle, paired bracts very different in size; in open bush land<br />
and dry rocky soils up to 1,500 m. Namibia to India. As the dry habitat<br />
typical for this taxon is not represented in the Nyika National Park, there is<br />
a possibility that the name is misapplied here. The situation requires investigation<br />
to assess the occurrence of P. paniculata and the identity of P.<br />
dewewrei. Similar to Dicliptera maculata.<br />
Malawi. Nyika National Park, without locality or collecting details (Mill<br />
1979).<br />
Peristrophe pumila (Lindau) Gilli<br />
Dicliptera pumila (Lindau) Dandy ex Brenan; Peristrophe usta<br />
C.B.Clarke<br />
Creeping or suberect, hairy pyrophyte, with stems 100 mm tall from a woody<br />
rootstock; leaves elliptic, to 40 mm long; flowers in erect spikes, bracts<br />
with dense stalked glandular hairs, petals white, mauve-tipped or magenta;<br />
in short or open and wooded grassland, usually flowering before the leaves<br />
are fully developed, soon after fire. Sudan to Zimbabwe.<br />
Source. Brummitt (1973).<br />
Malawi. Nyika National Park, valley N. of Nganda, 1972, Synge 387 (K,<br />
MAL).<br />
Phaulopsis imbricata (Forssk.) Sweet �<br />
Ruellia imbricata Forssk.; Aetheilema imbricatum (Forssk.) Spreng.;<br />
Micranthus imbricatus (Forssk.) O.Kuntze<br />
Glandular-hairy trailing herb with ascending stems and stalked leaves; leaves<br />
of a pair usually unequal, bases asymmetrical; flowers white with a bent tube,<br />
small, in clusters with overlapping, pale, ciliate bracts; in shade or semi-shade in<br />
woodland or forest margins. Widespread in eastern and southern Africa.<br />
Malawi. Nyika National Park, Fingira Rock, 2000, Winter 4219 (MAL).<br />
Thunbergia alata Bojer ex Sims �<br />
Thunbergia fuscata T.Anders. ex Lindau; Valentiana volubilis Raf.<br />
Climbing perennial herb up to 3 m, petiole winged; leaves arrow-shaped<br />
with five veins from the base; flowers yellow or orange, with a black or<br />
Anisotes nyassae Baden<br />
Found in the Karonga district of Malawi, S. Tanzania, and N.E. Zambia.<br />
Dowsett-Lemaire’s record appears to be a sight record, based on material<br />
in bud, which ‘appeared similar to A. nyassae collected in the nearby forests<br />
of the N. Viphya.’ Since she recorded the similar Metarungia pubinervia<br />
as well, it seems possible that both taxa could occur on the Nyika, but<br />
since there is no specimen to confirm its existence, it is excluded here.<br />
Malawi. Nyika Plateau, eastern escarpment submontane forests (Dowsett-<br />
Lemaire 1985: 354).<br />
Blepharis buchneri Lindau<br />
Large leaf pair in whorls more than (3–)5 times longer than smaller pair.<br />
Smaller pair rhomboid-triangular, 3(–7)-veined from the base, rarely<br />
pinnately veined. The correct application of this name is doubted, as B.<br />
buchneri is not recorded for Malawi in the recent revision, and as this<br />
name has often been used erroneously for what is actually B. grandis.<br />
Source. Vollesen (2000).<br />
Malawi. Nyika National Park, Runyina River, 2000, Phiri 3784 (MAL).<br />
No country given. Verboom s.n. (UZL).<br />
Dyschoriste albiflora Lindau<br />
The authenticity of this record could not be confirmed. Known only from<br />
54 Plants of the Nyika Plateau<br />
Excluded species<br />
deep purple throat; bush land and thicket. Widespread in eastern and W.<br />
Africa; also on Mauritius.<br />
Source. Brummitt (1973).<br />
Malawi. Nyika National Park, Vitinthiza Hill slope, 2000, Mwanyambo<br />
576 (MAL); Jalawe Viewpoint, 2000, Thera 3074 (MAL); North Rukuru Valley,<br />
2000, Winter 4116 (MAL, PRE); 8 km E. of Nganda, 1972, Brummitt,<br />
Munthali & Synge 125 (K, MAL, SRGH).<br />
Thunbergia kirkiana T.Anders.<br />
Low semi-procumbent perennial herb; leaf bases sometimes slightly winged,<br />
giving the leaf an arrow shape; flowers large, translucent white with a yellow<br />
throat; in miombo woodland, spring-flowering. D.R.C., Tanzania,<br />
Angola, Zambia, Malawi, Mozambique, and Zimbabwe.<br />
Malawi. Nyika National Park, without locality or collecting details (Patel 1999).<br />
Thunbergia lancifolia T.Anders.<br />
Herbaceous perennial up to 600 mm tall, with erect annual stems from a<br />
woody rootstock; leaves narrowly elliptic; flowers ca 50 mm diameter, pale<br />
to dark blue or tinged with violet, throat yellow, corolla tube slightly inflated,<br />
calyx covered with minute glistening hairs; in open grassland, or<br />
along the edge of Brachystegia woodland, spring-flowering. From Ethiopia<br />
to Zimbabwe.<br />
Malawi. Nyika National Park, without locality or collecting details (Mill 1979).<br />
Thunbergia mollis Lindau<br />
Stem with a rusty pubescence or glabrous; leaves ovate, cordate or hastate,<br />
60 x 36 mm, strigose on both sides, petiole rusty pubescent, 25 mm long;<br />
bracteoles 5-nerved, the middle one forming a marked keel; corolla large,<br />
purple above, tube 38 mm long; habitat unknown. Distribution range outside<br />
of Malawi is unknown.<br />
Malawi. Nyika National Park, without locality or collecting details (Mill<br />
1979).<br />
Thunbergia petersiana Lindau<br />
Stem-climbing, with scattered hairs; leaves cordate, long acuminate, 75 x<br />
57 mm, pilose on both sides, petiole more than 50 mm long; bracteoles 3nerved<br />
and 3-keeled, reticulated; corolla large, violet, tube 25 mm long. As<br />
this taxon was originally known from the lower Zambezi River basin in<br />
Mozambique, presumably in a different habitat, there is some doubt as to<br />
whether the Nyika taxon is the same.<br />
Source. Dowsett-Lemaire (1985).<br />
No country given. Nyika Plateau, cf. Dowsett-Lemaire 263 (K?).<br />
Tanzania.<br />
Malawi. Nyika National Park, without locality or collecting details (Patel<br />
1999).<br />
Justicia kirkiana T.Anders.<br />
Justicia fasciata Lindau<br />
Slender erect annual herb; leaves narrowly lanceolate, up to 62 x 5 mm;<br />
flowers yellow, with a dark stripe on each lobe of the lower lip, bracts and<br />
calyx with long, cottony hairs. Known only from sandy savanna at low<br />
altitudes (Flora of southern Africa; PRE specimens from other localities). A<br />
doubtful record perhaps confused with J. flava (Vahl) Vahl.<br />
Malawi. Nyika National Park, without locality or collecting details (Patel<br />
1999).<br />
Justicia petiolaris (Nees) T.Anders. subsp. bowiei<br />
(C.B.Clarke) Immelman<br />
Justicia mutica C.B.Clarke<br />
This species is known only from the Eastern Cape Province of South Africa.<br />
Confusion with J. linearispica, a species found on Nyika Plateau and<br />
from the same section, is suspected.<br />
Malawi. Nyika National Park, Mpopoti Peak, 2000, Izidine 62 (LMA), det:<br />
H. Patel.
Phaulopsis imbricata<br />
Del. Sandie Burrows<br />
20 mm<br />
10 mm<br />
Thunbergia alata<br />
Del. Sandie Burrows<br />
20 mm<br />
Alangium chinense<br />
Del. Sandie Burrows<br />
Plants of the Nyika Plateau<br />
55
ALANGIACEAE<br />
Alangium chinense (Lour.) Harms �<br />
Tree 9–30 m tall, leaves simple, base asymmetric, flowers white to yellow,<br />
?sweetly scented; in lowland and upland forest, next to streams; up to<br />
2,000 m in this area. Widespread in central Africa, extending through tropical<br />
Asia to Japan and Indonesia. A very rare tree in the Flora zambesiaca<br />
Note. It appears that this relatively large family is under-collected on the<br />
Nyika and many more than the three taxa listed below occur there.<br />
Achyranthes aspera L. var. pubescens (Moq.) Townsend �<br />
Erect or straggling perennial herb, stems 4-angled, leaves opposite, broadly<br />
elliptic, hairy; flowers in a slender terminal spike, minute and enclosed in a<br />
downward-pointing spiny bract; in many habitats, including evergreen forest,<br />
scrub, woodland, often as a weed in disturbed areas. Widespread<br />
throughout the tropics.<br />
Malawi. Nyika National Park, without locality or collecting details (Mill<br />
1979).<br />
Amaranthus hybridus L. subsp. cruentus (L.) Thell. �<br />
Amaranthus cruentus L.; Amaranthus paniculatus L.; Amaranthus<br />
chlorostachys auctt. non Willd.<br />
Robust annual herb to 2 m tall or more, leaves broadly lanceolate to rhomboid,<br />
more or less hairless; flowers in a dense terminal spike, red, green or<br />
Lannea discolor (Sond.) Engl.<br />
livelong; Chichewa: sidyatunga; Yao: chiumbu<br />
Deciduous tree to 15 m tall, leaves alternate, imparipinnate, leaflets in 4–<br />
10 pairs, dark green above, whitish below; fruit a red to purple berry 9–15<br />
x 7–10 mm, fleshy and edible; in open woodland of various types and on<br />
rocky hillsides. South Africa to D.R.C., Zambia, and Malawi. Bark used<br />
to cure diarrhoea, the poles used as a ‘live fence’. Fruit edible and relished.<br />
Malawi. Nyika National Park, without locality or collecting details (Mill<br />
1979).<br />
Lannea edulis (Sond.) Engl. �<br />
Chichewa: ufulukula; Yao: mndyankamba<br />
Suffrutex shrublet with annual stems arising from a large woody rootstock,<br />
leaves imparipinnate, leaflets in 1–3 pairs, hairy when young, glabrous<br />
later; fruit at ground level, in clusters, red, 9–11 x 6–7 mm, fleshy<br />
and edible; in open woodland, rocky hillsides, grassland, most evident after<br />
fires when it flowers. South Africa to Angola, D.R.C., and Uganda. A<br />
root extract is used to treat diarrhoea. Fruit is eaten.<br />
Malawi. Nyika National Park, without locality or collecting details (Mill<br />
1979).<br />
Ozoroa insignis Delile subsp. reticulata (Baker f.) Gillett �<br />
Much-branched evergreen tree to 15 m tall, leaves simple, alternate or in<br />
whorls of 3, variable in shape, hairy or not above, densely hairy below,<br />
veins prominently parallel; in various types of woodland, in savanna, rocky<br />
hillsides, and grassland. Three varieties of subsp. reticulata are recognized,<br />
but the identity of the Nyika material is unknown.<br />
Source. Flora zambesiaca 2(2): 576 (1966).<br />
Malawi. Nyika National Park, without locality or collecting details (Mill<br />
1979); Nyika National Park, without locality or collecting details (Patel<br />
1999).<br />
Rhus anchietae Ficalho & Hiern ex Hiern<br />
Dense shrub or small tree to 8 m tall; leaves 3-foliate, leaflets rhombic to<br />
elliptic, discolorous, hairy or not, veins prominent below; in riverine forest,<br />
swamps and savannas. Malawi to Zambia, Angola, D.R.C., and Tanzania.<br />
Source. Flora zambesiaca 2(2): 612 (1966)<br />
56 Plants of the Nyika Plateau<br />
ALANGIACEAE<br />
AMARANTHACEAE<br />
ANACARDIACEAE<br />
area; in Malawi known only from two other areas (Misuku Hills and<br />
Usumara on the Viphya).<br />
Source. Dowsett-Lemaire (1990: 75).<br />
Malawi. Nyika Plateau, Kasyaula Forest, Dowsett-Lemaire 335 (FHO).<br />
yellowish; in disturbed grassland, a weed of cultivation. This subspecies is<br />
presumed to be of (Central) American origin, but is now widespread in the<br />
tropical and subtropical regions of the world. However, if collected in the<br />
remote and almost uninhabited Nyika of 1903 (see specimen), then it seems<br />
unlikely to have been introduced.<br />
Source. Flora zambesiaca 9(1): 14 (1988).<br />
Malawi. Nyika Plateau, 1903, McClounie 157 (K).<br />
Cyathula cylindrica Moq. �<br />
Perennial straggling herb with opposite simple leaves; inflorescence distinctive,<br />
a congested cylindrical spike of creamy bristle-like bracts, the spikes becoming<br />
attached to animals’ fur and humans’ clothing. Scrambling in forests margins,<br />
montane scrub, and Brachystegia woodland; 1,150–2,300 m. Widespread<br />
in Africa from Cameroon and Sudan S. to South Africa (Cape).<br />
Source. Flora zambesiaca 9(1): 82 (1988).<br />
Malawi. Nyika Plateau, 2,180 m, 1975, Pawek 9968 (K, MAL, MO, SRGH,<br />
UC).<br />
Malawi. Nyika National Park, without precise locality, 1960, Chapman <strong>77</strong>1<br />
(COI, SRGH).<br />
Rhus longipes Engl. var. longipes �<br />
large-leaved rhus; Chichewa: kamwamadzi, mdima; Yao: mpilakuru,<br />
mtatu<br />
Shrub or small tree to 8 m, or scandent, leaves 3-foliate, leaflets elliptic or<br />
obovate, hairy when young, hairless with age, veins obscure above when<br />
young, but prominent below when older; in forests, thickets and woodlands.<br />
Zimbabwe to W. Africa and Kenya.<br />
Source. Brummitt (1973).<br />
Malawi. Nyika National Park, head of Mondwe Valley, 1972, Brummitt &<br />
Synge 237 (K, MAL, SRGH, LISC); Mbuzinandi, 2000, Mwanyambo 584<br />
(MAL); Sangule Peak, 2000, Izidine 95 (LMA).<br />
Zambia. Nyika Plateau, on path to N. Rukuru Waterfall, 1958, Robson 393A<br />
(K, LISC).<br />
Rhus ochracea Meikle var. ochracea<br />
Rhus squalida sensu White (1962), non Meikle<br />
Suffrutex or shrub to 2 m, from a large woody rootstock, leaves 3-foliate,<br />
leaflets elliptic to obovate, leathery, sparsely to densely hairy on both sides,<br />
discolorous, yellowish green to russety below; fruit small, to 4 mm in diameter,<br />
round, shiny; in miombo woodland, in grasslands among rocks, on<br />
termite mounds. Zambia, Malawi, and Tanzania. This is the first record<br />
for the Nyika Plateau.<br />
Malawi. Nyika National Park, Jalawe Viewpoint, 2000, Thera 3084 (MAL).<br />
Rhus pyroides Burch. var. pyroides<br />
Rhus vulgaris Meikle<br />
Shrub or tree to 9 m tall, leaves 3-foliate, leaflets ovate to obovate, apex<br />
rounded or obtuse, base tapering, lateral and tertiary nerves sunken above,<br />
prominently raised below, dark green above, paler and silvery hairy below;<br />
fruit red or brownish red, round, ca 5 mm diameter; on termite mounds<br />
and rocky hills, along stream banks, and on forest margins. South Africa to<br />
Ethiopia.<br />
Source. Flora zambesiaca 2(2): 604 (1966); Moffett (1993: 43).<br />
Malawi. Nyika Plateau, N. of Mpanda Summit, Lewis 104 (FHO); North<br />
Rukuru Valley, 2000, Winter 4121 (MAL).
Achyranthes aspera var. pubescens<br />
Del. Sandie Burrows<br />
20 mm<br />
20 mm<br />
Cyathula cylindrica<br />
Del. Sandie Burrows<br />
10 mm<br />
125 mm<br />
Lannea edulis<br />
Del. Sandie Burrows<br />
Amaranthus hybridus subsp. cruentus<br />
Del. Sandie Burrows<br />
20 mm<br />
100 mm<br />
Plants of the Nyika Plateau<br />
57
Ozoroa insignis subsp. reticulata<br />
Del. Sandie Burrows<br />
58 Plants of the Nyika Plateau<br />
20 mm<br />
Del. Sandie Burrows<br />
20 mm<br />
Anisophyllea boehmii<br />
20 mm<br />
Rhus longipes var. longipes<br />
Del. Sandie Burrows<br />
20 mm<br />
Annona senegalensis<br />
Del. Sandie Burrows
Anisophyllea boehmii Engl. �<br />
Anisophyllea pomifera Engl. & Brehmer<br />
Tumbuka: mangondo; Tonga: mfungo; Ngoni: mahota<br />
Small evergreen tree to 10 m, young branchlets crimson, leaves simple,<br />
ovate to broadly elliptic, leathery dark green, midrib yellowish; flowers<br />
small on long slender spikes; fruit a round to ellipsoid drupe, ca 35 x 20 mm,<br />
Annona senegalensis Pers. �<br />
wild custard-apple; Chichewa: mpoza, mneche, ulembe; Tumbuka:<br />
mdopa, munyele, mulolo; Yao: chiuta, mponjela, mposa<br />
Small, often spreading tree 1.5–8 m tall; leaves alternate, 60–185 x 30–<br />
115 mm, venation sunken above, prominent below; fruit single, 15–30 mm<br />
long, segmented, yellowish, edible; in various types of woodland or wooded<br />
grassland; 0–1,900 m. Widespread throughout tropical Africa.<br />
Malawi. Nyika National Park, without locality or collector (Mill 1979).<br />
Artabotrys monteiroae Oliv. �<br />
red hook-berry; Chichewa: kana; Yao: mtandaumbiri, ntandandumbili<br />
Shrub or liane 1.2–6 m tall, bearing small hooks on the branches; leaves<br />
alternate, oblong to ovate, 30–140 mm long, glossy, hairless above, finely<br />
hairy below; fruit in few-fruited clusters, small, red when ripe; in evergreen<br />
forest, riverine forest, woodland, and rocky outcrops; up to 2,200 m. South<br />
Africa to Uganda and D.R.C.<br />
Malawi. Nyika National Park, Kasyaula and Zovochipolo Forests (Dowsett-<br />
Lemaire 1985: 319).<br />
Zambia. Nyika Plateau, Manyenjere, Kasoma, and Chowo Forests (Dowsett-<br />
Lemaire 1985: 319).<br />
20 mm<br />
Artabotrys monteiroae<br />
Del. Sandie Burrows<br />
ANISOPHYLLACEAE<br />
ANNONACEAE<br />
plum-coloured, edible; in miombo woodland on sandy or rocky soils; 600–<br />
1,600 m. Malawi, Zambia, Tanzania, and D.R.C. Fruit much relished; the<br />
roots are used in Zambia as a snakebite remedy.<br />
Source. Flora zambesiaca 4: 97 (1978); Lebrun & Stork (1991: 167).<br />
Malawi. Nyika National Park, Thazima Gate, 2000, Burrows & Maroyi 6837<br />
(MAL, PRE).<br />
Artabotrys stolzii Diels<br />
Yao: chisakalawe<br />
Large woody climber to 15 m tall, very similar to A. monteiroae; in mist<br />
forest and evergreen forest: 1,500–2,100 m. Confined to the Nyika in<br />
Malawi and S. Tanzania. Regarded as a synonym of A. monteiroae by<br />
White et al. (2001).<br />
Source. Flora zambesiaca 1(1): 133 (1960).<br />
Malawi. Nyika National Park, without precise locality, 1955, Lees 66 (K).<br />
Friesodielsia obovata (Benth.) Verdc. �<br />
Popowia obovata (Benth.) Engl. & Diels<br />
northern dwaba-berry; Chichewa: mchinga, kombe, mufulafula (fruits);<br />
Yao: makatama, mkotamu<br />
Shrub to small tree 1–5 m tall; leaves alternate, oblong-obovate, 60–140 x<br />
35–95 mm, finely hairy or hairless, often glaucous below; fruit a cluster of<br />
3–6 carpels, each 1- to 4-seeded, pinkish to orange-yellow; in open woodland,<br />
thickets, and along rivers, often on granitic soils. Mainly a species of<br />
the Flora zambesiaca region.<br />
Source. Flora zambesiaca 1(1): 123 (1960).<br />
Malawi. Nyika Plateau, Mwenembwe, 1903, McClounie 148 (K).<br />
Friesodielsia obovata<br />
Del. Pat Halliday<br />
Plants of the Nyika Plateau<br />
59<br />
ANISOPHYLLACEAE
20 mm<br />
20 mm<br />
Alepidea peduncularis<br />
Del. Sandie Burrows<br />
60 Plants of the Nyika Plateau<br />
Afrocarum imbricatum<br />
Del. Sandie Burrows<br />
3,5 mm<br />
Agrocharis incognita<br />
Del. Sandie Burrows<br />
4 mm<br />
Centella asiatica<br />
Del. Sandie Burrows<br />
20 mm<br />
20 mm
Afrocarum imbricatum (Schinz) Rauschert �<br />
Baumiella imbricata (Schinz) H.Wolff.; Carum imbricatum Schinz<br />
Herb to 1 m high; leaves glabrous, pinnate, leaflets in 16–32(40) pairs,<br />
overlapping, sharply toothed; bracts commonly trisect or pinnate above;<br />
flowers cream-coloured; in wetlands, grassland, and woodland. Zimbabwe<br />
to Angola, C.A.R., and Tanzania. A monotypic genus endemic to highlands<br />
of the central African plateau. African relatives include Berula,<br />
Oenanthe, and Sium.<br />
Source. Brummitt (1973).<br />
Malawi. Nyika Plateau, road to Chelinda, Richards 10507 (K); W. foot of<br />
Nganda, 1972, Brummitt, Munthali & Synge 37 (K, MAL, SRGH, EA); Dembo<br />
River, 2000, Winter 4034 (MAL, SRGH, LMA, PRE, UZL).<br />
Agrocharis incognita (Norman) Heyw. & Jury �<br />
Caucalis incognita Norman; Torilis eminii Engl.; Torilis gracilis Engl.<br />
forma umbrosa Engl.<br />
Annual herbs (or short-lived perennial with a slender rootstock) set with<br />
bristly, tubercle-based hairs; leaves 2(–3)-pinnate, pinnae with broadly oblong-elliptic<br />
segments, more rarely (when tripinnate) the segments narrowly<br />
elliptic; umbels compound, involucre present beneath main umbel, petals<br />
greenish to white or creamy white; fruit with glochidiate spines, secondary<br />
ribs spinose, primary ribs pilose; in montane grassland, forest margins and<br />
clearings, or in deep shade in damp places, and as a weed of agriculture,<br />
900–3,000 m. Tropical eastern Africa to Sudan, D.R.C., Malawi, E. Zimbabwe,<br />
and adjacent Mozambique. The relationship between Agrocharis<br />
and Daucus is being investigated.<br />
Source. Patel (1999).<br />
Malawi. Nyika National Park, margin of Zovochipolo Forest, 2000, Winter<br />
4131 (MAL).<br />
Agrocharis melanantha Hochst.<br />
Agrocharis pedunculata (Baker f.) Heyw. & Jury; Caucalis melanantha<br />
(Hochst.) Hiern; Caucalis longisepala H.Wolff; Caucaliopsis stolzii<br />
H.Wolff.; Gynophyge tansaniensis Gilli<br />
Perennial pyrophytic herb with a thickened rootstock, 100 mm tall, set<br />
with bristly, tubercle-based hairs; leaves 3- to 4-pinnate into linear to linear-oblong<br />
segments, often subverticillate in appearance; umbels compound,<br />
with distinct and often long rays; involucre present beneath main umbel,<br />
petals greenish to yellowish or blackish purple; fruit with glochidiate spines,<br />
secondary ribs spinose, primary ribs pilose; in moist, short, montane grassland<br />
subject to frequent burning. Widespread in eastern Africa.<br />
Source. Brummitt (1973); Patel (1999).<br />
Malawi. Nyika National Park, between Nganda and Muzengapakweru,<br />
1972, Synge 359 (K, MAL, SRGH, EA); road to Chelinda bridge, near Chosi<br />
Hill turnoff, 2000, Winter 4052 (MAL, PRE); Nganda Hill crest, 2000, Winter<br />
40<strong>77</strong> (MAL, SRGH, PRE).<br />
Alepidea peduncularis A.Rich. �<br />
Alepidea gracilis Dümmer; Alepidea longifolia E.Mey. ex Dümmer;<br />
Alepidea propinqua Dümmer; Alepidea swynnertonii Dümmer<br />
Slender to robust herb to 1.2 m tall with a basal rosette of leaves; basal leaf<br />
margins barely to conspicuously ciliate; flowers in a large panicle of condensed<br />
simple umbels, involucral bracts prominent, white; in montane grassland<br />
and open miombo woodland. South Africa to Ethiopia. The huge variation<br />
in this aggregate species is not well understood and suspiciously simplified.<br />
Source. Cannon (1978); Townsend (1989); Lebrun & Stork (1992).<br />
Malawi. Nyika National Park, between Lake Kaulime and Zovochipolo<br />
Forest, 2000, Winter 4002 (MAL, SRGH, PRE); Nganda Hill, near crest,<br />
2000, Winter 4080 (MAL, SRGH, PRE, UZL); 2 km upstream of Wovwe<br />
River Gorge, 2000, Winter 4150 (MAL, PRE); Chisanga Falls footpath, 2000,<br />
Burrows & Burrows 6752 (MAL, PRE).<br />
Zambia. Nyika Plateau, N. of Zambian Rest House, ca 1 km E. of Sonkwe<br />
stream, 2000, Winter 4019 (UZL, PRE).<br />
Centella asiatica (L.) Urb. �<br />
Centella bupleurifolia (A.Rich.) Adamson; Centella coriacea Nannf.<br />
APIACEAE<br />
Low creeping perennial herb, stems rooting at nodes; leaves reniform; margins<br />
regularly crenate all round; umbels simple, axillary, petals greenish to<br />
purplish red; fruit strongly laterally compressed; often in moist places, grassland<br />
along rivers, among rocks, forest clearings. Pantropical, extending<br />
into some subtropical regions.<br />
Malawi. Nyika National Park, without locality or collecting (Patel 1999).<br />
*Cyclospermum leptophyllum (Pers.) Sprague �<br />
Apium leptophyllum (Pers.) Benth.<br />
Weak, glabrous annual herb; leaves tripinnate into filiform segments; petals<br />
minute, white; fruit almost round, laterally compressed, 1.25–1.75 mm<br />
long, with strongly protuberant, broad, pale primary ribs; a weed of cultivated<br />
and disturbed ground, sometimes in damp places, occasionally even<br />
in very shallow water. Probably Central American in origin, now widespread<br />
as a weed in the tropical regions of the world.<br />
Malawi. Nyika National Park, without locality or collecting details (Patel<br />
1999).<br />
Diplolophium buchananii (Benth. ex Oliv.) Norman �<br />
Physotrichia buchananii Benth. ex Oliv.<br />
A very robust perennial herb 1–2 m high, often glaucous, sometimes sub<br />
shrubby; leaf sheaths well-developed, leaf segments elliptic to subrotund;<br />
involucre and involucel very conspicuous, petals greenish white to creamy<br />
yellow; fruit strongly dorsally compressed, densely bristly pubescent; in<br />
moist places in rocky montane grassland and in Brachystegia woodland.<br />
Malawi and N. Mozambique.<br />
Malawi. Nyika National Park, without locality or collecting details (Patel<br />
1999).<br />
Frommia ceratophylloides H.Wolff �<br />
Perennial herb to 1 m tall; basal leaves forming a rosette above a scaly or<br />
fibrous collar formed by the decaying sheaths of previous years’ leaves;<br />
leaves pinnate, long and slender with each pinna very finely divided into<br />
very thin segments; flowers in much-branched inflorescences, petals yellow;<br />
fruit ovoid, 2–3 mm long, smooth and black, similar to that of<br />
Pimpinella; in thin soils associated with rock outcrops in grassland. A<br />
monotypic genus endemic to N.E. Zambia, N. Malawi, and S.W. Tanzania.<br />
Source. Mill (1979).<br />
Malawi. Nyika Plateau, Robinson 4<strong>49</strong>9 (K, SRGH); Chelinda Bridge rock<br />
outcrop, 2000, Winter 4054 (MAL, SRGH, PRE).<br />
Zambia. Malawi border, Richards 14398 (K); ca 3 km N. of Zambian Rest<br />
House, 2000, Koekemoer s.n. sub Winter 4017 (MAL, SRGH, PRE, UZL).<br />
Heracleum abyssinicum (Boiss.) Norman �<br />
Malabaila kirungae Engl.; Malabaila abyssinica Boiss.; Malabaila rivae<br />
Engl.<br />
Perennial or biennial herb to 1 m tall, stems solid, terete or ridged, often<br />
purplish at base, from woody or somewhat fleshy rootstock; leaves simply<br />
pinnate, to 300 x 70 mm; leaflets ovate, coarsely dentate; flowers white, in<br />
a compound umbel; fruits dorsally compressed and winged; along<br />
streamsides in grassland, 2,000–2,400 m. Widespread in eastern Africa from<br />
Ethiopia to Malawi.<br />
Source. Mill (1979).<br />
Malawi. Nyika National Park, Chelinda Bridge, wetland below forest, 2000,<br />
Winter 4056 (MAL).<br />
Heteromorpha arborescens (Spreng.) Cham. & Schldtl. var.<br />
abyssinica (A.Rich.) H.Wolff �<br />
Heteromorpha abyssinica Hochst. ex A.Rich.; Heteromorpha trifoliata<br />
(Wendl.) Eckl. & Zeyh.<br />
Chichewa: kapoloni, khozyo thengo<br />
Small tree 4(–12) m tall, bark peeling (birch-like) in transverse bands to<br />
expose smooth, bronze underbark between consecutive persistent encircling<br />
leaf scars; leaves shortly and softly pilose usually only on midrib above;<br />
umbel rays ca 25 mm long, flowers greenish yellow; fruit halves hetero-<br />
Plants of the Nyika Plateau<br />
61<br />
APIACEAE
2 mm<br />
2,5 mm<br />
62 Plants of the Nyika Plateau<br />
Cyclospermum leptophyllum<br />
Del. Sandie Burrows<br />
20 mm<br />
Del. Sandie Burrows<br />
20 mm<br />
Heracleum abyssinicum<br />
Diplolophium buchananii<br />
Del. Sandie Burrows<br />
20 mm<br />
Frommia ceratophylloides<br />
Del. Sandie Burrows<br />
20 mm
morphic, one with four ribs (two conspicuous) and the other with five<br />
(three conspicuous); in montane and riverine woodland, forest margins,<br />
and in secondary growth. South Africa (Cape) to Ethiopia and Yemen; also<br />
in Angola.<br />
Source. Brummitt (1973); Winter & Van Wyk (1996).<br />
Malawi. Nyika Plateau, without locality, 1946, Brass 17342 (BM, K, PRE,<br />
SRGH); 3 km W. of Muzengapakweru, 1972, Synge 379 (K); W. of Vitinthiza<br />
Hill, 2000, Salubeni & Mwanyambo 6750 (MAL); Jalawe Viewpoint, 2000,<br />
Thera 3081 (MAL); Juniper Forest, 2000, Burrows & Burrows 6924 (MAL,<br />
PRE, UNIN).<br />
Zambia. Nyika Plateau, Chowo Forest, Phillips 1720 (MO); ca 3 km N. of<br />
Zambian Rest House, 2000, Winter 4016 (PRE, UZL).<br />
Heteromorpha involucrata Conrath<br />
Suffrutex or woody shrub to 1.8 m tall; leaves short-petiolate and trifoliolate,<br />
with broad-based hairs predominantly on nerves below; terminal umbel<br />
often much larger than later umbels, rays up to 100 mm; petals creamcoloured<br />
to greenish yellow; in montane grassland, also open miombo or<br />
Protea woodland, particularly toward the plateau margins, 1,800–2,600 m.<br />
S. Tanzania to northern South Africa.<br />
Malawi. Nyika National Park, near Thazima Gate, 2000, Burrows sub Winter<br />
4051 (MAL, SRGH, PRE, UZL); Vitinthiza Hill slope, 2000, Salubeni &<br />
Mwanyambo 6811 (MAL); Lake Kaulime, 2000, Willis & Luhanga 106 (MAL).<br />
Zambia. Nyika Plateau, N. of Zambian Rest House, ca 1 km E. of Sonkwe<br />
stream, 2000, Winter 4018 (MAL, SRGH, PRE, UZL, LMA, UNIN, DSM).<br />
Heteromorpha kassneri H.Wolff<br />
Suffrutex or woody shrub similar to H. involucrata, to 1.8 m tall; leaves<br />
subsessile with pinnatifid or decurrent pinnae; in open miombo woodland,<br />
1,400–1,800 m. D.R.C., Zambia, and Malawi.<br />
Malawi. Mpora, 2 miles E. of Nyika, Phillips 1295 (MO); 10 km S. of Thazima<br />
Gate, 2000, Winter 4242 (MAL, UZL).<br />
Hydrocotyle mannii Hook.f. �<br />
Low creeping perennial herb, stems rooting at nodes; leaves reniform or<br />
almost round with 5–9 rounded to acute denticulate lobes, pilose on both<br />
surfaces; umbels simple, axillary, rounded capitate, petals greenish to white;<br />
fruit strongly laterally compressed; among grass in wet to somewhat damp<br />
situations at edges of lakes, swamps, and marshes, as well as disturbed<br />
places. Widespread in tropical Africa. H. sibthorpioides Lam. is expected<br />
to occur here, and the two species may have been confused previously.<br />
Malawi. Nyika National Park, without locality or collecting details (Patel<br />
1999).<br />
Lefebvrea abyssinica A.Rich. �<br />
Lefebvrea stuhlmannii Engl.; Peucedanum lefebvria Drude<br />
Slender virgate perennial herb to 3 m tall, characteristically overtopping<br />
surrounding vegetation; stems frequently bowed over with the weight of<br />
the fruiting umbels; leaves mostly confined to near base of stem, 2-pinnate<br />
(ternate-pinnate) with linear leaflets, leaflets with two veins parallel to the<br />
midvein; flowers pale yellowish cream, on long, thin, drooping pedicels;<br />
fruit strongly dorsally compressed as in Peucedanum, but deeply incised at<br />
apex with rounded, curved auricles overtopping the elongated, narrowly<br />
conical stylopodium; in montane grassland and open savanna woodland;<br />
1,000–2,300 m. Zimbabwe to tropical eastern Africa, D.R.C., and Angola.<br />
Source. Mill (1979).<br />
Malawi. Nyika National Park, Wovwe River Valley, 2000, Willis 168 (MAL).<br />
Zambia. Nyika National Park, N. of Zambia Rest House, ca 1 km E. of<br />
Sonkwe stream, 2000, Winter 4020 (MAL).<br />
Lefebvrea grantii (Hiern) Droop<br />
Lefebvrea microcarpa H.Wolff; Lefebvrea angolensis Welw. ex Ficalho;<br />
Lefebvrea welwitschii Engl.; Peucedanum buchananii Baker<br />
Robust biennial (? sometimes perennial) herb, glabrous, 0.6–2(3) m tall;<br />
leaflets with two veins parallel to the midvein in narrow-leaved forms, but<br />
these veins not noticeably parallel in broader-leaved forms; petals greenish<br />
yellow or cream; fruit strongly dorsally compressed as in Peucedanum, but<br />
deeply incised at apex with rounded, curved auricles overtopping the<br />
stylopodium; stylopodium not or only slightly longer than wide; fruits up<br />
to 12 mm long. Widespread in tropical Africa. It may be worth considering<br />
Lefebvrea microcarpa as a subspecies of L. grantii. It is identical in all<br />
respects, but with smaller fruit up to 8 mm long, leaflets very narrowly<br />
ovate to linear and a diffuse inflorescence, and seems to occupy a drier and<br />
warmer habitat at lower altitudes (1,250–1,750 m) in Zambia, Malawi,<br />
and southern Tanzania.<br />
Source. Cannon (1978); Patel (1999).<br />
Malawi. Nyika Plateau, 1896, Whyte s.n. (B, holo.†; BM, photo).<br />
Peucedanum articulatum C.C.Towns. �<br />
Perennial herb 0.5–0.65 m tall; leaves with hollow petioles and pronounced<br />
sheaths, ultimate leaf divisions filiform to narrowly linear; flowers white;<br />
fruit dorsally compressed with lateral wings, stylopodium exceeding the<br />
wings; in moist grassland, presumably temporarily waterlogged. Endemic<br />
to the Nyika Plateau.<br />
Source. Townsend (1987).<br />
Malawi. Nyika National Park, Chosi Hill, 2000, Winter 3988 (MAL, SRGH,<br />
PRE, UZL, LMA); 2000, Kasakula 199 (MAL, LMA, SRGH); 8 km E. of<br />
Chelinda on Kasaramba Road, 2,303 m, Pawek 11203 (K, holo.; UC, MO,<br />
SRGH, MAL, PRE, iso.); above Chelinda Bridge, 2,409 m, Droop 308 (K,<br />
RNG).<br />
Zambia. Nyika Plateau, Chowo Rock, ca 2,195 m, Droop 314 (K, RNG).<br />
Peucedanum claessensii Norman<br />
Erect coarse perennial herb 1–3.5 m tall, stem terete, hollow, to 18 mm<br />
wide at the base; leaves large, up to 0.45–1 m x 0.3–0.75 m; leaves divided<br />
into broad segments, 23–90 x 17–80 mm, margins scabrid with short hairs;<br />
flowers white or creamy, suffused with a green centre; fruit with 2 vittae on<br />
the commissural face; in montane grassland, forest margins, and other scrub<br />
associations, 1,070–2,300 m. S. Tanzania, D.R.C., E. Zambia, E. Zimbabwe,<br />
Malawi, and Mozambique.<br />
Source. Cannon (1978); Patel (1999).<br />
Zambia. Nyika Plateau, Richards 10402 (K, SRGH); N. of Zambian Rest<br />
House, ca 1 km E. of Sonkwe stream, 2000, Winter 4021 (MAL, PRE).<br />
Peucedanum eylesii Norman<br />
Erect perennial herb with stout, gnarled, woody rootstock, 0.9–2 m tall,<br />
stems pithy, angular, and sulcate-striate; lower leaves 190–700 x 80–<br />
180 mm, lower leaf segments broadly linear to narrowly oblong, upper<br />
leaves with the leaflets (especially the terminal) long and narrow, regularly<br />
sharply serrate, somewhat scabrid on midrib below, margins glabrous; flowering<br />
petals yellow to yellow-green; fruit dorsally compressed with lateral<br />
wings, obovate, shallowly truncate or notched at the base, the 2(–3) vittae<br />
on the commissural face superficial, stylopodium exceeding wings; in damp<br />
or seasonally flooded grassland or scrub near streams, 1,500–1,700 m. S.<br />
and W. Tanzania, Zambia, Zimbabwe, Malawi, and Mozambique. Can be<br />
confused with P. nyassicum, as well as Lefebvrea longipedicellata.<br />
Source. Mill (1979).<br />
Malawi. Nyika National Park, Thazima Gate, 2000, Burrows sub Winter<br />
4050 (MAL, UZL, PRE); ca 2 km N.W. of Lake Kaulime, 2000, Winter 4128<br />
(MAL).<br />
Zambia. Nyika Rest House, Richards 10402 (K).<br />
Peucedanum linderi Norman<br />
Peucedanum aberdaricum Chiov.<br />
Erect perennial herb similar to P. eylesii, 0.9–3 m tall, stem fistular, angular,<br />
and sulcate-striate; 8–19 mm wide at base; leaves divided into broad<br />
segments, 12–50 x 7–25 mm, leaflet margins serrate, glabrous; flowers white<br />
or creamy with a narrow greenish or brownish central streak along the<br />
single vitta; fruit broadly elliptical to round, deeply incised at the base, the<br />
2 vittae on the commissural face somewhat concealed by overlying tissue;<br />
in montane forest margins or clearings, tall grassland, scrub, along streams,<br />
and in boggy places, 1,860–3,422 m. Mountains of tropical eastern Africa,<br />
Rwanda, D.R.C., Malawi, Mozambique, and E. Zimbabwe.<br />
Source. Cannon (1978); Mill (1979).<br />
Malawi. Nyika National Park, 5 km S. of Chelinda, De Kruif 1325 (PRE);<br />
Mpopoti Hill, 2000, Salubeni & Mwanyambo 6788 (MAL); Mwanda, between<br />
road and ridge, 2000, Winter 4178 (MAL).<br />
Plants of the Nyika Plateau<br />
63<br />
APIACEAE
20 mm<br />
10 mm<br />
Lefebvrea abyssinica<br />
Del. Sandie Burrows<br />
64 Plants of the Nyika Plateau<br />
Hydrocotyle mannii<br />
Del. Sandie Burrows<br />
3 mm<br />
Peucedanum articulatum<br />
Del. Sandie Burrows<br />
Heteromorpha arborescens<br />
Del. Victoria Goaman<br />
20 mm
Peucedanum sp. nov., aff. P. harmsianum H.Wolff<br />
Glabrous, perennial herb 0.3–0.7 m tall, from rhizomatous rootstock; leaves<br />
less than five-jugate, segments 1–4(6) mm wide; flowering petals yellow, at<br />
least when dry, anthers yellow; fruit elliptic or slightly obovate, ca 7–9 x 3–<br />
5 mm, solitary vittae in valleculae, and two vittae on commissure distinguish<br />
this from P. harmsianum; in wetlands along streams in montane grassland,<br />
ca 2,300 m. Known only from the Nyika Plateau.<br />
Malawi. Nyika National Park, Dembo Bridge area, 2000, Winter 40<strong>49</strong> (MAL,<br />
SRGH, PRE, UZL)); Chelinda Stream, 7,500’, Pawek 7829 (UC, SRGH, PRE),<br />
wetland along stream, Pawek 10295 (UC, SRGH, PRE, K, MO, MAL).<br />
Physotrichia heracleoides H.Wolff �<br />
Perennial with a tough taproot, ca 0.3–0.9 m tall; lobes of lower leaves not<br />
separated into pinnae or, if so, then the pinnae not decurrent along the leafrachis;<br />
involucre and involucel very conspicuous, petals creamy-or greenish<br />
yellow, calyx teeth well-defined, ovary and young fruit not visibly ribbed,<br />
terete; fruit strongly dorsally compressed, with vesicular papillae; in rough,<br />
rather dry grassland, 1,500–2,100 m. From Burundi, S.W. Tanzania, and<br />
southern D.R.C. to Zambia and the Nyika Plateau.<br />
Source. Mill (1979).<br />
Malawi. Nyika National Park, ca 3 km N.W. of Lake Kaulime, 2000, Winter<br />
4123 (MAL); upstream of Wovwe Gorge, 2000, Winter 41<strong>49</strong> (MAL).<br />
Physotrichia muriculata (Hiern) Droop & C.C.Towns.<br />
Peucedanum heracleoides Baker [Type: Nyika Plateau, Whyte 224 (K)].<br />
Perennial herb 0.5–1.5 m tall, with a woody or fleshy taproot, stems ribbed<br />
or grooved, muriculate; leaves pinnatisect, with the pinnae broadly decurrent<br />
along the leaf-rachis, roughly hairy, to 250 x 180 mm; umbels compound,<br />
to 100 mm across; involucre and involucel very conspicuous, flowers<br />
yellowish, sometimes flushed purplish, calyx teeth well-defined; ovary<br />
and young fruit deeply sulcate and ribbed; fruit strongly dorsally compressed,<br />
with a papillate-muriculate surface; in montane grassland or<br />
wooded grassland up to 2,100 m. From Angola, through northern Zambia<br />
and southern D.R.C., to the Nyika Plateau and S. and W. Tanzania.<br />
Source. Cannon (1978).<br />
Malawi. Nyika Plateau, below Rest House, Boughey 1630 (K, SRGH);<br />
Domwe Viewpoint, 2000, Salubeni & Mwanyambo 6799 (MAL);<br />
Mbuzinandi, 2000, Salubeni & Mwanyambo 6816 (MAL); Chisanga Falls<br />
footpath, 2000, Winter 4012 (MAL, PRE, SRGH, UZL, LMA).<br />
Pimpinella buchananii H.Wolff subsp. buchananii var.<br />
buchananii<br />
Pimpinella stolzii H.Wolff; Pimpinella buchananii var. triradiata<br />
Norman; Pimpinella zernyi Gilli<br />
Slender to robust biennial to perennial herb, 0.35–1.5 m high; leaves variably<br />
pinnate, only rarely simple or trilobed; inflorescence much branched<br />
and rebranched, umbel rays (2)4–8(12); involucre and involucels absent,<br />
rarely a single bract present, petals white to creamy, univittate, stylopodia<br />
long-conical; in rocky grassland, miombo woodland. Angola, D.R.C., Zambia,<br />
Tanzania, Malawi, and Mozambique. The altitudinal limit of 1,880 m,<br />
reported in F.T.E.A. is extended to over 2,300 m by Nyika gatherings. At<br />
these altitudes, plants are restricted to rocky habitats, and resemble P.<br />
mulanjensis C.C.Towns., except for the lack of trivittate petals. An extremely<br />
variable species with an immense range in habit and leaf morphology,<br />
even within individual populations. Some of these collections appear<br />
to have an annual habit, but vary in the presence or absence of a tuberous<br />
taproot.<br />
Source. Mill (1979).<br />
Malawi. Nyika National Park, Chosi Hill, W. of summit, 2000, Winter 3990<br />
(MAL, SRGH, PRE, UZL); Fingira Rock, 2000, Willis & Luhanga 246 (MAL);<br />
Mwanda, E. of ridge base, 2000, Winter 41<strong>77</strong> (MAL, SRGH, PRE, LMA).<br />
Pimpinella caffra (Eckl. & Zeyh.) D.Dietr. subsp.<br />
conopodioides C.C.Towns.<br />
Pimpinella stadensis sensu Cannon, non (Eckl. & Zeyh.) D.Dietr., pro<br />
parte, excl. Pimpinella nyassica Norman<br />
Biennial or perhaps perennial herb, 0.22–0.85 m tall from a short and oblong<br />
to tapering, firm, tuber-like root; basal leaves deeply palmately divided<br />
into narrow segments which are pinnate or toothed; stem leaves sim-<br />
ply pinnate or bipinnate with very narrow segments not exceeding 2 mm<br />
wide and generally less; inflorescence simple or with few branches; umbels<br />
of 4–12(15) rays; petals white; fruit glabrous; stylopodial disk broad, concave,<br />
the stylopodia scarcely projecting; on mountain slopes and summits<br />
in grassland, among rocks, and on rocky outcrops and walls of gullies,<br />
2,310–2,760 m. Endemic to Nyika Plateau and S.W. Tanzania.<br />
Source. Patel (1999).<br />
Malawi. Nyika National Park, Nganda Hill, W.N.W. slope, 2,500 m, 2000,<br />
Winter 4065 (MAL, PRE); S. side of beacon, 2,600 m, 2000, Winter 4072<br />
(MAL, SRGH, PRE, LMA).<br />
Pimpinella duridentata C.C.Towns.<br />
Pyrophytic herbaceous perennial 0.4–0.65 m tall, with fibrous remains of<br />
petioles at base of stem; leaves coriaceous, simply pinnate with 2–4 pairs of<br />
sessile, very narrowly ovate pinnae, with firm cartilaginous-tipped teeth,<br />
lower lateral pinnae of at least the stem leaves deflexed; inflorescence moderately<br />
branched, umbel rays and pedicels glabrous, petals yellow; in grassland<br />
and damp, swampy ground, 2,250–2,600(2,800) m. Endemic to Tanzania<br />
(Southern Highlands) and the Nyika Plateau. Cannon’s (1978) note<br />
that ‘Pawek 2081 (UC) compares fairly well’ with the type of P. favifolia<br />
Norman from Lake Mweru seems to have been missed by Townsend (1985,<br />
1989), and this may yet prove to be the correct name for this species, in<br />
which case its range will extend to D.R.C.<br />
Source. Townsend (1985, 1989).<br />
Malawi. Nyika National Park, Chosi Hill, 2000, Winter 3992 (MAL, SRGH,<br />
PRE, UZL, LMA, DSM); Nganda Peak, Pawek 2081 (K, holo.).<br />
Pimpinella huillensis Engl.<br />
Pimpinella welwitschii Engl.; Pimpinella mechowii (Engl.) H.Wolff;<br />
Pimpinella platyphylla Hiern; Pimpinella gossweileri H.Wolff; Pimpinella<br />
robusta Norman; Pimpinella kassneri (H.Wolff) Cannon<br />
Slender to robust biennial or perennial herb, 0.6–2 m tall; basal leaves<br />
coriaceous, simple (entire) or at most ternate; inflorescence much-branched;<br />
bracts (from first flowering branch and upward) with reduced lamina shorter<br />
than firm sheath; flowers white to creamy, pedicels glabrous or with a few<br />
scattered hairs, involucels absent; fruit with dense silky, bristly or vesicular<br />
hairs, not much longer than broad; in Brachystegia–Uapaca woodland and<br />
in shallow boggy soil overlying laterite by streams, up to 1,500 m. Angola,<br />
Zambia, Zimbabwe, Malawi, S. and W. Tanzania, and N. Mozambique.<br />
Source. Cannon (1978); Patel (1999).<br />
Malawi. Nyika National Park, Nchenachena, Pawek 3370 (UC).<br />
Pimpinella ledermannii H.Wolff subsp. engleriana<br />
(H.Wolff) C.C.Towns. �<br />
Pimpinella engleriana H.Wolff; Pimpinella tomentosa sensu Engl., non<br />
C.B.Clarke; Pimpinella africana Hiroe<br />
Biennial or short-lived perennial herb, (0.2)0.5–1.8m tall; basal leaves<br />
coriaceous, entire or at most ternate; inflorescence simple or with few<br />
branches, bracts (from first flowering branch and upward) with reduced<br />
lamina shorter than firm sheath, flowers white, pedicels with dense, short,<br />
spreading hairs, involucels frequently present; fruit densely hairy, conspicuously<br />
longer than broad; in open or wooded (Brachystegia, Uapaca, Protea)<br />
grassland of plateaux and hill slopes, (810)1,400–1,900(2,400) m. Burundi,<br />
D.R.C. (Shaba Province), N. Zambia, S. and W. Tanzania, N. and C. Malawi,<br />
and N. Mozambique.<br />
Source. Brummitt (1973).<br />
Malawi. Nyika National Park, N.W. foot of Nganda, 1972, Synge 379 (K,<br />
MAL, SRGH); 8 km N.W. of Lake Kaulime, camp near North Rukuru River,<br />
2,020 m, Brummitt 10822 (PRE, K).<br />
Pimpinella nyassica Norman<br />
Pimpinella stadensis sensu Cannon, non (Eckl. & Zeyh.) D.Dietr., pro<br />
parte<br />
Slender herb 0.2–0.8 m tall, rhizome with thick side roots; leaves longpetiolate,<br />
lowermost leaf may be undivided, orbicular and deeply dentate,<br />
leaflets with long petiolules; inflorescence simple or with few branches; 2–<br />
3 umbels, rays many, petals creamy white; in wetlands along streams in<br />
grassland. This species is apparently endemic to wetlands on the Nyika<br />
Plateau. Its relationship needs to be clarified with plants from Zimbabwe<br />
Plants of the Nyika Plateau<br />
65<br />
APIACEAE
Physotrichia heracleoides<br />
Del. Sandie Burrows<br />
10 mm<br />
20 mm<br />
Sanicula elata<br />
Del. Sandie Burrows<br />
66 Plants of the Nyika Plateau<br />
Pimpinella ledermannii<br />
Del. Christine Grey-Wilson<br />
20 mm<br />
Steganotaenia araliacea<br />
Del. Sandie Burrows
to South Africa in the P. caffra/stadensis complex that occur in wetland<br />
habitats, for example, P. hydrophila H.Wolff, particularly concerning the<br />
rhizomatous habit and long petiolules.<br />
Source. Cannon (1978).<br />
Malawi. Nyika Mountains, 1932, Sanderson 58 (BM, holo.); Nyika National<br />
Park, wetland at Zovochipolo forest margin, 2000, Winter 4136 (MAL);<br />
tributary E. of Dembo Bridge, 2000, Winter 4041 (MAL, PRE, SRGH).<br />
Sanicula elata D.Don �<br />
Sanicula europaea L. var. elata (D.Don) Boiss.; Sanicula europaea L. var.<br />
partita (Kuntze) Hiroe<br />
Stoloniferous herb up to 0.8 m; leaves long-stalked, rosetted, palmate, 40–<br />
80 mm wide, leaf segments 3–7, margin serrate with a tooth arista or mucro<br />
1 mm long; umbels apparently simple; fruit 3–4 mm long, covered with hooked<br />
bristles; in evergreen forest. Widespread in Old World tropics and subtropics.<br />
Source. Brummitt (1973).<br />
Malawi. Nyika National Park, 8 km E. of Nganda, 1972, Brummitt, Munthali<br />
& Synge 130 (K, MAL, SRGH); Nyika Road, km 56.3, Chapman 181 (BM).<br />
Zambia. Nyika Plateau, upper slopes of Kangampande (Chowo Rocks),<br />
White 2760 (K).<br />
*Daucus carota L.<br />
Daucus abyssinicus auctt. Hochst. ex A.Rich., non C.A.Mey.<br />
carrot<br />
Unlikely, unless as an escape from cultivation in a disturbed situation. May<br />
have been confused with Agrocharis incognita.<br />
Malawi. Nyika National Park, without locality or collecting details (Patel<br />
1999).<br />
Excluded species<br />
Steganotaenia araliacea Hochst. �<br />
Peucedanum fraxinifolium Oliv.<br />
Chichewa: mporoni; Tumbuka: mnyongoloka<br />
Tree to 5 m tall, leafless when in flower; leaves imparipinnate with short,<br />
semi-amplexicaul basal sheath, margin serrate, tooth arista up to 3 mm<br />
long, apex acute to acuminate; umbels spherical, petals yellow; fruit strongly<br />
dorsally compressed, obovate, 8–15 x 5–8 mm; in drier woodland types.<br />
Widespread in the savannas of tropical Africa.<br />
Source. Brummitt (1973).<br />
Malawi. Nyika National Park, Sawi Valley, 1972, Synge 442 (K, MAL).<br />
Torilis arvensis (Huds.) Link �<br />
Erect annual herb to 1 m; leaves pinnately divided, hairy; involucre absent<br />
beneath main compound umbel, flowers white, petals setose-hairy; fruit<br />
with glochidiate spines, both secondary and primary ribs of fruit pilose; at<br />
high altitudes from the Cape (South Africa) to Europe and Asia; an introduced<br />
weed elsewhere in the world.<br />
Source. Cannon (1978).<br />
Malawi. Nyika Plateau, 1896, Whyte 158 (K).<br />
Peucedanum nyassicum H.Wolff<br />
The relationship of this sp. to P. eylesii is not clear, but Townsend (1989)<br />
considered this entity to be more or less confined to Mt. Mulanje, and<br />
quite different in its shorter habit (0.3–1.2 m) and leaf dissection.<br />
Malawi. Nyika National Park, without locality or collecting details (Patel<br />
1999).<br />
Species that are still expected, but have apparently not yet been<br />
collected on the Nyika Plateau<br />
Heteromorpha arborescens (Spreng.) Cham. & Schltdl. var.<br />
montana P.J.D.Winter<br />
The type locality of this variety is from Viphya (Vernal Pool Dome); also<br />
collected at Chikangawa (Viphya) and therefore probably also occurs on<br />
the Nyika although no specimens thus determined have been found there<br />
to date. Expected on the eastern slopes, in steep, open, precipitous habitats.<br />
Source. Winter & Van Wyk (1996).<br />
Lefebvrea longipedicellata Engl.<br />
Peucedanum longipedicellatum (Engl.) Drude; Lefebvrea brevipes<br />
H.Wolff; Lefebvrea naegeleana H.Wolff<br />
Acokanthera laevigata Kupicha �<br />
Shrub or tree to 12 m tall, all parts with milky sap, leaves leathery, flowers<br />
in axillary clusters, white; in mid-stratum of montane forest; in Juniperus<br />
forest, on Nkhonjera Hill, and at Mbuzinandi. N. Malawi and Tanzania.<br />
Source. Flora zambesiaca 7(2): 406 (1985); White et al. (2001).<br />
Malawi. Nyika National Park, in the Juniper Forest, 2,160 m, 1976, Pawek<br />
11818 (K).<br />
Ancylobotrys petersiana (Klotzsch) Pierre �<br />
Landolphia petersiana (Klotzsch) Dyer<br />
Scandent shrub or woody climber with a milky latex; leaves opposite, 40–<br />
110 mm long; flowers white or cream-white, corolla tube 6–14 mm long,<br />
lobes 11–24 mm long; on rocky hillsides in mixed woodland at low altitudes.<br />
South Africa to Tanzania and Kenya; also on Madagascar. Probably<br />
not a constituent of the Plateau flora.<br />
Malawi. Nyika National Park, Kasewerera, 2001, without collecting details<br />
(Patel & Overton 2002).<br />
APOCYNACEAE<br />
(including Asclepiadaceae and Periplocaceae)<br />
Distributed from tropical E. Africa to D.R.C., Zambia, Angola, southern<br />
Malawi, and N. Mozambique. The gap in the known distribution range<br />
seems to reflect missing data or misidentification of Nyika specimens rather<br />
than a true disjunction, as the Nyika National Park includes habitats which<br />
seem suitable for occupation by this species, and it is recorded from as<br />
close as Chisenga (Misuku Hills). It seems as if there may be difficulty<br />
distinguishing it from Peucedanum eylesii when mature fruit is not available,<br />
as both have more or less triplinerved pinnae. The number and length<br />
of pedicels may prove useful to distinguish them. Only a thorough comparison<br />
of specimens of both taxa can confirm this. If these species can be<br />
recognised in the vegetative state, this needs to be reflected in the literature,<br />
at least in the form of a diagnosis.<br />
Asclepias amabilis N.E.Br.<br />
Gomphocarpus amabilis (N.E.Br.) Bullock<br />
Slender erect single-stemmed tuberous herb with white latex; flowers in<br />
few-flowered umbels; in damp peaty soil in montane grassland; 2,250 m.<br />
Malawi, Zambia, and Tanzania.<br />
Malawi. Nyika Plateau, Lake Kaulime, 2,250 m, 1959, Richards 10459 (K).<br />
Asclepias pygmaea N.E.Br.<br />
Trachycalymma cucullatum sensu Bullock, in part<br />
Slender erect tuberous herb with white latex, leaves linear; flowers in white<br />
or purplish umbels; in burnt montane grassland; 2,200–2,500 m. Malawi<br />
and S. Tanzania.<br />
Source. Goyder (2001).<br />
Malawi. Nyika Plateau, Chelinda CDC Camp, 2,200 m, 1958, Robson &<br />
Angus 424 (K).<br />
Asclepias randii S.Moore �<br />
Ascending or erect tuberous herb with white latex, linear leaves and erect<br />
Plants of the Nyika Plateau<br />
67<br />
APOCYNACEAE
20 mm<br />
20 mm<br />
Ancylobotrys petersiana<br />
Del. Sandie Burrows<br />
68 Plants of the Nyika Plateau<br />
4 mm<br />
Torilis arvensis<br />
Del. Sandie Burrows<br />
10 mm<br />
Acokanthera laevigata<br />
Del. Sandie Burrows<br />
20 mm<br />
20 mm<br />
Asclepias randii<br />
Del. Sandie Burrows
20 mm<br />
10 mm<br />
Aspidoglossum breve<br />
Del. Sandie Burrows<br />
Del. Sandie Burrows<br />
20 mm<br />
Ceropegia schliebenii<br />
Carvalhoa campanulata<br />
Del. Sandie Burrows<br />
20 mm<br />
20 mm<br />
Cryptolepis oblongifolia<br />
Del. Sandie Burrows<br />
Plants of the Nyika Plateau<br />
69
APOCYNACEAE<br />
stellate flowers; in burnt montane grassland; 2,200–2,500 m. Also in Angola,<br />
Zambia, Zimbabwe, Mozambique, and southern Tanzania.<br />
Malawi. Nyika Plateau, top of ridge off road to Chelinda, 7,500’, 1967,<br />
Richards 22553 (K).<br />
Asclepias sp. [= Richards 57 (K)]<br />
Ascending or erect tuberous herb with white latex, oblong leaves and erect<br />
stellate flowers; in burnt montane grassland; 200–2,250 m. Also found in<br />
southern Tanzania and the Mbala region of northern Zambia.<br />
Malawi. Nyika Plateau, near Chelinda CDC Camp, 2,200 m, 1958, Robson<br />
& Angus 373 (K).<br />
Asclepias sp. [= Richards 6689 (K)]<br />
Ascending or erect tuberous herb with white latex, linear-lanceolate leaves and<br />
dull green or pinkish flowers; in burnt montane grassland; 2,200–2,500 m.<br />
Also in S. Tanzania on the Mbeya, Kitulo, and Poroto mountains.<br />
Malawi. Nyika Plateau, 30 km along road to Kasaramba Viewpoint, 1966,<br />
Pawek 745 (MAL).<br />
Aspidoglossum angustissimum (K.Schum.) Bullock<br />
Aspidoglossum whytei (N.E.Br.) Bullock; Schizoglossum angustissimum<br />
K.Schum.; Schizoglossum elatum K.Schum.; ?Schizoglossum ledermannii<br />
Schltr.; Schizoglossum whytei N.E.Br.; ?Schizoglossum zernyi Markgr.<br />
Slender erect single-stemmed tuberous herb to 1.5 m with white latex, clusters<br />
of brownish pubescent flowers; in montane grassland with Protea;<br />
2,000 m. Widespread in central and eastern parts of tropical Africa.<br />
Source. Patel (1999); synonymy from Kupicha (1994).<br />
Malawi. Nyika Plateau, N.E. of Nganda, Patel, Ludlow & David 5116 (K,<br />
MAL).<br />
Aspidoglossum breve Kupicha �<br />
Slender erect single-stemmed tuberous herb to 300 mm tall with white latex<br />
and clusters of brownish flowers; in montane grassland; 2,000–2,600 m.<br />
Restricted to the Nyika Plateau of northern Malawi and the Kitulo Plateau<br />
in southern Tanzania.<br />
Source. Kupicha (1994).<br />
Malawi. Nyika Plateau, Nganda Peak, 1992, Goyder, Paton & Tawakali<br />
3557 (BR, K, MAL, PRE).<br />
Carvalhoa campanulata K.Schum. �<br />
Carvalhoa macrophylla K.Schum.<br />
Shrub or small tree to 5 m tall, leaves opposite, thin, with milky latex;<br />
flowers in loose heads on a long stalk from leaf axils, cream to pale yellow<br />
with red streaks in throat; fruit of paired dehiscing carpels; in understorey of<br />
evergreen forest, 800–1,900 m. Mozambique, Malawi, Tanzania, and Kenya.<br />
Malawi. Nyika National Park, without locality or collecting details (Patel<br />
1999).<br />
Ceropegia claviloba Werderm.<br />
Slender twiner to 1.5 m from a globose tuber, latex clear, flowers tubular<br />
with lobes united at tip, small, white with black markings; in evergreen<br />
forest; 1,700–2,300 m. Tanzania, Zambia, Zimbabwe, and Malawi.<br />
Malawi. Nyika Plateau, Chisanga Forest on N. Rukuru River, Brummitt,<br />
Polhill & Banda 16189 (K, MAL).<br />
Ceropegia filipendula K.Schum.<br />
Ceropegia medoensis N.E.Br.; Ceropegia dichroantha K.Schum.<br />
Erect single-stemmed herb with clear latex and fleshy roots, flower tubular<br />
with lobes united at tip, large, yellow-green or brown; in Brachystegia<br />
woodland; 900–1,500 m. Also in Tanzania, Mozambique, Zambia, Zimbabwe,<br />
and Angola.<br />
Malawi. Nyika Plateau N.E. of Nganda, Patel, Ludlow, Scott & Creegan<br />
5087 (K, MAL).<br />
Ceropegia schliebenii Markgr. �<br />
Slender vine with clear latex, tuberous root, flowers purplish with a deep<br />
tube and spidery lobes; shallow soil over rock slabs; 2,000–2,250 m. Southern<br />
Tanzania and northern Malawi.<br />
Malawi. Nyika Plateau, Chelinda Bridge, Brummitt, Polhill & Banda 16148<br />
70 Plants of the Nyika Plateau<br />
(K, MAL); 6.2 km from Lake Kaulime on Zungwara Road, 2000, Burrows &<br />
Burrows 6880 (Buffelskloof Herbarium).<br />
Ceropegia sp. [= Goyder, Paton & Tawakali 3579]<br />
Slender erect herb with clear latex and a globose tuber, flower with narrow<br />
cylindrical tube and united lobes, yellowish with red streaks; in shallow<br />
soil over rock slabs; 1,750 m. Known only from the Nyika.<br />
Malawi. Nyika Plateau, escarpment below Jalawe Viewpoint, 1992, Goyder,<br />
Paton & Tawakali 3579 (K, MAL).<br />
Cryptolepis oblongifolia (Meisn.) Schltr. �<br />
Ectadiopsis nigritana Benth.; Ectadiopsis oblongifolia (Meisn.) Schltr.;<br />
Ectadiopsis welwitschii Baill.; Secamone acutifolia Sond.<br />
Erect subshrub with white latex, reddish stems and tubular white flowers;<br />
in Brachystegia woodland; 1,500–2,000 m. Widespread in eastern and<br />
southern Africa from Sudan to South Africa and Namibia.<br />
Malawi. Nyika National Park, escarpment below Jalawe Viewpoint, 1992,<br />
Goyder, Paton & Tawakali 35<strong>77</strong> (BR, K, MAL, PRE).<br />
Cynanchum praecox Schltr. ex S.Moore<br />
Cynanchum pygmaeum Schltr.<br />
Plant leafless at time of flowering, just an umbel of brownish yellow flowers<br />
at soil level, latex white; in burnt montane grassland; 2,000–2,300 m.<br />
Distribution scattered irregularly over W. and E. tropical Africa.<br />
Source. Liede (1996); Brummitt (1973).<br />
Malawi. Nyika National Park, Nganda, 1972, Synge 370 (K, MAL).<br />
Cynanchum rungweense Bullock �<br />
Vine climbing to 12 m with white latex, leaves heart-shaped, glaucous below<br />
with red veins; on forest fringes; 2,000–2,400 m. Northern Malawi,<br />
Zambia, and southern Tanzania.<br />
Source. White et al. (2001)<br />
Malawi. Nyika Plateau, central plateau, Dowsett-Lemaire 236 (K).<br />
Zambia. Nyika Plateau S. of Zambian Rest House, 1989, Goyder, Pope &<br />
Radcliffe-Smith 3269 (K, MAL, NDO).<br />
Glossostelma carsonii (N.E.Br.) Bullock �<br />
Asclepias carsonii (N.E.Br.) Schltr.; Gomphocarpus chlorojodina<br />
K.Schum.; Schizoglossum carsonii (N.E.Br.) N.E.Br.; Schizoglossum<br />
chlorojodinum (K.Schum.) N.E.Br.; Schizoglossum kassneri S.Moore;<br />
Xysmalobium carsonii N.E.Br.<br />
Erect single-stemmed tuberous herb with white latex, linear fleshy leaves<br />
and large, yellow-green flowers frequently marked with purple; in montane<br />
grassland and Brachystegia woodland; 1,700–2,250 m. From Rwanda to<br />
Angola and Mozambique.<br />
Source. Goyder (1995); Mill (1979).<br />
Malawi. Nyika Plateau, 5 km S.W. of Rest House, 2,150 m, 1958, Robson<br />
& Angus 206B (K).<br />
Zambia. Nyika Plateau, 2 km S.W. of Chowo Forest, 1992, Goyder, Paton<br />
& Tawakali 3538 (K, NDO).<br />
Glossostelma nyikense Goyder<br />
Low single-stemmed tuberous herb with white latex, linear fleshy leaves<br />
and small, yellow flowers; burnt montane grassland; 2,100–2,250 m. Endemic<br />
to the Nyika Plateau—known from just 6 collections.<br />
Source. Goyder (1995).<br />
Malawi. Nyika Plateau, near Chelinda CDC Camp, 2,200 m, 1958, Robson<br />
& Angus 452 (K, LISC, PRE, SRGH).<br />
Zambia. Nyika Plateau, Williamson 1009 (SRGH).<br />
Gomphocarpus praticola (S.Moore) Goyder & Nicholas<br />
Asclepias friesii Schltr.; Asclepias katangensis S.Moore; Asclepias moorei<br />
De Wild.; Asclepias praticola S.Moore<br />
Stout herb with 1–3 stems, with white latex, fleshy glaucous leaves and<br />
large, green and purple flowers; in Brachystegia or Uapaca woodland;<br />
1,700–2,000 m. Northern Malawi and southern Tanzania to Angola.<br />
Source. Goyder & Nicholas (2001).<br />
Malawi. Nyika Plateau, track to Rukuru River Falls, 1,800 m, 1959, Richards<br />
10529 (K).
Cynanchum rungweense<br />
Del. Sandie Burrows<br />
Gomphocarpus swynnertonii<br />
Del. Sandie Burrows<br />
20 mm<br />
20 mm<br />
20 mm<br />
Glossostelma carsonii<br />
Del. Sandie Burrows<br />
20 mm<br />
Landolphia buchananii<br />
Del. Sandie Burrows<br />
Plants of the Nyika Plateau<br />
71
Del. Sandie Burrows<br />
20 mm<br />
Pentarrhinum abyssinicum<br />
subsp. angolense<br />
72 Plants of the Nyika Plateau<br />
20 mm<br />
Margaretta rosea subsp. bidens<br />
Del. Sandie Burrows<br />
Pachycarpus lineolatus<br />
Del. Sandie Burrows<br />
20 mm<br />
20 mm<br />
Periploca linearifolia<br />
Del. Sandie Burrows
Gomphocarpus swynnertonii (S.Moore) Goyder &<br />
Nicholas �<br />
Asclepias nyikana Schltr.; Asclepias swynnertonii S.Moore<br />
Multi-stemmed herb with white latex, glaucous leaves, and small, greenish<br />
yellow flowers; in burnt montane grassland; 2,000–2,250 m. Southern Tanzania,<br />
Malawi, Mozambique, and Zimbabwe westwards to Angola.<br />
Source. Goyder & Nicholas (2001).<br />
Malawi. Nyika Plateau, Lake Kaulime, 1958, Robson & Angus 336 (K).<br />
Zambia. Nyika Plateau, near turn-off to Chelinda Camp, Lees 111 (K).<br />
Landolphia buchananii (Hall.f.) Stapf �<br />
Large liane up to 25 m, often with tendrils, leaves opposite, all parts with<br />
milky sap, flowers white, in terminal clusters; fruit round, to 100 mm diameter;<br />
in montane evergreen and riverine forest. Zimbabwe to D.R.C. and<br />
Tanzania. Fruit much loved by monkeys; stems used for making roofs of houses.<br />
Malawi. Nyika National Park, Kasyaula Forest (Dowsett-Lemaire 1985).<br />
Zambia. Nyika National Park, Manyenjere, Kasoma, and Chowo Forests<br />
(Dowsett-Lemaire 1985).<br />
Margaretta rosea Oliv. subsp. bidens Bullock �<br />
Margaretta cornetii Dewèvre; Margaretta decipiens Schltr.; Margaretta<br />
holstii K.Schum.; Margaretta orbicularis N.E.Br.; Margaretta pulchella<br />
Schltr.; Margaretta verdickii De Wild.<br />
Erect tuberous herb with white latex and flowers with a petaloid corona; in<br />
burnt grassland, usually in peaty soil; ?2,000–2,500 m. Also known from Kenya,<br />
Tanzania, the Shaba region of D.R.C., Zambia, N.W. Zimbabwe, and Nigeria.<br />
Source. Mwanyambo (1996).<br />
Malawi. Nyika Plateau, Mwenembwe, 1903, McClounie 63 (K).<br />
Pachycarpus goetzei (K.Schum.) Bullock<br />
Schizoglossum goetzei K.Schum.; Schizoglossum simulans N.E.Br.<br />
Robust erect tuberous herb with white latex, lanceolate leaves, and nodding<br />
white or cream flowers; in Brachystegia–Uapaca woodland; 1,500–<br />
2,000 m. Also occurs in southern Tanzania and Zambia.<br />
Source. Goyder (1998).<br />
Malawi. Nyika National Park, 15 km N. of M1, 19<strong>77</strong>, Pawek 12420 (K).<br />
Pachycarpus lineolatus (Decne.) Bullock �<br />
Asclepias browniana S.Moore; Asclepias conspicua N.E.Br.; Asclepias<br />
inflexa S.Moore; Asclepias schweinfurthii N.E.Br.; Gomphocarpus<br />
lineolatus Decne.; Pachycarpus mildbraedii Bullock; Pachycarpus<br />
schweinfurthii (N.E.Br.) Bullock<br />
Robust erect tuberous herb with white latex, broad leaves, and nodding<br />
white or pink flowers; in Brachystegia or Uapaca woodland; 1,600–1,800 m.<br />
Widespread in tropical Africa.<br />
Source. Goyder (1998); Mill (1979).<br />
Malawi. Nyika National Park, 12 km N. of M1, 1978, Pawek 13720 (K,<br />
MO, MAL); Juniper Forest Reserve, 2000, Chikuni 522 (MAL).<br />
Pachycarpus spurius (N.E.Br.) Bullock<br />
Schizoglossum debeersianum K.Schum.; ?Schizoglossum dolichoglossum<br />
(K.Schum.) N.E.Br.; Schizoglossum spurium (N.E.Br.) N.E.Br.;<br />
?Xysmalobium dolichoglossum K.Schum.; Xysmalobium spurium N.E.Br.<br />
Robust erect tuberous herb to ca 1 m with white latex, broad leaves, and<br />
nodding cream or pink flowers; in montane grassland or open disturbed<br />
woodland; 1,500–2,000 m. Also recorded from southern Tanzania and the<br />
Shaba plateaux of D.R.C.<br />
Malawi. Nyika Plateau N.E. of Nganda, Patel, Ludlow & Scott 5127 (K).<br />
Pentarrhinum abyssinicum Decne. subsp. angolense<br />
(N.E.Br.) Liede & Nicholas �<br />
Pentarrhinum abyssinicum Decne. var. angolense N.E.Br.<br />
Twiner with heart-shaped leaves; climbing over vegetation at forest margins;<br />
1,700–2,000 m. Scattered distribution from Cameroon, Uganda, and<br />
Kenya to Namibia, Zambia, and Tanzania.<br />
Malawi. Nyika Plateau, Kafwimba Forest, 1969, Pawek 2235 (K).<br />
Periploca linearifolia Quart.-Dill. & A.Rich. �<br />
Periploca linearis Hochst.<br />
Woody twiner with white latex, linear leaves, and bearded white flowers;<br />
scrambling over margins of wet forest; 1,900–2,400 m. Widespread on E.<br />
African mountains from Ethiopia to Malawi.<br />
Source. Brummitt (1973).<br />
Malawi. Nyika National Park, 8 km E. of Nganda, 1972, Brummitt, Munthali<br />
& Synge 147 (K, MAL).<br />
Zambia. Nyika Plateau, margins of Chowo Forest, Dowsett-Lemaire 98 (K).<br />
Raphionacme longituba E.A.Bruce �<br />
Tuberous plant with more or less decumbent stems, latex milky; leaves<br />
opposite; fruit a pair of comparatively large horn-like pods; geophyte in<br />
shallow soils overlying sheetrock in montane grassland. Zimbabwe, Zambia,<br />
Malawi, and Tanzania.<br />
Malawi. Nyika National Park, Mpopoti Peak, 2000, Willis & Luhanga 89<br />
(MAL, PRE).<br />
Rauvolfia caffra Sonder �<br />
Briefly deciduous tree to 30 m tall, leaves clustered towards branch ends,<br />
flowers white or yellowish in stalked clusters, fruit round or lobed, red when<br />
ripe; in rainforest and riverine forest. Widespread throughout tropical Africa.<br />
Malawi. Nyika Plateau, near Kasyaula Forest, without collecting details<br />
(Dowsett-Lemaire 1985: 313).<br />
Rauvolfia mannii Stapf<br />
Rauvolfia rosea K.Schum.<br />
Understorey shrub or small tree to 4 m tall; leaves in whorls of 3–6, flowers<br />
white or yellowish in stalked clusters, fruit of 1 or 2 ovoid or obovoid<br />
carpels, red when ripe; in wet situations in rainforest. From the Viphya<br />
Mountains of Malawi northwards throughout tropical Africa.<br />
Malawi. Nyika National Park, Mwenembwe, Chapman 2019 (White et al.<br />
2001); Eastern escarpment forests (Dowsett-Lemaire 1985: 354).<br />
Riocreuxia chrysochroma (Huber) A.R.Sm.<br />
Ceropegia chrysochroma Huber<br />
Herbaceous vine with clear latex and heart-shaped leaves; flowers orange<br />
within, lobes not united at tips; in scrub or forest patches; 2,000–2,400 m.<br />
Known only from the Nyika Plateau and the adjacent Kitulo and Kipengere<br />
ranges in southern Tanzania.<br />
Source. White et al. (2000).<br />
Malawi. Nyika Plateau, relic forest patch on E. side of Chelinda Valley,<br />
1958, Robson & Fanshawe 643 (K).<br />
Riocreuxia profusa N.E.Br. �<br />
Ceropegia burchellii (K.Schum.) Huber subsp. profusa (N.E.Br.) Huber;<br />
Riocreuxia torulosa Schltr.<br />
Herbaceous vine with clear latex and heart-shaped leaves, flowers pale orange<br />
or yellow, tubular, with slender lobes united at tips; on margins of<br />
montane forest; 1,900–2,400 m. Tanzania, Malawi, Zambia, Zimbabwe,<br />
and Angola.<br />
Source. White et al. (2001: 165)<br />
Zambia. Nyika Plateau, forest patch near Rest House, Dowsett-Lemaire 97<br />
(K).<br />
Secamone alpini Schult. �<br />
Periploca secamone L.; Secamone thunbergii E.Mey.<br />
Slender woody twiner with white latex, leathery leaves, and minute velvety<br />
white flowers; on forest margins; 2,000–2,400 m. Widespread over E. African<br />
mountains and coastal forests in South Africa.<br />
Source. Goyder (1992).<br />
Malawi. Nyika National Park, without locality or collector (Mill 1979).<br />
Zambia. Nyika Plateau, margins of Chowo Forest, Dowsett-Lemaire 99 (K).<br />
Stathmostelma fornicatum (N.E.Br.) Bullock subsp.<br />
fornicatum �<br />
Asclepias fornicata N.E.Br.<br />
Erect single-stemmed tuberous herb with white latex, linear leaves, and<br />
green or yellow flowers; in seasonally waterlogged grassland; 2,400 m.<br />
Found at lower altitude elsewhere in Malawi, Zambia, Zimbabwe, Botswana,<br />
and Angola.<br />
Plants of the Nyika Plateau<br />
73<br />
APOCYNACEAE
Raphionacme longituba<br />
Del. Sandie Burrows<br />
Riocreuxia profusa<br />
Del. Sandie Burrows<br />
74 Plants of the Nyika Plateau<br />
20 mm<br />
20 mm<br />
55 mm<br />
Secamone alpini<br />
Del. Sandie Burrows<br />
Rauvolfia caffra<br />
Del. Sandie Burrows<br />
20 mm<br />
20 mm<br />
20 mm
Source. Goyder (1988)<br />
Malawi. Nyika Plateau, 1903, McClounie 81 (K).<br />
Tabernaemontana stapfiana Britten �<br />
Tabernaemontana johnstonii (Stapf) Pichon; Conopharyngia stapfiana<br />
(Britten) Stapf<br />
Tree 5–25 m tall, leaves opposite, 120–400 x 30–140 mm, narrowly elliptic,<br />
glabrous; flowers white, large, sweetly scented; fruit in pairs or single,<br />
large, 100–200 mm across, green with yellowish dots; in montane forest,<br />
700–2,500 m. Zimbabwe and Mozambique to D.R.C. and Kenya.<br />
Source. Mill (1979).<br />
Malawi. Nyika National Park, eastern escarpment forests (Dowsett-Lemaire<br />
1985: 354).<br />
Tacazzea conferta N.E.Br. �<br />
Tacazzea floribunda K.Schum.; Tacazzea galactagoga Bullock<br />
Robust woody twiner with white latex; scrambling over forest margins;<br />
2,000–2,200 m. Widely distributed in tropical Africa from Ethiopia to<br />
Malawi.<br />
Malawi. Nyika Plateau, Zovochipolo Forest patches, Dowsett-Lemaire 251 (K).<br />
Zambia. Nyika Plateau, Chowo Forest, 1976, Pawek 11783 (K, MAL, MO).<br />
Trachycalymma buchwaldii (Schltr. & K.Schum.) Goyder �<br />
Asclepias affinis De Wild.; Asclepias buchwaldii (Schltr. & K.Schum.) De<br />
Wild.; Gomphocarpus buchwaldii Schltr. & K.Schum.<br />
Short, erect, single-stemmed, tuberous herb with white latex and nodding<br />
purple flowers; in montane grassland or Brachystegia woodland, generally<br />
on steep rocky hillsides; 900–2,400 m. Also found in Tanzania, Burundi,<br />
and D.R.C.<br />
Source. Goyder (2001).<br />
Malawi. Nyika National Park, escarpment below Jalawe Viewpoint, 1992,<br />
Goyder, Paton & Tawakali 3581 (K, MAL).<br />
Trachycalymma cristatum (Decne.) Bullock<br />
Asclepias cristata S.Moore; Asclepias palustris (K.Schum.) Schltr.;<br />
Gomphocarpus cristatus Decne.; Gomphocarpus palustris K.Schum.<br />
Short, erect, single-stemmed, tuberous herb with white latex and nodding<br />
white or pink flowers; montane grassland or open woodland; 2,000–<br />
2,600 m. Widespread in tropical sub-Saharan Africa from Nigeria to An-<br />
Landolphia kirkii Dyer<br />
Straggling shrub or liane to 18 m tall. This record is doubtful. Dowsett-<br />
Lemaire (1985) did not find it in her comprehensive survey of the Nyika<br />
forests. Both White et al. (2001) and Flora zambesiaca (1985) do not record<br />
this species from N. Malawi. Since Patel quoted no specimen and we have<br />
seen no specimens, it is excluded.<br />
Malawi. Nyika National Park, without locality or collecting details (Patel<br />
1999).<br />
Gomphocarpus glaucophyllus Schltr.<br />
This is almost certainly G. praticola.<br />
Malawi. Nyika National Park, without locality or collecting details (Mill 1979).<br />
Stathmostelma welwitschii Britt. & Rendle<br />
Asclepias welwitschii (Britt. & Rendle) Britt. & Rendle; Stathmostelma<br />
chironoides K.Schum. ex De Wild. & T.Durand; Stathmostelma<br />
laurentianum Dewevre<br />
Ilex mitis (L.) Radlk. �<br />
Cape holly; Chichewa: katenji; Yao: katenji, mulisanyundo<br />
Evergreen tree to 20 m, leaves simple, margins almost entire with a few<br />
minute teeth near the apex, fruit (female trees only) a small red berry; along<br />
streams and rivers in evergreen forest and riverine vegetation. From South<br />
Excluded species<br />
AQUIFOLIACEAE<br />
gola and Zimbabwe.<br />
Source. Goyder (2001).<br />
Malawi. Nyika Plateau, Circular Drive, 19<strong>77</strong>, Pawek 12271 (K, MO).<br />
Zambia. Nyika Plateau, Rest House, 2,100m, 1959, Richards 10410 (K).<br />
Trachycalymma foliosum (K.Schum.) Goyder<br />
Asclepias foliosa (K.Schum.) Hiern; Asclepias lepida S.Moore; Asclepias<br />
minuta A.Chev.; Asclepias modesta N.E.Br.; Gomphocarpus foliosus<br />
K.Schum; Trachycalymma pulchellum sensu Bullock, in part<br />
Erect, single-stemmed, tuberous herb with white latex and nodding white<br />
flowers; Brachystegia woodland; 1,100–2,000 m. Widespread in savanna<br />
regions of S. tropical Africa and W. Africa.<br />
Source. Goyder (2001).<br />
Malawi. Nyika, 1 mile in Nyika Road [from Rumphi?], 4,900’, 1969, Pawek<br />
32<strong>49</strong> (K).<br />
Tylophora anomala N.E.Br.<br />
Slender woody twiner with watery latex, large leathery ovate-lanceolate<br />
leaves and zig-zag inflorescences with green flowers in clusters; forest margins;<br />
1,500–2,000 m. Kenya, Tanzania, Zambia, Malawi, and Mozambique.<br />
Malawi. Nyika Plateau, Kasyaula Forest, Dowsett-Lemaire 243 (K).<br />
Tylophora erubescens (Liede & Meve) Liede ��<br />
Tylophoropsis erubescens Liede & Meve<br />
Slender twiner with clear latex, leaves ovate ca 10 mm long; flowers delicate<br />
maroon; in wet montane forest; 1,800–2,300 m. Also known from S.<br />
Tanzania.<br />
Malawi. Nyika National Park, Zovochipolo Forest, 2000, Winter & Burrows<br />
4143 (K, PRE).<br />
Xysmalobium undulatum (L.) Ait.f. �<br />
Asclepias undulata L.; Xysmalobium ensifolium Burch. ex Sc.-Elliot;<br />
Xysmalobium angolense Sc.-Elliot<br />
Stout erect herb with white latex, triangular leaves, and hairy, reddish purple<br />
flowers, fruit covered in hairy prickles; in damp hollows in montane<br />
grassland; 2,000–2,250 m. Widespread in sub-Saharan Africa.<br />
Source. Brummitt (1973).<br />
Malawi. Nyika National Park, W. foot of Nganda, 1972, Brummitt & Synge<br />
35 (K).<br />
This species does not occur in the area. The record may come from a<br />
misidentified specimen of Asclepias randii at K, MO, and SRGH labelled<br />
S. cf. welwitschii, or is S. fornicatum.<br />
Malawi. Nyika National Park, without locality or collecting details (Mill<br />
1979).<br />
Stenostelma sp.<br />
This is not Stenostelma; probably refers to Asclepias sp. (= Richards 57(K)).<br />
Malawi. Nyika National Park, without locality or collecting details (Patel<br />
1999).<br />
Trachycalymma pulchellum (Decne.) Bullock<br />
This is almost certainly T. foliosum.<br />
Malawi. Nyika National Park, without locality or collecting details (Mill<br />
1979).<br />
Africa (Western Cape) to Ethiopia and Guinea.<br />
Malawi. Nyika National Park, Lake Kaulime, 1958, Robson 2<strong>77</strong> (BM, K,<br />
LISC, PRE, SRGH); Dembo Bridge, 2000, Thera 3009 (MAL); near Chelinda<br />
Compound, 1959, Adlard 312 (FHO, SRGH).<br />
Plants of the Nyika Plateau<br />
75<br />
APOCYNACEAE
Stathmostelma fornicatum subsp. fornicatum<br />
Del. Sandie Burrows<br />
Tacazzea conferta<br />
Del. Sandie Burrows<br />
76 Plants of the Nyika Plateau<br />
20 mm<br />
20 mm<br />
20 mm<br />
20 mm<br />
Tabernaemontana stapfiana<br />
Del. Sandie Burrows<br />
Trachycalymma buchwaldii<br />
Del. Sandie Burrows<br />
10 mm<br />
50 mm
20 mm<br />
Ilex mitis<br />
Del. Sandie Burrows<br />
Tylophora erubescens<br />
Del. Sandie Burrows<br />
20 mm<br />
Xysmalobium undulatum<br />
Del. Sandie Burrows<br />
Cussonia arborea<br />
Del. Sandie Burrows<br />
Cussonia spicata<br />
20 mm<br />
200 mm<br />
200 mm<br />
Plants of the Nyika Plateau<br />
<strong>77</strong>