Type genera are listed alphabetically. For each nominal type genus, the name, author(s), year of publication, page number, current status and placement, type species, and typification, as well as information regarding the family-group name based on it, are given.

Type species are important nomenclaturally since they provide the objective standard of reference for the application of the genus-group name they represent (ICZN 1999a, Article 61.1). Species that can be fixed as type are nominal species originally included in the newly established genus or subgenus that are cited by an available name (excluded are nominal species that were doubtfully or conditionally included in the new taxon) (see ICZN 1999a, Article 67.2). Type species can either be fixed in the original publication (ICZN 1999a, Article 68), in a subsequent publication (ICZN 1999a, Article 69) or, when nomenclatural problems are resolved by the International Commission on Zoological Nomenclature (henceforth “Commission”), by a decision under its plenary power (ICZN 1999a, Article 81). In cases where several nominal species were originally included in a new genus or subgenus established before 1931, but none were fixed as the type species, the first author who subsequently designated one of the originally included nominal species as the type species fixed the type species and no later designation is valid (ICZN 1999a, Article 69.1). When an earlier type species fixation is discovered, it is to be accepted or a Case submitted to the Commission for a ruling if the overlooked fixation is considered to cause instability or confusion (ICZN 1999a, Article 70.2). When a type species is fixed, it is generally assumed that the identification of that species is correct (ICZN 1999a, Article 70.1); however, where evidence suggests that the type species was misidentified at the time it was fixed, we have selected the taxonomic species actually involved as type species accompanied with a reference to Article 70.3.2 (ICZN 1999a). We follow previous authors (e.g., Alonso-Zarazaga and Lyal 2009; Bousquet et al. 2015, 2018; Bouchard et al. 2021) and consider type-species designations in R. Lucas (1920) as potentially valid when a single species is listed under a particular genus-group name. Based on Recommendation 67B (ICZN 1999a), the name of each type species is cited by its original binomen and, when this is currently considered invalid, its valid synonym is given in parentheses (preceded by “=”). Also note that the type species (and fixation) for a new replacement name is the same as for the previously established name it replaces (ICZN 1999a, Article 67.8). Fossil taxa are indicated by the symbol “†”.

While the correct identification of a type genus is assumed when an author establishes a new nominal family-group taxon (ICZN 1999a, Articles 65.1), evidence suggests that in some cases the type genus was misidentified (i.e., interpreted in a sense other than that defined by its type species). The cases where these family-group names are not currently used as valid (summarized in Table 1) are likely to cause instability and confusion and therefore need to be referred to the Commission for a ruling (ICZN 1999a, Articles 65.2.1). For this reason, names in Table 1 are not included elsewhere in this work (for additional information see appendix 2 and appendix 3 in Bouchard et al. 2011, Bouchard and Bousquet 2020a). We follow Evenhuis (2023: 3) in using the Nomenclator Zoologicus (e.g., Neave 1939, 1940) as a potential source of First Reviser actions when more than one spelling was used to establish a new genus-group name.

The classification presented in Appendix 1 (down to the rank of subfamily) is based on the studies of Cai et al. (2022) unless stated otherwise. The synoptic classification in Bouchard et al. (2011) was used as the starting point for the classification of beetles below the rank of subfamily. While we tried to use the most up-to-date information to generate the classification of all beetles presented in Appendix 1, we realize that errors may remain and that there are often different points of view in the community regarding how taxa should be classified. We nevertheless hope that the information thus presented will adequately assist the retrieval of relevant information for the readers. We included type genera of all available family-group names known to us, up to the date of publication of this article; however, we also included the type genus of a small number of family-group names that were published after 2011 in an electronic-only work that failed to meet the criteria of availability as required in amendments of the International Code of Zoological Nomenclature (ICZN 2012f; henceforth “Code”). In these cases, the family-group name and its associated genus are included to highlight the generally unknown nomenclatural problems. The stem we include under each family-group name and use in Appendix 1 is the stem that we recommend to be used in order to provide greatest stability of the Coleoptera classification (see Bouchard et al. 2011, Bouchard and Bousquet 2020a). The spelling of an incorrectly formed stem is sometimes maintained following recommendations in Article 29 (ICZN 1999a).

It should be noted that a disproportionally large number of new family-group names have been proposed in recent years in the superfamily CURCULIONOIDEA Latreille, 1802 by a small number of individuals (e.g., Legalov, Poinar). While these taxa are included here for completeness, some were evidently proposed without adequate comparative morphological analysis (e.g., Oberprieler and Zimmerman 2020: 556–558; Wanat 2021: 54) and their validity remains to be tested in a phylogenetic framework. The recent proliferation of new family-group names, combined with the challenging diversity of the group, has unfortunately resulted in an unsatisfactory weevil classification that still requires much attention before a stable and globally accepted system can be agreed upon.

COLEOCATINIIDAE Ponomarenko & Prokin (2016): as pointed out by Bouchard and Bousquet (2020a: 70), this family-group name is unavailable since it was not based on an available genus-group name (ICZN 1999a, Article 11.7.1.1).

DONACOCIADAE Gistel (1856): this name, which was listed as available by Bouchard et al. (2011: 509), is based on Donacocia used by Gistel (1856: 377), which we consider to be an incorrect subsequent spelling of Donacia Fabricius, 1775. Consequently, the family-group name must be corrected (ICZN 1999a, Article 35.4.1) to DONACIADAE (see entry below under the type genus Donacia Fabricius, 1775).

PHILHYDRIDA J.E. LeConte (1849): the informal name Philhydrida, which was not based on an available type genus, was first coined by W.S. MacLeay (1825: 6). J.E. LeConte (1849: 27) was the first to include the genus-group name Philydrus Solier, 1834 under the name Philhydrida; however, we believe that he did not propose a new family-group name but rather used W.S. MacLeay’s informal name. Therefore, we do not treat Philhydrida, as used by J.E. LeConte (1849), as an available family- group name as previously done by Bouchard et al. (2011: 155).

RHYTIDORHINIDES Wollaston (1865): this name was listed as available by Bouchard et al. (2011: 603); however, it is based on Rhytidorhinus Agassiz, 1846, an unjustified emendation of Rhythirrinus Schönherr, 1823, and as such must be corrected (ICZN 1999a, Article 35.4.1) to RHYTHIRRINIDAE (see entry below under the type genus Rhythirrinus Schönherr, 1823). Rhytidorhinus Wollaston, 1864, incorrectly given as the type genus of RHYTIDORHINIDES Wollaston by Bouchard et al. (2011: 603), is an unavailable genus-group name not homonymous with Rhytidorhinus Agassiz, 1846.

Initials for the following authors were used in the text to avoid confusion: A. Abdullah, M. Abdullah, D.M. Anderson, R.S. Anderson, W.H. Anderson, M. Bai, X.L. Bai, F. Balfour-Browne, J. Balfour- Browne, F. Bates, H.W. Bates, J.R. Bell, K.L. Bell, R.T. Bell, T. Bell, A. Bezděk, J. Bezděk, W.F.H. Blandford, W.T. Blandford, A.E. Brown, F.M. Brown, C.H. Chen, S.H. Chen, Z.-Y. Chen, A. Costa, C. Costa, A.G. Desmarest, E. Desmarest, E.S. Dillon, L.S. Dillon, H.B. Douglas, J.W. Douglas, F. Dupuis, P. Dupuis, M.C. Ferreira, V.S. Ferreira, H. Franz, N.M. Franz, C. Girard, M. Girard, G.R. Gray, J.E. Gray, G.H. Horn, W. Horn, D. Huang, F. Huang, H. Huang, A. Janssens, E. Janssens, A.J. Johnson, C. Johnson, P.J. Johnson, W. Kirby, W.F. Kirby, C. Koch, C.L. Koch, J.D.W. Koch, J.E. LeConte, J.L. LeConte, J. Li, S. Li, X.-Y. Li, Y.-D. Li, L.Y. Liu, M. Liu, P.H. Lucas, R. Lucas, W.J. MacLeay, W.S. MacLeay, G.A.K. Marshall, T.A. Marshall, A. Matthews, A.[H.] Matthews, E.G. Matthews, G.S. Medvedev, L.N. Medvedev, S.I .Medvedev, B. Meyer, H. Meyer, J. Meyer, K.B. Miller, L. Miller, N.C.E. Miller, A. Milne-Edwards, H. Milne-Edwards, B.P. Moore, F. Moore, C. Müller, J. Müller, O.F. Müller, P.W.J. Müller, E. Olivier, G.-A. Olivier, C. d’Orbigny, H. d’Orbigny, M.F. Pacheco, T.L. Pacheco, J. Parker, J.B. Parker, H. Pang, X.-F. Pang, G. Pelletier, J. Pelletier, F. Philippi, R.A. Philippi, J. Redtenbacher, L. Redtenbacher, W. Redtenbacher, H.G.L. Reichenbach, L. Reichenbach, D. Ren, G.-D. Ren, L. Ren, P.O. Richter, R. Richter, C.V. Riley, N.D. Riley, C.R. Sahlberg, J.R. Sahlberg, R.F. Sahlberg, C. Schaufuss, L.W. Schaufuss, A. Schmidt, F.J. Schmidt, J. Schmidt, W.L.E. Schmidt, D.H. Schneider, J.G. Schneider, E.A. Schwarz, O. Schwarz, H. Scott, J. Scott, C. Shin, S. Shin, A.B.T. Smith, A.D. Smith, F. Smith, A. Solari, F. Solari, G. Strohmeyer, H. Strohmeyer, D.B. Thomas, M.C. Thomas, C.G. Thomson, J. Thomson, J.A. Thomson, B.D. Valentine, J.M. Valentine, A. Villa, G.B. Villa, B. Wang, W.I. Wang, C.O. Waterhouse, G.R. Waterhouse, J.B. Waterhouse, G. Wheeler, Q.D. Wheeler, W.M. Wheeler, A. White, B.E. White, R.E. White, A. Winkler, J.R. Winkler, Q. Yang, X.-K. Yang, F.N. Young, R.M. Young, H. Zhang, P. Zhang, W. Zhang, A. Zimmermann, C. Zimmermann. We follow Bouchard et al. (2021: 8) and use the surname “Fischer” for G. Fischer’s works up to the end of 1832 and “Fischer von Waldheim” starting from 1833, which is the year he became a member of the Russian nobility. Similarly, we use “Semenov” for works published by this author up to the end of 1906 and “Semenov-Tjan-Shansky” for works published after 1906 in order to account for the honorary title “Tjan-Schansky” added by the Emperor of All Russia Nikolay II to the surname of A.P. Semenov’s father (and all his family members) in 1906 (e.g., see Chigray et al. 2022: 4).

Because of the importance of the Principle of Priority in zoological nomenclature (ICZN 1999a, Article 23), we have tried to establish the date of publication of works listed herein as accurately as possible. In the References section, publication dates are listed after each reference in square brackets “ []” and preceded by DP (i.e., Date of Publication). These dates are either specific dates of publication (e.g., extracted from the works themselves or the journal wrappers, title pages, footers) or are the earliest known publication dates (e.g., extracted from recording journals or accounts of society meetings). In the latter case, the date is preceded by the word “by.”

For the Annali del Museo Civico di Storia Naturale di Genova, we originally compiled the dates on which the signatures were printed; however, following Poggi (2010: 48–49, 79–228), the publication date is that printed on the reprint, which is the date on which the compiled signatures were sent to the authors for distribution [although Poggi occasionally gave a reprint date earlier than the date of the last signature in the paper, in which case we cite the signature date]. The volume numbers listed in the articles published in this journal are for the entire sequence of volumes, ignoring the “Series” numbers.

Although we found it difficult to determine the date of publication for many Russian journals and books, we have often cited the date on which the article was “approved to print”, that is the date posted by the censor (“podpisano k pechati”), usually inside the front or back wrapper, or on the endleaf. With few exceptions, this date indicates the earliest possible date of publication, and the works were apparently usually published shortly thereafter (see Technical Translations 15 (12): Doklady Biological Sciences 154–156: B-5). All Julian dates for Russian publications have been converted to Gregorian dates.

Sources of information from recording journals (for further historical information see Bousquet 2016: 24–36), specific works and date stamps from various institutions are mentioned below.

Allg Anz Deutsch: Allgemeiner Anzeiger der Deutschen (Gotha, Germany).

Allgem Bibl: Allgemeine Bibliographie. Monatliches Verzeichniss der wichtigern neuen Erscheinungen der Deutschen und Ausländischen Literatur (Leipzig, Germany).

Allg Bibl Deutsch: Allgemeine Bibliographie für Deutschland (Leipzig, Germany).

Allg Lit Ztg: Allgemeine Literatur-Zeitung (Jena/Halle, Germany).

Amer Nat: The American Naturalist (Philadelphia, U.S.A.).

Ann Soc Ent Fr: Annales de la Société Entomologique de France (Paris, France).

Arch Naturg: Archiv für Naturgeschichte (Berlin, Germany).

Athenaeum: The Athenaeum. Journal of Literature, Science, and the Fine Arts (London, U.K.).

Berl Ent Zeit: Berliner Entomologische Zeitschrift (Berlin, Germany).

Bibl Fr: Bibliographie de la France (Paris, France).

Bulletin du Nord: Bulletin du Nord, journal scientifique et littéraire, contenant: des mémoires et notices, des analyses et extraits d’ouvrages nouveaux; des variétés et mélanges, des annonces bibliographiques, etc., etc. (Moscow, Russia).

Bull Acad R Sci Belg: Bulletins de l’Académie Royale des Sciences, des Lettres et des Beaux-Arts de Belgique (Bruxelles, Belgium).

Bull Acad Sci St-Pétersb: Bulletin de l’Académie Impériale des Sciences de St.-Péterbourg (St. Petersburg, Russia).

Bull Soc Imp Nat Mosc: Bulletin de la Société Impériale des Naturalistes de Moscou (Moscow, Russia).

Bull Soc Ent Fr: Bulletin de la Société Entomologique de France (Paris, France).

Bull Soc Hist Nat Toulouse: Bulletin de la Société d’Histoire Naturelle de Toulouse (Toulouse, France).

Bull Soc Imp Nat Mosc: Bulletin de la Société Impériale des Naturalistes de Moscou (Moscow, Russia).

Bull Zool Nomencl: The Bulletin of Zoological Nomenclature (London, U.K.).

Compt Rend Acad Sci Fr: Comptes Rendus (hebdomadaire) des séances de l’Académie des Sciences de France (Paris, France).

Compt Rend Soc Ent Belg: Comptes Rendus des Séances de la Société Entomologique de Belgique (Bruxelles, Belgium).

Ent Bericht: Entomologische Berichten (Amsterdam, Netherlands).

Ent Nachr: Entomologische Nachrichten (Berlin, Germany).

Ent News: Entomological News (Academy of Natural Sciences, Philadelphia, U.S.A.).

Ent Ztg Stettin: Entomologische Zeitung (Stettin, Germany [now in Poland]).

Feuil J Lib: Feuilleton du Journal de la Librairie Nationale de la France (issued with “Bibliographie de la France”; Paris, France).

Intell Allg Lit Ztg: Intelligenzblatt der Allgemeinen Literatur-Zeitung (Jena/Halle, Germany; published in conjunction with “Allgemeine Literatur-Zeitung”).

Leipz Repert: Leipziger Repertorium der Deutschen und Ausländischen Literatur (Leipzig, Germany).

Lit Centr Deutsch: Literarisches Centralblatt für Deutschland (Leipzig, Germany).

Lit Ztg: Literarische Zeitung (Berlin, Germany).

Lond Lit Gaz: The London Literary Gazette, and Journal of the Belles Lettres, Arts, Sciences (London, U.K.).

Merc Fr: Mercure de France (Paris, France).

Nat Nov: Naturae Novitates (Berlin, Germany).

Nature: Nature, a Weekly Illustrated Journal of Science (London, U.K. and New York, U.S.A.).

Öfvers Kongl Vetens Forh: Öfversigt af Kongliga Vetenskaps-Akademiens Förhandlingar (Stockholm, Sweden).

Pan-Pacific Ent: The Pan-Pacific Entomologist (San Francisco, U.S.A.).

Proc Acad Nat Sci Philad: Proceedings of the Academy of Natural Sciences, Philadelphia (Philadelphia, U.S.A.).

Proc Amer Phil Soc: Proceedings of the American Philosophical Society, held at Philadelphia, for promoting useful knowledge (Philadelphia, U.S.A.).

Proc Bost Soc Nat Hist: Proceedings of the Boston Society of Natural History (Boston, U.S.A.).

Proc Ent Acad Nat Sci: Proceedings of the Monthly Meetings of the Entomological Section of the Academy of Natural Sciences of Philadelphia (Philadelphia, U.S.A.).

Proc Ent Soc Lond: The Proceedings of the Entomological Society of London [and Journal of Proceedings] (London, U.K.).

Proc Ent Soc NSW: The Proceedings of the Entomological Society of New South Wales (Sydney, Australia; published with the Transactions).

Proc Linn Soc NSW: The Proceedings of the Linnean Society of New South Wales (Sydney, Australia).

Proc-verb Acad Sci Fr: Procès-verbaux des Séances de l’Académie des Sciences de France (Paris, France).

Rev Mag Zool: Revue et Magasin de Zoologie (Paris, France).

Rev Zool: Revue Zoologique par la Société Cuvierienne, Journal Mensuel (Paris, France).

Sitz-Ber Zool-Bot Ges Wien: Sitzungsberichte der Kaiserlich-Königlichen Zoologisch-Botanischen Gesellschaft in Wien (Vienna, Austria).

Soc Ent Fr, Bull Bibliogr: Société Entomologique de France, Bulletin Bibliographique (Paris, France).

Soc Ent Fr, Bull Ent: Société Entomologique de France, Bulletin Entomologique (Paris, France).

Tijdschr Ent: Tijdschrift voor Entomologie (Nederlandsche Entomologische Vereniging, Leiden, Netherlands).

Trans Ent Soc Lond: Transactions of the Entomological Society of London (London, U.K.).

Trans Proc New Zealand Inst: Transactions and Proceedings of the New Zealand Institute (Wellington, New Zealand).

Verh Ver Naturw Unt Hamb: Verhandlungen des Vereins für Naturwissenschaftliche Unterhaltung zu Hamburg (Hamburg, Germany).

Verz Bücher Landkarten: Verzeichniss der Bücher, Landkarten &c (Leipzig, Germany).

Verz neuer Bücher: Verzeichniss neuer Bücher (Leipzig, Germany).

Zool Rec: Zoological Record (London, U.K.).

Zoologist: The Zoologist: A Popular Miscellany of Natural History (London, U.K.).

The following works included important information for dating the references: Alonso-Zarazaga and Lyal (1999), Bellamy (2003), Bousquet (1993, 2012, 2016, 2017a, 2018), Bousquet and Bouchard (2017), Bousquet and Davies (2021), F.M. Brown (1964a, 1964b), Cowan (1969), Dawson (1958), Duncan (1937), Evenhuis (1994, 1997a-c, 2002, 2003a-b, 2008, 2012, 2014, 2015a, 2015b, 2019), Fox (1913), Griffin (1932), von Hayek (1989), Horn and Schenkling (1928–1929), Houston and Weir (1992), ICZN (1958b, 1999a), Kappel (1896), Kerzhner (1984), Lefèvre (1885, 1895), Merkl et al. (2008), Nagel and Schmidlin (2014), Oshanin (1910), Peavot (1913), Poggi (2010), Raphael (1970), Sclater (1893), Sherborn (1922, 1934, 1937), Silfverberg (1971), G. Wheeler (1912), and Zerche (1987).

Biblioth Nat Fr: Bibliothèque Nationale de France (Paris, France)

BMNH: The Natural History Museum (London, U.K.)

CNC: Canadian National Collection of Insects, Arachnids and Nematodes (Ottawa, Canada)

CUL: Cornell University Library (Ithaca, New York, U.S.A.)

Smithsonian: The Smithsonian Institution (Washington, D.C., U.S.A.)

We established the availability of electronic works published after 2011 that contained new names or new nomenclatural acts following the recommendations outlined in amendments to the Code (ICZN 2012f). Based on these requirements, electronic works are available when: 1) simultaneously obtainable copies of the work with fixed content and layout are widely accessible (Art. 8.1), and are not disclaimed (Arts 8.2, 8.3); 2) the work itself contains a date of publication (Art. 8.5.2); 3) the work was registered in ZooBank before it was published and contains evidence of such registration (Art. 8.5.3); 4) the ZooBank record indicates an intended online archive (Art. 8.5.3.1); and 5) the ZooBank record includes an ISSN/ISBN (Art. 8.5.3.2).

In many cases, works published after 2011 were issued electronically first and then in print later. According to Article 21.9 (ICZN 2012f) the “name or nomenclatural act published in a work issued in both print and electronic editions takes its date of publication from the edition that first fulfilled the criteria of publication of Article 8 and is not excluded by Article 9.” As recommended by Krell (2015), we use the final page numbers, since they are better finding aids, when pagination in an available electronic work differs from the pagination of the same work subsequently assigned to a particular journal issue. We report the first date of publication we could find for cited works published after 2011 that do not contain new names or new nomenclatural acts.

The following works, which were issued and distributed electronically only, are unavailable because they failed to meet one or more of the criteria of availability published in ICZN (2012f): Legalov (2017a, 2018c, but see Legalov 2023c), Kirejtshuk (2020), Rosa et al. (2020a, but see Rosa et al. 2020b), Szawaryn et al. (2020) and Motyka et al. (2023a). The new names and nomenclatural acts included in those works are therefore nomenclaturally unavailable. In order to promote stability (since the majority of the zoological community may not be aware of these facts), such unavailable names and acts are included in this work but preceded by an asterisk “*”. The Commission is currently exploring potential solutions to address the growing problem with new names and acts proposed only in electronic works since 2012 and that are not Code compliant (T. Pape, pers. comm. to P.B., 2022).

Bousquet (2016: 87) gave the dates “1871–1875” for this publication since he could find no evidence that the plates were issued in 1870, in spite of the fact that they all have the inscription “Tome X. 2e Partie (1870)”. Bonvouloir submitted the 42 plates and a preliminary version of the text to the Société Entomologique de France on 23.III.1870 (Bull Soc Ent Fr 1870: xvii), with the hope of having them published in Tome X. On 13.IV.1870 the Commission de Publication agreed to print the plates “au nombre ordinaire de notre tirage des Annales” (Bull Soc Ent Fr 1870: xxv–xxvi), but this apparently did not happen, since no collection of the plates was ever recorded with the Annales for 1870 or anywhere else, in spite of the fact that they were to be distributed “gratuitement à nos membres ainsi qu’aux diverses sociétés savantes”. Bonvouloir was the ‘Archiviste-Bibliothécaire’ for the Société at the time, so one would expect that if they were available in 1870, they would have been entered into the Bibliothèque at least in 1871 (the first notice of copies being available, as cahiers). The dates recorded in the Bulletin Bibliographique (see References) were very specific as to the number of pages and plates in each cahier, as well as to the “dates de parution”. Likewise, the text was not published “en deux fascicules: l’un publié dans le courant de 1870 et l’autre au commencement de 1871” as originally agreed. In fact, Bonvouloir made additions and changes to the text long after the original was submitted, citing (p. 765) a paper published at the end of 1871. Evidently he sent Gemminger a preliminary list of the new names and plates in 1869 because they were listed (as “Monogr. Bonvouloir. Ann. Fr. 1870” or “Bonv. Mon.”) with their provenances, in the Catalogue by Gemminger and Harold (1869b), but the page numbers were never cited, only the prospective plate and figure numbers for those illustrated, and some of the figures from this list were changed in the actual publication (e.g., Microrhagus pavidus “Bonv. Mon. t. 27 f. 2” was replaced with M. inconsultus, pl. 27 fig. 2; Nematodes buqueti (pl. 33 fig. 1) was not listed; Cephalodendron mozambicanum “Bonv. Mon. t. 36. f. 1” was changed to C. indigaceum var., etc.). Similarly, Gemminger included names published by Baudi the following year (1870), without page numbers. In the Nécrologie for Bonvouloir (Bull Soc Ent Fr 1914: 407) it was noted that the Monographie was issued “en 4 fascicules publiés de 1871 à 1875”, with no mention of a separate edition of the plates. We follow Muona (1987: 79–92) who, in his review of the generic names of Eucnemidae, did not include the plates as a separate publication, but cited species and genera from the plates as issued in the cahiers as 1871–1875.

Abacetus Dejean, 1828: 195. Current status: valid genus in CARABIDAE: HARPALINAE: ABACETINI: ABACETINA.

Type species: Abacetus gagates Dejean, 1828, by monotypy.

Family-group name: ABACÉTIDES Chaudoir, 1873. Stem: Abacet-.

Abacomorphus Chaudoir, 1878a: 14. Current status: valid genus in CARABIDAE: HARPALINAE: PTEROSTICHINI: ABACOMORPHINA.

Type species: Abax caledonicus Montrouzier, 1860, by monotypy.

Family-group name: ABACOMORPHINI Tschitschérine, 1902. Stem: Abacomorph-.

Abax Bonelli, 1810: Tab. Syn. Current status: valid genus in CARABIDAE: HARPALINAE: PTEROSTICHINI: PTEROSTICHINA.

Type species: Carabus striola Fabricius, 1792 (= Carabus parallelepipedus Piller & Mitterpacher, 1783), by subsequent designation (Westwood 1838c: 4). Note: no species were originally included in this genus by Bonelli (1810); the first ones were the two species listed by Panzer (1813: 68).

Family-group name: ABAXINI Schuler, 1970. Stem: Abac-.

Ablabera Dejean, 1833b: 159. Current status: valid genus in SCARABAEIDAE: SERICINAE: ABLABERINI.

Type species: Melolontha splendida Fabricius, 1781, by monotypy.

Family-group name: ABLABERITAE Blanchard, 1850. Stem: Ablaber-.

Abraeus Leach, 1817: 76. Current status: valid genus in HISTERIDAE: ABRAEINAE: ABRAEINI.

Type species: Hister globosus P.W.J. Müller, 1803 (= Hister perpusillus Marsham, 1802), by subsequent designation (Westwood 1838c: 22).

Family-group name: ABREIDAE W.S. MacLeay, 1819. Stem: Abrae-.

†Abrocar M. Liu & Ren, 2006: 62. Current status: valid genus in CARIDAE: †ABROCARINI. Note: Lower Cretaceous (China).

Type species: †Abrocar brachyorhinos M. Liu & Ren, 2006, by original designation.

Family-group name: †ABROCARINA Legalov, 2009. Stem: Abrocar-.

Abroteles Casey, 1890a: 190. Current status: valid genus in STAPHYLINIDAE: ALEOCHARINAE: COROTOCINI: ABROTELINA.

Type species: Abroteles beaumonti Casey, 1890, by monotypy.

Family-group name: ABROTELINA Seevers, 1957. Stem: Abrotel-.

Abryna Newman, 1842a: 289. Current status: valid genus in CERAMBYCIDAE: LAMIINAE: PTEROPLIINI.

Type species: Abryna coenosa Newman, 1842, by subsequent designation (J. Thomson 1864: 44).

Family-group name: ABRYNITAE J. Thomson, 1864. Stem: Abryn-.

Acalles Schönherr, 1825: 586. Current status: valid genus in CURCULIONIDAE: CRYPTORHYNCHINAE: CRYPTORHYNCHINI: TYLODINA.

Type species: Curculio camelus Fabricius, 1792, by original designation.

Family-group name: ACALLINAE Joy, 1932. Stem: Acall-.

Acallopistus Schönherr, 1826: 249. Current status: valid genus in CURCULIONIDAE: CURCULIONINAE: NERTHOPINI.

Type species: Acallopistus vellicosus Schönherr, 1826, by original designation.

Family-group name: ACALLOPISTIDES Lacordaire, 1865. Stem: Acallopist-.

Acalyptomerus Crowson, 1979: 615. Current status: valid genus in CLAMBIDAE: ACALYPTOMERINAE.

Type species: Acalyptomerus asiaticus Crowson, 1979, by original designation.

Family-group name: ACALYPTOMERINAE Crowson, 1979. Stem: Acalyptomer-.

Acalyptus Schönherr, 1833: 20. Current status: valid genus in CURCULIONIDAE: CURCULIONINAE: ACALYPTINI.

Type species: Curculio carpini Fabricius, 1792, by original designation.

Family-group name: ACALYPTINA C.G. Thomson, 1859. Stem: Acalypt-. Note: Gapon et al. (2019: 175) submitted an application to the Commission to remove the homonymy between this family-group name and ACALYPTINI Blatchley, 1926 (type genus Acalypta Westwood, 1840) in Hemiptera; Gapon et al. (2019: 175) recommended a change to the family-group name in Hemiptera, leaving ACALYPTINI C.G. Thomson, 1859 in Coleoptera unchanged.

Acamptus J.L. LeConte in J.L. LeConte and G.H. Horn, 1876: 238. Current status: valid genus in CURCULIONIDAE: COSSONINAE: ACAMPTINI.

Type species: Acamptus rigidus J.L. LeConte, 1876, by monotypy.

Family-group name: ACAMPTI J.L. LeConte, 1876. Stem: Acampt-.

Acangassu Galileo & Martins, 2001: 97. Current status: valid genus in CERAMBYCIDAE: CERAMBYCINAE: ACANGASSUINI.

Type species: Acangassu diminuta Galileo & Martins, 2001, by original designation.

Family-group name: ACANGASSUINI Galileo & Martins, 2001. Stem: Acangassu-.

Acanthinodera Hope, 1834a: 106. Current status: junior synonym of Amallopodes Lequien, 1833 in CERAMBYCIDAE: PRIONINAE: MACRODONTIINI.

Type species: Prionus cumingii Hope, 1834, by original designation.

Family-group name: ACANTHINODERITAE J. Thomson, 1864. Stem: Acanthinoder-.

Acanthinomerus Boheman, 1859: 141. Current status: valid genus in CURCULIONIDAE: COSSONINAE: ACANTHINOMERINI.

Type species: Acanthinomerus armatus Boheman, 1859, by monotypy.

Family-group name: ACANTHINOMERINI Voss, 1972. Stem: Acanthinomer-.

Acanthocerus W.S. MacLeay, 1819: 136. Current status: invalid senior synonym of Ceratocanthus A. White, 1842 in HYBOSORIDAE: CERATOCANTHINAE: CERATOCANTHINI. Note: junior homonym of Acanthocerus Palisot de Beauvois, 1818 [Hemiptera].

Type species: Acanthocerus aeneus W.S. MacLeay, 1819, by subsequent designation (Duponchel 1839: 30).

Family-group name: ACANTHOCERIDAE Streubel, 1846. Stem: Acanthocer-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

Acanthocinus Dejean, 1821: 106. Current status: valid genus in CERAMBYCIDAE: LAMIINAE: ACANTHOCININI.

Type species: Cerambyx aedilis Linnaeus, 1758, by subsequent designation (Blanchard in Audouin et al. 1841: pl. 67).

Family-group name: ACANTHOCINITES Blanchard, 1845. Stem: Acanthocin-.

Acanthocnemus Perris, 1866: 187. Current status: valid genus in ACANTHOCNEMIDAE.

Type species: Acanthocnemus ciliatus Perris, 1866 (= Dasytes nigricans Hope, 1843), by monotypy.

Family-group name: ACANTHOCNEMINAE Crowson, 1964. Stem: Acanthocnem-.

Acanthoderes Audinet-Serville, 1835a: 29. Current status: valid genus in CERAMBYCIDAE: LAMIINAE: ACANTHODERINI.

Type species: Cerambyx daviesii Swederus, 1787, by subsequent designation (Duponchel 1839: 32).

Family-group name: ACANTHODERITAE J. Thomson, 1860. Stem: Acanthoder-.

Acanthoglossa Kraatz, 1859: 144. Current status: valid genus in STAPHYLINIDAE: PAEDERINAE: LATHROBIINI: MEDONINA.

Type species: Acanthoglossa hirta Kraatz, 1859, by subsequent designation (R. Lucas 1920: 68).

Family-group name: ACANTHOGLOSSI Coiffait, 1982. Stem: Acanthogloss-.

Acanthomerosternoplon Tippmann, 1956: 10. Current status: junior synonym of Omosarotes Pascoe, 1860 in CERAMBYCIDAE: LAMIINAE: CYRTININI.

Type species: Acanthomerosternoplon paradoxum Tippmann, 1956, by original designation.

Family-group name: ACANTHOMEROSTERNOPLONINI Tippmann, 1956. Stem: Acanthomerosternopl-.

Acanthophorus Audinet-Serville, 1832: 152. Current status: valid genus in CERAMBYCIDAE: PRIONINAE: ACANTHOPHORINI.

Type species: Prionus serraticornis G.-A. Olivier, 1795, by subsequent designation (Duponchel 1839: 35).

Family-group name: ACANTHOPHORITAE J. Thomson, 1864. Stem: Acanthophor-.

Acanthopygus Montrouzier, 1861a: 869. Current status: valid genus in ANTHRIBIDAE: ANTHRIBINAE: ACANTHOPYGINI.

Type species: Acanthopygus metallicus Montrouzier, 1861, by subsequent designation (Kuschel 1998: 346).

Family-group name: ACANTHOPYGINI Legalov, 2023c: 22. Stem: Acanthopyg-. Note: ACANTHOPYGINI Legalov (2018c: 787) was published in a work that is nomenclaturally unavailable (ICZN 2012f, Article 8.5.3).

Acanthoscelides Schilsky, 1905: 95. Current status: valid genus in CHRYSOMELIDAE: BRUCHINAE: BRUCHINI: ACANTHOSCELIDINA. Note: two original spellings were used: Acanthocelides (pp. 41f, 41g, 41C) and Acanthoscelides (pp. 95, 96, 97, 98); Neave (1939: 11) acted as First Reviser (ICZN 1999a, Article 24.2.3) and selected Acanthocelides; an application to the Commission is necessary to retain the spelling Acanthoscelides, which is in prevailing usage.

Type species: Bruchus irresectus Fåhraeus, 1839 (= Bruchus obtectus Say, 1831), by subsequent designation (Bridwell 1929: 42).

Family-group name: ACANTHOSCELIDINI Bridwell, 1946. Stem: Acanthoscelid-.

Acanthoscelis Dejean, 1825: 402. Current status: valid genus in CARABIDAE: SCARITINAE: SCARITINI: ACANTHOSCELINA.

Type species: Scarites ruficornis Fabricius, 1801, by monotypy.

Family-group name: ACANTHOSCELINA Csiki, 1927. Stem: Acanthoscel-.

Acanthotrachelus Schönherr, 1842b: 236. Current status: valid genus in CURCULIONIDAE: ENTIMINAE: CYPHICERITAE: CYPHICERINI: ACANTHOTRACHELINA.

Type species: Acanthotrachelus ventricosus Boheman, 1842, by original designation.

Family-group name: ACANTHOTRACHELINI G.A.K. Marshall, 1944. Stem: Acanthotrachel-.

Acanthovalgus Kraatz, 1895: 444. Current status: valid genus in SCARABAEIDAE: CETONIINAE: VALGINI: VALGINA.

Type species: Acanthovalgus marquardi Kraatz, 1895, by monotypy.

Family-group name: ACANTHOVALGINAE Kolbe, 1904. Stem: Acanthovalg-.

Acentrus Duponchel, 1839: 61. Current status: valid genus in CURCULIONIDAE: CURCULIONINAE: ACENTRINI. Note: we consider that Acentrus, as used by Schönherr (1845: 57), is a subsequent use of Acentrus Duponchel, 1839 and not a new distinct taxon; the entry in the Dictionnaire universel d’histoire naturelle where the name Acentrus is made available is signed “(D.)” which corresponds to Duponchel (as indicated in the “Liste des auteurs par ordre de matières”), not to Desmarest as reported by some authors (e.g., Alonso-Zarazaga et al. 2017: 180).

Type species: Cryptorhynchus histrio Schönherr, 1837, by original designation.

Family-group name: ACENTRINA Seidlitz, 1890. Stem: Acentr-.

Aceraius Kaup, 1868a: 26. Current status: valid genus in PASSALIDAE: PASSALINAE: MACROLININI.

Type species: Passalus grandis Burmeister, 1847, by subsequent designation (Gravely 1918: 89).

Family-group name: ACERAIAE Kaup, 1871. Stem: Acerai-.

Acetalius Sharp, 1883b: 322. Current status: valid genus in STAPHYLINIDAE: PSELAPHINAE: EUPLECTITAE: TRICHONYCHINI: PANAPHANTINA.

Type species: Acetalius dubius Sharp, 1883, by monotypy.

Family-group name: ACETALIINI Jeannel, 1958. Stem: Acetali-.

Achaenops Suffrian, 1857: 61, 234. Current status: valid genus in CHRYSOMELIDAE: CRYPTOCEPHALINAE: CRYPTOCEPHALINI: ACHAENOPINA.

Type species: Achaenops dorsalis Suffrian, 1857, by monotypy.

Family-group name: ACHAENOPITES Chapuis, 1874. Stem: Achaenop-.

Acherusia Laporte & Gory, 1838: 1. Current status: invalid senior synonym of Mimicoclytrina Bellamy, 2003 in BUPRESTIDAE: POLYCESTINAE: TYNDARIDINI: MIMICOCLYTRININA. Note: junior homonym of Acherusia O.G. Costa, 1834 [Crustacea].

Type species: Acherusia childreni Laporte & Gory, 1838, by monotypy.

Family-group name: ACHERUSINI Cobos, 1955. Stem: Acherusi-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

Achloa Erichson, 1840a: 41. Current status: valid genus in SCARABAEIDAE: MELOLONTHINAE: TANYPROCTINI: TANYPROCTINA.

Type species: Achloa helvola Erichson, 1840, by present designation.

Family-group name: ACHLOIDAE Burmeister, 1855. Stem: Achlo-.

Achryson Audinet-Serville, 1834a: 572. Current status: valid genus in CERAMBYCIDAE: CERAMBYCINAE: ACHRYSONINI.

Type species: Stenocorus circumflexus Fabricius, 1787, by monotypy.

Family-group name: ACHRYSONIDES Lacordaire, 1868. Stem: Achryson-.

Acicnemis Fairmaire, 1849: 511. Current status: valid genus in CURCULIONIDAE: MOLYTINAE: TRACHODINI.

Type species: Acicnemis variegata Fairmaire, 1849, by subsequent designation (Fairmaire 1881: 299).

Family-group name: ACICNÉMIDES Lacordaire, 1865. Stem: Acicnemid-.

Acidocerus Klug, 1855: 649. Current status: valid genus in HYDROPHILIDAE: ACIDOCERINAE.

Type species: Acidocerus aphodioides Klug, 1855, by monotypy.

Family-group name: ACIDOCERINI Zaitzev, 1908. Stem: Acidocer-.

Acilius Leach, 1817: 69, 72. Current status: valid genus in DYTISCIDAE: DYTISCINAE: ACILIINI. Note: the senior homonym Acilius Rafinesque, 1815, an unnecessary new replacement name for Symethus Rafinesque, 1814 in Crustacea, was suppressed for both the Principles of Homonymy and Priority and Acilius Leach, 1817 placed on the Official List of Generic Names in Zoology (ICZN 1961e).

Type species: Dytiscus sulcatus Linnaeus, 1758, by monotypy (see ICZN 1961e). Note: name placed on the Official List of Specific Names in Zoology (ICZN 1961e).

Family-group name: ACILIIDES C.G. Thomson, 1867. Stem: Acili-.

Acinopus Dejean, 1821: 13. Current status: valid genus in CARABIDAE: HARPALINAE: HARPALINI: HARPALINA.

Type species: here fixed (under ICZN 1999a, Article 70.3.2) as Scarites picipes G.-A. Olivier, 1795, misidentified as Carabus megacephalus Rossi, 1794 (sensu Illiger, 1802) in the original designation by monotypy in Dejean (1821: 13).

Family-group name: ACINOPIDAE Laporte, 1834. Stem: Acinopod-.

Aclopus Erichson, 1835: 259. Current status: valid genus in SCARABAEIDAE: ACLOPINAE: ACLOPINI.

Type species: Aclopus vittatus Erichson, 1835, by subsequent designation (R. Lucas 1920: 70).

Family-group name: ACLOPITAE Blanchard, 1850. Stem: Aclop-.

Acmaeodera Eschscholtz, 1829b: 9. Current status: valid genus in BUPRESTIDAE: POLYCESTINAE: ACMAEODERINI: ACMAEODERINA. Note: name placed on the Official List of Generic Names in Zoology (ICZN 2005b).

Type species: Buprestis cylindrica Fabricius, 1775, by subsequent designation (Volkovitsh 1979: 339; see ICZN 2005b). Note: name placed on the Official List of Specific Names in Zoology (ICZN 2005b).

Family-group name: ACMAEODERINI Kerremans, 1893. Stem: Acmaeoder-.

Acmaeoderoides Van Dyke, 1942: 108. Current status: valid genus in BUPRESTIDAE: POLYCESTINAE: ACMAEODERINI: ACMAEODEROIDINA.

Type species: Acmaeodera insignis G.H. Horn, 1894, by original designation.

Family-group name: ACMAEODEROIDINI Cobos, 1955. Stem: Acmaeoderoid-.

Acmaeops J.L. LeConte, 1850a: 316, 321. Current status: valid genus in CERAMBYCIDAE: LEPTURINAE: ACMAEOPINI.

Type species: Leptura proteus W. Kirby, 1837, by subsequent designation (Casey 1913: 219).

Family-group name: ACMAEOPSINI Della Beffa, 1915. Stem: Acmaeop-.

Acmocera Dejean, 1835: 345. Current status: valid genus in CERAMBYCIDAE: LAMIINAE: ACMOCERINI.

Type species: Lamia compressa Fabricius, 1801, by monotypy.

Family-group name: ACMOCERITAE J. Thomson, 1864. Stem: Acmocer-.

Acoma Casey, 1890a: 165. Current status: valid genus in SCARABAEIDAE: MELOLONTHINAE: ACOMINI.

Type species: Acoma brunnea Casey, 1890, by monotypy.

Family-group name: ACOMINI Evans & A.B.T. Smith, 2020: 54. Stem: Acom-.

Acorynus Schönherr, 1833: 3, 123. Current status: valid genus in ANTHRIBIDAE: ANTHRIBINAE: TROPIDERINI.

Type species: Acorynus sulcirostris Boheman, 1833, by original designation.

Family-group name: ACORYNIDES Lacordaire, 1865. Stem: Acoryn-.

Acratus Lacordaire, 1865: 463. Current status: valid genus in BRENTIDAE: BRENTINAE: BRENTINI: ITHYSTENINA.

Type species: Brentus suturalis Lund, 1802, by original designation.

Family-group name: ACRATINI Alonso-Zarazaga, Lyal, Bartolozzi & Sforzi, 1999. Stem: Acrat-.

Acridocephala Chevrolat, 1855: 287. Current status: valid genus in CERAMBYCIDAE: LAMIINAE: ACRIDOCEPHALINI.

Type species: Acridocephala bistriata Chevrolat, 1855, by monotypy.

Family-group name: ACRIDOCEPHALIDI E.S. Dillon & L.S. Dillon, 1959. Stem: Acridocephal-.

Acritomorphus Wenzel, 1944: 56. Current status: valid genus in HISTERIDAE: ABRAEINAE: ACRITOMORPHINI.

Type species: Acritomorphus praecursor Wenzel, 1944, by original designation.

Family-group name: ACRITOMORPHINI Wenzel, 1944. Stem: Acritomorph-.

Acritorrhynchites Voss, 1941: 119, 212. Current status: valid genus in ATTELABIDAE: RHYNCHITINAE: RHYNCHITITAE: EUGNAMPTINI.

Type species: Rhynchites addendus Sharp, 1889, by monotypy.

Family-group name: ACRITORRHYNCHITINA Legalov, 2007. Stem: Acritorrhynchit-.

Acritosoma Pakaluk & Ślipiński, 1995: 330. Current status: valid genus in ANAMORPHIDAE.

Type species: Acritosoma elongatum Pakaluk & Ślipiński, 1995, by original designation.

Family-group name: ACRITOSOMATINAE Pakaluk & Ślipiński, 1995. Stem: Acritosomat-.

Acritus J.L. LeConte, 1853c: 288. Current status: valid genus in HISTERIDAE: ABRAEINAE: ACRITINI.

Type species: Hister nigricornis Hoffmann, 1803, by subsequent designation (C.G. Thomson 1859: 75).

Family-group name: ACRITINI Wenzel, 1944. Stem: Acrit-.

Acrobolbia Ohaus, 1912b: 316. Current status: valid genus in SCARABAEIDAE: DYNASTINAE: CYCLOCEPHALINI.

Type species: Acrobolbia macrophylla Ohaus, 1912, by monotypy.

Family-group name: ACROBOLBIINA Ohaus, 1918. Stem: Acrobolbi-.

Acrocinus Illiger, 1806: 247. Current status: valid genus in CERAMBYCIDAE: LAMIINAE: ACROCININI.

Type species: Cerambyx longimanus Linnaeus, 1758, by monotypy.

Family-group name: ACROCININAE Swainson, 1840. Stem: Acrocin-.

Acrognathus Erichson, 1839a: 607. Current status: invalid senior synonym of Manda Blackwelder, 1952 in STAPHYLINIDAE: OXYTELINAE: PLANEUSTOMINI. Note: junior homonym of Acrognathus Agassiz, 1836 [Pisces].

Type species: Omalium mandibulare Gyllenhal, 1827, by subsequent designation (Duponchel 1841b: 57).

Family-group name: ACROGNATHINI Reitter, 1909. Stem: Acrognath-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

Acroleptus Bourgeois, 1886: lxx. Current status: valid genus in LYCIDAE: LYCINAE: CALOPTERINI: ACROLEPTINA.

Type species: Acroleptus chevrolati Bourgeois, 1886, by monotypy.

Family-group name: ACROLEPTINA Bocáková, 2005. Stem: Acrolept-.

Acropis Burmeister, 1840a: [Heft 5]. Current status: valid genus in ZOPHERIDAE: COLYDIINAE: ACROPINI.

Type species: Acropis tuberculiferus Burmeister, 1840, by monotypy.

Family-group name: ACROPINAE Sharp, 1894. Stem: Acrop-.

Acropteron Perty, 1832: 64. Current status: junior synonym of Acropteryx Gistel, 1831 in TENEBRIONIDAE: TENEBRIONINAE: ACROPTERONINI.

Type species: Acropteron rufipes Perty, 1832 (= Acropteryx rufipes Gistel, 1831), by subsequent designation (Hope 1841a: 133).

Family-group name: ACROPTERONINI Doyen, 1989. Stem: Acropteron-.

Acrotona C.G. Thomson, 1859: 38. Current status: valid genus in STAPHYLINIDAE: ALEOCHARINAE: ATHETINI: ATHETINA. Note: name placed on the Official List of Generic Names in Zoology (ICZN 1961b).

Type species: Aleochara aterrima Gravenhorst, 1802, by original designation (see ICZN 1961b). Note: name placed on the Official List of Specific Names in Zoology (ICZN 1961b).

Family-group name: ACROTONAE Seevers, 1978. Stem: Acroton-.

Acrotrichis Motschulsky, 1848: 569, table. Current status: valid genus in PTILIIDAE: PTILIINAE: ACROTRICHINI.

Type species: Latridius fascicularis Herbst, 1793, by subsequent designation (C. Johnson 1969: 229).

Family-group name: ACROTRICHINI Reitter, 1909. Stem: Acrotrich-.

Actenodes Dejean, 1833a: 80. Current status: valid genus in BUPRESTIDAE: BUPRESTINAE: ACTENODINI. Note: name placed on the Official List of Generic Names in Zoology (ICZN 2002b).

Type species: Buprestis nobilis Linnaeus, 1758, by monotypy (see ICZN 2002b). Note: name placed on the Official List of Specific Names in Zoology (ICZN 2002b).

Family-group name: ACTENODEIDAE Gistel, 1848. Stem: Actenod-.

Actenonyx A. White, 1846: 2. Current status: valid genus in CARABIDAE: HARPALINAE: ODACANTHINI: ACTENONYCINA.

Type species: Actenonyx bembidioides A. White, 1846, by monotypy.

Family-group name: ACTENONYCINAE H.W. Bates, 1871. Stem: Actenonyc-.

Actidium A. Matthews, 1868: 12. Current status: valid genus in PTILIIDAE: PTILIINAE: PTILIINI.

Type species: Ptilium coarctatum Haliday, 1855, by original designation.

Family-group name: ACTIDIINI Portevin, 1929. Stem: Actidi-.

Actinophorus Creutzer, 1799: 79. Current status: junior synonym of Scarabaeus Linnaeus, 1758 in SCARABAEIDAE: SCARABAEINAE: SCARABAEINI.

Type species: Scarabaeus sacer Linnaeus, 1758, by subsequent designation (Ádám 2003: 130).

Family-group name: ACTINOPHORINI Ádám, 2003. Stem: Actinophor-.

Actizeta Pascoe, 1875: 214. Current status: valid genus in TENEBRIONIDAE: PIMELIINAE: CNEMEPLATIINI: ACTIZETINA.

Type species: Actizeta ammobioides Pascoe, 1875 (= Actizeta albata Pascoe, 1875), by original designation.

Family-group name: ACTIZETINA Watt, 1992. Stem: Actizet-.

Actobaena Gistel, 1856: 266, 355. Current status: junior synonym of Hyphydrus Illiger, 1802 in DYTISCIDAE: HYDROPORINAE: HYPHYDRINI.

Type species: Hyphydrus fabricii Cristofori & Jan, 1832 (= Dytiscus ovatus Linnaeus, 1761), by monotypy.

Family-group name: ACTOBAENIDAE Gistel, 1856. Stem: Actobaen-.

Actocharis Fauvel, 1871: 19. Current status: junior synonym of Actocharis Sharp, 1870 in STAPHYLINIDAE: ALEOCHARINAE: ACTOCHARINI. Note: junior homonym of Actocharis Sharp, 1870 [Coleoptera: STAPHYLINIDAE].

Type species: Actocharis marina Fauvel, 1871 (= Actocharis readingii Sharp, 1870), by monotypy.

Family-group name: ACTOCHARI Bernhauer & Schubert, 1911. Stem: Actochar-. Note: this family-group name was based on Actocharis “Fauvel, 1869” by Bernhauer and Schubert (1911: 91), assuming that the name had been issued prior to Actocharis Sharp, 1870; however, Fauvel’s name was not issued until 1871 (see Rye 1872: 30, 31) and is currently treated as a junior homonym of Sharp’s name; therefore, ACTOCHARINI Bernhauer & Schubert, 1911 is permanently invalid (ICZN 1999a, Article 39); an application to the Commission is necessary to change the type genus of this family-group name to Actocharis Sharp, 1870 or a new family-group name should be proposed for this tribe.

Aculagnathus Oke, 1932: 22. Current status: valid genus in CERYLONIDAE: CERYLONINAE.

Type species: Aculagnathus mirabilis Oke, 1932, by monotypy.

Family-group name: ACULAGNATHIDAE Oke, 1932. Stem: Aculagnath-.

Acupalpus Latreille, 1829a: 391. Current status: valid genus in CARABIDAE: HARPALINAE: HARPALINI: STENOLOPHINA.

Type species: Carabus meridianus Linnaeus, 1761, by subsequent designation (Westwood 1840b: 498).

Family-group name: ACUPALPINI Tschitschérine, 1900. Stem: Acupalp-.

Acylophorus Nordmann, 1837: 5, 127. Current status: valid genus in STAPHYLINIDAE: STAPHYLININAE: STAPHYLININI: ACYLOPHORINA.

Type species: Acylophorus ahrensii Nordmann, 1837 (= Staphylinus glaberrimus Herbst, 1784), by subsequent designation (Blackwelder 1943: 466).

Family-group name: ACYLOPHORINI Outerelo & Gamarra, 1986. Stem: Acylophor-.

Adalia Mulsant, 1846: [2 (Addenda et errata)]. Current status: valid genus in COCCINELLIDAE: COCCINELLINAE: COCCINELLINI. Note: new replacement name for Idalia Mulsant, 1846.

Type species: Coccinella bipunctata Linnaeus, 1758, by subsequent designation (Crotch 1874a: 99).

Family-group name: ADALIINA Hatch, 1962: 173. Stem: Adali-. Note: this name was overlooked in Bouchard et al. (2011) and Bouchard and Bousquet (2020a).

Adelaidia Blackburn, 1891: 130. Current status: valid genus in DERMESTIDAE: ATTAGENINAE: ADELAIDIINI.

Type species: Adelaidia rigua Blackburn, 1891, by monotypy.

Family-group name: ADELAIDIINA Háva, 2015: 1. Stem: Adelaidi-.

Adelina Dejean, 1835: 315. Current status: valid genus in TENEBRIONIDAE: DIAPERINAE: DIAPERINI: ADELININA.

Type species: Cucujus planus Fabricius, 1801, by monotypy.

Family-group name: ADELININI J.L. LeConte, 1862. Stem: Adelin-.

Adelium W. Kirby, 1819: 420. Current status: valid genus in TENEBRIONIDAE: LAGRIINAE: ADELIINI.

Type species: Adelium calosomoides W. Kirby, 1819, by subsequent designation (J. Meyer 1844: 180).

Family-group name: ADELIADAE W. Kirby, 1828. Stem: Adeli-. Note: senior homonym of ADELIINI Viereck, 1918 (type genus Adelius Haliday, 1833) in Hymenoptera.

Adelocera Latreille, 1829a: 451. Current status: valid genus in ELATERIDAE: AGRYPNINAE: AGRYPNINI.

Type species [suggested]: Elater ovalis Germar, 1823, by subsequent designation (Hyslop 1921: 623). Note: the valid type species fixation is that of Duponchel (1840: 120), who selected Elater fuscus Fabricius, 1801, but that species is currently included in the genus Agrypnus Eschscholtz, 1829 (see Kundrata et al. 2019c: 88); an application to the Commission is necessary to fix Elater ovalis Germar, 1823 as the type species of Adelocera Latreille, 1829.

Family-group name: ADELOCERAEIDAE Gistel, 1848. Stem: Adelocer-. Note: nomen oblitum (see Bouchard et al. 2011: 307).

Adelostoma Duponchel, 1827: 342. Current status: valid genus in TENEBRIONIDAE: PIMELIINAE: ADELOSTOMINI.

Type species: Adelostoma sulcatum Duponchel, 1827, by monotypy.

Family-group name: ADÉLOSTOMITES Solier, 1834. Stem: Adelostom-.

†Ademosyne Handlirsch, 1906: 402. Current status: valid genus in †ADEMOSYNIDAE. Note: Triassic (Argentina; Australia; Austria; Chile; South Africa).

Type species: †Ademosyne major Handlirsch, 1906, by subsequent designation (Tillyard 1916: 17).

Family-group name: †ADEMOSYNIDAE Ponomarenko, 1968. Stem: Ademosyn-.

Aderpas J. Thomson, 1864: 62. Current status: valid genus in CERAMBYCIDAE: LAMIINAE: ADERPASINI.

Type species: Crossotus griseus J. Thomson, 1858, by original designation.

Family-group name: ADERPASINI Breuning & Téocchi, 1978. Stem: Aderpas-.

Aderus Stephens, 1829b: 20. Current status: valid genus in ADERIDAE. Note: name placed on the Official List of Generic Names in Zoology (ICZN 1989d; original publication incorrectly given as Stephens 1829c: 255).

Type species: Lytta boleti Marsham, 1802 (= Notoxus populneus Creutzer, 1796), by subsequent designation (Westwood 1830a: 58; see ICZN 1989d). Note: Lytta boleti Marsham, 1802 was placed on the Official List of Specific Names in Zoology (ICZN 1989d).

Family-group name: ADERIDAE Csiki, 1909. Stem: Ader-.

Adesmia Fischer, 1822: 153. Current status: valid genus in TENEBRIONIDAE: PIMELIINAE: ADESMIINI.

Type species: Adesmia longipes Fischer, 1822 (= Pimelia anomala Fischer, 1820), by monotypy.

Family-group name: ADESMIIDES Lacordaire, 1859. Stem: Adesmi-. Note: nomen protectum (see Bouchard et al. 2007: 386, 2011: 861).

Adetus J.L. LeConte, 1852a: 161. Current status: valid genus in CERAMBYCIDAE: LAMIINAE: APOMECYNINI.

Type species: Polyopsia analis Melsheimer, 1847, by monotypy.

Family-group name: ADÉTIDES Lacordaire, 1872. Stem: Adet-.

Adimerus Sharp, 1894: 441. Current status: junior synonym of Monoedus G.H. Horn, 1882 in ZOPHERIDAE: COLYDIINAE: ADIMERINI.

Type species: Adimerus crispatus Sharp, 1894, by subsequent designation (Ivie and Ślipiński 1990: 15).

Family-group name: ADIMERIDAE Sharp, 1894. Stem: Adimer-.

Adinopsis Cameron, 1919: 242. Current status: valid genus in STAPHYLINIDAE: ALEOCHARINAE: GYMNUSINI.

Type species: Adinopsis rufobrunnea Cameron, 1919, by monotypy.

Family-group name: ADINOPSINI Cameron, 1919. Stem: Adinopse-.

Adoretus Dejean, 1833b: 157. Current status: valid genus in SCARABAEIDAE: RUTELINAE: ADORETINI: ADORETINA.

Type species: Melolontha nigrifrons Steven, 1809, by subsequent designation (S.I. Medvedev 1949: 313).

Family-group name: ADORETIDAE Burmeister, 1844. Stem: Adoret-.

Adorium Fabricius, 1801a: xx, 409. Current status: junior synonym of Oides Weber, 1801 in CHRYSOMELIDAE: GALERUCINAE: OIDINI.

Type species: Chrysomela bipunctata Fabricius, 1781 (= Oides andrewesi Jacoby, 1900), by subsequent designation (Latreille 1810: 432).

Family-group name: ADORIITES Chapuis, 1875. Stem: Adori-.

Adorodocia Brenske, 1893: 1. Current status: valid genus in SCARABAEIDAE: RUTELINAE: ADORETINI: ADORETINA.

Type species: Adorodocia maxima Brenske, 1893 (= Adoretus vittaticollis Fairmaire, 1883), by subsequent designation (Arrow 1901: 36).

Family-group name: ADORODOCIINA Ohaus, 1912. Stem: Adorodoci-.

Adoroleptus Brenske, 1893: 1. Current status: valid genus in SCARABAEIDAE: RUTELINAE: ADORETINI: ADORETINA.

Type species: Melolontha lanata Fabricius, 1801, by monotypy.

Family-group name: ADOROLEPTINA Ohaus, 1912. Stem: Adorolept-.

Adorrhinyptia Arrow, 1917: 273. Current status: valid genus in SCARABAEIDAE: RUTELINAE: ADORETINI: ADORRHINYPTIINA.

Type species: Rhinyptia dorsalis Burmeister, 1855, by original designation.

Family-group name: ADORRHINYPTIINI Arrow, 1917. Stem: Adorrhinypti-.

Adoxus W. Kirby, 1837: 209. Current status: junior synonym of Bromius Chevrolat, 1836 in CHRYSOMELIDAE: EUMOLPINAE: BROMIINI.

Type species: Cryptocephalus vitis Fabricius, 1775 (= Chrysomela obscura Linnaeus, 1758), by original designation.

Family-group name: ADOXINAE Baly, 1863. Stem: Adox-.

Adranes J.L. LeConte, 1849: 83. Current status: valid genus in STAPHYLINIDAE: PSELAPHINAE: CLAVIGERITAE: CLAVIGERINI.

Type species: Adranes coecus J.L. LeConte, 1849, by monotypy.

Family-group name: ADRANITES E. Desmarest, 1852. Stem: Adran-. Note: senior homonym of ADRANIDAE Scarlato & Starobogatov, 1985 (type genus Adrana H. Adams & A. Adams, 1858) in Mollusca.

Adrastus Eschscholtz, 1829a: 35. Current status: valid genus in ELATERIDAE: ELATERINAE: SYNAPTINI.

Type species: Elater limbatus Fabricius, 1777, by monotypy.

Family-group name: ADRASTITES Candèze, 1863. Stem: Adrast-.

Adzusa Kishii, 1957: 10. Current status: valid genus in ELATERIDAE incertae sedis.

Type species: Adzusa inexpecta Kishii, 1957, by monotypy.

Family-group name: ADZUSINI Kishii, 1989. Stem: Adzus-.

Aedemonus Schönherr, 1837: 243. Current status: valid genus in CURCULIONIDAE: CRYPTORHYNCHINAE: AEDEMONINI.

Type species: Aedemonus eminentepunctatus Boheman, 1837, by original designation.

Family-group name: ACDEMONINI Faust, 1898. Stem: Aedemon-.

Aedilis Audinet-Serville, 1835a: 32. Current status: junior synonym of Acanthocinus Dejean, 1821 in CERAMBYCIDAE: LAMIINAE: ACANTHOCININI.

Type species: Aedilis montana Audinet-Serville, 1835 (= Cerambyx aedilis Linneaus, 1758), by subsequent designation (Westwood 1838c: 40).

Family-group name: AEDILINAE Perrier, 1893. Stem: Aedil-.

Aegialia Latreille, 1806: 96. Current status: valid genus in SCARABAEIDAE: AEGIALIINAE: AEGIALIINI.

Type species: Scarabaeus globosus Kugelann, 1794 (= Scarabaeus arenarius Fabricius, 1787), by monotypy. Note: Scarabaeus arenarius Fabricius, 1787 was placed on the Official List of Specific Names in Zoology (ICZN 2006c).

Family-group name: AEGIALITES Laporte, 1840. Stem: Aegiali-.

Aegialites Mannerheim, 1853: 178. Current status: valid genus in SALPINGIDAE: AEGIALITINAE.

Type species: Aegialites debilis Mannerheim, 1853, by monotypy.

Family-group name: AEGIALITIDAE J.L. LeConte, 1862. Stem: Aegialit-.

Aegidinus Arrow, 1904: 739. Current status: valid genus in SCARABAEIDAE: ORPHNINAE: AEGIDIINI: AEGIDININA.

Type species: Aegidium guianense Westwood, 1846, by subsequent designation (Paulian 1984: 87).

Family-group name: AEGIDININA Frolov, Akhmetova & Neita-Moreno, 2023: 37. Stem: Aegidin-.

Aegidium Westwood, 1845a: 439, 440. Current status: valid genus in SCARABAEIDAE: ORPHNINAE: AEGIDIINI: AEGIDIINA.

Type species: Aegidium columbianum Westwood, 1845, by subsequent designation (Paulian 1984: 70).

Family-group name: AEGIDIINAE Paulian, 1984. Stem: Aegidi-.

Aegoprepes Pascoe, 1871c: 277. Current status: valid genus in CERAMBYCIDAE: LAMIINAE: AGAPANTHIINI.

Type species: Aegoprepes antennator Pascoe, 1871, by monotypy.

Family-group name: AEGOPREPINAE Pascoe, 1871. Stem: Aegoprep-.

Aegosoma Audinet-Serville, 1832: 162. Current status: valid genus in CERAMBYCIDAE: PRIONINAE: AEGOSOMATINI.

Type species: Cerambyx scabricornis Scopoli, 1763, by monotypy.

Family-group name: AEGOSOMITAE J. Thomson, 1861. Stem: Aegosomat-.

Aegostheta Dejean, 1833b: 159. Current status: valid genus in SCARABAEIDAE: MELOLONTHINAE: TANYPROCTINI: MACROPHYLLINA.

Type species: Melolontha longicornis Fabricius, 1787, by monotypy.

Family-group name: AEGOSTHETINI Lacroix, 2007. Stem: Aegosthet-.

Aegus W.S. MacLeay, 1819: 112. Current status: valid genus in LUCANIDAE: LUCANINAE: AEGINI.

Type species: Aegus chelifer W.S. MacLeay, 1819, by subsequent designation (Arrow 1950: 174).

Family-group name: AEGINI H. Huang & C.H. Chen, 2013: 68. Stem: Aeg-.

Aenictobia Seevers, 1953: 127. Current status: valid genus in STAPHYLINIDAE: ALEOCHARINAE: AENICTOTERATINI.

Type species: Aenictobia longicornis Seevers, 1953, by original designation.

Family-group name: AENICTOBIINA Kistner, 1997. Stem: Aenictobi-.

Aenictoteras W.M. Wheeler, 1932: 301. Current status: valid genus in STAPHYLINIDAE: ALEOCHARINAE: AENICTOTERATINI.

Type species: Aenictoteras chapmani W.M. Wheeler, 1932, by original designation.

Family-group name: AENICTOTERATINI Kistner, 1993. Stem: Aenictoterat-.

Aenigmaticum A. Matthews, 1888: 104. Current status: valid genus in CORYLOPHIDAE: CORYLOPHINAE: AENIGMATICINI.

Type species: Aenigmaticum ptilioides A. Matthews, 1888, by monotypy.

Family-group name: AENIGMATICINI Casey, 1900. Stem: Aenigmatic-.

Aephnidius W.S. MacLeay, 1825: 23. Current status: valid subgenus of Anaulacus W.S. MacLeay, 1825 in CARABIDAE: HARPALINAE: MASOREINI.

Type species: Anaulacus adelioides W.S. MacLeay, 1825, by monotypy.

Family-group name: AEPHNIDIINA Jakobson, 1907. Stem: Aephnidi-.

Aepus Leach in Samouelle, 1819: 149. Current status: valid genus in CARABIDAE: TRECHINAE: TRECHINI: TRECHINA. Note: this name is often attributed to Samouelle; however, in Opinion 1722 (ICZN 1993b) Leach was accepted as the author of the new genera attributed to him in Samouelle.

Type species: Aepus fulvescens Samouelle, 1819 (= Cicindela marina Strøm, 1783), by monotypy.

Family-group name: AËPYES Fowler, 1887. Stem: Aep-.

Aepyceratus Poinar, A.E. Brown & Legalov, 2017: 76. Current status: valid genus in MESOPHYLETIDAE: AEPYCERATINAE.

Type species: Aepyceratus hyperochus Poinar, A.E. Brown & Legalov, 2017, by original designation.

Family-group name: AEPYCERATINAE Poinar, A.E. Bown & Legalov, 2017: 75. Stem: Aepycerat-.

Aerenea J. Thomson, 1857: 298. Current status: valid genus in CERAMBYCIDAE: LAMIINAE: COMPSOSOMATINI. Note: the original spelling was Aerenaea but Aerenea, an incorrect subsequent spelling, is in prevailing usage, attributed to the publication of the original spelling, and therefore deemed to be the correct original spelling (ICZN 1999a, Article 33.3.1).

Type species: Aerenea posticalis J. Thomson, 1857, by monotypy.

Family-group name: AERENEITES J. Thomson, 1868. Stem: Aerene-.

Aerenica Dejean, 1835: 352. Current status: valid genus in CERAMBYCIDAE: LAMIINAE: AERENICINI.

Type species: Saperda canescens Klug, 1825, by monotypy.

Family-group name: AERÉNICIDES Lacordaire, 1872. Stem: Aerenic-.

Aesalus Fabricius, 1801a: xii. Current status: valid genus in LUCANIDAE: AESALINAE: AESALINI.

Type species: Lucanus scarabaeoides Panzer, 1795, by subsequent monotypy (Fabricius 1801b: 254).

Family-group name: AESALIDAE W.S. MacLeay, 1819. Stem: Aesal-.

Aferos Kazantsev, 1992: 44. Current status: valid genus in LYCIDAE: EROTINAE: SLIPINSKIINI.

Type species: Stadenus aethiops Kleine, 1933, by original designation.

Family-group name: AFEROTINI Kazantsev, 2004. Stem: Aferot-.

Afremus Levey, 1985: 420. Current status: valid genus in TENEBRIONOIDEA incertae sedis.

Type species: Afremus pickeri Levey, 1985, by original designation.

Family-group name: AFREMINAE Levey, 1985. Stem: Afrem-.

Afroapoderus Legalov, 2003: 483. Current status: valid genus in ATTELABIDAE: APODERINAE: HOPLAPODERINI: AFROAPODERINA.

Type species: Apoderus seminiger Faust, 1894, by original designation.

Family-group name: AFROAPODERINA Legalov, 2003. Stem: Afroapoder-.

Afrocorynus G.A.K. Marshall, 1955a: 21. Current status: valid genus in BELIDAE: OXYCORYNINAE: OXYCORYNITAE: AFROCORYNINI.

Type species: Afrocorynus turbatus G.A.K. Marshall, 1955, by original designation.

Family-group name: AFROCORYNINI Voss, 1957. Stem: Afrocoryn-.

Afroeubria Villiers, 1961: 439. Current status: valid genus in PSEPHENIDAE: AFROEUBRIINAE.

Type species: Afroeubria hydropetrica Villiers, 1961, by original designation.

Family-group name: AFROEUBRIINAE Lee, Satô, Shepard & Jäch, 2007. Stem: Afroeubri-.

Afroquedius Solodovnikov, 2006: 1075. Current status: valid genus in STAPHYLINIDAE: STAPHYLININAE: STAPHYLININI: AFROQUEDIINA.

Type species: Quedius sexpunctatus Bernhauer, 1917, by original designation.

Family-group name: AFROQUEDIINA Brunke, Żyła & Solodovnikov in Brunke et al., 2019: 179. Stem: Afroquedi-.

Agabetes Crotch, 1873d: 398, 401. Current status: valid genus in DYTISCIDAE: LACCOPHILINAE: AGABETINI.

Type species: Colymbetes acuductus Harris, 1828, by original designation.

Family-group name: AGABETINI Branden, 1884. Stem: Agabet-.

Agabinus Crotch, 1873d: 397. Current status: junior synonym of Platambus C.G. Thomson, 1859 in DYTISCIDAE: AGABINAE: AGABINI.

Type species: Colymbetes glabrellus Motschulsky, 1859, by original designation.

Family-group name: AGABININI Crotch, 1873. Stem: Agabin-.

Agabus Leach, 1817: 72. Current status: valid genus in DYTISCIDAE: AGABINAE: AGABINI.

Type species: Agabus paykullii Leach, 1817 (= Dytiscus serricornis Paykull, 1799), by monotypy.

Family-group name: AGABIDES C.G. Thomson, 1867. Stem: Agab-.

Agaeocera Saunders, 1871: 50. Current status: valid genus in BUPRESTIDAE: BUPRESTINAE: BUPRESTINI: AGAEOCERINA.

Type species: Anthaxia gigas Gory & Laporte, 1839, by monotypy.

Family-group name: AGAEOCERINI Nelson, 1982. Stem: Agaeocer-.

Agallissus Dalman, 1823: 66. Current status: valid genus in CERAMBYCIDAE: CERAMBYCINAE: AGALLISSINI.

Type species: Agallissus melaniodes Dalman, 1823, by monotypy.

Family-group name: AGALLISSINI J.L. LeConte, 1873. Stem: Agalliss-.

Agaocephala Lepeletier & Audinet-Serville, 1828: 370. Current status: valid genus in SCARABAEIDAE: DYNASTINAE: AGAOCEPHALINI. Note: the original spelling was Agacephala, but Agaocephala, an incorrect subsequent spelling, is in prevailing usage, attributed to the publication of the original spelling, and therefore deemed to be the correct original spelling (ICZN 1999a, Article 33.3.1).

Type species: Agaocephala cornigera Lepeletier & Audinet-Serville, 1828, by monotypy.

Family-group name: AGAOCEPHALIDAE Burmeister, 1847. Stem: Agaocephal-.

Agapanthia Audinet-Serville, 1835a: 35. Current status: valid genus in CERAMBYCIDAE: LAMIINAE: AGAPANTHIINI.

Type species: Cerambyx cardui Linnaeus, 1767, by subsequent designation (Westwood 1838c: 41).

Family-group name: AGAPANTHAIRES Mulsant, 1839. Stem: Agapanthi-.

Agapytho Broun, 1921: 542. Current status: valid genus in AGAPYTHIDAE.

Type species: Agapytho foveicollis Broun, 1921, by monotypy.

Family-group name: AGAPYTHINAE Sen Gupta & Crowson, 1969. Stem: Agapyth-.

Agathidium Panzer, 1796: no 13. Current status: valid genus in LEIODIDAE: LEIODINAE: ANISOTOMINI. Note: nomen protectum (see Wheeler and Miller 2005: 34).

Type species: Tetratoma globosa Herbst, 1791 (= Silpha seminulum Linnaeus, 1758), by monotypy.

Family-group name: AGATHIDIIDAE Westwood, 1838. Stem: Agathidi-.

Agelaea Gené, 1839: 49. Current status: valid genus in CARABIDAE: HARPALINAE: PLATYNINI.

Type species: Agelaea fulva Gené, 1839, by monotypy.

Family-group name: AGELAEINA Jakobson, 1907. Stem: Agelae-.

Agelastica Chevrolat in Dejean, 1836a: 381. Current status: valid genus in CHRYSOMELIDAE: GALERUCINAE: HYLASPINI.

Type species: Chrysomela alni Linnaeus, 1758, by monotypy.

Family-group name: AGELASTICITES Chapuis, 1875. Stem: Agelastic-.

Aglenus Erichson, 1845a: 285. Current status: valid genus in SALPINGIDAE: AGLENINAE.

Type species: Hypophlaeus brunneus Gyllenhal, 1813, by monotypy.

Family-group name: AGLENI G.H. Horn, 1878. Stem: Aglen-.

Aglycyderes Westwood, 1864a: 179. Current status: valid genus in BELIDAE: OXYCORYNINAE: AGLYCYDERITAE: AGLYCYDERINI.

Type species: Aglycyderes setifer Westwood, 1864, by monotypy.

Family-group name: AGLYCYDERIDAE Wollaston, 1864. Stem: Aglycyder-.

Aglymbus Sharp, 1880: cxlix. Current status: valid genus in DYTISCIDAE: COPELATINAE.

Type species: Agabus leprieurii Aubé, 1838, by monotypy.

Family-group name: AGLYMBINI Branden, 1884. Stem: Aglymb-.

Agnathus Germar, 1818: 232. Current status: valid genus in PYROCHROIDAE: AGNATHINAE. Note: this name, which is usually credited to “Germar, 1825”, was first published in an available work as a junior synonym and subsequently used as valid before 1961 by several authors, including in LaFerté-Sénectère (1849: 293), Agnathus was therefore made available from its first publication as a synonym (ICZN 1999a, Article 11.6.1).

Type species: Notoxus decoratus Germar, 1818, by monotypy.

Family-group name: AGNATHIDES Lacordaire, 1859. Stem: Agnath-.

Agnesiotis Pascoe, 1870: 474. Current status: valid genus in BELIDAE: BELINAE: PACHYURITAE: AGNESIOTIDINI.

Type species: Agnesiotis pilosula Pascoe, 1870, by monotypy.

Family-group name: AGNESIOTIDINI Zimmerman, 1994. Stem: Agnesiotid-.

Agnia Newman, 1842a: 291. Current status: valid genus in CERAMBYCIDAE: LAMIINAE: LAMIINI.

Type species: Agnia casta Newman, 1842, by subsequent designation (J. Thomson 1864: 86).

Family-group name: AGNITAE J. Thomson, 1864. Stem: Agni-.

Agonica Sloane, 1920: 174. Current status: valid genus in CARABIDAE: HARPALINAE: PELECIINI: AGONICINA.

Type species: Agonica simsoni Sloane, 1920, by subsequent designation (B.P. Moore 1963a: 22).

Family-group name: AGONICINI Sloane, 1920. Stem: Agonic-.

Agonum Bonelli, 1810: Tab. Syn. Current status: valid genus in CARABIDAE: HARPALINAE: PLATYNINI. Note: name placed on the Official List of Generic Names in Zoology (ICZN 1996d).

Type species: Carabus marginatus Linnaeus, 1758, by subsequent designation (Curtis 1827: pl. 183; see ICZN 1996d). Note: no species were originally included in this genus by Bonelli (1810); the first ones were the 12 species listed by Panzer (1813: 52–54); name placed on the Official List of Specific Names in Zoology (ICZN 1996d).

Family-group name: AGONIDAE W. Kirby, 1837. Stem: Agonum- (ICZN 1996d).

Agra Fabricius, 1801a: xxiii, 224. Current status: valid genus in CARABIDAE: HARPALINAE: LEBIINI: AGRINA.

Type species: Agra aenea Fabricius, 1801 (= Carabus cajennensis G.-A. Olivier, 1795), by subsequent designation (Latreille 1810: 426).

Family-group name: AGRIDAE W. Kirby, 1837. Stem: Agr-.

Agraphus Say, 1832: 13. Current status: valid genus in CURCULIONIDAE: ENTIMINAE: OTIORHYNCHITAE: AGRAPHINI. Note: see Prena (2018: 331) for notes on the authorship of this genus-group name.

Type species: Peritelus bellicus Say, 1832, by monotypy.

Family-group name: AGRAPHI G.H. Horn, 1876. Stem: Agraph-.

Agrilus Curtis, 1825: pl. 67. Current status: valid genus in BUPRESTIDAE: AGRILINAE: AGRILINI: AGRILINA.

Type species: Buprestis viridis Linnaeus, 1758, by original designation.

Family-group name: AGRILIDAE Laporte, 1835. Stem: Agril-.

Agriotes Eschscholtz, 1829a: 34. Current status: valid genus in ELATERIDAE: ELATERINAE: AGRIOTINI: AGRIOTINA.

Type species: Elater sputator Linnaeus, 1758, by subsequent designation (Curtis 1838a: pl. 694).

Family-group name: AGRIOTITES Laporte, 1840. Stem: Agriot-.

Agrodes Nordmann, 1837: 7, 161. Current status: junior synonym of Plochionocerus Dejean, 1833 in STAPHYLINIDAE: STAPHYLININAE: XANTHOLININI. Note: name placed on the Official List of Generic Names in Zoology (ICZN 1996c).

Type species: Agrodes elegans Nordmann, 1837, by monotypy (see ICZN 1996c). Note: name placed on the Official List of Specific Names in Zoology (ICZN 1996c).

Family-group name: AGRAEFORMES Nordmann, 1837. Stem: Agrod-. Note: name placed on the Official List of Family-Group Names in Zoology (ICZN 1996c, as AGRODINI Nordmann, 1837); younger name XANTHOLININI Erichson, 1839 given precedence over this name (ICZN 1996c).

Agronoma Gistel, 1848a: [2]. Current status: junior synonym of Amara Bonelli, 1810 in CARABIDAE: HARPALINAE: ZABRINI: AMARINA. Note: junior homonym of Agronoma Hübner, 1821 [Lepidoptera].

Type species: Carabus familiaris Duftschmid, 1812, by subsequent designation (Bouchard et al. 2011: 145).

Family-group name: AGRONOMAEIDAE Gistel, 1848. Stem: Agronom-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

Agrypnus Eschscholtz, 1829a: 32. Current status: valid genus in ELATERIDAE: AGRYPNINAE: AGRYPNINI.

Type species: Elater murinus Linnaeus, 1758, by subsequent designation (Westwood 1838c: 26).

Family-group name: AGRYPNIDES Candèze, 1857. Stem: Agrypn-. Note: nomen protectum (see Bouchard et al. 2011: 307).

Agyrtes Frölich, 1799: 15. Current status: valid genus in AGYRTIDAE: AGYRTINAE.

Type species: Mycetophagus castaneus Fabricius, 1792, by subsequent designation (Latreille 1810: 427).

Family-group name: AGYRTIDAE C.G. Thomson, 1859. Stem: Agyrt-.

Agyrtodes Portevin, 1907: 75. Current status: valid genus in LEIODIDAE: CAMIARINAE: AGYRTODINI.

Type species: Agyrtodes ovatus Portevin, 1907, by monotypy.

Family-group name: AGYRTODINI Jeannel, 1936. Stem: Agyrtod-.

Akalyptoischion Andrews, 1976: 3. Current status: valid genus in AKALYPTOISCHIIDAE.

Type species: Akalyptoischion tomeus Andrews, 1976, by original designation.

Family-group name: AKALYPTOISCHIIDAE Lord, Hartley, Lawrence, McHugh, Whiting & K.B. Miller, 2010. Stem: Akalyptoischi-.

Akatastopsis Pace, 2000a: 112. Current status: valid genus in STAPHYLINIDAE: ALEOCHARINAE: AKATASTOPSISINI.

Type species: Akatastopsis papuana Pace, 2000, by original designation.

Family-group name: AKATASTOPSISINI Pace, 2000. Stem: Akatastopsis- (ICZN 1999a, Article 29.4).

Akis Herbst, 1799: 124. Current status: valid genus in TENEBRIONIDAE: PIMELIINAE: AKIDINI.

Type species: Pimelia reflexa Fabricius, 1775, by subsequent designation (Latreille 1810: 429).

Family-group name: ACIDES Billberg, 1820. Stem: Akid-.

Alampes Champion, 1896: 463. Current status: invalid senior synonym of Peralampes P.J. Johnson, 2002 in ELATERIDAE: AGRYPNINAE: ANAISSINI. Note: this genus was first made available on page 463, issued on January 1896, by the association of two available species, Pyrophorus melanoxanthus Candèze, 1865 and Pyrophorus abnormis Candèze, 1863; the genus was described on page 474, issued on March 1896, with two species described, including Alampes vestitus Champion, 1896; the type species must be one of the species mentioned in January 1896 section of the publication; junior homonym of Alampes Horvath, 1884 [Hemiptera].

Type species: Pyrophorus melanoxanthus Candèze, 1865, by present designation. Note: the type species recognized in the literature, Alampes vestitus Champion, 1896, by subsequent designation (Hyslop 1921: 624), is not an originally available species as noted above.

Family-group name: ALAMPINA C. Costa, 1975. Stem: Alamp-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

Alanizus Di Iorio, 2003: 1. Current status: valid genus in CERAMBYCIDAE: CERAMBYCINAE: ALANIZINI.

Type species: Alanizus tortuosus Di Iorio, 2003, by original designation.

Family-group name: ALANIZINI Di Iorio, 2003. Stem: Alaniz-.

Alaudes G.H. Horn, 1870: 361. Current status: valid genus in TENEBRIONIDAE: PIMELIINAE: CNEMEPLATIINI: ALAUDINA.

Type species: Alaudes singularis G.H. Horn, 1870, by monotypy.

Family-group name: ALAUDINA Aalbu, Catarino & A.D. Smith, 2018: 250. Stem: Alaud-. Note: junior homonym of ALAUDIDAE Vigors, 1825 (type genus Alauda Linnaeus, 1758) in Aves.

Alaus Eschscholtz, 1829a: 33. Current status: valid genus in ELATERIDAE: AGRYPNINAE: HEMIRHIPINI: HEMIRHIPINA.

Type species: Elater oculatus Linnaeus, 1758, by subsequent designation (Duponchel 1840: 242).

Family-group name: ALAITES Candèze, 1874. Stem: Ala-.

Alceis Billberg, 1820: 45. Current status: valid genus in CURCULIONIDAE: ENTIMINAE: POLYDRUSITAE: NAUPACTINI: NAUPACTINA.

Type species: Curculio longimanus Fabricius, 1775 (= Siderodactylus ornatus Pascoe, 1879), by monotypy.

Family-group name: ALCEIDINI Pierce, 1913. Stem: Alceent-.

Alcides C.R. Sahlberg, 1823b: 47. Current status: invalid senior synonym of Alcidodes G.A.K. Marshall, 1939 in CURCULIONIDAE: MOLYTINAE: MECYSOLOBINI. Note: junior homonym of Alcides Hübner, 1822 [Lepidoptera].

Type species: Alcides senex C.R. Sahlberg, 1823, by monotypy.

Family-group name: ALCIDIDES Jekel, 1865. Stem: Alcid-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

Alcidodes G.A.K. Marshall, 1939a: 582. Current status: valid genus in CURCULIONIDAE: MOLYTINAE: MECYSOLOBINI. Note: new replacement name for Alcides Schönherr, 1823.

Type species: Alcides senex C.R. Sahlberg, 1823, by monotypy (type fixation for Alcides Schönherr).

Family-group name: ALCIDODINAE G.A.K. Marshall, 1939. Stem: Alcidod-.

Alegoria Laporte, 1840b: 221. Current status: valid genus in TENEBRIONIDAE: TENEBRIONINAE: ULOMINI.

Type species: Alegoria dilatata Laporte, 1840, by monotypy.

Family-group name: ALÉGORIIDES Lacordaire, 1859. Stem: Alegori-.

Aleochara Gravenhorst, 1802: 67. Current status: valid genus in STAPHYLINIDAE: ALEOCHARINAE: ALEOCHARINI: ALEOCHARINA.

Type species: Staphylinus curtulus Goeze, 1777, misidentified as Staphylinus fuscipes Linnaeus, 1758 (sensu Gravenhort, 1802) in the subsequent designation by Samouelle (1819: 177). Note: we follow Smetana (2004: 30) in using Article 69.2.4 (ICZN 1999a) to determine the identity of the type species for this genus; Goeze’s (1777) work is not consistently binominal (see ICZN 2021a) and therefore an application to the Commission is necessary to retain Staphylinus curtulus Goeze, 1777 as the valid name for the type species.

Family-group name: ALEOCHARIDAE Fleming, 1821. Stem: Aleochar-.

Alesia Reiche, 1848: pl. 26. Current status: junior synonym of Micraspis Chevrolat, 1836 in COCCINELLIDAE: COCCINELLINAE: COCCINELLINI. Note: this name is usually credited to Mulsant (1850: 343) in the literature but Reiche made it available earlier by the illustration of annulata on plate 26 of Ferret and Galinier’s ‘Voyage en Abyssinie’.

Type species: Alesia annulata Reiche, 1848, by monotypy.

Family-group name: ALESIAIRES Mulsant, 1850. Stem: Alesi-.

Alexia Stephens, 1833a: 23. Current status: junior synonym of Sphaerosoma Stephens, 1832 in ALEXIIDAE.

Type species: Tritoma piliferum P.W.J. Müller, 1821, by monotypy. Note: this species is usually listed as a junior synonym of Sphaerosoma quercus Samouelle, 1819 but S. quercus is a nomen nudum in Samouelle (1819: 394); the species was described by Stephens only in 1832.

Family-group name: ALEXIADAE Imhoff, 1856. Stem: Alexi-.

Algon Sharp, 1874a: 22. Current status: valid genus in STAPHYLINIDAE: STAPHYLININAE: STAPHYLININI: ALGONINA.

Type species: Algon grandicollis Sharp, 1874, by monotypy.

Family-group name: ALGONINA Schillhammer & Brunke in Chani-Posse et al., 2018: 30. Stem: Algon- (ICZN 1999a, Article 29.4).

†Alitrepanum Peng, Jiang, Engel & Wang in Peng et al., 2022: 2. Current status: valid genus in BOSTRICHIDAE: †ALITREPANINAE. Note: mid-Cretaceous (Myanmar).

Type species: †Alitrepanum aladelicatum Peng, Jiang, Engel & Wang, 2022, by original designation.

Family-group name: †ALITREPANINI Peng, Jiang, Engel & Wang in Peng et al., 2022: 2. Stem: Alitrepan-.

Allandrus J.L. LeConte in J.L. LeConte and G.H. Horn, 1876: 396. Current status: valid genus in ANTHRIBIDAE: ANTHRIBINAE: ALLANDRINI.

Type species: Allandrus bifasciatus J.L. LeConte, 1876, by monotypy.

Family-group name: ALLANDRINI Pierce, 1930. Stem: Allandr-.

Allapoderus Voss, 1927: 15. Current status: valid genus in ATTELABIDAE: APODERINAE: CLITOSTYLINI: ALLAPODERINA.

Type species: Apoderus dentipes Faust, 1883, by subsequent designation (Legalov 2003: 468).

Family-group name: ALLAPODERINA Legalov, 2003. Stem: Allapoder-.

Allara Britton, 1955: 124. Current status: valid genus in SCARABAEIDAE: SERICOIDINAE: LIPARETRINI.

Type species: Allara insularis Britton, 1955, by original designation.

Family-group name: ALLARINI Britton, 1955. Stem: Allar-.

Allecula Fabricius, 1801b: 21. Current status: valid genus in TENEBRIONIDAE: ALLECULINAE: ALLECULINI: ALLECULINA.

Type species: Cistela morio Fabricius, 1787, by subsequent designation (Duponchel 1840: 283).

Family-group name: ALLÉCULITES Laporte, 1840. Stem: Allecul-.

Alleuscelus Voss, 1937a: 159. Current status: valid genus in ATTELABIDAE: ATTELABINAE: ATTELABINI: EUSCELINA.

Type species: Alleuscelus violaceipennis Voss, 1937, by monotypy.

Family-group name: ALLEUSCELINA Legalov, 2003. Stem: Alleuscel-.

Allidiostoma Arrow, 1940: 16. Current status: valid genus in SCARABAEIDAE: ALLIDIOSTOMATINAE. Note: new replacement name for Idiostoma Arrow, 1904; the older but unused name Pytoderus Berg, 1880 (type species Orphnus strobeli Steinheil, 1873, currently placed in Allidiostoma) has priority over Allidiostoma Arrow, 1940; Reversal of Precedence (ICZN 1999a, Article 23.9) cannot be considered since Pytoderus has been used as valid at least once since 1900 (e.g., Ruiz 1924); an application to the Commission is necessary to conserve usage of the name Allidiostoma.

Type species: Idiostoma rufum Arrow, 1904, by original designation (type fixation for Idiostoma Arrow).

Family-group name: ALLIDIOSTOMIDAE Arrow, 1940. Stem: Allidiostomat-.

Allocorynus Sharp, 1890b: 46. Current status: junior synonym of Rhopalotria Chevrolat, 1878 in BELIDAE: OXYCORYNINAE: OXYCORYNITAE: ALLOCORYNINI.

Type species: Allocorynus mollis Sharp, 1890, by monotypy.

Family-group name: ALLOCORYNINAE Sharp, 1890. Stem: Allocoryn-.

†Alloioscarabaeus Bai, Ren & X.-K. Yang in Bai et al., 2012a: 163. Current status: valid genus in †ALLOIOSCARABAEIDAE. Note: Middle Jurassic (China).

Type species: †Alloioscarabaeus cheni Bai, Ren & X.-K. Yang, 2012, by original designation.

Family-group name: †ALLOIOSCARABAEIDAE Bai, Ren & X.-K. Yang in Bai et al., 2012a: 163. Stem: Alloioscarabae-.

Allomorphus Folwaczny, 1968: 41. Current status: valid genus in CURCULIONIDAE: COSSONINAE: ALLOMORPHINI.

Type species: Allomorphus franzi Folwaczny, 1968, by monotypy.

Family-group name: ALLOMORPHINI Folwaczny, 1973. Stem: Allomorph-.

Allopoda J.L. LeConte, 1866a: 144. Current status: valid genus in SCRAPTIIDAE: SCRAPTIINAE: ALLOPODINI.

Type species: Scraptia lutea Haldeman, 1848, by monotypy.

Family-group name: ALLOPODINI Franciscolo, 1964. Stem: Allopod-.

Allopogonia Cockerell, 1906: 241. Current status: valid genus in ARTEMATOPODIDAE: ALLOPOGONIINAE. Note: new replacement name for Allopogon G.H. Horn, 1880.

Type species: Allopogon villosus G.H. Horn, 1880, by monotypy (type fixation for Allopogon G.H. Horn).

Family-group name: ALLOPOGONINI Crowson, 1973. Stem: Allopogoni-.

Alloscelus Boucomont, 1923: 40. Current status: valid genus in SCARABAEIDAE: SCARABAEINAE: ONTHOPHAGINI.

Type species: Alloscelus paradoxus Boucomont, 1923, by monotypy.

Family-group name: ALLOSCELIDES A. Janssens, 1946. Stem: Alloscel-.

Allotrius Laporte, 1840a: 231. Current status: junior synonym of Senodonia Laporte, 1838 in ELATERIDAE: LISSOMINAE: PROTELATERINI. Note: junior homonym of Allotrius Temminck, 1835 [Aves].

Type species: Semiotus quadricollis Laporte, 1838, by monotypy.

Family-group name: ALLOTRIITES Candèze, 1863. Stem: Allotri-. Note: senior homonym of ALLOTRIINAE Förster, 1869 (type genus Allotria Westwood, 1833) in Hymenoptera; family-group name permanently invalid (ICZN 1999a, Article 39).

Alloxycorynus Voss, 1957: 101. Current status: valid genus in BELIDAE: OXYCORYNINAE: AGLYCYDERITAE: ALLOXYCORYNINI.

Type species: Oxycorynus bruchi Heller, 1911, by original designation.

Family-group name: ALLOXYCORYNINI Legalov, 2009. Stem: Alloxycoryn-.

Alniphagus Swaine, 1918: 73. Current status: valid genus in CURCULIONIDAE: SCOLYTINAE: HYLESININI.

Type species: Hylesinus aspericollis J.L. LeConte, 1876, by original designation.

Family-group name: ALNIPHAGINI Murayama, 1963. Stem: Alniphag-.

Alocorhinus C.R. Sahlberg, 1823a: 32. Current status: valid genus in CURCULIONIDAE: ENTIMINAE: ENTIMITAE: LORDOPINI.

Type species: Alocorhinus comprimatus C.R. Sahlberg, 1823, by subsequent designation (Jekel 1854: [VI] 12).

Family-group name: ALOCORHINI Jekel, 1856. Stem: Alocorhin-.

Alophus Schönherr, 1826: 166. Current status: junior synonym of Graptus Schönherr, 1823 in CURCULIONIDAE: ENTIMINAE: ENTIMITAE: BYRSOPAGINI: STRANGALIODINA.

Type species: Curculio triguttatus Fabricius, 1775, by original designation.

Family-group name: ALOPHINI J.L. LeConte, 1874. Stem: Aloph-.

Alphitobius Stephens, 1829b: 19. Current status: valid genus in TENEBRIONIDAE: TENEBRIONINAE: ALPHITOBIINI. Note: name placed on the Official List of Generic Names in Zoology (ICZN 1975).

Type species: Helops picipes Panzer, 1794 (= Opatrum laevigatum Fabricius, 1781), by monotypy (see ICZN 1975). Note: Opatrum laevigatum Fabricius, 1781 was placed on the Official List of Specific Names in Zoology (ICZN 1975).

Family-group name: ALPHITOBIINI Reitter, 1917. Stem: Alphitobi-.

Alphitophagus Stephens, 1832b: 12. Current status: valid genus in TENEBRIONIDAE: DIAPERINAE: DIAPERINI: ADELININA.

Type species: Alphitophagus quadripustulatus Stephens, 1832 (= Diaperis bifasciatus Say, 1824), by monotypy.

Family-group name: ALPHITOPHAGIDA Gistel, 1856. Stem: Alphitophag-.

Althanus Lewis, 1903: 420. Current status: valid genus in HISTERIDAE: HISTERINAE: PLATYSOMATINI.

Type species: Althanus teretrioides Lewis, 1903, by monotypy.

Family-group name: ALTHANINI Cooman, 1939. Stem: Althan-.

Altica Geoffroy, 1762: 244. Current status: valid genus in CHRYSOMELIDAE: GALERUCINAE: ALTICINI. Note: name placed on the Official List of Generic Names in Zoology (ICZN 1994a).

Type species: Chrysomela oleracea Linnaeus, 1758, by subsequent designation (Latreille 1810: 432; see ICZN 1994a). Note: name placed on the Official List of Specific Names in Zoology (ICZN 1994a).

Family-group name: HALTICITES Newman, 1834. Stem: Altic-.

Alurnus Fabricius, 1775: 94. Current status: valid genus in CHRYSOMELIDAE: CASSIDINAE: ALURNINI.

Type species: Alurnus grossus Fabricius, 1775, by monotypy.

Family-group name: ALURNITES Chapuis, 1875. Stem: Alurn-.

Alvarenganiella Viana & Martínez, 1971: 122. Current status: valid genus in PTINIDAE: ALVARENGANIELLINAE.

Type species: Alvarenganiella seabrai Viana & Martínez, 1971 (= Dasytanobium monstrosum Pic, 1910), by original designation.

Family-group name: ALVARENGANIELLINAE Viana & Martínez, 1971. Stem: Alvarenganiell-.

Alvarengius Frey, 1975: 84. Current status: junior synonym of Ottokelleria d’Andretta & Martínez, 1957 in SCARABAEIDAE: RUTELINAE: ALVARENGIINI.

Type species: Alvarengius silphoides Frey, 1975 (= Pachylus dispar Burmeister, 1847), by monotypy.

Family-group name: ALVARENGIINI Frey, 1975. Stem: Alvarengi-.

Alyculus Kazantsev, 1999: 252. Current status: valid genus in LYCIDAE: LYROPAEINAE: ALYCULINI.

Type species: Alyculus kurbatovi Kazantsev, 1999, by original designation.

Family-group name: ALYCULINI Bocák & Bocáková, 2008. Stem: Alycul-.

Alzadaesthetus Kistner, 1961: 217. Current status: valid genus in STAPHYLINIDAE: EUAESTHETINAE: ALZADAESTHETINI.

Type species: Alzadaesthetus chilensis Kistner, 1961, by original designation.

Family-group name: ALZADAESTHETINI Scheerpeltz, 1974. Stem: Alzadaesthet-.

Amalactus Schönherr, 1835: 251. Current status: valid genus in CURCULIONIDAE: MOLYTINAE: AMALACTINI.

Type species: Amalactus nigritus Gyllenhal, 1835, by original designation.

Family-group name: AMALACTIDES Lacordaire, 1863. Stem: Amalact-.

Amalthocus Fairmaire, 1886: 40. Current status: valid genus in MELYRIDAE: MALACHIINAE: AMALTHOCINI.

Type species: Amalthocus tetraspilus Fairmaire, 1886, by original designation.

Family-group name: AMALTHOCINAE Majer, 2002. Stem: Amalthoc-.

Amalus Schönherr, 1825: 583. Current status: valid genus in CURCULIONIDAE: CEUTORHYNCHINAE: AMALINI.

Type species: Curculio scortillum Herbst, 1795, by original designation.

Family-group name: AMALINA Wagner, 1936. Stem: Amal-.

Amara Bonelli, 1810: Tab. Syn. Current status: valid genus in CARABIDAE: HARPALINAE: ZABRINI: AMARINA.

Type species: here fixed (under ICZN 1999a, Article 70.3.2) as Amara lunicollis Schiødte, 1837, misidentified as Carabus vulgaris Linnaeus, 1758 (sensu Panzer, 1796), in the subsequent designation by Westwood (1838c: 4). Note: no species were originally included in this genus by Bonelli (1810); the first ones were the eight species listed by Panzer (1813: 57–59).

Family-group name: AMAROIDEN C. Zimmermann, 1832. Stem: Amar-.

Amarotypus H.W. Bates, 1872: 50. Current status: valid genus in CARABIDAE: MIGADOPINAE: AMAROTYPINI.

Type species: Amarotypus edwardsii H.W. Bates, 1872, by monotypy.

Family-group name: AMAROTYPINI Erwin, 1985. Stem: Amarotyp-.

Amarygmus Dalman, 1823: 60. Current status: valid genus in TENEBRIONIDAE: TENEBRIONINAE: AMARYGMINI.

Type species: Chrysomela micans Fabricius, 1794, by subsequent designation (Gebien 1921b: 410).

Family-group name: AMARYGMIIDAE Gistel, 1848. Stem: Amarygm-.

Amauronioides Champion, 1923a: 13. Current status: junior synonym of Pseudamauronia Pic, 1915 in MELYRIDAE: DASYTINAE: DANACEINI.

Type species: Amauronioides schilskyi Champion, 1923 (= Dasytiscus danacaeoides Reitter, 1885), by monotypy. Note: combined description by indication of a new genus and a single new species (ICZN 1999a, Article 12.2.6).

Family-group name: AMAURONOININI Majer, 1987. Stem: Amauronioid-.

Amaurops Fairmaire, 1851: 527. Current status: valid genus in STAPHYLINIDAE: PSELAPHINAE: BATRISITAE: AMAUROPINI.

Type species: Amaurops aubei Fairmaire, 1851, by monotypy.

Family-group name: AMAUROPSINI Jeannel, 1948. Stem: Amaurop-.

†Ambarticus Q. Yang, Z.-Y. Chen & Jia, 2019: 2. Current status: valid genus in DYTISCIDAE: DYTISCINAE: †AMBARTICINI. Note: mid-Cretaceous (Myanmar). Note: two original spellings were used, including Ambraticus (pp. 3, 4); as far as we know, no one has acted as First Reviser (ICZN 1999a, Article 24.2.3) and we here select Ambarticus as the correct original spelling.

Type species: †Ambarticus myanmaricus Q. Yang, Chen & Jia, 2019, by original designation.

Family-group name: †AMBARTICINI Q. Yang, Z.-Y. Chen & Jia, 2019: 2. Stem: Ambartic-. Note: two spellings were originally used for the name of this new tribe, including AMBRATICINI (p. 1); as far as we know no one has acted as First Reviser (ICZN 1999a, Article 24.2.3) so we here select AMBARTICINI as the correct original spelling.

Ambates Schönherr, 1835: 278. Current status: junior synonym of Embates Chevrolat, 1833 in CURCULIONIDAE: BARIDINAE: AMBATINI.

Type species: Ambates pictus Gyllenhal, 1835, by original designation.

Family-group name: AMBATIDES Lacordaire, 1863. Stem: Ambat-.

Ambicocerus Leleup, 1970: 314. Current status: valid genus in STAPHYLINIDAE: PSELAPHINAE: BATRISITAE: BATRISINI: AMBICOCERINA.

Type species: Ambicocerus carinaticollis Leleup, 1970, by original designation.

Family-group name: AMBICOCERINA Leleup, 1970. Stem: Ambicocer-.

Amblycerus Thunberg, 1815: 109. Current status: valid genus in CHRYSOMELIDAE: BRUCHINAE: AMBLYCERINI: AMBLYCERINA.

Type species [suggested]: Bruchus robiniae Fabricius, 1781, by subsequent designation (Bridwell in Pierce 1930: 29). Note: the valid type species fixation is that of Crotch (1870b: 227), who selected Anthribus nebulosus Forster, 1770 (see Bradley 1946: 97, 98), but that species is currently included in the genus Anthribus Geoffroy, 1762; Alonso-Zarazaga and Lyal (1999: 8, 23) mentioned that they would submit an application to the Commission to fix this problem but to our knowledge this has not been done, and an application is still necessary to fix Bruchus robiniae Fabricius, 1781 as the type species of Amblycerus Thunberg, 1815.

Family-group name: AMBLYCERINAE Bridwell, 1932. Stem: Amblycer-.

Amblycheila Say, 1830a: 67. Current status: valid genus in CICINDELIDAE: MANTICORINI.

Type species: Manticora cylindriformis Say, 1823, by original designation.

Family-group name: AMBLYCHILINAE Csiki, 1903. Stem: Amblycheil-.

Amblyderus LaFerté-Sénectère in P.H. Lucas, 1847b: 368. Current status: valid genus in ANTHICIDAE: ANTHICINAE: ANTHICINI. Note: the original spelling was Aneblyderes but the spelling Amblyderus, changed by LaFerté-Sénectère in Guérin-Méneville (1849 [no 23]: 1), is in prevailing usage, attributed to the publication of the original spelling, and therefore deemed to be the correct original spelling (ICZN 1999a, Article 33.2.3.1).

Type species: Anthicus scabricollis LaFerté-Sénectère, 1847, by monotypy.

Family-group name: AMBLYDERINI Desbrochers des Loges, 1899. Stem: Amblyder-.

Amblyopinus Solsky, 1875: 10. Current status: valid genus in STAPHYLINIDAE: STAPHYLININAE: STAPHYLININI: AMBLYOPININA.

Type species: Amblyopinus ielskii Solsky, 1875, by subsequent designation (R. Lucas 1920: 88).

Family-group name: AMBLYOPININAE Seevers, 1944. Stem: Amblyopin-.

Amblysterna Saunders, 1871: 2. Current status: valid genus in BUPRESTIDAE: JULODINAE.

Type species: Julodis natalensis Fåhraeus, 1851, by subsequent designation (Rye 1873: 271).

Family-group name: AMBLYSTERNINI Cobos, 1955. Stem: Amblystern-.

Amblystomus Erichson, 1837: 59. Current status: valid genus in CARABIDAE: HARPALINAE: HARPALINI: AMBLYSTOMINA.

Type species: Acupalpus mauritanicus Dejean, 1829, by subsequent designation (Andrewes 1938: 130).

Family-group name: AMBLYSTOMINI Fauvel, 1889. Stem: Amblystom-.

Amblytelus Erichson, 1842: 129. Current status: valid genus in CARABIDAE: PSYDRINAE: MORIOMORPHINI: AMBLYTELINA.

Type species: Carabus curtus Fabricius, 1801, by monotypy.

Family-group name: AMBLYTELIDES Blackburn, 1892. Stem: Amblytel-. Note: senior homonym of AMBLYTELINA Viereck, 1918 (type genus Amblyteles Wesmael, 1845) in Hymenoptera.

Amborotubus Leschen & Carlton, 2004: 443. Current status: valid genus in NITIDULIDAE: NITIDULINAE: CYLLODINI.

Type species: Amborotubus clarkei Leschen & Carlton, 2004, by original designation.

Family-group name: AMBOROTUBINI Leschen & Carlton, 2004. Stem: Amborotub-.

Ameris Dejean, 1821: 86. Current status: valid genus in CURCULIONIDAE: MOLYTINAE: CHOLINI: CHOLINA.

Type species: Rhynchaenus dufresnii W. Kirby, 1819, by monotypy.

Family-group name: AMERININAE Pierce, 1919. Stem: Amerin-.

Ametalla Hope, 1841a: 179. Current status: valid genus in CHRYSOMELIDAE: SAGRINAE: MEGAMERINI.

Type species: Ametalla spinolae Hope, 1841, by original designation.

Family-group name: AMÉTALLITES Chapuis, 1874. Stem: Ametall-.

Ametrocephala Blanchard, 1851: 480. Current status: junior synonym of Pseudocephalus Newman, 1842 in CERAMBYCIDAE: CERAMBYCINAE: PSEUDOCEPHALINI.

Type species: Ametrocephala monstrosa Blanchard, 1851, by monotypy.

Family-group name: AMETROCEPHALITAE J. Thomson, 1861. Stem: Ametrocephal-.

Aminyops Voss, 1956: 29. Current status: junior synonym of Niphadonyx Schenkling, 1932 in CURCULIONIDAE: MOLYTINAE: AMINYOPINI.

Type species: Aminyops castaneus Voss, 1956, by monotypy.

Family-group name: AMINYOPINI Voss, 1956. Stem: Aminyop-.

Ammobius Guérin-Méneville, 1844a: 121. Current status: valid genus in TENEBRIONIDAE: BLAPTINAE: OPATRINI: AMMOBIINA.

Type species: Ammobius rufus Guérin-Méneville, 1844, by monotypy. Note: combined description of a new genus and a single new species (ICZN 1999a, Article 12.2.6).

Family-group name: AMMOBIINI Desbrochers des Loges, 1902. Stem: Ammobi-.

Amomphus Schönherr, 1848: 359. Current status: junior synonym of Aspidiotes Schönherr, 1847 in CURCULIONIDAE: ENTIMINAE: POLYDRUSITAE: TANYMECINI: TAINOPHTHALMINA. Note: unnecessary new replacement name for Aspidiotes Schönherr, 1847.

Type species: Aspidiotes westringii Schönherr, 1847, by original designation (type fixation for Aspidiotes Schönherr).

Family-group name: AMOMPHI J.L. LeConte, 1874. Stem: Amomph-.

Amorphocephalus Schönherr, 1840a: 485. Current status: invalid senior synonym of Amorphocephala Damoiseau, 1966 in BRENTIDAE: BRENTINAE: BRENTINI: EREMOXENINA. Note: junior homonym of Amorphocephalus Bowdich, 1825 [Pisces].

Type species: Brentus coronatus Germar, 1817, by original designation.

Family-group name: AMORPHOCEPHALIDES Power, 1879. Stem: Amorphocephal-. Note: family- group name permanently invalid (ICZN 1999a, Article 39).

Amorphocerus Schönherr, 1826: 329. Current status: valid genus in CURCULIONIDAE: MOLYTINAE: AMORPHOCERINI.

Type species: Amorphocerus talpa Schönherr, 1826, by original designation.

Family-group name: AMORPHOCERINI Voss, 1939. Stem: Amorphocer-.

Amorphomerus Sloane, 1923: 249. Current status: valid genus in CARABIDAE: HARPALINAE: AMORPHOMERINI. Note: new replacement name for Trimerus Chaudoir, 1878.

Type species: Trimerus raffrayi Chaudoir, 1878, by monotypy (type fixation for Trimerus Chaudoir).

Family-group name: AMORPHOMERINI Sloane, 1923. Stem: Amorphomer-.

Amorphosoma Laporte, 1835: 166. Current status: valid genus in BUPRESTIDAE: AGRILINAE: CORAEBINI: AMORPHOSOMINA.

Type species: Buprestis penicillata Klug, 1825, by subsequent designation (Descarpentries and Villiers 1967: 488).

Family-group name: AMORPHOSOMINA Majer, 2000. Stem: Amorphosom- (ICZN 1999a, Article 29.4).

Amorphosternus Deyrolle, 1865: 114. Current status: valid genus in BUPRESTIDAE: AGRILINAE: AGRILINI: AMORPHOSTERNINA.

Type species: Amorphosternus cucullatus Deyrolle, 1865, by monotypy.

Family-group name: AMORPHOSTERNAE Cobos, 1974. Stem: Amorphostern-.

Ampedus Dejean, 1833a: 92. Current status: valid genus in ELATERIDAE: ELATERINAE: AMPEDINI.

Type species: Elater sanguineus Linnaeus, 1758, by subsequent designation (Curtis 1838a: pl. 694).

Family-group name: AMPEDIDAE Gistel, 1848. Stem: Amped-.

Amphicoma Latreille, 1806: 118. Current status: valid genus in GLAPHYRIDAE: AMPHICOMINAE.

Type species: Melolontha abdominalis Fabricius, 1781, by subsequent designation (Latreille 1810: 428).

Family-group name: AMPHICOMITES Blanchard, 1845. Stem: Amphicom-.

Amphicranus Erichson, 1836: 63. Current status: valid genus in CURCULIONIDAE: SCOLYTINAE: CORTHYLINI: CORTHYLINA.

Type species: Amphicranus thoracicus Erichson, 1836, by monotypy.

Family-group name: AMPHICRANIDAE Eichhoff, 1878. Stem: Amphicran-.

Amphicrossus Erichson, 1843a: 346. Current status: valid genus in NITIDULIDAE: AMPHICROSSINAE.

Type species: Nitidula ciliata G.-A. Olivier, 1811, by subsequent designation (E. Desmarest 1851: 283).

Family-group name: AMPHICROSSINI Kirejtshuk, 1986. Stem: Amphicross-.

Amphicyrta Erichson in Steffahny, 1842: 5, 39. Current status: valid genus in BYRRHIDAE: AMPHICYRTINAE.

Type species: Amphicyrta dentipes Erichson, 1842, by subsequent designation (Chevrolat 1843a: 657).

Family-group name: AMPHICYRTINI J.L. LeConte, 1861. Stem: Amphicyrt-.

Amphidora Eschscholtz, 1829d: 9. Current status: valid genus in TENEBRIONIDAE: BLAPTINAE: AMPHIDORINI.

Type species: Amphidora littoralis Eschscholtz, 1829, by monotypy.

Family-group name: AMPHIDORAE J.L. LeConte, 1862. Stem: Amphidor-.

Amphilabris Gistel, 1834: 12. Current status: junior synonym of Sericus Eschscholtz, 1829 in ELATERIDAE: ELATERINAE: ELATERINI.

Type species: Elater brunneus Linnaeus, 1758, by subsequent designation (Sánchez-Ruiz 1996: 155).

Family-group name: AMPHILABRIDAE Gistel, 1856. Stem: Amphilabr-.

Amphionycha Dejean, 1835: 352. Current status: junior synonym of Adesmus Lepeletier & Audinet- Serville, 1825 in CERAMBYCIDAE: LAMIINAE: HEMILOPHINI.

Type species: Saperda hemispila Germar, 1821, by subsequent designation (Marinoni 1977: 40).

Family-group name: AMPHIONYCHITAE J. Thomson, 1860. Stem: Amphionych-. Note: nomen oblitum (see Bousquet et al. 2009: 31).

Amphiops Erichson, 1843b: 229. Current status: valid genus in HYDROPHILIDAE: HYDROPHILINAE: AMPHIOPINI.

Type species: Hydrophilus gibbus Illiger, 1801, by subsequent designation (Knisch 1924: 259).

Family-group name: AMPHIOPITAE Kuwert, 1890. Stem: Amphiop-.

Amphiscolytus Mandelshtam & Beaver, 2003: 2. Current status: valid genus in CURCULIONIDAE: SCOLYTINAE: AMPHISCOLYTINI.

Type species: Dacryophthorus capensis Schedl, 1971, by original designation.

Family-group name: AMPHISCOLYTINI Mandelshtam & Beaver, 2003. Stem: Amphiscolyt-.

Amphisternus Germar, 1843: 85. Current status: valid genus in ENDOMYCHIDAE: LYCOPERDININAE.

Type species [suggested]: Amphisternus tuberculatus Germar, 1843, by subsequent designation (R. Lucas 1920: 91). Note: the valid type species fixation is that of Guérin-Méneville (1857: 579), who selected Amphisternus inaequalis Germar, 1843, but that species is currently included in the genus Cacodaemon J. Thomson, 1857; an application to the Commission is necessary to fix Amphisternus tuberculatus Germar, 1843 as the type species of Amphisternus Germar, 1843.

Family-group name: AMPHISTERNINI Strohecker, 1964. Stem: Amphistern-.

Amphix Laporte, 1840b: 522. Current status: junior synonym of Corynomalus Chevrolat, 1836 in ENDOMYCHIDAE: LYCOPERDININAE.

Type species: Eumorphus binotatus Laporte, 1840 (= Erotylus marginatus Fabricius, 1798), by subsequent designation (Strohecker 1980: 14).

Family-group name: AMPHICINI Csiki, 1910. Stem: Amphic-.

Amphizoa J.L. LeConte, 1853b: 227. Current status: valid genus in AMPHIZOIDAE.

Type species: Amphizoa insolens J.L. LeConte, 1853, by monotypy.

Family-group name: AMPHIZOIDAE J.L. LeConte, 1853. Stem: Amphizo-.

Amphoecus Montrouzier, 1861b: 274. Current status: valid genus in CERAMBYCIDAE: LAMIINAE: AMPHOECINI.

Type species: Amphoecus metallicus Montrouzier, 1861, by monotypy.

Family-group name: AMPHOECINI Breuning, 1951. Stem: Amphoec-.

Amplipalpa Harold, 1875: 185. Current status: junior synonym of Oediopalpa Baly, 1858 in CHRYSOMELIDAE: CASSIDINAE: OEDIOPALPINI. Note: unnecessary new replacement name for Oediopalpa Baly, 1858.

Type species: Hispa cyanipennis Fabricius, 1801, by subsequent designation (Monrós and Viana 1947: 151) (type fixation for Oediopalpa Baly).

Family-group name: AMPLIPALPINI Weise, 1910. Stem: Amplipalp-.

Amycterus Schönherr, 1823: 1144. Current status: valid genus in CURCULIONIDAE: CYCLOMINAE: AMYCTERINI.

Type species: Amycterus talpa Schönherr, 1823, by original designation.

Family-group name: AMYCTERIDAE G.R. Waterhouse, 1854. Stem: Amycter-.

Amydetes Illiger, 1807b: 342. Current status: valid genus in LAMPYRIDAE: AMYDETINAE.

Type species: Amydetes fastigiata Illiger, 1807, by monotypy.

Family-group name: AMYDETINI E. Olivier, 1907. Stem: Amydet-.

Amyia Saunders, 1871: 111. Current status: valid genus in BUPRESTIDAE: AGRILINAE: AGRILINI: AMYIINA.

Type species: Eumerus violaceus Gory & Laporte, 1839, by subsequent designation (Rye 1873: 271).

Family-group name: AMYIINA Holyński, 1993. Stem: Amyi-.

Amymoma Pascoe, 1866c: 329, 332. Current status: valid genus stat. nov. in CERAMBYCIDAE: LAMIINAE: DESMIPHORINI. Note: this name has been considered a junior homonym of Amymoma Latreille, 1797 [Crustacea] but Latreille’s name is an incorrect subsequent spelling of Amymone O.F. Müller, 1776 as noted by Neave (1939: 168) not in prevailing usage, and therefore the name is not available; the new replacement name Neoamymoma Marinoni, 1977 is therefore unnecessary.

Type species: Amymoma pulchella Pascoe, 1866, by monotypy.

Family-group name: AMYMOMIDES Lacordaire, 1872. Stem: Amymom-.

Anacaena C.G. Thomson, 1859: 18. Current status: valid genus in HYDROPHILIDAE: CHAETARTHRIINAE: ANACAENINI. Note: name placed on the Official List of Generic Names in Zoology (ICZN 1960).

Type species: Hydrophilus globulus Paykull, 1798, by original designation (see ICZN 1960, type fixation given as “by monotypy”). Note: name placed on the Official List of Specific Names in Zoology (ICZN 1960).

Family-group name: ANACAENINI Hansen, 1991. Stem: Anacaen-.

Anacanthus Audinet-Serville, 1832: 165. Current status: invalid senior synonym of Chorenta Gistel, 1848 in CERAMBYCIDAE: PRIONINAE: CALLIPOGONINI. Note: junior homonym of Anacanthus J.E. Gray, 1831 [Pisces].

Type species: Anacanthus costatus Audinet-Serville, 1832 (= Prionus reticulatus Dalman, 1817), by monotypy.

Family-group name: ANACANTHITAE J. Thomson, 1864. Stem: Anacanth-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

Anacolus Berthold, 1827: 395. Current status: valid genus in CERAMBYCIDAE: PRIONINAE: ANACOLINI.

Type species: Prionus sanguineus Lepeletier & Audinet-Serville, 1825, by subsequent designation (E. Desmarest 1860: 306). Note: no species were originally included in this genus by Berthold (1827); the first ones were the two species listed by Latreille (1829b: 108).

Family-group name: ANACOLITES J. Thomson, 1857. Stem: Anacol-.

Anaesthetis Dejean, 1835: 348. Current status: valid genus in CERAMBYCIDAE: LAMIINAE: DESMIPHORINI.

Type species: Saperda testacea Fabricius, 1781, by monotypy.

Family-group name: ANAESTHETINAE Perrier, 1893. Stem: Anaesthet-.

Anaglyptes Gistel, 1848b: 128. Current status: junior synonym of Chalcophora Solier, 1833 in BUPRESTIDAE: CHRYSOCHROINAE: CHRYSOCHROINI: CHALCOPHORINA.

Type species: Anaglyptes merianus Gistel, 1848 (= Buprestis mariana Linnaeus, 1758), by monotypy.

Family-group name: ANAGLYPTISIDAE Gistel, 1848. Stem: Anaglypt-. Note: nomen oblitum (see Bousquet et al. 2009: 41); senior homonym of ANAGLYPTINI Lacordaire, 1868 (type genus Anaglyptus Mulsant, 1839) in Coleoptera: CERAMBYCIDAE.

Anaglyptus Mulsant, 1839: 71, 91. Current status: valid genus in CERAMBYCIDAE: CERAMBYCINAE: ANAGLYPTINI. Note: two original spellings were used, including Anaclyptus (p. 71); subsequently, Mulsant (1863: 184, 185) used only Anaglyptus and therefore acted as First Reviser (ICZN 1999a, Article 24.2.4).

Type species: Leptura mystica Linnaeus, 1758, by subsequent designation (C.G. Thomson 1859: 150).

Family-group name: ANAGLYPTIDES Lacordaire, 1868. Stem: Anaglypt-. Note: nomen protectum (see Bousquet et al. 2009: 41); junior homonym of ANAGLYPTIDAE Gistel, 1848 (type genus Anaglyptes Gistel, 1848) in Coleoptera: BUPRESTIDAE.

Anaides Westwood, 1842a: 457. Current status: valid genus in HYBOSORIDAE: ANAIDINAE.

Type species: Anaides fossulatus Westwood, 1842, by monotypy.

Family-group name: ANAIDINI Nikolajev, 1996. Stem: Anaid-.

Anaissus Candèze, 1857: 187. Current status: valid genus in ELATERIDAE: AGRYPNINAE: ANAISSINI.

Type species: Anaissus tarsalis Candèze, 1857, by monotypy.

Family-group name: ANAISSINI Golbach, 1984. Stem: Anaiss-.

Anamorphus J.L. LeConte in E.A. Schwarz, 1878: 445. Current status: valid genus in ANAMORPHIDAE.

Type species: Anamorphus pusillus J.L. LeConte, 1878, by monotypy.

Family-group name: ANAMORPHINI Strohecker, 1953. Stem: Anamorph-.

Anapleus G.H. Horn, 1873b: 311. Current status: valid genus in HISTERIDAE: DENDROPHILINAE: ANAPLEINI.

Type species: Bacanius marginatus J.L. LeConte, 1853, by monotypy.

Family-group name: ANAPLEINI Olexa, 1982. Stem: Anaple-.

Anaplopus Blackburn, 1890: 311. Current status: valid genus in PYTHIDAE.

Type species: Anaplopus tuberculatus Blackburn, 1890, by monotypy.

Family-group name: ANAPLOPINAE M. Abdullah, 1966. Stem: Anaplopod-.

Anaspimorda Ermisch, 1950: 89. Current status: valid genus in SCRAPTIIDAE: ANASPIDINAE: ANASPIMORDINI.

Type species: Anaspimorda forticornis Ermisch, 1950, by original designation.

Family-group name: ANASPIMORDINI Franciscolo, 1954. Stem: Anaspimord-.

Anaspis Geoffroy, 1762: 315. Current status: valid genus in SCRAPTIIDAE: ANASPIDINAE: ANASPIDINI. Note: name attributed to Geoffroy, 1762 and placed on the Official List of Generic Names in Zoology (ICZN 1984a).

Type species: Mordella frontalis Linnaeus, 1758, by plenary power (see ICZN 1984a). Note: name placed on the Official List of Specific Names in Zoology (ICZN 1984a).

Family-group name: ANASPIENS Mulsant, 1856. Stem: Anaspid-.

Anatista Brême, 1845: 305. Current status: valid genus in SCARABAEIDAE: RUTELINAE: ANATISTINI.

Type species: Anatista lafertei Brême, 1845, by monotypy.

Family-group name: ANATISTIDES Lacordaire, 1855. Stem: Anatist-.

Anaulacus W.S. MacLeay, 1825: 22. Current status: valid genus in CARABIDAE: HARPALINAE: MASOREINI.

Type species: Anaulacus sericipennis W.S. MacLeay, 1825, by monotypy.

Family-group name: ANAULACINI Csiki, 1932. Stem: Anaulac-.

Anauxesis J. Thomson, 1857: 191. Current status: valid genus in CERAMBYCIDAE: LAMIINAE: AGAPANTHIINI.

Type species: Nemotragus calabaricus Chevrolat, 1855, by subsequent designation (J. Thomson 1864: 94).

Family-group name: ANAUXESITAE J. Thomson, 1864. Stem: Anauxese-.

†Anchineus Poinar & A.E. Brown, 2009a: 264. Current status: valid genus in †MESOPHYLETIDAE: †MESOPHYLETINAE. Note: mid-Cretaceous (Myanmar).

Type species: †Anchineus dolichobothris Poinar & A.E. Brown, 2009, by original designation.

Family-group name: †ANCHINEINI Poinar & Legalov in Legalov and Poinar, 2014: 558. Stem: Anchine-.

Anchisteus Kolbe, 1883: 185. Current status: valid genus in BRENTIDAE: BRENTINAE: TRACHELIZINI: HEPHEBOCERINA.

Type species: Anchisteus peregrinus Kolbe, 1883, by monotypy.

Family-group name: ANCHISTENINI Schedl, 1961. Stem: Anchiste-.

Anchomenus Bonelli, 1810: Tab. Syn. Current status: valid genus in CARABIDAE: HARPALINAE: PLATYNINI.

Type species: Carabus prasinus Thunberg, 1784 (= Carabus dorsalis Pontoppidan, 1763), by subsequent designation (Westwood 1838c: 3). Note: no nominal species were originally included in this genus by Bonelli (1810); the first ones were the three species listed by Panzer (1813: 54, 55).

Family-group name: ANCHOMENII Bonelli, 1810. Stem: Anchomen-.

Anchomma J.L. LeConte, 1858a: 63. Current status: valid genus in TENEBRIONIDAE: PIMELIINAE: STENOSINI: DICHILLINA.

Type species: Anchomma costatum J.L. LeConte, 1858, by monotypy.

Family-group name: ANCHOMMINI G.H. Horn, 1878. Stem: Anchommat-.

Anchonoderus Reiche, 1843a: 38. Current status: valid genus in CARABIDAE: HARPALINAE: LACHNOPHORINI: LACHNOPHORINA.

Type species: Platynus elegans Brullé, 1837, by original designation.

Family-group name: ANCHONODÉRIDES Lacordaire, 1854. Stem: Anchonoder-.

Anchonus Schönherr, 1825: 584. Current status: valid genus in CURCULIONIDAE: MOLYTINAE: ANCHONINI.

Type species: Curculio suillus Fabricius, 1792, by original designation.

Family-group name: ANCHONIDAE Imhoff, 1856. Stem: Anchon-.

Anchytarsus Guérin-Méneville, 1843a: 194. Current status: valid genus in PTILODACTYLIDAE: ANCHYTARSINAE.

Type species: Anchytarsus ater Guérin-Méneville, 1849 (= Atopa bicolor Mel­sheimer, 1845), by subsequent monotypy (Guérin-Méneville 1849 [no 15]: 1).

Family-group name: ANCHYTARSINI Champion, 1897. Stem: Anchytars-. Note: an application to the Commission is needed to conserve this family-group name as valid over COLOBODERINAE Erichson, 1847 (see Bouchard et al. 2011: 297).

Ancistria Erichson, 1845a: 305. Current status: valid genus in PASSANDRIDAE: PASSANDRINAE.

Type species: Colydium retusum Fabricius, 1801, by monotypy.

Family-group name: ANCISTRIINAE Sharp, 1899. Stem: Ancistri-.

Ancistrotus Audinet-Serville, 1832: 135. Current status: valid genus in CERAMBYCIDAE: PRIONINAE: MACRODONTIINI.

Type species: Ancistrotus hamaticollis Audinet-Serville, 1832 (= Prionus uncinatus Klug, 1825), by monotypy.

Family-group name: ANCISTROTIDES Lacordaire, 1868. Stem: Ancistrot-.

Ancita J. Thomson, 1864: 63. Current status: junior synonym of Hebecerus Dejean, 1835 in CERAMBYCIDAE: LAMIINAE: ANCITINI. Note: Hebecerus Dejean, 1835 has priority over the name Ancita J. Thomson, 1864, which has been used as valid in recent publications; we believe that a request should be submitted to the Commission to retain Ancita (and the tribe ANCITINI, instead of HEBESECIDINI Pascoe, 1871) as valid (see Bousquet and Bouchard 2013a: 83; Bouchard and Bousquet 2020a: 107).

Type species: Ancita crossotoides J. Thomson, 1864 (= Hebecerus niphonoides Pascoe, 1863), by original designation.

Family-group name: ANCITINI Aurivillius, 1917. Stem: Ancit-.

Ancylocera Audinet-Serville, 1834b: 107. Current status: valid genus in CERAMBYCIDAE: CERAMBYCINAE: TRACHYDERINI: ANCYLOCERINA.

Type species: Cerambyx cardinalis Dalman, 1823, by monotypy.

Family-group name: ANCYLOCERITAE J. Thomson, 1864. Stem: Ancylocer-.

Ancylocheira Eschscholtz, 1829b: 9. Current status: valid subgenus of Buprestis Linnaeus, 1758 in BUPRESTIDAE: BUPRESTINAE: BUPRESTINI: BUPRESTINA. Note: the original spelling was Ancylochira but Ancylocheira, an incorrect subsequent spelling, is in prevailing usage, attributed to the publication of the original spelling, and therefore deemed to be the correct original spelling (ICZN 1999a, Article 33.3.1).

Type species: Buprestis rustica Linnaeus, 1758, by subsequent designation (Westwood 1838c: 24).

Family-group name: ANCYLOCHIRINA Jakobson, 1913. Stem: Ancylocheir-.

Ancylocnemis G.A.K. Marshall, 1920: 387. Current status: valid genus in CURCULIONIDAE: CURCULIONINAE: ANCYLOCNEMIDINI.

Type species: Ancylocnemis fasciculata G.A.K. Marshall, 1920, by original designation.

Family-group name: ANCYLOCNEMINI Voss, 1962. Stem: Ancylocnemid-.

Ancylonotus Dejean, 1835: 335. Current status: valid genus in CERAMBYCIDAE: LAMIINAE: ANCYLONOTINI.

Type species: Lamia tribulus Fabricius, 1775, by monotypy.

Family-group name: ANCYLONOTIDES Lacordaire, 1869. Stem: Ancylonot-.

Ancylopoma Pascoe, 1871d: 354. Current status: valid genus in TENEBRIONIDAE incertae sedis.

Type species: Ancylopoma punctigerum Pascoe, 1871, by monotypy.

Family-group name: ANCYLOPOMINAE Pascoe, 1871. Stem: Ancylopomat-.

Ancyrona Reitter, 1876c: 51. Current status: valid genus in LOPHOCATERIDAE.

Type species: Ancyrona lewisi Reitter, 1876, by subsequent designation (Kolibáč 1993: 16).

Family-group name: ANCYRONINI Kolibáč, 2006. Stem: Ancyron-.

Ancyronyx Erichson, 1847a: 522. Current status: valid genus in ELMIDAE: ELMINAE: ANCYRONYCHINI.

Type species: Macronychus variegatus Germar, 1823, by monotypy.

Family-group name: ANCYRONYCHINI Ganglbauer, 1904. Stem: Ancyronych-.

Andotypus Spangler, 1979: 303. Current status: valid genus in HYDROPHILIDAE: CYLOMINAE.

Type species: Andotypus ashworthi Spangler, 1979, by original designation.

Family-group name: ANDOTYPINI Hansen, 1991. Stem: Andotyp-.

Androlyperus Crotch, 1873b: 55. Current status: valid genus in CHRYSOMELIDAE: GALERUCINAE: LUPERINI.

Type species: Androlyperus fulvus Crotch, 1873, by monotypy.

Family-group name: ANDROLYPERITES G.H. Horn, 1893. Stem: Androlyper-.

Androzelma Dostal, 1993: 117. Current status: valid genus in CARABIDAE: SCARITINAE: SALCEDIINI: ANDROZELMINA.

Type species: Androzelma gigas Dostal, 1993, by original designation.

Family-group name: ANDROZELMINA R.T. Bell, 1998. Stem: Androzelm-.

Anelastes W. Kirby, 1819: 384. Current status: valid genus in EUCNEMIDAE: PHYLLOCERINAE: ANELASTINI.

Type species: Anelastes drurii W. Kirby, 1819, by monotypy.

Family-group name: ANELASTINI Reitter, 1911. Stem: Anelast-.

Anelastidius Jacquelin du Val, 1859c: 117, 121. Current status: valid genus in EUCNEMIDAE: MACRAULACINAE: ANELASTIDINI.

Type species: Anelastidius ineditus Jacquelin du Val, 1859 (= Eucnemis feisthameli Graells, 1847), by original designation.

Family-group name: ANELASTIDINI Muona, 1993. Stem: Anelastid-.

Anemadus Reitter, 1884b: 38, 58. Current status: valid genus in LEIODIDAE: CHOLEVINAE: ANEMADINI: ANEMADINA.

Type species: Catops strigosus Kraatz, 1852, by subsequent designation (Jeannel 1922: 41).

Family-group name: ANEMADINA Hatch, 1927. Stem: Anemad-.

Anepius Blackburn, 1902: 29. Current status: valid genus in STAPHYLINIDAE: PROTEININAE: ANEPIINI.

Type species: Anepius koebelei Blackburn, 1902, by subsequent designation (R. Lucas 1920: 96).

Family-group name: ANEPIINI Steel, 1966. Stem: Anepi-.

Anepsius J.L. LeConte, 1851a: 147. Current status: valid genus in TENEBRIONIDAE: PIMELIINAE: ANEPSIINI.

Type species: Anepsius delicatulus J.L. LeConte, 1851, by monotypy.

Family-group name: ANEPSIINI J.L. LeConte, 1862. Stem: Anepsi-.

Angianus Sharp, 1919: 151. Current status: junior synonym of Vanapa Pouillaude, 1915 in CURCULIONIDAE: MOLYTINAE: PISSODINI: ORTHORHININA.

Type species: Angianus pratti Sharp, 1919 (= Vanapa oberthuri Pouillaude, 1915), by monotypy.

Family-group name: ANGIANIDES Sharp, 1919. Stem: Angian-.

Anilara Saunders, 1868: 19. Current status: valid genus in BUPRESTIDAE: BUPRESTINAE: CURIDINI: ANILARINA.

Type species: Anthaxia adelaidae Hope, 1846, by monotypy.

Family-group name: ANILARINI Bílý, 2000. Stem: Anilar-.

Anillus Jacquelin du Val, 1851: lxxiii. Current status: valid genus in CARABIDAE: TRECHINAE: ANILLINI.

Type species: Anillus coecus Jacquelin du Val, 1851, by monotypy.

Family-group name: ANILLINI Jeannel, 1937. Stem: Anill-.

Anisarthron Dejean, 1835: 331. Current status: valid genus in CERAMBYCIDAE: SPONDYLIDINAE: ANISARTHRINI.

Type species: Cerambyx barbipes Schrank, 1781, by monotypy.

Family-group name: ANISARTHRONINI Mamaev & Danilevsky, 1973. Stem: Anisarthr-.

Anischia Fleutiaux, 1896: 300. Current status: valid genus in EUCNEMIDAE: ANISCHIINAE.

Type species: Anischia boliviana Fleutiaux, 1896, by subsequent designation (Fleutiaux 1936: 292).

Family-group name: ANISCHINAE Fleutiaux, 1936. Stem: Anischi-.

Anisocerus Faldermann, 1837: 39. Current status: invalid senior synonym of Ceratanisus Gemminger, 1870 in TENEBRIONIDAE: PIMELIINAE: CERATANISINI. Note: junior homonym of Anisocerus Lacordaire, 1830 [Coleoptera: CERAMBYCIDAE].

Type species: Anisocerus tristis Faldermann, 1837, by monotypy.

Family-group name: ANISOCERINI Reitter, 1906. Stem: Anisocer-. Note: junior homonym of ANISOCERINI J. Thomson, 1860 (type genus Anisocerus Lacordaire, 1830) in Coleoptera: CERAMBYCIDAE; family-group name permanently invalid (ICZN 1999a, Article 39).

Anisocerus Lacordaire, 1830b: 181. Current status: valid genus in CERAMBYCIDAE: LAMIINAE: ANISOCERINI.

Type species: Anisocerus penicillatus Lacordaire, 1830 (= Lamia scopifera Germar, 1823), by monotypy.

Family-group name: ANISOCERITAE J. Thomson, 1860. Stem: Anisocer-.

Anisodactylus Dejean, 1829: 132. Current status: valid genus in CARABIDAE: HARPALINAE: HARPALINI: ANISODACTYLINA.

Type species: Carabus binotatus Fabricius, 1787, by subsequent designation (Westwood 1838c: 4).

Family-group name: ANISODACTYLIDES Lacordaire, 1854. Stem: Anisodactyl-. Note: nomen protectum (see Bouchard et al. 2011: 128).

Anisodera Chevrolat in Dejean, 1836a: 363. Current status: valid genus in CHRYSOMELIDAE: CASSIDINAE: ANISODERINI.

Type species: Alurnus ferrugineus Fabricius, 1801, by monotypy.

Family-group name: ANISODÉRITES Chapuis, 1875. Stem: Anisoder-.

Anisolemnia Crotch, 1874a: 146. Current status: valid genus in COCCINELLIDAE: COCCINELLINAE: COCCINELLINI.

Type species: Anisolemnia complicata Crotch, 1874, by original designation.

Family-group name: ANISOLEMNIINA Mader, 1954. Stem: Anisolemni-.

Anisolinus Sharp, 1889a: 113. Current status: valid genus in STAPHYLINIDAE: STAPHYLININAE: STAPHYLININI: ANISOLININA.

Type species: Anisolinus picticornis Sharp, 1889, by subsequent designation (R. Lucas 1920: 98).

Family-group name: ANISOLININA Hayashi, 1993. Stem: Anisolin-.

Anisomeria Brinck, 1943: 140. Current status: junior synonym of Rhantus Dejean, 1833 in DYTISCIDAE: COLYMBETINAE. Note: new replacement name for Anisomera Brullé, 1834.

Type species: Anisomera bistriata Brullé, 1834 (= Rhantus selkirki Jäch, Balke & Michat, 2014), by monotypy (type fixation for Anisomera Brullé).

Family-group name: ANISOMERIINI Brinck, 1948. Stem: Anisomeri-.

Anisomerinus Voss, 1933: 53. Current status: valid genus in ATTELABIDAE: RHYNCHITINAE: RHYNCHITITAE: RHYNCHITINI: TEMNOCERINA.

Type species: Anisomerinus proteae Voss, 1933, by original designation.

Family-group name: ANISOMERININA Legalov, 2003. Stem: Anisomerin-.

Anisomerus O. Schwarz, 1897: 13. Current status: valid genus in ELATERIDAE: ELATERINAE: ANISOMERINI.

Type species: Dicronychus bipectinatus O. Schwarz, 1896, by subsequent designation (Hyslop 1921: 627).

Family-group name: ANISOMERINI C. Girard, 2019: 413. Stem: Anisomer-. Note: junior homonym of ANISOMERINA Osten-Sacken, 1860 (type genus Anisomera Meigen, 1818) in Diptera.

Anisonychus Voss, 1927: 41. Current status: valid genus in ATTELABIDAE: APODERINAE: APODERINI.

Type species: Apoderus interstitialis Voss, 1922, by subsequent designation (Legalov 2003: 555).

Family-group name: ANISONYCHINA Legalov, 2003. Stem: Anisonych-. Note: junior homonym of ANISONYCHIDAE Burmeister, 1844 (type genus Anisonyx Latreille, 1806) in Coleoptera: SCARABAEIDAE; Burmeister’s name is currently considered a junior synonym.

Anisonyx Latreille, 1806: 119. Current status: valid genus in SCARABAEIDAE: MELOLONTHINAE: HOPLIINI: LEPITHRICHINA.

Type species: Melolontha crinita Fabricius, 1777 (= Scarabaeus longipes Linnaeus, 1764), by subsequent designation (Latreille 1810: 428).

Family-group name: ANISONYCHIDAE Burmeister, 1844. Stem: Anisonych-. Note: senior homonym of ANISONYCHINI Legalov, 2003 (type genus Anisonychus Voss, 1927) in Coleoptera: ATTELABIDAE.

Anisoplia Schönherr, 1817: 186. Current status: valid genus in SCARABAEIDAE: RUTELINAE: ANOMALINI: ANISOPLIINA.

Type species [suggested]: Scarabaeus agricola Poda von Neuhaus, 1761, by subsequent designation (S.I. Medvedev 1949: 239). Note: the valid type species fixation is that of Curtis (1834: pl. 526), who selected Scarabaeus horticola Linnaeus, 1758, but that is the type species of the genus Phyllopertha Stephens, 1830; a subsequent designation based on an originally included nominal species is that of Westwood (1838c: 24), who selected Melolontha austriaca Herbst, 1783, but that is the type species of the subgenus Autanisoplia S.I. Medvedev 1949, not the nominotypical subgenus; an application to the Commission is necessary to fix Scarabaeus agricola Poda von Neuhaus, 1761 as type species of Anisoplia Schönherr, 1817 (see Branco 2007: 5, 7).

Family-group name: ANISOPLIADAE Burmeister, 1844. Stem: Anisopli-.

Anisosphaera Tömösváry, 1883: 128. Current status: junior synonym of Cephennium P.W.J. Müller, 1822 in STAPHYLINIDAE: SCYDMAENINAE: CEPHENNIITAE: CEPHENNIINI.

Type species: Anisosphaera problematica Tömösváry, 1883 (= Cephennium majus Reitter, 1882), by monotypy. Note: Tömösváry’s species was described from larval stages as a Thysanura.

Family-group name: ANISOSPHAERIDAE Tömösváry, 1883. Stem: Anisosphaer-.

Anisosticta Chevrolat in Dejean, 1836a: 432. Current status: valid genus in COCCINELLIDAE: COCCINELLINAE: COCCINELLINI.

Type species: Coccinella novemdecimpunctata Linnaeus, 1758, by subsequent designation (Crotch 1874a: 93).

Family-group name: ANISOSTICTINA Jakobson, 1915. Stem: Anisostict-.

Anisotarsus Chaudoir, 1837: 41. Current status: valid subgenus of Notiobia Perty, 1830 in CARABIDAE: HARPALINAE: HARPALINI: ANISODACTYLINA.

Type species: Anisotarsus brevicollis Chaudoir, 1837, by subsequent designation (Casey 1914: 209).

Family-group name: ANISOTARSI Csiki, 1932. Stem: Anisotars-.

Anisotoma Panzer, 1796: no 8. Current status: valid genus in LEIODIDAE: LEIODINAE: ANISOTOMINI.

Type species [suggested]: Volvoxis glabra Kugelann, 1794, by subsequent designation (Hatch 1929: 50). Note: see Peck et al. (2020: 69) for interpretation of the type species; the valid type species fixation is that of C.G. Thomson (1859: 58), who selected Anisotoma picea Panzer, 1796, but that species is currently included in the genus Leiodes Latreille, 1797; an application to the Commission is necessary to fix Volvoxis glabra Kugelann, 1794 as the type species of Anisotoma Panzer, 1796.

Family-group name: ANISOTOMIDAE Horaninow, 1834. Stem: Anisotom-.

Anisoxiella Nikitsky, 1989: 42. Current status: valid genus in MELANDRYIDAE: MELANDRYINAE: ANISOXIELLINI.

Type species: Anisoxya ocularis Nomura & Katô, 1959, by original designation.

Family-group name: ANISOXIELLINI Nikitsky, 2007. Stem: Anisoxiell-.

Anobium Fabricius, 1775: 62. Current status: valid genus in PTINIDAE: ANOBIINAE. Note: name placed on the Official List of Generic Names in Zoology (ICZN 1976).

Type species: Ptinus punctatus DeGeer, 1774, by plenary power (see ICZN 1976). Note: name placed on the Official List of Specific Names in Zoology (ICZN 1976).

Family-group name: ANOBIUMEDAE Fleming, 1821. Stem: Anobi-.

Anochilia Burmeister, 1842: 558. Current status: valid genus in SCARABAEIDAE: CETONIINAE: STENOTARSIINI: ANOCHILIINA.

Type species: Cetonia laevigata Gory & Percheron, 1835, by subsequent designation (Sakai and Nagai 1998: 175).

Family-group name: ANOCHILIINA Krikken, 1984. Stem: Anochili-.

Anomala Leach in Samouelle, 1819: 191. Current status: valid genus in SCARABAEIDAE: RUTELINAE: ANOMALINI. Note: the senior homonym Anomala von Block, 1799 [Hymenoptera], and all other uses of the name Anomala prior to Anomala Leach in Samouelle, 1819, were suppressed for the purposes of the Principle of Priority and the Principle of Homonymy by the Commission and Anomala Leach in Samouelle, 1819 was placed on the Official List of Generic Names in Zoology (ICZN 1989c, as Anomala Samouelle, 1819).

Type species: Melolontha frischii Fabricius, 1775 (= Scarabaeus dubius Scopoli, 1763), by monotypy (see ICZN 1989c). Note: Melolontha frischii Fabricius, 1775 was placed on the Official List of Specific Names in Zoology (ICZN 1989c).

Family-group name: ANOMALIDAE Streubel, 1839. Stem: Anomal-. Note: nomen protectum (see A.B.T. Smith 2006: 173).

Anomalipus Guérin-Méneville, 1831b: pl. 29. Current status: valid genus in TENEBRIONIDAE: BLAPTINAE: PLATYNOTINI: PLATYNOTINA. Note: nomen protectum (see Bouchard and Bousquet 2020a: 101); the original spelling was Anomalipes but Anomalipus, an unjustified emendation by Lacordaire (1859a: 257), is in prevailing usage, attributed to the publication of the original spelling, and therefore deemed to be a justified emendation and the correct original spelling (ICZN 1999a, Article 33.2.3.1).

Type species: Blaps dentipes Fabricius, 1794, by monotypy.

Family-group name: ANOMALIPINA C. Koch, 1954. Stem: Anomalipod-.

Anommatus Wesmael, 1835: 338. Current status: valid genus in TEREDIDAE: ANOMMATINAE.

Type species: Anommatus terricola Wesmael, 1835 (= Lyctus duodecimstriatus P.W.J. Müller, 1821), by original designation.

Family-group name: ANOMMATINI Ganglbauer, 1899. Stem: Anommat-.

Anomophthalmus Fairmaire, 1884: 498. Current status: valid genus in CURCULIONIDAE: ENTIMINAE: ENTIMITAE: ANOMOPHTHALMINI.

Type species: Anomophthalmus insolitus Fairmaire, 1884, by monotypy.

Family-group name: ANOMOPHTHALMINA Morrone, 1998. Stem: Anomophthalm-.

Anomotarus Chaudoir, 1875: 48. Current status: valid genus in CARABIDAE: HARPALINAE: LEBIINI: CALLEIDINA.

Type species: Anomotarus olivaceus Chaudoir, 1875 (= Cymindis illawarrae W.J. MacLeay, 1873), by monotypy.

Family-group name: ANOMOTARINA Habu, 1967. Stem: Anomotar-.

Anopidium Jeannel & Paulian, 1945: 63. Current status: valid genus in TENEBRIONIDAE: DIAPERINAE: GNATHIDIINI: ANOPIDIINA.

Type species: Anopidium elgonicum Jeannel & Paulian, 1945, by original designation.

Family-group name: ANOPIDIINI Jeannel & Paulian, 1945. Stem: Anopidi-.

Anoploderma Guérin-Méneville, 1840a: 276. Current status: valid genus in VESPERIDAE: ANOPLODERMATINAE: ANOPLODERMATINI.

Type species: Anoploderma bicolor Guérin-Méneville, 1840, by monotypy.

Family-group name: ANOPLODERMIENS Guérin-Méneville, 1840. Stem: Anoplodermat-.

Anoplogenius Chaudoir, 1852: 88. Current status: junior synonym of Loxoncus Schmidt-Göbel, 1846 in CARABIDAE: HARPALINAE: HARPALINI: STENOLOPHINA.

Type species: Stenolophus alacer Dejean, 1831, by subsequent designation (Lacordaire 1854: 304).

Family-group name: ANOPLOGENII Schauberger, 1937. Stem: Anoplogeni-.

Anoplognathus Leach, 1815b: [Indexes]. Current status: valid genus in SCARABAEIDAE: RUTELINAE: ANOPLOGNATHINI: ANOPLOGNATHINA. Note: the name listed on page 43 is Rutela but it was changed in both the unpaginated Index Systematicus and the General Index to Anoplognathus.

Type species: Melolontha viridiaenea Donovan, 1805, by subsequent designation (Carne 1957a: 93).

Family-group name: ANOPLOGNATHIDAE W.S. MacLeay, 1819. Stem: Anoplognath-.

Anoplus Germar, 1820: 195. Current status: valid genus in CURCULIONIDAE: MOLYTINAE: ANOPLINI.

Type species: Curculio plantaris Naezén, 1794, by monotypy.

Family-group name: ANOPLINI Bedel, 1883. Stem: Anopl-.

Anopsilus Kirsch, 1870: 218. Current status: valid genus in CURCULIONIDAE: BARIDINAE: BARIDINI: BARIDINA.

Type species: Anopsilus bonvouloirii Kirsch, 1870, by monotypy.

Family-group name: ANOPSILINI Bondar, 1942. Stem: Anopsil-.

Antarctia Dejean, 1828: 525. Current status: invalid senior synonym of Metius Curtis, 1838 in CARABIDAE: HARPALINAE: PTEROSTICHINI: METIINA. Note: junior homonym of Antarctia Hübner, 1820 [Lepidoptera].

Type species: Carabus carnifex Fabricius, 1775, by subsequent designation (Hope 1838a: 86).

Family-group name: ANTARCTIIDES Lacordaire, 1854. Stem: Antarcti-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

Antennolycus Bocáková & Bocák, 1999: 105. Current status: valid genus in LYCIDAE: LYROPAEINAE: ANTENNOLYCINI.

Type species: Antennolycus constrictus Bocáková & Bocák, 1999, by original designation.

Family-group name: ANTENNOLYCINI Bocák & Bocáková, 2008. Stem: Antennolyc-.

Anthaxia Eschscholtz, 1829b: 9. Current status: valid genus in BUPRESTIDAE: BUPRESTINAE: ANTHAXIINI. Note: name placed on the Official List of Generic Names in Zoology (ICZN 2002c).

Type species: Buprestis nitida Rossi, 1792 (= Buprestis fulgurans Schrank, 1789), by plenary power (see ICZN 2002c). Note: Buprestis fulgurans Schrank, 1789 was placed on the Official List of Specific Names in Zoology (ICZN 2002c).

Family-group name: ANTHAXIDAE Gory & Laporte, 1839. Stem: Anthaxi-.

Anthaxomorphus Deyrolle, 1865: 219. Current status: valid genus in BUPRESTIDAE: AGRILINAE: APHANISTICINI: ANTHAXOMORPHINA.

Type species: Anthaxomorphus papuanus Deyrolle, 1865, by subsequent designation (Cobos 1979: 427).

Family-group name: ANTHAXOMORPHINA Holyński, 1993. Stem: Anthaxomorph-.

Antherophagus Dejean, 1821: 45. Current status: valid genus in CRYPTOPHAGIDAE: CRYPTOPHAGINAE: CRYPTOPHAGINI.

Type species: Tenebrio pallens Linnaeus, 1758, by subsequent designation (Curtis 1835: pl. 546). Note: as noted by C. Johnson et al. (2007: 514) the type species listed by Curtis (1835) was possibly misidentified for Antherophagus similis Curtis, 1835; however, in the absence of clear evidence to this effect, we prefer to recognize the species actually cited by Curtis (1835: pl. 546); both species are currently placed in the genus Antherophagus Dejean.

Family-group name: ANTHEROPHAGI J.L. LeConte, 1861. Stem: Antherophag-.

Anthia Weber, 1801: 17. Current status: valid genus in CARABIDAE: HARPALINAE: ANTHIINI.

Type species: Carabus sexguttatus Fabricius, 1775, by subsequent designation (Latreille 1810: 426).

Family-group name: ANTHIES Bonelli, 1813. Stem: Anthi-.

Anthicoclerus Schenkling, 1906: 264. Current status: valid genus in CLERIDAE: CLERINAE: CLERITAE: HYDNOCERINI: ANTHICOCLERINA.

Type species: Clerus anthicoides A. White, 1849, by monotypy.

Family-group name: ANTHICOCLERINAE Opitz, 2010. Stem: Anthicocler-.

Anthicoxenus Fairmaire & Germain, 1860a: 2. Current status: valid genus in MELOIDAE: ELETICINAE: SPASTICINI: ANTHICOXENINA.

Type species: Anthicoxenus nigroplagiatus Fairmaire & Germain, 1860, by monotypy.

Family-group name: ANTHICOXENINA Selander, 1966. Stem: Anthicoxen-.

Anthicus Paykull, 1798: 253. Current status: valid genus in ANTHICIDAE: ANTHICINAE: ANTHICINI. Note: name placed on the Official List of Generic Names in Zoology (ICZN 2016).

Type species: Meloe antherinus Linnaeus, 1761, by subsequent designation (Westwood 1830a: 59; see ICZN 2016, as Meloe antherinus Linnaeus, 1760). Note: name placed on the Official List of Specific Names in Zoology (ICZN 2016).

Family-group name: ANTHICITES Latreille, 1819. Stem: Anthic-.

Anthomanes Gistel, 1848b: xi, xiv. Current status: junior synonym of Pyrochroa Geoffroy, 1762 in PYROCHROIDAE: PYROCHROINAE. Note: unnecessary new replacement name for Pyrochroa Geoffroy, 1762.

Type species: Cantharis coccinea Linnaeus, 1761, by subsequent designation (Westwood 1838c: 30; see ICZN 1994a) (type fixation for Pyrochroa Geoffroy).

Family-group name: ANTHOMANISIDAE Gistel, 1848. Stem: Anthoman-.

Anthonomus Germar, 1817b: 340. Current status: valid genus in CURCULIONIDAE: CURCULIONINAE: ANTHONOMINI.

Type species: Curculio avarus Fabricius, 1798 (= Curculio pedicularius Linnaeus, 1758), by subsequent designation (Kojima and Morimoto 1994: 148).

Family-group name: ANTHONOMINA C.G. Thomson, 1859. Stem: Anthonom-.

Anthophagus Gravenhorst, 1802: 120. Current status: valid genus in STAPHYLINIDAE: OMALIINAE: ANTHOPHAGINI. Note: name placed on the Official List of Generic Names in Zoology (ICZN 2004e).

Type species: Staphylinus alpinus Fabricius, 1792 (= Staphylinus alpinus Paykull, 1790), by subsequent designation (C.G. Thomson 1859: 48; see ICZN 2004e). Note: name placed on the Official List of Specific Names in Zoology (ICZN 2004e).

Family-group name: ANTHOPHAGIDES C.G. Thomson, 1859. Stem: Anthophag-.

Anthotribus Gistel, 1856: 260, 375. Current status: junior synonym of Platystomos D.H. Schneider, 1791 in ANTHRIBIDAE: ANTHRIBINAE: PLATYSTOMINI. Note: junior homonym of Anthotribus Hoffmann, 1803 (unjustified emendation of Anthribus Geoffroy, 1862) [Coleoptera: ANTHRIBIDAE].

Type species: Curculio albinus Linnaeus, 1758, by monotypy.

Family-group name: ANTHOTRIBIDAE Gistel, 1856. Stem: Anthotrib-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

Anthracias Dejean, 1834: 205. Current status: junior synonym of Cryphaeus Klug, 1833 in TENEBRIONIDAE: TENEBRIONINAE: TOXICINI: TOXICINA.

Type species: Uloma cornutum Fischer, 1823, by monotypy.

Family-group name: ANTHRACIINI Houlbert, 1922. Stem: Anthraci-.

Anthracus Motschulsky, 1850: vii. Current status: valid genus-group name in CARABIDAE: HARPALINAE: HARPALINI: STENOLOPHINA.

Type species: Carabus consputus Duftschmid, 1812, by original designation.

Family-group name: ANTHRACINI Schuler, 1970. Stem: Anthrac-.

Anthrenocerus Arrow, 1915: 443. Current status: junior synonym of Eurhopalus Solier, 1849 in DERMESTIDAE: MEGATOMINAE: MEGATOMINI: TROGODERMINA. Note: generic synonymy established by Zhou et al. (2022: 19).

Type species: Anthrenus australis Hope, 1843, by original designation.

Family-group name: ANTHRENOCERINA Háva, 2015: 1. Stem: Anthrenocer-.

Anthrenus Geoffroy, 1762: 113. Current status: valid genus in DERMESTIDAE: MEGATOMINAE: ANTHRENINI. Note: name placed on the Official List of Generic Names in Zoology (ICZN 1994a).

Type species: Dermestes scrophulariae Linnaeus, 1758, by subsequent designation (Mroczkowski 1968: 139; see ICZN 1994a). Note: name placed on the Official List of Specific Names in Zoology (ICZN 1994a).

Family-group name: ANTHRENIDAE Gistel, 1848. Stem: Anthren-.

Anthribus Geoffroy, 1762: 306. Current status: valid genus in ANTHRIBIDAE: ANTHRIBINAE: ANTHRIBINI. Note: name placed on the Official List of Generic Names in Zoology (ICZN 1994a).

Type species: Anthribus fasciatus Forster, 1770, by subsequent designation (Jordan 1931: 285; see ICZN 1994a). Note: name placed on the Official List of Specific Names in Zoology (ICZN 1994a).

Family-group name: ANTHRIBIDES Billberg, 1820. Stem: Anthrib-.

Anthroherpon Reitter, 1889e: 294. Current status: valid genus in LEIODIDAE: CHOLEVINAE: LEPTODIRINI: ANTHROHERPONINA.

Type species: Leptoderus cylindricollis Apfelbeck, 1889, by subsequent designation (Jeannel 1910: 26).

Family-group name: ANTROHERPONA Jeannel, 1910. Stem: Anthroherpon-.

Anthypna Eschscholtz, 1818: 472. Current status: valid genus in GLAPHYRIDAE: AMPHICOMINAE.

Type species [suggested]: Melolontha cyanipennis Fabricius, 1801 (= Melolontha meles Fabricius, 1792), by subsequent designation (Chapin 1938: 80). Note: the valid type species fixation is that of Crotch (1870b: 231), who selected Melolontha ursus Fabricius, 1775, but that species is currently in the genus Anisonyx Latreille, 1805 in the tribe HOPLIINI (family SCARABAEIDAE); an application to the Commission is necessary to fix Melolontha cyanipennis Fabricius, 1801 as the type species of Anthypna Eschscholtz, 1818 in order to conserve the current usage of this genus-group name.

Family-group name: ANTHYPNIDAE Chalande, 1884. Stem: Anthypn-.

Anticheira Eschscholtz, 1818: 475. Current status: valid genus in SCARABAEIDAE: RUTELINAE: RUTELINI: RUTELINA.

Type species [suggested]: Scarabaeus virens Drury, 1773, by subsequent designation (Ohaus 1934: 146). Note: the valid type species fixation is that of Duponchel (1841a: 610), who selected Scarabaeus tetradactylus Linnaeus, 1771, but that species is currently placed in the genus Macraspis W.S. MacLeay, 1819; an application to the Commission is necessary to fix Scarabaeus virens Drury, 1773 as the type species of Anticheira Eschscholtz, 1818.

Family-group name: ANTICHIRIDES Lacordaire, 1855. Stem: Anticheir-.

†Antigracilus Zhou, Caterino, Ren & Ślipiński, 2020: 537. Current status: valid genus in HISTERIDAE: †ANTIGRACILINAE. Note: Lower Cretaceous (China).

Type species: †Antigracilus costatus Zhou, Caterino, Ren & Ślipiński, 2020, by original designation.

Family-group name: †ANTIGRACILINAE Zhou, Caterino, Ren & Ślipiński, 2020: 537. Stem: Antigracil-.

Antillusa Pace, 2012: 59. Current status: valid genus in STAPHYLINIDAE: ALEOCHARINAE: ANTILLUSINI.

Type species: Antillusa dominicana Pace, 2012, by original designation.

Family-group name: ANTILLUSINI Pace, 2012: 59. Stem: Antillus-.

Antimerus Fauvel, 1878: 549, 550. Current status: valid genus in STAPHYLINIDAE: STAPHYLININAE: STAPHYLININI: ANTIMERINA.

Type species: Antimerus smaragdinus Fauvel, 1878, by monotypy.

Family-group name: ANTIMERINA Brunke, Żyła & Solodovnikov in Brunke et al., 2019: 181. Stem: Antimer-.

Antipha Baly, 1865a: 251. Current status: invalid senior synonym of Dercetina Gressitt & Kimoto, 1963 in CHRYSOMELIDAE: GALERUCINAE: HYLASPINI. Note: junior homonym of Antipha Walker, 1855 [Lepidoptera].

Type species: Antipha picipes Baly, 1865, by original designation.

Family-group name: ANTIPHITES Chapuis, 1875. Stem: Antiph-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

†Antirhelus Kirejtshuk in Kirejtshuk et al., 2023a: 22. Current status: valid genus in KATERETIDAE: †ANTIRHELINAE. Note: Eocene (Rovno amber).

Type species: †Heterhelus buzina Kupryjanowicz, Lyubarsky & Perkovsky, 2021, by original designation.

Family-group name: †ANTIRHELINAE Kirejtshuk in Kirejtshuk et al., 2023a: 21. Stem: Antirhel-.

Antliarhis Billberg, 1820: 40. Current status: valid genus in BRENTIDAE: APIONINAE: ANTLIARHINITAE.

Type species: Curculio haustellatus G.-A. Olivier, 1791 (= Curculio zamiae Thunberg, 1784), by monotypy.

Family-group name: ANTLIARHINIDES Schönherr, 1823. Stem: Antliarhin-.

Anypotactus Schönherr, 1840b: 299. Current status: valid genus in CURCULIONIDAE: ENTIMINAE: POLYDRUSITAE: ANYPOTACTINI.

Type species: Anypotactus exilis Boheman, 1840, by original designation.

Family-group name: ANYPOTACTINA Champion, 1911. Stem: Anypotact-.

Aonychus Schönherr, 1844: 387. Current status: valid genus in CURCULIONIDAE: BRACHYCERINAE: ERIRHININI.

Type species: Aonychus hopei Boheman, 1844, by original designation.

Family-group name: AONYCHINI Zimmerman, 1993. Stem: Aonych-. Note: junior homonym of AONYCHINI Sokolov, 1973 (type genus Aonyx Lesson, 1827) in Mammalia; because of the homonymy, the stem of the genus was changed to Aonychus- and the family group name to AONYCHUSINI by Zimmerman in Alonso-Zarazaga and Lyal (1999: 72) but cases involving homonymy of family-group names based on similar genus-group names are to be referred to the Commission for a ruling (ICZN 1985g, 1999a).

Apalochrus Erichson, 1840a: 50. Current status: valid genus in MELYRIDAE: MALACHIINAE: APALOCHRINI.

Type species: Apalochrus femoralis Erichson, 1840, by subsequent designation (C.G. Thomson 1859: 112).

Family-group name: APALOCHRAIRES Mulsant & Rey, 1867. Stem: Apalochr-.

Apalus Fabricius, 1775: 127. Current status: valid genus in MELOIDAE: NEMOGNATHINAE: NEMOGNATHINI: SITARINA.

Type species: Meloe bimaculatus Linnaeus, 1761, by monotypy.

Family-group name: APALIDES J.B. Parker & Böving, 1924. Stem: Apal-.

Apate Fabricius, 1775: 54. Current status: valid genus in BOSTRICHIDAE: BOSTRICHINAE: APATINI. Note: nomen protectum (see Borowski and Węgrzynowicz 2009: 190).

Type species: here fixed (under ICZN 1999a, Article 70.3.2) as Ligniperda terebrans Pallas, 1772, misidentified as Dermestes muricatus Linnaeus, 1767 (sensu Fabricius, 1775), in the subsequent designation by Gorham (1883a: 212).

Family-group name: APATIDES Billberg, 1820. Stem: Apat-.

Apatetica Westwood, 1848a: 86. Current status: valid genus in STAPHYLINIDAE: APATETICINAE.

Type species: Apatetica lebioides Westwood, 1848, by monotypy.

Family-group name: APATETICAE Fauvel, 1895. Stem: Apatetic-.

Apatides Casey, 1898: 66, 70. Current status: valid genus in BOSTRICHIDAE: BOSTRICHINAE: BOSTRICHINI.

Type species: Amphicerus fortis J.L. LeConte, 1866, by subsequent designation (Lesne 1900: 47).

Family-group name: APATIDINI Bradley, 1930. Stem: Apatid-.

Apatophysis Chevrolat, 1860a: 95. Current status: valid genus in CERAMBYCIDAE: CERAMBYCINAE: DORCASOMINI.

Type species: Apatophysis toxotoides Chevrolat, 1860 (= Polyarthron barbarum P.H. Lucas, 1858), by monotypy. Note: combined description of a new genus and a single new species (ICZN 1999a, Article 12.2.6).

Family-group name: APATOPHYSIDES Lacordaire, 1869. Stem: Apatophyse-.

Apenes J.L. LeConte, 1851a: 174. Current status: valid genus in CARABIDAE: HARPALINAE: LEBIINI: APENINA. NOTE: nomen protectum (see Ball and Shpeley 2009: 95).

Type species: Cymindis lucidula Dejean, 1831, by subsequent designation (Lacordaire 1854: 110).

Family-group name: APENINA Ball, 1983. Stem: Apen-.

Aphaenostemmus Peyerimhoff, 1914: 245. Current status: valid genus in STAPHYLINIDAE: OMALIINAE: APHAENOSTEMMINI.

Type species: Aphaenostemmus bordei Peyerimhoff, 1914, by monotypy.

Family-group name: APHAENOSTEMMINI Peyerimhoff, 1914. Stem: Aphaenostemm-.

Aphanasium Dejean, 1835: 322. Current status: valid genus in CERAMBYCIDAE: CERAMBYCINAE: APHANASIINI.

Type species: Callidium australe Boisduval, 1835, by monotypy.

Family-group name: APHANASIIDES Lacordaire, 1868. Stem: Aphanasi-.

Aphanisticus Latreille, 1810: 169. Current status: valid genus in BUPRESTIDAE: AGRILINAE: APHANISTICINI: APHANISTICINA.

Type species: Buprestis emarginata Fabricius, 1792 (= Buprestis emarginata G.-A. Olivier, 1790), by monotypy.

Family-group name: APHANISTICITES Jacquelin du Val, 1859. Stem: Aphanistic-.

Aphanocephalus Wollaston, 1873a: 278. Current status: valid genus in DISCOLOMATIDAE: APHANOCEPHALINAE.

Type species: Aphaenocephalus hemisphaericus Wollaston, 1873, by monotypy.

Family-group name: APHANOCEPHALIDAE Jakobson, 1904. Stem: Aphanocephal-.

Aphela Pascoe, 1865b: 416. Current status: valid genus in CURCULIONIDAE: CYCLOMINAE: NOTIOMIMETINI.

Type species: Aphela helopoides Pascoe, 1865, by monotypy.

Family-group name: APHELI Csiki, 1936. Stem: Aphel-.

Aphneope Pascoe, 1863b: 567. Current status: valid genus in CERAMBYCIDAE: CERAMBYCINAE: APHNEOPINI.

Type species: Aphneope sericata Pascoe, 1863, by monotypy.

Family-group name: APHNÉOPIDES Lacordaire, 1868. Stem: Aphneop-.

Aphodius Hellwig, 1798: 101. Current status: valid genus in SCARABAEIDAE: APHODIINAE: APHODIINI: APHODIINA. Note: see Alonso-Zarazaga and Krell (2011: 67) for comments on the authorship of this genus-group name.

Type species: Scarabaeus fimetarius Linnaeus, 1758, by subsequent designation (Latreille 1810: 428). Note: name placed on the Official List of Specific Names in Zoology (ICZN 2014b).

Family-group name: APHODIDA Leach, 1815. Stem: Aphodi-.

Aphrastus Schönherr in Say, 1832: 9. Current status: valid genus in CURCULIONIDAE: ENTIMINAE: OTIORHYNCHITAE: PHYLLOBIINI. Note: see Prena (2018: 331) for notes on the authorship of this genus-group name.

Type species: Aphrastus taeniatus Say, 1832, by monotypy. Note: combined description of a new genus and a single new species (ICZN 1999a, Article 12.2.6).

Family-group name: APHRASTI J.L. LeConte, 1874. Stem: Aphrast-.

Aphricus J.L. LeConte, 1853f: 501. Current status: valid genus in ELATERIDAE: CARDIOPHORINAE.

Type species: Aphricus californicus J.L. LeConte, 1853, by monotypy.

Family-group name: APHRICI J.L. LeConte, 1861. Stem: Aphric-.

Aphthona Chevrolat in Dejean, 1836a: 391. Current status: valid genus in CHRYSOMELIDAE: GALERUCINAE: ALTICINI.

Type species: Haltica cyparissiae Knoch, 1803, by subsequent designation (Maulik 1926: 366).

Family-group name: APHTHONITES Chapuis, 1875. Stem: Aphthon-.

Aphyllura Reitter in Brenske and Reitter, 1884: 97. Current status: valid genus in CURCULIONIDAE: COSSONINAE: RHYNCOLINI: PSEUDOMIMINA.

Type species: Aphyllura brenskei Reitter, 1884, by monotypy.

Family-group name: APHYLLURINA Voss, 1955. Stem: Aphyllur-.

†Aphytocerus Zherichin, 1977: 131. Current status: valid genus in CEROPHYTIDAE. Note: Upper Cretaceous (Siberia, Russia).

Type species: †Aphytocerus communis Zherichin, 1977, by original designation.

Family-group name: †APHYTOCERINAE Kirejtshuk in Kirejtshuk and Azar, 2013: 128. Stem: Aphytocer-.

Aphytopus Sharp, 1886: 375. Current status: valid genus in STAPHYLINIDAE: ALEOCHARINAE: OXYPODINI: APHYTOPODINA.

Type species: Aphytopus gracilis Sharp, 1886, by monotypy.

Family-group name: APHYTOPI Bernhauer & Scheerpeltz, 1926. Stem: Aphytopod-.

Apiomorphus Wagner, 1912c: 33. Current status: valid genus in BRENTIDAE: APIONINAE: TANAITAE: APIOMORPHINI.

Type species: Apiomorphus cyaneus Wagner, 1912 (= Apion eximium Beguin-Billecocq, 1910), by monotypy.

Family-group name: APIOMORPHINI Legalov, 2023c: 30. Stem: Apiomorph-. Note: APIOMORPHINI Legalov (2018c: 810) was published in a work that is nomenclaturally unavailable (ICZN 2012f, Article 8.5.3).

Apion Herbst, 1797: 100. Current status: valid genus in BRENTIDAE: APIONINAE: APIONITAE: APIONINI: APIONINA.

Type species: Curculio frumentarius Linnaeus, 1758, by subsequent designation (Latreille 1810: 430).

Family-group name: APIONIDES Schönherr, 1823. Stem: Apion-.

Apistus Agassiz, 1846: 29. Current status: invalid senior synonym of Rhopalocerus W. Redtenbacher, 1842 in ZOPHERIDAE: COLYDIINAE: RHOPALOCERINI. Note: unjustified emendation of Apeistus Motschulsky, 1840; junior homonym of Apistus Cuvier, 1829 [Pisces]; Apistus Agassiz, 1846 and Apeistus Motschulsky, 1840 were suppressed for the Principle of Priority and placed on the Official Index of Rejected and Invalid Names in Zoology (ICZN 1986b).

Type species: Monotoma rondanii A. Villa & G.B. Villa, 1833, by monotypy (type fixation for Spartycerus Motschulsky, 1837). Note: name placed on the Official List of Specific Names in Zoology (ICZN 1986b).

Family-group name: APISTINI Ganglbauer, 1899. Stem: Apist-. Note: family-group name permanently invalid (ICZN 1999a, Article 39); name placed on the Official List of Rejected and Invalid Family-Group Names in Zoology (see ICZN 1986b).

Aplastus J.L. LeConte, 1859a: 73. Current status: valid genus in ELATERIDAE: ELATERINAE: CEBRIONINI: APLASTINA. Note: nomen protectum (see Appendix 2).

Type species: Aplastus speratus J.L. LeConte, 1859, by monotypy.

Family-group name: APLASTINAE Stibick, 1979. Stem: Aplast-.

Aplemonus Schönherr, 1847: 5. Current status: valid genus in BRENTIDAE: APIONINAE: APIONITAE: APLEMONINI.

Type species: Aplemonus gibbipennis Schönherr, 1847, by original designation. Note: combined description of a new genus-group taxon and a single new nominal species (ICZN 1999a, Article 12.2.6).

Family-group name: APLEMONINI Kissinger, 1968. Stem: Aplemon-.

Aplocnemus Stephens, 1830: 311, 316. Current status: valid genus in RHADALIDAE: RHADALINAE.

Type species: Crioceris impressa Marsham, 1802, by original designation.

Family-group name: HAPLOCNÉMATES Mulsant & Rey, 1868. Stem: Aplocnem-.

Aploderus Stephens, 1833b: 273. Current status: valid genus in STAPHYLINIDAE: OXYTELINAE: THINOBIINI.

Type species: Staphylinus brachypterus Marsham, 1802 (= Oxytelus caelatus Gravenhorst, 1802), by subsequent monotypy (Stephens 1834: 321).

Family-group name: APLODERINI Ádám, 2001. Stem: Aploder-.

Aploglossa Guérin-Méneville, 1849 [no 33]: 1. Current status: valid genus in PTILODACTYLIDAE: APLOGLOSSINAE.

Type species: Aploglossa sallei Guérin-Méneville, 1849, by present designation.

Family-group name: HAPLOGLOSSINI Champion, 1897. Stem: Aplogloss-.

Aplonycha Boisduval, 1835: 193. Current status: valid genus in SCARABAEIDAE: SERICOIDINAE: LIPARETRINI. Note: originally proposed as a synonym of Melolontha Fabricius, 1775 and subsequently treated as valid before 1961 by several authors, including Duponchel and Chevrolat (1841: 16).

Type species: Melolontha obesa Boisduval, 1835, by subsequent designation (Duponchel and Chevrolat 1841: 16). Note: Britton (1986: 4) mentioned that the name-bearing type of Melolontha obesa has been lost and the species is unidentifiable based on the brief original description.

Family-group name: HAPLONYCHIDAE Burmeister, 1855. Stem: Aplonych-. Note: senior homonym of APLONYCHINI De Stefani, 1908 (type genus Aplonyx De Stefani, 1908) in Diptera.

Aplothorax G.R. Waterhouse, 1841a: 145. Current status: valid genus in CARABIDAE: CARABINAE: CARABINI: CALOSOMATINA.

Type species: Aplothorax burchellii G.R. Waterhouse, 1841, by monotypy. Note: combined description of a new genus-group taxon and a single new nominal species (ICZN 1999a, Article 12.2.6).

Family-group name: APLOTHORACINA Lapouge, 1927. Stem: Aplothorac-.

Apocellus Erichson, 1839b: 30. Current status: valid genus in STAPHYLINIDAE: OXYTELINAE: OXYTELINI.

Type species: Lathrobium sphaericolle Say, 1831, by subsequent designation (Duponchel and Chevrolat 1841: 22). Note: no species were originally included in this genus by Erichson (1839b); the first ones were the three species described by Erichson (1840b: 813–814).

Family-group name: APOCELLARIA Lynch Arribálzaga, 1884. Stem: Apocell-.

Apocrypha Eschscholtz, 1831: 13. Current status: valid genus in TENEBRIONIDAE: TENEBRIONINAE: APOCRYPHINI.

Type species: Apocrypha anthicoides Eschscholtz, 1831, by monotypy.

Family-group name: APOCRYPHIDES Lacordaire, 1859. Stem: Apocryph-.

Apodasya Pascoe, 1863a: 54. Current status: valid genus in CERAMBYCIDAE: LAMIINAE: DESMIPHORINI. Note: name placed on the Official List of Generic Names in Zoology (ICZN 2011d).

Type species: Apodasya pilosa Pascoe, 1863 (= Chaetosoma pilosum Chevrolat, 1843), by original designation (see ICZN 2011d). Note: Chaetosoma pilosum Chevrolat, 1843 was placed on the Official List of Specific Names in Zoology (ICZN 2011d).

Family-group name: APODASYIDES Lacordaire, 1872. Stem: Apodasy-.

Apoderiger Wasmann, 1897: 263. Current status: valid genus in STAPHYLINIDAE: PSELAPHINAE: CLAVIGERITAE: CLAVIGERINI.

Type species: Apoderiger cervinus Wasmann, 1897, by monotypy.

Family-group name: APODERIGERINI Jeannel, 1954. Stem: Apoderiger-.

Apoderus G.-A. Olivier, 1807: 12. Current status: valid genus in ATTELABIDAE: APODERINAE: APODERINI.

Type species: Attelabus coryli Linnaeus, 1758, by subsequent designation (Latreille 1810: 430).

Family-group name: APODERIDAE Jekel, 1860. Stem: Apoder-.

Apodytapion Wanat, 2021: 23. Current status: valid genus in BRENTIDAE: APIONINAE incertae sedis.

Type species: Apodytapion stepniewskii Wanat, 2021, by original designation.

Family-group name: APODYTAPIINI Wanat, 2021: 56. Stem: Apodytapi-.

Apogonia W. Kirby, 1819: 401. Current status: valid genus in SCARABAEIDAE: MELOLONTHINAE: DIPLOTAXINI.

Type species: Apogonia gemellata W. Kirby, 1819 (= Melolontha rauca Fabricius, 1781), by monotypy.

Family-group name: APOGONIITAE Blanchard, 1851. Stem: Apogoni-.

Apolecta Pascoe, 1859c: 431. Current status: valid genus in ANTHRIBIDAE: ANTHRIBINAE: APOLECTINI.

Type species: Mecocerus parvulus J. Thomson, 1858, by original designation.

Family-group name: APOLECTIDES Lacordaire, 1865. Stem: Apolect-. Note: senior homonym of APOLECTIDAE Jordan, 1923 (type genus Apolectus Cuvier, 1832) in Pisces; Jordan’s name is permanently invalid (based on a preoccupied type genus).

Apoleon Gorham, 1885a: 51. Current status: valid genus in BOSTRICHIDAE: DYSIDINAE.

Type species: Apoleon edax Gorham, 1885, by monotypy.

Family-group name: APOLEONINAE Gardner, 1933. Stem: Apoleont-.

Apolites Jacquelin du Val, 1861: 324. Current status: invalid senior synonym of Ceratanisus Gemminger, 1870 in TENEBRIONIDAE: PIMELIINAE: CERATANISINI. Note: junior homonym of Apolites Sundevall, 1835 [Aves].

Type species: Helops mucoreus Waltl, 1838, by monotypy.

Family-group name: APOLITINA Seidlitz, 1895. Stem: Apolit-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

Apomecyna Dejean, 1821: 108. Current status: valid genus in CERAMBYCIDAE: LAMIINAE: APOMECYNINI.

Type species: Saperda alboguttata Megerle, 1802 (= Lamia histrio Fabricius, 1792), by monotypy. Note: Saperda alboguttata Megerle, 1802 was placed on the Official List of Specific Names in Zoology (ICZN 1993a, as Saperda alboguttata Megerle, 1803).

Family-group name: APOMECYNITAE J. Thomson, 1860. Stem: Apomecyn-.

Apophylia Duponchel & Chevrolat, 1841: 31. Current status: valid genus in CHRYSOMELIDAE: GALERUCINAE: GALERUCINI. Note: this genus is often credited to J. Thomson (1858b: 221) in the literature but it was first made available by Duponchel and Chevrolat (1841: 31) when they provided a description of this new taxon; the three species associated with the name Apophylia in Duponchel and Chevrolat (1841: 31) are nomina nuda and the first available species associated with the name was established by J. Thomson (1858b: 221).

Type species: Apophylia chloroptera J. Thomson, 1858, by subsequent monotypy (J. Thomson 1858b: 221).

Family-group name: APOPHYLIITES Chapuis, 1875. Stem: Apophyli-.

†Apophisandra Molino-Olmedo, 2017: 170. Current status: valid genus in †APOPHISANDRIDAE. Note: mid-Cretaceous (Myanmar).

Type species: †Apophisandra ammytae Molino-Olmedo, 2017, by original designation. Note: combined description of a new genus and a single new species (ICZN 1999a, Article 13.4).

Family-group name: †APOPHISANDRINI Molino-Olmedo, 2017: 173. Stem: Apophisandr-.

Apostasimerus Schönherr, 1844: 101. Current status: valid genus in CURCULIONIDAE: BARIDINAE: APOSTASIMERINI: APOSTASIMERINA.

Type species: Apotasimerus serrirostris Boheman, 1844, by original designation.

Family-group name: APOSTASIMERIDES Schönherr, 1844. Stem: Apostasimer-.

†Apotomoura Bao, Walczyńska, Moody, B. Wang & Rust, 2018: 16. Current status: valid genus in †APOTOMOURIDAE. Note: mid-Cretaceous (Myanmar).

Type species: †Apotomoura fortiscrura Bao, Walczyńska, Moody, B. Wang & Rust, 2018, by monotypy.

Family-group name: †APOTOMOURIDAE Bao, Walczyńska, Moody, B. Wang & Rust, 2018: 16. Stem: Apotomour-.

Apotomus Illiger, 1807b: 348. Current status: valid genus in CARABIDAE: APOTOMINAE.

Type species: Scarites rufus Rossi, 1790, by monotypy.

Family-group name: APOTOMI J.L. LeConte, 1853. Stem: Apotom-.

Apphianus Beal, 2006: 489. Current status: valid genus in DERMESTIDAE: ATTAGENINAE incertae sedis.

Type species: Apphianus yuccae Beal, 2006, by monotypy.

Family-group name: APPHIANINI Háva in Zahradnik and Háva, 2014: 306. Stem: Apphian-.

Aproida Pascoe, 1863a: 55. Current status: valid genus in CHRYSOMELIDAE: CASSIDINAE: APROIDINI.

Type species: Aproida balyi Pascoe, 1863, by monotypy.

Family-group name: APROIDINI Weise, 1911. Stem: Aproid-.

Aprosopus C. d’Orbigny, 1841: 40. Current status: valid genus in CERAMBYCIDAE: LAMIINAE: AGAPANTHIINI.

Type species: Aprosopus buquetii C. d’Orbigny, 1841, by original designation.

Family-group name: APROSOPITAE J. Thomson, 1864. Stem: Aprosop-.

Apteroessa Hope, 1838a: 13, 159. Current status: valid genus in CICINDELIDAE: CICINDELINI: APTEROESSINA.

Type species: Cicindela grossa Fabricius, 1781, by original designation.

Family-group name: APTEROESSINA Rivalier, 1971. Stem: Apteroess-.

Apterospasta J.L. LeConte, 1862: 272. Current status: junior synonym of Epicauta Dejean, 1834 in MELOIDAE: MELOINAE: EPICAUTINI.

Type species: Lytta segmenta Say, 1824, by subsequent designation (Wellman 1910a: 393).

Family-group name: APTEROSPASTIDES Wellman, 1910. Stem: Apterospast-.

Aptila Fåhraeus, 1870: 258. Current status: valid genus in TENEBRIONIDAE: BLAPTINAE: PEDININI: HELOPININA.

Type species: Aptila costata Fåhraeus, 1870, by subsequent designation (R. Lucas 1920: 115).

Family-group name: APTILINA C. Koch, 1958. Stem: Aptil-.

Aptinus Bonelli, 1810: Tab. Syn. Current status: valid genus in CARABIDAE: BRACHININAE: BRACHININI: APTININA.

Type species: Brachinus mutilatus Fabricius, 1801 (= Carabus bombarda Illiger, 1800), by subsequent monotypy (Panzer 1813: 71).

Family-group name: APTINIDAE Gistel, 1848. Stem: Aptin-.

Aptopus Eschscholtz, 1829a: 32. Current status: valid genus in ELATERIDAE: CARDIOPHORINAE.

Type species: Aptopus tibialis Eschscholtz, 1829, by subsequent designation (Hyslop 1921: 629).

Family-group name: APTOPINA Jakobson, 1913. Stem: Aptopod-.

Aquilex Moret, 1990: 246. Current status: valid genus in CARABIDAE: MIGADOPINAE: MIGADOPINI: AQUILICINA.

Type species: Aquilex diabolicola Moret, 1990, by original designation.

Family-group name: AQUILICINA Moret, 2005. Stem: Aquilic-.

Arachnopus Boisduval, 1835: 435. Current status: junior synonym of Arachnobas Boisduval, 1835 in CURCULIONIDAE: MOLYTINAE: ARACHNOPODINI. Note: this name has been credited to Guérin-Méneville (1838: 127) in the literature but it was first published in an available work by Boisduval (1835: 435) as a junior synonym of Arachnobas Boisduval, 1835; since the name has been treated as available before 1961 (e.g., Guérin-Méneville 1838: 127) it dates from its first publication as a synonym (ICZN 1999a, Article 11.6.1); we act as First Revisers (ICZN 1999a, Article 24.2.3) and select Arachnobas Boisduval, 1835 to have precedence over Arachnopus Boisduval, 1835.

Type species: Cryptorhynchus striga Guérin-Méneville, 1831, by subsequent designation (Lacordaire 1865: 160).

Family-group name: ARACHNOPIDES Lacordaire, 1865. Stem: Arachnopod-.

Araecerus Schönherr, 1823: 1135. Current status: valid genus in ANTHRIBIDAE: CHORAGINAE: ARAECERINI.

Type species: Anthribus coffeae Fabricius, 1801 (= Curculio fasciculatus DeGeer, 1775), by original designation.

Family-group name: ARAEOCÉRIDES Lacordaire, 1865. Stem: Araecer-.

Araeocnemus Nordmann, 1837: 7, 163. Current status: junior synonym of Plochionocerus Dejean, 1833 in STAPHYLINIDAE: STAPHYLININAE: XANTHOLININI.

Type species: Staphylinus fulgens Fabricius, 1792, by subsequent designation (Duponchel and Chevrolat 1841: 64).

Family-group name: ARAEOCNEMES Casey, 1906. Stem: Araeocnem-.

Araeopidius Cockerell, 1906: 241. Current status: valid genus in PTILODACTYLIDAE: ARAEOPIDIINAE. Note: new replacement name for Araeopus J.L. LeConte, 1874.

Type species: Araeopus monachus J.L. LeConte, 1874, by monotypy (type fixation for Araeopus J.L. LeConte).

Family-group name: ARAEOPIDIINAE Lawrence, 1991. Stem: Araeopidi-.

Araeoschizus J.L. LeConte, 1851a: 138. Current status: valid genus in TENEBRIONIDAE: PIMELIINAE: STENOSINI: ARAEOSCHIZINA.

Type species: Araeoschizus costipennis J.L. LeConte, 1851, by monotypy.

Family-group name: ARAEOSCHIZINI Casey, 1907. Stem: Araeoschiz-.

Araptus Eichhoff, 1872: 136. Current status: valid genus in CURCULIONIDAE: SCOLYTINAE: CORTHYLINI: PITYOPHTHORINA.

Type species: Araptus rufopalliatus Eichhoff, 1872, by monotypy.

Family-group name: ARAPTIDAE Eichhoff, 1878. Stem: Arapt-.

Aratea Lacordaire, 1848: 467. Current status: valid genus in CHRYSOMELIDAE: CRYPTOCEPHALINAE: CLYTRINI: ARATEINA.

Type species: Aratea costata Lacordaire, 1848, by monotypy.

Family-group name: ARATEINI Moldenke, 1981. Stem: Arate-.

Araucarius Kuschel, 1966: 7. Current status: valid genus in CURCULIONIDAE: COSSONINAE: ARAUCARIINI.

Type species: Araucarius medius Kuschel, 1966, by original designation.

Family-group name: ARAUCARIINI Kuschel, 1966. Stem: Araucari-.

Archaeoglenes Broun, 1893a: 188. Current status: valid genus in TENEBRIONIDAE: PHRENAPATINAE: ARCHAEOGLENINI.

Type species: Archaeoglenes costipennis Broun, 1893, by monotypy.

Family-group name: ARCHAEOGLENINI Watt, 1975. Stem: Archaeoglen-.

†Archaeoluprops Nabozhenko, Perkovsky & Nazarenko, 2023: 585. Current status: valid genus in TETRATOMIDAE: TETRATOMINAE: †ARCHAEOLUPROPINI. Note: combined description of a new family-group name and a single new genus (ICZN 1999a, Article 13.5). Note: Eocene (Baltic amber).

Type species: †Archaeoluprops groehni Nabozhenko, Perkovsky & Nazarenko, 2023, by monotypy.

Family-group name: †ARCHAEOLUPROPINI Nabozhenko, Perkovsky & Nazarenko, 2023: 585. Stem: Archaeoluprop-.

Archarias A. Villa & G.B. Villa, 1833: 23. Current status: invalid senior synonym of Balanobius Jekel, 1861 in CURCULIONIDAE: CURCULIONINAE: CURCULIONINI: BALANOBIINA. Note: junior homonym of Archarias Dejean, 1821 [Coleoptera: CURCULIONIDAE]; Archarias Dejean, 1821 was suppressed for the purposes of the Principle of Priority but not for those of the Principle of Homonymy (ICZN 1987d); see Alonso-Zarazaga et al. (2023: 8) for an alternative interpretation.

Type species: Curculio salicivorus Paykull, 1792, by subsequent designation (Alonso-Zarazaga and Lyal 1999: 21, 73).

Family-group name: ARCHARIINA Pelsue & O’Brien, 2011: 30. Stem: Archari-. Note: this family-group name was based on “Archarius Gistel, 1856”, which is an incorrect spelling of Archarias A. Villa & G.B. Villa, 1833, introduced as a valid name erroneously by Alonso-Zarazaga and Lyal (1999: 73); family-group name permanently invalid (ICZN 1999a, Article 39).

Archeocrypticus Kaszab, 1964: 360. Current status: valid genus in ARCHEOCRYPTICIDAE.

Type species: Archeocrypticus topali Kaszab, 1964, by original designation.

Family-group name: ARCHEOCRYPTICINI Kaszab, 1964. Stem: Archeo­cryptic-.

†Archescarabaeus Nikolajev, 2010: 69. Current status: valid genus in PLEOCOMIDAE: †ARCHESCARABAEINAE. Note: Lower Cretaceous (Transbai­kalia).

Type species: †Archescarabaeus mesozoicus Nikolajev, 2010, by original designation.

Family-group name: †ARCHESCARABAEINAE Nikolajev, 2010. Stem: Archescarabae-.

Archetypus J. Thomson, 1861: 292, 319. Current status: valid genus in CERAMBYCIDAE: PRIONINAE: MACROTOMINI: ARCHETYPINA.

Type species: Archetypus parandroides J. Thomson, 1861 (= Mallodon fulvipenne Pascoe, 1859), by subsequent designation (J. Thomson 1864: 307).

Family-group name: ARCHETYPI Lameere, 1912. Stem: Archetyp-.

Archeuops Legalov, 2003: 360. Current status: valid genus in ATTELABIDAE: ATTELABINAE: EUOPINI.

Type species: Euops vossi Heller, 1929, by original designation.

Family-group name: ARCHEUOPSINA Legalov, 2003. Stem: Archeuops- (ICZN 1999a, Article 29.4).

Archicorynus R.S. Anderson & Marvaldi, 2013: 66. Current status: valid genus in BELIDAE: OXYCORYNINAE: OXYCORYNITAE: ARCHICORYNINI.

Type species: Archicorynus kuscheli R.S. Anderson & Marvaldi, 2013, by original designation.

Family-group name: ARCHICORYNINI R.S. Anderson & Marvaldi, 2013: 66. Stem: Archicoryn-.

Archolabus Voss, 1929b: 200. Current status: junior synonym of Microstylus Schönherr, 1847 in CURCULIONIDAE: CURCULIONINAE: MICROSTYLINI.

Type species: Archolabus methneri Voss, 1929, by original designation.

Family-group name: ARCHOLABINAE Voss, 1929. Stem: Archolab-.

Ardella Paulsen, 2021: 2. Current status: valid genus in SCARABAEIDAE: MELOLONTHINAE: ARDELLINI.

Type species: Ardella magnaemirabilis Paulsen, 2021, by original designation.

Family-group name: ARDELLINI Paulsen, 2021: 5. Stem: Ardell-.

Ardistomis Putzeys, 1846: 2, 118. Current status: valid genus in CARABIDAE: SCARITINAE: CLIVININI: ARDISTOMINA.

Type species: Ardistomis fasciolata Putzeys, 1846, by subsequent designation (E. Desmarest 1851: 102).

Family-group name: ARDISTOMIDES Putzeys, 1867. Stem: Ardistom-.

Areoda W.S. MacLeay, 1819: 158. Current status: valid genus in SCARABAEIDAE: RUTELINAE: RUTELINI: AREODINA.

Type species: Areoda leachii W.S. MacLeay, 1819, by monotypy.

Family-group name: AREODIDAE Burmeister, 1844. Stem: Areod-.

Arescus Perty, 1832: 100. Current status: valid genus in CHRYSOMELIDAE: CASSIDINAE: ARESCINI.

Type species: Arescus labiatus Perty, 1832, by monotypy.

Family-group name: ARESCITES Chapuis, 1875. Stem: Aresc-.

Argentinomacer Legalov, 2023c: 21. Current status: valid genus in NEMONYCHIDAE: RHINORHYNCHINAE: ARGENTINOMACERINI. Note: Argentinomacer Legalov (2017a: 74) was published in a work that is nomenclaturally unavailable (ICZN 2012f, Article 8.5.3).

Type species: Argentinomacer unicus Legalov, 2023, by original designation. Note: Argentinomacer unicus Legalov, 2017 was published in a work that is nomenclaturally unavailable (ICZN 2012f, Article 8.5.3).

Family-group name: ARGENTINOMACERINI Legalov, 2023c: 21. Stem: Argentinomacer-. Note: ARGENTINOMACERINI Legalov (2017a: 73) was published in a work that is nomenclaturally unavailable (ICZN 2012f, Article 8.5.3).

Argentipilosa Gordon & de Almeida, 1991: 150. Current status: valid genus in COCCINELLIDAE: COCCINELLINAE: ARGENTIPILOSINI.

Type species: Argentipilosa nigra Gordon & de Almeida, 1991, by original designation.

Family-group name: ARGENTIPILOSINI Gordon & de Almeida, 1991. Stem: Argentipilos-.

Arhina Murray, 1867a: 178. Current status: invalid senior synonym of Arhinops Kirejtshuk & Bouchard, 2018 in NITIDULIDAE: CRYPTARCHINAE: ARHINOPINI. Note: junior homonym of Arhina Agassiz, 1846 [Diptera].

Type species: Arhina strongyloides Murray, 1867, by monotypy.

Family-group name: ARHININI Kirejtshuk, 1987. Stem: Arhin-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

Arhinops Kirejtshuk & Bouchard, 2018: 157. Current status: valid genus in NITIDULIDAE: CRYPTARCHINAE: ARHINOPINI. Note: new replacement name for Arhina Murray, 1867.

Type species: Arhina strongyloides Murray, 1867, by monotypy (type fixation for Arhina Murray).

Family-group name: ARHINOPINI Kirejtshuk & Bouchard, 2018: 157. Stem: Arhinop-.

Arhytodes Reitter, 1882a: 193, 209. Current status: junior synonym of Rhytus Westwood, 1870 in STAPHYLINIDAE: PSELAPHINAE: PSELAPHITAE: ARHYTODINI. Note: unnecessary new replacement name for Rhytus Westwood, 1870.

Type species: Rhytus vestitus Westwood, 1870, by monotypy (type fixation for Rhytus Westwood).

Family-group name: ARHYTODINI Raffray, 1890. Stem: Arhytod-.

Aristochroodes Marcilhac, 1993: 273. Current status: valid genus in CARABIDAE: HARPALINAE: PTEROSTICHINI: PTEROSTICHINA. Note: Marcilhac used two different spellings for this taxon, Aristochroodes (p. 271, 273, 274) and Aritochroodes (p. 274); we are not aware of anyone who acted as First Reviser (ICZN 1999a, Article 24.2.3) and we here select Aristochroodes as the correct original spelling.

Type species: Pterostichus reginae Marcilhac, 1993, by original designation.

Family-group name: ARISTOCHROODINI Sciaky, 1996. Stem: Aristochrood-.

Arneus Gistel, 1848b: xi, xiv. Current status: junior synonym of Sericus Eschscholtz, 1829 in ELATERIDAE: ELATERINAE: ELATERINI. Note: unnecessary new replacement name for Sericosomus Dejean, 1833.

Type species: Elater brunneus Linnaeus, 1758, by subsequent designation (Westwood 1838c: 25) (type fixation for Sericosomus Dejean).

Family-group name: ARNEIDAE Gistel, 1848. Stem: Arne-.

Arnyllium Reitter, 1884a: 391. Current status: valid genus in STAPHYLINIDAE: PSELAPHINAE: GONIACERITAE: ARNYLLIINI.

Type species: Arnyllium pectinatum Reitter, 1884, by subsequent designation (Jeannel 1952: 101).

Family-group name: ARNYLLIINI Jeannel, 1952. Stem: Arnylli-.

Arpediomimus Cameron, 1917b: 277. Current status: junior synonym of Crymus Fauvel, 1904 in STAPHYLINIDAE: OMALIINAE: OMALIINI. Note: new replacement name for Arpediopsis Cameron, 1917.

Type species: Arpediopsis falklandica Cameron, 1917 (= Crymus antarcticus Fauvel, 1904), by monotypy (type fixation for Arpediopsis Cameron).

Family-group name: ARPEDIOMIMI Cameron, 1917. Stem: Arpediomim-.

Arpediopsis Cameron, 1917a: 124. Current status: junior synonym of Crymus Fauvel, 1904 in STAPHYLINIDAE: OMALIINAE: OMALIINI. Note: junior homonym of Arpediopsis Ganglbauer, 1895 [Coleoptera: STAPHYLINIDAE].

Type species: Arpediopsis falklandica Cameron, 1917 (= Crymus antarticus Fauvel, 1904), by monotypy.

Family-group name: ARPEDIOPSINI Cameron, 1917. Stem: Arpediopse-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

Arrenodes Schönherr 1823: 1137. Current status: valid genus in BRENTIDAE: BRENTINAE: BRENTINI: ARRENODINA.

Type species: Brentus maxillosus G.-A. Olivier, 1807 (= Curculio minutus Drury, 1773), by original designation.

Family-group name: ARRHÉNODIDES Lacordaire, 1865. Stem: Arrenod-. Note: this family-group name is based on a misidentification of Arrenodes Schönherr, 1823 by Lacordaire (1865: 429), which corresponds to the genus Estenorhinus Lacordaire, 1865; Alonso- Zarazaga and Lyal (1999: 78) mentioned that they would apply to the Commission for the conservation of this tribe by designating Arrenodes Schönherr, 1823 as its type genus; meanwhile current usage is maintained.

Arrhipis Gemminger in Gemminger and Harold, 1869b: 1477. Current status: invalid senior synonym of Nematodinus Lea, 1919 in EUCNEMIDAE: MELASINAE: DIRHAGINI. Note: junior homonym of Arrhipis Agassiz, 1846 [Pisces]; this genus-group name is usually credited to Bonvouloir, 1871 but it was first made available by Gemminger in 1869 when he associated two available species with the name; Arrhipis Gemminger is currently recognized as a valid genus but since it is a junior homonym, Nematodinus Lea, 1919, stat. nov. is used as valid instead.

Type species: Hylochares subacutus Guérin-Méneville, 1843, by subsequent designation (Fleutiaux 1902: 659).

Family-group name: ARHIPINI Cobos, 1965. Stem: Arrhipid-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

Arrowinus Bernhauer, 1935: 215. Current status: valid genus in STAPHYLINIDAE: STAPHYLININAE: ARROWININI.

Type species: Arrowinus phaenomenalis Bernhauer, 1935, by original designation.

Family-group name: ARROWININI Solodovnikov & Newton, 2005. Stem: Arrowin-.

Arsipoda Erichson, 1842: 235. Current status: valid genus in CHRYSOMELIDAE: GALERUCINAE: ALTICINI.

Type species: Arsipoda bifrons Erichson, 1842, by monotypy.

Family-group name: ARSIPODITES Chapuis, 1875. Stem: Arsipod-.

Artaxus Gistel, 1848b: 216. Current status: junior synonym of Mycterus Clairville, 1798 in MYCTERIDAE: MYCTERINAE. Note: unnecessary new replacement name for Mycterus Clairville, 1798.

Type species: Mycterus griseus Clairville, 1798 (= Rhinomacer curculioides Fabricius, 1781), by monotypy (type fixation for Mycterus Clairville).

Family-group name: ARTAXIDAE Gistel, 1848. Stem: Artax-.

Artematopus Perty, 1832: 115. Current status: valid genus in ARTEMATOPODIDAE: ARTEMATOPODINAE: ARTEMATOPODINI.

Type species: Artematopus longicornis Perty, 1832, by monotypy.

Family-group name: ARTÉMATOPIDES Lacordaire, 1857. Stem: Artematopod-.

Arthrobrachus Solier, 1849: 414. Current status: valid genus in MELYRIDAE: MELYRINAE: ARTHROBRACHINI.

Type species: Arthrobrachus varians Solier, 1849 (= Zygia flavipennis Laporte, 1838), by subsequent designation (Majer 1987: 794).

Family-group name: ARTHROBRACHINI Majer, 1987. Stem: Arthrobrach-.

Arthrolips Wollaston, 1854: 475. Current status: valid genus in CORYLOPHIDAE: CORYLOPHINAE: PARMULINI.

Type species: Clypeaster piceus Comolli, 1837, by monotypy.

Family-group name: ARTHROLIPINAE Böving & Craighead, 1931. Stem: Arthrolip-.

†Arthropterites Wasmann, 1926: 28. Current status: valid genus in CARABIDAE: PAUSSINAE: PAUSSINI: †ARTHROPTERITINA. Note: Eocene (Baltic amber).

Type species: †Arthropterites klebsi Wasmann, 1926, by monotypy.

Family-group name: †ARTHROPTERITINA Luna de Carvalho, 1961. Stem: Arthropterit-.

Arthropterus W.S. MacLeay, 1838: 75. Current status: valid genus in CARABIDAE: PAUSSINAE: PAUSSINI: CERAPTERINA.

Type species: Cerapterus macleaii Donovan, 1805, by monotypy.

Family-group name: ARTHROPTERINI Wasmann, 1928. Stem: Arthropter-. Note: junior homonym of ARTHROPTERIDAE Jordan, 1923 (type genus Arthropterus Agassiz, 1843) in Pisces; Jordan’s name is permanently invalid (based on a preoccupied type genus).

Arthrostenus Schönherr, 1826: 287. Current status: valid genus in CURCULIONIDAE: BRACHYCERINAE: ERIRHININI.

Type species: Arthrostenus fullo Boheman, 1836, by subsequent designation (E. Desmarest 1860: 242). Note: no available species was originally included for this genus in Schönherr (1826); the first ones are the three species listed in Schönherr (1836: 534, 535).

Family-group name: ARTHROSTENINI Reitter, 1913. Stem: Arthrosten-.

Articerus Dalman, 1826: 394. Current status: valid genus in STAPHYLINIDAE: PSELAPHINAE: CLAVIGERITAE: CLAVIGERINI.

Type species: Articerus armatus Dalman, 1826, by monotypy.

Family-group name: ARTICERIDES E. Desmarest, 1852. Stem: Articer-. Note: this family-group name has been credited to L.W. Schaufuss (1872: 245) by most authors (e.g., Newton and Chandler 1989: 5) and its type genus is considered to be misidentified in the sense of Hope (1845a: 106); however, the family-group name was first made available by E. Desmarest (1852: 145) in the proper sense since E. Desmarest referred to the insect described by Dalman in copal.

Artipus C.R. Sahlberg, 1823a: 22. Current status: valid genus in CURCULIONIDAE: ENTIMINAE: POLYDRUSITAE: NAUPACTINI: NAUPACTINA. Note: we consider Artipus used by Schönherr (1823: 1141) a subsequent usage of Artipus C.R. Sahlberg, 1823.

Type species: Artipus corycaeus C.R. Sahlberg, 1823, by monotypy. Note: combined description of a new nominal genus and a single new nominal species (ICZN 1999a, Article 12.2.6).

Family-group name: ARTIPI G.H. Horn, 1876. Stem: Artipod-.

Asaphes W. Kirby, 1837: 146. Current status: invalid senior synonym of Hemicrepidius Germar, 1839 in ELATERIDAE: DENDROMETRINAE: DENDROMETRINI: HEMICREPIDIINA. Note: junior homonym of Asaphes Walker, 1834 [Hymenoptera].

Type species: Pedetes ruficornis W. Kirby, 1837, by monotypy.

Family-group name: ASAPHITES Candèze, 1863. Stem: Asaph-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

Asclera Dejean, 1834: 228. Current status: junior synonym of Ischnomera Stephens, 1832 in OEDEMERIDAE: OEDEMERINAE: ASCLERINI.

Type species: Necydalis sanguinicollis Fabricius, 1787, by subsequent designation (Arnett 1950: 219).

Family-group name: ASCLERAEIDAE Gistel, 1848. Stem: Ascler-.

Ascydmus Casey, 1897: 505. Current status: junior synonym of Euthiconus Reitter, 1882 in STAPHYLINIDAE: SCYDMAENINAE: CEPHENNIITAE: EUTHEIINI.

Type species: Ascydmus tener Casey, 1897, by monotypy.

Family-group name: ASCYDMINI Casey, 1897. Stem: Ascydm-.

Asemum Eschscholtz, 1830a: 66. Current status: valid genus in CERAMBYCIDAE: SPONDYLIDINAE: ASEMINI.

Type species: Cerambyx striatus Linnaeus, 1758, by subsequent designation (Westwood 1838c: 40). Note: name placed on the Official List of Specific Names in Zoology (ICZN 2017a).

Family-group name: ASEMITAE J. Thomson, 1861. Stem: Asem-.

Asida Latreille, 1802b: 167. Current status: valid genus in TENEBRIONIDAE: PIMELIINAE: ASIDINI.

Type species: Opatrum griseum Fabricius, 1781, by monotypy.

Family-group name: ASIDADAE Fleming, 1821. Stem: Asid-.

†Asiocoleus Rohdendorf, 1961: 396. Current status: valid genus in †ASIOCOLEIDAE. Note: lower Permian (Russia).

Type species: †Asiocoleus novojilovi Rohdendorf, 1961, by original designation.

Family-group name: †ASIOCOLEIDAE Rohdendorf, 1961. Stem: Asiocole-.

Asphalus Pascoe, 1868: xii. Current status: valid genus in TENEBRIONIDAE: TENEBRIONINAE: HELEINI: ASPHALINA.

Type species: Asphalus ebeninus Pascoe, 1868, by monotypy.

Family-group name: ASPHALINA E.G. Matthews & Lawrence, 2005. Stem: Asphal-.

Aspicela Dejean, 1836a: 387. Current status: valid genus in CHRYSOMELIDAE: GALERUCINAE: ALTICINI.

Type species: Altica unipunctata Latreille, 1813, by subsequent designation (Monrós and Bechyné 1956: 1134).

Family-group name: ASPICÉLITES Chapuis, 1875. Stem: Aspicel-.

Aspidapion Schilsky, 1901: 38G. Current status: valid genus in BRENTIDAE: APIONINAE: ASPIDAPIITAE: ASPIDAPIINI.

Type species: Apion validum Germar, 1817, by original designation.

Family-group name: ASPIDAPIINI Alonso-Zarazaga, 1990. Stem: Aspidapi-.

Aspidimerus Mulsant, 1850: 944. Current status: valid genus in COCCINELLIDAE: COCCINELLINAE: ASPIDIMERINI.

Type species: Aspidimerus spencii Mulsant, 1850, by monotypy.

Family-group name: ASPIDIMÉRAIRES Mulsant, 1850. Stem: Aspidimer-.

Aspidimorpha Hope, 1841a: 158, 159. Current status: valid genus in CHRYSOMELIDAE: CASSIDINAE: ASPIDIMORPHINI.

Type species: Cassida miliaris Fabricius 1775, by original designation.

Family-group name: ASPIDIMORPHITES Chapuis, 1875. Stem: Aspidimorph-.

Aspidiphorus Dejean, 1821: 47. Current status: valid genus in SPHINDIDAE: SPHINDINAE. Note: this genus was originally spelled Arpidiphorus but corrected to Aspidiphorus by the Commission and the name was placed (as Aspidiphorus Ziegler in Dejean, 1821) on the Official List of Generic Names in Zoology (ICZN 1997).

Type species: Nitidula orbiculata Gyllenhal, 1808, by monotypy (see ICZN 1997). Note: name placed on the Official List of Specific Names in Zoology (ICZN 1997).

Family-group name: ASPIDIPHORIDAE Kiesenwetter, 1877. Stem: Aspidiphor-.

Aspidytes Ribera, Beutel, Balke & Vogler, 2002: 2352. Current status: valid genus in ASPIDYTIDAE.

Type species: Aspidytes niobe Ribera, Beutel, Balke & Vogler, 2002, by original designation.

Family-group name: ASPIDYTIDAE Ribera, Beutel, Balke & Vogler, 2002. Stem: Aspidyt-.

Aspilus Schaum, 1848: 61. Current status: valid genus in SCARABAEIDAE: CETONIINAE: CREMASTOCHEILINI: ASPILINA.

Type species: Ptychophorus gambiensis Burmeister, 1842, by monotypy.

Family-group name: ASPILINA Krikken, 1984. Stem: Aspil-.

Astaena Erichson, 1847c: 101. Current status: valid genus in SCARABAEIDAE: SERICINAE: SERICINI: ASTAENINA.

Type species: Astaena tridentata Erichson, 1847, by monotypy.

Family-group name: ASTAENIDAE Burmeister, 1855. Stem: Astaen-.

Astathes Newman, 1842a: 299. Current status: junior synonym of Tetraophthalmus Dejean, 1835 in CERAMBYCIDAE: LAMIINAE: ASTATHINI.

Type species: Astathes perplexa Newman, 1842, by subsequent designation (J. Thomson 1864: 118).

Family-group name: ASTATHITAE J. Thomson, 1864. Stem: Astath-.

Astenus Dejean, 1833a: 65. Current status: valid genus in STAPHYLINIDAE: PAEDERINAE: LATHROBIINI: ASTENINA.

Type species: Staphylinus angustatus Paykull, 1789 (= Staphylinus gracilis Paykull, 1789), by subsequent designation (Westwood 1838c: 17).

Family-group name: ASTENINA Hatch, 1957. Stem: Asten-.

Asteriza Chevrolat in Dejean, 1836a: 372. Current status: valid genus in CHRYSOMELIDAE: CASSIDINAE: ISCHYROSONYCHINI.

Type species: Cassida flavicornis G.-A. Olivier, 1790, by monotypy.

Family-group name: ASTERIZINI Hincks, 1952. Stem: Asteriz-.

Astraeus Laporte & Gory, 1838: 1. Current status: valid genus in BUPRESTIDAE: POLYCESTINAE: ASTRAEINI. Note: the original spelling Asthraeus was emended to Astraeus by the Commission and Astraeus Laporte & Gory, 1838 placed on the Official List of Generic Names in Zoology (ICZN 1966b, as Astraeus Laporte & Gory, 1837).

Type species: Astraeus flavopictus Laporte & Gory, 1838, by monotypy (see ICZN 1966b, as Asthraeus flavopictus Laporte & Gory, 1837).

Family-group name: ASTRAEUSINI Cobos, 1980. Stem: Astrae-.

Astrotus J.L. LeConte, 1858b: 19. Current status: valid genus in TENEBRIONIDAE: PIMELIINAE: ASIDINI.

Type species: Microschatia contorta J.L. LeConte, 1853, by original designation.

Family-group name: ASTROTI G.H. Horn, 1870. Stem: Astrot-.

Astylus Laporte, 1838: 32. Current status: valid genus in MELYRIDAE: MELYRINAE: ASTYLINI.

Type species: Anobium lineatum Fabricius, 1775, by subsequent designation (Constantin 2011: 39).

Family-group name: ASTYLINI Pic, 1929. Stem: Astyl-.

Astynomus Dejean, 1835: 337. Current status: junior synonym of Aedilis Audinet-Serville, 1835 in CERAMBYCIDAE: LAMIINAE: ACANTHOCININI. Note: unnecessary new replacement name for Aedilis Audinet-Serville, 1835.

Type species: Aedilis montana Audinet-Serville, 1835 (= Cerambyx aedilis Linnaeus, 1758), by subsequent designation (Westwood 1838c: 40) (type fixation for Aedilis Audinet- Serville).

Family-group name: ASTYNOMINI Portevin, 1927. Stem: Astynom-.

Ataenius Harold, 1867a: 82. Current status: valid genus in SCARABAEIDAE: APHODIINAE: EUPARIINI. Note: this genus was described as new, with a single other new species, by von Harold (1867b: 100); name given precedence over Aphodinus Motschulsky, 1862 whenever the two are considered to be synonyms and placed on the Official List of Generic Names in Zoology (ICZN 2010a).

Type species: Ataenius scutellaris Harold, 1867, by subsequent designation (Chapin 1940: 12; see ICZN 2010a). Note: name placed on the Official List of Specific Names in Zoology (ICZN 2010a).

Family-group name: ATAENIDAE Harold, 1867. Stem: Ataeni-. Note: nomen oblitum (see A.B.T. Smith 2006: 158).

Atanygnathus Jakobson, 1909: 521. Current status: valid genus in STAPHYLINIDAE: STAPHYLININAE: STAPHYLININI: TANYGNATHININA. Note: new replacement name for Tanygnathus Erichson, 1839.

Type species: Tanygnathus terminalis Erichson, 1839, by monotypy (type fixation for Tanygnathus Erichson).

Family-group name: ATANYGNATHINI Lohse, 1964. Stem: Atanygnath-.

Ataxia Haldeman, 1847: 56. Current status: valid genus in CERAMBYCIDAE: LAMIINAE: PTEROPLIINI.

Type species: Ataxia sordida Haldeman, 1847 (= Lamia crypta Say, 1832), by monotypy.

Family-group name: ATAXIIDES Lacordaire, 1872. Stem: Ataxi-.

Ateledera Lacordaire, 1845: 607. Current status: valid genus in MEGALOPODIDAE: ATELEDERINAE.

Type species: Ateledera cygnoides Lacordaire, 1845, by monotypy.

Family-group name: ATELEDERINAE Rodríguez-Mirón in Rodríguez-Mirón et al., 2021: 694. Stem: Ateleder-.

Atelius C.O. Waterhouse, 1878: 104. Current status: valid genus in LYCIDAE: ATELIINAE: ATELIINI.

Type species: Lycus expansicornis Walker, 1858, by original designation.

Family-group name: ATELINAE Kleine, 1929. Stem: Ateli-.

Aterpus Schönherr, 1826: 156. Current status: junior synonym of Aades Schönherr, 1823 in CURCULIONIDAE: CYCLOMINAE: ATERPINI. Note: unnecessary new replacement name for Aades Schönherr, 1823.

Type species: Curculio cultratus Fabricius, 1775, by subsequent designation (Schönherr 1833: 13).

Family-group name: ATERPIDES Lacordaire, 1863. Stem: Aterp-. Note: nomen protectum (see Bouchard et al. 2011: 601).

Ateuchus Weber, 1801: 10. Current status: valid genus in SCARABAEIDAE: SCARABAEINAE: ATEUCHINI: ATEUCHINA. Note: this genus was described twice in the same year by Weber (1801: 10) and Fabricius (1801a: 54); bibliographic research suggests that Weber’s book (issued by March 1801, see Evenhuis 1997b: 809) was published before the first volume of Fabricius’ ‘Systema Eleutheratorum’ (issued by 26 April 1801, see Bousquet 2017a: 108; see also Chapin 1946: 79).

Type species: Ateuchus histeroides Weber, 1801, by monotypy.

Family-group name: ATEUCHIDAE Perty, 1830. Stem: Ateuch-. Note: Perty (1830: 38) did not provide the author to the genus Ateuchus; it is assumed here that he referred to Ateuchus Weber, not Ateuchus Fabricius.

Athertonium Crowson, 1990: 91, 92. Current status: valid genus in BOGANIIDAE: PARACUCUJINAE.

Type species: Athertonium parvum Crowson, 1990, by original designation.

Family-group name: ATHERTONIINI Crowson, 1990. Stem: Athertoni-.

Atheta C.G. Thomson, 1858: 36. Current status: valid genus in STAPHYLINIDAE: ALEOCHARINAE: ATHETINI: ATHETINA. Note: name placed on the Official List of Generic Names in Zoology (ICZN 1961b).

Type species: Aleochara graminicola Gravenhorst, 1806, by subsequent monotypy (see ICZN 1961b). Note: although the Commission (ICZN 1961b: 241) mentioned that the type species was designated by monotypy, this is incorrect since there is no species associated with the name Atheta in C.G. Thomson (1858: 36); Aleochara graminicola Gravenhorst was fixed as the type species by subsequent monotypy in C.G. Thomson (1859: 39); the Official List of Generic Names in Zoology needs to be updated to reflect the changes for the fixation of the type species mentioned above.

Family-group name: ATHETAE Casey, 1910. Stem: Athet-.

Athexenia Pace, 2000b: 336. Current status: valid genus in STAPHYLINIDAE: ALEOCHARINAE: LOMECHUSINI: MYRMEDONIINA.

Type species: Athexenia nilgiriensis Pace, 2000, by original designation.

Family-group name: ATHEXENINA Pace, 2000. Stem: Athexeni-.

Athlia Erichson, 1835: 266. Current status: valid genus in SCARABAEIDAE: MELOLONTHINAE: ATHLIINI.

Type species: Athlia rustica Erichson, 1835, by monotypy.

Family-group name: ATHLIINI A.B.T. Smith & Evans, 2018: 280. Stem: Athli-

Athoomorphus O. Schwarz, 1898: 156. Current status: valid genus in ELATERIDAE incertae sedis.

Type species: Athoomorphus cylindricus O. Schwarz, 1898, by monotypy.

Family-group name: ATHOOMORPHINAE Laurent, 1966. Stem: Athoomorph-.

Athous Eschscholtz, 1829a: 33. Current status: valid genus in ELATERIDAE: DENDROMETRINAE: DENDROMETRINI: DENDROMETRINA.

Type species: Elater vittatus Fabricius, 1792, by subsequent designation (Westwood 1838c: 26).

Family-group name: ATHOITES Candèze, 1859. Stem: Atho-.

Athyreus W.S. MacLeay, 1819: 123. Current status: valid genus in BOLBOCERATIDAE: ATHYREINAE. Note: name placed on the Official List of Generic Names in Zoology (ICZN 1985a).

Type species: Athyreus bifurcatus W.S. MacLeay, 1819, by subsequent designation (Howden and Martínez 1963: 350; see ICZN 1985a). Note: name placed on the Official List of Specific Names in Zoology (ICZN 1985a).

Family-group name: ATHYREITAE Lynch Arribálzaga, 1878. Stem: Athyre-.

Atimia Haldeman, 1847: 56. Current status: valid genus in CERAMBYCIDAE: SPONDYLIDINAE: ATIMIINI.

Type species: Atimia tristis Haldeman, 1847 (= Clytus confusus Say, 1826), by monotypy.

Family-group name: ATIMIINI J.L. LeConte, 1873. Stem: Atimi-.

Atomaria Stephens, 1829b: 8. Current status: valid genus in CRYPTOPHAGIDAE: ATOMARIINAE: ATOMARIINI.

Type species: Silpha evanescens Marsham, 1802 (= Dermestes pusillus Paykull, 1798), by subsequent designation (Partington 1834: 244). Note: the earliest type species designation previously known for this genus was Dermestes nigripennis Paykull, 1798 (= Latridius nigripennis Kugelann, 1794), as designated subsequently by Westwood (1838c: 14) (e.g., C. Johnson 2007a: 65, Otero and Pereira Martínez 2022: 3), but Partington’s fixation is older; Silpha evanescens Marsham, 1802, which was later nominally placed in the genus Ptenidium Erichson, 1845 in the family PTILIIDAE (e.g., Matthews 1872: 80) is actually a junior synonym of Atomaria pusilla (Paykull, 1798) (see Gemminger and von Harold 1868c: 890; Ganglbauer 1899: 303); the synonymy of the type species has been confirmed by examination of type photographs (M.L. Gimmel, pers. comm. to P.B., 2023); this change of type species will result in the subgenus previously known under the name Anchicera C.G. Thomson, 1863 changing to Atomaria Stephens, 1829 and the subgenus previously known under the name Atomaria Stephens, 1829 changing to Agathengis Gozis, 1886 (M.L. Gimmel, pers. comm. to P.B., 2023).

Family-group name: ATOMARIINI J.L. LeConte, 1861. Stem: Atomari-.

Atopa Paykull, 1799: 116. Current status: junior synonym of Dascillus Latreille, 1797 in DASCILLIDAE: DASCILLINAE: DASCILLINI.

Type species: Chrysomela cervina Linnaeus, 1758, by monotypy.

Family-group name: ATOPITES Laporte, 1834. Stem: Atop-.

Atopida A. White, 1846: 8. Current status: valid genus in SCIRTIDAE: SCIRTINAE.

Type species: Atopida castanea A. White, 1846, by monotypy.

Family-group name: ATOPIDINI Pic, 1914. Stem: Atopid-.

Atopobrentus Damoiseau, 1965: 3. Current status: valid genus in BRENTIDAE: BRENTINAE: TRACHELIZINI: ATOPOBRENTINA.

Type species: Atopobrentus allardi Damoiseau, 1965, by original designation.

Family-group name: ATOPOBRENTINI Damoiseau, 1965. Stem: Atopobrent-.

Atossa J. Thomson, 1864: 100. Current status: synonym of Grammoechus J. Thomson, 1864 in CERAMBYCIDAE: LAMIINAE: PTEROPLIINI. Note: the First Reviser (ICZN 1999a, Article 24.2.2) for Grammoechus J. Thomson, 1864 versus Atossa J. Thomson, 1864 is Breuning (1961: 232).

Type species: Atossa strenua J. Thomson, 1864, by original designation.

Family-group name: ATOSSIDES Lacordaire, 1872. Stem: Atoss-.

Atractocerus Palisot de Beauvois, 1801: 516. Current status: valid genus in LYMEXYLIDAE: ATRACTOCERINAE. Note: this genus-group name was spelled Atractourus in the original article but corrected to Atractocerus on page 576 of the same publication.

Type species: Atractocerus necydaloides Palisot de Beauvois, 1801 (= Necydalis brevicornis Linnaeus, 1767), by monotypy.

Family-group name: ATRACTOCÉRITES Laporte, 1840. Stem: Atractocer-.

Atranopsis Baehr, 1982: 261. Current status: valid genus in CARABIDAE: HARPALINAE: SPHODRINI: ATRANOPSINA.

Type species: Atranopsis scheuerni Baehr, 1982, by original designation.

Family-group name: ATRANOPSINA Baehr, 1982. Stem: Atranops-.

Atranus J.L. LeConte, 1847: 438. Current status: valid genus in CARABIDAE: HARPALINAE: ATRANINI.

Type species: Anchomenus pubescens Dejean, 1828, by monotypy.

Family-group name: ATRANI G.H. Horn, 1881. Stem: Atran-.

Atrecus Jacquelin du Val, 1857b: 31. Current status: valid genus in STAPHYLINIDAE: STAPHYLININAE: OTHIINI.

Type species: Staphylinus pilicornis Paykull, 1790, by original designation.

Family-group name: ATRECINI Hatch, 1957. Stem: Atrec-.

Attagenus Latreille, 1802b: 121. Current status: valid genus in DERMESTIDAE: ATTAGENINAE: ATTAGENINI.

Type species: Dermestes trifasciatus Fabricius, 1787, by subsequent designation (des Gozis 1886: 10). Note: Dermestes pellio Linnaeus, 1758 is usually listed as type species of the genus (e.g., Háva 2007: 307) but it is not an originally included species.

Family-group name: ATTAGÉNITES Laporte, 1840. Stem: Attagen-.

Attalomimus Wittmer, 1976a: 288. Current status: valid genus in MELYRIDAE: MALACHIINAE: LEMPHINI.

Type species: Attalomimus cephalotes Wittmer, 1976, by original designation.

Family-group name: ATTALOMIMIDAE Majer, 1995. Stem: Attalomim-.

Attalus Erichson, 1840a: 89. Current status: valid genus in MELYRIDAE: MALACHIINAE: MALACHIINI.

Type species: Attalus lusitanicus Erichson, 1840, by subsequent designation (Champion 1914a: 42).

Family-group name: ATTALAIRES Abeille de Perrin, 1890. Stem: Attal-.

Attapsenius Bruch, 1933a: 27. Current status: valid genus in STAPHYLINIDAE: PSELAPHINAE: PSELAPHITAE: ATTAPSENIINI.

Type species: Attapsenius chernosvitovi Bruch, 1933, by original designation.

Family-group name: ATTAPSENIINI Bruch, 1933. Stem: Attapseni-.

Attavicinus Philips & K.L. Bell, 2008: 68. Current status: valid genus in SCARABAEIDAE: SCARABAEINAE: ONITICELLINI: ATTAVICINA.

Type species: Oniticellus monstrosus H.W. Bates, 1887, by original designation.

Family-group name: ATTAVICINA Philips, 2016: 41. Stem: Attavicin-.

Attelabus Linnaeus, 1758: 342, 387. Current status: valid genus in ATTELABIDAE: ATTELABINAE: ATTELABINI: ATTELABINA. Note: name placed on the Official List of Generic Names in Zoology (ICZN 1983a).

Type species: Attelabus curculionoides Linnaeus, 1767 (= Curculio nitens Scopoli, 1763), by plenary power (see ICZN 1983a). Note: Curculio nitens Scopoli, 1763 was placed on the Official List of Specific Names in Zoology (ICZN 1983a).

Family-group name: ATTELABIDES Billberg, 1820. Stem: Attelab-.

Atysa Baly, 1864a: 238. Current status: valid genus in CHRYSOMELIDAE: GALERUCINAE: GALERUCINI.

Type species: Atysa terminalis Baly, 1864, by original designation.

Family-group name: ATYSITES Chapuis, 1875. Stem: Atys-.

Aubehydrus Guignot, 1942: 11. Current status: junior synonym of Notaticus A. Zimmermann, 1928 in DYTISCIDAE: DYTISCINAE: AUBEHYDRINI.

Type species: Aubehydrus speciosissimus Guignot, 1942 (= Notaticus fasciatus A. Zimmermann, 1928), by original designation.

Family-group name: AUBEHYDRINAE Guignot, 1942. Stem: Aubehydr-.

Auchmeresthes Kraatz, 1862a: 119. Current status: valid genus in CURCULIONIDAE: ENTIMINAE: CYPHICERITAE: METACINOPINI.

Type species: Auchmeresthes kiesenwetteri Kraatz, 1862, by monotypy.

Family-group name: AUCHMERESTHINAE Reitter, 1913. Stem: Auchmeresthet-.

Auchmobius J.L. LeConte, 1851a: 139. Current status: valid genus in TENEBRIONIDAE: PIMELIINAE: EDROTINI.

Type species: Auchmobius sublaevis J.L. LeConte, 1851, by monotypy.

Family-group name: AUCHMOBII J.L. LeConte & G.H. Horn, 1883. Stem: Auchmobi-.

Augyles Schiødte, 1866: 159, 166. Current status: valid genus in HETEROCERIDAE: HETEROCERINAE: AUGYLINI.

Type species: Heterocerus hispidulus Kiesenwetter, 1843, by monotypy.

Family-group name: AUGYLIINI M.F. Pacheco, 1964. Stem: Augyl-.

Aulacocerus A. White, 1853: 13. Current status: junior synonym of Braderochus Buquet, 1852 in CERAMBYCIDAE: PRIONINAE: PRIONINI.

Type species: Aulacocerus mundus A. White, 1853, by monotypy.

Family-group name: AULACOCERITAE J. Thomson, 1864. Stem: Aulacocer-.

Aulacocyclus Kaup, 1868a: 4. Current status: valid genus in PASSALIDAE: AULACOCYCLINAE: AULACOCYCLINI.

Type species: Passalus edentulus W.S. MacLeay, 1826, by subsequent designation (Gravely 1918: 17).

Family-group name: AULACOCYCLINAE Kaup, 1868. Stem: Aulacocycl-.

Aulacophora Chevrolat in Dejean, 1836a: 378. Current status: valid genus in CHRYSOMELIDAE: GALERUCINAE: LUPERINI.

Type species: Galleruca quadraria G.-A. Olivier, 1808, by subsequent designation (Duponchel and Chevrolat 1842a: 337).

Family-group name: AULACOPHORITES Chapuis, 1875. Stem: Aulacophor-.

Aulacopus Audinet-Serville, 1832: 144. Current status: valid genus in CERAMBYCIDAE: PRIONINAE: MACROTOMINI: MACROTOMINA.

Type species: Aulacopus reticulatus Audinet-Serville, 1832, by monotypy.

Family-group name: AULACOPINAE Kolbe, 1897. Stem: Aulacopod-.

Aulacoscelis Duponchel & Chevrolat, 1842a: 342. Current status: valid genus in ORSODACNIDAE: AULACOSCELIDINAE.

Type species: Aulacoscelis melanocera Duponchel & Chevrolat, 1842, by monotypy.

Family-group name: AULACOSCÉLITES Chapuis, 1874. Stem: Aulacoscelid-.

Aulacotrachelus Benick, 1921: 1. Current status: junior synonym of Megalopinus Eichelbaum, 1915 in STAPHYLINIDAE: MEGALOPSIDIINAE. Note: new replacement name for Megalops Erichson, 1839.

Type species: Oxyporus caelatus Gravenhorst, 1802, by subsequent designation (E. Desmarest 1852: 85) (type fixation for Megalops Erichson).

Family-group name: AULACOTRACHELINAE L. Benick, 1921. Stem: Aulacotrachel-.

Auletanus Voss, 1922a: 17, 20. Current status: valid genus in ATTELABIDAE: RHYNCHITINAE: RHYNCHITITAE: AULETINI: AULETOBIINA.

Type species: Auletobius ascendens Heller, 1915, by subsequent designation (Voss 1932: 101).

Family-group name: AULETANINI Legalov, 2003. Stem: Auletan-.

Auletes Schönherr, 1826: 46. Current status: valid genus in ATTELABIDAE: RHYNCHITINAE: RHYNCHITITAE: AULETINI: AULETINA.

Type species: Auletes tubicen Schönherr, 1826, by original designation. Note: combined description of a new nominal genus and a single new nominal species (ICZN 1999a, Article 12.2.6).

Family-group name: AULETINIDAE Desbrochers des Loges, 1908. Stem: Aulet-.

Auletobius Desbrochers des Loges, 1869: 396. Current status: valid genus in ATTELABIDAE: RHYNCHITINAE: RHYNCHITITAE: AULETINI: AULETOBIINA.

Type species: Auletes basilaris Gyllenhal, 1839 (= Involvulus sanguisorbae Schrank, 1798), by subsequent designation (Brauer 1870: 133).

Family-group name: AULETOBINI Voss, 1930. Stem: Auletobi-.

Auletorhinus Voss, 1935: 509. Current status: valid genus in ATTELABIDAE: RHYNCHITINAE: RHYNCHITITAE: AULETINI: AULETOBIINA.

Type species: Auletorhinus hirtellus Voss, 1935, by original designation.

Family-group name: AULETORHININI Voss, 1935. Stem: Auletorhin-.

Aulonocnemis Klug, 1837: 53. Current status: valid genus in SCARABAEIDAE: AULONOCNEMINAE.

Type species: Aulonocnemis opatrina Klug, 1837, by subsequent designation (R. Lucas 1920: 129).

Family-group name: AULONOCNEMINAE A. Janssens, 1946. Stem: Aulonocnem-.

Aulonogyrus Motschulsky, 1853c: 9. Current status: valid genus in GYRINIDAE: GYRININAE: DINEUTINI: ENHYDRUSINA.

Type species: Gyrinus strigipennis Suffrian, 1842 (= Gyrinus concinnus Klug, 1834), by subsequent designation (Ochs 1930: 3).

Family-group name: AULONOGYRINI Ochs, 1953. Stem: Aulonogyr-.

Aurelius Kuwert, 1891: 168. Current status: valid genus in PASSALIDAE: PASSALINAE: MACROLININI.

Type species: Aurelius dohrnii Kuwert, 1891, by monotypy.

Family-group name: AURELIINAE Kuwert, 1896. Stem: Aureli-.

Australestes Pace, 2015: 66. Current status: valid genus in STAPHYLINIDAE: ALEOCHARINAE: AUSTRALESTESINI.

Type species: Australestes crassum Pace, 2015, by original designation.

Family-group name: AUSTRALESTESINI Pace, 2015: 65. Stem: Australestes- (ICZN 1999a, Article 29.4).

Australiotyphlus Pace, 2013a: 99. Current status: valid genus in STAPHYLINIDAE: LEPTOTYPHLINAE: AUSTRALIOTYPHLINI.

Type species: Australiotyphlus cabbagensis Pace, 2013, by original designation.

Family-group name: AUSTRALIOTYPHLINI Pace, 2013a: 99. Stem: Australiotyphl-.

Australobolbus Howden & Cooper, 1977: 19. Current status: valid genus in BOLBOCERATIDAE: BOLBOCERATINAE: BOLBOCERATINI.

Type species: Bolboceras obscurius Blackburn, 1904, by original designation.

Family-group name: AUSTRALOBOLBINI Nikolajev, 1996. Stem: Australobolb-.

Austroesthetus Oke, 1933: 112. Current status: valid genus in STAPHYLINIDAE: EUAESTHETINAE: AUSTROESTHETINI. Note: the original spelling was Austroesthethus, but Austroesthetus, an incorrect subsequent spelling, is in prevailing usage, attributed to the publication of the original spelling, and therefore deemed to be the correct original spelling (ICZN 1999a, Article 33.3.1).

Type species: Austroesthetus passerculus Oke, 1933, by original designation.

Family-group name: AUSTROAESTHETINI Cameron, 1944. Stem: Austroesthet-.

Austropassalus Mjöberg, 1917: 11. Current status: valid genus in PASSALIDAE: PASSALINAE: MACROLININI.

Type species: Austropassalus hultgreni Mjöberg, 1917, by monotypy.

Family-group name: AUSTROPASSALINAE Mjöberg, 1917. Stem: Austropassal-.

Austrorhysus Steel, 1966: 297. Current status: valid genus in STAPHYLINIDAE: PROTEININAE: AUSTRORHYSINI.

Type species: Anepius raucus Blackburn, 1902, by original designation.

Family-group name: AUSTRORHYSINI Newton & Thayer, 1995. Stem: Austrorhys-.

Autalia Leach in Samouelle, 1819: 177. Current status: valid genus in STAPHYLINIDAE: ALEOCHARINAE: AUTALIINI.

Type species: Staphylinus impressus G.-A. Olivier, 1795, by subsequent designation (Westwood 1838c: 20).

Family-group name: AUTALIIDES C.G. Thomson, 1859. Stem: Autali-.

Automolius Britton, 1978: 7. Current status: valid genus in SCARABAEIDAE: SERICOIDINAE: AUTOMOLIINI. Note: new replacement name for Automolus Burmeister, 1855.

Type species: Automolus angustulus Burmeister, 1855, by monotypy (type fixation for Automolus Burmeister).

Family-group name: AUTOMOLIINI Britton, 1978. Stem: Automoli-.

Automolus Burmeister, 1855: 192. Current status: invalid senior synonym of Automolius Britton, 1978 in SCARABAEIDAE: SERICOIDINAE: AUTOMOLIINI. Note: junior homonym of Automolus L. Reichenbach, 1853 [Aves].

Type species: Automolus angustulus Burmeister, 1855, by monotypy.

Family-group name: AUTOMOLINI Britton, 1957. Stem: Automol-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

Auxenocerus Jeannel, 1962a: 324. Current status: valid genus in STAPHYLINIDAE: PSELAPHINAE: EUPLECTITAE: JUBINI.

Type species: Auxenocerus bispina Jeannel, 1962, by original designation.

Family-group name: AUXENOCERINI Jeannel, 1962. Stem: Auxenocer-.

Auxesis J. Thomson, 1858b: 196. Current status: valid genus in CERAMBYCIDAE: CERAMBYCINAE: AUXESINI.

Type species: Auxesis gabonicus J. Thomson, 1858, by monotypy.

Family-group name: AUXÉSIDES Lacordaire, 1872. Stem: Auxes-.

†Avitortor Ponomarenko, 1977: 42. Current status: valid genus in CROWN COLEOPTERA incertae sedis. Note: Lower Cretaceous (Russia).

Type species: †Avitortor primitivus Ponomarenko, 1977, by original designation.

Family-group name: †AVITORTORINAE Nikolajev, 2007. Stem: Avitortor-.

†Axelrodiellus Zherikhin & Gratshev, 2004: 66. Current status: valid genus in BRENTIDAE: EURHYNCHINAE: †AXELRODIELLINI. Note: Lower Cretaceous (Brazil).

Type species: †Axelrodiellus ruptus Zherikhin & Gratshev, 2004, by original designation.

Family-group name: †AXEIRODIELLINI Legalov, 2009. Stem: Axelrodiell-.

Axinidium Sturm, 1843: 327. Current status: valid genus in CARABIDAE: SCARITINAE: PROMECOGNATHINI.

Type species: Axinidium africanum Sturm, 1843, by monotypy.

Family-group name: AXINIDIINI Basilewsky, 1963. Stem: Axinidi-.

Axonya Andrewes, 1923b: 679. Current status: junior synonym of Broscodes Bolívar y Pieltain, 1914 in CARABIDAE: BROSCINAE: BROSCINI: AXONYINA.

Type species: Axonya championi Andrewes, 1923, by monotypy.

Family-group name: AXONYINA Roig-Juñent, 2000. Stem: Axony-.

Azya Mulsant, 1850: 928. Current status: valid genus in COCCINELLIDAE: COCCINELLINAE: COCCIDULINI.

Type species: Azya luteipes Mulsant, 1850, by subsequent designation (Crotch 1874a: 279).

Family-group name: AZYAIRES Mulsant, 1850. Stem: Azy-.

Babia Chevrolat in Dejean, 1836a: 417. Current status: valid genus in CHRYSOMELIDAE: CRYPTOCEPHALINAE: CLYTRINI: BABIINA.

Type species: Clytra quadriguttata G.-A. Olivier, 1791, by subsequent designation (Monrós 1953: 46).

Family-group name: BABIDEAE Lacordaire, 1848. Stem: Babi-.

Bacanius J.L. LeConte, 1853c: 291. Current status: valid genus in HISTERIDAE: DENDROPHILINAE: BACANIINI.

Type species: Bacanius tantillus J.L. LeConte, 1853, by subsequent designation (Bickhardt 1916: 71).

Family-group name: BACANIINI Kryzhanovskij, 1976. Stem: Bacani-.

Badister Clairville, 1806: 90, 91. Current status: valid genus in CARABIDAE: HARPALINAE: LICININI: LICININA.

Type species: Carabus bipustulatus Fabricius, 1792 (= Carabus bullatus Schrank, 1798), by monotypy.

Family-group name: BADISTIDAE Gistel, 1856. Stem: Badister-.

Baeocera Erichson, 1845a: 4. Current status: valid genus in STAPHYLINIDAE: SCAPHIDIINAE: SCAPHISOMATINI. Note: name placed on the Official List of Generic Names in Zoology (ICZN 1982).

Type species: Baeocera falsata Achard, 1920, by plenary power (see ICZN 1982). Note: name placed on the Official List of Specific Names in Zoology (ICZN 1982).

Family-group name: BAEOCERITAE Achard, 1924. Stem: Baeocer-.

Baeoceridium Reitter, 1889a: 6. Current status: valid genus in STAPHYLINIDAE: SCAPHIDIINAE: SCAPHISOMATINI.

Type species: Baeoceridium depressipes Reitter, 1889, by monotypy.

Family-group name: BAEOCERIDIITAE Achard, 1924. Stem: Baeoceridi-.

Bagous Germar, 1817b: 340. Current status: valid genus in CURCULIONIDAE: BAGOINAE.

Type species: Curculio binodulus Herbst, 1795, by subsequent designation (Schönherr 1826: 21).

Family-group name: BAGOINA C.G. Thomson, 1859. Stem: Bago-. Note: nomen protectum (see Bouchard et al. 2011: 586).

†Baissorhynchus Zherikhin, 1977a: 176. Current status: valid genus in CARIDAE: †BAISSORHYNCHINAE: †BAISSORHYNCHINI. Note: Lower Cretaceous (Russia).

Type species: †Baissorhynchus tarsalis Zherikhin, 1977, by original designation.

Family-group name: †BAISSORHYNCHINI Zherikhin, 1993. Stem: Baissorhynch-. Note: more research is needed to clarify the correct position of †BAISSORHYNCHINAE as well as the tribe therein before the priority problem with the older the name †NANOPHYDINI can be addressed.

Balaninus Germar, 1817b: 340. Current status: junior synonym of Curculio Linnaeus, 1758 in CURCULIONIDAE: CURCULIONINAE: CURCULIONINI: CURCULIONINA.

Type species: Curculio nucum Linnaeus, 1758, by subsequent designation (Partington 1834: 292).

Family-group name: BALANINIDAE Gistel, 1848. Stem: Balanin-.

Balanobius Jekel, 1861: 267. Current status: valid genus in CURCULIONIDAE: CURCULIONINAE: CURCULIONINI: BALANOBIINA. Note: this genus was listed as a junior synonym of “Archarius Gistel, 1856” by Alonso-Zarazaga and Lyal (1999: 73); however, “Archarius Gistel, 1856” is an incorrect spelling of Archarias A. Villa & G.B. Villa, 1833 (a junior homonym of Archarias Dejean, 1821 [Coleoptera: CURCULIONIDAE]) and therefore cannot be used as valid.

Type species: Curculio crux Fabricius, 1777, by subsequent designation (Voss 1958: 88).

Family-group name: BALANOBIINA Prena, 2021: 752. Stem: Balanobi-. Note: new replacement name for ARCHARIINA Pelsue & O’Brien, 2011.

Balgus Fleutiaux, 1920: 95, 97. Current status: valid genus in ELATERIDAE: THYLACOSTERNINAE. Note: this name was proposed simultaneously as a new replacement name for Galba Latreille, 1829 in EUCNEMIDAE, a junior homonym of Galba Schrank, 1803 [Mollusca], and as a new replacement name for Pterotarsus sensu Latreille, 1834 in ELATERIDAE (non Guérin-Méneville, 1829); this taxon is considered as valid in the family ELATERIDAE (e.g., Kundrata et al. 2019d).

Type species: Melasis tuberculosa Dalman, 1823, by original designation.

Family-group name: BALGINAE Fleutiaux, 1926. Stem: Balg-.

†Baltognathus Brunke, Żyła & Solodovnikov in Brunke et al., 2019: 179. Current status: valid genus in STAPHYLINIDAE: STAPHYLININAE: STAPHYLININI: †BALTOGNATHINA. Note: mid- Eocene (Baltic amber).

Type species: †Baltognathus aenigmaticus Brunke, Żyła & Solodovnikov, 2019, by original designation.

Family-group name: †BALTOGNATHINA Brunke, Żyła & Solodovnikov in Brunke et al., 2019: 175. Stem: Baltognath-.

†Baltostigus Jałoszyński, 2016: 1060. Current status: valid genus in STAPHYLINIDAE: SCYDMAENINAE: MASTIGITAE: †BALTOSTIGINI. Note: Eocene (Lithuania; Poland; Russia; Ukraine).

Type species: †Baltostigus antennatus Jałoszyński, 2016, by original designation.

Family-group name: †BALTOSTIGINI Jałoszyński & Brunke in Jałoszyński et al., 2018: 638. Stem: Baltostig-.

Barada Raffray, 1891: 314. Current status: junior synonym of Euphalepsus Reitter, 1883 in STAPHYLINIDAE: PSELAPHINAE: GONIACERITAE: BRACHYGLUTINI: BARADINA.

Type species: Barada mucronata Raffray, 1891, by monotypy.

Family-group name: BARADIINI Park, 1951. Stem: Barad-.

Baraeus J. Thomson, 1858b: 163. Current status: valid genus in CERAMBYCIDAE: LAMIINAE: PTEROPLIINI.

Type species: Baraeus aurisecator J. Thomson, 1858, by monotypy.

Family-group name: BAROEIDES Lacordaire, 1872. Stem: Barae-.

Baripus Dejean, 1828: 24. Current status: valid genus in CARABIDAE: BROSCINAE: BROSCINI: BARIPODINA.

Type species: Molops rivalis Germar, 1823, by monotypy.

Family-group name: BARYPITAE Jeannel, 1941. Stem: Baripod-.

Baris Germar, 1817b: 340. Current status: valid genus in CURCULIONIDAE: BARIDINAE: BARIDINI: BARIDINA.

Type species: Curculio artemisiae Herbst, 1795 (= Curculio artemisiae Panzer, 1794), by subsequent designation (Curtis 1839: pl. 766).

Family-group name: BARIDIDES Schönherr, 1836. Stem: Barid-.

Barrosellus Jeannel, 1951a: 107. Current status: valid genus in STAPHYLINIDAE: PSELAPHINAE: GONIACERITAE: BARROSELLINI.

Type species: Barrosellus sphaeroides Jeannel, 1951, by original designation.

Family-group name: BARROSELLINI Leleup, 1973. Stem: Barrosell-.

Barymela Weise, 1910a: 459. Current status: valid genus in CHRYSOMELIDAE: CHRYSOMELINAE: CHRYSOMELINI.

Type species: Barymela scutellaris Weise, 1910, by monotypy.

Family-group name: BARYMELINI Bechyné, 1948. Stem: Barymel-.

Barymerus Lacordaire, 1865: 259. Current status: valid genus in CURCULIONIDAE: BARIDINAE: MADARINI: BARYMERINA. Note: new replacement name for Physomerus Schönherr, 1844.

Type species: Physomerus calandroides Boheman, 1844, by original designation (type fixation for Physomerus Schönherr).

Family-group name: BARYMÉRIDES Lacordaire, 1865. Stem: Barymer-.

Barynotus Germar, 1817b: 341. Current status: valid genus in CURCULIONIDAE: ENTIMINAE: POLYDRUSITAE: GEONEMINI.

Type species: Curculio obscurus Fabricius, 1775, by subsequent designation (Schönherr 1823: 1142). Note: nomen protectum (see Alonso-Zarazaga 2013a: 68).

Family-group name: BARYNOTIDES Lacordaire, 1863. Stem: Barynot-.

Baryodirus Perreau, 2000: 19. Current status: valid genus in LEIODIDAE: CHOLEVINAE: PTOMAPHAGINI: BARYODIRINA.

Type species: Baryodirus hammondi Perreau, 2000, by original designation.

Family-group name: BARYODIRINA Perreau, 2000. Stem: Baryodir-.

Barypeithes Jacquelin du Val, 1854: 13. Current status: valid genus in CURCULIONIDAE: ENTIMINAE: CYPHICERITAE: OMIINI.

Type species: Barypeithes rufipes Jacquelin du Val, 1854 (= Omias sulcifrons Boheman, 1843), by subsequent designation (Lacordaire 1863: 59).

Family-group name: BARYPEITHINI Pic, 1897: 298. Stem: Barypeith-. Note: this name was noted by Alonso-Zarazaga (2013b: 71) but overlooked in Bouchard et al. (2011), Bouchard and Bousquet (2020a).

Bascanus Péringuey, 1896: 539, 540. Current status: valid genus in CARABIDAE: HARPALINAE: BASCANINI.

Type species: Bascanus gracilis Péringuey, 1896, by subsequent designation (Basilewsky 1953: 47).

Family-group name: BASCANINI Basilewsky, 1953. Stem: Bascan-.

Basilepta Baly, 1860: 23. Current status: valid genus in CHRYSOMELIDAE: EUMOLPINAE: TYPOPHORINI.

Type species: Basilepta longipes Baly, 1860, by original designation.

Family-group name: BASILEPTINI Chûjô, 1956. Stem: Basilept-.

Basiprionota Chevrolat in Dejean, 1836a: 367. Current status: valid genus in CHRYSOMELIDAE: CASSIDINAE: BASIPRIONOTINI.

Type species: Cassida octopunctata Fabricius, 1787, by monotypy.

Family-group name: BASIPRIONOTINI Gressitt, 1952. Stem: Basiprionot-.

Basipta Chevrolat, 1842a: 489. Current status: valid genus in CHRYSOMELIDAE: CASSIDINAE: CASSIDINI.

Type species: Basipta glauca Chevrolat, 1842, by monotypy. Note: combined description of a new nominal genus and a single new nominal species (ICZN 1999a, Article 12.2.6).

Family-group name: BASIPTITES Chapuis, 1875. Stem: Basipt-.

Basiptera J. Thomson, 1864: 204. Current status: valid genus in CERAMBYCIDAE: CERAMBYCINAE: BASIPTERINI.

Type species: Basiptera castaneipennis J. Thomson, 1864, by original designation.

Family-group name: BASIPTERINI Fragoso, Monné & Campos Seabra, 1987. Stem: Basipter-.

Basitoxus Audinet-Serville, 1832: 174. Current status: valid genus in CERAMBYCIDAE: PRIONINAE: MACROTOMINI: BASITOXINA.

Type species: Basitoxus armatus Audinet-Serville, 1832 (= Prionus megacephalus Germar, 1823), by subsequent designation (E. Desmarest 1860: 307).

Family-group name: BASITOXI Lameere, 1912. Stem: Basitox-.

Basitropis Jekel, 1855: 90. Current status: valid genus in ANTHRIBIDAE: ANTHRIBINAE: BASITROPINI.

Type species: Basitropis nitidicutis Jekel, 1855, by original designation. Note: the type species was listed as nitidicollis on page 91, which was corrected to nitidicutis on page 237 in the same work (Jekel 1855).

Family-group name: BASITROPIDES Lacordaire, 1865. Stem: Basitrop-.

Bathyris J.L. LeConte, 1874a: 462. Current status: junior synonym of Colecerus Schönherr, 1840 in CURCULIONIDAE: ENTIMINAE: ENTIMITAE: EUDIAGOGINI.

Type species: Bathyris dispar J.L. LeConte, 1874, by monotypy.

Family-group name: BATHYRINI J.L. LeConte, 1874. Stem: Bathyrin-.

Bathyscia Schiødte, 1848: 79. Current status: valid genus in LEIODIDAE: CHOLEVINAE: LEPTODIRINI: BATHYSCIINA.

Type species: Bathyscia montana Schiødte, 1848, by subsequent designation (Jeannel 1910: 16).

Family-group name: BATHYSCIAE G.H. Horn, 1880. Stem: Bathysci-.

Bathysciotes Jeannel, 1910: 15. Current status: valid genus in LEIODIDAE: CHOLEVINAE: LEPTODIRINI: BATHYSCIOTINA.

Type species: Adelops khevenhuelleri L. Miller, 1852, by original designation.

Family-group name: BATHYSCIOTINA Guéorguiev, 1974. Stem: Bathysciot-.

Batobius Fairmaire & Germain, 1863: 268. Current status: valid genus in MYCTERIDAE: EURYPINAE.

Type species: Batobius ruficollis Fairmaire & Germain, 1863, by subsequent designation (Spilman 1954: 87).

Family-group name: BATOBIINA Seidlitz, 1917. Stem: Batobi-.

Batocera Dejean, 1835: 341. Current status: valid genus in CERAMBYCIDAE: LAMIINAE: BATOCERINI.

Type species: Cerambyx rubus Linnaeus, 1758, by subsequent designation (Blanchard 1845a: 175).

Family-group name: BATOCERITAE J. Thomson, 1864. Stem: Batocer-.

Batonota Hope, 1839: 98. Current status: junior synonym of Dorynota Chevrolat, 1836 in CHRYSOMELIDAE: CASSIDINAE: DORYNOTINI.

Type species: Cassida bidens Fabricius, 1781, by original designation.

Family-group name: BATONOTITAE Spaeth, 1923. Stem: Batonot-.

Batrisus Aubé, 1833: 12, 45. Current status: valid genus in STAPHYLINIDAE: PSELAPHINAE: BATRISITAE: BATRISINI: BATRISINA.

Type species [suggested]: Batrisus formicarius Aubé, 1833, by subsequent designation (Duponchel 1842a: 506). Note: the valid type species fixation is that of Westwood (1838c: 21), who selected Bryaxis nigriventris Denny, 1825 (= Pselaphus venustus H.G.L. Reichenbach, 1816), a species currently placed in the genus Batrisodes Reitter, 1882 (e.g., see Newton and Chandler 1989: 8); an application to the Commission is necessary to fix Batrisus formicarius Aubé, 1833 as the type species of Batrisus Aubé, 1833.

Family-group name: BATRISINI Reitter, 1882. Stem: Batris-.

Batulius J.L. LeConte, 1851a: 148. Current status: valid genus in TENEBRIONIDAE: PIMELIINAE: ANEPSIINI.

Type species: Batulius setosus J.L. LeConte, 1851, by subsequent designation (Casey 1907: 497).

Family-group name: BATULIINI G.H. Horn, 1870. Stem: Batuli-.

Beccaria Gorham, 1885b: 521. Current status: invalid senior synonym of Beccariola Arrow, 1943 in ENDOMYCHIDAE: LYCOPERDININAE. Note: junior homonym of Beccaria Trinchese, 1870 [Mollusca].

Type species: Beccaria papuensis Gorham, 1885, by monotypy.

Family-group name: BECCARIINI Arrow, 1925. Stem: Beccari-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

Belionota Eschscholtz, 1829b: 8, 9. Current status: valid genus in BUPRESTIDAE: BUPRESTINAE: ACTENODINI.

Type species: Belionota sagittaria Eschscholtz, 1829, by monotypy.

Family-group name: BELIONOTINA Jakobson, 1913. Stem: Belionot-.

Beliophorus Eschscholtz, 1829a: 34. Current status: valid genus in ELATERIDAE incertae sedis.

Type species [suggested]: Beliophorus cebrionoides Eschscholtz, 1829, by subsequent designation (Lacordaire 1857: 162). Note: the valid type species fixation is that of Duponchel (1842a: 532), who selected Elater mucronatus G.-A. Olivier, 1792, but that species is the type species of Oxynopterus Hope, 1842; an application to the Commission is necessary to fix Beliophorus cebrionoides Eschscholtz, 1829 as the type species of Beliophorus Eschscholtz, 1829.

Family-group name: BELIOPHORINA Jakobson, 1913. Stem: Beliophor-.

Belohina Paulian, 1959: 42. Current status: valid genus in BELOHINIDAE.

Type species: Belohina inexpectata Paulian, 1959, by original designation.

Family-group name: BELOHININAE Paulian, 1959. Stem: Belohin-.

Belopherus Schönherr, 1833: 6, 334. Current status: junior synonym of Belorhynchus Berthold, 1827 in BRENTIDAE: BRENTINAE: BRENTINI: ARRENODINA.

Type species: Brentus militaris G.-A. Olivier, 1807, by original designation.

Family-group name: BÉLOPHÉRIDES Lacordaire, 1865. Stem: Belopher-.

Belopus Gebien, 1911: 459. Current status: valid genus in TENEBRIONIDAE: LAGRIINAE: BELOPINI. Note: nomen protectum (see Bouchard and Bousquet 2020b: 6); new replacement name for Calcar Dejean, 1821.

Type species: Tenebrio elongatus Herbst, 1797, by monotypy (type fixation for Calcar Dejean).

Family-group name: BELOPINAE Reitter, 1917. Stem: Belop-.

Belorhynchus Berthold, 1827: 384. Current status: valid genus in BRENTIDAE: BRENTINAE: BRENTINI: ARRENODINA.

Type species: Curculio nasutus Fabricius, 1781, by subsequent monotypy (Latreille 1828b: 595).

Family-group name: BÉLORHYNCHIDES Lacordaire, 1865. Stem: Belorhynch-.

Belus Schönherr, 1823: 1137. Current status: junior synonym of Rhinotia W. Kirby, 1819 in BELIDAE: BELINAE: BELITAE: BELINI: BELINA.

Type species: Curculio semipunctatus Fabricius, 1775, by original designation.

Family-group name: BELIDES Schönherr, 1826. Stem: Bel-.

Bembidarenas Erwin, 1972: 8. Current status: valid genus in CARABIDAE: TRECHINAE: BEMBIDARENINI.

Type species: Bembidion reicheellum Csiki, 1929, by original designation.

Family-group name: BEMBIDARENINI Maddison & Erwin in Maddison et al., 2019: 173. Stem: Bembidaren-.

Bembidion Latreille, 1802b: 82. Current status: valid genus in CARABIDAE: TRECHINAE: BEMBIDIINI.

Type species: Carabus quadriguttatus Fabricius, 1775 [p. 248] (= Cicindela quadrimaculata Linnaeus, 1761), by subsequent designation (Andrewes 1935: 17). Note: Andrewes (1935: 17) designated Cicindela quadrimaculata Linnaeus, 1761 as type species of Bembidion Latreille, 1802, a species not originally included, but since he listed that species in synonymy with Carabus quadriguttatus Fabricius, 1775, a species originally included in Bembidion, he is deemed to have designated the latter species as type species (ICZN 1999a, Article 69.2.2).

Family-group name: BEMBIDIIDAE Stephens, 1827. Stem: Bembidi-.

†Berendtimirus J.R. Winkler, 1987: 52. Current status: valid genus in †BERENDTIMIRIDAE: †BERENDTIMIRINAE. Note: [geological time not mentioned] (Baltic amber).

Type species: †Berendtimirus progenitor J.R. Winkler, 1987, by original designation.

Family-group name: †BERENDTIMIRIDAE J.R. Winkler, 1987. Stem: Berendtimir-.

Berginus Erichson, 1846: 405. Current status: valid genus in MYCETOPHAGIDAE: BERGININAE.

Type species: Berginus tamarisci Erichson, 1846, by monotypy. Note: this species is usually credited to Wollaston (1854: 195) in the literature but Erichson (1846), when describing the genus Berginus in his key, listed one species by name, providing therefore a combined description of a new nominal genus and a new nominal species (ICZN 1999a, Article 12.2.6).

Family-group name: BERGINI Leng, 1920. Stem: Bergin-.

Berosus Leach, 1817: 92. Current status: valid genus in HYDROPHILIDAE: HYDROPHILINAE: BEROSINI. Note: name placed on the Official List of Generic Names in Zoology (ICZN 1990a).

Type species: Dytiscus luridus Linnaeus, 1761, by monotypy (see ICZN 1990a). Note: name placed on the Official List of Specific Names in Zoology (ICZN 1990a).

Family-group name: BÉROSAIRES Mulsant, 1844. Stem: Beros-.

Betschia Dajoz, 1980a: 135. Current status: valid genus in TENEBRIONIDAE: DIAPERINAE: GNATHIDIINI: BETSCHIINA.

Type species: Betschia minuta Dajoz, 1980, by original designation.

Family-group name: BETSCHIINI Dajoz, 1980. Stem: Betschi-.

Bibloplectus Reitter, 1882b: 529. Current status: valid genus in STAPHYLINIDAE: PSELAPHINAE: EUPLECTITAE: TRICHONYCHINI: PANAPHANTINA.

Type species: Pselaphus ambiguus H.G.L. Reichenbach, 1816, by subsequent designation (R. Lucas 1920: 137).

Family-group name: BIBLOPLECTINA Jeannel, 1959. Stem: Bibloplect-.

Bibloporellus Jeannel, 1949a: 47. Current status: valid genus in STAPHYLINIDAE: PSELAPHINAE: EUPLECTITAE: TRICHONYCHINI: BIBLOPORINA.

Type species: Bibloporellus microphthalmus Jeannel, 1949, by original designation.

Family-group name: BIBLOPORELLINA Jeannel, 1952. Stem: Bibloporell-.

Bibloporus C.G. Thomson, 1859: 53. Current status: valid genus in STAPHYLINIDAE: PSELAPHINAE: EUPLECTITAE: TRICHONYCHINI: BIBLOPORINA.

Type species: Euplectus bicolor Denny, 1825, by original designation.

Family-group name: BIBLOPORINI Park, 1951. Stem: Biblopor-.

Bidessus Sharp, 1880: cxlviii. Current status: valid genus in DYTISCIDAE: HYDROPORINAE: BIDESSINI. Note: name placed on the Official List of Generic Names in Zoology (ICZN 2021a).

Type species: Dytiscus unistriatus Schrank, 1781, by plenary power (see ICZN 2021a). Note: name placed on the Official List of Specific Names in Zoology (ICZN 2021a).

Family-group name: BIDESSINI Sharp, 1880. Stem: Bidess-.

Bimia A. White, 1850: 13. Current status: valid genus in CERAMBYCIDAE: CERAMBYCINAE: BIMIINI.

Type species: Bimia bicolor A. White, 1850, by monotypy.

Family-group name: BIMIIDES Lacordaire, 1868. Stem: Bimi-.

Biophytus Guillebeau, 1894: 279, 295. Current status: junior synonym of Pseudolibrus Flach, 1889 in PHALACRIDAE: PHALACRINAE.

Type species: Biophytus grouvellei Guillebeau, 1894, by original designation.

Family-group name: BIOPHYTINI Guillebeau, 1894. Stem: Biophyt-.

Bioplanes Mulsant, 1854: 144. Current status: valid genus in TENEBRIONIDAE: BLAPTINAE: DENDARINI: DENDARINA.

Type species: Bioplanes meridionalis Mulsant, 1854, by monotypy.

Family-group name: BIOPLANINA Reikhardt, 1936. Stem: Bioplanet-.

Biphyllus Dejean, 1821: 102. Current status: valid genus in BIPHYLLIDAE.

Type species: Dermestes lunatus Fabricius, 1787, by monotypy.

Family-group name: DIPHYLLIDAE J.L. LeConte, 1861. Stem: Biphyll-.

Bitoma Herbst, 1793: 25. Current status: valid genus in ZOPHERIDAE: COLYDIINAE: SYNCHITINI.

Type species: Tritoma crenata Fabricius, 1775, by subsequent designation (Latreille 1810: 431). Note: even if Latreille’s designation was for Ditoma (as “Ditome”), an unjustified emendation of Bitoma by Illiger (1807b: 320), both available genus-group names have the same type species (ICZN 1999a, Article 67.8).

Family-group name: DITOMIDAE Imhoff, 1856. Stem: Bitom-.

Bius Dejean, 1834: 205. Current status: valid genus in TENEBRIONIDAE: TENEBRIONINAE: TENEBRIONINI.

Type species: Trogosita thoracicus Fabricius, 1792, by monotypy.

Family-group name: BIUINI Skopin, 1978. Stem: Bi-.

Blaesia Burmeister, 1842: 615. Current status: valid genus in SCARABAEIDAE: CETONIINAE: GYMNETINI: BLAESIINA.

Type species: Blaesia atra Burmeister, 1842, by monotypy.

Family-group name: BLAESIAE Schoch, 1895. Stem: Blaesi-.

Blaps Fabricius, 1775: 254. Current status: valid genus in TENEBRIONIDAE: BLAPTINAE: BLAPTINI: BLAPTINA.

Type species: Tenebrio mortisagus Linnaeus, 1758, by subsequent designation (Latreille 1810: 429).

Family-group name: BLAPSIDA Leach, 1815. Stem: Blapt-.

Blapstinus Dejean, 1821: 66. Current status: valid genus in TENEBRIONIDAE: BLAPTINAE: OPATRINI: BLAPSTININA.

Type species: Blaps punctata Fabricius, 1792, by monotypy.

Family-group name: BLAPSTINITES Mulsant & Rey, 1853. Stem: Blapstin-.

Blastophila Gistel, 1856: 370. Current status: junior synonym of Trachodes Germar, 1823 in CURCULIONIDAE: MOLYTINAE: TRACHODINI.

Type species: Curculio hispidus Linnaeus, 1758, by monotypy.

Family-group name: BLASTOPHILADAE Gistel, 1856. Stem: Blastophil-.

Bledius Leach in Samouelle, 1819: 174. Current status: valid genus in STAPHYLINIDAE: OXYTELINAE: BLEDIINI.

Type species: Staphylinus armatus Panzer, 1799 (= Staphylinus tricornis Herbst, 1784), by monotypy.

Family-group name: BLEDIINI Ádám, 2001. Stem: Bledi-.

Blepharhymenus Solier, 1849: 339. Current status: valid genus in STAPHYLINIDAE: ALEOCHARINAE: OXYPODINI: DINARDINA.

Type species: Blepharhymenus sulcicollis Solier, 1849, by monotypy.

Family-group name: BLEPHARRHYMENI Klimaszewski & Peck, 1986. Stem: Blepharhymen-.

Blepharida Chevrolat in Dejean, 1836a: 394. Current status: valid genus in CHRYSOMELIDAE: GALERUCINAE: ALTICINI.

Type species: Chrysomela meticulosus G.-A. Olivier, 1808 (= Chrysomela rhois Forster, 1771 [as rhoi]), by subsequent designation (Chevrolat 1842a: 606).

Family-group name: BLÉPHARIDITES Chapuis, 1875. Stem: Blepharid-.

Bleusea Bedel, 1897: 345. Current status: valid genus in CARABIDAE: HARPALINAE: HARPALINI: HARPALINA.

Type species: Bleusea deserticola Bedel, 1897, by monotypy.

Family-group name: BLEUSEI Antoine, 1959. Stem: Bleuse-.

Blosyrus Schönherr, 1823: 1139. Current status: valid genus in CURCULIONIDAE: ENTIMINAE: POLYDRUSITAE: BLOSYRINI.

Type species: Curculio oniscus G.-A. Olivier, 1807, by original designation.

Family-group name: BLOSYRIDES Lacordaire, 1863. Stem: Blosyr-.

Boganium Sen Gupta & Crowson, 1966: 70. Current status: valid genus in BOGANIIDAE: BOGANIINAE.

Type species: Boganium armstrongi Sen Gupta & Crowson, 1966, by original designation.

Family-group name: BOGANIIDAE Sen Gupta & Crowson, 1966. Stem: Bogani-.

Bolbelasmus Boucomont, 1911: 335. Current status: valid genus in BOLBOCERATIDAE: BOLBOCERATINAE: BOLBELASMINI.

Type species: Bolbocerus bocchus Erichson, 1841, by subsequent designation (Sharp 1912: 205). Note: the type species usually listed for this genus is Bolboceras gallicum Mulsant, 1842 (e.g., Branco 2007: 8; Nikolajev et al. 2016: 33) by subsequent designation in Cartwright (1953: 97) but Sharp’s fixation is older; both nominal species are currently included in the nominotypical subgenus of Bolbelasmus.

Family-group name: BOLBELASMINI Iablokoff-Khnzorian, 1977. Stem: Bolbelasm-.

Bolboceras W. Kirby, 1819: 459. Current status: valid genus in BOLBOCERATIDAE: BOLBOCERATINAE: BOLBOCERATINI. Note: name not given priority over Odonteus Samouelle, 1819 whenever the two are considered synonyms and Bolboceras W. Kirby, 1819 placed on the Official List of Generic Names in Zoology (ICZN 2006a).

Type species: Scarabaeus quadridens Fabricius, 1781, by plenary power (see ICZN 2006a). Note: name placed on the Official List of Specific Names in Zoology (ICZN 2006a).

Family-group name: BOLBOCÉRAIRES Mulsant, 1842. Stem: Bolbocerat-.

Bolbochromus Boucomont, 1909: 117. Current status: valid genus in BOLBOCERATIDAE: BOLBOCERATINAE: BOLBOCHROMINI.

Type species: Bolboceras laetus Westwood, 1852, by subsequent designation (Sharp 1911: 231).

Family-group name: BOLBOCHROMINI Nikolajev, 1970. Stem: Bolbochrom-.

†Boleopsis Kirejtshuk & Nel, 2013: 189. Current status: valid genus in †BOLEOPSIDAE. Note: lower Eocene (France).

Type species: †Boleopsis polinae Kirejtshuk & Nel, 2013, by original designation.

Family-group name: †BOLEOPSIDAE Kirejtshuk & Nel, 2013: 185. Stem: Boleops-.

Bolitobius Leach in Samouelle, 1819: 176. Current status: valid genus in STAPHYLINIDAE: MYCETOPORINAE.

Type species: fixed by Schülke (1999: 982) according to Article 70.3.2 (ICZN 1999a) as Megacronus castaneus Stephens, 1832, the taxonomic species involved in the misidentification of Staphylinus analis Fabricius, 1787 (sensu Paykull, 1789) in the original designation by Samouelle (1819: 176). Note: the fixation of the type species by Schülke (1999: 982), although published before the start of the fourth edition of the Code, is valid (see ICZN 1999a, Article 86.1.1).

Family-group name: BOLITOBII G.H. Horn, 1877. Stem: Bolitobi-.

Bolitochara Mannerheim, 1830: 75. Current status: valid genus in STAPHYLINIDAE: ALEOCHARINAE: HOMALOTINI: BOLITOCHARINA. Note: name placed on the Official List of Generic Names in Zoology (ICZN 1961a).

Type species: Aleochara pulchra Gravenhorst, 1806, by plenary power (see ICZN 1961a). Note: name placed on the Official List of Specific Names in Zoology (ICZN 1961a).

Family-group name: BOLITOCHARIDES C.G. Thomson, 1859. Stem: Bolitochar-.

Bolitophagus Illiger, 1798: 100. Current status: valid genus in TENEBRIONIDAE: TENEBRIONINAE: BOLITOPHAGINI.

Type species [suggested]: Silpha reticulata Linnaeus, 1767, by subsequent designation (C.G. Thomson 1859: 115). Note: the valid type species fixation is that of Curtis (1836: pl. 586), who selected Opatrum agricola Herbst, 1783, which is the type species of the genus Eledona Latreille, 1797 (see Bouchard and Bousquet 2020b: 8); an application to the Commission is necessary to fix Silpha reticulata Linnaeus, 1767 as the type species of Bolitophagus Illiger, 1798.

Family-group name: BOLITOPHAGIDAE W. Kirby, 1837. Stem: Bolitophag-. Note: nomen protectum (see Bouchard et al. 2011: 415).

Bonesia Baly, 1865b: 100. Current status: valid genus in CHRYSOMELIDAE: GALERUCINAE: HYLASPINI.

Type species: Bonesia clarkii Baly, 1865, by original designation.

Family-group name: BONESIITES Laboissière, 1926. Stem: Bonesi-.

Borborophorus Hansen, 1990: 321, 326. Current status: valid genus in HYDROPHILIDAE: CYLOMINAE.

Type species: Borborophorus pubescens Hansen, 1990, by original designation.

Family-group name: BORBOROPHORINI Hansen, 1991. Stem: Borborophor-.

Boreaphilus C.R. Sahlberg, 1832: 433. Current status: valid genus in STAPHYLINIDAE: OMALIINAE: CORYPHIINI: BOREAPHILINA.

Type species: Boreaphilus henningianus C.R. Sahlberg, 1832, by monotypy.

Family-group name: BOREAPHILINA Zerche, 1990. Stem: Boreaphil-.

Boreocypha Klimaszewski & Langor in Klimaszewski et al., 2011: 79. Current status: valid genus in STAPHYLINIDAE: ALEOCHARINAE: BOREOCYPHINI.

Type species: Boreocypha websteri Klimaszewski & Langor, 2011, by original designation.

Family-group name: BOREOCYPHINI Klimaszewski & Langor in Klimaszewski et al., 2011: 79. Stem: Boreocyph-.

Boromorphus Wollaston, 1854: 492. Current status: valid genus in TENEBRIONIDAE: PIMELIINAE: BOROMORPHINI.

Type species: Boromorphus maderae Wollaston, 1854 (= Boros tagenioides P.H. Lucas, 1846), by monotypy.

Family-group name: BOROMORPHINI Skopin, 1978. Stem: Boromorph-.

Boros Herbst, 1797: 318. Current status: valid genus in BORIDAE: BORINAE.

Type species: Boros elongatus Herbst, 1797 (= Helops schneideri Panzer, 1796), by monotypy.

Family-group name: BORIDAE C.G. Thomson, 1859. Stem: Bor-.

Bostrichus Geoffroy, 1762: 301. Current status: valid genus in BOSTRICHIDAE: BOSTRICHINAE: BOSTRICHINI. Note: name placed on the Official List of Generic Names in Zoology (ICZN 1994a).

Type species: Dermestes capucinus Linnaeus, 1758, by subsequent designation (Latreille 1810: 431; see ICZN 1994a). Note: name placed on the Official List of Specific Names in Zoology (ICZN 1994a).

Family-group name: BOSTRICHINI Latreille, 1802. Stem: Bostrich-.

Bostrychopsis Lesne, 1899: 444, 524. Current status: valid genus in BOSTRICHIDAE: BOSTRICHINAE: BOSTRICHINI.

Type species: fixed by Ivie (2010: 29) according to Article 70.3.2 (ICZN 1999a) as Bostrychopsis villosula Lesne, 1905, the taxonomic species involved in the misidentification of Bostrichus cephalotes G.-A. Olivier, 1790 (sensu Lesne, 1899), as designated subsequently by Chûjô (1937: 52). Note: see Lesne (1905: 298) for a discussion about the misidentification of the type species.

Family-group name: BOSTRYCHOPSINI Lesne, 1921. Stem: Bostrychopse-.

Bothrideres Dejean, 1835: 312. Current status: valid genus in BOTHRIDERIDAE.

Type species: Dermestes contractus Fourcroy, 1785, by monotypy. Note: the species listed by Dejean (1835: 312) is “contractus Fabr.”, which corresponds to Lyctus contractus Fabricius, 1792 where there is a reference to Ips contracta G.-A. Olivier, 1790, where, in turn, there is a reference to Dermestes contractus Fourcroy, 1785; therefore, Fourcroy’s species is the type species.

Family-group name: BOTHRIDERINI Erichson, 1845. Stem: Bothrider-.

Bothriophorus Mulsant & Rey, 1853a: 19. Current status: valid genus in LIMNICHIDAE: LIMNICHINAE: LIMNICHINI.

Type species: Bothriophorus atomus Mulsant & Rey, 1853, by monotypy.

Family-group name: BOTRIOPHORATES Mulsant & Rey, 1869. Stem: Bothriophor-.

Bothriospila Aurivillius, 1924: 1. Current status: valid genus in CERAMBYCIDAE: CERAMBYCINAE: BOTHRIOSPILINI.

Type species: Bothriospila elegans Aurivillius, 1924, by monotypy.

Family-group name: BOTHRIOSPILINAE Lane, 1950. Stem: Bothriospil-.

Bothrorrhina Burmeister, 1842: 200. Current status: junior synonym of Plaesiorrhina Westwood, 1842 in SCARABAEIDAE: CETONIINAE: GOLIATHINI: RHOMBORHININA.

Type species: Cetonia reflexa Gory & Percheron, 1835, by subsequent designation (Pouillaude 1916: 56).

Family-group name: BOTHRORRHINAE Schoch, 1894. Stem: Bothrorrhin-.

Bothrosternus Eichhoff, 1868a: 150. Current status: valid genus in CURCULIONIDAE: SCOLYTINAE: BOTHROSTERNINI.

Type species: Bothrosternus truncatus Eichhoff, 1868, by monotypy.

Family-group name: BOTHROSTERNINI W.F.H. Blandford, 1896. Stem: Bothrostern-.

Bothynus Hope, 1837: 95. Current status: valid genus in SCARABAEIDAE: DYNASTINAE: PENTODONTINI: PENTODONTINA. Note: name placed on the Official List of Generic Names in Zoology (ICZN 2008a).

Type species: Scarabaeus ascanius W. Kirby, 1819, by subsequent designation (Lacordaire 1855: 413; see ICZN 2008a). Note: name placed on the Official List of Specific Names in Zoology (ICZN 2008a).

Family-group name: BOTHYNIDAE Burmeister, 1847. Stem: Bothyn-.

Botryonopa Guérin-Méneville, 1840b: 332. Current status: valid genus in CHRYSOMELIDAE: CASSIDINAE: BOTRYONOPINI. Note: the original spelling was Bothryonopa but the incorrect subsequent spelling Botryonopa is in prevailing usage, attributed to the original publication, and therefore deemed to be the valid name (ICZN 1999a, Article 33.3.1).

Type species: Botryonopa sanguinea Guérin-Méneville, 1840, by subsequent designation (Baly 1858: 92).

Family-group name: BOTRYONOPITES Chapuis, 1875. Stem: Botryonop-.

Brachiacantha Chevrolat in Dejean, 1836a: 434. Current status: valid genus in COCCINELLIDAE: COCCINELLINAE: HYPERASPIDINI: BRACHIACANTHINA.

Type species: Coccinella dentipes Fabricius, 1801, by subsequent designation (Crotch 1873c: 377).

Family-group name: BRACHYACANTHAIRES Mulsant, 1850. Stem: Brachiacanth-.

Brachinus Weber, 1801: 22. Current status: valid genus in CARABIDAE: BRACHININAE: BRACHININI: BRACHININA.

Type species: Carabus crepitans Linnaeus, 1758, by subsequent designation (Latreille 1810: 426).

Family-group name: BRACHINII Bonelli, 1810. Stem: Brachin-.

Brachonyx Schönherr, 1825: 583. Current status: valid genus in CURCULIONIDAE: CURCULIONINAE: ANTHONOMINI.

Type species: Curculio indigena Herbst, 1795 (= Curculio pineti Paykull, 1792), by original designation.

Family-group name: BRACHONYCHINAE Joy, 1932. Stem: Brachonych-.

†Brachycamacus Poinar, Legalov & A.E. Brown, 2013: 2. Current status: valid genus in CURCULIONIDAE: ENTIMINAE: POLYDRUSITAE: EUSTYLINI: †BRACHYCAMACINA. Note: lower Miocene (Dominican Republic)

Type species: †Brachycamacus gyrommatus Poinar, Legalov & A.E. Brown, 2013, by original designation.

Family-group name: †BRACHYCAMACINA Poinar, Legalov & A.E. Brown, 2013: 2. Stem: Brachycamac-.

Brachyceropsis Aurivillius, 1886: 24. Current status: valid genus in CURCULIONIDAE: MOLYTINAE: BRACHYCEROPSEINI.

Type species: Brachycerus tuberculosus Gyllenhal, 1833, by original designation.

Family-group name: BRACHYCEROPSEINAE Aurivillius, 1926. Stem: Brachyceropse-.

Brachycerus G.-A. Olivier, 1789a: 36. Current status: valid genus in CURCULIONIDAE: BRACHYCERINAE: BRACHYCERINI.

Type species: Curculio barbarus Linnaeus, 1758, by subsequent designation (Latreille 1810: 430). Note: no species were originally included in this genus by G.-A. Olivier (1789a: 36); the first ones were the 20 species listed by G.-A. Olivier (1790b: 181, 182).

Family-group name: BRACHYCERIDES Billberg, 1820. Stem: Brachycer-.

Brachyderes Schönherr, 1823: 1140. Current status: valid genus in CURCULIONIDAE: ENTIMINAE: POLYDRUSITAE: BRACHYDERINI. Note: name placed on the Official List of Generic Names in Zoology (ICZN 1987a).

Type species: Curculio incanus Linnaeus, 1758, by original designation (see ICZN 1987a). Note: name placed on the Official List of Specific Names in Zoology (ICZN 1987a).

Family-group name: BRACHYDERIDES Schönherr, 1826. Stem: Brachyder-.

Brachygluta C.G. Thomson, 1859: 54. Current status: valid genus in STAPHYLINIDAE: GONIACERITAE: BRACHYGLUTINI: BRACHYGLUTINA.

Type species: Pselaphus fossulatus H.G.L. Reichenbach, 1816, by original designation.

Family-group name: BRACHYGLUTINI Raffray, 1904. Stem: Brachyglut-.

Brachygnathus Perty, 1830: 6. Current status: valid genus in CARABIDAE: HARPALINAE: PANAGAEINI: BRACHYGNATHINA.

Type species: Brachygnathus muticus Perty, 1830, by subsequent designation (Lorenz 1998: 55).

Family-group name: BRACHYGNATHINI Basilewsky, 1946. Stem: Brachygnath-.

Brachylobus Chaudoir, 1876a: 287. Current status: valid subgenus of Chlaenius Bonelli, 1810 in CARABIDAE: HARPALINAE: CHLAENIINI: CHLAENIINA.

Type species: Chlaenius lithophilus Say, 1823, by monotypy.

Family-group name: BRACHYLOBINI Basilewsky & Grundmann, 1955. Stem: Brachylob-.

Brachypsectra J.L. LeConte, 1874b: 55. Current status: valid genus in BRACHYPSECTRIDAE.

Type species: Brachypsectra fulva J.L. LeConte, 1874, by monotypy.

Family-group name: BRACHYPSECTRINI G.H. Horn, 1881. Stem: Brachypsectr-.

Brachypteroma Heyden, 1863: 128. Current status: valid genus in CERAMBYCIDAE: CERAMBYCINAE: BRACHYPTEROMINI.

Type species: Brachypteroma ottomanum Heyden, 1863, by monotypy.

Family-group name: BRACHYPTEROMINI Sama, 2008. Stem: Brachypterom- (ICZN 1999a, Article 29.4).

Brachypterus Kugelann, 1794: 560. Current status: valid genus in KATERETIDAE: KATERETINAE. Note: name placed on the Official List of Generic Names in Zoology (ICZN 1999b).

Type species: Dermestes urticae Fabricius, 1792, by subsequent designation (C.G. Thomson 1859: 67; see ICZN 1999b). Note: name placed on the Official List of Specific Names in Zoology (ICZN 1999b).

Family-group name: BRACHYPTERINAE Erichson, 1845. Stem: Brachypter-.

Brachypus Schönherr, 1826: 217. Current status: invalid senior synonym of Brachygyius G.A.K. Marshall, 1939 in CURCULIONIDAE: BRACHYCERINAE: TANYSPHYRINI. Note: junior homonym of Brachypus B. Meyer, 1814 [Aves]; Schönherr’s name was placed on the Official Index of Rejected and Invalid Generic Names in Zoology (see ICZN 1958a).

Type species: Brachypus lixoides Schönherr, 1826, by original designation.

Family-group name: BRACHYPI J.L. LeConte, 1876. Stem: Brachypod-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

Brachyrhinus Latreille, 1802b: 200. Current status: invalid senior synonym of Cryphiphorus Stierlin, 1883 in CURCULIONIDAE: ENTIMINAE: OTIORHYNCHITAE: OTIORHYNCHINI. Note: name suppressed for the Principle of Priority and placed on the Official Index of Rejected and Invalid Generic Names in Zoology (ICZN 1972).

Type species: Curculio ligustici Linnaeus, 1758, by subsequent designation (Pierce 1913: 422).

Family-group name: BRACHYRRHINIDAE Bedel, 1883. Stem: Brachyrhin-. Note: family-group name permanently invalid (ICZN 1999a, Article 39); name placed on the Official Index of Rejected and Invalid Family-group Names in Zoology (see ICZN 1972).

Brachys Dejean, 1833a: 83. Current status: valid genus in BUPRESTIDAE: AGRILINAE: TRACHEINI: BRACHINA.

Type species: Trachys tesselatus Fabricius, 1801, by monotypy.

Family-group name: BRACHES J.L. LeConte, 1861. Stem: Brach-.

Brachysternus Guérin-Méneville, 1831a: pl. 3. Current status: valid genus in SCARABAEIDAE: RUTELINAE: ANOPLOGNATHINI: BRACHYSTERNINA.

Type species: Brachysternus prasinus Guérin-Méneville, 1831, by monotypy.

Family-group name: BRACHYSTERNIDAE Burmeister, 1844. Stem: Brachystern-.

Brachytarsus Schönherr, 1823: 1135. Current status: junior synonym of Anthribus Geoffroy, 1762 in ANTHRIBIDAE: ANTHRIBINAE: ANTHRIBINI.

Type species: Bruchus varius Fabricius, 1787 (= Anthribus nebulosus Forster, 1770), by original designation.

Family-group name: BRACHYTARSINA C.G. Thomson, 1859. Stem: Brachytars-.

Bradybaenus Dejean, 1829: 160. Current status: valid genus in CARABIDAE: HARPALINAE: HARPALINI: HARPALINA.

Type species: Carabus scalaris G.-A. Olivier, 1790, by subsequent designation (Hope 1838a: 84).

Family-group name: BRADYBAENI Csiki, 1932. Stem: Bradybaen-. Note: senior homonym of BRADYBAENIDAE Pilsbry, 1934 (type genus Bradybaena Beck, 1837) in Gastropoda.

Bradybatus Germar, 1823: 305. Current status: valid genus in CURCULIONIDAE: CURCULIONINAE: ANTHONOMINI.

Type species: Bradybatus creutzeri Germar, 1823, by monotypy.

Family-group name: BRADYBATINI Pierce, 1919. Stem: Bradybat-. Note: junior homonym of BRADYBATINA Bonaparte, 1850 (type genus Bradybates Tschudi, 1838) in Amphibia.

Bradycellus Erichson, 1837: 64. Current status: valid genus in CARABIDAE: HARPALINAE: HARPALINI: STENOLOPHINA.

Type species [suggested]: Carabus collaris Paykull, 1798 (= Acupalpus caucasicus Chaudoir, 1846), by subsequent designation (Andrewes 1935: 20). Note: the valid type species fixation is that of Westwood (1838c: 6), who selected Harpalus placidus Gyllenhal, 1827, a species currently included in the genus Trichocellus Ganglbauer, 1892; an application to the Commission is necessary to fix Carabus collaris Paykull, 1798 as the type species of Bradycellus Erichson, 1837.

Family-group name: BRADYCELLINI Jeannel, 1942. Stem: Bradycell-.

Branchus J.L. LeConte, 1862: 222. Current status: valid genus in TENEBRIONIDAE: PIMELIINAE: BRANCHINI.

Type species: Branchus floridanus J.L. LeConte, 1862, by monotypy.

Family-group name: BRANCHINI J.L. LeConte, 1862. Stem: Branch-.

Brarus Kuschel in Kuschel and May, 1997: 16. Current status: valid genus in NEMONYCHIDAE: RHINORHYNCHINAE: MECOMACERINI: BRARINA.

Type species: Brarus mystes Kuschel, 1997, by original designation.

Family-group name: BRARINA Legalov, 2009. Stem: Brar-.

Brasilucanus Vulcano & Pereira, 1961: 473. Current status: valid genus in LUCANIDAE: LUCANINAE: LUCANINI.

Type species: Brasilucanus alvarengai Vulcano & Pereira, 1961, by original designation.

Family-group name: BRASILUCANINI Nikolajev, 1999. Stem: Brasilucan-.

Brathinus J.L. LeConte, 1852f: 156. Current status: valid genus in STAPHYLINIDAE: OMALIINAE: ANTHOPHAGINI.

Type species: Brathinus nitidus J.L. LeConte, 1852, by subsequent designation (Blackwelder 1952: 411). Note: R. Lucas (1920: 149) listed two species as type species and this is considered as a typification made in an ambiguous manner and is not available under the Code (ICZN 1999a, Article 67.5.3).

Family-group name: BRATHINIDAE J.L. LeConte, 1861. Stem: Brathin-.

†Brenthorrhinoides Gratshev & Zherikhin, 1996: 119. Current status: valid genus in NEMONYCHIDAE: †BRENTHORRHININAE: †BRENTHORRHININI: †BRENTHORRHINOIDINA. Note: Upper Jurassic (Kazakhstan).

Type species: †Brenthorrhinoides mandibulatus Gratshev & Zherikhin, 1996, by original designation.

Family-group name: †BRENTHORRHINOIDINI Legalov, 2003. Stem: Brenthorrhinoid-.

†Brenthorrhinus Arnoldi, 1977: 172. Current status: valid genus in NEMONYCHIDAE: †BRENTHORRHININAE: †BRENTHORRHININI: †BRENTHORRHININA. Note: Upper Jurassic (Kazakhstan).

Type species: †Brenthorrhinus mirabilis Arnoldi, 1977, by original designation.

Family-group name: †BRENTHORRHININAE Arnoldi, 1977. Stem: Brenthorrhin-.

Brentus Fabricius, 1787: 95. Current status: valid genus in BRENTIDAE: BRENTINAE: BRENTINI: BRENTINA.

Type species: Curculio anchorago Linnaeus, 1758, by subsequent designation (Latreille 1810: 430).

Family-group name: BRENTHIDES Billberg, 1820. Stem: Brent-.

Brises Pascoe, 1869a: 145. Current status: valid genus in TENEBRIONIDAE: TENEBRIONINAE: HELEINI: HELEINA.

Type species: Brises trachynotoides Pascoe, 1869, by monotypy.

Family-group name: BRISEINAE Carter, 1924. Stem: Bris-.

Brittona G.S. Medvedev & Lawrence, 1986: 574. Current status: valid genus in TENEBRIONIDAE: DIAPERINAE: HYOCIINI: BRITTONINA.

Type species: Brittona minuta G.S. Medvedev & Lawrence, 1986, by original designation.

Family-group name: BRITTONINA G.S. Medvedev & Lawrence, 1986. Stem: Britton-.

†Brochocoleus Hong, 1982: 103. Current status: valid genus in OMMATIDAE: †BROCHOCOLEINAE. Note: Lower Jurassic/Late Cretaceous (China; Kazakhstan; Mongolia; Myanmar; Russia; Spain; United Kingdom).

Type species: †Brochocoleus punctatus Hong, 1982, by original designation.

Family-group name: †BROCHOCOLEIDAE Hong, 1982. Stem: Brochocole-.

Bromius Chevrolat in Dejean, 1836a: 412. Current status: valid genus in CHRYSOMELIDAE: EUMOLPINAE: BROMIINI. Note: name placed on the Official List of Generic Names in Zoology (ICZN 2012b).

Type species: Chrysomela obscura Linnaeus, 1758, by subsequent designation (Monrós and Bechyné 1956: 1127; see ICZN 2012b). Note: name hereby placed on the Official List of Specific Names in Zoology (ICZN 2012b).

Family-group name: BROMIINAE Baly, 1865. Stem: Bromi-.

Brontes Fabricius, 1801a: xix. Current status: junior synonym of Uleiota Latreille, 1797 in SILVANIDAE: BRONTINAE: BRONTINI.

Type species: Cucujus flavipes Fabricius, 1779 (= Cerambyx planatus Linnaeus, 1761), by subsequent designation (Partington 1835: 632). Note: no species were originally included in this genus by Fabricius (1801a); the first ones were the five species listed in Fabricius (1801b: 97, 98).

Family-group name: BRONTITES Blanchard, 1845. Stem: Bront-.

Broscus Panzer, 1813: 62. Current status: valid genus in CARABIDAE: BROSCINAE: BROSCINI: BROSCINA. Note: new replacement name for Cephalotes Bonelli, 1810.

Type species: Carabus cephalotes Linnaeus, 1758, by monotypy.

Family-group name: BROSCHIDAE Hope, 1838. Stem: Brosc-.

Brotheus Stephens, 1829b: 13. Current status: valid genus in CURCULIONIDAE: BRACHYCERINAE: BRACHYCERINI.

Type species: Curculio porcatus Marsham, 1802 (= Brachycerus praemorsus Thunberg, 1799), by monotypy.

Family-group name: BROTHEINAE G.A.K. Marshall, 1907. Stem: Brothe-. Note: junior homonym of BROTHEINI Simon, 1879 (type genus Brotheas C.L. Koch, 1837) in Scorpiones; because of the homonymy, the stem of the genus was changed to Brotheus- and the family group name to BROTHEUSINI by Lyal and Alonso-Zarazaga (2006: 24); however, cases involving homonymy of family-group names based on similar genus- group names must be referred to the Commission unless the senior homonym is a nomen oblitum (ICZN 1999a, Article 55.3.1).

Bruchela Dejean, 1821: 78. Current status: valid genus in ANTHRIBIDAE: URODONTINAE.

Type species: Anthribus sericeus Fabricius, 1801 (= Bruchus rufipes G.-A. Olivier, 1790), by subsequent designation (Schönherr 1823: 1134) (type fixation for Urodon Schönherr, 1823, an unnecessary new replacement name for Bruchela Dejean).

Family-group name: BRUCHELIDAE Pierce, 1916. Stem: Bruchel-.

Bruchidius Schilsky, 1905: 41B. Current status: valid genus in CHRYSOMELIDAE: BRUCHINAE: BRUCHINI: ACANTHOSCELIDINA.

Type species: Bruchus quinqueguttatus G.-A. Olivier, 1795, by subsequent designation (Bridwell 1932: 104).

Family-group name: BRUCHIDIINI Bridwell, 1946. Stem: Bruchidi-.

Bruchus Linnaeus, 1767: 604. Current status: valid genus in CHRYSOMELIDAE: BRUCHINAE: BRUCHINI: BRUCHINA. Note: the senior homonym Bruchus Geoffroy (1762: 163) was suppressed for both Principles of Priority and Homonymy and placed on the Official Index of Generic Names in Zoology while Bruchus Linnaeus, 1767 was placed on the Official List of Generic Names in Zoology (ICZN 1995b).

Type species: Dermestes pisorum Linnaeus, 1758, by subsequent designation (Latreille 1810: 430; see ICZN 1995b). Note: name placed on the Official List of Specific Names in Zoology (ICZN 1995b).

Family-group name: BRUCHELAE Latreille, 1802. Stem: Bruch-.

Bryaxis Kugelann, 1794: 580. Current status: valid genus in STAPHYLINIDAE: PSELAPHINAE: GONIACERITAE: BYTHININI: BYTHININA. Note: name placed on the Official List of Generic Names in Zoology (ICZN 1969b).

Type species: Pselaphus bulbifer H.G.L. Reichenbach, 1816, by plenary power (see ICZN 1969b). Note: name placed on the Official List of Specific Names in Zoology (ICZN 1969b).

Family-group name: BRYAXINA Jakobson, 1910. Stem: Bryaxe-.

Bryaxis Leach, 1817: 81. Current status: invalid senior synonym of Rybaxis Saulcy, 1876 in STAPHYLINIDAE: PSELAPHINAE: GONIACERITAE: BRACHYGLUTINI: BRACHYGLUTINA. Note: junior homonym of Bryaxis Kugelann, 1794; name placed on the Official Index of Rejected and Invalid Generic Names in Zoology (see ICZN 1969b).

Type species: Staphylinus sanguineus Linnaeus, 1758, by subsequent designation (Curtis 1830: pl. 315).

Family-group name: BRYAXES J.L. LeConte, 1861. Stem: Bryaxe-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

Brychius C.G. Thomson, 1859: 11. Current status: valid genus in HALIPLIDAE.

Type species: Dytiscus elevatus Panzer, 1793, by original designation.

Family-group name: BRYCHIINI Ádám, 1996. Stem: Brychi-.

Bubastes Laporte & Gory, 1838: 1. Current status: valid genus in BUPRESTIDAE: BUPRESTINAE: BUBASTINI.

Type species: Bubastes sphaenoida Laporte & Gory, 1838, by monotypy.

Family-group name: BUBASTINI Obenberger, 1920. Stem: Bubast-.

Bulaea Mulsant, 1850: 36, 69. Current status: valid genus in COCCINELLIDAE: COCCINELLINAE: COCCINELLINI.

Type species: Coccinella lichatschovii Hummel, 1827, by subsequent designation (Crotch 1874a: 105).

Family-group name: BULAEINI Savoyskaya, 1969. Stem: Bulae-.

Bulis Laporte & Gory, 1838: 1. Current status: valid genus in BUPRESTIDAE: POLYCESTINAE: BULINI.

Type species: Buprestis bivittata Fabricius, 1801, by subsequent designation (Lacordaire 1857: 42).

Family-group name: BULISINA Bellamy, 1995. Stem: Bul-.

Bumetopia Pascoe, 1858: 252. Current status: valid genus in CERAMBYCIDAE: LAMIINAE: HOMONOEINI.

Type species: Bumetopia oscitans Pascoe, 1858, by monotypy.

Family-group name: BUMÉTOPIDES Lacordaire, 1872. Stem: Bumetopi-.

Bunyaeus Kuschel, 1994: 612. Current status: valid genus in NEMONYCHIDAE: RHINORHYNCHINAE: MECOMACERINI: BUNYAEINA.

Type species: Bunyaeus monteithi Kuschel, 1994, by original designation.

Family-group name: BUNYAEINA Legalov, 2023c: 21. Stem: Bunyae-. Note: BUNYAEINA Legalov (2017a: 69) was published in a work that is nomenclaturally unavailable (ICZN 2012f, Article 8.5.3).

Buprestis Linnaeus, 1758: 342, 408. Current status: valid genus in BUPRESTIDAE: BUPRESTINAE: BUPRESTINI: BUPRESTINA. Note: name placed on the Official List of Generic Names in Zoology (ICZN 1994f).

Type species: Buprestis octoguttata Linnaeus, 1758, by plenary power (see ICZN 1994f). Note: name placed on the Official List of Specific Names in Zoology (ICZN 1994f).

Family-group name: BUPRESTIDES Leach, 1815. Stem: Buprest-.

†Burmocorynus Legalov, 2020c: 132. Current status: valid genus in †MESOPHYLETIDAE: †MESOPHYLETINAE. Note: mid-Cretaceous (Myanmar).

Type species: †Burmocorynus jarzembowskii Legalov, 2020, by original designation.

Family-group name: †BURMOCORYNINI Legalov, 2020c: 132. Stem: Burmocoryn-.

†Burmolycus Bocák, Li & Ellenberger, 2019: 152. Current status: valid genus in LYCIDAE: DEXORINAE: DEXORINI: †BURMOLYCINA. Note: mid-Cretaceous (Myanmar).

Type species: †Burmolycus compactus Bocák, Li & Ellenberger, 2019, by original designation.

Family-group name: †BURMOLYCINI Bocák, Li & Ellenberger, 2019: 150. Stem: Burmolyc-.

†Burmomacer Legalov, 2020c: 129. Current status: valid genus in NEMONYCHIDAE: RHINORHYNCHINAE: †BURMOMACERATINI. Note: mid-Cretaceous (Myanmar).

Type species: †Burmomacer kirejtshuki Legalov, 2020, by original designation.

Family-group name: †BURMOMACERATINI Legalov, 2020c: 128. Stem: Burmomacerat-. Note: the names Burmomacer and BURMOMACERATINI were established in the printed version of this article (Legalov 2020c) and not in the electronic version that was made public in 2018 since it was not registered in Official Register of Zoological Nomenclature (ZooBank) before the electronic work was published (ICZN 2012f, Article 8.5.3); Clarke et al. (2018: 39) pre-emptively synonymized the unavailable BURMOMACERATINI Legalov, 2018 (as BURMOMACRINI) with MECOMACERINI; the correct spelling and status of this family-group name (which could be preserved as originally proposed if Article 29.4 is used (ICZN 1999a)) needs further attention.

Byctiscus C.G. Thomson, 1859: 130. Current status: valid genus in ATTELABIDAE: RHYNCHITINAE RHYNCHITITAE: BYCTISCINI: BYCTISCINA.

Type species: Curculio populi Linnaeus, 1758, by original designation.

Family-group name: BYCTISCINI Voss, 1923. Stem: Byctisc-.

Byrrhidium Harold, 1869a: 96. Current status: treated as a valid genus in SCARABAEIDAE: SCARABAEINAE: BYRRHIDIINI [but see notes below]. Note: junior homonym of Byrrhidium Heer, 1864¹ in Coleoptera: BYRRHIDAE; this name was first used by Harold in Gemminger and von Harold (1869a: 997) but it is not available from that publication since there is no description or indication, the sole species associated with the name being a nomen nudum; a substitute name will be proposed by the authors of BYRRHIDIINI in the near future (C. Deschodt, pers. comm. to P.B., 2022).

Type species: Byrrhidium ovale Harold, 1869, by monotypy.

Family-group name: BYRRHIDIINI Davis, Deschodt & Scholtz, 2019: 134. Stem: Byrrhidi-. Note: family-group name permanently invalid (ICZN 1999a, Article 39); a substitute name will be proposed by the authors of BYRRHIDIINI in the near future (C. Deschodt, pers. comm. to P.B., 2022).

Byrrhus Linnaeus, 1767: 568. Current status: valid genus in BYRRHIDAE: BYRRHINAE: BYRRHINI. Note: the senior homonym Byrrhus Geoffroy, 1762 (and all other uses of the name prior to the publication of Byrrhus Linnaeus, 1767) was suppressed for both Principles of Priority and Homonymy and placed on the Official Index of Rejected and Invalid Generic Names in Zoology while Byrrhus Linnaeus, 1767 was placed on the Official List of Generic Names in Zoology (ICZN 1994a).

Type species: Dermestes pilula Linnaeus, 1758, by subsequent designation (Latreille 1810: 428; see ICZN 1994a). Note: name placed on the Official List of Specific Names in Zoology (ICZN 1994a).

Family-group name: BYRRHII Latreille, 1804. Stem: Byrrh-.

Byrsopages Schönherr, 1842a: 208. Current status: valid genus in CURCULIONIDAE: ENTIMINAE: ENTIMITAE: BYRSOPAGINI: BYRSOPAGINA.

Type species: Byrsopages villosus Bohemam, 1842, by original designation.

Family-group name: BYRSOPAGIDES Lacordaire, 1863. Stem: Byrsopag-.

Byrsops Germar, 1829: 358. Current status: valid genus in CURCULIONIDAE: BRACHYCERINAE: BRACHYCERINI.

Type species: Brachycerus quadratus Wiedemann, 1821, by subsequent designation (Alonso- Zarazaga and Lyal 1999: 21, 62).

Family-group name: BYRSOPTIDES Germar, 1829. Stem: Byrsop-.

Bythinoplectus Reitter, 1882a: 195. Current status: valid genus in STAPHYLINIDAE: PSELAPHINAE: EUPLECTITAE: BYTHINOPLECTINI: BYTHINOPLECTINA.

Type species: Bythinoplectus foveatus Reitter, 1883, by subsequent monotypy (Reitter 1883a: 37).

Family-group name: BYTHINOPLECTINI L.W. Schaufuss, 1890. Stem: Bythinoplect-.

Bythinus Leach, 1817: 82. Current status: valid genus in STAPHYLINIDAE: PSELAPHINAE: GONIACERITAE: BYTHININI: BYTHININA. Note: name placed on the Official List of Generic Names in Zoology (ICZN 1969b).

Type species: Pselaphus securiger H.G.L. Reichenbach, 1816, by subsequent designation (Westwood 1838c: 21; see ICZN 1969b). Note: name placed on the Official List of Specific Names in Zoology (ICZN 1969b).

Family-group name: BYTHININI Raffray, 1890. Stem: Bythin-.

Byturus Latreille, 1797: 69. Current status: valid genus in BYTURIDAE: BYTURINAE.

Type species: Dermestes tomentosus DeGeer, 1774, by subsequent designation (Latreille 1810: 427). Note: no species were originally included in this genus by Latreille (1797); the first ones were the three species listed by Latreille (1802b: 134).

Family-group name: BYTURIDAE Gistel, 1848. Stem: Bytur-.

Cachiporra Chamorro & Konstantinov, 2011: 50. Current status: valid genus in CHRYSOMELIDAE: LAMPROSOMATINAE: CACHIPORRINI.

Type species: Cachiporra extremaglobosa Chamorro & Konstantinov, 2011, by original designation.

Family-group name: CACHIPORRINI Chamorro & Konstantinov, 2011: 49. Stem: Cachiporr-.

†Cacomorphocerus C. Schaufuss, 1891a: 57. Current status: valid genus in CANTHARIDAE: CANTHARINAE: †CACOMORPHOCERINI. Note: Eocene (Baltic and Rovno ambers).

Type species: †Cacomorphocerus cerambyx C. Schaufuss, 1891, by monotypy.

Family-group name: †CACOMORPHOCERINI Fanti & Kupryjanowicz, 2018: 150. Stem: Cacomorphocer-.

Cacosceles Newman, 1838: 491. Current status: valid genus in CERAMBYCIDAE: PRIONINAE: CACOSCELINI.

Type species: Cacosceles oedipus Newman, 1838, by monotypy.

Family-group name: CACOSCELITAE J. Thomson, 1861. Stem: Cacoscel-.

Cacoscelis Chevrolat in Dejean, 1836a: 389. Current status: valid genus in CHRYSOMELIDAE: GALERUCINAE: ALTICINI.

Type species: Chrysomela famelica Fabricius, 1787 (= Altica marginata Fabricius, 1775), by subsequent designation (Monrós and Bechyné 1956: 1133).

Family-group name: CACOSCELINI Bechyné, 1968. Stem: Cacoscelid-.

Cactopinus O. Schwarz, 1899: 11. Current status: valid genus in CURCULIONIDAE: SCOLYTINAE: CACTOPININI.

Type species: Cactopinus hubbardi O. Schwarz, 1899, by monotypy.

Family-group name: CACTOPINAE Chamberlin, 1939. Stem: Cactopin-.

Caecossonus Gilbert, 1955: 193. Current status: valid genus in CURCULIONIDAE: MOLYTINAE: LYMANTINI: CAECOSSONINA.

Type species: Caecossonus dentipes Gilbert, 1955, by monotypy.

Family-group name: CAECOSSONINA Osella, 1980. Stem: Caecosson-.

Caelostomus W.S. MacLeay, 1825: 23. Current status: valid genus in CARABIDAE: HARPALINAE: DRIMOSTOMATINI.

Type species: Anaulacus picipes W.S. MacLeay, 1825, by monotypy.

Family-group name: CAELOSTOMINA Burgeon, 1935. Stem: Caelostom-.

Caenocara C.G. Thomson, 1859: 90. Current status: valid genus in PTINIDAE: DORCATOMINAE.

Type species: Dorcatoma bovistae P.W.J. Müller, 1803, by original designation. Note: the authorship of the type species is credited to Hoffmann in the literature but Niehuis (2012: 2, 3, 42) showed that P.W.J. Müller is the author.

Family-group name: CAENOCARINI Böving, 1927. Stem: Caenocar-.

Caenocrypticus Gebien, 1920: 139. Current status: valid genus in TENEBRIONIDAE: PIMELIINAE: CAENOCRYPTICINI.

Type species: Caenocrypticus uncinatus Gebien, 1920, by monotypy.

Family-group name: CAENOCRYPTICINI C. Koch, 1958. Stem: Caenocryptic-.

Caenoscelis C.G. Thomson, 1863: 266. Current status: valid genus in CRYPTOPHAGIDAE: CRYPTOPHAGINAE: CAENOSCELINI.

Type species: Cryptophagus ferrugineus C.R. Sahlberg, 1820, by monotypy.

Family-group name: CAENOSCELINI Casey, 1900. Stem: Caenoscel-.

Calandra Clairville, 1798: 62, pl. 2. Current status: invalid senior synonym of Sphenophorus Schönherr, 1838 in CURCULIONIDAE: DRYOPHTHORINAE: SPHENOPHORINI: SPHENOPHORINA. Note: two original spellings were used, including Calendra (p. 62); Latreille (1806: 270) acted as First Reviser (ICZN 1999a, Article 24.2.3) and selected the spelling Calandra; the name Calendra is suppressed for the Principle of Priority and placed on the Official Index of Rejected and Invalid Generic Names in Zoology (ICZN 1959b, as Calandra Clairville & Schellenberg, 1798).

Type species: Curculio abbreviatus Fabricius, 1787, by subsequent designation (Latreille 1810: 431). Note: name validated by the Commission (i.e., not a primary homonym of Curculio abbreviatus Linnaeus, 1758) and placed on the Official List of Specific Names in Zoology (ICZN 1959b).

Family-group name: CALANDRAEDES Billberg, 1820. Stem: Calandr-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

Calandra Gistel, 1848b: 136. Current status: junior synonym of Sitophilus Schönherr, 1838 in CURCULIONIDAE: DRYOPHTHORINAE: LITOSOMINI: SITOPHILINA. Note: unnecessary new replacement name for Sitophilus Schönherr, 1838; name placed on the Official Index of Rejected and Invalid Generic Names in Zoology (see ICZN 1959b).

Type species: Curculio oryzae Linnaeus, 1763, by original designation (type fixation for Sitophilus Schönherr).

Family-group name: CALANDRINA C.G. Thomson, 1859. Stem: Calandr-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

Calathus Bonelli, 1810: Tab. Syn. Current status: valid genus in CARABIDAE: HARPALINAE: SPHODRINI: CALATHINA.

Type species: Carabus cisteloides Panzer, 1793 (= Carabus fuscipes Goeze, 1777), by subsequent designation (Curtis 1827: pl. 184). Note: no species were originally included in this genus by Bonelli (1810); the first ones were the two species listed by Panzer (1813: 60); Goeze’s (1777) work is not consistently binominal (see ICZN 2021a) and therefore an application to the Commission is necessary to retain Carabus fuscipes Goeze, 1777 as the valid name for the type species.

Family-group name: CALATHIDAE Laporte, 1834. Stem: Calath-.

Calcar Dejean, 1821: 67. Current status: invalid senior synonym of Belopus Gebien, 1911 in TENEBRIONIDAE: LAGRIINAE: BELOPINI. Note: junior homonym of Calcar de Montfort, 1810 [Mollusca].

Type species: Tenebrio elongatus Herbst, 1797, by monotypy.

Family-group name: CALCARIENS Mulsant, 1854. Stem: Calcar-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

Calcodes Westwood, 1834: 118. Current status: valid genus in LUCANIDAE: LUCANINAE: LUCANINI. Note: name also referred to as “Chalcodes”, an incorrect subsequent spelling of Calcodes, in the literature.

Type species: Lucanus aeratus Hope, 1833, by monotypy.

Family-group name: CHALCODINAE Didier & Séguy, 1953. Stem: Calcod-.

Caledonica Chaudoir, 1861: 312. Current status: valid genus in CICINDELIDAE: CICINDELINI: DROMICINA.

Type species: Distipsidera mniszechii J. Thomson, 1856, by subsequent designation (W. Horn 1915: 438).

Family-group name: CALEDONICINI W. Horn, 1893. Stem: Caledonic-.

Calicnemis Laporte, 1832a: pl. 7. Current status: valid genus in SCARABAEIDAE: DYNASTINAE: PENTODONTINI: PENTODONTINA.

Type species: Calicnemis latreillei Laporte, 1832 [as latreillii], by monotypy.

Family-group name: CALICNÉMIENS Mulsant, 1842. Stem: Calicnemid-.

Calitys C.G. Thomson, 1859: 71. Current status: valid genus in TROGOSSITIDAE: CALITYINAE.

Type species: Silpha dentata Fabricius, 1787 (= Hispa scabra Thunberg, 1784), by original designation.

Family-group name: CALITYNI Houlbert, 1922. Stem: Cality-.

Calleida Latreille in Latreille and Dejean, 1824: 132. Current status: valid genus in CARABIDAE: HARPALINAE: LEBIINI: CALLEIDINA.

Type species: Carabus decorus Fabricius, 1801, by subsequent designation (E. Desmarest 1851: 72).

Family-group name: CALLIDIDES Chaudoir, 1873. Stem: Calleid-.

Callia Audinet-Serville, 1835a: 60. Current status: valid genus in CERAMBYCIDAE: LAMIINAE: CALLIINI.

Type species: Callia azurea Audinet-Serville, 1835, by monotypy.

Family-group name: CALLITAE J. Thomson, 1864. Stem: Calli-.

Callicerus Gravenhorst, 1802: 65. Current status: valid genus in STAPHYLINIDAE: ALEOCHARINAE: GEOSTIBINI. Note: name placed on the Official List of Generic Names in Zoology (ICZN 2012e).

Type species: Callicerus obscurus Gravenhorst, 1802, by monotypy (see ICZN 2012e). Note: name placed on the Official List of Specific Names in Zoology (see ICZN 2012e).

Family-group name: CALLICERINA Jakobson, 1908. Stem: Callicer-. Note: suppressed for both the Principle of Priority and Principle of Homonymy and placed on the Official Index of Rejected and Invalid Family-group Names in Zoology (ICZN 2012e).

Callichroma Latreille, 1816: 341. Current status: valid genus in CERAMBYCIDAE: CERAMBYCINAE: CALLICHROMATINI.

Type species [suggested]: Cerambyx suturalis Fabricius, 1781 (= Cerambyx auricomus Linnaeus, 1767), by subsequent designation (E. Desmarest 1860: 315). Note: the valid type species fixation is that of Audouin (1823: 52), who selected Cerambyx alpinus Linnaeus, 1758, but that species is the type species of the genus Rosalia Audinet- Serville, 1834; a subsequent designation based on an originally included nominal species is that of G.R. Waterhouse (1840: 228), who selected Cerambyx moschatus Linnaeus, 1758, but that species is the type species of the genus Aromia Audinet- Serville, 1834; an application to the Commission is necessary to fix Cerambyx suturalis Fabricius, 1781 as the type species of Callichroma Latreille, 1816 to conserve current usage (e.g., Monné and Nearns 2022: 16).

Family-group name: CALLICHROMINAE Swainson, 1840. Stem: Callichromat-.

Callidiopis A. White, 1855: 328. Current status: valid genus in CERAMBYCIDAE: CERAMBYCINAE: CALLIDIOPINI.

Type species: Callidium scutellare Fabricius, 1801, by subsequent designation (J. Thomson 1864: 237).

Family-group name: CALLIDIOPSIDES Lacordaire, 1868. Stem: Callidiop-.

Callidium Fabricius, 1775: 187. Current status: valid genus in CERAMBYCIDAE: CERAMBYCINAE: CALLIDIINI.

Type species [suggested]: Cerambyx violaceus Linnaeus, 1758, by subsequent designation (C.G. Thomson 1859: 150). Note: the valid type species fixation is that of Latreille (1810: 431), who selected Cerambyx bajulus Linnaeus, 1758, but that species is the type species of the genus Hylotrupes Audinet-Serville, 1834; a subsequent designation based on an originally included nominal species is that of Lefebvre (1838a: 46), who designated Cerambyx variabilis Linnaeus, 1761, but that species is the type species of the genus Phymatodes Mulsant, 1839; yet another designation based on an originally included nominal species is that of Blanchard in Audouin et al. (1843: pl. 66), who selected Leptura arietis Linnaeus, 1758, but that species is the type species of the genus Clytus Laicharting, 1784; an application to the Commission is necessary to fix Cerambyx violaceus Linnaeus, 1758 as the type species of Callidium Fabricius, 1775.

Family-group name: CALLIDIADAE W. Kirby, 1837. Stem: Callidi-.

Callimerus Gorham, 1876: 60, 64. Current status: valid genus in CLERIDAE: CLERINAE: CLERITAE: HYDNOCERINI: CALLIMERINA. Note: this name, published on 7 August 1876 (p. 57 footer of the journal), is a senior homonym of Callimerus Stebbing, 1876 [Crustacea], issued in volume 18 (fourth series) of the ‘Annals and Magazine of Natural History’ on 1 December 1876 (Evenhuis 2003a: 27).

Type species: Clerus dulcis Westwood, 1852, by original designation.

Family-group name: CALLIMERINI Kolibáč, 1998. Stem: Callimer-.

Callipogon Audinet-Serville, 1832: 125, 140. Current status: valid genus in CERAMBYCIDAE: PRIONINAE: CALLIPOGONINI.

Type species: Prionus barbatus Fabricius, 1781, by monotypy.

Family-group name: CALLIPOGONITAE J. Thomson, 1861. Stem: Callipogon-.

Calliprason A. White in A. White and Doubleday, 1843: 277. Current status: valid genus in CERAMBYCIDAE: CERAMBYCINAE: RHAGIOMORPHINI.

Type species: Callichroma sinclairi A. White, 1843, by monotypy.

Family-group name: CALLIPRASONINI McKeown, 1947. Stem: Callipras-.

Callirhipis Latreille, 1829a: 459. Current status: valid genus in CALLIRHIPIDAE.

Type species: Callirhipis dejeanii Latreille, 1829, by original designation.

Family-group name: CALLIRHIPINI Emden, 1924. Stem: Callirhip-.

Callirhopalus Hochhuth, 1851: 54. Current status: valid genus in CURCULIONIDAE: ENTIMINAE: POLYDRUSITAE: CNEORHININI.

Type species: Callirhopalus sedakowii Hochhuth, 1851, by monotypy.

Family-group name: CALLIRHOPALINI Voss, 1956. Stem: Callirhopal-.

Callisina Baly, 1860: 30. Current status: valid genus in CHRYSOMELIDAE: EUMOLPINAE: TYPOPHORINI.

Type species: Callisina fasciata Baly, 1860, by original designation.

Family-group name: CALLISINITES Chapuis, 1874. Stem: Callisin-.

Callispa Baly, 1858: 4. Current status: valid genus in CHRYSOMELIDAE: CASSIDINAE: CALLISPINI.

Type species: Callispa fortunii Baly, 1858, by original designation.

Family-group name: CALLISPITES Chapuis, 1875. Stem: Callisp-.

Callisthenes Fischer, 1820: pl. 7. Current status: valid subgenus of Calosoma Weber, 1801 in CARABIDAE: CARABINAE: CARABINI: CALOSOMATINA.

Type species: Callisthenes panderi Fischer, 1820, by monotypy.

Family-group name: CALLISTHENISIDAE Gistel, 1848. Stem: Callisthen-.

Callistoides Motschulsky, 1865: 334. Current status: valid genus in CARABIDAE: HARPALINAE: CHLAENIINI: CHLAENIINA.

Type species: Callistoides malachinus Motschulsky, 1865, by monotypy.

Family-group name: CALLISTOIDINI Basilewsky, 1950. Stem: Callistoid-.

Callistus Bonelli, 1810: Tab. Syn. Current status: valid genus in CARABIDAE: HARPALINAE: CHLAENIINI: CALLISTINA.

Type species: Carabus lunatus Fabricius, 1775, by subsequent monotypy (Panzer 1813: 52).

Family-group name: CALLISTIDAE Laporte, 1834. Stem: Callist-.

Callohispa Uhmann, 1960: 61. Current status: valid genus in CHRYSOMELIDAE: CASSIDINAE: CALLOHISPINI.

Type species: Callohispa mirifica Uhmann, 1960, by original designation.

Family-group name: CALLOHISPINI Uhmann, 1960. Stem: Callohisp-.

Callopistus Deyrolle, 1865: 9. Current status: invalid senior synonym of Eucallopistus Bellamy, 2003 in BUPRESTIDAE: CHRYSOCHROINAE: CHRYSOCHROINI: EUCALLOPISTINA. Note: junior homonym of Callopistus Schönherr, 1832 [Coleoptera: CURCULIONIDAE].

Type species: Callopistus castelnaudii Deyrolle, 1865, by original designation.

Family-group name: CALLOPISTINA Kurosawa, 1990. Stem: Callopist-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

Calocerus Fauvel, 1891: 88. Current status: junior synonym of Glyptoma Erichson, 1839 in STAPHYLINIDAE: OSORIINAE: THORACOPHORINI: GLYPTOMINA. Note: junior homonym of Calocerus J.L. LeConte, 1853 [Coleoptera: ELATERIDAE].

Type species: Glyptoma crassicorne Erichson, 1840, by subsequent designation (R. Lucas 1920: 159).

Family-group name: CALOCERI Blackwelder, 1942. Stem: Calocer-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

Calochromus Guérin-Méneville, 1833b: 158. Current status: valid genus in LYCIDAE: CALOCHROMINAE.

Type species: Calochromus glaucopterus Guérin-Méneville, 1833, by monotypy.

Family-group name: CALOCHROMIDES Lacordaire, 1857. Stem: Calochrom-.

Calocomus Audinet-Serville, 1832: 194. Current status: valid genus in CERAMBYCIDAE: PRIONINAE: CALOCOMINI.

Type species: Calocomus hamatiferus Audinet-Serville, 1832 (= Prionus desmarestii Guérin- Méneville, 1831), by monotypy.

Family-group name: CALOCOMINI Galileo & Martins, 1993. Stem: Calocom-.

Calodera Mannerheim, 1830: 85. Current status: valid genus in STAPHYLINIDAE: ALEOCHARINAE: OXYPODINI: OXYPODINA.

Type species: Calodera nigrita Mannerheim, 1830, by subsequent designation (Westwood 1838c: 20).

Family-group name: CALODÉRATES Mulsant & Rey, 1874. Stem: Caloder-.

Calodromus Guérin-Méneville in Guérin-Méneville and Gory, 1832: pl. 34. Current status: valid genus in BRENTIDAE: BRENTINAE: CYPHAGOGINI.

Type species: Calodromus mellyi Guérin-Méneville & Gory, 1832, by monotypy.

Family-group name: CALODROMINI Kleine, 1922. Stem: Calodrom-.

Calognathus Guérin-Méneville, 1837: pl. 172. Current status: valid genus in TENEBRIONIDAE: PIMELIINAE: CRYPTOCHILINI: CALOGNATHINA.

Type species: Calognathus chevrolatii Guérin-Méneville, 1837, by monotypy.

Family-group name: CALOGNATHIDES Lacordaire, 1859. Stem: Calognath-.

Calonecrus J. Thomson, 1857: 117. Current status: valid genus in NITIDULIDAE: CALONECRINAE.

Type species: Calonecrus wallacei J. Thomson, 1857, by monotypy.

Family-group name: CALONECRINAE Kirejtshuk, 1982. Stem: Calonecr-.

Calophaena Klug, 1821: 295. Current status: valid genus in CARABIDAE: HARPALINAE: CALOPHAENINI.

Type species: Carabus acuminatus G.-A. Olivier, 1790, by subsequent designation (Hope 1838a: 104).

Family-group name: CALOPHAENIDAE Jeannel, 1948. Stem: Calophaen-.

Calopteron Laporte, 1838: 25. Current status: valid genus in LYCIDAE: LYCINAE: CALOPTERINI: CALOPTERINA.

Type species [suggested]: fixed by Bocáková (2003: 215) according to Article 70.3.2 (ICZN 1999a) as Lycus apicalis Guérin-Méneville, 1838, the taxonomic species involved in the misidentification of Lycus limbatus Fabricius, 1801 (sensu Kleine, 1933), as designated subsequently by Bocák (1998: 249). Note: the valid type species fixation is that of Bescherelle (1845: 520), who selected Cantharis bicolor Linnaeus, 1763, but that species is currently placed in the genus Thonalmus Bourgeois, 1884; an application to the Commission is necessary to retain Lycus apicalis Guérin-Méneville, 1838 as the type species of Calopteron Laporte, 1838.

Family-group name: CALOPTERINI Green, 1949. Stem: Calopter-.

Calopus Fabricius, 1775: 182. Current status: valid genus in OEDEMERIDAE: CALOPODINAE.

Type species: Cerambyx serraticornis Linnaeus, 1758, by subsequent designation (Latreille 1810: 430).

Family-group name: CALOPINI A. Costa, 1852. Stem: Calopod-. Note: nomen protectum (see Bouchard et al. 2011: 435).

Calosoma Weber, 1801: 20. Current status: valid genus in CARABIDAE: CARABINAE: CARABINI: CALOSOMATINA. Note: this genus was described in 1801 by both Weber (p. 20) and Fabricius (p. 211); bibliographic research suggests that Weber’s book (issued by March 1801, see Evenhuis 1997b: 809) was published before Fabricius first volume of his Systema Eleutheratorum (issued by 26 April 1801, see Bousquet 2017a: 108; see also Chapin 1946: 79).

Type species: Carabus sycophanta Linnaeus, 1758, by subsequent designation (Latreille 1810: 426).

Family-group name: CALOSOMII Bonelli, 1810. Stem: Calosomat-.

Calospasta J.L. LeConte, 1862: 273. Current status: junior synonym of Eupompha J.L. LeConte, 1858 in MELOIDAE: MELOINAE: EUPOMPHINI.

Type species: Epicauta elegans J.L. LeConte, 1851, by monotypy.

Family-group name: CALOSPASTIDES Wellman, 1910. Stem: Calospast-.

Calydus Reitter, 1896: 192. Current status: valid genus in MELOIDAE: MELOINAE: MYLABRINI. Note: new replacement name for Caloenas Reitter, 1889.

Type species: Caloenas pulcher Reitter, 1889, by monotypy (type fixation for Caloenas Reitter).

Family-group name: CALYDINA Kaszab, 1960. Stem: Calyd-.

Calyptillus G.H. Horn in J.L. LeConte and G.H. Horn, 1876: 27. Current status: valid genus in CURCULIONIDAE: ENTIMINAE: POLYDRUSITAE: PSALLIDIINI.

Type species: Calyptillus cryptops G.H. Horn, 1876, by monotypy.

Family-group name: CALYPTILLI G.H. Horn, 1876. Stem: Calyptill-.

Calyptocephalus G.R. Gray in Griffith and Pidgeon, 1831: 370, pls 39, 61. Current status: valid genus in LAMPYRIDAE: LAMPYRINAE: PLEOTOMINI.

Type species: Calyptocephalus fasciatus G.R. Gray, 1831, by monotypy.

Family-group name: CALYPTOCEPHALINA Jakobson, 1911. Stem: Calyptocephal-.

Calyptocerus Guérin-Méneville, 1843b: 177. Current status: valid genus in EUCNEMIDAE: MELASINAE: CALYPTOCERINI.

Type species: Calyptocerus leboucherii Guérin-Méneville, 1843, by monotypy.

Family-group name: CALYPTOCERINI Muona, 1993. Stem: Calyptocer-.

Calyptomerus L. Redtenbacher, 1847a: 159. Current status: valid genus in CLAMBIDAE: CALYPTOMERINAE.

Type species: Calyptomerus alpestris L. Redtenbacher, 1847, by monotypy.

Family-group name: CALYPTOMERIDAE Crowson, 1955. Stem: Calyptomer-.

Camaragnathus Bertrand-Bocandé, 1849: 460. Current status: junior synonym of Hiletus Schiødte, 1847 in CARABIDAE: HILETINAE.

Type species: Camaragnathus guerinii Bertrand-Bocandé, 1849 (= Hiletus versutus Schiødte, 1847), by subsequent designation (E. Desmarest 1851: 158).

Family-group name: CAMARAGNATHINI Csiki, 1927. Stem: Camaragnath-.

Camaria Lepeletier & Audinet-Serville, 1828: 454. Current status: valid genus in TENEBRIONIDAE: STENOCHIINAE: CNODALONINI.

Type species: Camaria nitida Lepeletier & Audinet-Serville, 1828 (= Tenebrio nitens G.-A. Olivier, 1795), by monotypy.

Family-group name: CAMARIINAE Anonymous, 1924. Stem: Camari-.

Camarotus Germar in Schönherr, 1833: 185. Current status: valid genus in CURCULIONIDAE: CURCULIONINAE: CAMAROTINI: CAMAROTINA.

Type species: Camarotus coccinelloides Germar, 1833, by original designation (p. 4).

Family-group name: CAMAROTIDES Schönherr, 1833. Stem: Camarot-.

Cambaia Lane, 1951: 12. Current status: junior synonym of Paraleptidea Gounelle, 1913 in CERAMBYCIDAE: CERAMBYCINAE: PSEBIINI.

Type species: Cambaia longitarsis Lane, 1951, by original designation.

Family-group name: CAMBAIINAE Lane, 1951. Stem: Cambai-.

†Camelomorpha Kirejtshuk, Azar & Telnov in Kirejtshuk and Azar, 2008: 41. Current status: valid genus in ANTHICIDAE: MACRATRIINAE: †CAMELOMORPHINI. Note: Lower Cretaceous (Lebanon).

Type species: †Camelomorpha longicervix Kirejtshuk, Azar & Telnov, 2008, by original designation.

Family-group name: †CAMELOMORPHINI Kirejtshuk, Azar & Telnov, 2008. Stem: Camelomorph-.

Camenta Erichson, 1847b: 695. Current status: valid genus in SCARABAEIDAE: SERICINAE: ABLABERINI.

Type species: Camenta obesa Burmeister, 1855, by present designation. Note: no species were originally included in this genus by Erichson (1847b); the first ones were the five species described by Burmeister (1855: 144–146).

Family-group name: CAMENTINI Machatschke, 1959. Stem: Cament-.

Camiarus Sharp, 1878b: 36. Current status: valid genus in LEIODIDAE: CAMIARINAE: CAMIARINI. Note: new replacement name for Camirus Sharp, 1876.

Type species [suggested]: Camirus thoracicus Sharp, 1876, by subsequent designation (Hatch 1928: 210). Note: the valid type species fixation is that of Bertkau (1879: 448), who selected Camirus convexus Sharp, 1876, but that species is the type species of the genus Camiarites Jeannel, 1957; an application to the Commission is necessary to fix Camirus thoracicus Sharp, 1876 as the type species of Camiarus Sharp, 1878.

Family-group name: CAMIARINAE Jeannel, 1911. Stem: Camiar-.

Campsosternus Latreille, 1834: 141. Current status: valid genus in ELATERIDAE: DENDROMETRINAE: OXYNOPTERINI.

Type species: Elater fulgens G.-A. Olivier, 1790 (= Elater auratus Drury, 1773), by monotypy.

Family-group name: CAMPSOSTERNINAE Fleutiaux, 1927. Stem: Campsostern-.

Camptocerus Dejean, 1821: 100. Current status: valid genus in CURCULIONIDAE: SCOLYTINAE: SCOLYTINI.

Type species: Hylesinus aeneipennis Fabricius, 1801, by monotypy.

Family-group name: CAMPTOCÉRIDES Lacordaire, 1865. Stem: Camptocer-. Note: senior homonym of CAMPTOCERINI Wagner, 1961 (type genus Camptocera Jakovlev, 1877) in Hemiptera.

Camptorhinus Schönherr, 1825: 585. Current status: valid genus in CURCULIONIDAE: CRYPTORHYNCHINAE: CAMPTORHININI.

Type species: Curculio statua Fabricius, 1792 (= Curculio statua Rossi, 1790), by original designation.

Family-group name: CAMPTORHINIDES Lacordaire, 1865. Stem: Camptorhin-.

Campyloscelus Schönherr, 1845: 197. Current status: valid genus in CURCULIONIDAE: CONODERINAE: CAMPYLOSCELINI: CAMPYLOSCELINA.

Type species: Campyloscelus westermanni Boheman, 1845, by original designation.

Family-group name: CAMPYLOSCELIDES Schönherr, 1845. Stem: Campyloscel-.

Campyloxenus Fairmaire & Germain, 1860a: 6. Current status: valid genus in ELATERIDAE: PITYOBIINAE: CAMPYLOXENINI.

Type species: Campyloxenus pyrothorax Fairmaire & Germain, 1860, by monotypy.

Family-group name: CAMPYLOXENINAE C. Costa, 1975. Stem: Campyloxen-.

Campylus Fischer, 1823a: pl. 24. Current status: junior synonym of Denticollis Piller & Mitterpacher, 1783 in ELATERIDAE: DENDROMETRINAE: DENDROMETRINI: DENTICOLLINA.

Type species: Elater mesomelus Linnaeus, 1758 (= Elater linearis Linnaeus, 1758), by subsequent designation (Latreille 1830: 76). Note: Latreille (1830: 76) selected Elater linearis Linnaeus, 1758 as type species of Campylus Fischer, 1823, which is not an originally included species, but since he listed it in synonymy with Elater mesomelus Linnaeus, 1758, an originally included species, he is deemed to have designated the latter species as type species (ICZN 1999a, Article 69.2.2).

Family-group name: CAMPYLIDAE Gistel, 1848. Stem: Campyl-.

Canephorotomus Voss, 1954: 212. Current status: junior synonym of Amitrus Schönherr, 1840 in CURCULIONIDAE: ENTIMINAE: POLYDRUSITAE: NAUPACTINI: NAUPACTINA.

Type species: Canephorus jelskyi Kirsch, 1890, by original designation.

Family-group name: CANEPHOROTOMINA Voss, 1954. Stem: Canephorotom-.

Canonopsis C.O. Waterhouse, 1875: 54. Current status: valid genus in CURCULIONIDAE: ENTIMINAE: OTIORHYNCHITAE: ECTEMNORHININI.

Type species: Canonopsis sericea C.O. Waterhouse, 1875, by monotypy.

Family-group name: CANONOPSINI Dreux & Voisin, 1989. Stem: Canonopse-.

Cantharis Linnaeus, 1758: 342, 400. Current status: valid genus in CANTHARIDAE: CANTHARINAE: CANTHARINI.

Type species: Cantharis fusca Linnaeus, 1758, by subsequent designation (C.G. Thomson 1859: 110).

Family-group name: CANTHARIDAE Imhoff, 1856. Stem: Canthar-. Note: junior homonym of CANTHARINI Bonaparte, 1831 (type genus Cantharus Cuvier, 1816) in Pisces; Bonaparte’s name is permanently invalid (based on a preoccupied type genus).

Cantharocnemis Audinet-Serville, 1832: 132. Current status: valid genus in CERAMBYCIDAE: PRIONINAE: CANTHAROCNEMINI.

Type species: Cantharocnemis spondyloides Audinet-Serville, 1832, by monotypy.

Family-group name: CANTHAROCNEMITAE J. Thomson, 1861. Stem: Cantharocnem-.

Canthon Hoffmannsegg, 1817: 38. Current status: valid genus in SCARABAEIDAE: SCARABAEINAE: DELTOCHILINI.

Type species: Scarabaeus volvens Fabricius, 1792 (= Scarabaeus pilularius Linnaeus, 1758), by subsequent designation (Duponchel 1842b: 131). Note: the type species has been cited in recent literature (e.g., Cupello and Vaz-de-Mello 2018) as Scarabaeus pilularius Linnaeus, 1758 by subsequent designation of Paulian (1939: 17); however, Duponchel’s fixation is older.

Family-group name: CANTHONIDES Van Lansberge, 1874. Stem: Canthon-.

Capnisa Dejean, 1836b: 197. Current status: junior synonym of Gnathosia Fischer, 1821 in TENEBRIONIDAE: PIMELIINAE: TENTYRIINI.

Type species: Bradyus karelini Faldermann, 1836, by monotypy. Note: see Bousquet and Bouchard (2013b: 231) regarding the date of publication of Bradyus karelini Faldermann, 1836.

Family-group name: CAPNISINI Casey, 1907. Stem: Capnis-.

Capnodis Eschscholtz, 1829b: 9. Current status: valid genus in BUPRESTIDAE: CHRYSOCHROINAE: DICERCINI: DICERCINA.

Type species: Buprestis tenebricosa G.-A. Olivier, 1790, by subsequent designation (Duponchel 1842b: 136).

Family-group name: CAPNODINI Csiki, 1909. Stem: Capnod-.

Cappadox Alonso-Zarazaga & Lyal, 1999: 29. Current status: junior synonym of Phloeophilus Schönherr, 1833 in ANTHRIBIDAE: ANTHRIBINAE: PHLOEOPHILINI.

Type species: Phloeophilus sulcifrons Fåhraeus, 1839, by original designation.

Family-group name: CAPPADOCINI Alonso-Zarazaga & Lyal, 1999. Stem: Cappadoc-.

Capreolucanus Didier, 1929: 65. Current status: valid genus in LUCANIDAE: LUCANINAE: CAPREOLUCANINI.

Type species: Capreolucanus sicardi Didier, 1929, by original designation.

Family-group name: CAPREOLUCANINI H. Huang & C.H. Chen, 2013: 69. Stem: Capreolucan-.

Capula Jakobson, 1925: 145. Current status: valid genus in CHRYSOMELIDAE: GALERUCINAE: HYLASPINI.

Type species: Capula metallica Jakobson, 1925, by original designation.

Family-group name: CAPULINI Ogloblin, 1936. Stem: Capul-. Note: junior homonym of CAPULIDAE Fleming, 1822 (type genus Capulus Montfort, 1810) in Mollusca.

Car Blackburn, 1897: 35. Current status: valid genus in CARIDAE: CARINAE.

Type species: Car condensatus Blackburn, 1897, by monotypy.

Family-group name: CARINAE Thompson, 1992. Stem: Car-.

Carabdytes Balke, Hendrich & Wewalka, 1992: 94. Current status: valid genus in DYTISCIDAE: COLYMBETINAE.

Type species: Carabdytes upin Balke, Hendrich & Wewalka, 1992, by original designation.

Family-group name: CARABDYTINI Pederzani, 1995. Stem: Carabdyt-.

Carabhydrus Watts, 1978: 26. Current status: valid genus in DYTISCIDAE: HYDROPORINAE: HYDROPORINI: STERNOPRISCINA.

Type species: Carabhydrus monteithi Watts, 1978, by original designation.

Family-group name: CARABHYDRINI Watts, 1978. Stem: Carabhydr-.

Carabidomemnus Kolbe, 1924a: 346. Current status: valid genus in CARABIDAE: PAUSSINAE: PAUSSINI: CARABIDOMEMNINA. Note: this name was also proposed later in the same year by Kolbe (1924b: 1).

Type species: Arthropterus pallidus Raffray, 1886, by subsequent designation (Wasmann 1928: 271).

Family-group name: CARABIDOMEMNINAE Wasmann, 1928. Stem: Carabidomemn-.

Carabus Linnaeus, 1758: 342, 413. Current status: valid genus in CARABIDAE: CARABINAE: CARABINI: CARABINA. Note: name placed on the Official List of Generic Names in Zoology (ICZN 1954a).

Type species: Carabus granulatus Linnaeus, 1758, by plenary power (see ICZN 1954a). Note: name placed on the Official List of Specific Names in Zoology (ICZN 1954a).

Family-group name: CARABICI Latreille, 1802. Stem: Carab-.

Caraphia Gahan, 1906: 75. Current status: valid genus in CERAMBYCIDAE: LEPTURINAE: CARAPHIINI.

Type species: Caraphia cribrata Gahan, 1906, by original designation.

Family-group name: CARAPHIINI Ohbayashi, Lin & Yamasako, 2016: 189. Stem: Caraphi-.

Carcilia Roelofs, 1875: 152. Current status: valid genus in CURCULIONIDAE: MESOPTILIINAE: CARCILIINI.

Type species: Carcilia strigicollis Roelofs, 1875, by monotypy.

Family-group name: CARCILIINAE Pierce, 1916. Stem: Carcili-.

Cardiophorus Eschscholtz, 1829a: 34. Current status: valid genus in ELATERIDAE: CARDIOPHORINAE.

Type species: Elater thoracicus Fabricius, 1775 (= Elater gramineus Scopoli, 1763), by subsequent designation (Curtis 1838a: pl. 694).

Family-group name: CARDIOPHORITES Candèze, 1859. Stem: Cardiophor-.

Cardiorhinus Eschscholtz, 1829a: 34. Current status: valid genus in ELATERIDAE: ELATERINAE: AGRIOTINI: CARDIORHININA.

Type species: Elater plagiatus Germar, 1823, by subsequent designation (Duponchel 1842b: 168).

Family-group name: CARDIORHINITES Candèze, 1863. Stem: Cardiorhin-.

Carenum Bonelli, 1813: 479. Current status: valid genus in CARABIDAE: SCARITINAE: CARENINI.

Type species: here fixed (under ICZN 1999a, Article 70.3.2) as Carenum bonellii Westwood, 1842, misidentified as Scarites cyaneus Fabricius, 1775 (sensu Bonelli, 1813), in the original designation by Bonelli (1813: 479). Note: see Westwood (1842b: 83) regarding the misidentification of the type species.

Family-group name: CARENIDES W.J. MacLeay, 1887. Stem: Caren-.

Carilia Mulsant, 1863: 489. Current status: valid genus in CERAMBYCIDAE: LEPTURINAE: ACMAEOPINI.

Type species: Leptura virginea Linnaeus, 1758, by monotypy.

Family-group name: CARILIINI Zamoroka, 2022: 308, syn. nov. Stem: Carili-. Note: this family-group name is unnecessary since the name ACMAEOPINI Della Beffa, 1915 is already available for this taxon.

Carinodula Gordon, Pakaluk & Ślipiński, 1989: 360. Current status: valid genus in COCCINELLIDAE: MICROWEISEINAE: CARINODULINI.

Type species: Carinodula campbelli Gordon, Pakaluk & Ślipiński, 1989, by original designation.

Family-group name: CARINODULINI Gordon, Pakaluk & Ślipiński, 1989. Stem: Carinodul-.

Carodes Zimmerman, 1994: 511. Current status: valid genus in CARIDAE: CHILECARINAE: CARODESINI.

Type species: Carodes revelatus Zimmerman, 1994, by original designation.

Family-group name: CARODESINA Legalov, 2009. Stem: Carodes- (ICZN 1999a, Article 29.4).

Carpelimus Leach in Samouelle, 1819: 174. Current status: valid genus in STAPHYLINIDAE: OXYTELINAE: THINOBIINI.

Type species: Oxytelus fuliginosus Gravenhorst, 1802, by subsequent monotypy (Stephens 1829b: 24, as Carpalimus).

Family-group name: CARPELIMINI Hatch, 1957. Stem: Carpelim-.

Carphoborus Eichhoff, 1864: 27, 44. Current status: valid genus in CURCULIONIDAE: SCOLYTINAE: POLYGRAPHINI.

Type species: Hylesinus minimus Fabricius, 1801, by subsequent designation (Lacordaire 1865: 362).

Family-group name: CARPHOBORINAE Nüsslin, 1911. Stem: Carphobor-.

Carphodicticus Wood, 1971: 19. Current status: valid genus in CURCULIONIDAE: SCOLYTINAE: CARPHODICTICINI.

Type species: Carphodicticus cristatus Wood, 1971, by original designation.

Family-group name: CARPHODICTICINI Wood, 1971. Stem: Carphodictic-.

Carphurus Erichson, 1840a: 132. Current status: valid genus in MELYRIDAE: MALACHIINAE: CARPHURINI.

Type species: Carphurus dispar Erichson, 1840, by subsequent designation (Champion 1923b: 9).

Family-group name: CARPHURINAE Champion, 1923. Stem: Carphur-.

Carpophagus W.S. MacLeay, 1826: 447. Current status: valid genus in CHRYSOMELIDAE: SAGRINAE: CARPOPHAGINI.

Type species: Carpophagus banksiae W.S. MacLeay, 1826, by monotypy.

Family-group name: CARPOPHAGITES Chapuis, 1874. Stem: Carpophag-. Note: junior homonym of CARPOPHAGINAE Selby, 1835 (type genus Carpophaga Selby, 1835) in Aves.

Carpophilus Stephens, 1829b: 7. Current status: valid genus in NITIDULIDAE: CARPOPHILINAE.

Type species: Nitidula flexuosa Herbst, 1790 (= Dermestes hemipterus Linnaeus, 1758), by monotypy.

Family-group name: CARPOPHILINAE Erichson, 1842. Stem: Carpophil-.

Caryedon Schönherr, 1823: 1134. Current status: valid genus in CHRYSOMELIDAE: BRUCHINAE: PACHYMERINI: CARYEDONTINA.

Type species: Bruchus serratus G.-A. Olivier, 1790, by original designation.

Family-group name: CARYEDINI Bridwell, 1929. Stem: Caryedont-.

Caryonoda Bechyné, 1951: 265. Current status: valid genus in CHRYSOMELIDAE: EUMOLPINAE: CARYONODINI.

Type species: Caryonoda kuscheli Bechyné, 1951, by original designation.

Family-group name: CARYONODINI Bechyné, 1951. Stem: Caryonod-.

Caryopemon Jekel, 1855: 25. Current status: valid genus in CHRYSOMELIDAE: BRUCHINAE: PACHYMERINI: CARYOPEMINA. Note: two original spellings were used, including Carpopemon (p. 26); as far as we know no one has acted as First Reviser (ICZN 1999a, Article 24.2.3) and we here select Caryopemon, the spelling in prevailing usage.

Type species: Caryopemon hieroglyphicus Jekel, 1855, by original designation.

Family-group name: CARYOPEMINI Bridwell, 1929. Stem: Caryopem-.

Casignetus W.S. MacLeay, 1819: 98. Current status: valid genus in LUCANIDAE: LUCANINAE: CASIGNETINI.

Type species: Casignetus geotrupoides W.S. MacLeay, 1819 (= Lamprima humboldti Gyllenhal, 1817), by monotypy.

Family-group name: CASIGNETINI Reid, 1999. Stem: Casignet-. Note: junior homonym of CASIGNETINI Falkovitsh, 1972 (type genus Casigneta Wallengren, 1881) in Lepidoptera; Falkovitsh’s name is permanently invalid (i.e., its type genus is preoccupied by Casigneta Brunner von Wattenwyl, 1878, in Orthoptera).

Cassida Linnaeus, 1758: 342, 362. Current status: valid genus in CHRYSOMELIDAE: CASSIDINAE: CASSIDINI. Note: name placed on the Official List of Generic Names in Zoology (ICZN 1974b).

Type species: Cassida nebulosa Linnaeus, 1758, by subsequent designation (Spaeth 1914: 140; see ICZN 1974b). Note: name placed on the Official List of Specific Names in Zoology (ICZN 1974b).

Family-group name: CASSIDEAE Gyllenhal, 1813. Stem: Cassid-.

Catachirotes Gistel, 1856: 384. Current status: junior synonym of Calopus Fabricius, 1775 in OEDEMERIDAE: CALOPODINAE.

Type species: Cerambyx serraticornis Linnaeus, 1758, by monotypy.

Family-group name: CATACHIROTIDAE Gistel, 1856. Stem: Catachirotet-.

Catadromus W.S. MacLeay, 1825: 18. Current status: valid genus in CARABIDAE: HARPALINAE: PTEROSTICHINI: PTEROSTICHINA.

Type species: Carabus tenebrioides G.-A. Olivier, 1790, by monotypy.

Family-group name: CATADROMIENS Brullé, 1834. Stem: Catadrom-.

Cataphronetis P.H. Lucas, 1846: pl. 30. Current status: junior synonym of Phtora Germar, 1836 in TENEBRIONIDAE: DIAPERINAE: PHALERIINI.

Type species: Cataphronetis levaillantii P.H. Lucas, 1846 (= Phtora crenata Germar, 1836), by monotypy.

Family-group name: CATAPHRONETINI Reitter, 1917. Stem: Cataphronet-.

Catapiesis Solier in Brullé, 1835: 42. Current status: valid genus in CARABIDAE: HARPALINAE: CATAPIESEINI.

Type species: Catapiesis nitida Solier, 1835, by subsequent designation (Hope 1838a: 108).

Family-group name: CATAPIESINAE H.W. Bates, 1882. Stem: Catapiese-.

Catapiestus Perty, 1831: xxxviii. Current status: valid genus in TENEBRIONIDAE: STENOCHIINAE: CNODALONINI.

Type species: Catapiestus piceus Perty, 1831, by monotypy.

Family-group name: CATAPIESTIDES Lacordaire, 1859. Stem: Catapiest-.

Catapion Schilsky, 1906: vi. Current status: valid genus in BRENTIDAE: APIONINAE: APIONITAE: CATAPIINI.

Type species: Apion seniculus W. Kirby, 1808, by original designation.

Family-group name: CATAPIINA Alonso-Zarazaga, 1990. Stem: Catapi-.

Catascopus W. Kirby, 1823a: 94. Current status: valid genus in CARABIDAE: HARPALINAE: LEBIINI: PERICALINA.

Type species: Catascopus hardwickii W. Kirby, 1823, by monotypy.

Family-group name: CATASCOPI Csiki, 1932. Stem: Catascop-.

Cathormiocerus Schönherr, 1842b: 120. Current status: valid genus in CURCULIONIDAE: ENTIMINAE: CYPHICERITAE: TRACHYPHLOEINI: TRACHYPHLOEINA.

Type species: Trachyphloeus horrens Gyllenhal, 1834, by original designation.

Family-group name: CATHORMIOCERINI Reitter, 1913. Stem: Cathormiocer-.

†Catinius Ponomarenko, 1968: 137. Current status: valid genus in †CATINIIDAE. Note: Jurassic and Early Cretaceous (Kazakhstan; China).

Type species: †Catinius pelta Ponomarenko, 1968, by original designation.

Family-group name: †CATINIIDAE Ponomarenko, 1968. Stem: Catini-. Note: junior homonym of CATINIIDAE Bocquet & Stock, 1957 (type genus Catinia Bocquet & Stock, 1957) in Crustacea.

Catogenus Westwood, 1830b: 221. Current status: valid genus in PASSANDRIDAE: PASSANDRINAE.

Type species: Cucujus rufus Fabricius, 1798, by original designation.

Family-group name: CATOGENINI Grouvelle, 1916. Stem: Catogen-.

Catopocerus Motschulsky, 1870: 351. Current status: valid genus in LEIODIDAE: CATOPOCERINAE: CATOPOCERINI.

Type species: Catopocerus politus Motschulsky, 1870, by monotypy.

Family-group name: CATOPOCERINI Hatch, 1927. Stem: Catopocer-.

Catopochrotus Reitter, 1889d: 290. Current status: valid genus in CRYPTOPHAGIDAE: CRYPTOPHAGINAE: CRYPTOPHAGINI.

Type species: Catopochrotus crematogastri Reitter, 1889, by monotypy.

Family-group name: CATOPOCHROTIDAE Reitter, 1889. Stem: Catopochrot-.

Catops Paykull, 1798: 342. Current status: valid genus in LEIODIDAE: CHOLEVINAE: CHOLEVINI: CATOPINA.

Type species: Helops fuscus Panzer, 1794, misidentified as Tritoma sericea Fabricius, 1787 (sensu Paykull, 1798) in the subsequent designation by Jeannel (1922: 44). Note: we follow Peck et al. (2020: 36) in using Article 69.2.4 (ICZN 1999a) to determine the identity of the type species for this genus.

Family-group name: CATOPIDES Chaudoir, 1845. Stem: Catop-.

Catoxantha Solier, 1833: 261, 266. Current status: invalid senior synonym of Xanthocata Kubáň, 2016 in BUPRESTIDAE: CHRYSOCHROINAE: CHRYSOCHROINI: CHRYSOCHROINA. Note: junior homonym of Catoxantha Dejean, 1833 [Coleoptera: BUPRESTIDAE] (see Bousquet and Bouchard 2013a: 16, 17).

Type species: Buprestis opulenta Gory, 1832, by monotypy.

Family-group name: CATOXANTHINA Jakobson, 1913. Stem: Catoxanth-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

Catypnes Pascoe, 1864a: 243. Current status: valid genus in CERAMBYCIDAE: PRIONINAE: CATYPNINI.

Type species: Catypnes macleayi Pascoe, 1864, by monotypy.

Family-group name CATYPNIDES Lacordaire, 1868. Stem: Catypn-. Note: this name was treated as unavailable by Bouchard et al. (2011: 454) but was accepted as valid and attributed to Lacordaire, 1868 by Jin et al. (2023: 510) and is therefore available from that date.

Caulobius Le Guillou, 1844: 224. Current status: invalid senior synonym of Deuterocaulobius Dalla Torre, 1912 in SCARABAEIDAE: SERICOIDINAE: AUTOMOLIINI. Note: junior homonym of Caulobius Duponchel, 1838 [Lepidoptera].

Type species: Caulobius villosus Le Guillou, 1844, by monotypy.

Family-group name: CAULOBIINA Burmeister, 1855. Stem: Caulobi-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

Cautires C.O. Waterhouse, 1880a: 36. Current status: valid genus in LYCIDAE: METRIORRHYNCHINAE: METRIORRHYNCHINI: CAUTIRINA.

Type species: Lycus excellens C.O. Waterhouse, 1878, by subsequent designation (Kleine 1936: 4).

Family-group name: CAUTIRINA Sklenarova, Kubecek & Bocák, 2014: 52. Stem: Cautir-.

Cavicoxum Pic, 1928a: 21. Current status: junior synonym of Agraeus Candèze, 1857 in ELATERIDAE: AGRYPNINAE: AGRYPNINI.

Type species: Cavicoxum monstrosum Pic, 1928, by monotypy.

Family-group name: CAVICOXUMIDAE Pic, 1928. Stem: Cavicox-.

Cavognatha Crowson, 1965: 241. Current status: junior synonym of Taphropiestes Reitter, 1875 in CAVOGNATHIDAE.

Type species: Cavognatha pullivora Crowson, 1965, by original designation.

Family-group name: CAVOGNATHINAE Sen Gupta & Crowson, 1966. Stem: Cavognath-.

Ceballosmelasis Cobos, 1965: 381. Current status: valid genus in EUCNEMIDAE: MELASINAE: CEBALLOSMELASINI.

Type species: Ceballosmelasis elongatissimus Cobos, 1965, by monotypy.

Family-group name: CEBALLOSMELASINI Muona, 1993. Stem: Ceballosmelas-.

Cebrio G.-A. Olivier, 1790a: [30bis] 1, 4. Current status: valid genus in ELATERIDAE: ELATERINAE: CEBRIONINI: CEBRIONINA.

Type species: Cistela gigas Fabricius, 1787, by subsequent designation (Latreille 1810: 426).

Family-group name: CEBRIONATES Latreille, 1802. Stem: Cebrion-.

Cebriognathus Chobaut, 1899: 22. Current status: junior synonym of Analestesa Leach, 1824 in ELATERIDAE: CARDIOPHORINAE.

Type species: Cebriognathus desertorum Chobaut, 1899 (= Analestesa testacea Leach, 1824), by monotypy.

Family-group name: CEBRIOGNATHINAE Paulus, 1981. Stem: Cebriognath-.

Cechenus Fischer, 1822: 110. Current status: invalid senior synonym of Cechenochilus Motschulsky, 1850 in CARABIDAE: CARABINAE: CARABINI: CARABINA. Note: junior homonym of Cechenus Illiger, 1807 [Hymenoptera].

Type species: Carabus boeberi Adams, 1817, by subsequent designation (Froriep 1825: 348).

Family-group name: CECHENOGENICI Csiki, 1927. Stem: Cechen-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

Celaenephes Schmidt-Göbel, 1846: 77. Current status: valid genus in CARABIDAE: HARPALINAE: LEBIINI: CELAENEPHINA.

Type species: Celaenephes parallelus Schmidt-Göbel, 1846, by monotypy.

Family-group name: CELAENEPHINA Habu, 1982. Stem: Celaeneph-.

Celetes Schönherr, 1836: 634. Current status: valid genus in CURCULIONIDAE: CURCULIONINAE: DERELOMINI: DERELOMINA.

Type species: Celetes binotatus Gyllenhal, 1836, by original designation.

Family-group name: CELETINI Voss, 1954. Stem: Celet-.

Celeuthetes Schönherr, 1842b: 250. Current status: valid genus in CURCULIONIDAE: ENTIMINAE: OTIORHYNCHITAE: CELEUTHETINI: CELEUTHETINA. Note: unnecessary new replacement name for Sphaeromus Schönherr, 1834.

Type species: Sphaeromus australis Gyllenhal, 1834 (= Curculio echinatus Fabricius, 1801), by original designation (type fixation for Sphaeromus Schönherr).

Family-group name: CÉLEUTHÉTIDES Lacordaire, 1863. Stem: Celeuthet-.

Celina Aubé, 1837: 218, 219. Current status: valid genus in DYTISCIDAE: HYDROPORINAE: METHLINI.

Type species: Hydroporus latipes Brullé, 1836, by monotypy.

Family-group name: CELININI Branden, 1884. Stem: Celin-.

Celioschesis Tschitschérine, 1898: 93. Current status: junior synonym of Aristopus LaFerté-Sénectère, 1853 in CARABIDAE: HARPALINAE: ABACETINI: ABACETINA.

Type species: Distrigus bipustulatus Brullé, 1834, by subsequent designation (Jeannel 1948: 442).

Family-group name: CELIOSCHESINI Jeannel, 1948. Stem: Celioschese-.

Cenocephalus Chapuis, 1865: 325. Current status: valid genus in CURCULIONIDAE: PLATYPODINAE: TESSEROCERINI: TESSEROCERINA.

Type species: Cenocephalus thoracicus Chapuis, 1865, by monotypy.

Family-group name: CENOCEPHALARIAE H. Strohmeyer, 1914. Stem: Cenocephal-.

Centrinus Schönherr, 1825: 586. Current status: valid genus in CURCULIONIDAE: BARIDINAE: APOSTASIMERINI: DIORYMERINA.

Type species: Baris bicuspis Germar, 1823, by subsequent designation (Schönherr 1826: 22).

Family-group name: CENTRINIDES Jekel, 1865. Stem: Centrin-. Note: junior homonym of CENTRINIDAE Swainson, 1838 (type genus Centrina Cuvier, 1816) in Pisces.

Centrioptera Mannerheim, 1843: 279. Current status: junior synonym of Cryptoglossa Solier, 1837 in TENEBRIONIDAE: PIMELIINAE: CRYPTOGLOSSINI.

Type species: Centrioptera caraboides Mannerheim, 1843, by monotypy.

Family-group name: CENTRIOPTÉRIDES Lacordaire, 1859. Stem: Centriopter-.

Centrocorynus Jekel, 1860: 167. Current status: valid genus in ATTELABIDAE: APODERINAE: APODERINI.

Type species: Apoderus scutellaris Gyllenhal, 1833, by original designation.

Family-group name: CENTROCORYNINA Legalov, 2003. Stem: Centrocoryn-.

Centronopus Solier, 1848b: 154, 258. Current status: valid genus in TENEBRIONIDAE: TENEBRIONINAE: CENTRONOPINI.

Type species: Centronopus extensicollis Solier, 1848 (= Tenebrio suppressus Say, 1835), by original designation.

Family-group name: CENTRONOPINI Doyen, 1989. Stem: Centronop-.

Centrophthalmus Schmidt-Göbel, 1838: 7. Current status: valid genus in STAPHYLINIDAE: PSELAPHINAE: PSELAPHITAE: TYRINI: CENTROPHTHALMINA.

Type species: Centrophthalmus paria Schmidt-Göbel, 1838, by monotypy.

Family-group name: CENTROPHTHALMINA Jeannel, 1949. Stem: Centrophthalm-.

Ceophyllus J.L. LeConte, 1849: 73. Current status: valid genus in STAPHYLINIDAE: PSELAPHINAE: PSELAPHITAE: TYRINI: TYRINA.

Type species: Ceophyllus monilis J.L. LeConte, 1849, by monotypy.

Family-group name: CEOPHYLLINI Park, 1951. Stem: Ceophyll-.

Cephacerus Rafinesque, 1815: 113. Current status: junior synonym of Erodius Fabricius, 1775 in TENEBRIONIDAE: PIMELIINAE: ERODIINI. Note: unnecessary new replacement name for Erodius Fabricius, 1775.

Type species: Erodius gibbus Fabricius, 1775, by subsequent designation (Latreille 1810: 429) (type fixation for Erodius Fabricius).

Family-group name: CEPHACERIA Rafinesque, 1815. Stem: Cephacer-.

Cephalobyrrhus Pic, 1923: 4. Current status: valid genus in LIMNICHIDAE: CEPHALOBYRRHINAE.

Type species: Cephalobyrrhus latus Pic, 1923, by subsequent designation (Satô 1965: 123).

Family-group name: CEPHALOBYRRHINAE Champion, 1925. Stem: Cephalobyrrh-.

Cephalodonta Chevrolat, 1842b: 272. Current status: junior synonym of Sceloenopla Chevrolat, 1836 in CHRYSOMELIDAE: CASSIDINAE: SCELOENOPLINI.

Type species: Chalepus goniapterus Perty, 1832, by monotypy.

Family-group name: CEPHALODONTINAE Gestro, 1906. Stem: Cephalodont-.

Cephaloleia Chevrolat in Dejean, 1836a: 366. Current status: valid genus in CHRYSOMELIDAE: CASSIDINAE: CEPHALOLEIINI.

Type species: Hispa nigricornis Fabricius, 1792, by subsequent designation (Staines 1991: 247).

Family-group name: CÉPHALOLÉITES Chapuis, 1875. Stem: Cephalolei-.

Cephaloon Newman, 1838: 376. Current status: valid genus in STENOTRACHELIDAE: CEPHALOINAE.

Type species: Cephaloon lepturides Newman, 1838, by monotypy.

Family-group name: CEPHALOIDAE J.L. LeConte, 1862. Stem: Cephalo-.

Cephalophanus John, 1940: 82. Current status: valid genus in DISCOLOMATIDAE: CEPHALOPHANINAE.

Type species: Cephalophanus clipeoexcisus John, 1940, by original designation.

Family-group name: CEPHALOPHANINAE John, 1954. Stem: Cephalophan-.

Cephaloplectus Sharp, 1883a: 295. Current status: valid genus in PTILIIDAE: PTILIINAE: CEPHALOPLECTINI.

Type species: Cephaloplectus godmani Sharp, 1883, by monotypy.

Family-group name: CEPHALOPLECTINAE Sharp, 1883. Stem: Cephaloplect-.

Cephaloscymnus Crotch, 1873c: 382. Current status: valid genus in COCCINELLIDAE: COCCINELLINAE: CEPHALOSCYMNINI.

Type species: Cephaloscymnus zimmermanni Crotch, 1873, by monotypy.

Family-group name: CEPHALOSCYMNINI Gordon, 1985. Stem: Cephaloscymn-.

Cephalotes Bonelli, 1810: Tab Syn. Current status: invalid senior synonym of Broscus Panzer, 1813 in CARABIDAE: BROSCINAE: BROSCINI: BROSCINA. Note: junior homonym of Cephalotes Latreille, 1802 [Hymenoptera].

Type species: Carabus cephalotes Linnaeus, 1758, by subsequent monotypy in Panzer (1813: 62) (type fixation for Broscus Panzer).

Family-group name: CEPHALOTIDA Heer, 1838. Stem: Cephalot-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

Cephalotoma Lesne, 1911: 207. Current status: valid genus in BOSTRICHIDAE: LYCTINAE: CEPHALOTOMINI.

Type species: Cephalotoma singularis Lesne, 1911, by monotypy.

Family-group name: CEPHALOTOMINI L.Y. Liu in L.Y. Liu and Schönitzer, 2011: 99, 120. Stem: Cephalotom-.

Cephalotrichia Hope, 1837: 102. Current status: invalid senior synonym of Sparrmannia Laporte, 1840 (nomen protectum) in SCARABAEIDAE: MELOLONTHINAE: TANYPROCTINI: TANYPROCTINA. Note: nomen oblitum (see Bouchard et al. 2011: 255).

Type species: Melolontha alopex Fabricius, 1787, by original designation.

Family-group name: CEPHALOTRICHIADAE Burmeister, 1855. Stem: Cephalotrichi-.

Cephalotyphlus Coiffait, 1955: 68. Current status: valid genus in STAPHYLINIDAE: LEPTOTYPHLINAE: CEPHALOTYPHLINI.

Type species: Leptotyphlus revelierei Saulcy, 1878, by original designation.

Family-group name: CEPHALOTYPHLINI Coiffait, 1963. Stem: Cephalotyphl-.

Cephennium P.W.J. Müller in P.W.J. Müller and Kunze, 1822: 188. Current status: valid genus in STAPHYLINIDAE: SCYDMAENINAE: CEPHENNIITAE: CEPHENNIINI. Note: this name was first proposed by P.W.J. Müller in P.W.J. Müller and Kunze (1822: 188) in synonymy with Scydmaenus thoracicus P.W.J. Müller & Kunze, 1822; it is available from that date since it was treated as an available name before 1961 by several authors (e.g., Casey 1897: 500; Ganglbauer 1899: 11).

Type species: Scydmaenus thoracicus P.W.J. Müller & Kunze, 1822, by subsequent designation (E. Desmarest 1852: 148). Note: the type species usually listed for this genus is “Cephennium truncorum P.W.J. Müller, 1822 (= Scydmaenus thoracicus P.W.J. Müller & Kunze, 1822)”; however, Cephennium truncorum was first proposed by P.W.J. Müller in P.W.J. Müller and Kunze (1822: 188) in synonymy with Scydmaenus thoracicus P.W.J. Müller & Kunze, 1822 but, as far as we know, it is unavailable since it was not treated as an available name before 1961 and either adopted as the name of a taxon or treated as a senior homonym (ICZN 1999a, Article 11.6.1); the first available species associated with the name Cephennium are the three species, Scydmaenus thoracicus P.W.J. Müller & Kunze, 1822, Scydmaenus laticollis Aubé, 1843, and Scydmaenus minutissimus Aubé, 1843, listed by Erichson (1845b: 99).

Family-group name: CEPHENNIINI Reitter, 1882. Stem: Cephenni-.

Cepurus Schönherr, 1834b: 347. Current status: valid genus in CURCULIONIDAE: HYPERINAE: HYPERINI: CEPURINA.

Type species: Curculio torridus G.-A. Olivier, 1807, by original designation.

Family-group name: CÉPURIDES Capiomont, 1867. Stem: Cepur-.

Ceracupes Kaup, 1871: 16. Current status: valid genus in PASSALIDAE: AULACOCYCLINAE: CERACUPEDINI.

Type species: Passalus fronticornis Westwood, 1841, by monotypy.

Family-group name: CERACUPINI Boucher, 2006. Stem: Ceracup- (ICZN 1999a, Article 29.4).

†Ceracyclus Boucher & Bai in Boucher et al., 2017: 7. Current status: valid genus in PASSALIDAE: AULACOCYCLINAE: CYLINDROCAULINI. Note: mid-Cretaceous (Myanmar).

Type species: †Ceracyclus lotus Boucher, Bai & Montreuil, 2017, by original designation.

Family-group name: †CERACYCLINI Boucher & Bai in Boucher et al., 2017: 4. Stem: Ceracycl-.

Cerallus Jacquelin du Val, 1859a: 43. Current status: valid genus in MELYRIDAE: MELYRINAE: CERALLINI.

Type species: Dasytes rubidus Gyllenhal, 1817, by original designation.

Family-group name: CERALLINI Pic, 1929. Stem: Cerall-.

Cerambyciscapha Pic, 1915a: 30. Current status: valid genus in STAPHYLINIDAE: SCAPHIDIINAE: SCAPHIDIINI.

Type species: Cerambyciscapha dohertyi Pic, 1915, by monotypy.

Family-group name: CERAMBYCISCAPHINI Pic, 1915. Stem: Cerambyciscaph-.

Cerambyx Linnaeus, 1758: 342, 388. Current status: valid genus in CERAMBYCIDAE: CERAMBYCINAE: CERAMBYCINI: CERAMBYCINA.

Type species: Cerambyx cerdo Linnaeus, 1758, by subsequent designation (Latreille 1810: 431).

Family-group name: CERAMBICINI Latreille, 1802. Stem: Cerambyc-.

Cerapterus Swederus, 1788: 203. Current status: valid genus in CARABIDAE: PAUSSINAE: PAUSSINI: CERAPTERINA.

Type species: Cerapterus latipes Swederus, 1788, by monotypy.

Family-group name: CERAPTERIDES Billberg, 1820. Stem: Cerapter-.

Cerasommatidia Brèthes, 1925: 199. Current status: valid genus in CERASOMMATIDIIDAE.

Type species: Cerasommatidia arrowi Brèthes, 1925, by monotypy.

Family-group name: CERASOMMATIDIIDAE Brèthes, 1925. Stem: Cerasommatidi-.

Cerasphorus Audinet-Serville, 1834b: 10. Current status: valid genus in CERAMBYCIDAE: CERAMBYCINAE: HESPEROPHANINI: HESPEROPHANINA.

Type species [suggested]: Cerasphorus hirticornis Audinet-Serville, 1834, by subsequent designation (J. Thomson 1861: 236). Note: the valid type species fixation is that of E. Desmarest (1860: 312), who selected Stenocorus garganicus Fabricius, 1775, but that species is currently included as a junior synonym of Cerambyx cinctus Drury, 1773 in the genus Chion Newman, 1840 (see Bousquet 2008a: 619); an application to the Commission is necessary to fix Cerasphorus hirticornis Audinet-Serville, 1834 as the type species of Cerasphorus Audinet-Serville, 1834.

Family-group name: CERASPHORITAE J. Thomson, 1861. Stem: Cerasphor-.

Ceraspis Lepeletier & Audinet-Serville, 1828: 370. Current status: valid genus in SCARABAEIDAE: MELOLONTHINAE: MACRODACTYLINI.

Type species: Ceraspis pruinosa Lepeletier & Audinet-Serville, 1828 (= Melolontha bivulnerata Germar, 1823), by subsequent designation (Duponchel 1842b: 287).

Family-group name: CERASPIDIDAE Burmeister, 1855. Stem: Ceraspid-.

Ceratanisus Gemminger in Gemminger and Harold, 1870: 1818. Current status: valid genus in TENEBRIONIDAE: PIMELIINAE: CERATANISINI. Note: new replacement name for Anisocerus Faldermann, 1837.

Type species: Anisocerus tristis Faldermann, 1837, by monotypy (type fixation for Anisocerus Faldermann).

Family-group name: CERATANISINI Gebien, 1937. Stem: Ceratanis-.

Ceratapion Schilsky, 1901: 38F. Current status: valid genus in BRENTIDAE: APIONINAE: ASPIDAPIITAE: CERATAPIINI.

Type species: Apion carduorum W. Kirby, 1808, by original designation.

Family-group name: CERATAPIINI Alonso-Zarazaga, 1990. Stem: Ceratapi-.

Ceratocanthus A. White, 1842: 93. Current status: valid genus in HYBOSORIDAE: CERATOCANTHINAE: CERATOCANTHINI. Note: new replacement name for Acanthocerus W.S. MacLeay, 1819; the valid name for this genus should be Sphaeromorphus Dejean, 1833 (see Bousquet and Bouchard 2013a: 41); an application has been submitted to the Commission by A. Ballerio (Case 3766) to retain Ceratocanthus A. White, 1842 as valid.

Type species: Acanthocerus aeneus W.S. MacLeay, 1819, by subsequent designation (Duponchel 1839: 30) (type fixation for Acanthocerus W.S. MacLeay).

Family-group name: CERATOCANTHINI Martínez, 1968. Stem: Ceratocanth-.

Ceratoderus Westwood, 1841b: 112. Current status: valid genus in CARABIDAE: PAUSSINAE: PAUSSINI: PAUSSINA.

Type species: Paussus bifasciatus Kollar, 1836, by monotypy.

Family-group name: CERATODERINA Darlington, 1950. Stem: Ceratoder-.

Ceratognathus Westwood, 1838a: 260. Current status: valid genus in LUCANIDAE: AESALINAE: CERATOGNATHINI.

Type species: Ceratognathus niger Westwood, 1838, by original designation.

Family-group name: CERATOGNATHINI Sharp, 1899. Stem: Ceratognath-.

Ceratopus Schönherr, 1843: 120. Current status: valid genus in CURCULIONIDAE: CURCULIONINAE: CERATOPODINI.

Type species: Ceratopus bisignatus Boheman, 1843, by original designation.

Family-group name: CÉRATOPIDES Lacordaire, 1863. Stem: Ceratopod-.

Ceratorhina Westwood, 1843a: 170. Current status: junior synonym of Eudicella A. White, 1839 in SCARABAEIDAE: CETONIINAE: GOLIATHINI: RHOMBORHININA.

Type species: Goliathus grallii Buquet, 1936, by subsequent designation (Marais and Holm 1992: 14).

Family-group name: CERATORRHINIDAE Kraatz, 1880. Stem: Ceratorhin-.

Ceratotrupes Jekel, 1866: 540. Current status: valid genus in GEOTRUPIDAE: GEOTRUPINAE: CERATOTRUPINI.

Type species: Geotrupes fronticornis Erichson, 1847, by original designation.

Family-group name: CERATOTRUPINI Zunino, 1984. Stem: Ceratotrup-.

Cercus Latreille, 1797: 68. Current status: junior synonym of Kateretes Herbst, 1793 in KATERETIDAE: KATERETINAE.

Type species: Dermestes pedicularius Linnaeus, 1758, by subsequent monotypy (Latreille 1802b: 135).

Family-group name: CERCIDAE E. Desmarest, 1851. Stem: Cerc-.

Cercyon Leach, 1817: 95. Current status: valid genus in HYDROPHILIDAE: SPHAERIDIINAE: MEGASTERNINI: MEGASTERNINA.

Type species: Dermestes melanocephalus Linnaeus, 1758, by subsequent designation (C.G. Thomson 1859: 19).

Family-group name: CERCYONES G.H. Horn, 1890. Stem: Cercyon-.

Cerenopus J.L. LeConte, 1851a: 143. Current status: valid genus in TENEBRIONIDAE: TENEBRIONINAE: CERENOPINI.

Type species: Cerenopus concolor J.L. LeConte, 1851, by subsequent designation (R. Lucas 1920: 173).

Family-group name: CERENOPI G.H. Horn, 1870. Stem: Cerenop-.

Cerocoma Geoffroy, 1762: 357. Current status: valid genus in MELOIDAE: MELOINAE: CEROCOMINI. Note: name placed on the Official List of Generic Names in Zoology (ICZN 1994a).

Type species: Meloe schaefferi Linnaeus, 1758, by subsequent monotypy (Fabricius 1775: 262; see ICZN 1994a). Note: name placed on the Official List of Specific Names in Zoology (ICZN 1994a).

Family-group name: CEROCOMATIDA Leach, 1815. Stem: Cerocom-.

Cerocosmus Gemminger, 1873: 146. Current status: valid genus in PTINIDAE: ERNOBIINAE: ERNOBIINI. Note: new replacement name for Cosmocerus Solier, 1849.

Type species: Cosmocerus cinereus Solier, 1849, by monotypy (type fixation for Cosmocerus Solier).

Family-group name: CEROCOSMINAE Pic, 1912. Stem: Cerocosm-.

Ceroglossus Solier, 1848a: 58. Current status: valid genus in CARABIDAE: CARABINAE: CEROGLOSSINI.

Type species: Carabus chilensis Eschscholtz, 1829, by original designation.

Family-group name: CEROGLOSSINA Lapouge, 1927. Stem: Cerogloss-.

Cerophysa Chevrolat in Dejean, 1836a: 379. Current status: valid genus in CHRYSOMELIDAE: GALERUCINAE: LUPERINI.

Type species: Galleruca nodicornis Wiedemann, 1823, by monotypy.

Family-group name: CEROPHYSITES Chapuis, 1875. Stem: Cerophys-.

Cerophytum Latreille, 1809a: 375. Current status: valid genus in CEROPHYTIDAE.

Type species: Melasis elateroides Latreille, 1804, by monotypy.

Family-group name: CEROPHYTIDES Latreille, 1834. Stem: Cerophyt-.

Ceroplesis Audinet-Serville, 1835a: 93. Current status: valid genus in CERAMBYCIDAE: LAMIINAE: CEROPLESINI: CEROPLESINA. Note: name placed on the Official List of Generic Names in Zoology (ICZN 1986c).

Type species: Lamia aethiops Fabricius, 1775, by plenary power (see ICZN 1986c). Note: name placed on the Official List of Specific Names in Zoology (ICZN 1986c).

Family-group name: CEROPLESITAE J. Thomson, 1860. Stem: Ceroples-.

Cerotoma Chevrolat in Dejean, 1836a: 379. Current status: valid genus in CHRYSOMELIDAE: GALERUCINAE: LUPERINI.

Type species: Crioceris caminea Fabricius, 1801 (= Crioceris ruficornis G.-A. Olivier, 1791), by subsequent designation (Chapuis 1875: 230).

Family-group name: CÉROTOMITES Chapuis, 1875. Stem: Cerotom-.

Certallum Dejean, 1821: 111. Current status: valid genus in CERAMBYCIDAE: CERAMBYCINAE: CERTALLINI.

Type species: Saperda ruficollis Fabricius, 1787 (= Cerambyx ebulinus Linnaeus, 1767), by monotypy.

Family-group name: CARTALLITES Fairmaire, 1864. Stem: Certall-.

†Ceruchites Statz, 1952: 5. Current status: valid genus in LUCANIDAE: †CERUCHITINAE. Note: Oligocene (Germany).

Type species: †Ceruchites hahnei Statz, 1952, by monotypy.

Family-group name: †CERUCHITINAE Nikolajev, 2006. Stem: Ceruchit-.

Ceruchus W.S. MacLeay, 1819: 96, 115. Current status: valid genus in LUCANIDAE: SYNDESINAE: CERUCHINI.

Type species: Lucanus tenebroides Fabricius, 1787 (= Lucanus chrysomelinus Hochenwarth, 1785), by monotypy.

Family-group name: CERUCHITES Jacquelin du Val, 1859. Stem: Ceruch-.

Cerylon Latreille, 1802b: 205. Current status: valid genus in CERYLONIDAE: CERYLONINAE. Note: name placed on the Official List of Generic Names in Zoology (ICZN 1995d).

Type species: Lyctus histeroides Fabricius, 1792, by plenary power (see ICZN 1995d). Note: name placed on the Official List of Specific Names in Zoology (ICZN 1995d).

Family-group name: CERYLONIDES Billberg, 1820. Stem: Cerylon-.

Cesauletes Hamilton, 1983: 68. Current status: valid genus in ATTELABIDAE: RHYNCHITINAE: RHYNCHITITAE: CESAULETINI.

Type species: Auletes nasalis J.L. LeConte, 1876, by original designation.

Family-group name: CESAULETINI Legalov, 2003. Stem: Cesaulet-.

Cetonia Fabricius, 1775: 42. Current status: valid genus in SCARABAEIDAE: CETONIINAE: CETONIINI: CETONIINA.

Type species: Scarabaeus auratus Linnaeus, 1758, by subsequent designation (Latreille 1810: 429).

Family-group name: CETONIDA Leach, 1815. Stem: Cetoni-.

Ceutocerus Germar, 1823: 85. Current status: valid genus in MURMIDIIDAE.

Type species: Ceutocerus advena Germar, 1823 (= Hister ovalis Beck, 1817), by subsequent designation (Hope 1841a: 108).

Family-group name: CEUTHOCERA Mannerheim, 1852. Stem: Ceutocer-. Note: nomen oblitum (see Bouchard and Bousquet 2020a: 128).

Ceutorhynchus Germar, 1823: 217. Current status: valid genus in CURCULIONIDAE: CEUTORHYNCHINAE: CEUTORHYNCHINI. Note: name placed on the Official List of Generic Names in Zoology (ICZN 1989b, as Ceutorhynchus Germar, 1824).

Type species: Curculio assimilis Paykull, 1792, by subsequent designation (C.G. Thomson 1859: 140; see ICZN 1989b). Note: the senior homonym Curculio assimilis Fabricius, 1775 and all other uses of that name prior to Curculio assimilis Paykull, 1792 were suppressed for both the Principles of Priority and Homonymy and Curculio assimilis Paykull, 1792 was placed on the Official List of Specific Names in Zoology (ICZN 1989b).

Family-group name: CENTORHYNCHIDAE Gistel, 1848. Stem: Ceutorhynch-.

Chaeridiona Baly, 1869: 380. Current status: valid genus in CHRYSOMELIDAE: CASSIDINAE: ONCOCEPHALINI.

Type species: Chaeridiona metallica Baly, 1869, by subsequent designation (Maulik 1919: 85).

Family-group name: CHOERIDIONINI Weise, 1911. Stem: Chaeridion-.

Chaerodes A. White, 1846: 12. Current status: valid genus in TENEBRIONIDAE: LAGRIINAE: CHAERODINI.

Type species: Chaerodes trachyscelides A. White, 1846, by monotypy.

Family-group name: CHAERODINI Doyen, E.G. Matthews & Lawrence, 1990. Stem: Chaerod-.

Chaetarthria Stephens, 1833a: 22. Current status: valid genus in HYDROPHILIDAE: CHAETARTHRIINAE: CHAETARTHRIINI.

Type species: Hydrophilus seminulum Paykull, 1798 (= Hydrophilus seminulum Herbst, 1797), by monotypy.

Family-group name: CHAETARTHRIINI Bedel, 1881. Stem: Chaetarthri-.

Chaetocanthus Péringuey, 1901: 492, 495. Current status: valid genus in OCHODAEIDAE: CHAETOCANTHINAE: CHAETOCANTHINI.

Type species: Chaetocanthus insuetus Péringuey, 1901, by monotypy.

Family-group name: CHAETOCANTHINAE Scholtz, 1988. Stem: Chaetocanth-.

Chaetocnema Stephens, 1831b: 325. Current status: valid genus in CHRYSOMELIDAE: GALERUCINAE: ALTICINI.

Type species [suggested]: Galeruca aridella Paykull, 1799 (= Altica hortensis Geoffroy, 1785), by subsequent designation (C.G. Thomson 1859: 156). Note: the valid type species fixation is that of Westwood (1838c: 42), who selected Chrysomela concinna Marsham, 1802, but that species is currently included in the subgenus Tlanoma Motschulsky, 1845, not in the nominotypical subgenus; an application to the Commission is necessary to fix Galeruca aridella Paykull, 1799 as the type species of Chaetocnema Stephens, 1831.

Family-group name: CHAETOCNEMAE J.L. LeConte & G.H. Horn, 1883. Stem: Chaetocnem-.

Chaetodactyla Tschitschérine, 1897: 271. Current status: valid genus in CARABIDAE: HARPALINAE: CHAETODACTYLINI.

Type species: Chaetodactyla mirabilis Tschitschérine, 1897, by monotypy.

Family-group name: CHAETODACTYLINI Tschitschérine, 1903. Stem: Chaetodactyl-. Note: senior homonym of CHAETODACTYLIDAE Zachvatkin, 1941 (type genus Chaetodactylus Rondani, 1866) in Acari: Astigmata.

Chaetogenys Emden, 1958: 19. Current status: valid genus in CARABIDAE: HARPALINAE: CHAETOGENYINI.

Type species: Chaetogenys straneoi Emden, 1958, by original designation.

Family-group name: CHAETOGENYINA Emden, 1958. Stem: Chaetogeny-.

Chaetomalachius Kraatz, 1882b: 96. Current status: valid genus in MELYRIDAE: DASYTINAE: CHAETOMALACHIINI.

Type species: Chaetomalachius dasytoides Kraatz, 1882, by monotypy.

Family-group name: CHAETOMALACHIINI Majer, 1987. Stem: Chaetomalachi-.

Chaetophloeus J.L. LeConte in J.L. LeConte and G.H. Horn, 1876: 382. Current status: valid genus in CURCULIONIDAE: SCOLYTINAE: CHAETOPHLOEINI.

Type species: Hylesinus hystrix J.L. LeConte, 1858, by monotypy.

Family-group name: CHAETOPHLOEINI Jordal, 2021: 108. Stem: Chaetophloe-. Note: this family-group name was attributed to Schedl (1966: 361) in Bouchard et al. (2011: 635) but we agree with Alonso-Zarazaga and Lyal (2009: 74) and Jordal (2021: 108) that the family-group name is not available from Schedl (1966: 361) since a description was not provided.

Chaetosoma Westwood, 1851: 171. Current status: valid genus in CHAETOSOMATIDAE. Note: the senior homonym Chaetosoma Chevrolat, 1843 [Coleoptera: CERAMBYCIDAE] was suppressed for the purposes of both the Principles of Priority and Homonymy and placed on the Official Index of Rejected and Invalid Generic Names in Zoology while Chaetosoma Westwood, 1851 was placed on the Official List of Generic Names in Zoology (ICZN 2011d).

Type species: Chaetosoma scaritides Westwood, 1851, by monotypy (see ICZN 2011d). Note: name placed on the Official List of Specific Names in Zoology (ICZN 2011d).

Family-group name: CHAETOSOMATIDAE Crowson, 1952. Stem: Chaetosomat-. Note: the senior homonym CHAETOSOMATIDAE Claus, 1872 (type genus Chaetosoma Claparède, 1863) in Nematoda was suppressed for the purposes of both the Principles of Priority and Homonymy and placed on the Official Index of Rejected and Invalid Family-Group Names in Zoology (ICZN 2011d).

Chalcodrya L. Redtenbacher, 1868: 138. Current status: valid genus in CHALCODRYIDAE.

Type species: Chalcodrya variegata L. Redtenbacher, 1868, by monotypy.

Family-group name: CHALCODRYIDAE Watt, 1974. Stem: Chalcodry-.

Chalcolepidius Eschscholtz, 1829a: 32. Current status: valid genus in ELATERIDAE: AGRYPNINAE: HEMIRHIPINI: HEMIRHIPINA.

Type species: Elater sulcatus Fabricius, 1777, by subsequent designation (Duponchel 1843a: 371). Note: the type species often reported in the literature is Chalcolepidius zonatus Eschscholtz, 1829 by subsequent designation of Hyslop (1921: 634) but Duponchel’s typification is earlier (see Kundrata et al. 2019c: 103); both species are currently included in the genus Chalcolepidius.

Family-group name: CHALCOLÉPIDIIDES Candèze, 1857. Stem: Chalcolepidi-.

Chalcophana Chevrolat in Dejean, 1836a: 407. Current status: valid genus in CHRYSOMELIDAE: EUMOLPINAE: EUMOLPINI.

Type species: Colaspis hilaris Germar, 1823, by subsequent designation (Monrós and Bechyné 1956: 1125).

Family-group name: CHALCOPHANITES Chapuis, 1874. Stem: Chalcophan-. Note: junior homonym of CHALCOPHANINAE Sundevall, 1872 (type genus Chalcophanes Wagler, 1827) in Aves.

Chalcophora Dejean, 1833a: 77. Current status: valid genus in BUPRESTIDAE: CHRYSOCHROINAE: CHRYSOCHROINI: CHALCOPHORINA.

Type species: Buprestis mariana Linnaeus, 1758, by subsequent designation (Duponchel 1843a: 372).

Family-group name: CHALCOPHORIDES Lacordaire, 1857. Stem: Chalcophor-.

Chalcophorella Kerremans, 1903a: 79. Current status: valid genus in BUPRESTIDAE: CHRYSOCHROINAE: CHRYSOCHROINI: CHALCOPHORINA.

Type species: Buprestis stigmatica Dalman, 1817, by subsequent designation (Obenberger 1942: 4).

Family-group name: CHALCOPHORELLINI Tôyama, 1986. Stem: Chalcophorell-.

Chalcoplectus Oke, 1925: 13. Current status: valid genus in STAPHYLINIDAE: PSELAPHINAE: PSELAPHITAE: TYRINI: TYRINA.

Type species: Chalcoplectus depressus Oke, 1925, by monotypy.

Family-group name: CHALCOPLECTINI Oke, 1925. Stem: Chalcoplect-.

Chalcosoma Hope, 1837: 86. Current status: valid genus in SCARABAEIDAE: DYNASTINAE: DYNASTINI: CHALCOSOMINA.

Type species: Scarabaeus atlas Linnaeus, 1758, by original designation.

Family-group name: CHALCOSOMINA Rowland & K.B. Miller, 2012: 11. Stem: Chalcosom- (ICZN 1999a, Article 29.4).

Chalcothea Burmeister, 1842: 319. Current status: valid genus in SCARABAEIDAE: CETONIINAE: TAENIODERINI: CHALCOTHEINA.

Type species: Macronota resplendens Gory & Percheron, 1834, by subsequent designation (Mikšić 1976: 379)

Family-group name: CHALCOTHEINA Mikšić, 1976. Stem: Chalcothe-.

Chalepus W.S. MacLeay, 1819: 149. Current status: invalid senior synonym of Dyscinetus Harold, 1869 in SCARABAEIDAE: DYNASTINAE: CYCLOCEPHALINI. Note: junior homonym of Chalepus Thunberg, 1805 [Coleoptera: CHRYSOMELIDAE].

Type species: Melolontha geminata Fabricius, 1801 (= Melolontha dubia G.-A. Olivier, 1789), by original designation.

Family-group name: CHALEPIDAE Streubel, 1846. Stem: Chalep-. Note: senior homonym of CHALEPINI Weise, 1910 (type genus Chalepus Thunberg, 1805) in Coleoptera: CHRYSOMELIDAE; family-group name permanently invalid (ICZN 1999a, Article 39).

Chalepus Thunberg, 1805: 282. Current status: valid genus in CHRYSOMELIDAE: CASSIDINAE: CHALEPINI.

Type species: Hispa sanguinicollis Linnaeus, 1771, by subsequent designation (Crotch 1870b: 218).

Family-group name: CHALEPINI Weise, 1910. Stem: Chalep-. Note: nomen protectum (see Bouchard and Bousquet 2020a: 108); junior homonym of CHALEPIDAE Streubel, 1846 (type genus Chalepus W.S. MacLeay, 1819) in Coleoptera: SCARABAEIDAE; Streubel’s name is permanently invalid (based on a preoccupied type genus).

Chaltenia Roig-Juñent & Cicchino, 2001: 663. Current status: valid genus in CARABIDAE: TRECHINAE: SINOZOLINI: CHALTENIINA.

Type species: Chaltenia patagonia Roig-Juñent & Cicchino, 2001, by original designation.

Family-group name: CHALTENIINA Roig-Juñent & Cicchino, 2001. Stem: Chalteni-.

Chanopterus Boheman, 1858: 98. Current status: valid genus in PROMECHEILIDAE.

Type species: Chanopterus paradoxus Boheman, 1858, by monotypy.

Family-group name: CHANOPTERINAE Borchmann, 1915. Stem: Chanopter-.

Chapuisia Dugès, 1886: 56. Current status: invalid senior synonym of Schedlarius Wood, 1957 in CURCULIONIDAE: PLATYPODINAE: SCHEDLARIINI. Note: junior homonym of Chapuisia Duvivier, 1885 [Coleoptera: CHRYSOMELIDAE].

Type species: Chapuisia mexicana Dugès, 1886, by monotypy.

Family-group name: CHAPUISIIDES W.F.H. Blandford, 1895. Stem: Chapuisi-. Note: family- group name permanently invalid (ICZN 1999a, Article 39).

Charidotis Boheman, 1855: 7. Current status: valid genus in CHRYSOMELIDAE: CASSIDINAE: CASSIDINI.

Type species: Charidotis miniata Boheman, 1855, by subsequent designation (Hincks 1952: 343).

Family-group name: CHARIDOTITAE Spaeth, 1942. Stem: Charidotid-.

†Charonoscapha Ponomarenko, 1977: 32. Current status: valid genus in †COPTOCLAVIDAE: †CHARONOSCAPHINAE. Note: Upper Jurassic (Kazakhstan).

Type species: †Charonoscapha grossa Ponomarenko, 1977, by original designation.

Family-group name: †CHARONOSCAPHINAE Ponomarenko, 1977. Stem: Charonoscaph-.

Chasmatopterus Dejean, 1821: 60. Current status: valid genus in SCARABAEIDAE: MELOLONTHINAE: CHASMATOPTERINI.

Type species: Melolontha villosula Illiger, 1803, by subsequent designation (Blanchard in Audouin et al. 1843: pl. 44). Note: the type species usually listed for this genus is Melolontha hirtula Illiger, 1803 (e.g., Branco 2007: 9; Branco and A. Bezdĕk 2016: 211) by subsequent designation of Baraud (1965: 264) but Blanchard’s type fixation is earlier; both nominal species are currently placed in the genus Chasmatopterus.

Family-group name: CHASMATOPTÉRIDES Lacordaire, 1855. Stem: Chasmatopter-.

Chasme Lepeletier & Audinet-Serville, 1828: 378. Current status: valid genus in SCARABAEIDAE: MELOLONTHINAE: HOPLIINI: HOPLIINA.

Type species: Chasme decora Lepeletier & Audinet-Serville, 1828 (= Melolontha decora Wiedemann, 1823), by monotypy.

Family-group name: CHASMIDAE Streubel, 1846. Stem: Chasm-.

Chasmodia W.S. MacLeay, 1819: 155. Current status: junior synonym of Lagochile Hoffmannsegg, 1817 in SCARABAEIDAE: RUTELINAE: RUTELINI: RUTELINA.

Type species: Chasmodia viridis W.S. MacLeay, 1819 (= Rutela emarginata Gyllenhal, 1817), by subsequent designation (Duponchel 1843a: 405).

Family-group name: CHASMODIIDAE Burmeister, 1844. Stem: Chasmodi-.

Chauliognathus Hentz, 1830: 460. Current status: valid genus in CANTHARIDAE: CHAULIOGNATHINAE: CHAULIOGNATHINI.

Type species: Cantharis bimaculata Fabricius, 1781 (= Telephorus pensylvanicus DeGeer, 1774), by subsequent designation (Calder 1998: 186). Note: Calder (1998: 186) selected Telephorus pensylvanicus DeGeer, 1774 as type species, which is not an originally included species, but since he listed it in synonymy with Cantharis bimaculata Fabricius, 1881, an originally included species, he is deemed to have designated the latter species as type species (ICZN 1999a, Article 69.2.2); Fanti (2019: 29) remarked that Telephorus pensylvanicus DeGeer, 1774 is a junior synonym of Cantharis americana Forster, 1771, as noted already by some early authors, such as J.L. LeConte (1851b: 338); an application to the Commission is necessary to conserve usage of the well-known name Chauliognathus pensylvanicus (DeGeer, 1774).

Family-group name: CHAULIOGNATHINI J.L. LeConte, 1861. Stem: Chauliognath-.

Cheguevaria Kazantsev, 2007: 370. Current status: valid genus in LAMPYRIDAE: CHEGUEVARIINAE.

Type species: Cheguevaria taino Kazantsev, 2007, by original designation.

Family-group name: CHEGUEVARIINI Kazantsev, 2007. Stem: Cheguevari-.

Cheilomenes Chevrolat in Dejean, 1836a: 435. Current status: valid genus in COCCINELLIDAE: COCCINELLINAE: COCCINELLINI.

Type species: Coccinella lunata Fabricius, 1775, by subsequent designation (Crotch 1874a: 179).

Family-group name: CHEILOMENINI F.A. Schilder & M. Schilder, 1928. Stem: Cheilomen-.

Cheiridea Baly, 1878a: 253. Current status: valid genus in CHRYSOMELIDAE: EUMOLPINAE: TYPOPHORINI.

Type species: Cheiridea chapuisi Baly, 1878, by original designation.

Family-group name: CHEIRIDEITAE Lefèvre, 1885. Stem: Cheiride-.

Cheiroplatys Hope, 1837: 84. Current status: valid genus in SCARABAEIDAE: DYNASTINAE: PENTODONTINI: CHEIROPLATINA.

Type species: here fixed (under ICZN 1999a, Article 70.3.2) as Scarabaeus latipes Guérin- Méneville, 1838, misidentified as Scarabaeus truncatus Fabricius, 1775, in the original designation by Hope (1837: 84). Note: see Carne (1957b: 83) regarding the misidentification of the type species.

Family-group name: CHEIROPLATINA Carne, 1957. Stem: Cheiroplat-.

Chelonarium Fabricius, 1801a: xviii, 101. Current status: valid genus in CHELONARIIDAE.

Type species: Chelonarium atrum Fabricius, 1801, by subsequent designation (T. Bell 1830: 509).

Family-group name: CHELONARIITES Blanchard, 1845. Stem: Chelonari-.

Chennium Latreille, 1807: 77. Current status: valid genus in STAPHYLINIDAE: PSELAPHINAE: PSELAPHITAE: CTENISTINI.

Type species: Chennium bituberculatum Latreille, 1807, by monotypy.

Family-group name: CHENNIIDES Reitter, 1882. Stem: Chenni-.

Cherrocrius Berg, 1898: 31. Current status: valid genus in VESPERIDAE: ANOPLODERMATINAE: ANOPLODERMATINI.

Type species: Cherrocrius bruchi Berg, 1898, by original designation.

Family-group name: CHERROCRIINAE Prosen, 1960. Stem: Cherrocri-.

Chespirito V.S. Ferreira, Keller & Branham, 2020: 4. Current status: valid genus in LAMPYRIDAE: CHESPIRITOINAE.

Type species: Chespirito zaragozai V.S. Ferreira, Keller & Branham, 2020, by original designation.

Family-group name: CHESPIRITOINAE V.S. Ferreira, Keller & Branham, 2020: 4. Stem: Chespirito-.

Chevrolatia Jacquelin du Val, 1850: 45. Current status: valid genus in STAPHYLINIDAE: SCYDMAENINAE: SCYDMAENITAE: CHEVROLATIINI.

Type species: Chevrolatia insignis Jacquelin du Val, 1850, by monotypy.

Family-group name: CHEVROLATINI Reitter, 1882. Stem: Chevrolati-.

Chiasognathus Stephens, 1831a: 5. Current status: valid genus in LUCANIDAE: LUCANINAE: CHIASOGNATHINI.

Type species: Chiasognathus grantii Stephens, 1831, by original designation.

Family-group name: CHIASOGNATHIDAE Burmeister, 1847. Stem: Chiasognath-.

Chilapion Kissinger, 1968: 19. Current status: valid genus in BRENTIDAE: APIONINAE: CHILAPIITAE.

Type species: Apion macilentum Blanchard, 1851, by original designation.

Family-group name: CHILAPIITAE Wanat, 2001. Stem: Chilapi-.

Chilecar Kuschel, 1992: 205. Current status: valid genus in CARIDAE: CHILECARINAE: CHILECARINI.

Type species: Chilecar pilgerodendri Kuschel, 1992, by original designation.

Family-group name: CHILECARINI Legalov, 2009. Stem: Chilecar-.

Chilenius Lesne, 1921: 287. Current status: valid genus in BOSTRICHIDAE: PSOINAE: CHILENIINI.

Type species: Exopioides spinicollis Fairmaire & Germain, 1861, by original designation.

Family-group name: CHILENIIDAE Lesne, 1921. Stem: Chileni-.

Chilocorus Leach, 1815a: 116. Current status: valid genus in COCCINELLIDAE: COCCINELLINAE: CHILOCORINI.

Type species: Coccinella cacti Linnaeus, 1767, by monotypy.

Family-group name: CHILOCORIENS Mulsant, 1846. Stem: Chilocor-.

Chiloea Dajoz, 1973: 173. Current status: junior synonym of Sosteamorphus Hinton, 1936 in DRYOPIDAE.

Type species: Chiloea chilensis Dajoz, 1973 (= Sosteamorphus verrucatus Hinton, 1936), by original designation.

Family-group name: CHILOEIDAE Dajoz, 1973. Stem: Chiloe-.

Chiron W.S. MacLeay, 1819: 96, 107. Current status: valid genus in SCARABAEIDAE: CHIRONINAE.

Type species: Sinodendron digitatum Fabricius, 1801 (= Scarites cylindrus Fabricius, 1798), by monotypy.

Family-group name: CHIRIDITES Blanchard, 1845. Stem: Chiron-.

Chiroscelis Lamarck, 1804: 261. Current status: valid genus in TENEBRIONIDAE: LAGRIINAE: PYCNOCERINI.

Type species: Chiroscelis bifenestra Lamarck, 1804 (= Tenebrio digitatus Fabricius, 1801), by monotypy.

Family-group name: CHIROSCELIDAE Hope, 1841. Stem: Chiroscelid-.

Chlaenioctenus H.W. Bates, 1892: 309. Current status: valid genus in CARABIDAE: HARPALINAE: CHLAENIINI: CHLAENIINA.

Type species: Chlaenius pectinipes H.W. Bates, 1892, by subsequent designation (Basilewsky and Grundmann 1954: 243).

Family-group name: CHLAENIOCTENINI Basilewsky & Grundmann, 1955. Stem: Chlaeniocten-.

Chlaeniodus Jeannel, 1949b: 778. Current status: valid subgenus of Chlaenius Bonelli, 1810 in CARABIDAE: HARPALINAE: CHLAENIINI: CHLAENIINA.

Type species: Chlaenius perrieri Fairmaire, 1901 (= Chlaenius perrieranus Lorenz, 1998), by original designation.

Family-group name: CHLAENIODINI Jeannel, 1949. Stem: Chlaeniod-.

Chlaenionus Kuntzen, 1913: 33. Current status: valid subgenus of Chlaenius Bonelli, 1810 in CARABIDAE: HARPALINAE: CHLAENIINI: CHLAENIINA.

Type species: Rembus dohrnii Bertoloni, 1857, by subsequent designation (Basilewsky 1950: 44).

Family-group name: CHLAENIONINI Jeannel, 1949. Stem: Chlaenion-.

Chlaenius Bonelli, 1810: Tab. Syn. Current status: valid genus in CARABIDAE: HARPALINAE: CHLAENIINI: CHLAENIINA.

Type species [suggested]: Carabus festivus Panzer, 1796, by subsequent designation (Madge 1975: 581). Note: no species were originally included in this genus by Bonelli (1810); the first ones were the five species listed by Panzer (1813: 55, 56); the valid type species fixation is that of Hope (1838a: 75), who selected Carabus spoliatus Rossi, 1792, but that species is currently included in the subgenus Chlaenites Motschulsky, 1860, not in the nominotypical subgenus; an application to the Commission is necessary to fix Carabus festivus Panzer, 1796 as the type species of Chlaenius Bonelli, 1810.

Family-group name: CHLAENIDES Brullé, 1834. Stem: Chlaeni-.

Chlamisus Rafinesque, 1815: 116. Current status: valid genus in CHRYSOMELIDAE: CRYPTOCEPHALINAE: FULCIDACINI. Note: new replacement name for Chlamys Knoch, 1801.

Type species [suggested]: Chlamys foveolata Knoch, 1801, by subsequent designation (Navajas 1946: 244) (type fixation for Chlamys Knoch). Note: see “Note” in the entry for Chlamys Knoch, 1801 below.

Family-group name: CHLAMISINAE Gressitt, 1946. Stem: Chlamis-.

Chlamydopsis Westwood, 1869a: 317. Current status: valid genus in HISTERIDAE: CHLAMYDOPSINAE.

Type species: Chlamydopsis striatella Westwood, 1869, by subsequent designation (Lewis 1903: 428).

Family-group name: CHLAMYDOPSINI Bickhardt, 1914. Stem: Chlamydops-.

Chlamys Knoch, 1801: 122. Current status: invalid senior synonym of Chlamisus Rafinesque, 1815 in CHRYSOMELIDAE: CRYPTOCEPHALINAE: FULCIDACINI. Note: junior homonym of Chlamys Röding, 1798 [Mollusca].

Type species [suggested]: Chlamys foveolata Knoch, 1801, by subsequent designation (Navajas 1946: 244). Note: the valid type species fixation is that of S.H. Chen (1940: 191), who selected Chlamys tuberosa Knoch, 1801 (as “Bruchus gibbosus F. (= Chlamys tuberosa Kn.)”), one of the two originally included species (see ICZN 1999a, Article 69.2.2); however, this type species is a junior synonym of Bruchus gibbosus Fabricius, 1777 (now in Neochlamisus Karren, 1972; see C. Shin et al. 2012: 120); an application to the Commission is necessary to fix Chlamys foveolata Knoch, 1801 as the type species of Chlamys Knoch, 1801 in order to conserve the concept for this type genus and its new replacement name.

Family-group name: CHLAMYDEAE Lacordaire, 1848. Stem: Chlamyd-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

Chlidones C.O. Waterhouse, 1879a: 320. Current status: valid genus in CERAMBYCIDAE: CERAMBYCINAE: CHLIDONINI.

Type species: Chlidones lineolatus C.O. Waterhouse, 1879, by monotypy.

Family-group name: CHLIDONINAE C.O. Waterhouse, 1879. Stem: Chlidon-.

Chloemela Gistel, 1856: 379. Current status: invalid senior synonym of Chrysolina Motschulsky, 1860 (nomen protectum) in CHRYSOMELIDAE: CHRYSOMELINAE: CHRYSOMELINI. Note: nomen oblitum (see J. Bezděk 2020: 180).

Type species: Chrysomela staphylaea Linnaeus, 1758, by subsequent designation (J. Bezděk 2020: 180).

Family-group name: CHLOËMELADAE Gistel, 1856. Stem: Chloemel-.

Chnaunanthus Burmeister, 1844: 6, 31. Current status: valid genus in SCARABAEIDAE: MELOLONTHINAE: CHNAUNANTHINI.

Type species: Chnaunanthus discolor Burmeister, 1844, by monotypy.

Family-group name: CHNAUNANTHINI Evans & A.B.T. Smith, 2020: 63. Stem: Chnaunanth-.

Chnoodes Chevrolat, 1843a: 612. Current status: valid genus in COCCINELLIDAE: COCCINELLINAE: COCCIDULINI.

Type species: Chnoodes chaudoiri Mulsant, 1850, by subsequent designation (Gordon 2007: 1). Note: no species were originally included in this genus by Chevrolat (1843a); the first species included in the genus are the 13 species listed by Mulsant (1850: 908–915).

Family-group name: CHNOODIENS Mulsant, 1850. Stem: Chnood-.

Choeridium Lepeletier & Audinet-Serville, 1828: 356. Current status: junior synonym of Ateuchus Weber, 1801 in SCARABAEIDAE: SCARABAEINAE: ATEUCHINI: ATEUCHINA. Note: see Cupello et al. (2021: 7) for details regarding the original spelling of this genus-group name.

Type species: Choeridium simplex Lepeletier & Audinet-Serville, 1828, by subsequent designation (Chevrolat 1843a: 613).

Family-group name: CHOERIDIDAE Harold, 1867. Stem: Choeridi-.

Choerorhinus Fairmaire, 1858: 742. Current status: valid genus in CURCULIONIDAE: COSSONINAE: RHYNCOLINI: CHOERORHININA.

Type species: Choerorhinus squalidus Fairmaire, 1858, by monotypy.

Family-group name: CHOERORHINI Folwaczny, 1973. Stem: Choerorhin-.

Choleva Latreille, 1797: 14. Current status: valid genus in LEIODIDAE: CHOLEVINAE: CHOLEVINI: CHOLEVINA.

Type species [suggested]: Luperus cisteloides Frölich, 1799, by subsequent designation of Jeannel (1922: 43). Note: no species were originally included in Choleva; the valid type species fixation is that of Latreille (1802b: 122), who first associated Tritoma sericea Fabricius, 1787 (as “Catops sericea. F.”) with this genus-group name, but that species is the type species of Ptomaphagus Hellwig, 1795; an application to the Commission is needed to fix Luperus cisteloides Frölich, 1799 as the type species of Choleva Latreille, 1797.

Family-group name: CHOLEVIDAE W. Kirby, 1837. Stem: Cholev-.

Cholomus Roelofs, 1880: xl. Current status: valid genus in CURCULIONIDAE: MOLYTINAE: CHOLINI: CHOLOMINA.

Type species: Cholomus villei Roelofs, 1880, by monotypy.

Family-group name: CHOLOMINI Vaurie, 1974. Stem: Cholom-.

Cholus Germar, 1823: 212. Current status: valid genus in CURCULIONIDAE: MOLYTINAE: CHOLINI: CHOLINA. Note: name placed on the Official List of Generic Names in Zoology (ICZN 1987d, as Cholus Germar, 1824).

Type species: Cholus albicinctus Germar, 1823, by subsequent designation (Schönherr 1825: 584; see ICZN 1987d). Note: name placed on the Official List of Specific Names in Zoology (ICZN 1987d, as Cholus albicinctus Germar, 1824).

Family-group name: CHOLIDES Schönherr, 1825. Stem: Chol-.

Chonostropheus Prell, 1924: 162. Current status: valid genus in ATTELABIDAE: RHYNCHITINAE: RHYNCHITITAE: DEPORAINI.

Type species: Attelabus tristis Fabricius, 1794, by monotypy.

Family-group name: CHONOSTROPHEINA Morimoto, 1962. Stem: Chonostrophe-.

Choragus W. Kirby, 1819: 447. Current status: valid genus in ANTHRIBIDAE: CHORAGINAE: CHORAGINI. Note: name placed on the Official List of Generic Names in Zoology (ICZN 1994b).

Type species: Choragus sheppardi W. Kirby, 1819, by monotypy (see ICZN 1994b). Note: name placed on the Official List of Specific Names in Zoology (ICZN 1994b).

Family-group name: CHORAGIDAE W. Kirby, 1819. Stem: Chorag-.

Chorina Baly, 1866: 471. Current status: valid genus in CHRYSOMELIDAE: GALERUCINAE: GALERUCINI.

Type species: Monocesta cincta Clark, 1865, by original designation.

Family-group name: CHORINI Weise, 1923. Stem: Chorin-. Note: junior homonym of CHORININAE Dana, 1851 (type genus Chorinus Latreille, 1825) in Crustacea.

Chrestomera Jeannel, 1962a: 344. Current status: valid genus in STAPHYLINIDAE: PSELAPHINAE: EUPLECTITAE: TRICHONYCHINI: TRICHONYCHINA.

Type species: Chrestomera longicollis Jeannel, 1962, by original designation.

Family-group name: CHRESTOMERINA Jeannel, 1962. Stem: Chrestomer-.

Chromogeotrupes Bovo & Zunino, 1983: 404. Current status: valid subgenus of Phelotrupes Jekel, 1866 in GEOTRUPIDAE: GEOTRUPINAE: ENOPLOTRUPINI.

Type species: Geotrupes auratus Motschulsky, 1858, by original designation.

Family-group name: CHROMOGEOTRUPINI Zunino, 1984. Stem: Chromogeotrup-.

Chromoptilia Westwood, 1842b: 128. Current status: valid genus in SCARABAEIDAE: CETONIINAE: STENOTARSIINI: CHROMOPTILIINA.

Type species: Chromoptilia diversipes Westwood, 1842, by monotypy.

Family-group name: CHROMOPTILIINA Krikken, 1984. Stem: Chromoptili-.

Chrysobothris Eschscholtz, 1829b: 9. Current status: valid genus in BUPRESTIDAE: BUPRESTINAE: CHRYSOBOTHRINI. Note: the original spelling was Chrysobotris but Chrysobothris was ruled the correct spelling by the Commission and the name was placed on the Official List of Generic Names in Zoology (ICZN 1994f).

Type species: Buprestis chrysostigma Linnaeus, 1758, by subsequent designation (Westwood 1838c: 24; see ICZN 1994f). Note: name placed on the Official List of Specific Names in Zoology (ICZN 1994f).

Family-group name: CHRYSOBOTHRIDAE Gory & Laporte, 1836. Stem: Chrysobothr-.

Chrysochroa Dejean, 1833a: 75. Current status: valid genus in BUPRESTIDAE: CHRYSOCHROINAE: CHRYSOCHROINI: CHRYSOCHROINA. Note: this name is a nomen protectum and its senior synonym Euchloris Billberg, 1820 is a nomen oblitum (Bílý 2001: 149).

Type species: Buprestis fulminans Fabricius, 1787, by subsequent designation (Kurosawa 1982: 185).

Family-group name: CHRYSOCHROIDAE Laporte, 1835. Stem: Chrysochro-.

Chrysodema Laporte & Gory, 1837: 1. Current status: valid genus in BUPRESTIDAE: CHRYSOCHROINAE: CHRYSOCHROINI: CHALCOPHORINA. Note: name placed on the Official List of Generic Names in Zoology (ICZN 2004c, as Chrysodema Laporte & Gory, 1835).

Type species: Chrysodema sonnerati Laporte & Gory, 1837, by plenary power (see ICZN 2004c). Note: name placed on the Official List of Specific Names in Zoology (ICZN 2004c, as Chrysodema sonnerati Laporte & Gory, 1835).

Family-group name: CHRYSODÉMIDES Kerremans, 1892. Stem: Chrysodem-.

Chrysodina Baly, 1864b: 221. Current status: junior synonym of Spintherophyta Dejean, 1836 in CHRYSOMELIDAE: EUMOLPINAE: EUMOLPINI.

Type species: Chrysodina igneicollis Baly, 1864, by monotypy.

Family-group name: CHRYSODINITAE Lefèvre, 1885. Stem: Chrysodin-.

Chrysolina Motschulsky, 1860b: 210. Current status: valid genus in CHRYSOMELIDAE: CHRYSOMELINAE: CHRYSOMELINI. Note: given precedence over Atechna Chevrolat, 1837 whenever the two names are considered synonyms and name placed on the Official List of Generic Names in Zoology (ICZN 1984c); nomen protectum (see J. Bezdĕk 2020: 180).

Type species: Chrysomela staphylaea Linnaeus, 1758, by original designation (see ICZN 1984c). Note: name placed on the Official List of Specific Names in Zoology (ICZN 1984c).

Family-group name: CHRYSOLININA S.H. Chen, 1936. Stem: Chrysolin-.

Chrysomela Linnaeus, 1758: 342, 368. Current status: valid genus in CHRYSOMELIDAE: CHRYSOMELINAE: CHRYSOMELINI. Note: name placed on the Official List of Generic Names in Zoology (ICZN 1984c).

Type species: Chrysomela populi Linnaeus, 1758, by subsequent designation (Latreille 1810: 432; see ICZN 1984c). Note: name placed on the Official List of Specific Names in Zoology (ICZN 1984c).

Family-group name: CHRYSOMELINAE Latreille, 1802. Stem: Chrysomel-.

Chrysophora Dejean, 1821: 60. Current status: valid genus in SCARABAEIDAE: RUTELINAE: RUTELINI: RUTELINA.

Type species: Melolontha chrysochlora Latreille, 1809, by monotypy.

Family-group name: CHRYSOPHORIDAE Burmeister, 1844. Stem: Chrysophor-.

Ciceronius Kaup, 1871: 29. Current status: valid genus in PASSALIDAE: PASSALINAE: SOLENOCYCLINI.

Type species: Passalus morbillosus Klug, 1833, by monotypy.

Family-group name: CICERONINAE Kuwert, 1891. Stem: Ciceroni-.

Cicindela Linnaeus, 1758: 342, 407. Current status: valid genus in CICINDELIDAE: CICINDELINI: CICINDELINA.

Type species: Cicindela campestris Linnaeus, 1758, by subsequent designation (Latreille 1810: 425).

Family-group name: CICINDELETAE Latreille, 1802. Stem: Cicindel-.

Cicindis Bruch, 1908: 497. Current status: valid genus in CARABIDAE: CICINDINAE.

Type species: Cicindis horni Bruch, 1908, by monotypy.

Family-group name: CICINDINI Csiki, 1927. Stem: Cicind-.

Cillaeus Laporte, 1834c: 133. Current status: valid genus in NITIDULIDAE: CILLAEINAE: CILLAEINI.

Type species: Cillaeus castaneus Laporte, 1834, by subsequent designation (E. Desmarest 1851: 290).

Family-group name: CILLAEINAE Kirejtshuk & Audisio, 1986. Stem: Cillae-.

Cimberis Gozis, 1881: 150. Current status: valid genus in CIMBERIDIDAE: CIMBERIDINI. Note: name placed on the Official List of Generic Names in Zoology (ICZN 2005c).

Type species: Rhinomacer attelaboides Fabricius, 1787, by subsequent designation (Kuschel 1959: 234; see ICZN 2005c). Note: name placed on the Official List of Specific Names in Zoology (ICZN 2005c).

Family-group name: CIMBERIDAE Gozis, 1882. Stem: Cimberid-.

Cinnabarium Fairmaire, 1894: ccvi. Current status: junior synonym of Coptocera Murray, 1868 in DASCILLIDAE: DASCILLINAE: CINNABARIINI.

Type species: Cinnabarium seminigrum Fairmaire, 1894, by monotypy.

Family-group name: CINNABARIINI Pic, 1914. Stem: Cinnabari-.

Cinyra Laporte & Gory, 1837: 4. Current status: valid genus in BUPRESTIDAE: CHRYSOCHROINAE: PARALEPTODEMINI: PARALEPTODEMINA.

Type species: Buprestis elongata Laporte & Gory, 1837, by subsequent designation (Casey 1909: 53).

Family-group name: CINYRINI Cobos, 1979. Stem: Cinyr-.

Cionus Clairville, 1798: 56, 64. Current status: valid genus in CURCULIONIDAE: CURCULIONINAE: CIONINI.

Type species: Curculio blattariae Fabricius, 1792 (= Curculio alauda Herbst, 1784), by subsequent designation (Crotch 1870a: 50).

Family-group name: CIONIDES Schönherr, 1825. Stem: Cion-.

†Circaeus Iablokoff-Khnzorian, 1961: 209. Current status: valid genus in ADERIDAE. Note: Lower Tertiary (Baltic amber).

Type species: †Circaeus borisjaki Iablokoff-Khnzorian, 1961, by original designation.

Family-group name: †CIRCAEIDAE Iablokoff-Khnzorian, 1961. Stem: Circae-.

Cis Latreille, 1797: 50. Current status: valid genus in CIIDAE: CIINAE: CIINI.

Type species: Dermestes boleti Scopoli, 1763, by subsequent monotypy (Latreille 1802b: 205).

Family-group name: CISIDAE Leach, 1819. Stem: Ci-.

Cisanthribus Zimmerman, 1938: 247. Current status: valid genus in ANTHRIBIDAE: CHORAGINAE: CISANTHRIBINI.

Type species: Cisanthribus convexus Zimmerman, 1938, by original designation.

Family-group name: CISANTHRIBINI Zimmerman, 1994. Stem: Cisanthrib-.

Cisseis Gory & Laporte, 1839: 1. Current status: junior synonym of Diphucrania Dejean, 1833 in BUPRESTIDAE: AGRILINAE: CORAEBINI: CISSEINA.

Type species: Buprestis duodecimguttata Guérin-Méneville, 1838 (= Buprestis duodecimmaculata Fabricius, 1801), by subsequent designation (Duponchel 1843a: 719).

Family-group name: CISSEINA Majer, 2000. Stem: Cisse-.

Cissites Latreille, 1804a: 154, 199. Current status: valid genus in MELOIDAE: NEMOGNATHINAE: HORIINI.

Type species: Cucujus maculatus Swederus, 1787, by monotypy.

Family-group name: CISSITINI Kaszab, 1966. Stem: Cissit-.

Cistela Fabricius, 1775: 116. Current status: invalid senior synonym of Cteniopus Solier, 1835 in TENEBRIONIDAE: ALLECULINAE: CTENIOPODINI. Note: junior homonym of Cistela Geoffroy, 1762 [Coleoptera: BYRRHIDAE], which was suppressed for the Principle of Priority but not for the Principle of Homonymy (ICZN 1994a).

Type species: Chrysomela sulphurea Linnaeus, 1758, by subsequent designation (Latreille 1810: 429).

Family-group name: CISTELENIAE Latreille, 1802. Stem: Cistel-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

Cladodes Solier, 1849: 444. Current status: valid genus in LAMPYRIDAE: CLADODINAE.

Type species: Cladodes flabellatus Solier, 1849, by monotypy.

Family-group name: CLADODINAE Bocáková, Campello-Gonçalves & Silveira, 2022: 1186. Stem: Cladod-. Note: this name could be a junior homonym of CLADODIDAE (type genus Cladodus Egerton, 1841) in Pisces, although more research is needed to confirm this; if a family-group name based on Cladodus is indeed available in Pisces then the case will need to be referred to the Commission for a ruling to remove the homonymy ICZN (1999a, Article 55.3.1).

Cladognathus Burmeister, 1847: 364. Current status: junior synonym of Prosopocoilus Hope, 1845 in LUCANIDAE: LUCANINAE: DORCINI. Note: new replacement name for Macrognathus Hope, 1845.

Type species: Lucanus girafa G.-A. Olivier, 1789, by subsequent designation (Lacordaire 1855: 29) (type fixation for Macrognathus Hope).

Family-group name: CLADOGNATHIDAE Parry, 1870. Stem: Cladognath-.

Cladophorus Guérin-Méneville, 1830: pl. 2. Current status: valid genus in LYCIDAE: METRIORRHYNCHINAE: METRIORRHYNCHINI: METRIORRHYNCHINA. Note: this genus-group name was first made available by Guérin-Méneville (1829a: pl. 14) when he illustrated the new species Cladophorus ruficollis; however, that species is currently placed in the valid genus Pseudodontocerus Pic, 1921; in order to preserve nomenclatural stability, we recommend that an application be submitted to the Commission to suppress the name Cladophorus Guérin-Méneville, 1829 for both the Principles of Homonymy and Priority.

Type species: Cladophorus dimidiatus Guérin-Méneville, 1830, by subsequent designation (Bocák 1998: 247). Note: Bocák (1998: 247) credited the type species designation to C.O. Waterhouse (1878: 103); however, C.O. Waterhouse listed Cladophorus formosus Guérin-Méneville, 1838, which is not a species originally included in Cladophorus, as type species of this genus.

Family-group name: CLADOPHORINAE Kleine, 1929. Stem: Cladophor-.

Cladotoma Westwood, 1836: 254. Current status: valid genus in PTILODACTYLIDAE: CLADOTOMINAE.

Type species: Cladotoma ovalis Westwood, 1836, by monotypy.

Family-group name: CLADOTOMINI Pic, 1914. Stem: Cladotom-.

Cladoxena Motschulsky, 1866a: 428. Current status: valid genus in EROTYLIDAE: LANGURIINAE: LANGURIINI.

Type species: Cladoxena rufipes Motschulsky, 1866, by subsequent designation (Crotch 1876: 396).

Family-group name: CLADOXENINAE Arrow, 1925. Stem: Cladoxen-.

Clambus Fischer, 1821b: 52. Current status: valid genus in CLAMBIDAE: CLAMBINAE.

Type species: Dermestes armadillo DeGeer, 1774 [as armadillus], by subsequent designation (Westwood 1838c: 10).

Family-group name: CLAMBINI Fischer, 1821. Stem: Clamb-.

Clanis Mulsant, 1850: 999. Current status: invalid senior synonym of Jauravia Motschulsky, 1858 in COCCINELLIDAE: COCCINELLINAE: STICHOLOTIDINI. Note: junior homonym of Clanis Hübner, 1819 [Lepidoptera].

Type species: Coccinella pubescens Fabricius, 1798, by monotypy.

Family-group name: CLANINI Weise, 1901. Stem: Clan-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

Claviger Preyssler, 1790: 68. Current status: valid genus in STAPHYLINIDAE: PSELAPHINAE: CLAVIGERITAE: CLAVIGERINI.

Type species: Claviger testaceus Preyssler, 1790, by monotypy.

Family-group name: CLAVIGERIDES Leach, 1815. Stem: Claviger-. Note: senior homonym of CLAVIGERINAE J.B. Waterhouse, 1975 (type genus Clavigera J.A. Thomson, 1913) in Brachiopoda.

Clavigerodes Raffray, 1877: 279. Current status: valid genus in STAPHYLINIDAE: PSELAPHINAE: CLAVIGERITAE: CLAVIGERINI.

Type species: Clavigerodes abyssinicus Raffray, 1877, by monotypy.

Family-group name: CLAVIGERODINI L.W. Schaufuss, 1882. Stem: Clavigerod-.

Clavigeropsis Raffray, 1882: 3. Current status: valid genus in STAPHYLINIDAE: PSELAPHINAE: CLAVIGERITAE: CLAVIGERINI.

Type species: Clavigeropsis formicarius Raffray, 1882, by monotypy.

Family-group name: CLAVIGEROPSINI L.W. Schaufuss, 1890. Stem: Clavigeropse-.

Clavilispinus Bernhauer, 1926: 255. Current status: valid genus in STAPHYLINIDAE: OSORIINAE: THORACOPHORINI: CLAVILISPININA.

Type species: Paralispinus siargaoanus Bernhauer, 1926, by monotypy.

Family-group name: CLAVILISPININA Newton & Thayer, 1992. Stem: Clavilispin-.

Clavipalpus Laporte, 1833b: 406. Current status: valid genus in SCARABAEIDAE: MELOLONTHINAE: MACRODACTYLINI.

Type species: Clavipalpus dejeanii Laporte, 1833, by monotypy.

Family-group name: CLAVIPALPIDES Lacordaire, 1855. Stem: Clavipalp-.

Cleidecosta P.J. Johnson, 2002: 16. Current status: valid genus in ELATERIDAE: AGRYPNINAE: EUPLINTHINI: CLEIDECOSTINA. Note: new replacement name for Heligmus Candèze, 1865.

Type species: Heligmus glyphoderus Candèze, 1865, by monotypy (type fixation for Heligmus Candèze).

Family-group name: CLEIDECOSTINI P.J. Johnson, 2002. Stem: Cleidecost-.

Cleidostethus Arrow, 1929a: 464. Current status: valid genus in CORYLOPHIDAE: CORYLOPHINAE: CLEIDOSTETHINI.

Type species: Cleidostethus meliponae Arrow, 1929, by monotypy.

Family-group name: CLEIDOSTETHINI Bowestead, Booth, Ślipiński & Lawrence, 2001. Stem: Cleidosteth-.

Clema Semenov, 1900: 590. Current status: valid genus in BUPRESTIDAE: AGRILINAE: CORAEBINI: CLEMINA.

Type species: Clema deserti Semenov, 1900, by original designation.

Family-group name: CLEMINA Majer, 2000. Stem: Clem- (ICZN 1999a, Article 29.4).

Cleogonus Schönherr, 1825: 586. Current status: valid genus in CURCULIONIDAE: MOLYTINAE: CLEOGONINI.

Type species: Curculio rubetra Fabricius, 1787, by original designation.

Family-group name: CLEOGONIDAE Gistel, 1848. Stem: Cleogon-.

Cleomenes J. Thomson, 1864: 161. Current status: valid genus in CERAMBYCIDAE: CERAMBYCINAE: CLEOMENINI.

Type species: Cleomenes dihammaphoroides J. Thomson, 1864, by original designation.

Family-group name: CLÉOMÉNIDES Lacordaire, 1868. Stem: Cleomen-.

Cleonis Dejean, 1821: 96. Current status: valid genus in CURCULIONIDAE: LIXINAE: CLEONINI.

Type species: Curculio sulcirostris Linnaeus, 1767 (= Curculio piger Scopoli, 1763), by subsequent designation (Chevrolat 1843b: 6).

Family-group name: CLEONIDAE W. Kirby, 1837. Stem: Cleon-. Note: junior homonym of CLEONIDES Schönherr, 1826 (type genus Cleonus Schönherr, 1826) in Coleoptera: CURCULIONIDAE.

Cleonus Schönherr, 1826: 145. Current status: junior synonym of Cleonis Dejean, 1821 in CURCULIONIDAE: LIXINAE: CLEONINI. Note: unnecessary new replacement name for Geomorus Schönherr, 1823.

Type species: Curculio sulcirostris Linnaeus, 1767 (= Curculio piger Scopoli, 1763), by original designation (type fixation for Geomorus Schönherr).

Family-group name: CLEONIDES Schönherr, 1826. Stem: Cleon-. Note: nomen protectum (see Bouchard et al. 2011: 620); senior homonym of CLEONIDAE W. Kirby, 1837 (type genus Cleonis Dejean, 1821) in Coleoptera: CURCULIONIDAE.

Cleopterium Gistel, 1856: 360. Current status: junior synonym of Acrotrichis Motschulsky, 1848 in PTILIIDAE: PTILIINAE: ACROTRICHINI.

Type species: Latridius fascicularis Herbst, 1793, by subsequent designation (Biström and Silfverberg 1979: 13).

Family-group name: CLEOPTERIIDAE Gistel, 1856. Stem: Cleopteri-.

Cleropiestus Fairmaire, 1889: xcii. Current status: valid genus in CLERIDAE: CLERINAE: OPILONITAE: OPILONINI: CLEROPIESTINA.

Type species: Cleropiestus oberthurii Fairmaire, 1890, by monotypy.

Family-group name: CLEROPIESTINAE J.R. Winkler, 1980. Stem: Cleropiest-.

Clerus Geoffroy, 1762: 303. Current status: valid genus in CLERIDAE: CLERINAE: CLERITAE: CLERINI. Note: name attributed to Geoffroy, 1762 and placed on the Official List of Generic Names in Zoology (ICZN 1984a).

Type species: Clerus mutillarius Fabricius, 1775, by plenary power (see ICZN 1984a). Note: name placed on the Official List of Specific Names in Zoology (ICZN 1984a).

Family-group name: CLERII Latreille, 1802. Stem: Cler-.

†Clessidromma Jarzembowski, B. Wang & Zheng, 2018: 112. Current status: valid genus in OMMATIDAE incertae sedis. Note: mid-Cretaceous (Myanmar).

Type species: †Clessidromma palmeri Jarzembowski, B. Wang & Zheng, 2018, by original designation.

Family-group name: †CLESSIDROMMATINI Jarzembowski, B. Wang & Zheng, 2018: 112. Stem: Clessidrommat-.

Clidicus Laporte, 1833b: 396. Current status: valid genus in STAPHYLINIDAE: SCYDMAENINAE: MASTIGITAE: CLIDICINI.

Type species: Clidicus grandis Laporte, 1833, by monotypy.

Family-group name: CLIDICINI Casey, 1897. Stem: Clidic-.

Clinidium W. Kirby, 1830: 6. Current status: valid genus in CARABIDAE: RHYSODINAE: CLINIDIINI.

Type species: Clinidium guildingii W. Kirby, 1830, by monotypy.

Family-group name: CLINIDIINA R.T. Bell & J.R. Bell, 1978. Stem: Clinidi-.

Clinolabus Jekel, 1860: 196. Current status: valid genus in ATTELABIDAE: ATTELABINAE: ATTELABINI: EUSCELINA.

Type species: Attelabus angulatus Fabricius, 1787, by original designation. Note: Jekel (1860) designated two type species separately in his work, one (Attelabus angulatus Fabricius, 1787) on page 197, the other (Attelabus melancoryphus Germar, 1823) in the table [unpaginated page]; Alonso-Zarazaga and Lyal (1999: 45) acted as First Revisers (ICZN 1999a, Article 24.2.2) for the type species selection.

Family-group name: CLINOLABINA Legalov, 2003. Stem: Clinolab-.

Clinteria Burmeister, 1842: 299. Current status: valid genus in SCARABAEIDAE: CETONIINAE: GYMNETINI: GYMNETINA.

Type species: Clinteria guttifera Burmeister, 1842 [by 6 December 1842] (= Gymnetis auronotata Blanchard, 1842 [November 1842]), by subsequent designation (Duponchel 1843b: 10).

Family-group name: CLINTERIIDAE Kraatz, 1882. Stem: Clinteri-.

Clitostylus Voss, 1929b: 193. Current status: junior synonym of Trachelismus Motschulsky, 1870 in ATTELABIDAE: APODERINAE: CLITOSTYLINI: CLITOSTYLINA.

Type species: Apoderus badeni Faust, 1883 (= Apoderus macrostylus Motschulsky, 1861), by subsequent designation (Legalov 2003: 476). Note: Legalov (2003: 476) selected Apoderus macrostylus Motschulsky, 1861 as type species of Clitostylus Voss, 1929, which is not an originally included species, but since he listed it in synonymy with Apoderus badeni Faust, 1883, an originally included species, he is deemed to have designated the latter species as type species (ICZN 1999a, Article 69.2.2).

Family-group name: CLITOSTYLINI Voss, 1929. Stem: Clitostyl-.

Clivina Latreille, 1802b: 96. Current status: valid genus in CARABIDAE: SCARITINAE: CLIVININI: CLIVININA.

Type species: Scarites arenarius Fabricius, 1775 (= Tenebrio fossor Linnaeus, 1758), by monotypy.

Family-group name: CLIVINIDIA Rafinesque, 1815. Stem: Clivin-.

Cloniocerus Dejean, 1835: 340. Current status: valid genus in CERAMBYCIDAE: LAMIINAE: CLONIOCERINI.

Type species: Lamia hystrix Fabricius, 1781, by monotypy.

Family-group name: CLONIOCÉRIDES Lacordaire, 1872. Stem: Cloniocer-.

Closterus Audinet-Serville, 1832: 193. Current status: valid genus in CERAMBYCIDAE: PRIONINAE: MEROSCELISINI.

Type species: Closterus flabellicornis Audinet-Serville, 1832, by monotypy.

Family-group name: CLOSTÉRIDES Lacordaire, 1868. Stem: Closter-.

Clypeaster Dejean, 1821: 129. Current status: invalid senior synonym of Clypastraea Haldeman, 1842 in CORYLOPHIDAE: CORYLOPHINAE: PARMULINI. Note: junior homonym of Clypeaster Lamarck, 1801 [Echinodermata].

Type species: Cossyphus pusillus Gyllenhal, 1810, by monotypy.

Family-group name: CLYPEASTRINA Perty, 1840. Stem: Clypeaster-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

Clytra Laicharting, 1781: 165. Current status: valid genus in CHRYSOMELIDAE: CRYPTOCEPHALINAE: CLYTRINI: CLYTRINA.

Type species: Chrysomela quadripunctata Linnaeus, 1758, by subsequent designation (Latreille 1810: 432).

Family-group name: CLYTHRIDAE W. Kirby, 1837. Stem: Clytr-.

Clytus Laicharting, 1784: 88. Current status: valid genus in CERAMBYCIDAE: CERAMBYCINAE: CLYTINI.

Type species: Leptura arietis Linnaeus, 1758, by subsequent designation (Curtis 1828: pl. 199).

Family-group name: CLYTAIRES Mulsant, 1839. Stem: Clyt-.

Cnemalobus Guérin-Méneville, 1839a: pl. 226 (p. 9). Current status: valid genus in CARABIDAE: HARPALINAE: CNEMALOBINI.

Type species: Cnemalobus desmarestii Guérin-Méneville, 1839, by subsequent designation (Roig-Juñent 1993: 10).

Family-group name: CNEMALOBINI Germain, 1911. Stem: Cnemalob-.

Cnemeplatia A. Costa, 1847: 146. Current status: valid genus in TENEBRIONIDAE: PIMELIINAE: CNEMEPLATIINI: CNEMEPLATIINA.

Type species: Cnemeplatia atropos A. Costa, 1847, by monotypy.

Family-group name: CNÉMÉPLATIITES Jacquelin du Val, 1861. Stem: Cnemeplati-.

Cnemidontus Schenkling, 1935: 29. Current status: valid genus in CURCULIONIDAE: MESOPTILIINAE: MESOPTILIINI. Note: new replacement name for Cnemidophorus Schönherr, 1835.

Type species: Cnemidophorus fasciculatus Boheman, 1835, by original designation (type fixation for Cnemidophorus Schönherr).

Family-group name: CNEMIDONTINI Kuschel, 1955. Stem: Cnemidont-.

Cnemidophorus Schönherr, 1835: 276. Current status: invalid senior synonym of Cnemidontus Schenkling, 1935 in CURCULIONIDAE: MESOPTILIINAE: MESOPTILIINI. Note: junior homonym of Cnemidophorus Wagler, 1830 [Reptilia].

Type species: Cnemidophorus fasciculatus Boheman, 1835, by original designation.

Family-group name: CNEMIDOPHORINI Hustache, 1937. Stem: Cnemidophor-. Note: family- group name permanently invalid (ICZN 1999a, Article 39).

Cnemodinus Cockerell, 1906: 242. Current status: valid genus in TENEBRIONIDAE: PIMELIINAE: CNEMODININI. Note: new replacement name for Cnemodus G.H. Horn, 1870.

Type species: Cnemodus testaceus G.H. Horn, 1870, by monotypy (type fixation for Cnemodus G.H. Horn).

Family-group name: CNEMODININAE Gebien, 1910. Stem: Cnemodin-.

Cnemodus G.H. Horn, 1870: 266. Current status: invalid senior synonym of Cnemodinus Cockerell, 1906 in TENEBRIONIDAE: PIMELIINAE: CNEMODININI. Note: junior homonym of Cnemodus Herrich- Schäffer, 1850 [Hemiptera].

Type species: Cnemodus testaceus G.H. Horn, 1870, by monotypy.

Family-group name: CNEMODINI G.H. Horn, 1870. Stem: Cnemod-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

Cnemogonus J.L. LeConte in J.L. LeConte and G.H. Horn, 1876: 269. Current status: valid genus in CURCULIONIDAE: CEUTORHYNCHINAE: CNEMOGONINI.

Type species: here fixed (under ICZN 1999a, Article 70.3.2) as Cnemogonus lecontei Dietz, 1896, misidentified as Curculio epilobii Paykull, 1800 (sensu J.L. LeConte, 1876) in the designation by monotypy by J.L. LeConte in J.L. LeConte and G.H. Horn (1876: 269).

Family-group name: CNEMOGONINI Colonnelli, 1979. Stem: Cnemogon-.

Cnemoplites Newman, 1842b: 351. Current status: valid genus in CERAMBYCIDAE: PRIONINAE: MACROTOMINI: MACROTOMINA.

Type species: Cnemoplites edulis Newman, 1842, by subsequent designation (J. Thomson 1864: 301).

Family-group name: CNEMOPLITINAE Schröder, 1905. Stem: Cnemoplit-.

Cneoglossa Guérin-Méneville, 1843a: 194. Current status: valid genus in CNEOGLOSSIDAE.

Type species: Cneoglossa collaris Guérin-Méneville, 1849, by subsequent monotypy (Guérin- Méneville 1849: [no 36] 1).

Family-group name: CNEOGLOSSINI Champion, 1897. Stem: Cneogloss-.

Cneorhinus Schönherr, 1823: 1139. Current status: valid genus in CURCULIONIDAE: ENTIMINAE: POLYDRUSITAE: CNEORHININI.

Type species: Curculio barcelonicus Herbst, 1797, by original designation.

Family-group name: CNÉORHINIDES Lacordaire, 1863. Stem: Cneorhin-.

Cnides Motschulsky, 1862: 38. Current status: valid genus in CARABIDAE: TRECHINAE: TRECHINI: TRECHODINA.

Type species: Cnides rostratus Motschulsky, 1862, by monotypy.

Family-group name: CNIDINA Jeannel, 1958. Stem: Cnid-.

Cnodalon Latreille, 1797: 23. Current status: valid genus in TENEBRIONIDAE: STENOCHIINAE: CNODALONINI. Note: name placed on the Official List of Generic Names in Zoology (ICZN 2023).

Type species: Cnodalon viride Latreille, 1804, by plenary power (see ICZN 2023). Note: no species were originally included in this genus by Latreille (1797); Bousquet and Bouchard (2019: 114, 116) submitted a request to the Commission to retain Cnodalon viride Latreille, 1804, the currently accepted type species of the genus, after the discovery that this was not an originally included species; name placed on the Official List of Specific Names in Zoology (ICZN 2023).

Family-group name: NODALIDEN Oken, 1843. Stem: Cnodalon-.

Cnopus Champion, 1893: 460. Current status: valid genus in ADERIDAE.

Type species: Cnopus flohri Champion, 1893, by monotypy.

Family-group name: CNOPINA Báguena Corella, 1948. Stem: Cnop-.

Coccidula Kugelann in Illiger, 1798: 421. Current status: valid genus in COCCINELLIDAE: COCCINELLINAE: COCCIDULINI.

Type species: Dermestes rufus Herbst, 1783, by subsequent designation (C.G. Thomson 1859: 161). Note: Chrysomela scutellata Herbst, 1783 is usually listed as the type species of this genus (e.g., Kovář 2007: 572; Szawaryn et al. 2021: 64) by subsequent designation of Crotch (1874a: 300) but C.G. Thomson’s designation is earlier; both species are currently included in the genus Coccidula.

Family-group name: COCCIDULIENS Mulsant, 1846. Stem: Coccidul-.

Coccinella Linnaeus, 1758: 342, 364. Current status: valid genus in COCCINELLIDAE: COCCINELLINAE: COCCINELLINI.

Type species: Coccinella septempunctata Linnaeus, 1758, by subsequent designation (Latreille 1810: 432).

Family-group name: COCCINELLIDAE Latreille, 1807. Stem: Coccinell-.

Coelaenomenodera Blanchard, 1845a: 181. Current status: valid genus in CHRYSOMELIDAE: CASSIDINAE: COELAENOMENODERINI.

Type species: Acentroptera cucullata Guérin-Méneville, 1844, by subsequent designation (Baly 1858: 119). Note: no species were originally included in this genus by Blanchard (1845a); the first ones were the four species listed by Baly (1858: 119, 120).

Family-group name: COELAENOMENODERINI Weise, 1911. Stem: Coelaenomenoder-.

Coeliodes Schönherr, 1837: 282. Current status: valid genus in CURCULIONIDAE: CEUTORHYNCHINAE: CEUTORHYNCHINI. Note: name placed on the Official List of Generic Names in Zoology (ICZN 1989b).

Type species: Curculio quercus Linnaeus, 1758 (sensu Fabricius, 1787) (= Curculio rana Fabricius, 1787), by original designation (see ICZN 1989b). Note: the Commission (ICZN 1989b) ruled that the type species of Coeliodes Schönherr, 1837 was “Curculio quercus Fabricius, 1787 (a junior primary homonym replaced by Curculio dryados Gmelin, 1790)” and placed Curculio dryados Gmelin, 1790 on the Official List of Specific Names in Zoology; however, Curculio quercus Fabricius, 1787 is not available as Fabricius misidentified Curculio quercus Linnaeus, 1758 (currently Orchestes quercus (Linnaeus, 1758)); the taxon Curculio quercus Linnaeus, 1758 (sensu Fabricius, 1787) is currently known under the valid name Curculio rana Fabricius, 1787 (see Alonso-Zarazaga et al. 2017: 141); an application to the Commission is necessary to change the entry for Coeliodes Schönherr, 1837 and its type species on the Official Lists.

Family-group name: COELIODES J.L. LeConte, 1876. Stem: Coeliod-.

Coelomera Chevrolat in Dejean, 1836a: 375. Current status: valid genus in CHRYSOMELIDAE: GALERUCINAE: GALERUCINI.

Type species: Chrysomela cajennensis Fabricius, 1787, by subsequent designation (Weise 1924: 51).

Family-group name: COELOMÉRITES Chapuis, 1875. Stem: Coelomer-.

Coelometopon E. Janssens, 1972: 386. Current status: valid genus in HYDRAENIDAE: PROSTHETOPINAE: COELOMETOPONINI.

Type species: Coelometopon leleupi E. Janssens, 1972, by original designation.

Family-group name: COELOMETOPONINI Perkins, 2005. Stem: Coelometopon-.

Coelometopus Solier, 1848b: 154, 278. Current status: valid genus in TENEBRIONIDAE: STENOCHIINAE: CNODALONINI.

Type species: Coelometopus clypeatus Solier, 1848 (= Blaps clypeata Germar, 1813), by monotypy.

Family-group name: COELOMETOPIDAE Schaum, 1859. Stem: Coelometop-.

Coelonertus A. Solari & F. Solari, 1906: 441. Current status: valid genus in CURCULIONIDAE: BARIDINAE: APOSTASIMERINI: COELONERTINA.

Type species: Coelonertus squamulosus A. Solari & F. Solari, 1906, by subsequent designation (Champion 1908: 335).

Family-group name: COELONERTINI Casey, 1922. Stem: Coelonert-.

Coelophora Mulsant, 1850: 390. Current status: valid genus in COCCINELLIDAE: COCCINELLINAE: COCCINELLINI.

Type species: Coccinella inaequalis Fabricius, 1775, by subsequent designation (Crotch 1874a: 148).

Family-group name: COELOPHORAIRES Mulsant, 1850. Stem: Coelophor-.

Coelopterus Mulsant & Rey, 1853c: 89. Current status: valid genus in COCCINELLIDAE: COCCINELLINAE: STICHOLOTIDINI. Note: this genus is often attributed to Mulsant & Rey, 1852 (p. 224); however, that version of the article (in the ‘Mémoires de l’Académie Nationale des Sciences…’) was evidently not published until after the version in the ‘Opuscules Entomologiques’, in 1853 (see Reference 1853c cited above).

Type species: Coelopterus salinus Mulsant & Rey, 1853, by monotypy.

Family-group name: COELOPTERINI Weise, 1906. Stem: Coelopter-.

Coelorrhina Hope, 1841b: 302. Current status: valid genus in SCARABAEIDAE: CETONIINAE: GOLIATHINI: RHOMBORHININA. Note: the original spelling was Caelorrhina but Coelorrhina, an incorrect subsequent spelling, is in prevailing usage, attributed to the publication of the original spelling, and therefore deemed to be the correct original spelling (ICZN 1999a, Article 33.3.1).

Type species: Coelorrhina concolor Hope, 1841, by monotypy.

Family-group name: COELORRHINAE Schoch, 1895. Stem: Coelorrhin-.

Coelostoma Brullé, 1835: 293. Current status: valid genus in HYDROPHILIDAE: SPHAERIDIINAE: COELOSTOMATINI.

Type species: Hydrophilus orbicularis Fabricius, 1775, by monotypy.

Family-group name: COELOSTOMITAE Heyden, 1891. Stem: Coelostomat-.

Coelus Eschscholtz, 1829d: 5. Current status: valid genus in TENEBRIONIDAE: PIMELIINAE: CONIONTINI.

Type species: Coelus ciliatus Eschscholtz, 1829, by monotypy.

Family-group name: COELINI Casey, 1907. Stem: Coel-.

Coenochilus Schaum in Burmeister and Schaum, 1841: 250, 268. Current status: valid genus in SCARABAEIDAE: CETONIINAE: CREMASTOCHEILINI: COENOCHILINA.

Type species: Cremastocheilus paulus Gory & Percheron, 1833 (= Cetonia maura Fabricius, 1801), by subsequent designation (Duponchel 1843b: 78).

Family-group name: COENOCHILIDAE Burmeister, 1842. Stem: Coenochil-.

Colaphus Dejean, 1836a: 411. Current status: junior synonym of Colaspidema Laporte, 1833 in CHRYSOMELIDAE: CHRYSOMELINAE: CHRYSOMELINI. Note: unnecessary new replacement name for Colaspidema Laporte, 1833; Colaphus has been attributed to L. Redtenbacher (1845: 116) in the literature and considered a junior homonym of Colaphus Dejean, 1836; however, both names apply to the same taxon; the earlier usage of the name Colaphus by Dahl (1823: 46) was suppressed for the purposes of zoological nomenclature (see ICZN 1964).

Type species: Colaspis barbara Fabricius, 1801, by monotypy (type fixation for Colaspidema Laporte).

Family-group name: COLAPHIDAE Siegel, 1866. Stem: Colaph-.

Colaspis Fabricius, 1801a: xxi, 411. Current status: valid genus in CHRYSOMELIDAE: EUMOLPINAE: EUMOLPINI.

Type species: Chrysomela flavicornis Fabricius, 1787 (= Chrysomela occidentalis Linnaeus, 1758), by subsequent designation (Latreille 1810: 432).

Family-group name: COLASPIDAE Hope, 1841. Stem: Colaspid-.

Colaspoides Laporte, 1833a: 20. Current status: valid genus in CHRYSOMELIDAE: EUMOLPINAE: EUMOLPINI.

Type species: Colaspis limbata G.-A. Olivier, 1808, by subsequent designation (Baly 1864c: 134). Note: Moseyko and Sprecher-Uebersax (2010b: 632) listed Cryptocephalus limbatus Fabricius, 1781 as type species of this genus; however, Baly (1864c: 134) designated “Colaspoides limbata, Oliv.” as type species, an originally included species, and there is no reference to Cryptocephalus limbatus Fabricius, 1781 in G.-A. Olivier’s text; the species described by G.-A. Olivier and Fabricius are both from Cayenne and whether or not they are synonymous has yet to be documented.

Family-group name: COLASPOIDINI S.H. Chen, 1940. Stem: Colaspoid-.

Colasposoma Laporte, 1833a: 22. Current status: valid genus in CHRYSOMELIDAE: EUMOLPINAE: EURYOPINI.

Type species: Colasposoma senegalensis Laporte, 1833, by monotypy.

Family-group name: COLASPOSOMINI Špringlová de Bechyné, 1960. Stem: Colasposomat-.

Colecerus Schönherr, 1840a: 927. Current status: valid genus in CURCULIONIDAE: ENTIMINAE: ENTIMITAE: EUDIAGOGINI.

Type species: Colecerus setosus Boheman, 1840, by original designation.

Family-group name: COLEOCERINI Jekel, 1875. Stem: Colecer-.

Coleomerus Schönherr, 1836: 817. Current status: valid genus in CURCULIONIDAE: BARIDINAE: BARIDINI: COLEOMERINA.

Type species: Coleomerus ebeninus Gyllenhal, 1836, by original designation.

Family-group name: COLEOMERINI Casey, 1922. Stem: Coleomer-.

†Coleopsis Kirejtshuk, Poschmann & Nel in Kirejtshuk et al., 2013: 579. Current status: valid genus in †COLEOPSIDAE. Note: lower Permian (Germany).

Type species: †Coleopsis archaica Kirejtshuk, Poschmann & Nel, 2013, by original designation.

Family-group name: †COLEOPSIDAE Kirejtshuk & Nel, 2016a: 71. Stem: Coleops- (ICZN 1999a, Article 29.4). Note: Kirejtshuk (2020: 10) proposed the emendation COLEOPSEIDAE for this family-group name but the work is nomenclaturally unavailable (ICZN 2012f, Article 8.5.3.1) and, consequently, the emendation is unavailable as well.

Colilodion Besuchet, 1991: 500. Current status: valid genus in STAPHYLINIDAE: PSELAPHINAE: PSELAPHITAE: COLILODIONINI.

Type species: Colilodion inopinatus Besuchet, 1991, by original designation.

Family-group name: COLILODIONINI Besuchet, 1991. Stem: Colilodion-.

Colliuris DeGeer, 1774: 79. Current status: valid genus in CARABIDAE: HARPALINAE: ODACANTHINI: ODACANTHINA.

Type species: Attelabus surinamensis Linnaeus, 1758, by monotypy.

Family-group name: COLLIURINI Bedel, 1910. Stem: Colliur-.

Collops Erichson, 1840a: 54. Current status: valid genus in MELYRIDAE: MALACHIINAE: MALACHIINI.

Type species: Malachius bipunctatus Say, 1823, by subsequent designation (Mayor 2007a: 60).

Family-group name: COLLOPINA Hatch, 1962. Stem: Collop-.

Collyris Fabricius, 1801a: xxiii, 226. Current status: valid genus in CICINDELIDAE: COLLYRIDINI: COLLYRIDINA.

Type species: Cicindela longicollis Fabricius, 1787, by subsequent designation (Latreille 1810: 425).

Family-group name: COLLYRIENS Brullé, 1834. Stem: Collyrid-.

Colobodera Klug, 1837: 53. Current status: senior synonym of Daemon Laporte, 1838 in PTILODACTYLIDAE: ANCHYTARSINAE. Note: this name is considered a junior synonym of Daemon Laporte, 1838 (a name usually incorrectly listed as published in 1836); an application to the Commission is needed to retain Daemon Laporte, 1838 as the valid name for this genus.

Type species: Colobodera ovata Klug, 1837, by present designation.

Family-group name: COLOBODERIDES Erichson, 1847. Stem: Coloboder-.

Colobodes Schönherr, 1837: 465. Current status: valid genus in CURCULIONIDAE: MOLYTINAE: ITHYPORINI: COLOBODINA.

Type species: Colobodes billbergii Boheman, 1837, by original designation.

Family-group name: COLOBODINI Voss, 1958. Stem: Colobod-.

Colobothea Lepeletier & Audinet-Serville, 1825: 336. Current status: valid genus in CERAMBYCIDAE: LAMIINAE: COLOBOTHEINI. Note: the original spelling was Colobotea but Colobothea, an incorrect subsequent spelling, is in prevailing usage, attributed to the publication of the original spelling, and therefore deemed to be the correct original spelling (ICZN 1999a, Article 33.3.1).

Type species: Saperda cassandra Dalman, 1823, by subsequent designation (Duponchel 1843b: 120).

Family-group name: COLOBOTHEITAE J. Thomson, 1860. Stem: Colobothe-.

Colon Herbst, 1797: 224. Current status: valid genus in COLONIDAE. Note: the original spelling Kolon was changed to Colon and Colon Herbst, 1797 placed on the Official List of Generic Names in Zoology (ICZN 1995c).

Type species [suggested]: Colon viennense Herbst, 1797, by subsequent designation (C.G. Thomson 1859: 60; see ICZN 1995c, as Kolon viennensis). Note: name placed on the Official List of Specific Names in Zoology (ICZN 1995c); an earlier type species designation is that of Colon clavigerum Herbst, 1797 (as “Colon claviger Herbst”) by Duponchel (1843b: 124); since the ruling by the ICZN (1995c) did not include a statement such as “all type species designations for prior to that of C.G. Thomson (1859) are hereby set aside”, Duponchel’s designation should be considered the valid type species fixation; however, Colon clavigerum is not currently classified in the nominotypical subgenus and using it would cause instability; an application should be submitted to the Commission to maintain Colon viennense Herbst, 1797 as the type species and dismiss Duponchel’s and any other type species designations before that of C.G. Thomson (1859: 60).

Family-group name: COLONES G.H. Horn, 1880. Stem: Colon-.

Colotes Erichson, 1840a: 129. Current status: valid genus in MELYRIDAE: MALACHIINAE: MALACHIINI.

Type species: Colotes trinotatus Erichson, 1840 (= Malachius maculatus Laporte, 1838), by subsequent designation (Jacquelin du Val 1859a: 47).

Family-group name: CAULAUTAIRES Abeille de Perrin, 1890. Stem: Colot-.

Colpochila Erichson, 1843c: 195. Current status: valid genus in SCARABAEIDAE: SERICOIDINAE: LIPARETRINI.

Type species: Sericesthis gouldii Hope, 1842, by monotypy.

Family-group name: COLPOCHILINI Britton, 1957. Stem: Colpochil-.

Colpoderus Audinet-Serville, 1832: 178. Current status: junior synonym of Notophysis Audinet- Serville, 1832 in CERAMBYCIDAE: PRIONINAE: CACOSCELINI. Note: First Reviser (ICZN 1999a, Article 24.2.2) for Notophysis Audinet-Serville, 1832 versus Colpoderus Audinet-Serville, 1832 is Lameere (1903: 20).

Type species: Colpoderus caffer Audinet-Serville, 1832, by monotypy.

Family-group name: COLPODÉRIDES Lacordaire, 1868. Stem: Colpoder-.

Colpodes W.S. MacLeay, 1825: 17. Current status: valid genus in CARABIDAE: HARPALINAE: PLATYNINI.

Type species: Rembus brunneus W.S. MacLeay, 1825, by monotypy.

Family-group name: COLPODIDAS Chaudoir, 1872. Stem: Colpod-. Note: senior homonym of COLPODIDAE Poche, 1913 (type genus Colpoda O.F. Müller, 1773) in Protista.

Colpothis Gistel, 1848b: 216. Current status: junior synonym of Dasytes Paykull, 1798 in MELYRIDAE: DASYTINAE: DASYTINI. Note: unnecessary new replacement name for Dasytes Paykull, 1798.

Type species: Hispa coerulea Fabricius, 1775 (= Telephorus caeruleus DeGeer, 1774), by subsequent designation (Westwood 1838c: 28) (type fixation for Dasytes Paykull).

Family-group name: COLPOTHISIDAE Gistel, 1848. Stem: Colpoth-.

Colydiopeltis Ślipiński, 1992: 444. Current status: valid genus in LOPHOCATERIDAE.

Type species: Colydiopeltis burckhardti Ślipiński, 1992, by original designation.

Family-group name: COLYDIOPELTINI Kolibáč, 2006. Stem: Colydiopelt-.

Colydium Fabricius, 1792b: 495. Current status: valid genus in ZOPHERIDAE: COLYDIINAE: COLYDIINI. Note: name placed on the Official List of Generic Names in Zoology (ICZN 1995d).

Type species: Bostrichus elongatus Fabricius, 1787, by subsequent designation (Latreille 1810: 431; see ICZN 1995d). Note: name placed on the Official List of Specific Names in Zoology (ICZN 1995d).

Family-group name: COLYDIIDES Billberg, 1820. Stem: Colydi-.

Colymbetes Clairville, 1806: 188. Current status: valid genus in DYTISCIDAE: COLYMBETINAE.

Type species: Dytiscus striatus Linnaeus, 1758, by subsequent designation (Curtis 1828: pl. 207).

Family-group name: COLYMBETINI Erichson, 1837. Stem: Colymbet-.

Colymbomorpha Blanchard in H. Milne-Edwards et al., 1850: 98. Current status: valid genus in SCARABAEIDAE: MELOLONTHINAE: COLYMBOMORPHINI.

Type species: Colymbomorpha lineata Blanchard, 1850, by monotypy.

Family-group name: COLYMBOMORPHITAE Blanchard, 1850. Stem: Colymbomorph-.

†Colymbotethis Ponomarenko, 1994: 189. Current status: valid genus in †COLYMBOTETHIDAE. Note: Upper Triassic (Kazakhstan).

Type species: †Colymbotethis antecessor Ponomarenko, 1994, by original designation.

Family-group name: †COLYMBOTETHIDAE Ponomarenko, 1994. Stem: Colymboteth-.

Commatocerus Raffray, 1882: 1. Current status: valid genus in STAPHYLINIDAE: PSELAPHINAE: CLAVIGERITAE: CLAVIGERINI.

Type species: Commatocerus elegantulus Raffray, 1882, by monotypy.

Family-group name: COMMATOCERINI L.W. Schaufuss, 1882. Stem: Commatocer-.

Comophorina Strand, 1928: 3. Current status: valid genus in SCARABAEIDAE: MELOLONTHINAE: COMOPHINI. Note: new replacement name for Comophorus Blanchard, 1850.

Type species: Comophorus testaceipennis Blanchard, 1850, by monotypy (type fixation for Comophorus Blanchard).

Family-group name: COMOPHORININI Britton, 1987. Stem: Comophorin-.

Comophorus Blanchard in H. Milne-Edwards et al., 1850: 106. Current status: invalid senior synonym of Comophorina Strand, 1928 in SCARABAEIDAE: MELOLONTHINAE: COMOPHINI. Note: junior homonym of Comophorus Agassiz, 1846 [Pisces].

Type species: Comophorus testaceipennis Blanchard, 1850, by monotypy.

Family-group name: COMOPHORINI Britton, 1957. Stem: Comophor-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

Comophus Britton, 1978: 7. Current status: junior synonym of Comophorina Strand, 1928 in SCARABAEIDAE: MELOLONTHINAE: COMOPHINI. Note: new replacement name for Comophorus Blanchard, 1850.

Type species: Comophorus testaceipennis Blanchard, 1850, by monotypy (type fixation for Comophorus Blanchard).

Family-group name: COMOPHINI Britton, 1978. Stem: Comoph-.

Compactopedia Kistner, 1970c: 18. Current status: valid genus in STAPHYLINIDAE: ALEOCHARINAE: ALEOCHARINI: COMPACTOPEDIINA.

Type species: Compactopedia compactus Kistner, 1970, by original designation.

Family-group name: COMPACTOPEDINA Kistner, 1970. Stem: Compactopedi-.

Compsa Perty, 1832: 92. Current status: valid genus in CERAMBYCIDAE: CERAMBYCINAE: TROPIDINI: COMPSINA.

Type species: Compsa albopicta Perty, 1832, by subsequent designation (R. Lucas 1920: 198).

Family-group name: COMPSINA Martins & Galileo, 2007. Stem: Comps-. Note: junior homonym of COMPSINI Pierce, 1913 (type genus Compsus Schönherr, 1823) in Coleoptera: CURCULIONIDAE.

Compsocerus Audinet-Serville, 1834b: 62. Current status: valid genus in CERAMBYCIDAE: CERAMBYCINAE: COMPSOCERINI. Note: this genus-group name was first made available by Lacordaire (1830b: 175) when he listed Saperda barbicornis Fabricius, 1792 (as “C. barbicornis (Saperda plumigera, Oliv.)”) within it; however, that species is the type species of Paromoeocerus Gounelle, 1910, and a request should be submitted to the Commission to suppress Compsocerus Lacordaire, 1830 for the purposes of the Principles of Priority and Homonymy (see Bousquet et al. 2009: 44).

Type species: Compsocerus barbicornis Audinet-Serville, 1834, by monotypy.

Family-group name: COMPSOCERITAE J. Thomson, 1864. Stem: Compsocer-.

Compsocnemis Bonvouloir, 1871: 72 [in key]. Current status: valid genus in EUCNEMIDAE: MELASINAE: MELASINI: COMPSOCNEMINA. Note: this name was first used by Gemminger in Gemminger and von Harold (1869b: 1479) but is unavailable from that date as the three species listed under the name were nomina nuda; the relevant plate of Bonvouloir’s work on EUCNEMIDAE referred to by Gemminger was not issued until 1872 (1872b).

Type species: Compsocnemis maculata Bonvouloir, 1872, by subsequent designation (Muona 1987: 82). Note: no species were originally included in this genus by Bonvouloir (1871); the first ones were the species illustrated by Bonvouloir (1872b: plate 33).

Family-group name: COMPSOCNEMINA Muona, 1993. Stem: Compsocnem-.

Compsoplinthus C. Costa, 1975: 71. Current status: valid genus in ELATERIDAE: AGRYPNINAE: EUPLINTHINI: COMPSOPLINTHINA.

Type species: Pyrophorus ruber Candèze, 1878, by original designation.

Family-group name: COMPSOPLINTHINA C. Costa, 1975. Stem: Compsoplinth-.

Compsosoma Lacordaire, 1830b: 183. Current status: valid genus in CERAMBYCIDAE: LAMIINAE: COMPSOSOMATINI.

Type species: Compsosoma niveosignata Audinet-Serville, 1835, by subsequent designation (E. Desmarest 1860: 325). Note: no available species was originally included for this genus in Lacordaire (1830b); the first ones are the three species listed by Audinet-Serville (1835a: 56, 57).

Family-group name: COMPSOSOMITES J. Thomson, 1857. Stem: Compsosomat-.

Compsus Schönherr, 1823: 1140. Current status: valid genus in CURCULIONIDAE: ENTIMINAE: POLYDRUSITAE: EUSTYLINI: EUSTYLINA.

Type species: Curculio elegans G.-A. Olivier, 1807 (= Curculio argyreus Linnaeus, 1758), by monotypy.

Family-group name: COMPSI Pierce, 1913. Stem: Comps-. Note: senior homonym of COMPSINA Martins & Galileo, 2007 (type genus Compsa Perty, 1832) in Coleoptera: CERAMBYCIDAE.

Conalia Mulsant & Rey, 1858: 313. Current status: valid genus in MORDELLIDAE: MORDELLINAE: CONALIINI.

Type species: Conalia baudii Mulsant & Rey, 1858, by monotypy.

Family-group name: CONALIINI Ermisch, 1956. Stem: Conali-.

Conderis C.O. Waterhouse, 1880a: 59. Current status: valid genus in LYCIDAE: LYCINAE: CONDERINI.

Type species: Calopteron signicolle Kirsch, 1875, by original designation.

Family-group name: CONDERINI Bocák & Bocáková, 1990. Stem: Conder-.

Coniatus Germar, 1817b: 340. Current status: valid genus in CURCULIONIDAE: HYPERINAE: HYPERINI: CONIATINA.

Type species: Curculio tamarisci Fabricius, 1787, by subsequent designation (Schönherr 1823: 1143).

Family-group name: CONIATINA Legalov, 2007. Stem: Coniat-.

Coniontis Eschscholtz, 1829d: 7. Current status: valid genus in TENEBRIONIDAE: PIMELIINAE: CONIONTINI.

Type species: Coniontis viatica Eschscholtz, 1829, by subsequent designation (Casey 1908: 57).

Family-group name: CONIONTIDAE G.R. Waterhouse, 1858. Stem: Coniont-.

Coniporus C.G. Thomson, 1859: 90. Current status: junior synonym of Aspidiphorus Dejean, 1821 in SPHINDIDAE: SPHINDINAE. Note: name placed on the Official Index of Rejected and Invalid Generic Names in Zoology (see ICZN 1997).

Type species: Nitidula orbiculata Gyllenhal, 1808, by original designation.

Family-group name: CONIPORINA C.G. Thomson, 1859. Stem: Conipor-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

†Conjunctia Ponomarenko, 1977: 87. Current status: valid genus in CARABIDAE: †CONJUNCTIINAE. Note: Lower Cretaceous (Russia; China).

Type species: †Conjunctia prodroma Ponomarenko, 1977, by original designation.

Family-group name: †CONJUNCTIINI Ponomarenko, 1977. Stem: Conjuncti-.

Conoderes Schönherr, 1833: 26. Current status: valid genus in CURCULIONIDAE: CONODERINAE: CONODERINI.

Type species: Conoderes albidus Schönherr, 1833, by original designation.

Family-group name: CONODERIDES Schönherr, 1833. Stem: Conoder-. Note: senior homonym of CONODERINAE Fleutiaux, 1919 (type genus Conoderus Eschscholtz, 1829) in Coleoptera: ELATERIDAE.

Conoderus Eschscholtz, 1829a: 31. Current status: junior synonym of Monocrepidius Eschscholtz, 1829 in ELATERIDAE: AGRYPNINAE: OOPHORINI. Note: the First Reviser (ICZN 1999a, Article 24.2.2) for Conoderus Eschscholtz, 1829 versus Monocrepidius Eschscholtz, 1829 is Candèze (1859: 187).

Type species: Conoderus fuscofasciatus Eschscholtz, 1829, by subsequent designation (Duponchel 1843b: 188).

Family-group name: CONODERINAE Fleutiaux, 1919. Stem: Conoder-. Note: junior homonym of CONODERINAE Schönherr, 1833 (type genus Conoderes Schönherr, 1833) in Coleoptera: CURCULIONIDAE.

Cononotus J.L. LeConte, 1851a: 137. Current status: valid genus in PYROCHROIDAE: AGNATHINAE.

Type species: Cononotus sericans J.L. LeConte, 1851, by subsequent designation (Spilman 1954: 88).

Family-group name: CONONOTINI J.L. LeConte, 1862. Stem: Cononot-.

Conopalpus Gyllenhal, 1810: 547. Current status: valid genus in MELANDRYIDAE: OSPHYINAE.

Type species: Conopalpus flavicollis Gyllenhal, 1810 (= Melyris testacea G.-A. Olivier, 1790), by monotypy.

Family-group name: CONOPALPIENS Mulsant, 1856. Stem: Conopalp-.

Conophorus Schönherr, 1838: 719. Current status: junior synonym of Conoderes Schönherr, 1833 in CURCULIONIDAE: CONODERINAE: CONODERINI. Note: junior homonym of Conophorus Meigen, 1803 [Diptera]; unnecessary new replacement name for Conoderes Schönherr, 1833.

Type species: Conoderes albidus Schönherr, 1833, by original designation (type fixation for Conoderes Schönherr).

Family-group name: CONOPHORIDES Schönherr, 1838. Stem: Conophor-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

Conosoma Kraatz, 1857b: 431. Current status: junior synonym of Tachinus Gravenhorst, 1802 in STAPHYLINIDAE: TACHYPORINAE: TACHYPORINI: EUCONOSOMATINA. Note: junior homonym of Conosoma Lenz, 1802 [Diptera]; new replacement name for Conurus Stephens, 1829.

Type species: Oxyporus bipustulatus Fabricius, 1792, by subsequent designation (Westwood 1838c: 19) (type fixation for Conurus Stephens).

Family-group name: CONOSOMINI Jeannel & Jarrige, 1949. Stem: Conosomat-. Note: family- group name permanently invalid (ICZN 1999a, Article 39).

Conotelus Erichson, 1843a: 249. Current status: valid genus in NITIDULIDAE: CILLAEINAE: CONOTELINI.

Type species: Stenus conicus Fabricius, 1801, by subsequent designation (E. Desmarest 1851: 291).

Family-group name: CONOTELINI Kirejtshuk & Kovalev, 2022: 12. Stem: Conotel-.

Conotrachelus Dejean, 1835: 296. Current status: valid genus in CURCULIONIDAE: MOLYTINAE: CLEOGONINI.

Type species: Balaninus diaconitus Klug, 1829, by subsequent designation (Schönherr 1837: 392).

Family-group name: CONOTRACHELIDES Jekel, 1865. Stem: Conotrachel-.

Coomania Cameron, 1939: 22. Current status: valid genus in STAPHYLINIDAE: STAPHYLININAE: COOMANIINI.

Type species: Coomania tonkinensis Cameron, 1939, by monotypy.

Family-group name: COOMANIINAE Żyła & Solodovnikov, 2019: 124. Stem: Coomani-.

Coomaniella Bourgoin, 1924: 178. Current status: valid genus in BUPRESTIDAE: BUPRESTINAE: COOMANIELLINI.

Type species: Coomaniella modesta Bourgoin, 1924, by subsequent designation (Bílý 1974: 30).

Family-group name: COOMANIELLINI Bílý, 1974. Stem: Coomaniell-.

Copelatus Erichson, 1832: 18, 38. Current status: valid genus in DYTISCIDAE: COPELATINAE.

Type species: Dytiscus posticatus Fabricius, 1801, by monotypy.

Family-group name: COPELATINI Branden, 1884. Stem: Copelat-.

Copobaenus Fairmaire & Germain, 1863: 236. Current status: valid genus in ANTHICIDAE: COPOBAENINAE.

Type species: Copobaenus tristis Fairmaire & Germain, 1863, by subsequent designation (R. Lucas 1920: 201).

Family-group name: COPOBAENINAE M. Abdullah, 1969. Stem: Copobaen-.

†Coprinisphaera Sauer, 1955: 129. Current status: valid genus in †COPRINISPHAERIDAE. Note: Ichnotaxon; Upper Cretaceous to Pleistocene (Antarctica, Argentina, Ecuador, Kenya, Pakistan, Somalia, Tanzania, Uruguay, and USA); this name was given precedence over Fontanai Roselli, 1939, a rarely used senior subjective synonym whenever the two are considered to be synonyms, and Coprinisphaera Sauer, 1955 was placed on the Official List of Generic Names in Zoology (ICZN 2008b).

Type species: †Coprinisphaera ecuadoriensis Sauer, 1955, by monotypy (see ICZN 2008b). Note: name placed on the Official List of Specific Names in Zoology ICZN 2008b).

Family-group name: †COPRINISPHAERIDAE Genise, 2004. Stem: Coprinisphaer-.

Copris Geoffroy, 1762: 87. Current status: valid genus in SCARABAEIDAE: SCARABAEINAE: COPRINI. Note: name placed on the Official List of Generic Names in Zoology (ICZN 1994a).

Type species: Scarabaeus lunaris Linnaeus, 1758, by subsequent designation (Latreille 1810: 428; see ICZN 1994a). Note: name placed on the Official List of Specific Names in Zoology (ICZN 1994a).

Family-group name: COPRIDES Leach, 1815. Stem: Copr-.

Coprobius Latreille, 1829a: 535. Current status: junior synonym of Canthon Hoffmansegg, 1817 in SCARABAEIDAE: SCARABAEINAE: DELTOCHILINI.

Type species: Scarabaeus volvens Fabricius, 1792 (= Scarabaeus pilularius Linnaeus, 1758), by subsequent designation (Reiche 1841: 213).

Family-group name: COPROBIADAE Burmeister, 1873. Stem: Coprobi-.

Coprophilus Latreille, 1829a: 439. Current status: valid genus in STAPHYLINIDAE: OXYTELINAE: COPROPHILINI. Note: name placed on the Official List of Generic Names in Zoology (ICZN 1993b).

Type species: Staphylinus striatulus Fabricius, 1792, by plenary power (see ICZN 1993b). Note: name placed on the Official List of Specific Names in Zoology (ICZN 1993b).

Family-group name: COPROPHILINA Heer, 1839. Stem: Coprophil-.

†Coptoclava Ping, 1928: 39. Current status: valid genus in †COPTOCLAVIDAE: †COPTOCLAVINAE. Note: Lower Cretaceous (China; Mongolia; Russia; South Korea).

Type species: †Coptoclava longipoda Ping, 1928, by original designation.

Family-group name: †COPTOCLAVIDAE Ponomarenko, 1961. Stem: Coptoclav-.

†Coptoclavisca Ponomarenko, 1987: 84. Current status: valid genus in †COPTOCLAVIDAE: †COPTOCLAVISCINAE. Note: Lower Cretaceous (China; Mongolia).

Type species: †Coptoclavisca nigricollinus Ponomarenko, 1987, by original designation.

Family-group name: †COPTOCLAVISCINAE Soriano, Ponomarenko & Delclòs, 2007. Stem: Coptoclavisc-.

Coptocycla Chevrolat in Dejean, 1836a: 372. Current status: valid genus in CHRYSOMELIDAE: CASSIDINAE: CASSIDINI.

Type species: Cassida undecimpunctata Fabricius, 1781, by subsequent designation (Duponchel and Chevrolat 1842b: 211).

Family-group name: COPTOCYCLITAE Spaeth & Reitter, 1926. Stem: Coptocycl-.

Coptodactyla Burmeister, 1846: [Copris 5. Subgen., in key]. Current status: valid genus in SCARABAEIDAE: SCARABAEINAE: COPRINI.

Type species: Coptodactyla emana Burmeister, 1846, by monotypy. Note: combined description of a new genus-group taxon and a single new nominal species (ICZN 1999a, Article 12.2.6).

Family-group name: COPTODACTYLINI Paulian, 1933. Stem: Coptodactyl-.

Coptodera Dejean, 1825: 169, 273. Current status: valid genus in CARABIDAE: HARPALINAE: LEBIINI: PERICALINA.

Type species: Coptodera festiva Dejean, 1825, by subsequent designation (Hope 1838a: 85).

Family-group name: COSTODÉRIDES Chaudoir, 1848. Stem: Coptoder-.

Coptomia Burmeister, 1842: 549. Current status: valid genus in SCARABAEIDAE: CETONIINAE: STENOTARSIINI: COPTOMIINA.

Type species: Cetonia mauritania Gory & Percheron, 1833, by subsequent designation (Duponchel 1843b: 202).

Family-group name: COPTOMIINI Schenkling, 1921. Stem: Coptomi-.

Coptomma Newman, 1840a: 18. Current status: valid genus in CERAMBYCIDAE: CERAMBYCINAE: COPTOMMATINI.

Type species: Coptomma virgatum Newman, 1840 (= Callidium variegatum Fabricius, 1775), by subsequent designation (Lacordaire 1869: 223). Note: Lacordaire (1869) selected Callidium variegatum Fabricius, 1775, not an originally included nominal species, as type species but since he listed it in synonymy with Coptomma virgatum Newman, 1840, an originally included species, he is deemed to have designated the latter species as type species (ICZN 1999a, Article 69.2.2).

Family-group name: COPTOMMIDES Lacordaire, 1869. Stem: Coptommat-.

Coptonotus Chapuis, 1869: 11. Current status: valid genus in CURCULIONIDAE: SCOLYTINAE: COPTONOTINI.

Type species: Coptonotus cyclopus Chapuis, 1869, by monotypy.

Family-group name: COPTONOTIDAE Chapuis, 1869. Stem: Coptonot-.

Coptotermoecia Oke, 1933: 135. Current status: valid genus in STAPHYLINIDAE: ALEOCHARINAE: ATHETINI: COPTOTERMOECIINA.

Type species: Coptotermoecia alutacia Oken, 1933, by original designation.

Family-group name: COPTOTERMOECIINA Kistner & Pasteels, 1970. Stem: Coptotermoeci-.

Coptotomus Say, 1830b: 29. Current status: valid genus in DYTISCIDAE: COPTOTOMINAE.

Type species: Coptotomus serripalpus Say, 1830, by monotypy.

Family-group name: COPTOTOMINI Branden, 1884. Stem: Coptotom-.

Copturus Schönherr, 1825: 586. Current status: valid genus in CURCULIONIDAE: CONODERINAE: LECHRIOPINI.

Type species: Poecilma papaveratum Germar, 1823, by original designation.

Family-group name: COPTURIDAE Desbrochers des Loges, 1892. Stem: Coptur-.

Coraebus Gory & Laporte, 1839: 1. Current status: valid genus in BUPRESTIDAE: AGRILINAE: CORAEBINI: CORAEBINA.

Type species: Buprestis undatus Fabricius, 1787, by subsequent designation (Descarpentries and Villiers 1967: 471).

Family-group name: CORAEBINI Bedel, 1921. Stem: Coraeb-.

Cordobanus Bernhauer, 1910: 386. Current status: valid genus in STAPHYLINIDAE: ALEOCHARINAE: CORDOBANINI.

Type species: Cordobanus mirabilis Bernhauer, 1910, by monotypy.

Family-group name: CORDOBANINI Bernhauer, 1910. Stem: Cordoban-.

Cordylospasta G.H. Horn, 1876: 152. Current status: valid genus in MELOIDAE: MELOINAE: EUPOMPHINI.

Type species: Cordylospasta fulleri G.H. Horn, 1876, by monotypy.

Family-group name: CORDYLOSPASTIDES Wellman, 1910. Stem: Cordylospast-.

Coriacephilus Schedl, 1939: 339. Current status: valid genus in CURCULIONIDAE: SCOLYTINAE: CORIACEPHILINI.

Type species: Stephanoderes coriaceus Eichhoff, 1878, by original designation.

Family-group name: CORIACEPHILINI A.J. Johnson in A.J. Johnson et al., 2020: 12. Stem: Coriacephil-.

Corigetus Desbrochers des Loges, 1872: 662. Current status: valid genus in CURCULIONIDAE: ENTIMINAE: CYPHICERITAE: CYPHICERINI: CYPHICERINA.

Type species: Corigetus marmoratus Desbrochers des Loges, 1873, by subsequent monotypy (Desbrochers des Loges 1873a: 746).

Family-group name: CORIGETINI Faust, 1885. Stem: Coriget-.

Corimalia Gozis, 1885: 129. Current status: valid genus in BRENTIDAE: NANOPHYINAE: CORIMALIINI.

Type species: Nanophyes tamarisci Gyllenhal, 1838, by subsequent designation (Pajni and Bhateja 1982: 433).

Family-group name: CORIMALIINI Alonso-Zarazaga, 1989. Stem: Corimali-.

Corintascaris Basilewsky, 1952a: 227. Current status: valid genus in CARABIDAE: SCARITINAE: SCARITINI: CORINTASCARINA.

Type species: Corintascaris ferreirae Basilewsky, 1952, by original designation.

Family-group name: CORINTASCARINI Basilewsky, 1973. Stem: Corintascar-.

Cormodes Pascoe, 1860b: 46. Current status: valid genus in CLERIDAE: CLERINAE: NATALIINI: CORMODINA.

Type species: Cormodes darwini Pascoe, 1860, by monotypy.

Family-group name: CORMODINA Bartlett, 2021: 750. Stem: Cormod-. Note: two spellings were originally used for the name of this new subtribe, including CORMODIINA (p. 764); as far as we know no one has acted as First Reviser (ICZN 1999a, Article 24.2.3) and we here select CORMODINA as the correct spelling.

Corneolabium Steel, 1950: 54. Current status: valid genus in STAPHYLINIDAE: OMALIINAE: CORNEOLABIINI.

Type species: Corneolabium mandibularis Steel, 1950, by original designation.

Family-group name: CORNEOLABIINI Steel, 1950. Stem: Corneolabi-.

Corotoca Schiødte, 1854: 102. Current status: valid genus in STAPHYLINIDAE: ALEOCHARINAE: COROTOCINI: COROTOCINA.

Type species: Corotoca melantho Schiødte, 1854, by subsequent designation (Fenyes 1919: 21).

Family-group name: COROTOCINI Fenyes, 1919. Stem: Corotoc-.

Corrhecerus Schönherr, 1826: 40. Current status: valid genus in ANTHRIBIDAE: ANTHRIBINAE: CORRHECERINI.

Type species: Anthribus mixtus Germar, 1823, by original designation.

Family-group name: CORRHÉCÉRIDES Lacordaire, 1865. Stem: Corrhecer-.

Corsyra Dejean, 1825: 168, 326. Current status: valid genus in CARABIDAE: HARPALINAE: CORSYRINI.

Type species: Cymindis fusula Fischer, 1820, by monotypy.

Family-group name: CORSYRINI Ganglbauer, 1891. Stem: Corsyr-.

Corthylus Erichson, 1836: 64. Current status: valid genus in CURCULIONIDAE: SCOLYTINAE: CORTHYLINI: CORTHYLINA.

Type species: Bostrichus compressicornis Fabricius, 1801, by subsequent designation (E. Desmarest 1860: 275).

Family-group name: CORTHYLI J.L. LeConte, 1876. Stem: Corthyl-.

Corticaria Marsham, 1802: 106. Current status: valid genus in LATRIDIIDAE: CORTICARIINAE. Note: name placed on the Official List of Generic Names in Zoology (ICZN 2011e).

Type species: Corticaria ferruginea Marsham, 1802, by subsequent designation (Dajoz 1975: 205; see ICZN 2011e). Note: name placed on the Official List of Specific Names in Zoology (ICZN 2011e).

Family-group name: CORTICARIDAE Curtis, 1829. Stem: Corticari-.

Corticeus Piller & Mitterpacher, 1783: 87. Current status: valid genus in TENEBRIONIDAE: DIAPERINAE: HYPOPHLAEINI.

Type species: Corticeus unicolor Piller & Mitterpacher, 1783, by monotypy.

Family-group name: CORTICEINI Boddy, 1965. Stem: Cortice-.

Corticus Germar, 1823: 146. Current status: valid genus in ZOPHERIDAE: COLYDIINAE: SYNCHITINI. Note: name first published as a synonym of Sarrotrium Illiger, 1798 and subsequently used as valid before 1961 by several authors, including Latreille (1829b: 24); see ICZN (1999a, Article 11.6.1).

Type species: Sarrotrium celtis Germar, 1823, by monotypy.

Family-group name: CORTICIDES Reitter, 1882. Stem: Cortic-.

Corylophodes A. Matthews, 1885: 160. Current status: junior synonym of Holopsis Broun, 1883 in CORYLOPHIDAE: CORYLOPHINAE: PELTINODINI.

Type species: Corylophus marginicollis J.L. LeConte, 1852, by present designation. Note: Corylophodes punctipennis A. Matthews, 1899 is often listed as the type species of this genus (e.g., Bowestead 1999: 23) by subsequent designation of Paulian (1950: 22) but that species is not an originally included species.

Family-group name: CORYLOPHODINI Paulian, 1950. Stem: Corylophod-.

Corylophus Stephens, 1833a: 26. Current status: valid genus in CORYLOPHIDAE: CORYLOPHINAE: CORYLOPHINI.

Type species: Dermestes cassidoides Marsham, 1802, by monotypy.

Family-group name: CORYLOPHI J.L. LeConte, 1852. Stem: Coryloph-.

Corymbites Latreille, 1834: 150. Current status: junior synonym of Ctenicera Latreille, 1829 in ELATERIDAE: DENDROMETRINAE: PROSTERNINI.

Type species: Elater pectinicornis Linnaeus, 1758, by subsequent designation (C.G. Thomson 1859: 102).

Family-group name: CORYMBITINI J.L. LeConte, 1861. Stem: Corymbit-.

Corynemerus Fåhraeus, 1871: 281. Current status: valid genus in CURCULIONIDAE: CONODERINAE: CORYNEMERINI.

Type species: Corynemerus femoralis Fåhraeus, 1871, by subsequent designation (Faust 1898b: 78).

Family-group name: CORYNEMERINAE Hustache, 1929. Stem: Corynemer-.

Corynodes Hope, 1841a: 162. Current status: junior synonym of Platycorynus Chevrolat, 1836 in CHRYSOMELIDAE: EUMOLPINAE: EUMOLPINI.

Type species: Chrysomela cyanea Fabricius, 1792, by original designation. Note: the type species listed by Hope (1841a: 162) is “E[umolpus]. Cyaneus, Fabricius” and Fabricius (1801a: 419) referred this name to his Chrysomela cyanea Fabricius, 1792 described from “America meridionali”; it seems that the locality listed by Fabricius (1792a: 324; 1801a: 419) could be erroneous as G.-A. Olivier (1808: 900) recorded the species from “Bengale”, more research is needed to ascertain all the facts; the type species usually listed for this genus is Eumolpus compressicornis Fabricius, 1801 described from Guinea (e.g., Jacoby 1908: 492; Moseyko and Sprecher-Uebersax 2010b: 633).

Family-group name: CORYNODINA T.A. Marshall, 1865. Stem: Corynod-.

Corynofrea Aurivillius, 1911a: 37. Current status: valid genus in CERAMBYCIDAE: LAMIINAE: CEROPLESINI: CROSSOTINA.

Type species: Corynofrea mirabilis Aurivillius, 1911, by monotypy.

Family-group name: CORYNOFREINAE Aurivillius, 1911. Stem: Corynofre-.

Corynomalus Chevrolat in Dejean, 1836a: 439. Current status: valid genus in ENDOMYCHIDAE: LYCOPERDININAE. Note: this name is often credited to Erichson (1847c: 181) (e.g., Strohecker 1980: 14; Tomaszewska 2005: 17) but it was first made available by Chevrolat in 1836.

Type species: Eumolpus limbatus G.-A. Olivier, 1808 (= Erotylus marginatus Fabricius, 1798), by subsequent designation (Bousquet and Bouchard 2013a: 139).

Family-group name: CORYNOMALIDAE Gorham, 1873. Stem: Corynomal-.

Coryphium Stephens, 1834: 344. Current status: valid genus in STAPHYLINIDAE: OMALIINAE: CORYPHIINI: CORYPHIINA. Note: name placed on the Official List of Generic Names in Zoology (ICZN 1990b).

Type species: Coryphium angusticolle Stephens, 1834, by monotypy (see ICZN 1990b). Note: name placed on the Official List of Specific Names in Zoology (ICZN 1990b).

Family-group name: CORYPHIINA Jakobson, 1908. Stem: Coryphi-.

Coryphocera Burmeister, 1842: 220. Current status: valid subgenus of Dicheros Gory & Percheron, 1833 in SCARABAEIDAE: CETONIINAE: GOLIATHINI: RHOMBORHININA.

Type species: Dicheros cuvera Newman, 1838, by subsequent designation (Mikšić 1974: 753).

Family-group name: CORYPHOCERIDAE Burmeister, 1842. Stem: Coryphocer-.

Corypticus Sturm, 1843: 347. Current status: junior synonym of Xiphodontus Westwood, 1838 in LUCANIDAE: LUCANINAE: LUCANINI. Note: two original spellings were used, including Coryptius (p. 136); Bouchard et al. (2011: 233) acted as First Revisers (ICZN 1999a, Article 24.2.3) and selected Corypticus.

Type species: Corypticus capensis Sturm, 1843 (= Xiphodontus antilope Westwood, 1838), by monotypy.

Family-group name: CORYPTICIDAE Gistel, 1848. Stem: Coryptic-.

Coryssomerus Schönherr, 1825: 583. Current status: junior synonym of Poecilma Germar, 1821 in CURCULIONIDAE: CONODERINAE: CORYSSOMERINI. Note: see Anzaldo (2022: 339) regarding the synonymy of the type genus and usage of Poecilma as valid, however, Alonso-Zarazaga et al. (2023: 8) advocate for usage of Coryssomerus as valid while an application to the Commission is being prepared.

Type species: Rhynchaenus capucinus Beck, 1817, by original designation.

Family-group name: CORYSSOMERINA C.G. Thomson, 1859. Stem: Coryssomer-.

Coryssopus Schönherr, 1826: 303. Current status: valid genus in CURCULIONIDAE: CONODERINAE: CORYSSOPODINI.

Type species: Zygops hexastichus Schönherr, 1826, by original designation.

Family-group name: CORYSSOPIDES Lacordaire, 1865. Stem: Coryssopod-.

Corythoderus Klug, 1845: [Decas 5], sign. b. Current status: valid genus in SCARABAEIDAE: APHODIINAE: CORYTHODERINI.

Type species: Corythoderus loripes Klug, 1845, by monotypy.

Family-group name: CORYTHODERINA A. Schmidt, 1910. Stem: Corythoder-.

Coscinia Dejean, 1831: 478. Current status: invalid senior synonym of Cymbionotum Baudi di Selve, 1864 in CARABIDAE: MELAENINAE. Note: junior homonym of Coscinia Hübner, 1819 [Lepidoptera].

Type species: Siagona schuppelii Dejean, 1825, by subsequent designation (Hope 1838a: 81).

Family-group name: COSCINIDES Chaudoir, 1876. Stem: Coscini-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

Cosmovalgus Kolbe, 1897a: 190, 204. Current status: valid genus in SCARABAEIDAE: CETONIINAE: VALGINI: VALGINA.

Type species: Cosmovalgus conradti Kolbe, 1897, by original designation.

Family-group name: COSMOVALGINAE Kolbe, 1904. Stem: Cosmovalg-.

Cosnania Dejean, 1821: 2. Current status: valid subgenus of Colliuris DeGeer, 1774 in CARABIDAE: HARPALINAE: ODACANTHINI. Note: this name was used in the literature for a long time under the spelling Casnonia and attributed to Latreille in Latreille and Dejean (1824: 130) or Dejean (1825: 170) but the name was first made available by Dejean (1821) under the spelling Cosnania, which is currently in prevailing usage; this name is a junior objective synonym of Macrotrachelus Latreille, 1818 (see Bousquet 2002: 30), which was not used as valid after 1899 to our knowledge; however, we are unable to provide 25 references to conserve usage of Cosnania (or the variant spelling Casnonia) as the valid name (ICZN 1999a, Article 23.9.1.2); nevertheless we believe that Cosnania Dejean, 1821 should be retained as the valid name and the case submitted to the Commission for a ruling.

Type species: Attelabus pensylvanicus Linnaeus, 1758, by monotypy.

Family-group name: CASNONIAE J.L. LeConte, 1861. Stem: Cosnani-.

Cossonus Clairville, 1798: 57, 58. Current status: valid genus in CURCULIONIDAE: COSSONINAE: COSSONINI.

Type species: Curculio linearis Fabricius, 1775, by subsequent designation (Latreille 1810: 431).

Family-group name: COSSONIDES Schönherr, 1825. Stem: Cosson-.

Cossyphodes Westwood, 1851: 168. Current status: valid genus in TENEBRIONIDAE: PIMELIINAE: COSSYPHODINI: COSSYPHODINA.

Type species: Cossyphodes wollastonii Westwood, 1851, by monotypy.

Family-group name: COSSYPHODIDAE Wasmann, 1899. Stem: Cossyphod-.

Cossyphodites Brauns, 1901: 91. Current status: valid genus in TENEBRIONIDAE: PIMELIINAE: COSSYPHODINI: COSSYPHODITINA.

Type species: Cossyphodes woodrooffei Péringuey, 1885, by monotypy.

Family-group name: COSSYPHODITINAE Basilewsky, 1950. Stem: Cossyphodit-.

Cossyphus G.-A. Olivier, 1791: 121. Current status: valid genus in TENEBRIONIDAE: LAGRIINAE: COSSYPHINI.

Type species: Lampyris depressa Fabricius, 1781, by monotypy.

Family-group name: COSSYPHORES Latreille, 1802. Stem: Cossyph-. Note: this family-group name is a junior homonym of COSSYPHINA Vigors, 1825 (type genus Cossypha Vigors, 1825) in Aves.

Cotaster Motschulsky, 1851: 425. Current status: valid genus in CURCULIONIDAE: COSSONINAE: DRYOTRIBINI.

Type species: Phloeophagus uncipes Boheman, 1838, by subsequent designation (Lacordaire 1865: 330).

Family-group name: COTASTERINEN Faust, 1886. Stem: Cotaster-.

Cotasteromimus Chûjô & Voss, 1960: 10. Current status: valid genus in CURCULIONIDAE: MOLYTINAE: PISSODINI: COTASTEROMIMINA.

Type species: Cotasteromimus morimotoi Chûjô & Voss, 1960, by original designation.

Family-group name: COTASTEROMIMINI Morimoto, 1962. Stem: Cotasteromim-.

Cotasterosoma Konishi, 1962: 3. Current status: valid genus in CURCULIONIDAE: COSSONINAE: RHYNCOLINI: RHYNCOLINA.

Type species: Cotasterosoma omogoense Konishi, 1962, by original designation.

Family-group name: COTASTEROSOMINI Konishi, 1962. Stem: Cotasterosomat-.

Coxelus Dejean, 1821: 67. Current status: valid genus in ZOPHERIDAE: COLYDIINAE: SYNCHITINI.

Type species: Bolitophagus pictus Sturm, 1807, by monotypy.

Family-group name: COXELINI Seidlitz, 1872. Stem: Coxel-.

Craniotus J.L. LeConte, 1851a: 142. Current status: valid genus in TENEBRIONIDAE: PIMELIINAE: ASIDINI.

Type species: Craniotus pubescens J.L. LeConte, 1851, by monotypy.

Family-group name: CRANIOTINI J.L. LeConte & G.H. Horn, 1883. Stem: Craniot-.

Cranophorus Mulsant, 1850: 940. Current status: valid genus in COCCINELLIDAE: COCCINELLINAE: COCCIDULINI.

Type species: Cranophorus quadrinotatus Mulsant, 1850, by subsequent designation (Crotch 1874a: 293).

Family-group name: CRANOPHORAIRES Mulsant, 1850. Stem: Cranophor-.

Cranopoeus G.A.K. Marshall, 1931: 274. Current status: valid genus in CURCULIONIDAE: CURCULIONINAE: CRANOPOEINI.

Type species: Cranopoeus turritus G.A.K. Marshall, 1931, by original designation.

Family-group name: CRANOPOEINI Kuschel, 2009. Stem: Cranopoe-.

Craspedomerus Bernhauer, 1911: 88. Current status: valid genus in STAPHYLINIDAE: STAPHYLININAE: STAPHYLININI: PHILONTHINA.

Type species: Philonthus glenoides Schubert, 1908, by monotypy.

Family-group name: CRASPEDOMERI Bernhauer, 1911. Stem: Craspedomer-.

Cratacanthus Dejean, 1829: 4, 40. Current status: valid genus in CARABIDAE: HARPALINAE: HARPALINI: HARPALINA.

Type species: Cratacanthus pensylvanicus Dejean, 1829 (= Harpalus dubius Palisot de Beauvois, 1812), by monotypy.

Family-group name: CRATACANTHI Lindroth, 1968. Stem: Cratacanth-.

Cratocara J.L. LeConte, 1863a: 11. Current status: junior synonym of Polpochila Solier, 1849 in CARABIDAE: HARPALINAE: HARPALINI: STENOLOPHINA. Note: new replacement name for Melanotus Dejean, 1831.

Type species: Melanotus flavipes Dejean, 1831, by subsequent designation (Hope 1838a: 111) (type fixation for Melanotus Dejean).

Family-group name: CRATOCARINI Casey, 1914. Stem: Cratocar-.

Cratocerus Dejean, 1829: 3, 12. Current status: valid genus in CARABIDAE: HARPALINAE: CRATOCERINI.

Type species: Cratocerus monilicornis Dejean, 1829, by monotypy.

Family-group name: CRATOCÉRIDES Lacordaire, 1854. Stem: Cratocer-.

Cratogaster Blanchard, 1843: pl. 2. Current status: valid genus in CARABIDAE: HARPALINAE: PTEROSTICHINI: PTEROSTICHINA.

Type species: Cratogaster sulcatus Blanchard, 1843, by monotypy.

Family-group name: CRATOGASTRI Csiki, 1929. Stem: Cratogastr-.

Cratognathus Dejean, 1829: 4, 46. Current status: valid genus in CARABIDAE: HARPALINAE: HARPALINI: HARPALINA.

Type species: Cratognathus mandibularis Dejean, 1829, by monotypy.

Family-group name: CRATOGNATHIDAE Laporte, 1834. Stem: Cratognath-.

Cratomorphus Motschulsky, 1853a: 35. Current status: valid genus in LAMPYRIDAE: LAMPYRINAE: CRATOMORPHINI.

Type species: Photinus fabricii Laporte, 1840, by original designation.

Family-group name: CRATOMORPHINAE E. Olivier, 1911. Stem: Cratomorph-.

Cratonychus Dejean, 1833a: 87. Current status: junior synonym of Melanotus Eschscholtz, 1829 in ELATERIDAE: ELATERINAE: MELANOTINI. Note: unnecessary new replacement name for Melanotus Eschscholtz, 1829.

Type species: Elater fulvipes Herbst, 1806 (= Elater villosus Geoffroy, 1785), by subsequent designation (Westwood 1838c: 26) (type fixation for Melanotus Eschscholtz).

Family-group name: CRATONYCHIDAE Gistel, 1848. Stem: Cratonych-.

Cratoparis Dejean, 1834: 235. Current status: junior synonym of Euparius Schönherr, 1823 in ANTHRIBIDAE: ANTHRIBINAE: CRATOPARINI. Note: unnecessary new replacement name for Euparius Schönherr, 1823.

Type species: Anthribus lunatus Fabricius, 1801 (= Macrocephalus marmoreus G.-A. Olivier, 1800), by monotypy (type fixation for Euparius Schönherr).

Family-group name: CRATOPARES J.L. LeConte, 1876. Stem: Cratopar-.

Cratopus Schönherr, 1823: 1140. Current status: valid genus in CURCULIONIDAE: ENTIMINAE: POLYDRUSITAE: CRATOPODINI.

Type species: Curculio melanocephalus Fabricius, 1798, by original designation.

Family-group name: CRATOPINI Hustache, 1919. Stem: Cratopod-.

Cremastocheilus Knoch, 1801: 115. Current status: valid genus in SCARABAEIDAE: CETONIINAE: CREMASTOCHEILINI: CREMASTOCHEILINA.

Type species: Cremastocheilus castaneae Knoch, 1801, by monotypy.

Family-group name: CREMASTOCHILIDAE Burmeister & Schaum, 1841. Stem: Cremastocheil-.

Crematoxenus Mann, 1921: 547. Current status: valid genus in STAPHYLINIDAE: ALEOCHARINAE: CREMATOXENINI.

Type species: Crematoxenus aenigma Mann, 1921, by original designation.

Family-group name: CREMATOXENINI Mann, 1921. Stem: Crematoxen-.

Creobius Guérin-Méneville, 1839a: pl. 225 (p. 4). Current status: valid genus in CARABIDAE: BROSCINAE: BROSCINI: CREOBIINA.

Type species: Feronia eydouxii Guérin-Méneville, 1839, by monotypy.

Family-group name: CREOBITAE Jeannel, 1941. Stem: Creobi-.

Creophilus Leach in Samouelle, 1819: 172. Current status: valid genus in STAPHYLINIDAE: STAPHYLININAE: STAPHYLININI: STAPHYLININA. Note: name placed on the Official List of Generic Names in Zoology (ICZN 1959a).

Type species: Staphylinus maxillosus Linnaeus, 1758, by original designation (see ICZN 1959a). Note: name placed on the Official List of Specific Names in Zoology (ICZN 1959a).

Family-group name: CREOPHILIDAE W. Kirby, 1837. Stem: Creophil-.

Crepidodera Chevrolat in Dejean, 1836a: 391. Current status: valid genus in CHRYSOMELIDAE: GALERUCINAE: ALTICINI.

Type species: Chrysomela nitidula Linnaeus, 1758, by subsequent designation (Maulik 1926: 234).

Family-group name: CRÉPIDODÉRITES Chapuis, 1875. Stem: Crepidoder-.

Crepidogaster Boheman, 1848: 68. Current status: valid genus in CARABIDAE: BRACHININAE: CREPIDOGASTRINI.

Type species: Crepidogaster bimaculatus Boheman, 1848, by monotypy.

Family-group name: CREPIDOGASTRITAE Jeannel, 1949. Stem: Crepidogastr-.

Crepidomenus Erichson, 1842: 140. Current status: valid genus in ELATERIDAE: DENDROMETRINAE: CREPIDOMENINI.

Type species: Crepidomenus fulgidus Erichson, 1842, by subsequent designation (Hyslop 1921: 637).

Family-group name: CRÉPIDOMÉNITES Candèze, 1863. Stem: Crepidomen-.

Crepidotus Schönherr, 1838: 859. Current status: valid genus in CURCULIONIDAE: DRYOPHTHORINAE: LITOSOMINI: CREPIDOTINA.

Type species: Crepidotus audouini Gyllenhal, 1838 (= Calandra variolosa Klug, 1833), by original designation.

Family-group name: CREPIDOTINA Legalov, 2020a: 323. Stem: Crepidot-.

†Cretakarenni Peris & Delclòs, 2015: 335. Current status: valid genus in MONOTOMIDAE: MONOTOMINAE: †CRETAKARENNIINI. Note: Cretaceous (Myanmar; Spain).

Type species: †Cretakarenni birmanicus Peris & Delclòs, 2015, by original designation.

Family-group name: †CRETAKARENNIINI Peris & Delclòs, 2015: 335. Stem: Cretakarenni-.

†Cretanthribus Legalov, 2009a: 292. Current status: valid genus in ANTHRIBIDAE: ANTHRIBINAE: †CRETANTHRIBINI. Note: Lower Cretaceous (Russia).

Type species: †Anthribites cretaceus Zherikhin, 1993, by original designation.

Family-group name: †CRETANTHRIBINI Legalov, 2009. Stem: Cretanthrib-.

†Cretochodaeus Nikolajev, 1995a: 79. Current status: valid genus in OCHODAEIDAE: †CRETOCHODAEINAE. Note: Lower Cretaceous (Mongolia).

Type species: †Cretochodaeus mongolicus Nikolajev, 1995, by original designation.

Family-group name: †CRETOCHODAEINI Nikolajev, 1995. Stem: Cretochodae-.

†Cretochoragus Soriano, Gratshev & Delclòs, 2006: 557. Current status: valid genus in ANTHRIBIDAE: CHORAGINAE: †CRETOCHORAGINI. Note: Lower Cretaceous (Spain).

Type species: †Cretochoragus pygmaeus Soriano, Gratshev & Delclòs, 2006, by original designation.

Family-group name: †CRETOCHORAGINI Legalov, 2016: 1460. Stem: Cretochorag-.

†Cretocoma Nikolajev, 2002: 54. Current status: valid genus in PLEOCOMIDAE: †CRETOCOMINAE. Note: Lower Cretaceous (Mongolia).

Type species: †Cretocoma tologoica Nikolajev, 2002, by original designation.

Family-group name: †CRETOCOMINI Nikolajev, 2002. Stem: Cretocom-.

†Cretodermestes Deng, Ślipiński, Ren & H. Pang, 2017a: 3. Current status: valid genus in DERMESTIDAE: ATTAGENINAE: †CRETODERMESTINI. Note: mid-Cretaceous (Myanmar).

Type species: †Cretodermestes palpalis Deng, Ślipiński, Ren & H. Pang, 2017, by original designation.

Family-group name: †CRETODERMESTINI Deng, Ślipiński, Ren & H. Pang, 2017: 3. Stem: Cretodermest-.

†Cretogeotrupes Nikolajev, 1992: 80. Current status: valid genus in GEOTRUPIDAE: GEOTRUPINAE: †CRETOGEOTRUPINI. Note: Lower Cretaceous (China; Russia).

Type species: †Cretogeotrupes convexus Nikolajev, 1992, by original designation.

Family-group name: †CRETOGEOTRUPINI Nikolajev, 1996. Stem: Cretogeotrup-.

†Cretoglaphyrus Nikolajev, 2005a: 70. Current status: valid genus in GLAPHYRIDAE: †CRETOGLAPHYRINAE. Note: Lower Cretaceous (Transbaikalia).

Type species: †Cretoglaphyrus rohdendorfi Nikolajev, 2005, by original designation.

Family-group name: †CRETOGLAPHYRINI Nikolajev, 2005. Stem: Cretoglaphyr-.

†Cretohister Zhou, Caterino, Ślipiński & Cai, 2018: 722. Current status: valid genus in †CRETOHISTERIDAE. Note: Lower Cretaceous (China).

Type species: †Cretohister sinensis Zhou, Caterino, Ślipiński & Cai, 2018, by original designation.

Family-group name: †CRETOHISTERIDAE Zhou, Caterino, Ślipiński & Cai, 2018: 721. Stem: Cretohister-.

†Cretolycus Tihelka, D. Huang & Cai, 2019a: 263. Current status: valid genus in LYCIDAE: DEXORINAE: †CRETOLYCINI. Note: mid-Cretaceous (Myanmar).

Type species: †Cretolycus praecursor Tihelka, D. Huang & Cai, 2019, by original designation.

Family-group name: †CRETOLYCINI Tihelka, D. Huang & Cai, 2019: 263. Stem: Cretolyc-.

†Cretomelolontha Nikolajev, 1998: 81. Current status: valid genus in SCARABAEIDAE: MELOLONTHINAE: †CRETOMELOLONTHINI. Note: Lower Cretaceous (Russia).

Type species: †Cretomelolontha transbaikalica Nikolajev, 1998, by original designation.

Family-group name: †CRETOMELOLONTHINI Nikolajev, 1998. Stem: Cretomelolonth-.

†Cretonemonyx Gratshev & Legalov, 2009: 412. Current status: valid genus in NEMONYCHIDAE: †CRETONEMONYCHINAE: †CRETONEMONYCHINI. Note: Lower Cretaceous (Russia).

Type species: †Cretonemonyx profligatus Gratshev & Legalov, 2009, by original designation.

Family-group name: †CRETONEMONYCHINAE Gratshev & Legalov, 2009. Stem: Cretonemonych-.

†Cretonodes Kirejtshuk & Azar in Kirejtshuk et al., 2009a: 121. Current status: valid genus in DERMESTIDAE: TRINODINAE: †CRETONODINI. Note: Lower Cretaceous (Lebanon).

Type species: †Cretonodes antounazari Kirejtshuk & Azar, 2009, by original designation.

Family-group name: †CRETONODINI Kirejtshuk & Azar, 2009. Stem: Cretonod-.

†Cretoparacucujus Cai & Escalona in Cai et al., 2018: 2806. Current status: valid genus in †PARANDREXIDAE: †CRETOPARACUCUJINAE. Note: mid-Cretaceous (Myanmar).

Type species: †Cretoparacucujus cycadophilus Cai & Escalona, 2018, by original designation. Note: combined description of a new genus and a single new species (ICZN 1999a, Article 13.4).

Family-group name: †CRETOPARACUCUJINAE Kirejtshuk in Kirejtshuk et al., 2023a: 24. Stem: Cretoparacucu-.

†Cretophengodes Y.-D. Li, Kundrata, Tihelka & Cai in Y.-D. Li et al., 2021: 2. Current status: valid genus in †CRETOPHENGODIDAE. Note: mid-Cretaceous (Myanmar).

Type species: †Cretophengodes azari Y.-D. Li, Kundrata, Tihelka & Cai, 2021, by original designation.

Family-group name: †CRETOPHENGODIDAE Y.-D. Li, Kundrata, Tihelka & Cai in Y.-D. Li et al., 2021: 2. Stem: Cretophengod-.

†Cretoscarabaeus Nikolajev, 1995b: 147. Current status: valid genus in SCARABAEIDAE: †CRETOSCARABAEINAE. Note: Lower Cretaceous (Russia).

Type species: †Cretoscarabaeus gibbosus Nikolajev, 1995, by original designation.

Family-group name: †CRETOSCARABAEINAE Nikolajev, 1995. Stem: Cretoscarabae-.

†Cretulio Zherikhin, 1993: 33. Current status: valid genus in CARIDAE: CHILECARINAE: †CRETULIINI. Note: Lower Cretaceous (Russia).

Type species: †Cretulio nucula Zherikhin, 1993, by original designation.

Family-group name: †CRETULIINI Legalov, 2009. Stem: Cretuli-.

Crinotarsus Blanchard, 1846: pl. 16. Current status: valid genus in CERAMBYCIDAE: LAMIINAE: DESMIPHORINI.

Type species: Crinotarsus plagiatus Blanchard, 1846, by monotypy.

Family-group name: CRINOTARSIDES Lacordaire, 1872. Stem: Crinotars-.

Criocephalum Dejean, 1835: 328. Current status: junior synonym of Arhopalus Audinet-Serville, 1834 in CERAMBYCIDAE: SPONDYLIDINAE: ASEMINI.

Type species: Cerambyx rusticus Linnaeus, 1758, by monotypy.

Family-group name: CRIOCEPHALINAE Perrier, 1893. Stem: Criocephal-.

Crioceris Geoffroy, 1762: 237. Current status: valid genus in CHRYSOMELIDAE: CRIOCERINAE: CRIOCERINI. Note: name placed on the Official List of Generic Names in Zoology as Crioceris O.F. Müller, 1764 (ICZN 1970a); however, the author of this name was subsequently changed to Geoffroy, 1762 (ICZN 1994a).

Type species: Chrysomela asparagi Linnaeus, 1758, by plenary power (see ICZN 1970a). Note: name placed on the Official List of Specific Names in Zoology (ICZN 1970a, 1994a).

Family-group name: CRIOCERIDES Latreille, 1804. Stem: Criocer-.

Criomorphus Mulsant, 1839: 58. Current status: junior synonym of Tetropium W. Kirby, 1837 in CERAMBYCIDAE: SPONDYLIDINAE: TETROPIINI. Note: junior homonym of Criomorphus Curtis, 1831 [Hemiptera].

Type species: Callidium aulicum Fabricius, 1775 (= Cerambyx castaneus Linnaeus, 1758), by monotypy.

Family-group name: CRIOMORPHINI Portevin, 1927. Stem: Criomorph-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

Crossotarsus Chapuis, 1865: 22, 44. Current status: valid genus in CURCULIONIDAE: PLATYPODINAE: PLATYPODINI.

Type species: Platypus wallacei J. Thomson, 1858, by subsequent designation (Hopkins 1914: 119).

Family-group name: CROSSOTARSARIAE H. Strohmeyer, 1914. Stem: Crossotars-.

Crossotus Audinet-Serville, 1835a: 52. Current status: valid genus in CERAMBYCIDAE: LAMIINAE: CEROPLESINI: CROSSOTINA.

Type species: Crossotus plumicornis Audinet-Serville, 1835, by monotypy.

Family-group name: CROSSOTITAE J. Thomson, 1864. Stem: Crossot-.

Crowsoniella Pace, 1975: 446. Current status: valid genus in CROWSONIELLIDAE.

Type species: Crowsoniella relicta Pace, 1975, by original designation.

Family-group name: CROWSONIELLIDAE Iablokoff-Khnzorian, 1983. Stem: Crowsoniell-.

Cryphalus Erichson, 1836: 61. Current status: valid genus in CURCULIONIDAE: SCOLYTINAE: CRYPHALINI.

Type species [suggested]: Bostrichus asperatus Gyllenhal, 1813, by subsequent designation (C.G. Thomson 1859: 146). Note: the valid type species fixation is that of Laporte (1840b: 370), who selected Apate tiliae Panzer, 1793, but that species is the type species of the genus Ernoporus C.G. Thomson, 1859; an application to the Commission is necessary to fix Bostrichus asperatus Gyllenhal, 1813 as the type species of Cryphalus Erichson, 1836.

Family-group name: CRYPHALOIDEAE Lindemann, 1876. Stem: Cryphal-.

Cryptafricus Leschen, 1996: 624. Current status: valid genus in CRYPTOPHAGIDAE: ATOMARIINAE: CRYPTAFRICINI.

Type species: Cryptafricus leleupi Leschen, 1996, by original designation.

Family-group name: CRYPTAFRICINI Leschen, 1996. Stem: Cryptafric-.

Cryptamorpha Wollaston, 1854: 156. Current status: valid genus in SILVANIDAE: BRONTINAE: TELEPHANINI.

Type species: Cryptomorpha musae Wollaston, 1854 (= Psammoecus desjardinsii Guérin- Méneville, 1844), by monotypy.

Family-group name: CRYPTAMORPHINI Casey, 1884. Stem: Cryptamorph-.

Cryptarcha Shuckard, 1839: 165. Current status: valid genus in NITIDULIDAE: CRYPTARCHINAE: CRYPTARCHINI.

Type species: Nitidula strigata Fabricius, 1787, by subsequent designation (Hope 1841a: 154).

Family-group name: CRYPTARCHINA C.G. Thomson, 1859. Stem: Cryptarch-.

Crypticus Latreille, 1816: 298. Current status: valid genus in TENEBRIONIDAE: DIAPERINAE: CRYPTICINI.

Type species: Helops glaber Fabricius, 1775 (= Tenebrio quisquilius Linnaeus, 1761), by monotypy.

Family-group name: CRYPTICITES Brullé, 1832. Stem: Cryptic-.

Cryptobium Mannerheim, 1830: 38. Current status: junior synonym of Ochthephilum Stephens, 1829 in STAPHYLINIDAE: PAEDERINAE: PAEDERINI: CRYPTOBIINA.

Type species: Paederus fracticornis Paykull, 1800, by monotypy.

Family-group name: CRYPTOBIA Casey, 1905. Stem: Cryptobi-. Note: senior homonym of CRYPTOBIIDAE Poche, 1913 (type genus Cryptobia Leidy, 1846) in Kinetoplastida.

†Cryptocardius Dolin, 1980: 74. Current status: valid genus in ELATERIDAE: AGRYPNINAE: †CRYPTOCARDIINI. Note: Jurassic (Kazakhstan).

Type species: †Cryptocardius mirabilis Dolin, 1980, by original designation.

Family-group name: †CRYPTOCARDIINI Dolin, 1980. Stem: Cryptocardi-.

Cryptocephalus Geoffroy, 1762: 231. Current status: valid genus in CHRYSOMELIDAE: CRYPTOCEPHALINAE: CRYPTOCEPHALINI: CRYPTOCEPHALINA. Note: name placed on the Official List of Generic Names in Zoology (ICZN 1994a).

Type species: Chrysomela sericea Linnaeus, 1758, by subsequent designation (Latreille 1810: 432; see ICZN 1994a). Note: name placed on the Official List of Specific Names in Zoology (ICZN 1994a).

Family-group name: CRYPTOCEPHALOIDEAE Gyllenhal, 1813. Stem: Cryptocephal-.

Cryptochile Latreille, 1828a: 576. Current status: valid genus in TENEBRIONIDAE: PIMELIINAE: CRYPTOCHILINI: CRYPTOCHILINA. Note: the older but unusued name Phymatium Billberg, 1820 has priority over Cryptochile Latreille, 1828; an application to the Commission is needed to conserve usage of the well-established name Cryptochile (see Bouchard et al. 2021: 152).

Type species: Pimelia maculata Fabricius, 1781, by monotypy.

Family-group name: CRYPTOCHILITES Solier, 1841. Stem: Cryptochil-. Note: senior homonym of CRYPTOCHILIDAE Berger, 1979 (type genus Cryptochilum Maupas, 1883) in Ciliophora.

Cryptodacne Sharp, 1878c: 82. Current status: valid genus in EROTYLIDAE: EROTYLINAE: DACNINI.

Type species: Cryptodacne synthetica Sharp, 1878, by monotypy.

Family-group name: CRYPTODACNINI Sen Gupta, 1970. Stem: Cryptodacn-.

Cryptoderma Ritsema, 1885: 54. Current status: valid genus in CURCULIONIDAE: DRYOPHTHORINAE: CRYPTODERMATINI. Note: new replacement name for Oxyrhynchus Schönherr, 1826; Nosoxylon Gistel, 1848 is an older name for Cryptoderma Ritsema, 1885 and an application to the Commission or usage of Reversal of Precedence (ICZN 1999a, Article 23.9) is necessary to retain Ritsema’s name as valid; Alonso-Zarazaga and Lyal (1999: 63) mentioned that they would submit an application to the Commission to suppress the older name Nosoxylon Gistel, 1848 but this has not taken place to our knowledge.

Type species: Calandra discors Fabricius, 1801, by original designation (type fixation for Oxyrhynchus Schönherr).

Family-group name: CRYPTODERMINAE Bovie, 1908. Stem: Cryptodermat-.

Cryptodontes Burmeister, 1847: 292. Current status: valid genus in SCARABAEIDAE: CETONIINAE: CRYPTODONTINI.

Type species: Leptognathus latreillianus Westwood, 1841, by monotypy.

Family-group name: CRYPTODONTIDES Lacordaire, 1855. Stem: Cryptodont-. Note: senior homonym of CRYPTODONTIDAE Dall, 1895 (type genus Cryptodon Turton, 1822) in Mollusca: Bivalvia.

Cryptodus W.S. MacLeay, 1819: 138. Current status: valid genus in SCARABAEIDAE: DYNASTINAE: PHILEURINI: CRYPTODINA.

Type species: Cryptodus paradoxus W.S. MacLeay, 1819, by monotypy.

Family-group name: CRYPTODINAE Burmeister & Schaum, 1840. Stem: Cryptod-.

Cryptogenius Westwood, 1842a: 457. Current status: valid genus in HYBOSORIDAE: ANAIDINAE. Note: see additional information on the date of publication and page number for the establishment of this genus-group name in the entry for Westwood (1842a) in the References section, which differs slightly from data recently provided in Costa-Silva and Vaz-de-Mello (2023: 178).

Type species: Cryptogenius miersianus Westwood, 1842, by monotypy.

Family-group name: CRYPTOGENIINI Howden, 2001. Stem: Cryptogeni-.

Cryptoglossa Solier, 1837: 638, 680. Current status: valid genus in TENEBRIONIDAE: PIMELIINAE: CRYPTOGLOSSINI.

Type species: Cryptoglossa bicostata Solier, 1837, by monotypy.

Family-group name: CRYPTOGLOSSINI J.L. LeConte, 1862. Stem: Cryptogloss-.

Cryptognatha Mulsant, 1850: 497. Current status: valid genus in COCCINELLIDAE: COCCINELLINAE: CRYPTOGNATHINI.

Type species: Cryptognatha auriculata Mulsant, 1850, by subsequent designation (Crotch 1874a: 206).

Family-group name: CRYPTOGNATHAIRES Mulsant, 1850. Stem: Cryptognath-. Note: senior homonym of CRYPTOGNATHIDAE Oudemans, 1902 (type genus Cryptognathus Kramer, 1879) in Acari: Trombidiformes.

Cryptohypnus Eschscholtz, 1830b: 17. Current status: junior synonym of Hypnoidus Dillwyn, 1829 in ELATERIDAE: HYPNOIDINAE.

Type species: Elater riparius Fabricius, 1792, by subsequent designation (Duponchel 1844a: 432).

Family-group name: CRYPTOHYPNITES Candèze, 1860. Stem: Cryptohypn-.

Cryptolarynx Schalkwyk, 1966: 745. Current status: valid genus in CURCULIONIDAE: BRACHYCERINAE: CRYPTOLARYNGINI. Note: new replacement name for Cryptopharynx G.A.K. Marshall, 1957.

Type species: Cryptopharynx vitis G.A.K. Marshall, 1957, by original designation (type fixation for Cryptopharynx G.A.K. Marshall).

Family-group name: CRYPTOLARYNGINAE Schalkwyk, 1966. Stem: Cryptolaryng-.

Cryptomera Broun, 1893b: 1358. Current status: junior synonym of Eucinetus Germar, 1818 in EUCINETIDAE. Note: junior homonym of Cryptomera Rafinesque 1820 [Chilopoda].

Type species: Cryptomera nigra Broun, 1893, by monotypy.

Family-group name: CRYPTOMERIDAE Broun, 1893. Stem: Cryptomer-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

Cryptommata Wollaston, 1877: 156. Current status: valid genus in CURCULIONIDAE: COSSONINAE: CRYPTOMMATINI.

Type species: Cryptommata cucullata Wollaston, 1877, by monotypy.

Family-group name: CRYPTOMMATINI Voss, 1972. Stem: Cryptommat-.

Cryptonotopsis Pace, 2003: 39. Current status: valid genus in STAPHYLINIDAE: ALEOCHARINAE: CRYPTONOTOPSISINI.

Type species: Cryptonotopsis rougemonti Pace, 2003, by original designation.

Family-group name: CRYPTONOTOPSISINI Pace, 2003. Stem: Cryptonotopsis- (ICZN 1999a, Article 29.4).

Cryptonychus Gyllenhal in Schönherr, 1817: 7. Current status: valid genus in CHRYSOMELIDAE: CASSIDINAE: CRYPTONYCHINI.

Type species: Hispa porrecta Gyllenhal, 1817, by monotypy.

Family-group name: CRYPTONYCHITES Chapuis, 1875. Stem: Cryptonych-.

Cryptophagus Herbst, 1792: 172. Current status: valid genus in CRYPTOPHAGIDAE: CRYPTOPHAGINAE: CRYPTOPHAGINI. Note: the original spelling Kryptophagus was changed to Cryptophagus and Cryptophagus Herbst, 1792 placed on the Official List of Generic Names in Zoology (ICZN 1995c).

Type species: Dermestes cellaris Scopoli, 1763, by subsequent designation (Westwood 1838c: 13; see ICZN 1995c). Note: name placed on the Official List of Specific Names in Zoology (ICZN 1995c).

Family-group name: CRYPTOPHAGIDAE W. Kirby, 1826. Stem: Cryptophag-.

Cryptopharynx G.A.K. Marshall, 1957: 17. Current status: invalid senior synonym of Cryptolarynx Schalkwyk, 1966 in CURCULIONIDAE: BRACHYCERINAE: CRYPTOLARYNGINI. Note: junior homonym of Cryptopharynx Kahl, 1928 [Protozoa].

Type species: Cryptopharynx vitis G.A.K. Marshall, 1957, by original designation.

Family-group name: CRYPTOPHARYNGINAE G.A.K. Marshall, 1957. Stem: Cryptopharyng-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

Cryptophilus Reitter, 1875a: 381. Current status: valid genus in EROTYLIDAE: CRYPTOPHILINAE: CRYPTOPHILINI.

Type species: fixed by Gimmel et al. (2019a: 529) according to Article 70.3.2 (ICZN 1999a) as Typhaea angusta Rosenhauer, 1856, the taxonomic species involved in the misidentification of Cryptophagus integer Heer, 1841 (sensu Reitter, 1875), as subsequently designated by Chûjô (1969: 277).

Family-group name: CRYPTOPHILINI Casey, 1900. Stem: Cryptophil-.

Cryptoplus Erichson, 1842: 198. Current status: valid genus in CURCULIONIDAE: CURCULIONINAE: CRYPTOPLINI.

Type species: Cryptoplus perdix Erichson, 1842, by monotypy.

Family-group name: CRYPTOPLIDES Lacordaire, 1863. Stem: Cryptopl-.

Cryptops Wiedemann, 1823b: 164. Current status: invalid senior synonym of Byrsops Germar, 1829 in CURCULIONIDAE: BRACHYCERINAE: BRACHYCERINI. Note: junior homonym of Cryptops Leach, 1814 [Chilopoda]; Alonso-Zarazaga and Lyall (1999: 62) consider the type genus to have been proposed independently also by Schönherr (1823: 1138) but we consider that the two names apply to the same taxon.

Type species: Brachycerus quadratus Wiedemann, 1821, by subsequent designation (Schönherr 1823: 1138).

Family-group name: CRYPTOPSIDES Schönherr, 1826. Stem: Cryptop-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

Cryptorhynchidius Pierce, 1919: 25. Current status: junior synonym of Cryptorhynchus Illiger, 1807 in CURCULIONIDAE: CRYPTORHYNCHINAE: CRYPTORHYNCHINI: CRYPTORHYNCHINA.

Type species: Curculio lapathi Linnaeus, 1758, by monotypy.

Family-group name: CRYPTORHYNCHIDIINI Bradley, 1930. Stem: Cryptorhynchidi-.

Cryptorhynchus Illiger, 1807b: 330. Current status: valid genus in CURCULIONIDAE: CRYPTORHYNCHINAE: CRYPTORHYNCHINI: CRYPTORHYNCHINA. Note: name placed on the Official List of Generic Names in Zoology (ICZN 1967b).

Type species: Curculio lapathi Linnaeus, 1758, by plenary power (see ICZN 1967b). Note: name placed on the Official List of Specific Names in Zoology (ICZN 1967b).

Family-group name: CRYPTORHYNCHIDES Schönherr, 1825. Stem: Cryptorhynch-.

Cryptosomatula Bruce, 1940: 683. Current status: valid genus in CRYPTOPHAGIDAE: CRYPTOPHAGINAE: PICROTINI.

Type species: Cryptosomatula longicornis Bruce, 1940, by monotypy.

Family-group name: CRYPTOSOMATULINI Leschen, 1996. Stem: Cryptosomatul-.

Cryptostoma Dejean, 1821: 34. Current status: invalid senior synonym of Ceratogonys Perty, 1830 in EUCNEMIDAE: MACRAULACINAE: ORODOTINI. Note: junior homonym of Cryptostoma de Blainville, 1818 [Mollusca].

Type species: Elater spinicornis Fabricius, 1801, by monotypy.

Family-group name: CRYPTOSTOMIDAE Laporte, 1835. Stem: Cryptostomat-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

Crypturgus Erichson, 1836: 60. Current status: valid genus in CURCULIONIDAE: SCOLYTINAE: CRYPTURGINI.

Type species: Bostrichus cinereus Herbst, 1794, by subsequent designation (Laporte 1840b: 370). Note: the type species usually listed for this genus is Bostrichus pusillus Gyllenhal, 1813 (e.g., Alonso-Zarazaga and Lyal 2009: 48; Knížek 2011: 227) by subsequent designation of C.G. Thomson (1859: 147) but Laporte’s designation is earlier; both species are currently included in the genus Crypturgus Erichson, 1836.

Family-group name: CRYPTURGI J.L. LeConte, 1876. Stem: Crypturg-.

Ctenicera Latreille, 1829a: 454. Current status: valid genus in ELATERIDAE: DENDROMETRINAE: PROSTERNINI.

Type species: Elater pectinicornis Linnaeus, 1758, by subsequent designation (Westwood 1838c: 26, as Ctenicerus).

Family-group name: CTENICERINA Jakobson, 1913. Stem: Ctenicer-.

Ctenidia Laporte, 1840b: 264. Current status: valid genus in MORDELLIDAE: CTENIDIINAE.

Type species: Ctenidia mordelloides Laporte, 1840, by monotypy.

Family-group name: CTENIDIINAE Franciscolo, 1951. Stem: Ctenidi-.

Cteniopus Solier, 1835b: 245, 246. Current status: valid genus in TENEBRIONIDAE: ALLECULINAE: CTENIOPODINI.

Type species: Chrysomela sulphurea Linnaeus, 1758, by subsequent designation (Westwood 1838c: 32).

Family-group name: CTÉNIOPITES Solier, 1835. Stem: Cteniopod-.

Ctenistes H.G.L. Reichenbach, 1816: 75. Current status: valid genus in STAPHYLINIDAE: PSELAPHINAE: PSELAPHITAE: CTENISTINI.

Type species: Ctenistes palpalis H.G.L. Reichenbach, 1816, by monotypy.

Family-group name: CTÉNISTITES Blanchard, 1845. Stem: Ctenist-.

Ctenodactyla Dejean, 1825: 169, 226. Current status: valid genus in CARABIDAE: HARPALINAE: CTENODACTYLINI.

Type species: Ctenodactyla chevrolatii Dejean, 1825, by monotypy.

Family-group name: CTENODACTYLIDAE Laporte, 1834. Stem: Ctenodactyl-. Note: senior homonym of CTENODACTYLINA C. Girard, 1851 (type genus Ctenodactylus J.E. Gray, 1830) in Mammalia.

Ctenophorus Chapuis, 1869: 49. Current status: junior synonym of Scolytodes Ferrari, 1867 in CURCULIONIDAE: SCOLYTINAE: HEXACOLINI. Note: junior homonym of Ctenophorus Fitzinger, 1843 [Reptilia].

Type species: Ctenophorus laevigatus Chapuis, 1869 (= Scolytodes chapuisi Wood, 1977), by monotypy.

Family-group name: CTENOPHORIDAE Chapuis, 1869. Stem: Ctenophor-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

Ctenoscelis Audinet-Serville, 1832: 134. Current status: valid genus in CERAMBYCIDAE: PRIONINAE: CALLIPOGONINI.

Type species: Prionus ater G.-A. Olivier, 1795, by subsequent designation (J. Thomson 1864: 297).

Family-group name: CTENOSCELITAE J. Thomson, 1864. Stem: Ctenoscelid-.

Ctenostoma Klug, 1821: 302. Current status: valid genus in CICINDELIDAE: CTENOSTOMATINI.

Type species: here fixed (under ICZN 1999a, Article 70.3.2) as Ctenostoma jekelii Chevrolat, 1858, misidentified as Collyris formicaria Fabricius, 1801 (sensu Klug, 1821) in the original designation by monotypy in Klug (1821: 302).

Family-group name: CTENOSTOMIDAE Laporte, 1834. Stem: Ctenostomat-.

Ctesias Stephens, 1830: 124. Current status: valid genus in DERMESTIDAE: MEGATOMINAE: CTESIINI.

Type species: Dermestes serra Fabricius, 1792, by monotypy.

Family-group name: CTESIINI Rees, 1943. Stem: Ctesi-.

Ctesibius Champion, 1897: 592. Current status: valid genus in ARTEMATOPODIDAE: ARTEMATOPODINAE: CTESIBIINI.

Type species: Ctesibius eumolpoides Champion, 1897, by monotypy.

Family-group name: CTESIBIINAE Crowson, 1973. Stem: Ctesibi-.

Cubispa Barber, 1946: 21. Current status: valid genus in CHRYSOMELIDAE: EUMOLPINAE: CUBISPINI.

Type species: Cubispa turquino Barber, 1946, by original designation.

Family-group name: CUBISPINI Monrós, 1954. Stem: Cubisp-.

Cucujus Fabricius, 1775: 204. Current status: valid genus in CUCUJIDAE: CUCUJINAE. Note: the senior homonym Cucujus Geoffroy, 1762 (and all other uses of the name prior to the publication of Cucujus Fabricius, 1775) was suppressed for both Principles of Priority and Homonymy and placed on the Official Index of Rejected and Invalid Generic Names in Zoology while Cucujus Fabricius, 1775 was placed on the Official List of Generic Names in Zoology (ICZN 1994a).

Type species: Cucujus depressus Fabricius, 1775 (= Meloe cinnaberinus Scopoli, 1763), by monotypy (see ICZN 1994a). Note: Meloe cinnaberinus Scopoli, 1763 was placed on the Official List of Specific Names in Zoology (ICZN 1994a).

Family-group name: CUCUJIPES Latreille, 1802. Stem: Cucuj-.

Cuneipectus Sloane, 1907: 358. Current status: valid genus in CARABIDAE: HARPALINAE: CUNEIPECTINI.

Type species: Cuneipectus frenchi Sloane, 1907, by monotypy.

Family-group name: CUNEIPECTINI Sloane, 1907. Stem: Cuneipect-.

Cupes Fabricius, 1801a: xvii. Current status: valid genus in CUPEDIDAE: CUPEDINAE.

Type species: Cupes capitata Fabricius, 1801, by subsequent monotypy (Fabricius 1801b: 60).

Family-group name: CUPESIDAE Laporte, 1838. Stem: Cuped-.

Curculio Linnaeus, 1758: 342, 377. Current status: valid genus in CURCULIONIDAE: CURCULIONINAE: CURCULIONINI: CURCULIONINA.

Type species: Curculio nucum Linnaeus, 1758, by subsequent designation (Latreille 1810: 430).

Family-group name: CURCULIONITES Latreille, 1802. Stem: Curculion-.

†Curculiopsis Martynov, 1937: 39. Current status: invalid senior synonym of Rossocoleus Rohdendorf, 1961 in †SCHIZOCOLEIDAE. Note: junior homonym of Curculiopsis Handlirsch, 1907 [Coleoptera: CURCULIONOIDEA]; Permian (Russia).

Type species: †Curculiopsis ellipticus Martynov, 1937, by original designation.

Family-group name: †CURCULIOPSIDAE Martynov, 1937. Stem: Curculiopse-. Note: family- group name permanently invalid (ICZN 1999a, Article 39).

Curis Laporte & Gory, 1837: 47. Current status: junior synonym of Selagis Dejean, 1836 in BUPRESTIDAE: BUPRESTINAE: CURIDINI: CURIDINA.

Type species: Stigmodera peroni Laporte & Gory, 1837, by subsequent designation (Rye 1873: 271). Note: the type species usually recognized for this genus is Buprestis caloptera Boisduval, 1835, by subsequent designation of Bellamy (1997a: 376) but Rye’s designation is earlier; both species are currently included in the genus Curis.

Family-group name: CURIDINA Holyński, 1988. Stem: Curid-.

Curius Newman, 1840d: 17. Current status: valid genus in CERAMBYCIDAE: CERAMBYCINAE: CURIINI.

Type species: Curius dentatus Newman, 1840, by monotypy.

Family-group name: CURII J.L. LeConte, 1873. Stem: Curi-.

Curtimorda Méquignon, 1946: 58. Current status: valid genus in MORDELLIDAE: MORDELLINAE: CURTIMORDINI.

Type species: Mordella bisignata L. Redtenbacher, 1849, by original designation.

Family-group name: CURTIMORDINI Odnosum, 2010. Stem: Curtimord-.

Curtos Motschulsky, 1845b: 36. Current status: valid genus in LAMPYRIDAE: LUCIOLINAE: CURTOSINI.

Type species: Curtos mongolicus Motschulsky, 1845, by monotypy.

Family-group name: CURTOSINI McDermott, 1964. Stem: Curtos-.

Cussolenis Fleutiaux, 1918a: 59. Current status: valid genus in ELATERIDAE: THYLACOSTERNINAE. Note: new replacement name for Soleniscus Bonvouloir, 1871.

Type species: Soleniscus mutabilis Bonvouloir, 1875, by subsequent monotypy (de Bonvouloir 1875: 828; type fixation for Soleniscus Bonvouloir).

Family-group name: CUSSOLENITAE Cobos, 1961. Stem: Cussolen-.

Cyathiger King, 1865: 174. Current status: junior synonym of Plagiophorus Motschulsky, 1851 in STAPHYLINIDAE: PSELAPHINAE: GONIACERITAE: CYATHIGERINI.

Type species: Cyathiger punctatus King, 1865, by monotypy.

Family-group name: CYATHIGERINI L.W. Schaufuss, 1872. Stem: Cyathiger-.

Cyathocerus Sharp, 1882c: 142. Current status: junior synonym of Lepicerus Motschulsky, 1855 in LEPICERIDAE.

Type species: Cyathocerus horni Sharp, 1882 (= Lepicerus inaequalis Motschulsky, 1855), by monotypy.

Family-group name: CYATHOCERIDAE Sharp, 1882. Stem: Cyathocer-.

Cybebus Schönherr, 1839: 447. Current status: valid genus in BRENTIDAE: APIONINAE: CYBEBITAE.

Type species: Cybebus rufipennis Boheman, 1839 (= Curculio dimidiatus G.-A. Olivier, 1791), by original designation.

Family-group name: CYBÉBIDES Lacordaire, 1863. Stem: Cybeb-.

Cybister Curtis, 1827: pl. 151. Current status: valid genus in DYTISCIDAE: CYBISTERINAE. Note: new replacement name for Trogus Leach, 1817.

Type species: Dytiscus lateralis Fabricius, 1798 (= Dytiscus tripunctatus G.-A. Olivier, 1795), by monotypy (type fixation for Trogus Leach).

Family-group name: CYBISTRINI Sharp, 1880. Stem: Cybister-.

Cybocephalus Erichson, 1844: 441. Current status: valid genus in NITIDULIDAE: CYBOCEPHALINAE.

Type species: Anisotoma exigua C.R. Sahlberg, 1833 (= Nitidula polita Gyllenhal, 1813), by subsequent designation (E. Desmarest 1851: 278).

Family-group name: CYBOCÉPHALITES Jacquelin du Val, 1858. Stem: Cybocephal-.

Cychramptodes Reitter, 1878b: 383. Current status: valid genus in NITIDULIDAE: NITIDULINAE: CYCHRAMPTODINI.

Type species: Cychramptodes murrayi Reitter, 1878, by monotypy.

Family-group name: CYCHRAMPTODINI Kirejtshuk & Lawrence, 1992. Stem: Cychramptod-.

Cychramus Kugelann, 1794: 543. Current status: valid genus in NITIDULIDAE: NITIDULINAE: CYCHRAMINI.

Type species: Sphaeridium luteum Fabricius, 1787, by subsequent designation (Duponchel 1844a: 486). Note: the type species usually recognized for Cychramus Kugelann, 1794 is Strongylus quadripunctatus Herbst, 1792 (e.g., Jelínek and Audisio 2003: 161; Jelínek and Audisio 2007: 471) by subsequent designation of C.G. Thomson (1859: 69) but Duponchel’s designation is earlier; both luteus (Fabricius) and quadripunctatus (Herbst) are currently included in the genus Cychramus.

Family-group name: CYCHRAMIDAE Gistel, 1848. Stem: Cychram-.

Cychrus Fabricius, 1794: 440. Current status: valid genus in CARABIDAE: CARABINAE: CYCHRINI. Note: this name was proposed by Fabricius in 1794 under two spellings in two different publications, as Cychrus in Tome 4 of his ‘Entomologia systematica’, issued by 5 October 1794 (see Bousquet 2017a: 107), and as Cychrys in Volume 3 of the journal ‘Skrivter af Naturhistorie-Selskabet’, for which only the year of publication is currently known; therefore, Cychrus is the correct original spelling and the one in use today.

Type species: Tenebrio rostratus Linnaeus, 1761 (= Tenebrio caraboides Linnaeus, 1758), by subsequent designation (Latreille 1810: 426).

Family-group name: CYCHRII Perty, 1830. Stem: Cychr-.

Cyclaxyra Broun, 1893b: 1076. Current status: valid genus in CYCLAXYRIDAE. Note: new replacement name for Cyclomorpha Broun, 1881; name placed on the Official List of Generic Names in Zoology (ICZN 1988a).

Type species: Cyclomorpha politula Broun, 1881, by monotypy (see ICZN 1988a) (type fixation for Cyclomorpha Broun). Note: name placed on the Official List of Specific Names in Zoology (ICZN 1988a).

Family-group name: CYCLAXYRIDAE Gimmel, Leschen & Ślipiński, 2009. Stem: Cyclaxyr-.

Cyclocephala Dejean, 1821: 57. Current status: valid genus in SCARABAEIDAE: DYNASTINAE: CYCLOCEPHALINI.

Type species [suggested]: Melolontha signata Fabricius, 1781, by subsequent designation (Casey 1915: 112). Note: the valid type species fixation is that of Blanchard in Audouin et al. (1843: pl. 41), who selected Melolontha geminata Fabricius, 1801, but that nominal species is currently placed in the genus Dyscinetus Harold, 1869; an application to the Commission is necessary to fix Melolontha signata Fabricius, 1781 as the type species of Cyclocephala Dejean, 1821.

Family-group name: CYCLOCEPHALITES Laporte, 1840. Stem: Cyclocephal-.

Cycloderes C.R. Sahlberg, 1823a: 21. Current status: valid genus in CURCULIONIDAE: ENTIMINAE: POLYDRUSITAE: TANYMECINI: TANYMECINA. Note: name placed on the Official List of Generic Names in Zoology (ICZN 1987a).

Type species: Cycloderes catarractus C.R. Sahlberg, 1823 (= Curculio mus Herbst, 1797), by monotypy (see ICZN 1987a). Note: combined description of a new genus-group taxon and a single new species (ICZN 1999a, Article 12.2.6); Cycloderes catarractus C.R. Sahlberg, 1823 was placed on the Official List of Specific Names in Zoology (ICZN 1987a).

Family-group name: CYCLODERINI Hoffmann, 1950. Stem: Cycloder-.

Cyclommatus Parry, 1863: 449. Current status: valid genus in LUCANIDAE: LUCANINAE: CYCLOMMATINI. Note: new replacement name for Megaloprepes J. Thomson, 1862.

Type species: Lucanus tarandus Thunberg, 1806, by original designation (type fixation for Megaloprepes J. Thomson).

Family-group name: CYCLOMMATINI H. Huang & C.H. Chen, 2013: 70. Stem: Cyclommat-.

Cyclomus Schönherr, 1826: 198. Current status: junior synonym of Epicthonius Schönherr, 1823 in CURCULIONIDAE: CYCLOMINAE: CYCLOMINI.

Type species: Curculio simus Wiedemann, 1823, by monotypy.

Family-group name: CYCLOMIDES Schönherr, 1826. Stem: Cyclom-.

Cyclonotum Erichson, 1837: 212, 214. Current status: junior synonym of Coelostoma Brullé, 1835 in HYDROPHILIDAE: SPHAERIDIINAE: COELOSTOMATINI. Note: this genus-group name was first made available by Dejean (1833b: 134) with Sphaeridium abdominale Fabricius, 1792 as type species by monotypy, a species currently placed in the genus Dactylosternum Wollaston, 1854; a request should be submitted to the Commission to suppress Dejean’s name for the purposes of both the Principles of Priority and Homonymy (see Bousquet and Bouchard 2013a: 30) in order to conserve stability.

Type species: Hydrophilus orbicularis Fabricius, 1775, by subsequent designation (Westwood 1838c: 10).

Family-group name: CYCLONOTIDES G.H. Horn, 1890. Stem: Cyclonot-.

Cyclopterus Marseul, 1872: 30. Current status: invalid senior synonym of Nucterocephalus Desbrochers des Loges, 1897 in CURCULIONIDAE: ENTIMINAE: OTIORHYNCHITAE: HOLCORHININI. Note: junior homonym of Cyclopterus Linnaeus, 1758 [Pisces].

Type species: Cyclopterus spinifer Marseul, 1872, by monotypy.

Family-group name: CYCLOPTERINI Reitter, 1913. Stem: Cyclopter-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

Cyclosomus Latreille, 1829a: 394. Current status: valid genus in CARABIDAE: HARPALINAE: CYCLOSOMINI.

Type species: Carabus flexuosus Fabricius, 1775, by monotypy.

Family-group name: CYCLOSOMIDAE Laporte, 1834. Stem: Cyclosom-.

Cycloteres Schönherr, 1843: 383. Current status: valid genus in CURCULIONIDAE: MOLYTINAE: CYCLOTERINI: CYCLOTERINA.

Type species: Cycloteres audouini Boheman, 1843, by original designation.

Family-group name: CYCLOTÉRIDES Lacordaire, 1863. Stem: Cycloter-.

Cyclotoma Mulsant, 1851a: 71. Current status: valid genus in ENDOMYCHIDAE: CYCLOTOMINAE.

Type species: Cyclotoma testudinaria Mulsant, 1851, by monotypy.

Family-group name: CYCLOTOMIDAE Imhoff, 1856. Stem: Cyclotom-.

Cycloxenus Arrow, 1925: 152. Current status: valid genus in EUXESTIDAE.

Type species: Cycloxenus hispidus Arrow, 1925, by original designation.

Family-group name: CYCLOXENINI Jeannel & Paulian, 1945. Stem: Cyclo­xen-.

Cycnotrachelus Jekel, 1860: 160. Current status: valid genus in ATTELABIDAE: APODERINAE: APODERINI.

Type species: Apoderus flavotuberosus Jekel, 1860, by subsequent designation (Voss 1925: 5).

Family-group name: CYCNOTRACHELINA Legalov, 2003. Stem: Cycnotrachel-.

Cydistus Bourgeois, 1885: xxxvii. Current status: valid genus in PHENGODIDAE: CYDISTINAE.

Type species: Cydistus reitteri Bourgeois, 1885, by monotypy.

Family-group name: CYDISTINAE Paulus, 1972. Stem: Cydist-.

Cydonia Mulsant, 1850: 429, 430. Current status: junior synonym of Cheilomenes Chevrolat, 1836 in COCCINELLIDAE: COCCINELLINAE: COCCINELLINI.

Type species: Cydonia quadrilineata Mulsant, 1850 (= Cydonia propinqua Mulsant, 1850), by subsequent designation (Casey 1899: 84).

Family-group name: CYDONIAIRES Mulsant, 1850. Stem: Cydoni-.

Cylas Latreille, 1802b: 196. Current status: valid genus in BRENTIDAE: BRENTINAE: CYLADINI: CYLADINA.

Type species: Brentus brunneus G.-A. Olivier, 1790, by monotypy.

Family-group name: CYLADES Schönherr, 1823. Stem: Cylad-.

Cylidrus Latreille, 1817a: 43. Current status: valid genus in CLERIDAE: TILLINAE. Note: this name is usually credited to Latreille (1829a: 476) in the literature but it was first made available by Latreille in 1817 (see Bouchard and Bousquet 2020a: 92).

Type species: Clerus cyaneus Fabricius, 1787, by monotypy.

Family-group name: CYLIDRINA Reitter, 1894. Stem: Cylidr-.

Cylindrinotus Faldermann, 1837: 73. Current status: valid genus in TENEBRIONIDAE: TENEBRIONINAE: HELOPINI: CYLINDRINOTINA.

Type species: Cylindrinotus lugubris Faldermann, 1837 (= Helops femoratus Faldermann, 1837), by subsequent designation (Gebien 1943: 425). Note: Gebien (1943: 425) selected Helops femoratus Faldermann, 1837 as type species, a species not originally included, but since he listed it in synonymy with Cylindrinotus lugubris Faldermann, 1837, a species originally included, he is deemed to have designated the latter species as type (ICZN 1999a, Article 69.2.2).

Family-group name: CYLINDRONOTINI Español, 1956. Stem: Cylindrinot-.

†Cylindrobrotus Kirejtshuk, Azar, Beaver, Mandelshtam & Nel, 2009: 105. Current status: valid genus in CURCULIONIDAE: SCOLYTINAE: †CYLINDROBROTINI. Note: Lower Cretaceous (Lebanese amber).

Type species: †Cylindrobrotus pectinatus Kirejtshuk, Azar, Beaver, Mandelshtam & Nel, 2009, by original designation.

Family-group name: †CYLINDROBROTINI Kirejtshuk, Azar, Beaver, Mandelshtam & Nel, 2009. Stem: Cylindrobrot-.

Cylindrocaulus Fairmaire, 1880: 164. Current status: valid genus in PASSALIDAE: AULACOCYCLINAE: CYLINDROCAULINI.

Type species: Cylindrocaulus bucerus Fairmaire, 1880, by monotypy.

Family-group name: CYLINDROCAULINAE Zang, 1905. Stem: Cylindrocaul-.

Cylindrocopturus Heller, 1895: 4, 56. Current status: valid genus in CURCULIONIDAE: CONODERINAE: ZYGOPINI.

Type species: Zygops quercus Say, 1832, by subsequent designation (Sleeper 1963: 217).

Family-group name: CYLINDROCOPTURINI Böving, 1927. Stem: Cylindrocoptur-.

Cylindrocorynus Schönherr, 1837: 231. Current status: valid genus in CURCULIONIDAE: CRYPTORHYNCHINAE: CRYPTORHYNCHINI: MECISTOSTYLINA.

Type species: Cylindrocorynus imaginarius Boheman, 1837, by original designation.

Family-group name: CYLINDROCORYNINAE Pascoe, 1875. Stem: Cylindrocoryn-.

Cylindromorphoides Kerremans, 1903c: 301. Current status: valid genus in BUPRESTIDAE: AGRILINAE: APHANISTICINI: CYLINDROMORPHOIDINA.

Type species: Taphrocerus agriliformis Kerremans, 1897, by monotypy.

Family-group name: CYLINDROMORPHOIDINI Cobos, 1979. Stem: Cylindromorphoid-.

Cylindromorphus Kiesenwetter, 1856: 159. Current status: valid genus in BUPRESTIDAE: AGRILINAE: APHANISTICINI: CYLINDROMORPHINA.

Type species: Buprestis filum Gyllenhal, 1817, by subsequent designation (E. Desmarest 1860: 42).

Family-group name: CYLINDROMORPHINI Portevin, 1931. Stem: Cylindromorph-.

Cylindroxystus Bierig, 1943: 158. Current status: valid genus in STAPHYLINIDAE: PAEDERINAE: LATHROBIINI: CYLINDROXYSTINA.

Type species: Cylindroxystus longulus Bierig, 1943, by original designation.

Family-group name: CYLINDROXYSTINI Bierig, 1943. Stem: Cylindroxyst-.

Cyllene Newman, 1840c: 7. Current status: invalid senior synonym of Megacyllene Casey, 1912 in CERAMBYCIDAE: CERAMBYCINAE: CLYTINI. Note: junior homonym of Cyllene J.E. Gray, 1834 [Mollusca].

Type species: Cyllene spinifera Newman, 1840 (= Clytus nebulosus Laporte & Gory, 1838), by monotypy.

Family-group name: CYLLENITAE J. Thomson, 1864. Stem: Cyllen-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

Cyllodes Erichson, 1843a: 342. Current status: valid genus in NITIDULIDAE: NITIDULINAE: CYLLODINI.

Type species: Strongylus ater Herbst, 1792, by subsequent designation (Duponchel 1844a: 515).

Family-group name: CYLLODINI Everts, 1898. Stem: Cyllod-.

Cyloma Sharp, 1872: 152. Current status: valid genus in HYDROPHILIDAE: CYLOMINAE.

Type species: Cyloma lawsonus Sharp, 1872, by monotypy.

Family-group name: CYLLOMINA Zaitzev, 1908. Stem: Cylom-.

Cylydrorhinus Guérin-Méneville, 1838: 119. Current status: valid genus in CURCULIONIDAE: ENTIMINAE: CYLYDRORHINITAE: CYLYDRORHININI.

Type species: Curculio lemniscatus Quoy & Gaimard, 1825, by monotypy.

Family-group name: CYLINDRORHINIDES Lacordaire, 1863. Stem: Cylydrorhin-.

Cymatopterus Dejean, 1833a: 54. Current status: junior synonym of Colymbetes Clairville, 1806 in DYTISCIDAE: COLYMBETINAE.

Type species: Dytiscus fuscus Linnaeus, 1758, by subsequent designation (C.G. Thomson 1859: 13).

Family-group name: CYMATOPTERINI Portevin, 1929. Stem: Cymatopter-.

Cymbionotum Baudi di Selve, 1864: 211. Current status: valid genus in CARABIDAE: MELAENINAE.

Type species: Cymbionotum collare Baudi di Selve, 1864 (= Coscinia semelederi Chaudoir, 1861), by monotypy.

Family-group name: CYMBIONOTINI Andrewes, 1933. Stem: Cymbionot-.

Cymindis Latreille, 1805: 190, 228. Current status: valid genus in CARABIDAE: HARPALINAE: LEBIINI: CYMINDIDINA.

Type species: Buprestis humeralis Geoffroy, 1785, by monotypy.

Family-group name: CYMINDIDAE Laporte, 1834. Stem: Cymindid-.

Cymophorus W. Kirby, 1827: 153. Current status: valid genus in SCARABAEIDAE: CETONIINAE: CREMASTOCHEILINI: CYMOPHORINA.

Type species: Cymophorus undatus W. Kirby, 1827, by monotypy.

Family-group name: CYMOPHORINA Krikken, 1984. Stem: Cymophor-.

Cynegetis Chevrolat in Dejean, 1836a: 437. Current status: valid genus in COCCINELLIDAE: COCCINELLINAE: EPILACHNINI.

Type species: Coccinella impunctata Linnaeus, 1767, by subsequent designation (C.G. Thomson 1859: 160).

Family-group name: CYNEGETIDES C.G. Thomson, 1866. Stem: Cyneget-.

Cyno T.A. Marshall, 1865: 350. Current status: valid genus in CHRYSOMELIDAE: EUMOLPINAE: BROMIINI.

Type species: Cyno mordicans T.A. Marshall, 1865, by monotypy.

Family-group name: CYNOINI Clavareau, 1914. Stem: Cynoi-.

Cyparella Achard, 1924: 28. Current status: junior synonym of Baeocera Erichson, 1845 in STAPHYLINIDAE: SCAPHIDIINAE: SCAPHISOMATINI.

Type species: Scaphisoma rufoguttatum Fairmaire, 1898, by original designation.

Family-group name: CYPARELLINI Achard, 1924. Stem: Cyparell-.

Cyparium Erichson, 1845a: 3. Current status: valid genus in STAPHYLINIDAE: SCAPHIDIINAE: CYPARIINI.

Type species: Cyparium palliatum Erichson, 1845, by monotypy.

Family-group name: CYPARIINI Achard, 1924. Stem: Cypari-.

Cyphagogus Parry, 1849: 182. Current status: valid genus in BRENTIDAE: BRENTINAE: CYPHAGOGINI.

Type species: Cyphagogus westwoodii Parry, 1849, by subsequent designation (R. Lucas 1920: 219).

Family-group name: CYPHAGOGINAE Kolbe, 1892. Stem: Cyphagog-.

Cyphaleus Westwood, 1841a: 43. Current status: valid genus in TENEBRIONIDAE: TENEBRIONINAE: HELEINI: CYPHALEINA.

Type species: Helops rugosus J.E. Gray, 1831, by subsequent designation (Duponchel 1844a: 547).

Family-group name: CYPHALÉIDES Lacordaire, 1859. Stem: Cyphale-.

Cyphicerus Schönherr, 1823: 1144. Current status: valid genus in CURCULIONIDAE: ENTIMINAE: CYPHICERITAE: CYPHICERINI: CYPHICERINA.

Type species: Curculio orbitalis Wiedemann, 1819 (= Curculio novemlineatus G.-A. Olivier, 1807), by original designation.

Family-group name: CYPHICÉRIDES Lacordaire, 1863. Stem: Cyphicer-.

Cypholoba Chaudoir, 1850: 43. Current status: valid genus in CARABIDAE: HARPALINAE: ANTHIINI.

Type species: Anthia alveolata Brême, 1845, by monotypy.

Family-group name: CYPHOLOBINI G. Strohmeyer, 1928. Stem: Cypholob-.

Cyphon Paykull, 1799: 117. Current status: junior synonym of Elodes Latreille, 1797 in SCIRTIDAE: SCIRTINAE.

Type species: Cistela pallida Fabricius, 1775 (= Lampyris minuta Linnaeus, 1767), by subsequent designation (Westwood 1838c: 27).

Family-group name: CYPHONIDAE Stephens, 1829. Stem: Cyphon-.

Cyphonocerus Kiesenwetter, 1879: 311. Current status: valid genus in LAMPYRIDAE: CYPHONOCERINAE.

Type species: Cyphonocerus ruficollis Kiesenwetter, 1879, by monotypy.

Family-group name: CYPHONOCERINAE Crowson, 1972. Stem: Cyphonocer-.

Cyphosoma Hope, 1842a: 426. Current status: invalid senior synonym of Cratogaster Blanchard, 1853 in CARABIDAE: HARPALINAE: PTEROSTICHINI: PTEROSTICHINA. Note: junior homonym of Cyphosoma Mannerheim, 1837 [Coleoptera: BUPRESTIDAE].

Type species: Cyphosoma unicolor Hope, 1842, by monotypy.

Family-group name: CYPHOSOMATINI Tschitschérine, 1902. Stem: Cyphosomat-. Note: family- group name permanently invalid (ICZN 1999a, Article 39).

Cyphus Germar, 1823: 427. Current status: junior synonym of Cyrtomon Schönherr, 1823 in CURCULIONIDAE: ENTIMINAE: POLYDRUSITAE: NAUPACTINI: NAUPACTINA.

Type species: Curculio gibber Pallas, 1781, by subsequent designation (Schönherr 1826: 11).

Family-group name: CYPHIDES Lacordaire, 1863. Stem: Cyph-.

Cyrionyx Faust, 1896: 97. Current status: valid genus in CURCULIONIDAE: BARIDINAE: OPTATINI.

Type species: Cyrionyx biplagiatus Faust, 1896, by subsequent designation (Casey 1922: 10).

Family-group name: CYRIONICHINI Casey, 1922. Stem: Cyrionych-.

Cyrtinus J.L. LeConte, 1852a: 166. Current status: valid genus in CERAMBYCIDAE: LAMIINAE: CYRTININI.

Type species: Clytus pygmaeus Haldeman, 1847, by monotypy.

Family-group name: CYRTINITAE J. Thomson, 1864. Stem: Cyrtin-. Note: senior homonym of CYRTININAE Fredericks, 1912 (type genus Cyrtina Davidson, 1858) in Brachiopoda.

Cyrtoderus Hope, 1842a: 427. Current status: valid genus in CARABIDAE: HARPALINAE: PTEROSTICHINI: PTEROSTICHINA.

Type species: Cyrtoderus australasiae Hope, 1842, by monotypy.

Family-group name: CYRTODERINI Tschitschérine, 1902. Stem: Cyrtoder-.

Cyrtognathus Dejean, 1835: 316. Current status: valid genus in CERAMBYCIDAE: PRIONINAE: PRIONINI.

Type species: Prionus paradoxus Faldermann, 1833, by monotypy.

Family-group name: CYRTHOGNATHITAE J. Thomson, 1861. Stem: Cyrtognath-.

Cyrtolaus H.W. Bates, 1882: 99. Current status: valid genus in CARABIDAE: HARPALINAE: DRIMOSTOMATINI.

Type species: Cyrtolaus furculifer H.W. Bates, 1882, by subsequent designation (K.W. Kirby 1884: 32).

Family-group name: CYRTOLAINA Whitehead & Ball, 1975. Stem: Cyrtola-.

Cyrtonops A. White, 1853: 32. Current status: valid genus in DISTENIIDAE: CYRTONOPINI.

Type species: Cyrtonops punctipennis A. White, 1853, by monotypy.

Family-group name: CYRTONOPINI Gressitt, 1940. Stem: Cyrtonop-.

Cyrtoquedius Bernhauer, 1917: 92. Current status: valid genus in STAPHYLINIDAE: STAPHYLININAE: STAPHYLININI: CYRTOQUEDIINA.

Type species: Quedius basiventris Sharp, 1884, by subsequent designation (Blackwelder 1952: 115).

Family-group name: CYRTOQUEDIINA Brunke & Solodovnikov in Brunke et al., 2016: 445. Stem: Cyrtoquedi-.

Cyrtoscydmus Motschulsky, 1869b: 260. Current status: junior synonym of Stenichnus C.G. Thomson, 1859 in STAPHYLINIDAE: SCYDMAENINAE: SCYDMAENITAE: STENICHNINI.

Type species: Scydmaenus godarti Latreille, 1805, by subsequent designation (Atkinson 1891: 183).

Family-group name: CYRTOSCYDMINI L.W. Schaufuss, 1889. Stem: Cyrtoscydm-.

Cyrtotriplax Crotch, 1873a: 189. Current status: junior synonym of Tritoma Fabricius, 1775 in EROTYLIDAE: EROTYLINAE: TRITOMINI. Note: new replacement name for Tritoma Fabricius, 1775.

Type species: Tritoma bipustulata Fabricius, 1775, by subsequent designation (Latreille 1810: 432; see ICZN 1994a) (type fixation for Tritoma Fabricius).

Family-group name: CYRTOTRIPLACINA Chûjô, 1969. Stem: Cyrtotriplac-.

Cysteodemus J.L. LeConte, 1851a: 158. Current status: valid genus in MELOIDAE: MELOINAE: EUPOMPHINI.

Type species: Cysteodemus armatus J.L. LeConte, 1851, by subsequent designation (Wellman 1910a: 391).

Family-group name: CYSTEODEMIDES Wellman, 1910. Stem: Cysteodem-.

Dacne Latreille, 1797: 12. Current status: valid genus in EROTYLIDAE: EROTYLINAE: DACNINI. Note: this genus was first described without nominal species by Latreille (1797: 12).

Type species: Ips humeralis Fabricius, 1787 (= Dermestes bipustulatus Thunberg, 1781), by subsequent monotypy (Latreille 1802b: 132). Note: the type species used in recent literature (e.g., Skelley 2023: 2) is Dermestes bipustulatus Thunberg, 1781 by subsequent designation in Latreille (1810: 427).

Family-group name: DACNEIDAE Gistel, 1848. Stem: Dacn-.

Dacoderus J.L. LeConte, 1858a: 74. Current status: valid genus in SALPINGIDAE: DACODERINAE.

Type species: Dacoderus striaticeps J.L. LeConte, 1858, by monotypy.

Family-group name: DACODERINI J.L. LeConte, 1862. Stem: Dacoder-.

Dactylipalpus Chapuis, 1869: 12. Current status: valid genus in CURCULIONIDAE: SCOLYTINAE: HYLESININI.

Type species: Dactylipalpus transversus Chapuis, 1869, by subsequent designation (Hopkins 1914: 120).

Family-group name: DACTYLIPALPI W.F.H. Blandford, 1893. Stem: Dactylipalp-.

Dactylocalcar Gebien, 1938a: 46. Current status: valid subgenus of Zophosis Latreille, 1802 in TENEBRIONIDAE: PIMELIINAE: ZOPHOSINI.

Type species: Dactylocalcar caecus Gebien, 1938, by monotypy.

Family-group name: DACTYLOCALCARINI Gebien, 1938. Stem: Dactylocalcar-.

Dadophora Duponchel, 1844a: 574. Current status: valid genus in LAMPYRIDAE: LAMPYRINAE: LUCIDOTINI: DADOPHORINA. Note: the genus Dadophora is credited to E. Olivier (1907: 27) in the literature but the name was first made available by Duponchel (1844a: 574) when he described the species Dadophora hyalina (see Bouchard and Bousquet 2020a: 89); combined description of a new genus-group taxon and a single new species (ICZN 1999a, Article 12.2.6).

Type species: Dadophora hyalina Duponchel, 1844, by monotypy.

Family-group name: DADOPHORINI E. Olivier, 1907. Stem: Dadophor-.

Daira Gistel, 1848a: [5]. Current status: junior synonym of Synaptus Eschscholtz, 1829 in ELATERIDAE: ELATERINAE: SYNAPTINI. Note: junior homonym of Daira H. Milne- Edwards, 1830 [Crustacea].

Type species: Elater filiformis Fabricius, 1781, by monotypy.

Family-group name: DAIRAEIDAE Gistel, 1848. Stem: Dair-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

Dalmodes Reitter, 1882a: 197. Current status: valid genus in STAPHYLINIDAE: PSELAPHINAE: GONIACERITAE: INIOCYPHINI: INIOCYPHINA.

Type species: Dalmodes rybaxides Reitter, 1883, by subsequent designation (Newton and Chandler 1989: 50). Note: no species were originally included in this genus by Reitter (1882a); the first ones were the two species listed by Reitter (1883c: 382, 383); R. Lucas (1920: 225) mentioned two species as type species of Dalmodes, but this is considered as an ambiguous typification and is rejected (ICZN 1999a, Article 67.5.3).

Family-group name: DALMODIINI Park, 1951. Stem: Dalmod-.

Dalyat Mateu, 2002: 67. Current status: valid genus in CARABIDAE: SCARITINAE: DALYATINI.

Type species: Dalyat mirabilis Mateu, 2002, by original designation.

Family-group name: DALYATINAE Mateu, 2002. Stem: Dalyat-.

Danacaeomimus Champion, 1922b: 130. Current status: valid genus in MELYRIDAE: DASYTINAE: DANACEINI.

Type species: Danacaeomimus nigropectus Champion, 1922, by original designation.

Family-group name: DANACAEOMIMINI Majer, 1987. Stem: Danacaeomim-.

Danacea Laporte, 1838: 31. Current status: valid genus in MELYRIDAE: DASYTINAE: DANACEINI.

Type species: Lagria pallipes Panzer, 1793, by monotypy.

Family-group name: DANACAEINA C.G. Thomson, 1859. Stem: Danace-.

Danascelis Tomaszewska, 1999: 279. Current status: valid genus in ENDOMYCHIDAE: DANASCELINAE.

Type species: Danascelis elongata Tomaszewska, 1999, by original designation.

Family-group name: DANASCELINAE Tomaszewska, 2000. Stem: Danascel-.

Dapsa Latreille, 1829b: 159. Current status: valid genus in ENDOMYCHIDAE: LYCOPERDININAE.

Type species [suggested]: Endomychus denticollis Germar & Kaulfuss, 1816, by subsequent designation (R. Lucas 1920: 225). Note: the valid type species fixation is that of Latreille (1829b: 159), who included a single species in his new genus, Eumorphus kirbyanus Latreille, 1807, but that is the type species of the genus Indalmus Gerstaecker, 1858; an application to the Commission is necessary to fix Endomychus denticollis Germar & Kaulfuss, 1816 as the type species of Dapsa Latreille, 1829 as is currently accepted in the literature.

Family-group name: DAPSINI Gerstaecker, 1858. Stem: Daps-.

Daptus Fischer, 1823a: pl. 46. Current status: valid genus in CARABIDAE: HARPALINAE: HARPALINI: HARPALINA.

Type species: Daptus vittatus Fischer, 1823, by subsequent designation (Hope 1838a: 88).

Family-group name: DAPTINI J.L. LeConte, 1847. Stem: Dapt-.

Daramus Fairmaire, 1892: 121. Current status: valid genus in CERAMBYCIDAE: CERAMBYCINAE: HESPEROPHANINI: DARAMINA.

Type species: Daramus serricornis Fairmaire, 1892, by monotypy.

Family-group name: DARAMINA Sama, 2008. Stem: Daram-.

Darodilia Laporte, 1867: 71. Current status: valid genus in CARABIDAE: HARPALINAE: PTEROSTICHINI: PTEROSTICHINA.

Type species: Darodilia mandibularis Laporte, 1867, by monotypy.

Family-group name: DARODILIINI Tschitschérine, 1902. Stem: Darodili-.

Dascillus Latreille, 1797: 43. Current status: valid genus in DASCILLIDAE: DASCILLINAE: DASCILLINI.

Type species: Chrysomela cervina Linnaeus, 1758, by subsequent monotypy (Latreille 1802b: 98).

Family-group name: DASCILLIDAE Guérin-Méneville, 1843. Stem: Dascill-.

Dastarcus Walker, 1858: 209. Current status: valid genus in BOTHRIDERIDAE.

Type species: Dastarcus porosus Walker, 1858, by monotypy.

Family-group name: DASTARCINI Reitter, 1922. Stem: Dastarc-.

Dasycerus Brongniart, 1800: 115. Current status: valid genus in STAPHYLINIDAE: DASYCERINAE.

Type species: Dasycerus sulcatus Brongniart, 1800, by monotypy.

Family-group name: DASYCERINI Reitter, 1887. Stem: Dasycer-.

Dasytes Paykull, 1799: 156. Current status: valid genus in MELYRIDAE: DASYTINAE: DASYTINI.

Type species: Hispa coerulea Fabricius, 1775 (= Telephorus caeruleus DeGeer, 1774), by subsequent designation (Westwood 1838c: 28).

Family-group name: DASYDITES Laporte, 1840. Stem: Dasyt-.

Dasyvalgus Kolbe, 1904: 25, 34. Current status: valid genus in SCARABAEIDAE: CETONIINAE: VALGINI: VALGINA.

Type species: Valgus vethi Ritsema, 1879, by subsequent designation (Arrow 1910: 233).

Family-group name: DASYVALGINAE Kolbe, 1904. Stem: Dasyvalg-.

†Davidibelus Zherikhin & Gratshev, 2004: 63. Current status: valid genus in BELIDAE: BELINAE: BELITAE: †DAVIDIBELINI. Note: Lower Cretaceous (Brazil).

Type species: †Davidibelus cearensis Zherikhin & Gratshev, 2004, by original designation.

Family-group name: †DAVIDIBELINI Legalov, 2016: 1475. Stem: Davidibel-.

Decamerus Solier, 1849: 369. Current status: valid genus in THYMALIDAE: DECAMERINAE.

Type species: Decamerus haemorrhoidalis Solier, 1849, by monotypy.

Family-group name: DECAMERINAE Crowson, 1964. Stem: Decamer-. Note: senior homonym of DECAMERIDAE Rasmussen, 1978 (type genus Decameros C. d’Orbigny, 1850) in Echinodermata.

Decarthrocera Laboissière, 1937: 27. Current status: valid genus in CHRYSOMELIDAE: GALERUCINAE: DECARTHROCERINI. Note: two original spellings were used, including Decarthrocerina (p. 29); as far as we know, no one has acted as First Reviser (ICZN 1999a, Article 24.2.3) and we here select Decarthrocera as the correct spelling.

Type species: Decarthrocera viberti Laboissière, 1937, by original designation.

Family-group name: DECARTHROCERINA Laboissière, 1937. Stem: Decarthrocer-.

Decarthron Brendel, 1865: 30. Current status: valid genus in STAPHYLINIDAE: PSELAPHINAE: GONIACERITAE: BRACHYGLUTINI: DECARTHRINA.

Type species: Bryaxis formiceti J.L. LeConte, 1850, by subsequent designation (Bowman 1934: 144).

Family-group name: DECARTHRONINI Park, 1951. Stem: Decarthr-.

Decataphanes Labram & Imhoff, 1840: [Heft 7]. Current status: valid genus in ANTHRIBIDAE: ANTHRIBINAE: DECATAPHANINI.

Type species: Decataphanes gracilis Labram & Imhoff, 1840, by present designation.

Family-group name: DÉCATAPHANIDES Lacordaire, 1865. Stem: Decataphan-.

Declinia Nikitsky, Lawrence, Kirejshuk & Gratshev, 1994: 4. Current status: valid genus in DECLINIIDAE.

Type species: Declinia relicta Nikitsky, Lawrence, Kirejshuk & Gratshev, 1994, by monotypy.

Family-group name: DECLINIIDAE Nikitsky, Lawrence, Kirejtshuk & Gratshev, 1994. Stem: Declini-.

Dectes J.L. LeConte, 1852a: 144. Current status: valid genus in CERAMBYCIDAE: LAMIINAE: ACANTHOCININI.

Type species: Lamia spinosa Say, 1826 (= Dectes sayi L.S. Dillon & E.S. Dillon, 1953), by monotypy.

Family-group name: DECTITAE J. Thomson, 1860. Stem: Dect-.

Decusa Casey, 1900: 54. Current status: valid genus in STAPHYLINIDAE: ALEOCHARINAE: OXYPODINI: DINARDINA.

Type species: Homoeusa expansa J.L. LeConte, 1866, by original designation.

Family-group name: DECUSINI Fenyes, 1919. Stem: Decus-.

Deilus Audinet-Serville, 1834b: 73. Current status: valid genus in CERAMBYCIDAE: CERAMBYCINAE: DEILINI.

Type species: Callidium fugax G.-A. Olivier, 1790, by monotypy.

Family-group name: DÉILATES Fairmaire, 1864. Stem: Deil-.

Deinopsis A.[H.] Matthews, 1838: 193. Current status: valid genus in STAPHYLINIDAE: ALEOCHARINAE: GYMNUSINI.

Type species: Deinopsis fuscatus A. Matthews, 1838 (= Aleochara erosa Stephens, 1832), by monotypy.

Family-group name: DEINOPSINI Sharp, 1883. Stem: Deinops-.

Dejanira J. Thomson, 1864: 134. Current status: valid genus in CERAMBYCIDAE: CERAMBYCINAE: DEJANIRINI.

Type species: Dejanira quadripunctata J. Thomson, 1864, by original designation.

Family-group name: DÉJANIRIDES Lacordaire, 1868. Stem: Dejanir-.

Deleaster Erichson, 1839a: 610. Current status: valid genus in STAPHYLINIDAE: OXYTELINAE: DELEASTERINI.

Type species: Anthophagus dichrous Gravenhorst, 1802, by monotypy.

Family-group name: DELEASTERINI Reitter, 1909. Stem: Deleaster-.

Delevea Reichardt, 1976: 209. Current status: valid genus in TORRIDINCOLIDAE: DELEVEINAE.

Type species: Delevea bertrandi Reichardt, 1976, by original designation.

Family-group name: DELEVEINAE Endrödy-Younga, 1997. Stem: Deleve-.

Delinius Westwood, 1864b: 3. Current status: valid genus in CARABIDAE: HARPALINAE: PTEROSTICHINI: PTEROSTICHINA.

Type species: Delinius essingtonii Westwood, 1864, by monotypy.

Family-group name: DELINIINI Tschitschérine, 1902. Stem: Delini-.

Delocheilus J. Thomson, 1861: 309. Current status: valid genus in CERAMBYCIDAE: PRIONINAE: ANACOLINI.

Type species: Delocheilus prionoides J. Thomson, 1861, by monotypy.

Family-group name: DELOCHILI Lameere, 1913. Stem: Delocheil-.

Delocrania Guérin-Méneville, 1844b: pl. 131. Current status: valid genus in CHRYSOMELIDAE: CASSIDINAE: DELOCRANIINI.

Type species: Delocrania cossyphoides Guérin-Méneville, 1844, by monotypy.

Family-group name: DELOCRANIITAE Spaeth, 1929. Stem: Delocrani-.

Deltochilum Eschscholtz, 1822: 37. Current status: valid genus in SCARABAEIDAE: SCARABAEINAE: DELTOCHILINI.

Type species: Deltochilum dentipes Eschscholtz, 1822, by subsequent designation (Thon 1832a: 424).

Family-group name: DELTOCHILIDES Lacordaire, 1855. Stem: Deltochil-.

Deltomerodes Deuve, 1992: 82. Current status: valid genus in CARABIDAE: PATROBINAE: PATROBINI: DELTOMERODINA.

Type species: Deltomerodes memorabilis Deuve, 1992, by original designation.

Family-group name: DELTOMERODINA Zamotajlov, 2002. Stem: Deltomerod-.

Deltomerus Motschulsky, 1850: 71. Current status: valid genus in CARABIDAE: TRECHINAE: PATROBINI: DELTOMERINA.

Type species: Platynus fulvipes Motschulsky, 1839, by subsequent designation (Enderlein 1909: 487).

Family-group name: DELTOMERIDAE Chaudoir, 1871. Stem: Deltomer-.

Demarziella Balthasar, 1961: 178. Current status: valid genus in SCARABAEIDAE: SCARABAEINAE: ATEUCHINI incertae sedis.

Type species: Demarziella mirifica Balthasar, 1961, by original designation.

Family-group name: DEMARZIELLINI Balthasar, 1961. Stem: Demarziell-.

Demetrias Bonelli, 1810: Tab. Syn. Current status: valid genus in CARABIDAE: HARPALINAE: LEBIINI: DEMETRIADINA.

Type species: Carabus atricapillus Linnaeus, 1758, by subsequent designation (Latreille 1817a: 246). Note: no species were originally included in this genus by Bonelli (1810); the first ones were the three species listed by Panzer (1813: 73).

Family-group name: DEMETRIINAE H.W. Bates, 1886. Stem: Demetriad-.

Demimaea Pascoe, 1870: 440. Current status: valid genus in CURCULIONIDAE: CURCULIONINAE: TYCHIINI: DEMIMAEINA.

Type species: Demimaea luctuosa Pascoe, 1870, by monotypy.

Family-group name: DEMIMAEINI Voss, 1937. Stem: Demimae-.

Dendarus Dejean, 1821: 65. Current status: valid genus in TENEBRIONIDAE: BLAPTINAE: DENDARINI: DENDARINA.

Type species: Helops tristis Rossi, 1790, by subsequent designation (Blanchard in Audouin et al. 1844: pl. 48). Note: see Bouchard et al. (2021: 159) and Kamiński et al. (2021: 576) for comments on the type species of this genus.

Family-group name: PANDARITES Mulsant & Rey, 1854. Stem: Dendar-.

Dendezia Basilewsky, 1952b: 36. Current status: valid genus in LUCANIDAE: LUCANINAE: LUCANINI.

Type species: Dendezia renieri Basilewsky, 1952, by original designation.

Family-group name: DENDEZIINI Benesh, 1955. Stem: Dendezi-.

Dendrocharis Guérin-Méneville, 1843b: 193. Current status: valid genus in EUCNEMIDAE: EUCNEMINAE: DENDROCHARINI.

Type species: Galba flavicornis Guérin-Méneville, 1843, by monotypy.

Family-group name: DENDROCHARINI Fleutiaux, 1920. Stem: Dendrochar-.

Dendroctonus Erichson, 1836: 52. Current status: valid genus in CURCULIONIDAE: SCOLYTINAE: HYLURGINI. Note: name placed on the Official List of Generic Names in Zoology (ICZN 1963a).

Type species: Bostrichus micans Kugelann, 1794, by plenary power (see ICZN 1963a). Note: name placed on the Official List of Specific Names in Zoology (ICZN 1963a).

Family-group name: DENDROCTONIDES Nüsslin, 1912. Stem: Dendrocton-.

Dendrometrus Gistel, 1848b: xi. Current status: junior synonym of Limonius Eschscholtz, 1829 in ELATERIDAE: DENDROMETRINAE: DENDROMETRINI: DENDROMETRINA. Note: unnecessary new replacement name for Limonius Eschscholtz, 1829.

Type species: Elater minutus Linnaeus, 1758, by subsequent designation (Curtis 1838a: pl. 694) (type fixation for Limonius Eschscholtz).

Family-group name: DENDROMETRIDAE Gistel, 1848. Stem: Dendrometr-.

Dendrophagus Schönherr, 1809: 50. Current status: valid genus in SILVANIDAE: BRONTINAE: BRONTINI.

Type species: Cucujus crenatus Paykull, 1799, by monotypy.

Family-group name: DENDROPHAGIDAE Gistel, 1848. Stem: Dendrophag-.

Dendrophilus Leach, 1817: 77. Current status: valid genus in HISTERIDAE: DENDROPHILINAE: DENDROPHILINI.

Type species: Hister punctatus Herbst, 1791, by monotypy.

Family-group name: DENDROPHILINI Reitter, 1909. Stem: Dendrophil-.

Dendroplanetes Gistel, 1856: 368. Current status: junior synonym of Opilo Latreille, 1802 in CLERIDAE: CLERINAE: OPILONITAE: OPILONINI: OPILONINA.

Type species: Attelabus mollis Linnaeus, 1758, by present designation.

Family-group name: DENDROPLANETEIDAE Gistel, 1856. Stem: Dendroplanet-.

Denticollis Piller & Mitterpacher, 1783: 86. Current status: valid genus in ELATERIDAE: DENDROMETRINAE: DENDROMETRINI: DENTICOLLINA.

Type species: Denticollis rubens Piller & Mitterpacher, 1783, by subsequent designation (Hyslop 1921: 639).

Family-group name: DENTICOLLINI Stein & Weise, 1877. Stem: Denticoll-.

Depasophilus Voss, 1922b: 410. Current status: valid genus in ATTELABIDAE: RHYNCHITINAE: RHYNCHITITAE: DEPORAINI.

Type species: Depasophilus bakeri Voss, 1922, by monotypy.

Family-group name: DEPASOPHILINA Legalov, 2003. Stem: Depasophil-.

Deporaus Leach in Samouelle, 1819: 201. Current status: valid genus in ATTELABIDAE: RHYNCHITINAE: RHYNCHITITAE: DEPORAINI. Note: this name is currently considered valid but it is a junior synonym of Platyrynchus Thunberg, 1815; Alonso-Zarazaga and Lyal (1999: 43) mentioned that they would submit an application to the Commission to suppress the older name Platyrynchus Thunberg, 1815, but this has not taken place; Deporaus is listed as a nomen protectum in more recent literature (e.g., Alonso-Zarazaga 2011: 117) but the 25 references needed to establish its status as a nomen protectum (see ICZN 1999a, Article 23.9.1.2) have not been published to our knowledge; an application to the Commission (or publication of evidence to support the nomen protectum status) is still required to conserve the status of Deporaus Leach, 1819 as valid.

Type species: Attelabus betulae Linnaeus, 1758, by monotypy.

Family-group name: DEPORAINI Voss, 1929. Stem: Depora-.

Deracanthus Schönherr, 1823: 1139. Current status: valid genus in CURCULIONIDAE: ENTIMINAE: ENTIMITAE: OPHRYASTINI: DERACANTHINA.

Type species: Curculio spinifex Fabricius, 1787 (= Curculio inderiensis Pallas, 1771), by original designation.

Family-group name: DERACANTHINA Legalov, 2020b: 339. Stem: Deracanth-. Note: junior homonym of DERACANTHIDAE Jacobson, 1905 (type genus Deracantha Fischer von Waldheim, 1833) in Orthoptera.

Derallus Sharp, 1882c: 77. Current status: valid genus in HYDROPHILIDAE: HYDROPHILINAE: BEROSINI.

Type species: Derallus angustus Sharp, 1882, by monotypy.

Family-group name: DERALLINI Bertrand, 1967. Stem: Derall-.

Derancistrus Audinet-Serville, 1832: 181. Current status: valid genus in CERAMBYCIDAE: PRIONINAE: SOLENOPTERINI.

Type species: Prionus elegans Palisot de Beauvois, 1819, by monotypy.

Family-group name: DERANCISTRINI Lameere, 1912. Stem: Derancistr-.

Derbyiella Lea, 1907: 430. Current status: valid genus in CURCULIONIDAE: CRYPTORHYNCHINAE: AEDEMONINI. Note: new replacement name for Derbyia Lea, 1900.

Type species: Derbyia laminatus Lea, 1900, by monotypy (type fixation for Derbyia Lea).

Family-group name: DERBYIELLINA Zimmerman, 1994. Stem: Derbyiell-.

Dercylus Laporte, 1833b: 392. Current status: valid genus in CARABIDAE: HARPALINAE: DERCYLINI.

Type species: Dercylus ater Laporte, 1833, by monotypy.

Family-group name: DERCYLINI Sloane, 1923. Stem: Dercyl-.

Derelomus Schönherr, 1825: 583. Current status: valid genus in CURCULIONIDAE: CURCULIONINAE: DERELOMINI: DERELOMINA.

Type species: Curculio chamaeropis Fabricius, 1798, by original designation.

Family-group name: DÉRÉLOMIDES Lacordaire, 1865. Stem: Derelom-.

Derema Fauvel, 1899: 41. Current status: valid genus in STAPHYLINIDAE: ALEOCHARINAE: MIMANOMMATINI: DORYLOPHILINA.

Type species: Derema foveicollis Fauvel, 1899, by subsequent designation (Fenyes 1912: 24).

Family-group name: DEREMINI Seevers, 1965. Stem: Derem-.

Deretaphrus Newman, 1842c: 403. Current status: valid genus in BOTHRIDERIDAE.

Type species: Deretaphrus fossus Newman, 1842, by subsequent designation (Wollaston in Wollaston and Newman 1855: ccviii).

Family-group name: DERETAPHRINI G.H. Horn, 1878. Stem: Deretaphr-.

Deridea Westwood, 1875: 226. Current status: valid genus in MELOIDAE: ELETICINAE: DERIDEINI.

Type species: Deridea curculionides Westwood, 1875, by monotypy.

Family-group name: DERIDEIDES Wellman, 1910. Stem: Deride-.

Dermatodes Schönherr, 1840a: 895. Current status: valid genus in CURCULIONIDAE: ENTIMINAE: POLYDRUSITAE: DERMATODINI. Note: new replacement name for Lagostomus Schönherr, 1833.

Type species: Lagostomus paganus Gyllenhal, 1833, by original designation (type fixation for Lagostomus Schönherr).

Family-group name: DERMATODINI Emden, 1936. Stem: Dermatod-.

Dermatohomoeus Hlisnikovský, 1963: 302. Current status: valid genus in LEIODIDAE: LEIODINAE: PSEUDOLIODINI.

Type species: Dermatohomoeus guineensis Hlisnikovský, 1963, by original designation.

Family-group name: DERMATOHOMOEINI Hlisnikovský, 1963. Stem: Dermatohomoe-.

Dermeanthrenus Háva, 2008: 62. Current status: valid genus in DERMESTIDAE: MEGATOMINAE: ANTHRENINI.

Type species: Dermeanthrenus pretiosus Háva, 2008, by original designation.

Family-group name: DERMEANTHRENINA Háva, 2011: 6. Stem: Dermeanthren-.

Dermestes Linnaeus, 1758: 342, 354. Current status: valid genus in DERMESTIDAE: DERMESTINAE: DERMESTINI.

Type species: Dermestes lardarius Linnaeus, 1758, by subsequent designation (Latreille 1810: 428).

Family-group name: DERMESTINI Latreille, 1804. Stem: Dermest-.

Dermestoides Schäffer, 1777: pl. cxxxviii. Current status: valid genus in CLERIDAE: KORYNETINAE.

Type species [suggested]: Dermestes sanguinicollis Fabricius, 1787, by subsequent designation (J.R. Winkler 1979: 311). Note: no species were originally included in this genus by Schäffer (1777); the first ones were the two species listed by Herbst (1783: 40), Dermestoides unipunctatus Herbst, 1783 (= Dermestes linearis Goeze, 1777) and Dermestoides bipunctatus Herbst, 1783 (possibly a senior synonym of Rhizophagus bipunctatus Herbst, 1793); acceptance of either of these two as type species of Dermestoides will threaten the status of this type genus since the former is now included in Lyctus Fabricius, 1792 (BOSTRICHIDAE) and the latter possibly in MONOTOMIDAE (more research is needed to ascertain the identity of Dermestoides bipunctatus Herbst, 1783); an application to the Commission is necessary to fix Dermestes sanguinicollis Fabricius, 1787 as the type species of Dermestoides Schäffer, 1777 in order to conserve its currently accepted concept.

Family-group name: DERMESTOIDINI Jakobson, 1911. Stem: Dermestoid-.

Derobrachus Audinet-Serville, 1832: 154. Current status: valid genus in CERAMBYCIDAE: PRIONINAE: PRIONINI.

Type species: Derobrachus brevicollis Audinet-Serville, 1832, by monotypy.

Family-group name: DEROBRACHITAE J. Thomson, 1864. Stem: Derobrach-.

Derodontus J.L. LeConte, 1861: 100. Current status: valid genus in DERODONTIDAE: DERODONTINAE.

Type species: Cryptophagus maculatus Melsheimer, 1844, by subsequent designation (R. Lucas 1920: 232).

Family-group name: DERODONTIDAE J.L. LeConte, 1861. Stem: Derodont-.

Derolathrus Sharp in Sharp and H. Scott, 1908: 430. Current status: valid genus in JACOBSONIIDAE.

Type species: Derolathrus atomus Sharp, 1908, by monotypy.

Family-group name: DEROLATHRIINAE Sen Gupta, 1979. Stem: Derolathr-.

Deronectes Sharp, 1882b: 390, 418. Current status: valid genus in DYTISCIDAE: HYDROPORINAE: HYDROPORINI: DERONECTINA. Note: this genus was first made available by Sharp (1880: cxlviii) when he listed two available species, Dytiscus duodecimpustulatus Fabricius, 1792 and Dytiscus elegans Panzer, 1794, but the first species is the type species of Stictotarsus A. Zimmermann, 1919 and the second is currently included in the genus Nebrioporus Régimbart, 1906; an application to the Commission is necessary to suppress the older but unused name Deronectes Sharp, 1880 in order to conserve the current usage of Deronectes Sharp, 1882.

Type species: Hydroporus latus Stephens, 1829, by subsequent designation (F. Balfour-Browne 1934: 179).

Family-group name: DERONECTINI Galewski, 1994. Stem: Deronect-.

Derops Sharp, 1889a: 418. Current status: valid genus in STAPHYLINIDAE: TACHYPORINAE: DEROPINI.

Type species: Derops longicornis Sharp, 1889, by monotypy.

Family-group name: DEROPSINI Smetana, 1983. Stem: Derop-.

†Desmatus Dolin, 1975: 60. Current status: valid genus in ELATERIDAE: †PROTAGRYPNINAE: †DESMATINI. Note: Jurassic (Kazakhstan).

Type species: †Desmatus lapidarius Dolin, 1975, by original designation.

Family-group name: †DESMATINI Dolin, 1975. Stem: Desmat-.

Desmidophorus Dejean, 1835: 296. Current status: valid genus in CURCULIONIDAE: BRACHYCERINAE: ERIRHININI.

Type species: Curculio hebes Fabricius, 1781, by monotypy.

Family-group name: DESMIDOPHORINAE Morimoto, 1962. Stem: Desmidophor-.

Desmiphora Audinet-Serville, 1835a: 62. Current status: valid genus in CERAMBYCIDAE: LAMIINAE: DESMIPHORINI.

Type species: Lamia fasciculata G.-A. Olivier, 1792, by subsequent designation (Drapiez 1838: 456).

Family-group name: DESMIPHORITAE J. Thomson, 1860. Stem: Desmiphor-.

Desmocerus Dejean, 1821: 111. Current status: valid genus in CERAMBYCIDAE: LEPTURINAE: DESMOCERINI.

Type species: Stenocorus cyaneus Fabricius, 1775 (= Cerambyx palliatus Forster, 1771), by monotypy.

Family-group name: DESMOCÉRITES Blanchard, 1845. Stem: Desmocer-.

Desmonyx Arrow, 1907: 355. Current status: valid genus in SCARABAEIDAE: RUTELINAE: RUTELINI: DESMONYCHINA.

Type species: Desmonyx humeralis Arrow, 1907, by monotypy.

Family-group name: DESMONYCHINAE Arrow, 1917. Stem: Desmonych-.

Desmoris J.L. LeConte in J.L. LeConte and G.H. Horn, 1876: 167. Current status: junior synonym of Smicronyx Schönherr, 1843 in CURCULIONIDAE: CURCULIONINAE: SMICRONYCHINI.

Type species: Rhynchaenus constrictus Say, 1824, by subsequent designation (D.M. Anderson 1962: 322).

Family-group name: DESMORHINES J.L. LeConte, 1876. Stem: Desmorin-. Note: nomen oblitum (see Bouchard et al. 2011: 584).

Dexoris C.O. Waterhouse, 1878: 105. Current status: valid genus in LYCIDAE: DEXORINAE: DEXORINI: DEXORINA.

Type species: Dexoris insignis C.O. Waterhouse, 1878, by original designation.

Family-group name: DEXORINAE Bocák & Bocáková, 1989. Stem: Dexor-.

Dhysores Grouvelle, 1903: 90, 92. Current status: valid genus in CARABIDAE: RHYSODINAE: DHYSORINI.

Type species: Rhysodes thoreyi Grouvelle, 1903, by subsequent designation (Hincks 1950a: 4).

Family-group name: DHYSORINA R.T. Bell & J.R. Bell, 1978. Stem: Dhysor-.

Diabathrarius Schönherr, 1840b: 464. Current status: valid genus in CURCULIONIDAE: MOLYTINAE: LITHININI: LITHININA.

Type species: Diabathrarius variegatus Boheman, 1840, by original designation.

Family-group name: DIABATHRARIIDES Lacordaire, 1863. Stem: Diabathrari-.

Diabrotica Chevrolat in Dejean, 1836a: 380. Current status: valid genus in CHRYSOMELIDAE: GALERUCINAE: LUPERINI.

Type species: Crioceris fucata Fabricius, 1787, by subsequent designation (Barber 1947a: 151).

Family-group name: DIABROTICITES Chapuis, 1875. Stem: Diabrotic-.

Diacantha Chevrolat in Dejean, 1836a: 378. Current status: valid genus in CHRYSOMELIDAE: GALERUCINAE: LUPERINI.

Type species: Galleruca unifasciata G.-A. Olivier, 1808, by subsequent designation (Hincks 1949: 613).

Family-group name: DIACANTHINI Weise, 1902a: 273. Stem: Diacanth-. Note: this name was overlooked in Bouchard et al. (2011) and Bouchard and Bousquet (2020a); junior homonym of the nomen oblitum DIACANTHIDAE Gistel, 1848 (type genus Diacanthus Latreille, 1834) in Coleoptera: ELATERIDAE.

Diacanthus Latreille, 1834: 151. Current status: junior synonym of Selatosomus Stephens, 1830 in ELATERIDAE: DENDROMETRINAE: PROSTERNINI.

Type species: Elater aeneus Linnaeus, 1758, by subsequent designation (C.G. Thomson 1859: 102).

Family-group name: DIACANTHIDAE Gistel, 1848. Stem: Diacanth-. Note: nomen oblitum (see Bouchard et al. 2011: 315); senior homonym of DIACANTHINI Weise, 1902 (type genus Diacantha Chevrolat, 1836) in Coleoptera: CHRYSOMELIDAE.

Diamerus Erichson, 1836: 57. Current status: valid genus in CURCULIONIDAE: SCOLYTINAE: DIAMERINI.

Type species: Hylesinus hispidus Klug, 1833, by monotypy.

Family-group name: DIAMERINAE Hagedorn, 1909. Stem: Diamer-.

Diaperis Geoffroy, 1762: 337. Current status: valid genus in TENEBRIONIDAE: DIAPERINAE: DIAPERINI: DIAPERINA. Note: name placed on the Official List of Generic Names in Zoology (ICZN 1994a).

Type species: Chrysomela boleti Linnaeus, 1758, by subsequent monotypy (O.F. Müller 1776: 74; see ICZN 1994a). Note: name placed on the Official List of Specific Names in Zoology (ICZN 1994a).

Family-group name: DIAPERIALAE Latreille, 1802. Stem: Diaper-.

Diaphanops Schönherr, 1845: 342. Current status: valid genus in CHRYSOMELIDAE: SAGRINAE: DIAPHANOPSIDINI.

Type species: Diaphanops westermanni Boheman, 1845, by original designation.

Family-group name: DIAPHANOPSIDINI Monrós, 1958. Stem: Diaphanopsid-.

Diaphonia Newman, 1840b: 366. Current status: valid genus in SCARABAEIDAE: CETONIINAE: SCHIZORHININI: SCHIZORHININA.

Type species: Diaphonia dispar Newman, 1840, by monotypy.

Family-group name: DIAPHONIADAE Kraatz, 1880. Stem: Diaphoni-.

Diapus Chapuis, 1865: 43, 329. Current status: valid genus in CURCULIONIDAE: PLATYPODINAE: TESSEROCERINI: DIAPODINA.

Type species: Diapus quadrispinatus Chapuis, 1865, by subsequent designation (Hopkins 1914: 121).

Family-group name: DIAPODARIAE H. Strohmeyer, 1914. Stem: Diapod-.

Diatelium Pascoe, 1863a: 27. Current status: valid genus in STAPHYLINIDAE: SCAPHIDIINAE: SCAPHIDIINI.

Type species: Diatelium wallacei Pascoe, 1863, by monotypy.

Family-group name: DIATELIITAE Achard, 1924. Stem: Diateli-.

Dibolia Latreille, 1829b: 155. Current status: valid genus in CHRYSOMELIDAE: GALERUCINAE: ALTICINI.

Type species: Haltica occultans J.D.W. Koch, 1803, by subsequent designation (Curtis 1833a: pl. 435 [p. 2]).

Family-group name: DIBOLIITES Chapuis, 1875. Stem: Diboli-.

Dicaelus Bonelli, 1813: 446. Current status: valid genus in CARABIDAE: HARPALINAE: LICININI: DICAELINA.

Type species: Dicaelus violaceus Bonelli, 1813 (= Dicaelus purpuratus Bonelli, 1813), by subsequent designation (Hope 1838a: 82).

Family-group name: DICOELIDAE Laporte, 1834. Stem: Dicael-.

Dicax Fauvel, 1878: 512, 518. Current status: valid genus in STAPHYLINIDAE: PAEDERINAE: PAEDERINI: DICAXINA.

Type species: Lathrobium longiceps Fauvel, 1877, by subsequent designation (Bertkau 1879: 441).

Family-group name: DICAXINA Schomann & Solodovnikov, 2017: 10. Stem: Dicax-.

Dicerca Eschscholtz, 1829b: 9. Current status: valid genus in BUPRESTIDAE: CHRYSOCHROINAE: DICERCINI: DICERCINA. Note: the original spelling was Dicerea but Dicerca was ruled by the Commission to be the correct original spelling and the name was placed on the Official List of Generic Names in Zoology (ICZN 1994f).

Type species: Buprestis aenea Linnaeus, 1761, by subsequent designation (Westwood 1838c: 24; see ICZN 1994f). Note: name placed on the Official List of Specific Names in Zoology (ICZN 1994f).

Family-group name: DICERCAEIDAE Gistel, 1848. Stem: Dicerc-.

Dichelonyx Harris, 1827: 7. Current status: valid genus in SCARABAEIDAE: MELOLONTHINAE: DICHELONYCHINI.

Type species: Melolontha linearis Gyllenhal, 1817 (= Melolontha elongatula Schönherr, 1817), by monotypy.

Family-group name: DICHELONYCHIDAE Burmeister, 1855. Stem: Dichelonych-.

Dichelus Lepeletier & Audinet-Serville, 1828: 373. Current status: valid genus in SCARABAEIDAE: MELOLONTHINAE: HOPLIINI: HOPLIINA.

Type species: Melolontha dentipes Fabricius, 1781, by monotypy.

Family-group name: DICHELIDEN Oken, 1843. Stem: Dichel-.

Dichillus Jacquelin du Val, 1860b: 253. Current status: valid genus in TENEBRIONIDAE: PIMELIINAE: STENOSINI: DICHILLINA.

Type species: Tagenia minuta Solier, 1838, by original designation.

Family-group name: DICHILLINA Reitter, 1916. Stem: Dichill-.

Dichophyia Gistel, 1848b: xi, 131. Current status: junior synonym of Mallosoma Audinet-Serville, 1834 in CERAMBYCIDAE: CERAMBYCINAE: DICHOPHYIINI. Note: unnecessary new replacement name for Mallosoma Audinet-Serville, 1834.

Type species: Mallosoma elegans Audinet-Serville, 1834 (= Cerambyx zonatus C.R. Sahlberg, 1823), by monotypy (type fixation for Mallosoma Audinet-Serville).

Family-group name: DICHOPHYIAEIDAE Gistel, 1848. Stem: Dichophyi-.

Dichotomius Hope, 1838b: 321. Current status: valid genus in SCARABAEIDAE: SCARABAEINAE: DICHOTOMIINI.

Type species: Scarabaeus boreus G.-A. Olivier, 1789, by original designation.

Family-group name: DICHOTOMIINI Pereira, 1954. Stem: Dichotomi-.

Dichotrachelus Stierlin, 1853: 171. Current status: valid genus in CURCULIONIDAE: CYCLOMINAE: DICHOTRACHELINI.

Type species: Dichotrachelus sulcipennis Stierlin, 1853, by monotypy.

Family-group name: DICHOTRACHELINI Hoffmann, 1957. Stem: Dichotrachel-.

Dicrania Lepeletier & Audinet-Serville, 1828: 371. Current status: valid genus in SCARABAEIDAE: MELOLONTHINAE: MACRODACTYLINI.

Type species: Dicrania rubricollis Lepeletier & Audinet-Serville, 1828, by subsequent designation (Blanchard in Audouin et al. 1843: pl. 44).

Family-group name: DICRANIADAE Burmeister, 1855. Stem: Dicrani-.

Dicranosterna Motschulsky, 1860b: 193. Current status: valid genus in CHRYSOMELIDAE: CHRYSOMELINAE: CHRYSOMELINI.

Type species: Paropsis picea G.-A. Olivier, 1807, by original designation.

Family-group name: DICRANOSTERNINI Weise, 1915. Stem: Dicranostern-.

Dicrepidius Eschscholtz, 1829a: 31. Current status: valid genus in ELATERIDAE: ELATERINAE: DICREPIDIINI.

Type species: Dicrepidius pectinicornis Eschscholtz, 1829 (= Elater ramicornis Palisot de Beauvois, 1805), by subsequent designation (Hyslop 1921: 640).

Family-group name: DICREPIDIITAE J. Thomson, 1858. Stem: Dicrepidi-.

Dicrochile Guérin-Méneville, 1847: ciii. Current status: valid genus in CARABIDAE: HARPALINAE: LICININI: DICROCHILINA. Note: this genus was first described without nominal species by Guérin-Méneville (1846: 428) under the spelling Dicronochilus; the spelling Dicrochile, which is in prevailing usage, is an incorrect subsequent spelling of Dicronochilus but since Dicrochile is not attributed to the publication of the original spelling, Article 33.3.1 (ICZN 1999a) cannot be used; an application to the Commission is necessary to conserve the currently used spelling Dicrochile.

Type species: Dicrochile anchomenoides Guérin-Méneville, 1847, by subsequent designation (Larochelle and Larivière 2001: 114).

Family-group name: DICROCHILINA Ball, 1992. Stem: Dicrochil-.

Dicrodontus Chaudoir, 1872a: 139. Current status: valid genus in CARABIDAE: HARPALINAE: ZUPHIINI: DICRODONTINA.

Type species: Polistichus brunneus Dejean, 1831, by monotypy.

Family-group name: DICRODONTINI Machado, 1992. Stem: Dicrodont-.

Dicronocephalus Hope, 1831: 24. Current status: valid genus in SCARABAEIDAE: CETONIINAE: GOLIATHINI: DICRONOCEPHALINA.

Type species: Dicronocephalus wallichii Hope, 1831, by monotypy.

Family-group name: DICRONOCEPHALINA Krikken, 1984. Stem: Dicronocephal-.

Dictyoptera Latreille, 1829a: 464. Current status: valid genus in LYCIDAE: EROTINAE: DICTYOPTERINI.

Type species [suggested]: Pyrochroa aurora Herbst, 1784, misidentified as Cantharis sanguineus Linnaeus, 1761 (sensu Latreille, 1829) in the subsequent designation by Blanchard in Audouin et al. (1842: pl. 32). Note: the valid type species fixation is that of Westwood (1838c: 27), who selected Pyrochroa minuta Fabricius, 1787, but that species is the type species of Platycis C.G. Thomson, 1859; an application to the Commission is necessary to fix Pyrochroa aurora Herbst, 1784 as the type species of Dictyoptera Latreille, 1829.

Family-group name: DICTYOPTERINI Houlbert, 1922. Stem: Dictyopter-.

Didactylia H. d’Orbigny, 1896: 247. Current status: valid genus in SCARABAEIDAE: APHODIINAE: APHODIINI: DIDACTYLIINA.

Type species: Aphodius pallicolor Fairmaire, 1886, by monotypy. Note: H. d’Orbigny (1896: 248) proposed the unjustified emendation pallidicolor when he fixed the type species of his new genus.

Family-group name: DIDACTYLIINA Pittino, 1985. Stem: Didactyli-.

Didrepanephorus Wood-Mason, 1878: 423. Current status: valid genus in SCARABAEIDAE: RUTELINAE: RUTELINI: DIDREPANEPHORINA.

Type species: Didrepanephorus bifalcifer Wood-Mason, 1878, by original designation.

Family-group name: DIDREPANOPHORINA Ohaus, 1918. Stem: Didrepanephor-.

Didymonycha Aurivillius, 1922a: 31. Current status: junior synonym of Amillarus J. Thomson, 1857 in CERAMBYCIDAE: LAMIINAE: AGAPANTHIINI.

Type species: Didymonycha singularis Aurivillius, 1922, by monotypy.

Family-group name: DIDYMONYCHINI Aurivillius, 1922. Stem: Didymonych-.

Dieropsis Gahan, 1908: 95. Current status: valid genus in CLERIDAE: CLERINAE: OPILONITAE: DIEROPSEINI.

Type species: Dieropsis quadriplagiata Gahan, 1908, by monotypy.

Family-group name: DIEROPSINAE J.R. Winkler, 1964. Stem: Dieropse-. Note: see Bouchard et al. (2011: 347) regarding the recommended stem for this family-group name.

Diestota Mulsant & Rey, 1870: 194. Current status: valid genus in STAPHYLINIDAE: ALEOCHARINAE: DIESTOTINI.

Type species: Diestota mayeti Mulsant & Rey, 1870 (= Bolitochara testacea Kraatz, 1859), by monotypy.

Family-group name: DIESTOTATES Mulsant & Rey, 1871. Stem: Diestot-.

Diglossa Haliday, 1837: 252. Current status: invalid senior synonym of Diglotta Champion, 1899 in STAPHYLINIDAE: ALEOCHARINAE: DIGLOTTINI. Note: junior homonym of Diglossa Wagler, 1832 [Aves].

Type species: Diglossa mersa Haliday, 1837, by monotypy.

Family-group name: DIGLOSSAIRES Mulsant & Rey, 1873. Stem: Digloss-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

Diglotta Champion, 1899: 265. Current status: valid genus in STAPHYLINIDAE: ALEOCHARINAE: DIGLOTTINI. Note: new replacement name for Diglossa Haliday, 1837.

Type species: Diglossa mersa Haliday, 1837, by monotypy (type fixation for Diglossa Haliday).

Family-group name: DIGLOTTINA Jakobson, 1909. Stem: Diglott-.

Dignamptus J.L. LeConte, 1878a: 421. Current status: junior synonym of Talanus Jacquelin du Val, 1857 in TENEBRIONIDAE: STENOCHIINAE: TALANINI.

Type species: Dignamptus stenochinus J.L. LeConte, 1878, by subsequent designation (Bousquet et al. 2018: 325).

Family-group name: DIGNAMPTINI J.L. LeConte & G.H. Horn, 1883. Stem: Dignampt-.

Digrammus Fauvel, 1900a: 123. Current status: valid genus in STAPHYLINIDAE: ALEOCHARINAE: DIGRAMMINI.

Type species: Digrammus miricollis Fauvel, 1900, by monotypy.

Family-group name: DIGRAMMINI Fauvel, 1900. Stem: Digramm-.

Dihammatus C.O. Waterhouse, 1880a: 29. Current status: valid genus in LYCIDAE: METRIORRHYNCHINAE: DIHAMMATINI.

Type species: Dihammatus cribripennis C.O. Waterhouse, 1880, by subsequent designation (Bourgeois 1891: 349).

Family-group name: DIHAMMATINI Bocák & Bocáková, 2008. Stem: Dihammat-.

Dila Fischer von Waldheim, 1844: 111. Current status: valid genus in TENEBRIONIDAE: BLAPTINAE: BLAPTINI: BLAPTINA.

Type species: Blaps laevicollis Gebler, 1841, by subsequent designation (Motschulsky 1860: 530).

Family-group name: DILINA Ren in Ren et al., 2016: 85, 431. Stem: Dil-.

Dilolycus Kleine, 1926: 186. Current status: junior synonym of Metriorrhynchus Gemminger, 1869 in LYCIDAE: METRIORRHYNCHINAE: METRIORRHYNCHINI: METRIORRHYNCHINA.

Type species: Dilolycus lamellatus Kleine, 1926, by original designation.

Family-group name: DILOLYCINAE Kleine, 1926. Stem: Dilolyc-.

Dilophochila H.W. Bates, 1888: 261. Current status: valid genus in SCARABAEIDAE: RUTELINAE: ANOMALINI: ANOMALINA.

Type species: Dilophochila bolacoides H.W. Bates, 1888, by monotypy.

Family-group name: DILOPHOCHILINA Ohaus, 1918. Stem: Dilophochil-.

Dilophotes C.O. Waterhouse, 1880a: 75. Current status: valid genus in LYCIDAE: METRIORRHYNCHINAE: DILOPHOTINI.

Type species: Lycus exilis C.O. Waterhouse, 1878, by original designation.

Family-group name: DILOPHOTINAE Kleine, 1929. Stem: Dilophot-.

Dima Charpentier, 1825: 191. Current status: valid genus in ELATERIDAE: DENDROMETRINAE: DIMINI.

Type species: Dima elateroides Charpentier, 1825, by monotypy.

Family-group name: DIMITES Candèze, 1863. Stem: Dim-.

Dimerometopus Célis, 1970: 245. Current status: valid genus in STAPHYLINIDAE: PSELAPHINAE: CLAVIGERITAE: CLAVIGERINI.

Type species: Hadrophorus bicephalus Jeannel, 1954, by original designation.

Family-group name: DIMEROMETOPINI Célis, 1970. Stem: Dimerometop-.

Dimerus Fiori, 1900: 103. Current status: junior synonym of Octomicrus L.W. Schaufuss, 1877 in STAPHYLINIDAE: PSELAPHINAE: EUPLECTITAE: DIMERINI.

Type species: Dimerus staphylinoides Fiori, 1900, by monotypy.

Family-group name: DIMERINI Raffray, 1908. Stem: Dimer-.

Dimetrota Mulsant & Rey, 1873: 165. Current status: valid subgenus of Atheta C.G. Thomson, 1858 in STAPHYLINIDAE: ALEOCHARINAE: ATHETINI: ATHETINA.

Type species: Homalota tristicula Mulsant & Rey, 1873 (= Homalota cadaverina Brisout de Barneville, 1860), by subsequent designation (Blackwelder 1952: 125).

Family-group name: DIMETROTAE Seevers, 1978. Stem: Dimetrot-.

Dimonomera Cameron, 1933: 103. Current status: valid genus in STAPHYLINIDAE: ALEOCHARINAE: MYLLAENINI.

Type species: Dimonomera indica Cameron, 1933, by monotypy.

Family-group name: DIMONOMERINI Cameron, 1933. Stem: Dimonomer-.

Dinapate G.H. Horn, 1886: 1. Current status: valid genus in BOSTRICHIDAE: BOSTRICHINAE: APATINI.

Type species: Dinapate wrightii G.H. Horn, 1886, by monotypy.

Family-group name: DINAPATINAE Lesne, 1910. Stem: Dinapat-.

Dinarda Leach in Samouelle, 1819: 177. Current status: valid genus in STAPHYLINIDAE: ALEOCHARINAE: OXYPODINI: DINARDINA.

Type species: Lomechusa dentata Gravenhorst, 1806, by original designation.

Family-group name: DINARDAIRES Mulsant & Rey, 1873. Stem: Dinard-.

Dinardopsis Bruch, 1917: 257. Current status: valid genus in STAPHYLINIDAE: ALEOCHARINAE: HOMALOTINI: DINARDOPSINA.

Type species: Dinardopsis solenopsidicola Bruch, 1917, by original designation.

Family-group name: DINARDOPSES Bernhauer & Scheerpeltz, 1926. Stem: Dinardops-.

Dineutus W.S. MacLeay, 1825: 30. Current status: valid genus in GYRINIDAE: GYRININAE: DINEUTINI: DINEUTINA.

Type species: Dineutus politus W.S. MacLeay, 1825, by monotypy.

Family-group name: DINEUTIDES E. Desmarest, 1851. Stem: Dineut-.

Dinoderus Stephens, 1830: 349, 352. Current status: valid genus in BOSTRICHIDAE: DINODERINAE.

Type species: Dinoderus ocellaris Stephens, 1830, by monotypy. Note: two species were originally listed by Stephens (1830) but one of them is conditionally included and therefore cannot be considered as an originally included species (ICZN 1999a, Article 67.2.5).

Family-group name: DINODERINA C.G. Thomson, 1863. Stem: Dinoder-.

Dinomorphus Perty, 1832: 71. Current status: valid genus in CURCULIONIDAE: MOLYTINAE: DINOMORPHINI.

Type species: Dinomorphus pimelioides Perty, 1832, by monotypy.

Family-group name: DINOMORPHIDES Lacordaire, 1863. Stem: Dinomorph-.

Dinorhopala Pascoe, 1860b: 61. Current status: valid genus in CURCULIONIDAE: CURCULIONINAE: RHAMPHINI: DINORHOPALINA.

Type species: Dinorhopala spinosa Pascoe, 1860, by monotypy.

Family-group name: DINORRHOPALINA Voss, 1936. Stem: Dinorhopal-.

Diocalandra Faust, 1894a: 353. Current status: valid genus in CURCULIONIDAE: DRYOPHTHORINAE: LITOSOMINI: DIOCALANDRINA.

Type species: Calandra frumenti Fabricius, 1801, by monotypy.

Family-group name: DIOCALANDRINI Zimmerman, 1993. Stem: Diocalandr-.

Diochus Erichson, 1839b: 300. Current status: valid genus in STAPHYLINIDAE: STAPHYLININAE: DIOCHINI.

Type species: Diochus nanus Erichson, 1839, by monotypy.

Family-group name: DIOCHI Casey, 1906. Stem: Dioch-.

Diodesma Latreille, 1829b: 97. Current status: valid genus in ZOPHERIDAE: COLYDIINAE: SYNCHITINI.

Type species: Diodesma subterranea Latreille, 1829, by monotypy.

Family-group name: DIODESMINI Reitter, 1911. Stem: Diodesmat-.

Diomus Mulsant, 1850: 951. Current status: valid genus in COCCINELLIDAE: COCCINELLINAE: DIOMINI.

Type species: Coccinella thoracica Fabricius, 1801, by subsequent designation (Korschefsky 1931: 116).

Family-group name: DIOMINI Gordon, 1999. Stem: Diom-.

Diorus A. White, 1853: 135. Current status: valid genus in CERAMBYCIDAE: CERAMBYCINAE: DIORINI.

Type species: Diorus biapiculatus A. White, 1853, by monotypy.

Family-group name: DIORINAE Lane, 1950. Stem: Dior-.

Dioryche W.S. MacLeay, 1825: 21. Current status: valid genus in CARABIDAE: HARPALINAE: HARPALINI: HARPALINA.

Type species: Harpalus torta W.S. MacLeay, 1825, by monotypy.

Family-group name: DIORYCHI Csiki, 1932. Stem: Diorych-.

Diorymerus Schönherr, 1825: 586. Current status: valid genus in CURCULIONIDAE: BARIDINAE: APOSTASIMERINI: DIORYMERINA.

Type species: Orobitis altus Germar, 1823, by original designation.

Family-group name: DIORYMERIDES Jekel, 1865. Stem: Diorymer-.

Dipelicus Hope, 1843a: 62. Current status: valid genus in SCARABAEIDAE: DYNASTINAE: PENTODONTINI: DIPELICINA.

Type species: Dipelicus cantori Hope, 1843, by monotypy.

Family-group name: DIPELICINA Carne, 1957. Stem: Dipelic-.

Diphaulaca Chevrolat in Dejean, 1836a: 388. Current status: valid genus in CHRYSOMELIDAE: GALERUCINAE: ALTICINI.

Type species: Altica aulica G.-A. Olivier, 1808, by subsequent designation (Chevrolat 1844: 46).

Family-group name: DIPHAULACINI Bechyné & Špringlová de Bechyné, 1975. Stem: Diphaulac-.

Diphucephala Dejean, 1821: 58. Current status: valid genus in SCARABAEIDAE: SERICINAE: DIPHUCEPHALINI.

Type species: Melolontha sericea W. Kirby, 1818, by monotypy.

Family-group name: DIPHUCÉPHALITES Laporte, 1840. Stem: Diphucephal-.

Diphycerus Deyrolle & Fairmaire, 1878: 100. Current status: valid genus in SCARABAEIDAE: MELOLONTHINAE: DIPHYCERINI.

Type species: Diphycerus davidis Fairmaire, 1878, by monotypy.

Family-group name: DIPHYCERINI S.I. Medvedev, 1952. Stem: Diphycer-.

Diphyllostoma Fall, 1901: 324. Current status: valid genus in DIPHYLLOSTOMATIDAE. Note: new replacement name for Phyllostoma Fall, 1901.

Type species: Phyllostoma fimbriatum Fall, 1901, by monotypy (type fixation for Phyllostoma Fall).

Family-group name: DIPHYLLOSTOMATIDAE Holloway, 1972. Stem: Diphyllostomat-.

Diplocyrtus Quedenfeldt, 1887: 257. Current status: valid genus in TENEBRIONIDAE: TENEBRIONINAE: APOCRYPHINI.

Type species: Diplocyrtus floccosus Quedenfeldt, 1887, by monotypy.

Family-group name: DIPLOCYRTHINI Escalera, 1914. Stem: Diplocyrt-.

Diplognatha Gory & Percheron, 1833: 19, 31, 51. Current status: valid genus in SCARABAEIDAE: CETONIINAE: DIPLOGNATHINI. Note: the spelling Diplognata used on page 51 was corrected to Diplognatha in the “Errata” of the same work.

Type species: Scarabaeus gagates Forster, 1771, by subsequent designation (Duponchel 1843a: 360).

Family-group name: DIPLOGNATHIDAE Burmeister, 1842. Stem: Diplognath-.

Diplotaxis W. Kirby, 1837: 129. Current status: valid genus in SCARABAEIDAE: MELOLONTHINAE: DIPLOTAXINI.

Type species: Diplotaxis tristis W. Kirby, 1837, by monotypy.

Family-group name: DIPLOTAXIDAE W. Kirby, 1837. Stem: Diplotax-.

Dircaea Fabricius, 1798: 6. Current status: valid genus in MELANDRYIDAE: MELANDRYINAE: DIRCAEINI.

Type species: Dircaea quadriguttata Fabricius, 1798 (= Hypulus quadriguttatus Paykull, 1798), by subsequent designation (Blanchard in Audouin et al. 1844: pl. 52). Note: based on currently available dates of publication, it is possible that Fabricius’ species, issued by 29 April 1798 (Bousquet 2017a: 107), may be older than Paykull’s species, issued by 20 June 1798 (Bousquet 2016: 409) although the date of the introduction in Paykull’s work predates that of Fabricius’ work by about two weeks; at this time we prefer to maintain the status quo.

Family-group name: DIRCAEIDAE W. Kirby, 1837. Stem: Dircae-.

Dirhagus Latreille, 1834: 130. Current status: junior synonym of Microrhagus Dejean, 1833 in EUCNEMIDAE: MELASINAE: DIRHAGINI.

Type species [suggested]: Elater pygmaeus Fabricius, 1792, by subsequent designation (Muona 1987: 82). Note: the type species for this genus is usually given as “Elater pygmaeus Fabricius, 1792, by monotypy”; however, contrary to comments in Muona (1987: 82), the species Dirhagus longulus was made available by Latreille (1834: 131) and unambiguously combined with Dirhagus, as such, it should be treated as a second species originally included in Dirhagus; additionally, D. longulus, the identity of which needs further investigation, was selected as type species of Dirhagus by Guérin- Méneville (1843b: 181).

Family-group name: DIRRHAGINI Reitter, 1911. Stem: Dirhag-.

Dirotognathus G.H. Horn in J.L. LeConte and G.H. Horn, 1876: 79. Current status: valid genus in CURCULIONIDAE: ENTIMINAE: ENTIMITAE: BYRSOPAGINI: BYRSOPAGINA.

Type species: Dirotognathus sordidus G.H. Horn, 1876, by monotypy.

Family-group name: DIROTOGNATHINI G.H. Horn, 1876. Stem: Dirotognath-.

Disarthricerus Raffray, 1895: 78. Current status: valid genus in STAPHYLINIDAE: PSELAPHINAE: CLAVIGERITAE: DISARTHRICERINI.

Type species: Disarthricerus integer Raffray, 1895, by monotypy.

Family-group name: DIARTHRICERINI Jeannel, 1949. Stem: Disarthricer-.

Discheramocephalus C. Johnson, 2007b: 223. Current status: valid genus in PTILIIDAE: PTILIINAE: DISCHERAMOCEPHALINI.

Type species: Discheramocephalus semisulcatus C. Johnson, 2007, by original designation.

Family-group name: DISCHERAMOCEPHALINI Grebennikov, 2009. Stem: Discheramocephal-.

Discoloma Erichson, 1845a: 292. Current status: valid genus in DISCOLOMATIDAE: DISCOLOMATINAE.

Type species: Discoloma parmula Erichson, 1845, by monotypy.

Family-group name: DISCOLOMIDAE G.H. Horn, 1878. Stem: Discolomat-.

Discoptera Semenov, 1889a: 396. Current status: valid genus in CARABIDAE: HARPALINAE: CORSYRINI.

Type species: Discoptera komarowi Semenov, 1889, by subsequent designation (Kryzhanovskij et al. 1995: 160).

Family-group name: DISCOPTERINI Jedlička, 1941. Stem: Discopter-.

Discotenes Labram & Imhoff, 1841: [Heft 8]. Current status: valid genus in ANTHRIBIDAE: ANTHRIBINAE: DISCOTENINI.

Type species: Discotenes coelebs Labram & Imhoff, 1841, by monotypy.

Family-group name: DISCOTÉNIDES Lacordaire, 1865. Stem: Discoten-.

Discotoma Mulsant, 1850: 215. Current status: valid genus in COCCINELLIDAE: COCCINELLINAE: COCCINELLINI.

Type species: Discotoma ornata Mulsant, 1850, by monotypy.

Family-group name: DISCOTOMAIRES Mulsant, 1850. Stem: Discotom-.

Disker Dreux & Voisin, 1987: 314. Current status: valid genus in CURCULIONIDAE: ENTIMINAE: OTIORHYNCHITAE: ECTEMNORHININI.

Type species: Antarctonesiodes tenuicornis Jeannel, 1940, by original designation.

Family-group name: DISKERINI Voisin, Chapelin-Viscardi, Ponel & Rapp, 2017: 134. Stem: Disker-.

Dismorpha Gistel, 1848b: x. Current status: valid genus in BUPRESTIDAE: AGRILINAE: CORAEBINI: DISMORPHINA. Note: new replacement name for Stenogaster Solier, 1833.

Type species: Buprestis atomaria Fabricius, 1792, by monotypy (type fixation for Stenogaster Solier).

Family-group name: DISMORPHINA Cobos, 1990. Stem: Dismorph-.

Disonycha Chevrolat in Dejean, 1836a: 390. Current status: valid genus in CHRYSOMELIDAE: GALERUCINAE: ALTICINI.

Type species: Crioceris collata Fabricius, 1801, by subsequent designation (Blake 1933: 1).

Family-group name: DISONYCHAE J.L. LeConte & G.H. Horn, 1883. Stem: Disonych-.

Disphericus G.R. Waterhouse, 1841b: 298. Current status: valid genus in CARABIDAE: HARPALINAE: PELECIINI: PELECIINA.

Type species: Disphericus gambianus G.R. Waterhouse, 1841, by monotypy.

Family-group name: DISPHAERICINI Sloane, 1923. Stem: Dispheric-.

Dissonomus Jacquelin du Val, 1861: 280. Current status: valid genus in TENEBRIONIDAE: TENEBRIONINAE: DISSONOMINI. Note: new replacement name for Heterophylus Mulsant & Rey, 1859.

Type species: Heliopates picipes Faldermann, 1837, by original designation. Note: the type fixation for for this genus was erroneously given as “by subsequent designation (Gebien 1938b: 397)” in Bouchard et al. (2021: 166).

Family-group name: DISSONOMINI G.S. Medvedev, 1968. Stem: Dissonom-.

Distenia Lepeletier & Audinet-Serville, 1828: 485. Current status: valid genus in DISTENIIDAE: DISTENIINI.

Type species: Distenia columbina Lepeletier & Audinet-Serville, 1828, by monotypy.

Family-group name: DISTENITAE J. Thomson, 1861. Stem: Disteni-.

†Distenorrhinoides Gratshev & Zherikhin, 2000a: 39. Current status: valid genus in NEMONYCHIDAE: †EOBELINAE: †DISTENORRHINOIDINI. Note: Lower Cretaceous (Spain).

Type species: †Distenorrhinoides simulator Gratshev & Zherikhin, 2000, by original designation.

Family-group name: †DISTENORRHINOIDINI Legalov, 2009. Stem: Distenorrhinoid-.

†Distenorrhinus Arnoldi, 1977: 170. Current status: valid genus in NEMONYCHIDAE: †BRENTHORRHININAE: †DISTENORRHININI. Note: Upper Jurassic (Kazakhstan).

Type species: †Distenorrhinus angulatus Arnoldi, 1977, by original designation.

Family-group name: †DISTENORRHININI Arnoldi, 1977. Stem: Distenorrhin-.

Disterna J. Thomson, 1864: 88. Current status: valid genus in CERAMBYCIDAE: LAMIINAE: ZYGOCERINI.

Type species: Zygocera bifasciata Pascoe, 1859, by monotypy.

Family-group name: DISTERNINAE Pascoe, 1871. Stem: Distern-.

Distichocera W. Kirby, 1819: 471. Current status: valid genus in CERAMBYCIDAE: CERAMBYCINAE: DISTICHOCERINI.

Type species: Distichocera maculicollis W. Kirby, 1819, by monotypy.

Family-group name: DISTICHOCERINAE Pascoe, 1867. Stem: Distichocer-.

Distipsidera Westwood, 1836: 251. Current status: valid genus in CICINDELIDAE: CICINDELINI: IRESIINA.

Type species: Distipsidera undulata Westwood, 1836, by monotypy.

Family-group name: DISTYPSIDERINI W. Horn, 1893. Stem: Distipsider-.

Ditomus Bonelli, 1810: Tab. Syn. Current status: valid genus in CARABIDAE: HARPALINAE: HARPALINI: DITOMINA.

Type species: Scarites calydonius Rossi, 1790, by subsequent designation (Hope 1838a: 81). Note: no species were originally included in this genus by Bonelli (1810); the first ones were the five species listed by Latreille (1816: 190) without question marks under the new replacement name Aristus Latreille, 1816 (see Madge 1975: 582).

Family-group name: DITOMICI Bonelli, 1810. Stem: Ditom-.

Ditropalia Casey, 1906: 263. Current status: junior synonym of Bolitochara Mannerheim, 1830 in STAPHYLINIDAE: ALEOCHARINAE: HOMALOTINI: BOLITOCHARINA.

Type species: Bolitochara bella Märkel, 1844, by subsequent designation (Fenyes 1919: 22).

Family-group name: DITROPALIINI Hatch, 1957. Stem: Ditropali-.

Ditylus Fischer, 1817: 469. Current status: valid genus in OEDEMERIDAE: OEDEMERINAE: DITYLINI.

Type species: Helops laevis Fabricius, 1787, by subsequent monotypy (Dejean 1821: 72). Note: Ditylus helopioides Fischer is often listed as type species of this genus (e.g., Švihla 2008: 361; Kubisz and Iwan 2020: 487) but that species was not described until 1821 [November] (not in 1817 as noted, but after Dejean (1821 [May])) and therefore is not an originally included species.

Family-group name: DITYLATES Mulsant, 1858. Stem: Dityl-.

Docohammus Aurivillius, 1908: 217. Current status: valid genus in CERAMBYCIDAE: LAMIINAE: LAMIINI.

Type species: Docohammus bennigseni Aurivillius, 1908, by monotypy.

Family-group name: DOCOHAMMIDI E.S. Dillon & L.S. Dillon, 1959. Stem: Docohamm-.

Dodecosis H.W. Bates, 1867: 27. Current status: valid genus in CERAMBYCIDAE: CERAMBYCINAE: DODECOSINI.

Type species: Dodecosis saperdina H.W. Bates, 1867, by monotypy.

Family-group name: DODECOSINI Aurivillius, 1912. Stem: Dodecos-.

Dolicaon Laporte, 1834c: 119. Current status: valid genus in STAPHYLINIDAE: PAEDERINAE: PAEDERINI: DOLICAONINA.

Type species: Dolicaon lathrobioides Laporte, 1834, by monotypy.

Family-group name: DOLICAONES Casey, 1905. Stem: Dolicaon-.

Dolichus Bonelli, 1810: Tab. Syn. Current status: valid genus in CARABIDAE: HARPALINAE: SPHODRINI: DOLICHINA.

Type species: Carabus flavicornis Fabricius, 1787 (= Carabus halensis Schaller, 1783), by subsequent designation (Latreille 1817a: 532). Note: no species were originally included in this genus by Bonelli (1810); the first ones were the two species listed in Latreille (1816: 194).

Family-group name: DOLICHIENS Brullé, 1834. Stem: Dolich-.

Doliema Pascoe, 1860b: 50. Current status: junior synonym of Adelina Dejean, 1835 in TENEBRIONIDAE: DIAPERINAE: DIAPERINI: ADELININA.

Type species: Doliema platisoides Pascoe, 1860, by monotypy.

Family-group name: DOLIEMINI Reitter, 1917. Stem: Doliem-.

Dolocerus Mulsant, 1863: 223, 230. Current status: junior synonym of Brachypteroma Heyden, 1863 in CERAMBYCIDAE: CERAMBYCINAE: BRACHYPTEROMINI.

Type species: Dolocerus reichii Mulsant, 1863, by monotypy.

Family-group name: DOLOCERINI Özdikmen in Albayati et al., 2016: 665. Stem: Dolocer-. Note: Özdikmen in Albayati et al. (2016: 665) proposed DOLOCERINI as a replacement for BRACHYPTEROMINI Sama, 2008 because the type genus of the latter name, Brachypteroma Heyden, 1863, was considered a junior synonym of Dolocerus Mulsant, 1863; however, Heyden’s Brachypteroma was published in June 1863 while Mulsant’s name was issued late in 1863 as it was recorded on 4 January 1864 by the ‘Bibliographie de la France’ (Bousquet 2016: 377); also, such nomenclatural action is contrary to Article 40.1 (ICZN 1999a), which states that when the name of a type genus of a nominal family-group taxon is considered to be a junior synonym of the name of another nominal genus, the family-group name is not to be replaced on that account alone.

Dolosus Dajoz, 1963: 91. Current status: junior synonym of Thyroderus Sharp, 1885 in CERYLONIDAE: CERYLONINAE.

Type species: Dolosus leleupi Dajoz, 1963, by original designation.

Family-group name: DOLOSIDAE Dajoz, 1963. Stem: Dolos-.

†Dominibrentus Poinar, 2009a: 51. Current status: valid genus in BRENTIDAE: BRENTINAE: TAPHRODERINI. Note: Miocene (Dominican amber).

Type species: †Dominibrentus leptus Poinar, 2009, by original designation.

Family-group name: †DOMINIBRENTINI Poinar, 2009. Stem: Dominibrent-.

Dominopteron Kazantsev, 2013a: 15. Current status: valid genus in LYCIDAE: LYCINAE: LEPTOLYCINI: DOMINOPTERINA.

Type species: Dominopteron hispaniolum Kazantsev, 2013, by original designation.

Family-group name: DOMINOPTERINI Kazantsev, 2013: 28. Stem: Dominopter-.

Donacia Fabricius, 1775: 195. Current status: valid genus in CHRYSOMELIDAE: DONACIINAE: DONACIINI.

Type species: Donacia crassipes Fabricius, 1775, by subsequent designation (Curtis 1834: pl. 494).

Family-group name: DONACIADAE W. Kirby, 1837. Stem: Donaci-.

Dorcacerus Dejean, 1821: 105. Current status: valid genus in CERAMBYCIDAE: CERAMBYCINAE: TRACHYDERINI: TRACHYDERINA.

Type species: Cerambyx barbatus Fabricius, 1775, by monotypy.

Family-group name: DORCACERINAE H.W. Bates, 1870. Stem: Dorcacer-.

Dorcadion Dalman in Schönherr, 1817: 397. Current status: valid genus in CERAMBYCIDAE: LAMIINAE: DORCADIONINI.

Type species [suggested]: Cerambyx glicyrrhizae Pallas, 1773, by subsequent designation (J. Thomson 1864: 43). Note: as pointed out by Bousquet (2008a: 619), the valid type species fixation is that of Blanchard in Audouin et al. (1841: pl. 68), who selected Cerambyx fuliginator Linnaeus, 1758, but that species is the type species of the genus Iberodorcadion Breuning, 1943; an application to the Commission is necessary to fix Cerambyx glicyrrhizae Pallas, 1773 as the type species of Dorcadion Dalman, 1817 to conserve current usage (e.g., Danilevsky 2020: 337).

Family-group name: DORCADIONINAE Swainson, 1840. Stem: Dorcadion-.

Dorcadodium Gistel, 1856: 263, 406. Status: senior synonym of the subgenus Carinatodorcadion Breuning, 1943 of the genus Dorcadion Dalman, 1817 in CERAMBYCIDAE: LAMIINAE: DORCADIONINI. Note: two spellings were originally used for this new taxon, including Dorcadodion (pp. 293, 296, 470); we act as First Revisers and selected Dorcadodium as the correct original spelling; some authors (e.g., Sama 2002: 88, Özdikmen 2012: 56) treat Dorcadodium Gistel, 1856 as a nomen oblitum, however, they did not specifically give evidence that the conditions of Article 23.9.1.2 were met (see such requirement in Article 23.9.2, ICZN 1999a); an application to the Commission is necessary to maintain Carinatodorcadion Breuning, 1943 as valid.

Type species: Lamia morio Fabricius, 1787 (= Cerambyx aethiops Scopoli, 1763), by subsequent designation (Vives and Alonso-Zarazaga 2000: 659).

Family-group name: DORCADODIIDAE Gistel, 1856. Stem: Dorcadodi-.

Dorcaschema Chevrolat, 1844: 110. Current status: valid genus in CERAMBYCIDAE: LAMIINAE: DORCASCHEMATINI.

Type species: Dorcaschema leptocera Chevrolat, 1844 (= Saperda alternata Say, 1824), by monotypy.

Family-group name: DORCASCHEMITAE J. Thomson, 1860. Stem: Dorcaschemat-.

Dorcasomus Audinet-Serville, 1834b: 12. Current status: valid genus in CERAMBYCIDAE: CERAMBYCINAE: DORCASOMINI.

Type species: Cerambyx ebulinus Fabricius, 1787, by monotypy.

Family-group name: DORCASOMIDES Lacordaire, 1868. Stem: Dorcasom-.

Dorcatoma Herbst, 1791: 103. Current status: valid genus in PTINIDAE: DORCATOMINAE. Note: the original spelling Dorkatoma was changed to Dorcatoma and Dorcatoma Herbst, 1791 was placed on the Official List of Generic Names in Zoology (ICZN 1995c, as Dorcatoma Herbst, 1792).

Type species: Dorcatoma dresdensis Herbst, 1791, by monotypy (see ICZN 1995c, as Dorkatoma dresdensis Herbst, 1792). Note: name placed on the Official List of Specific Names in Zoology (ICZN 1995c).

Family-group name: DORCATOMINA C.G. Thomson, 1859. Stem: Dorcatom-.

Dorcus W.S. MacLeay, 1819: 111. Current status: valid genus in LUCANIDAE: LUCANINAE: DORCINI.

Type species: Scarabaeus parallelipipedus Linnaeus, 1758, by subsequent designation (Thon 1836: 87).

Family-group name: DORCIDAE Parry, 1864. Stem: Dorc-.

Dorylogaster Wasmann, 1904: 625. Current status: valid genus in STAPHYLINIDAE: ALEOCHARINAE: DORYLOGASTRINI.

Type species: Dorylogaster longipes Wasmanm, 1904, by monotypy.

Family-group name: DORYLOGASTRINI Wasmann, 1916. Stem: Dorylogastr-.

Dorylomimus Wasmann, 1904: 620. Current status: valid genus in STAPHYLINIDAE: ALEOCHARINAE: DORYLOMIMINI.

Type species: Dorylomimus kohli Wasmann, 1904, by monotypy.

Family-group name: DORYLOMIMINI Wasmann, 1916. Stem: Dorylomim-.

Dorylophila Wasmann, 1904: 632. Current status: valid genus in STAPHYLINIDAE: ALEOCHARINAE: MIMANOMMATINI: DORYLOPHILINA.

Type species: Dorylophila rotundicollis Wasmann, 1904, by monotypy.

Family-group name: DORYLOPHILINI Fenyes, 1921. Stem: Dorylophil-.

Dorynota Chevrolat in Dejean, 1836a: 370. Current status: valid genus in CHRYSOMELIDAE: CASSIDINAE: DORYNOTINI.

Type species: Cassida bidens Fabricius, 1781, by subsequent designation (Duponchel and Chevrolat 1842b: 211).

Family-group name: DORYNOTINI Monrós & Viana, 1949. Stem: Dorynot-.

Doryphora Illiger, 1807b: 331. Current status: valid genus in CHRYSOMELIDAE: CHRYSOMELINAE: CHRYSOMELINI.

Type species: Chrysomela punctatissima G.-A. Olivier, 1791, by subsequent designation (Latreille 1810: 432).

Family-group name: DORYPHORES Motschulsky, 1860. Stem: Doryphor-. Note: junior homonym of DORYPHORI Fitzinger, 1843 (type genus Doryphorus Cuvier, 1829) in Reptilia.

Doryscelis Dejean, 1836b: 189. Current status: valid genus in SCARABAEIDAE: CETONIINAE: STENOTARSIINI: DORYSCELINA.

Type species: Cetonia calcarata Klug, 1833, by monotypy.

Family-group name: DORYSCELINA Schenkling, 1921. Stem: Doryscel-.

Dorytomus Germar, 1817b: 340. Current status: valid genus in CURCULIONIDAE: CURCULIONINAE: ELLESCINI.

Type species: Curculio vorax Fabricius, 1792 (= Curculio longimanus Forster, 1771), by subsequent designation (Westwood 1838c: 37).

Family-group name: DORYTOMINI Bedel, 1886. Stem: Dorytom-.

Doydirhynchus Dejean, 1821: 79. Current status: valid genus in CIMBERIDIDAE: DOYDIRHYNCHINI.

Type species: Rhynchites austriacus G.-A. Olivier, 1807, by monotypy.

Family-group name: DOYDIRHYNCHOIDEA Pierce, 1916. Stem: Doydirhynch-.

Drapetes Dejean, 1821: 34. Current status: valid genus in ELATERIDAE: LISSOMINAE: LISSOMINI.

Type species: Elater equestris Fabricius, 1798 (= Elater mordelloides Host, 1789), by monotypy.

Family-group name: DRAPETINI J.L. LeConte, 1863. Stem: Drapet-. Note: the name DRAPETINI Collin, 1961 (type genus Drapetis Meigen, 1822) in Diptera has been used with the correct stem (as DRAPETIDINI) in recent literature (e.g., Sinclair et al. 2022).

Drasterius Eschscholtz, 1829a: 33. Current status: valid genus in ELATERIDAE: AGRYPNINAE: OOPHORINI. Note: name placed on the Official List of Generic Names in Zoology (ICZN 1987b).

Type species: Elater bimaculatus Rossi, 1790, by subsequent designation (Curtis 1838a: 694, see Sánchez-Ruiz 1996: 48). Note: name placed on the Official List of Specific Names in Zoology (ICZN 1987b); the subsequent designation was attributed to Westwood (1838c: 26) in ICZN (1987b); however, it was published later than the fixation by (Curtis 1838a: 694) and as a result, an application to the Commission is necessary to change the entry on the Official List of Generic Names in Zoology.

Family-group name: DRASTERIINI Houlbert, 1912. Stem: Drasteri-. Note: senior homonym of DRASTERIINI Wiltshire, 1976 (type genus Drasteria Hübner, 1818) in Lepidoptera.

Drepanocerus W. Kirby, 1828: 521. Current status: valid genus in SCARABAEIDAE: SCARABAEINAE: ONITICELLINI: DREPANOCERINA.

Type species: Drepanocerus kirbyi Kirby, 1828 [as kirbii], by monotypy.

Family-group name: DRÈPANOCÉRIDES Van Lansberge, 1875. Stem: Drepanocer-.

Drepanoxenus Kistner & Watson, 1972: 2. Current status: valid genus in STAPHYLINIDAE: ALEOCHARINAE: DREPANOXENINI.

Type species: Drepanoxenus physogastra Kistner & Watson, 1972, by original designation.

Family-group name: DREPANOXENINI Kistner & Watson, 1972. Stem: Drepanoxen-.

Drilonius Kiesenwetter, 1874: 282. Current status: valid genus in OMETHIDAE: DRILONIINAE.

Type species: Drilonius striatulus Kiesenwetter, 1874, by monotypy.

Family-group name: DRILONIINAE Crowson, 1972. Stem: Driloni-.

Drilus G.-A. Olivier, 1790a: [23] 1. Current status: valid genus in ELATERIDAE: AGRYPNINAE: DRILINI.

Type species: Ptilinus flavescens Geoffroy, 1785, by subsequent designation (Latreille 1810: 426).

Family-group name: DRILITES Blanchard, 1845. Stem: Dril-.

Drimostoma Dejean, 1831: 694, 745. Current status: junior synonym of Caelostomus W.S. MacLeay, 1825 in CARABIDAE: HARPALINAE: DRIMOSTOMATINI.

Type species: Drimostoma striatocolle Dejean, 1831, by subsequent designation (Drapiez 1838: 579).

Family-group name: DRIMOSTOMIDAS Chaudoir, 1872. Stem: Drimostomat-.

Dromaeolus Kiesenwetter, 1858: 197. Current status: valid genus in EUCNEMIDAE: MACRAULACINAE: MACRAULACINI.

Type species: Eucnemis barnabita Comolli, 1837, by monotypy. Note: the author of the type species is usually given as “A. Villa & G.B. Villa” in the literature (e.g., Muona 2007: 86) because the name “Villa” is attached to the species’ name in Comolli (1837: 15); however, the contents of Comolli’s paper do not reveal that the brothers Villa were responsible for satisfying the criteria of availability (ICZN 1999a, Article 50.1.1).

Family-group name: DROMAEOLINI Beaulieu, 1919. Stem: Dromaeol-. Note: as pointed out by Bouchard and Bousquet (2020a: 87) the names FORNACINI Beaulieu, 1919 and DROMAEOLINI Beaulieu, 1919 take precedence over MACRAULACINAE/-INI Fleutiaux, 1923 and an application to the Commission is necessary to preserve usage of MACRAULACINAE/-INI as valid.

Dromica Dejean, 1826: 434. Current status: valid genus in CICINDELIDAE: CICINDELINI: DROMICINA.

Type species: Cicindela coarctata Dejean, 1822, by monotypy.

Family-group name: DROMICITAE J. Thomson, 1859. Stem: Dromic-. Note: senior homonym of DROMICINAE Cope, 1896 (type genus Dromicus Cocteau & Bibron, 1843) in Reptilia.

Dromius Bonelli, 1810: Tab. Syn. Current status: valid genus in CARABIDAE: HARPALINAE: LEBIINI: DROMIUSINA. Note: name placed on the Official List of Generic Names in Zoology (ICZN 2006b).

Type species: Carabus quadrimaculatus Linnaeus, 1758, by subsequent designation (Hope 1838a: 63; see ICZN 2006b). Note: name placed on the Official List of Specific Names in Zoology (ICZN 2006b).

Family-group name: DROMIEI Bonelli, 1810. Stem: Dromius- (ICZN 2006b).

Drosochrus Erichson, 1843b: 243. Current status: valid genus in TENEBRIONIDAE: BLAPTINAE: PEDININI: HELOPININA.

Type species: Drosochrus brunnipes Erichson, 1843, by subsequent designation (Gebien 1943: 910).

Family-group name: DROSOCHRINI C. Koch, 1958. Stem: Drosochr-.

Dryobia Gistel, 1856: 368. Current status: junior synonym of Dryophilus Chevrolat, 1832 in PTINIDAE: DRYOPHILINAE: DRYOPHILINI.

Type species: Dryophilus anobioides Chevrolat, 1832, by present designation.

Family-group name: DRYOBIADAE Gistel, 1856. Stem: Dryobi-. Note: nomen oblitum (see Bousquet et al. 2009: 45); senior homonym of DRYOBIINI Arnett, 1962 (type genus Dryobius J.L. LeConte, 1850) in Coleoptera: CERAMBYCIDAE.

Dryobius J.L. LeConte, 1850c: 23. Current status: valid genus in CERAMBYCIDAE: CERAMBYCINAE: DRYOBIINI.

Type species: Callidium sexfasciatum Say, 1824 (= Dryobius sexnotatus Linsley, 1957), by monotypy.

Family-group name: DRYOBIINI Arnett, 1962. Stem: Dryobi-. Note: junior homonym of DRYOBIIDAE Gistel, 1856 (type genus Dryobia Gistel, 1856) in Coleoptera: PTINIDAE.

Dryocoetes Eichhoff, 1864: 38, 45. Current status: valid genus in CURCULIONIDAE: SCOLYTINAE: DRYOCOETINI. Note: name placed on the Official List of Generic Names in Zoology (ICZN 1979b).

Type species: Bostrichus autographus Ratzeburg, 1837, by subsequent designation (Hopkins 1914: 121; see ICZN 1979b). Note: name placed on the Official List of Specific Names in Zoology (ICZN 1979b).

Family-group name: DRYOCOETOIDEAE Lindemann, 1877. Stem: Dryocoet-.

Dryoctenes Audinet-Serville, 1835a: 27. Current status: valid genus in CERAMBYCIDAE: LAMIINAE: ACANTHODERINI.

Type species: Dryoctenes caliginosus Audinet-Serville, 1835 (= Lamia scrupulosa Germar, 1823), by monotypy.

Family-group name: DRYOCTENITAE J. Thomson, 1860. Stem: Dryocten-.

Dryophilus Chevrolat, 1832a: pl. 3. Current status: valid genus in PTINIDAE: DRYOPHILINAE: DRYOPHILINI.

Type species: Dryophilus anobioides Chevrolat, 1832, by monotypy.

Family-group name: DRYOPHILIDAE Gistel, 1848. Stem: Dryophil-.

Dryophthorus Germar, 1823: 302. Current status: valid genus in CURCULIONIDAE: DRYOPHTHORINAE: DRYOPHTHORINI. Note: name placed on the Official List of Generic Names in Zoology (ICZN 1987c).

Type species: Curculio lymexylon Fabricius, 1792 (= Curculio corticalis Paykull, 1792), by monotypy (see ICZN 1987c). Note: Curculio corticalis Paykull, 1792 was placed on the Official List of Specific Names in Zoology (ICZN 1987c).

Family-group name: DRYOPHTHORIDES Schönherr, 1825. Stem: Dryophthor-.

Dryops G.-A. Olivier, 1791: 297. Current status: valid genus in DRYOPIDAE.

Type species: Dermestes auriculatus Geoffroy, 1785, by subsequent designation (Froriep 1827: 483).

Family-group name: DRYOPIDES Billberg, 1820. Stem: Dryop-.

Dryotribus G.H. Horn, 1873c: 432. Current status: valid genus in CURCULIONIDAE: COSSONINAE: DRYOTRIBINI.

Type species: Dryotribus mimeticus G.H. Horn, 1873, by monotypy.

Family-group name: DRYOTRIBI J.L. LeConte, 1876. Stem: Dryotrib-.

Drypta Latreille, 1797: 75. Current status: valid genus in CARABIDAE: HARPALINAE: DRYPTINI.

Type species: Carabus emarginatus Gmelin, 1790 (= Carabus dentatus Rossi, 1790), by subsequent designation (Latreille 1810: 426). Note: no species were originally included in this genus by Latreille (1797); the first ones were the two species listed by Fabricius (1801a: 230, 231).

Family-group name: DRYPTINAE Bonelli, 1810. Stem: Drypt-.

Dynamopus Semenov, 1896: 336. Current status: junior synonym of Orubesa Reitter, 1895 in SCARABAEIDAE: DYNAMOPODINAE: DYNAMOPODINI.

Type species: Dynamopus athleta Semenov, 1896, by monotypy.

Family-group name: DYNAMOPINAE Arrow, 1911. Stem: Dynamopod-.

Dynamostes Pascoe, 1857: 90. Current status: valid genus in DISTENIIDAE: DYNAMOSTINI.

Type species: Dynamostes audax Pascoe, 1857, by monotypy.

Family-group name: DYNAMOSTIDES Lacordaire, 1868. Stem: Dynamost-.

Dynastes W.S. MacLeay, 1819: 66. Current status: valid genus in SCARABAEIDAE: DYNASTINAE: DYNASTINI: DYNASTINA. Note: name placed on the Official List of Generic Names in Zoology (ICZN 2014a).

Type species: Scarabaeus hercules Linnaeus, 1758, by subsequent designation (K.W. Kirby 1825: 567; see ICZN 2014a). Note: name placed on the Official List of Specific Names in Zoology (ICZN 2014a).

Family-group name: DYNASTIDAE W.S. MacLeay, 1819. Stem: Dynast-.

Dysantes Pascoe, 1869a: 31. Current status: valid genus in TENEBRIONIDAE: TENEBRIONINAE: TOXICINI: DYSANTINA. Note: Dysantes Forster, 1869 [published in May] in Hymenoptera is a junior homonym of Dysantes Pascoe, 1869 [published 1 January] and not the opposite as noted in the literature (see Bouchard and Bousquet 2020a: 101).

Type species: Diceroderes elongatus L. Redtenbacher, 1868, by monotypy.

Family-group name: DYSANTINAE Gebien, 1922. Stem: Dysant-.

Dyscharachthis Blackburn, 1900: 216, 217. Current status: valid genus in EUCNEMIDAE: EUCNEMINAE: DYSCHARACHTHINI.

Type species: Dyscharachthis brevipennis Blackburn, 1900, by monotypy.

Family-group name: DYSCHARACHTHINI Muona, 1993. Stem: Dyscharachth-.

Dyscherus Chaudoir, 1855: 4, 21. Current status: valid genus in CARABIDAE: SCARITINAE: SCARITINI: DYSCHERINA.

Type species: Scarites costatus Klug, 1833, by monotypy.

Family-group name: DYSCHERINA Basilewsky, 1973. Stem: Dyscher-.

Dyschirius Bonelli, 1810: Tab. Syn. Current status: valid genus in CARABIDAE: SCARITINAE: DYSCHIRIINI.

Type species: Scarites thoracicus Rossi, 1790, by subsequent designation (Curtis 1831: pl. 354). Note: no species were originally included in this genus by Bonelli (1810); the first ones were the two species listed by Panzer (1813: 67).

Family-group name: DYSCHIRIINI Kolbe, 1880. Stem: Dyschiri-.

Dysides Perty, 1832: 113. Current status: valid genus in BOSTRICHIDAE: DYSIDINAE.

Type species: Dysides obscurus Perty, 1832, by monotypy.

Family-group name: DYSIDINI Lesne, 1894. Stem: Dysid-.

Dyslobus J.L. LeConte, 1869a: 380. Current status: valid genus in CURCULIONIDAE: ENTIMINAE: ENTIMITAE: BYRSOPAGINI: STRANGALIODINA.

Type species: Otiorhynchus segnis J.L. LeConte, 1857, by subsequent designation (G.H. Horn in J.L. LeConte and G.H. Horn 1876: 41).

Family-group name: DYSLOBINI J.L. LeConte, 1874. Stem: Dyslob-.

Dysmorphocerus Solier, 1849: 451. Current status: valid genus in CANTHARIDAE: DYSMORPHOCERINAE.

Type species: Dysmorphocerus blanchardii Solier, 1849, by monotypy.

Family-group name: DYSMORPHOCERINAE Brancucci, 1980. Stem: Dysmorphocer-.

Dystaxia J.L. LeConte, 1867: 385. Current status: valid genus in SCHIZOPODIDAE: DYSTAXIINI.

Type species: Dystaxia murrayi J.L. LeConte, 1867, by monotypy.

Family-group name: DYSTAXINI Théry, 1929. Stem: Dystaxi-.

Dystheamon Grouvelle, 1927: 141. Current status: valid genus in DISCOLOMATIDAE: NOTIOPHYGINAE: DYSTHEAMONINI.

Type species: Dystheamon aeneum Grouvelle, 1927, by monotypy.

Family-group name: DYSTHEAMONINI John, 1954. Stem: Dystheamon-.

Dytiscus Linnaeus, 1758: 342, 411. Current status: valid genus in DYTISCIDAE: DYTISCINAE: DYTISCINI. Note: name placed on the Official List of Generic Names in Zoology (ICZN 1961e).

Type species: Dytiscus marginalis Linnaeus, 1758, by subsequent designation (Latreille 1810: 426; see ICZN 1961e). Note: name placed on the Official List of Specific Names in Zoology (ICZN 1961e).

Family-group name: DYTICIDES Leach, 1815. Stem: Dytisc-.

Earophilus Gistel, 1856: 375. Current status: junior synonym of Cucujus Fabricius, 1775 in CUCUJIDAE: CUCUJINAE.

Type species: Cantharis sanguinolenta Linnaeus, 1767 (= Meloe cinnaberinus Scopoli, 1763), by monotypy.

Family-group name: EAROPHILIDAE Gistel, 1856. Stem: Earophil-.

Ebaeus Erichson, 1840b: 113. Current status: valid genus in MELYRIDAE: MALACHIINAE: MALACHIINI.

Type species: Cantharis pedicularia Linnaeus, 1758, by subsequent designation (C.G. Thomson 1859: 112).

Family-group name: EBAEINI Portevin, 1931. Stem: Ebae-.

Eboo Reid, 1993: 62. Current status: valid genus in CHRYSOMELIDAE: EUMOLPINAE: BROMIINI. Note: new replacement name for Odontionopa Erichson, 1842.

Type species: Colaspis viridula Erichson, 1842, by subsequent designation (Reid 1993: 62) (type fixation for Odontionopa Erichson).

Family-group name: EBOOINA Reid, 1993. Stem: Eboo-.

Eburia Lacordaire, 1830b: 177. Current status: valid genus in CERAMBYCIDAE: CERAMBYCINAE: EBURIINI.

Type species: Cerambyx quadrimaculatus Linnaeus, 1767, by subsequent designation (Hope 1844: 189).

Family-group name: ÉBURIITES Blanchard, 1845. Stem: Eburi-.

Eburniogaster Seevers, 1938: 424. Current status: valid genus in STAPHYLINIDAE: ALEOCHARINAE: COROTOCINI: EBURNIOGASTRINA.

Type species: Eburniogaster termitocolus Seevers, 1938, by original designation.

Family-group name: EBURNIOGASTRINA Jacobson, Kistner & Pasteels, 1986. Stem: Eburniogastr-.

†Eccoptarthrus Arnoldi, 1977: 169. Current status: valid genus in NEMONYCHIDAE: †ECCOPTARTHRINAE. Note: Upper Jurassic (Kazakhstan).

Type species: †Eccoptarthrus crassipes Arnoldi, 1977, by original designation.

Family-group name: †ECCOPTARTHRINI Arnoldi, 1977. Stem: Eccoptarthr-.

Eccoptomenus Chaudoir, 1850: 409. Current status: valid genus in CARABIDAE: HARPALINAE: CHLAENIINI: CHLAENIINA.

Type species: Chlaenius eximius Dejean, 1831, by subsequent designation (Basilewsky and Grundmann 1954: 247).

Family-group name: ECCOPTOMENINI Jeannel, 1949. Stem: Eccoptomen-.

Eccoptopterus Motschulsky, 1863: 515. Current status: valid genus in CURCULIONIDAE: SCOLYTINAE: XYLEBORINI.

Type species: Eccoptopterus sexspinosus Motschulsky, 1863 (= Scolytus spinosus G.-A. Olivier, 1795), by monotypy.

Family-group name: ECCOPTOPTERINI Kalshoven, 1959. Stem: Eccoptopter-.

†Eccoptothorax Arnoldi, 1977: 158. Current status: junior synonym of Procurculio Arnoldi, 1977 in NEMONYCHIDAE: †ECCOPTARTHRINAE. Note: Upper Jurassic (Kazakhstan).

Type species: †Eccoptothorax latipennis Arnoldi, 1977 (= Procurculio fortipes Arnoldi, 1977), by original designation.

Family-group name: †ECCOPTOTHORACINI Arnoldi, 1977. Stem: Eccoptothorac-.

Eccoptus Dejean, 1821: 86. Current status: invalid senior synonym of Zygops Schönherr, 1825 in CURCULIONIDAE: CONODERINAE: ZYGOPINI. Note: name suppressed for the Principle of Priority and placed on the Official Index of Rejected and Invalid Generic Names in Zoology (see ICZN 1987e).

Type species: Curculio strix G.-A. Olivier, 1791, by monotypy.

Family-group name: ECCOPTINAE Pierce, 1919. Stem: Eccopt-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

Ecelonerus Schönherr, 1839: 163. Current status: junior synonym of Dendropemon Schönherr, 1839 in ANTHRIBIDAE: ANTHRIBINAE: ECELONERINI. Note: First Reviser (ICZN 1999a, Article 24.2.2) for Dendropemon Schönherr, 1839 versus Ecelonerus Schönherr, 1839 is Zimmerman (1994: 83).

Type species: Ecelonerus subfasciatus Fåhraeus, 1839, by original designation.

Family-group name: ECÉLONÉRIDES Lacordaire, 1865. Stem: Eceloner-.

Echiaster Erichson, 1839b: 29. Current status: valid genus in STAPHYLINIDAE: PAEDERINAE: LATHROBIINI: ECHIASTERINA.

Type species: Echiaster longicollis Erichson, 1840, by subsequent designation (Duponchel 1841b: 57). Note: no species were originally included in this genus by Erichson (1839b); the first ones were the two species described by Erichson (1840a: 637).

Family-group name: ECHIASTERES Casey, 1905. Stem: Echiaster-.

Echinaspis Blatchley, 1922: 122. Current status: junior synonym of Microhyus J.L. LeConte, 1876 in CURCULIONIDAE: MOLYTINAE: CLEOGONINI. Note: junior homonym of Echinaspis Haeckel, 1881 [Protista].

Type species: Echinaspis wolcotti Blatchley, 1922 (= Microhyus setiger J.L. LeConte, 1876), by monotypy.

Family-group name: ECHINASPINI Blatchley, 1922. Stem: Echinaspid-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

Echinocnemus Schönherr, 1843: 315. Current status: valid genus in CURCULIONIDAE: BRACHYCERINAE: ERIRHININI.

Type species: Erirhinus squameus Boheman, 1836, by original designation.

Family-group name: ECHINOCNEMINA Legalov, 2020a: 321. Stem: Echinocnem-.

Echthrogaster Blackburn, 1900: 213. Current status: valid genus in EUCNEMIDAE: MACRAULACINAE: ECHTHROGASTERINI.

Type species: Echthrogaster lugubris Blackburn, 1900, by monotypy.

Family-group name: ECHTHROGASTERINI Cobos, 1965. Stem: Echthro­gaster-.

Ecitochara Wasmann, 1887: 404. Current status: valid genus in STAPHYLINIDAE: ALEOCHARINAE: ATHETINI: ECITOCHARINA.

Type species: Ecitochara fusicornis Wasmann, 1887, by monotypy.

Family-group name: ECITOCHARINI Seevers, 1965. Stem: Ecitochar-.

Ecitoclimax Borgmeier, 1934: 452. Current status: valid genus in STAPHYLINIDAE: OXYTELINAE: OXYTELINI.

Type species: Ecitoclimax tarsalis Borgmeier, 1934, by original designation.

Family-group name: ECITOCLIMACINI Borgmeier, 1934. Stem: Ecitoclimac-.

Ecitogaster Wasmann, 1900a: 404. Current status: valid genus in STAPHYLINIDAE: ALEOCHARINAE: ECITOGASTRINI.

Type species: Ecitogaster schmalzi Wasmann, 1900, by subsequent monotypy (Wasmann 1900b: 224).

Family-group name: ECITOGASTRINI Fenyes, 1919. Stem: Ecitogastr-.

†Eckfeldattagenus Háva & Wappler, 2014: 68. Current status: valid genus in DERMESTIDAE: ATTAGENINAE: †ECKFELDATTAGENINI. Note: Middle Eocene (Germany).

Type species: †Eckfeldattagenus eocenicus Háva & Wappler, 2014, by original designation.

Family-group name: †ECKFELDATTAGENINI Háva, 2015: 1. Stem: Eckfeldattagen-.

Ectatosia Pascoe, 1857: 109. Current status: valid genus in CERAMBYCIDAE: LAMIINAE: APOMECYNINI.

Type species: Ectatosia moorei Pascoe, 1857, by monotypy.

Family-group name: ECTATOSIIDES Lacordaire, 1872. Stem: Ectatosi-.

Ectemnorhinus G.R. Waterhouse, 1853: 184. Current status: valid genus in CURCULIONIDAE: ENTIMINAE: OTIORHYNCHITAE: ECTEMNORHININI.

Type species: Ectemnorhinus viridis G.R. Waterhouse, 1853, by monotypy.

Family-group name: ECTEMNORHINIDES Lacordaire, 1863. Stem: Ectemnorhin-.

Ectenessa H.W. Bates, 1885: 257. Current status: valid genus in CERAMBYCIDAE: CERAMBYCINAE: ECTENESSINI.

Type species: Ectenessa nitida H.W. Bates, 1885, by subsequent designation (Napp and Martins 1982: 371).

Family-group name: ECTENESSINI Martins, 1998. Stem: Ecteness-.

Ectrephes Pascoe, 1866d: xvi. Current status: valid genus in PTINIDAE: PTININAE: ECTREPHINI.

Type species: Ectrephes formicarum Pascoe, 1866, by monotypy.

Family-group name: ECTREPHIDAE Wasmann, 1894. Stem: Ectreph-.

Ectyche Pascoe, 1869a: 143. Current status: valid genus in TENEBRIONIDAE: DIAPERINAE: ECTYCHINI.

Type species: Ectyche erebea Pascoe, 1869, by monotypy.

Family-group name: ECTYCHINI Doyen, E.G. Matthews & Lawrence, 1990. Stem: Ectych-.

Ecyroschema J. Thomson, 1864: 48. Current status: valid genus in CERAMBYCIDAE: LAMIINAE: CEROPLESINI: CROSSOTINA.

Type species: Ecyroschema favosa J. Thomson, 1864, by original designation.

Family-group name: ÉCYROSCHÉMIDES Lacordaire, 1872. Stem: Ecyroschemat-.

Edrotes J.L. LeConte, 1851a: 140. Current status: valid genus in TENEBRIONIDAE: PIMELIINAE: EDROTINI.

Type species: Edrotes ventricosus J.L. LeConte, 1851, by monotypy.

Family-group name: ÉDROTIDES Lacordaire, 1859. Stem: Edrot-.

Edusa Chevrolat in Dejean, 1836a: 408. Current status: junior synonym of Edusella Chapuis, 1874 in CHRYSOMELIDAE: EUMOLPINAE: EUMOLPINI. Note: this name has precedence over Edusella Chapuis, which is currently used as valid (see Bousquet and Bouchard 2013a: 111); an application to the Commission is needed to conserve Edusella Chapuis, 1874 as valid.

Type species: Colaspis varipes Boisduval, 1835, by monotypy.

Family-group name: EDUSITES Chapuis, 1874. Stem: Edus-.

Edusella Chapuis, 1874: 309. Current status: valid genus in CHRYSOMELIDAE: EUMOLPINAE: EUMOLPINI.

Type species: Edusa suturalis Chapuis, 1874, by monotypy.

Family-group name: EDUSELLINI Clavareau, 1914. Stem: Edusell-.

Ega Laporte, 1834b: 93. Current status: valid genus in CARABIDAE: HARPALINAE: LACHNOPHORINI: LACHNOPHORINA.

Type species: Ega formicaria Laporte, 1834, by monotypy.

Family-group name: EGINI G.H. Horn, 1881. Stem: Eg-.

Egidyella Reitter, 1899: 284. Current status: valid genus in DERMESTIDAE: ATTAGENINAE incertae sedis.

Type species: Egidyella prophetea Reitter, 1899, by monotypy.

Family-group name: EPIDYELLINI Semenov-Tjan-Shansky, 1914. Stem: Egidyell-.

Egolia Erichson, 1842: 150. Current status: valid genus in TROGOSSITIDAE: EGOLIINAE.

Type species: Egolia variegata Erichson, 1842, by monotypy.

Family-group name: ÉGOLIIDES Lacordaire, 1854. Stem: Egoli-.

Egrius Pascoe, 1865b: 428. Current status: valid genus in CURCULIONIDAE: CEUTORHYNCHINAE: EGRIINI.

Type species: Egrius camelus Pascoe, 1865, by monotypy.

Family-group name: EGRIINI Pajni & Kohli, 1982. Stem: Egri-.

Eicolyctus J.R. Sahlberg, 1919: 4. Current status: junior synonym of Zavaljus Reitter, 1880 in EROTYLIDAE: XENOSCELINAE.

Type species: Cryptophagus brunneus Gyllenhal, 1808, by monotypy.

Family-group name: EICOLYCTINI Vogt, 1967. Stem: Eicolyct-.

Eidoreus Sharp in Blackburn and Sharp, 1885: 146. Current status: valid genus in EUPSILOBIIDAE.

Type species: Eidoreus minutus Sharp, 1885, by monotypy.

Family-group name: EIDOREINAE Sasaji, 1986. Stem: Eidore-.

Ekkoptogaster Herbst, 1793: 124. Current status: junior synonym of Scolytus Geoffroy, 1762 in CURCULIONIDAE: SCOLYTINAE: SCOLYTINI. Note: this name, as well as its unjustified emendation Eccoptogaster Erichson, 1836, were placed on the Official Index of Rejected and Invalid Generic Names in Zoology (ICZN 1963c).

Type species: Bostrichus scolytus Fabricius, 1775, by subsequent designation (Hopkins 1914: 121).

Family-group name: ECCOPTOGASTRIDAE Gistel, 1848. Stem: Ekkoptogastr-. Note: family- group name permanently invalid (ICZN 1999a, Article 39).

Elacatis Pascoe, 1860b: 52. Current status: valid genus in SALPINGIDAE: OTHNIINAE.

Type species: Elacatis delusa Pascoe, 1860, by monotypy.

Family-group name: ELACATIDAE Reitter, 1879. Stem: Elacat-. Note: junior homonym of ELACATINAE Gill, 1861 (type genus Elacate Cuvier, 1832) in Pisces.

Elachistarthron Notman, 1920: 715. Current status: junior synonym of Apheloglossa Casey, 1894 in STAPHYLINIDAE: ALEOCHARINAE: DIESTOTINI.

Type species: Elachistarthron ambiguum Notman, 1920, by monotypy.

Family-group name: ELACHISTARTHRONINI Notman, 1920. Stem: Elachistarthr-.

Elaphidion Audinet-Serville, 1834b: 66. Current status: valid genus in CERAMBYCIDAE: CERAMBYCINAE: ELAPHIDIINI.

Type species: Cerambyx spinicornis Drury, 1773, by subsequent designation (Newman 1840c: 6).

Family-group name: ELAPHIDIONITAE J. Thomson, 1864. Stem: Elaphidi-.

Elaphinis Burmeister, 1842: 595. Current status: valid genus in SCARABAEIDAE: CETONIINAE: CETONIINI: CETONIINA.

Type species: Cetonia cinerascens Fabricius, 1798 (= Scarabaeus cinereonebulosus DeGeer, 1778), by subsequent designation (Duponchel 1844b: 227).

Family-group name: ELAPHINI Schoch, 1894. Stem: Elaphin-.

Elaphocera Gené, 1836a: iii. Current status: valid genus in SCARABAEIDAE: MELOLONTHINAE: TANYPROCTINI: TANYPROCTINA. Note: this name was also introduced in the same year by Gené (1836b: 188) in volume 39 of ‘Memorie della Reale Accademia delle Scienze di Torino’; volume 5 (p. iii) of the ‘Annales de la Société Entomologique de France’ was published earlier.

Type species: Elaphocera obscura Gené, 1836 (= Melolontha emarginata Gyllenhal, 1817), by monotypy.

Family-group name: ELAPHOCERITAE Blanchard, 1851. Stem: Elaphocer-. Note: nomen oblitum (see A.B.T. Smith 2006: 168).

Elaphrus Fabricius, 1775: 227. Current status: valid genus in CARABIDAE: ELAPHRINAE.

Type species: Cicindela riparia Linnaeus, 1758, by subsequent designation (Latreille 1810: 425).

Family-group name: ELAPHRII Latreille, 1802. Stem: Elaphr-.

Elater Linnaeus, 1758: 342, 404. Current status: valid genus in ELATERIDAE: ELATERINAE: ELATERINI.

Type species: Elater ferrugineus Linnaeus, 1758, by subsequent designation (Latreille 1810: 426).

Family-group name: ELATERIDES Leach, 1815. Stem: Elater-.

†Electrapate Iablokoff-Khnzorian, 1962: 87. Current status: valid genus in SCHIZOPODIDAE: †ELECTRAPATINI. Note: Eocene (Lithuania).

Type species: †Electrapate martynovi Iablokoff-Khnzorian, 1962, by original designation.

Family-group name: †ELECTRAPATIDAE Iablokoff-Khnzorian, 1962. Stem: Electrapat-.

Electribius Crowson, 1973: 231. Current status: valid genus in ARTEMATOPODIDAE: ELECTRIBIINAE.

Type species: Electribius oligocenicus Crowson, 1973, by original designation.

Family-group name: ELECTROPOGONINI Crowson, 1975. Stem: Electribi-. Note: ELECTROPOGONINI used by Crowson (1975: 77) is a lapsus calami for ELECTRIBIIDAE (see Lawrence and Newton 1995: 850).

†Electrocoryssopus Legalov, 2023a: 50. Current status: valid genus in CURCULIONIDAE: CONODERINAE: †ELECTROCORYSSOPINI. Note: upper Eocene (Baltic amber).

Type species: †Electrocoryssopus andrushchenkoi Legalov, 2023, by original designation.

Family-group name: †ELECTROCORYSSOPINI Legalov, 2023a: 49. Stem: Electrocoryssop-.

†Electropteron Kazantsev, 2012a: 1. Current status: valid genus in LYCIDAE: LYCINAE: LEPTOLYCINI: †ELECTROPTERINA. Note: Miocene (Dominican amber).

Type species: †Electropteron avus Kazantsev, 2012, by original designation.

Family-group name: †ELECTROPTERINI Kazantsev, 2013a: 28. Stem: Electropter-.

†Electrorubesopsis Bai & B. Wang in Li et al., 2018: 348. Current status: valid genus in SCARABAEIDAE: †ELECTRORUBESOPSINAE. Note: mid-Cretaceous (Myanmar).

Type species: †Electrorubesopsis beuteli Bai & B. Wang, 2018, by original designation.

Family-group name: †ELECTRORUBESOPSINAE Bai & B. Wang in Li et al., 2018: 348. Stem: Electrorubesops-.

Eledona Latreille, 1797: 19. Current status: valid genus in TENEBRIONIDAE: TENEBRIONINAE: BOLITOPHAGINI.

Type species: Opatrum agricola Herbst, 1783, by subsequent monotypy (Latreille 1802b: 162).

Family-group name: ELEDONAEDES Billberg, 1820. Stem: Eledon-. Note: nomen oblitum (see Bouchard et al. 2011: 415); senior homonym of ELEDONINAE Grimpe, 1921 (type genus Eledone Leach, 1817) in Mollusca.

Eleia Gistel, 1848b: viii, xiv. Current status: junior synonym of Chrysomela Linnaeus, 1758 in CHRYSOMELIDAE: CHRYSOMELINAE: CHRYSOMELINI. Note: unnecessary new replacement name for Chrysomela Linnaeus, 1758.

Type species: Chrysomela populi Linnaeus, 1758, by subsequent designation (Latreille 1810: 432; see ICZN 1984c) (type fixation for Chrysomela Linnaeus).

Family-group name: ELEIAEIDAE Gistel, 1848. Stem: Elei-.

Elenophorus Dejean, 1821: 64. Current status: junior synonym of Leptoderis Billberg, 1820 in TENEBRIONIDAE: PIMELIINAE: ELENOPHORINI.

Type species: Tenebrio collaris Linnaeus, 1767, by monotypy.

Family-group name: ELÉNOPHORITES Solier, 1837. Stem: Elenophor-.

Eleodes Eschscholtz, 1829d: 8. Current status: valid genus in TENEBRIONIDAE: BLAPTINAE: AMPHIDORINI.

Type species: Eleodes dentipes Eschscholtz, 1829, by subsequent designation (Hope 1841a: 124).

Family-group name: ELEODIINI Blaisdell, 1909. Stem: Eleod-.

Eleodopsis Blaisdell, 1939: 52. Current status: junior synonym of Blapylis G.H. Horn, 1870 in TENEBRIONIDAE: BLAPTINAE: AMPHIDORINI.

Type species: Eleodopsis subvestita Blaisdell, 1939, by original designation.

Family-group name: ELEODOPSINAE Blaisdell, 1939. Stem: Eleodopse-.

Eletica Dejean, 1834: 224. Current status: valid genus in MELOIDAE: ELETICINAE: ELETICINI: ELETICINA.

Type species: Lytta rufa Fabricius, 1801 (= Lytta bipustulata Fabricius, 1801), by monotypy.

Family-group name: ELETICIDES Wellman, 1910. Stem: Eletic-.

Eleusis Laporte, 1834c: 131. Current status: valid genus in STAPHYLINIDAE: OSORIINAE: ELEUSININI.

Type species: Eleusis tibialis Laporte, 1834, by monotypy.

Family-group name: ELEUSININA Sharp, 1887. Stem: Eleusin-.

Eligmoderma J. Thomson, 1864: 246. Current status: valid genus in CERAMBYCIDAE: CERAMBYCINAE: ELIGMODERMINI.

Type species: Eligmoderma ibidionoides J. Thomson, 1864, by original designation.

Family-group name: ÉLIGMODERMIDES Lacordaire, 1868. Stem: Eligmoderm-.

Ellescus Dejean, 1821: 87. Current status: valid genus in CURCULIONIDAE: CURCULIONINAE: ELLESCINI.

Type species: Curculio scanicus Paykull, 1792, by subsequent designation (Schönherr 1835: 321).

Family-group name: ELLESCHINA C.G. Thomson, 1859. Stem: Ellesc-.

Elmis Latreille, 1802a: 398. Current status: valid genus in ELMIDAE: ELMINAE: ELMINI: ELMINA. Note: name placed on the Official List of Generic Names in Zoology (ICZN 1995e).

Type species: Elmis maugetii Latreille, 1802, by monotypy (see ICZN 1995e). Note: name placed on the Official List of Specific Names in Zoology (1995e).

Family-group name: ELMIDAE Curtis, 1830. Stem: Elm- (see ICZN 1995e).

Elodes Latreille, 1797: 44. Current status: valid genus in SCIRTIDAE: SCIRTINAE.

Type species: Cistela pallida Fabricius, 1775 (= Lampyris minuta Linnaeus, 1767), by subsequent designation (Latreille 1810: 426). Note: no species were originally included in this genus by Latreille (1797); the first ones were the two species listed by Latreille (1802b: 98).

Family-group name: ELODIIDAE Shuckard, 1839. Stem: Elod-.

†Elodophthalmus Kirejtshuk & Azar, 2008: 25. Current status: valid genus in †ELODOPHTHALMIDAE. Note: Lower Cretaceous (Lebanese amber).

Type species: †Elodophthalmus harmonicus Kirejtshuk & Azar, 2008, by original designation.

Family-group name: †ELODOPHTHALMIDAE Kirejtshuk & Azar, 2008. Stem: Elodophthalm-.

Elosoma Motschulsky, 1845b: 33. Current status: valid genus in MELYRIDAE: MALACHIINAE incertae sedis.

Type species: Elosoma persica Motschulsky, 1845, by monotypy.

Family-group name: ELOSOMATIDAE Jakobson, 1915. Stem: Elosomat-.

Elpidus Péringuey, 1907: 314, 318. Current status: valid genus in SCARABAEIDAE: CETONIINAE: TRICHIINI.

Type species: Xiphoscelis hopei Burmeister, 1842, by monotypy.

Family-group name: ELPIDIDES Péringuey, 1907. Stem: Elpid-.

Elythomerus C.O. Waterhouse, 1874: 535. Current status: valid genus in HETEROCERIDAE: ELYTHOMERINAE.

Type species: Elythomerus elongatulus C.O. Waterhouse, 1874, by monotypy.

Family-group name: ELYTHOMERINI M.F. Pacheco, 1964. Stem: Elythomer-.

Elytracantha Lane, 1955: 281. Current status: invalid senior synonym of Elytracanthina Monné, 2005 in CERAMBYCIDAE: LAMIINAE: ELYTRACANTHININI. Note: junior homonym of Elytracantha Kleine, 1915 [Coleoptera: BRENTIDAE].

Type species: Elytracantha pugionata Lane, 1955, by monotypy.

Family-group name: ELYTRACANTHINAE Lane, 1955. Stem: Elytracanth-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

Elytracanthina Monné, 2005: 428. Current status: valid genus in CERAMBYCIDAE: LAMIINAE: ELYTRACANTHININI. Note: new replacement name for Elytracantha Lane, 1955.

Type species: Elytracantha pugionata Lane, 1955, by monotypy (type fixation for Elytracantha Lane).

Family-group name: ELYTRACANTHININI Bousquet, 2009. Stem: Elytracanthin-.

Elytroxys Jekel, 1856: 9bis. Current status: valid genus in CURCULIONIDAE: ENTIMINAE: ENTIMITAE: LORDOPINI.

Type species: Elytroxys inflatus Lacordaire, 1863, by subsequent monotypy (Lacordaire 1863: 275). Note: no species were originally included in this genus by Jekel (1856); the first one was included by Lacordaire (1863: 275).

Family-group name: ELYTROXYSI Jekel, 1856. Stem: Elytroxe-.

Elytrurus Boisduval, 1835: 400. Current status: valid genus in CURCULIONIDAE: ENTIMINAE: OTIORHYNCHITAE: ELYTRURINI.

Type species: Elytrurus lapeyrousei Boisduval, 1835, by subsequent designation (Schönherr 1842b: 239).

Family-group name: ELYTRURINI G.A.K. Marshall, 1956. Stem: Elytrur-.

Embrithes Schönherr, 1842b: 392. Current status: valid genus in CURCULIONIDAE: ENTIMINAE: CYPHICERITAE: EMBRITHINI.

Type species: Embrithes agnatus Boheman, 1842, by original designation.

Family-group name: EMBRITHINI G.A.K. Marshall, 1942. Stem: Embrith-.

Emelinus Casey, 1895: 773, 777. Current status: valid genus in ADERIDAE.

Type species: Xylophilus melsheimeri J.L. LeConte, 1855, by subsequent designation (Werner 1990: 212).

Family-group name: EMELININI Báguena Corella, 1948. Stem: Emelin-.

Emma Escalera, 1913: 318. Current status: invalid senior synonym of Emmita Escalera, 1914 in DASCILLIDAE: KARUMIINAE: EMMITINI. Note: junior homonym of Emma J.E. Gray, 1843 [Bryozoa].

Type species: Emma serricornis Escalera, 1913, by monotypy.

Family-group name: EMMINAE Escalera, 1913. Stem: Emm-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

Emmalus Erichson, 1843b: 251. Current status: valid genus in TENEBRIONIDAE: BLAPTINAE: OPATRINI: AMMOBIINA.

Type species: Emmalus pilosus Erichson, 1843, by monotypy.

Family-group name: EMMALLINA C. Koch, 1956. Stem: Emmal-.

Emmita Escalera, 1914: 226. Current status: valid genus in DASCILLIDAE: KARUMIINAE: EMMITINI. Note: new replacement name for Emma Escalera 1913.

Type species: Emma serricornis Escalera, 1913, by monotypy (type fixation for Emma Escalera).

Family-group name: EMMITINAE Escalera, 1914. Stem: Emmit-.

Empelus J.L. LeConte, 1861: 52. Current status: valid genus in STAPHYLINIDAE: EMPELINAE.

Type species: Litochrus brunnipennis Mannerheim, 1852, by original designation.

Family-group name: EMPELINAE Newton & Thayer, 1992. Stem: Empel-.

Emphyastes Mannerheim, 1852: 353. Current status: valid genus in CURCULIONIDAE: MOLYTINAE: EMPHYASTINI.

Type species: Emphyastes fucicola Mannerheim, 1852, by monotypy.

Family-group name: EMPHIASTIDES Lacordaire, 1863. Stem: Emphyast-.

Emphylus Erichson, 1846: 346. Current status: junior synonym of Spavius Motschulsky, 1844 in CRYPTOPHAGIDAE: CRYPTOPHAGINAE: CRYPTOPHAGINI.

Type species: Cryptophagus glaber Gyllenhal, 1808, by monotypy.

Family-group name: EMPHYLI Casey, 1900. Stem: Emphyl-.

Emphytoecia Fairmaire & Germain, 1860b: 529. Current status: valid genus in CERAMBYCIDAE: LAMIINAE: PTEROPLIINI.

Type species: Agapanthia suturella Blanchard, 1851, by subsequent designation (J. Thomson 1864: 115).

Family-group name: EMPHYTOECIIDES Lacordaire, 1872. Stem: Emphyto­eci-.

Empocryptus Sharp, 1900: 593. Current status: valid genus in EROTYLIDAE: CRYPTOPHILINAE: EMPOCRYPTINI.

Type species: Empocryptus ovalis Sharp, 1900, by monotypy.

Family-group name: EMPOCRYPTINI Leschen, 2003. Stem: Empocrypt-.

Enaria Erichson, 1847b: 657. Current status: valid genus in SCARABAEIDAE: MELOLONTHINAE: MELOLONTHINI: ENARIINA.

Type species: Melolontha melanictera Klug, 1833, by subsequent monotypy (Blanchard in H. Milne-Edwards et al. 1851: 152).

Family-group name: ENARINA Dewailly, 1950. Stem: Enari-.

Encaustes Lacordaire, 1842a: 30, 33. Current status: valid genus in EROTYLIDAE: EROTYLINAE: ENCAUSTINI.

Type species: Engis verticalis W.S. MacLeay, 1825, by subsequent designation (Arrow 1925: 69).

Family-group name: ENCAUSTINI Crotch, 1876. Stem: Encaust-.

Enceladus Bonelli, 1813: 460. Current status: valid genus in CARABIDAE: SIAGONINAE: ENCELADINI.

Type species: Enceladus gigas Bonelli, 1813 [as gygas], by monotypy.

Family-group name: ENCELADINI G.H. Horn, 1881. Stem: Encelad-.

Enchoptera Saunders, 1850: 76. Current status: valid genus in CERAMBYCIDAE: CERAMBYCINAE: MACRONINI.

Type species: Enchoptera apicalis Saunders, 1850, by subsequent designation (J. Thomson 1864: 139).

Family-group name: ENCHAPTERITAE J. Thomson, 1861. Stem: Enchopter-.

Encrates Gistel, 1848b: ix, 127. Current status: junior synonym of Lebia Latreille, 1802 in CARABIDAE: HARPALINAE: LEBIINI: LEBIINA. Note: unnecessary new replacement name for Lebia Latreille, 1802.

Type species: Carabus haemorrhoidalis Fabricius, 1787 (= Buprestis marginatus Geoffroy, 1785), by subsequent designation (E. Desmarest 1851: 76) (type fixation for Lebia Latreille).

Family-group name: ENCRATIDAE Gistel, 1856. Stem: Encrat-.

Encyclops Newman, 1838: 392. Current status: valid genus in CERAMBYCIDAE: LEPTURINAE: ENCYCLOPINI.

Type species: Encyclops pallipes Newman, 1838 (= Leptura caerulea Say, 1826), by monotypy.

Family-group name: ENCYCLOPINI J.L. LeConte, 1873. Stem: Encyclop-.

Endaeus Schönherr, 1826: 250. Current status: valid genus in CURCULIONIDAE: CURCULIONINAE: OCHYROMERINI.

Type species: Endaeus castus Schönherr, 1826, by original designation.

Family-group name: ENDAEINI Voss, 1958. Stem: Endae-.

Endecatomus Mellié, 1847: 108. Current status: valid genus in BOSTRICHIDAE: ENDECATOMINAE.

Type species: Anobium reticulatum Herbst, 1793, by monotypy.

Family-group name: ENDECATOMINI J.L. LeConte, 1861. Stem: Endecatom-.

Endocephalus Chevrolat in Dejean, 1836a: 412. Current status: valid genus in CHRYSOMELIDAE: EUMOLPINAE: EUMOLPINI.

Type species: Eumolpus bigatus Germar, 1823 (= Cryptocephalus lineatus Fabricius, 1775), by subsequent designation (Monrós and Bechyné 1956: 1127).

Family-group name: ENDOCÉPHALITES Chapuis, 1874. Stem: Endocephal-.

Endomia Laporte, 1840b: 259. Current status: valid genus in ANTHICIDAE: ANTHICINAE: ENDOMIINI.

Type species: Anthicus elongatissimus Laporte, 1840 (= Notoxus tenuicollis Rossi, 1792), by monotypy.

Family-group name: ENDOMIINI Kaszab, 1956. Stem: Endomi-.

Endomychus Panzer, 1795: 175. Current status: valid genus in ENDOMYCHIDAE: ENDOMYCHINAE.

Type species: Chrysomela coccinea Linnaeus, 1758, by subsequent designation (Latreille 1810: 432).

Family-group name: ENDOMYCHIDES Leach, 1815. Stem: Endomych-.

Endophloeus Dejean, 1834: 195. Current status: valid genus in ZOPHERIDAE: COLYDIINAE: SYNCHITINI.

Type species: Eledona spinosula Latreille, 1807 (= Silpha markovichiana Piller & Mitterpacher, 1783), by monotypy.

Family-group name: ENDOPHLOEIDAE Wollaston, 1868. Stem: Endophloe-.

Endroedyolus Scholtz & Howden, 1987: 92. Current status: valid genus in SCARABAEIDAE: SCARABAEINAE: ENDROEDYOLINI.

Type species: Endroedyolus paradoxus Scholtz & Howden, 1987, by original designation.

Family-group name: ENDROEDYOLINI Davis, Deschodt & Scholtz, 2019: 138. Stem: Endroedyol-.

Engis W.S. MacLeay, 1825: 41. Current status: invalid senior synonym of Encaustes Lacordaire, 1842 in EROTYLIDAE: EROTYLINAE: ENCAUSTINI. Note: junior homonym of Engis Paykull, 1800 [Coleoptera: EROTYLIDAE]; W.S. MacLeay (1825: 41) added “Nobis” after the name Engis indicating that he was creating a new genus despite the fact that he was aware of Engis Paykull, 1800; therefore, we consider Engis W.S. MacLeay, 1825 as a junior homonym of Engis Paykull, 1800 as done by Węgrzynowicz (2007: 539), and not as a misidentification of Paykull’s genus as done by Bouchard et al. (2011: 358).

Type species: Engis verticalis W.S. MacLeay, 1825, by subsequent designation (Węgrzynowicz 2007: 539).

Family-group name: ENGIDAE W.S. MacLeay, 1825. Stem: Engid-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

Enhydrus Laporte, 1834c: 110. Current status: valid genus in GYRINIDAE: GYRININAE: DINEUTINI: ENHYDRUSINA. Note: the older genus-group name Enhydrus W.S. MacLeay, 1825 was suppressed for the Principles of Priority and Homonymy and placed on the Official Index of Rejected and Invalid Generic Names in Zoology while Enhydrus Laporte, 1834 was placed on the Official List of Generic Names in Zoology (ICZN 1964).

Type species: Gyrinus sulcatus Wiedemann, 1821, by monotypy (see ICZN 1964). Note: name placed on the Official List of Specific Names in Zoology (ICZN 1964).

Family-group name: ENHYDRINI Branden, 1882. Stem: Enhydrus- (ICZN 2012a).

Enicocerus Stephens, 1829a: 196. Current status: valid subgenus of Ochthebius Leach, 1815 in HYDRAENIDAE: OCHTHEBIINAE: OCHTHEBIINI.

Type species: Enicocerus viridiaeneus Stephens, 1829 (= Ochthebius exsculptus Germar, 1823), by monotypy.

Family-group name: ENICOCERINA Perkins, 1997. Stem: Enicocer-.

Enicodes G.R. Gray in Griffith and Pidgeon, 1831: pls 65, 73. Current status: valid genus in CERAMBYCIDAE: LAMIINAE: ENICODINI.

Type species: Cerambyx fichtelii Schreber, 1802, by monotypy.

Family-group name: ENICODITAE J. Thomson, 1864. Stem: Enicod-.

Enicopus Stephens, 1830: 318. Current status: valid genus in MELYRIDAE: DASYTINAE: ENICOPINI.

Type species: Lagria atra Fabricius, 1787, by monotypy. Note: the type species often reported for this genus (e.g., Mayor 2007b: 407) is Dermestes hirtus Linnaeus, 1767 as selected by Westwood (1838c: 28), but that species is not an originally included species; Enicopus ater (Fabricius, 1787) is currently included in the nominotypical subgenus of Enicopus Stephens, 1830.

Family-group name: HENICOPINI Escalera, 1927. Stem: Enicop- (ICZN 1999a, Article 29.5).

Enneapaussus Jeannel, 1946: 62. Current status: junior synonym of Anapaussus Wasmann, 1929 in CARABIDAE: PAUSSINAE: PAUSSINI: PAUSSINA.

Type species: Paussus howa Dohrn, 1881, by original designation.

Family-group name: ENNEAPAUSSINI Jeannel, 1946. Stem: Enneapauss-.

Ennearabdus Van Lansberge, 1874: 20. Current status: valid genus in SCARABAEIDAE: SCARABAEINAE: EUCRANIINI.

Type species: Onthophagus lobocephalus Harold, 1868, by monotypy.

Family-group name: ENNEARABDINI Pereira & Martínez, 1956. Stem: Ennearabd-.

Ennearthron Mellié, 1847: 110. Current status: valid genus in CIIDAE: CIINAE: CIINI.

Type species: Cis cornutus Gyllenhal, 1827, by subsequent designation (E. Desmarest 1860: 261). Note: no available species was originally included in this genus by Mellié (1847); the first ones are the species listed by Mellié (1849: 361, 362).

Family-group name: ENNEARTHRONINAE Chûjô, 1939. Stem: Ennearthr-.

Enochrus C.G. Thomson, 1859: 18. Current status: valid genus in HYDROPHILIDAE: ENOCHRINAE.

Type species: here fixed (under ICZN 1999a, Article 70.3.2) as Hydrophilus melanocephalus G.-A. Olivier, 1793, misidentified as Hydrophilus bicolor Fabricius, 1792 (sensu Gyllenhal, 1808) in the original designation by C.G. Thomson (1859: 18).

Family-group name: ENOCHRINAE Short & Fikáček, 2013: 741. Stem: Enochr-.

Enodognathus Benderitter, 1920: 112. Current status: valid genus in OCHODAEIDAE: OCHODAEINAE: ENODOGNATHINI.

Type species: Enodognathus gilleti Benderitter, 1920, by monotypy.

Family-group name: ENDOGNATHINI Scholtz, 1988. Stem: Enodognath-.

Enoicus Péringuey, 1896: 568, 581. Current status: valid genus in CARABIDAE: HARPALINAE: ENOICINI.

Type species: Euleptus fallax Péringuey, 1896, by monotypy.

Family-group name: ENOICINI Basilewsky, 1985. Stem: Enoic-.

Enoplium Latreille, 1802b: 111. Current status: valid genus in CLERIDAE: KORYNETINAE.

Type species: Tillus serraticornis G.-A. Olivier, 1790, by monotypy.

Family-group name: ENOPLIIDAE Gistel, 1848. Stem: Enopli-.

Enoplocerus Audinet-Serville, 1832: 146. Current status: valid genus in CERAMBYCIDAE: PRIONINAE: CALLIPOGONINI.

Type species: Cerambyx armillatus Linnaeus, 1767, by monotypy.

Family-group name: ENOPLOCERITAE J. Thomson, 1864. Stem: Enoplocer-.

Enoplopus Solier, 1848b: 151, 158. Current status: junior synonym of Accanthopus Dejean, 1821 in TENEBRIONIDAE: TENEBRIONINAE: HELOPINI: ENOPLOPODINA. Note: unnecessary new replacement name for Accanthopus Dejean, 1821.

Type species: fixed by Bouchard et al. (2021: 174) according to Article 70.3.2 (ICZN 1999a) as Tenebrio velikensis Piller & Mitterpacher, 1783, the taxonomic species involved in the misidentification of Tenebrio caraboides Linnaeus, 1758 (sensu Germar, 1817), as designated by monotypy by Dejean (1821: 71) (type fixation for Accanthopus Dejean).

Family-group name: ENOPLOPINI Reitter, 1917. Stem: Enoplopod-.

Enoplotrupes P.H. Lucas, 1869: xiii. Current status: valid genus in GEOTRUPIDAE: GEOTRUPINAE: ENOPLOTRUPINI.

Type species: Enoplotrupes sinensis P.H. Lucas, 1869, by monotypy.

Family-group name: ENOPLOTRUPINI Paulian, 1945. Stem: Enoplotrup-.

Enoptes Gistel, 1848b: x. Current status: junior synonym of Pytho Latreille, 1797 in PYTHIDAE. Note: unnecessary new replacement name for Pytho Latreille, 1797.

Type species: Cucujus coeruleus Fabricius, 1787 (= Tenebrio depressus Linnaeus, 1767), by subsequent designation (Latreille 1810: 429) (type fixation for Pytho Latreille).

Family-group name: ENOPTISIDAE Gistel, 1848. Stem: Enopt-.

Entimus Germar, 1817b: 341. Current status: valid genus in CURCULIONIDAE: ENTIMINAE: ENTIMITAE: ENTIMINI: ENTIMINA.

Type species: Curculio imperialis Forster, 1771, by subsequent designation (Schönherr 1823: 1138).

Family-group name: ENTIMIDES Schönherr, 1823. Stem: Entim-.

Entomoculia Croissandeau, 1891: 150. Current status: valid genus in STAPHYLINIDAE: LEPTOTYPHLINAE: ENTOMOCULIINI.

Type species: fixed by Herman (2001: 6) according to Article 70.3.2 (ICZN 1999a) as Entomoculia jeanneli Coiffait, 1955, the taxonomic species involved in the misidentification of Leptotyphlus grouvellei Fauvel, 1890 (sensu Croissandeau, 1891), as designated subsequently by Blackwelder (1952: 149).

Family-group name: ENTOMOCULINI Coiffait, 1957. Stem: Entomoculi-.

Entomosatopus Bonvouloir, 1871: 67, 279. Current status: valid genus in EUCNEMIDAE: EUCNEMINAE: ENTOMOSATOPINI.

Type species: Entomosatopus curtus Bonvouloir, 1871, by monotypy.

Family-group name: ENTOMOSATOPINI Muona, 1993. Stem: Entomosatop-.

Entomoscelis Chevrolat in Dejean, 1836a: 402. Current status: valid genus in CHRYSOMELIDAE: CHRYSOMELINAE: CHRYSOMELINI.

Type species: Chrysomela adonidis Pallas, 1771, by subsequent designation (Chevrolat 1843a: 654).

Family-group name: ENTOMOSCÉLITES Chapuis, 1874. Stem: Entomoscelid-.

†Eobelus Arnoldi, 1977: 147. Current status: valid genus in NEMONYCHIDAE: †EOBELINAE: †EOBELINI. Note: Upper Jurassic (Kazakhstan).

Type species: †Eobelus longipes Arnoldi, 1977, by original designation.

Family-group name: †EOBELIDAE Arnoldi, 1977. Stem: Eobel-.

†Eocaenonemonyx Legalov, 2013a: 58. Current status: valid genus in NEMONYCHIDAE: †CRETONEMONYCHINAE: †EOCAENONEMONYCHINI. Note: middle Eocene (USA).

Type species: †Eocaenonemonyx kuscheli Legalov, 2013, by original designation.

Family-group name: †EOCAENONEMONYCHINI Legalov, 2013a: 58. Stem: Eocaenonemonych-.

Eocatops Peyerimhoff, 1924: 63. Current status: valid genus in LEIODIDAE: CHOLEVINAE: ANEMADINI: EOCATOPINA.

Type species: Eocatops ambiguus Peyerimhoff, 1924, by original designation.

Family-group name: EOCATOPINA Jeannel, 1936. Stem: Eocatop-.

†Eocenesibinia Legalov, 2017b: 984. Current status: valid genus in CURCULIONIDAE: CURCULIONINAE: TYCHIINI: †EOCENESIBINIINA. Note: upper Eocene (Baltic amber).

Type species: †Eocenesibinia prussica Legalov, 2017, by original designation.

Family-group name: †EOCENESIBINIINA Legalov, 2017b: 984. Stem: Eocenesibini-.

Eoclytra Monrós, 1958: 36. Current status: junior synonym of Diapericera Lacordaire, 1848 in CHRYSOMELIDAE: CRYPTOCEPHALINAE: CLYTRINI: EOCLYTRINA.

Type species: Eoclytra freudei Monrós, 1958, by original designation.

Family-group name: EOCLYTRINI Monrós, 1958. Stem: Eoclytr-.

†Eodromeus Ponomarenko, 1977: 66. Current status: valid genus in TRACHYPACHIDAE: †EODROMEINAE. Note: Middle-Upper Jurassic/Lower Cretaceous (China; Kazakhstan; Mongolia; Russia).

Type species: †Eodromeus dissectus Ponomarenko, 1977 [as Eodromeus fasciatus], by original designation. Note: see Bouchard et al. (2011: 101) for a discussion regarding the type species.

Family-group name: †EODROMEINAE Ponomarenko, 1977. Stem: Eodrome-.

Eohomopterus Wasmann, 1920: 110. Current status: valid genus in CARABIDAE: PAUSSINAE: PAUSSINI: CARABIDOMEMNINA.

Type species: Homopterus aequatoriensis Wasmann, 1899, by monotypy.

Family-group name: EOHOMOPTERINAE Wasmann, 1929. Stem: Eohomopter-.

†Eopaussus Wasmann, 1926: 29. Current status: valid genus in CARABIDAE: PAUSSINAE: PAUSSINI: †EOPAUSSINA. Note: Eocene (Russia).

Type species: †Eopaussus balticus Wasmann, 1926, by monotypy.

Family-group name: †EOPAUSSINAE Luna de Carvalho, 1951. Stem: Eopauss-.

Eorhipidius Iablokoff-Khnzorian, 1986: 89. Current status: junior synonym of Pterydrias Reitter, 1895 in RIPIPHORIDAE: RIPIDIINAE: EORHIPIDIINI.

Type species: Eorhipidius januschevi Iablokoff-Khnzorian, 1986, by monotypy.

Family-group name: EORHIPIDIINI Iablokoff-Khnzorian, 1986. Stem: Eorhipidi-.

Eosalacus Legalov, 2007: 30. Current status: junior synonym of Pseudominurus Voss, 1956 in ATTELABIDAE: RHYNCHITINAE: RHYNCHITITAE: AULETINI: MANDELSCHTAMIINA.

Type species: Eosalacus reunionensis Legalov, 2007, by original designation.

Family-group name: EOSALACINA Legalov, 2007. Stem: Eosalac-.

Eospasta Selander, 1966: 474. Current status: valid genus in MELOIDAE: ELETICINAE: ELETICINI: EOSPASTINA.

Type species: Spastica inconstans Fairmaire & Germain, 1863, by original designation.

Family-group name: EOSPASTINA Selander, 1966. Stem: Eospast-.

Epactius D.H. Schneider in Fabricius, 1791: 23. Current status: senior synonym of Omophron Latreille, 1802 in CARABIDAE: OMOPHRONINAE. Note: new replacement name for Scolytus Fabricius, 1791; this name should have priority over Omophron Latreille, 1802 and an application to the Commission is needed to retain Latreille’s well-established name; Reversal of Precedence (ICZN 1999a, Article 23.9) cannot be considered since Epactius has been used as valid at least once since 1900 (e.g., Semenov 1903a).

Type species: Carabus limbatus Fabricius, 1777, by subsequent designation (Latreille 1810: 426) (type fixation for Omophron Latreille).

Family-group name: EPACTIINI Fauvel, 1888. Stem: Epacti-.

Epactoides Olsouffief, 1947: 171. Current status: valid genus in SCARABAEIDAE: SCARABAEINAE: EPACTOIDINI.

Type species: Phacosoma madecassus Paulian, 1936, by monotypy.

Family-group name: EPACTOIDINI Rossini, Grebennikov, Merrien, Miraldo, Viljanen & Tarasov 2022: 661. Stem: Epactoid-.

Eparchium Bernhauer, 1934: 481. Current status: junior synonym of Apterodasytes Champion, 1923 in MELYRIDAE: DASYTINAE: DASYTINI.

Type species: Eparchium paradoxum Bernhauer, 1934, by original designation.

Family-group name: EPARCHIINI Bernhauer, 1934: 481. Stem: Eparchi-. Note: this name was overlooked in Bouchard et al. (2011), Bouchard and Bousquet (2020a).

Ephistemus Stephens, 1829a: 167. Current status: valid genus in CRYPTOPHAGIDAE: ATOMARIINAE: ATOMARIINI.

Type species: Dermestes gyrinoides Marsham, 1802 (= Dermestes globulus Paykull, 1798), by subsequent designation (Westwood 1838c: 10).

Family-group name: EPHISTEMINI Casey, 1900. Stem: Ephistem-.

Epicaerus Schönherr, 1834a: 323. Current status: valid genus in CURCULIONIDAE: ENTIMINAE: POLYDRUSITAE: GEONEMINI.

Type species: Epicaerus mexicanus Boheman, 1834, by original designation.

Family-group name: EPICAERI G.H. Horn, 1876. Stem: Epicaer-.

Epicasta J. Thomson, 1864: 90. Current status: valid genus in CERAMBYCIDAE: LAMIINAE: DESMIPHORINI.

Type species: Epicasta ocellata J. Thomson, 1864, by original designation.

Family-group name: ÉPICASTIDES Lacordaire, 1872. Stem: Epicast-.

Epicauta Dejean, 1834: 224. Current status: valid genus in MELOIDAE: MELOINAE: EPICAUTINI.

Type species [suggested]: Meloe erythrocephalus Pallas, 1771, by subsequent designation (Werner 1945: 425). Note: the valid type species fixation is that of Duponchel (1844b: 356), who selected Lytta gigas Fabricius, 1792 (= Cantharis gigas G.-A. Olivier, 1790), but that species is currently included in the genus Cyaneolytta Péringuey, 1909; an application to the Commission is necessary to fix Meloe erythrocephalus Pallas, 1771 as the type species of Epicauta Dejean, 1834.

Family-group name: EPICAUTINI J.B. Parker & Böving, 1924. Stem: Epicaut-.

Epiclines Chevrolat in Guérin-Méneville, 1839a: 40. Current status: valid genus in CLERIDAE: EPICLININAE.

Type species: Epiclines gayi Chevrolat, 1839, by monotypy.

Family-group name: EPICLININAE Gunter, Leavengood & Bartlett in Gunter et al., 2013: 633. Stem: Epiclin-.

Epilachna Chevrolat in Dejean, 1836a: 436. Current status: valid genus in COCCINELLIDAE: COCCINELLINAE: EPILACHNINI.

Type species: Coccinella borealis Fabricius, 1775, by subsequent designation (Hope 1841a: 157).

Family-group name: EPILACHNIENS Mulsant, 1846. Stem: Epilachn-.

Epilissus Dejean, 1836b: 151. Current status: valid genus in SCARABAEIDAE: SCARABAEINAE: EPILISSINI.

Type species: Canthon prasinus Klug, 1833, by subsequent designation (Reiche 1841: 212).

Family-group name: EPILISSIDES Van Lansberge, 1874. Stem: Epiliss-.

Epimetopus Lacordaire, 1854: 465, 467. Current status: valid genus in EPIMETOPIDAE. Note: new replacement name for Ceratoderus Mulsant, 1851.

Type species: Ceratoderus graniger Mulsant, 1851, by monotypy (type fixation for Ceratoderus Mulsant).

Family-group name: EPIMETOPINA Zaitzev, 1908. Stem: Epimetop-.

Epipedocera Chevrolat, 1863: 339. Current status: valid genus in CERAMBYCIDAE: CERAMBYCINAE: TILLOMORPHINI.

Type species: Epipedocera zona Chevrolat, 1863, by subsequent designation (Pascoe 1869b: 640).

Family-group name: EPIPEDOCERINI Gahan, 1906. Stem: Epipedocer-.

Epipedophyes G.A.K. Marshall, 1946: 97. Current status: valid genus in CURCULIONIDAE: MOLYTINAE: PETALOCHILINI. Note: new replacement name for Epipedus Schönherr, 1842.

Type species: Epipedus squamifer Boheman, 1842, by original designation (type fixation for Epipedus Schönherr).

Family-group name: EPIPEDOPHYINAE Kuschel, 1955. Stem: Epipedophy-.

Epipedus Schönherr, 1842a: 462. Current status: invalid senior synonym of Epipedophyes G.A.K. Marshall, 1946 in CURCULIONIDAE: MOLYTINAE: PETALOCHILINI. Note: junior homonym of Epipedus Spinola, 1837 [Hemiptera].

Type species: Epipedus squamifer Boheman, 1842, by original designation.

Family-group name: ÉPIPÉDIDES Lacordaire, 1865. Stem: Epiped-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

Epiphanis Eschscholtz, 1829a: 35. Current status: valid genus in EUCNEMIDAE: MELASINAE: EPIPHANINI.

Type species: Epiphanis cornutus Eschscholtz, 1829, by monotypy.

Family-group name: EPIPHANINI Muona, 1993. Stem: Epiphan-. Note: junior homonym of EPIPHANIDAE Harring, 1913 (type genus Epiphanes Ehrenberg, 1832) in Rotifera.

Epiphloeus Spinola, 1841: 75. Current status: valid genus in CLERIDAE: KORYNETINAE.

Type species: Epiphloeus pantherinus Spinola, 1845, by subsequent designation (E. Desmarest 1852: 265). Note: no species were originally included in this genus by Spinola (1841); the first ones were the six species listed by Spinola (1845b: 8, as Epiphlaeus).

Family-group name: EPIPHLÖINEN Kuwert, 1893. Stem: Epiphloe-.

Epiphysa Dejean, 1834: 178. Current status: valid genus in TENEBRIONIDAE: PIMELIINAE: ADESMIINI.

Type species: Pimelia flavicollis Fabricius, 1794, by monotypy.

Family-group name: ÉPIPHYSIDES Lacordaire, 1859. Stem: Epiphys-.

Epipocus Chevrolat in Dejean, 1836a: 439. Current status: valid genus in ENDOMYCHIDAE: EPIPOCINAE.

Type species: Endomychus rufitarsis Chevrolat, 1835, by subsequent designation (R. Lucas 1920: 270). Note: the type species often reported for Epipocus is Endomychus tibialis Guérin-Méneville, 1834 by subsequent designation of Strohecker (1953: 66) but R. Lucas’ typification is earlier; both species are currently placed in the genus Epipocus.

Family-group name: EPIPOCIDAE Gorham, 1873. Stem: Epipoc-.

Epirinus Dejean, 1833b: 137. Current status: valid genus in SCARABAEIDAE: SCARABAEINAE: EPIRININI.

Type species: Scarabaeus granulatus G.-A. Olivier, 1789 (= Scarabaeus flagellatus Fabricius, 1775), by subsequent designation (M.C. Ferreira 1972: 136). Note: M.C. Ferreira (1972: 136) designated Scarabaeus flagellatus Fabricius, 1775 as type species, not an originally included species, but since she listed it in synonymy with Scarabaeus granulatus G.-A. Olivier, 1789, a species originally included, she is deemed to have designated the latter species as type species (ICZN 1999a, Article 69.2.2; see Daniel 2019: 64).

Family-group name: EPIRINIDES Van Lansberge, 1874. Stem: Epirin-.

Epirrhynchus Schönherr, 1823: 1139. Current status: valid genus in CURCULIONIDAE: LIXINAE: CLEONINI.

Type species: Curculio argus Sparrman, 1785, by original designation.

Family-group name: EPIRHYNCHINI Petri, 1914. Stem: Epirrhynch-.

Episomus Schönherr, 1823: 1143. Current status: valid genus in CURCULIONIDAE: ENTIMINAE: OTIORHYNCHITAE: EPISOMINI.

Type species: Curculio lacerta Fabricius, 1781, by original designation.

Family-group name: EPISOMIDES Lacordaire, 1863. Stem: Episom-.

Epistictina Hincks, 1950b: 509. Current status: valid genus in CHRYSOMELIDAE: CASSIDINAE: BASIPRIONOTINI. Note: new replacement name for Epistictia Boheman, 1850.

Type species: Epistictia viridimaculata Boheman, 1850, by subsequent designation (Maulik 1919: 318) (type fixation for Epistictia Boheman).

Family-group name: EPISTICTININI Hincks, 1952. Stem: Epistictin-.

Epistomentis Solier, 1849: 479. Current status: valid genus in BUPRESTIDAE: BUPRESTINAE: EPISTOMENTINI.

Type species: Chrysochroa picta Gory, 1841, by monotypy.

Family-group name: EPISTOMENTINI Levey, 1978. Stem: Epistoment-.

Epistrophus Kirsch, 1869: 200. Current status: valid genus in CURCULIONIDAE: MOLYTINAE: HYLOBIINI: EPISTROPHINA. Note: this taxon is a junior homonym of Epistrophus Gistel, 1834, which is a junior objective synonym of Drasterius Eschscholtz, 1829 [Coleoptera: ELATERIDAE]; Alonso-Zarazaga and Lyal (1999: 202) mentioned that an application would be submitted to the Commission for the suppression of Gistel’s name and recommended that current usage be maintained; an application to the Commission has yet to be submitted to fix this problem.

Type species: Epistrophus tumidus Kirsch, 1869, by monotypy.

Family-group name: EPISTROPHINA G.A.K. Marshall, 1932. Stem: Epistroph-.

Epitragus Latreille, 1802b: 165. Current status: valid genus in TENEBRIONIDAE: PIMELIINAE: EPITRAGINI.

Type species: Epitragus fuscus Latreille, 1804, by subsequent monotypy (Latreille 1804b: 322). Note: Latreille (1802b) originally included “Helops variegatus? Fabricius” (= Tenebrio variegatus Fabricius, 1781) in his genus but since the species was cited as a species inquirenda, it is deemed not to be originally included (ICZN 1999a, Article 67.2.5).

Family-group name: ÉPITRAGITES Blanchard, 1845. Stem: Epitrag-.

Epiverta Dieke, 1947: 169. Current status: valid genus in COCCINELLIDAE: COCCINELLINAE: EPILACHNINI.

Type species: Solanophila chelonia Mader, 1933, by original designation.

Family-group name: EPIVERTINI X.-F. Pang & Mao, 1979. Stem: Epivert-.

Epuraea Erichson, 1843a: 267. Current status: valid genus in NITIDULIDAE: EPURAEINAE: EPURAEINI.

Type species: Nitidula silacea Herbst, 1783, by subsequent designation (C.G. Thomson 1859: 68).

Family-group name: EPURAEINI Kirejtshuk, 1986. Stem: Epurae-.

Eremazus Mulsant, 1851b: 139. Current status: valid genus in SCARABAEIDAE: EREMAZINAE: EREMAZINI.

Type species: Eremazus unistriatus Mulsant, 1851, by monotypy.

Family-group name: EREMAZINA Stebnicka, 1977. Stem: Eremaz-.

Eremnus Schönherr, 1823: 1145. Current status: valid genus in CURCULIONIDAE: ENTIMINAE: ENTIMITAE: TANYRHYNCHINI.

Type species: Eremnus exaratus Schönherr, 1823, by original designation.

Family-group name: ÉREMNIDES Lacordaire, 1863. Stem: Eremn-.

Eremochilus Weise, 1913: 117. Current status: valid genus in COCCINELLIDAE: COCCINELLINAE: EPILACHNINI.

Type species: Eremochilus peregrinus Weise, 1913, by monotypy.

Family-group name: EREMOCHILINI Gordon & Vanderberg, 1987. Stem: Eremochil-.

Eremosaprinus Ross, 1939: 39. Current status: valid genus in HISTERIDAE: SAPRININAE: EREMOSAPRININI.

Type species: Saprinus unguiculatus Ross, 1939, by original designation.

Family-group name: EREMOSAPRININI Lackner in Lackner et al. 2024: 59. Stem: Eremosaprin-.

Eremotes Wollaston, 1861c: 364. Current status: junior synonym of Rhyncolus Germar, 1817 in CURCULIONIDAE: COSSONINAE: RHYNCOLINI: RHYNCOLINA.

Type species: Hylurgus crassicornis Brullé, 1839, by monotypy.

Family-group name: EREMOTINI Voss, 1939. Stem: Eremot-.

Eremoxenus Semenov, 1892: 438. Current status: valid genus in BRENTIDAE: BRENTINAE: BRENTINI: EREMOXENINA.

Type species: Eremoxenus chan Semenov, 1892, by monotypy.

Family-group name: EREMOXENIDAE Semenov, 1892. Stem: Eremoxen-.

Eretes Laporte, 1833b: 397. Current status: valid genus in DYTISCIDAE: DYTISCINAE: ERETINI.

Type species: Dytiscus griseus Fabricius, 1781, by original designation.

Family-group name: ERETINI Crotch, 1873. Stem: Eret-.

Ergania Pascoe, 1882: 445. Current status: valid genus in CURCULIONIDAE: CURCULIONINAE: CURCULIONINI: ERGANIINA.

Type species: Ergania gibba Pascoe, 1882, by monotypy.

Family-group name: ERGANIINA Pelsue & O’Brien, 2011: 29. Stem: Ergani-.

Ergates Audinet-Serville, 1832: 143. Current status: valid genus in CERAMBYCIDAE: PRIONINAE: ERGATINI.

Type species: Prionus serrarius Panzer, 1793 (= Cerambyx faber Linnaeus, 1761), by monotypy.

Family-group name: ERGATITES Fairmaire, 1864. Stem: Ergat-.

Erichsonia Westwood, 1849: 210. Current status: valid genus in CERAMBYCIDAE: PRIONINAE: PARANDRINI: ERICHSONIINA. Note: Erichsonia Dana, 1849 [Crustacea] was published in November 1849 while Erichsonia Westwood, 1849 was published on 5 November 1849 (G. Wheeler 1912: 754), therefore, Westwood’s name has priority.

Type species: Erichsonia dentifrons Westwood, 1849, by monotypy.

Family-group name: ERICHSONITAE J. Thomson, 1861. Stem: Erichsoni-. Note: senior homonym of ERICHSONIINA Brunke & Solodovnikov, 2016 (type genus Erichsonius Fauvel, 1874) in Coleoptera: STAPHYLINIDAE.

Erichsonius Fauvel, 1874: 394, 427. Current status: valid genus in STAPHYLINIDAE: STAPHYLININAE: STAPHYLININI: ERICHSONIINA.

Type species: Staphylinus cinerascens Gravenhorst, 1802, by subsequent designation (R. Lucas 1920: 73) (type fixation for Actobius Fauvel, 1875, an unnecessary new replacement name for Erichsonius Fauvel).

Family-group name: ERICHSONIINA Brunke & Solodovnikov in Brunke et al., 2016: 447. Stem: Erichsoni-. Note: junior homonym of ERICHSONITAE J. Thomson, 1861 (type genus Erichsonia Westwood, 1849) in Coleoptera: CERAMBYCIDAE.

Erineophilus Hopkins, 1902: 34. Current status: junior synonym of Scolytodes Ferrari, 1867 in CURCULIONIDAE: SCOLYTINAE: HEXACOLINI.

Type species: Erineophilus schwarzi Hopkins, 1902, by original designation.

Family-group name: ERINEOPHILIDES Hopkins, 1902. Stem: Erineophil-.

Eriocnemis Kaup, 1868a: 21. Current status: invalid senior synonym of Pelopides Kuwert, 1896 in PASSALIDAE: PASSALINAE: MACROLININI. Note: junior homonym of Eriocnemis L. Reichenbach, 1849 [Aves].

Type species: Passalus tridens Wiedemann, 1823, by subsequent designation (Hincks and Dibb 1935: 85).

Family-group name: ERIOCNEMIAE Kaup, 1871. Stem: Eriocnemid-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

Erionomus Kaup, 1868a: 16. Current status: valid genus in PASSALIDAE: PASSALINAE: SOLENOCYCLINI.

Type species: Passalus planiceps Eschscholtz, 1828, by monotypy.

Family-group name: ERIONOMINAE Kuwert, 1891. Stem: Erionom-.

Eriotomus Piochard de la Brûlerie, 1873: 11, 72. Current status: junior synonym of Oedesis Motschulsky, 1850 in CARABIDAE: HARPALINAE: HARPALINI: DITOMINA.

Type species: Ditomus tomentosus Dejean, 1831, by subsequent designation (Tschitschérine 1901: 225).

Family-group name: ERIOTOMI Antoine, 1959. Stem: Eriotom-.

Eriphus Audinet-Serville, 1834b: 88. Current status: valid genus in CERAMBYCIDAE: CERAMBYCINAE: TRACHYDERINI: TRACHYDERINA. Note: Santos-Silva et al. (2023: 3) treated this name as a nomen protectum over Eryphus Klug, 1829; however, they did not specifically give evidence that the conditions of Article 23.9.1.2 were met (see such requirement in Article 23.9.2, ICZN 1999a); Santos-Silva et al. (2023: 3) also mention that the name Eriphus was made available earlier by Audinet-Serville (1834a: 541); however, the name in the key used by Audinet-Serville (1834a: 541) was a French vernacular name and as such is not a scientific name in the sense of Article 1 (ICZN 1999a, also see Glossary therein).

Type species: Callidium bisignatum Germar, 1823, by subsequent designation (Chevrolat 1844: 404).

Family-group name: ERIPHITAE J. Thomson, 1864. Stem: Eriph-.

Erirhinus Schönherr, 1825: 582. Current status: junior synonym of Notaris Germar, 1817 in CURCULIONIDAE: BRACHYCERINAE: ERIRHININI.

Type species: Curculio aethiops Fabricius, 1792, by subsequent designation (Schönherr 1826: 229).

Family-group name: ERIRHINIDES Schönherr, 1825. Stem: Erirhin-.

Erlandia Aurivillius, 1904: 205. Current status: valid genus in CERAMBYCIDAE: CERAMBYCINAE: ERLANDIINI.

Type species: Erlandia inopinata Aurivillius, 1904, by monotypy.

Family-group name: ERLANDIINI Aurivillius, 1912. Stem: Erlandi-.

Ernobius C.G. Thomson, 1859: 88. Current status: valid genus in PTINIDAE: ERNOBIINAE: ERNOBIINI.

Type species: Dermestes mollis Linnaeus, 1758, by original designation.

Family-group name: ERNOBIINAE Pic, 1912. Stem: Ernobi-.

Ernoporus C.G. Thomson, 1859: 147. Current status: valid genus in CURCULIONIDAE: SCOLYTINAE: ERNOPORINI.

Type species: Apate tiliae Panzer, 1793, by original designation.

Family-group name: ERNOPORINAE Nüsslin, 1911. Stem: Ernopor-.

Erodiscus Schönherr, 1825: 583. Current status: valid genus in CURCULIONIDAE: CURCULIONINAE: ERODISCINI.

Type species: Lixus attenuatus Fabricius, 1801, by original designation.

Family-group name: ERODISCIDES Lacordaire, 1863. Stem: Erodisc-. Note: this name is based on a misidentification of Erodiscus Schönherr, 1825 by Lacordaire (1863: 567) since Lacordaire excluded the type species of Erodiscus, placing it in Toxeutes Schönherr, 1843; Alonso-Zarazaga and Lyal (1999: 78) mentioned that they will apply to the Commission for the conservation of this tribe by designating Erodiscus Schönherr, 1825 as its type genus; meanwhile current usage is maintained.

Erodius Fabricius, 1775: 258. Current status: valid genus in TENEBRIONIDAE: PIMELIINAE: ERODIINI.

Type species: Erodius gibbus Fabricius, 1775, by subsequent designation (Latreille 1810: 429).

Family-group name: ERODIIDES Billberg, 1820. Stem: Erodi-.

Eros Newman, 1838: 382. Current status: valid genus in LYCIDAE: EROTINAE: EROTINI.

Type species: Lycus humeralis Fabricius, 1801, by original designation.

Family-group name: EROTES J.L. LeConte, 1881. Stem: Erot-.

Eroschema Pascoe, 1859a: 17. Current status: valid genus in CERAMBYCIDAE: CERAMBYCINAE: EROSCHEMINI.

Type species: Eroschema poweri Pascoe, 1859, by monotypy.

Family-group name: ÉROSCHÉMIDES Lacordaire, 1868. Stem: Eroschem-.

Erotylus Fabricius, 1775: 123. Current status: valid genus in EROTYLIDAE: EROTYLINAE: EROTYLINI.

Type species: Coccinella gigantea Linnaeus, 1758, by subsequent designation (J.E. Gray 1831a: 611).

Family-group name: EROTILENAE Latreille, 1802. Stem: Erotyl-.

Ertlia Borchmann, 1942: 702. Current status: junior synonym of Morphozonitis Pic, 1922 in MELOIDAE: ELETICINAE: ERTLIANINI. Note: junior homonym of Ertlia Aurivillius, 1907 [Coleoptera: CERAMBYCIDAE].

Type species: Ertlia fasciata Borchmann, 1942, by monotypy.

Family-group name: ERTLIINI Kaszab, 1959. Stem: Ertli-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

Ertliana Selander, 1964: 11. Current status: junior synonym of Morphozonitis Pic, 1922 in MELOIDAE: ELETICINAE: ERTLIANINI. Note: new replacement name for Ertlia Borchmann, 1942.

Type species: Ertlia fasciata Borchmann, 1942, by monotypy (type fixation for Ertlia Borchmann).

Family-group name: ERTLIANINA Selander, 1966. Stem: Ertlian-.

Erythraenus H.W. Bates, 1875: 52. Current status: valid genus in CERAMBYCIDAE: PRIONINAE: ANACOLINI.

Type species: Erythraenus borneensis H.W. Bates, 1875, by monotypy.

Family-group name: ERYTHRAENINAE H.W. Bates, 1875. Stem: Erythraen-.

Erythrus A. White, 1853: 142. Current status: valid genus in CERAMBYCIDAE: CERAMBYCINAE: PSEUDOLEPTURINI. Note: the older name Erythrus Walker in Curtis, 1829 [Hymenoptera] is a nomen nudum (see Heffern 2013: [76]).

Type species: Erythrus championi A. White, 1853, by subsequent designation (J. Thomson 1864: 158) (designation for Pseudoleptura J. Thomson, 1861).

Family-group name: ERYTHRINAE Pascoe, 1866. Stem: Erythr-.

Esarcus Reiche, 1864: 238. Current status: valid genus in MYCETOPHAGIDAE: ESARCINAE.

Type species: Esarcus leprieurii Reiche, 1864, by monotypy.

Family-group name: ESARCINI Reitter, 1882. Stem: Esarc-.

Escalerina Bolívar y Pieltain, 1926: 196, 199. Current status: junior synonym of Karumia Escalera, 1913 in DASCILLIDAE: KARUMIINAE: ESCALERININI.

Type species: Karumia microcephala Escalera, 1913, by original designation.

Family-group name: ESCALERINI Paulus, 1972. Stem: Escalerin-.

Eschatoporis Blaisdell, 1906: 76. Current status: valid genus in TENEBRIONIDAE: LAGRIINAE: ESCHATOPORIINI.

Type species: Eschatoporis nunenmacheri Blaisdell, 1906, by monotypy.

Family-group name: ESCHATOPORINI Blaisdell, 1906. Stem: Eschatopori-.

†Escheria Heer, 1847: 57. Current status: valid genus in HYDROPHILIDAE incertae sedis. Note: late Miocene (Germany).

Type species: †Escheria ovalis Heer, 1847, by monotypy.

Family-group name: †ESCHERIIDAE Kolbe, 1880. Stem: Escheri-.

Esemephe Steiner, 1980: 385. Current status: valid genus in TENEBRIONIDAE: PIMELIINAE: COSSYPHODINI: ESEMEPHINA.

Type species: Esemephe tumi Steiner, 1980, by original designation.

Family-group name: ESEMEPHINI Steiner, 1980. Stem: Esemeph-.

Essisus Pascoe, 1866a: 90. Current status: valid genus in CERAMBYCIDAE: LAMIINAE: DESMIPHORINI.

Type species: Essisus dispar Pascoe, 1866, by monotypy.

Family-group name: ESSISINI Aurivillius, 1917. Stem: Essis-.

Estadia Fairmaire, 1903: 183. Current status: junior synonym of Dietta Sharp, 1876 in LEIODIDAE: LEIODINAE: ESTADIINI.

Type species: Estadia capito Fairmaire, 1903, by original designation.

Family-group name: ESTADIINI Portevin, 1914. Stem: Estadi-.

Esthesopus Eschscholtz, 1829a: 32. Current status: valid genus in ELATERIDAE: CARDIOPHORINAE.

Type species: Esthesopus castaneus Eschscholtz, 1829, by monotypy.

Family-group name: ESTHESOPINAE Fleutiaux, 1919. Stem: Esthesopod-.

Estola Fairmaire & Germain, 1860b: 524. Current status: valid genus in CERAMBYCIDAE: LAMIINAE: DESMIPHORINI.

Type species: Estola hirsuta Fairmaire & Germain, 1860 (= Estola hirsutella Aurivillius, 1922), by subsequent designation (J. Thomson 1861: 348).

Family-group name: ESTOLIDES Lacordaire, 1872. Stem: Estol-.

Ethonion Kubáň in Kubáň et al. 2001: 202. Current status: valid genus in BUPRESTIDAE: AGRILINAE: CORAEBINI: ETHONIINA.

Type species: Buprestis fissiceps W. Kirby, 1818, by original designation.

Family-group name: ETHONIINA Majer, 2001. Stem: Ethoni-.

Euaesthetus Gravenhorst, 1806: 201. Current status: valid genus in STAPHYLINIDAE: EUAESTHETINAE: EUAESTHETINI.

Type species: Euaesthetus scaber Gravenhorst, 1806 (= Stenus bipunctatus Ljungh, 1804), by monotypy.

Family-group name: EUAESTHETINA C.G. Thomson, 1859. Stem: Euaesthet-.

Euanoma Reitter, 1889b: 98. Current status: valid genus in ELATERIDAE: OMALISINAE.

Type species: Euanoma starcki Reitter, 1889, by monotypy.

Family-group name: EUANOMINI Kazantsev, 2010. Stem: Euanom-.

Eubaptus Lacordaire, 1845: 226, 605. Current status: valid genus in CHRYSOMELIDAE: BRUCHINAE: EUBAPTINI.

Type species: Eubaptus palliatus Lacordaire, 1845, by monotypy.

Family-group name: EUBAPTINAE Bridwell, 1932. Stem: Eubapt-.

Eubolbitus Reitter, 1892: 125. Current status: valid genus in BOLBOCERATIDAE: BOLBOCERATINAE: EUBOLBITINI.

Type species: Bolboceras radoszkovskii Solsky, 1876, by monotypy.

Family-group name: EUBOLBITINI Nikolajev, 1970. Stem: Eubolbit-.

Eubria Germar, 1818: 239. Current status: valid genus in PSEPHENIDAE: EUBRIINAE. Note: notwithstanding Löbl’s (2016: 32) statement, this name was first proposed as a junior synonym of Cyphon Paykull, 1799 and subsequently treated as the name of a valid taxon before 1961 by several authors, including Latreille (1829a: 462); therefore, it dates from its first publication as a synonym (ICZN 1999a, Article 11.6.1).

Type species: Cyphon palustris Germar, 1818, by monotypy. Note: name placed on the Official List of Specific Names in Zoology (ICZN 2011b).

Family-group name: EUBRIADES Lacordaire, 1857. Stem: Eubri-.

Eubrianax Kiesenwetter, 1874: 246. Current status: valid genus in PSEPHENIDAE: EUBRIANACINAE.

Type species: Eubrianax ramicornis Kiesenwetter, 1874, by monotypy.

Family-group name: EUBRIANACINI Jakobson, 1913. Stem: Eubrianac-.

Eucaerus J.L. LeConte, 1853e: 384, 386. Current status: valid genus in CARABIDAE: HARPALINAE: LACHNOPHORINI: LACHNOPHORINA.

Type species: Eucaerus varicornis J.L. LeConte, 1853, by monotypy.

Family-group name: EUCAERI J.L. LeConte, 1861. Stem: Eucaer-.

Eucallia Guérin-Méneville, 1844c: pl. 144. Current status: junior synonym of Callidema Guérin- Méneville, 1843 in CICINDELIDAE: CICINDELINI: IRESIINA.

Type species: Callidema boussingaultii Guérin-Méneville, 1843, by monotypy.

Family-group name: EUCALLIINI W. Horn, 1893. Stem: Eucalli-.

Eucallopistus Bellamy, 2003: 32. Current status: junior synonym of Pseudocallopistus Obenberger, 1942 in BUPRESTIDAE: CHRYSOCHROINAE: CHRYSOCHROINI: EUCALLOPISTINA. Note: new replacement name for Callopistus Deyrolle, 1865.

Type species: Callopistus castelnaudii Deyrolle, 1865, by original designation (type fixation for Callopistus Deyrolle).

Family-group name: EUCALLOPISTINA Bellamy, 2003. Stem: Eucallopist-.

Eucalosphaera Jelínek, 1978: 217. Current status: valid genus in NITIDULIDAE: CRYPTARCHINAE: EUCALOSPHAERINI. Note: new replacement name for Calosphaera Jelínek, 1974.

Type species: Cryptarcha ocularis Reitter, 1875, by original designation (type fixation for Calosphaera Jelínek).

Family-group name: EUCALOSPHAERINI Kirejtshuk, 1987. Stem: Eucalosphaer-.

Eucanthus Westwood, 1848c: 387. Current status: valid genus in BOLBOCERATIDAE: BOLBOCERATINAE: EUCANTHINI.

Type species: Scarabaeus meliboeus Fabricius, 1792 (= Scarabaeus lazarus Fabricius, 1775), by monotypy.

Family-group name: EUCANTHINI Nikolajev, 2003. Stem: Eucanth-.

Eucatops Portevin, 1903: 157, 162. Current status: valid genus in LEIODIDAE: CHOLEVINAE: EUCATOPINI.

Type species: Eucatops curvipes Portevin, 1903, by subsequent designation (Jeannel 1922: 39).

Family-group name: EUCATOPINI Jeannel, 1921. Stem: Eucatop-.

Eucheila Dejean in Dejean and Boisduval, 1829: 60. Current status: valid genus in CARABIDAE: HARPALINAE: LEBIINI: PERICALINA. Note: the original spelling was Eucheyla but Eucheila, an incorrect subsequent spelling, is in prevailing usage, attributed to the publication of the original spelling, and therefore deemed to be the correct original spelling (ICZN 1999a, Article 33.3.1).

Type species: Eucheila flavilabris Dejean, 1829, by subsequent monotypy (Dejean and Boisduval, 1830: 178).

Family-group name: EUCHEILINAE H.W. Bates, 1883. Stem: Eucheil-.

Euchirus W. Kirby in W. Kirby and Spence 1828b: 636. Current status: valid genus in SCARABAEIDAE: MELOLONTHINAE: EUCHIRINI.

Type species: Scarabaeus longimanus Linnaeus, 1758, by monotypy.

Family-group name: EUCHEIRIDAE Hope, 1841. Stem: Euchir-.

Euchlora W.S. MacLeay, 1819: 147. Current status: junior synonym of Anomala Leach, 1819 in SCARABAEIDAE: RUTELINAE: ANOMALINI: ANOMALINA.

Type species: Melolontha viridis Fabricius, 1775, by subsequent designation (Hope 1837: 71).

Family-group name: EUCHLORIDAE Hope, 1839. Stem: Euchlor-. Note: nomen oblitum (see A.B.T. Smith 2006: 173).

Euchroa Brullé, 1834: 330, 335. Current status: valid genus in CARABIDAE: HARPALINAE: PTEROSTICHINI: EUCHROINA.

Type species: Euchroa nitidicollis Brullé, 1834, by monotypy.

Family-group name: EUCHROIDES Chaudoir, 1874. Stem: Euchro-.

Euchroea Burmeister, 1842: 571. Current status: valid genus in SCARABAEIDAE: CETONIINAE: STENOTARSIINI: EUCHROEINA.

Type species: Euchroea aurora Burmeister, 1842, by subsequent designation (Pouillaude 1916: 66).

Family-group name: EUCHROEINA Paulian & Descarpentries, 1982. Stem: Euchroe-. Note: junior homonym of EUCHROEIDAE Dahlbom, 1854 (type genus Euchroeus Latreille, 1809) in Hymenoptera.

Euchroma Dejean, 1833a: 76. Current status: valid genus in BUPRESTIDAE: CHRYSOCHROINAE: PARALEPTODEMINI: EUCHROMATINA.

Type species: Buprestis gigantea Linnaeus, 1758, by monotypy.

Family-group name: EUCHROMATINA Holyński, 1993. Stem: Euchromat-.

Eucibdelus Kraatz, 1859: 70. Current status: valid genus in STAPHYLINIDAE: STAPHYLININAE: STAPHYLININI: STAPHYLININA.

Type species: Eucibdelus gracilis Kraatz, 1859, by monotypy.

Family-group name: EUCIBDELINI Sharp, 1889. Stem: Eucibdel-.

Eucinetus Germar, 1818: 255. Current status: valid genus in EUCINETIDAE.

Type species: Scaphidium haemorrhoidale Germar, 1818, by monotypy.

Family-group name: EUCINÉTIDES Lacordaire, 1857. Stem: Eucinet-.

Eucnemis Ahrens, 1812: 38. Current status: valid genus in EUCNEMIDAE: EUCNEMINAE: EUCNEMINI.

Type species: Eucnemis capucinus Ahrens, 1812, by monotypy.

Family-group name: EUCNEMIDES Eschscholtz, 1829. Stem: Eucnem-.

Euconnus C.G. Thomson, 1859: 61. Current status: valid genus in STAPHYLINIDAE: SCYDMAENINAE: SCYDMAENITAE: STENICHNINI.

Type species: Pselaphus hirticollis Illiger, 1798, by original designation.

Family-group name: EUCONNINI Casey, 1897. Stem: Euconn-.

Euconosoma Cameron, 1918: 215, 216. Current status: valid genus in STAPHYLINIDAE: TACHYPORINAE: TACHYPORINI: EUCONOSOMATINA. Note: two original spellings were used, including Eucononosoma (p. 215) but this spelling was changed in the errata (p. xix) to Euconosoma.

Type species: Euconosoma elegans Cameron, 1918 (= Conosoma pictum Bernhauer, 1903), by original designation.

Family-group name: EUCONOSOMINI Cameron, 1918. Stem: Euconosomat-.

Eucrada J.L. LeConte, 1861: 202. Current status: valid genus in PTINIDAE: EUCRADINAE: EUCRADINI.

Type species: Hedobia humeralis Melsheimer, 1845, by monotypy.

Family-group name: EUCRADINI J.L. LeConte, 1861. Stem: Eucrad-.

Eucranium Dejean, 1833: 135. Current status: valid genus in SCARABAEIDAE: SCARABAEINAE: EUCRANIINI. Note: name previously credited to Brullé, 1838 with Eucranium arachnoides Reiche, 1842 as its type species (by subsequent designation in Reiche 1842: 65); however, see below regarding the earlier establishment of the name of the type species.

Type species: Ateuchus arachnoides Lacordaire, 1830, by monotypy. Note: this species name was first established by Lacordaire (1830a: 261).

Family-group name: EUCRANIADAE Burmeister, 1873. Stem: Eucrani-.

Eudactylus Sallé, 1855: 266. Current status: invalid senior synonym of Platycrepidius Candèze, 1859 in ELATERIDAE: AGRYPNINAE: PLATYCREPIDIINI. Note: junior homonym of Eudactylus Fitzinger, 1843 [Reptilia].

Type species: Eudactylus wapleri Sallé, 1855, by monotypy.

Family-group name: EUDACTYLITES Candèze, 1859. Stem: Eudactyl-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

Euderes Schönherr, 1825: 582. Current status: valid genus in CURCULIONIDAE: MOLYTINAE: EUDERINI.

Type species: Rhynchaenus lineicollis Wiedemann, 1823, by original designation.

Family-group name: EUDÉRIDES Lacordaire, 1865. Stem: Euder-. Note: senior homonym of EUDERINI Erdös, 1956 (type genus Euderus Haliday, 1844) in Hymenoptera.

Euderia Broun, 1880: 344. Current status: valid genus in BOSTRICHIDAE: EUDERIINAE.

Type species: Euderia squamosa Broun, 1880, by monotypy.

Family-group name: EUDERIITAE Lesne, 1934. Stem: Euderi-.

Eudiagogus Schönherr, 1840b: 307. Current status: valid genus in CURCULIONIDAE: ENTIMINAE: ENTIMITAE: EUDIAGOGINI.

Type species: Promecops episcopalis Gyllenhal, 1834, by original designation.

Family-group name: EUDIAGOGINI J.L. LeConte, 1874. Stem: Eudiagog-.

Eudicronychus Méquignon, 1931: 207. Current status: valid genus in ELATERIDAE: ELATERINAE: EUDICRONYCHINI.

Type species: Dicronychus serraticornis Laporte, 1840, by original designation.

Family-group name: EUDICRONYCHINAE C. Girard, 1971. Stem: Eudicronych-.

Eudyasmus Pascoe, 1885: 275. Current status: valid genus in CURCULIONIDAE: MOLYTINAE: EUDYASMINI.

Type species: Eudyasmus albertisii Pascoe, 1885, by monotypy.

Family-group name: EUDYASMINI Legalov, 2018d: 347. Stem: Eudyasm-.

Eudysantes Bouchard, Lawrence, Davies & Newton, 2005: 508. Current status: junior synonym of Dysantes Pascoe, 1869 in TENEBRIONIDAE: TENEBRIONINAE: TOXICINI: DYSANTINA. Note: unnecessary new replacement name for Dysantes Pascoe, 1869.

Type species: Diceroderes elongatus L. Redtenbacher, 1868, by monotypy (type fixation for Dysantes Pascoe).

Family-group name: EUDYSANTINA Bouchard, Lawrence, Davies & Newton, 2005. Stem: Eudysant-.

Eugenysa Chevrolat in Dejean, 1836a: 368. Current status: valid genus in CHRYSOMELIDAE: CASSIDINAE: MESOMPHALIINI.

Type species: Cassida grossa Linnaeus, 1758, by monotypy.

Family-group name: EUGENYSINI Hincks, 1952. Stem: Eugenys-.

Euglenes Westwood, 1830a: 59. Current status: valid genus in ADERIDAE. Note: name placed on the Official List of Generic Names in Zoology (ICZN 1989d).

Type species: Anthicus oculatus Paykull, 1798, by subsequent designation (Pic 1900: 3; see ICZN 1989d). Note: name placed on the Official List of Specific Names in Zoology (ICZN 1989d).

Family-group name: EUGLENINI Seidlitz, 1875. Stem: Euglenes- (ICZN 1989d).

Euglochis Gistel, 1856: 382. Current status: junior synonym of Orthocerus Latreille, 1797 in ZOPHERIDAE: COLYDIINAE: ORTHOCERINI.

Type species: Dermestes clavicornis Linnaeus, 1758, by monotypy.

Family-group name: EUGLOCHIDAE Gistel, 1856. Stem: Euglochin-.

Eugnamptus Schönherr, 1839: 339. Current status: valid genus in ATTELABIDAE: RHYNCHITINAE: RHYNCHITITAE: EUGNAMPTINI.

Type species: Anthribus collaris Fabricius, 1801 (= Rhynchites angustatus Herbst, 1797), by original designation.

Family-group name: EUGNAMPTINA Voss, 1930. Stem: Eugnampt-.

Eugnomus Schönherr, 1847: 45. Current status: valid genus in CURCULIONIDAE: CURCULIONINAE: EUGNOMINI: EUGNOMINA.

Type species: Eugnomus durvillei Schönherr, 1847, by original designation.

Family-group name: EUGNOMIDES Lacordaire, 1863. Stem: Eugnom-.

Eugonus Schönherr, 1833: 144. Current status: valid genus in ANTHRIBIDAE: ANTHRIBINAE: EUGONINI.

Type species: Eugonus virgatus Gyllenhal, 1833, by original designation.

Family-group name: EUGONIDES Lacordaire, 1865. Stem: Eugon-.

Eulabis Eschscholtz, 1829d: 14. Current status: valid genus in TENEBRIONIDAE: TENEBRIONINAE: EULABINI.

Type species: Eulabis bicarinata Eschscholtz, 1829, by subsequent designation (Blaisdell 1932: 44).

Family-group name: EULABES G.H. Horn, 1870. Stem: Eulab-.

Eulichas Jakobson, 1913: 727. Current status: valid genus in EULICHADIDAE. Note: new replacement name for Lichas Westwood, 1853.

Type species: Lichas funebris Westwood, 1853, by monotypy (type fixation for Lichas Westwood).

Family-group name: EULICHADIDAE Crowson, 1973. Stem: Eulichad-.

Eumichthus J.L. LeConte, 1873: 190. Current status: valid genus in CERAMBYCIDAE: CERAMBYCINAE: EUMICHTHINI.

Type species: Eumichthus oedipus J.L. LeConte, 1873, by monotypy.

Family-group name: EUMICHTHINI Linsley, 1940. Stem: Eumichth-.

Eumicrus Laporte, 1833b: 395. Current status: junior synonym of Scydmaenus Latreille, 1802 in STAPHYLINIDAE: SCYDMAENINAE: SCYDMAENITAE: SCYDMAENINI.

Type species: Scydmaenus tarsatus P.W.J. Müller & Kunze, 1822, by subsequent designation (Duponchel 1844b: 496).

Family-group name: EUMICRINI Reitter, 1882. Stem: Eumicr-.

Eumolpus Weber, 1801: 28, 52. Current status: valid genus in CHRYSOMELIDAE: EUMOLPINAE: EUMOLPINI. Note: the senior homonym Eumolpus Illiger, 1798 was suppressed for the purposes of both Principles of Priority and Homonymy and placed on the Official Index of Rejected and Invalid Names in Zoology while Eumolpus Weber, 1801 was placed on the Official List of Generic Names in Zoology (ICZN 2012b).

Type species: Chrysomela ignita Fabricius, 1787, by subsequent designation (Hope 1841a: 162; see ICZN 2012b). Note: name placed on the Official List of Specific Names in Zoology (ICZN 2012b).

Family-group name: EUMOLPIDAE Hope, 1841. Stem: Eumolp-.

Eumorphus Weber, 1801: 31. Current status: valid genus in ENDOMYCHIDAE: LYCOPERDININAE.

Type species: Eumorphus sumatrae Weber, 1801 (= Erotylus quadriguttatus Illiger, 1800), by monotypy.

Family-group name: EUMORPHIDAE Gistel, 1848. Stem: Eumorph-. Note: senior homonym of EUMORPHIDAE Tutt, 1904 (type genus Eumorpha Hübner, 1807) in Lepidoptera.

Eunectes Erichson, 1832: 17, 23. Current status: invalid senior synonym of Eretes Laporte, 1833 in DYTISCIDAE: DYTISCINAE: ERETINI. Note: junior homonym of Eunectes Wagler, 1830 [Reptilia].

Type species: Dytiscus sticticus Linnaeus, 1767, by monotypy.

Family-group name: EUNECTINI Portevin, 1929. Stem: Eunect-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

Eunemadus Portevin, 1914: 192. Current status: valid genus in LEIODIDAE: CHOLEVINAE: ANEMADINI: EUNEMADINA.

Type species: Eunemadus chilensis Portevin, 1914, by subsequent designation (Hatch 1928: 208).

Family-group name: EUNEMADINA Newton, 1998. Stem: Eunemad-.

Eunidia Erichson, 1843b: 261. Current status: valid genus in CERAMBYCIDAE: LAMIINAE: EUNIDIINI.

Type species: Eunidia nebulosa Erichson, 1843, by monotypy.

Family-group name: EUNIDIINI Téocchi, Sudre & Jiroux, 2010. Stem: Eunidi-.

Euomus Schönherr, 1847: 52. Current status: valid genus in CURCULIONIDAE: CYCLOMINAE: AMYCTERINI.

Type species: Amycterus insculptus Boheman, 1843, by original designation.

Family-group name: EUOMIDES Lacordaire, 1863. Stem: Euom-.

Euops Schönherr, 1839: 318. Current status: valid genus in ATTELABIDAE: ATTELABINAE: EUOPINI.

Type species: Euops australasiae Fåhraeus, 1839 (= Attelabus falcatus Guérin-Méneville, 1833), by original designation.

Family-group name: EUOPSINI Voss, 1925. Stem: Euop-.

Eupages Schönherr, 1834b: 413. Current status: valid genus in CURCULIONIDAE: CYCLOMINAE: RHYTHIRRININI.

Type species: Eupages tuberculosus Gyllenhal, 1834, by original designation.

Family-group name: EUPAGIDESp>Family-group name: EUPAGIDES Lacordaire, 1863. Stem: Eupag-.

Euparia Lepeletier & Audinet-Serville, 1828: 357. Current status: valid genus in SCARABAEIDAE: APHODIINAE: EUPARIINI.

Type species: Euparia castanea Lepeletier & Audinet-Serville, 1828, by monotypy.

Family-group name: EUPARIINA A. Schmidt, 1910. Stem: Eupari-. Note: nomen protectum (see A.B.T. Smith 2006: 158); senior homonym of EUPARIINI B.D. Valentine, 1960 (type genus Euparius Schönherr, 1823) in Coleoptera: ANTHRIBIDAE.

Euparius Schönherr, 1823: 1135. Current status: valid genus in ANTHRIBIDAE: ANTHRIBINAE: CRATOPARINI.

Type species: Anthribus lunatus Fabricius, 1801 (= Macrocephalus marmoreus G.-A. Olivier, 1800), by monotypy.

Family-group name: EUPARIINI B.D. Valentine, 1960. Stem: Eupari-. Note: junior homonym of EUPARIINI A. Schmidt, 1910 (type genus Euparia Lepeletier & Serville, 1828) in Coleoptera: SCARABAEIDAE.

Euphanias Fairmaire & Laboulbène, 1856: 657. Current status: valid genus in STAPHYLINIDAE: OXYTELINAE: EUPHANIINI.

Type species: Euphanias insignicornis Fairmaire & Laboulbène, 1856 (= Pholidus insignis Mulsant & Rey, 1856), by monotypy.

Family-group name: EUPHANIAE Reitter, 1909. Stem: Euphani-.

Eupholus Boisduval, 1835: 363. Current status: valid genus in CURCULIONIDAE: ENTIMINAE: POLYDRUSITAE: EUPHOLINI. Note: we consider Eupholus used by Guérin-Méneville (1838: 114) a subsequent usage of Eupholus Boisduval, 1835.

Type species: Geonemus geoffroyii Guérin-Méneville, 1831, by subsequent designation (Porion 1993: 9).

Family-group name: EUPHOLINI Günther, 1943. Stem: Euphol-.

Euphoria Burmeister, 1842: 370. Current status: valid genus in SCARABAEIDAE: CETONIINAE: CETONIINI: EUPHORIINA.

Type species: Cetonia sepulcralis Fabricius, 1801, by subsequent designation (Casey 1915: 298).

Family-group name: EUPHORIAE G.H. Horn, 1880. Stem: Euphori-.

Euplectalecia Obenberger, 1924: 10. Current status: valid genus in BUPRESTIDAE: CHRYSOCHROINAE: PARALEPTODEMINI: EUPLECTALECIINA.

Type species: Halecia pyropus Kerremans, 1893 (= Buprestis erythropus Gory, 1841), by original designation.

Family-group name: EUPLECTALECIINA Holyński, 1993. Stem: Euplectaleci-.

Euplectroscelis Crotch, 1873b: 75. Current status: valid genus in CHRYSOMELIDAE: GALERUCINAE: ALTICINI.

Type species: Euplectroscelis xanti Crotch, 1873, by monotypy.

Family-group name: EUPLECTROSCELES G.H. Horn, 1889. Stem: Euplectroscelid-.

Euplectus Leach, 1817: 80, 82. Current status: valid genus in STAPHYLINIDAE: PSELAPHINAE: EUPLECTITAE: EUPLECTINI.

Type species: Euplectus reichenbachii Leach, 1817 (= Pselaphus nanus H.G.L. Reichenbach, 1816), by monotypy.

Family-group name: EUPLECTIDAE Streubel, 1839. Stem: Euplect-.

Euplinthus C. Costa, 1975: 67. Current status: valid genus in ELATERIDAE: AGRYPNINAE: EUPLINTHINI: EUPLINTHINA.

Type species: Elater ophthalmicus Perty, 1830, by original designation.

Family-group name: EUPLINTHINA C. Costa, 1975. Stem: Euplinth-.

Eupoecila Burmeister, 1842: 538. Current status: valid genus in SCARABAEIDAE: CETONIINAE: SCHIZORHININI: SCHIZORHININA.

Type species: Cetonia australasiae Donovan, 1805, by subsequent designation (Kraatz 1880: 190).

Family-group name: EUPOECILIDAE Kraatz, 1880. Stem: Eupoecil-.

Eupogonius J.L. LeConte, 1852a: 159. Current status: valid genus in CERAMBYCIDAE: LAMIINAE: DESMIPHORINI.

Type species: Desmiphora tomentosa Haldeman, 1847, by subsequent designation (J. Thomson 1861: 346).

Family-group name: EUPOGONII J.L. LeConte, 1873. Stem: Eupogoni-.

Eupompha J.L. LeConte, 1858b: 21. Current status: valid genus in MELOIDAE: MELOINAE: EUPOMPHINI.

Type species: Eupompha fissiceps J.L. LeConte, 1858, by monotypy.

Family-group name: EUPOMPHAE J.L. LeConte, 1862. Stem: Eupomph-.

Eupromera Westwood, 1845b: 209. Current status: valid genus in CERAMBYCIDAE: LAMIINAE: EUPROMERINI.

Type species: Eupromera spryana Westwood, 1845, by monotypy.

Family-group name: EUPROMERINI Galileo & Martins, 1995. Stem: Eupromer-.

Euprosopus W.S. MacLeay, 1825: 9. Current status: valid genus in CICINDELIDAE: CICINDELINI: IRESIINA. Note: this name is attributed to Dejean (1825: 4, 150 [5 September]) in the literature but it was first made available by W.S. MacLeay (1825 [1 June]) when he associated Cicindela quadrinotata Latreille & Dejean, 1822 (as Euprosopus quadrinotatus Lat.) with it.

Type species: Cicindela quadrinotata Latreille & Dejean, 1822, by monotypy.

Family-group name: EUPROSOPINI W. Horn, 1893. Stem: Euprosop-.

Eupsenius J.L. LeConte, 1849: 90. Current status: valid genus in STAPHYLINIDAE: PSELAPHINAE: GONIACERITAE: BRACHYGLUTINI: EUPSENIINA.

Type species: Eupsenius glaber J.L. LeConte, 1849, by monotypy.

Family-group name: EUPSENIINI Park, 1951. Stem: Eupseni-.

Eupsilobius Casey, 1895: 454. Current status: junior synonym of Eidoreus Sharp, 1885 in EUPSILOBIIDAE.

Type species: Eupsilobius politus Casey, 1895, by monotypy.

Family-group name: EUPSILOBIINI Casey, 1895. Stem: Eupsilobi-.

Eurhinus W. Kirby, 1819: 427. Current status: invalid senior synonym of Eurhynchus W. Kirby, 1828 in BRENTIDAE: EURHYNCHINAE: EURHYNCHINI. Note: two original spellings were used, including Eurhynus (p. 428); Zimmerman (1994: 255) acted as First Reviser (ICZN 1999a, Article 24.2.3) and selected the spelling Eurhinus; junior homonym of Eurhinus Illiger, 1807 [Coleoptera: CURCULIONIDAE].

Type species: Eurhinus scabrior W. Kirby, 1819, by subsequent designation (Schönherr 1833: 248; see ICZN 1985e).

Family-group name: EURHININI Kissinger, 1968. Stem: Eurhin-. Note: junior homonym of EURHININA Lacordaire, 1865 (based on Eurhinus Illiger, 1807) in Coleoptera: CURCULIONIDAE; family-group name permanently invalid (ICZN 1999a, Article 39).

Eurhinus Illiger, 1807a: 309. Current status: valid genus in CURCULIONIDAE: BARIDINAE: BARIDINI: EURHININA. Note: this name was originally proposed under the spelling Eurhin but this spelling was placed on the Official Index of Rejected and Invalid Generic Names in Zoology, Eurhinus was considered as the correct original spelling and Eurhinus Illiger, 1807 placed on the Official List of Generic Names in Zoology (ICZN 1985e).

Type species: Eurhinus cupratus Illiger, 1807, by monotypy (see ICZN 1985e). Note: name placed on the Official List of Specific Names in Zoology (ICZN 1985e).

Family-group name: EURHINIDES Lacordaire, 1865. Stem: Eurhin-. Note: senior homonym of EURHININI Kissinger, 1968 (type genus Eurhinus W. Kirby, 1819) in Coleoptera: BRENTIDAE; Kissinger’s taxon is permanently invalid (based on a preoccupied type genus).

Eurhynchus W. Kirby in W. Kirby and Spence, 1828a: 324. Current status: valid genus in BRENTIDAE: EURHYNCHINAE: EURHYNCHINI. Note: new replacement name for Eurhinus W. Kirby, 1819; the senior homonym Eurhynchus Berthold, 1827 was suppressed for the purposes of both the Principle of Priority and Homonymy and placed on the Official Index of Rejected and Invalid Generic Names in Zoology (see ICZN 1985e).

Type species: Eurhinus scabrior W. Kirby, 1819, by subsequent designation (Schönherr 1833: 248; see ICZN 1985e).

Family-group name: EURHYNCHIDES Lacordaire, 1863. Stem: Eurhynch-.

Eurispa Baly, 1858: 85. Current status: valid genus in CHRYSOMELIDAE: CASSIDINAE: EURISPINI.

Type species: Eurispa vittata Baly, 1858, by original designation.

Family-group name: EURISPITES Chapuis, 1875. Stem: Eurisp-.

Europs Wollaston, 1854: 149. Current status: valid genus in MONOTOMIDAE: MONOTOMINAE: EUROPINI.

Type species: Europs impressicollis Wollaston, 1854, by monotypy.

Family-group name: EUROPINI Sen Gupta, 1988. Stem: Europ-.

Eurrhacus C.O. Waterhouse, 1880a: 24. Current status: valid genus in LYCIDAE: LYCINAE: EURRHACINI.

Type species: Eurrhacus tristis C.O. Waterhouse, 1880, by monotypy.

Family-group name: EURRHACINA Bocáková, 2005. Stem: Eurrhac-.

Eurychora Thunberg, 1789: 9. Current status: valid genus in TENEBRIONIDAE: PIMELIINAE: ADELOSTOMINI. Note: the original spelling was Evrychora but Eurychora, an incorrect subsequent spelling, is in prevailing usage and attributed to the original publication, and therefore deemed to be the correct original spelling (ICZN 1999a, Article 33.3.1).

Type species: Pimelia ciliata Fabricius, 1781, by monotypy.

Family-group name: EURYCHORIDAE Hope, 1841. Stem: Eurychor-.

Eurygenius LaFerté-Sénectère in Guérin-Méneville, 1849: [No 17] 1. Current status: valid genus in ANTHICIDAE: EURYGENIINAE: EURYGENIINI.

Type species: Eurygenius reichei LaFerté-Sénectère, 1849, by monotypy.

Family-group name: EURYGENII J.L. LeConte, 1862. Stem: Eurygeni-.

Eurylobus Schönherr, 1826: 152. Current status: valid genus in CURCULIONIDAE: ENTIMINAE: ENTIMITAE: EURYLOBINI.

Type species: Hypsonotus cingulatus Germar, 1823, by original designation (p. 13).

Family-group name: EURYLOBI Jekel, 1856. Stem: Eurylob-.

Eurymetopon Eschscholtz, 1831: 5, 8. Current status: valid genus in TENEBRIONIDAE: PIMELIINAE: EDROTINI.

Type species: Eurymetopon rufipes Eschscholtz, 1831, by subsequent designation (Casey 1907: 288).

Family-group name: EURYMETOPONINI Casey, 1907. Stem: Eurymetop-.

Eurymycter J.L. LeConte in J.L. LeConte and G.H. Horn, 1876: 394. Current status: valid genus in ANTHRIBIDAE: ANTHRIBINAE: TROPIDERINI.

Type species: Macrocephalus fasciatus A.-G. Olivier, 1800, by monotypy.

Family-group name: EURYMYCTERINI Pierce, 1930. Stem: Eurymycter-.

Eurynotus W. Kirby, 1819: 418. Current status: valid genus in TENEBRIONIDAE: BLAPTINAE: PLATYNOTINI: EURYNOTINA.

Type species: Eurynotus muricatus W. Kirby, 1819 (= Helops capensis Fabricius, 1794), by monotypy.

Family-group name: EURYNOTAIRES Mulsant & Rey, 1854. Stem: Eurynot-.

Euryoda Lacordaire, 1842b: 12, 27. Current status: junior synonym of Heptodonta Hope, 1838 in CICINDELIDAE: CICINDELINI: CICINDELINA. Note: unnecessary new replacement name for Heptodonta Hope, 1838.

Type species: Cicindela analis Fabricius, 1801, by original designation (type fixation for Heptodonta Hope).

Family-group name: EURYODINI W. Horn, 1899. Stem: Euryod-.

Euryope Dalman, 1824: 17. Current status: valid genus in CHRYSOMELIDAE: EUMOLPINAE: EURYOPINI.

Type species: Eumolpus ruber Latreille, 1807 (= Cryptocephalus subserricornis Latreille, 1805), by subsequent designation (Blanchard in Audouin et al. 1841: pl. 72). Note: the type species usually recognized is Cryptocephalus rubrifrons Fabricius, 1787 by subsequent designation of Moseyko and Sprecher-Uebersax (2010a: 83) but Blanchard’s typification is earlier; both species are currently included in the genus Euryope.

Family-group name: EURYOPITES Chapuis, 1874. Stem: Euryop-.

Eurypalpus J.L. LeConte, 1850b: 241. Current status: invalid senior synonym of Psephenus Haldeman, 1853 in PSEPHENIDAE: PSEPHENINAE. Note: junior homonym of Eurypalpus Macquart, 1835 [Diptera].

Type species: Eurypalpus lecontei J.L. LeConte, 1850, by monotypy.

Family-group name: EURYPALPINI J.L. LeConte, 1852. Stem: Eurypalp-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

Eurypoda Saunders, 1853: 109. Current status: valid genus in CERAMBYCIDAE: PRIONINAE: EURYPODINI.

Type species: Eurypoda antennata Saunders, 1853, by monotypy.

Family-group name: EURYPODINI Gahan, 1906. Stem: Eurypod-.

Eurypogon Motschulsky, 1860a: 358, 363. Current status: valid genus in ARTEMATOPODIDAE: ARTEMATOPODINAE: MACROPOGONINI.

Type species: Ochina nigra Melsheimer, 1845, by monotypy.

Family-group name: EURYPOGONIDAE Böving & Craighead, 1931. Stem: Eurypogon-.

Euryptychus J.L. LeConte, 1852c: 45, 46. Current status: valid genus in EUCNEMIDAE: MACRAULACINAE: EURYPTYCHINI.

Type species: Eucnemis heterocerus Say, 1839, by monotypy.

Family-group name: EURYPTYCHINI Mamaev, 1976. Stem: Euryptych-.

Eurypus W. Kirby, 1819: 389. Current status: valid genus in MYCTERIDAE: EURYPINAE.

Type species: Eurypus rubens W. Kirby, 1819, by monotypy.

Family-group name: EURYPITAE J. Thomson, 1860. Stem: Euryp-.

Eurysphindus J.L. LeConte, 1878b: 602. Current status: valid genus in SPHINDIDAE: SPHINDINAE.

Type species: Eurysphindus hirtus J.L. LeConte, 1878, by monotypy.

Family-group name: EURYSPHINDINAE Sen Gupta & Crowson, 1979. Stem: Eurysphind-.

Eurysternus Dalman, 1824: 8. Current status: valid genus in SCARABAEIDAE: SCARABAEINAE: EURYSTERNINI.

Type species: Eurysternus planus Dalman, 1824 (= Scarabaeus caribaeus Herbst, 1789), by subsequent designation (Jessop 1985: 1089).

Family-group name: EURYSTERNINI Vulcano, Martínez & Pereira, 1961. Stem: Eurystern-. Note: junior homonym of EURYSTERNIDAE Dollo, 1886 (type genus Eurysternum J. Meyer, 1839) in Reptilia.

Eurystethes Seidlitz, 1916: 127. Current status: junior synonym of Aegialites Mannerheim, 1853 in SALPINGIDAE: AEGIALITINAE. Note: unnecessary new replacement name for Aegialites Mannerheim, 1853.

Type species: Aegialites debilis Mannerheim, 1853, by monotypy (type fixation for Aegialites Mannerheim).

Family-group name: EURYSTETHIDAE Seidlitz, 1916. Stem: Eurysteth-.

Eurytrichus J.L. LeConte, 1847: 387. Current status: junior synonym of Anisotarsus Chaudoir, 1837 in CARABIDAE: HARPALINAE: HARPALINI: ANISODACTYLINA.

Type species: Feronia terminata Say, 1823, by subsequent designation (Emden 1953: 525).

Family-group name: EURYTRICHINI J.L. LeConte, 1847. Stem: Eurytrich-. Note: nomen oblitum (see Bouchard et al. 2011: 128).

Euryusa Erichson, 1837: 371. Current status: valid genus in STAPHYLINIDAE: ALEOCHARINAE: HOMALOTINI: BOLITOCHARINA.

Type species: Euryusa sinuata Erichson, 1837, by monotypy.

Family-group name: EURYUSIDES C.G. Thomson, 1859. Stem: Euryus-.

Eusattus J.L. LeConte, 1851a: 131. Current status: valid genus in TENEBRIONIDAE: PIMELIINAE: CONIONTINI.

Type species: Eusattus difficilis J.L. LeConte, 1851, by subsequent designation (Casey 1908: 56).

Family-group name: EUSATTI Casey, 1908. Stem: Eusatt-.

Euscelophilus Voss, 1925: 30. Current status: valid genus in ATTELABIDAE: ATTELABINAE: ATTELABINI: EUSCELOPHILINA.

Type species: Euscelus chinensis Schilsky, 1906, by subsequent designation (Ter-Minasian 1950: 162).

Family-group name: EUSCELOPHILINA Voss, 1925. Stem: Euscelophil-.

Euscelus Schönherr, 1833: 4, 205. Current status: valid genus in ATTELABIDAE: ATTELABINAE: ATTELABINI: EUSCELINA.

Type species: Attelabus scutellatus Gyllenhal, 1833 (= Attelabus scutellatus Klug, 1829), by original designation.

Family-group name: EUSCELINA Voss, 1925. Stem: Euscel-.

Eusphalerum Kraatz, 1857c: 1003. Current status: valid genus in STAPHYLINIDAE: OMALIINAE: EUSPHALERINI.

Type species: Anthobium triviale Erichson, 1839 (= Omalium primulae Stephens, 1834), by subsequent designation (Blackwelder 1952: 161).

Family-group name: EUSPHALERINI Hatch, 1957. Stem: Eusphaler-.

Euspilotus Lewis, 1907: 320. Current status: valid genus in HISTERIDAE: SAPRININAE: EUSPILOTINI.

Type species: Euspilotus zonalis Lewis, 1907, by original designation.

Family-group name: EUSPILOTINI Lackner in Lackner et al. 2024: 56. Stem: Euspilot-.

Eusteniamorpha Cameron, 1920: 253. Current status: valid genus in STAPHYLINIDAE: ALEOCHARINAE: EUSTENIAMORPHINI.

Type species: Eusteniamorpha rufa Cameron, 1920, by monotypy.

Family-group name: EUSTENIAMORPHINI Bernhauer & Scheerpeltz, 1926. Stem: Eusteniamorph-.

Eustilbus Sharp, 1888: 253. Current status: junior synonym of Stilbus Seidlitz, 1872 in PHALACRIDAE: PHALACRINAE. Note: unnecessary new replacement name for Stilbus Seidlitz, 1872.

Type species: Anisotoma testacea Panzer, 1797, by subsequent designation (Guillebeau 1894: 283).

Family-group name: EUSTILBINI Guillebeau, 1892. Stem: Eustilb-.

Eustra Schmidt-Göbel, 1846: 65. Current status: valid genus in CARABIDAE: PAUSSINAE: OZAENINI.

Type species: Eustra plagiata Schmidt-Göbel, 1846, by monotypy.

Family-group name: EUSTRINI Jeannel, 1946. Stem: Eustr-.

Eustrophus Illiger, 1802: 301. Current status: valid genus in TETRATOMIDAE: EUSTROPHINAE: EUSTROPHINI.

Type species: Mycetophagus dermestoides Fabricius, 1792, by monotypy.

Family-group name: EUSTROPHIDAE Gistel, 1848. Stem: Eustroph-.

Eustylus Schönherr, 1842b: 40. Current status: valid genus in CURCULIONIDAE: ENTIMINAE: POLYDRUSITAE: EUSTYLINI: EUSTYLINA.

Type species: Curculio puber G.-A. Olivier, 1807, by original designation.

Family-group name: EUSTYLIDES Lacordaire, 1863. Stem: Eustyl-.

Eusyneta Gistel, 1856: 356. Current status: junior synonym of Callistus Bonelli, 1810 in CARABIDAE: HARPALINAE: CHLAENIINI: CALLISTINA.

Type species: Carabus lunatus Fabricius, 1775, by monotypy.

Family-group name: EUSYNETADAE Gistel, 1856. Stem: Eusynet-.

Eutelus Solier, 1843: 56. Current status: invalid senior synonym of Eutelonotus Alluaud, 1902 in TENEBRIONIDAE: STENOCHIINAE: CNODALONINI. Note: two original spellings were used, including Lutelus (p. 4); Bouchard et al. (2021: 186) acted as First Revisers (ICZN 1999a, Article 24.2.3) and chose Eutelus as the correct original spelling; junior homonym of Eutelus Walker, 1834 [Hymenoptera].

Type species: Eutelus requieni Solier, 1843, by subsequent designation (R. Lucas 1920: 291).

Family-group name: EUTÉLIDES Lacordaire, 1859. Stem: Eutel-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

Eutheia Stephens, 1830: 59. Current status: valid genus in STAPHYLINIDAE: SCYDMAENINAE: CEPHENNIITAE: EUTHEIINI.

Type species: Eutheia scydmaenoides Stephens, 1830, by subsequent monotypy (Stephens 1830: 115).

Family-group name: EUTHEIINI Casey, 1897. Stem: Euthei-.

Eutomus Lacordaire, 1865: 369. Current status: junior synonym of Rhipidandrus J.L. LeConte, 1862 in TENEBRIONIDAE: TENEBRIONINAE: BOLITOPHAGINI.

Type species: Eutomus micrographus Lacordaire, 1865, by subsequent designation (Barber 1914: 191).

Family-group name: EUTOMIDES Lacordaire, 1866. Stem: Eutom-.

Eutoxus Schönherr, 1844: 103. Current status: valid genus in CURCULIONIDAE: BARIDINAE: EUTOXINI: EUTOXINA.

Type species: Eutoxus reflexus Boheman, 1844, by original designation.

Family-group name: EUTOXIDES Champion, 1908. Stem: Eutox-.

Eutrachelus Berthold, 1827: 383. Current status: valid genus in BRENTIDAE: BRENTINAE: BRENTINI: ARRENODINA. Note: the original spelling Entrachelus (p. 383) was corrected to Eutrachelus (p. 575).

Type species: Brentus temminckii Latreille, 1825, by monotypy.

Family-group name: EUTRACHÉLIDES Lacordaire, 1865. Stem: Eutrachel-.

Eutryptes Gistel, 1856: 375. Current status: valid subgenus of Mycterus Clairville, 1798 in MYCTERIDAE: MYCTERINAE.

Type species: Bruchus umbellatarum Fabricius, 1787, by subsequent designation (Löbl 2008b: 412).

Family-group name: FUTRYPTEIDAE Gistel, 1856. Stem: Eutrypt-.

Euxestus Wollaston, 1858: 411. Current status: valid genus in EUXESTIDAE.

Type species: Euxestus parkii Wollaston, 1858, by monotypy.

Family-group name: EUXESTINAE Grouvelle, 1908. Stem: Euxest-. Note: senior homonym of EUXESTINAE Hennig, 1940 (type genus Euxesta Loew, 1868) in Diptera.

Evambates Gistel, 1856: 365. Current status: junior synonym of Trichius Fabricius, 1775 in SCARABAEIDAE: CETONIINAE: TRICHIINI.

Type species: Scarabaeus fasciatus Linnaeus, 1758, by subsequent designation (Bouchard and Bousquet 2020a: 84). Note: name placed on the Official List of Specific Names in Zoology (ICZN 2004d).

Family-group name: EVAMBATEIDAE Gistel, 1856. Stem: Evambat-.

Evaniocera Guérin-Méneville in Guérin-Méneville and Percheron, 1835: pl. 2. Current status: junior synonym of Ptilophorus Dejean, 1834 in RIPIPHORIDAE: PTILOPHORINAE. Note: two original spellings were used, including Evanicocera (p. [4]); Mannerheim (1838: 76) acted as First Reviser (ICZN 1999a, Article 24.2.3) and selected Evaniocera.

Type species: Pelecotoma dufourii Latreille, 1817, by original designation.

Family-group name: ÉVANIOCÉRIDES Lacordaire, 1859. Stem: Evaniocer-.

Evaniosomus Guérin-Méneville, 1834b: pl. 109bis (p. 14). Current status: valid genus in TENEBRIONIDAE: PIMELIINAE: EVANIOSOMINI.

Type species: Evaniosomus orbignianus Guérin-Méneville, 1834, by monotypy.

Family-group name: ÉVANIOSOMIDES Lacordaire, 1859. Stem: Evaniosom-.

Evaspistes Gistel, 1856: 381. Current status: junior synonym of Cassida Linnaeus, 1758 in CHRYSOMELIDAE: CASSIDINAE: CASSIDINI.

Type species: Cassida viridis Linnaeus, 1758, by present designation.

Family-group name: EVASPIDOTIDAE Gistel, 1856. Stem: Evaspist-.

Evides Dejean, 1833a: 77. Current status: valid genus in BUPRESTIDAE: CHRYSOCHROINAE: EVIDINI.

Type species: Buprestis elegans Fabricius, 1781, by subsequent designation (Bellamy 1997a: 370).

Family-group name: EVIDINI Tôyama, 1987. Stem: Evid-.

Evodinus LeConte, 1850a: 325. Current status: valid genus in CERAMBYCIDAE: LEPTURINAE: EVODININI.

Type species: Leptura monticola Randall, 1838, by monotypy.

Family-group name: EVODININI Zamoroka, 2022: 303. Stem: Evodin-. Note: two original spellings were used, EVODININI (pp. 297, 301) and EVODINIINI (pp. 302, 303); we here select EVODININI as the correct original spelling.

Evotus J.L. LeConte, 1874a: 459. Current status: valid genus in CURCULIONIDAE: ENTIMINAE: POLYDRUSITAE: EVOTINI.

Type species: Otiorhynchus naso J.L. LeConte, 1857, by monotypy.

Family-group name: EVOTINI J.L. LeConte, 1874. Stem: Evot-.

Exagistus Deyrolle, 1865: 65. Current status: valid genus in BUPRESTIDAE: BUPRESTINAE: EXAGISTINI.

Type species: Exagistus igniceps Deyrolle, 1865, by monotypy.

Family-group name: EXAGISTINI Tôyama, 1987. Stem: Exagist-.

Exapion Bedel, 1887: 360. Current status: valid genus in BRENTIDAE: APIONINAE: APIONITAE: EXAPIINI.

Type species: Curculio fuscirostris Fabricius, 1775, by subsequent designation (Schilsky 1901: 38F).

Family-group name: EXAPIINI Alonso-Zarazaga, 1990. Stem: Exapi-.

Exocentrus Dejean, 1835: 339. Current status: valid genus in CERAMBYCIDAE: LAMIINAE: EXOCENTRINI.

Type species: here fixed (under ICZN 1999a, Article 70.3.2) as Cerambyx lusitanus Linnaeus, 1767, misidentified as Cerambyx balteus Linnaeus, 1767 (sensu Dejean, 1835), in the original designation by monotypy in Dejean (1835: 339). Note: the nominal species Cerambyx balteus Linnaeus, 1767 is a valid species in the genus Parmena Dejean, 1821 of the tribe PARMENINI Mulsant, 1839 (see Bousquet and Bouchard 2013a: 82).

Family-group name: EXOCENTRINAE Pascoe, 1864. Stem: Exocentr-.

Exomella Casey, 1914: 378. Current status: valid genus in BYRRHIDAE: BYRRHINAE: EXOMELLINI. Note: new replacement name for Exoma Casey, 1908.

Type species: Exoma pleuralis Casey, 1908, by original designation (type fixation for Exoma Casey).

Family-group name: EXOMELLINI Casey, 1914. Stem: Exomell-.

Exophthalmodes Pierce, 1916: 464. Current status: junior synonym of Exophthalmus Schönherr, 1823 in CURCULIONIDAE: ENTIMINAE: POLYDRUSITAE: EUSTYLINI: EUSTYLINA. Note: unnecessary new replacement name for Exophthalmus Schönherr, 1823.

Type species: Curculio quadrivittatus G.-A. Olivier, 1807, by original designation (type fixation for Exophthalmus Schönherr).

Family-group name: EXOPHTHALMODINA Voss, 1954. Stem: Exophthalmod-.

Exophthalmus Schönherr, 1823: 1140. Current status: valid genus in CURCULIONIDAE: ENTIMINAE: EUSTYLINI: EUSTYLINA.

Type species: Curculio quadrivittatus G.-A. Olivier, 1807, by original designation.

Family-group name: EXOPHTHALMINI G.H. Horn, 1876. Stem: Exophthalm-.

Exoplectra Chevrolat in Dejean, 1836a: 437. Current status: valid genus in COCCINELLIDAE: COCCINELLINAE: COCCIDULINI.

Type species: Coccinella coccinea Fabricius, 1801, by subsequent designation (Korschefsky 1932: 227).

Family-group name: EXOPLECTRIDES Crotch, 1874. Stem: Exoplectr-.

Exosternus Lewis, 1902: 233. Current status: valid genus in HISTERIDAE: HISTERINAE: EXOSTERNINI.

Type species: Exosternus manicatus Lewis, 1902, by subsequent designation (Bickhardt 1919: 206).

Family-group name: EXOSTERNINI Bickhardt, 1914. Stem: Exostern-.

Exothispa Kolbe, 1897a: 354. Current status: valid genus in CHRYSOMELIDAE: CASSIDINAE: EXOTHISPINI.

Type species: Exothispa reimeri Kolbe, 1897, by monotypy.

Family-group name: EXOTHISPINI Weise, 1911. Stem: Exothisp-.

Fabia Martínez & Viana, 1964: 8. Current status: invalid senior synonym of Fabrasia Martínez & Viana, 1964 in PTINIDAE incertae sedis. Note: junior homonym of Fabia Dana, 1851 [Crustacea].

Type species: Fabia alvarengai Martínez & Viana, 1964, by original designation.

Family-group name: FABIINAE Martínez & Viana, 1964. Stem: Fabi-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

Fabrasia Martínez & Viana, 1965: 18. Current status: valid genus in PTINIDAE incertae sedis. Note: new replacement name for Fabia Martínez & Viana, 1964.

Type species: Fabia alvarengai Martínez & Viana, 1964, by original designation (type fixation for Fabia Martínez & Viana).

Family-group name: FABRASIINAE Lawrence & Reichardt, 1966. Stem: Fabrasi-.

Falagria Leach in Samouelle, 1819: 177. Current status: valid genus in STAPHYLINIDAE: ALEOCHARINAE: FALAGRIINI.

Type species: Staphylinus sulcatus Paykull, 1789 (= Falagria caesa Erichson, 1837), by original designation.

Family-group name: FALAGRIATES Mulsant & Rey, 1873. Stem: Falagri-.

Falklandius Enderlein, 1907: 65. Current status: valid genus in CURCULIONIDAE: CYCLOMINAE: LISTRODERINI: FALKLANDIINA.

Type species: Falklandius brachyomma Enderlein, 1907 (= Otiorhynchus antarcticus Stierlin, 1903), by original designation.

Family-group name: FALKLANDIINA Morrone, 2013: 42. Stem: Falklandi-.

Falsamblesthis Breuning, 1959: 85. Current status: valid genus in CERAMBYCIDAE: LAMIINAE: FORSTERIINI.

Type species: Amblesthis seriepilosa Kirsch, 1890, by original designation.

Family-group name: FALSAMBLESTHIINI Gilmour, 1961. Stem: Falsamblesth-.

Falsocossyphus Pic, 1916a: 4. Current status: valid genus in TENEBRIONIDAE: TENEBRIONINAE: FALSOCOSSYPHINI.

Type species: Falsocossyphus pilosus Pic, 1916, by monotypy.

Family-group name: FALSOCOSSYPHINI Ferrer, 2006. Stem: Falsocossyph-.

Falsomycterus Pic, 1907: 127. Current status: valid genus in TENEBRIONIDAE: PIMELIINAE: FALSOMYCTERINI.

Type species: Falsomycterus diversipes Pic, 1907, by monotypy.

Family-group name: FALSOMYCTERINAE Gebien, 1910. Stem: Falsomycter-.

Faronus Aubé, 1844: 157. Current status: valid genus in STAPHYLINIDAE: PSELAPHINAE: FARONITAE.

Type species: Faronus lafertei Aubé, 1844, by monotypy.

Family-group name: FARONIDES Reitter, 1882. Stem: Faron-.

Febra Clark, 1864: 261. Current status: valid genus in CHRYSOMELIDAE: GALERUCINAE: ALTICINI.

Type species: Febra venusta Clark, 1864, by original designation.

Family-group name: FEBRAINA Nadein, 2013: 499. Stem: Febra- (ICZN 1999a, Article 29.4).

Felda Blackwelder, 1952: 165. Current status: junior synonym of Termitobiella Wasmann, 1916 in STAPHYLINIDAE: ALEOCHARINAE: FELDINI. Note: new replacement name for Asticta Wasmann, 1916.

Type species: Asticta butteli Wasmann, 1916, by monotypy (type fixation for Asticta Wasmann).

Family-group name: FELDINI Kistner, 1972. Stem: Feld-.

Fenderia Hatch, 1957: 245. Current status: valid genus in STAPHYLINIDAE: EUAESTHETINAE: STICTOCRANIINI.

Type species: Fenderia capizzii Hatch, 1957, by original designation.

Family-group name: FENDERIINI Scheerpeltz, 1974. Stem: Fenderi-.

Feronia Latreille, 1816: 191. Current status: junior synonym of Poecilus Bonelli, 1810 in CARABIDAE: HARPALINAE: PTEROSTICHINI: PTEROSTICHINA.

Type species: Carabus cupreus Linnaeus, 1758, by subsequent designation (Blanchard in Audouin et al. 1842: pl. 22).

Family-group name: FÉRONIENS Dejean, 1825. Stem: Feroni-.

Figulus W.S. MacLeay, 1819: 96, 109. Current status: valid genus in LUCANIDAE: LUCANINAE: FIGULINI.

Type species: Lucanus striatus G.-A. Olivier, 1789, by monotypy.

Family-group name: FIGULIDAE Burmeister, 1847. Stem: Figul-.

Flagrax Kazantsev, 1992: 38. Current status: valid genus in LYCIDAE: EROTINAE: SLIPINSKIINI.

Type species: Stadenus auberti Bourgeois, 1881, by original designation.

Family-group name: FLAGRAXINA Kazantsev, 2004. Stem: Flagrax-.

Flaminius Kuwert, 1891: 185. Current status: valid genus in PASSALIDAE: PASSALINAE: SOLENOCYCLINI.

Type species: Flaminius nonfriedi Kuwert, 1891, by monotypy.

Family-group name: FLAMININAE Kuwert, 1891. Stem: Flamini-.

Foadia Pakaluk, 1985: 406. Current status: valid genus in CORYLOPHIDAE: CORYLOPHINAE: FOADIINI.

Type species: Foadia maculata Pakaluk, 1985, by original designation.

Family-group name: FOADIINI Ślipiński, Tomaszewska & Lawrence, 2009. Stem: Foadi-.

Foranotum Nabozhenko & Sadeghi, 2017: 165. Current status: valid genus in TENEBRIONIDAE: KUHITANGIINAE: FORANOTINI.

Type species: Foranotum perforatum Nabozhenko & Sadeghi, 2017, by original designation.

Family-group name: FORANOTINI Nabozhenko & Sadeghi, 2017: 165. Stem: Foranot-.

Forcipator Maindron, 1904: 265. Current status: valid genus in CARABIDAE: SCARITINAE: FORCIPATORINI. Note: new replacement name for Oxystomus Dejean, 1825.

Type species: Oxystomus cylindricus Dejean, 1825, by monotypy (type fixation for Oxystomus Dejean).

Family-group name: FORCIPATORINA Bänninger, 1938. Stem: Forcipator-.

Formicomus LaFerté-Sénectère in Guérin-Méneville, 1849 [No 25]: 1. Current status: junior synonym of Anthelephila Hope, 1833 in ANTHICIDAE: ANTHICINAE: FORMICOMINI. Note: unjustified emendation of Formicoma Motschulsky, 1845 (see Kejval 2003: 382).

Type species: Carabus pedestris Rossi, 1790, by original designation (type fixation for Formicoma Motschulsky).

Family-group name: FORMICOMINI Bonadona, 1974. Stem: Formicom-.

Fornax Laporte, 1835: 167, 172. Current status: valid genus in EUCNEMIDAE: MACRAULACINAE: MACRAULACINI.

Type species: Fornax ruficollis Laporte, 1835, by monotypy.

Family-group name: FORNACINI Beaulieu, 1919. Stem: Fornac-. Note: as pointed out by Bouchard and Bousquet (2020a: 87) the names FORNACINI Beaulieu, 1919 and DROMAEOLINI Beaulieu, 1919 take precedence over MACRAULACINAE/-INI Fleutiaux, 1923 and an application to the Commission is necessary to preserve usage of MACRAULACINAE/-INI as valid.

Forsteria Tippmann, 1960: 211. Current status: junior synonym of Falsamblesthis Breuning, 1955 in CERAMBYCIDAE: LAMIINAE: FORSTERIINI.

Type species: Forsteria unguicularis Tippmann, 1960, by original designation.

Family-group name: FORSTERIINI Tippmann, 1960. Stem: Forsteri-.

Fruhstorferia Kolbe, 1894: 3. Current status: valid genus in SCARABAEIDAE: RUTELINAE: RUTELINI: RUTELINA.

Type species: Fruhstorferia javana Kolbe, 1894, by monotypy.

Family-group name: FRUHSTORFERIINA Ohaus, 1918. Stem: Fruhstorferi-.

Fulcidax Crotch, 1870b: 214. Current status: junior synonym of Poropleura Lacordaire, 1848 in CHRYSOMELIDAE: CRYPTOCEPHALINAE: FULCIDACINI. Note: this name was first proposed by Voet (1806: 33) but it is not available because Voet’s work is not consistently binominal (ICZN 1999a, Article 11.4); the name was subsequently listed in synonymy by Crotch (1870b: 214) and considered as valid by Clavareau (1913: 222), therefore, the name Fulcidax is available and dates from its first publication as a synonym (ICZN 1999a, Article 11.6.1).

Type species: Clytra monstrosa Fabricius, 1798, by subsequent designation (Chamorro-Lacayo and Konstantinov 2009: 76). Note: no species were originally included in this genus by Crotch (1870b); the first ones were the six species listed by Clavareau (1913: 223).

Family-group name: FULCIDACINA Jakobson, 1924. Stem: Fulcidac-.

Fustiger J.L. LeConte, 1866c: 108, 109. Current status: valid genus in STAPHYLINIDAE: PSELAPHINAE: CLAVIGERITAE: CLAVIGERINI.

Type species: Articerus brasiliensis Westwood, 1856, by subsequent designation (Newton and Chandler 1989: 64).

Family-group name: FUSTIGERINI Jeannel, 1949. Stem: Fustiger-.

Gahania Distant, 1907: 423. Current status: valid genus in CERAMBYCIDAE: CERAMBYCINAE: GAHANIINI.

Type species: Gahania simmondsi Distant, 1907, by monotypy.

Family-group name: GAHANIINI Quentin & Villiers, 1969. Stem: Gahani-.

Galba Latreille, 1829a: 451. Current status: invalid senior synonym of Galbites Fleutiaux, 1918 in EUCNEMIDAE: EUCNEMINAE: GALBITINI. Note: junior homonym of Galba Schrank, 1803 [Mollusca].

Type species: Galba marmorata Guérin-Méneville, 1829, by subsequent monotypy (Guérin- Méneville 1829a: pl. 12).

Family-group name: GABINI Beaulieu, 1919. Stem: Galb-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

Galbella Westwood, 1848a: 83. Current status: valid genus in BUPRESTIDAE: GALBELLINAE.

Type species: Galbella violacea Westwood, 1848, by monotypy.

Family-group name: GALBELLINAE Reitter, 1911. Stem: Galbell-.

Galbites Fleutiaux, 1918a: 59. Current status: valid genus in EUCNEMIDAE: EUCNEMINAE: GALBITINI. Note: new replacement name for Galba Latreille, 1829.

Type species: Galba marmorata Guérin-Méneville, 1829, by subsequent monotypy (Guérin- Méneville 1829a: pl. 12) (type fixation for Galba Latreille).

Family-group name: GALBITINI Muona, 1991. Stem: Galbit-.

Galerita Fabricius, 1801a: xxiii, 214. Current status: valid genus in CARABIDAE: HARPALINAE: GALERITINI: GALERITINA. Note: the senior homonym Galerita Gouan, 1770 [Aves] was suppressed both for the Principles of Priority and Homonymy and placed on the Official Index of Rejected and Invalid Names in Zoology while Galerita Fabricius, 1801 was placed on the Official List of Generic Names in Zoology (ICZN 1968b).

Type species: Carabus americanus Linnaeus, 1758, by subsequent designation (Latreille 1810: 426; see ICZN 1968b). Note: name placed on the Official List of Specific Names in Zoology (ICZN 1968b).

Family-group name: GALERITIDAE W. Kirby, 1825. Stem: Galerit-.

Galeritina Jeannel, 1949b: 1058. Current status: junior synonym of Galerita Fabricius, 1801 in CARABIDAE: HARPALINAE: GALERITINI: GALERITINA. Note: new replacement name for Galerita Fabricius, 1801.

Type species: Carabus americanus Linnaeus, 1758, by subsequent designation (Latreille 1810: 426) (type fixation for Galerita Fabricius).

Family-group name: GALERITININI Jeannel, 1949. Stem: Galeritin-. Note: name placed on the Official List of Rejected and Invalid Family-Group Names in Zoology (ICZN 1968b).

Galeritula Strand, 1936: 168. Current status: junior synonym of Galerita Fabricius, 1801 in CARABIDAE: HARPALINAE: GALERITINI: GALERITINA. Note: new replacement name for Galerita Fabricius, 1801.

Type species: Carabus americanus Linnaeus, 1758, by subsequent designation (Latreille 1810: 426) (type fixation for Galerita Fabricius).

Family-group name: GALERITULINI Jedlička, 1963. Stem: Galeritul-. Note: name placed on the Official List of Rejected and Invalid Family-Group Names in Zoology (ICZN 1968b).

Galeruca Geoffroy, 1762: 251. Current status: valid genus in CHRYSOMELIDAE: GALERUCINAE: GALERUCINI. Note: name placed on the Official List of Generic Names in Zoology (ICZN 1994a).

Type species: Chrysomela tanaceti Linnaeus, 1758, by subsequent designation (Latreille 1810: 432; see ICZN 1994a). Note: name placed on the Official List of Specific Names in Zoology (ICZN 1994a).

Family-group name: GALERUCAE Latreille, 1802. Stem: Galeruc-.

Gallerucida Motschulsky, 1861: 24. Current status: valid genus in CHRYSOMELIDAE: GALERUCINAE: HYLASPINI.

Type species: Gallerucida bifasciata Motschulsky, 1861, by monotypy.

Family-group name: GALLERUCIDINI Gressitt & Kimoto, 1963. Stem: Gallerucid-.

Gallerucidia Chaudoir, 1872b: 415, 417. Current status: valid genus in CARABIDAE: HARPALINAE: LEBIINI: GALLERUCIDIINA.

Type species: Gallerucidia octonotata Chaudoir, 1872, by subsequent designation (Lorenz 1998: 89).

Family-group name: GALLERUCIDIAE Chaudoir, 1872. Stem: Gallerucidi-.

Galloisia Hustache, 1920a: 493. Current status: valid genus in CURCULIONIDAE: CURCULIONINAE: MICROSTYLINI.

Type species: Galloisia inflata Hustache, 1920, by original designation.

Family-group name: GALLOISIINAE Morimoto, 1962. Stem: Galloisi-.

Ganglbaueria Semenov, 1891: 378. Current status: valid genus in OEDEMERIDAE: OEDEMERINAE: ASCLERINI.

Type species: Ganglbaueria collaris Semenov, 1891, by monotypy.

Family-group name: GANGLBAUERIIDAE Semenov, 1894. Stem: Ganglbaueri-.

Gasterocercus Laporte & Brullé, 1828: 198. Current status: valid genus in CURCULIONIDAE: CRYPTORHYNCHINAE: GASTEROCERCINI.

Type species: Gasterocercus dumerilii Laporte & Brullé, 1828 (= Curculio depressirostris Fabricius, 1792), by subsequent designation (Chevrolat 1845: 30). Note: Chevrolat (1845) selected “G. depressicornis Fab.” (a lapsus for G. depressirostris Fab.) as type species, a species not originally included, but since he listed it as a synonym of Gasterocercus dumerilii Laporte & Brullé, 1828, a species originally included, he is deemed to have designated the latter species as the type (ICZN 1999a, Article 69.2.2).

Family-group name: GASTEROCERCINI Zherikhin, 1991. Stem: Gasterocerc-.

Gastraulacus Guérin-Méneville, 1843b: 188. Current status: valid genus in EUCNEMIDAE: EUCNEMINAE: EUCNEMINI.

Type species: Galba bisulcatus Latreille, 1834, by original designation.

Family-group name: GASTRAULACI Fleutiaux, 1902. Stem: Gastraulac-.

Gastrophysa Chevrolat in Dejean, 1836a: 405. Current status: valid genus in CHRYSOMELIDAE: CHRYSOMELINAE: CHRYSOMELINI.

Type species: Chrysomela polygoni Linnaeus, 1758, by subsequent designation (Chevrolat 1845: 34).

Family-group name: GASTROPHYSINA Kippenberg, 2010. Stem: Gastrophys-. Note: junior homonym of GASTROPHYSINI Harting, 1864 (type genus Gastrophysus J. Müller, 1843) in Pisces.

Gehringia Darlington, 1933: 110. Current status: valid genus in CARABIDAE: GEHRINGIINAE: GEHRINGIINI: GEHRINGIINA.

Type species: Gehringia olympica Darlington, 1933, by original designation.

Family-group name: GEHRINGIINI Darlington, 1933. Stem: Gehringi-.

Gempylodes Pascoe, 1863c: 123, 132. Current status: valid genus in ZOPHERIDAE: COLYDIINAE: GEMPYLODINI.

Type species: Gempylodes macer Pascoe, 1863, by monotypy.

Family-group name: GEMPYLODINI Sharp, 1893. Stem: Gempylod-.

Genecerus Walker, 1871: 13. Current status: valid genus in DASCILLIDAE: KARUMIINAE: GENECERINI.

Type species: Genecerus cervinus Walker, 1871, by monotypy.

Family-group name: GENECERINI Pic, 1914. Stem: Genecer-.

Geniates W. Kirby, 1819: 401. Current status: valid genus in SCARABAEIDAE: RUTELINAE: GENIATINI.

Type species: Geniates barbatus W. Kirby, 1819, by monotypy.

Family-group name: GENIATIDAE Burmeister, 1844. Stem: Geniat-.

Genuchus W. Kirby, 1825: 569. Current status: valid genus in SCARABAEIDAE: CETONIINAE: CREMASTOCHEILINI: GENUCHINA.

Type species: Cetonia cruenta Fabricius, 1787 (= Cetonia hottentotta Fabricius, 1775), by original designation.

Family-group name: GENUCHINA Krikken, 1984. Stem: Genuch-.

Genyocerus Motschulsky, 1858a: 68. Current status: valid genus in CURCULIONIDAE: PLATYPODINAE: TESSEROCERINI: DIAPODINA.

Type species: Genyocerus albipennis Motschulsky, 1858, by monotypy.

Family-group name: GENYOCERINAE Hopkins, 1915. Stem: Genyocer-.

Geobaenus Dejean, 1829: 5, 402. Current status: valid genus in CARABIDAE: HARPALINAE: GEOBAENINI.

Type species: Geobaenus lateralis Dejean, 1829, by monotypy.

Family-group name: GEOBAENIDES Péringuey, 1896. Stem: Geobaen-.

Geochus Broun, 1882a: 409. Current status: valid genus in CURCULIONIDAE: MOLYTINAE: PHRYNIXINI: GEOCHINA. Note: new replacement name for Geophilus Broun, 1880; the same name was also proposed subsequently by Broun (1882b: 128) on [31] May, as opposed to 1 May in Broun (1882a).

Type species: Geophilus inaequalis Broun, 1880, by monotypy (type fixation for Geophilus Broun).

Family-group name: GEOCHINA Legalov, 2020a: 328. Stem: Geoch-.

Geomorus Schönherr, 1823: 1141. Current status: junior synonym of Cleonis Dejean, 1821 in CURCULIONIDAE: LIXINAE: CLEONINI.

Type species: Curculio sulcirostris Linnaeus, 1767, by original designation.

Family-group name: GEOMORIDES Schönherr, 1823. Stem: Geomor-. Note: nomen oblitum (see Bouchard et al. 2011: 620).

Geonemus Schönherr, 1833: 13. Current status: valid genus in CURCULIONIDAE: ENTIMINAE: POLYDRUSITAE: GEONEMINI. Note: this genus-group name is a junior homonym of Geonemus Guérin-Méneville, 1831 [Coleoptera: CURCULIONIDAE]; Alonso Zarazaga and Lyal (1999: 159) mentioned that an application will be submitted to the Commission to suppress the senior homonym and that meanwhile current usage should be maintained; name placed on the Official List of Generic Names in Zoology (ICZN 1988d).

Type species: Curculio flabellipes G.-A. Olivier, 1807, by plenary power (see ICZN 1988d). Note: name placed on the Official List of Specific Names in Zoology (ICZN 1988d).

Family-group name: GEONEMIDAE Gistel, 1848. Stem: Geonem-.

Geopaederus Gistel, 1848b: x, xi. Current status: junior synonym of Paederus Fabricius, 1775 in STAPHYLINIDAE: PAEDERINAE: PAEDERINI: PAEDERINA. Note: unnecessary new replacement name for Paederus Fabricius, 1775.

Type species: Staphylinus riparius Linnaeus, 1758, by subsequent designation (Latreille 1810: 427) (type fixation for Paederus Fabricius).

Family-group name: GEOPAEDERIDAE Gistel, 1848. Stem: Geopaeder-.

Georissus Latreille, 1809a: 377. Current status: valid genus in GEORISSIDAE. Note: name placed on the Official List of Generic Names in Zoology (ICZN 1998).

Type species: Pimelia pygmaea Fabricius, 1798 (= Byrrhus crenulatus Rossi, 1794), by monotypy (see ICZN 1998). Note: Byrrhus crenulatus Rossi, 1794 placed on the Official List of Specific Names in Zoology (ICZN 1998).

Family-group name: GÉORISSITES Laporte, 1840. Stem: Georiss-. Note: senior homonym of GEORISSINAE W.T. Blandford, 1864 (type genus Georissa W.T. Blandford, 1864) in Mollusca.

Geostiba C.G. Thomson, 1858: 33. Current status: valid genus in STAPHYLINIDAE: ALEOCHARINAE: GEOSTIBINI. Note: Evanystes Gistel, 1856, a senior synonym of Geostiba, has been suppressed for the purposes of the Principle of Priority but not for the Principle of Homonymy and Geostiba C.G. Thomson, 1858 placed on the Official List of Generic Names in Zoo1ogy (ICZN 2005a).

Type species: Aleochara circellaris Gravenhorst, 1806, by monotypy (see ICZN 2005a). Note: name placed on the Officia1 List of Specific Names in Zoo1ogy (ICZN 2005a).

Family-group name: GEOSTIBAE Seevers, 1978. Stem: Geostib-.

Geotrupes Latreille, 1797: 6. Current status: valid genus in GEOTRUPIDAE: GEOTRUPINAE: GEOTRUPINI. Note: name placed on the Official List of Generic Names in Zoology (ICZN 1955b).

Type species: Scarabaeus stercorarius Linnaeus, 1758, by plenary power (see ICZN 1955b). Note: name placed on the Official List of Specific Names in Zoology (ICZN 1955b).

Family-group name: GEOTRUPINI Latreille, 1802. Stem: Geotrup-.

Geralius Harold, 1869b: 123. Current status: valid genus in BUPRESTIDAE: AGRILINAE: CORAEBINI: GERALIINA. Note: new replacement name for Acanthopygus Deyrolle, 1864.

Type species: Stenogaster furciventris Chevrolat, 1838, by monotypy (type fixation for Acanthopygus Deyrolle).

Family-group name: GERALIINA Cobos, 1988. Stem: Gerali-.

Gerania Audinet-Serville, 1835a: 70. Current status: valid genus in CERAMBYCIDAE: LAMIINAE: LAMIINI.

Type species: Saperda boscii Fabricius, 1801, by monotypy.

Family-group name: GERANITAE J. Thomson, 1864. Stem: Gerani-.

Germarica Blackburn, 1888: 257. Current status: valid genus in BUPRESTIDAE: AGRILINAE: APHANISTICINI: GERMARICINA.

Type species: Germarica casuarinae Blackburn, 1888, by monotypy.

Family-group name: GERMARICINI Cobos, 1979. Stem: Germaric-.

Ghanius Ahmad, 1973: 449. Current status: valid genus in COCCINELLIDAE: COCCINELLINAE: SHIROZUELLINI.

Type species: Ghanius karachiensis Ahmad, 1973, by monotypy.

Family-group name: GHANIINI Ahmad, 1973. Stem: Ghani-.

Ghidinia Pavan, 1939: 106. Current status: junior synonym of Boldoria Jeannel, 1924 in LEIODIDAE: CHOLEVINAE: LEPTODIRINI: PHOLEUINA.

Type species: Ghidinia morettii Pavan, 1939, by monotypy.

Family-group name: GHIDINIINA Guéorguiev, 1974. Stem: Ghidini-.

Gibbium Scopoli, 1777: 505. Current status: valid genus in PTINIDAE: PTININAE: GIBBIINI.

Type species: Ptinus scotias Fabricius, 1781 (= Scotias psylloides Czenpinski, 1778), by subsequent monotypy (Latreille 1804d: 37).

Family-group name: GIBBIITES Jacquelin du Val, 1860. Stem: Gibbi-.

Gietella Constantin & Menier, 1987: 53. Current status: valid genus in RHADALIDAE: GIETELLINAE.

Type species: Gietella fortunata Constantin & Menier, 1987, by original designation.

Family-group name: GIETELLINAE Constantin & Menier, 1987. Stem: Gietell-.

Gilletinus Boucomont, 1932: 264. Current status: valid genus in BOLBOCERATIDAE: BOLBOCERATINAE: GILLETININI.

Type species: Bolboceras multicostatus Van Lansberge, 1885, by original designation.

Family-group name: GILLETININI Nikolajev, 1990. Stem: Gilletin-.

Ginema Ball & Shpeley, 2002: 83. Current status: valid genus in CARABIDAE: HARPALINAE: GINEMINI.

Type species: Ginema thomasi Ball & Shpeley, 2002, by original designation.

Family-group name: GINEMINI Ball & Shpeley, 2002. Stem: Ginem-.

Glacicavicola Westcott, 1968: 3. Current status: valid genus in LEIODIDAE: CATOPOCERINAE: GLACICAVICOLINI.

Type species: Glacicavicola bathyscioides Westcott, 1968, by original designation.

Family-group name: GLACICAVICOLINAE Westcott, 1968. Stem: Glacicavicol-.

Glandularia L.W. Schaufuss, 1889: 3. Current status: junior synonym of Napochus C.G. Thomson, 1859 in STAPHYLINIDAE: SCYDMAENINAE: SCYDMAENITAE: STENICHNINI.

Type species: Glandularia fricatoris L.W. Schaufuss, 1889, by original designation.

Family-group name: GLANDULARIIDAE L.W. Schaufuss, 1889. Stem: Glandulari-.

Glaphyrometopus Pierce, 1913: 406. Current status: valid genus in CURCULIONIDAE: ENTIMINAE: POLYDRUSITAE: NAUPACTINI: NAUPACTINA.

Type species: Glaphyrometopus ornithodorus Pierce, 1913, by monotypy.

Family-group name: GLAPHYROMETOPI Pierce, 1913. Stem: Glaphyro­metop-.

†Glaphyroptera Heer, 1852: 13. Current status: valid genus in BUPRESTIDAE: BUPRESTINAE: †GLAPHYROPTERINI. Note: Lower Jurassic (Switzerland).

Type species: †Glaphyroptera insignis Heer, 1852, by subsequent designation (Handlirsch 1906: 452).

Family-group name: †GLAPHYROPTERIDAE Pongrácz, 1935. Stem: Glaphyropter-.

Glaphyrus Latreille, 1802b: 150. Current status: valid genus in GLAPHYRIDAE: GLAPHYRINAE.

Type species: Melolontha serratulae Fabricius, 1792, by monotypy. Note: the type species often listed for this genus is Scarabaeus maurus Linnaeus, 1758 (e.g., Nikodým and A. Bezděk 2016: 95) but Latreille (1802b: 151) listed a single species, serratulae Fabricius in his new genus; both species are currently included in the nominotypical subgenus of Glaphyrus.

Family-group name: GLAPHYRIDAE W.S. MacLeay, 1819. Stem: Glaphyr-.

Glaresis Erichson, 1848: 925. Current status: valid genus in GLARESIDAE.

Type species: Glaresis rufa Erichson, 1848, by monotypy.

Family-group name: GLARESINI Prudhomme de Borre, 1886. Stem: Glares-.

Glaucytes J. Thomson, 1858a: 423. Current status: valid genus in CERAMBYCIDAE: CERAMBYCINAE: GLAUCYTINI. Note: new replacement name for Leptocera Dejean, 1821.

Type species: Cerambyx scriptus Fabricius, 1798 (= Cerambyx interruptus G.-A. Olivier, 1790), by monotypy (type fixation for Leptocera Dejean).

Family-group name: GLAUCYTIDES Lacordaire, 1868. Stem: Glaucyt-.

Glenea Newman, 1842a: 301. Current status: valid genus in CERAMBYCIDAE: LAMIINAE: SAPERDINI. Note: new replacement name for Sphenura Dejean, 1835.

Type species: Saperda novemguttata Guérin-Méneville, 1825, by subsequent designation (Breuning 1956: 2). Note: this species is not a nominal species originally included in the senior synonym Sphenura Dejean (see Article 67.8.1, ICZN 1999a); however, the three originally included nominal species are currently placed in other genera; a request should be submitted to the Commission to retain Breuning’s type species fixation (see Bousquet et al. 2009: 37).

Family-group name: GLENEÏTAE J. Thomson, 1864. Stem: Glene-.

Glischrochilus Reitter, 1873: 162. Current status: valid genus in NITIDULIDAE: CRYPTARCHINAE: CRYPTARCHINI.

Type species: Silpha quadripustulata Linnaeus, 1761 (= Silpha quadripunctata Linnaeus, 1758), by subsequent designation (Parsons 1943: 264).

Family-group name: GLISCHROCHILINI Chagnon, 1934. Stem: Glischrochil-.

Globa Raffray, 1887: 37. Current status: valid genus in STAPHYLINIDAE: PSELAPHINAE: GONIACERITAE: BRACHYGLUTINI: BRACHYGLUTINA.

Type species: Globa longipes Raffray, 1887, by monotypy.

Family-group name: GLOBINA Jeannel, 1959. Stem: Glob-.

Globulosis García, 2001: 153. Current status: valid genus in HYDROPHILIDAE: ACIDOCERINAE.

Type species: Globulosis hemisphericus García, 2001, by original designation.

Family-group name: GLOBULINA García, 2001. Stem: Globulos-.

Gloeosoma Wollaston, 1854: 480. Current status: valid genus in CORYLOPHIDAE: CORYLOPHINAE: RYPOBIINI.

Type species: Gloeosoma velox Wollaston, 1854, by monotypy.

Family-group name: GLOEOSOMATINI Bowestead, 1999. Stem: Gloeosomat-.

Glostatus Schedl, 1939: 386. Current status: valid genus in CURCULIONIDAE: SCOLYTINAE: XYLOCTONINI: GLOSTATINA.

Type species: Glostatus declividepressus Schedl, 1939, by monotypy.

Family-group name: GLOSTATINA Jordal, 2023: 203. Stem: Glostat-.

Glycyphana Burmeister, 1842: 345. Current status: valid genus in SCARABAEIDAE: CETONIINAE: CETONIINI: CETONIINA.

Type species [suggested]: Cetonia horsfieldii Hope, 1831, by subsequent designation (Arrow 1910: 120). Note: the valid type species fixation is that of Duponchel (1867: 535), who selected Cetonia tricolor G.-A. Olivier, 1789, but that nominal species is the type species of the genus Glycosia Schoch, 1896; an application to the Commission is necessary to fix Cetonia horsfieldii Hope, 1831 as the type species of Glycyphana Burmeister, 1842 to conserve the current usage of the genus-group name.

Family-group name: GLYCYPHANAE Schoch, 1894. Stem: Glycyphan-.

Glypholoma Jeannel, 1962b: 482. Current status: valid genus in STAPHYLINIDAE: GLYPHOLOMATINAE.

Type species: Glypholoma pustuliferum Jeannel, 1962, by original designation.

Family-group name: GLYPHOLOMINI Jeannel, 1962. Stem: Glypholomat-.

Glyptoma Erichson, 1839b: 32. Current status: valid genus in STAPHYLINIDAE: OSORIINAE: THORACOPHORINI: GLYPTOMINA.

Type species: Glyptoma crassicorne Erichson, 1840, by subsequent designation (Duponchel 1841b: 57). Note: no species were originally included in this genus by Erichson (1839a); the first ones were the six species listed by Erichson (1840a: 908–910).

Family-group name: GLYPTOMINA Newton & Thayer, 1992. Stem: Glyptom-.

Glyptoscelimorpha G.H. Horn, 1893b: 137. Current status: valid genus in SCHIZOPODIDAE: DYSTAXIINI.

Type species: Glyptoscelimorpha marmorata G.H. Horn, 1893, by monotypy.

Family-group name: GLYPTOSCELIMORPHINI Cobos, 1963. Stem: Glyptoscelimorph-.

Glyptus Brullé, 1835: 83. Current status: valid genus in CARABIDAE: HARPALINAE: GLYPTINI. Note: the senior homonym Glyptus Hoffmannsegg, 1818 (and all uses of that name prior to the publication of Glyptus Brullé, 1835) was suppressed for the purposes both the Principle of Priority and Homonymy and placed on the Official Index of Rejected and Invalid Names in Zoology while Glyptus Brullé, 1835 was placed on the Official List of Generic Names in Zoology (ICZN 1985a).

Type species: Glyptus sculptilis Brullé, 1835, by monotypy (see ICZN 1985a). Note: name placed on the Official List of Specific Names in Zoology (ICZN 1985a).

Family-group name: GLYPTI G.H. Horn, 1881. Stem: Glypt-. Note: senior homonym of GLYPTINI Cushman & Rohwer, 1920 (type genus Glypta Gravenhorst, 1829) in Hymenoptera.

Gnaphalocnemis Heller, 1901: 10. Current status: junior synonym of Pelopides Kuwert, 1896 in PASSALIDAE: PASSALINAE: MACROLININI. Note: new replacement name for Eriocnemis Kaup, 1868.

Type species: Passalus tridens Wiedemann, 1823, by subsequent designation (Hincks and Dibb 1935: 85).

Family-group name: GNAPHALOCNEMINAE Heller, 1901. Stem: Gnaphalocnemid-.

Gnaptor Brullé, 1831: 254. Current status: valid genus in TENEBRIONIDAE: BLAPTINAE: BLAPTINI: GNAPTORINA.

Type species: Tenebrio spinimanus Pallas, 1781, by monotypy.

Family-group name: GNAPTORINA G.S. Medvedev, 2001. Stem: Gnaptor-.

Gnaptorina Reitter, 1887: 364. Current status: valid genus in TENEBRIONIDAE: BLAPTINAE: BLAPTINI: GNAPTORININA.

Type species: Gnaptorina felicitana Reitter, 1887, by monotypy.

Family-group name: GNAPTORININA G.S. Medvedev, 2001. Stem: Gnaptorin-.

Gnathidium Gebien, 1921a: 41. Current status: valid genus in TENEBRIONIDAE: DIAPERINAE: GNATHIDIINI: GNATHIDIINA.

Type species: Gnathidium cephalotes Gebien, 1921, by monotypy.

Family-group name: GNATHIDIINAE Gebien, 1921. Stem: Gnathidi-.

Gnathocera W. Kirby, 1825: 571. Current status: valid genus in SCARABAEIDAE: CETONIINAE: GOLIATHINI: RHOMBORHININA.

Type species: Gnathocera vitticollis W. Kirby, 1825 (= Scarabaeus trivittatus Swederus, 1787), by original designation. Note: Gnathocera immaculata W. Kirby, 1825 is usually listed as type species of this genus (e.g., J. Bezděk 2016: 394) but W. Kirby (1825: 572) unambiguously designated Gnathocera vitticollis as type species; both nominal species are currently included in Gnathocera W. Kirby, 1825.

Family-group name: GNATHOCERIDAE Schoch, 1894. Stem: Gnathocer-.

Gnathosia Fischer, 1821a: 13. Current status: valid genus in TENEBRIONIDAE: PIMELIINAE: TENTYRIINI.

Type species: Gnathosia glabra Fischer, 1821, by monotypy.

Family-group name: GNATHOSIIDES Lacordaire, 1859. Stem: Gnathosi-.

Gnathotrichus Eichhoff, 1869: 275. Current status: valid genus in CURCULIONIDAE: SCOLYTINAE: CORTHYLINI: CORTHYLINA.

Type species: Gnathotrichus corthyloides Eichhoff, 1869 (= Tomicus materiarius Fitch, 1858), by monotypy.

Family-group name: GNATHOTRICHINA Balachowsky, 1949. Stem: Gnathotrich-.

Gnathymenus Solier, 1849: 326. Current status: valid genus in STAPHYLINIDAE: PAEDERINAE: PAEDERINI: DOLICAONINA.

Type species: Gnathymenus apterus Solier, 1849, by monotypy.

Family-group name: GNATIMENITOS Solier, 1849. Stem: Gnathymen-.

Gnatocerus Thunberg, 1814: 47. Current status: valid genus in TENEBRIONIDAE: DIAPERINAE: DIAPERINI: ADELININA. Note: name placed on the Official List of Generic Names in Zoology (ICZN 1975).

Type species: Gnatocerus ruber Thunberg, 1814 (= Trogossita cornuta Fabricius, 1798), by monotypy (see ICZN 1975). Note: Trogossita cornuta Fabricius, 1798 was placed on the Official List of Specific Names in Zoology (ICZN 1975).

Family-group name: GNATHOCERINI Skopin, 1978. Stem: Gnatocer-.

Gnoma Fabricius, 1801a: xxii. Current status: valid genus in CERAMBYCIDAE: LAMIINAE: LAMIINI.

Type species: Cerambyx longicollis Fabricius, 1787, by subsequent designation (J. Thomson 1864: 82). Note: no species were originally included in this genus by Fabricius (1801a); the first ones were the four species listed by Fabricius (1801b: 315–317).

Family-group name: GNOMITAE J. Thomson, 1860. Stem: Gnom-.

Gnostus Westwood, 1855a: 90. Current status: valid genus in PTINIDAE: PTININAE: GNOSTINI.

Type species: Gnostus formicicola Westwood, 1855, by monotypy.

Family-group name: GNOSTIDAE King, 1866. Stem: Gnost-.

†Gobicar Gratshev & Zherikhin, 1999: 40. Current status: valid genus in CARIDAE: †BAISSORHYNCHINAE: †GOBICARINI. Note: Upper Jurassic (Mongolia).

Type species: †Gobicar ponomarenkoi Gratshev & Zherikhin, 1999, by original designation.

Family-group name: †GOBICARINI Legalov, 2009. Stem: Gobicar-.

Goes J.L. LeConte, 1852a: 150. Current status: valid genus in CERAMBYCIDAE: LAMIINAE: LAMIINI.

Type species: Cerambyx tigrinus DeGeer, 1775, by subsequent designation (J. Thomson 1864: 76).

Family-group name: GOES J.L. LeConte, 1873. Stem: Goet-.

Goliathopsis Janson, 1881: 609. Current status: valid genus in SCARABAEIDAE: CETONIINAE: CREMASTOCHEILINI: GOLIATHOPSIDINA.

Type species: Goliathopsis cervus Janson, 1881, by monotypy. Note: Goliathopsis cervus is often listed as a junior synonym of Pilinurgus despectus Westwood, 1874, but Xu and Qiu (2020: 95) consider Janson’s name as valid.

Family-group name: GOLIATHOPSIDINA Krikken, 1984. Stem: Goliathopsid-.

Goliathus Lamarck, 1801: 209. Current status: valid genus in SCARABAEIDAE: CETONIINAE: GOLIATHINI: GOLIATHINA.

Type species: Scarabaeus goliatus Linnaeus, 1771, by subsequent designation (Latreille 1810: 429).

Family-group name: GOLIATHIDES Latreille, 1829. Stem: Goliath-.

Gompelia Alonso-Zarazaga, 2010: 299. Current status: valid genus in ADERIDAE. Note: new replacement name for Olotelus Mulsant & Rey, 1866.

Type species: Xylophilus neglectus Jacquelin du Val, 1863, by subsequent designation (Alonso- Zarazaga 2010: 299) (type fixation for Olotelus Mulsant & Rey).

Family-group name: GOMPELIINA Bouchard, 2011. Stem: Gompeli-.

Gonatas Kaup, 1871: 50. Current status: valid genus in PASSALIDAE: PASSALINAE: MACROLININI.

Type species: Passalus naviculator Percheron, 1844, by subsequent designation (Gravely 1918: 108).

Family-group name: GONATINAE Kuwert, 1891. Stem: Gonat-. Note: junior homonym of GONATIDAE Hoyle, 1886 (type genus Gonatus J.E. Gray, 1849) in Cephalopoda.

Goniacerus Motschulsky, 1855a: 17. Current status: valid genus in STAPHYLINIDAE: PSELAPHINAE: GONIACERITAE: GONIACERINI.

Type species: Goniacerus gibbus Motschulsky, 1855, by monotypy.

Family-group name: GONIACERIDES Reitter, 1882. Stem: Goniacer-.

Goniadera Perty, 1832: 62. Current status: valid genus in TENEBRIONIDAE: LAGRIINAE: GONIADERINI. Note: two original spellings were used, including Gonyodera (p. 63); subsequently, Perty (1833: 14) used Goniadera and therefore acted as First Reviser (ICZN 1999a, Article 24.2.4).

Type species: Goniadera crenata Perty, 1832, by monotypy.

Family-group name: GONIADÉRIDES Lacordaire, 1859. Stem: Goniader-.

†Gonialaena Nabozhenko, Bukejs & Telnov, 2019: 254. Current status: valid genus in TENEBRIONIDAE: LAGRIINAE: †GONIALAENINI. Note: Eocene (Kaliningrad, Russia).

Type species: †Gonialaena groehni Nabozhenko, Bukejs & Telnov, 2019, by original designation.

Family-group name: †GONIALAENINI Nabozhenko, Bukejs & Telnov, 2019: 254. Stem: Gonialaen-.

Goniastes Westwood, 1870: 125. Current status: valid genus in STAPHYLINIDAE: PSELAPHINAE: GONIACERITAE: GONIACERINI.

Type species: Goniastes sulcifrons Westwood, 1870, by monotypy.

Family-group name: GONIASTINI L.W. Schaufuss, 1872. Stem: Goniast-.

Goniochenia Weise, 1896: 10. Current status: valid genus in CHRYSOMELIDAE: CASSIDINAE: GONIOCHENIINI.

Type species: Mesomphalia quadraticollis Boheman, 1850, by subsequent designation (Spaeth 1914: 29).

Family-group name: GONIOCHENIITAE Spaeth, 1942. Stem: Goniocheni-.

Gonioctena Chevrolat in Dejean, 1836a: 403. Current status: valid genus in CHRYSOMELIDAE: CHRYSOMELINAE: CHRYSOMELINI.

Type species: Chrysomela viminalis Linnaeus, 1758, by subsequent designation (C.G. Thomson 1859: 158).

Family-group name: GONIOCTÈNES Motschulsky, 1860. Stem: Goniocten-.

Gonipterus Schönherr, 1833: 8, 456. Current status: valid genus in CURCULIONIDAE: HYPERINAE: GONIPTERINI.

Type species: Gonipterus lepidotus Gyllenhal, 1833, by original designation.

Family-group name: GONIPTÉRIDES Lacordaire, 1863. Stem: Gonipter-.

Gonocephalum Solier, 1834bc: 498. Current status: valid genus in TENEBRIONIDAE: BLAPTINAE: OPATRINI: OPATRINA.

Type species: Opatrum fuscum Herbst, 1793 (= Opatrum rusticum G.-A. Olivier, 1812), by subsequent designation (Gebien 1939: 443).

Family-group name: GONOCÉPHALATES Mulsant & Revelière, 1861. Stem: Gonocephal-.

Gonodera Mulsant, 1856a: 41. Current status: valid genus in TENEBRIONIDAE: ALLECULINAE: ALLECULINI: GONODERINA.

Type species: Cistela fulvipes Fabricius, 1792 (= Cistela luperus Herbst, 1783), by monotypy.

Family-group name: GONODERINA Seidlitz, 1896. Stem: Gonoder-.

Gonophora Chevrolat in Dejean, 1836a: 366. Current status: valid genus in CHRYSOMELIDAE: CASSIDINAE: GONOPHORINI.

Type species: Hispa haemorrhoidalis Weber, 1801, by monotypy.

Family-group name: GONOPHORITES Chapuis, 1875. Stem: Gonophor-.

Gonopus Latreille, 1828a: 580. Current status: valid genus in TENEBRIONIDAE: BLAPTINAE: PLATYNOTINI: PLATYNOTINA.

Type species: Blaps tibialis Fabricius, 1798, by monotypy.

Family-group name: GONOPIDES Lacordaire, 1859. Stem: Gonopod-.

Gracilia Audinet-Serville, 1834b: 81. Current status: valid genus in CERAMBYCIDAE: CERAMBYCINAE: GRACILIINI.

Type species: Callidium pygmaeum Fabricius, 1792 (= Saperda minuta Fabricius, 1781), by monotypy.

Family-group name: GRACILIAIRES Mulsant, 1839. Stem: Gracili-.

Grammoptera Audinet-Serville, 1835b: 215. Current status: valid genus in CERAMBYCIDAE: LEPTURINAE: LEPTURINI.

Type species: Leptura praeusta Fabricius, 1787 (= Leptura ustulata Schaller, 1783), by subsequent designation (Westwood 1838c: 41).

Family-group name: GRAMMOPTERINI Della Beffa, 1915. Stem: Grammopter-.

Graniger Motschulsky, 1864: 197. Current status: valid genus in CARABIDAE: HARPALINAE: HARPALINI: HARPALINA.

Type species: Graniger algirinus Motschulsky, 1864 (= Carterus femoralis Coquerel, 1858), by monotypy.

Family-group name: GRANIGERI Antoine, 1959. Stem: Graniger-.

Graphipterus Latreille, 1802b: 83. Current status: valid genus in CARABIDAE: HARPALINAE: GRAPHIPTERINI.

Type species: Carabus variegatus Fabricius, 1781 (= Carabus serrator Forsskål, 1775), by subsequent designation (Latreille 1810: 426).

Family-group name: GRAPHIPTERIDES Latreille, 1802. Stem: Graphipter-.

Graphisurus W. Kirby, 1837: 169. Current status: valid genus in CERAMBYCIDAE: LAMIINAE: ACANTHOCININI.

Type species: Cerambyx fasciatus DeGeer, 1775, by original designation.

Family-group name: GRAPHISURINI Leng, 1920. Stem: Graphisur-.

Graphoderus Dejean, 1833a: 54. Current status: valid genus in DYTISCIDAE: DYTISCINAE: ACILIINI. Note: name placed on the Official List of Generic Names in Zoology (ICZN 1961d).

Type species: Dytiscus cinereus Linnaeus, 1758, by subsequent designation (Westwood 1838c: 8; see ICZN 1961d). Note: name placed on the Official List of Specific Names in Zoology (ICZN 1961d).

Family-group name: GRAPHODERINI Houlbert, 1934. Stem: Graphoder-.

†Gratshevbelus Soriano, 2009: 100. Current status: valid genus in BELIDAE: †MONTSECBELINAE: †GRATSHEVBELINI. Note: Lower Cretaceous (France).

Type species: †Gratshevbelus erici Soriano, 2009, by original designation.

Family-group name: †GRATSHEVIBELINI Legalov, 2016: 1474. Stem: Gratshevbel-. Note: Legalov (2016: 1474) used the incorrect subsequent spelling Gratshevibelus Soriano to form the stem of his new family-group name.

Gromphas Dejean, 1836: 143. Current status: valid genus in SCARABAEIDAE: SCARABAEINAE: PHANAEINI: GROMPHADINA.

Type species: Coprobius lacordairii Oken, 1834, by monotypy. Note: see Cupello (2024: 32) for information on the type species.

Family-group name: GROMPHINA Zunino, 1985. Stem: Gromphad-.

Gronops Schönherr, 1823: 1142. Current status: valid genus in CURCULIONIDAE: CYCLOMINAE: HIPPORHININI.

Type species: Curculio lunatus Fabricius, 1775, by subsequent designation (Schönherr 1826: 13).

Family-group name: GRONOPINI Bedel, 1884. Stem: Gronop-.

Gryllica J. Thomson, 1860a: 120. Current status: valid genus in CERAMBYCIDAE: LAMIINAE: CALLIINI.

Type species: Onocephala picta Pascoe, 1858, by subsequent designation (L.S. Dillon and E.S. Dillon 1946: 156).

Family-group name: GRYLLICIDES Lacordaire, 1872. Stem: Gryllic-.

Guineauletes Legalov, 2003: 110. Current status: valid genus in ATTELABIDAE: RHYNCHITINAE: RHYNCHITITAE: AULETINI: AULETOBIINA.

Type species: Guineauletes mirabilis Legalov, 2003, by original designation.

Family-group name: GUINEAULETINA Legalov, 2003. Stem: Guineaulet-.

Guioperus Perty, 1832: 78. Current status: valid genus in CURCULIONIDAE: MOLYTINAE: GUIOPERINI.

Type species: Guioperus griseus Perty, 1832, by monotypy.

Family-group name: GUIOPÉRIDES Lacordaire, 1865. Stem: Guioper-.

Gyaritus Pascoe, 1858: 244. Current status: valid genus in CERAMBYCIDAE: LAMIINAE: GYARITINI.

Type species: Gyaritus hamatus Pascoe, 1858, by monotypy.

Family-group name: GYARITINI Breuning, 1950. Stem: Gyarit-.

Gymnetis W.S. MacLeay, 1819: 152. Current status: valid genus in SCARABAEIDAE: CETONIINAE: GYMNETINI: GYMNETINA. Note: name placed on the Official List of Generic Names in Zoology (ICZN 1967a).

Type species: Scarabaeus lanius Linnaeus, 1758, by plenary power (see ICZN 1967a). Note: name placed on the Official List of Specific Names in Zoology (ICZN 1967a).

Family-group name: GYMNETIDAE W. Kirby, 1827. Stem: Gymnet-.

Gymnetron Schönherr, 1825: 587. Current status: valid genus in CURCULIONIDAE: CURCULIONINAE: MECININI.

Type species: Curculio beccabungae Linnaeus, 1761, by subsequent designation (Schönherr 1826: 23).

Family-group name: GYMNETRINA C.G. Thomson, 1859. Stem: Gymnetr-. Note: junior homonym of GYMNETRINAE Swainson, 1839 (type genus Gymnetrus Bloch, 1795) in Pisces.

Gymnochila Erichson, 1844: 454. Current status: junior synonym of Gymnocheilis Dejean, 1835 in TROGOSSITIDAE: TROGOSSITINAE: TROGOSSITINI.

Type species: Trogosita vestita Griffith, 1832 (= Trogosita varia Fabricius, 1801), by monotypy.

Family-group name: GYMNOCHILIDES Lacordaire, 1854. Stem: Gymnochil-.

Gymnognathus Schönherr, 1826: 37. Current status: valid genus in ANTHRIBIDAE: ANTHRIBINAE: GYMNOGNATHINI.

Type species: Anthribus ancora Germar, 1823, by original designation.

Family-group name: GYMNOGNATHINI B.D. Valentine, 1960. Stem: Gymnognath-.

Gymnoloma Dejean, 1833b: 167. Current status: valid genus in SCARABAEIDAE: MELOLONTHINAE: HOPLIINI: HOPLIINA.

Type species: Melolontha atomaria Fabricius, 1781, by monotypy.

Family-group name: GYMNOLOMIDAE Burmeister, 1844. Stem: Gymnolomat-.

Gymnopleurus Illiger, 1803: 199. Current status: valid genus in SCARABAEIDAE: SCARABAEINAE: GYMNOPLEURINI.

Type species: Scarabaeus geoffroyi Fuesslin, 1775 (cited as “Ateuchus pilularius of Fabricius”), by subsequent designation (Partington 1836a: 693). Note: we follow Branco (2007: 12) in using Article 69.2.4 (ICZN 1999a) to determine the identity of the type species for this genus, although our data associated with the fixation of the type species (i.e., Partington 1836a: 693) predates the designation given by Reiche (1841: 212), which is typically reported in the literature.

Family-group name: GYMNOPLEURIDAE Streubel, 1846. Stem: Gymnopleur-.

Gymnusa Gravenhorst, 1806: 173. Current status: valid genus in STAPHYLINIDAE: ALEOCHARINAE: GYMNUSINI.

Type species: Gymnusa sinuata Gravenhorst, 1806 (= Staphylinus brevicollis Paykull, 1800), by monotypy.

Family-group name: GYMNUSIDA Heer, 1839. Stem: Gymnus-.

Gynaecomeloe Wellman, 1910b: 217. Current status: junior synonym of Cordylospasta G.H. Horn, 1876 in MELOIDAE: MELOINAE: EUPOMPHINI.

Type species: Megetra opaca G.H. Horn, 1868, by original designation.

Family-group name: GYNAECOMELODIDES Wellman, 1910. Stem: Gynaecomelo-.

Gyrinus Geoffroy, 1762: 193. Current status: valid genus in GYRINIDAE: GYRININAE: GYRININI. Note: name placed on the Official List of Generic Names in Zoology (ICZN 1994a).

Type species: Dytiscus natator Linnaeus, 1758, by subsequent designation (Latreille 1810: 426; see ICZN 1994a). Note: name placed on the Official List of Specific Names in Zoology (ICZN 1994a).

Family-group name: GYRINITES Latreille, 1810. Stem: Gyrin-.

Gyrohypnus Leach in Samouelle, 1819: 172. Current status: valid genus in STAPHYLINIDAE: STAPHYLININAE: XANTHOLININI. Note: name placed on the Official List of Generic Names in Zoology (ICZN 1983c, as Gyrohypnus Samouelle, 1819).

Type species: Staphylinus fracticornis O.F. Müller, 1776, by plenary power (see ICZN 1983c). Note: name placed on the Official List of Specific Names in Zoology (ICZN 1983c).

Family-group name: GYROHYPNIDAE W. Kirby, 1837. Stem: Gyrohypn-. Note: placed on the Official List of Family-Group Names in Zoology (ICZN 1996c, as GYROHYPNINI Kirby, 1837); younger name XANTHOLININI Erichson, 1839 given precedence over this name (ICZN 1996c).

Gyrophaena Mannerheim, 1830: 74. Current status: valid genus in STAPHYLINIDAE: ALEOCHARINAE: HOMALOTINI: GYROPHAENINA.

Type species: Staphylinus nanus Paykull, 1800, by subsequent designation (Westwood 1838c: 20).

Family-group name: GYROPHAENINI Kraatz, 1856. Stem: Gyrophaen-.

Habrobates Semenov, 1903b: 11. Current status: valid genus in TENEBRIONIDAE: PIMELIINAE: PIMELIINI: HABROBATINA.

Type species: Habrobates vernalis Semenov, 1903, by monotypy.

Family-group name: HABROBATINA Nabozhenko & Chigray, 2022 in Chigray et al. 2022: 5. Stem: Habrobat-.

Habrocerus Erichson, 1839a: 400. Current status: valid genus in STAPHYLINIDAE: HABROCERINAE.

Type species: Tachyporus capillaricornis Gravenhorst, 1806, by monotypy.

Family-group name: HABROCÉRIENS Mulsant & Rey, 1876. Stem: Habrocer-.

Habrophora Erichson, 1847c: 163. Current status: valid genus in CHRYSOMELIDAE: EUMOLPINAE: HABROPHORINI.

Type species: Habrophora lateralis Erichson, 1847, by subsequent designation (Baly 1863: 156).

Family-group name: HABROPHORINI Bechyné & Špringlová de Bechyné, 1969. Stem: Habrophor-.

Hadesia J. Müller, 1911: 175. Current status: valid genus in LEIODIDAE: CHOLEVINAE: LEPTODIRINI: ANTHROHERPONINA.

Type species: Hadesia vasiceki J. Müller, 1911, by monotypy.

Family-group name: HADESIINI Absolon, 1913. Stem: Hadesi-.

Hadrognathus Schaum, 1852: 31. Current status: valid genus in STAPHYLINIDAE: OMALIINAE: HADROGNATHINI. Note: new replacement name for Eugnathus Mulsant & Rey, 1851.

Type species: Eugnathus longipalpis Mulsant & Rey, 1851, by monotypy (type fixation for Eugnathus Mulsant & Rey).

Family-group name: HADROGNATHINI Portevin, 1929. Stem: Hadrognath-.

Hadromerus Schönherr, 1834: 127. Current status: invalid senior synonym of Hadromeropsis Pierce, 1913 in CURCULIONIDAE: ENTIMINAE: POLYDRUSITAE: TANYMECINI: TANYMECINA. Note: junior homonym of Hadromerus Schönherr, 1823 [Coleoptera: CURCULIONIDAE].

Type species: Hadromerus nobilitatus Gyllenhal, 1834, by monotypy.

Family-group name: HADROMERIDES Jekel, 1865. Stem: Hadromer-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

Haemonia Dejean, 1821: 114. Current status: junior synonym of Macroplea Samouelle, 1819 in CHRYSOMELIDAE: DONACIINAE: HAEMONIINI.

Type species: Donacia equiseti Fabricius, 1798 (= Donacia appendiculata Panzer, 1794), by monotypy. Note: the type species often reported for Haemonia Dejean is Donacia zosterae Fabricius, 1801 (= Rhagium muticum Fabricius, 1792) (e.g., Silfverberg 2010: 357) but zosterae is listed as a junior synonym of equiseti, the sole valid species listed by Dejean (1821: 114) in the genus Haemonia; Article 68.3 (ICZN 1999a) denotes that when an author establishes a new nominal genus-group taxon for a single taxonomic species, that species is the type species by monotypy regardless of any cited synonyms.

Family-group name: HAEMONIINI S.H. Chen, 1941. Stem: Haemoni-.

Haeterius Dejean, 1833b: 128. Current status: valid genus in HISTERIDAE: HAETERIINAE: HAETERIINI.

Type species: Hister quadratus Kugelann, 1794 (= Hister ferrugineus G.-A. Olivier, 1789), by monotypy.

Family-group name: HÉTÉRIENS Marseul, 1857. Stem: Haeteri-.

Haliplus Latreille, 1802b: 77. Current status: valid genus in HALIPLIDAE.

Type species: Dytiscus impressus Fabricius, 1787 (= Dytiscus ruficollis DeGeer, 1774), by subsequent designation (Latreille 1810: 426).

Family-group name: HALIPLIDES Aubé, 1836. Stem: Halipl-.

Hallomenus Panzer, 1793: no 17. Current status: valid genus in TETRATOMIDAE: HALLOMENINAE.

Type species: Hallomenus humeralis Panzer, 1793 (= Chrysomela binotata Quensel, 1790), by subsequent designation (Latreille 1810: 429).

Family-group name: HALLOMENIDAE Gistel, 1848. Stem: Hallomen-.

Halorabyxis Jeannel, 1954a: 338. Current status: junior synonym of Physoplectus Reitter, 1882 in STAPHYLINIDAE: PSELAPHINAE: GONIACERITAE: BRACHYGLUTINI: BRACHYGLUTINA.

Type species: Halorabyxis vinsoni Jeannel, 1954, by original designation.

Family-group name: HALORABYXINA Leleup, 1969. Stem: Halorabyx-.

Halyzia Mulsant, 1846: 148. Current status: valid genus in COCCINELLIDAE: COCCINELLINAE: COCCINELLINI.

Type species: Coccinella sedecimguttata Linnaeus, 1758 [as sexdecimguttata], by monotypy.

Family-group name: HALYZIAIRES Mulsant, 1846. Stem: Halyzi-.

Hameedia M. Abdullah & Qadri, 1968: 311. Current status: valid genus in STAPHYLINOIDEA incertae sedis.

Type species: Hameedia batoolae M. Abdullah & Qadri, 1968, by original designation.

Family-group name: HAMEEDINI M. Abdullah & Qadri, 1968. Stem: Hameedi-.

Hamotus Aubé, 1844: 91. Current status: valid genus in STAPHYLINIDAE: PSELAPHINAE: PSELAPHITAE: TYRINI: SOMATIPIONINA.

Type species: Hamotus latericius Aubé, 1844, by subsequent designation (R. Lucas 1920: 314).

Family-group name: HAMOTINI Park, 1951. Stem: Hamot-.

Hapalips Reitter, 1877: 122. Current status: valid genus in EROTYLIDAE: LANGURIINAE: HAPALIPINI.

Type species: Hapalips mexicanus Reitter, 1877, by subsequent designation (Sen Gupta 1968: 6).

Family-group name: HAPALIPINI Leschen, 2003. Stem: Hapalip-.

Hapatesus Candèze, 1863: 76, 188. Current status: valid genus in ELATERIDAE: HAPATESINAE.

Type species: Hapatesus hirtus Candèze, 1863, by monotypy.

Family-group name: HAPATESINAE Kusy, Motyka & Bocák, 2021: 8. Stem: Hapates-.

†Haplochelus Kirejtshuk & Poinar, 2006: 157. Current status: junior synonym of Lepicerus Motschulsky, 1855 in LEPICERIDAE. Note: mid-Cretaceous (Myanmar).

Type species: †Haplochelus georissoides Kirejtshuk & Poinar, 2006, by original designation.

Family-group name: †HAPLOCHELIDAE Kirejtshuk & Poinar, 2006. Stem: Haplochel-.

Haplonyx Schönherr, 1836: 606. Current status: valid genus in CURCULIONIDAE: CURCULIONINAE: CRYPTOPLINI.

Type species: Haplonyx spencei Gyllenhal, 1836, by original designation.

Family-group name: HAPLONYCIDES Lacordaire, 1865. Stem: Haplonych-.

Haplostethus J.L. LeConte, 1860c: 253. Current status: junior synonym of Mastogenius Solier, 1849 in BUPRESTIDAE: POLYCESTINAE: HAPLOSTETHINI.

Type species: Haplostethus subcyaneus J.L. LeConte, 1860, by monotypy.

Family-group name: HAPLOSTETHINI J.L. LeConte, 1861. Stem: Haplosteth-.

Haplotrinchus Kerremans, 1903b: 126. Current status: valid genus in BUPRESTIDAE: CHRYSOCHROINAE: DICERCINI: HAPLOTRINCHINA.

Type species: Buprestis viridula G.-A. Olivier, 1790, by subsequent designation (Bellamy 1997b: 76).

Family-group name: HAPLOTRINCHINA Holyński, 1993. Stem: Haplotrinch-.

Hapsodrilus C. Costa, 1975: 90. Current status: valid genus in ELATERIDAE: AGRYPNINAE: PYROPHORINI: HAPSODRILINA.

Type species: Pyrophorus ignifer Germar, 1841, by original designation.

Family-group name: HAPSODRILINA C. Costa, 1975. Stem: Hapsodril-.

†Hapsomela Poinar & A.E. Brown, 2004: 790. Current status: valid genus in STAPHYLINIDAE: SCYDMAENINAE: †HAPSOMELITAE. Note: Cretaceous (Burmese amber).

Type species: †Hapsomela burmitis Poinar & A.E. Brown, 2004, by original designation.

Family-group name: †HAPSOMELINAE Poinar & A.E. Brown, 2004. Stem: Hapsomel-.

Haraia García & Jiménez-Ramos, 2020: 461. Current status: valid genus in HETEROCERIDAE: HETEROCERINAE: HARAIAINI.

Type species: Haraia cerromachensis García & Jiménez-Ramos, 2020, by original designation.

Family-group name: Tribe HARAIAINI García & Jiménez-Ramos, 2020: 460. Stem: Haraia- (ICZN 1999a, Article 29.4).

Harpaglossus Motschulsky, 1858b: 173. Current status: valid genus in CARABIDAE: HARPALINAE: CHLAENIINI: CHLAENIINA. Note: new replacement name for Ceroglossus Chaudoir, 1856.

Type species: Feronia laevigata Dejean, 1828, by subsequent designation (Andrewes 1938: 135).

Family-group name: HARPAGLOSSINI Basilewsky, 1950. Stem: Harpagloss-.

Harpalus Latreille, 1802b: 92. Current status: valid genus in CARABIDAE: HARPALINAE: HARPALINI: HARPALINA.

Type species [suggested]: Carabus proteus Paykull, 1790 (= Carabus affinis Schrank, 1781), by subsequent designation (Andrewes 1935: 19). Note: the valid type species fixation is that of Latreille (1810: 426), who selected Carabus ruficornis Fabricius, 1775 (= Carabus rufipes DeGeer, 1774), but that species is currently included in the subgenus Pseudoophonus Motschulsky, 1844, not the nominotypical subgenus of Harpalus Latreille; an application to the Commission is necessary to fix Carabus proteus Paykull, 1790 as the type species of Harpalus Latreille, 1802; a first request was postponed (see ICZN 1950); Andrewes (1935: 19) designated Carabus affinis Schrank, 1781 as type species of Harpalus, a species not originally included, but since he listed it at the same time as a synonym of Carabus proteus Paykull, 1790, a species originally included, he is deemed to have designated the latter species as type species (ICZN, 1999a, Article 69.2.2).

Family-group name: HARPALII Bonelli, 1810. Stem: Harpal-.

Haruspex J. Thomson, 1864: 221. Current status: valid genus in CERAMBYCIDAE: CERAMBYCINAE: PIEZOCERINI: HARUSPICINA.

Type species: Ozodes brevipes A. White, 1855, by original designation.

Family-group name: HARUSPICINA Martins, 1976. Stem: Haruspic-.

Harvengia Ferrer, 2004: 367. Current status: valid genus in TENEBRIONIDAE: PIMELIINAE: STENOSINI: HARVENGIINA.

Type species: Harvengia vietnamita Ferrer, 2004, by original designation.

Family-group name: HARVENGINA Ferrer, 2004. Stem: Harvengi-.

Hebesecis Pascoe, 1865a: 353. Current status: junior synonym of Hebecerus Dejean, 1835 in CERAMBYCIDAE: LAMIINAE: ANCITINI. Note: unnecessary new replacement name for Hebecerus Dejean, 1835.

Type species: Acanthocinus marginicollis Boisduval, 1835, by subsequent designation (Bousquet and Bouchard 2013a: 83) (type fixation for Hebecerus Dejean).

Family-group name: HEBESECINAE Pascoe, 1871. Stem: Hebesecid-.

Hebestola Blanchard, 1851: 513. Current status: invalid senior synonym of Neohebestola Marinoni, 1977 in CERAMBYCIDAE: LAMIINAE: FORSTERIINI. Note: junior homonym of Hebestola Haldeman, 1847 [Coleoptera: CERAMBYCIDAE].

Type species: Hebestola parvula Blanchard, 1851, by subsequent designation (J. Thomson 1864: 108).

Family-group name: HEBESTOLITAE J. Thomson, 1864. Stem: Hebestol-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

Hecyrida J. Thomson, 1860a: 39. Current status: junior synonym of Hecyra J. Thomson, 1857 in CERAMBYCIDAE: LAMIINAE: CEROPLESINI: CROSSOTINA. Note: unnecessary new replacement name for Hecyra J. Thomson, 1857.

Type species: Hecyra improba J. Thomson, 1857, by monotypy (type fixation for Hecyra J. Thomson).

Family-group name: HÉCYRIDIDES Lacordaire, 1872. Stem: Hecyrid-.

Hedobia Dejean, 1821: 41. Current status: valid genus in PTINIDAE: EUCRADINAE: HEDOBIINI.

Type species: Ptinus pubescens G.-A. Olivier, 1790, by monotypy.

Family-group name: HÉDOBIAIRES Mulsant & Rey, 1868. Stem: Hedobi-.

Hedyphanes Fischer, 1820: pl. 15. Current status: valid genus in TENEBRIONIDAE: TENEBRIONINAE: HELOPINI: HELOPINA.

Type species: Hedyphanes coerulescens Fischer, 1820, by monotypy.

Family-group name: HEDYPHANINA Reitter, 1922. Stem: Hedyphan-.

Hegemona Laporte, 1840b: 230. Current status: valid genus in TENEBRIONIDAE: STENOCHIINAE: CNODALONINI.

Type species: Hegemona resplendens Laporte, 1840, by monotypy.

Family-group name: HEGEMONINI Reitter, 1922. Stem: Hegemon-.

Heilipus Germar, 1823: 399. Current status: valid genus in CURCULIONIDAE: MOLYTINAE: HYLOBIINI: HYLOBIINA.

Type species: Heilipus lactarius Germar, 1823, by subsequent designation (Schönherr 1825: 582).

Family-group name: HEILIPINAE Faust, 1892. Stem: Heilipod-.

Helea Latreille, 1804b: 326. Current status: valid genus in TENEBRIONIDAE: TENEBRIONINAE: HELEINI: HELEINA. Note: the older name Helea Meigen, 1800 [Diptera] was published in a work that has been suppressed for the purposes of zoological nomenclature (see ICZN 1963b); the name Helea Osten Sacken, 1882 is available in Diptera (see Evenhuis and Pape 2017: 37).

Type species: Helea perforatus Latreille, 1817, by subsequent monotypy (Latreille 1817b: 261).

Family-group name: HELEADAE Fleming, 1821. Stem: Hele-.

Helenaea Schatzmayr & Koch, 1934: 23, 24. Current status: valid genus in CARABIDAE: GEHRINGIINAE: GEHRINGIINI: HELENAEINA.

Type species: Helenaea torretassoi Schatzmayr & C. Koch, 1934, by monotypy.

Family-group name: HELENAEINA Deuve, 2007. Stem: Helenae-.

Helictopleurus H. d’Orbigny, 1915: 402. Current status: valid genus in SCARABAEIDAE: SCARABAEINAE: ONITICELLINI: HELICTOPLEURINA.

Type species: Scarabaeus marsyas G.-A. Olivier, 1789, by subsequent designation (Paulian 1937: 132).

Family-group name: HELICTOPLEURIDES A. Janssens, 1946. Stem: Helictopleur-.

Heligmus Candèze, 1865: 52. Current status: invalid senior synonym of Cleidecosta P.J. Johnson, 2002 in ELATERIDAE: AGRYPNINAE: EUPLINTHINI: CLEIDECOSTINA. Note: junior homonym of Heligmus Dujardin, 1844 [Nematoda].

Type species: Heligmus glyphoderus Candèze, 1865, by monotypy.

Family-group name: HELIGMINI C. Costa, 1975. Stem: Heligm-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

Heliolus Fauvel, 1907: 70. Current status: valid genus in CERAMBYCIDAE: LAMIINAE: HELIOLINI. Note: new replacement name for Helius Fauvel, 1906.

Type species: Helius brevicornis Fauvel, 1906, by monotypy (type fixation for Helius Fauvel).

Family-group name: HELIOLINI Breuning, 1951. Stem: Heliol-.

Heliomene Gistel, 1848b: xi, 134. Current status: junior synonym of Chrysolopus Germar, 1817 in CURCULIONIDAE: CYCLOMINAE: ATERPINI: ATERPINA. Note: unnecessary new replacement name for Chrysolopus Germar, 1817.

Type species: Curculio spectabilis Fabricius, 1775, by subsequent designation (Schönherr 1823: 1142) (type fixation for Chrysolopus Germar).

Family-group name: HELIOMENEIDAE Gistel, 1848. Stem: Heliomen-. Note: nomen oblitum (see Bouchard et al. 2011: 601).

Heliopates Dejean, 1834: 191. Current status: valid genus in TENEBRIONIDAE: BLAPTINAE: DENDARINI: DENDARINA.

Type species [suggested]: Tenebrio lusitanicus Herbst, 1797, by subsequent designation (Gebien 1938b: 304). Note: as noted by Bousquet and Bouchard (2013a: 46), Heliopates is a new replacement name for Heliophilus Dejean, 1821 and as such should have the same type species, Pedinus hybridus Latreille, 1804 (= Tenebrio abbreviatus G.-A. Olivier, 1795) by subsequent designation of Duponchel (1845: 517), which is currently included in the genus Phylan Dejean, 1821; however, currently Heliopates Dejean is recognized as a valid genus and subgenus (Iwan and Löbl 2020: 302, Bouchard et al. 2021: 202); an application to the Commission is needed to resolve this nomenclatural issue.

Family-group name: HÉLIOPATHAIRES Mulsant & Rey, 1854. Stem: Heliopat-.

Helluo Bonelli, 1813: 450, 453. Current status: valid genus in CARABIDAE: HARPALINAE: HELLUONINI: HELLUONINA.

Type species: Helluo costatus Bonelli, 1813, by original designation.

Family-group name: HELLUONIDAE Hope, 1838. Stem: Helluon-.

Helluodes Westwood, 1846: 354. Current status: valid genus in CARABIDAE: HARPALINAE: PHYSOCROTAPHINI.

Type species: Helluodes taprobanae Westwood, 1846, by monotypy.

Family-group name: HELLUODINI Csiki, 1932. Stem: Helluod-.

Helluomorpha Laporte, 1834b: 52. Current status: valid genus in CARABIDAE: HARPALINAE: HELLUONINI: OMPHRINA.

Type species: Helluo heros Gory, 1833, by subsequent designation (Hope 1838a: 110).

Family-group name: HELLUOMORPHINA Reichardt, 1974. Stem: Helluomorph-.

Helochares Mulsant, 1844: [197]. Current status: valid genus in HYDROPHILIDAE: ACIDOCERINAE. Note: name placed on the Official List of Generic Names in Zoology (ICZN 1964).

Type species: Dytiscus lividus Forster, 1771, by subsequent designation (C.G. Thomson 1859: 18; see ICZN 1964). Note: name placed on the Official List of Specific Names in Zoology (ICZN 1964).

Family-group name: HELOCHARAE Orchymont, 1919. Stem: Helochar-.

Helopeltis G.H. Horn, 1873a: 118, 137. Current status: invalid senior synonym of Helobata Bergroth, 1888 in HYDROPHILIDAE: ACIDOCERINAE. Note: junior homonym of Helopeltis Signoret, 1858 [Hemiptera].

Type species: Helopeltis larvalis G.H. Horn, 1873, by monotypy.

Family-group name: HELOPELTINI G.H. Horn, 1873: 118. Stem: Helopelt-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

Helophorus Fabricius, 1775: 66. Current status: valid genus in HELOPHORIDAE. Note: the original spelling Elophorus was emended to Helophorus by the Commission and Helophorus Fabricius, 1775 was placed on the Official List of Generic Names in Zoology (ICZN 1993c).

Type species: Silpha aquatica Linnaeus, 1758, by subsequent designation (Latreille 1810: 428; see ICZN 1993c). Note: name placed on the Official List of Specific Names in Zoology (ICZN 1993c).

Family-group name: HELOPHERIDA Leach, 1815. Stem: Helophor-.

Helopinus Solier, 1848b: 152, 197. Current status: valid genus in TENEBRIONIDAE: BLAPTINAE: PEDININI: HELOPININA.

Type species: Helopinus costatus Solier, 1848, by monotypy.

Family-group name: HÉLOPINIDES Lacordaire, 1859. Stem: Helopin-.

Helops Fabricius, 1775: 257. Current status: valid genus in TENEBRIONIDAE: TENEBRIONINAE: HELOPINI: HELOPINA. Note: name placed on the Official List of Generic Names in Zoology ICZN 2009c).

Type species: Tenebrio caeruleus Linnaeus, 1758, by plenary power (see ICZN 2009c). Note: as mentioned in Bousquet et al. (2018: 183) and Vela et al. (2020: 63) Tenebrio caeruleus Linnaeus, 1758, the type species recognized by the Commission for the genus Helops (ICZN 2009c), is in fact a leaf beetle species which belongs to the genus Timarcha Samouelle, 1819 [Coleoptera: CHRYSOMELIDAE]; obviously, Fabricius (1775: 257) misidentified Linnaeus’s Tenebrio caeruleus; a request to the Commission is necessary to change the type species of Helops to Tenebrio caeruleus Linnaeus, 1758 (sensu Fabricius, 1775) (= Helops chalibaeus Rossi, 1790).

Family-group name: HELOPII Latreille, 1802. Stem: Helop-.

Helota W.S. MacLeay, 1825: 42. Current status: valid genus in HELOTIDAE.

Type species: Helota vigorsii W.S. MacLeay, 1825, by monotypy.

Family-group name: HÉLOTIDES Chapuis, 1876. Stem: Helot-. Note: junior homonym of HELOTIDAE Adams et al., 1854 (type genus Helotes Cuvier, 1829) in Pisces.

Hemiconderis Kleine, 1926: 162. Current status: junior synonym of Falsolucidota Pic, 1921 in LYCIDAE: METRIORRHYNCHINAE: METRIORRHYNCHINI: METRIORRHYNCHINA.

Type species: Hemiconderis explicatus Kleine, 1926, by monotypy.

Family-group name: HEMICONDERININA Bocák & Bocáková, 1990. Stem: Hemiconderin-.

Hemicrepidius Germar, 1839: 206, 212. Current status: valid genus in ELATERIDAE: DENDROMETRINAE: DENDROMETRINI: HEMICREPIDIINA.

Type species: Hemicrepidius thomasi Germar, 1839 (= Elater memnonius Herbst, 1806), by monotypy. Note: the original spelling of the valid name of the type species was nemnonius; however, the incorrect subsequent spelling memnonius is in prevailing usage, attributed to the publication of the original spelling, and therefore deemed to be the correct original spelling (ICZN 1999a, Article 33.3.1).

Family-group name: HEMICREPIDIINI Champion, 1896. Stem: Hemicrepidi-.

Hemilophus Audinet-Serville, 1835a: 49. Current status: valid genus in CERAMBYCIDAE: LAMIINAE: HEMILOPHINI.

Type species: Hemilophus dimidiaticornis Audinet-Serville, 1835, by monotypy.

Family-group name: HEMILOPHITAE J. Thomson, 1868. Stem: Hemiloph-. Note: nomen protectum (see Bousquet et al. 2009: 31).

Hemiops Eschscholtz in Laporte, 1838: table. Current status: valid genus in ELATERIDAE: HEMIOPINAE: HEMIOPINI.

Type species: Hemiops flava Laporte, 1838, by subsequent designation (Hyslop 1921: 648).

Family-group name: HEMIOPINAE Fleutiaux, 1941. Stem: Hemiop-.

Hemipeplus Latreille, 1829b: 53. Current status: valid genus in MYCTERIDAE: HEMIPEPLINAE. Note: this name is sometimes credited to Berthold (1827: 394) (e.g., Löbl et al. 2020: 563) but the genus is not made available by a description or an indication in Berthold’s work.

Type species: Hemipeplus hemipterus Lacordaire, 1854, by subsequent monotypy (Lacordaire 1854: 454).

Family-group name: HÉMIPÉPLIDES Lacordaire, 1854. Stem: Hemipepl-.

Hemipharis Burmeister, 1842: 531. Current status: valid genus in SCARABAEIDAE: CETONIINAE: SCHIZORHININI: SCHIZORHININA.

Type species: Schyzorhina insularis Gory & Percheron, 1833, by subsequent designation (Kraatz 1880: 183).

Family-group name: HEMIPHARIDAE Kraatz, 1880. Stem: Hemiphar-.

Hemirhipidius Heller, 1920: 168. Current status: valid genus in RIPIPHORIDAE: HEMIRHIPIDIINAE.

Type species: Hemirhipidius nigroapicalis Heller, 1920, by original designation.

Family-group name: HEMIRHIPIDIINI Heller, 1920. Stem: Hemirhipidi-.

Hemirhipus Berthold, 1827: 336. Current status: valid genus in ELATERIDAE: AGRYPNINAE: HEMIRHIPINI: HEMIRHIPINA.

Type species: Elater lineatus G.-A. Olivier, 1790, by monotypy.

Family-group name: HÉMIRHIPIDES Candèze, 1857. Stem: Hemirhip-.

Hemisphaerota Chevrolat in Dejean, 1836a: 367. Current status: valid genus in CHRYSOMELIDAE: CASSIDINAE: HEMISPHAEROTINI.

Type species: Cassida erythrocera Germar, 1823, by monotypy. Note: this species is generally listed as a junior synonym of Imatidium cyaneum Say, 1824, but Germar’s name is older and should be used as the valid name for the species; an application to the Commission is necessary to conserve usage of Hemisphaerota cyanea (Say, 1824) as valid.

Family-group name: HEMISPHAEROTINI Monrós & Viana, 1951. Stem: Hemisphaerot-.

Hemydacne Jacoby, 1897: 244. Current status: valid genus in CHRYSOMELIDAE: EUMOLPINAE: HEMYDACNINI.

Type species: Hemydacne maculicollis Jacoby, 1897, by monotypy.

Family-group name: HEMYDACNINI Bechyné, 1951. Stem: Hemydacn-.

Henicolabus Voss, 1925: 224. Current status: valid genus in ATTELABIDAE: ATTELABINAE: ATTELABINI: HENICOLABINA.

Type species: Attelabus giganteus Faust, 1882, by original designation.

Family-group name: HENICOLABINA Legalov, 2007. Stem: Henicolab-.

Hephebocerus Schönherr, 1840a: 501. Current status: valid genus in BRENTIDAE: BRENTINAE: TRACHELIZINI: HEPHEBOCERINA.

Type species: Brenthus nanus Boheman, 1833, by original designation.

Family-group name: HÉPHÉBOCÉRIDES Lacordaire, 1865. Stem: Hephebocer-.

Heptaphylla Friedenreich, 1883: 375. Current status: junior synonym of Rhipidandrus J.L. LeConte, 1862 in TENEBRIONIDAE: TENEBRIONINAE: BOLITOPHAGINI.

Type species: Heptaphylla fungicola Friedenreich, 1883, by monotypy.

Family-group name: HEPTAPHYLLINI Prudhomme de Borre, 1886. Stem: Heptaphyll-.

Heptophylla Motschulsky, 1857a: 32. Current status: valid genus in SCARABAEIDAE: MELOLONTHINAE: HEPTOPHYLLINI.

Type species: Heptophylla picea Motschulsky, 1857, by monotypy.

Family-group name: HEPTOPHYLLINI S.I. Medvedev, 1951. Stem: Heptophyll-.

Hermaeophaga Foudras, 1860: 35, 299. Current status: valid genus in CHRYSOMELIDAE: GALERUCINAE: ALTICINI.

Type species: Haltica cicatrix Illiger, 1807, by subsequent designation (Maulik 1926: 411).

Family-group name: HERMAEOPHAGINA Bechyné, 1997. Stem: Hermaeophag-.

Hesperophanes Dejean, 1835: 328. Current status: valid genus in CERAMBYCIDAE: CERAMBYCINAE: HESPEROPHANINI: HESPEROPHANINA.

Type species [suggested]: Callidium sericeum Fabricius, 1787, by subsequent designation (J. Thomson 1864: 253). Note: this species is a species inquirenda in Dejean (1835: 328) and therefore is not an originally included species (ICZN 1999a, Article 67.2.5) but all originally included species belong to other cerambycid genera (see Bousquet and Bouchard 2013a: 83); we believe that a request should be submitted to the Commission to fix Callidium sericeum Fabricius, 1787 as the type species of Hesperophanes Dejean, 1835.

Family-group name: HESPÉROPHANAIRES Mulsant, 1839. Stem: Hesperophan-.

Hesthesis Newman, 1840a: 17. Current status: valid genus in CERAMBYCIDAE: CERAMBYCINAE: HESTHESINI.

Type species: Leptura variegata Fabricius, 1775, by subsequent designation (J. Thomson 1864: 162).

Family-group name: HESTHESINAE Pascoe, 1867. Stem: Hesthes-.

Hetairotermes Cameron, 1920: 223. Current status: valid genus in STAPHYLINIDAE: ALEOCHARINAE: TERMITOHOSPITINI: HETAIROTERMITINA. Note: new replacement name for Termophila Lea, 1910.

Type species: Termophila latebricola Cameron, 1920, by subsequent designation (Fenyes 1919: 25) (type fixation for Termophila Lea).

Family-group name: HETAIROTERMITINA Seevers, 1957. Stem: Hetairotermit-.

Heteraspis Chevrolat in Dejean, 1836a: 413. Current status: valid genus in CHRYSOMELIDAE: EUMOLPINAE: BROMIINI.

Type species: Eumolpus vittatus G.-A. Olivier, 1808, by monotypy.

Family-group name: HETERASPINAE Baly, 1863. Stem: Heteraspid-.

Heterocerus Fabricius, 1792a: 262. Current status: valid genus in HETEROCERIDAE: HETEROCERINAE: HETEROCERINI.

Type species: Apate marginatus Fabricius, 1787, by subsequent designation (Latreille 1810: 428).

Family-group name: HETEROCERIDAE W.S. MacLeay, 1825. Stem: Heterocer-.

Heterocheira Dejean, 1836b: 220. Current status: valid genus in TENEBRIONIDAE: BLAPTINAE: OPATRINI: HETEROTARSINA.

Type species: Uloma australe Boisduval, 1835, by monotypy.

Family-group name: HETEROCHEIRINI C. Koch, 1956. Stem: Heterocheir-.

Heterochelus Burmeister, 1844: 87. Current status: valid genus in SCARABAEIDAE: MELOLONTHINAE: HOPLIINI: HOPLIINA.

Type species: to be determined. Note: Burmeister (1844) placed the 54 species of the genus Heterochelus in several informal groups and for one of the groups he designated Melolontha dentipes Fabricius, 1781 as the type on page 108; however, we do not consider that this represents a valid type species fixation for Heterochelus (also, Melolontha dentipes Fabricius, 1781 is the type species of Dichelus Lepeletier & Audinet-Serville, 1828); as far as we know, no type species has been proposed for Heterochelus and we refrain from suggesting one since the taxonomy of the African Hopliini is in need of a thorough revision (H. Dombrow, pers. comm. to P.B., 2021).

Family-group name: HETEROCHELIDAE Burmeister, 1844. Stem: Heterochel-.

Heterogenius Moser, 1911: 143. Current status: valid genus in SCARABAEIDAE: CETONIINAE: CREMASTOCHEILINI: HETEROGENIINA.

Type species: Heterogenius albomaculatus Moser, 1911, by monotypy.

Family-group name: HETEROGENIINA Krikken, 1984. Stem: Heterogeni-.

Heterogyrus Legros, 1953: 63. Current status: valid genus in GYRINIDAE: HETEROGYRINAE.

Type species: Heterogyrus milloti Legros, 1953, by original designation.

Family-group name: HETEROGYRINI Brinck, 1956. Stem: Heterogyr-.

Heteromastix Boheman, 1858: 81. Current status: valid genus in CANTHARIDAE: HETEROMASTIGINAE.

Type species: Heteromastix bicolor Boheman, 1858, by monotypy.

Family-group name: HETEROMASTIGINAE Motyka, Biffi & Bocák in Motyka et al., 2023b: 559. Stem: Heteromastig-.

Heteronyx Guérin-Méneville, 1831a: pl. 3. Current status: valid genus in SCARABAEIDAE: SERICOIDINAE: HETERONYCHINI.

Type species: Heteronyx australis Guérin-Méneville, 1831, by monotypy.

Family-group name: HÉTÉRONYCIDES Lacordaire, 1855. Stem: Heteronych-. Note: senior homonym of HETERONYCHIDAE André, 1891 (type genus Heteronyx Saussure, 1887) in Hymenoptera; André’s name is permanently invalid (based on a preoccupied type genus).

Heteropalpus Buquet, 1843: pl. 118. Current status: valid genus in DISTENIIDAE: HETEROPALPINI.

Type species: Heteropalpus pretiosus Buquet,1843, by monotypy.

Family-group name: HETEROPALPINI Villiers, 1961. Stem: Heteropalp-.

Heteropaussus J. Thomson, 1860c: 68, 70. Current status: valid genus in CARABIDAE: PAUSSINAE: PAUSSINI: HETEROPAUSSINA.

Type species: Cerapterus alternans Westwood, 1849, by monotypy.

Family-group name: HETEROPAUSSINES E. Janssens, 1950. Stem: Heteropauss-.

Heterophana Burmeister, 1842: 602. Current status: valid genus in SCARABAEIDAE: CETONIINAE: STENOTARSIINI: HETEROPHANINA.

Type species: Cetonia canaliculata Gory & Percheron, 1833, by subsequent designation (Sakai and Nagai 1998: 182).

Family-group name: HETEROPHANAE Schoch, 1894. Stem: Heterophan-.

Heterops Blanchard, 1842a: 51. Current status: valid genus in CERAMBYCIDAE: CERAMBYCINAE: EBURIINI.

Type species: Purpuricenus loreyi Duponchel, 1837, by subsequent designation (J. Thomson 1864: 203).

Family-group name: HÉTÉROPSIDES Lacordaire, 1868. Stem: Heterops-. Note: nomen protectum (see Bousquet et al. 2009: 48).

Heterorhina Westwood, 1842b: 132. Current status: valid genus in SCARABAEIDAE: CETONIINAE: GOLIATHINI: RHOMBORHININA. Note: two original spellings were used, including Hetororhina (p. 134); Neave (1940: 644) acted as First Reviser (ICZN 1999a, Article 24.2.3) and selected the spelling Heterorhina, the spelling currently in prevailing usage in publications dealing with CETONIINAE or SCARABAEIDAE classification and nomenclature.

Type species: Cetonia nigritarsis Hope, 1831, by subsequent designation (Duponchel 1845: 603).

Family-group name: HETERORRHINIDAE Kraatz, 1880. Stem: Heterorhin-.

Heteroscapha Achard, 1914a: 394. Current status: junior synonym of Bironium Csiki, 1909 in STAPHYLINIDAE: SCAPHIDIINAE: SCAPHISOMATINI.

Type species: Heteroscapha feai Achard, 1914, by original designation.

Family-group name: HETEROSCAPHINI Achard, 1914. Stem: Heteroscaph-.

Heterosoma Schaum, 1845: 390, 426. Current status: valid genus in SCARABAEIDAE: CETONIINAE: STENOTARSIINI: HETEROSOMATINA.

Type species: Cetonia collata Gory & Percheron, 1835, by monotypy.

Family-group name: HETEROSOMATINA Krikken, 1984. Stem: Heterosomat-.

Heterosternus Dupont, 1832: pl. 10. Current status: valid genus in SCARABAEIDAE: RUTELINAE: RUTELINI: HETEROSTERNINA.

Type species: Heterosternus buprestoides Dupont, 1832, by monotypy.

Family-group name: HETEROSTERNINAE H.W. Bates, 1888. Stem: Heterostern-. Note: nomen protectum (see A.B.T. Smith 2006: 171).

Heterota Mulsant & Rey, 1873: 162. Current status: valid genus in STAPHYLINIDAE: ALEOCHARINAE: HOMALOTINI: BOLITOCHARINA.

Type species: Homalota plumbea G.R. Waterhouse, 1858, by monotypy.

Family-group name: HETEROTAE Fenyes, 1919. Stem: Heterot-.

Heterotarsus Latreille, 1829b: 26. Current status: valid genus in TENEBRIONIDAE: BLAPTINAE: OPATRINI: HETEROTARSINA.

Type species: Heterotarsus tenebrioides Guérin-Méneville, 1831, by subsequent monotypy (Guérin-Méneville 1831b: pl. 30).

Family-group name: HÉTÉROTARSITES Blanchard, 1845. Stem: Heterotars-.

Heterotaxus Bernhauer, 1915a: 313. Current status: valid genus in STAPHYLINIDAE: ALEOCHARINAE: OXYPODININI.

Type species: Heterotaxus bihastatus Bernhauer, 1915, by monotypy.

Family-group name: HETEROTAXINI Fenyes, 1921. Stem: Heterotax-.

Heterothops Stephens, 1829b: 23. Current status: valid genus in STAPHYLINIDAE: STAPHYLININAE: STAPHYLININI: AMBLYOPININA.

Type species: Staphylinus binotatus Gravenhorst, 1802, by monotypy. Note: name placed on the Official List of Specific Names in Zoology (ICZN 2003b).

Family-group name: HETEROTHOPSI Coiffait, 1978. Stem: Heterothop-.

Hexacolus Eichhoff, 1868b: 399. Current status: junior synonym of Scolytodes Ferrari, 1867 in CURCULIONIDAE: SCOLYTINAE: HEXACOLINI.

Type species: Hexacolus glaber Eichhoff, 1868, by monotypy.

Family-group name: HEXACOLIDAE Eichhoff, 1878. Stem: Hexacol-.

Hexagonia W. Kirby, 1825: 563. Current status: valid genus in CARABIDAE: HARPALINAE: HEXAGONIINI.

Type species: Hexagonia terminata W. Kirby, 1825, by monotypy.

Family-group name: HEXAGONIAE G.H. Horn, 1881. Stem: Hexagoni-.

Hexaplatarthrus Jeannel, 1955: 181. Current status: valid genus in CARABIDAE: PAUSSINAE: PAUSSINI: PENTAPLATARTHRINA.

Type species: Hexaplatarthrus vadoni Jeannel, 1955, by original designation.

Family-group name: HEXAPLATARTHRINA Luna de Carvalho, 1961. Stem: Hexaplatarthr-.

Hexatricha A. White, 1846: 21. Current status: valid genus in CERAMBYCIDAE: LAMIINAE: PARMENINI.

Type species: Lamia pulverulenta Westwood, 1843, by monotypy.

Family-group name: HEXATHRICITAE J. Thomson, 1864. Stem: Hexatrich-.

Hexodon G.-A. Olivier, 1789b: [07] 3. Current status: valid genus in SCARABAEIDAE: DYNASTINAE: HEXODONTINI.

Type species: Hexodon reticulatum G.-A. Olivier, 1789, by subsequent designation (Latreille 1810: 428).

Family-group name: HEXODONTIDES Lacordaire, 1855. Stem: Hexodont-.

Hexoplon J. Thomson, 1864: 219. Current status: valid genus in CERAMBYCIDAE: CERAMBYCINAE: HEXOPLONINI.

Type species: Hexoplon venus J. Thomson, 1864, by original designation.

Family-group name: HEXOPLONINI Martins, 2006. Stem: Hexoplon- (ICZN 1999a, Article 29.4).

Hiletus Schiødte, 1847: 346. Current status: valid genus in CARABIDAE: HILETINAE.

Type species: Hiletus versutus Schiødte, 1847, by monotypy.

Family-group name: HILETINI Schiødte, 1847. Stem: Hilet-.

Himalusa Pace, 2006: 357. Current status: valid genus in STAPHYLINIDAE: ALEOCHARINAE: HIMALUSINI.

Type species: Himalusa annapurnensis Pace, 2006, by original designation.

Family-group name: HIMALUSINI Klimaszewski, Pace & Center, 2010. Stem: Himalus-.

Himasthlophallus Egorov & Zherikhin in Zherikhin and Egorov, 1991: 32. Current status: valid genus in CURCULIONIDAE: BRACHYCERINAE: HIMASTHLOPHALLINI.

Type species: Himasthlophallus flagellifer Egorov & Zherikhin, 1991, by original designation.

Family-group name: HIMASTHLOPHALLINI Zherikhin, 1991. Stem: Himasthlophall-.

Himatinum Cockerell, 1906: 243. Current status: junior synonym of Himatium Wollaston, 1873 in CURCULIONIDAE: COSSONINAE: HIMATININI. Note: unnecessary new replacement name for Himatium Wollaston, 1873.

Type species: Himatium pubescens Wollaston, 1873, by monotypy (type fixation for Himatium Wollaston).

Family-group name: HIMATININI Konishi, 1962. Stem: Himatin-.

Himatismus Erichson, 1843b: 253. Current status: valid genus in TENEBRIONIDAE: PIMELIINAE: TENTYRIINI.

Type species: Himatismus mandibularis Erichson, 1843, by subsequent designation (Gebien 1937: 574).

Family-group name: HIMATISMINA Skopin, 1979. Stem: Himatism-.

Himatolabus Jekel, 1860: 189. Current status: valid genus in ATTELABIDAE: ATTELABINAE: ATTELABINI: HIMATOLABINA.

Type species: Attelabus vestitus Gyllenhal, 1839, by original designation.

Family-group name: HIMATOLABINA Legalov, 2003. Stem: Himatolab-.

Hippodamia Chevrolat in Dejean, 1836a: 432. Current status: valid genus in COCCINELLIDAE: COCCINELLINAE: COCCINELLINI.

Type species: Coccinella mutabilis Scriba, 1791 (= Coccinella variegata Goeze, 1777), by subsequent designation (Chevrolat 1845: 623). Note: Goeze’s (1777) work is not consistently binominal (see ICZN 2021a) and therefore an application to the Commission is necessary to retain Coccinella variegata Goeze, 1777 as the valid name for the type species.

Family-group name: HIPPODAMIINI Weise, 1885. Stem: Hippodami-.

Hippoloetis Laporte, 1834c: 152. Current status: valid genus in CARABIDAE: HARPALINAE: HARPALINI: STENOLOPHINA. Note: Laporte used two original spellings, including Hipollaetis (p. 153); Laporte (1840a: 95) subsequently used only Hippoloetis and therefore acted as First Reviser (ICZN 1999a, Article 24.2.4).

Type species: Hippoloetis rufa Laporte, 1834, by monotypy.

Family-group name: HIPPOLAETINA Basilewsky, 1951. Stem: Hippoloet-.

Hippomelas Laporte & Gory, 1837: 3. Current status: valid genus in BUPRESTIDAE: CHRYSOCHROINAE: DICERCINI: HIPPOMELANINA.

Type species: Buprestis mexicana Laporte & Gory, 1837, by monotypy.

Family-group name: HIPPOMELANINA Holyński, 1993. Stem: Hippomelan-.

Hippopsicon J. Thomson, 1858b: 195. Current status: valid genus in CERAMBYCIDAE: LAMIINAE: AGAPANTHIINI.

Type species: Hippopsicon lacteolum J. Thomson, 1858, by monotypy.

Family-group name: HIPPOPSICONINI L.S. Dillon & E.S. Dillon, 1945. Stem: Hippopsicon-.

Hippopsis Lepeletier & Audinet-Serville, 1825: 336. Current status: valid genus in CERAMBYCIDAE: LAMIINAE: AGAPANTHIINI.

Type species: Hippopsis lineolatus Lepeletier & Audinet-Serville, 1825, by subsequent designation (J. Thomson 1864: 97). Note: although Thomson (1864: 97) listed Saperda lemniscata Fabricius, 1801, not an originally included nominal species, as type species of Hippopsis, he listed “Hippopsis lineolata [sic] Lepeletier & Audinet-Serville”, one of the two originally included species in Hippopsis, at the same time in synonymy with Saperda lemniscata, he is therefore deemed to have designed Hippopsis lineolata Lepeletier & Audinet-Serville as type species (ICZN 1999a, Article 69.2.2).

Family-group name: HIPPOPSITAE J. Thomson, 1860. Stem: Hippopse-.

Hipporhis Billberg, 1820: 45. Current status: junior synonym of Bronchus Germar, 1817 in CURCULIONIDAE: CYCLOMINAE: HIPPORHININI. Note: unnecessary new replacement name for Bronchus Germar, 1817.

Type species: Curculio capensis Linnaeus, 1764, by subsequent designation (Alonso-Zarazaga and Lyal 1999: 21, 138) (type fixation for Bronchus Germar).

Family-group name: HIPPORHINIDES Lacordaire, 1863. Stem: Hipporhin-.

Hispa Linnaeus, 1767: 603. Current status: valid genus in CHRYSOMELIDAE: CASSIDINAE: HISPINI.

Type species: Hispa atra Linnaeus, 1767, by subsequent designation (Latreille 1810: 432).

Family-group name: HISPOIDEAE Gyllenhal, 1813. Stem: Hisp-.

†Hispanoclavina Soriano, Ponomarenko & Delclòs, 2007: 527. Current status: valid genus in †COPTOCLAVIDAE: †HISPANOCLAVINAE. Note: Lower Cretaceous (Spain).

Type species: †Hispanoclavina diazromerali Soriano, Ponomarenko & Delclòs, 2007, by original designation.

Family-group name: †HISPANOCLAVINAE Soriano, Ponomarenko & Delclòs, 2007. Stem: Hispanoclav- (ICZN 1999a, Article 29.4).

Hispodes G.A.K. Marshall, 1955a: 23. Current status: valid genus in BELIDAE: OXYCORYNINAE: OXYCORYNITAE: HISPODINI.

Type species: Hispodes spicatus G.A.K. Marshall, 1955, by original designation.

Family-group name: HISPODINI Voss, 1957. Stem: Hispod-.

Hispoleptis Baly, 1864d: 262. Current status: valid genus in CHRYSOMELIDAE: CASSIDINAE: HISPOLEPTINI.

Type species: Promecotheca diluta Guérin-Méneville, 1840, by original designation.

Family-group name: HISPOLEPTITES Chapuis, 1875. Stem: Hispolept-.

Hispostoma Weise, 1907: 134. Current status: valid genus in CHRYSOMELIDAE: CHRYSOMELINAE: CHRYSOMELINI.

Type species: Hispostoma marginatum Weise, 1907, by monotypy.

Family-group name: HISPOSTOMINI Daccordi, 1981. Stem: Hispostomat-.

Hister Linnaeus, 1758: 342, 358. Current status: valid genus in HISTERIDAE: HISTERINAE: HISTERINI.

Type species: Hister unicolor Linnaeus, 1758, by subsequent designation (Latreille 1810: 428).

Family-group name: HISTEROIDES Gyllenhal, 1808. Stem: Hister-.

Hobartius Sen Gupta & Crowson, 1966: 74. Current status: valid genus in HOBARTIIDAE.

Type species: Hobartius tasmanicus Sen Gupta & Crowson, 1966 (= Atomaria eucalypti Blackburn, 1892), by original designation.

Family-group name: HOBARTIINI Sen Gupta & Crowson, 1966. Stem: Hobarti-.

Hodoxenus Kistner, 1970b: 12. Current status: valid genus in STAPHYLINIDAE: ALEOCHARINAE: ALEOCHARINI: HODOXENINA.

Type species: Hodoxenus sheasbyi Kistner, 1970, by original designation.

Family-group name: HODOXENINA Kistner, 1970. Stem: Hodoxen-.

Holcorhinus Schönherr, 1826: 194. Current status: valid genus in CURCULIONIDAE: ENTIMINAE: OTIORHYNCHITAE: HOLCORHININI.

Type species: Holcorhinus hispidulus Thon, 1833, by subsequent monotypy (Thon 1833: 13).

Family-group name: HOLCORHINIDAE Desbrochers des Loges, 1898. Stem: Holcorhin-.

†Holcorobeus Nikritin, 1977: 127. Current status: valid genus in SCARABAEIDAE: ACLOPINAE: †HOLCOROBEINI. Note: Upper Jurassic/Lower Cretaceous (China; Kazakhstan; Mongolia; Russia); two original spellings were used, including Holcoribeus (p. 127); Krell (2000: 889) acted as First Reviser (ICZN 1999a, Article 24.2.3) and selected Holcorobeus.

Type species: †Holcorobeus vittatus Nikritin, 1977, by original designation.

Family-group name: †HOLCOROBEINI Nikolajev, 1992. Stem: Holcorobe-.

Holisus Erichson, 1839b: 298. Current status: valid genus in STAPHYLINIDAE: STAPHYLININAE: STAPHYLININI: HYPTIOMINA.

Type species: Holisus analis Erichson, 1839, by subsequent designation (Duponchel 1841b: 57).

Family-group name: HOLISINA Newton, 1988. Stem: Holis-.

Hololepta Paykull, 1811: 101. Current status: valid genus in HISTERIDAE: HISTERINAE: HOLOLEPTINI.

Type species [suggested]: Hister planus Sulzer, 1776, by subsequent designation (Hope 1841a: 105). Note: the valid type species fixation is that of J.E. Gray (1831b: 323), who selected Hister depressus Fabricius, 1787, but that species is currently a junior synonym of Hister compressus Herbst, 1783 in the genus Platysoma Leach, 1817; an application to the Commission is necessary to fix Hister planus Sulzer, 1776 as the type species of Hololepta Paykull, 1811.

Family-group name: HOLOLEPTIDAE Hope, 1841. Stem: Hololept-.

Holoparamecus Curtis, 1833b: 186. Current status: valid genus in ENDOMYCHIDAE: MEROPHYSIINAE.

Type species: Holoparamecus depressus Curtis, 1833, by monotypy.

Family-group name: HOLOPARAMECINI Seidlitz, 1888. Stem: Holoparamec-.

Holopleura J.L. LeConte, 1873: 193. Current status: valid genus in CERAMBYCIDAE: CERAMBYCINAE: HOLOPLEURINI.

Type species: Holopleura marginata J.L. LeConte, 1873, by subsequent designation (Linsley 1962: 181).

Family-group name: HOLOPLEURINI Chemsak & Linsley, 1974. Stem: Holopleur-.

Holopterus Blanchard, 1851: 475. Current status: valid genus stat. nov. in CERAMBYCIDAE: CERAMBYCINAE: HOLOPTERINI.

Type species: Holopterus chilensis Blanchard, 1851, by monotypy.

Family-group name: HOLOPTÉRIDES Lacordaire, 1868. Stem: Holopter-.

Holostrophus G.H. Horn, 1888: 32, 36. Current status: valid genus in TETRATOMIDAE: EUSTROPHINAE: HOLOSTROPHINI.

Type species: Eustrophus bifasciatus Say, 1824, by subsequent designation (Nikitsky 1998: 48).

Family-group name: HOLOSTROPHINI Nikitsky, 1998. Stem: Holostroph-.

Holozodus Fairmaire, 1898: 346. Current status: valid genus in STAPHYLINIDAE: PSELAPHINAE: PSELAPHITAE: ARHYTODINI. Note: new replacement name for Hologlyptus Fairmaire, 1898.

Type species: Hologlyptus raffrayi Fairmaire, 1898, by monotypy (type fixation for Hologlyptus Fairmaire).

Family-group name: HOLOZODINI Raffray, 1900. Stem: Holozod-.

Homalocerus Schönherr, 1839: 358. Current status: valid genus in BELIDAE: BELINAE: BELITAE: BELINI: HOMALOCERINA.

Type species: Rhinotia lyciformis Germar, 1833, by original designation.

Family-group name: HOMALOCERINA Legalov, 2009. Stem: Homalocer-.

Homalota Mannerheim, 1830: 73. Current status: valid genus in STAPHYLINIDAE: ALEOCHARINAE: HOMALOTINI.

Type species: Aleochara plana Gyllenhal, 1810, by monotypy.

Family-group name: HOMALOTIDA Heer, 1839. Stem: Homalot-.

Homebius Endrödy-Younga, 1989: 124. Current status: valid genus in TENEBRIONIDAE: PIMELIINAE: CRYPTOCHILINI: HOMEBIINA.

Type species: Homebius kaszabi Endrödy-Younga, 1989, by original designation.

Family-group name: HOMEBIINA Endrödy-Younga, 1989. Stem: Homebi-.

Homethes Newman, 1842c: 402. Current status: valid genus in CARABIDAE: HARPALINAE: ODACANTHINI: HOMETHINA.

Type species: Homethes elegans Newman, 1842, by monotypy.

Family-group name: HOMETHINA Liebherr, 2016: 223. Stem: Hometh-.

Homoeodera Wollaston, 1870: 23. Current status: valid genus in ANTHRIBIDAE: CHORAGINAE: HOMOEODERINI.

Type species: Homoeodera rotundipennis Wollaston, 1870, by subsequent designation (Basilewsky 1972: 274).

Family-group name: HOMOEODERIDES Wollaston, 1870. Stem: Homoeoder-.

Homoeoplastus Gistel, 1856: 360. Current status: junior synonym Byturus Latreille, 1797 in BYTURIDAE: BYTURINAE.

Type species: Dermestes tomentosus DeGeer, 1774, by subsequent designation (Bouchard and Bousquet 2020a: 93).

Family-group name: HOMOEOPLASTIDAE Gistel, 1856. Stem: Homoeoplast-.

Homonoea Newman, 1842a: 319. Current status: valid genus in CERAMBYCIDAE: LAMIINAE: HOMONOEINI.

Type species: Homonoea patrona Newman, 1842, by subsequent designation (J. Thomson 1864: 35).

Family-group name: HOMONAEITAE J. Thomson, 1864. Stem: Homonoe-.

Homopterus Westwood, 1840a: 75. Current status: valid genus in CARABIDAE: PAUSSINAE: PAUSSINI: HOMOPTERINA.

Type species: Cerapterus brasiliensis Westwood, 1840, by original designation.

Family-group name: HOMOPTERINAE Wasmann, 1920. Stem: Homopter-. Note: junior homonym of HOMOPTERINAE Boisduval, 1852 (type genus Homoptera Boisduval, 1852) in Lepidoptera.

Homorhythmus Bedel, 1883: 38. Current status: junior synonym of Simo Dejean, 1821 in CURCULIONIDAE: ENTIMINAE: CYPHICERITAE: PERITELINI: SIMONINA.

Type species: Curculio hirticornis Herbst, 1795, by monotypy.

Family-group name: HOMORYTHMINI Hoffmann, 1950. Stem: Homorhythm-.

Hoplandria Kraatz, 1857a: 4. Current status: valid genus in STAPHYLINIDAE: ALEOCHARINAE: HOPLANDRIINI: HOPLANDRIINA.

Type species: Hoplandria ochracea Kraatz, 1857 (= Gyrophaena lateralis Melsheimer, 1844), by subsequent designation (Casey 1910: 170).

Family-group name: HOPLANDRIAE Casey, 1910. Stem: Hoplandri-.

Hoplapoderus Jekel, 1860: 171. Current status: valid genus in ATTELABIDAE: APODERINAE: HOPLAPODERINI: HOPLAPODERINA.

Type species: Attelabus gemmatus Thunberg, 1784, by subsequent designation (Voss 1925: 5).

Family-group name: HOPLAPODERINI Voss, 1926. Stem: Hoplapoder-.

Hoplia Illiger, 1803: 226. Current status: valid genus in SCARABAEIDAE: MELOLONTHINAE: HOPLIINI: HOPLIINA.

Type species: Scarabaeus farinosus Linnaeus, 1761 (= Scarabaeus argenteus Poda von Neuhaus, 1761), by subsequent designation (Latreille 1810: 428).

Family-group name: HOPLIDES Latreille, 1829. Stem: Hopli-.

Hoplideres Audinet-Serville, 1832: 147. Current status: valid genus in CERAMBYCIDAE: PRIONINAE: HOPLIDERINI.

Type species: Hoplideres spinipennis Audinet-Serville, 1832, by monotypy.

Family-group name: HOPLIDERITAE J. Thomson, 1864. Stem: Hoplider-.

Hoplionota Hope, 1841a: 153. Current status: junior synonym of Notosacantha Chevrolat, 1836 in CHRYSOMELIDAE: CASSIDINAE: NOTOSACANTHINI.

Type species: Cassida echinata Fabricius, 1801, by original designation.

Family-group name: HOPLIONOTITAE Spaeth, 1929. Stem: Hoplionot-.

Hoplitoxenus Jeannel, 1960: 73, 173. Current status: valid genus in STAPHYLINIDAE: PSELAPHINAE: CLAVIGERITAE: CLAVIGERINI.

Type species: Hoplitoxenus joannae Jeannel, 1960, by original designation.

Family-group name: HOPLITOXENINI Célis, 1969. Stem: Hoplitoxen-.

Hoplopisthius Senna, 1892: 451. Current status: valid genus in BRENTIDAE: BRENTINAE: TRACHELIZINI: HOPLOPISTHIINA.

Type species: Hoplopisthius trichemerus Senna, 1892, by monotypy.

Family-group name: HOPLOPISTHI Senna & Calabresi, 1919. Stem: Hoplopisthi-.

Hoplorhinus Chevrolat, 1878a: 163. Current status: junior synonym of Celetes Schönherr, 1836 in CURCULIONIDAE: CURCULIONINAE: DERELOMINI: DERELOMINA.

Type species: Hoplorhinus melanocephalus Chevrolat, 1878, by present designation.

Family-group name: HOPLORRHININA Champion, 1903. Stem: Hoplorhin-.

Horatoma Solier, 1841a: 210, 264. Current status: valid genus in TENEBRIONIDAE: PIMELIINAE: CRYPTOCHILINI: CRYPTOCHILINA.

Type species: Horatoma parvula Solier, 1841, by original designation.

Family-group name: HORATOMINA C. Koch, 1955. Stem: Horatom-.

Horelophopsis Hansen, 1997: 109. Current status: valid genus in HYDROPHILIDAE: ACIDOCERINAE.

Type species: Horelophopsis avita Hansen, 1997, by original designation.

Family-group name: HORELOPHOPSINAE Hansen, 1997. Stem: Horelophops-.

Horelophus Orchymont, 1913a: 94, 99. Current status: valid genus in HYDROPHILIDAE: CHAETARTHRIINAE: ANACAENINI.

Type species: Horelophus walkeri Orchymont, 1913, by monotypy.

Family-group name: HORELOPHINAE Hansen, 1991. Stem: Horeloph-.

Horia Fabricius, 1787: 164. Current status: valid genus in MELOIDAE: NEMOGNATHINAE: HORIINI. Note: name placed on the Official List of Generic Names in Zoology (ICZN 1999c).

Type species: Lymexylon testaceum Fabricius, 1781 (= Horia fabriciana Betrem, 1929), by subsequent designation (Betrem 1929: xxvii; see ICZN 1999c, as Horia fabriciana Betrem, 1929). Note: Horia fabriciana Betrem, 1929 placed on the Official List of Specific Names in Zoology (ICZN 1999c).

Family-group name: HORIALES Latreille, 1802. Stem: Hori-.

Hormops J.L. LeConte in J.L. LeConte and G.H. Horn, 1876: 321. Current status: valid genus in CURCULIONIDAE: MOLYTINAE: PETALOCHILINI.

Type species: Hormops abducens J.L. LeConte, 1876, by monotypy.

Family-group name: HORMOPINI J.L. LeConte, 1876. Stem: Hormop-.

Hormorus G.H. Horn in J.L. LeConte and G.H. Horn, 1876: 23. Current status: valid genus in CURCULIONIDAE: ENTIMINAE: OTIORHYNCHITAE: HORMORINI.

Type species: Chlorophanus undulatus Uhler, 1855, by monotypy.

Family-group name: HORMORI G.H. Horn, 1876. Stem: Hormor-.

Hornia C.V. Riley, 1877: 564, 565. Current status: valid genus in MELOIDAE: NEMOGNATHINAE: NEMOGNATHINI: NEMOGNATHINA.

Type species: Hornia minutipennis C.V. Riley, 1877, by monotypy.

Family-group name: HORNII Dugès, 1889. Stem: Horni-.

Hornibius Fairmaire, 1888a: 60, 61. Current status: junior synonym of Hornius Fairmaire, 1885 in CHRYSOMELIDAE: SPILOPYRINAE. Note: unnecessary new replacement name for Hornius Fairmaire, 1885.

Type species: Hornius sulcifrons Fairmaire, 1885, by monotypy (type fixation for Hornius Fairmaire).

Family-group name: HORNIBIINI Crowson, 1946. Stem: Hornibi-.

Hornietus Stebnicka, 2000: 287. Current status: valid genus in SCARABAEIDAE: APHODIINAE: HORNIETIINI.

Type species: Pleurophorus ventralis G.H. Horn, 1887, by original designation.

Family-group name: HORNIETIINI Minkina, 2020: 444. Stem: Hornieti- (ICZN 1999a, Article 29.3.3).

Horologion J.M. Valentine, 1932: 2. Current status: valid genus in CARABIDAE: TRECHINAE: HOROLOGIONINI.

Type species: Horologion speokoites J.M. Valentine, 1932, by monotypy.

Family-group name: HOROLOGIONIDAE Jeannel, 1949. Stem: Horologion-.

Hybalus Dejean, 1833b: 149. Current status: valid genus in SCARABAEIDAE: ORPHNINAE: ORPHNINI.

Type species: Geobius cornifrons Brullé, 1832, by monotypy.

Family-group name: HYBALIDAE Marseul, 1857. Stem: Hybal-.

Hybocephalus Motschulsky, 1852: 482. Current status: valid genus in STAPHYLINIDAE: PSELAPHINAE: PSELAPHITAE: HYBOCEPHALINI.

Type species: Hybocephalus minimus Motschulsky, 1852, by subsequent designation (R. Lucas 1920: 339).

Family-group name: HYBOCEPHALINI Raffray, 1890. Stem: Hybocephal-.

Hybodera J.L. LeConte, 1873: 191. Current status: valid genus in CERAMBYCIDAE: CERAMBYCINAE: HYBODERINI.

Type species: Hybodera tuberculata J.L. LeConte, 1873, by monotypy.

Family-group name: HYBODERINI Linsley, 1940. Stem: Hyboder-.

Hybolabus Jekel, 1860: 195. Current status: valid genus in ATTELABIDAE: ATTELABINAE: ATTELABINI: HYBOLABINA.

Type species: Attelabus variabilis Gyllenhal, 1833 (= Attelabus ater G.-A. Olivier, 1789), by original designation.

Family-group name: HYBOLABINA Voss, 1925. Stem: Hybolab-.

Hybomorphus Saunders & Jekel, 1855: 301. Current status: valid genus in CURCULIONIDAE: CURCULIONINAE: STOREINI.

Type species: Hybomorphus melanosomus Saunders & Jekel, 1855, by original designation.

Family-group name: HYBOMORPHIDES Lacordaire, 1865. Stem: Hybomorph-.

Hyborhabdus Aurivillius, 1911b: 22. Current status: valid genus in CERAMBYCIDAE: LAMIINAE: HYBORHABDINI.

Type species: Hyborhabdus singularis Aurivillius, 1911, by monotypy.

Family-group name: HYBORHABDINAE Aurivillius, 1911. Stem: Hyborhabd-.

Hybosispa Weise, 1910b: 96. Current status: valid genus in CHRYSOMELIDAE: CASSIDINAE: HYBOSISPINI.

Type species: Hybosispa melanura Weise, 1910, by monotypy.

Family-group name: HYBOSITES Weise, 1910. Stem: Hybosisp-.

Hybosorus W.S. MacLeay, 1819: 120. Current status: valid genus in HYBOSORIDAE: HYBOSORINAE.

Type species: here fixed (under ICZN 1999a, Article 70.3.2) as Hybosorus illigeri Reiche, 1853, misidentified as Scarabaeus arator Fabricius, 1775 (sensu W.S. MacLeay, 1819) in the original designation by monotypy in W.S. MacLeay (1819: 120). Note: see Branco (2007: 13) for interpretation of the type species; Hybosorus illigeri Reiche, 1853 was placed on the Official List of Specific Names in Zoology (ICZN 2021c).

Family-group name: HYBOSORIDAE Erichson, 1847. Stem: Hybosor-.

Hydaticus Leach, 1817: 69, 72. Current status: valid genus in DYTISCIDAE: DYTISCINAE: HYDATICINI.

Type species: Dytiscus transversalis Pontoppidan, 1763, by subsequent designation (Curtis 1825: pl. 95).

Family-group name: HYDATICINI Sharp, 1880. Stem: Hydatic-.

Hydatophilus Gistel, 1856: 353. Current status: junior synonym of Hydrophilus Geoffroy, 1762 in HYDROPHILIDAE: HYDROPHILINAE: HYDROPHILINI.

Type species: Dytiscus piceus Linnaeus, 1758, by monotypy.

Family-group name: HYDATOPHILIDAE Gistel, 1856. Stem: Hydatophil-.

Hyderodes Hope, 1838a: 131, 166. Current status: valid genus in DYTISCIDAE: DYTISCINAE: DYTISCINI.

Type species: Hyderodes shuckardi Hope, 1838, by original designation.

Family-group name: HYDERODINI K.B. Miller, 2000. Stem: Hyderod-.

Hydnobius W.L.E. Schmidt, 1841: 193. Current status: valid genus in LEIODIDAE: LEIODINAE: SOGDINI.

Type species: Anisotoma punctatum Sturm, 1807, by subsequent designation (C.G. Thomson 1859: 58).

Family-group name: HYDNOBIINI Jeannel, 1962. Stem: Hydnobi-.

Hydnocera Newman, 1838: 379. Current status: junior synonym of Phyllobaenus Dejean, 1833 in CLERIDAE: CLERINAE: CLERITAE: HYDNOCERINI: HYDNOCERINA.

Type species: Hydnocera serrata Newman, 1838 (= Clerus pallipennis Say, 1825), by monotypy.

Family-group name: HYDNOCÉROÏDES Spinola, 1844. Stem: Hydnocer-.

Hydraena Kugelann, 1794: 578. Current status: valid genus in HYDRAENIDAE: HYDRAENINAE: HYDRAENINI.

Type species: Hydraena riparia Kugelann, 1794, by monotypy.

Family-group name: HYDRAENAIRES Mulsant, 1844. Stem: Hydraen-.

Hydraenida Germain, 1901: 531, 537. Current status: valid genus in HYDRAENIDAE: HYDRAENINAE: HYDRAENIDINI. Note: the original spelling was Hydroenida, but Hydraenida, an incorrect subsequent spelling introduced by d’ Orchymont (1913b: 103), is in prevailing usage, attributed to the publication of the original spelling, and therefore deemed to be the correct original spelling (ICZN 1999a, Article 33.3.1).

Type species: Hydraenida ocellata Germain, 1901, by monotypy.

Family-group name: HYDRAENIDINI Perkins, 1980. Stem: Hydraenid-.

Hydrobius Leach, 1815a: 96. Current status: valid genus in HYDROPHILIDAE: HYDROPHILINAE: HYDROBIUSINI. Note: name placed on the Official List of Generic Names in Zoology (ICZN 1990a).

Type species: Dytiscus fuscipes Linnaeus, 1758, by subsequent designation (Hope 1838a: 125 [December]; see ICZN 1990a). Note: name placed on the Official List of Specific Names in Zoology (ICZN 1990a); Westwood (1838c: 10 [May]) is the first author who selected that species as type species of Hydrobius Leach, 1815; however, the Commission (ICZN 1990a) set aside all fixations of type species for the nominal genus Hydrobius Leach, 1815 before the designation of Hope (1838a).

Family-group name: HYDROBIAIRES Mulsant, 1844. Stem: Hydrobius- (ICZN 2003a).

Hydrocanthus Say, 1823: 105. Current status: valid genus in NOTERIDAE: NOTERINAE: NOTERINI.

Type species: Hydrocanthus iricolor Say, 1823, by monotypy.

Family-group name: HYDROCANTHINI Sharp, 1880. Stem: Hydrocanth-.

Hydrochus Leach, 1817: 90. Current status: valid genus in HYDROCHIDAE.

Type species: Silpha elongata Schaller, 1783, by subsequent designation (Curtis 1831: pl. 359).

Family-group name: HYDROCHIDAE C.G. Thomson, 1859. Stem: Hydroch-.

Hydrocoptus Motschulsky, 1853c: 5. Current status: junior synonym of Hydroporus Clairville, 1806 in DYTISCIDAE: HYDROPORINAE: HYDROPORINI: HYDROPORINA.

Type species: Dytiscus tristis Paykull, 1798, by subsequent designation (Nilsson et al. 1989: 297).

Family-group name: HYDROCOPTINI Kolbe, 1883. Stem: Hydrocopt-.

Hydrodytes K.B. Miller, 2001: 77. Current status: valid genus in DYTISCIDAE: HYDRODYTINAE.

Type species: Agaporomorphus opalinus A. Zimmermann, 1921, by original designation.

Family-group name: HYDRODYTINAE K.B. Miller, 2001. Stem: Hydrodyt-.

Hydronebrius Jakowlew, 1897: 37. Current status: valid genus in DYTISCIDAE: AGABINAE: AGABINI.

Type species: Agabus cordaticollis Reitter, 1896, by original designation.

Family-group name: HYDRONEBRIINI Guignot, 1948. Stem: Hydronebri-.

Hydronomus Schönherr, 1825: 583. Current status: junior synonym of Bagous Germar, 1817 in CURCULIONIDAE: BAGOINAE.

Type species: Curculio alismatis Marsham, 1802, by original designation.

Family-group name: HYDRONOMIDES Lacordaire, 1863. Stem: Hydronom-.

Hydrophilus Geoffroy, 1762: 180. Current status: valid genus in HYDROPHILIDAE: HYDROPHILINAE: HYDROPHILINI. Note: name placed on the Official List of Generic Names in Zoology (ICZN 1994a).

Type species: Dytiscus piceus Linnaeus, 1758, by subsequent designation (Latreille 1810: 428; see ICZN 1994a). Note: name placed on the Official List of Specific Names in Zoology (ICZN 1994a).

Family-group name: HYDROPHILII Latreille, 1802. Stem: Hydrophil-.

Hydroporus Clairville, 1806: 182. Current status: valid genus in DYTISCIDAE: HYDROPORINAE: HYDROPORINI: HYDROPORINA.

Type species: Dytiscus fusculus Schrank, 1781 (= Dytiscus planus Fabricius, 1781), by subsequent designation (Ádám 1996: 20). Note: Dytiscus planus Fabricius, 1781 was placed on the Official List of Specific Names in Zoology (ICZN 1989e).

Family-group name: HYDROPORIDES Aubé, 1836. Stem: Hydropor-.

Hydroscapha J.L. LeConte, 1874b: 45. Current status: valid genus in HYDROSCAPHIDAE: HYDROSCAPHINAE.

Type species: Hydroscapha natans J.L. LeConte, 1874, by monotypy.

Family-group name: HYDROSCAPHIDAE J.L. LeConte, 1874. Stem: Hydroscaph-.

Hydrotrupes Sharp, 1882b: 491, 492. Current status: valid genus in DYTISCIDAE: AGABINAE: HYDROTRUPINI.

Type species: Hydrotrupes palpalis Sharp, 1882, by monotypy.

Family-group name: HYDROTRUPINAE Roughley, 2000. Stem: Hydrotrup-.

Hydrovatus Motschulsky, 1853c: 4. Current status: valid genus in DYTISCIDAE: HYDROPORINAE: HYDROVATINI.

Type species: Hyphydrus cuspidatus Kunze, 1818, by monotypy.

Family-group name: HYDROVATINI Sharp, 1880. Stem: Hydrovat-.

Hygrobia Latreille, 1804a: 139. Current status: valid genus in HYGROBIIDAE. Note: new replacement name for Hydrachna Fabricius, 1801; this name was originally spelled Hygriobia but subsequently emended to Hygrobia by the Commission and placed on the Official List of Generic Names in Zoology (ICZN 1954b).

Type species: Dytiscus hermanni Fabricius, 1775 [as herrmanni], by monotypy (see ICZN 1954b). Note: name placed on the Official List of Specific Names in Zoology (ICZN 1954b).

Family-group name: HYGROBIINAE Régimbart, 1879. Stem: Hygrobi-.

Hygronoma Erichson, 1837: 312. Current status: valid genus in STAPHYLINIDAE: ALEOCHARINAE: HYGRONOMINI: HYGRONOMINA.

Type species: Aleochara dimidiata Gravenhorst, 1806, by monotypy.

Family-group name: HYGRONOMIDES C.G. Thomson, 1859. Stem: Hygronom-.

Hygrotus Stephens, 1828: 38, 46. Current status: valid genus in DYTISCIDAE: HYDROPORINAE: HYGROTINI.

Type species: Dytiscus inaequalis Fabricius, 1777, by subsequent designation (Curtis 1835: pl. 531).

Family-group name: HYGROTINI Portevin, 1929. Stem: Hygrot-.

Hylaspes Baly, 1865c: 436. Current status: junior synonym of Gallerucida Motschulsky, 1861 in CHRYSOMELIDAE: GALERUCINAE: HYLASPINI.

Type species: Hylaspes longicornis Baly, 1865, by original designation.

Family-group name: HYLASPITES Chapuis, 1875. Stem: Hylasp-.

Hylastes Erichson, 1836: 47. Current status: valid genus in CURCULIONIDAE: SCOLYTINAE: HYLASTINI.

Type species [suggested]: Bostrichus ater Paykull, 1800, by subsequent designation (Westwood 1838c: 39). Note: as pointed out by Alonso-Zarazaga et al. (2016: 5), “Bostrichus ater Paykull, 1800” is a misidentification of Bostrichus ater Fabricius, 1792 (= Bostrichus bidentatus Herbst, 1783, currently in the genus Pityogenes Bedel, 1888) and is therefore an unavailable name; however, because “Bostrichus ater Paykull, 1800” is the accepted name of a major pest, we believe that an application be submitted to the Commission to treat Paykull’s name as available and the type species of Hylastes Erichson, 1836.

Family-group name: HYLASTES J.L. LeConte, 1876. Stem: Hylast-.

Hylecoetus Latreille, 1805: 266. Current status: junior synonym of Elateroides Schäffer, 1777 in LYMEXYLIDAE: HYLECOETINAE.

Type species: Cantharis dermestoides Linnaeus, 1761, by monotypy.

Family-group name: HYLECOETI Germar, 1818. Stem: Hylecoet-.

Hylepnigalio Gistel, 1856: 384. Current status: junior synonym of Melandrya Fabricius, 1801 in MELANDRYIDAE: MELANDRYINAE: MELANDRYINI.

Type species: Chrysomela caraboides Linnaeus, 1761, by subsequent designation (Bouchard et al. 2011: 388).

Family-group name: HYLEPNIGALIONIDAE Gistel, 1856. Stem: Hylepnigalion-.

Hylesinus Fabricius, 1801a: xii. Current status: valid genus in CURCULIONIDAE: SCOLYTINAE: HYLESININI.

Type species: Hylesinus crenatus Fabricius, 1801, by subsequent designation (Latreille 1810: 431). Note: no species were originally included in this genus by Fabricius (1801a); the first ones were the 30 species listed by Fabricius (1801b: 390–395).

Family-group name: HYLESINEN Erichson, 1836. Stem: Hylesin-.

Hylobius Germar, 1817b: 340. Current status: valid genus in CURCULIONIDAE: MOLYTINAE: HYLOBIINI: HYLOBIINA.

Type species: Curculio pineti Fabricius, 1792 (= Curculio excavatus Laicharting, 1781), by subsequent designation (Schönherr 1823: 1143).

Family-group name: HYLOBIDAE W. Kirby, 1837. Stem: Hylobi-.

Hylochares Latreille, 1834: 127. Current status: valid genus in EUCNEMIDAE: MELASINAE: HYLOCHARINI.

Type species: Elater cruentatus Gyllenhal, 1808, by subsequent designation (Lacordaire 1857: 114).

Family-group name: HYLOCHARITES Jacquelin du Val, 1859. Stem: Hylochar-.

Hylocurus Eichhoff, 1872: 133. Current status: valid genus in CURCULIONIDAE: SCOLYTINAE: MICRACIDINI.

Type species: Hylocurus elegans Eichhoff, 1872, by monotypy.

Family-group name: HYLOCURIDAE Eichhoff, 1878. Stem: Hylocur-.

Hylophilus Berthold, 1827: 375. Current status: invalid senior synonym of Aderus Stephens, 1829 in ADERIDAE. Note: junior homonym of Hylophilus Temminck, 1822 [Aves].

Type species: Notoxus populneus Panzer, 1796, by original designation.

Family-group name: HYLOPHILIDAE Streubel, 1846. Stem: Hylophil-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

Hylotorus Dalman, 1823: 103. Current status: valid genus in CARABIDAE: PAUSSINAE: PAUSSINI: PAUSSINA.

Type species: Paussus bucephalus Gyllenhal, 1817, by monotypy.

Family-group name: HYLOTORINI Luna de Carvalho, 1951. Stem: Hylotor-.

Hylotrupes Audinet-Serville, 1834b: 77. Current status: valid genus in CERAMBYCIDAE: CERAMBYCINAE: HYLOTRUPINI.

Type species: Cerambyx bajulus Linnaeus, 1758, by monotypy.

Family-group name: HYLOTRUPINI Rose, 1983. Stem: Hylotrup-.

Hylurgops J.L. LeConte in J.L. LeConte and G.H. Horn, 1876: 389. Current status: valid genus in CURCULIONIDAE: SCOLYTINAE: HYLASTINI.

Type species: Hylastes pinifex Fitch, 1858 (= Hylurgus rugipennis Mannerheim, 1843), by subsequent designation (Hopkins 1914: 123).

Family-group name: HYLURGOPINA Balachowsky, 1949. Stem: Hylurgop-.

Hylurgus Latreille, 1806: 274. Current status: valid genus in CURCULIONIDAE: SCOLYTINAE: HYLURGINI.

Type species: Bostrichus ligniperda Fabricius, 1787 [as Bostricilus ligniperda], by monotypy.

Family-group name: HYLURGIDAE Gistel, 1848. Stem: Hylurg-.

Hymaea Pascoe, 1869a: 35. Current status: valid genus in PHLOEOSTICHIDAE.

Type species: Hymaea succinifera Pascoe, 1869, by monotypy.

Family-group name: HYMAEINAE Sen Gupta & Crowson, 1966. Stem: Hymae-.

Hyocis Pascoe, 1866b: 457. Current status: valid genus in TENEBRIONIDAE: DIAPERINAE: HYOCIINI: HYOCIINA.

Type species: Hyocis bakewellii Pascoe, 1866, by original designation.

Family-group name: HYOCINI G.S. Medvedev & Lawrence, 1982. Stem: Hyoci-.

Hyorrhynchus W.F.H. Blandford, 1894: 58. Current status: valid genus in CURCULIONIDAE: SCOLYTINAE: HYORRHYNCHINI.

Type species: Hyorrhynchus lewisi W.F.H. Blandford, 1894, by monotypy.

Family-group name: HYORRHYNCHINAE Hopkins, 1915. Stem: Hyorrhynch-.

Hypasclera Kirsch, 1866: 210. Current status: valid genus in OEDEMERIDAE: OEDEMERINAE: ASCLERINI.

Type species: Hypasclera schistacea Kirsch, 1866, by subsequent designation (Arnett 1950: 221).

Family-group name: HYPASCLERINI Arnett, 1984. Stem: Hypascler-.

Hypera Germar, 1817b: 340. Current status: valid genus in CURCULIONIDAE: HYPERINAE: HYPERINI: HYPERINA.

Type species: Curculio nigrirostris Fabricius, 1775, by subsequent designation (Samouelle 1819: 205).

Family-group name: HYPÉRIDES Lacordaire, 1863. Stem: Hyper-.

Hyperacantha Chapuis, 1879: 18. Current status: valid genus in CHRYSOMELIDAE: GALERUCINAE: LUPERINI.

Type species: Hyperacantha fenestrata Chapuis, 1879, by subsequent designation (Hincks 1949: 612).

Family-group name: HYPERACANTHITES Laboissière, 1924. Stem: Hyperacanth-.

Hyperaspis Chevrolat in Dejean, 1836a: 435. Current status: valid genus in COCCINELLIDAE: COCCINELLINAE: HYPERASPIDINI: HYPERASPIDINA.

Type species: Coccinella reppensis Herbst, 1783, by subsequent designation (C.G. Thomson 1859: 161).

Family-group name: HYPÉRASPIENS Mulsant, 1846. Stem: Hyperaspid-.

Hyphalus Britton, 1971: 88. Current status: valid genus in LIMNICHIDAE: HYPHALINAE.

Type species: Hyphalus insularis Britton, 1971, by original designation.

Family-group name: HYPHALINAE Britton, 1971. Stem: Hyphal-.

Hyphydrus Illiger, 1802: 299. Current status: valid genus in DYTISCIDAE: HYDROPORINAE: HYPHYDRINI.

Type species: Dytiscus gibbus Fabricius, 1777 (= Dytiscus ovatus Linnaeus, 1761), by subsequent designation (Latreille 1810: 426).

Family-group name: HYPHYDRIIDAE Gistel, 1848. Stem: Hyphydr-.

Hypnoidus Dillwyn, 1829: 32. Current status: valid genus in ELATERIDAE: HYPNOIDINAE.

Type species: Elater riparius Fabricius, 1792, by subsequent designation (Westwood 1838c: 26).

Family-group name: HYPNOIDINI O. Schwarz, 1906. Stem: Hypnoid-.

†Hypnomorphus Dolin, 1975: 54. Current status: valid genus in ELATERIDAE: †PROTAGRYPNINAE: †HYPNOMORPHINI. Note: Jurassic (Kazakhstan).

Type species: †Hypnomorphus rohdendorfi Dolin, 1975, by original designation.

Family-group name: †HYPNOMORPHINI Dolin, 1975. Stem: Hypnomorph-.

Hypoborus Erichson, 1836: 62. Current status: valid genus in CURCULIONIDAE: SCOLYTINAE: HYPOBORINI: HYPOBORINA.

Type species: Hypoborus ficus Erichson, 1836, by monotypy.

Family-group name: HYPOBORINAE Nüsslin, 1911. Stem: Hypobor-.

Hypocaccus C.G. Thomson, 1867: 400. Current status: valid genus in HISTERIDAE: SAPRININAE: HYPOCACCINI.

Type species: here fixed (under ICZN 1999a, Article 70.3.2) as Hister rugiceps Duftschmid, 1805, misidentified as Hister quadristriatus Thunberg, 1794 (sensu Hoffmann, 1803) in the subsequent designation by Lewis (1899: 3).

Family-group name: HYPOCACCINI Lackner in Lackner et al. 2024: 60. Stem: Hypocacc-.

Hypocephalus A.G. Desmarest, 1832: pl. 24. Current status: valid genus in VESPERIDAE: ANOPLODERMATINAE: HYPOCEPHALINI.

Type species: Hypocephalus armatus A.G. Desmarest, 1832, by monotypy.

Family-group name: HYPOCÉPHALIENS Blanchard, 1845. Stem: Hypocephal-.

Hypocoprus Motschulsky, 1839: 72. Current status: valid genus in CRYPTOPHAGIDAE: ATOMARIINAE: HYPOCOPRINI.

Type species: Hypocoprus latridioides Motschulsky, 1839, by monotypy.

Family-group name: HYPOCOPRINI Reitter, 1879. Stem: Hypocopr-.

Hypocyphtus Gyllenhal, 1827: 294. Current status: junior synonym of Cypha Leach, 1819 in STAPHYLINIDAE: ALEOCHARINAE: HYPOCYPHTINI.

Type species: Scaphidium longicorne Paykull, 1800, by monotypy.

Family-group name: HYPOCYPHTIDAE Laporte, 1835. Stem: Hypocypht-.

Hypodacne J.L. LeConte, 1876: 170. Current status: valid genus in EUXESTIDAE.

Type species: Hypodacne punctata J.L. LeConte, 1876, by monotypy.

Family-group name: HYPODACNINAE Dajoz, 1976. Stem: Hypodacn-.

Hypodesis Latreille, 1834: 156. Current status: valid genus in ELATERIDAE: ELATERINAE: ELATERINI.

Type species: Hypodesis sericea Latreille, 1834, by monotypy.

Family-group name: HYPODÉSITES Candèze, 1863. Stem: Hypodese-.

Hypohypurus Hustache, 1920b: 334. Current status: valid genus in CURCULIONIDAE: CEUTORHYNCHINAE: HYPOHYPURINI.

Type species: Hypohypurus perrieri Hustache, 1920, by subsequent designation (Colonelli 1997: 387).

Family-group name: HYPOHYPURINI Colonnelli, 2004. Stem: Hypohypur-.

Hypolithus Eschscholtz, 1829a: 33. Current status: valid genus in ELATERIDAE: HYPNOIDINAE. Note: this name, issued by 10 May 1829, is a senior homonym of Hypolithus Dejean, 1829 [Coleoptera: CARABIDAE], published by 23 November, 1829.

Type species: Hypolithus littoralis Eschscholtz, 1829, by subsequent designation (Hyslop 1921: 650).

Family-group name: HYPOLITHINAE Fleutiaux, 1928. Stem: Hypolith-.

Hypomelus Solier, 1843: 4, 93. Current status: valid genus in TENEBRIONIDAE: PIMELIINAE: SEPIDIINI: HYPOMELINA.

Type species: Hypomelus bicolor Solier, 1843 (= Helops peronatus Germar, 1823), by original designation.

Family-group name: HYPOMELINA C. Koch, 1955. Stem: Hypomel-.

Hypophlaeus Fabricius, 1790: 222. Current status: junior synonym of Corticeus Piller & Mitterpacher, 1783 in TENEBRIONIDAE: DIAPERINAE: HYPOPHLAEINI.

Type species: Hypophlaeus castaneus Fabricius, 1790 (= Corticeus unicolor Piller & Mitterpacher, 1783), by subsequent designation (Curtis 1832: pl. 430).

Family-group name: HYPOPHLAEIDES Billberg, 1820. Stem: Hypophlae-.

Hypoprasis Fairmaire & Germain, 1864: 260. Current status: valid genus in BUPRESTIDAE: CHRYSOCHROINAE: PARALEPTODEMINI: HYPOPRASINA.

Type species: Hypoprasis harpagon Fairmaire & Germain, 1864, by monotypy.

Family-group name: HYPOPRASINA Holyński, 1993. Stem: Hypopras-.

Hypselogenia Burmeister, 1840b: [Heft 6]. Current status: valid genus in SCARABAEIDAE: CETONIINAE: GOLIATHINI: GOLIATHINA.

Type species: Diplognatha concava Gory & Percheron, 1833, by subsequent designation (Krikken 1979: 114).

Family-group name: HYPSELOGENIAE Schoch, 1894. Stem: Hypselogeni-.

Hypselomus Perty, 1832: 95. Current status: valid genus in CERAMBYCIDAE: LAMIINAE: ONCIDERINI.

Type species: Hypselomus cristatus Perty, 1832, by monotypy.

Family-group name: HYPSELOMINAE Pascoe, 1864. Stem: Hypselom-.

Hypsioma Audinet-Serville, 1835a: 38. Current status: valid genus in CERAMBYCIDAE: LAMIINAE: ONCIDERINI.

Type species: Hypsioma gibberum Audinet-Serville, 1835, by monotypy.

Family-group name: HYPSIOMITAE J. Thomson, 1860. Stem: Hypsiomat-.

Hypsonotus Germar, 1823: 367. Current status: valid genus in CURCULIONIDAE: ENTIMINAE: ENTIMITAE: LORDOPINI.

Type species: Hypsonotus clavulus Germar, 1823, by subsequent designation (Schönherr 1826: 13).

Family-group name: HYPSONOTIDAE Jekel, 1853. Stem: Hypsonot-.

Hyptioma Casey, 1906: 362. Current status: junior synonym of Holisus Erichson, 1839 in STAPHYLINIDAE: STAPHYLININAE: STAPHYLININI: HYPTIOMINA.

Type species: Hyptioma cubensis Casey, 1906, by monotypy.

Family-group name: HYPTIOMAE Casey, 1906. Stem: Hyptiom-.

Hypulus Paykull, 1798: 251. Current status: valid genus in MELANDRYIDAE: MELANDRYINAE: HYPULINI.

Type species: Elater quercinus Quensel, 1790, by subsequent designation (Curtis 1829: pl. 255).

Family-group name: HYPULIDAE Gistel, 1848. Stem: Hypul-. Note: junior homonym of HYPULIA Rafinesque, 1815 (type genus Hypulus Rafinesque, 1815) in Coleoptera: TENEBRIONIDAE; Rafinesque’s name is permanently invalid (based on a preoccupied type genus).

Hypulus Rafinesque, 1815: 114. Current status: junior synonym of Helops Fabricius, 1775 in TENEBRIONIDAE: TENEBRIONINAE: HELOPINI: HELOPINA. Note: unnecessary new replacement name for Helops Fabricius, 1775; junior homonym of Hypulus Paykull, 1798 [Coleoptera: MELANDRYIDAE].

Type species: Tenebrio caeruleus Linnaeus, 1758, by subsequent designation (Hope 1841a: 133) (type fixation for Helops Fabricius).

Family-group name: HYPULIA Rafinesque, 1815. Stem: Hypul-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

Hypurus Rey, 1882: 187, 188. Current status: valid genus in CURCULIONIDAE: CEUTORHYNCHINAE: HYPURINI.

Type species: Ceutorhynchus bertrandi Perris, 1852, by subsequent designation (K.W. Kirby 1884: 105).

Family-group name: HYPURIDAE Schultze, 1902. Stem: Hypur-.

Iberobaenia Bocák, Kundrata, Andújar & Vogler, 2016: 4. Current status: valid genus in IBEROBAENIIDAE. Note: combined description of a new family-group name and a single new genus (ICZN 1999a, Article 13.5).

Type species: Iberobaenia minuta Bocák, Kundrata, Andújar & Vogler, 2016, by original designation.

Family-group name: IBEROBAENIIDAE Bocák, Kundrata, Andújar & Vogler, 2016: 5. Stem: Iberobaeni-.

Ibidion Audinet-Serville, 1834b: 103. Current status: invalid senior synonym of Neoibidion Monné, 2012 in CERAMBYCIDAE: CERAMBYCINAE: TROPIDINI: NEOIBIDIONINA. Note: junior homonym of Ibidion Gory, 1833 [Coleoptera: CERAMBYCIDAE].

Type species: Ibidion comatum Audinet-Serville, 1834, by subsequent designation (J. Thomson 1864: 215).

Family-group name: IBIDIONITAE J. Thomson, 1861. Stem: Ibidion-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

Ichnea Laporte, 1838: 55. Current status: valid genus in CLERIDAE: KORYNETINAE.

Type species: Ichnea lycoides Laporte, 1838, by monotypy.

Family-group name: ICHNOÏDES Spinola, 1841. Stem: Ichne-.

Ichnestoma Gory & Percheron, 1833: 19, 41. Current status: valid genus in SCARABAEIDAE: CETONIINAE: GOLIATHINI: ICHNESTOMATINA.

Type species: Ichnestoma heteroclyta Gory & Percheron, 1833, by monotypy. Note: this species is made available by the combined description of a new genus and a single new species (ICZN 1999a, Article 12.2.6); the other species in the genus were described by Gory & Percheron in another livraison of the work issued in 1834.

Family-group name: ISCHNOSTOMIDAE Burmeister, 1842. Stem: Ichnestomat-. Note: see Perissinotto (2022: 279) regarding the recommendation to use the incorrectly formed ICHNESTOMINA.

Ichthyurus Westwood, 1848a: 83. Current status: valid genus in CANTHARIDAE: CHAULIOGNATHINAE: ICHTHYURINI.

Type species: Ichthyurus lateralis Westwood, 1848, by subsequent designation (Delkeskamp 1977: 468).

Family-group name: ICHTHYURINI Champion, 1915. Stem: Ichthyur-. Note: junior homonym of ICHTHYURINAE Packard, 1895 (type genus Ichthyura Hübner, 1819) in Lepidoptera.

Icthyosoma Boisduval, 1835: 468. Current status: junior synonym of Tmesisternus Latreille, 1829 in CERAMBYCIDAE: LAMIINAE: TMESISTERNINI. Note: this taxon was originally proposed as a synonym of Tmesisternus Latreille, 1829 and subsequently treated as valid before 1961 by Montrouzier (1855: 58, 59).

Type species: Tmesisternus oblongus Boisduval, 1835, by present designation. Note: Bousquet et al. (2009: 39) designated Icthyosoma armatum Montrouzier, 1855 as type species of Icthyosoma but that species is not an originally included species.

Family-group name: ICHTHYOSOMITAE J. Thomson, 1864. Stem: Icthyosomat-.

Ictistygna Pascoe, 1866b: 491. Current status: valid genus in ANTHICIDAE: EURYGENIINAE: ICTISTYGNINI.

Type species: Ictistygna adusta Pascoe, 1866, by subsequent designation (Borchmann 1936: 537).

Family-group name: ICTISTYGNINAE Borchmann, 1936. Stem: Ictistygn-.

Idacantha Fairmaire, 1869: 252. Current status: junior synonym of Diacantha Chevrolat, 1836 in CHRYSOMELIDAE: GALERUCINAE: LUPERINI. Note: unnecessary new replacement name for Diacantha Chevrolat, 1836.

Type species: Galleruca unifasciata G.-A. Olivier, 1808, by subsequent designation (Brauer 1870: 141).

Family-group name: IDACANTHITES Laboissière, 1921. Stem: Idacanth-.

Ideratus J. Thomson, 1864: 183. Current status: valid genus in CERAMBYCIDAE: CERAMBYCINAE: IDERATINI.

Type species: Ideratus cyanipennis J. Thomson, 1864, by monotypy.

Family-group name: IDERATINI Martins & Napp, 2009. Stem: Iderat-.

Idiobius Gistel, 1856: 383. Current status: junior synonym of Olibrus Erichson, 1845 in PHALACRIDAE: PHALACRINAE.

Type species: Phalacrus flavicornis Sturm, 1807, by subsequent designation (Pakaluk et al. 1994: 229).

Family-group name: IDIOBIIDAE Gistel, 1856. Stem: Idiobi-.

Idiomacer Kuschel in Kuschel and Leschen, 2011: 592. Current status: valid genus in NEMONYCHIDAE: RHINORHYNCHINAE: IDIOMACERINI.

Type species: Idiomacer basicornis Kuschel, 2011, by original designation.

Family-group name: IDIOMACERINAE Legalov, 2011a: 163. Stem: Idiomacer-.

Idiomorphus Chaudoir, 1847: 515. Current status: valid genus in CARABIDAE: HARPALINAE: IDIOMORPHINI.

Type species: Idiomorphus guerinii Chaudoir, 1847, by monotypy.

Family-group name: IDIOMORPHINAE H.W. Bates, 1891. Stem: Idiomorph-.

Idiostoma Arrow, 1904: 740. Current status: invalid senior synonym of Allidiostoma Arrow, 1940 in SCARABAEIDAE: ALLIDIOSTOMATINAE. Note: junior homonym of Idiostoma Walsingham, 1882 [Lepidoptera].

Type species: Idiostoma rufum Arrow, 1904, by original designation.

Family-group name: IDIOSTOMINAE Arrow, 1904. Stem: Idiostomat-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

Idisia Pascoe, 1866b: 452. Current status: valid genus in TENEBRIONIDAE: PIMELIINAE: IDISIINI.

Type species: Idisia ornata Pascoe, 1866, by monotypy.

Family-group name: IDISIINI G.S. Medvedev, 1973. Stem: Idisi-.

Illops Erichson, 1840b: 87. Current status: valid genus in MELYRIDAE: MALACHIINAE: MALACHIINI.

Type species: Illops corniculatus Erichson, 1840, by monotypy.

Family-group name: ILLOPINA Jakobson, 1911. Stem: Illop-.

Ilybius Erichson, 1832: 18, 34. Current status: valid genus in DYTISCIDAE: AGABINAE: AGABINI.

Type species: Dytiscus fenestratus Fabricius, 1781, by subsequent designation (Westwood 1838c: 8).

Family-group name: ILYBII Acloque, 1895. Stem: Ilybi-.

Imatidium Fabricius, 1801a: xvii, 345. Current status: valid genus in CHRYSOMELIDAE: CASSIDINAE: IMATIDIINI.

Type species: Imatidium thoracicum Fabricius, 1801, by subsequent designation (Latreille 1810: 432).

Family-group name: IMATIDIIDAE Hope, 1841. Stem: Imatidi-.

Imirus Reitter, 1885: 228. Current status: valid genus in STAPHYLINIDAE: PSELAPHINAE: GONIACERITAE: IMIRINI. Note: new replacement name for Mirus Saulcy, 1877.

Type species: Mirus permirus Saulcy, 1877, by monotypy (type fixation for Mirus Saulcy).

Family-group name: IMIRINI Jeannel, 1949. Stem: Imir-.

Inca Lepeletier & Audinet-Serville, 1828: 380. Current status: valid genus in SCARABAEIDAE: CETONIINAE: INCINI.

Type species: Cetonia clathrata G.-A. Olivier, 1792, by subsequent designation (E. Desmarest 1860: 59). Note: the type species usually listed for this genus is Cetonia ynca Weber, 1801 (e.g., Siedel et al. 2018: 396) by subsequent designation in Howden (1968: 12), but Demarest’s fixation is older.

Family-group name: INCADAE Burmeister, 1842. Stem: Inc-.

Indoquedius Blackwelder, 1952: 199. Current status: valid genus in STAPHYLINIDAE: STAPHYLININAE: STAPHYLININI: INDOQUEDIINA. Note: this name was originally proposed by Cameron (1932: 281) who failed to fix a type species as required for all genus-group names published after 1930 (ICZN 1999a, Article 13.3); Blackwelder (1952: 1999) was the first author to do so.

Type species: Quedius oculatus Fauvel, 1895, by original designation.

Family-group name: INDOQUEDIINA Brunke & Solodovnikov in Brunke et al., 2016: 447. Stem: Indoquedi-.

Iniocyphus Raffray, 1912: 435. Current status: valid genus in STAPHYLINIDAE: PSELAPHINAE: GONIACERITAE: INIOCYPHINI: INIOCYPHINA.

Type species: Iniocyphus iheringi Raffray, 1912, by monotypy.

Family-group name: INIOCYPHINI Park, 1951. Stem: Iniocyph-.

Inopeplus F. Smith, 1851: 4. Current status: valid genus in SALPINGIDAE: INOPEPLINAE. Note: new replacement name for Ino Laporte, 1835.

Type species: Ino picta Laporte, 1835, by monotypy (type fixation for Ino Laporte).

Family-group name: INOPEPLINI Grouvelle, 1908. Stem: Inopepl-.

Iphimeis Baly, 1864c: 133. Current status: valid genus in CHRYSOMELIDAE: EUMOLPINAE: EUMOLPINI.

Type species: Iphimeis fulvipes Baly, 1864, by original designation.

Family-group name: IPHIMÉITES Chapuis, 1874. Stem: Iphime-.

Iphius Schönherr, 1823: 1140. Current status: junior synonym of Alceis Billberg, 1820 in CURCULIONIDAE: ENTIMINAE: POLYDRUSITAE: NAUPACTINI: NAUPACTINA. Note: unnecessary new replacement name for Alceis Billberg, 1820.

Type species: Curculio longimanus Fabricius, 1775 (= Siderodactylus ornatus Pascoe, 1879), by monotypy (type fixation for Alceis Billberg).

Family-group name: IPHIIDES Schönherr, 1823. Stem: Iphi-. Note: nomen oblitum (see Bouchard et al. 2011: 611).

Ips DeGeer, 1775: 190, 193. Current status: valid genus in CURCULIONIDAE: SCOLYTINAE: IPINI: IPINA.

Type species: Dermestes typographus Linnaeus, 1758, by subsequent designation (Crotch 1870a: 44).

Family-group name: IPINI Bedel, 1888. Stem: Ip-.

Iresia Dejean in Dejean and Boisduval, 1829: 5, 8. Current status: valid genus in CICINDELIDAE: CICINDELINI: IRESIINA.

Type species: Iresia lacordairei Dejean, 1829, by monotypy.

Family-group name: IRESINA Rivalier, 1971. Stem: Iresi-.

Iridotaenia Deyrolle, 1865: 25. Current status: valid genus in BUPRESTIDAE: CHRYSOCHROINAE: CHRYSOCHROINI: CHALCOPHORINA. Note: the original spelling was Iridotoenia but Iridotaenia, an incorrect subsequent spelling, is in prevailing usage, attributed to the publication of the original spelling, and therefore deemed to be the correct original spelling (ICZN 1999a, Article 33.3.1); name placed on the Official List of Generic Names in Zoology (ICZN 2004c).

Type species: Chrysodema sumptuosa Laporte & Gory, 1837, by subsequent designation (Kurosawa 1982: 192; see ICZN 2004c). Note: name placed on the Official List of Specific Names in Zoology (ICZN 2004c, Chrysodema sumptuosa Laporte & Gory, 1835).

Family-group name: IRIDOTAENINI Tôyama, 1987. Stem: Iridotaeni-.

Isaniris J. Thomson, 1858b: 129. Current status: valid genus in CURCULIONIDAE: ENTIMINAE: OTIORHYNCHITAE: ISANIRINI.

Type species: Isaniris viridimicans J. Thomson, 1858, by subsequent designation (G.A.K. Marshall 1944: 337).

Family-group name: ISANIRINI Legalov, 2020b: 340. Stem: Isanir-.

Ischalia Pascoe, 1860b: 54. Current status: valid genus in ISCHALIIDAE.

Type species: Ischalia indigacea Pascoe, 1860, by monotypy.

Family-group name: ISCHALIINAE Blair, 1920. Stem: Ischali-.

Ischioloncha J. Thomson, 1860a: 122. Current status: valid genus in CERAMBYCIDAE: LAMIINAE: APOMECYNINI.

Type species: Ischioloncha wollastonii J. Thomson, 1860, by monotypy.

Family-group name: ISCHIOLONCHIDES Lacordaire, 1872. Stem: Ischiolonch-.

Ischiopachys Chevrolat in Dejean, 1836a: 416. Current status: valid genus in CHRYSOMELIDAE: CRYPTOCEPHALINAE: CLYTRINI: BABIINA.

Type species: Clytra bicolor G.-A. Olivier, 1791, by monotypy.

Family-group name: ISCHIOPACHITES Chapuis, 1874. Stem: Ischiopache-.

Ischnocerus Schönherr, 1839: 191. Current status: junior synonym of Meconemus Labram & Imhoff, 1838 in ANTHRIBIDAE: ANTHRIBINAE: MECOCERINI.

Type species: Ischnocerus infuscatus Fåhraeus, 1839 (= Meconemus tuberculatus Labram & Imhoff, 1838), by original designation.

Family-group name: ISCHNOCÉRIDES Lacordaire, 1865. Stem: Ischnocer-.

Ischnomerus Labram & Imhoff, 1838: [Heft 2]. Current status: valid genus in BRENTIDAE: BRENTINAE: BRENTINI: ITHYSTENINA.

Type species: Ischnomerus erythroderes Labram & Imhoff, 1838, by monotypy.

Family-group name: ISCHNOMERI Kolbe, 1883. Stem: Ischnomer-. Note: junior homonym of ISCHNOMÉRIDES Lacordaire, 1865 (type genus Ischnomerus Schönherr, 1840) in Coleoptera: BRENTIDAE; Lacordaire’s name is permanently invalid (based on a preoccupied type genus).

Ischnomerus Schönherr, 1840a: 571. Current status: junior synonym of Aulacoderes Chevrolat, 1839 in BRENTIDAE: BRENTINAE: TAPHRODERINI. Note: two original spellings were used, including Ischnocerus (p. 572); subsequently, Schönherr (1842a: 492) used Ischnomerus and therefore acted as First Reviser (ICZN 1999a, Article 24.2.4); junior homonym of Ischnomerus Labram & Imhoff, 1838 [Coleoptera: BRENTIDAE].

Type species: Ischnomerus linearis Boheman, 1840 (= Aulacoderes immotus Chevrolat, 1839), by original designation.

Family-group name: ISCHNOMÉRIDES Lacordaire, 1865. Stem: Ischnomer-. Note: senior homonym of ISCHNOMERI Kolbe, 1883 (type genus Ischnomerus Labram & Imhoff, 1838) in Coleoptera: BRENTIDAE; family-group name permanently invalid (ICZN 1999a, Article 39).

Ischnoscelis Burmeister, 1842: 179. Current status: valid genus in SCARABAEIDAE: CETONIINAE: GOLIATHINI: RHOMBORHININA.

Type species: Goliathus hoepfneri Gory & Percheron, 1833, by monotypy. Note: the original spelling of the type species was hoepfner; however, the incorrect subsequent spelling hoepfneri is in prevailing usage, attributed to the publication of the original spelling, and therefore deemed to be the correct original spelling (ICZN 1999a, Article 33.3.1).

Family-group name: ISCHNOSCELI Schoch, 1894. Stem: Ischnoscelid-.

Ischnovalgus Kolbe, 1897b: 185, 190. Current status: valid genus in SCARABAEIDAE: CETONIINAE: VALGINI: VALGINA.

Type species: Ischnovalgus gracilis Kolbe, 1897, by subsequent designation (Krikken 1978: 158).

Family-group name: ISCHNOVALGINAE Kolbe, 1904. Stem: Ischnovalg-.

Ischyomius Chevrolat, 1878b: 98. Current status: valid genus in PYTHIDAE.

Type species: Ischyomius singularis Chevrolat, 1878, by monotypy.

Family-group name: ISCHYOMIIDES Champion, 1886. Stem: Ischyomi-.

Ischyrosonyx Sturm, 1843: 273. Current status: junior synonym of Eurypedus Gistel, 1834 in CHRYSOMELIDAE: CASSIDINAE: ISCHYROSONYCHINI.

Type species: Cassida oblonga Thon, 1826 (= Ischyrosonyx thoni Barber, 1946), by monotypy.

Family-group name: ISCHYROSONYCHITES Chapuis, 1875. Stem: Ischyrosonych-.

Isoclerus Lewis, 1892: 191. Current status: valid genus in THANEROCLERIDAE: THANEROCLERINAE: ISOCLERINI.

Type species: Isoclerus pictus Lewis, 1892, by monotypy.

Family-group name: ISOCLERINA Kolibáč, 1992. Stem: Isocler-.

Isolabus Voss, 1925: 214. Current status: valid genus in ATTELABIDAE: ATTELABINAE: ATTELABINI: ISOLABINA.

Type species: Attelabus caeruleus Jekel, 1860 (= Isolabus jekeli Legalov, 2002), by original designation.

Family-group name: ISOLABINA Legalov, 2007. Stem: Isolab-.

Isonychus Mannerheim, 1828: 69. Current status: valid genus in SCARABAEIDAE: MELOLONTHINAE: MACRODACTYLINI.

Type species: Isonychus sulphureus Mannerheim, 1828, by original designation.

Family-group name: ISONYCHIDAE Burmeister, 1855. Stem: Isonych-.

Isopleurus W. Kirby, 1837: 49. Current status: junior synonym of Celia C. Zimmermann, 1832 in CARABIDAE: HARPALINAE: ZABRINI: AMARINA.

Type species: Isopleurus nitidus W. Kirby, 1837 (= Celia sinuosa Casey, 1918), by monotypy.

Family-group name: ISOPLEURIDAE W. Kirby, 1837. Stem: Isopleur-.

Isoplia Burmeister, 1855: 487. Current status: valid genus in SCARABAEIDAE: RUTELINAE: ANOMALINI: ISOPLIINA.

Type species: Isoplia lasiosoma Burmeister, 1855, by monotypy.

Family-group name: ISOPLIINI Péringuey, 1902. Stem: Isopli-.

Isopterus Faust, 1895: 4. Current status: valid genus in CURCULIONIDAE: ENTIMINAE: OTIORHYNCHITAE: CELEUTHETINI: ISOPTERINA.

Type species: Isopterus signatus Faust, 1895, by subsequent designation (Yunakov 2013: 65).

Family-group name: ISOPTERINA Morimoto & Kojima, 2001. Stem: Isopter-.

Isorhynchus Schönherr, 1833: 22. Current status: valid genus in CURCULIONIDAE: CEUTORHYNCHINAE: CEUTORHYNCHINI.

Type species: Curculio pudicus Sparrman, 1785, by original designation.

Family-group name: ISORHYNCHIDES Lacordaire, 1865. Stem: Isorhynch-.

†Isothea Scudder, 1893: 20. Current status: valid genus in ATTELABIDAE: RHYNCHITINAE: RHYNCHITITAE: RHYNCHITINI: RHYNCHITINA. Note: Oligocene (USA).

Type species: †Isothea alleni Scudder,1893, by monotypy.

Family-group name: †ISOTHEINAE Scudder, 1893. Stem: Isothe-.

Istrisia Lewis, 1895a: 254. Current status: valid genus in SALPINGIDAE: SALPINGINAE.

Type species: Istrisia rufobrunnea Lewis, 1895, by monotypy.

Family-group name: ISTRISIINI Nikitsky, 1992. Stem: Istrisi-.

Ita Tournier, 1878: xviii. Current status: valid genus in CURCULIONIDAE: MOLYTINAE: ITINI.

Type species: Ita crassirostris Tournier, 1878, by subsequent designation (Alonso-Zarazaga and Lyal 1999: 21, 79).

Family-group name: ITINI Reitter, 1913. Stem: It-. Note: senior homonym of ITINI Lepesme, 1943 (type genus Ites C.O. Waterhouse, 1880) in Coleoptera: CERAMBYCIDAE.

Italodytes G. Müller, 1938: 136, 139. Current status: valid genus in CARABIDAE: SCARITINAE: CLIVININI: CLIVININA.

Type species: Italodytes stammeri G. Müller, 1938, by original designation.

Family-group name: ITALODYTINA Jeannel, 1957. Stem: Italodyt-.

Ites C.O. Waterhouse, 1880b: 297. Current status: valid genus in CERAMBYCIDAE: LAMIINAE: HEMILOPHINI.

Type species: Ites plagiatus C.O. Waterhouse, 1880, by monotypy.

Family-group name: ITESINI Lepesme, 1943. Stem: It-. Note: junior homonym of ITINI Reitter, 1913 (type genus Ita Tournier, 1878) in Coleoptera: CURCULIONIDAE.

Ithaura Pascoe, 1871b: 215. Current status: valid genus in CURCULIONIDAE: MOLYTINAE: LYMANTINI: LYMANTINA.

Type species: Ithaura strangulata Pacoe, 1871, by monotypy.

Family-group name: ITHAURINAE Kuschel, 1959. Stem: Ithaur-.

Ithycerus Schönherr, 1823: 1136. Current status: valid genus in BRENTIDAE: ITHYCERINAE: ITHYCERINI.

Type species: Rhynchites curculionoides Herbst, 1797 (= Curculio noveboracensis Forster, 1771), by original designation.

Family-group name: ITHYCERIDES Schönherr, 1823. Stem: Ithycer-.

Ithyporus Schönherr, 1836: 550. Current status: valid genus in CURCULIONIDAE: MOLYTINAE: ITHYPORINI: ITHYPORINA.

Type species: Ithyporus capensis Boheman, 1836 (= Curculio stolidus Fabricius, 1781), by original designation.

Family-group name: ITHYPORIDES Lacordaire, 1865. Stem: Ithypor-.

Ithystenus Pascoe, 1862: 390. Current status: valid genus in BRENTIDAE: BRENTINAE: BRENTINI: ITHYSTENINA. Note: new replacement name for Leptorhynchus Boisduval, 1835.

Type species: Brenthus angustatus Guérin-Méneville, 1831, by monotypy (type fixation for Leptorhynchus Boisduval).

Family-group name: ITHYSTÉNIDES Lacordaire, 1865. Stem: Ithysten-.

Ivieolus Howden & Gill, 1988: 2077. Current status: valid genus in HYBOSORIDAE: CERATOCANTHINAE: SCARABATERMITINI.

Type species: Ivieolus pseudoscutellatus Howden & Gill, 1988, by original designation.

Family-group name: IVIEOLINI Howden & Gill, 2000. Stem: Ivieol-.

Ixalma Pascoe, 1871b: 214. Current status: valid genus in CURCULIONIDAE: CURCULIONINAE: RHAMPHINI: IXALMINA.

Type species: Ixalma rufescens Pascoe, 1871, by monotypy.

Family-group name: IXALMINA Voss, 1936. Stem: Ixalm-.

Ixapion Roudier & Tempère, 1973: 80. Current status: valid genus in BRENTIDAE: APIONINAE: APIONITAE: IXAPIINI. Note: new replacement name for Ixias Sainte Claire-Deville, 1924.

Type species: Apion variegatum Wencker, 1864, by monotypy (type fixation for Ixias Sainte Claire-Deville).

Family-group name: IXAPIINI Alonso-Zarazaga, 1990. Stem: Ixapi-.

Jacobsonium Heller, 1926: 127. Current status: junior synonym of Sarothrias Grouvelle, 1918 in JACOBSONIIDAE.

Type species: Jacobsonium dimerum Heller, 1926, by original designation.

Family-group name: JACOBSONIIDAE Heller, 1926. Stem: Jacobsoni-.

Jamwonus Harold, 1879: 158. Current status: valid genus in CERAMBYCIDAE: PRIONINAE: AEGOSOMATINI.

Type species: Jamwonus subcostatus Harold, 1879, by monotypy.

Family-group name: JAMWONINAE Kolbe, 1897. Stem: Jamwon-.

Janssenius Luna de Carvalho, 1951: 19. Current status: junior synonym of Heteropaussus J. Thomson, 1860 in CARABIDAE: PAUSSINAE: PAUSSINI: HETEROPAUSSINA. Note: new replacement name for Pleuropterus Westwood, 1841.

Type species: Cerapterus westermanni Westwood, 1840, by monotypy (type fixation for Pleuropterus Westwood).

Family-group name: JANSSENINI Luna de Carvalho, 1951. Stem: Jansseni-.

Janusculus Cerruti, 1970: 117. Current status: valid genus in STAPHYLINIDAE: PSELAPHINAE: PSELAPHITAE: TYRINI: JANUSCULINA.

Type species: Janusculus leleupi Cerruti, 1970, by monotypy.

Family-group name: JANUSCULINA Cerruti, 1970. Stem: Januscul-.

Jenibuntor Muona, 1993: 21. Current status: valid genus in EUCNEMIDAE: MACRAULACINAE: JENIBUNTORINI.

Type species: Jenibuntor tabang Muona, 1993, by original designation.

Family-group name: JENIBUNTORINI Muona, 1993. Stem: Jenibuntor-.

Jordanthribus Zimmerman, 1938: 236. Current status: valid genus in ANTHRIBIDAE: ANTHRIBINAE: PROSCOPORHININI.

Type species: Jordanthribus planifacietus Zimmerman, 1938, by original designation.

Family-group name: JORDANTHRIBINI Morimoto, 1980. Stem: Jordanthrib-.

Juanorhinus Aurivillius, 1926: 466. Current status: valid genus in CURCULIONIDAE: MOLYTINAE: JUANORHININI.

Type species: Juanorhinus robinsoni Aurivillius, 1926, by monotypy.

Family-group name: JUANORHININI Aurivillius, 1931. Stem: Juanorhin-.

Jubus L.W. Schaufuss, 1872: 247. Current status: valid genus in STAPHYLINIDAE: PSELAPHINAE: EUPLECTITAE: JUBINI.

Type species: Jubus spinicollis L.W. Schaufuss, 1877, by subsequent designation (R. Lucas 1920: 355). Note: no species were originally included in this genus by L.W. Schaufuss (1872); the first ones were the four species listed by L.W. Schaufuss (1877b: 455–456).

Family-group name: JUBINI Raffray, 1898. Stem: Jub-.

Julodimorpha Harold in Gemminger and Harold, 1869b: 1397. Current status: valid genus in BUPRESTIDAE: BUPRESTINAE: JULODIMORPHINI.

Type species: Stigmodera bakewellii A. White, 1859, by monotypy.

Family-group name: JULODIMORPHITES Kerremans, 1903. Stem: Julodimorph-.

Julodis Eschscholtz, 1829b: 9. Current status: valid genus in BUPRESTIDAE: JULODINAE. Note: the original spelling was Jalodis but Julodis, an unjustified emendation by von Mannerheim (1837: 11), is in prevailing usage, attributed to the original author and date, and therefore deemed to be a justified emendation and the correct original spelling (ICZN 1999a, Article 33.2.3.1).

Type species: Buprestis fascicularis Linnaeus, 1758, by subsequent designation (Gussmann 1997: 338).

Family-group name: JULODIDES Lacordaire, 1857. Stem: Julod-.

Jumnos Saunders, 1839: 176. Current status: valid genus in SCARABAEIDAE: CETONIINAE: GOLIATHINI: RHOMBORHININA.

Type species: Jumnos ruckeri Saunders, 1839, by monotypy.

Family-group name: JUMNIDAE Burmeister, 1842. Stem: Jumn-.

†Juranthribus Legalov, 2011b: 35. Current status: valid genus in NEMONYCHIDAE: †BRENTHORRHININAE: †JURANTHRIBINI. Note: Upper Jurassic (Karatau).

Type species: †Juranthribus thompsoni Legalov, 2011, by original designation.

Family-group name: †JURANTHRIBINAE Legalov, 2011b: 34. Stem: Juranthrib-.

Jurasai Rosa, C. Costa, Kramp & Kundrata, 2020b: [1]. Current status: valid genus in JURASAIDAE. Note: this genus-group name was first proposed by Rosa et al. (2020a: [4]) but the work was not available nomenclaturally because the electronic-only version was not registered in ZooBank before it was published (ICZN 2012f, Article 8.5.3).

Type species: Jurasai itajubense Rosa, C. Costa, Kramp & Kundrata, 2020, by original designation.

Family-group name: JURASAIDAE Rosa, C. Costa, Kramp & Kundrata, 2020b: [3]. Stem: Jurasa-. Note: this family-group name was first proposed by Rosa et al. (2020a: [7]) but the work was not available nomenclaturally because the electronic-only version was not registered in ZooBank before it was published (ICZN 2012f, Article 8.5.3).

†Jurodes Ponomarenko, 1985: 54. Current status: valid genus in JURODIDAE. Note: Middle/Upper Jurassic (Russia; China; Mongolia).

Type species: †Jurodes ignoramus Ponomarenko, 1985, by monotypy.

Family-group name: †JURODIDAE Ponomarenko, 1985. Stem: Jurod-.

Kalcapion Schilsky, 1906: iv. Current status: valid genus in BRENTIDAE: APIONINAE: ASPIDAPIITAE: KALCAPIINI.

Type species: Apion pallipes W. Kirby, 1808, by original designation.

Family-group name: KALCAPIINI Alonso-Zarazaga, 1990. Stem: Kalcapi-.

†Kaltanocoleus Rohdendorf, 1961: 397. Current status: valid genus in †PERMOCUPEDIDAE: †PERMOCUPEDINAE. Note: lower Permian (Russia).

Type species: †Kaltanocoleus pospelovi Rohdendorf, 1961, by original designation.

Family-group name: †KALTANOCOLEIDAE Rohdendorf, 1961. Stem: Kaltanocole-.

†Kararhynchus Zherikhin & Gratshev, 1994: 60. Current status: valid genus in †OBRIENIIDAE: KARARHYNCHINAE: †KARARHYNCHINI. Note: Upper Jurassic (Kazakhstan).

Type species: †Kararhynchus occiduus Zherikhin & Gratshev, 1994, by original designation.

Family-group name: †KARARHYNCHINAE Zherikhin & Gratshev, 1994. Stem: Kararhynch-.

†Karataucar Legalov, 2009a: 286, 288. Current status: junior synonym of Ampliceps Arnoldi, 1977 in NEMONYCHIDAE: †EOBELINAE: †KARATAUCARINI. Note: Upper Jurassic (Kazakhstan).

Type species: †Oxycorynoides progressivus Arnoldi, 1977, by original designation.

Family-group name: †KARATAUCARINI Legalov, 2009. Stem: Karataucar-.

Karumia Escalera, 1913: 320. Current status: valid genus in DASCILLIDAE: KARUMIINAE: KARUMIINI.

Type species: Karumia estafilinoides Escalera, 1913, by subsequent designation (R. Lucas 1920: 356).

Family-group name: KARUMINAE Escalera, 1913. Stem: Karumi-.

Kateretes Herbst, 1793: 11. Current status: valid genus in KATERETIDAE: KATERETINAE. Note: name placed on the Official List of Generic Names in Zoology (ICZN 1999b).

Type species: Dermestes pedicularius Linnaeus, 1758, by subsequent designation (Hope 1841a: 155; see ICZN 1999b). Note: name placed on the Official List of Specific Names in Zoology (ICZN 1999b).

Family-group name: CATHERETIDAE W. Kirby, 1837. Stem: Kateret-.

Kaupiolus Zang, 1903: 418. Current status: junior synonym of Labienus Kaup, 1871 in PASSALIDAE: PASSALINAE: MACROLININI. Note: new replacement name for Vellejus Kaup, 1871.

Type species: Passalus compergus Boisduval, 1835, by subsequent designation (Hincks and Dibb 1935: 93).

Family-group name: KAUPIOLINAE Zang, 1905. Stem: Kaupiol-.

†Kenderlyka Zherikhin & Gratshev, 1994: 59. Current status: valid genus in †OBRIENIIDAE: †KARARHYNCHINAE: †KENDERLYKANINI. Note: Upper Triassic (Kazakhstan).

Type species: †Kenderlyka consobrina Zherikhin & Gratshev, 1994, by original designation.

Family-group name: †KENDERLYKANINI Legalov, 2009. Stem: Kenderlykan- (ICZN 1999a, Article 29.4).

Kisanthobia Marseul, 1865: 195, 200. Current status: valid genus in BUPRESTIDAE: BUPRESTINAE: KISANTHOBIINI.

Type species: Anthaxia ariasi Robert, 1858, by monotypy.

Family-group name: KISANTHOBIINI Richter, 1949. Stem: Kisanthobi-.

Klewaria Reitter, 1910: 20. Current status: valid genus in TENEBRIONIDAE: PIMELIINAE: KLEWARIINI.

Type species: Klewaria colydiiformis Reitter, 1910, by monotypy.

Family-group name: KLEWARIINAE Gebien, 1910. Stem: Klewari-.

Korynetes Herbst, 1792: 148. Current status: valid genus in CLERIDAE: KORYNETINAE. Note: name placed on the Official List of Generic Names in Zoology (ICZN 1961c).

Type species: Clerus caeruleus DeGeer, 1775, by subsequent designation (Lacordaire 1857: 489; see ICZN 1961c). Note: name placed on the Official List of Specific Names in Zoology (ICZN 1961c).

Family-group name: CORYNETIDAE Laporte, 1838. Stem: Korynet-.

†Kresnikus Tihelka, D. Huang & Cai, 2019b: 507. Current status: valid genus in TROGIDAE: †KRESNIKINAE. Note: mid-Cretaceous (Myanmar).

Type species: †Kresnikus beynoni Tihelka, D. Huang & Cai, 2019, by original designation.

Family-group name: †KRESNIKINAE Tihelka, D. Huang & Cai, 2019: 507. Stem: Kresnik-.

†Kryzhanovskiana Kataev & Kirejtshuk in Kataev et al. 2019: 2. Current status: valid genus in CARABIDAE: PAUSSINAE: KRYZHANOVSKIANINI. Note: mid-Cretaceous (Myanmar).

Type species: †Kryzhanovskiana olegi Kataev & Kirejtshuk, 2019, by original designation.

Family-group name: †KRYZHANOVSKIANINAE Deuve, 2020: 203. Stem: Kryzhanovskian-.

Kuhitangia G.S. Medvedev, 1962: 1184. Current status: valid genus in TENEBRIONIDAE: KUHITANGIINAE: KUHITANGIINI.

Type species: Kuhitangia kryzhanovskii G.S. Medvedev, 1962, by original designation.

Family-group name: KUHITANGIINAE G.S. Medvedev, 1962. Stem: Kuhitangi-.

Kuschelinus Straneo, 1963: 124. Current status: valid genus in CARABIDAE: HARPALINAE: PTEROSTICHINI: METIINA.

Type species: Kuschelinus insularis Straneo, 1963, by original designation.

Family-group name: KUSCHELININI Jeannel, 1967. Stem: Kuschelin-.

†Kuschelomacer Riedel, 2010: 31. Current status: valid genus in CIMBERIDIDAE: †KUSCHELOMACERINI. Note: Eocene (Baltic amber).

Type species: †Kuschelomacer kerneggeri Riedel, 2010, by original designation.

Family-group name: †KUSCHELOMACERINI Riedel, 2010. Stem: Kuschelomacer- (ICZN 1999a, Article 29.4).

Kytorhinus Fischer, 1809: 298, 300. Current status: valid genus in CHRYSOMELIDAE: BRUCHINAE: KYTORHININI. Note: two original spellings were used, including Kytorhynus (p. 298); subsequently, Fischer (1821b: 73) used only Kytorhinus and therefore acted as First Reviser (ICZN 1999a, Article 24.2.4).

Type species: Kytorhinus karasini Fischer, 1809, by subsequent designation (Crotch 1870b: 222).

Family-group name: KYTORHININAE Bridwell, 1932. Stem: Kytorhin-.

Labaninus Morimoto, 1981: 110. Current status: valid genus in CURCULIONIDAE: CURCULIONINAE: CURCULIONINI: LABANININA.

Type species: Curponinus plicatulus Heller, 1925, by original designation.

Family-group name: LABANININA Pelsue & O’Brien, 2011: 30. Stem: Labanin-.

Labidopullus Borgmeier, 1958: 240. Current status: valid genus in STAPHYLINIDAE: ALEOCHARINAE: MIMECITINI: LABIDOPULLINA.

Type species: Labidopullus appendiculatus Borgmeier, 1958, by original designation.

Family-group name: LABIDOPULLINA Jacobson & Kistner, 1991. Stem: Labidopull-.

†Labradorocoleus Ponomarenko, 1969b: 309. Current status: valid genus in †LABRADOROCOLEIDAE. Note: Upper Cretaceous (Labrador, Canada).

Type species: †Labradorocoleus carpenteri Ponomarenko, 1969, by original designation.

Family-group name: †LABRADOROCOLEIDAE Ponomarenko, 1969. Stem: Labradorocole-.

Laccobius Erichson, 1837: 202. Current status: valid genus in HYDROPHILIDAE: HYDROPHILINAE: LACCOBIINI.

Type species: Chrysomela minuta Linnaeus, 1758, by subsequent designation (Drapiez 1839: 222).

Family-group name: LACCOBIINI Houlbert, 1922. Stem: Laccobi-.

Lacconectus Motschulsky, 1855b: 83. Current status: valid genus in DYTISCIDAE: COPELATINAE.

Type species: Lacconectus fulvescens Motschulsky, 1855, by monotypy.

Family-group name: LACCONECTINI Branden, 1884. Stem: Lacconect-.

Lacconotus J.L. LeConte, 1862: 255. Current status: valid genus in MYCTERIDAE: EURYPINAE.

Type species: Lacconotus punctatus J.L. LeConte, 1862, by monotypy.

Family-group name: LACCONOTINI J.L. LeConte, 1862. Stem: Lacconot-.

Laccophilus Leach, 1817: 69, 72. Current status: valid genus in DYTISCIDAE: LACCOPHILINAE: LACCOPHILINI.

Type species: Dytiscus minutus Linnaeus, 1758, by monotypy.

Family-group name: LACCOPHILIDAE Gistel, 1848. Stem: Laccophil-.

Laccornellus Roughley and Wolfe, 1987: 1347. Current status: valid genus in DYTISCIDAE: HYDROPORINAE: LACCORNELLINI.

Type species: Hydroporus lugubris Aubé, 1838, by original designation.

Family-group name: LACCORNELLINI K.B. Miller & Bergsten, 2014: 139. Stem: Laccornell-.

Laccornis Gozis, 1914: 111. Current status: valid genus in DYTISCIDAE: HYDROPORINAE: LACCORNINI.

Type species: Hydroporus oblongus Stephens, 1835, by monotypy.

Family-group name: LACCORNINI Wolfe & Roughley, 1990. Stem: Laccorn-.

Laches Kaup, 1871: 48. Current status: junior synonym of Episphenus Kaup, 1871 in PASSALIDAE: PASSALINAE: MACROLININI. Note: junior homonym of Laches Gistel, 1848 [Hymenoptera] and Laches Thorell, 1869 [Arachnida]; the First Reviser (ICZN 1999a, Article 24.2.2) for Laches Kaup, 1871 versus Episphenus Kaup, 1871 is Gravely (1914: 281).

Type species: Aceraius comptoni Kaup, 1868, by subsequent designation (Hincks and Dibb 1935: 77).

Family-group name: LACHINAE Kuwert, 1896. Stem: Lach-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

Lachna Billberg, 1820: 35. Current status: junior synonym of Lagria Fabricius, 1775 in TENEBRIONIDAE: LAGRIINAE: LAGRIINI: LAGRIINA.

Type species: Chrysomela hirta Linnaeus, 1758, by subsequent designation (Merkl 2006: 223).

Family-group name: LACHNAEDES Billberg, 1820. Stem: Lachn-. Note: nomen oblitum (see Favret and Bouchard 2016: 647); senior homonym of LACHNIDAE Herrich- Schäffer, 1854 (type genus Lachnus Burmeister 1835) in Hemiptera.

Lachnodactylus Seidlitz, 1898: 837, 838. Current status: valid genus in TENEBRIONIDAE: PIMELIINAE: LACHNOGYINI: LACHNODACTYLINA. Note: new replacement name for Lachnopus Seidlitz, 1894.

Type species: Lachnopus digitalis Seidlitz, 1894, by monotypy (type fixation for Lachnopus Seidlitz).

Family-group name: LACHNODACTYLINA Reitter, 1904. Stem: Lachnodactyl-.

Lachnogya Ménétriés, 1849b: 228. Current status: valid genus in TENEBRIONIDAE: PIMELIINAE: LACHNOGYINI: LACHNOGYINA.

Type species: Lachnogya squamosa Ménétriés, 1849, by monotypy.

Family-group name: LACHNOGYINI Seidlitz, 1894. Stem: Lachnogy-.

Lachnophorus Dejean, 1831: 2, 28. Current status: valid genus in CARABIDAE: HARPALINAE: LACHNOPHORINI.

Type species: Lachnophorus pilosus Dejean, 1831, by subsequent designation (E. Desmarest 1851: 189).

Family-group name: LACHNOPHORI J.L. LeConte, 1853. Stem: Lachnophor-.

Lacon Laporte, 1838: 11. Current status: valid genus in ELATERIDAE: AGRYPNINAE: AGRYPNINI.

Type species: Elater atomarius Fabricius, 1798 (= Elater punctatus Herbst, 1779), by subsequent designation (Hyslop 1921: 652).

Family-group name: LACONINI Dajoz, 1964. Stem: Lacon-.

Lactica Erichson, 1847c: 173. Current status: junior synonym of Monomacra Chevrolat, 1836 in CHRYSOMELIDAE: GALERUCINAE: ALTICINI.

Type species: Lactica melaleuca Erichson, 1847, by monotypy.

Family-group name: LACTICITES Chapuis, 1875. Stem: Lactic-.

Laemophloeus Dejean, 1835: 315. Current status: valid genus in LAEMOPHLOEIDAE.

Type species: Cucujus muticus Fabricius, 1781, by subsequent designation (C.G. Thomson 1859: 83).

Family-group name: LAEMOPHLOEINI Ganglbauer, 1899. Stem: Laemophloe-.

Laemosaccus Schönherr, 1823: 1136. Current status: valid genus in CURCULIONIDAE: MESOPTILIINAE: LAEMOSACCINI.

Type species: Curculio plagiatus Fabricius, 1798 (= Curculio nephele Herbst, 1797), by original designation.

Family-group name: LÉMOSACIDES Lacordaire, 1865. Stem: Laemosacc-.

Laemotmetus Gerstaecker, 1871: 45. Current status: junior synonym of Aulonosoma Motschulsky, 1858 in PASSANDRIDAE: PASSANDRINAE.

Type species: Laemotmetus ferrugineus Gerstaecker, 1871 (= Aulonosoma tenebrioides Motschulsky, 1858), by monotypy.

Family-group name: LAEMOTMETINI Kessel, 1921. Stem: Laemotmet-.

Laena Dejean, 1821: 64. Current status: valid genus in TENEBRIONIDAE: LAGRIINAE: LAENINI.

Type species: fixed by Bouchard et al. (2021: 225) according to Article 70.3.2 (ICZN 1999a) as Scaurus viennensis Sturm, 1807, the taxonomic species involved in the misidentification of Helops pimelia Fabricius, 1787 (sensu Dejean, 1821), as originally designated by monotypy by Dejean (1821: 64). Note: see Bouchard and Bousquet (2020b: 7) for comments regarding the identity of the type species.

Family-group name: LAENINA Seidlitz, 1895. Stem: Laen-.

Lagenoderus A. White, 1841: 181. Current status: valid genus in ATTELABIDAE: ATTELABINAE: ATTELABINI: LAGENODERINA.

Type species: Attelabus gnomoides A. White, 1841, by monotypy.

Family-group name: LAGENODERINA Voss, 1925. Stem: Lagenoder-.

Lagocheirus Dejean, 1835: 336. Current status: valid genus in CERAMBYCIDAE: LAMIINAE: ACANTHOCININI.

Type species: Cerambyx araneiformis Linnaeus, 1767, by monotypy.

Family-group name: LAGOCHEIRINAE H.W. Bates, 1863. Stem: Lagocheir-.

Lagria Fabricius, 1775: 124. Current status: valid genus in TENEBRIONIDAE: LAGRIINAE: LAGRIINI: LAGRIINA.

Type species: Chrysomela hirta Linnaeus, 1758, by subsequent designation (Latreille 1810: 429).

Family-group name: LAGRIARIAE Latreille, 1825. Stem: Lagri-.

Lagrioida Fairmaire & Germain, 1860a: 3. Current status: valid genus in LAGRIOIDIDAE.

Type species: Lagrioida rufula Fairmaire & Germain, 1860, by subsequent designation (Spilman 1954: 89).

Family-group name: LAGRIOIDINI M. Abdullah & A. Abdullah, 1968. Stem: Lagrioid-.

Laius Guérin-Méneville, 1830: pl. 2. Current status: valid genus in MELYRIDAE: MALACHIINAE: MALACHIINI.

Type species: Laius cyaneus Guérin-Méneville, 1830, by monotypy.

Family-group name: LAIINA Jakobson, 1911. Stem: Lai-.

Lamia Fabricius, 1775: 170. Current status: valid genus in CERAMBYCIDAE: LAMIINAE: LAMIINI.

Type species: Cerambyx textor Linnaeus, 1758, by subsequent designation (Latreille 1810: 431).

Family-group name: LAMIARIAE Latreille, 1825. Stem: Lami-.

Lamingtonium Sen Gupta & Crowson, 1969: 125. Current status: valid genus in LAMINGTONIIDAE.

Type species: Lamingtonium binnaburrense Sen Gupta & Crowson, 1969, by original designation.

Family-group name: LAMINGTONIIDAE Sen Gupta & Crowson, 1969. Stem: Lamingtoni-.

Lamprias Bonelli, 1810: Tab. Syn. Current status: valid subgenus of Lebia Latreille, 1802 in CARABIDAE: HARPALINAE: LEBIINI: LEBIINA.

Type species: Carabus cyanocephalus Linnaeus, 1758, by subsequent monotypy (Panzer 1813: 71).

Family-group name: LAMPRIADAE Chaudoir, 1871. Stem: Lampri-.

Lamprigera Motschulsky, 1853a: 47. Current status: valid genus in LAMPYRIDAE incertae sedis.

Type species: Lamprigera boyei Motschulsky, 1853, by original designation.

Family-group name: LAMPRIGERINA McDermott, 1964. Stem: Lampriger-.

Lamprima Latreille, 1804a: 150. Current status: valid genus in LUCANIDAE: LAMPRIMINAE: LAMPRIMINI.

Type species: Lethrus aeneus Fabricius, 1792, by monotypy.

Family-group name: LAMPRIMIDAE W.S. MacLeay, 1819. Stem: Lamprim-.

Lamprocera Laporte, 1833c: 129. Current status: valid genus in LAMPYRIDAE: LAMPYRINAE: LAMPROCERINI.

Type species: Homalisus grandis Sturm, 1826, by monotypy.

Family-group name: LAMPROCERINI E. Olivier, 1907. Stem: Lamprocer-.

Lamprocheila Saunders, 1871: 15. Current status: valid genus in BUPRESTIDAE: BUPRESTINAE: BUPRESTINI: LAMPROCHEILINA.

Type species: Chrysodema maillei Laporte & Gory, 1837, by monotypy.

Family-group name: LAMPROCHEILINA Holyński, 1993. Stem: Lamprocheil-.

Lamprohiza Motschulsky, 1853a: 47. Current status: valid genus in LAMPYRIDAE: LAMPROHIZINAE.

Type species: Lampyris splendidula Linnaeus, 1767, by original designation.

Family-group name: LAMPROHIZINI Kazantsev, 2010. Stem: Lamprohiz-.

Lamprolabus Jekel, 1860: 189. Current status: valid genus in ATTELABIDAE: ATTELABINAE: ATTELABINI: LAMPROLABINA.

Type species: Attelabus bispinosus Gyllenhal, 1833, by original designation.

Family-group name: LAMPROLABINA Voss, 1925. Stem: Lamprolab-.

Lampropepla Fairmaire, 1904a: 231. Current status: junior synonym of Madecassia Kerremans, 1903 in BUPRESTIDAE: CHRYSOCHROINAE: CHRYSOCHROINI: CHALCOPHORINA. Note: unnecessary new replacement name for Madecassia Kerremans, 1903.

Type species: Chalcophoropsis rothschildi Gahan, 1893, by monotypy (type fixation for Madecassia Kerremans).

Family-group name: LAMPROPEPLINA Holyński, 1993. Stem: Lampropepl-.

Lamprosoma W. Kirby, 1819: 445. Current status: valid genus in CHRYSOMELIDAE: LAMPROSOMATINAE: LAMPROSOMATINI.

Type species: Lamprosoma bicolor W. Kirby, 1819, by subsequent designation (Hope 1841a: 162).

Family-group name: LAMPROSOMIDEAE Lacordaire, 1848. Stem: Lamprosomat-.

Lampyris Geoffroy, 1762: 165. Current status: valid genus in LAMPYRIDAE: LAMPYRINAE: LAMPYRINI. Note: name attributed to Geoffroy, 1762 and placed on the Official List of Generic Names in Zoology (ICZN 1984a).

Type species: Cantharis noctiluca Linnaeus, 1758, by plenary power (see ICZN 1984a). Note: name placed on the Official List of Specific Names in Zoology (ICZN 1984a).

Family-group name: LAMPYRIA Rafinesque, 1815. Stem: Lampyr-.

Lampyrolycus Burgeon, 1937: 152. Current status: valid genus in LYCIDAE: DEXORINAE: MIMOLIBNETINI: LAMPYROLYCINA.

Type species: Lampyrolycus hulstaerti Burgeon, 1937, by monotypy.

Family-group name: LAMPYROLYCINI Kazantsev, 2018: 149. Stem: Lampyrolyc-.

Lancetes Sharp, 1882b: 562, 602. Current status: valid genus in DYTISCIDAE: LANCETINAE.

Type species: Dytiscus varius Fabricius, 1775, by subsequent designation (Guignot 1946: 117).

Family-group name: LANCETINI Branden, 1884. Stem: Lancet-.

Langbianella Prokofiev, 2015: 21. Current status: valid genus in SCARABAEIDAE: MELOLONTHINAE: LANGBIANELLINI.

Type species: Langbianellina auricomes Prokofiev, 2015, by original designation.

Family-group name: LANGBIANELLINI Prokofiev, 2015: 20. Stem: Langbianell-.

Langelandia Aubé, 1843: 227. Current status: valid genus in ZOPHERIDAE: COLYDIINAE: SYNCHITINI.

Type species: Langelandia anophthalma Aubé, 1843, by monotypy.

Family-group name: LANGELANDIINA Fowler, 1889. Stem: Langelandi-.

Languria Latreille, 1802b: 209. Current status: valid genus in EROTYLIDAE: LANGURIINAE: LANGURIINI.

Type species: Languria ruficollis Latreille, 1802 (= Trogosita bicolor Fabricius, 1798), by monotypy.

Family-group name: LANGUIRIDAE Hope, 1841. Stem: Languri-.

Laogenia Pascoe, 1874: 75. Current status: valid genus in CURCULIONIDAE: DRYOPHTHORINAE: LITOSOMINI: LAOGENINA.

Type species: Laogenia sorex Pascoe, 1874, by subsequent designation (Setliff 2007: 73).

Family-group name: LAOGENINA Legalov, 2020a: 324. Stem: Laogen-.

Laparocerus Schönherr, 1834b: 530. Current status: valid genus in CURCULIONIDAE: ENTIMINAE: OTIORHYNCHITAE: LAPAROCERINI.

Type species: Laparocerus morio Boheman, 1834, by original designation.

Family-group name: LAPAROCÉRIDES Lacordaire, 1863. Stem: Laparocer-.

Lapethus Casey, 1890b: 317, 321. Current status: junior synonym of Mychocerus Erichson, 1845 in CERYLONIDAE: CERYLONINAE.

Type species: Lapethus discretus Casey, 1890, by monotypy.

Family-group name: LAPETHINAE Sharp, 1894. Stem: Lapeth-.

Lara J.L. LeConte, 1852b: 42. Current status: valid genus in ELMIDAE: LARAINAE: LARAINI. Note: Lara used by Drapiez (1819: 54) is an incorrect subsequent spelling of Larra Fabricius, 1792 in Hymenoptera (see Jäch et al. 2016: 104) not in prevailing usage; name placed on the Official List of Generic Names in Zoology (ICZN 1988g).

Type species: Lara avara J.L. LeConte, 1852, by monotypy (see ICZN 1988g). Note: name placed on the Official List of Specific Names in Zoology (ICZN 1988g).

Family-group name: LARINI J.L. LeConte, 1861. Stem: Lara- (ICZN 1988g).

Laria Scopoli, 1763: 21. Current status: invalid senior synonym of Bruchus Linnaeus, 1767 in CHRYSOMELIDAE: BRUCHINAE: BRUCHINI: BRUCHINA. Note: name suppressed for the Principle of Priority and placed on the Official Index of Rejected and Invalid Generic Names in Zoology (ICZN 1995b).

Type species: Laria salicis Scopoli, 1763, by subsequent designation (Pierce 1916: 463).

Family-group name: LARIIDAE Bedel, 1901. Stem: Lari-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

Laricobius Rosenhauer, 1846: 5. Current status: valid genus in DERODONTIDAE: LARICOBIINAE.

Type species: Laricobius erichsonii Rosenhauer, 1846, by monotypy.

Family-group name: LARICOBIENS Mulsant & Rey, 1864. Stem: Laricobi-.

Larinotus Carter & Zeck, 1937: 182, 186. Current status: valid genus in TROGOSSITIDAE: LARINOTINAE.

Type species: Larinotus umbilicatus Carter & Zeck, 1937, by monotypy.

Family-group name: LARINOTINAE Ślipiński, 1992. Stem: Larinot-.

Larinus Dejean, 1821: 97. Current status: valid genus in CURCULIONIDAE: LIXINAE: LIXINI. Note: name first proposed as a junior synonym of Rhinobatus Germar, 1817 and subsequently used as valid before 1961 (e.g., Schönherr 1823: 1146; Germar 1823: 379), it is available and dates from its first publication as a synonym (ICZN 1999a, Article 11.6.1).

Type species: Curculio cynarae Fabricius, 1787, by subsequent designation (Schönherr 1823: 1146). Note: nomen protectum (see Gültekin 2013: 108).

Family-group name: LARINIDAE Gistel, 1848. Stem: Larin-. Note: this name was listed as permanently invalid by Bouchard et al. (2011: 621) on the incorrect belief that it was proposed for Larinus Germar, 1823, a junior homonym of Larinus Dejean, 1821; we believe that Germar’s (1823: 379) name is simply a subsequent use of Dejean’s name.

Lariversius Blaisdell, 1947: 59. Current status: valid genus in TENEBRIONIDAE: BLAPTINAE: AMPHIDORINI.

Type species: Lariversius tibialis Blaisdell, 1947, by original designation.

Family-group name: LARIVERSIINA La Rivers, 1948. Stem: Lariversi-.

Lasiocala Blanchard in H. Milne-Edwards et al., 1851: 220. Current status: valid genus in SCARABAEIDAE: RUTELINAE: RUTELINI: LASIOCALINA.

Type species: Lasiocala fulvohirta Blanchard, 1851, by monotypy.

Family-group name: LASIOCALINA Ohaus, 1918. Stem: Lasiocal-.

Lasiocera Dejean, 1831: 278, 283. Current status: valid genus in CARABIDAE: HARPALINAE: ODACANTHINI: ODACANTHINA.

Type species: Lasiocera nitidula Dejean, 1831, by monotypy.

Family-group name: LASIOCERINI Jeannel, 1948. Stem: Lasiocer-.

Lasioderma Stephens, 1835a: 417. Current status: valid genus in PTINIDAE: XYLETININAE: LASIODERMINI.

Type species: Lasioderma testaceum Stephens, 1835 (= Ptinus serricornis Fabricius, 1792), by monotypy.

Family-group name: LASIODERMINI Böving, 1927. Stem: Lasioderm-.

Lasiopus J.L. LeConte, 1856b: 282. Current status: invalid senior synonym of Podolasia Harold, 1869 in SCARABAEIDAE: PODOLASIINAE. Note: junior homonym of Lasiopus Schönherr, 1823 [Coleoptera: CURCULIONIDAE].

Type species: Lasiopus ferrugineus J.L. LeConte, 1856, by monotypy.

Family-group name: LASIOPODES J.L. LeConte, 1856. Stem: Lasiopod-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

Lasiorhynchites Jekel, 1860: 227. Current status: valid genus in ATTELABIDAE: RHYNCHITINAE: RHYNCHITITAE: RHYNCHITINI: LASIORHYNCHITINA.

Type species: here fixed (under ICZN 1999a, Article 70.3.2) as Rhynchites cavifrons Gyllenhal, 1833, misidentified as Curculio pubescens Fabricius, 1775 (sensu Herbst, 1797) by original designation in Jekel (1860: 227).

Family-group name: LASIORHYNCHITINA Legalov, 2003. Stem: Lasiorhynchit-.

†Lasiosyne Tan, Ren & Shih, 2007: 237. Current status: valid genus in †LASIOSYNIDAE. Note: Middle/Upper Jurassic and Lower Cretaceous (Russia, Mongolia, China).

Type species: †Lasiosyne euglyphea Tan, Ren & Shih, 2007, by original designation.

Family-group name: †LASIOSYNIDAE Kirejtshuk, Chang, Ren & Kun, 2010. Stem: Lasiosyn-.

Lathrobium Gravenhorst, 1802: 51. Current status: valid genus in STAPHYLINIDAE: PAEDERINAE: LATHROBIINI: LATHROBIINA.

Type species: Staphylinus elongatus Linnaeus, 1767, by subsequent designation (Latreille 1810: 427).

Family-group name: LATHROBIDAE Laporte, 1835. Stem: Lathrobi-.

Lathropus Erichson, 1845a: 307. Current status: valid genus in LAEMOPHLOEIDAE.

Type species: Trogossita sepicola P.W.J. Müller, 1821, by subsequent monotypy (Erichson 1846: 328).

Family-group name: LATHROPINI Casey, 1916: 117. Stem: Lathrop-. Note: this name was overlooked in Bouchard et al. (2011) and Bouchard and Bousquet (2020a).

Laticranium Lane, 1959: 312. Current status: valid genus in CERAMBYCIDAE: LAMIINAE: LATICRANIINI.

Type species: Laticranium mandibulare Lane, 1959, by original designation.

Family-group name: LATICRANIINAE Lane, 1959. Stem: Laticrani-.

Latometus Erichson, 1842: 213. Current status: valid genus in ZOPHERIDAE: ZOPHERINAE: LATOMETINI.

Type species: Latometus pubescens Erichson, 1842, by monotypy.

Family-group name: LATOMETINI Ślipiński & Lawrence, 1999. Stem: Latomet-.

Latridius Herbst, 1793: 3. Current status: valid genus in LATRIDIIDAE: LATRIDIINAE. Note: name placed on the Official List of Generic Names in Zoology (ICZN 2011e).

Type species: Latridius porcatus Herbst, 1793, by subsequent designation (Latreille 1810: 431; see ICZN 2011e).

Family-group name: LATHRIDIEN Erichson, 1842. Stem: Latridi-.

Lawrencerosus Kirejtshuk, 1991: 866. Current status: valid genus in NITIDULIDAE: NITIDULINAE: LAWRENCEROSINI.

Type species: Lawrencerosus mirator Kirejtshuk, 1991, by original designation.

Family-group name: LAWRENCEROSINI Kirejtshuk, 1991. Stem: Lawrenceros-.

†Lebanophytum Kirejtshuk & Azar, 2008: 31. Current status: valid genus in CRYPTOPHAGIDAE. Note: Lower Cretaceous (Lebanon).

Type species: †Lebanophytum excellens Kirejtshuk & Azar, 2008, by original designation.

Family-group name: †LEBANOPHYTIDAE Kirejtshuk in Kirejtshuk and Azar, 2013: 114. Stem: Lebanophyt-.

Lebia Latreille, 1802b: 85. Current status: valid genus in CARABIDAE: HARPALINAE: LEBIINI: LEBIINA.

Type species [suggested]: Carabus haemorrhoidalis Fabricius, 1787 (= Buprestis marginatus Geoffroy, 1785), by subsequent designation (E. Desmarest 1851: 76). Note: the valid type species fixation is that of Latreille (1810: 426), who selected Carabus quadrimaculatus Linnaeus, 1758, but that species is the type species of Dromius Bonelli, 1810; a subsequent designation is that of Drapiez (1839: 303), who selected Carabus cyanocephalus Linnaeus, 1758, but that species is currently included in the subgenus Lamprias Bonelli, 1810, not in the nominotypical subgenus; a new application to the Commission is necessary to fix Carabus haemorrhoidalis Fabricius, 1787 as the type species of Lebia Latreille, 1802 since a previous request was postponed (ICZN 1950: 447).

Family-group name: LEBIOTAE Bonelli, 1810. Stem: Lebi-.

Lebidia Morawitz, 1862: 322. Current status: valid genus in CARABIDAE: HARPALINAE: LEBIINI: GALLERUCIDIINA.

Type species: Lebidia octoguttata Morawitz, 1862, by monotypy.

Family-group name: LEBIDIINA Jakobson, 1907. Stem: Lebidi-.

Lechriops Schönherr, 1825: 586. Current status: valid genus in CURCULIONIDAE: CONODERINAE: LECHRIOPINI.

Type species: Rhynchaenus sciurus Fabricius, 1801, by original designation.

Family-group name: LÉCHRIOPIDES Lacordaire, 1865. Stem: Lechriop-.

†Leehermania Chatzimanolis, Grimaldi & Engel in Chatzimanolis et al., 2012: 6. Current status: valid genus in HYDROSCAPHIDAE: †LEEHERMANIINAE. Note: Upper Triassic (USA).

Type species: †Leehermania prorova Chatzimanolis et al., 2012, by original designation.

Family-group name: †LEEHERMANIINAE Kirejtshuk, Prokin & Ponomarenko, 2023b: 7. Stem: Leehermani-.

Leichenum Dejean, 1834: 194. Current status: valid genus in TENEBRIONIDAE: BLAPTINAE: PEDININI: LEICHENINA.

Type species: Opatrum pictum Fabricius, 1801, by monotypy.

Family-group name: LEICHENAIRES Mulsant, 1854. Stem: Leichen-.

Leiestes Chevrolat in Dejean, 1836a: 440. Current status: valid genus in ENDOMYCHIDAE: LEIESTINAE.

Type species: Cryptophagus seminiger Gyllenhal, 1808, by monotypy.

Family-group name: LEIESTINA C.G. Thomson, 1863. Stem: Leiest-.

Leiochrinus Westwood, 1883: 68. Current status: valid genus in TENEBRIONIDAE: DIAPERINAE: LEIOCHRININI.

Type species: Leiochrinus fulvicollis Westwood, 1883, by subsequent designation (K.W. Kirby 1885: 89).

Family-group name: LEIOCHRININAE Lewis, 1894. Stem: Leiochrin-.

Leiodes Latreille, 1797: 22. Current status: valid genus in LEIODIDAE: LEIODINAE: LEIODINI.

Type species: Sphaeridium ferrugineum Fabricius, 1787, by subsequent monotypy (Latreille 1802b: 163).

Family-group name: LEIODESIDAE Fleming, 1821. Stem: Leiod-.

Leiopleura Deyrolle, 1865: 219. Current status: valid genus in BUPRESTIDAE: AGRILINAE: TRACHEINI: LEIOPLEURINA.

Type species: Brachys concinna Gory, 1841, by original designation.

Family-group name: LEIOPLEURINA Holyński, 1993. Stem: Leiopleur-.

Leiopus Audinet-Serville, 1835a: 86. Current status: valid genus in CERAMBYCIDAE: LAMIINAE: ACANTHOCININI.

Type species: Cerambyx nebulosus Linnaeus, 1758, by subsequent designation (Westwood 1838c: 41).

Family-group name: LIOPI J.L. LeConte, 1873. Stem: Leiopod-. Note: senior homonym of LEIOPIDAE [should be LEIOPODIDAE] Lang, 1970 (type genus Leiopus Beddard, 1886) in Crustacea; Lang’s name is permanently invalid (based on a preoccupied type genus).

Leiosoma Stephens, 1829b: 14. Current status: valid genus in CURCULIONIDAE: MOLYTINAE: MOLYTINI: LEIOSOMATINA.

Type species: Curculio punctatus Marsham, 1802 (= Curculio deflexus Panzer, 1795), by monotypy.

Family-group name: LIOSOMINA Reitter, 1913. Stem: Leiosomat-.

Leleupaussus Luna de Carvalho, 1962: 66, 68. Current status: valid genus in CARABIDAE: PAUSSINAE: PAUSSINI: PAUSSINA.

Type species: Leleupaussus tetramerus Luna de Carvalho, 1962, by original designation.

Family-group name: LELEUPAUSSINA Luna de Carvalho, 1989. Stem: Leleupauss-.

Leleupidia Basilewsky, 1951: 176. Current status: valid genus in CARABIDAE: HARPALINAE: ZUPHIINI: LELEUPIDIINA.

Type species: Leleupidia luvubuana Basilewsky, 1951, by original designation.

Family-group name: LELEUPIDIINI Basilewsky, 1951. Stem: Leleupidi-.

Lema Fabricius, 1798: 4. Current status: valid genus in CHRYSOMELIDAE: CRIOCERINAE: LEMINI. Note: name placed on the Official List of Generic Names in Zoology (ICZN 1970a).

Type species: Lema cyanea Fabricius, 1798, by plenary power (see ICZN 1970a). Note: name placed on the Official List of Specific Names in Zoology (ICZN 1970a).

Family-group name: LEMOIDEAE Gyllenhal, 1813. Stem: Lem-.

Lemidia Spinola, 1841: 75. Current status: valid genus in CLERIDAE: CLERINAE: CLERITAE: HYDNOCERINI: LEMIDIINA.

Type species: Hydnocera nitens Newman, 1841, by subsequent monotypy (Erichson 1843c: 179).

Family-group name: LEMIDIINI Kolibáč, 1998. Stem: Lemidi-.

Lemodes Boheman, 1858: 103. Current status: valid genus in ANTHICIDAE: LEMODINAE.

Type species: Lemodes coccinea Boheman, 1858, by monotypy.

Family-group name: LEMODINAE E.G. Matthews, 1987. Stem: Lemod-.

Lemphus Erichson, 1840b: 131. Current status: valid genus in MELYRIDAE: MALACHIINAE: LEMPHINI.

Type species: Lemphus mancus Erichson, 1840, by monotypy.

Family-group name: LEMPHINI Wittmer, 1976. Stem: Lemph-.

Lenax Sharp, 1877b: 265, 269. Current status: valid genus in MONOTOMIDAE: MONOTOMINAE: LENACINI.

Type species: Lenax mirandus Sharp, 1877, by monotypy.

Family-group name: LENACINAE Crowson, 1952. Stem: Lenac-.

Lendomus Casey, 1924: 185. Current status: junior synonym of Agyrtecanus Reitter, 1901 in AGYRTIDAE: AGYRTINAE.

Type species: Lendomus politus Casey, 1924 (= Necrophilus longulus J.L. LeConte, 1859), by original designation.

Family-group name: LENDOMINI Bradley, 1930. Stem: Lendom-.

Leoglymmius R.T. Bell & J.R. Bell, 1978: 53. Current status: valid genus in CARABIDAE: RHYSODINAE: LEOGLYMMIINI.

Type species: Rhysodes lignarius Olliff, 1886, by original designation.

Family-group name: LEOGLYMMIINA R.T. Bell & J.R. Bell, 1978. Stem: Leoglymmi-.

Lepanomus J. Balfour-Browne, 1945: 59. Current status: valid genus in BRENTIDAE: APIONINAE incertae sedis.

Type species: Lepanomus crinalis J. Balfour-Browne, 1945, by original designation.

Family-group name: LEPANOMINI Wanat, 2021: 56. Stem: Lepanom-.

Leperina Erichson, 1844: 453. Current status: valid genus in TROGOSSITIDAE: TROGOSSITINAE: TROGOSSITINI.

Type species: Trogossita decorata Erichson, 1842, by subsequent designation (Kolibáč 2005: 65).

Family-group name: LEPERINI Reitter, 1876. Stem: Leperin-.

Lepicerus Motschulsky, 1855a: 14. Current status: valid genus in LEPICERIDAE.

Type species: Lepicerus inaequalis Motschulsky, 1855, by monotypy.

Family-group name: LEPICERIDAE Hinton, 1936. Stem: Lepicer-.

Lepisia Lepeletier & Audinet-Serville, 1828: 374. Current status: valid genus in SCARABAEIDAE: MELOLONTHINAE: HOPLIINI: HOPLIINA.

Type species: Melolontha rupicola Fabricius, 1775, by subsequent designation (Blanchard in Audouin et al. 1843: pl. 43).

Family-group name: LEPISIIDAE Burmeister, 1844. Stem: Lepisi-.

Lepithrix Lepeletier & Audinet-Serville, 1828: 381. Current status: valid genus in SCARABAEIDAE: MELOLONTHINAE: HOPLIINI: LEPITHRICHINA. Note: although originally established as Lepitrix, the spelling Lepithrix, an unjustified emendation by von Dalla Torre (1913: 342), is in prevailing usage, attributed to the original publication (see Cupello and Ribeiro-Costa 2019: 140, 141), and therefore deemed to be a justified emendation and the correct original spelling (ICZN 1999a, Article 33.2.3.1).

Type species: Trichius lineatus Fabricius, 1775, by subsequent designation (Blanchard in Audouin et al. 1843: pl. 44).

Family-group name: LEPITRICHINA Perty, 1840. Stem: Lepithrich-.

Leprotes Baly, 1863: 148, 158. Current status: junior synonym of Fidia Motschulsky, 1860 in CHRYSOMELIDAE: EUMOLPINAE: BROMIINI.

Type species: Endoxus gracilicornis Baly, 1861, by original designation.

Family-group name: LEPROTITES Chapuis, 1874. Stem: Leprotet-.

Leptanillophilus Holmgren, 1908: 340. Current status: valid genus in STAPHYLINIDAE: ALEOCHARINAE: MIMECITINI: LEPTANILLOPHILINA.

Type species: Leptanillophilus similis Holmgren, 1908, by monotypy.

Family-group name: LEPTANILLOPHILINI Fenyes, 1919. Stem: Leptanillophil-.

Leptaulax Kaup, 1868a: 11. Current status: valid genus in PASSALIDAE: PASSALINAE: LEPTAULACINI.

Type species: Passalus dentatus Fabricius, 1792, by subsequent designation (Gravely 1918: 112).

Family-group name: LEPTAULACEAE Kaup, 1871. Stem: Leptaulac-.

Leptidea Mulsant, 1839: 105. Current status: invalid senior synonym of Nathrius Brèthes, 1916 in CERAMBYCIDAE: CERAMBYCINAE: PSEBIINI. Note: junior homonym of Leptidea Billberg, 1820 [Lepidoptera].

Type species: Leptidea brevipennis Mulsant, 1839, by monotypy.

Family-group name: LEPTIDEINAE Perrier, 1893. Stem: Leptide-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

Leptinopterus Hope, 1838b: 316. Current status: valid genus in LUCANIDAE: LUCANINAE: LUCANINI. Note: the original spelling was Leptynopterus but Leptinopterus, an unjustified emendation by Hope (1845b: 5), is in prevailing usage, attributed to the original author and date, and therefore deemed to be a justified emendation (ICZN 1999a, Article 33.2.3.1); Psalicerus Dejean, 1833 is a senior objective synonym of this genus (see Bousquet and Bouchard 2013a: 40) and an application to the Commission or Reversal of Precedence (ICZN 1999a, Article 23.9) is needed to conserve Leptinopterus Hope as a valid name.

Type species: Lucanus femoratus G.-A. Olivier, 1789, by subsequent designation (R. Lucas 1920: 369).

Family-group name: LEPTINOPTERINI Jakobson, 1911. Stem: Leptinopter-.

Leptinus P.W.J. Müller, 1817: 266. Current status: valid genus in LEIODIDAE: PLATYPSYLLINAE.

Type species: Leptinus testaceus P.W.J. Müller, 1817, by monotypy.

Family-group name: LEPTINIDAE J.L. LeConte, 1872. Stem: Leptin-.

Leptispa Baly, 1858: 1. Current status: valid genus in CHRYSOMELIDAE: CASSIDINAE: LEPTISPINI. Note: new replacement name for Leptomorpha Germar, 1842.

Type species: Leptomorpha filiformis Germar, 1842, by monotypy (type fixation for Leptomorpha Germar).

Family-group name: LEPTISPITES Fairmaire, 1868. Stem: Leptisp-.

Leptobium Casey, 1905: 57. Current status: valid genus in STAPHYLINIDAE: PAEDERINAE: PAEDERINI: DOLICAONINA.

Type species: Lathrobium biguttulum Lacordaire, 1835 (= Lathrobium gracile Gravenhorst, 1802), by monotypy.

Family-group name: LEPTOBII Bordoni, 1980. Stem: Leptobi-.

Leptochirus Germar, 1823: 35. Current status: valid genus in STAPHYLINIDAE: OSORIINAE: LEPTOCHIRINI.

Type species: Leptochirus scoriaceus Germar, 1823, by monotypy.

Family-group name: LEPTOCHIRINA Sharp, 1887. Stem: Leptochir-.

Leptochromus Motschulsky, 1855a: 12. Current status: valid genus in STAPHYLINIDAE: SCYDMAENINAE: MASTIGITAE: LEPTOCHROMINI.

Type species: Leptochromus fulvescens Motschulsky, 1855, by monotypy.

Family-group name: LEPTOCHROMINI Jałoszyński & Brunke in Jałoszyński et al., 2018: 635. Stem: Leptochrom-.

Leptodes Dejean, 1834: 181. Current status: valid genus in TENEBRIONIDAE: PIMELIINAE: LEPTODINI.

Type species: Sepidium boisduvalii Zoubkoff, 1833, by monotypy.

Family-group name: LEPTODIDES Lacordaire, 1859. Stem: Leptod-. Note: senior homonym of LEPTODINAE Schuchert, 1913 (type genus Leptodus Kayser, 1883) in Brachiopoda.

Leptodinodes Jeannel, 1949b: 786. Current status: valid genus in CARABIDAE: HARPALINAE: CHLAENIINI: CHLAENIINA.

Type species: Chlaenius parallelus Dejean, 1831, by original designation.

Family-group name: LEPTODINODINI Basilewsky & Grundmann, 1955. Stem: Leptodinod-.

Leptodirus F.J. Schmidt, 1832a: 9. Current status: valid genus in LEIODIDAE: CHOLEVINAE: LEPTODIRINI: LEPTODIRINA. Note: the name was also proposed by F.J. Schmidt subsequently the same year (1832b: 83).

Type species: Leptodirus hochenwartii F.J. Schmidt, 1832, by monotypy.

Family-group name: LEPTODÉRIDES Lacordaire, 1854. Stem: Leptodir-.

Leptohoplia Saylor, 1935: 132. Current status: valid genus in SCARABAEIDAE: RUTELINAE: ANOMALINI: LEPTOHOPLIINA.

Type species: Leptohoplia testaceipennis Saylor, 1935, by original designation.

Family-group name: LEPOTHOPLIINI Potts, 1974. Stem: Leptohopli-.

Leptolycus Leng & Mutchler, 1922: 430. Current status: valid genus in LYCIDAE: LYCINAE: LEPTOLYCINI: LEPTOLYCINA.

Type species: Leptolycus heterocornis Leng & Mutchler, 1922, by monotypy.

Family-group name: LEPTOLYCINI Leng & Mutchler, 1922. Stem: Leptolyc-.

Leptomastax Pirazzoli, 1855: 3. Current status: valid genus in STAPHYLINIDAE: SCYDMAENINAE: MASTIGITAE: LEPTOMASTACINI.

Type species: Leptomastax hypogeum Pirazzoli, 1855, by monotypy.

Family-group name: LEPTOMASTACINI Casey, 1897. Stem: Leptomastac-.

Leptonota J. Thomson, 1861: 353. Current status: valid genus in CERAMBYCIDAE: LAMIINAE: ENICODINI.

Type species: Enicodes comitessa A. White, 1855, by monotypy.

Family-group name: LEPTONOTITAE J. Thomson, 1864. Stem: Leptonot-.

Leptopalpus Guérin-Méneville, 1834a: pl. 35. Current status: valid genus in MELOIDAE: NEMOGNATHINAE: NEMOGNATHINI: ZONITIDINA.

Type species: Leptopalpus chevrolatii Guérin-Méneville, 1834 (= Zonitis rostrata Fabricius, 1792), by monotypy.

Family-group name: LEPTOPALPINA Kaszab, 1963. Stem: Leptopalp-.

Leptopius Oke, 1951: 24. Current status: valid genus in CURCULIONIDAE: ENTIMINAE: ENTIMITAE: EUDIAGOGINI. Note: new replacement name for Leptops Schönherr, 1834.

Type species: Curculio robustus G.-A. Olivier, 1807, by original designation (type fixation for Leptops Schönherr).

Family-group name: LEPTOPIINAE Oke, 1951. Stem: Leptopi-.

†Leptopodocoleus Hong, 1982: 119. Current status: valid genus in TRACHYPACHIDAE: †EODROMEINAE. Note: Lower Cretaceous (China).

Type species: †Leptopodocoleus sarmentosus Hong, 1982, by original designation.

Family-group name: †LEPTOPODOCOLEIDAE Hong, 1982. Stem: Leptopodocole-.

Leptops Schönherr, 1834a: 297. Current status: invalid senior synonym of Leptopius Oke, 1951 in CURCULIONIDAE: ENTIMINAE: ENTIMITAE: EUDIAGOGINI. Note: junior homonym of Leptops Rafinesque, 1820 [Pisces].

Type species: Curculio robustus G.-A. Olivier, 1807, by original designation.

Family-group name: LEPTOPSIDES Lacordaire, 1863. Stem: Leptop-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

Leptorhynchus Boisduval, 1835: 318. Current status: invalid senior synonym of Ithystenus Pascoe, 1862 in BRENTIDAE: BRENTINAE: BRENTINI: ITHYSTENINA. Note: junior homonym of Leptorhynchus Clift, 1828 [Reptilia].

Type species: Brenthus angustatus Guérin-Méneville, 1831, by monotypy.

Family-group name: LEPTORRHYNCHIDAE Schoenfeldt, 1908. Stem: Leptorhynch-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

Leptoschoinus Dejean, 1835: 291. Current status: valid genus in CURCULIONIDAE: BARIDINAE: MADARINI: MADARINA.

Type species: Baris fucata Klug, 1829, by monotypy.

Family-group name: LEPTOSCHOÏNIDES Lacordaire, 1865. Stem: Leptoschoin-.

Leptoscydmus Casey, 1897: 518. Current status: valid genus in STAPHYLINIDAE: SCYDMAENINAE: SCYDMAENITAE: LEPTOSCYDMINI.

Type species: Leptoscydmus cavifrons Casey, 1897, by subsequent designation (H. Franz 1985: 177).

Family-group name: LEPTOSCYDMINI Casey, 1897: 518. Stem: Leptoscydm-.

Leptosonyx Weise, 1885b: 315. Current status: junior synonym of Theone Gistel, 1857 in CHRYSOMELIDAE: GALERUCINAE: GALERUCINI.

Type species: Galleruca silphoides Dalman, 1823, by original designation.

Family-group name: LEPTOSONYCHINI Weise, 1924. Stem: Leptosonych-.

Leptostethus G.R. Waterhouse, 1853: 175. Current status: valid genus in CURCULIONIDAE: ENTIMINAE: ENTIMITAE: LEPTOSTETHINI.

Type species: Leptostethus marginatus G.R. Waterhouse, 1853, by original designation.

Family-group name: LEPTOSTÉTHIDES Lacordaire, 1863. Stem: Leptosteth-.

Leptosus G.A.K. Marshall, 1952: 265. Current status: junior synonym of Leptopius Oke, 1951 in CURCULIONIDAE: ENTIMINAE: ENTIMITAE: BYRSOPAGINI: STRANGALIODINA. Note: unnecessary new replacement name for Leptops Schönherr, 1833.

Type species: Curculio robustus G.-A. Olivier, 1807, by original designation (type fixation for Leptops Schönherr).

Family-group name: LEPTOSINAE G.A.K. Marshall, 1952. Stem: Leptos-.

Leptotyphlus Fauvel, 1874: [2^e supplément] 36. Current status: valid genus in STAPHYLINIDAE: LEPTOTYPHLINAE: LEPTOTYPHLINI.

Type species: Leptotyphlus sublaevis Fauvel, 1874, by monotypy.

Family-group name: LEPTOTYPHLI Fauvel, 1874. Stem: Leptotyphl-.

Leptura Linnaeus, 1758: 342, 397. Current status: valid genus in CERAMBYCIDAE: LEPTURINAE: LEPTURINI.

Type species [suggested]: Leptura quadrifasciata Linnaeus, 1758, by subsequent designation (Westwood 1838c: 41). Note: the valid type species fixation is that of Partington (1836b: 45), who selected Leptura melanura Linnaeus, 1758, but that species is the type species of the genus Stenurella Villiers, 1974; an application to the Commission is necessary to fix Leptura quadrifasciata Linnaeus, 1758 as the type species of Leptura Linnaeus, 1758.

Family-group name: LEPTURETAE Latreille, 1802. Stem: Leptur-. Note: senior homonym of LEPTURINAE Gill, 1860 (type genus Lepturus Günther, 1854) in Pisces.

Lepturoides Herbst, 1784: 103. Current status: junior synonym of Denticollis Piller & Mitterpacher, 1783 in ELATERIDAE: DENDROMETRINAE: DENDROMETRINI: DENTICOLLINA.

Type species: Lepturoides linearis Herbst, 1784 (= Elater linearis Linnaeus, 1758), by monotypy.

Family-group name: LEPTUROIDINI O. Schwarz, 1906. Stem: Lepturoid-.

Leptusa Kraatz, 1856: 60. Current status: valid genus in STAPHYLINIDAE: ALEOCHARINAE: HOMALOTINI: BOLITOCHARINA. Note: name placed on the Official List of Generic Names in Zoology (ICZN 2005d).

Type species: Bolitochara pulchella Mannerheim, 1830, by subsequent designation (Gusarov and Herman 2003: 115; see ICZN 2005d). Note: name placed on the Official List of Specific Names in Zoology (ICZN 2005d).

Family-group name: LEPTUSAE Fenyes, 1919. Stem: Leptus-.

Lepyrus Germar, 1817b: 340. Current status: valid genus in CURCULIONIDAE: MOLYTINAE: LEPYRINI.

Type species [suggested]: Curculio colon Linnaeus, 1771 (= Curculio palustris Scopoli, 1763), by subsequent designation (Schönherr 1823: 1143). Note: the valid type species fixation is that of Samouelle (1819: 204), who selected Curculio triguttatus Fabricius, 1775, but that species is the type species of the genus Graptus Schönherr, 1823; Alonso-Zarazaga and Lyal (1999: 204) mentioned that they would submit an application to the Commission to fix Curculio colon Linnaeus, 1771 as the type species of this type genus but this has not taken place; an application to the Commission is still necessary to fix this problem.

Family-group name: LEPYRIDAE W. Kirby, 1837. Stem: Lepyr-.

Lesteva Latreille, 1797: 75. Current status: valid genus in STAPHYLINIDAE: OMALIINAE: ANTHOPHAGINI. Note: name placed on the Official List of Generic Names in Zoology (ICZN 2004e).

Type species: Lesteva punctulata Latreille, 1804 (= Staphylinus longoelytratus Goeze, 1777), by plenary power (see ICZN 2004e). Note: Lesteva punctulata Latreille, 1804 was placed on the Official List of Specific Names in Zoology (ICZN 2004e); Goeze’s (1777) work is not consistently binominal (see ICZN 2021a) and therefore an application to the Commission is necessary to retain Staphylinus longoelytratus Goeze, 1777 as the valid name for the type species.

Family-group name: LESTEVINA Jakobson, 1908. Stem: Lestev-.

Lestignathus Erichson, 1842: 132. Current status: valid genus in CARABIDAE: HARPALINAE: LICININI: LESTIGNATHINA.

Type species: Lestignathus cursor Erichson, 1842, by monotypy.

Family-group name: LESTIGNATHINA Ball, 1992. Stem: Lestignath-.

Lethrus Scopoli, 1777: 439. Current status: valid genus in GEOTRUPIDAE: GEOTRUPINAE: LETHRINI.

Type species: here fixed (under ICZN 1999a, Article 70.3.2) as Lucanus apterus Laxmann, 1770, misidentified as Scarabaeus cephalotes Pallas, 1771 (sensu Acharius, 1781), by subsequent monotypy in Fabricius (1787: 2). Note: the nominal species Scarabaeus cephalotes Pallas, 1771 is currently placed in the subgenus Ceratodirus Fischer von Waldheim, 1845.

Family-group name: LETHREN Oken, 1843. Stem: Lethr-.

Leucastea Stål, 1855: 344. Current status: valid genus in MEGALOPODIDAE: MEGALOPODINAE: LEUCASTEINI. Note: the original spelling was Lucastea but Leucastea, an incorrect subsequent spelling, is in prevailing usage, attributed to the publication of the original spelling, and therefore deemed to be the correct original spelling (ICZN 1999a, Article 33.3.1).

Type species: Leucastea dohrni Stål, 1855, by subsequent designation (Jacoby and Clavareau 1905: 15).

Family-group name: LEUCASTEINI Rodríguez-Mirón in Rodríguez-Mirón et al., 2021: 694. Stem: Leucaste-.

Leucocelis Burmeister, 1842: 421. Current status: valid genus in SCARABAEIDAE: CETONIINAE: CETONIINI: LEUCOCELINA.

Type species: Cetonia haemorrhoidalis Fabricius, 1775, by subsequent designation (Arrow 1910: 175).

Family-group name: LEUCOCELIDEN Kraatz, 1882. Stem: Leucocel-.

Leucocraspedum Kraatz, 1859: 51. Current status: valid genus in STAPHYLINIDAE: ALEOCHARINAE: LEUCOCRASPEDINI.

Type species: Leucocraspedum pulchellum Kraatz, 1859, by monotypy.

Family-group name: LEUCOCRASPEDINI Fenyes, 1921. Stem: Leucocrasped-.

Leucolaephus P.H. Lucas, 1859: xxii. Current status: valid genus in TENEBRIONIDAE: PIMELIINAE: PIMELIINI: PIMELIINA. Note: the original spelling was Leucoloephus but Leucolaephus, an incorrect subsequent spelling, is in prevailing usage, attributed to the publication of the original spelling, and therefore deemed to be the correct original spelling (ICZN 1999a, Article 33.3.1).

Type species: Leucolaephus perrisii P.H. Lucas, 1859 (= Pimelia liliputana P.H. Lucas, 1857), by subsequent designation (Gebien 1937: 803).

Family-group name: LEUCOLAEPHUSINI Pierre, 1961. Stem: Leucolaeph-.

Leucopholis Dejean, 1833b: 160. Current status: valid genus in SCARABAEIDAE: MELOLONTHINAE: LEUCOPHOLINI.

Type species: Melolontha rorida Fabricius, 1801, by subsequent designation (J. Bezděk 2006: 33).

Family-group name: LEUCOPHOLIDAE Burmeister, 1855. Stem: Leucophol-.

Leucothyreus W.S. MacLeay, 1819: 145. Current status: valid genus in SCARABAEIDAE: RUTELINAE: GENIATINI.

Type species: Leucothyreus kirbyanus W.S. MacLeay, 1819, by monotypy.

Family-group name: LEUCOTHYREIDAE Burmeister, 1844. Stem: Leucothyre-.

Leupelia Jeannel, 1954b: 106. Current status: valid genus in STAPHYLINIDAE: PSELAPHINAE: BATRISITAE: BATRISINI: LEUPELIINA.

Type species: Leupelia longicornis Jeannel, 1954, by original designation.

Family-group name: LEUPELIINA Jeannel, 1954. Stem: Leupeli-.

Lexiphanes Gistel, 1848b: 123. Current status: valid genus in CHRYSOMELIDAE: CRYPTOCEPHALINAE: CRYPTOCEPHALINI: MONACHULINA. Note: new replacement name for Monachus Chevrolat, 1836.

Type species: Cryptocephalus saponatus Fabricius, 1801, by monotypy (type fixation for Monachus Chevrolat).

Family-group name: LEXIPHANINI Wilcox, 1954. Stem: Lexiphan-.

†Liadytes Ponomarenko, 1963: 129. Current status: valid genus in †LIADYTIDAE. Note: Jurassic (Asia).

Type species: †Liadytes avus Ponomarenko, 1963, by original designation.

Family-group name: †LIADYTIDAE Ponomarenko, 1977. Stem: Liadyt-.

†Liadytiscus Prokin & Ren, 2010: 48. Current status: valid genus in DYTISCIDAE: †LIADYTISCINAE: †LIADYTISCINI. Note: Lower Cretaceous (China).

Type species: †Liadytiscus cretaceus Prokin & Ren, 2010, by original designation.

Family-group name: †LIADYTISCINAE Prokin & Ren, 2010. Stem: Liadytisc-.

†Liaoximordella W.I. Wang, 1993: 87. Current status: valid genus in †LIAOXIMORDELLIDAE. Note: Lower Cretaceous (China).

Type species: †Liaoximordella hongi W.I. Wang, 1993, by original designation.

Family-group name: †LIAOXIMORDELLIDAE W.I. Wang, 1993. Stem: Liaoximordell-.

Liatongus Reitter, 1892: 167. Current status: valid genus in SCARABAEIDAE: SCARABAEINAE: ONITICELLINI: LIATONGINA. Note: this name is a junior synonym of Onthosphaenus Motschulsky, 1860 but Liatongus is well established and should be maintained until a case is submitted to the Commission (see J. Bezděk 2016: 13).

Type species: Onthophagus phanaeoides Westwood, 1839, by subsequent designation (Arrow 1931: 362).

Family-group name: LIATONGINA Philips, 2016: 41. Stem: Liatong-.

†Libanopsis Kirejtshuk in Kirejtshuk et al., 2015: 466. Current status: valid genus in SPHINDIDAE: †LIBANOPSINAE. Note: Lower Cretaceous (Lebanon).

Type species: †Libanopsis poinari Kirejtshuk, 2015, by original designation.

Family-group name: †LIBANOPSINAE Kirejtshuk in Kirejtshuk et al., 2015: 462. Stem: Libanops-.

Libnetis C.O. Waterhouse, 1878: 104. Current status: valid genus in LYCIDAE: METRIORRHYNCHINAE: LIBNETINI. Note: the original spelling was Libnetus but Libnetis, an unjustified emendation by Bourgeois (1883: 647), is in prevailing usage, attributed to the original author and date, and therefore deemed to be a justified emendation (ICZN 1999a, Article 33.2.3.1).

Type species: Libnetis pumilio C.O. Waterhouse, 1878, by original designation.

Family-group name: LIBNETININA Bocák & Bocáková, 1990. Stem: Libnet-.

Lichas Westwood, 1853: 237. Current status: invalid senior synonym of Eulichas Jakobson, 1913 in EULICHADIDAE. Note: junior homonym of Lichas Dalman, 1827 [Trilobita].

Type species: Lichas funebris Westwood, 1853, by monotypy.

Family-group name: LICHADIDAE Csiki, 1902. Stem: Lichad-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

Lichenophanes Lesne, 1899: 443, 457. Current status: valid genus in BOSTRICHIDAE: BOSTRICHINAE: BOSTRICHINI.

Type species: Bostrichus tristis Fåhraeus, 1871, by subsequent designation (Chûjô 1937: 44).

Family-group name: LICHENOPHANINI Portevin, 1931. Stem: Lichenophan-.

Lichnasthenus J. Thomson, 1858b: 35. Current status: valid genus in CARABIDAE: HARPALINAE: LEBIINI: DROMIUSINA. Note: Thomson originally used the spelling Lichnastenus (p. 35, 37, pl. 1), which he intentionnally changed to Lichnasthenus in the errata (p. [471]).

Type species: Lichnasthenus armiventris J. Thomson, 1858, by monotypy.

Family-group name: LICHNASTHENITAE J. Thomson, 1858. Stem: Lichnasthen-.

Lichnia Erichson, 1835: 269. Current status: valid genus in SCARABAEIDAE: MELOLONTHINAE: LICHNIINI.

Type species: Lichnia limbata Erichson, 1835, by monotypy.

Family-group name: LICHNIADAE Burmeister, 1844. Stem: Lichni-.

Licinus Latreille, 1802b: 92. Current status: valid genus in CARABIDAE: HARPALINAE: LICININI: LICININA.

Type species: Carabus cassideus Fabricius, 1792, by subsequent designation (Latreille 1810: 426).

Family-group name: LICINII Bonelli, 1810. Stem: Licin-. Note: senior homonym of LICININA Edward, 1857 (type genus Licina J.E. Gray, 1850) in Mollusca.

†Lidryops Kirejtshuk & Nel, 2016b: 142. Current status: valid genus in BYRRHIDAE: †LIDRYOPINAE. Note: Lower Cretaceous (France).

Type species: †Lidryops occultus Kirejtshuk & Nel, 2016, by original designation.

Family-group name: †LIDRYOPINAE Kirejtshuk & Nel, 2016b: 141. Stem: Lidryop-.

Ligniperda Pallas, 1772: 6. Current status: invalid senior synonym of Apate Fabricius, 1775 (nomen protectum) in BOSTRICHIDAE: BOSTRICHINAE: APATINI. Note: nomen oblitum (see Borowski and Węgrzynowicz 2009: 190).

Type species: Ligniperda terebrans Pallas, 1772, by subsequent designation (Borowski and Węgrzynowicz 2007: 19, 155).

Family-group name: LIGNIPERDIDAE Jacobi, 1906. Stem: Ligniperd-.

Lignyodes Dejean, 1835: 278. Current status: valid genus in CURCULIONIDAE: CURCULIONINAE: TYCHIINI: LIGNYODINA.

Type species: Curculio enucleator Panzer, 1798, by monotypy.

Family-group name: LIGNYODINI Bedel, 1883. Stem: Lignyod-.

Limnebius Leach, 1815a: 96. Current status: valid genus in HYDRAENIDAE: HYDRAENINAE: LIMNEBIINI.

Type species: Hydrophilus picinus Marsham, 1802 (= Hydrophilus nitidus Marsham, 1802), by monotypy.

Family-group name: LIMNÉBIAIRES Mulsant, 1844. Stem: Limnebi-.

Limnichopharus Miyatake, 1994: 269. Current status: valid genus in COCCINELLIDAE: COCCINELLINAE: LIMNICHOPHARINI.

Type species: Limnichopharus hainanensis Miyatake, 1994, by original designation.

Family-group name: LIMNICHOPHARINI Miyatake, 1994. Stem: Limnichophar-.

Limnichus Dejean, 1821: 49. Current status: valid genus in LIMNICHIDAE: LIMNICHINAE: LIMNICHINI.

Type species: Byrrhus pygmaeus Sturm, 1807, by monotypy.

Family-group name: LIMNICHINI Erichson, 1846. Stem: Limnich-.

Limnius Illiger, 1802: 297. Current status: valid genus in ELMIDAE: ELMINAE: ELMINI: ELMINA.

Type species: Dytiscus volckmari Panzer, 1793, by monotypy.

Family-group name: LIMNIIDAE C.G. Thomson, 1859. Stem: Limni-.

Limnobaris Bedel, 1885: 183. Current status: valid genus in CURCULIONIDAE: BARIDINAE: APOSTASIMERINI: DIORYMERINA.

Type species: Curculio t-album Linnaeus, 1758, by original designation.

Family-group name: LIMNOBARINI Casey, 1922. Stem: Limnobarid-.

Limonius Eschscholtz, 1829a: 33. Current status: valid genus in ELATERIDAE: DENDROMETRINAE: DENDROMETRINI: DENDROMETRINA.

Type species: Elater minutus Linnaeus, 1758, by subsequent designation (Curtis 1838a: pl. 694). Note: Curtis’ typification, issued 1 June 1838 (see Evenhuis 1997a: 166), is earlier than Westwood’s typification, issued July 1838 (ICZN 1957a), of Elater bructeri Panzer, 1795 (= Elater aeneoniger DeGeer, 1774), currently the type species of Pheletes Kiesenwetter, 1858.

Family-group name: LIMONIINA Jakobson, 1913. Stem: Limoni-. Note: junior homonym of LIMONIIDAE Speiser, 1909 (type genus Limonia Meigen, 1803) in Diptera.

Limulodes A. Matthews, 1867: 409. Current status: valid genus in PTILIIDAE: PTILIINAE: CEPHALOPLECTINI.

Type species: Limulodes paradoxus A. Matthews, 1867, by monotypy.

Family-group name: LIMULODINAE Ganglbauer, 1898. Stem: Limulod-.

Lina Latreille, 1829b: 151. Current status: junior synonym of Chrysomela Linnaeus, 1758 in CHRYSOMELIDAE: CHRYSOMELINAE: CHRYSOMELINI.

Type species: Chrysomela populi Linnaeus, 1758, by monotypy.

Family-group name: LINAEINES Motschulsky, 1860. Stem: Lin-.

Liogenys Guérin-Méneville, 1831a: pl. 3. Current status: valid genus in SCARABAEIDAE: MELOLONTHINAE: DIPLOTAXINI.

Type species: Liogenys castaneus Guérin-Méneville, 1831 (= Melolontha palpalis Eschscholtz, 1822), by monotypy.

Family-group name: LIOGENITAE Blanchard, 1851. Stem: Liogeny-.

Lionychus Wissmann, 1846: 25. Current status: valid genus in CARABIDAE: HARPALINAE: LEBIINI: DROMIUSINA.

Type species: Lebia quadrillum Duftschmid, 1812, by monotypy.

Family-group name: LIONYCHIDAE Jeannel, 1948. Stem: Lionych-.

Lioon Casey, 1912: 67. Current status: valid genus in BYRRHIDAE: BYRRHINAE: SIMPLOCARIINI.

Type species: Lioon speculare Casey, 1912 (= Amphicyrta simplicipes Mannerheim, 1852), by subsequent designation (R. Lucas 1920: 377).

Family-group name: LIOONINI Casey, 1912. Stem: Lioon-.

Liophloeus Germar, 1817b: 341. Current status: valid genus in CURCULIONIDAE: ENTIMINAE: POLYDRUSITAE: POLYDRUSINI.

Type species: Curculio nubilus Fabricius, 1777 (= Curculio tessulatus O.F. Müller, 1776), by monotypy.

Family-group name: LIOPHLOEIDAE Gistel, 1848. Stem: Liophloe-.

Lioxyonyx Hustache, 1933: 100. Current status: valid genus in CURCULIONIDAE: CEUTORHYNCHINAE: LIOXYONYCHINI.

Type species: Lioxyonyx uniseriatus Hustache, 1933, by original designation.

Family-group name: LIOXYONYXINI Colonnelli, 1984. Stem: Lioxyonych-.

Liparetrus Guérin-Méneville, 1831a: pl. 3. Current status: valid genus in SCARABAEIDAE: SERICOIDINAE: LIPARETRINI.

Type species: Liparetrus discipennis Guérin-Méneville, 1831, by monotypy.

Family-group name: LIPARETRIDAE Burmeister, 1855. Stem: Liparetr-.

Liparocephalus Mäklin in Mannerheim, 1853: 191. Current status: valid genus in STAPHYLINIDAE: ALEOCHARINAE: LIPAROCEPHALINI.

Type species: Liparocephalus brevipennis Mäklin, 1853, by monotypy.

Family-group name: LIPAROCEPHALI Fenyes, 1919. Stem: Liparocephal-.

Liparochrus Erichson, 1848: 925. Current status: valid genus in HYBOSORIDAE: LIPAROCHRINAE.

Type species: Liparochrus geminatus Westwood, 1852, by subsequent designation (Blackburn 1912: 41). Note: no species were originally included in this genus by Erichson (1848); the first included species were the three species described by Westwood (1852: 69, 70).

Family-group name: LIPAROCHRINAE Ocampo, 2006. Stem: Liparochr-.

Liparoderus LaFerté-Sénectère in Guérin-Méneville, 1849 [No 29]: 85. Current status: valid genus in ANTHICIDAE: ANTHICINAE: MICROHORIINI.

Type species: Anthicus insignis P.H. Lucas, 1843, by original designation.

Family-group name: LIPARODERINI Bonadona, 1990. Stem: Liparoder-.

Liparus G.-A. Olivier, 1807: [83] 283. Current status: valid genus in CURCULIONIDAE: MOLYTINAE: MOLYTINI: MOLYTINA.

Type species: Curculio germanus Linnaeus, 1758, by subsequent designation (Latreille 1810: 430).

Family-group name: LIPARIDES Latreille, 1828. Stem: Lipar-. Note: senior homonym of LIPARIDAE Gill, 1861 (type genus Liparis Scopoli, 1777) in Pisces.

Lispinus Erichson, 1839b: 31. Current status: valid genus in STAPHYLINIDAE: OSORIINAE: THORACOPHORINI: LISPININA.

Type species: Lispinus attenuatus Erichson, 1840, by subsequent designation (Duponchel 1841b: 57). Note: no species were originally included in this genus by Erichson (1839b); the first ones were the seven species listed by Erichson (1840a: 828–830).

Family-group name: LISPINI Bernhauer & Schubert, 1910. Stem: Lispin-.

Lispotherium Faust, 1891: 194. Current status: valid genus in BRENTIDAE: APIONINAE: MYRMACICELITAE: LISPOTHERIINI.

Type species: Lispotherium hildebrandti Faust, 1891, by monotypy.

Family-group name: LISPOTHERIINI Wanat, 2001. Stem: Lispotheri-.

Lissauchenius W.S. MacLeay, 1825: 13. Current status: valid subgenus of Chlaenius Bonelli, 1810 in CARABIDAE: HARPALINAE: CHLAENIINI: CHLAENIINA.

Type species: Lissauchenius rufifemoratus W.S. MacLeay, 1825, by monotypy.

Family-group name: LISSAUCHENIIDAE Gistel, 1848. Stem: Lissaucheni-.

Lissodema Curtis, 1833b: 187. Current status: valid genus in SALPINGIDAE: SALPINGINAE.

Type species: Lissodema heyana Curtis, 1833 (= Salpingus cursor Gyllenhal, 1813), by monotypy.

Family-group name: LISSODEMINA Seidlitz, 1916. Stem: Lissodem-.

Lissogenius Schaum, 1845: 420. Current status: valid genus in SCARABAEIDAE: CETONIINAE: CREMASTOCHEILINI: LISSOGENIINA.

Type species: Lissogenius planicollis Schaum, 1845, by monotypy.

Family-group name: LISSOGENIINA Krikken, 1984. Stem: Lissogeni-.

Lissomus Dalman, 1824: 13. Current status: valid genus in ELATERIDAE: LISSOMINAE: LISSOMINI.

Type species: Lissomus punctulatus Dalman, 1824, by subsequent designation (Fleutiaux 1947: 138).

Family-group name: LISSOMIDAE Laporte, 1835. Stem: Lissom-.

Lissonotus Dalman in Schönherr, 1817: 159. Current status: valid genus in CERAMBYCIDAE: CERAMBYCINAE: LISSONOTINI.

Type species: Cerambyx biguttatus Dalman, 1817, by monotypy.

Family-group name: LISSONOTINAE Swainson, 1840. Stem: Lissonot-. Note: senior homonym of LISSONOTINI Förster, 1869 (type genus Lissonota Gravenhorst, 1829) in Hymenoptera.

Lissopogonus Andrewes, 1923a: 213. Current status: valid genus in CARABIDAE: PATROBINAE: LISSOPOGONINI.

Type species: Lissopogonus glabellus Andrewes, 1923, by monotypy.

Family-group name: LISSOPOGONINI Zamotajlov, 2000. Stem: Lissopogon-.

Lissorhoptrus J.L. LeConte in J.L. LeConte and G.H. Horn, 1876: 183. Current status: valid genus in CURCULIONIDAE: BRACHYCERINAE: TANYSPHYRINI.

Type species: Bagous simplex Say, 1832, by subsequent designation (Kuschel 1952: 28).

Family-group name: LISSORHOPTRINAE Böving & Craighead, 1931. Stem: Lissorhoptr-.

Lissotes Westwood, 1855b: 213. Current status: valid genus in LUCANIDAE: LUCANINAE: SCLEROSTOMINI.

Type species: Lucanus cancroides Fabricius, 1787, by subsequent designation (R. Lucas 1920: 378). Note: the type species usually listed in the literature is Lissotes menalcas Westwood, 1855, by subsequent designation of Didier and Séguy (1953: 172) but R. Lucas’ designation is earlier; both nominal species are currently placed in the genus Lissotes Westwood, 1855.

Family-group name: LISSOTINI Benesh, 1955. Stem: Lissotet-.

Listriophorus L.W. Schaufuss, 1872: 245. Current status: valid genus in STAPHYLINIDAE: PSELAPHINAE: GONIACERITAE: GONIACERINI.

Type species: Listriophorus felix L.W. Schaufuss, 1874, by subsequent monotypy (L.W. Schaufuss 1874: 289).

Family-group name: LISTRIOPHORINI L.W. Schaufuss, 1890. Stem: Listriophor-.

Listrobyctiscus Voss, 1923: 511. Current status: valid genus in ATTELABIDAE: RHYNCHITINAE: RHYNCHITITAE: BYCTISCINI: LISTROBYCTISCINA.

Type species: Rhynchites corvinus Pascoe, 1885, by original designation.

Family-group name: LISTROBYCTISCINA Legalov, 2003. Stem: Listrobyctisc-.

Listroderes Schönherr, 1826: 158. Current status: valid genus in CURCULIONIDAE: CYCLOMINAE: LISTRODERINI: LISTRODERINA.

Type species: Listroderes costirostris Schönherr, 1826, by original designation.

Family-group name: LISTRODERI J.L. LeConte, 1876. Stem: Listroder-.

Listrus Motschulsky, 1860a: 389. Current status: valid genus in MELYRIDAE: DASYTINAE: LISTRINI.

Type species: Listrus punctatus Motschulsky, 1860, by subsequent designation (Blaisdell 1938: 27).

Family-group name: LISTRINI Majer, 1990. Stem: Listr-.

Lithinus Klug, 1833: 108. Current status: valid genus in CURCULIONIDAE: MOLYTINAE: LITHININI: LITHININA.

Type species: Lithinus superciliosus Klug, 1833, by monotypy. Note: combined description of a new genus-group taxon and a single new species (ICZN 1999a, Article 12.2.6).

Family-group name: LITHINIDES Lacordaire, 1863. Stem: Lithin-. Note: senior homonym of LITHININI Forbes, 1948 (type genus Lithina Hübner, 1825) in Lepidoptera.

Lithocharis Dejean, 1833a: 65. Current status: valid genus in STAPHYLINIDAE: PAEDERINAE: LATHROBIINI: MEDONINA.

Type species: Paederus ochraceus Gravenhorst, 1802, by subsequent designation (C.G. Thomson 1859: 28).

Family-group name: LITHOCHARES Casey, 1905. Stem: Lithochar-.

†Lithocupes Ponomarenko, 1966: 65. Current status: valid genus in ARCHOSTEMATA incertae sedis. Note: Upper Triassic (Kyrgystan).

Type species: †Lithocupes incertus Ponomarenko, 1966, by original designation.

Family-group name: †LITHOCUPEDINI Ponomarenko, 1969. Stem: Lithocuped-.

†Litholamprima Nikolajev & Ren, 2015: 16. Current status: valid genus in LUCANIDAE: †LITHOLAMPRIMINAE. Note: Lower Cretaceous (China).

Type species: †Litholamprima longimana Nikolajev & Ren, 2015, by original designation.

Family-group name: †LITHOLAMPRIMINAE Nikolajev in Nikolajev and Ren, 2015: 16. Stem: Litholamprim-.

Lithophilus Frölich, 1799: 11. Current status: invalid senior synonym of Tetrabrachys Kapur, 1948 in COCCINELLIDAE: COCCINELLINAE: COCCIDULINI. Note: junior homonym of Lithophilus D.H. Schneider, 1791 [Coleoptera: CARABIDAE].

Type species: Lithophilus ruficollis Frölich, 1799 (= Tritoma connata Panzer, 1796), by monotypy.

Family-group name: LITHOPHILIDAE Imhoff, 1856. Stem: Lithophil-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

†Lithoscarabaeus Nikolajev, 1992: 78. Current status: valid genus in SCARABAEIDAE: †LITHOSCARABAEINAE. Note: Lower Cretaceous (Russia).

Type species: †Proteroscarabaeus baissensis Nikritin,1977, by original designation.

Family-group name: †LITHOSCARABAEINAE Nikolajev, 1992. Stem: Lithoscarabae-.

†Lithostoma Martynov, 1926: 13. Current status: valid genus in TROGOSSITIDAE: TROGOSSITINAE: †LITHOSTOMINI. Note: Upper Jurassic (Kazakhstan).

Type species: †Lithostoma expansum Martynov, 1926, by monotypy.

Family-group name: †LITHOSTOMINI Kolibáč & D.-Y. Huang, 2008. Stem: Lithostom- (ICZN 1999a, Article 29.4).

Litoborus Mulsant & Rey, 1854: 124, 126. Current status: valid genus in TENEBRIONIDAE: BLAPTINAE: DENDARINI: MELAMBIINA.

Type species: Philax moreletii P.H. Lucas, 1846, by subsequent designation (Antoine 1931: 181).

Family-group name: LITOBORINAE Antoine, 1941. Stem: Litobor-.

Litosomus Lacordaire, 1865: 305. Current status: synonym of Toxorhinus Lacordaire, 1865 in CURCULIONIDAE: DRYOPHTHORINAE: LITOSOMINI: LITOSOMINA. Note: First Reviser (ICZN 1999a, Article 24.2.2) for Toxorhinus Lacordaire, 1865 versus Litosomus Lacordaire, 1865 is Wibmer and O’Brien (1986: 364).

Type species: Litosomus grallarius Lacordaire, 1865, by original designation.

Family-group name: LITOSOMIDES Lacordaire, 1865. Stem: Litosom-. Note: junior homonym of LITOSOMIDAE J.W. Douglas & J. Scott, 1865 (type genus Litosoma J.W. Douglas & J. Scott, 1865) in Hemiptera.

Lixus Fabricius, 1801a: xi. Current status: valid genus in CURCULIONIDAE: LIXINAE: LIXINI.

Type species: Curculio paraplecticus Linnaeus, 1758, by subsequent designation (Latreille 1810: 430). Note: no species were originally included in this genus by Fabricius (1801a); the first ones were the 43 species listed by Fabricius (1801b: 498–507).

Family-group name: LIXIDES Schönherr, 1823. Stem: Lix-.

Ljudmilinius Legalov, 2003: 362. Current status: valid genus in ATTELABIDAE: ATTELABINAE: EUOPINI.

Type species: Euops femoralis Voss, 1924, by original designation.

Family-group name: LJUDMILININA Legalov, 2007. Stem: Ljudmilin- (ICZN 1999a, Article 29.4).

†Lobanoviella Kirejtshuk & Reid, 2021: 1895. Current status: valid genus in MEGALOPODIDAE: PALOPHAGINAE: †LOBANOVIELLINI. Note: Eocene (Baltic amber).

Type species: †Lobanoviella andreyi Kirejtshuk & Reid, 2021, by original designation.

Family-group name: †LOBANOVIELLINI Kirejtshuk & Reid, 2021: 1895. Stem: Lobanoviell-.

Loberonotha Sen Gupta & Crowson, 1969: 127. Current status: valid genus in EROTYLIDAE: XENOSCELINAE.

Type species: Telmatophilus olivascens Broun, 1893, by original designation.

Family-group name: LOBERONOTHINI Sen Gupta & Crowson, 1969. Stem: Loberonoth-.

Loberus J.L. LeConte, 1861: 98. Current status: valid genus in EROTYLIDAE: LOBERINAE.

Type species: Loberus impressus J.L. LeConte, 1863, by subsequent monotypy (J.L. LeConte 1863b: 70).

Family-group name: LOBERINAE Bruce, 1951. Stem: Lober-.

Lobodontus Chaudoir, 1842: 841. Current status: valid genus in CARABIDAE: HARPALINAE: LEBIINI: PERICALINA.

Type species: Lobodontus trisignatus Chaudoir, 1842, by monotypy.

Family-group name: LOBODONTINI Jeannel, 1949. Stem: Lobodont-.

Lobonyx Jacquelin du Val, 1859a: 42. Current status: valid genus in PRIONOCERIDAE.

Type species: Lagria aenea Fabricius, 1798, by original designation. Note: this species, described from “Tanger”, is not the same as Lagria aenea Fabricius, 1787 (= Lagria aenea Fabricius, 1775) described from “insulis Americae.”

Family-group name: LOBONYCHINI Majer, 1987. Stem: Lobonych-.

Loborhynchus Dejean, 1821: 90. Current status: invalid senior synonym of Otiorhynchus Germar, 1822 in CURCULIONIDAE: ENTIMINAE: OTIORHYNCHITAE: OTIORHYNCHINI. Note: name first published in an available work as a junior synonym and subsequently used as valid before 1961 (e.g., Schönherr 1823: 1145); name suppressed for the Principle of Priority and placed on the Official Index of Rejected and Invalid Generic Names in Zoology (as Loborhynchus Schönherr, 1823) in Opinion 982 (ICZN 1972) but the original work, author, and date of publication for this name were corrected in Opinion 2299 (ICZN 2012c).

Type species: to be determined (see ICZN 2012c).

Family-group name: LOBORHYNCHIDES Schönherr, 1823. Stem: Loborhynch-. Note: name placed on the Official Index of Rejected and Invalid Family-group Names in Zoology (see ICZN 1972, 2012c); family-group name permanently invalid (ICZN 1999a, Article 39).

Lobotrachelus Schönherr, 1838: 711. Current status: valid genus in CURCULIONIDAE: CONODERINAE: LOBOTRACHELINI.

Type species: Lobotrachelus vestitus Rosenschoeld, 1838, by original designation.

Family-group name: LOBOTRACHÉLIDES Lacordaire, 1865. Stem: Lobotrachel-.

Loebliorylon Ślipiński, 1990: 81. Current status: valid genus in CERYLONIDAE: LOEBLIORYLONINAE.

Type species: Loebliorylon carinatus Ślipiński, 1990, by original designation.

Family-group name: LOEBLIORYLININAE Ślipiński, 1990. Stem: Loebliorylon-.

Loebliquasis Dolin, 1997: 845. Current status: valid genus in ELATERIDAE: NEGASTRIINAE: QUASIMUSINI: LOEBLIQUASINA.

Type species: Loebliquasis burkhardti Dolin, 1997, by original designation.

Family-group name: LOEBLIQUASINA Schimmel & Tarnawski, 2009. Stem: Loebliquas-.

Loensus R. Lucas, 1920: 380, 489. Current status: valid genus in TENEBRIONIDAE: BLAPTINAE: PEDININI: PEDININA. Note: new replacement name for Pedinopsis Gebien, 1910.

Type species: Pedinopsis pilipes Gebien, 1910, by monotypy (type fixation for Pedinopsis Gebien).

Family-group name: LOENSINI C. Koch, 1956. Stem: Loens-.

Lomanoxia Martínez, 1951: 29. Current status: valid genus in SCARABAEIDAE: APHODIINAE: EUPARIINI.

Type species: Euparia costulata Harold, 1867, by original designation.

Family-group name: LOMANOXIINI Stebnicka, 1999. Stem: Lomanoxi-.

Lomaptera Gory & Percheron, 1833: 19, 43. Current status: valid genus in SCARABAEIDAE: CETONIINAE: SCHIZORHININI: LOMAPTERINA.

Type species [suggested]: Cetonia papua Guérin-Méneville, 1831, by subsequent designation (Valck Lucassen 1961: 1). Note: Gory and Percheron’s work on CETONIINAE was issued in livraisons published on different dates and the only species of Lomaptera mentioned in the first livraison (1833), where the genus is described, is latreillei [as latreille] and that species is the type species by monotypy since it is a combined description of a new genus and a single new species (ICZN 1999a, Article 12.2.6); that species; however, is currently included in the genus Ischiopsopha Gestro, 1874; Chevrolat (1846a: 428) selected Lomaptera fasciata Burmeister, 1842 as the type species, but it is not an originally included nominal species; the current concept of the genus is based on the type species Cetonia papua Guérin-Méneville, 1831, which is not an originally included nominal species, as designated by Valck Lucassen (1961: 1); an application to the Commission is necessary to fix Cetonia papua Guérin-Méneville, 1831 as the type species of Lomaptera Gory & Percheron, 1833 in order to promote stability.

Family-group name: LOMAPTERIDAE Burmeister, 1842. Stem: Lomapter-.

Lomechusa Gravenhorst, 1806: 178. Current status: valid genus in STAPHYLINIDAE: ALEOCHARINAE: LOMECHUSINI: LOMECHUSINA.

Type species: Staphylinus emarginatus Paykull, 1789, by subsequent designation (Latreille 1810: 427).

Family-group name: LOMECHUSIDAE Fleming, 1821. Stem: Lomechus-.

Loncophorus Chevrolat, 1832c: 215. Current status: valid genus in CURCULIONIDAE: CURCULIONINAE: ANTHONOMINI.

Type species: Loncophorus obliquus Chevrolat, 1832, by subsequent designation (Pierce 1916: 467).

Family-group name: LONCOPHORINI Pierce, 1916. Stem: Loncophor-.

Longitarsus Latreille, 1829b: 155. Current status: valid genus in CHRYSOMELIDAE: GALERUCINAE: ALTICINI.

Type species: Chrysomela atricilla Linnaeus, 1761, by subsequent designation (Maulik 1926: 333).

Family-group name: LONGITARSES Maehler, 1850. Stem: Longitars-.

Lopheros J.L. LeConte, 1881b: 23. Current status: valid genus in LYCIDAE: EROTINAE: EROTINI.

Type species: Omalisus fraternus Randall, 1838, by monotypy.

Family-group name: LOPHEROTINI Kazantsev, 2012b: 399. Stem: Lopherot-.

Lophioderus Casey, 1897: 356. Current status: valid genus in STAPHYLINIDAE: SCYDMAENINAE: SCYDMAENITAE: STENICHNINI.

Type species: Lophioderus arcifer Casey, 1897, by subsequent designation (H. Franz 1985: 150).

Family-group name: LOPHIODERINI Casey, 1897. Stem: Lophioder-.

Lophocateres Olliff, 1883: 180. Current status: valid genus in LOPHOCATERIDAE.

Type species: Lophocateres nanus Olliff, 1883 (= Peltis pusilla Klug, 1833), by monotypy.

Family-group name: LOPHOCATERINAE Crowson, 1964. Stem: Lophocater-. Note: nomen protectum (see Bouchard et al. 2011: 343).

Lophotus Schönherr, 1834a: 314. Current status: junior synonym of Psuchocephalus Latreille, 1828 in CURCULIONIDAE: CYCLOMINAE: ATERPINI: ATERPINA. Note: junior homonym of Lophotus Giorna, 1809 [Pisces].

Type species: Lophotus eschscholtzi Boheman, 1834, by original designation.

Family-group name: LOPHOTIDES Jekel, 1865. Stem: Lophot-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

Lordops Schönherr, 1823: 1142. Current status: valid genus in CURCULIONIDAE: ENTIMINAE: ENTIMITAE: LORDOPINI.

Type species: Curculio schoenherri Dalman, 1823, by original designation.

Family-group name: LORDOPIDES Schönherr, 1823. Stem: Lordop-.

Loricera Latreille, 1802b: 88. Current status: valid genus in CARABIDAE: LORICERINAE.

Type species: Carabus pilicornis Fabricius, 1775, by monotypy.

Family-group name: LORICERIDES Bonelli, 1810. Stem: Loricer-.

Loubacantus Bonadona, 1959: 1033. Current status: junior synonym of Entypodera Gerstaecker, 1871 in TENEBRIONIDAE: LAGRIINAE: LAGRIINI: STATIRINA.

Type species: Loubacantus giganteus Bonadona, 1959, by original designation.

Family-group name: LOUBACANTINI Bonadona, 1959. Stem: Loubacant-.

Lovricia Pretner, 1979: 383. Current status: valid genus in CARABIDAE: TRECHINAE: LOVRICIINI.

Type species: Lovricia jalzini Pretner, 1979, by monotypy.

Family-group name: LOVRICIINA Giachino, Guéorguiev & Vailati, 2011: 68. Stem: Lovrici-.

Loxandrus J.L. LeConte, 1853a: 226, 250. Current status: valid genus in CARABIDAE: HARPALINAE: ABACETINI: LOXANDRINA. Note: new replacement name for Megalostylus Chaudoir, 1842.

Type species: Feronia erratica Dejean, 1828, by present designation. Note: the type species currently recognized for this genus, Feronia recta Say, 1823 by subsequent designation of Casey (1918: 325), is not an originally included species in Megalostylus Chaudoir, 1842 and thus cannot be the type species of Loxandrus since this name is a new replacement name for Megalostylus.

Family-group name: LOXANDRINA Erwin & Sims, 1984. Stem: Loxandr-.

Loxomerus Chaudoir, 1842: 851. Current status: valid genus in CARABIDAE: MIGADOPINAE: MIGADOPINI: MIGADOPINA.

Type species: Loxomerus nebrioides Chaudoir, 1842, by monotypy.

Family-group name: LOXOMERIFORMES Erwin, 1985. Stem: Loxomer-.

Lucanus Scopoli, 1763: 1. Current status: valid genus in LUCANIDAE: LUCANINAE: LUCANINI.

Type species: Scarabaeus cervus Linnaeus, 1758, by subsequent designation (Latreille 1810: 429).

Family-group name: LUCANIDES Latreille, 1804. Stem: Lucan-.

Lucidota Laporte, 1833c: 136. Current status: valid genus in LAMPYRIDAE: LAMPYRINAE: LUCIDOTINI: LUCIDOTINA.

Type species: Lucidota banoni Laporte, 1833, by subsequent designation (de Motschulsky 1853a: 41).

Family-group name: LUCIDOTIDES Lacordaire, 1857. Stem: Lucidot-.

Luciola Laporte, 1833c: 146. Current status: valid genus in LAMPYRIDAE: LUCIOLINAE: LUCIOLINI.

Type species: to be determined. Note: the type species frequently listed for this genus (e.g., McDermott 1964: 43, Calder 1998: 178, Ballantyne et al. 2019: 87) is “Luciola pedemontana Bonelli” as designated subsequently by de Motschulsky (1853a: 53); however, “Luciola pedemontana Bonelli” (= Luciola pedemontana Curtis, 1846) was not an available name when originally included in Luciola by Laporte and cannot be selected as the type species (ICZN 1999a, Article 67.2.1); other authors (e.g., Fanti 2022a: 169) have used Cantharis italica Linnaeus, 1758 as the type species of this genus, as designated by de Motschulsky (1853a: 53); however, Motschulsky clearly selected the unavailable name “Luciola pedemontana Bonelli” as the type of Luciola; E. Desmarest (1860: 14) subsequently designated “Luciola italica Fab.” as the type species for Luciola; however, the species selected by E. Desmarest is a misidentification (Cantharis italica Linnaeus, 1758 sensu Fabricius, 1775 = Lampyris lusitanica Charpentier, 1825); accordingly, following the recommendations in Article 70.3 (ICZN 1999a), authors working on this issue will determine if the nominal species previously cited as the type by E. Desmarest or the taxonomic species actually involved should be fixed as the type species of Luciola Laporte (L. Ballantyne, pers. comm. to P.B., 2023).

Family-group name: LUCIOLIDES Lacordaire, 1857. Stem: Luciol-.

Ludioctenus Fairmaire, 1893: lxviii. Current status: junior synonym of Tetrigus Candèze, 1857 in ELATERIDAE: AGRYPNINAE: HEMIRHIPINI: LUDIOCTENINA.

Type species: Ludioctenus akbesianus Fairmaire, 1893 (= Tetrigus cyprius Baudi di Selve, 1871), by monotypy.

Family-group name: LUDIOCTENINA Jakobson, 1913. Stem: Ludiocten-.

Ludius Berthold, 1827: 336. Current status: junior synonym of Elater Linnaeus, 1758 in ELATERIDAE: ELATERINAE: ELATERINI.

Type species: Elater ferrugineus Linnaeus, 1758, by monotypy.

Family-group name: LUDIIDES Lacordaire, 1857. Stem: Ludi-. Note: senior homonym of LUDIINI Aurivillius, 1904 (type genus Ludia Wallengren, 1865) in Lepidoptera.

Lulua Burgeon, 1931: 92. Current status: valid genus in CERAMBYCIDAE: PRIONINAE: MEROSCELISINI.

Type species: Lulua squamosa Burgeon, 1931, by monotypy.

Family-group name: LULUINA Gilmour, 1956. Stem: Lulu-.

Lunilla Jeannel, 1957: 110. Current status: valid genus in STAPHYLINIDAE: PSELAPHINAE: CLAVIGERITAE: LUNILLINI.

Type species: Lunilla saprinoides Jeannel, 1957, by original designation.

Family-group name: LUNILLINI Célis, 1969. Stem: Lunill-.

Luperaltica Crotch, 1873b: 70. Current status: valid genus in CHRYSOMELIDAE: GALERUCINAE: ALTICINI.

Type species: Altica senilis Say, 1824, by subsequent designation (Wilcox 1953: 56).

Family-group name: LUPERALTICINI Leng, 1920. Stem: Luperaltic-.

Luperca Laporte, 1840a: 63. Current status: valid genus in CARABIDAE: SIAGONINAE: LUPERCINI.

Type species: Siagona herculeana Laporte, 1835 (= Carabus laevigatus Fabricius, 1781), by monotypy.

Family-group name: LUPERCINI Lecordier, 1977. Stem: Luperc-.

Luperus Geoffroy, 1762: 230. Current status: valid genus in CHRYSOMELIDAE: GALERUCINAE: LUPERINI. Note: name attributed to Geoffroy, 1762 and placed on the Official List of Generic Names in Zoology (ICZN 1984a).

Type species: Chrysomela flavipes Linnaeus, 1767, by plenary power (see ICZN 1984a). Note: name placed on the Official List of Specific Names in Zoology (ICZN 1984a).

Family-group name: LUPERIIDAE Gistel, 1848. Stem: Luper-.

Luprops Hope, 1833: 63. Current status: valid genus in TENEBRIONIDAE: LAGRIINAE: LUPROPINI.

Type species: Luprops chrysophthalmus Hope, 1833 (= Tagenia indica Wiedemann, 1823), by monotypy.

Family-group name: LUPROPINI Lesne, 1926. Stem: Luprop-.

Luscosmodicum Martins, 1970: 47. Current status: valid genus in CERAMBYCIDAE: CERAMBYCINAE: LUSCOSMODICINI.

Type species: Luscosmodicum beaveri Martins, 1970, by original designation.

Family-group name: LUSCOSMODICINI Martins, 2003. Stem: Luscosmodic-.

Lutrochus Erichson, 1847a: 509. Current status: valid genus in LUTROCHIDAE.

Type species: Lutrochus pilula Erichson, 1847, by monotypy.

Family-group name: LUTROCHIDAE Kasap & Crowson, 1975. Stem: Lutroch-.

Lycoperdina Latreille, 1807: 73. Current status: valid genus in ENDOMYCHIDAE: LYCOPERDININAE.

Type species: Galleruca bovistae Fabricius, 1792, by subsequent designation (Latreille 1810: 432).

Family-group name: LYCOPERDINADAE Bromhead, 1838. Stem: Lycoperdin-.

Lycoprogenthes Pic, 1915b: 6. Current status: valid genus in LYCIDAE: METRIORRHYNCHINAE: LYCOPROGENTHINI. Note: the original spelling was Lycoprogentes but the incorrect subsequent spelling Lycoprogenthes is in prevailing usage, attributed to the original publication, and therefore is to be treated as the correct original spelling (ICZN 1999a, Article 33.3.1).

Type species: Lycoprogenthes pouilloni Pic, 1915, by monotypy.

Family-group name: LYCOPROGENTHINI Bocák & Bocáková, 2008. Stem: Lycoprogenth-.

Lycoptis Casey, 1890b: 311. Current status: valid genus in LOPHOCATERIDAE.

Type species: Lycoptis villosa Casey, 1890 (= Peltis americana Motschulsky, 1863), by monotypy.

Family-group name: LYCOPTINI Casey, 1890. Stem: Lycopt-. Note: nomen oblitum (see Bouchard et al. 2011: 343).

Lyctus Fabricius, 1792b: 502. Current status: valid genus in BOSTRICHIDAE: LYCTINAE: LYCTINI.

Type species: Lyctus canaliculatus Fabricius, 1792 (= Dermestes linearis Goeze, 1777), by subsequent designation (Latreille 1810: 431). Note: Goeze’s (1777) work is not consistently binominal (see ICZN 2021a) and therefore an application to the Commission is necessary to retain Dermestes linearis Goeze, 1777 as the valid name for the type species.

Family-group name: LYCTIDES Billberg, 1820. Stem: Lyct-.

Lycus Fabricius, 1787: 163. Current status: valid genus in LYCIDAE: LYCINAE: LYCINI.

Type species: Lampyris latissima Linnaeus, 1767, by subsequent designation (Blanchard in Audouin et al. 1842: pl. 32).

Family-group name: LYCUSIDAE Laporte, 1838. Stem: Lyc-.

Lydus Dejean, 1821: 75. Current status: valid genus in MELOIDAE: MELOINAE: LYTTINI.

Type species: Meloe algiricus Linnaeus, 1758, by subsequent designation (Blanchard in Audouin et al. 1841: pl. 54).

Family-group name: LYDINA Kaszab, 1959. Stem: Lyd-. Note: junior homonym of LYDIDAE Newman, 1834 (type genus Lyda Fabricius, 1805) in Hymenoptera.

Lygistopterus Dejean, 1833b: 98. Current status: valid genus in LYCIDAE: CALOCHROMINAE. Note: this name is often credited to Mulsant (1838: 79) in the literature (e.g., Bocáková and Bocák 2007: 224) but it was made available earlier by Dejean (1833b) (see Bousquet and Bouchard 2013a: 26).

Type species: Cantharis sanguineus Linnaeus, 1758, by subsequent designation (Chevrolat 1846a: 515).

Family-group name: LYGISTOPTERI J.L. LeConte, 1881. Stem: Lygistopter-.

Lygophilus Rafinesque, 1815: 114. Current status: junior synonym of Epitragus Latreille, 1802 in TENEBRIONIDAE: PIMELIINAE: EPITRAGINI. Note: unnecessary new replacement name for Epitragus Latreille, 1802.

Type species: Epitragus fuscus Latreille, 1804, by subsequent monotypy (Latreille 1804b: 322) (type fixation for Epitragus Latreille).

Family-group name: LYGOPHILIA Rafinesque, 1815. Stem: Lygophil-.

Lygrus Fåhraeus, 1872: 55. Current status: junior synonym of Pelossus J. Thomson, 1864 in CERAMBYCIDAE: CERAMBYCINAE: LYGRINI.

Type species: Lygrus apicalis Fåhraeus, 1872, by monotypy.

Family-group name: LYGRINI Sama, 2008. Stem: Lygr-.

Lymantes Schönherr, 1838: 1085. Current status: valid genus in CURCULIONIDAE: MOLYTINAE: LYMANTINI: LYMANTINA.

Type species: Lymantes scrobicollis Gyllenhal, 1838, by original designation.

Family-group name: LYMANTIDES Lacordaire, 1865. Stem: Lymant-.

Lymexylon Fabricius, 1775: 204. Current status: valid genus in LYMEXYLIDAE: LYMEXYLINAE.

Type species: Cantharis navalis Linnaeus, 1758, by subsequent designation (Latreille 1810: 427).

Family-group name: LYMOXYLONIDAE Fleming, 1821. Stem: Lymexyl-.

Lymnastis Motschulsky, 1862: 27. Current status: valid genus in CARABIDAE: TRECHINAE: TACHYINI: TACHYINA. Note: Motschulsky used two original spellings, including Lymastis (p. 31); subsequently, de Motschulsky (1868c: 9) used only the spelling Lymnastis and therefore acted as First Reviser (ICZN 1999a, Article 24.2.4).

Type species: Lymnaeum indicum Motschulsky, 1851, by subsequent designation (Casey 1918: 3).

Family-group name: LIMNASTINI Jeannel, 1937. Stem: Lymnast-.

Lypesthes Baly, 1863: 152. Current status: junior synonym of Fidia Motschulsky, 1860 in CHRYSOMELIDAE: EUMOLPINAE: BROMIINI.

Type species: Fidia atra Motschulsky, 1861, by original designation.

Family-group name: LYPESTHINI Chûjô, 1956. Stem: Lypesthet-.

Lypnea Baly, 1876: 446. Current status: valid genus in CHRYSOMELIDAE: GALERUCINAE: ALTICINI.

Type species: Lypnea flava Baly, 1876, by monotypy.

Family-group name: LYPNEINA Bechyné, 1968. Stem: Lypne-.

Lyponia C.O. Waterhouse, 1878: 99. Current status: valid genus in LYCIDAE: ATELIINAE: LYPONIINI.

Type species: Lyponia debilis C.O. Waterhouse, 1878, by original designation.

Family-group name: LYPONIININA Bocák & Bocáková, 1990. Stem: Lyponi-.

Lyprus Schönherr, 1826: 21, 288. Current status: junior synonym of Bagous Germar, 1817 in CURCULIONIDAE: BAGOINAE.

Type species: Curculio cylindrus Paykull, 1800 (= Bagous tubulus Caldara & O’Brien, 1994), by original designation.

Family-group name: LYPRIIDAE Gistel, 1848. Stem: Lypr-. Note: nomen oblitum (see Bouchard et al. 2011: 586).

Lyropaeus C.O. Waterhouse, 1878: 104. Current status: valid genus in LYCIDAE: LYROPAEINAE: LYROPAEINI.

Type species: Lycus fallax Walker, 1858, by original designation.

Family-group name: LYROPAEINI Bocák & Bocáková, 1989. Stem: Lyropae-.

Lyrosoma Mannerheim, 1853: 174. Current status: valid genus in AGYRTIDAE: AGYRTINAE.

Type species: Pteroloma pallidum Eschscholtz, 1829, by subsequent designation (Hatch 1928: 71).

Family-group name: LYROSOMINI G.H. Horn, 1880. Stem: Lyrosomat-.

Lystronychus Latreille, 1829b: 41. Current status: valid genus in TENEBRIONIDAE: ALLECULINAE: ALLECULINI: XYSTROPODINA. Note: the original spelling was Lystronichus but the incorrect subsequent spelling Lystronychus is in prevailing usage, attributed to the publication of the original spelling, and therefore deemed to be the correct original spelling (ICZN 1999a, Article 33.3.1).

Type species: Helops equestris Fabricius, 1775, by subsequent designation (Saunders 1837: 153).

Family-group name: LYSTRONYCHIDES Lacordaire, 1859. Stem: Lystronych-.

Lytta Fabricius, 1775: 260. Current status: valid genus in MELOIDAE: MELOINAE: LYTTINI.

Type species: Meloe vesicatorius Linnaeus, 1758, by subsequent designation (C.G. Thomson 1859: 124).

Family-group name: LYTTIDAE Streubel, 1846. Stem: Lytt-.

Machadous Jeannel, 1951a: 105. Current status: valid genus in STAPHYLINIDAE: PSELAPHINAE: GONIACERITAE: MACHADOINI.

Type species: Machadous punctatus Jeannel, 1951, by original designation.

Family-group name: MACHADOINI Jeannel, 1951. Stem: Machado-.

Machaerites L. Miller, 1855: 509. Current status: valid genus in STAPHYLINIDAE: PSELAPHINAE: GONIACERITAE: BYTHININI: MACHAERITINA.

Type species: Machaerites spelaeus L. Miller, 1855, by monotypy.

Family-group name: MACHAERITINA Jeannel, 1950. Stem: Machaerit-.

Machla Herbst, 1799: 152. Current status: valid genus in TENEBRIONIDAE: PIMELIINAE: ASIDINI. Note: the senior homonym Machla Lichtenstein, 1796 was published in a work that was suppressed for nomenclatural purposes (ICZN 1995f).

Type species: Opatrum villosum G.-A. Olivier, 1795, by subsequent designation (R. Lucas 1920: 386).

Family-group name: MACHLINI Chatanay, 1914. Stem: Machl-.

Machozetus Chaudoir, 1850: 448. Current status: valid genus in CARABIDAE: HARPALINAE: HARPALINI: DITOMINA. Note: new replacement name for Harpactes Ménétriés, 1848.

Type species: Harpactes lehmanni Ménétriés, 1848, by monotypy (type fixation for Harpactes Ménétriés).

Family-group name: MACHOZETI Schauberger, 1934. Stem: Machozet-.

Macraspis W.S. MacLeay, 1819: 156. Current status: valid genus in SCARABAEIDAE: RUTELINAE: RUTELINI: RUTELINA.

Type species: Cetonia quadrivittata G.-A. Olivier, 1789 (= Scarabaeus cincta Drury, 1782), by subsequent designation (Crotch 1870b: 235).

Family-group name: MACRASPIDA Perty, 1840. Stem: Macraspid-.

Macratria Newman, 1838: 377. Current status: valid genus in ANTHICIDAE: MACRATRIINAE: MACRATRIINI.

Type species: Macratria linearis Newman, 1838 (= Dircaea murina Fabricius, 1801), by monotypy.

Family-group name: MACRATRIINI J.L. LeConte, 1862. Stem: Macratri-.

Macraulacus Bonvouloir, 1871: 71. Current status: valid genus in EUCNEMIDAE: MACRAULACINAE: MACRAULACINI. Note: this name was first used by Gemminger in Gemminger and von Harold (1869b: 1474) but is unavailable from that date as the species listed under the name is a nomen nudum; Bonvouloir’s plate 25 referred to by Gemminger was not issued until 1872 (1872a), but before the species description (1872b: 509).

Type species: Macraulacus excavatus Bonvouloir, 1872, by subsequent monotypy (Bonvouloir 1872b: 509).

Family-group name: MACRAULACINAE Fleutiaux, 1923. Stem: Macraulac-.

Macrobasis J.L. LeConte, 1862: 272. Current status: valid subgenus of Epicauta Dejean, 1834 in MELOIDAE: MELOINAE: EPICAUTINI.

Type species: Lytta albida Say, 1824, by subsequent designation (Wellman 1910a: 396). Note: no species were originally included in this genus by J.L. LeConte (1862); the first ones were the species listed by J.L. LeConte (1866b: 68).

Family-group name: MACROBASES J.L. LeConte, 1862. Stem: Macrobase-.

Macrodactylus Dejean, 1821: 58. Current status: valid genus in SCARABAEIDAE: MELOLONTHINAE: MACRODACTYLINI.

Type species: Melolontha subspinosa Fabricius, 1775, by monotypy.

Family-group name: MACRODACTYLIDAE W. Kirby, 1837. Stem: Macrodactyl-.

Macrodontia Lacordaire, 1830b: 171. Current status: valid genus in CERAMBYCIDAE: PRIONINAE: MACRODONTIINI.

Type species: Cerambyx cervicornis Linnaeus, 1758, by monotypy.

Family-group name: MACRODONTITAE J. Thomson, 1861. Stem: Macrodonti-.

Macrolinus Kaup, 1868a: 18. Current status: valid genus in PASSALIDAE: PASSALINAE: MACROLININI.

Type species: Passalus latipennis Percheron, 1841, by subsequent designation (Gravely 1918: 80).

Family-group name: MACROLINEAE Kaup, 1871. Stem: Macrolin-.

Macrolopha Weise, 1902b: 119. Current status: valid genus in MEGALOPODIDAE: MEGALOPODINAE: MEGALOPODINI: MACROLOPHINA.

Type species: Macrolopha rustica Weise, 1902, by subsequent designation (Rodríguez-Mirón, 2018: 277).

Family-group name: MACROLOPHINA Rodríguez-Mirón in Rodríguez-Mirón et al., 2021: 695. Stem: Macroloph-.

Macrolycinella Pic, 1922: 21. Current status: valid genus in LYCIDAE incertae sedis.

Type species: Macrolycinella bicolor Pic, 1922, by monotypy.

Family-group name: MACROLYCINELLINI Kazantsev, 2012b: 399. Stem: Macrolycinell-.

Macrolycus C.O. Waterhouse, 1878: 96. Current status: valid genus in LYCIDAE: ATELIINAE: MACROLYCINI.

Type species: Macrolycus bowringii C.O. Waterhouse, 1878, by monotypy. Note: two original spellings were used by C.O. Waterhouse (1878) for this species, M. coccineus (p. 96) and M. bowringii (p. 105); subsequently, C.O. Waterhouse (1880a: pl. I, fig. 1) used only M. bowringii and therefore acted as First Reviser (ICZN 1999a, Article 24.2.4).

Family-group name: MACROLYCINAE Kleine, 1929. Stem: Macrolyc-.

Macroma Gory & Percheron, 1833: 19, 35, 53. Current status: junior synonym of Campsiura Hope, 1831 in SCARABAEIDAE: CETONIINAE: CREMASTOCHEILINI: MACROMINA.

Type species: Cetonia scutellata Fabricius, 1801, by subsequent designation (Hope 1837: 63).

Family-group name: MACROMINAE Burmeister & Schaum, 1840. Stem: Macrom-.

Macrones Newman, 1841: 33. Current status: valid genus in CERAMBYCIDAE: CERAMBYCINAE: MACRONINI.

Type species: Macrones exilis Newman, 1841, by monotypy.

Family-group name: MACRONIDES Lacordaire, 1868. Stem: Macron-.

Macronota Hoffmannsegg, 1817: 15. Current status: valid genus in SCARABAEIDAE: CETONIINAE: SCHIZORHININI: LOMAPTERINA.

Type species: Macronota anthracina Wiedemann, 1823 (= Cetonia nigrita Fabricius, 1775), by subsequent designation (Krikken 2018: 21, valid name of type species given as “Cetonia nigrita Fröhlich, 1792”). Note: no species were originally included in this genus by Hoffmannsegg (1817); the first ones were the three species listed by Wiedemann (1823a: 83, 84).

Family-group name: MACRONOTIDAE Burmeister, 1842. Stem: Macronot-.

Macronychus P.W.J. Müller, 1806: 209. Current status: valid genus in ELMIDAE: ELMINAE: MACRONYCHINI.

Type species: Macronychus quadrituberculatus P.W.J. Müller, 1806, by monotypy.

Family-group name: MACRONYCHIDAE Gistel, 1848. Stem: Macronych-.

Macrophylla Hope, 1837: 103. Current status: junior synonym of Aegostheta Dejean, 1833 in SCARABAEIDAE: MELOLONTHINAE: TANYPROCTINI: MACROPHYLLINA.

Type species: Melolontha longicornis Fabricius, 1787, by original designation.

Family-group name: MACROPHYLLIDAE Burmeister, 1855. Stem: Macrophyll-. Note: senior homonym of MACROPHYLLINA J.E. Gray, 1866 (type genus Macrophyllum J.E. Gray, 1838) in Mammalia.

Macroplea Samouelle, 1819: 211. Current status: valid genus in CHRYSOMELIDAE: DONACIINAE: HAEMONIINI.

Type species: Donacia zosterae Fabricius, 1801 (= Rhagium muticum Fabricius, 1792), by subsequent designation (Curtis 1830: pl. 318).

Family-group name: MACROPLEINI Lopatin, 1977. Stem: Macrople-.

Macropnus G.H. Horn, 1867: 397. Current status: junior synonym of Macropoides Guérin-Méneville, 1844 in SCARABAEIDAE: RUTELINAE: RUTELINI: HETEROSTERNINA.

Type species: Macropnus crassipes G.H. Horn, 1867, by monotypy.

Family-group name: MACROPNI G.H. Horn, 1867. Stem: Macropn-. Note: nomen oblitum (see A.B.T. Smith 2006: 171).

Macropoda Solier, 1835d: 512, 515. Current status: valid genus in TENEBRIONIDAE: PIMELIINAE: ADESMIINI.

Type species: Pimelia variolaris G.-A. Olivier, 1795, by subsequent designation (Hope 1841a: 118).

Family-group name: MACROPODOIDAE Erichson, 1846. Stem: Macropod-. Note: the authorship of this family-group name was credited to “Agassiz” in Bouchard et al. (2011: 401) but here we follow Bouchard et al. (2021: 9) in crediting new nomenclatural acts in the fascicle on Coleoptera of Agassiz’ ‘Nomenclator Zoologicus’ to the beetle specialist W.F. Erichson, whose contribution to the publication was acknowledged on the fascicle’s title page; nomen oblitum (see Bouchard et al. 2011: 401); junior homonym of MACROPODIDAE J.E. Gray, 1821 (type genus Macropus Shaw, 1790) in Mammalia.

Macropogon Motschulsky, 1845b: 38. Current status: valid genus in ARTEMATOPODIDAE: ARTEMATOPODINAE: MACROPOGONINI.

Type species: Macropogon sibiricum Motschulsky, 1845, by original designation.

Family-group name: MACROPOGONINI J.L. LeConte, 1861. Stem: Macropogon-.

Macrosiagon Hentz, 1830: 462. Current status: valid genus in RIPIPHORIDAE: RIPIPHORINAE: MACROSIAGONINI.

Type species: Mordella dimidiata Fabricius, 1781, by monotypy.

Family-group name: MACROSIAGONINI Heyden, 1908. Stem: Macrosiagon-.

Macrostylus Boheman in Schönherr, 1840a: 921. Current status: valid genus in CURCULIONIDAE: ENTIMINAE: POLYDRUSITAE: NAUPACTINI: NAUPACTINA.

Type species: Macrostylus crinitus Boheman, 1840, by original designation.

Family-group name: MACROSTYLES J.L. LeConte, 1874. Stem: Macrostyl-.

Macrostyphlus Kirsch, 1890: 25. Current status: valid genus in CURCULIONIDAE: CYCLOMINAE: LISTRODERINI: MACROSTYPHLINA.

Type species: Macrostyphlus gualcalae Kirsch, 1890, by monotypy.

Family-group name: MACROSTYPHLINA Morrone, 2013: 27. Stem: Macrostyphl-.

Macrotarrhus Bedel, 1906: 93. Current status: valid genus in CURCULIONIDAE: HYPERINAE: HYPERINI: MACROTARRHUSINA. Note: new replacement name for Macrotarsus Schönherr, 1842.

Type species: Macrotarsus bartelsii Schönherr, 1842, by original designation (type fixation for Macrotarsus Schönherr).

Family-group name: MACROTARRHUSINA Legalov, 2007. Stem: Macrotarrhus- (ICZN 1999a, Article 29.4).

Macrotoma Audinet-Serville, 1832: 137. Current status: valid genus in CERAMBYCIDAE: PRIONINAE: MACROTOMINI: MACROTOMINA. Note: nomen protectum (see Heffern et al. 2006).

Type species: Prionus palmatus Fabricius, 1792, by subsequent designation (E. Desmarest 1860: 307). Note: the type species often listed in the literature is Prionus serripes Fabricius, 1781 (e.g., Drumont and Komiya 2010: 90, 2020: 111) by subsequent designation of J. Thomson (1864: 299) but E. Desmarest’s designation is earlier; both species are currently included in the genus Macrotoma Audinet-Serville, 1832.

Family-group name: MACROTOMITAE J. Thomson, 1861. Stem: Macrotom-.

Mada Mulsant, 1850: 1095. Current status: valid genus in COCCINELLIDAE: COCCINELLINAE: EPILACHNINI.

Type species: Epilachna fraterna Mulsant, 1850, by subsequent designation (Korschefsky 1931: 68).

Family-group name: MADAINI Gordon, 1975. Stem: Mad-.

Madagaster Perkins, 1997: 179. Current status: valid genus in HYDRAENIDAE: HYDRAENINAE: MADAGASTRINI.

Type species: Madagaster steineri Perkins, 1997, by original designation.

Family-group name: MADAGASTRINI Perkins, 1997. Stem: Madagastr-.

Madarus Schönherr, 1825: 584. Current status: valid genus in CURCULIONIDAE: BARIDINAE: MADARINI: MADARINA.

Type species: Calandra corvina Fabricius, 1801, by original designation.

Family-group name: MADARIDES Jekel, 1865. Stem: Madar-.

*Madeirodula Szawaryn, Větrovec & Tomaszewska, 2020: 7. Current status: valid genus in COCCINELLIDAE: MICROWEISEINAE: *MADEIRODULINI. Note: published in a work that is nomenclaturally unavailable (ICZN 2012f, Article 8.5.3).

Type species: *Madeirodula atlantica Szawaryn, Větrovec & Tomaszewska, 2020, by original designation. Note: published in a work that is nomenclaturally unavailable (ICZN 2012f, Article 8.5.3).

Family-group name: *MADEIRODULINI Szawaryn, Větrovec & Tomaszewska, 2020: 5. Stem: Madeirodul-. Note: published in a work that is nomenclaturally unavailable (ICZN 2012f, Article 8.5.3).

Madopterus Schönherr, 1836: 734. Current status: valid genus in CURCULIONIDAE: BARIDINAE: BARIDINI: MADOPTERINA.

Type species: Madapterus talpa Gyllenhal, 1836, by original designation.

Family-group name: MADOPTÉRIDES Lacordaire, 1865. Stem: Madopter-.

Maechidius W.S. MacLeay, 1819: 140. Current status: valid genus in SCARABAEIDAE: SERICOIDINAE: MAECHIDIINI.

Type species: Trox spurius W. Kirby, 1819, by monotypy.

Family-group name: MAECHIDIINA Burmeister, 1855. Stem: Maechidi-.

Magdalinus Germar in Schönherr, 1843: 135. Current status: junior synonym of Magdalis Germar, 1817 in CURCULIONIDAE: MESOPTILIINAE: MAGDALIDINI. Note: unnecessary new replacement name for Thamnophilus Schönherr, 1823 (an unnecessary replacement name for Magdalis Germar, 1817); name placed on the Official Index of Rejected and Invalid Names in Zoology (see ICZN 1955a).

Type species: Curculio violaceus Linnaeus, 1758, by plenary power (see ICZN 1955a) (type fixation for Magdalis Germar).

Family-group name: MAGDALINIDAE Gistel, 1856. Stem: Magdalin-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

Magdalis Germar, 1817b: 340. Current status: valid genus in CURCULIONIDAE: MESOPTILIINAE: MAGDALIDINI. Note: name placed on the Official List of Generic Names in Zoology (ICZN 1955a).

Type species: Curculio violaceus Linnaeus, 1758, by plenary power (see ICZN 1955a). Note: name placed on the Official List of Specific Names in Zoology (ICZN 1955a).

Family-group name: MAGDALINAE Pascoe, 1870. Stem: Magdalid-. Note: nomen protectum (see Bouchard et al. 2011: 622).

†Magnocoleus Hong, 1998: 41. Current status: valid genus in †MAGNOCOLEIDAE. Note: Lower Cretaceous (China); type genus placed as incertae sedis within CUPEDOIDEA by Cai et al. (2022).

Type species: †Magnocoleus huangjiapuensis Hong, 1998, by original designation.

Family-group name: †MAGNOCOLEIDAE Hong, 1998. Stem: Magnocole-.

Malachius Fabricius, 1775: 207. Current status: valid genus in MELYRIDAE: MALACHIINAE: MALACHIINI.

Type species: Cantharis aenea Linnaeus, 1758, by subsequent designation (Latreille 1810: 426).

Family-group name: MALACHIUSIDAE Fleming, 1821. Stem: Malachi-.

Malchinus Kiesenwetter, 1863: 726. Current status: junior synonym of Macrocerus Motschulsky, 1845 in CANTHARIDAE: MALTHININAE: MALCHININI.

Type species: Malthodes sinuatocollis Kiesenwetter, 1852, by subsequent designation (Delkeskamp 1977: 356).

Family-group name: MALCHININI Brancucci, 1980. Stem: Malchin-.

Mallaspis Audinet-Serville, 1832: 188. Current status: valid genus in CERAMBYCIDAE: PRIONINAE: MALLASPINI.

Type species: Prionus scutellaris G.-A. Olivier, 1795, by monotypy.

Family-group name: MALLASPITAE J. Thomson, 1861. Stem: Mallasp-.

Mallodon Lacordaire, 1830b: 171. Current status: valid genus in CERAMBYCIDAE: PRIONINAE: MACROTOMINI: MALLODONINA.

Type species: Cerambyx spinibarbis Linnaeus, 1758, by monotypy.

Family-group name: MALLODONITAE J. Thomson, 1861. Stem: Mallodon-.

Mallodrya G.H. Horn, 1888: 42. Current status: valid genus in SYNCHROIDAE.

Type species: Mallodrya subaenea G.H. Horn, 1888, by monotypy.

Family-group name: MALLODRYINI G.H. Horn, 1888. Stem: Mallodry-.

Malthinus Latreille, 1805: 261. Current status: valid genus in CANTHARIDAE: MALTHININAE: MALTHININI.

Type species [suggested]: Malthinus flavus Latreille, 1805 (= Cantharis flaveola Herbst, 1786), by subsequent designation (Hope 1841a: 141). Note: the valid type species fixation is that of J.E. Gray (1832: 723), who selected Malthinus marginatus Latreille, 1805, but that species is currently placed in the genus Malthodes Kiesenwetter, 1852; an application to the Commission is necessary to fix Malthinus flavus Latreille, 1805 as the type species of Malthinus Latreille, 1805.

Family-group name: MALTHINEN Kiesenwetter, 1852. Stem: Malthin-.

Malthodes Kiesenwetter, 1852: 242, 265. Current status: valid genus in CANTHARIDAE: MALTHININAE: MALTHODINI.

Type species [suggested]: Malthinus marginatus Latreille, 1805, by subsequent designation (Brancucci 1980: 307). Note: the valid type species fixation is that of E. Desmarest (1860: 5), who selected Cantharis sanguinolenta Linnaeus, 1767 (= Meloe cinnaberinus Scopoli, 1763), but that species is currently placed in the genus Cucujus Fabricius, 1775; presumably E. Desmarest’s type species was based on a misidentification but the taxonomic species actually involved is unknown; an application to the Commission is necessary to fix Malthinus marginatus Latreille, 1805 as the type species of Malthodes Kiesenwetter, 1852.

Family-group name: MALTHODINAE Böving & Craighead, 1931. Stem: Malthod-.

Malvapion Hoffmann, 1958: 1471, 1503. Current status: valid genus in BRENTIDAE: APIONINAE: ASPIDAPIITAE: MALVAPIINI.

Type species: Curculio malvae Fabricius, 1775, by monotypy.

Family-group name: MALVAPIINI Alonso-Zarazaga, 1990. Stem: Malvapi-.

Manda Blackwelder, 1952: 230. Current status: valid genus in STAPHYLINIDAE: OXYTELINAE: PLANEUSTOMINI. Note: new replacement name for Acrognathus Erichson, 1839.

Type species: Omalium mandibulare Gyllenhal, 1827, by subsequent designation (Duponchel 1841b: 57) (type fixation for Acrognathus Erichson).

Family-group name: MANDINI Gildenkov, 2003. Stem: Mand-.

Mandelschtamius Legalov, 2003: 107. Current status: valid genus in ATTELABIDAE: RHYNCHITINAE: RHYNCHITITAE: AULETINI: MANDELSCHTAMIINA.

Type species: Mandelschtamius turneroides Legalov, 2003, by original designation.

Family-group name: MANDELSCHTAMIINA Legalov, 2003. Stem: Mandelschtami-.

Manobia Jacoby, 1885: 73. Current status: valid genus in CHRYSOMELIDAE: GALERUCINAE: ALTICINI.

Type species: Manobia nigripennis Jacoby, 1885, by subsequent designation (Maulik 1926: 407).

Family-group name: MANOBIINI Bechyné & Špringlová de Bechyné, 1975. Stem: Manobi-.

Manticora Fabricius, 1781: 320. Current status: valid genus in CICINDELIDAE: MANTICORINI.

Type species: Manticora maxillosa Fabricius, 1781 (= Carabus tuberculatus DeGeer, 1778), by monotypy.

Family-group name: MANTICORIDAE Laporte, 1834. Stem: Manticor-.

Maoraxia Obenberger, 1937: 1449. Current status: valid genus in BUPRESTIDAE: BUPRESTINAE: MAORAXIINI. Note: new replacement name for Maoriella Obenberger, 1924.

Type species: Maoriella novaezeelandiae Obenberger, 1924, by original designation (type fixation for Maoriella Obenberger).

Family-group name: MAORAXIINI Holyński, 1984. Stem: Maoraxi-.

Maorothius Assing, 2000: 22. Current status: valid genus in STAPHYLINIDAE: STAPHYLININAE: MAOROTHIINI.

Type species: Othius adustus Broun, 1880, by original designation.

Family-group name: MAOROTHIINI Assing, 2000. Stem: Maorothi-.

Marcapatia Bechyné, 1958: 653. Current status: valid genus in CHRYSOMELIDAE: GALERUCINAE: ALTICINI.

Type species: Crepidodera longicornis Jacoby, 1904, by original designation.

Family-group name: MARCAPATIINI Bechyné & Špringlová de Bechyné, 1975. Stem: Marcapati-.

Marcepania Jałoszyński, 2013: 177. Current status: valid genus in STAPHYLINIDAE: SCYDMAENINAE: CEPHENNIITAE: MARCEPANIINI.

Type species: Marcepania semengohensis Jałoszyński, 2013, by original designation.

Family-group name: MARCEPANIINI Jałoszyński, 2013: 177. Stem: Marcepani-.

Mariouta Pic, 1898: 73. Current status: valid genus in DERMESTIDAE: DERMESTINAE: MARIOUTINI.

Type species: Mariouta letourneuxi Pic, 1898, by monotypy.

Family-group name: MARIOUTINI Jakobson, 1913. Stem: Mariout-.

Marolia Mulsant, 1856b: 84, 92. Current status: valid genus in MELANDRYIDAE: MELANDRYINAE: HYPULINI.

Type species: Serropalpus variegatus Bosc, 1791, by monotypy.

Family-group name: MAROLIINI Portevin, 1934. Stem: Maroli-.

Masoreus Dejean, 1821: 15. Current status: valid genus in CARABIDAE: HARPALINAE: MASOREINI.

Type species: Harpalus luxatus Audinet-Serville, 1821 (= Harpalus wetterhallii Gyllenhal, 1813), by monotypy.

Family-group name: MAZORÉIDES Chaudoir, 1871. Stem: Masore-.

Mastachilus Kaup, 1868a: 19. Current status: valid genus in PASSALIDAE: PASSALINAE: MACROLININI. Note: the original spelling Mastachilus was changed to Mastochilus in the next issue of the journal ‘Coleopterologische Hefte’ (vol. 4, p. 31); therefore, Article 32.5.1.1 (ICZN 1999a) does not apply and the original is to be considered the correct spelling unless Mastochilus is in prevailing usage, which is not the case; Mastochilus is an unjustified emendation of Mastachilus.

Type species: Passalus polyphyllus W.S. MacLeay, 1826, by subsequent designation (Gravely 1918: 97).

Family-group name: MASTACHILINAE Kuwert, 1891. Stem: Mastachil-.

Mastax Fischer, 1828: 111. Current status: valid genus in CARABIDAE: BRACHININAE: BRACHININI: MASTACINA.

Type species: Brachinus thermarum Steven, 1806, by monotypy.

Family-group name: MASTACINA Erwin, 1970. Stem: Mastac-.

Mastiger Motschulsky, 1852: 501. Current status: valid genus in STAPHYLINIDAE: PSELAPHINAE: CLAVIGERITAE: MASTIGERINI.

Type species: Mastiger abruptus Motschulsky, 1852, by monotypy.

Family-group name: MASTIGERINI Jeannel, 1954. Stem: Mastiger-.

Mastigocoleus Tihelka & Cai in Tihelka et al., 2022: 6. Current status: valid genus in MASTIGOCOLEIDAE.

Type species: Mastigocoleus resinicola Tihelka & Cai, 2022, by original designation.

Family-group name: MASTIGOCOLEIDAE Tihelka, Jäch, Kundrata & Cai in Tihelka et al., 2022: 5. Stem: Mastigocole-.

Mastigus Latreille, 1802b: 115. Current status: valid genus in STAPHYLINIDAE: SCYDMAENINAE: MASTIGITAE: MASTIGINI.

Type species: Ptinus spinicornis Fabricius, 1787, by monotypy.

Family-group name: MASTIGOIDAE Fleming, 1821. Stem: Mastig-.

Mastinocerus Solier, 1849: 440. Current status: valid genus in PHENGODIDAE: MASTINOCERINAE.

Type species: Mastinocerus brevipennis Solier, 1849, by monotypy.

Family-group name: MASTINOCERINI J.L. LeConte, 1881. Stem: Mastinocer-.

Mastogenius Solier, 1849: 507. Current status: valid genus in BUPRESTIDAE: POLYCESTINAE: HAPLOSTETHINI.

Type species: Mastogenius parallelus Solier, 1849, by monotypy.

Family-group name: MASTOGENINI J.L. LeConte & G. H. Horn, 1883. Stem: Mastogeni-.

Masuria Cameron, 1928: 51. Current status: valid genus in STAPHYLINIDAE: ALEOCHARINAE: MASURIINI.

Type species: Masuria plumbea Cameron, 1928, by original designation.

Family-group name: MASURIINI Cameron, 1939. Stem: Masuri-.

Matheteus J.L. LeConte, 1874b: 58. Current status: valid genus in OMETHIDAE: MATHETEINAE.

Type species: Matheteus theveneti J.L. LeConte, 1874, by monotypy.

Family-group name: MATHETEI J.L. LeConte, 1881. Stem: Mathete-.

Matus Aubé, 1836: 42. Current status: valid genus in DYTISCIDAE: MATINAE.

Type species: Colymbetes bicarinatus Say, 1823, by subsequent monotypy (Aubé 1837: 189).

Family-group name: MATINI Branden, 1884. Stem: Mat-.

Mauesia Lane, 1956: 19. Current status: valid genus in CERAMBYCIDAE: LAMIINAE: MAUESIINI.

Type species: Mauesia cornuta Lane, 1956, by monotypy.

Family-group name: MAUESINAE Lane, 1956. Stem: Mauesi-.

Mauia Blackburn in Blackburn and Sharp, 1885: 194. Current status: valid genus in ANTHRIBIDAE: ANTHRIBINAE: MAUIINI.

Type species: Mauia satelles Blackburn, 1885 (= Araeocerus subnotatus Boheman, 1859), by monotypy.

Family-group name: MAUIINI B.D. Valentine, 1990. Stem: Maui-.

Mauroniscus Bourgeois, 1911: 211. Current status: valid genus in MAURONISCIDAE.

Type species: Mauroniscus roberti Bourgeois, 1911, by monotypy.

Family-group name: MAURONISCIDAE Majer, 1995. Stem: Mauronisc-.

Mayetia Mulsant & Rey, 1875: 87. Current status: valid genus in STAPHYLINIDAE: PSELAPHINAE: EUPLECTITAE: MAYETIINI.

Type species: Mayetia sphaerifer Mulsant & Rey, 1875, by monotypy.

Family-group name: MAYETIINI Scheerpeltz, 1925. Stem: Mayeti-.

Maynipeplus Kirejtshuk, 1998: 551. Current status: valid genus in NITIDULIDAE: MAYNIPEPLINAE.

Type species: Maynipeplus lomechusoides Kirejtshuk, 1998, by original designation.

Family-group name: MAYNIPEPLINAE Kirejtshuk, 1998. Stem: Maynipepl-.

Mechistocerus Fauvel, 1863: 159. Current status: valid genus in CURCULIONIDAE: CRYPTORHYNCHINAE: AEDEMONINI.

Type species: Coelosternus impressus Montrouzier, 1860, by monotypy.

Family-group name: MECISTOCERINI Morimoto, 1978. Stem: Mechistocer-.

Mecinus Germar, 1821: 315. Current status: valid genus in CURCULIONIDAE: CURCULIONINAE: MECININI. Note: nomen protectum (see Caldara 2013: 53); see Alonso-Zarazaga et al. (2023: 8) for an alternative interpretation regarding the author and type species of Mecinus, an application to the Commission may be necessary to unambiguously determine the correct author and type species for this genus-group name.

Type species: Curculio pyraster Herbst, 1795, by subsequent designation (Schönherr 1825: 587).

Family-group name: MECINIDAE Gistel, 1848. Stem: Mecin-.

Mecistostylus Lacordaire, 1865: 136. Current status: valid genus in CURCULIONIDAE: CRYPTORHYNCHINAE: CRYPTORHYNCHINI: MECISTOSTYLINA.

Type species: Mecistostylus douei Lacordaire, 1865, by original designation.

Family-group name: MÉCISTOSTYLIDES Lacordaire, 1865. Stem: Mecistostyl-.

Mecocerus Schönherr, 1833: 115. Current status: junior synonym of Acanthothorax Gaede, 1832 in ANTHRIBIDAE: ANTHRIBINAE: MECOCERINI. Note: the senior homonym Mecocerus Billberg, 1820 [Coleoptera: CURCULIONIDAE] was suppressed for the purposes of both the Principles of Priority and Homonymy and Mecocerus Schönherr, 1833 was placed on the Official List of Generic Names in Zoology (ICZN 1972).

Type species: Mecocerus gazella Gyllenhal, 1833 (= Acanthothorax longicornis Gaede, 1832), by original designation (see ICZN 1972). Note: name placed on the Official List of Specific Names in Zoology (ICZN 1972).

Family-group name: MÉCOCÉRIDES Lacordaire, 1865. Stem: Mecocer-.

Mecolenus Schönherr, 1847: 7. Current status: valid genus in BRENTIDAE: APIONINAE: MECOLENITAE.

Type species: Mecolenus wahlbergi Schönherr, 1847, by original designation.

Family-group name: MECOLENINAE Wanat, 2001. Stem: Mecolen-.

Mecomacer Kuschel, 1954a: 108. Current status: valid genus in NEMONYCHIDAE: RHINORHYNCHINAE: MECOMACERINI: MECOMACERINA.

Type species: Mecomacer scambus Kuschel, 1954, by original designation.

Family-group name: MECOMACERINI Kuschel, 1994. Stem: Mecomacer-.

Meconemus Labram & Imhoff, 1839: [Heft 4]. Current status: valid genus in ANTHRIBIDAE: ANTHRIBINAE: MECOCERINI.

Type species: Meconemus tuberculatus Labram & Imhoff, 1839, by monotypy.

Family-group name: MECONEMINI Pierce, 1930. Stem: Meconem-. Note: junior homonym of MECONEMINI Burmeister, 1838 (type genus Meconema Audinet-Serville, 1831) in Orthoptera.

Mecopelmus Blackman, 1944: 78. Current status: valid genus in CURCULIONIDAE: PLATYPODINAE: MECOPELMINI.

Type species: Mecopelmus zeteki Blackman, 1944, by original designation.

Family-group name: MECOPELMINAE Thompson, 1992. Stem: Mecopelm-.

Mecopus Schönherr, 1825: 586. Current status: valid genus in CURCULIONIDAE: CONODERINAE: MECOPINI.

Type species: Rhynchaenus bispinosus Weber, 1801, by original designation.

Family-group name: MÉCOPIDES Lacordaire, 1865. Stem: Mecop-.

Mecosarthron Buquet, 1840a: 172. Current status: valid genus in CERAMBYCIDAE: PRIONINAE: MACROTOMINI: BASITOXINA.

Type species: Mecosarthron buphagus Buquet, 1840, by monotypy.

Family-group name: MECOSARTHRINI Melzer, 1919. Stem: Mecosarthr-.

Mecyclothorax Sharp, 1903: 243. Current status: valid genus in CARABIDAE: PSYDRINAE: MORIOMORPHINI: AMBLYTELINA.

Type species: Cyclothorax montivagus Blackburn, 1878, by subsequent designation (Andrewes 1938: 135).

Family-group name: MECYCLOTHORACITAE Jeannel, 1940. Stem: Mecyclothorac-.

Mecynodera Hope, 1841a: 181. Current status: valid genus in CHRYSOMELIDAE: SAGRINAE: MEGAMERINI.

Type species: Mecynodera picta Hope, 1841 (= Lema coxalgica Boisduval, 1835), by original designation.

Family-group name: MÉCYNODÉRITES Chapuis, 1874. Stem: Mecynoder-.

Mecynorhina Hope, 1837: 60, 119. Current status: valid genus in SCARABAEIDAE: CETONIINAE: GOLIATHINI: RHOMBORHININA.

Type species: Scarabaeus polyphemus Fabricius, 1781, by original designation.

Family-group name: MECYNORRHININA Schenkling, 1921. Stem: Mecynorhin-.

Mecysmoderes Schönherr, 1837: 596. Current status: valid genus in CURCULIONIDAE: CEUTORHYNCHINAE: MECYSMODERINI.

Type species: Mecysmoderes euglyptus Gyllenhal, 1837, by original designation.

Family-group name: MECYSMODERINA Wagner, 1938. Stem: Mecysmoder-.

Mecysolobus Reitter, 1905: 248. Current status: junior synonym of Merus Gistel, 1857 in CURCULIONIDAE: MOLYTINAE: MECYSOLOBINI.

Type species: Myecysolobus lixoides Reitter, 1905 (= Alcides flavosignatus Roelofs, 1875), by monotypy.

Family-group name: MECYSLOBINI Reitter, 1913. Stem: Mecysolob-.

Medisores R.T. Bell & J.R. Bell, 1987: 288. Current status: valid genus in CARABIDAE: RHYSODINAE: MEDISORINI.

Type species: Medisores abditus R.T. Bell & J.R. Bell, 1987, by original designation.

Family-group name: MEDISORINA R.T. Bell & J.R. Bell, 1987. Stem: Medisor-.

†Mediumiuga Peris & Ruzzier, 2013: 2. Current status: valid genus in MORDELLIDAE: MORDELLINAE: †MEDIUMIUGINI. Note: Lower Cretaceous (Spain).

Type species: †Mediumiuga sinespinis Peris & Ruzzier, 2013, by original designation.

Family-group name: †MEDIUMIUGAMIINI Peris & Ruzzier, 2013: 2. Stem: Mediumiug- (stem corrected to Mediumiug- by Peris and Ruzzier (2015: 178)).

†Medmetrioxenoides Gratshev & Legalov, 2009: 414. Current status: junior synonym of Ampliceps Arnoldi, 1977 in NEMONYCHIDAE: †EOBELINAE: †KARATAUCARINI. Note: Jurassic (Kazakhstan).

Type species: †Medmetrioxenoides prodromus Gratshev & Legalov, 2009 (= Ampliceps furcitibia Arnoldi, 1977), by original designation.

Family-group name: †MEDMETRIOXENOIDESINI Legalov, 2010. Stem: Medmetrioxenoides- (ICZN 1999a, Article 29.4).

Medon Stephens, 1833b: 272, 273. Current status: valid genus in STAPHYLINIDAE: PAEDERINAE: LATHROBIINI: MEDONINA.

Type species: Medon ruddii Stephens, 1833 (= Paederus castaneus Gravenhorst, 1802), by monotypy.

Family-group name: MEDONES Casey, 1905. Stem: Medon-.

Megabasis Audinet-Serville, 1835a: 53. Current status: valid genus in CERAMBYCIDAE: LAMIINAE: MEGABASINI.

Type species: Megabasis speculifer Audinet-Serville, 1835 (= Lamia speculifera W. Kirby, 1818), by monotypy.

Family-group name: MEGABASITAE J. Thomson, 1860. Stem: Megabas-.

Megacantha Westwood, 1843c: 121. Current status: valid genus in TENEBRIONIDAE: TENEBRIONINAE: AMARYGMINI.

Type species: Megacantha tenebrosa Westwood, 1843 (= Helops dentatus Fabricius, 1801), by monotypy.

Family-group name: MÉGACANTHIDES Lacordaire, 1859. Stem: Megacanth-.

Megacephala Latreille, 1802b: 79. Current status: valid genus in CICINDELIDAE: MEGACEPHALINI.

Type species: Cicindela megacephala G.-A. Olivier, 1790, by absolute tautonymy.

Family-group name: MEGACEPHALIDAE Laporte, 1834. Stem: Megacephal-.

Megaceras Hope, 1837: 29, 82. Current status: valid genus in SCARABAEIDAE: DYNASTINAE: ORYCTINI.

Type species: to be determined. Note: Hope (1837: 82) originally fixed “Geotrupes chorinaeus, Fab.” (= Scarabaeus chorinaeus Fabricius, 1775) as the type species of Megaceras but that species is currently considered a junior synonym of Scarabaeus pan Fabricius, 1775 in the genus Enema Hope, 1837 (Ratcliffe 2003: 300); Scarabaeus chorinaeus G.-A. Olivier, 1789, considered as a member of Megaceras and sometimes its type species, for which F. Dupuis and Mantilleri (2012: 176) designated a lectotype, is not an available name since G.-A. Olivier (1789b: [3] 15) referred to the same species described by Fabricius (1775: 5); an application to the Commission is necessary to fix one of the approximately twenty nominal species currently included in Megaceras Hope (e.g., Scarabaeus jason Fabricius, 1775) as type species.

Family-group name: MEGACERIDAE Burmeister, 1847. Stem: Megacerat-.

Megacerus Fåhraeus in Schönherr, 1839: 34. Current status: valid genus in CHRYSOMELIDAE: BRUCHINAE: BRUCHINI: MEGACERINA. Note: this name was first proposed as a synonym of Bruchus Linnaeus, 1767, and subsequently used as valid before 1961 by Drapiez (1841: 203) and others, it is available and dates from its first publication as a synonym (ICZN 1999a, Article 11.6.1).

Type species: Bruchus pescaprae Fåhraeus, 1839, by monotypy.

Family-group name: MEGACERINI Bridwell, 1946. Stem: Megacer-. Note: senior homonym of MEGACERINI Viret, 1961 (type genus Megaceros Owen, 1844) in Mammalia.

Megacoelus Lacordaire, 1868: 458. Current status: valid genus in CERAMBYCIDAE: CERAMBYCINAE: MEGACOELINI. Note: new replacement name for Megaproctus Chevrolat, 1840.

Type species: Megaproctus didelphis Chevrolat, 1840, by monotypy (type fixation for Megaproctus Chevrolat).

Family-group name: MEGACOELINI Quentin & Villiers, 1969. Stem: Megacoel-.

Megaderus Dejean, 1821: 106. Current status: invalid senior synonym of Neomegaderus Monné, 2006 in CERAMBYCIDAE: CERAMBYCINAE: TRACHYDERINI: TRACHYDERINA. Note: we consider the name Megaderus Dejean, 1821 as available and a junior homonym of Megaderus Rafinesque, 1815 [Pisces], notwithstanding the status recently given by Monné and Monné (2013: 360), who treat the cerambycid name as valid (as “Megaderus Germar, 1824”) as opposed to a junior homonym (see Monné 2006: 4, 179).

Type species: Cerambyx stigma Linnaeus, 1758, by monotypy.

Family-group name: MEGADERITAE J. Thomson, 1861. Stem: Megader-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

Megagenius Solier, 1835d: 512, 513. Current status: valid genus in TENEBRIONIDAE: PIMELIINAE: TENTYRIINI.

Type species: Megagenius frioli Solier, 1835, by monotypy.

Family-group name: MEGAGENIANOS Solier, 1851. Stem: Megageni-. Note: nomen oblitum (see Bouchard et al. 2011: 401).

Megalodacne Crotch, 1873c: 352. Current status: valid genus in EROTYLIDAE: EROTYLINAE: MEGALODACNINI.

Type species: Ips fasciata Fabricius, 1777, by original designation.

Family-group name: MEGALODACNINI Sen Gupta, 1970. Stem: Megalodacn-.

Megalognatha Baly, 1878b: 416. Current status: valid genus in CHRYSOMELIDAE: GALERUCINAE: LUPERINI.

Type species: Megalognatha elegans Baly, 1878, by original designation.

Family-group name: MEGALOGNATHITES Laboissière, 1926. Stem: Megalognath-.

Megalopaussus Lea, 1906: 217. Current status: valid genus in CARABIDAE: PAUSSINAE: PAUSSINI: CERAPTERINA.

Type species: Megalopaussus amplipennis Lea, 1906, by monotypy.

Family-group name: MEGALOPAUSSINAE Wasmann, 1920. Stem: Megalopauss-.

Megalophthalmus G.R. Gray in Griffith and Pidgeon, 1831: 371, pls 39, 61. Current status: invalid senior synonym of Magnoculus McDermott, 1964 in LAMPYRIDAE: AMYDETINAE. Note: three spellings were used in the original work, Megalophthalmus (pl. 61), Megolophthalmus (pl. 39) and Megalopthalmus (p. 371); we act as First Revisers (ICZN 1999a, Article 24.2.3) and select Megalophthalmus as the correct original spelling; junior homonym of Megalophthalmus Leach, 1830 [Crustacea].

Type species: Megalophthalmus bennettii G.R. Gray, 1831, by monotypy.

Family-group name: MEGALOPHTHALMINI E. Olivier, 1907. Stem: Megalophthalm-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

Megalopinus Eichelbaum, 1915: 104. Current status: valid genus in STAPHYLINIDAE: MEGALOPSIDIINAE. Note: new replacement name for Megalops Erichson, 1839.

Type species: Oxyporus caelatus Gravenhorst, 1802, by subsequent designation (E. Desmarest 1852: 85) (type fixation for Megalops Erichson).

Family-group name: MEGALOPININAE Naomi, 1986. Stem: Megalopin-.

Megalops Erichson, 1839b: 30. Current status: invalid senior synonym of Megalopinus Eichelbaum, 1915 in STAPHYLINIDAE: MEGALOPSIDIINAE. Note: junior homonym of Megalops Lacepède, 1803 [Pisces].

Type species: Oxyporus caelatus Gravenhorst, 1802, by subsequent designation (E. Desmarest 1852: 85). Note: no species were originally included in this genus by Erichson (1839b); the first ones were the three species listed by Erichson (1840a: 751, 752).

Family-group name: MEGALOPINI Erichson, 1839. Stem: Megalop-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

Megalopsidia Leng, 1918: 204. Current status: junior synonym of Megalopinus Eichelbaum, 1915 in STAPHYLINIDAE: MEGALOPSIDIINAE. Note: new replacement name for Megalops Erichson, 1839.

Type species: Oxyporus caelatus Gravenhorst, 1802, by subsequent designation (E. Desmarest 1852: 85) (type fixation for Megalops Erichson).

Family-group name: MEGALOPSIDIINI Leng, 1920. Stem: Megalopsidi-.

Megalopus Fabricius, 1801a: xvi. Current status: valid genus in MEGALOPODIDAE: MEGALOPODINAE: MEGALOPODINI: MEGALOPODINA.

Type species: Megalopus nigricornis Fabricius, 1801, by subsequent designation (Latreille 1810: 431). Note: no species were originally included in this genus by Fabricius (1801a); the first ones were the two species listed by Fabricius (1801b: 367, 368).

Family-group name: MEGALOPIDES Latreille, 1802. Stem: Megalopod-.

Megalostomis Chevrolat in Dejean, 1836a: 416. Current status: valid genus in CHRYSOMELIDAE: CRYPTOCEPHALINAE: CLYTRINI: MEGALOSTOMIDINA.

Type species: Clytra boopis Germar, 1823 (= Clytra grossa Forsberg, 1821), by subsequent designation (Monrós 1953: 46).

Family-group name: MEGALOSTOMIDEAE Lacordaire, 1848. Stem: Megalostomid-.

Megamerus W.S. MacLeay, 1826: 448. Current status: valid genus in CHRYSOMELIDAE: SAGRINAE: MEGAMERINI.

Type species: Megamerus kingii W.S. MacLeay, 1826, by monotypy.

Family-group name: MÉGAMÉRITES Chapuis, 1874. Stem: Megamer-.

†Megametrioxenoides Gratshev & Legalov, 2009: 413. Current status: valid genus in NEMONYCHIDAE: †METRIOXENOIDINAE. Note: Early Cretaceous (Mongolia).

Type species: †Megametrioxenoides longus Gratshev & Legalov, 2009, by original designation.

Family-group name: †MEGAMETRIOXENOIDESINI Legalov, 2010. Stem: Megametrioxenoides- (ICZN 1999a, Article 29.4).

Megapalpus Guillebeau, 1893: 297. Current status: invalid senior synonym of Megistopalpus Guillebeau, 1895 in PHALACRIDAE: PHALACRINAE. Note: junior homonym of Megapalpus Macquart, 1834 [Diptera].

Type species: Megapalpus simoni Guillebeau, 1893, by monotypy.

Family-group name: MEGAPALPINI Guillebeau, 1894. Stem: Megapalp-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

Megapenthes Kiesenwetter, 1858: 353. Current status: valid genus in ELATERIDAE: ELATERINAE: MEGAPENTHINI.

Type species: Elater lugens W. Redtenbacher, 1842, by subsequent designation (Hyslop 1921: 655).

Family-group name: MEGAPENTHINI Gurjeva, 1973. Stem: Megapenth-.

Megarhinus Schönherr, 1835: 397. Current status: invalid senior synonym of Trigonocolus Lacordaire, 1863 in CURCULIONIDAE: MOLYTINAE: TRIGONOCOLINI. Note: junior homonym of Megarhinus Rafinesque, 1820 [Pisces].

Type species: Megarhinus firmus Gyllenhal, 1835, by original designation.

Family-group name: MEGARHININAE Faust, 1888. Stem: Megarhin-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

Megarthropsis Cameron, 1919: 231. Current status: valid genus in STAPHYLINIDAE: TACHYPORINAE: TACHINUSINI.

Type species: Megarthropsis decorata Cameron, 1919, by monotypy.

Family-group name: MEGARTHROPSINI Cameron, 1919. Stem: Megarthrops-.

Megarthrus Stephens, 1829b: 24. Current status: valid genus in STAPHYLINIDAE: PROTEININAE: PROTEININI.

Type species: Staphylinus depressus Paykull, 1789, by subsequent designation (Westwood 1838c: 18).

Family-group name: MEGARTHRINI Joy, 1932. Stem: Megarthr-.

Megascelis Sturm, 1826: 80. Current status: valid genus in CHRYSOMELIDAE: EUMOLPINAE: MEGASCELIDINI.

Type species: Megascelis aenea Sturm, 1826, by monotypy.

Family-group name: MÉGASCÉLIDES Chapuis, 1874. Stem: Megascelid-.

Megasoma W. Kirby, 1825: 566. Current status: valid genus in SCARABAEIDAE: DYNASTINAE: DYNASTINI: DYNASTINA.

Type species: Scarabaeus actaeon Linnaeus, 1758, by original designation.

Family-group name: MEGASOMINAE Swainson, 1840. Stem: Megasomat-.

Megasternum Mulsant, 1844: 187. Current status: valid genus in HYDROPHILIDAE: SPHAERIDIINAE: MEGASTERNINI: MEGASTERNINA. Note: name placed on the Official List of Generic Names in Zoology (ICZN 1981b).

Type species: Dermestes obscurus Marsham, 1802 (= Dermestes concinnus Marsham, 1802), by plenary power (see ICZN 1981b). Note: Dermestes obscurus Marsham, 1802 was placed on the Official List of Specific Names in Zoology (ICZN 1981b).

Family-group name: MÉGASTERNAIRES Mulsant, 1844. Stem: Megastern-.

Megataphrus Casey, 1890b: 309. Current status: valid genus in ZOPHERIDAE: COLYDIINAE: SYNCHITINI.

Type species: Megataphrus tenuicornis Casey, 1890, by monotypy.

Family-group name: MEGATAPHRINI Casey, 1890. Stem: Megataphr-.

Megatoma Herbst, 1791: 92. Current status: valid genus in DERMESTIDAE: MEGATOMINAE: MEGATOMINI: MEGATOMINA.

Type species: Dermestes undatus Linnaeus, 1758, by subsequent designation (Curtis 1829: pl. 244).

Family-group name: MEGATOMIDA Leach, 1815. Stem: Megatom-.

Megelenophorus Gebien, 1910: 121. Current status: valid genus in TENEBRIONIDAE: PIMELIINAE: MEGELENOPHORINI. Note: new replacement name for Cacicus Dejean, 1834.

Type species: Elenophorus americanus Lacordaire, 1830, by monotypy (type fixation for Cacicus Dejean).

Family-group name: MEGALENOPHORINA Ferrer, 2015: 24. Stem: Megelenophor-.

Megetra J.L. LeConte, 1859c: 127. Current status: valid genus in MELOIDAE: MELOINAE: EUPOMPHINI.

Type species: Meloe cancellatus Brandt & Erichson, 1832, by subsequent designation (Wellman 1910a: 392).

Family-group name: MEGETRINA Kaszab, 1959. Stem: Megetr-.

Megodontus Solier, 1848a: 58. Current status: valid subgenus of Carabus Linnaeus, 1758 in CARABIDAE: CARABINAE: CARABINI: CARABINA.

Type species: Carabus caelatus Fabricius, 1801, by original designation.

Family-group name: MEGADONTICI Csiki, 1927. Stem: Megodont-.

Megopis Audinet-Serville, 1832: 161. Current status: valid genus in CERAMBYCIDAE: PRIONINAE: AEGOSOMATINI.

Type species: Megopis mutica Audinet-Serville, 1832, by monotypy.

Family-group name: MEGOPIDES Lameere, 1912. Stem: Megopid-.

†Mekorhamphus Poinar, A.E. Brown & Legalov, 2016: 158. Current status: valid genus in †MESOPHYLETIDAE: †MESOPHYLETINAE. Note: Upper Cretaceous (Myanmar).

Type species: †Mekorhamphus gyralommus Poinar, A.E. Brown & Legalov, 2016, by original designation.

Family-group name: †MEKORHAMPHINI Poinar, A.E. Brown & Legalov, 2016: 158. Stem: Mekorhamph-.

Melaenus Dejean, 1831: 471, 481. Current status: valid genus in CARABIDAE: MELAENINAE.

Type species: Melaenus elegans Dejean, 1831, by monotypy.

Family-group name: MELAENINAE Csiki, 1933. Stem: Melaen-.

Melambius Mulsant & Rey, 1854: 124. Current status: valid genus in TENEBRIONIDAE: BLAPTINAE: DENDARINI: MELAMBIINA.

Type species: Opatrum barbarum Erichson, 1841, by monotypy.

Family-group name: MELAMBIATES Mulsant & Rey, 1854. Stem: Melambi-.

Melanactes J.L. LeConte, 1853f: 493. Current status: valid genus in ELATERIDAE: DENDROMETRINAE: OXYNOPTERINI.

Type species: Melanactes densus J.L. LeConte, 1853, by subsequent designation (Hyslop 1921: 656).

Family-group name: MÉLANACTIDES Candèze, 1857. Stem: Melanact-.

Melanchiton Andrewes, 1940: 536. Current status: valid genus in CARABIDAE: HARPALINAE: MELANCHITONINI. Note: new replacement name for Melanodes Chaudoir, 1876.

Type species: Chlaenius aterrimus LaFerté-Sénectère, 1853, by subsequent designation (Alluaud 1916: 227) (type fixation for Melanodes Chaudoir).

Family-group name: MELANCHITONITAE Jeannel, 1948. Stem: Melanchiton-.

Melandrya Fabricius, 1801a: xxi, 163. Current status: valid genus in MELANDRYIDAE: MELANDRYINAE: MELANDRYINI.

Type species: Helops serratus Fabricius, 1775 (= Chrysomela caraboides Linnaeus, 1761), by subsequent designation (Latreille 1810: 429).

Family-group name: MELYANDRIDA Leach, 1815. Stem: Melandry-.

Melaneros Fairmaire, 1877: 173. Current status: valid genus in LYCIDAE incertae sedis.

Type species: Melaneros acuticollis Fairmaire, 1877, by subsequent designation (Bourgeois 1891: 349). Note: the type species sometimes listed for this genus is Melaneros atroviolaceus Fairmaire, 1877 by subsequent designation of Blair (1928: 99) (e.g., Calder 1998: 149; Bocák and Bocáková 1992: 255) but, as pointed out by Bocáková (2001: 77), Bourgeois’ designation is older.

Family-group name: MELANEROTINI Kazantsev, 2010. Stem: Melanerot-.

Melanimon Steven, 1828: 98. Current status: valid genus in TENEBRIONIDAE: TENEBRIONINAE: MELANIMONINI.

Type species: Opatrum tibiale Fabricius, 1781, by monotypy.

Family-group name: MELANIMONINA Seidlitz, 1894. Stem: Melanimon-.

Melanodes Chaudoir, 1876b: 360. Current status: invalid senior synonym of Melanchiton Andrewes, 1940 in CARABIDAE: HARPALINAE: MELANCHITONINI. Note: junior homonym of Melanodes Guenée, 1857 [Lepidoptera].

Type species: Chlaenius aterrimus LaFerté-Sénectère, 1853, by subsequent designation (Alluaud 1916: 227).

Family-group name: MELANODINI Alluaud, 1916. Stem: Melanod-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

Melanophila Eschscholtz, 1829b: 9. Current status: valid genus in BUPRESTIDAE: BUPRESTINAE: MELANOPHILINI. Note: name placed on the Official List of Generic Names in Zoology (ICZN 1996b).

Type species: Buprestis acuminata DeGeer, 1774, by plenary power (see ICZN 1996b). Note: name placed on the Official List of Specific Names in Zoology (ICZN 1996b).

Family-group name: MELANOPHILINI Bedel, 1921. Stem: Melanophil-.

Melanophthalma Motschulsky, 1866b: 269. Current status: valid genus in LATRIDIIDAE: CORTICARIINAE.

Type species: Latridius transversalis Gyllenhal, 1827, by subsequent designation (Dajoz 1967: 606).

Family-group name: MELANOPHTHALMIDAE Stickney, 1923. Stem: Melanophthalm-.

Melanotus Eschscholtz, 1829a: 32. Current status: valid genus in ELATERIDAE: ELATERINAE: MELANOTINI.

Type species: Elater fulvipes Herbst, 1806 (= Elater villosus Geoffroy, 1785), by subsequent designation (Curtis 1838a: pl. 694).

Family-group name: MÉLANOTITES Candèze, 1859. Stem: Melanot-.

Melasis G.-A. Olivier, 1790a: [30] 1, 3. Current status: valid genus in EUCNEMIDAE: MELASINAE: MELASINI: MELASINA.

Type species: Elater buprestoides Linnaeus, 1761, by monotypy.

Family-group name: MELASIDAE Fleming, 1821. Stem: Melas-.

Meleagros Kirschenhofer, 1999: 68, 72. Current status: valid genus in CARABIDAE: HARPALINAE: PLATYNINI.

Type species: Meleagros coeruleus Kirschenhofer, 1999, by original designation.

Family-group name: MELEAGROSINI Morvan, 2004. Stem: Meleagros- (ICZN 1999a, Article 29.4).

Meliboeus Deyrolle, 1865: 132. Current status: valid genus in BUPRESTIDAE: AGRILINAE: CORAEBINI: MELIBOEINA.

Type species: Buprestis aeneicollis Villers, 1789 (= Coraebus fulgidicollis P.H. Lucas, 1846), by subsequent designation (Bedel 1921: 193).

Family-group name: MELIBOEINA Majer, 2000. Stem: Meliboe-.

Meligethes Stephens, 1829b: 7. Current status: valid genus in NITIDULIDAE: MELIGETHINAE. Note: this genus-group name is sometimes attributed to Stephens (1830: 30) in the literature (e.g., Jelínek and Audisio 2007: 478, Audisio et al. 2009: 456) but it was made available one year earlier by Stephens (1829b: 7).

Type species: to be determined. Note: Nitidula atrata G.-A. Olivier, 1790 is sometimes listed as the type species of this genus (e.g., Jelínek and Audisio 2007: 478, Audisio et al. 2014: 19), but it is not an originally included species; Audisio et al. (2009: 456) used the following information for the type species of Meligethes “Nitidula rufipes Marsham, 1802 (= Nitidula atrata A. G. Olivier, 1790), by subsequent designation (C.G. Thomson 1859: 67)” but Marsham (1802: 130) did not propose a new species; instead he referred to Nitidula rufipes Fabricius, 1775, which is considered a senior invalid synonym of Brassicogethes viridescens (Fabricius, 1787); the valid type species fixation for Meligethes is that of Westwood (1838c: 12), who selected Nitidula viridescens Fabricius, 1787, but that species is currently included in the genus Brassicogethes Audisio & Cline, 2009; Hope (1841a: 154) subsequently selected Nitidula rufipes Fabricius, 1775 as the type species of Meligethes; more research is needed to establish the necessary facts and determine the best possible solution, which may be to ask the Commission to fix Nitidula atrata G.-A. Olivier, 1790 as the type species of Meligethes to conserve the current usage of the name.

Family-group name: MELIGETHINA C.G. Thomson, 1859. Stem: Meligeth-.

†Meligethiella L.N. Medvedev, 1969: 119. Current status: valid genus in NITIDULOIDEA incertae sedis. Note: Upper Jurassic/Lower Cretaceous (Kazakhstan; eastern Siberia).

Type species: †Meligethiella soroniiformis L.N. Medvedev, 1969, by original designation.

Family-group name: †MELIGETHIELLINAE Kirejtshuk & Ponomarenko, 1990. Stem: Meligethiell-.

Melisodera Westwood, 1835: pl. 132. Current status: valid genus in CARABIDAE: PSYDRINAE: MORIOMORPHINI: MORIOMORPHINA.

Type species: Melisodera piceipennis Westwood, 1835, by monotypy.

Family-group name: MELISODERIDES Sloane, 1898. Stem: Melisoder-.

Melittomma Murray, 1867b: 314. Current status: valid genus in LYMEXYLIDAE: MELITTOMMATINAE.

Type species: Hylecoetus brasiliensis Laporte, 1833, by original designation.

Family-group name: MELITTOMMINAE Q.D. Wheeler, 1986. Stem: Melittommat-.

Melobasis Laporte & Gory, 1837: 3. Current status: valid genus in BUPRESTIDAE: BUPRESTINAE: MELOBASINI.

Type species: Buprestis cupriceps W. Kirby, 1819, by subsequent designation (Bellamy 1998: 11).

Family-group name: MELOBASINI Bílý, 2000. Stem: Melobas- (ICZN 1999a, Article 29.4).

Meloe Linnaeus, 1758: 342, 419. Current status: valid genus in MELOIDAE: MELOINAE: MELOINI. Note: name placed on the Official List of Generic Names in Zoology (ICZN 1999c).

Type species: Meloe proscarabaeus Linnaeus, 1758, by subsequent designation (Latreille, 1810: 430; see ICZN 1999c). Note: name placed on the Official List of Specific Names in Zoology (ICZN 1999c).

Family-group name: MELOOIDES Gyllenhal, 1810. Stem: Melo-. Note: family-group name given precedence over HORIIDAE Latreille, 1802 (ICZN 1999c).

Meloetyphlus C.O. Waterhouse, 1872: 31. Current status: valid genus in MELOIDAE: TETRAONYCINAE.

Type species: Meloetyphlus fuscatus C.O. Waterhouse, 1872, by monotypy.

Family-group name: MELOETYPHLINI Borgmeier, 1937. Stem: Meloetyphl-.

Melolontha Fabricius, 1775: 31. Current status: valid genus in SCARABAEIDAE: MELOLONTHINAE: MELOLONTHINI: MELOLONTHINA. Note: the senior homonym Melolontha Geoffroy, 1762 (and all other uses of the name prior to the publication of Melolontha Fabricius, 1775) was suppressed for both Principles of Priority and Homonymy and placed on the Official Index of Rejected and Invalid Generic Names in Zoology while Melolontha Fabricius, 1775 was placed on the Official List of Generic Names in Zoology (ICZN 1994a).

Type species: Scarabaeus melolontha Linnaeus, 1758, by absolute tautonymy (see ICZN 1994a). Note: name placed on the Official List of Specific Names in Zoology (ICZN 1994a).

Family-group name: MELOLONTHIDAE Leach, 1819. Stem: Melolonth-. Note: senior homonym of MELOLONTHINI Bedel, 1891 (type genus Melolontha Geoffroy, 1762) in Coleoptera: CHRYSOMELIDAE; Bedel’s name is permanently invalid (based on a suppressed type genus).

Melolontha Geoffroy, 1762: 195. Current status: suppressed name in CHRYSOMELIDAE: CRYPTOCEPHALINAE: CLYTRINI: CLYTRINA. Note: usually listed as a synonym of Clytra Laicharting, 1781; name suppressed for both the Principles of Priority and Homonymy and placed on the Official Index of Rejected and Invalid Names in Zoology (see ICZN 1994a).

Type species: to be determined.

Family-group name: MELOLONTHINI Bedel, 1891. Stem: Melolonth-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

Melyris Fabricius, 1775: 58. Current status: valid genus in MELYRIDAE: MELYRINAE: MELYRINI.

Type species: Melyris viridis Fabricius, 1775, by monotypy.

Family-group name: MELYRIDES Leach, 1815. Stem: Melyr-.

Mendizabalia Cobos, 1957: 233. Current status: valid genus in BUPRESTIDAE: BUPRESTINAE: MENDIZABALIINI.

Type species: Agrilus germaini Germain & Kerremans, 1906, by original designation.

Family-group name: MENDIZABALIINI Cobos, 1968. Stem: Mendizabali-.

Menemachus Schönherr, 1843: 266. Current status: valid genus in CURCULIONIDAE: CONODERINAE: MENEMACHINI.

Type species: Menemachus naevus Boheman, 1843, by original designation.

Family-group name: MÉNÉMACHIDES Lacordaire, 1865. Stem: Menemach-.

Menephilus Mulsant, 1854: 291. Current status: valid genus in TENEBRIONIDAE: STENOCHIINAE: CNODALONINI.

Type species: Tenebrio curvipes Fabricius, 1792 (= Tenebrio cylindricus Herbst, 1784), by monotypy.

Family-group name: MENEPHILINI Reitter, 1920. Stem: Menephil-.

Menoetius Dejean, 1821: 94. Current status: invalid senior synonym of Lachnopus Schönherr, 1840 in CURCULIONIDAE: ENTIMINAE: POLYDRUSITAE: EUSTYLINI: EUSTYLINA. Note: name suppressed for the Principle of Priority and placed on the Official Index of Rejected and Invalid Generic Names in Zoology (see ICZN 1987f).

Type species: Curculio valgus Fabricius, 1775, by subsequent designation (O’Brien and Wibmer 1982: 8).

Family-group name: MENOETIINI Pierce, 1913. Stem: Menoeti-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

Mentophilus Laporte, 1840b: 74. Current status: valid genus in SCARABAEIDAE: SCARABAEINAE: DELTOCHILINI. Note: three original spellings were used, including Mintophilus (pl. 23) and Minthophilus (p. 63); as far as we know, no one has acted as First Reviser (ICZN 1999a, Article 24.2.3) and we here select Mentophilus, the spelling currently in prevailing usage, as the correct original spelling; this genus is a junior synonym of Aulacium Dejean, 1833 (see Bousquet and Bouchard 2013a: 33), and an application to the Commission is necessary to conserve Mentophilus as a valid name.

Type species: here fixed (under ICZN 1999a, Article 70.3.2) as Aulacium carinatum Reiche, 1841, misidentified as Scarabaeus novaehollandiae Fabricius, 1775 (sensu Dejean, 1833), in the original designation by monotypy in Laporte (1840b: 74). Note: Laporte (1840b: 74) listed the species as “hollandiae” but from the references given, it is clear that he was referring to Scarabaeus novaehollandiae Fabricius, 1775; the nominal species Scarabaeus novaehollandiae Fabricius, 1775 is the type species of the genus Tesserodon Hope, 1837; see Duponchel and Chevrolat (1842a: 334, 335) regarding the misidentification of the type species of Mentophilus.

Family-group name: MINTHOPHILIDES Lacordaire, 1855. Stem: Mentophil-.

Menuthianaspis Franciscolo, 1972: 124. Current status: valid genus in SCRAPTIIDAE: ANASPIDINAE: MENUTHIANASPIDINI.

Type species: Anaspis sicardi Pic, 1942, by original designation.

Family-group name: MENUTHIANASPIDINI Franciscolo, 1972. Stem: Menuthianaspid-.

Meonis Laporte, 1867: 69. Current status: valid genus in CARABIDAE: PSYDRINAE: MORIOMORPHINI: AMBLYTELINA.

Type species: Meonis niger Laporte, 1867, by subsequent designation (B.P. Moore 1963b: 288).

Family-group name: MEONIDES Sloane, 1898. Stem: Meon-.

Meotica Mulsant & Rey, 1873: 174. Current status: valid genus in STAPHYLINIDAE: ALEOCHARINAE: OXYPODINI: MEOTICINA.

Type species: Meotica parasita Mulsant & Rey, 1873, by subsequent designation (Blackwelder 1952: 237).

Family-group name: MEOTICAE Seevers, 1978. Stem: Meotic-.

Meracantha W. Kirby, 1837: 237. Current status: valid genus in TENEBRIONIDAE: TENEBRIONINAE: AMARYGMINI.

Type species: Meracantha canadensis W. Kirby, 1837 (= Helops contractus Palisot de Beauvois, 1812), by monotypy.

Family-group name: MÉRACANTHIDES Lacordaire, 1859. Stem: Meracanth-.

Meriphus Erichson, 1842: 199. Current status: valid genus in CURCULIONIDAE: CURCULIONINAE: EUGNOMINI: MERIPHINA.

Type species: Meriphus fullo Erichson, 1842, by monotypy.

Family-group name: MERIPHINAE G.A.K. Marshall, 1937. Stem: Meriph-.

Merismoderus Westwood, 1847c: 130. Current status: junior synonym of Melanospilus Westwood, 1847 in CARABIDAE: PAUSSINAE: PAUSSINI: PAUSSINA. Note: Merismoderus was originally described as a subgenus of the new genus Melanospilus, but it is also an objective synonym of Dimeroderus Westwood (1847b: 469), which was also published as a subgenus of Melanospilus; see Nagel et al. (2017a: 12) regarding the need to submit an application to the Commission in order to maintain usage of the valid genus name Melanospilus Westwood, 1847 over the unused name Leiomatocerus Benson, 1846.

Type species: Melanospilus bensoni Westwood, 1847, by monotypy.

Family-group name: MERISMODERINI Wasmann, 1929. Stem: Merismoder-.

Merizodus Solier, 1849: 185. Current status: valid genus in CARABIDAE: TRECHINAE: ZOLINI.

Type species: Merizodus angusticollis Solier, 1849, by monotypy.

Family-group name: MERIZODINI Sloane, 1920. Stem: Merizodont-.

Meroda Baly, 1860: 29. Current status: valid genus in CHRYSOMELIDAE: EUMOLPINAE: MERODINI.

Type species: Meroda costata Baly, 1860, by original designation.

Family-group name: MERODITES Chapuis, 1874. Stem: Merod-.

Merodontus Jekel, 1856: 9bis. Current status: valid genus in CURCULIONIDAE: ENTIMINAE: ENTIMITAE: LORDOPINI.

Type species: Merodontus derosus Jekel, 1856, by monotypy.

Family-group name: MERODONTI Jekel, 1856. Stem: Merodont-.

Meropathus Enderlein, 1901: 121. Current status: valid genus in HYDRAENIDAE: OCHTHEBIINAE: OCHTHEBIINI.

Type species: Meropathus chuni Enderlein, 1901, by monotypy.

Family-group name: MEROPATHINA Perkins, 1997. Stem: Meropath-.

Merophysia P.H. Lucas, 1852: xxix. Current status: valid genus in ENDOMYCHIDAE: MEROPHYSIINAE.

Type species: Merophysia formicaria P.H. Lucas, 1852, by monotypy.

Family-group name: MEROPHYSIINI Seidlitz, 1872. Stem: Merophysi-.

Meroscelisus Audinet-Serville, 1832: 157. Current status: valid genus in CERAMBYCIDAE: PRIONINAE: MEROSCELISINI.

Type species: Meroscelisus violaceus Audinet-Serville, 1832, by monotypy.

Family-group name: MEROSCELISITAE J. Thomson, 1861. Stem: Meroscelis-.

Meru Spangler & Steiner, 2005: 340. Current status: valid genus in MERUIDAE.

Type species: Meru phyllisae Spangler & Steiner, 2005, by original designation.

Family-group name: MERUIDAE Spangler & Steiner, 2005. Stem: Meru-.

Meryx Latreille, 1802b: 208. Current status: valid genus in ULODIDAE.

Type species: Meryx rugosa Latreille, 1802, by monotypy.

Family-group name: MERYCIDAE Crowson, 1953. Stem: Meryc-.

Mesagroicus Schönherr, 1840b: 281. Current status: valid genus in CURCULIONIDAE: ENTIMINAE: POLYDRUSITAE: NAUPACTINI: MESAGROICINA.

Type species: Thylacites piliferus Boheman, 1833, by original designation.

Family-group name: MESAGROICINA Legalov, 2020b: 341. Stem: Mesagroic-.

†Mesernobius Engel, 2010: 32. Current status: valid genus in SCIRTIDAE: †MESERNOBIINAE. Note: mid-Cretaceous (Burmese amber).

Type species: †Mesernobius anawrahtai Engel, 2010, by original designation.

Family-group name: †MESERNOBIINAE Engel, 2010. Stem: Mesernobi-. Note: combined description of a new family-group taxon and a single new nominal genus (ICZN 1999a, Article 13.5).

†Mesocinetus Ponomarenko, 1986: 96. Current status: valid genus in †MESOCINETIDAE. Note: Lower Cretaceous (Mongolia).

Type species: †Mesocinetus mongolicus Ponomarenko, 1986, by original designation.

Family-group name: †MESOCINETIDAE Kirejtshuk & Ponomarenko, 2010. Stem: Mesocinet-.

Mesocoelopus Jacquelin du Val, 1860a: 143. Current status: valid genus in PTINIDAE: MESOCOELOPODINAE: MESOCOELOPODINI.

Type species: Ptilinus niger P.W.J. Müller, 1821, by original designation.

Family-group name: MÉSOCOELOPAIRES Mulsant & Rey, 1864. Stem: Mesocoelopod-.

†Mesocupes Martynov, 1926: 6. Current status: valid genus in CUPEDIDAE: †MESOCUPEDINAE. Note: Middle/Upper Jurassic (Kazakhstan; China).

Type species: †Mesocupes primitivus Martynov, 1926, by monotypy.

Family-group name: †MESOCUPEDINI Ponomarenko, 1969. Stem: Mesocuped-.

†Mesoderus Prokin & Ren, 2010: 50. Current status: valid genus in DYTISCIDAE: †LIADYTISCINAE: †MESODERINI. Note: Mesozoic (China).

Type species: †Mesoderus magnus Prokin & Ren, 2010, by original designation.

Family-group name: †MESODERINI Prokin, Petrov, B. Wang & Ponomarenko, 2013: 145. Stem: Mesoder-.

Mesogenus Gemminger in Gemminger and Harold, 1869b: 1485. Current status: valid genus in EUCNEMIDAE: EUCNEMINAE: MESOGENINI.

Type species: Fornax austrocaledonicus Perroud & Montrouzier, 1865, by monotypy. Note: Since Mesogenus is credited to Bonvouloir’s work (1871–1875) on EUCNEMIDAE in the literature, the type species was considered to be Mesogenus mellyi Bonvouloir, 1871, by subsequent designation (Fleutiaux 1902: 651) (e.g., Calder 1998: 16); however, this genus-group name is available from Gemminger and Harold’s catalogue, with only one available species listed under it, F. austroacledonicus, which is therefore the type species by monotypy.

Family-group name: MESOGENINI Muona, 1993. Stem: Mesogen-.

†Mesolpinus Kirejtshuk, Moseyko & Ren, 2016: 1426. Current status: valid genus in CHRYSOMELIDAE: CHRYSOMELINAE: †MESOLPININI. Note: Lower Cretaceous (China).

Type species: †Mesolpinus antennatus Kirejtshuk, Moseyko & Ren, 2016, by original designation.

Family-group name: †MESOLPININI Kirejtshuk, Moseyko & Ren, 2016: 1425. Stem: Mesolpin-.

Mesomphalia Hope, 1839: 94. Current status: valid genus in CHRYSOMELIDAE: CASSIDINAE: MESOMPHALIINI.

Type species: Cassida gibbosa Fabricius, 1781, by original designation.

Family-group name: MESOMPHALIDAE Hope, 1841. Stem: Mesomphali-.

†Mesopassandra Jin, Ślipiński, Zhou & H. Pang, 2019: 1951. Current status: valid genus in PASSANDRIDAE: †MESOPASSANDRINAE. Note: Late Cretaceous (Myanmar).

Type species: †Mesopassandra keyao Jin, Ślipiński, Zhou & H. Pang, 2019, by original designation.

Family-group name: †MESOPASSANDRINAE Jin, Ślipiński, Zhou & H. Pang, 2019: 1951. Stem: Mesopassandr-.

†Mesophyletis Poinar, 2006: 879. Current status: valid genus in †MESOPHYLETIDAE: †MESOPHYLETINAE. Note: mid-Cretaceous (Burmese amber).

Type species: †Mesophyletis calhouni Poinar, 2006, by original designation.

Family-group name: †MESOPHYLETINAE Poinar, 2006. Stem: Mesophylet-.

Mesoporus Cameron, 1959: 120. Current status: valid genus in STAPHYLINIDAE: ALEOCHARINAE: MESOPORINI.

Type species: Mesoporus rufoflavus Cameron, 1959, by monotypy.

Family-group name: MESOPORINAE Cameron, 1959. Stem: Mesopor-.

Mesoptilius Labram & Imhoff, 1845: [Heft 13] 24. Current status: valid genus in CURCULIONIDAE: MESOPTILIINAE: MESOPTILIINI.

Type species: Mesoptilius apicalis Labram & Imhoff, 1845, by monotypy.

Family-group name: MÉSOPTILIDES Lacordaire, 1863. Stem: Mesoptili-.

Mesosa Latreille, 1829b: 124. Current status: valid genus in CERAMBYCIDAE: LAMIINAE: MESOSINI.

Type species [suggested]: Cerambyx curculionoides Linnaeus, 1761, by subsequent designation (C.G. Thomson 1859: 153). Note: the valid type species fixation is that of Westwood (1838c: 41), who selected Cerambyx nubilus Gmelin, 1789 (= Lamia nebulosa Fabricius, 1781), but that species is currently in the subgenus Aplocnemia Stephens, 1831, not the nominotypical subgenus; an application to the Commission is necessary to fix Cerambyx curculionoides Linnaeus, 1761 as the type species of Mesosa Latreille, 1829.

Family-group name: MÉSOSAIRES Mulsant, 1839. Stem: Mesos-.

Mesostyloides Bouchard & Bousquet, 2020a: 114. Current status: valid genus in CURCULIONIDAE: ENTIMINAE: OTIORHYNCHITAE: MESOSTYLOIDINI. Note: new replacement name for Mesostylus Faust, 1894.

Type species: Mesostylus hauseri Faust, 1894, by subsequent designation (Faust 1898b: 231) (type fixation for Mesostylus Faust).

Family-group name: MESOSTYLOIDINI Bouchard & Bousquet, 2020a: 114. Stem: Mesostyloid-.

Mesostylus Faust in Hauser, 1894: 57. Current status: invalid senior synonym of Mesostyloides Bouchard & Bousquet, 2020 in CURCULIONIDAE: ENTIMINAE: OTIORHYNCHITAE: MESOSTYLOIDINI. Note: junior homonym of Mesostylus Bronn & Roemer, 1852 [Crustacea].

Type species: Mesostylus hauseri Faust, 1894, by subsequent designation (Faust 1898b: 231).

Family-group name: MESOSTYLINI Reitter, 1913. Stem: Mesostyl-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

Mesothes Mulsant & Rey, 1864b: 311. Current status: valid genus in PTINIDAE: MESOCOELOPODINAE: MESOCOELOPODINI.

Type species: Xyletinus ferrugineus Mulsant & Rey, 1861, by monotypy.

Family-group name: MESOTHINI Portevin, 1931. Stem: Mesothet-.

Mestogaster Schmidt-Göbel, 1838: 9. Current status: valid genus in STAPHYLINIDAE: PSELAPHINAE: PSELAPHITAE: HYBOCEPHALINI.

Type species: Mestogaster crassicornis Schmidt-Göbel, 1838, by monotypy.

Family-group name: MESTOGASTRINA Jakobson, 1910. Stem: Mestogastr-.

Metacerylon Grouvelle, 1906: 110. Current status: valid genus in EUXESTIDAE.

Type species: Metacerylon parallelum Grouvelle, 1906, by monotypy.

Family-group name: METACERYLINI Heinze, 1944. Stem: Metacerylon-.

Metachroma Chevrolat in Dejean, 1836a: 412. Current status: valid genus in CHRYSOMELIDAE: EUMOLPINAE: BROMIINI: TYPOPHORINI.

Type species: Colaspis quercata Fabricius, 1801, by subsequent designation (Crotch 1873b: 41).

Family-group name: MÉTACHROMITES Chapuis, 1874. Stem: Metachromat-.

Metacinops Kraatz, 1862a: 117. Current status: valid genus in CURCULIONIDAE: ENTIMINAE: CYPHICERITAE: METACINOPINI.

Type species: Metacinops rhinomacer Kraatz, 1862, by monotypy.

Family-group name: METACINOPINAE Reitter, 1913. Stem: Metacinop-.

Metaclisa Jacquelin du Val, 1861: 296. Current status: valid genus in TENEBRIONIDAE: TENEBRIONINAE: METACLISINI. Note: nomen protectum (see Bouchard et al. 2007: 393).

Type species: Platydema parallela Fairmaire, 1855 (= Diaperis azurea Waltl, 1838), by original designation.

Family-group name: METACLISINI Steiner, 2016: 542. Stem: Metaclis-.

Metacycla Baly, 1861: 206. Current status: valid genus in CHRYSOMELIDAE: GALERUCINAE: METACYCLINI.

Type species: Metacycla sallei Baly, 1861, by original designation.

Family-group name: MÉTACYCLITES Chapuis, 1875. Stem: Metacycl-.

Metadromius Bedel, 1907: 279. Current status: valid genus in CARABIDAE: HARPALINAE: LEBIINI: DROMIUSINA.

Type species: Dromius myrmidon Fairmaire, 1859, by subsequent designation (Jeannel 1942: 1078).

Family-group name: METADROMIINA Basilewsky, 1984. Stem: Metadromi-.

Metagnoma Aurivillius, 1925: 13. Current status: valid genus in CERAMBYCIDAE: LAMIINAE: PTEROPLIINI.

Type species: Metagnoma singularis Aurivillus, 1925, by monotypy.

Family-group name: METAGNOMINI Aurivillius, 1925. Stem: Metagnom-.

Metallica Chaudoir, 1873: 175. Current status: valid genus in CARABIDAE: HARPALINAE: LEBIINI: METALLICINA.

Type species: Metallica viridipennis Chaudoir, 1873, by subsequent designation (Shpeley 1986: 273).

Family-group name: METALLICINI Basilewsky, 1984. Stem: Metallic-.

Metanastes Arrow, 1911: 166. Current status: valid genus in SCARABAEIDAE: DYNASTINAE: PENTODONTINI: PENTODONTINA.

Type species: Heteronychus australis Fauvel, 1863, by original designation.

Family-group name: METANASTINA Carne, 1957. Stem: Metanast-.

Metanoeus C.O. Waterhouse, 1880a: 73. Current status: valid genus in LYCIDAE: METRIORRHYNCHINAE: METRIORRHYNCHINI: METANOEINA.

Type species: Lycus conformis C.O. Waterhouse, 1878, by subsequent designation (Bourgeois 1891: 347).

Family-group name: METANOEINA Sklenarova, Kubecek & Bocák, 2014: 49. Stem: Metanoe-.

Metapion Schilsky, 1906: v. Current status: valid genus in BRENTIDAE: APIONINAE: ASPIDAPIITAE: METAPIINI.

Type species: Apion candidum Wencker, 1864, by original designation.

Family-group name: METAPIINI Alonso-Zarazaga, 1990. Stem: Metapi-.

Metatyges Pascoe, 1865b: 424. Current status: junior synonym of Omophorus Schönherr, 1835 in CURCULIONIDAE: MOLYTINAE: METATYGINI.

Type species: Metatyges turritus Pascoe, 1865 (= Omophorus stomachosus Boheman, 1835), by monotypy.

Family-group name: METATYGINAE Pascoe, 1888. Stem: Metatyg-.

Metaxina Broun, 1909: 407. Current status: valid genus in CHAETOSOMATIDAE.

Type species: Metaxina ornata Broun, 1909, by monotypy.

Family-group name: METAXINIDAE Kolibáč, 2004. Stem: Metaxin-.

Metaxymorphus Chaudoir, 1850: 370. Current status: valid genus in CARABIDAE: HARPALINAE: LEBIINI: DROMIUSINA.

Type species: Dromius frenatus Dejean, 1831, by original designation.

Family-group name: METAXYMORPHINA Basilewsky, 1984. Stem: Metaxymorph-.

Metazuphium Mateu, 1992: 198. Current status: valid genus in CARABIDAE: HARPALINAE: ZUPHIINI: METAZUPHIINA.

Type species: Metazuphium spinangulis Mateu, 1992, by original designation.

Family-group name: METAZUPHIINA Mateu, 1992. Stem: Metazuphi-.

Methia Newman, 1842d: 418. Current status: valid genus in CERAMBYCIDAE: CERAMBYCINAE: METHIINI. Note: new replacement name for Thia Newman, 1842.

Type species: Thia pusilla Newman, 1840 (= Saperda necydalea Fabricius, 1798), by monotypy (type fixation for Thia Newman).

Family-group name: METHIITAE J. Thomson, 1860. Stem: Methi-.

Methioides Chemsak & Linsley, 1967: 29, 31. Current status: valid genus in CERAMBYCIDAE: CERAMBYCINAE: OEMINI: METHIOIDINA.

Type species: Methioides cicatricosa Chemsak & Linsley, 1967, by original designation.

Family-group name: METHIOIDINA Martins, 1997. Stem: Methioid-.

Methles Sharp, 1882b: 317, 489. Current status: valid genus in DYTISCIDAE: HYDROPORINAE: METHLINI.

Type species: Methles punctipennis Sharp, 1882 (= Hydroporus cribratellus Fairmaire, 1880), by subsequent designation (Guignot 1946: 113).

Family-group name: METHLIDAE Branden, 1884. Stem: Methl-.

Metholcus Jacquelin du Val, 1860a: 142. Current status: valid genus in PTINIDAE: XYLETININAE: METHOLCINI.

Type species: Ptilinus cylindricus Germar, 1817, by original designation. Note: Trypopitys phoenicis Fairmaire, 1859 is often listed as the valid name for Ptilinus cylindricus Germar, 1817 (e.g., Zahradník 2007: 361) based on the belief that Germar’s name is a primary homonym of Ptilinus cylindricus O.F. Müller, 1776; however, O.F. Müller’s name was suppressed for both the Principles of Priority and Homonymy (ICZN 1994a) and therefore Ptilinus cylindricus Germar, 1817 can be used as the valid name for the type species.

Family-group name: METHOLCINI Zahradník, 2009. Stem: Metholc-.

Metialma Pascoe, 1871a: 217. Current status: valid genus in CURCULIONIDAE: CONODERINAE: CORYSSOMERINI.

Type species: Metialma scenica Pascoe, 1871, by subsequent designation (G.A.K. Marshall 1939b: 10).

Family-group name: METIALMINI Hustache, 1932. Stem: Metialm-.

Metius Curtis, 1838b: 329. Current status: valid genus in CARABIDAE: HARPALINAE: PTEROSTICHINI: METIINA.

Type species: Metius harpalioides Curtis, 1838, by monotypy.

Family-group name: METIINI Straneo, 1951. Stem: Meti-.

Metocalolabus Legalov, 2003: 433. Current status: valid genus in ATTELABIDAE: ATTELABINAE: ATTELABINI: METOCALOLABINA.

Type species: Metocalolabus orientalis Legalov, 2003, by original designation.

Family-group name: METOCALOLABINA Legalov, 2003. Stem: Metocalolab-.

Meton Pascoe, 1859a: 42. Current status: valid genus in CERAMBYCIDAE: LAMIINAE: DESMIPHORINI.

Type species: Meton granulicollis Pascoe, 1859, by monotypy.

Family-group name: MÉTONIDES Lacordaire, 1869. Stem: Metont-.

Metopias Gory, 1832: pl. 42. Current status: valid genus in STAPHYLINIDAE: PSELAPHINAE: EUPLECTITAE: METOPIASINI: METOPIASINA.

Type species: Metopias curculionoides Gory, 1832, by monotypy (see ICZN 1994e).

Family-group name: METOPIINI Raffray, 1904. Stem: Metopias- (ICZN 1994e).

Metopocoilus Audinet-Serville, 1832: 170. Current status: valid genus in CERAMBYCIDAE: CERAMBYCINAE: TRACHYDERINI: TRACHYDERINA.

Type species: Metopocoilus maculicollis Audinet-Serville, 1832, by monotypy.

Family-group name: METOPOCOÏLITAE J. Thomson, 1864. Stem: Metopocoil-.

Metoponcus Kraatz, 1857b: 651. Current status: junior synonym of Zeteotomus Jacquelin du Val, 1856 in STAPHYLINIDAE: STAPHYLININAE: XANTHOLININI.

Type species: Leptacinus brevicornis Erichson, 1839, by subsequent designation (R. Lucas 1920: 410). Note: two species were originally included in this genus, including Leptacinus filarius Erichson, 1839.

Family-group name: METOPONCI Casey, 1906. Stem: Metoponc-.

Metopsia Wollaston, 1854: 616. Current status: valid genus in STAPHYLINIDAE: PROTEININAE: PROTEININI.

Type species: Metopsia ampliata Wollaston, 1854, by monotypy.

Family-group name: METOPSIINAE Tottenham, 1954. Stem: Metopsi-.

Metriorrhynchus Gemminger in Gemminger and Harold, 1869c: 1629. Current status: valid genus in LYCIDAE: METRIORRHYNCHINAE: METRIORRHYNCHINI: METRIORRHYNCHINA. Note: this name is an unjustified emendation of Metriorhynchus Guérin-Méneville, 1838 (a junior homonym of Metriorhynchus H. Meyer, 1830 [Reptilia]) since two or more names in Gemminger in Gemminger and Harold (1869c) are treated in the same way (ICZN 1999a, Article 33.2.1) by the inclusion of an additional “r” to the original spelling to form “rrhynchus” (rostrum) instead of “rhynchus,” e.g., Psilorrhynchus (p. 1657) and Metriorrhynchus (see Biffi 2017: 132).

Type species: Lycus parallelus Guérin-Méneville, 1838, by subsequent designation (C.O. Waterhouse 1878: 101).

Family-group name: METRIORRHYNCHINAE Kleine, 1926. Stem: Metriorrhynch-.

Metrioxena Pascoe, 1870: 442. Current status: valid genus in BELIDAE: OXYCORYNINAE: OXYCORYNITAE: METRIOXENINI: METRIOXENINA.

Type species: Metrioxena serricollis Pascoe, 1870, by monotypy.

Family-group name: METRIOXENINI Voss, 1953. Stem: Metrioxen-.

†Metrioxenoides Gratshev, Zherikhin & Jarzembowski, 1998: 323. Current status: valid genus in NEMONYCHIDAE: †METRIOXENOIDINAE. Note: Lower Cretaceous (England).

Type species: †Metrioxenoides pusillus Gratshev, Zherikhin & Jarzembowski 1998, by original designation.

Family-group name: †METRIOXENOIDINAE Legalov, 2009. Stem: Metrioxenoid-.

Metrius Eschscholtz, 1829b: 8. Current status: valid genus in CARABIDAE: PAUSSINAE: METRIINI.

Type species: Metrius contractus Eschscholtz, 1829, by monotypy.

Family-group name: METRII J.L. LeConte, 1853. Stem: Metri-.

Metrotyphlus Coiffait, 1959: 328. Current status: valid genus in STAPHYLINIDAE: LEPTOTYPHLINAE: METROTYPHLINI.

Type species: Megatyphlus hervei Coiffait, 1957, by original designation.

Family-group name: METROTYPHLINI Coiffait, 1963. Stem: Metrotyphl-.

Mezium Leach in Samouelle, 1819: 180. Current status: valid genus in PTINIDAE: PTININAE: MEZIINI. Note: this name is credited to Curtis (1828: pl. 232) in the literature but it was first made available by Leach in 1819.

Type species: Ptinus sulcatus Fabricius, 1781, by original designation.

Family-group name: MEZIINI Bellés, 1985. Stem: Mezi-.

Miarus Schönherr, 1826: 320. Current status: valid genus in CURCULIONIDAE: CURCULIONINAE: MECININI.

Type species: Curculio campanulae Linnaeus, 1767, by original designation.

Family-group name: MIARIDES Tournier, 1874. Stem: Miar-.

Miccotrogus Schönherr, 1825: 583. Current status: junior synonym of Tychius Germar, 1817 in CURCULIONIDAE: CURCULIONINAE: TYCHIINI: TYCHIINA.

Type species: Curculio cuprifer Panzer, 1799, by original designation.

Family-group name: MICCOTROGIDAE Gistel, 1856. Stem: Miccotrog-.

Micholaemus Viana, 1971: 70. Current status: valid genus in RIPIPHORIDAE: PELECOTOMINAE.

Type species: Micholaemus gerstaeckeri Viana, 1971, by original designation.

Family-group name: MICHOLAEMINAE Viana, 1971. Stem: Micholaem-.

Michthisoma J.L. LeConte, 1850c: 30. Current status: valid genus in CERAMBYCIDAE: SPONDYLIDINAE: SAPHANINI.

Type species: Michthisoma heterodoxus J.L. LeConte, 1850, by monotypy.

Family-group name: MICHTHYSOMINI J.L. LeConte, 1873. Stem: Michthisomat-.

Micilus Mulsant & Rey, 1872: 15. Current status: valid genus in HETEROCERIDAE: HETEROCERINAE: MICILINI. Note: two original spellings were used, including Mirulus (p. 40); Neave (1940: 145) acted as First Reviser (ICZN 1999a, Article 24.2.3) and selected the spelling Micilus, the spelling currently in prevailing usage.

Type species: Heterocerus murinus Kiensewetter, 1843, by monotypy.

Family-group name: MICILINI M.F. Pacheco, 1964. Stem: Micil-.

Micracis J.L. LeConte in C. Zimmermann, 1869: 164. Current status: valid genus in CURCULIONIDAE: SCOLYTINAE: MICRACIDINI.

Type species: Micracis suturalis J.L. LeConte, 1869, by subsequent designation (Hopkins 1914: 125).

Family-group name: MICRACIDES J.L. LeConte, 1876. Stem: Micracid-.

Micragyrtes Champion, 1918: 46. Current status: junior synonym of Microsilpha Broun, 1886 in STAPHYLINIDAE: MICROSILPHINAE.

Type species: Micragyrtes ocelligerus Champion, 1918, by original designation.

Family-group name: MICRAGYRTINI Jeannel, 1962. Stem: Micragyrt-.

Micralymma Westwood, 1837: 129. Current status: valid genus in STAPHYLINIDAE: OMALIINAE: OMALIINI.

Type species: Micralymma johnstonis Westwood, 1837 (= Staphylinus marinus Strøm, 1783), by monotypy.

Family-group name: MICRALYMMATES Mulsant & Rey, 1880. Stem: Micralymmat-.

Micrantereus Solier, 1848b: 151, 175. Current status: valid genus in TENEBRIONIDAE: BLAPTINAE: PEDININI: HELOPININA.

Type species: Acanthomerus anomalus Guérin-Méneville, 1834, by original designation.

Family-group name: MICRANTEREINI Reitter, 1917. Stem: Micrantere-.

Micratopus Casey, 1914: 42. Current status: valid genus in CARABIDAE: TRECHINAE: TACHYINI: TACHYINA.

Type species: Micratopus fusciceps Casey, 1914 (= Blemus aenescens J.L. LeConte, 1848), by monotypy.

Family-group name: MICRATOPINI Casey, 1914. Stem: Micratop-.

Microcephalus Dejean, 1828: 4, 198. Current status: valid genus in CARABIDAE: HARPALINAE: PTEROSTICHINI: MICROCEPHALINA.

Type species: Microcephalus depressicollis Dejean, 1828, by monotypy.

Family-group name: MICROCEPHALIDES Tschitschérine, 1898. Stem: Microcephal-.

Microceroxenus Kistner, 1970a: 10. Current status: valid genus in STAPHYLINIDAE: ALEOCHARINAE: ATHETINI: MICROCEROXENINA.

Type species: Microceroxenus alzadae Kistner, 1970, by original designation.

Family-group name: MICROCEROXENINA Kistner, 1970. Stem: Microceroxen-.

Microcerus Schönherr, 1833: 8. Current status: valid genus in BRENTIDAE: MICROCERINAE.

Type species: Curculio retusus Fabricius, 1781, by original designation.

Family-group name: MICROCÉRIDES Lacordaire, 1863. Stem: Microcer-.

Microchaetes Hope, 1834b: 12. Current status: valid genus in BYRRHIDAE: SYNCALYPTINAE: MICROCHAETINI.

Type species: Microchaetes sphaericus Hope, 1834, by monotypy.

Family-group name: MICROCHAETINI Paulus, 1973. Stem: Microchaet-. Note: junior homonym of MICROCHAETINI Beddard, 1895 (type genus Microchaetus Rapp, 1849) in Oligochaeta.

Microcheila Brullé, 1834: 330, 336. Current status: valid genus in CARABIDAE: HARPALINAE: MICROCHEILINI.

Type species: Microcheila picea Brullé, 1834, by monotypy.

Family-group name: MICROCHILITAE Jeannel, 1948. Stem: Microcheil-.

Microcrania Burmeister, 1855: 11, 75. Current status: junior synonym of Barybas Blanchard, 1850 in SCARABAEIDAE: MELOLONTHINAE: MACRODACTYLINI.

Type species: Philochloenia compacta Erichson, 1847, by subsequent designation (Evans 2003: 230).

Family-group name: MICROCRANIADAE Burmeister, 1855. Stem: Microcrani-.

Microcymatura Breuning, 1951: 386. Current status: valid genus in CERAMBYCIDAE: LAMIINAE: MICROCYMATURINI.

Type species: Microcymatura antennalis Breuning, 1951, by original designation.

Family-group name: MICROCYMATURINI Breuning & Téocchi, 1985. Stem: Microcymatur-.

Microglotta Kraatz, 1862b: 300. Current status: junior synonym of Haploglossa Kraatz, 1856 in STAPHYLINIDAE: ALEOCHARINAE: OXYPODINI: MICROGLOTTINA. Note: unnecessary new replacement name for Haploglossa Kraatz, 1856.

Type species: fixed by Smetana (2004: 31) according to Article 70.3.2 (ICZN 1999a) as Aleochara villosula Stephens, 1832, misidentified as Aleochara pulla Gravenhorst, 1802 (sensu Gyllenhal, 1827), in the subsequent designation by C.G. Thomson (1859: 33) (type fixation for Haploglossa Kraatz).

Family-group name: MICROGLOTTAE Fenyes, 1919. Stem: Microglott-.

Microhoria Chevrolat, 1877: 168. Current status: valid genus in ANTHICIDAE: ANTHICINAE: MICROHORIINI. Note: two original spellings were used, including Michrohoria (p. 168); Microhoria was placed on the Official List of Generic Names in Zoology (ICZN 2016).

Type species: Anthicus oedipus Chevrolat, 1860, by subsequent designation (Bonadona 1952: 234; see ICZN 2016). Note: name placed on the Official List of Specific Names in Zoology (ICZN 2016).

Family-group name: MICROHORINI Bonadona, 1974. Stem: Microhori-.

Microjulistus Reitter, 1889c: 111. Current status: valid genus in RHADALIDAE: RHADALINAE.

Type species: Microjulistus fulvus Reitter, 1889, by monotypy.

Family-group name: MICROJULISTINI Majer, 1987. Stem: Microjulist-.

Micromalthus J.L. LeConte, 1878b: 613. Current status: valid genus in MICROMALTHIDAE.

Type species: Micromalthus debilis J.L. LeConte, 1878, by monotypy.

Family-group name: MICROMALTHIDAE Barber, 1913. Stem: Micromalth-.

Micropeplus Latreille, 1809a: 377. Current status: valid genus in STAPHYLINIDAE: MICROPEPLINAE.

Type species: Staphylinus porcatus Paykull, 1789, by monotypy.

Family-group name: MICROPEPLIDA Leach, 1815. Stem: Micropepl-.

Micropsalis Burmeister, 1861: 314. Current status: invalid senior synonym of Apterocaulus Fairmaire, 1864 in CERAMBYCIDAE: PRIONINAE: PRIONINI. Note: junior homonym of Micropsalis H. Meyer 1859 [Crustacea].

Type species: Micropsalis heterogama Burmeister, 1861, by monotypy.

Family-group name: MICROPSALINI Gounelle, 1911. Stem: Micropsalid-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

Microrhagus Dejean, 1833a: 85. Current status: valid genus in EUCNEMIDAE: MELASINAE: DIRHAGINI.

Type species: Elater pygmaeus Fabricius, 1792, by subsequent designation (Westwood 1838c: 25).

Family-group name: MICRORHAGINAE Fleutiaux, 1919. Stem: Microrhag-.

Microrhopala Chevrolat in Dejean, 1836a: 365. Current status: valid genus in CHRYSOMELIDAE: CASSIDINAE: CHALEPINI.

Type species: Hispa vittata Fabricius, 1798, by subsequent designation (Baly 1864d: 268).

Family-group name: MICRORHOPALIDES G.H. Horn, 1883. Stem: Microrhopal-. Note: nomen oblitum (see Bouchard and Bousquet 2020a: 108).

Microsilpha Broun, 1886: 889. Current status: valid genus in STAPHYLINIDAE: MICROSILPHINAE.

Type species: Microsilpha litorea Broun, 1886, by monotypy.

Family-group name: MICROSILPHINAE Crowson, 1950. Stem: Microsilph-.

Micrositus Mulsant & Rey, 1854: 131, 148. Current status: valid genus in TENEBRIONIDAE: BLAPTINAE: DENDARINI: DENDARINA.

Type species: Micrositus orbicularis Mulsant & Rey, 1854, by subsequent designation (Gebien 1938b: 415).

Family-group name: MICROSITATES Mulsant & Rey, 1854. Stem: Microsit-.

Microsporus Kolenati, 1846: 64. Current status: junior synonym of Sphaerius Waltl, 1838 in SPHAERIUSIDAE. Note: name placed on the Official List of Generic Names in Zoology (ICZN 1985d).

Type species: Microsporus obsidianus Kolenati, 1846 (= Sphaerius acaroides Waltl, 1838), by monotypy (see ICZN 1985d). Note: Microsporus obsidianus Kolenati, 1846 was placed on the Official List of Specific Names in Zoology (ICZN 1985d).

Family-group name: MICROSPORIDAE Crotch, 1873. Stem: Microspor-.

Microstylus Schönherr, 1847: 15. Current status: valid genus in CURCULIONIDAE: CURCULIONINAE: MICROSTYLINI.

Type species: Microstylus rufus Schönherr, 1847, by original designation. Note: combined description of a new nominal genus and a single new nominal species (ICZN 1999a, Article 12.2.6).

Family-group name: MICROSTYLIDES Lacordaire, 1865. Stem: Microstyl-. Note: this family- group name was treated as unavailable by Bouchard et al. (2011: 582) but has since been latinized, accepted as valid, and as dating from its first publication in vernacular form (e.g., Caldara et al. 2014: 611, Alonso-Zarazaga et al. 2017: 204), MICROSTYLINI Lacordaire, 1865 is therefore treated as available (ICZN 1999a, Article 11.7.2).

Microtrachelizus Senna, 1893: 315. Current status: valid genus in BRENTIDAE: BRENTINAE: TRACHELIZINI: MICROTRACHELIZINA.

Type species: Trachelizus lyratus Perroud & Montrouzier, 1865, by subsequent designation (R. Lucas 1920: 417).

Family-group name: MICROTRACHELIZINA Zimmerman, 1994. Stem: Microtracheliz-.

Microvalgus Kraatz, 1883: 374. Current status: valid genus in SCARABAEIDAE: CETONIINAE: VALGINI: MICROVALGINA.

Type species: Valgus lapeyrouse Gory & Percheron, 1833, by subsequent designation (W.F. Kirby 1885: 71).

Family-group name: MICROVALGINAE Kolbe, 1904. Stem: Microvalg-.

Microweisea Cockerell, 1903: 38. Current status: valid genus in COCCINELLIDAE: MICROWEISEINAE: MICROWEISEINI. Note: new replacement name for Smilia Weise, 1892.

Type species: Smilia felschei Weise, 1892, by monotypy (type fixation for Smilia Weise).

Family-group name: MICROWEISEINI Leng, 1920. Stem: Microweise-.

Microxylobius Chevrolat, 1835a: 98. Current status: valid genus in CURCULIONIDAE: COSSONINAE: MICROXYLOBIINI.

Type species: Microxylobius westwoodii Chevrolat, 1835, by monotypy.

Family-group name: MICROXYLOBIINI Voss, 1972. Stem: Microxylobi-.

Microzaena Fairmaire, 1901: 106. Current status: valid genus in CARABIDAE: PAUSSINAE: MICROZAENINI.

Type species: Microzaena madecassa Fairmaire, 1901, by monotypy.

Family-group name: MICROZAENINI Deuve, 2020: 203. Stem: Microzaen-.

Microzoum Dejean, 1834: 193. Current status: junior synonym of Melanimon Steven, 1828 in TENEBRIONIDAE: TENEBRIONINAE: MELANIMONINI.

Type species: Opatrum tibiale Fabricius, 1781, by monotypy.

Family-group name: MICROZOUMATES Mulsant, 1854. Stem: Microzo-.

Migadops G.R. Waterhouse, 1842a: 136. Current status: valid genus in CARABIDAE: MIGADOPINAE: MIGADOPINI: MIGADOPINA.

Type species: Migadops virescens G.R. Waterhouse, 1842 (= Antarctia lata Guérin-Méneville, 1841), by subsequent designation (Jeannel 1938: 27).

Family-group name: MIGADOPIDAE Chaudoir, 1861. Stem: Migadop-.

Mimanomma Wasmann, 1912: 478. Current status: valid genus in STAPHYLINIDAE: ALEOCHARINAE: MIMANOMMATINI: MIMANOMMATINA.

Type species: Mimanomma spectrum Wasmann, 1912, by monotypy.

Family-group name: MIMANOMMATINAE Wasmann, 1912. Stem: Mimanommat-.

†Mimaphodius Nikolajev, 2007a: 47. Current status: valid genus in HYBOSORIDAE: †MIMAPHODIINAE. Note: Lower Cretaceous (Russia).

Type species: †Mimaphodius pusillus Nikolajev, 2007, by original designation.

Family-group name: †MIMAPHODIINAE Nikolajev, 2007. Stem: Mimaphodi-.

Mimastra Baly, 1865a: 253. Current status: valid genus in CHRYSOMELIDAE: GALERUCINAE: LUPERINI.

Type species: Mimastra arcuata Baly, 1865, by original designation.

Family-group name: MIMASTRITES Chapuis, 1875. Stem: Mimastr-.

Mimeciton Wasmann, 1893a: 97. Current status: valid genus in STAPHYLINIDAE: ALEOCHARINAE: MIMECITINI: MIMECITINA.

Type species: Mimeciton pulex Wasmann, 1893, by monotypy.

Family-group name: MIMECITONINI Wasmann, 1909. Stem: Mimecit-.

Mimela W. Kirby, 1823a: 101. Current status: valid genus in SCARABAEIDAE: RUTELINAE: ANOMALINI: ANOMALINA.

Type species: Mimela chinensis W. Kirby, 1823, by monotypy.

Family-group name: MIMELIDAE Theobald, 1882. Stem: Mimel-.

Mimicoclytrina Bellamy, 2003: 26. Current status: valid genus in BUPRESTIDAE: POLYCESTINAE: TYNDARIDINI: MIMICOCLYTRININA. Note: new replacement name for Acherusia Laporte & Gory, 1838.

Type species: Acherusia childreni Laporte & Gory, 1838, by monotypy (type fixation for Acherusia Laporte & Gory).

Family-group name: MIMICOCLYTRININA Bellamy, 2003. Stem: Mimicoclytrin-.

Mimolibnetis Pic, 1936: 33 [suggested]. Current status: valid genus in LYCIDAE: DEXORINAE: MIMOLIBNETINI: MIMOLIBNETINA. Note: this name was first proposed by Pic (1936: 33) without a type species designation, a mandatory requirement for genus-group names proposed after 1930 (ICZN 1999a, Article 13.3) and it is therefore unavailable from that date; Kazantsev (2000: 12) designated Mimolibnetis apicalis Pic, 1936 as type species of the genus but since he did not indicate he was proposing a new taxon, a mandatory requirement for names published after 1999 (ICZN 1999a, Article 16.1), Mimolibnetis is also unavailable from that date; in order to promote stability, we believe that a request should be submitted to the Commission to accept Mimolibnetis Pic, 1936 as available, as well as the family-group name formed from it.

Type species [suggested]: Mimolibnetis apicalis Pic, 1936, by original designation.

Family-group name: MIMOLIBNETINAE Kazantsev, 2013a: 28. Stem: Mimolibnet-. Note: family- group name used as valid in recent classifications but it is not an available name since it was not formed from an available genus-group name (ICZN 1999a, Article 13.2.1; see comments above in the entry for Mimolibnetis Kazantsev, 2000).

Mimonilla Wasmann, 1913: 380. Current status: valid genus in STAPHYLINIDAE: ALEOCHARINAE: MIMECITINI: MIMONILLINA.

Type species: Mimonilla ecitonis Wasmann, 1913, by monotypy.

Family-group name: MIMONILLAE Bernhauer & Scheerpeltz, 1926. Stem: Mimonill-.

†Mimoplatycis Kazantsev, 2013b: 288. Current status: valid genus in CANTHARIDAE: MALTHININAE: †MIMOPLATYCINI. Note: Eocene (Baltic amber)

Type species: †Mimoplatycis notha Kazantsev, 2013, by original designation.

Family-group name: †MIMOPLATYCINI Kazantsev, 2013b: 288. Stem: Mimoplatyc-.

Mimoscymnus Gordon, 1994: 692, 712. Current status: valid genus in COCCINELLIDAE: COCCINELLINAE: HYPERASPIDINI: MIMOSCYMNINA.

Type species: Mimoscymnus aquilerai Gordon, 1994, by original designation.

Family-group name: MIMOSCYMNINA Vandenberg & Hanson, 2019: 262. Stem: Mimoscymn-.

Miniduliticola Kazantsev, 2003: 16. Current status: valid genus in LYCIDAE: LYROPAEINAE: MINIDULITICOLINI.

Type species: Miniduliticola nelsoni Kazantsev, 2003, by original designation.

Family-group name: MINIDULITICOLINI Kazantsev, 2003. Stem: Miniduliticol-.

Minulus Eggers, 1912: 206. Current status: junior synonym of Cnemonyx Eichhoff, 1868 in CURCULIONIDAE: SCOLYTINAE: SCOLYTINI. Note: this name is not a junior homonym of Minulus Rafinesque, 1820 [Pisces] since Neave (1940: 183) rejected the spelling of Rafinesque’s name in favor of the alternative original spelling Minnilus (see Alonso- Zarazaga and Lyal 2009: 24).

Type species: Minulus barbatus Eggers, 1912 (= Cnemonyx galeritus Eichhoff, 1868), by monotypy.

Family-group name: MINULINI Reitter, 1913. Stem: Minul-.

Minurus G.R. Waterhouse, 1842b: 68. Current status: valid genus in ATTELABIDAE: SAYREVILLEINAE: MINURINI.

Type species: Minurus testaceus G.R. Waterhouse, 1842, by monotypy.

Family-group name: MINURINI Legalov, 2003. Stem: Minur-.

Minyomerus G.H. Horn in J.L. LeConte and G.H. Horn, 1876: 17. Current status: valid genus in CURCULIONIDAE: ENTIMINAE: POLYDRUSITAE: TANYMECINI: TANYMECINA.

Type species: Minyomerus innocuus G.H. Horn, 1876, by subsequent designation (Pierce 1913: 400).

Family-group name: MINYOMERI G.H. Horn, 1876. Stem: Minyomer-.

Minyops Schönherr, 1823: 1143. Current status: valid genus in CURCULIONIDAE: MOLYTINAE: MOLYTINI: PLINTHINA.

Type species: Curculio carinatus Linnaeus, 1767, by original designation.

Family-group name: MINYOPIDAE Marseul, 1863. Stem: Minyop-.

†Miocenocylas Legalov, 2018a: 32. Current status: valid genus in BRENTIDAE: BRENTINAE: CYLADINI: †MIOCENOCYLADINA. Note: mid-Miocene (Germany).

Type species: †Miocenocylas heeri Legalov, 2018, by original designation.

Family-group name: †MIOCENOCYLADINI Legalov, 2018a: 35. Stem: Miocenocylad-.

Miroclaviger Wasmann, 1893a: 108, 111. Current status: valid genus in STAPHYLINIDAE: PSELAPHINAE: CLAVIGERITAE: CLAVIGERINI.

Type species: Miroclaviger cervicornis Wasmann, 1893, by monotypy.

Family-group name: MIROCLAVIGERINI Jeannel, 1949. Stem: Miroclaviger-.

Mirus Saulcy, 1877: 169. Current status: invalid senior synonym of Imirus Reitter, 1885 in STAPHYLINIDAE: PSELAPHINAE: GONIACERITAE: IMIRINI. Note: junior homonym of Mirus Albers, 1850 [Mollusca].

Type species: Mirus permirus Saulcy, 1877, by monotypy.

Family-group name: MIRINI Raffray, 1917. Stem: Mir-. Note: junior homonym of MIRIDAE Hahn, 1833 (type genus Miris Fabricius, 1794) in Hemiptera; family-group name permanently invalid (ICZN 1999a, Article 39).

Miscelus Klug, 1834: 82. Current status: valid genus in CARABIDAE: HARPALINAE: LEBIINI: PERICALINA.

Type species: Miscelus javanus Klug, 1834, by monotypy.

Family-group name: MISCELINI Sloane, 1907. Stem: Miscel-.

Mischocephalus Chaudoir, 1863a: 318. Current status: valid genus in CARABIDAE: HARPALINAE: ZUPHIINI: MISCHOCEPHALINA.

Type species: Mischocephalus spinicollis Chaudoir, 1863, by monotypy.

Family-group name: MISCHOCEPHALINA Mateu, 1992. Stem: Mischocephal-.

Misolampidius Solsky, 1876: 292. Current status: valid genus in TENEBRIONIDAE: STENOCHIINAE: CNODALONINI.

Type species: Misolampidius tentyrioides Solskly, 1876, by monotypy.

Family-group name: MISOLAMPIDIINI Reitter, 1917. Stem: Misolampidi-.

Misolampus Latreille, 1806: 160. Current status: valid genus in TENEBRIONIDAE: STENOCHIINAE: CNODALONINI.

Type species: Misolampus hoffmanssegii Latreille, 1806 (= Pimelia gibbula Herbst, 1799), by monotypy.

Family-group name: MISOLAMPIDES Lacordaire, 1859. Stem: Misolamp-.

Mitracephala Raffray, 1890a: 104, 107. Current status: invalid senior synonym of Mitrametopus Raffray, 1911 in STAPHYLINIDAE: PSELAPHINAE: EUPLECTITAE: EUPLECTINI. Note: junior homonym of Mitracephala J. Thomson, 1859 [Coleoptera: SCARABAEIDAE].

Type species: Mitracephala longipennis Raffray, 1890, by monotypy. Note: combined description of a new nominal genus and a single new nominal species (ICZN 1999a, Article 12.2.6).

Family-group name: MITRACEPHALINI Park, 1951. Stem: Mitracephal-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

Mitraelabrus Solier, 1851: 259. Current status: valid genus in ANTHICIDAE: EURYGENIINAE: MITRAELABRINI.

Type species: Mitraelabrus obscurus Solier, 1851, by subsequent designation (M. Abdullah 1969: 351).

Family-group name: MITRAELABRINI M. Abdullah & A. Abdullah, 1968. Stem: Mitraelabr-.

Mitrametopus Raffray, 1911: 50. Current status: valid genus in STAPHYLINIDAE: PSELAPHINAE: EUPLECTITAE: EUPLECTINI. Note: new replacement name for Mitracephala Raffray, 1890.

Type species: Mitracephala longipennis Raffray, 1890, by monotypy (type fixation for Mitracephala Raffray).

Family-group name: MITRAMETOPINA Park, 1952. Stem: Mitrametop-.

Mitrorhinus Kaup, 1871: 79. Current status: valid genus in PASSALIDAE: PASSALINAE: PASSALINI.

Type species: Mitrorhinus punctifrons Kaup, 1871 (= Passalus cayor Percheron, 1834), by monotypy.

Family-group name: MITRORHINAE Kuwert, 1891. Stem: Mitrorhin-.

Mniophila Stephens, 1831b: 330. Current status: valid genus in CHRYSOMELIDAE: GALERUCINAE: ALTICINI.

Type species: Altica muscorum J.D.W. Koch, 1803, by monotypy.

Family-group name: MNIOPHILITES Chapuis, 1875. Stem: Mniophil-.

Molops Bonelli, 1810: Tab. Syn. Current status: valid genus in CARABIDAE: HARPALINAE: PTEROSTICHINI: PTEROSTICHINA.

Type species: Carabus elatus Fabricius, 1801, by subsequent designation (Stephens 1832c: 248). Note: no species were originally included in this genus by Bonelli (1810); the first ones were the three species listed by Panzer (1813: 69); the species often listed as type species of this genus is Carabus terricola Herbst, 1784 sensu Fabricius, 1792 (= Scarites piceus Panzer, 1793) (e.g., Bousquet 2017b: 680) as designated by Hope (1838a: 71) but Stephens’ designation is earlier; both nominal species are currently included in the nominotypical subgenus of Molops.

Family-group name: MOLOPIDES Bonelli, 1810. Stem: Molop-.

Molorchus Fabricius, 1792b: 356. Current status: valid genus in CERAMBYCIDAE: CERAMBYCINAE: MOLORCHINI.

Type species: Necydalis umbellatarum Schreber, 1759, by subsequent designation (Curtis 1824: pl. 11).

Family-group name: MOLORCHIDAE Gistel, 1848. Stem: Molorch-.

Moluris Latreille, 1802b: 169. Current status: valid genus in TENEBRIONIDAE: PIMELIINAE: SEPIDIINI: MOLURINA.

Type species: Pimelia gibba Fabricius, 1787 (= Tenebrio gibbus Pallas, 1781), by monotypy.

Family-group name: MOLURITES Solier, 1834. Stem: Molur-.

Molytes Schönherr, 1823: 1143. Current status: junior synonym of Liparus G.-A. Olivier, 1807 in CURCULIONIDAE: MOLYTINAE: MOLYTINI: MOLYTINA.

Type species: Curculio germanus Linnaeus, 1758, by original designation.

Family-group name: MOLYTIDES Schönherr, 1823. Stem: Molyt-.

Mombasica Fairmaire, 1888b: 363. Current status: valid genus in CHRYSOMELIDAE: GALERUCINAE: GALERUCINI. Note: unnecessary new replacement name for Mombasa Fairmaire, 1884.

Type species: Mombasa armicollis Fairmaire, 1884, by monotypy (type fixation for Mombasa Fairmaire).

Family-group name: MOMBASICITES Laboissière, 1922. Stem: Mombasic-.

Monachulus Leng, 1918: 208. Current status: junior synonym of Lexiphanes Gistel, 1848 in CHRYSOMELIDAE: CRYPTOCEPHALINAE: CRYPTOCEPHALINI: MONACHULINA. Note: new replacement name for Monachus Chevrolat, 1836.

Type species: Cryptocephalus saponatus Fabricius, 1801, by monotypy (type fixation for Monachus Chevrolat).

Family-group name: MONACHULINI Leng, 1920. Stem: Monachul-.

Monachus Chevrolat in Dejean, 1836a: 425. Current status: invalid senior synonym of Lexiphanes Gistel, 1848 in CHRYSOMELIDAE: CRYPTOCEPHALINAE: CRYPTOCEPHALINI: MONACHULINA. Note: junior homonym of Monachus Fleming, 1822 [Mammalia].

Type species: Cryptocephalus saponatus Fabricius, 1801, by monotypy.

Family-group name: MONACHITES Chapuis, 1874. Stem: Monach-. Note: senior homonym of MONACHINA J.E. Gray, 1869 (type genus Monachus Fleming, 1822) in Mammalia; the Coleoptera family-group name permanently invalid (ICZN 1999a, Article 39).

Monardita Bechyné, 1948: 534. Current status: valid genus in CHRYSOMELIDAE: CHRYSOMELINAE: CHRYSOMELINI.

Type species: Chrysomela opulenta Reiche, 1848 (= Sphaeratrix latifrons Gistel, 1848), by original designation.

Family-group name: MONARDITINI Bechyné, 1953. Stem: Monardit-.

Moneilema Say, 1824: 403. Current status: valid genus in CERAMBYCIDAE: LAMIINAE: ACANTHOCININI.

Type species: Moneilema annulata Say, 1824, by monotypy.

Family-group name: MONEILEMITAE J. Thomson, 1864. Stem: Moneilem-.

†Mongolobrenthorrhinus Gratshev & Legalov in Legalov, 2011c: 64. Current status: valid genus in CARIDAE: †BAISSORHYNCHINAE: †MONGOLOBRENTHORRHININI. Note: Lower Cretaceous (Mongolia).

Type species: †Mongolobrenthorrhinus pusillus Gratshev & Legalov, 2011, by original designation.

Family-group name: †MONGOLOBRENTHORRHININI Gratshev & Legalov in Legalov, 2011a: 64. Stem: Mongolobrenthorrhin-.

†Mongolocar Gratshev & Legalov, 2011: 69. Current status: valid genus in BRENTIDAE: ITHYCERINAE: †MONGOLOCARINI. Note: Lower Cretaceous (Mongolia).

Type species: †Mongolocar orcinus Gratshev & Legalov, 2011, by original designation.

Family-group name: †MONGOLOCARINAE Gratshev & Legalov, 2011: 69. Stem: Mongolocar-.

Monochamus Dejean, 1821: 106. Current status: valid genus in CERAMBYCIDAE: LAMIINAE: LAMIINI.

Type species: Cerambyx sutor Linnaeus, 1758, by subsequent designation (Curtis 1828: pl. 219).

Family-group name: MONOHAMMIDAE Gistel, 1848. Stem: Monocham-.

Monochirus Chapuis, 1875: 330. Current status: invalid senior synonym of Hispellinus Weise, 1898 in CHRYSOMELIDAE: CASSIDINAE: HISPINI. Note: junior homonym of Monochirus Rafinesque, 1814 [Pisces].

Type species: Hispa callicantha H.W. Bates, 1866, by subsequent monotypy (Chapuis 1876: xxv).

Family-group name: MONOCHIRINI Weise, 1905. Stem: Monochir-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

Monocoryna Gorham, 1885b: 527. Current status: valid genus in COCCINELLIDAE: MONOCORYNINAE.

Type species: Monocoryna decempunctata Gorham, 1885, by monotypy.

Family-group name: MONOCORYNINI Miyatake, 1988. Stem: Monocoryn-. Note: junior homonym of MONOCORYNINAE Rees, 1956 (type genus Monocoryne Broch, 1910) in Cnidaria.

Monocrepidius Eschscholtz, 1829a: 31. Current status: valid genus in ELATERIDAE: AGRYPNINAE: OOPHORINI. Note: the First Reviser (ICZN 1999a, Article 24.2.2) for Monocrepidius Eschscholtz, 1829 versus Conoderus Eschscholtz, 1829 is Candèze (1859: 187).

Type species: Monocrepidius pallipes Eschscholtz, 1829, by subsequent designation (Hyslop 1921: 657).

Family-group name: MONOCRÉPIDIITES Candèze, 1859. Stem: Monocrepidi-.

Monodesmus Audinet-Serville, 1832: 160. Current status: valid genus in CERAMBYCIDAE: PRIONINAE: MEROSCELISINI.

Type species: Monodesmus callidioides Audinet-Serville, 1832, by monotypy.

Family-group name: MONODESMINAE Gahan, 1890. Stem: Monodesm-.

Monoedus G.H. Horn, 1882: 116. Current status: valid genus in ZOPHERIDAE: COLYDIINAE: ADIMERINI.

Type species: Monoedus guttatus G.H. Horn, 1882, by monotypy.

Family-group name: MONOEDIDAE Schaeffer, 1911. Stem: Monoed-.

Monolepta Chevrolat in Dejean, 1836a: 383. Current status: valid genus in CHRYSOMELIDAE: GALERUCINAE: LUPERINI.

Type species: Crioceris bioculata Fabricius, 1781, by subsequent designation (Chevrolat 1845: 5).

Family-group name: MONOLEPTITES Chapuis, 1875. Stem: Monolept-.

Monolobus Solier, 1849: 187. Current status: valid genus in CARABIDAE: MIGADOPINAE: MIGADOPINI: MIGADOPINA.

Type species: Monolobus testaceus Solier, 1849, by monotypy.

Family-group name: MONOLOBINAE Jeannel, 1938. Stem: Monolob-.

Monomacra Chevrolat in Dejean, 1836a: 389. Current status: valid genus in CHRYSOMELIDAE: GALERUCINAE: ALTICINI.

Type species [suggested]: Altica inermis Klug, 1829, by subsequent designation (Monrós and Bechyné 1956: 1133). Note: the valid type species fixation is that of Chevrolat (1845: 6), who selected Altica tibialis G.-A. Olivier, 1808, but that species is currently included in the genus Parchicola Bechyné & Springlovà de Bechyné, 1975; an application to the Commission is necessary to fix Altica inermis Klug, 1829 as the type species of Monomacra Chevrolat, 1836.

Family-group name: MONOMACRINA Bechyné & Špringlová de Bechyné, 1975. Stem: Monomacr-.

Monomma Klug, 1833: 94. Current status: valid genus in ZOPHERIDAE: ZOPHERINAE: MONOMMATINI.

Type species: Monomma irroratum Klug, 1833, by monotypy. Note: combined description of a new genus-group taxon and a single new species (ICZN 1999a, Article 12.2.6).

Family-group name: MONOMMITES Blanchard, 1845. Stem: Monommat-.

Mononychus Germar, 1823: 241. Current status: valid genus in CURCULIONIDAE: CEUTORHYNCHINAE: MONONYCHINI. Note: name placed on the Official List of Generic Names in Zoology (ICZN 1989b).

Type species: Curculio pseudacori Fabricius, 1792 (= Curculio punctumalbum Herbst, 1784), by subsequent designation (Schönherr 1825: 586; see ICZN 1989b). Note: Curculio punctumalbum Herbst, 1784 placed on the Official List of Specific Names in Zoology (ICZN 1989b).

Family-group name: MONONYCHI J.L. LeConte, 1876. Stem: Mononych-. Note: junior homonym of MONONYCHIDAE Fieber, 1851 (type genus Mononyx Laporte, 1832) in Hemiptera.

Monoplatus Chevrolat, 1846b: 333. Current status: junior synonym of Sphaeronychus Dejean, 1836 in CHRYSOMELIDAE: GALERUCINAE: ALTICINI.

Type species: Monoplatus nigripes Clark, 1860, by subsequent designation (Scherer 1962: 501). Note: no species were originally included in this genus by Chevrolat (1846b); the first ones were the 20 species listed by Clark (1860: 5–21, 278–280).

Family-group name: MONOPLATITES Chapuis, 1875. Stem: Monoplat-.

Monotoma Herbst, 1793: 22. Current status: valid genus in MONOTOMIDAE: MONOTOMINAE: MONOTOMINI. Note: nomen protectum (see Bouchard et al. 2011: 359).

Type species: Monotoma picipes Herbst, 1793, by subsequent designation (Stephens 1832c: 264).

Family-group name: MONOTOMITES Laporte, 1840. Stem: Monotom-.

†Montsecanomalus Soriano, Gratshev & Delclòs, 2006: 558. Current status: valid genus in BRENTIDAE: ITHYCERINAE: †MONTSECANOMALINI. Note: Lower Cretaceous (Spain).

Type species: †Montsecanomalus zherikhini Soriano, Gratshev & Delclòs, 2006, by original designation.

Family-group name: †MONTSECANOMALINI Legalov, 2016: 1466. Stem: Montsecanomal-.

†Montsecbelus Zherikhin & Gratshev, 1997: 626. Current status: valid genus in BELIDAE: †MONTSECBELINAE: †MONTSECBELINI. Note: Lower Cretaceous (Spain).

Type species: †Eobelus solutus Whalley & Jarzembowski, 1985, by original designation.

Family-group name: †MONTSECBELINAE Legalov, 2016: 1474. Stem: Montsecbeli-.

†Moravocoleus Kukalová, 1969: 141. Current status: valid genus in †TSHEKARDOCOLEIDAE. Note: lower Permian (Czech Republic).

Type species: †Moravocoleus permianus Kukalová, 1969, by original designation.

Family-group name: †MORAVOCOLEIDAE Kukalová-Peck & Beutel, 2012: 642. Stem: Moravocole-.

Mordella Linnaeus, 1758: 342, 420. Current status: valid genus in MORDELLIDAE: MORDELLINAE: MORDELLINI.

Type species: Mordella aculeata Linnaeus, 1758, by subsequent designation (Latreille 1810: 430).

Family-group name: MORDELLONAE Latreille, 1802. Stem: Mordell-.

Mordellistena A. Costa, 1854: 16. Current status: valid genus in MORDELLIDAE: MORDELLINAE: MORDELLISTENINI.

Type species: Mordellistena confinis A. Costa, 1854, by subsequent designation (Chûjô 1935: 80).

Family-group name: MORDELLISTENINI Ermisch, 1941. Stem: Mordellisten-.

Mordellochroa Emery, 1876: 80. Current status: valid genus in MORDELLIDAE: MORDELLINAE: MORDELLOCHROINI.

Type species: Mordella abdominalis Fabricius, 1775, by subsequent designation (Ermisch 1954: 133).

Family-group name: MORDELLOCHROINI Odnosum, 2010. Stem: Mordellochro-.

Morimonella Podaný, 1979: 43. Current status: valid genus in CERAMBYCIDAE: LAMIINAE: MORIMONELLINI.

Type species: Morimonella bednariki Podaný, 1979, by original designation.

Family-group name: MORIMONELLINI Lobanov, Danilevsky & Murzin, 1981. Stem: Morimonell-.

Morimopsis J. Thomson, 1857: 182. Current status: valid genus in CERAMBYCIDAE: LAMIINAE: MORIMOPSINI.

Type species: Morimopsis lacrymans J. Thomson, 1857, by monotypy.

Family-group name: MORIMOPSIDES Lacordaire, 1869. Stem: Morimops-.

Morimus Brullé, 1832: 258. Current status: valid genus in CERAMBYCIDAE: LAMIINAE: LAMIINI. Note: name proposed as Morinus, but the spelling was corrected to Morimus in the Errata issued in 1833 in a subsequent part of the same volume.

Type species: Lamia lugubris Fabricius, 1792 (= Cerambyx asper Sulzer, 1776), by subsequent designation (J. Thomson 1864: 77).

Family-group name: MORIMITAE J. Thomson, 1864. Stem: Morim-.

Moriomorpha Laporte, 1867: 37. Current status: valid genus in CARABIDAE: PSYDRINAE: MORIOMORPHINI: MORIOMORPHINA.

Type species: Moriomorpha victoriae Laporte, 1867, by subsequent designation (B.P. Moore 1963b: 281).

Family-group name: MORIOMORPHINI Sloane, 1890. Stem: Moriomorph-.

Morion Latreille, 1810: 159. Current status: valid genus in CARABIDAE: HARPALINAE: MORIONINI.

Type species: Harpalus monilicornis Latreille, 1805, by monotypy.

Family-group name: MORIONIENS Brullé, 1835. Stem: Morion-.

Mormolyce Hagenbach, 1825: 3. Current status: valid genus in CARABIDAE: HARPALINAE: LEBIINI: PERICALINA.

Type species: Mormolyce phyllodes Hagenbach, 1825, by monotypy.

Family-group name: MORMOLYCIDAE E. Desmarest, 1851. Stem: Mormolyc-.

Morostoma Candèze, 1879: 485. Current status: valid genus in ELATERIDAE: MOROSTOMINAE.

Type species: Morostoma palpale Candèze, 1879, by monotypy.

Family-group name: MOROSTOMINAE Dolin, 2000. Stem: Morostom- (ICZN 1999a, Article 29.4).

Morphozonitis Pic, 1922: 16. Current status: valid genus in MELOIDAE: ELETICINAE: ERTLIANINI.

Type species: Morphozonitis atripennis Pic, 1922, by monotypy.

Family-group name: MORPHOZONITINI Kaszab, 1969. Stem: Morphozonitid-.

Morychus Erichson, 1846: 468. Current status: valid genus in BYRRHIDAE: BYRRHINAE: MORYCHINI.

Type species: Byrrhus aeneus Fabricius, 1775, by subsequent designation (C.G. Thomson 1859: 74). Note: no species were originally included in this genus by Erichson (1846); the first ones were the three species listed by Erichson (1847a: 492, 493).

Family-group name: MORYCHINI El Moursy, 1961. Stem: Morych-.

Mosotalesus Kishii, 1977: 31, 32. Current status: valid genus in ELATERIDAE: DENDROMETRINAE: SELATOSOMINI: MOSOTALESINA.

Type species: Elater impressus Fabricius, 1792, by original designation.

Family-group name: MOSOTALESINA Schimmel, Tarnawski, Han & Platia, 2015: 31. Stem: Mosotales-.

Muonaja Özdikmen, 2008: 675. Current status: valid genus in EUCNEMIDAE: EUCNEMINAE: MUONAJINI. Note: new replacement name for Yanga Muona, 1993.

Type species: Yanga yonde Muona, 1993, by original designation (type fixation for Yanga Muona).

Family-group name: MUONAJINI Özdikmen, 2008. Stem: Muonaj-.

Murmidius Leach, 1821: 41. Current status: valid genus in MURMIDIIDAE.

Type species: Murmidius ferrugineus Leach, 1821 (= Hister ovalis Beck, 1817), by monotypy.

Family-group name: MURMIDIIDES Jacquelin du Val, 1858. Stem: Murmidi-. Note: nomen protectum (see Bouchard and Bousquet 2020a: 128).

†Myanmarops Legalov & Kirejtshuk in Legalov et al., 2020a: 2. Current status: valid genus in CHRYSOMELIDAE: BRUCHINAE: †MYANMAROPINI. Note: mid-Cretaceous (Myanmar).

Type species: †Myanmarops damgaardi Legalov & Kirejtshuk, 2020, by original designation.

Family-group name: †MYANMAROPINI Legalov & Kirejtshuk in Legalov et al., 2020a: 2. Stem: Myanmarop-.

Myas Sturm, 1826: 6, 171. Current status: valid genus in CARABIDAE: HARPALINAE: PTEROSTICHINI: PTEROSTICHINA.

Type species: Abax chalybaeus Palliardi, 1825, by monotypy.

Family-group name: MYADINA Jakobson, 1907. Stem: Myad-.

Mycetaea Stephens, 1829b: 8. Current status: valid genus in MYCETAEIDAE.

Type species: here fixed (under ICZN 1999a, Article 70.3.2) as Dermestes subterraneus Fabricius, 1801, misidentified as Dermestes fumatus Linnaeus, 1767 (sensu Marsham, 1802), in the original designation by monotypy in Stephens (1829b: 8).

Family-group name: MYCETÉIDES Jacquelin du Val, 1857. Stem: Mycetae-.

Mycetina Mulsant, 1846: 15. Current status: valid genus in ENDOMYCHIDAE: LYCOPERDININAE.

Type species: Chrysomela cruciata Schaller, 1783, by monotypy.

Family-group name: MYCETININA Lafer, 1992: 330. Stem: Mycetin-. Note: this name was overlooked in Bouchard et al. (2011) and Bouchard and Bousquet (2020a).

Mycetochara Guérin-Méneville, 1827: 346. Current status: valid genus in TENEBRIONIDAE: ALLECULINAE: ALLECULINI: MYCETOCHARINA. Note: new replacement name for Mycetophila Gyllenhal, 1810.

Type species [suggested]: Cistela flavipes Fabricius, 1792, by subsequent designation (C.G. Thomson 1859: 118; typification for Mycetophila Gyllenhal). Note: there are three earlier type species designations based on an originally included nominal species that threaten the stability of this genus: Guérin-Méneville (1827: 346), who selected Cistela humeralis Fabricius, 1787, Stephens (1832c: 375), who selected Cistela linearis Illiger, 1794 (= Cistela maura Fabricius, 1792) for Mycetocharis Gyllenhal, 1827 (another new replacement name for Mycetophila Gyllenhal, 1810), and Westwood (1838c: 32), who selected Cistela scapularis Illiger, 1805 (= Cistela humeralis Fabricius, 1787) for Mycetochara (as Mycetocharus); all these species are currently placed in the subgenus Ernocharis C.G. Thomson, 1859, not the nominotypical subgenus; an application to the Commission is necessary to fix Cistela flavipes Fabricius, 1792 as the type species of Mycetophila Gyllenhal, 1810, for which Mycetochara Guérin-Méneville, 1827 is a new replacement name.

Family-group name: MYCETOCHARISIDAE Gistel, 1848. Stem: Mycetochar-.

Mycetoma Dejean, 1834: 201. Current status: valid genus in TETRATOMIDAE: HALLOMENINAE.

Type species: Dryops suturalis Panzer, 1797, by monotypy.

Family-group name: MYCÉTOMIENS Mulsant, 1856. Stem: Mycetomat-.

Mycetophagus Fabricius, 1792b: 497. Current status: valid genus in MYCETOPHAGIDAE: MYCETOPHAGINAE: MYCETOPHAGINI.

Type species: Silpha quadrimaculata Schaller, 1783, by subsequent designation (Latreille 1810: 431).

Family-group name: MYCETOPHAGIDA Leach, 1815. Stem: Mycetophag-.

Mycetoporus Mannerheim, 1830: 62. Current status: valid genus in STAPHYLINIDAE: MYCETOPORINAE. Note: name placed on the Official List of Generic Names in Zoology (ICZN 1993d, as Mycetoporus Mannerheim, 1831).

Type species: Tachinus punctus Gravenhorst, 1806, by subsequent designation (C.G. Thomson 1859: 47; see ICZN 1993d). Note: name placed on the Official List of Specific Names in Zoology (ICZN 1993d).

Family-group name: MYCETOPORIDES C.G. Thomson, 1859. Stem: Mycetopor-.

Mychothenus Strohecker, 1953: 24. Current status: valid genus in ANAMORPHIDAE. Note: new replacement name for Mychophilus Frivaldsky, 1877.

Type species: Mychophilus minutus Frivaldsky, 1877, by monotypy (type fixation for Mychophilus Frivaldsky).

Family-group name: MYCHOTHENINAE Sasaji, 1978. Stem: Mychothen-.

Mycteis Pascoe, 1860a: 44. Current status: valid genus in ANTHRIBIDAE: ANTHRIBINAE: MYCTEINI.

Type species: Mycteis marginicollis Pascoe, 1860, by subsequent designation (Trýzna and Valentine 2011: 94).

Family-group name: MYCTEINI Morimoto, 1972. Stem: Mycte-.

Mycterus Clairville, 1798: 56, 124. Current status: valid genus in MYCTERIDAE: MYCTERINAE.

Type species: Mycterus griseus Clairville, 1798 (= Rhinomacer curculioides Fabricius, 1781), by monotypy.

Family-group name: MYCTERINA Perty, 1840. Stem: Mycter-.

Mylabris Fabricius, 1775: 261. Current status: valid genus in MELOIDAE: MELOINAE: MYLABRINI. Note: the senior homonym Mylabris Geoffroy (1762: 163) was suppressed for both Principles of Priority and Homonymy and placed on the Official Index of Generic Names in Zoology while Mylabris Fabricius, 1775 was placed on the Official List of Generic Names in Zoology (ICZN 1995b).

Type species: Meloe cichorii Linnaeus, 1758, by subsequent designation (Latreille 1810: 430; see ICZN 1995b). Note: Meloe cichorii Linnaeus, 1758 was placed on the Official List of Specific Names in Zoology (ICZN 1995b); currently, that species is placed in the genus Hycleus Latreille, 1817 (Bologna 2020: 523); the Linnean syntypes consist of two species, one corresponding to Hycleus chicorii (Linnaeus, 1758) and the second to Mylabris variabilis (Pallas, 1781); a lectotype needs to be designated in order to clarify the application of the name Meloe cichorii Linnaeus, 1758 (see Löbl 2008a: 45).

Family-group name: MYLABRIA Rafinesque, 1815. Stem: Mylabr-.

Mylacorrhina Reitter, 1913a: 13. Current status: junior synonym of Altonomus Desbrochers des Loges, 1907 in CURCULIONIDAE: ENTIMINAE: CYPHICERITAE: CYPHICERINI: MYLACORRHININA.

Type species: Mylacus villosus Reitter, 1906, by subsequent designation (Yunakov 2013: 65).

Family-group name: MYLACORRHYNCHINA Reitter, 1913. Stem: Mylacorrhin-.

Mylacus Boheman in Schönherr, 1842b: 144. Current status: junior synonym of Omias Germar, 1817 in CURCULIONIDAE: ENTIMINAE: CYPHICERITAE: OMIINI.

Type species: Mylacus murinus Boheman, 1842, by original designation.

Family-group name: MYLACINI Reitter, 1913. Stem: Mylac-.

Mylassa Stål, 1857: 60. Current status: valid genus in CHRYSOMELIDAE: CRYPTOCEPHALINAE: MYLASSINI.

Type species: Mylassa fasciatipennis Stål, 1857 (= Pachybrachis crassicollis Blanchard, 1851), by subsequent designation (Jakobson 1924: 258). Note: Jakobson (1924: 257) selected Pachybrachis crassicollis Blanchard, 1851 as type species, which was not an originally included nominal species, but since he listed it in synonymy with Mylassa fasciatipennis Stål, 1857, one of the two nominal species originally included in Mylassa, he is deemed to have designated the latter species as type species (ICZN 1999a, Article 69.2.2).

Family-group name: MYLASSINI Gómez-Zurita & Cardoso, 2021: 605. Stem: Mylass-.

Myllaena Erichson, 1837: 382. Current status: valid genus in STAPHYLINIDAE: ALEOCHARINAE: MYLLAENINI.

Type species: Aleochara dubia Gravenhorst, 1806, by subsequent designation (Duponchel 1841b: 57, as Myllana).

Family-group name: MYLLAENINI Ganglbauer, 1895. Stem: Myllaen-.

Myllocerus Schönherr, 1823: 1144. Current status: valid genus in CURCULIONIDAE: ENTIMINAE: CYPHICERITAE: CYPHICERINI: MYLLOCERINA.

Type species: Curculio curvicornis Fabricius, 1792, by original designation.

Family-group name: MYLLOCERINI Pierce, 1913. Stem: Myllocer-.

Myloechus Latreille, 1806: 30. Current status: valid subgenus of Colon Herbst, 1797 in COLONIDAE.

Type species: Myloechus brunneus Latreille, 1806, by monotypy.

Family-group name: MYLOECHINA C.G. Thomson, 1859. Stem: Myloech-.

Myochrous Erichson, 1847c: 164. Current status: valid genus in CHRYSOMELIDAE: EUMOLPINAE: BROMIINI.

Type species: Myochrous immundus Erichson, 1847, by monotypy.

Family-group name: MYOCHROINAE Baly, 1865. Stem: Myochro-.

Myoderma Dejean, 1833b: 168. Current status: valid genus in SCARABAEIDAE: CETONIINAE: TRICHIINI. Note: this genus-group name is often listed as “Myodermum Burmeister & Schaum, 1840” in the literature but Dejean’s name has precedence (see Bousquet and Bouchard 2013a: 38).

Type species: Stripsypher sordidus Gory & Percheron, 1833 (= Trichius alutaceum Afzelius, 1817), by monotypy.

Family-group name: MYODERMINI Péringuey, 1907. Stem: Myodermat-.

Myodites Latreille, 1819b: 303. Current status: junior synonym of Ripiphorus Bosc, 1791 in RIPIPHORIDAE: RIPIPHORINAE: RIPIPHORINI. Note: new replacement name for Myodes Latreille, 1818; the original spelling was Myodite but Myodites, an incorrect subsequent spelling, is in prevailing usage, attributed to the publication of the original spelling, and therefore deemed to be the correct original spelling (ICZN 1999a, Article 33.3.1).

Type species: Myodes dorthesii Latreille, 1818 (= Ripiphorus subdipterus Fabricius, 1792), by monotypy (type fixation for Myodes Latreille).

Family-group name: MYODITINI Gerstaecker, 1855. Stem: Myodit-.

Myorhinus Schönherr, 1826: 213. Current status: junior synonym of Apsis Germar, 1820 in CURCULIONIDAE: ENTIMINAE: OTIORHYNCHITAE: MYORHININI.

Type species: Myorhinus albolineatus Guérin-Méneville, 1833, by subsequent monotypy (Guérin-Méneville 1833a: pl. 38).

Family-group name: MYORHINIDAE Marseul, 1863. Stem: Myorhin-. Note: senior homonym of MYORHININI Baranov, 1942 (type genus Myorhina Robineau-Desvoidy, 1830) in Diptera.

Myrabolia Reitter, 1876b: 55. Current status: valid genus in MYRABOLIIDAE.

Type species: Myrabolia haroldiana Reitter, 1876 (= Silvanus brevicornis Erichson, 1842), by monotypy.

Family-group name: MYRABOLIINAE Lawrence & Britton, 1991. Stem: Myraboli-.

Myrmacicelus Chevrolat, 1833: 358. Current status: junior synonym of Rhynolaccus Guérin- Méneville, 1831 in BRENTIDAE: APIONINAE: MYRMACICELITAE: MYRMACICELINI.

Type species: Myrmacicelus bistriatus Chevrolat, 1833 (= Rhynolaccus formicarius Guérin- Méneville, 1831), by monotypy. Note: the type species was first listed as Myrmacicelus formicarius Guérin-Méneville but it was changed in the errata and addenda [p. xcix] to Myrmacicelus bistriatus.

Family-group name: MYRMACICELINAE Zimmerman, 1994. Stem: Myrmacicel-.

Myrmechixenus Chevrolat, 1835b: 267. Current status: valid genus in TENEBRIONIDAE: DIAPERINAE: MYRMECHIXENINI.

Type species: Myrmechixenus subterraneus Chevrolat, 1835, by monotypy.

Family-group name: MYRMÉCHIXÉNITES Jacquelin du Val, 1858. Stem: Myrmechixen-.

Myrmedonia Erichson, 1837: 286. Current status: junior synonym of Drusilla Leach, 1819 in STAPHYLINIDAE: ALEOCHARINAE: LOMECHUSINI: MYRMEDONIINA.

Type species: Staphylinus canaliculatus Fabricius, 1787, by subsequent designation (Duponchel 1841b: 57).

Family-group name: MYRMEDONIIDES C.G. Thomson, 1867. Stem: Myrmedoni-.

Myrmetes Marseul, 1862: 499, 511. Current status: valid genus in HISTERIDAE: SAPRININAE: MYRMETINI.

Type species: Hister piceus Paykull, 1809 (= Myrmetes paykulli Kanaar, 1979), by monotypy.

Family-group name: MYRMETINI Portevin, 1929. Stem: Myrmet-.

Myrmex Sturm, 1826: 32, 172. Current status: valid genus in CURCULIONIDAE: CURCULIONINAE: OTIDOCEPHALINI.

Type species: Curculio myrmex Herbst, 1797, by monotypy.

Family-group name: MYRMECINAE Tanner, 1966. Stem: Myrmec-.

Myrtonymus Kuschel, 1990: 80. Current status: valid genus in CURCULIONIDAE: BRACHYCERINAE: MYRTONYMINI.

Type species: Myrtonymus zelandicus Kuschel, 1990, by original designation.

Family-group name: MYRTONYMINA Kuschel, 1990. Stem: Myrtonym-.

Mysteria J. Thomson, 1861: 278. Current status: valid genus in VESPERIDAE: ANOPLODERMATINAE: MYSTERIINI.

Type species: Mysteria cylindripennis J. Thomson, 1861, by monotypy.

Family-group name: MYSTERINAE Prosen, 1960. Stem: Mysteri-.

†Mysteriomorphus Alekseev & Ellenberger, 2019: 485. Current status: valid genus in †MYSTERIOMORPHIDAE. Note: Cretaceous (Myanmar).

Type species: †Mysteriomorphus pelevini Alekseev & Ellenberger, 2019, by original designation.

Family-group name: †MYSTERIOMORPHIDAE Alekseev & Ellenberger, 2019: 484. Stem: Mysteriomorph-.

Mystropomus Chaudoir, 1848: 107. Current status: valid genus in CARABIDAE: PAUSSINAE: MYSTROPOMINI.

Type species: Mystropomus subcostatus Chaudoir, 1848, by monotypy.

Family-group name: MYSTROPOMINI G.H. Horn, 1881. Stem: Mystropom-.

Mystrops Erichson, 1843a: 234. Current status: valid genus in NITIDULIDAE: NITIDULINAE: MYSTROPINI.

Type species: Mystrops debilis Erichson, 1843, by subsequent designation (Kirejtshuk and Couturier 2011: 372).

Family-group name: MYSTROPIDAE Murray, 1864. Stem: Mystrop-.

Mythodes J. Thomson, 1864: 139. Current status: valid genus in CERAMBYCIDAE: CERAMBYCINAE: MYTHODINI.

Type species: Mythodes plumosa J. Thomson, 1864, by original designation.

Family-group name: MYTHODIDES Lacordaire, 1868. Stem: Mythod-.

Myzia Mulsant, 1846: 129. Current status: valid genus in COCCINELLIDAE: COCCINELLINAE: COCCINELLINI. Note: as pointed out by Bouchard and Bousquet (2020a: 95–96) two original spellings were used, Mysia (p. 129) and Myzia (p. 277); subsequently, Mulsant (1850: 137) used the spelling Mysia and therefore acted as First Reviser (Article 24.2.4). Mysia is therefore the correct original spelling but this name is a junior homonym of Mysia Lamarck, 1818 [Mollusca]; Myzia, an incorrect original spelling, has been used as valid in the literature and is in prevailing usage; an application to the Commission is necessary to retain Myzia Mulsant, 1846 as a valid name.

Type species: Coccinella oblongoguttata Linnaeus, 1758, by monotypy.

Family-group name: MYSIINA Hatch, 1962: 175. Stem: Myzi-. Note: a family-group name based on this type genus was attributed to Mulsant, 1846 (as MYSIATES) and treated as unavailable by Bouchard et al. (2011: 376).

Nacerdes Dejean, 1834: 228. Current status: valid genus in OEDEMERIDAE: OEDEMERINAE: NACERDINI.

Type species: Necydalis notata Fabricius, 1792 (= Cantharis melanura Linnaeus, 1758), by subsequent designation (Arnett 1950: 222).

Family-group name: NACERDATES Mulsant, 1858. Stem: Nacerd-.

Nama Borovec & Meregalli, 2013: 502. Current status: valid genus in CURCULIONIDAE: ENTIMINAE incertae sedis.

Type species: Nama richtersveldiana Borovec & Meregalli, 2013, by original designation.

Family-group name: NAMAINI Borovec & Meregalli in Meregalli et al., 2021: 102. Stem: Nama-.

Nanoglossa Fauvel, 1868: 379. Current status: valid genus in STAPHYLINIDAE: ALEOCHARINAE: HOMALOTINI: BOLITOCHARINA. Note: new replacement name for Microglossa Fauvel, 1866.

Type species: Microglossa chilensis Fauvel, 1866, by subsequent designation (Fenyes 1919: 23) (type fixation for Microglossa Fauvel).

Family-group name: NANOGLOSSAE Fenyes, 1919. Stem: Nanogloss-.

†Nanophydes Arnoldi, 1977: 173. Current status: valid genus in CARIDAE: †BAISSORHYNCHINAE: †NANOPHYDINI. Note: Upper Jurassic (Kazakhstan).

Type species: †Nanophydes ovatus Arnoldi, 1977, by original designation.

Family-group name: †NANOPHYDINAE Arnoldi, 1977. Stem: Nanophyd-. Note: the name †NANOPHYDINI, as currently classified, is older than the subfamily †BAISSORHYNCHINAE; however, more research is needed to clarify the correct position of †BAISSORHYNCHINAE as well as the tribe therein before the priority problem can be addressed.

Nanophyes Schönherr, 1838: 780. Current status: valid genus in BRENTIDAE: NANOPHYINAE: NANOPHYINI. Note: new replacement name for Nanodes Schönherr, 1825, which was suppressed for the Principle of Priority and placed on the Official Index of Rejected and Invalid Names in Zoology while Nanophyes Schönherr, 1838 was placed on the Official List of Generic Names in Zoology (ICZN 1989a).

Type species: Curculio lythri Fabricius, 1787 (= Curculio marmoratus Goeze, 1777), by original designation (see ICZN 1989a) (type fixation for Nanodes Schönherr). Note: Curculio marmoratus Goeze, 1777, which has been published in a work that is not consistently binominal, was placed on the Official List of Specific Names in Zoology (ICZN 1989a).

Family-group name: NANOPHYEIDAE Gistel, 1848. Stem: Nanophy-.

Nanosella Motschulsky, 1869a: 172, 187. Current status: valid genus in PTILIIDAE: PTILIINAE: NANOSELLINI. Note: two original spellings were used, including Nonosella (p. 187); Neave (1940: 265, 345) acted as First Reviser (ICZN 1999a, Article 24.2.3) and selected Nanosella, the spelling currently in prevailing usage.

Type species: Nanosella fungi Motschulsky, 1869 (= Ptilium fungi J.L. LeConte, 1863), by monotypy.

Family-group name: NANOSELLINAE Barber, 1924. Stem: Nanosell-.

Narthecius J.L. LeConte, 1861: 95. Current status: valid genus in LAEMOPHLOEIDAE.

Type species: Narthecius grandiceps J.L. LeConte, 1863, by subsequent monotypy (J.L. LeConte 1863b: 70).

Family-group name: NARTHECIINI Casey, 1890. Stem: Nartheci-.

Nascio Laporte & Gory, 1838: 1. Current status: valid genus in BUPRESTIDAE: BUPRESTINAE: NASCIONINI. Note: name placed on the Official List of Generic Names in Zoology (ICZN 2002b).

Type species: Buprestis vetusta Boisduval, 1835, by monotypy (see ICZN 2002b). Note: name placed on the Official List of Specific Names in Zoology (ICZN 2002b).

Family-group name: NASCIONINA Holyński, 1988. Stem: Nascion-.

Nastus Schönherr, 1842b: 405. Current status: valid genus in CURCULIONIDAE: ENTIMINAE: OTIORHYNCHITAE: NASTINI.

Type species: Nastus goryi Boheman, 1842, by original designation.

Family-group name: NASTINI Reitter, 1913. Stem: Nast-.

Nasutiphilus Kistner, 1970d: 500. Current status: valid genus in STAPHYLINIDAE: ALEOCHARINAE: ATHETINI: NASUTIPHILINA.

Type species: Nasutiphilus niger Kistner, 1970, by original designation.

Family-group name: NASUTIPHILINA Kistner, 1970. Stem: Nasutiphil-.

Nasutitella Pasteels, 1967: 54. Current status: valid genus in STAPHYLINIDAE: ALEOCHARINAE: COROTOCINI: NASUTITELLINA.

Type species: Nasutitella puncticollis Pasteels, 1967, by original designation.

Family-group name: NASUTITELLINA Jacobson, Kistner & Pasteels, 1986. Stem: Nasutitell-.

Natalis Laporte, 1838: 40 Current status: valid genus in CLERIDAE: CLERINAE: NATALIINI: NATALIINA.

Type species: Natalis laplacei Laporte, 1838, by monotypy.

Family-group name: NATALIINI Bartlett, 2021: 750. Stem: Natali-. Note: the correct stem based on Natalis Laporte, a Latin adjective, is Natal- (M.A. Alonso-Zarazaga, pers. comm. to P.B., 2022); however, since using the correct stem would have led to the creation of a homonym with NATALIDAE Gray, 1866 (type genus Natalus Gray, 1838) in Chiroptera, we assume that Bartlett (2021: 750) used the stem Natalii- in order to avoid creating a homonymous family-group name (ICZN 1999a, Article 29.6).

Nathrius Brèthes, 1916: 76. Current status: valid genus in CERAMBYCIDAE: CERAMBYCINAE: PSEBIINI.

Type species: Nathrius porteri Brèthes, 1916 (= Leptidea brevipennis Mulsant, 1839), by monotypy.

Family-group name: NATHRIINI Arnett, 1962. Stem: Nathri-.

Natypleurus Newton & Thayer, 1992: 20. Current status: valid genus in STAPHYLINIDAE: PSELAPHINAE: GONIACERITAE: INIOCYPHINI: NATYPLEURINA. Note: new replacement name for Tanypleurus Raffray, 1890.

Type species: Tanypleurus malaianus Raffray, 1890, by monotypy (type fixation for Tanypleurus Raffray).

Family-group name: NATYPLEURINA Newton & Thayer, 1992. Stem: Natypleur-.

Naupactus Dejean, 1821: 94. Current status: valid genus in CURCULIONIDAE: ENTIMINAE: POLYDRUSITAE: NAUPACTINI: NAUPACTINA.

Type species: Curculio rivulosus G.-A. Olivier, 1790, by subsequent designation (Champion 1911: 232).

Family-group name: NAUPACTIDAE Gistel, 1848. Stem: Naupact-. Note: nomen protectum (see Bouchard et al. 2011: 611).

Navomorpha A. White, 1855: 334. Current status: valid genus in CERAMBYCIDAE: CERAMBYCINAE: COPTOMMATINI.

Type species: Callidium lineatum Fabricius, 1775, by subsequent designation (J. Thomson 1864: 38).

Family-group name: NAVOMORPHIDES Lacordaire, 1869. Stem: Navomorph-.

†Nebrenthorrhinus Legalov, 2003: 89. Current status: valid genus in NEMONYCHIDAE: †PALEOCARTINAE: †PALEOCARTINI: †NEBRENTHORRHININA. Note: Lower Cretaceous (Spain).

Type species: †Brenthorrhinoides lacasei Gratshev & Zherikhin, 2000, by original designation.

Family-group name: †NEBRENTHORRHININA Legalov, 2007. Stem: Nebrenthorrhin-.

Nebria Latreille, 1802b: 89. Current status: valid genus in CARABIDAE: NEBRIINAE: NEBRIINI.

Type species: Carabus brevicollis Fabricius, 1792, by subsequent designation (Latreille 1810: 426).

Family-group name: NEBRIIDAE Laporte, 1834. Stem: Nebri-.

Necrobia Latreille, 1797: 35. Current status: valid genus in CLERIDAE: KORYNETINAE. Note: this name is usually credited to “G.-A. Olivier, 1795” in the literature but it was first made available by Latreille in 1797 (see Bouchard and Bousquet 2020a: 93); sections 67 (part)–80 in G.-A. Olivier’s work where Necrobia appears (i.e., in section 76: 4) were not issued until 1800 (see Bousquet 2018: 140); name placed on the Official List of Generic Names in Zoology (ICZN 1961c, as Necrobia G.-A. Olivier, 1795).

Type species: Dermestes violaceus Linnaeus, 1758, by subsequent designation (Latreille 1810: 356; see ICZN 1961c). Note: no species were originally included in this genus by Latreille (1797); the first ones were the three species listed by G.-A. Olivier (1800: [76bis] 5, 6); name placed on the Official List of Specific Names in Zoology (ICZN 1961c).

Family-group name: NECROBIAEIDAE Gistel, 1848. Stem: Necrobi-.

Necrodes Leach, 1815a: 88. Current status: valid genus in STAPHYLINIDAE: SILPHINAE: SILPHINI.

Type species: Silpha littoralis Linnaeus, 1758, by subsequent designation (Curtis 1830: pl. 334).

Family-group name: NECRODISIDAE Gistel, 1848. Stem: Necrod-.

†Necronectes Ponomarenko, 1977: 22. Current status: junior synonym of Timarchopsis Brauer, J. Redtenbacher & Ganglbauer, 1889 in †COPTOCLAVIDAE: †TIMARCHOPSINAE. Note: junior homonym of Necronectes A. Milne-Edwards, 1881 [Crustacea].

Type species: †Necronectes aquaticus Ponomarenko, 1977, by original designation.

Family-group name: †NECRONECTINAE Ponomarenko, 1977. Stem: Necronect-. Note: family- group name permanently invalid (ICZN 1999a, Article 39).

Necrophilus Latreille, 1829a: 500. Current status: valid genus in AGYRTIDAE: NECROPHILINAE.

Type species: Silpha subterranea Dahl, 1807, by monotypy.

Family-group name: NECROPHILINAE Newton, 1997. Stem: Necrophil-.

Necydalis Linnaeus, 1758: 342, 421. Current status: valid genus in CERAMBYCIDAE: LEPTURINAE: NECYDALINI.

Type species: Necydalis major Linnaeus, 1758, by subsequent designation (Latreille 1829b: 120).

Family-group name: NECYDALIDES Latreille, 1825. Stem: Necydal-.

Necydalopsis Blanchard, 1851: 473. Current status: valid genus in CERAMBYCIDAE: CERAMBYCINAE: NECYDALOPSINI.

Type species: Necydalopsis trizonatus Blanchard, 1851, by monotypy.

Family-group name: NÉCYDALOPSIDES Lacordaire, 1868. Stem: Necydalops-.

Nedine J. Thomson, 1864: 27. Current status: valid genus in CERAMBYCIDAE: LAMIINAE: DESMIPHORINI.

Type species: Nedine longipes J. Thomson, 1864, by original designation.

Family-group name: NÉDINIDES Lacordaire, 1872. Stem: Nedin-.

Negastrius C.G. Thomson, 1859: 106. Current status: valid genus in ELATERIDAE: NEGASTRIINAE: NEGASTRIINI.

Type species: Elater pulchellus Linnaeus, 1761, by original designation.

Family-group name: NEGASTRIINAE Nakane & Kishii, 1956. Stem: Negastri-.

Neleides Harold in Gemminger and Harold, 1868c: 978. Current status: valid genus in PASSALIDAE: PASSALINAE: PASSALINI. Note: this name is attributed to Kaup (1869: 33) in the literature; however, it was first made available by Harold in 1868 when he included four nominal species under it.

Type species: Passalus affinis Percheron, 1834, by subsequent designation (Brauer 1870: 127).

Family-group name: NELIDINAE Kuwert, 1896. Stem: Neleid-.

Neleus Harold in Gemminger and Harold, 1868c: 977. Current status: junior synonym of Passalus Fabricius, 1792 in PASSALIDAE: PASSALINAE: PASSALINI. Note: junior homonym of Neleus Desbonne & Schramm, 1867 [Crustacea]; this name is attributed to Kaup (1869: 30) in the literature; however, it was first made available by Harold in 1868 when he included several nominal species under this genus.

Type species: Lucanus interruptus Linnaeus, 1758, by subsequent designation (Hincks and Dibb 1935: 38).

Family-group name: NELEINAE Kaup, 1869. Stem: Nele-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

Nemadus C.G. Thomson, 1867: 351. Current status: valid genus in LEIODIDAE: CHOLEVINAE: ANEMADINI: NEMADINA.

Type species: Catops colonoides Kraatz, 1851, by monotypy.

Family-group name: NEMADINAE Jeannel, 1936. Stem: Nemad-.

Nemaschema J. Thomson, 1861: 354. Current status: valid genus in CERAMBYCIDAE: LAMIINAE: ENICODINI.

Type species: Navomorpha sanguinicollis Chevrolat, 1857, by monotypy.

Family-group name: NEMASCHEMITAE J. Thomson, 1864. Stem: Nemaschemat-.

Nematidium Erichson, 1845a: 275. Current status: valid genus in ZOPHERIDAE: COLYDIINAE: NEMATIDIINI.

Type species: Colydium cylindricum Fabricius, 1801, by monotypy.

Family-group name: NEMATIDII G.H. Horn, 1878. Stem: Nematidi-.

Nematodes Guérin-Méneville, 1827: 498. Current status: valid genus in EUCNEMIDAE: MACRAULACINAE: NEMATODINI. Note: this genus is often credited to Berthold (1827: 335) (e.g., Muona 2007: 87) but it was first made available by Guérin-Méneville (1827: 498) (see Bouchard and Bousquet 2020a: 87).

Type species: Elater filum Fabricius, 1801, by original designation.

Family-group name: NEMATODINI Leiler, 1976. Stem: Nematod-.

Nematoplus J.L. LeConte, 1855: 270, 275. Current status: valid genus in STENOTRACHELIDAE: NEMATOPLINAE.

Type species: Nematoplus collaris J.L. LeConte, 1855, by monotypy.

Family-group name: NEMATOPLI J.L. LeConte, 1862. Stem: Nematopl-.

Nematoscelis Wollaston, 1868: 231. Current status: valid genus in STAPHYLINIDAE: ALEOCHARINAE: HYPOCYPHTINI.

Type species: Nematoscelis filipes Wollaston, 1868, by monotypy.

Family-group name: NEMATOSCELINI Fenyes, 1921. Stem: Nematoscelid-.

Nemognatha Illiger, 1807b: 333. Current status: valid genus in MELOIDAE: NEMOGNATHINAE: NEMOGNATHINI: NEMOGNATHINA. Note: name placed on the Official List of Generic Names in Zoology (ICZN 1999c).

Type species: Zonitis vittata Fabricius, 1801 (= Zonitis piazata Fabricius, 1798), by monotypy (see ICZN 1999c). Note: Zonitis vittata Fabricius, 1801 placed on the Official List of Specific Names in Zoology (ICZN 1999c).

Family-group name: NEMOGNATHITES Laporte, 1840. Stem: Nemognath-. Note: family-group name given precedence over HORIIDAE Latreille, 1802 (ICZN 1999c).

Nemonyx L. Redtenbacher, 1845: 96, 152. Current status: valid genus in NEMONYCHIDAE: NEMONYCHINAE. Note: name placed on the Official List of Generic Names in Zoology (ICZN 2005c).

Type species: Rhinomacer lepturoides Fabricius, 1801, by monotypy (see ICZN 2005c). Note: name placed on the Official List of Specific Names in Zoology (ICZN 2005c).

Family-group name: NEMONYCHIDAE Bedel, 1882. Stem: Nemonych-.

Nemotarsus J.L. LeConte, 1853e: 377. Current status: valid genus in CARABIDAE: HARPALINAE: LEBIINI: NEMOTARSINA.

Type species: Nemotarsus elegans J.L. LeConte, 1853, by monotypy.

Family-group name: NEMOTARSINAE H.W. Bates, 1883. Stem: Nemotars-.

Nemotragus Westwood, 1843b: 57. Current status: valid genus in CERAMBYCIDAE: LAMIINAE: AGAPANTHIINI.

Type species: Nemotragus helvolus Westwood, 1843, by monotypy.

Family-group name: NEMOTRAGITAE J. Thomson, 1864. Stem: Nemotrag-.

Nemozoma Latreille, 1804a: 157. Current status: valid genus in TROGOSSITIDAE: TROGOSSITINAE: TROGOSSITINI.

Type species: Dermestes elongatus Linnaeus, 1761, by subsequent monotypy (Latreille 1804c: 239).

Family-group name: NEMOSOMIDA Leach, 1815. Stem: Nemozomat-.

Neoceratopsis Jeannel, 1956: 50. Current status: valid genus in STAPHYLINIDAE: PSELAPHINAE: CLAVIGERITAE: CLAVIGERINI.

Type species: Neoceratopsis pauliani Jeannel, 1956, by original designation.

Family-group name: NEOCERATOPSINI Célis, 1970. Stem: Neoceratopse-.

Neocerus Wasmann, 1893a: 105, 111. Current status: valid genus in STAPHYLINIDAE: PSELAPHINAE: CLAVIGERITAE: CLAVIGERINI.

Type species: Neocerus compressicornis Wasmann, 1893, by monotypy.

Family-group name: NEOCERINI Jeannel, 1954. Stem: Neocer-. Note: junior homonym of NEOCERINI Salmon, 1941 (type genus Neocerus Salmon, 1941) in Collembola: TOMOCERIDAE; Salmon’s name is permanently invalid (based on a preoccupied type genus).

Neocharis Sharp, 1877c: 485. Current status: valid genus in EUCNEMIDAE: MELASINAE: NEOCHARINI.

Type species: Neocharis varia Sharp, 1877, by subsequent designation (Fleutiaux 1921: 294).

Family-group name: NEOCHARINI Muona, 1993. Stem: Neochar-.

Neochlamys Jacoby, 1882: 51. Current status: valid genus in CHRYSOMELIDAE: LAMPROSOMATINAE: NEOCHLAMYSINI.

Type species: Neochlamys strigicollis Jacoby, 1882, by original designation.

Family-group name: NEOCHLAMYSINI Monrós, 1959. Stem: Neochlamys-.

Neochthebius Orchymont, 1932: 43. Current status: junior synonym of Ochthebius Leach, 1815 in HYDRAENIDAE: OCHTHEBIINAE: OCHTHEBIINI.

Type species: Ochthebius vandykei Knisch, 1924, by monotypy.

Family-group name: NEOCHTHEBINA Perkins, 1997. Stem: Neochthebi-.

Neoclytus J. Thomson, 1860b: 67. Current status: valid genus in CERAMBYCIDAE: CERAMBYCINAE: CLYTINI. Note: new replacement name for Rhopalomerus Chevrolat, 1860; as pointed out by Bousquet et al. (2017: 26) the genus-group name Rhopalopachys Chevrolat (1860b: 457; type species Clytus morosus Chevrolat, 1860 (= Clytus irroratus LeConte, 1858) by original designation) is older than Neoclytus J. Thomson, 1860 and should be used as valid; an application to the Commission is necessary to retain Neoclytus J. Thomson, 1860 as the valid name.

Type species: Callidium erythrocephalum Fabricius, 1787 (= Callidium acuminatum Fabricius, 1775), by original designation.

Family-group name: NEOCLYTITAE J. Thomson, 1861. Stem: Neoclyt-.

Neocorus J. Thomson, 1864: 220. Current status: valid genus in CERAMBYCIDAE: CERAMBYCINAE: NEOCORINI.

Type species: Stenygra ibidionoides Audinet-Serville, 1834, by original designation.

Family-group name: NEOCORINI Martins, 2005. Stem: Neocor-.

Neocuris Saunders, 1868: 19. Current status: valid genus in BUPRESTIDAE: BUPRESTINAE: CURIDINI: NEOCURIDINA.

Type species: Stigmodera guerinii Hope, 1843, by subsequent designation (Rye 1873: 271). Note: the type species usually recognized for this genus is Anthaxia fortnumi Hope, 1846, by subsequent designation of Bellamy (1997c: 221) but Rye’s designation is earlier; both species are currently included in the genus Neocuris.

Family-group name: NEOCURIDINA Holyński, 1988. Stem: Neocurid-.

Neocyphus Bedel, 1883: 23. Current status: junior synonym of Cyrtomon Schönherr, 1823 in CURCULIONIDAE: ENTIMINAE: POLYDRUSITAE: NAUPACTINI: NAUPACTINA. Note unnecessary new replacement name for Cyphus Germar, 1823.

Type species: Curculio gibber Pallas, 1781, by subsequent designation (Schönherr 1826: 11) (type fixation for Cyphus Germar).

Family-group name: NEOCYPHINI Hustache, 1919. Stem: Neocyph-.

Neohydrocoptus Satô, 1972: 144. Current status: valid genus in NOTERIDAE: NOTERINAE: NOTERINI.

Type species: Hydrocoptus bivittis Motschulsky, 1859, by original designation.

Family-group name: NEOHYDROCOPTINI Zalat, Saleh, Angus & Kaschef, 2000. Stem: Neohydrocopt-.

Neoibidion Monné, 2012: 36. Current status: valid genus in CERAMBYCIDAE: CERAMBYCINAE: TROPIDINI: NEOIBIDIONINA. Note: new replacement name for Ibidion Audinet-Serville, 1834.

Type species: Ibidion comatum Audinet-Serville, 1834, by subsequent designation (J. Thomson 1864: 215) (type fixation for Ibidion Audinet-Serville).

Family-group name: NEOIBIDIONINI Monné, 2012: 35. Stem: Neoibidion-.

Neopachypterus Bouchard, Löbl & Merkl, 2007: 386. Current status: valid genus in TENEBRIONIDAE: BLAPTINAE: OPATRINI: NEOPACHYPTERINA. Note: new replacement name for Pachypterus P.H. Lucas, 1847.

Type species: Pachypterus mauritanicus P.H. Lucas, 1847, by monotypy (type fixation for Pachypterus P.H. Lucas).

Family-group name: NEOPACHYPTERINA Bouchard, Löbl & Merkl, 2007. Stem: Neopachypter-.

Neopedilus M. Abdullah, 1969: 358. Current status: junior synonym of Pedilus Fischer, 1820 in PYROCHROIDAE: PEDILINAE.

Type species: Pedilus parvicollis Fall, 1919, by original designation.

Family-group name: NEOPEDILINI M. Abdullah, 1969. Stem: Neopedil-.

Neopelatops Jeannel, 1936: 12. Current status: valid genus in LEIODIDAE: CAMIARINAE: NEOPELATOPINI.

Type species: Neopelatops edwardsi Jeannel, 1936, by monotypy.

Family-group name: NEOPELATOPINI Jeannel, 1962. Stem: Neopelatop-.

Neophonus Fauvel, 1905: 99. Current status: valid genus in STAPHYLINIDAE: NEOPHONINAE.

Type species: Neophonus bruchi Fauvel, 1905, by monotypy.

Family-group name: NEOPHONI Fauvel, 1905. Stem: Neophon-.

Neopsectropus Kaszab, 1941: 30. Current status: valid genus in TENEBRIONIDAE: TENEBRIONINAE: ULOMINI.

Type species: Neopsectropus gebieni Kaszab, 1941, by original designation.

Family-group name: NEOPSECTROPINAE Kaszab, 1941. Stem: Neopsectropod-.

Neorthopleura Barr, 1976: 2. Current status: valid genus in CLERIDAE: KORYNETINAE.

Type species: Orthopleura texana Bland, 1863, by original designation.

Family-group name: NEORTHOPLEURINAE Opitz, 2009. Stem: Neorthopleur-.

Neosharpia Hoffmann, 1956: 247. Current status: valid genus in CURCULIONIDAE: BARIDINAE: MADARINI: NEOSHARPIINA.

Type species: Neosharpia artemisiae Hoffmann, 1956, by original designation.

Family-group name: NEOSHARPINI Hoffmann, 1956. Stem: Neosharpi-.

Neostenus Pascoe, 1857: 91. Current status: valid genus in CERAMBYCIDAE: CERAMBYCINAE: URACANTHINI.

Type species: Neostenus saundersii Pascoe, 1857, by monotypy.

Family-group name: NÉOSTÉNIDES Lacordaire, 1868. Stem: Neosten-.

Neotropospeonella Pace, 1985: 542. Current status: junior synonym of Oryotus L. Miller, 1856 in LEIODIDAE: CHOLEVINAE: LEPTODIRINI: BATHYSCIINA.

Type species: Neotropospeonella decui Pace, 1985 (= Oryotus ravasinii J. Müller, 1922), by original designation.

Family-group name: NEOTROPOSPEONELLINA Perkovsky, 1997. Stem: Neotropospeonell-.

Neotyphlus Coiffait, 1959: 387. Current status: valid genus in STAPHYLINIDAE: LEPTOTYPHLINAE: NEOTYPHLINI.

Type species: Neotyphlus californicus Coiffait, 1959, by original designation.

Family-group name: NEOTYPHLINI Coiffait, 1963. Stem: Neotyphl-.

Nephanes C.G. Thomson, 1859: 62. Current status: valid genus in PTILIIDAE: PTILIINAE: ACROTRICHINI. Note: name placed on the Official List of Generic Names in Zoology (ICZN 1985b).

Type species: Trichopteryx abbreviatella Heer, 1841 (= Trichopteryx titan Newman, 1834), by original designation (see ICZN 1985b). Note: Trichopteryx titan Newman, 1834 was placed on the Official List of Specific Names in Zoology (ICZN 1985b).

Family-group name: NEPHANINI Portevin, 1929. Stem: Nephan-.

Nephodes Blanchard, 1845a: 34. Current status: invalid senior synonym of Nephodinus Gebien, 1943 in TENEBRIONIDAE: TENEBRIONINAE: HELOPINI: HELOPINA. Note: junior homonym of Nephodes Schönherr, 1840 [Coleoptera: CURCULIONIDAE].

Type species: Nephodes villiger Rosenhauer, 1856, by subsequent monotypy (Rosenhauer 1856: 218).

Family-group name: NEPHODINI Reitter, 1917. Stem: Nephod-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

Nephrites Shuckard, 1838: 512. Current status: valid genus in RIPIPHORIDAE: HEMIRHIPIDIINAE.

Type species: Nephrites nitidus Shuckard, 1838, by monotypy.

Family-group name: NEPHRITINAE Selander, 1957. Stem: Nephrit-.

Nerissus Chapuis, 1874: 286. Current status: valid genus in CHRYSOMELIDAE: EUMOLPINAE: BROMIINI.

Type species: Nerissus strigosus Chapuis, 1874, by monotypy.

Family-group name: NERISSINI Kuntzen, 1912. Stem: Neriss-.

Nerthops Schönherr, 1826: 60. Current status: valid genus in CURCULIONIDAE: CURCULIONINAE: NERTHOPINI.

Type species: Nerthops multiguttatus Schönherr, 1826 (= Bruchus punctatus Fabricius, 1798), by original designation.

Family-group name: NERTHOPIDES Lacordaire, 1865. Stem: Nerthop-.

Nertinus G.A.K. Marshall, 1943: 118. Current status: valid genus in CURCULIONIDAE: BARIDINAE: APOSTASIMERINI: NERTININA. Note: new replacement name for Nertus Schönherr, 1844.

Type species: Nertus mannerheimi Boheman, 1844, by original designation (type fixation for Nertus Schönherr).

Family-group name: NERTININA Voss, 1954. Stem: Nertin-.

Nertus Schönherr, 1844: 76. Current status: invalid senior synonym of Nertinus G.A.K. Marshall, 1943 in CURCULIONIDAE: BARIDINAE: APOSTASIMERINI: NERTININA. Note: junior homonym of Nertus Boie, 1828 [Aves].

Type species: Nertus mannerheimi Boheman, 1844, by original designation.

Family-group name: NERTINI Bondar, 1949. Stem: Nert-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

Nesiobius G.A.K. Marshall, 1943: 118. Current status: valid genus in CURCULIONIDAE: COSSONINAE: NESIOBIINI. Note: new replacement name for Nesiotes Wollaston, 1861.

Type species: Nesiotes squamosus Wollaston, 1861, by monotypy (type fixation for Nesiotes Wollaston).

Family-group name: NESIOBIINI Alonso-Zarazaga & Lyal, 1999. Stem: Nesiobi-.

Nesiotes Wollaston, 1861d: 211. Current status: invalid senior synonym of Nesiobius G.A.K. Marshall, 1943 in CURCULIONIDAE: COSSONINAE: NESIOBIINI. Note: junior homonym of Nesiotes Martens, 1860 [Mollusca].

Type species: Nesiotes squamosus Wollaston, 1861, by monotypy.

Family-group name: NESIOTINAE Voss, 1972. Stem: Nesiot-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

Nesoneus Bernhauer, 1939: 205. Current status: valid genus in STAPHYLINIDAE: PROTEININAE: NESONEINI.

Type species: Nesoneus acuticeps Bernhauer, 1939, by monotypy.

Family-group name: NESONEINI Steel, 1966. Stem: Nesone-.

Nesotrinchus Obenberger, 1924: 12. Current status: valid genus in BUPRESTIDAE: CHRYSOCHROINAE: POECILONOTINI: NESOTRINCHINA.

Type species: Nesotrinchus simondsi Obenberger, 1924 (= Haplotrinchus australicus Kerremans, 1903), by original designation.

Family-group name: NESOTRINCHINA Bílý, Kubáň & Volkovitsh, 2009. Stem: Nesotrinch-.

Nessiara Pascoe, 1860b: 60. Current status: valid genus in ANTHRIBIDAE: ANTHRIBINAE: ZYGAENODINI. Note: new replacement name for Nessia Pascoe, 1859.

Type species: Nessia didyma Pascoe, 1859, by subsequent designation (Jordan 1894b: 630).

Family-group name: NESSIARINI Morimoto, 1972. Stem: Nessiar-.

Nettarhinus Schönherr, 1826: 269. Current status: valid genus in CURCULIONIDAE: MOLYTINAE: NETTARHININI.

Type species: Nettarhinus anthribiformis Schönherr, 1826, by original designation.

Family-group name: NETTARHINIDES Lacordaire, 1865. Stem: Nettarhin-.

Netuschilia Reitter, 1904: 34, 35. Current status: valid genus in TENEBRIONIDAE: PIMELIINAE: LACHNOGYINI: NETUSCHILIINA.

Type species: Lachnopus hauseri Reitter, 1897, by monotypy.

Family-group name: NETUSCHILIINA Ferrer & Yvinec, 2004. Stem: Netuschili-.

Neuglenes C.G. Thomson, 1859: 63. Current status: junior synonym of Ptinella Motschulsky, 1844 in PTILIIDAE: PTILIINAE: PTINELLINI.

Type species: Ptilium apterum Guérin-Méneville, 1839, by original designation.

Family-group name: NEUGLENINI Reitter, 1891. Stem: Neuglen-.

Neumatora Normand, 1920: 24. Current status: valid genus in CURCULIONIDAE: COSSONINAE: NEUMATORINI.

Type species: Neumatora scillae Normand, 1920, by subsequent designation (Hlaváč and Maughan 2013: 59).

Family-group name: NEUMATORINI Folwaczny, 1973. Stem: Neumator-.

Neuraphes C.G. Thomson, 1859: 61. Current status: valid genus in STAPHYLINIDAE: SCYDMAENINAE: SCYDMAENITAE: STENICHNINI. Note: the original spelling was Nevraphes but Neuraphes, an incorrect subsequent spelling, is in prevailing usage, attributed to the publication of the original spelling, and therefore deemed to be the correct original spelling (ICZN 1999a, Article 33.3.1).

Type species: Scydmaenus angulatus P.W.J. Müller & Kunze, 1822, by original designation.

Family-group name: NEURAPHINI Csiki, 1909. Stem: Neuraph-.

Nicagus J.L. LeConte, 1861: 130. Current status: valid genus in LUCANIDAE: AESALINAE: NICAGINI.

Type species: Ochodaeus obscurus J.L. LeConte, 1847, by monotypy.

Family-group name: NICAGINI J.L. LeConte, 1861. Stem: Nicag-.

Nicrophorus Fabricius, 1775: 71. Current status: valid genus in STAPHYLINIDAE: SILPHINAE: NICROPHORINI.

Type species: Silpha vespillo Linnaeus, 1758, by subsequent designation (Latreille 1810: 427).

Family-group name: NECROPHORIDAE W. Kirby, 1837. Stem: Nicrophor-.

Nigidius W.S. MacLeay, 1819: 96, 108. Current status: valid genus in LUCANIDAE: LUCANINAE: FIGULINI.

Type species: Nigidius cornutus W.S. MacLeay, 1819, by monotypy.

Family-group name: NIGIDIINI Jakobson, 1911. Stem: Nigidi-.

Nilio Latreille, 1802b: 179. Current status: valid genus in TENEBRIONIDAE: NILIONINAE. Note: the original spelling of the genus is Nilion but Nilio, an incorrect subsequent spelling, is in prevailing usage, attributed to the publication of the original spelling, and therefore deemed to be the correct original spelling (ICZN 1999a, Article 33.3.1).

Type species: Coccinella villosa Fabricius, 1787, by monotypy.

Family-group name: NILIONIDEN Oken, 1843. Stem: Nilion-.

Niphades Pascoe, 1871b: 174. Current status: valid genus in CURCULIONIDAE: MOLYTINAE: AMINYOPINI.

Type species: Niphades pardalotus Pascoe, 1871, by subsequent designation (Voss 1958: 47).

Family-group name: NIPHADINI Voss, 1963. Stem: Niphad-.

Niphadonothus Voss, 1965: 340. Current status: valid genus in CURCULIONIDAE: MOLYTINAE: AMINYOPINI.

Type species: Niphadonothus gentilis Voss, 1965, by original designation.

Family-group name: NIPHADONOTHINA Voss, 1965. Stem: Niphadonoth-.

Niphona Mulsant, 1839: 169. Current status: valid genus in CERAMBYCIDAE: LAMIINAE: PTEROPLIINI.

Type species: Niphona picticornis Mulsant, 1839, by monotypy.

Family-group name: NIPHONINAE Pascoe, 1864. Stem: Niphon-.

Niponius Lewis, 1885: 333. Current status: valid genus in HISTERIDAE: NIPONIINAE.

Type species: Niponius impressicollis Lewis, 1885, by subsequent designation (Gardner 1935: 4).

Family-group name: NIPONIIDAE Fowler, 1912. Stem: Niponi-.

Nipponocyphon Lawrence & Yoshitomi, 2007: 509. Current status: valid genus in SCIRTIDAE: NIPPONOCYPHONINAE.

Type species: Nipponocyphon nakanei Lawrence & Yoshitomi, 2007, by original designation.

Family-group name: NIPPONOCYPHONINAE Lawrence & Yoshitomi, 2007. Stem: Nipponocyphon-.

Nitidula Fabricius, 1775: 77. Current status: valid genus in NITIDULIDAE: NITIDULINAE: NITIDULINI.

Type species: Silpha bipustulata Linnaeus, 1761 (= Silpha bipunctata Linnaeus, 1758), by subsequent designation (Latreille 1810: 427).

Family-group name: NITIDULARIAE Latreille, 1802. Stem: Nitidul-.

Noda Chevrolat in Dejean, 1836a: 410. Current status: invalid senior synonym of Brachypnoea Gistel, 1848 in CHRYSOMELIDAE: EUMOLPINAE: TYPOPHORINI. Note: junior homonym of Noda Schellenberg, 1803 [Diptera].

Type species: Colaspis tristis G.-A. Olivier, 1808, by subsequent designation (Monrós and Bechyné 1956: 1124).

Family-group name: NODINI Selman, 1965. Stem: Nod-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

Nodina Motschulsky, 1858a: 108. Current status: valid genus in CHRYSOMELIDAE: EUMOLPINAE: TYPOPHORINI.

Type species: Nodina pusilla Motschulsky, 1858, by subsequent designation (R. Lucas 1920: 441).

Family-group name: NODININI S.H. Chen, 1940. Stem: Nodin-.

Nodostoma Motschulsky, 1860b: 176. Current status: junior synonym of Basilepta Baly, 1860 in CHRYSOMELIDAE: EUMOLPINAE: TYPOPHORINI.

Type species: Nodostoma fulvipes Motschulsky, 1860, by subsequent designation (Jacoby 1908: 301).

Family-group name: NODOSTOMITES Chapuis, 1874. Stem: Nodostomat-.

Nodotelus C. Koch, 1950a: 67. Current status: junior synonym of Eutelonotus Alluaud, 1902 in TENEBRIONIDAE: STENOCHIINAE: CNODALONINI. Note: new replacement name for Eutelus Solier, 1843.

Type species: Eutelus requieni Solier, 1843, by subsequent designation (R. Lucas 1920: 291) (type fixation for Eutelus Solier).

Family-group name: NODOTELINI C. Koch, 1950. Stem: Nodotel-.

Nomius Laporte, 1834c: 144. Current status: valid genus in CARABIDAE: PSYDRINAE: PSYDRINI. Note: Morio pygmaeus Dejean, 1831 was placed on the Official List of Generic Names in Zoology (ICZN 2011a).

Type species: Nomius graecus Laporte, 1834 (= Morio pygmaeus Dejean, 1831), by monotypy (see ICZN 2011a). Note: name placed on the Official List of Specific Names in Zoology (ICZN 2011a).

Family-group name: NOMIIDAE Gozis, 1875. Stem: Nomius- (ICZN 2011a).

Nordenskioldia J.R. Sahlberg, 1880: 96. Current status: valid genus in STAPHYLINIDAE: EUAESTHETINAE: NORDENSKIOLDIINI.

Type species: Nordenskioldia glacialis J.R. Sahlberg, 1880, by monotypy.

Family-group name: NORDENSKIOELDIINI Bernhauer & Schubert, 1911. Stem: Nordenskioldi-.

Noseolucanus Araya & Tanaka, 1998: 334. Current status: valid genus in LUCANIDAE: LUCANINAE: NOSEOLUCANINI.

Type species: Noseolucanus rugosus Araya & Tanaka, 1998 (= Pseudolucanus denticulus Boucher, 1996), by original designation.

Family-group name: NOSEOLUCANINI H. Huang & C.H. Chen, 2013: 67. Stem: Noseolucan-.

Nosodendron Latreille, 1804a: 146. Current status: valid genus in NOSODENDRIDAE.

Type species: Byrrhus fascicularis G.-A. Olivier, 1790, by monotypy.

Family-group name: NOSODENDRINI Erichson, 1846. Stem: Nosodendr-.

Nossidium Erichson, 1845a: 17. Current status: valid genus in PTILIIDAE: NOSSIDIINAE.

Type species: Dermestes pilosellus Marsham, 1802, by subsequent designation (E. Desmarest 1852: 199).

Family-group name: NOSSIDIINAE Sörensson & Delgado, 2019: 791. Stem: Nossidi-.

Notapion Zimmerman, 1994: 330. Current status: valid genus in BRENTIDAE: APIONINAE: RHADINOCYBITAE: NOTAPIONINI.

Type species: Apion trilobicolle Lea, 1926, by original designation.

Family-group name: NOTAPIONINI Zimmerman, 1994. Stem: Notapion-.

Noterapion Kissinger, 2002: 315. Current status: valid genus in BRENTIDAE: APIONINAE: NOTERAPIONITAE.

Type species: Apion meorrhynchum R.A. Philippi & F. Philippi, 1864, by original designation.

Family-group name: NOTERAPIONINI Kissinger, 2004. Stem: Noterapion- (ICZN 1999a, Article 29.4).

Noterus Clairville, 1806: 222. Current status: valid genus in NOTERIDAE: NOTERINAE: NOTERINI.

Type species: Dytiscus crassicornis O.F. Müller, 1776, by monotypy.

Family-group name: NOTERIDES C.G. Thomson, 1860. Stem: Noter-.

Nothobroscus Roig-Juñent & Ball, 1995: 303, 306. Current status: valid genus in CARABIDAE: BROSCINAE: BROSCINI: NOTHOBROSCINA.

Type species: Nothobroscus chilensis Roig-Juñent & Ball, 1995, by original designation.

Family-group name: NOTHOBROSCINA Roig-Juñent, 2000. Stem: Nothobrosc-.

Nothochodaeus Nikolajev, 2005b: 219. Current status: valid genus in OCHODAEIDAE: OCHODAEINAE: NOTHOCHODAEINI. Note: two original spellings were used, including Notochodaeus (p. 219); Masumoto et al. (2013: 107) acted as First Revisers (ICZN 1999a, Article 24.2.3) and selected Nothochodaeus.

Type species: Ochodaeus maculatus C.O. Waterhouse, 1875, by original designation.

Family-group name: NOTHOCHODAEINI Nikolajev, 2015a: 25. Stem: Nothochodae-.

Nothognathus G.A.K. Marshall, 1916: 250. Current status: valid genus in CURCULIONIDAE: ENTIMINAE: OTIORHYNCHITAE: NOTHOGNATHINI.

Type species: Nothognathus schoutedeni G.A.K. Marshall, 1916, by original designation.

Family-group name: NOTHOGNATHIDES G.A.K. Marshall, 1916. Stem: Nothognath-.

Nothomorpha Saunders, 1871: 78. Current status: valid genus in BUPRESTIDAE: POLYCESTINAE: ACMAEODERINI: NOTHOMORPHINA.

Type species: Amorphosoma verrucosum Gory & Laporte, 1839, by subsequent designation (Rye 1873: 271).

Family-group name: NOTOMORPHINI Cobos, 1955. Stem: Nothomorph-.

Nothorhina L. Redtenbacher, 1845: 109, 153. Current status: valid genus in CERAMBYCIDAE: SPONDYLIDINAE: NOTHORHININI.

Type species: Callidium muricatum Dalman, 1817, by subsequent designation (J. Thomson 1864: 267).

Family-group name: NOTHORHININI Zagajkevich, 1991. Stem: Nothorhin-.

†Nothotytthonyx Y.-D. Li, Biffi, Kundrata & Cai, 2022: 264. Current status: valid genus in CANTHARIDAE: MALTHININAE: †NOTHOTYTTHONYCHINI. Note: mid-Cretaceous (Myanmar).

Type species: †Nothotytthonyx serratus Y.-D. Li, Biffi, Kundrata & Cai, 2022, by original designation.

Family-group name: †NOTHOTYTTHONYCHINI Fanti, 2022: 321. Stem: Nothotytthonych-.

Nothus G.-A. Olivier, 1812: 383, 384. Current status: junior synonym of Osphya Illiger, 1807 in MELANDRYIDAE: OSPHYINAE.

Type species: Nothus clavipes G.-A. Olivier, 1812 (= Cantharis bipunctata Fabricius, 1775), by subsequent designation (Westwood 1838c: 31). Note: the type species currently recognized for this genus is Nothus praeustus G.-A. Olivier, 1812 (e.g., Nikitsky 2020: 58), which is also a junior synonym of Cantharis bipunctata Fabricius, 1775, therefore there is no change in generic concept with the change of type species.

Family-group name: NOTHIDAE Shuckard, 1839. Stem: Noth-.

Notiokasis Kavanaugh & Nègre, 1983: 551. Current status: valid genus in CARABIDAE: NEBRIINAE: NOTIOKASIINI.

Type species: Notiokasis chaudoiri Kavanaugh & Nègre, 1983, by original designation.

Family-group name: NOTIOKASIINI Kavanaugh & Nègre, 1983. Stem: Notiokasi-.

Notiomimetes Wollaston, 1873b: 440, 504. Current status: valid genus in CURCULIONIDAE: CYCLOMINAE: NOTIOMIMETINI.

Type species: Notiomimetes pascoei Wollaston, 1873, by monotypy.

Family-group name: NOTIOMIMETIDES Wollaston, 1873. Stem: Notiomimet-.

Notiophilus Duméril, 1805: 194. Current status: valid genus in CARABIDAE: NEBRIINAE: NOTIOPHILINI.

Type species: Cicindela aquatica Linnaeus, 1758, by subsequent designation (Curtis 1829: pl. 254).

Family-group name: NOTIOPHILI Motschulsky, 1850. Stem: Notiophil-.

Notiophygus Gory, 1834: 453. Current status: valid genus in DISCOLOMATIDAE: NOTIOPHYGINAE: NOTIOPHYGINI.

Type species: Notiophygus nigropunctatus Gory, 1834, by subsequent designation (John 1959: 20).

Family-group name: NOTIOPHYGIDAE Jakobson, 1915. Stem: Notiophyg-.

Notioxenus Wollaston, 1861d: 212. Current status: invalid senior synonym of Valenfriesia Alonso-Zarazaga & Lyal, 1999 in ANTHRIBIDAE: CHORAGINAE: TYPOPHORINI. Note: junior homonym of Notioxenus Motschulsky, 1858 [Coleoptera: CARABIDAE].

Type species: Notioxenus bewickii Wollaston, 1861, by subsequent designation (Jordan 1907: 380).

Family-group name: NOTIOXÉNIDES Lacordaire, 1865. Stem: Notioxen-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

†Notocupes Ponomarenko, 1964: 61. Current status: valid genus in ARCHOSTEMATA incertae sedis. Note: Triassic/Late Cretaceous (China; Kazakhstan; Mongolia; Russia).

Type species: †Notocupes picturatus Ponomarenko, 1964, by original designation.

Family-group name: †NOTOCUPEDINI Ponomarenko, 1966. Stem: Notocuped-.

Notodermus Desbrochers des Loges, 1875: 19. Current status: junior synonym of Procas Stephens, 1831 in CURCULIONIDAE: BRACHYCERINAE: ERIRHININI.

Type species: Notodermus bruleriei Desbrochers des Loges, 1875, by monotypy.

Family-group name: NOTODERMINA Voss, 1952. Stem: Notoderm-.

Notolomus J.L. LeConte in J.L. LeConte and G.H. Horn, 1876: 222. Current status: valid genus in CURCULIONIDAE: CURCULIONINAE: DERELOMINI: NOTOLOMINA.

Type species: Notolomus basalis J.L. LeConte, 1876, by subsequent designation (Lepesme et al. 1947: 587).

Family-group name: NOTOLOMINA N.M. Franz, 2006. Stem: Notolom-.

Notomicrus Sharp, 1882b: 260. Current status: valid genus in NOTERIDAE: NOTOMICRINAE: NOTOMICRINI.

Type species: Notomicrus brevicornis Sharp, 1882, by subsequent designation (Guignot 1946: 115).

Family-group name: NOTOMICRINI Branden, 1884. Stem: Notomicr-.

Notophysis Audinet-Serville, 1832: 158. Current status: valid genus in CERAMBYCIDAE: PRIONINAE: CACOSCELINI.

Type species: Notophysis lucanoides Audinet-Serville, 1832, by monotypy.

Family-group name: NOTHOPHYSINI Lameere, 1903. Stem: Notophyse-.

Notosacantha Chevrolat in Dejean, 1836a: 367. Current status: valid genus in CHRYSOMELIDAE: CASSIDINAE: NOTOSACANTHINI.

Type species: Cassida echinata Fabricius, 1801, by monotypy.

Family-group name: NOTOSACANTHINA Gressitt, 1952. Stem: Notosacanth-.

Nototylus Gemminger & Harold, 1868a: 161. Current status: valid genus in CARABIDAE: NOTOTYLINAE. Note: new replacement name for Tylonotus Schaum, 1863.

Type species: Tylonotus fryi Schaum, 1863, by monotypy (type fixation for Tylonotus Schaum).

Family-group name: NOTOTYLINI Bänninger, 1927. Stem: Nototyl-.

Notoxus Geoffroy, 1762: 356. Current status: valid genus in ANTHICIDAE: NOTOXINAE. Note: name placed on the Official List of Generic Names in Zoology (ICZN 1994a).

Type species: Attelabus monoceros Linnaeus, 1761, by subsequent designation (Latreille 1810: 430; see ICZN 1994a). Note: name placed on the Official List of Specific Names in Zoology (ICZN 1994a).

Family-group name: NOTOXIDAE Stephens, 1829. Stem: Notox-.

Novius Mulsant, 1846: [4]. Current status: valid genus in COCCINELLIDAE: COCCINELLINAE: NOVIINI. Note: the spelling Nomius (p. 213) was changed to Novius in the Addenda and Errata (p. [4]).

Type species: Novius cruentatus Mulsant, 1846, by monotypy.

Family-group name: NOVIAIRES Mulsant, 1846. Stem: Novi-.

Nucleotops Perkins & J. Balfour-Browne, 1994: 13. Current status: valid genus in HYDRAENIDAE: PROSTHETOPINAE: NUCLEOTOPINI.

Type species: Nucleotops nimbaceps Perkins & J. Balfour-Browne, 1994, by original designation.

Family-group name: NUCLEOTOPINI Perkins, 1994. Stem: Nucleotop-.

Nyassinus Westwood, 1879: 199. Current status: valid genus in SCARABAEIDAE: CETONIINAE: CREMASTOCHEILINI: NYASSININA.

Type species: Nyassinus maculipes Westwood, 1879, by subsequent designation (Marais and Holm 1992: 53).

Family-group name: NYASSININA Krikken, 1984. Stem: Nyassin-.

Nyctelia Berthold, 1827: 367. Current status: valid genus in TENEBRIONIDAE: PIMELIINAE: NYCTELIINI. Note: nomen protectum (see Silvestro and Flores 2016: 658).

Type species: Zophosis nodosa Germar, 1823 (= Zophosis picipes Billberg, 1815), by monotypy.

Family-group name: NYCTÉLITES Solier, 1834. Stem: Nycteli-.

Nycteropus Klug, 1833: 89. Current status: valid genus in TENEBRIONIDAE: TENEBRIONINAE: TOXICINI: NYCTEROPINA.

Type species: Nycteropus ebeninus Klug, 1833, by subsequent designation (Hope 1841a: 124).

Family-group name: NYCTÉROPIDES Lacordaire, 1859. Stem: Nycterop-.

Nyctimene J. Thomson, 1857: 314. Current status: invalid senior synonym of Nyctimenius Gressitt, 1951 in CERAMBYCIDAE: LAMIINAE: NYCTIMENIINI. Note: junior homonym of Nyctimene Borkenhausen, 1797 [Mammalia].

Type species: Nyctimene agriloides J. Thomson, 1857 (= Saperda varicornis Fabricius, 1801), by monotypy.

Family-group name: NYCTIMENITAE J. Thomson, 1864. Stem: Nyctimen-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

Nyctimenius Gressitt, 1951: 629. Current status: valid genus in CERAMBYCIDAE: LAMIINAE: NYCTIMENIINI. Note: new replacement name for Nyctimene J. Thomson, 1857.

Type species: Nyctimene agriloides J. Thomson, 1857 (= Saperda varicornis Fabricius, 1801), by monotypy (type fixation for Nyctimene J. Thomson).

Family-group name: NYCTIMENIINI Gressitt, 1951. Stem: Nyctimeni-.

Nyctophyxis C. Costa, 1975: 86. Current status: valid genus in ELATERIDAE: AGRYPNINAE: PYROPHORINI: NYCTOPHYXINA. Note: two original spellings were used, including Nyctophixis (p. 86); Sáiz et al. (1993: 70) acted as First Revisers (ICZN 1999a, Article 24.2.3) and selected Nyctophyxis, the spelling currently in prevailing usage.

Type species: Pyrophorus ocellatus Germar, 1841, by original designation.

Family-group name: NYCTOPHYXINA C. Costa, 1975. Stem: Nyctophyx-.

Nyctoporis Eschscholtz, 1831: 10, 11. Current status: valid genus in TENEBRIONIDAE: PIMELIINAE: NYCTOPORINI.

Type species: Nyctoporis cristata Eschscholtz, 1831, by subsequent designation (Hope 1841a: 124).

Family-group name: NYCTOPORIDES Lacordaire, 1859. Stem: Nyctopor-. Note: senior homonym of NYCTOPORINAE Hill, 1951 (type genus Nyctopora Nicholson, 1879) in Coelenterata.

Nyctor Semenov-Tjan-Shansky & Pjatakova, 1936: 102. Current status: valid genus in ELATERIDAE: CARDIOPHORINAE.

Type species: Nyctor expallidus Semenov-Tjan-Shansky & Pjatakova, 1936, by original designation.

Family-group name: NYCTORINI Semenov-Tjan-Shansky & Pjatakova, 1936. Stem: Nyctor-.

Nyctozoilus Guérin-Méneville, 1831a: pl. 4. Current status: valid genus in TENEBRIONIDAE: TENEBRIONINAE: HELEINI: CYPHALEINA.

Type species: Nyctozoilus obesus Guérin-Méneville, 1831, by monotypy.

Family-group name: NYCTOZOÏLIDES Lacordaire, 1859. Stem: Nyctozoil-.

Nymphister Reichensperger, 1933: 188. Current status: valid genus in HISTERIDAE: HAETERIINAE: NYMPHISTERINI.

Type species: Nymphister simplicissimus Reichensperger, 1933, by original designation.

Family-group name: NYMPHISTERINI Tishechkin, 2007. Stem: Nymphister- (ICZN 1999a, Article 29.4).

Oberea Dejean, 1835: 351. Current status: valid genus in CERAMBYCIDAE: LAMIINAE: SAPERDINI.

Type species: Cerambyx linearis Linnaeus, 1761, by subsequent designation (C.G. Thomson 1859: 153).

Family-group name: OBEREITAE J. Thomson, 1864. Stem: Obere-.

†Oborocoleus Kukalová, 1969: 155. Current status: valid genus in †OBOROCOLEIDAE. Note: Permian (Czech Republic).

Type species: †Oborocoleus rohdendorfi Kukalová, 1969, by original designation.

Family-group name: †OBOROCOLEIDAE Kukalová, 1969. Stem: Oborocole-.

†Obrienia Zherikhin & Gratshev, 1994: 52. Current status: valid genus in †OBRIENIIDAE: †OBRIENIINAE. Note: Middle/Upper Triassic (Kyrgyzstan).

Type species: †Obrienia kuscheli Zherikhin & Gratshev, 1994, by original designation.

Family-group name: †OBRIENIIDAE Zherikhin & Gratshev, 1994. Stem: Obrieni-.

Obrium Dejean, 1821: 110. Current status: valid genus in CERAMBYCIDAE: CERAMBYCINAE: OBRIINI.

Type species: Cerambyx cantharinus Linnaeus, 1767, by subsequent designation (Curtis 1825: pl. 91).

Family-group name: OBRIAIRES Mulsant, 1839. Stem: Obri-.

Ocalea Erichson, 1837: 298. Current status: valid genus in STAPHYLINIDAE: ALEOCHARINAE: OXYPODINI: OXYPODINA.

Type species: Ocalea castanea Erichson, 1837 (= Aleochara picata Stephens, 1832), by subsequent designation (Weimar 1839: 506).

Family-group name: OCALEIDES C.G. Thomson, 1859. Stem: Ocale-.

Ochina Dejean, 1821: 40. Current status: valid genus in PTINIDAE: ERNOBIINAE: OCHININI.

Type species: Ptilinus hederae P.W.J. Müller, 1821 (= Crioceris ptinoides Marsham, 1802), by subsequent designation (Blanchard 1845a: 91).

Family-group name: OCHININI Zahradník in Zahradník and Háva, 2014: 307. Stem: Ochin-.

Ochodaeus Dejean, 1821: 56. Current status: valid genus in OCHODAEIDAE: OCHODAEINAE: OCHODAEINI.

Type species: Melolontha chrysomelina Fabricius, 1792 (= Scarabaeus chrysomeloides Schrank, 1781), by monotypy.

Family-group name: OCHODAEIDAE Streubel, 1846. Stem: Ochodae-.

Ochrolitus Sharp, 1889b: 264. Current status: valid genus in PHALACRIDAE: PHALACRINAE.

Type species: Ochrolitus optatus Sharp, 1889, by monotypy.

Family-group name: OCHROLITINI Guillebeau, 1894. Stem: Ochrolit-.

Ochthebius Leach, 1815a: 95. Current status: valid genus in HYDRAENIDAE: OCHTHEBIINAE: OCHTHEBIINI. Note: name placed on the Official List of Generic Names in Zoology (ICZN 1991a).

Type species: Helophorus marinus Paykull, 1798, by subsequent designation (d’ Orchymont 1942: 2; see ICZN 1991a). Note: name placed on the Official List of Specific Names in Zoology ICZN 1991a).

Family-group name: OCHTEBIIDAE C.G. Thomson, 1859. Stem: Ochthebi-.

Ochtheosus Perkins, 1997: 124. Current status: valid genus in HYDRAENIDAE: OCHTHEBIINAE: OCHTHEOSINI.

Type species: Ochtheosus fungicolus Perkins, 1997, by original designation.

Family-group name: OCHTHEOSINI Perkins, 1997. Stem: Ochtheos-.

Ochyra Pascoe, 1871c: 273. Current status: valid genus in CERAMBYCIDAE: CERAMBYCINAE: OCHYRINI.

Type species: Ochyra coarctata Pascoe, 1871, by monotypy.

Family-group name: OCHYRINAE Pascoe, 1871. Stem: Ochyr-.

Ochyromera Pascoe, 1874: 31, 33. Current status: valid genus in CURCULIONIDAE: CURCULIONINAE: OCHYROMERINI.

Type species: Ochyromera rufescens Pascoe, 1874, by subsequent designation (G.A.K. Marshall 1926: 361).

Family-group name: OCHYROMERINA Voss, 1935. Stem: Ochyromer-.

Ochyropus Schiødte, 1847: 350. Current status: valid genus in CARABIDAE: SCARITINAE: SCARITINI: OCHYROPINA.

Type species: Ochyropus gigas Schiødte, 1847, by monotypy.

Family-group name: OCHYROPINI Basilewsky, 1973. Stem: Ochyrop-.

Ocladius Schönherr, 1825: 587. Current status: valid genus in CURCULIONIDAE: BRACHYCERINAE: ERIRHININI.

Type species: Rhynchaenus salicorniae G.-A. Olivier, 1807, by original designation.

Family-group name: OCLADIIDES Lacordaire, 1865. Stem: Ocladi-.

Ocnoscelis Erichson, 1847c: 174. Current status: valid genus in CHRYSOMELIDAE: GALERUCINAE: ALTICINI.

Type species: Ocnoscelis cyanoptera Erichson, 1847, by present designation.

Family-group name: OCNOSCELINA Bechyné, 1997. Stem: Ocnoscelid-.

Ochthephilus Nietner, 1857: 136. Current status: invalid senior synonym of Perileptus Schaum, 1860 in CARABIDAE: TRECHINAE: TRECHINI: TRECHODINA. Note: junior homonym of Ochthephilus Mulsant & Rey, 1856 [Coleoptera: STAPHYLINIDAE].

Type species: Ochthephilus ceylanicus Nietner, 1857, by monotypy.

Family-group name: OCHTHEPHILINI Jeannel, 1922. Stem: Ochthephil-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

Octocryptus Candèze, 1892: 486. Current status: valid genus in ELATERIDAE: AGRYPNINAE: AGRYPNINI.

Type species: Octocryptus cardoni Candèze, 1892, by monotypy.

Family-group name: OCTOCRYPTINI O. Schwarz, 1906. Stem: Octocrypt-.

Octogonotes Drapiez, 1820: 191. Current status: valid genus in CHRYSOMELIDAE: GALERUCINAE: ALTICINI.

Type species: Octogonotes banoni Drapiez, 1820, by monotypy.

Family-group name: OCTOGONOTINI Weise, 1921. Stem: Octogonot-.

Octomicrus L.W. Schaufuss, 1877a: 14. Current status: valid genus in STAPHYLINIDAE: PSELAPHINAE: EUPLECTITAE: DIMERINI.

Type species: Octomicrus longulus L.W. Schaufuss, 1877, by monotypy.

Family-group name: OCTOMICRINI Jeannel, 1952. Stem: Octomicr-.

Octotemnus Mellié, 1847: 110. Current status: valid genus in CIIDAE: CIINAE: OROPHIINI.

Type species: Cis glabriculus Gyllenhal, 1827, by subsequent designation (Jacquelin du Val 1860b: 239).

Family-group name: OCTOTEMNIDAE Reitter, 1878. Stem: Octotemn-.

Octotoma Dejean, 1836a: 366. Current status: valid genus in CHRYSOMELIDAE: CASSIDINAE: UROPLATINI.

Type species: Hispa plicatula Fabricius, 1801, by monotypy.

Family-group name: OCTOTOMIDES G.H. Horn, 1884. Stem: Octotom-.

Ocularia Jordan, 1894a: 202. Current status: valid genus in CERAMBYCIDAE: LAMIINAE: OCULARIINI.

Type species: Ocularia apicalis Jordan, 1894, by original designation.

Family-group name: OCULARIINI Breuning, 1950. Stem: Oculari-.

Ocypus Leach in Samouelle, 1819: 172. Current status: valid genus in STAPHYLINIDAE: STAPHYLININAE: STAPHYLININI: STAPHYLININA.

Type species: Staphylinus cyaneus Paykull, 1789 (= Staphylinus ophthalmicus Scopoli, 1763), by original designation.

Family-group name: OCYPINA Hatch, 1957. Stem: Ocypod-. Note: junior homonym of OCYPODIDAE Rafinesque, 1815 (type genus Ocypode Weber, 1795) in Crustacea.

Ocyusa Kraatz, 1856: 156. Current status: valid genus in STAPHYLINIDAE: ALEOCHARINAE: OXYPODINI: OXYPODINA.

Type species: Oxypoda maura Erichson, 1837, by subsequent designation (C.G. Thomson 1859: 36).

Family-group name: OCYUSATES Mulsant & Rey, 1874. Stem: Ocyus-.

Odacantha Paykull, 1798: 169. Current status: valid genus in CARABIDAE: HARPALINAE: ODACANTHINI.

Type species: Attelabus melanurus Linnaeus, 1767, by monotypy.

Family-group name: ODACANTHIDAE Laporte, 1834. Stem: Odacanth-.

Odochilus Harold, 1877: 97. Current status: valid genus in SCARABAEIDAE: APHODIINAE: ODOCHILINI.

Type species: Odochilus syntheticus Harold, 1877, by monotypy.

Family-group name: ODOCHILINI Rakovič, 1987. Stem: Odochil-.

Odontalgus Raffray, 1877: 286. Current status: valid genus in STAPHYLINIDAE: PSELAPHINAE: PSELAPHITAE: ODONTALGINI.

Type species: Odontalgus tuberculatus Raffray, 1877, by subsequent designation (Jeannel 1949a: 178).

Family-group name: ODONTALGINI Jeannel, 1949. Stem: Odontalg-.

Odonteus Samouelle, 1819: 189. Current status: valid genus in BOLBOCERATIDAE: BOLBOCERATINAE: ODONTEINI. Note: name given priority over Bolboceras W. Kirby, 1819 whenever the two are considered synonyms and Odonteus Samouelle, 1819 placed on the Official List of Generic Names in Zoology (ICZN 2006a).

Type species: Scarabaeus mobilicornis Fabricius, 1775 (= Scarabaeus armiger Scopoli, 1772), by monotypy (see ICZN 2006a).

Family-group name: ODONTAEIDAE Streubel, 1846. Stem: Odonte-.

Odontionopa Chevrolat in Dejean, 1836a: 408. Current status: valid genus in CHRYSOMELIDAE: EUMOLPINAE: EURYOPINI.

Type species: Colaspis sericea Gyllenhal, 1808, by monotypy.

Family-group name: ODONTIONOPITES Lefèvre, 1876. Stem: Odontionop-.

Odontionopa Erichson, 1842: 232. Current status: invalid senior synonym of Eboo Reid, 1993 in CHRYSOMELIDAE: EUMOLPINAE: BROMIINI. Note: junior homonym of Odontionopa Chevrolat, 1836 in Coleoptera: CHRYSOMELIDAE.

Type species: Colaspis viridula Erichson, 1842, by subsequent designation (Reid 1993: 62).

Family-group name: ODONTIONOPINI Clavareau, 1914. Stem: Odontionop-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

Odontocheila Laporte, 1834a: 34. Current status: valid genus in CICINDELIDAE: CICINDELINI: DROMICINA. Note: Laporte used two original spellings, including Odontacheila (p. 35); Laporte (1840a: 20) subsequently used only Odontocheila and therefore acted as First Reviser (ICZN 1999a, Article 24.2.4).

Type species [suggested]: Odontocheila desmarestii Laporte, 1834 (= Cicindela fulgens Klug, 1834), by subsequent designation (Bousquet 2002: 35). Note: the valid type species fixation is that of Hope (1838a: 15), who selected Cicindela lacordairei Guérin- Méneville, 1833, but that species is currently included in the subgenus Poecilochila Rivalier, 1969 of the genus Pentacomia H.W. Bates, 1872; an application to the Commission is necessary to fix Odontocheila desmarestii Laporte, 1834 as the type species of Odontocheila Laporte, 1834.

Family-group name: ODONTOCHILINI W. Horn, 1899. Stem: Odontocheil-.

Odontolabis Hope, 1842b: 127. Current status: valid genus in LUCANIDAE: LUCANINAE: ODONTOLABIDINI.

Type species: Odontolabis cuvera Hope, 1842, by subsequent designation (Benesh 1960: 112).

Family-group name: ODONTOLABIDAE Parry, 1870. Stem: Odontolabid-.

Odontolochus A. Schmidt, 1917: 99. Current status: valid genus in SCARABAEIDAE: APHODIINAE: ODONTOLOCHINI. Note: new replacement name for Odontoderus Clouët des Pesruches, 1900.

Type species: Ataenius spinicollis Harold, 1871, by subsequent designation (R. Lucas 1920: 450) (type fixation for Odontoderus Clouët des Pesruches).

Family-group name: ODONTOLOCHINI Stebnicka & Howden, 1996. Stem: Odontoloch-.

Odontoloma Boheman, 1857: 202. Current status: valid genus in SCARABAEIDAE: SCARABAEINAE: ODONTOLOMINI.

Type species: Odontoloma pauxillum Boheman, 1857, by monotypy.

Family-group name: ODONTOLOMINI Davis, Deschodt & Scholtz, 2019: 136. Stem: Odontolom-

Odontonychus Candèze, 1897: 26. Current status: valid genus in ELATERIDAE: ELATERINAE: ODONTONYCHINI.

Type species: Odontonychus granulatus Candèze, 1897, by monotypy.

Family-group name: ODONTONYCHINI C. Girard, 1973. Stem: Odontonych-.

Odontosphindus J.L. LeConte, 1878b: 601. Current status: valid genus in SPHINDIDAE: ODONTOSPHINDINAE.

Type species: Odontosphindus denticollis J.L. LeConte, 1878, by monotypy.

Family-group name: ODONTOSPHINDINI Sen Gupta & Crowson, 1979. Stem: Odontosphind-.

Oedemera G.-A. Olivier, 1789a: 31. Current status: valid genus in OEDEMERIDAE: OEDEMERINAE: OEDEMERINI.

Type species: Necydalis caerulea Linnaeus, 1767 (= Cantharis nobilis Scopoli, 1763), by subsequent designation (Latreille 1810: 430). Note: no species were originally included in this genus by G.-A. Olivier (1789a); the first ones were the 20 species listed by G.-A. Olivier (1795: [50] 4–15).

Family-group name: OEDEMERITES Latreille, 1810. Stem: Oedemer-.

Oedenoderus Chevrolat, 1858: 245. Current status: valid genus in CERAMBYCIDAE: CERAMBYCINAE: OEDENODERINI.

Type species: Oedenoderus pupa Chevrolat, 1858, by subsequent designation (J. Thomson 1861: 251).

Family-group name: OEDENODERINI Aurivillius, 1912. Stem: Oedenoder-.

Oedionychis Latreille, 1829b: 154. Current status: valid genus in CHRYSOMELIDAE: GALERUCINAE: ALTICINI.

Type species: Galeruca marginella Fabricius, 1801 (= Chrysomela cincta Fabricius, 1787), by subsequent designation (Heikertinger 1922: 46).

Family-group name: OEDIONYCHITES Chapuis, 1875. Stem: Oedionych-.

Oediopalpa Baly, 1858: 16. Current status: valid genus in CHRYSOMELIDAE: CASSIDINAE: OEDIOPALPINI.

Type species: Hispa cyanipennis Fabricius, 1801, by subsequent designation (Monrós and Viana 1947: 151).

Family-group name: OEDIOPALPINI Monrós & Viana, 1947. Stem: Oediopalp-.

Oeme Newman, 1840c: 8. Current status: valid genus in CERAMBYCIDAE: CERAMBYCINAE: OEMINI: OEMINA.

Type species: Oeme indecora Newman, 1840 (= Stenocorus rigidus Say, 1826), by monotypy.

Family-group name: OEMIDES Lacordaire, 1868. Stem: Oem-.

Oeneis Mulsant, 1850: 500. Current status: invalid senior synonym of Calloeneis Grote, 1873 in COCCINELLIDAE: COCCINELLINAE: CRYPTOGNATHINI. Note: junior homonym of Oeneis Hübner, 1819 [Lepidoptera].

Type species: Oeneis nigrans Mulsant, 1850, by subsequent designation (Crotch 1874a: 206).

Family-group name: OENEINI Casey, 1899. Stem: Oene-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

Oestodes J.L. LeConte, 1853f: 422, 424. Current status: valid genus in ELATERIDAE: OESTODINAE.

Type species: Elater tenuicollis Randall, 1838, by subsequent designation (Hyslop 1921: 659).

Family-group name: OESTODINI Hyslop, 1917. Stem: Oestod-.

Oides Weber, 1801: 26, 53. Current status: valid genus in CHRYSOMELIDAE: GALERUCINAE: OIDINI.

Type species: Chrysomela bipunctata Fabricius, 1781 (= Oides andrewesi Jacoby, 1900), by subsequent designation (Laboissière 1922: 195).

Family-group name: OÏDITES Laboissière, 1921. Stem: Oid-.

Oileus Kaup, 1868b: 19. Current status: valid genus in PASSALIDAE: PASSALINAE: PROCULINI.

Type species: Passalus rimator Truqui, 1857, by monotypy. Note: Kaup (1868b: 19) listed four species-group names in this genus but only one was available at the time.

Family-group name: OILEINAE Kuwert, 1891. Stem: Oile-.

†Oiserhinus Legalov, Kirejtshuk & Nel, 2020b: 730. Current status: valid genus in ANTHRIBIDAE: ANTHRIBINAE: †OISERHININI. Note: lower Eocene (France).

Type species: †Oiserhinus insolitus Legalov, Kirejtshuk & Nel, 2020, by original designation.

Family-group name: †OISERHININI Legalov, Kirejtshuk & Nel, 2020b: 730. Stem: Oiserhin-.

Oisocerus Bonvouloir in Murray, 1868: 103. Current status: valid genus in EUCNEMIDAE: MACRAULACINAE: OISOCERINI.

Type species: Oisocerus murrayi Bonvouloir, 1868, by monotypy.

Family-group name: OISOCERINI Muona, 1993. Stem: Oisocer-.

Olexandrella Zajciw, 1960: 606. Current status: valid genus in CERAMBYCIDAE: CERAMBYCINAE: DODECOSINI.

Type species: Olexandrella serotina Zajciw, 1960, by original designation.

Family-group name: OLEXANDRELLAEINI Zajciw, 1960. Stem: Olexandrell-.

Olibrus Erichson, 1845a: 108, 113. Current status: valid genus in PHALACRIDAE: PHALACRINAE.

Type species: Sphaeridium aeneum Fabricius, 1792, by subsequent designation (C.G. Thomson 1859: 66). Note: the type species usually recognized is Sphaeridium bicolor Fabricius, 1792 (e.g., Švec 2007: 508) by subsequent designation of Guillebeau (1892: 147) but C.G. Thomson’s typification is earlier; both species are currently included in the genus Olibrus.

Family-group name: OLIBRINI Guillebeau, 1892. Stem: Olibr-.

Oligocys Gistel, 1856: 373. Current status: junior synonym of Liophloeus Germar, 1817 in CURCULIONIDAE: ENTIMINAE: POLYDRUSITAE: POLYDRUSINI.

Type species: Curculio nubilus Fabricius, 1777 (= Curculio tessulatus O.F. Müller, 1776), by monotypy.

Family-group name: OLIGOCYIDAE Gistel, 1856. Stem: Oligoce-.

Oligota Mannerheim, 1830: 72. Current status: valid genus in STAPHYLINIDAE: ALEOCHARINAE: HYPOCYPHTINI.

Type species: Aleochara pusillima Gravenhorst, 1806, by monotypy.

Family-group name: OLIGOTIDES C.G. Thomson, 1859. Stem: Oligot-.

Olisthaerus Dejean, 1833a: 69. Current status: valid genus in STAPHYLINIDAE: OLISTHAERINAE.

Type species: Staphylinus substriatus Paykull, 1790, by monotypy.

Family-group name: OLISTHAERINI C.G. Thomson, 1858. Stem: Olisthaer-.

Olocrates Mulsant, 1854: 147, 150, 383. Current status: junior synonym of Phylan Sturm, 1826 in TENEBRIONIDAE: BLAPTINAE: DENDARINI: DENDARINA. Note: this name was spelled Omocrates (pp. 147, 150) but changed to Olocrates in the errata of the same work (p. 383).

Type species: Opatrum gibbum Fabricius, 1775, by subsequent designation (C.G. Thomson 1859: 115, as Omocrates).

Family-group name: OLOCRATARII Baudi di Selve, 1875. Stem: Olocrat-.

Olotelus Mulsant & Rey, 1866: 22. Current status: invalid senior synonym of Gompelia Alonso-Zarazaga, 2010 in ADERIDAE. Note: junior homonym of Olotelus Solier, 1851 [Coleoptera: ELATERIDAE].

Type species: Xylophilus neglectus Jacquelin du Val, 1863, by subsequent designation (Alonso- Zarazaga 2010: 299).

Family-group name: OLOTELINA Báguena Corella, 1948. Stem: Olotel-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

Omalisus Geoffroy, 1762: 179. Current status: valid genus in ELATERIDAE: OMALISINAE. Note: name placed on the Official List of Generic Names in Zoology (ICZN 1994a).

Type species: Omalisus fontisbellaquaei Geoffroy, 1785, by subsequent monotypy (de Fourcroy 1785: 64; see ICZN 1994a). Note: name placed on the Official List of Specific Names in Zoology (ICZN 1994a).

Family-group name: HOMALISIDES Lacordaire, 1857. Stem: Omalis-.

Omalium Gravenhorst, 1802: 111. Current status: valid genus in STAPHYLINIDAE: OMALIINAE: OMALIINI. Note: name placed on the Official List of Generic Names in Zoology (ICZN 2015b).

Type species: Staphylinus rivularis Paykull, 1789, by subsequent designation (Latreille 1810: 427; see ICZN 2015b).

Family-group name: OMALIDAE W.S. MacLeay, 1825. Stem: Omali-. Note: the spelling of the family-group name OMALIIDAE Handlirsch, 1904 [Archaeorthoptera] was ruled to be OMALIAIDAE by the Commission (see ICZN 2015b) to remove the homonymy with W.S. MacLeay’s name.

Omalodera Blanchard, 1842b: pl. 1. Current status: valid genus in CARABIDAE: TRECHINAE: TRECHINI: TRECHINA.

Type species: Omalodera limbata Blanchard, 1842, by subsequent designation (Bousquet 2002: 36).

Family-group name: HOMALODERINI Jeannel, 1926. Stem: Omaloder-.

Omalodes Dejean, 1833b: 128. Current status: valid genus in HISTERIDAE: HISTERINAE: OMALODINI.

Type species: Hister omega W. Kirby, 1818, by subsequent designation (Hope 1841a: 105).

Family-group name: OMALODINI Kryzhanovskij, 1972. Stem: Omalod-.

Omaloplia Schönherr, 1817: 178. Current status: valid genus in SCARABAEIDAE: SERICINAE: SERICINI: SERICINA.

Type species [suggested]: Melolontha ruricola Fabricius, 1775, by subsequent designation (Stephens 1832c: 628). Note: the valid type species fixation is that of Thon (1832b: 347), who selected Scarabaeus brunneus Linnaeus, 1758, but that nominal species is currently the type species of the genus Serica W.S. MacLeay, 1819; an application to the Commission is necessary to fix Melolontha ruricola Fabricius, 1775 as the type species of Omaloplia Schönherr, 1817.

Family-group name: OMALOPLIITES Blanchard, 1845. Stem: Omalopli-.

Omethes J.L. LeConte, 1861: 187. Current status: valid genus in OMETHIDAE: OMETHINAE.

Type species: Omethes marginatus J.L. LeConte, 1861, by monotypy.

Family-group name: OMETHES J.L. LeConte, 1861. Stem: Ometh-.

Ometis Latreille, 1829a: 554. Current status: junior synonym of Lagochile Hoffmannsegg, 1817 in SCARABAEIDAE: RUTELINAE: RUTELINI: RUTELINA.

Type species: Rutela cetonioides Lepeletier & Audinet-Serville, 1825, by subsequent designation (Brullé 1838: 349).

Family-group name: OMETIDAE Streubel, 1846. Stem: Omet-.

Omias Germar, 1817b: 341. Current status: valid genus in CURCULIONIDAE: ENTIMINAE: CYPHICERITAE: OMIINI.

Type species: Curculio rotundatus Fabricius, 1792 (= Omias puberulus Boheman, 1834), by subsequent designation (Schönherr 1823: 1144).

Family-group name: OMIADAE Shuckard, 1839. Stem: Omi-. Note: senior homonym of OMIINI Beck, 1996 (type genus Omia Hübner, 1821) in Lepidoptera.

Omicrus Sharp, 1879: 81. Current status: valid genus in HYDROPHILIDAE: SPHAERIDIINAE: OMICRINI.

Type species: Omicrus brevipes Sharp, 1879, by monotypy.

Family-group name: OMICRINI Smetana, 1975. Stem: Omicr-.

Omileus G.H. Horn in J.L. LeConte and G.H. Horn, 1876: 102. Current status: valid genus in CURCULIONIDAE: ENTIMINAE: POLYDRUSITAE: EUSTYLINI: EUSTYLINA.

Type species: Omileus epicaeroides G.H. Horn, 1876, by monotypy.

Family-group name: OMILEI G.H. Horn, 1876. Stem: Omile-.

Omma Newman, 1839: 303. Current status: valid genus in OMMATIDAE: OMMATINAE.

Type species: Omma stanleyi Newman, 1839, by monotypy.

Family-group name: OMMADIDAE Sharp & Muir, 1912. Stem: Ommat-.

Ommatolampes Schönherr, 1838: 837. Current status: valid genus in CURCULIONIDAE: DRYOPHTHORINAE: OMMATOLAMPINI.

Type species: Calandra haemorrhoidalis Wiedemann, 1819, by original designation.

Family-group name: OMMATOLAMPIDES Lacordaire, 1865. Stem: Ommatolamp-.

Omocerus Chevrolat, 1835c: no 119. Current status: valid genus in CHRYSOMELIDAE: CASSIDINAE: OMOCERINI. Note: the spelling Omocerus was changed to Omocera in the corrections ([p. 7] at the end of the work) issued in the same volume; therefore, the correct original spelling is Omocera but Omocerus, an incorrect subsequent spelling used subsequently by several authors, is in prevailing usage, attributed to the publication of the original spelling, and therefore deemed to be the correct original spelling (ICZN 1999a, Article 33.3.1).

Type species: Cassida bicornis Linnaeus, 1763, by subsequent designation (Duponchel and Chevrolat 1842b: 210).

Family-group name: OMOCERINI Hincks, 1952. Stem: Omocer-.

Omoglymmius Ganglbauer, 1891: 533, 534. Current status: valid genus in CARABIDAE: RHYSODINAE: OMOGLYMMIINI.

Type species: Rhysodes germari Ganglbauer, 1891, by monotypy.

Family-group name: OMOGLYMMIINA R.T. Bell & J.R. Bell, 1978. Stem: Omoglymmi-.

Omolabus Jekel, 1860: 191. Current status: valid genus in ATTELABIDAE: ATTELABINAE: ATTELABINI: OMOLABINA.

Type species: Attelabus bifoveatus Jekel, 1860, by original designation.

Family-group name: OMOLABINA Legalov, 2003. Stem: Omolab-.

Omophlus Dejean, 1834: 213. Current status: valid genus in TENEBRIONIDAE: ALLECULINAE: CTENIOPODINI. Note: the earlier usage of the name Omophlus by Dahl (1823: 46) was suppressed for the purposes of zoological nomenclature (see ICZN 1964).

Type species: Cistela lepturoides Fabricius, 1787, by subsequent designation (Solier 1835b: 246).

Family-group name: OMOPHLIENS Mulsant, 1856. Stem: Omophl-.

Omophorus Schönherr, 1835: 479. Current status: valid genus in CURCULIONIDAE: MOLYTINAE: METATYGINI.

Type species: Omophorus stomachosus Boheman, 1835, by original designation.

Family-group name: OMOPHORINAE G.A.K. Marshall, 1917. Stem: Omophor-.

Omophron Latreille, 1802b: 89. Current status: valid genus in CARABIDAE: OMOPHRONINAE. Note: new replacement name for Scolytus Fabricius, 1790.

Type species: Carabus limbatus Fabricius, 1777, by subsequent designation (Latreille 1810: 426).

Family-group name: OMOPHRONII Bonelli, 1810. Stem: Omophron-.

Omorgus Erichson, 1847c: 111. Current status: valid genus in TROGIDAE: OMORGINAE.

Type species: Trox suberosus Fabricius, 1775, by subsequent designation (Lacordaire 1855: 151).

Family-group name: OMORGINI Nikolajev, 2005. Stem: Omorg-.

Omphra Dejean, 1825: 285. Current status: valid genus in CARABIDAE: HARPALINAE: HELLUONINI: OMPHRINA. Note: this name was first published as a junior synonym of Helluo Bonelli, 1813 and, because it was treated as an available and valid name prior to 1961 (e.g., Reiche 1843b: 330), it is available and dates from its first publication as a synonym (ICZN 1999a, Article 11.6.1).

Type species: Galerita hirta Fabricius, 1801, by monotypy.

Family-group name: OMPHRINI Jedlička, 1941. Stem: Omphr-.

Omphreus Dejean, 1828: 4, 93. Current status: valid genus in CARABIDAE: HARPALINAE: OMPHREINI.

Type species: Omphreus morio Dejean, 1828, by monotypy.

Family-group name: OMPHREINI Ganglbauer, 1891. Stem: Omphre-.

Omus Eschscholtz, 1829b: 3, 4. Current status: valid genus in CICINDELIDAE: MANTICORINI.

Type species: Omus californicus Eschscholtz, 1829, by monotypy.

Family-group name: OMITES W. Horn, 1907. Stem: Om-.

Oncerus J.L. LeConte, 1856b: 283. Current status: valid genus in SCARABAEIDAE: ONCERINAE.

Type species: Oncerus floralis J.L. LeConte, 1856, by monotypy.

Family-group name: ONCERINI J.L. LeConte, 1861. Stem: Oncer-.

Oncideres Lacordaire, 1830b: 183. Current status: valid genus in CERAMBYCIDAE: LAMIINAE: ONCIDERINI. Note: the original spelling was Oncyderes but Oncideres, an incorrect subsequent spelling, is in prevailing usage, attributed to the publication of the original spelling, and therefore deemed to be the correct original spelling (ICZN 1999a, Article 33.3.1).

Type species: Lamia vomicosa Germar, 1823 (= Lamia saga Dalman, 1823), by subsequent designation (J. Thomson 1864: 104). Note: no species were originally included in this genus by Lacordaire (1830b); the first ones were the two species listed by Audinet- Serville (1835a: 68).

Family-group name: ONCIDERITAE J. Thomson, 1860. Stem: Oncider-.

Oncideropsis Aurivillius, 1922b: 165. Current status: junior synonym of Phelipara Pascoe, 1866 in CERAMBYCIDAE: LAMIINAE: ONCIDEROPSIDINI.

Type species: Oncideropsis nebulosa Aurivillius, 1922, by monotypy.

Family-group name: ONCIDEROPSIDINI Aurivillius, 1922. Stem: Oncideropsid-.

Oncocephala Agassiz, 1846: 259. Current status: junior synonym of Onchocephala Guérin-Méneville, 1844 in CHRYSOMELIDAE: CASSIDINAE: ONCOCEPHALINI. Note: unjustified emendation of Onchocephala Guérin-Méneville, 1844; Bouchard and Bousquet (2020a: 109) erroneously treated Onchocephala Guérin-Méneville, 1844 as a junior homonym of Onchocephala de Blainville, 1828 [Annelides], the name in Annelides is a suprageneric name and does not enter into homonymy at the genus-group level.

Type species: Oncocephala quadrilobata Guérin-Méneville, 1844, by subsequent designation (Gressitt 1957: 265).

Family-group name: ONCOCÉPHALITES Chapuis, 1875. Stem: Oncocephal-.

Oncomera Stephens, 1829b: 20. Current status: valid subgenus of Oedemera G.-A. Olivier, 1789 in OEDEMERIDAE: OEDEMERINAE: OEDEMERINI. Note: name placed on the Official List of Generic Names in Zoology (ICZN 1988f).

Type species: Dryops femorata Fabricius, 1792 (= Oedemera femoralis G.-A. Olivier, 1803), by plenary power (see ICZN 1988f). Note: Dryops femorata Fabricius, 1792 was placed on the Official List of Specific Names in Zoology (ICZN 1988f).

Family-group name: ONCOMERADAE Gistel, 1856. Stem: Oncomer-.

Oncosoma Westwood, 1843c: 121. Current status: junior synonym of Amatodes Dejean, 1834 in TENEBRIONIDAE: BLAPTINAE: PEDININI: HELOPININA. Note: the original spelling was Ogcosoma but the unjustified emendation Oncosoma, introduced by Agassiz (1846: 257, 259), is in prevailing usage, attributed to the original publication, and therefore deemed to be a justified emendation and the correct spelling (ICZN 1999a, Article 33.2.3.1).

Type species: Oncosoma granulare Westwood, 1843 (= Pimelia gemmata Fabricius, 1801), by monotypy.

Family-group name: ONCOSOMINA C. Koch, 1958. Stem: Oncosom-.

Oncotus Blanchard, 1845a: 13, 24. Current status: valid genus in TENEBRIONIDAE: BLAPTINAE: PLATYNOTINI: EURYNOTINA. Note: this genus was described under the French vernacular name Oncote (p. 13) but listed in the same work under the scientific name Oncotus (p. 24).

Type species: Oncotus farctus Solier, 1848, by subsequent designation (Gebien 1938b: 72). Note: no species were originally included in this genus by Blanchard (1845a); the first ones were the five species described by Solier (1848b: 216–222).

Family-group name: ONCOTINI C. Koch, 1953. Stem: Oncot-.

Oniticellus Dejean, 1821: 53. Current status: valid genus in SCARABAEIDAE: SCARABAEINAE: ONITICELLINI: ONITICELLINA.

Type species [suggested]: Scarabaeus cinctus Fabricius, 1775, by subsequent designation (Arrow 1931: 375). Note: the valid type species fixation is that of Thon (1832b: 466), who selected Scarabaeus flavipes Fabricius, 1792, but that species is currently placed in the genus Euoniticellus A. Janssens, 1953; an application to the Commission is necessary to fix Scarabaeus cinctus Fabricius, 1775 as the type species of Oniticellus Dejean, 1821.

Family-group name: ONITICELLINI Kolbe, 1905. Stem: Oniticell-.

Onitis Fabricius, 1798: 2. Current status: valid genus in SCARABAEIDAE: SCARABAEINAE: ONITINI.

Type species: Scarabaeus sphinx Fabricius, 1775 [p. 25] (= Onitis belial Fabricius, 1798), by subsequent designation (Latreille 1810: 428). Note: Fabricius (1798: 25–27) listed eight species in his new genus Onitis including O. sphinx (p. 26), which he referred to the Scarabaeus sphinx of his work of 1792 (p. 53); in 1792, Fabricius referred his S. sphinx to the Scarabaeus sphinx of G.-A. Olivier (1789b: [3] 135) who in turn referred the species to Fabricius (1775: 25); Fabricius (1775) described two species under the name Scarabaeus sphinx, one on page 14 (from “Sierra Leon Africae”), the other on page 25 (from “America”), and the type species of Onitis is, following the sequence described above, the Scarabaeus sphinx described by Fabricius (1775: 25) from “America,” which is, according to recent literature (e.g., Branco 2007: 16), a senior synonym of Onitis belial Fabricius, 1798 described from “Cajennae”; the genus Onitis, except for adventive species in the Americas and Australia, is restricted to the southern Palaearctic, Afrotropical and Oriental Regions (Palestrini et al. 2002: 97) suggesting that the provenance given by Fabricius (1775 and 1798) for Scarabaeus sphinx (p. 25) and Onitis belial (p. 27) is incorrect.

Family-group name: ONITIDES Laporte, 1840. Stem: Onit-.

Onocephala Sturm, 1843: 262. Current status: valid genus in CERAMBYCIDAE: LAMIINAE: ONCIDERINI.

Type species: Saperda diophthalma Perty, 1830, by monotypy.

Family-group name: ONOCEPHALITAE J. Thomson, 1860. Stem: Onocephal-.

Onthophagus Latreille, 1802b: 141. Current status: valid genus in SCARABAEIDAE: SCARABAEINAE: ONTHOPHAGINI.

Type species: Scarabaeus taurus Schreber, 1759, by monotypy.

Family-group name: ONTHOPHAGIDAE Streubel, 1846. Stem: Onthophag-.

Onthophilus Leach, 1817: 76. Current status: valid genus in HISTERIDAE: ONTHOPHILINAE.

Type species: Hister striatus Forster, 1771, by subsequent designation (Curtis 1828: pl. 220). Note: the type species often listed for this genus is Hister sulcatus Moll, 1784 (= Scolytus punctatus O.F. Müller, 1776) by subsequent designation of Westwood (1838c: 22) (e.g., Mazur 1997: 7; Bousquet and Laplante 2006: 423; Lackner et al. 2015: 110) but Curtis’ designation is older; both species are currently included in the genus Onthophilus Leach, 1817.

Family-group name: ONTHOPHILIDAE W.S. MacLeay, 1819. Stem: Onthophil-.

Onychius Chapuis, 1869: 48. Current status: valid genus in CURCULIONIDAE: COSSONINAE: ONYCHIINI.

Type species: Onychius nitidus Chapuis, 1869, by monotypy.

Family-group name: ONYCHIIDAE Chapuis, 1869. Stem: Onychi-. Note: junior homonym of ONYCHIINAE Forster, 1869 (type genus Onychia Haliday, 1830) in Hymenoptera.

Onychocerus Lacordaire, 1830b: 181. Current status: valid genus in CERAMBYCIDAE: LAMIINAE: ANISOCERINI.

Type species: Cerambyx scorpio Fabricius, 1781, by monotypy.

Family-group name: ONYCHOCERITAE J. Thomson, 1864. Stem: Onychocer-.

Onychoglenea Aurivillius, 1922a: 28. Current status: valid genus in CERAMBYCIDAE: LAMIINAE: ONYCHOGLENEINI.

Type species: Onychoglenea brunnea Aurivillius, 1922, by monotypy.

Family-group name: ONYCHOGLENEINI Aurivillius, 1923. Stem: Onychoglene-.

Onycholips Wollaston, 1861c: 389. Current status: valid genus in CURCULIONIDAE: COSSONINAE: ONYCHOLIPINI.

Type species: Onycholips bifurcatus Wollaston, 1861, by monotypy.

Family-group name: ONYCHOLIPIDES Wollaston, 1873. Stem: Onycholip-.

Oochrotus P.H. Lucas, 1852: xxix. Current status: junior synonym of Phtora Germar, 1836 in TENEBRIONIDAE: DIAPERINAE: CRYPTICINI.

Type species: Oochrotus unicolor P.H. Lucas, 1852, by monotypy.

Family-group name: OOCHROTINI Desbrochers des Loges, 1901. Stem: Oochrot-.

Oocyclus Sharp, 1882c: 61. Current status: valid genus in HYDROPHILIDAE: HYDROPHILINAE: LACCOBIINI.

Type species: Oocyclus brevicornis Sharp, 1882, by subsequent designation (Knisch 1924: 181).

Family-group name: OOCYCLINI Hansen, 1991. Stem: Oocycl-.

Oodes Bonelli, 1810: Tab. Syn. Current status: valid genus in CARABIDAE: HARPALINAE: OODINI.

Type species: Carabus helopioides Fabricius, 1792, by subsequent monotypy (Panzer 1813: 51).

Family-group name: OODITES LaFerté-Sénectère, 1851. Stem: Ood-.

Oophorus Dejean, 1833a: 93. Current status: junior synonym of Aeolus Eschscholtz, 1829 in ELATERIDAE: AGRYPNINAE: OOPHORINI.

Type species: Elater elegans Fabricius, 1792, by subsequent designation (Hyslop 1921: 659).

Family-group name: OOPHORIDAE Gistel, 1848. Stem: Oophor-.

Oopterus Guérin-Méneville, 1841: 123. Current status: valid genus in CARABIDAE: TRECHINAE: ZOLINI.

Type species: Oopterus clivinoides Guérin-Méneville, 1841, by monotypy.

Family-group name: OOPTERINI Jeannel, 1940. Stem: Oopter-.

Oosomus Schönherr, 1826: 195. Current status: valid genus in CURCULIONIDAE: ENTIMINAE: CYPHICERITAE: OOSOMINI.

Type species: Peritelus hariolus Boheman, 1834, by subsequent designation (Alonso-Zarazaga and Lyal 1999: 22, 167).

Family-group name: OOSOMIDES Lacordaire, 1863. Stem: Oosom-.

Oosternum Sharp, 1882c: 112. Current status: valid genus in HYDROPHILIDAE: SPHAERIDIINAE: MEGASTERNINI: OOSTERNINA. Note: name placed on the Official List of Generic Names in Zoology (ICZN 1998).

Type species: Oosternum costatum Sharp, 1882, by monotypy. Note: name placed on the Official List of Specific Names in Zoology (ICZN 1998).

Family-group name: OOSTERNINA Arriaga-Varela & Fikáček in Arriaga-Varela et al. 2021: 577. Stem: Oostern-.

Opanassenkovius Legalov, 2003: 555. Current status: valid genus in ATTELABIDAE: APODERINAE: APODERINI.

Type species: Apoderus verrucosus Pascoe, 1881, by original designation.

Family-group name: OPANASSENKOVIINA Legalov, 2003. Stem: Opanassenkovi-.

Opatrinus Dejean, 1821: 66. Current status: valid genus in TENEBRIONIDAE: BLAPTINAE: PLATYNOTINI: PLATYNOTINA.

Type species: Opatrum clathratum Fabricius, 1787, by monotypy.

Family-group name: OPATRINAIRES Mulsant & Rey, 1853. Stem: Opatrin-.

Opatrum Fabricius, 1775: 76. Current status: valid genus in TENEBRIONIDAE: BLAPTINAE: OPATRINI: OPATRINA.

Type species: Silpha sabulosa Linnaeus, 1758, by subsequent designation (Latreille 1810: 429).

Family-group name: OPATRITES Brullé, 1832. Stem: Opatr-.

Ophioglossa Fauvel, 1866: 255, 259. Current status: valid genus in STAPHYLINIDAE: ALEOCHARINAE: AUTALIINI.

Type species: Ophioglossa araucana Fauvel, 1866, by monotypy.

Family-group name: OPHIOGLOSSAE Fenyes, 1919. Stem: Ophiogloss-.

Ophonus Dejean, 1821: 13. Current status: valid genus in CARABIDAE: HARPALINAE: HARPALINI: HARPALINA. Note: name placed on the Official List of Generic Names in Zoology (ICZN 1990c).

Type species: Carabus sabulicola Panzer, 1796, by plenary power (see ICZN 1990c). Note: name placed on the Official List of Specific Names in Zoology (ICZN 1990c).

Family-group name: OPHONIDAE Laporte, 1834. Stem: Ophon-.

Ophryastes Germar, 1829: 358. Current status: valid genus in CURCULIONIDAE: ENTIMINAE: ENTIMITAE: OPHRYASTINI: OPHRYASTINA.

Type species: Liparus vittatus Say, 1824, by monotypy.

Family-group name: OPHRYASTIDES Lacordaire, 1863. Stem: Ophryast-.

Ophtalmorrhynchus Hoffmann, 1965: 1408. Current status: valid genus in CURCULIONIDAE: ENTIMINAE: POLYDRUSITAE: DERMATODINI.

Type species: Ophtalmorrhynchus descarpentriesi Hoffmann, 1965, by original designation.

Family-group name: OPHTALMORYNCHINI Hoffmann, 1965. Stem: Ophtalmorrhynch-.

Opilo Latreille, 1802b: 111. Current status: valid genus in CLERIDAE: CLERINAE: OPILONITAE: OPILONINI: OPILONINA.

Type species: Attelabus mollis Linnaeus, 1758, by monotypy.

Family-group name: OPILONIDAE Gistel, 1848. Stem: Opilon-.

Opisthius W. Kirby, 1837: 60. Current status: valid genus in CARABIDAE: NEBRIINAE: OPISTHIINI.

Type species: Opisthius richardsoni W. Kirby, 1837, by monotypy.

Family-group name: OPISTHIINAE P. Dupuis, 1912. Stem: Opisthi-.

Oplosia Mulsant, 1863: 300. Current status: valid genus in CERAMBYCIDAE: LAMIINAE: ACANTHODERINI.

Type species: Cerambyx fennicus Paykull, 1800 (= Exocentrus cinereus Mulsant, 1839), by monotypy.

Family-group name: HOPLOSIAE J.L. LeConte & G.H. Horn, 1883. Stem: Oplosi-.

Oplostomus W.S. MacLeay, 1838: 20. Current status: valid genus in SCARABAEIDAE: CETONIINAE: CREMASTOCHEILINI: OPLOSTOMINA.

Type species: Cetonia fuliginea G.-A. Olivier, 1789, by monotypy.

Family-group name: OPLOSTOMINA Krikken, 1984. Stem: Oplostom-.

Opresus Casey, 1897: 493, 494. Current status: junior synonym of Microscydmus Saulcy & Croissandeau, 1893 in STAPHYLINIDAE: SCYDMAENINAE: SCYDMAENITAE: STENICHNINI.

Type species: Scydmaenus misellus J.L. LeConte, 1852, by subsequent designation (H. Franz 1985: 175).

Family-group name: OPRESINI Casey, 1897. Stem: Opres-.

Opsimus Mannerheim, 1843: 305. Current status: valid genus in CERAMBYCIDAE: CERAMBYCINAE: OPSIMINI.

Type species: Opsimus quadrilineatus Mannerheim, 1843, by monotypy.

Family-group name: OPSIMI J.L. LeConte, 1873. Stem: Opsim-.

Optatus Pascoe, 1889: 322, 327. Current status: valid genus in CURCULIONIDAE: BARIDINAE: OPTATINI.

Type species: Centrinus palmaris Pascoe, 1889, by original designation.

Family-group name: OPTATIDES Champion, 1907. Stem: Optat-.

Orchesia Latreille, 1806: 194. Current status: valid genus in MELANDRYIDAE: MELANDRYINAE: ORCHESIINI.

Type species: Hallomenus micans Panzer, 1793, by monotypy.

Family-group name: ORCHÉSIENS Mulsant, 1856. Stem: Orchesi-.

Orchestes Illiger, 1798: 498. Current status: valid genus in CURCULIONIDAE: CURCULIONINAE: RHAMPHINI: RHAMPHINA.

Type species: Orchestes signifer Creutzer, 1799 (= Curculio hortorum Fabricius, 1792), by subsequent monotypy (Creutzer 1799: 125).

Family-group name: ORCHESTIDES Latreille, 1828. Stem: Orchest-.

Orchymontia Broun, 1919: 108. Current status: valid genus in HYDRAENIDAE: ORCHYMONTIINAE.

Type species: Orchymontia spinipennis Broun, 1919, by monotypy.

Family-group name: ORCHYMONTINAE Perkins, 1997. Stem: Orchymonti-.

Orectochilus Dejean, 1833a: 59. Current status: valid genus in GYRINIDAE: GYRININAE: ORECTOCHILINI.

Type species: Gyrinus villosus O.F. Müller, 1776, by subsequent designation (Westwood 1838c: 8).

Family-group name: ORECTOCHILINI Kolbe, 1880. Stem: Orectochil-.

Oreina Chevrolat in Dejean, 1836a: 402. Current status: valid genus in CHRYSOMELIDAE: CHRYSOMELINAE: CHRYSOMELINI.

Type species: Chrysomela speciosa Linnaeus, 1767, by subsequent designation (Chevrolat 1843a: 656).

Family-group name: OREININI Bechyné, 1958. Stem: Orein-. Note: junior homonym of OREININI Bleeker, 1863 (type genus Oreinus McClelland, 1839) in Pisces.

Oreodera Audinet-Serville, 1835a: 19. Current status: valid genus in CERAMBYCIDAE: LAMIINAE: ACANTHODERINI.

Type species: Cerambyx glaucus Linnaeus, 1758, by subsequent designation (E. Desmarest 1860: 321).

Family-group name: OREODERITAE J. Thomson, 1860. Stem: Oreoder-.

Oritocatops Jeannel, 1921: 235, 236. Current status: valid genus in LEIODIDAE: CHOLEVINAE: ORITOCATOPINI.

Type species: Oritocatops kenyensis Jeannel, 1921, by original designation.

Family-group name: ORITOCATOPINI Jeannel, 1936. Stem: Oritocatop-.

Ormiscus G.R. Waterhouse, 1845: 37. Current status: valid genus in ANTHRIBIDAE: ANTHRIBINAE: ZYGAENODINI.

Type species: Ormiscus variegatus G.R. Waterhouse, 1845, by monotypy.

Family-group name: HORMISCI J.L. LeConte, 1876. Stem: Ormisc-.

Ornithognathus J. Thomson, 1858b: 215. Current status: valid genus in CHRYSOMELIDAE: GALERUCINAE: LUPERINI.

Type species: Ornithognathus generosus J. Thomson, 1858, by monotypy.

Family-group name: ORNITHOGNATHITES Chapuis, 1875. Stem: Ornithognath-.

Orobitis Germar, 1817b: 340. Current status: valid genus in CURCULIONIDAE: OROBITIDINAE.

Type species: Attelabus globosus Fabricius, 1792 (= Curculio cyaneus Linnaeus, 1758), by monotypy.

Family-group name: OROBITINA C.G. Thomson, 1859. Stem: Orobitid-.

Orodotes Bonvouloir, 1871: 74. Current status: valid genus in EUCNEMIDAE: MACRAULACINAE: ORODOTINI. Note: this name was first used by Gemminger in Gemminger and von Harold (1869b: 1481) but is unavailable from that date since it was not accompanied by a description or indication (ICZN 1999a, Article 12.1).

Type species: Orodotes jansoni Bonvouloir, 1875, by subsequent monotypy (de Bonvouloir 1875: 717).

Family-group name: ORODOTINI Muona, 1993. Stem: Orodot-.

Orophius L. Redtenbacher, 1847c: 350. Current status: junior synonym of Octotemnus Mellié, 1847 in CIIDAE: CIINAE: OROPHIINI.

Type species: Cis mandibularis Gyllenhal, 1813, by monotypy.

Family-group name: OROPHIINA C.G. Thomson, 1863. Stem: Orophi-. Note: senior homonym of OROPHIINI Lvovsky, 1974 (type genus Orophia Hübner, 1825) in Lepidoptera; the lepidopteran name was recently replaced with EUTORNINI Lvovsky, 2019 but such action is contrary to Article 55.3.1 (ICZN 1999a), which requires that the case be referred to the Commission for a ruling to remove the homonymy.

†Oropsis Legalov & Kirejtshuk in Legalov et al., 2017: 112. Current status: valid genus in NEMONYCHIDAE: †PALEOCARTINAE: †OROPSINI. Note: Lower Cretaceous (Lebanon).

Type species: †Oropsis marinae Legalov & Kirejtshuk, 2017, by original designation.

Family-group name: †OROPSINI Legalov & Kirejtshuk in Legalov et al., 2017: 112. Stem: Orops-.

Orphilus Erichson, 1846: 424, 461. Current status: valid genus in DERMESTIDAE: ORPHILINAE.

Type species: Anthrenus glabratus Fabricius, 1801 (= Byrrhus niger Rossi, 1790), by subsequent designation (Lacordaire 1854: 473).

Family-group name: ORPHILI J.L. LeConte, 1861. Stem: Orphil-.

Orphinus Motschulsky, 1858a: 48. Current status: valid genus in DERMESTIDAE: MEGATOMINAE: MEGATOMINI: ORPHININA.

Type species: Orphinus haemorrhoidalis Motschulsky, 1858, by subsequent designation (Arrow 1915: 437). Note: the spelling haemorrhoidalis, which is an incorrect subsequent spelling of the original spelling haemorhoidalis, is in prevailing usage, attributed to the publication of the original spelling, and therefore deemed to be the correct original spelling (ICZN 1999a, Article 33.3.1).

Family-group name: ORPHININA Zhou, Nicholls, Liu, Hartley, Szito, Ślipiński & Zwick, 2022: 18. Stem: Orphin-.

Orphnus W.S. MacLeay, 1819: 119. Current status: valid genus in SCARABAEIDAE: ORPHNINAE: ORPHNINI.

Type species: Geotrupes bicolor Fabricius, 1801, by monotypy.

Family-group name: ORPHNIDAE Erichson, 1847. Stem: Orphn-.

Orropygia Raffray, 1910: 419. Current status: valid genus in STAPHYLINIDAE: PSELAPHINAE: BATRISITAE: BATRISINI: BATRISINA.

Type species: Orropygia myrmecophila Raffray, 1910, by monotypy.

Family-group name: OROPYGIINA Jeannel, 1949. Stem: Orropygi-.

Orsodacne Latreille, 1802b: 223. Current status: valid genus in ORSODACNIDAE: ORSODACNINAE. Note: name placed on the Official List of Generic Names in Zoology (ICZN 2002a).

Type species: Chrysomela cerasi Linnaeus, 1758, by plenary power (see ICZN 2002a). Note: name placed on the Official List of Specific Names in Zoology (ICZN 2002a).

Family-group name: ORSODACHNIDAE C.G. Thomson, 1859. Stem: Orsodacn-.

Ortalia Mulsant, 1850: 893. Current status: valid genus in COCCINELLIDAE: COCCINELLINAE: ORTALIINI.

Type species: Ortalia variata Mulsant, 1850, by subsequent designation (Crotch 1874a: 275).

Family-group name: ORTALIENS Mulsant, 1850. Stem: Ortali-.

Orthocerus Latreille, 1797: 16. Current status: valid genus in ZOPHERIDAE: COLYDIINAE: ORTHOCERINI. Note: name placed on the Official List of Generic Names in Zoology (ICZN 1995d, as Orthocerus Latreille, 1796).

Type species: Hispa mutica Linnaeus, 1767 (= Dermestes clavicornis Linnaeus, 1758), by subsequent monotypy (Latreille 1802b: 173, as Sarrotrium muticum F.). Note: in Opinion 1811 (ICZN 1995d), the Commission listed the type species of Orthocerus as Tenebrio hirticornis DeGeer, 1775 (= Dermestes clavicornis Linnaeus, 1758) by subsequent monotypy in Latreille (1807: 172); however, Latreille (1802b: 173) listed a different nominal species by name (also considered a junior synonym of Dermestes clavicornis Linnaeus, 1758) under Orthocerus in an earlier publication, hence the change of type species as reported here; the original spelling Hispa muticus Linnaeus, 1797 was corrected to H. mutica in the “Errata” section of the same work on page [37]; the valid name of the type species, Dermestes clavicornis Linnaeus, 1758, was already placed on the Official List of Specific Names in Zoology (ICZN 1995d).

Family-group name: ORTHOCÉRITES Blanchard, 1845. Stem: Orthocer-.

Orthochaetes Germar, 1823: 302. Current status: valid genus in CURCULIONIDAE: CURCULIONINAE: STYPHLINI.

Type species: Rhynchaenus setiger Beck, 1817, by monotypy.

Family-group name: ORTHOCHAETINAE Joy, 1932. Stem: Orthochaet-.

Orthognathus Schönherr, 1838: 812. Current status: valid genus in CURCULIONIDAE: DRYOPHTHORINAE: ORTHOGNATHINI.

Type species: Orthognathus lividus Gyllenhal, 1838, by original designation.

Family-group name: ORTHOGNATHIDES Lacordaire, 1865. Stem: Orthognath-.

Orthogonius W.S. MacLeay, 1825: 26. Current status: valid genus in CARABIDAE: HARPALINAE: ORTHOGONIINI. Note: this name was also proposed in the same year by Dejean (1825: 169, 279 [by 5 September]) subsequently to W.S. MacLeay’s publication dated 1 June.

Type species: Carabus duplicatus Wiedemann, 1819, by subsequent designation (Hope 1838a: 85).

Family-group name: ORTHOGONIDEN Schaum, 1857. Stem: Orthogoni-.

Orthomegas Audinet-Serville, 1832: 149. Current status: valid genus in CERAMBYCIDAE: PRIONINAE: CALLIPOGONINI.

Type species: Cerambyx corticinus G.-A. Olivier, 1790 (= Cerambyx cinnamomeus Linnaeus, 1758), by subsequent designation (E. Desmarest 1860: 307).

Family-group name: ORTHOMEGITAE J. Thomson, 1864. Stem: Orthomegal-.

Orthoperus Stephens, 1829a: 158. Current status: valid genus in CORYLOPHIDAE: CORYLOPHINAE: ORTHOPERINI.

Type species: Dermestes punctum Marsham, 1802, by subsequent designation (Westwood 1838c: 11). Note: no species were originally included in this genus by Stephens (1829a); the first ones were the six species listed by Stephens (1829a: 186, 187); Bowestead (2007: 634) listed the type species as a synonym of Cryptophagus atomus Gyllenhal, 1808 but mentioned that the assignment is doubtful.

Family-group name: ORTHOPÉRITES Jacquelin du Val, 1857. Stem: Orthoper-.

Orthopleura Spinola, 1845b: 80. Current status: junior synonym of Dermestoides Schäffer, 1777 in CLERIDAE: KORYNETINAE. Note: three original spellings were used, Orthoplevra (pp. 80–83, 151, 167), Orthopleura (pl. xxxii), and Orthopleuva (pl. xlii); as far as we know, no one has acted as First Reviser (ICZN 1999a, Article 24.2.3) and therefore we here select Orthopleura as the correct original spelling, i.e., the spelling in prevailing usage.

Type species: Dermestes sanguinicollis Fabricius, 1787, by subsequent designation (Audinet- Serville 1845: 430).

Family-group name: ORTHOPLEURINAE Böving & Craighead, 1931. Stem: Orthopleur-.

Orthorhinus Schönherr, 1825: 582. Current status: valid genus in CURCULIONIDAE: MOLYTINAE: PISSODINI: ORTHORHININA.

Type species: Curculio cylindrirostris Fabricius, 1775, by original designation.

Family-group name: ORTHORHINIDES Jekel, 1865. Stem: Orthorhin-.

Orthosoma Audinet-Serville, 1832: 155. Current status: valid genus in CERAMBYCIDAE: PRIONINAE: PRIONINI.

Type species: Prionus cylindricus Fabricius, 1775 (= Cerambyx brunneus Forster, 1771 [as brunnus]), by monotypy.

Family-group name: ORTHOSOMITAE J. Thomson, 1864. Stem: Orthosomat-.

Orvoenia Dajoz, 1980b: 190. Current status: junior synonym of Megauchenia W.S. MacLeay, 1825 in NITIDULIDAE: NITIDULINAE: NITIDULINI.

Type species: Orvoenia borneensis Dajoz, 1980, by original designation.

Family-group name: ORVOENINI Dajoz, 1980. Stem: Orvoeni-.

Orychodes Pascoe, 1862: 389. Current status: valid genus in BRENTIDAE: BRENTINAE: BRENTINI: ARRENODINA.

Type species: Brentus serrirostris Lund, 1802, by subsequent designation (Lacordaire 1865: 433).

Family-group name: ORYCHODI Senna, 1895. Stem: Orychod-.

Oryctes Hellwig, 1798: 100. Current status: valid genus in SCARABAEIDAE: DYNASTINAE: ORYCTINI. Note: see Alonso-Zarazaga and Krell (2011: 68) for comment regarding the authorship of this name.

Type species: Scarabaeus nasicornis Linnaeus, 1758, by subsequent designation (Latreille 1810: 428).

Family-group name: ORYCTÉSAIRES Mulsant, 1842. Stem: Oryct-.

Oryctoderus Boisduval, 1835: 160. Current status: valid genus in SCARABAEIDAE: DYNASTINAE: ORYCTODERINI.

Type species: Oryctoderus latitarsis Boisduval, 1835, by monotypy.

Family-group name: ORYCTODERINI Endrödi, 1966. Stem: Oryctoder-.

Oryctomorphus Guérin-Méneville, 1831a: pl. 3. Current status: valid genus in SCARABAEIDAE: RUTELINAE: RUTELINI: ORYCTOMORPHINA.

Type species: Oryctomorphus bimaculatus Guérin-Méneville, 1831, by monotypy.

Family-group name: ORYCTOMORPHIDAE Burmeister, 1847. Stem: Oryctomorph-.

Oryotus L. Miller, 1856: 627. Current status: valid genus in LEIODIDAE: CHOLEVINAE: LEPTODIRINI: BATHYSCIINA.

Type species: Oryotus schmidtii L. Miller, 1856, by monotypy.

Family-group name: ORIOTINI Reitter, 1889. Stem: Oryot-.

Oryssomus Mulsant, 1850: 939. Current status: valid genus in COCCINELLIDAE: COCCINELLINAE: COCCIDULINI.

Type species: Oryssomus subterminatus Mulsant, 1850, by monotypy.

Family-group name: ORYSSOMINI Gordon, 1974. Stem: Oryssom-.

Osmoderma Lepeletier & Audinet-Serville, 1828: 702. Current status: valid genus in SCARABAEIDAE: CETONIINAE: OSMODERMATINI. Note: name placed on the Official List of Generic Names in Zoology (ICZN 2007).

Type species: Scarabaeus eremita Scopoli, 1763, by subsequent designation (W.S. MacLeay 1838: 16; see ICZN 2007). Note: name placed on the Official List of Specific Names in Zoology (ICZN 2007).

Family-group name: OSMODERMINI Schenkling, 1922. Stem: Osmodermat-.

Osorius Latreille, 1829a: 438. Current status: valid genus in STAPHYLINIDAE: OSORIINAE: OSORIINI: OSORIINA.

Type species: Osorius brasiliensis Guérin-Méneville, 1829, by subsequent monotypy (Guérin- Méneville 1829a: pl. 9).

Family-group name: OSORINI Erichson, 1839. Stem: Osori-.

Osphryon Pascoe, 1969b: 662. Current status: valid genus in CERAMBYCIDAE: PRIONINAE: OSPHRYONINI.

Type species: Osphryon adustus Pascoe, 1869, by monotypy.

Family-group name: OSPHRYONINI Jin, De Keyzer, Ashman, Zwick & Ślipiński, 2023: 507. Stem: Osphryon-.

Osphya Illiger, 1807b: 370. Current status: valid genus in MELANDRYIDAE: OSPHYINAE. Note: in Volume 6 of ‘Magazin für Insektenkunde’, the name Pelecina was used by Illiger (1807a: 300) and Illiger (1807b: 333) but the unnecessary new replacement name Osphya was proposed instead in the ‘Verbesserung’ on page 370 by Illiger (1807c); both Pelecina and Osphya have the same publication date; Mulsant (1856b: 108–109) acted as First Reviser (ICZN 1999a, Article 24.2.2) when he used Osphya as valid.

Type species: Cantharis bipunctata Fabricius, 1775, by monotypy.

Family-group name: OSPHYENS Mulsant, 1856. Stem: Osphy-.

Osphyoplesius A. Winkler, 1915: 331. Current status: valid genus in PYTHIDAE.

Type species: Osphyoplesius anophthalmus A. Winkler, 1915, by monotypy.

Family-group name: OSPHYOPLESIINI A. Winkler, 1915. Stem: Osphyoplesi-.

Ostoma Laicharting, 1781: 102. Current status: valid genus in PELTIDAE. Note: this name became a senior synonym of Peltis, which has incorrectly been credited to O.F. Müller (1764: 13) in the literature (e.g., Kolibáč 2007: 366; Kolibáč et al. 2016: 683) since the Commission (ICZN 1994a) ruled that Peltis Geoffroy, 1762 and all other uses of the name prior to the publication of Peltis Kugelann, 1792 are suppressed for the purposes of both the Principle of Priority and the Principle of Homonymy.

Type species: Silpha ferruginea Linnaeus, 1758, by subsequent designation (Stephens 1830: 368).

Family-group name: OSTOMINI Harold, 1876. Stem: Ostomat-.

Ostomopsis H. Scott, 1922: 250. Current status: valid genus in CERYLONIDAE: OSTOMOPSINAE.

Type species: Ostomopsis solitaria H. Scott, 1922, by monotypy.

Family-group name: OSTOMOPSINI Sen Gupta & Crowson, 1973. Stem: Ostomops-.

Othippia Pascoe, 1874: 49. Current status: valid genus in CURCULIONIDAE: CONODERINAE: OTHIPPIINI.

Type species: Othippia distigma Pascoe, 1874, by subsequent designation (Setliff 2007: 94).

Family-group name: OTHIPPIINI Morimoto, 1962. Stem: Othippi-.

Othius Stephens, 1829b: 23. Current status: valid genus in STAPHYLINIDAE: STAPHYLININAE: OTHIINI. Note: name placed on the Official List of Generic Names in Zoology (ICZN 1983c).

Type species: Staphylinus punctulatus Goeze, 1777, by plenary power (see ICZN 1983c). Note: name placed on the Official List of Specific Names in Zoology (ICZN 1983c); this species name was published in a work that is not consistently binominal (see ICZN 2021a).

Family-group name: OTHIIDES C.G. Thomson, 1859. Stem: Othi-.

Othnius J.L. LeConte, 1861: 103. Current status: junior synonym of Elacatis Pascoe, 1860 in SALPINGIDAE: OTHNIINAE.

Type species: Othnius umbrosus J.L. LeConte, 1861, by subsequent designation (Packard 1869: 447).

Family-group name: OTHNIIDAE J.L. LeConte, 1861. Stem: Othni-.

Otidocephalus Chevrolat, 1832b: 100. Current status: junior synonym of Myrmex Sturm, 1826 in CURCULIONIDAE: CURCULIONINAE: OTIDOCEPHALINI. Note: the original spelling Otiocephalus was intentionally changed to Otidocephalus (p. 442) in a subsequent part (issued in 1833) of the same volume of the journal and therefore the original spelling is to be corrected (ICZN 1999a, Article 32.5.1.1).

Type species: Otidocephalus mexicanus Chevrolat, 1832, by subsequent designation (Schönherr 1835: 364).

Family-group name: OTIDOCÉPHALIDES Lacordaire, 1863. Stem: Otidocephal-.

Otiorhynchus Germar, 1822b: no 12. Current status: valid genus in CURCULIONIDAE: ENTIMINAE: OTIORHYNCHITAE: OTIORHYNCHINI. Note: this name was credited to “Germar, 1824” and placed on the Official List of Generic Names in Zoology in Opinion 982 (ICZN 1972) but the original work and date of publication for this name were corrected in Opinion 2299 (ICZN 2012c).

Type species: Curculio rhacusensis Germar, 1822, by monotypy (see ICZN 2012c, type fixation previously recorded as Curculio clavipes Bonsdorff, 1785, by subsequent designation in Schönherr 1826 in Opinion 982 (ICZN 1972)). Note: name placed on the Official List of Specific Names in Zoology (ICZN 2012c).

Family-group name: OTIORHYNCHIDES Schönherr, 1826. Stem: Otiorhynch-.

Ototreta E. Olivier, 1900: 237. Current status: junior synonym of Drilaster Kiesenwetter, 1879 in LAMPYRIDAE: OTOTRETINAE.

Type species: Ototreta fornicata E. Olivier, 1900, by subsequent designation (R. Lucas 1920: 468). Note: the type species often listed for this genus-group name is Ototreta weyersi E. Olivier, 1900 (e.g., Geisthardt and Satô 2007: 233) by subsequent designation of McDermott (1964: 47) but R. Lucas’ designation is earlier; both species are currently included in the genus Drilaster Kiesenwetter, 1879.

Family-group name: OTOTRETINAE McDermott, 1964. Stem: Ototret-.

Ototretadrilus Pic, 1921: 13. Current status: valid genus in LAMPYRIDAE: OTOTRETINAE.

Type species: Ototretadrilus atritarsis Pic, 1921, by subsequent designation (N.D. Riley 1923: 124). Note: the same type species was designated again by Janisova and Bocakova (2013: 6).

Family-group name: OTOTRETADRILINAE Crowson, 1972. Stem: Ototretadril-.

Ottistira Pascoe, 1872: 440. Current status: valid genus in CURCULIONIDAE: ENTIMINAE: POLYDRUSITAE: OTTISTIRINI.

Type species: Ottistira bispinosa Pascoe, 1872, by subsequent designation (Heller 1925a: 55).

Family-group name: OTTISTIRINI Heller, 1925. Stem: Ottistir-.

Ovosoma Motschulsky, 1860b: 214. Current status: valid genus in CHRYSOMELIDAE: CHRYSOMELINAE: CHRYSOMELINI.

Type species: Chrysomela vernalis Brullé, 1832, by original designation.

Family-group name: OVOSOMES Motschulsky, 1860. Stem: Ovosomat-.

Oxacis J.L. LeConte, 1866a: 164, 165. Current status: valid genus in OEDEMERIDAE: OEDEMERINAE: ASCLERINI.

Type species: Asclera cana J.L. LeConte, 1854, by subsequent designation (Arnett 1950: 223).

Family-group name: OXACIINI Arnett, 1984. Stem: Oxacid-.

Oxeostomum Gistel, 1856: 374. Current status: junior synonym of Apion Herbst, 1797 in BRENTIDAE: APIONINAE: APIONITAE: APIONINI: APIONINA.

Type species: Curculio frumentarius Linnaeus, 1758, by subsequent designation (Alonso- Zarazaga and Lyal 1999: 21, 57).

Family-group name: OXEOSTOMATIDAE Gistel, 1856. Stem: Oxeostom-.

Oxura W. Kirby, 1819: 413. Current status: valid genus in TENEBRIONIDAE: PIMELIINAE: SEPIDIINI: OXURINA.

Type species: Oxura setosa W. Kirby, 1819, by monotypy.

Family-group name: OXURINA C. Koch, 1955. Stem: Oxur-.

Oxycheila Dejean, 1825: 4, 15. Current status: valid genus in CICINDELIDAE: OXYCHEILINI.

Type species: Cicindela tristis Fabricius, 1775, by subsequent designation (Hope 1838a: 15).

Family-group name: OXYCHEILITES J. Thomson, 1857. Stem: Oxycheil-.

Oxycoleus Lacordaire, 1868: 484. Current status: valid genus in CERAMBYCIDAE: CERAMBYCINAE: OXYCOLEINI.

Type species: Oxycoleus clavipes Lacordaire, 1868, by monotypy.

Family-group name: OXYCOLEINI Martins & Galileo, 2003. Stem: Oxycole-.

Oxycopis Arnett, 1951: 305. Current status: valid genus in OEDEMERIDAE: OEDEMERINAE: ASCLERINI.

Type species: Necydalis notoxoides Fabricius, 1801, by original designation.

Family-group name: OXYCOPIINI Arnett, 1984. Stem: Oxycopid-.

†Oxycorynoides Arnoldi, 1977: 159. Current status: valid genus in NEMONYCHIDAE: †EOBELINAE: †OXYCORYNOIDINI. Note: Upper Jurassic (Kazakhstan).

Type species: †Oxycorynoides similis Arnoldi, 1977, by original designation.

Family-group name: †OXYCORYNOIDINAE Arnoldi, 1977. Stem: Oxycorynoid-.

Oxycorynus Chevrolat, 1832c: 212. Current status: valid genus in BELIDAE: OXYCORYNINAE: OXYCORYNITAE: OXYCORYNINI.

Type species: Oxycorynus melanocerus Chevrolat, 1832, by subsequent designation (Schönherr 1840a: 582).

Family-group name: OXYCORYNIDES Schönherr, 1840. Stem: Oxycoryn-.

Oxycraspedus Kuschel, 1955: 309. Current status: valid genus in BELIDAE: OXYCORYNINAE: OXYCORYNITAE: OXYCRASPEDINI.

Type species: Oxycorynus minutus R.A. Philippi & F. Philippi, 1864, by original designation.

Family-group name: OXYCRASPEDINA Marvaldi & Oberprieler, 2006. Stem: Oxycrasped-.

Oxygona Chevrolat, 1847a: 368. Current status: junior synonym of Platiprosopus Chevrolat, 1834 in CHRYSOMELIDAE: GALERUCINAE: ALTICINI.

Type species: Platiprosopus acutangulus Chevrolat, 1834, by monotypy.

Family-group name: OXYGONITES Chapuis, 1875. Stem: Oxygon-.

Oxylobus Chaudoir, 1855: 4, 5. Current status: valid genus in CARABIDAE: SCARITINAE: SCARITINI: OXYLOBINA.

Type species: Scarites lateralis Dejean, 1825, by subsequent designation (Motschulsky 1857b: 96).

Family-group name: OXYLOBIDES Andrewes, 1929. Stem: Oxylob-.

Oxymirus Mulsant, 1863: 464. Current status: valid genus in CERAMBYCIDAE: LEPTURINAE: OXYMIRINI.

Type species: Cerambyx cursor Linnaeus, 1758, by monotypy.

Family-group name: OXYMIRINI Danilevsky, 1997. Stem: Oxymir-.

Oxynopterus Hope, 1842c: 77. Current status: valid genus in ELATERIDAE: DENDROMETRINAE: OXYNOPTERINI.

Type species: Elater mucronatus G.-A. Olivier, 1792, by subsequent designation (Lacordaire 1857: 160).

Family-group name: OXYNOPTÉRIDES Candèze, 1857. Stem: Oxynopter-.

Oxyonyx Faust, 1885: 192. Current status: valid genus in CURCULIONIDAE: CEUTORHYNCHINAE: CEUTORHYNCHINI.

Type species: Coeliodes brisouti Faust, 1885, by subsequent designation (Hustache 1931: 246).

Family-group name: OXYONYXINA Hoffmann, 1957. Stem: Oxyonych-.

Oxypeltus Blanchard, 1851: 459. Current status: valid genus in OXYPELTIDAE.

Type species: Oxypeltus quadrispinosus Blanchard, 1851, by monotypy.

Family-group name: OXYPELTIDES Lacordaire, 1868. Stem: Oxypelt-.

Oxypoda Mannerheim, 1830: 69. Current status: valid genus in STAPHYLINIDAE: ALEOCHARINAE: OXYPODINI: OXYPODINA. Note: name placed on the Official List of Generic Names in Zoology (ICZN 1957b, as Oxypoda Mannerheim, 1831).

Type species: Oxypoda spectabilis Märkel, 1844, by plenary power (see ICZN 1957b). Note: name placed on the Official List of Specific Names in Zoology (ICZN 1957b).

Family-group name: OXYPODIDES C.G. Thomson, 1859. Stem: Oxypod-.

Oxypodinus Bernhauer, 1902: 174. Current status: valid genus in STAPHYLINIDAE: ALEOCHARINAE: OXYPODININI.

Type species: Oxypodinus anxius Bernhauer, 1902, by subsequent designation (Fenyes 1919: 24).

Family-group name: OXYPODININI Fenyes, 1919. Stem: Oxypodin-.

Oxyporus Fabricius, 1775: 267. Current status: valid genus in STAPHYLINIDAE: OXYPORINAE.

Type species: Staphylinus rufus Linnaeus, 1758, by subsequent designation (Latreille 1810: 427).

Family-group name: OXYPORIDAE Fleming, 1821. Stem: Oxypor-.

Oxyrhynchus Schönherr, 1823: 1137. Current status: invalid senior synonym of Cryptoderma Ritsema, 1885 in CURCULIONIDAE: DRYOPHTHORINAE: CRYPTODERMATINI. Note: junior homonym of Oxyrhynchus Leach, 1818 [Pisces].

Type species: Calandra discors Fabricius, 1801, by original designation.

Family-group name: OXYRHYNCHIDES Schönherr, 1823. Stem: Oxyrhynch-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

Oxystoma Duméril, 1805: 226. Current status: valid genus in BRENTIDAE: APIONINAE: APIONITAE: APIONINI: TOXORHYNCHINA.

Type species: Attelabus pomonae Fabricius, 1798, by subsequent designation (Schilsky 1901: 38G).

Family-group name: OXYSTOMATINI Alonso-Zarazaga, 1990. Stem: Oxystomat-.

Oxystomus Dejean, 1825: 410. Current status: invalid senior synonym of Forcipator Maindron, 1904 in CARABIDAE: SCARITINAE: FORCIPATORINI. Note: junior homonym of Oxystomus Fischer, 1803 [Mammalia].

Type species: Oxystomus cylindricus Dejean, 1825, by monotypy.

Family-group name: OXYSTOMINA Csiki, 1927. Stem: Oxystom-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

Oxytelus Gravenhorst, 1802: 101. Current status: valid genus in STAPHYLINIDAE: OXYTELINAE: OXYTELINI.

Type species: Staphylinus piceus Linnaeus, 1767, by subsequent designation (Latreille 1810: 427).

Family-group name: OXYTELIDAE Fleming, 1821. Stem: Oxytel-.

Ozaena G.-A. Olivier, 1812: 619. Current status: valid genus in CARABIDAE: PAUSSINAE: OZAENINI.

Type species: Ozaena dentipes G.-A. Olivier, 1812, by monotypy.

Family-group name: OZAENIDAE Hope, 1838. Stem: Ozaen-.

Ozodecerus Chevrolat, 1839: 175. Current status: valid genus in BRENTIDAE: BRENTINAE: BRENTINI: ITHYSTENINA.

Type species: Ozodecerus forficulatus Chevrolat, 1839, by subsequent designation (Kleine 1939: 130).

Family-group name: OZODOCERINI Jakobson, 1911. Stem: Ozodecer-.

Ozotomerus Perroud, 1853: 406. Current status: valid genus in ANTHRIBIDAE: ANTHRIBINAE: OZOTOMERINI.

Type species: Ozotomerus maculosus Perroud, 1853, by monotypy.

Family-group name: OZOTOMERINI Morimoto, 1972. Stem: Ozotomer-.

Pachnaeus Schönherr, 1826: 121. Current status: valid genus in CURCULIONIDAE: ENTIMINAE: POLYDRUSITAE: TANYMECINI: TANYMECINA. Note: unnecessary new replacement name for Docorhinus Schönherr, 1823; Reily and N.M. Franz (2019) submitted an application to the Commission to suppress the older name Docorhinus Schönherr, 1823 and to place Pachnaeus Schönherr, 1826 on the Official List of Generic Names in Zoology.

Type species: Curculio opalus G.-A. Olivier, 1807, by original designation (type fixation for Docorhinus Schönherr).

Family-group name: PACHNAEI J.L. LeConte, 1874. Stem: Pachnae-.

Pachnoda Burmeister, 1842: 511. Current status: valid genus in SCARABAEIDAE: CETONIINAE: CETONIINI: CETONIINA.

Type species: Scarabaeus marginatus Drury, 1773, by subsequent designation (Rigout 1989: 15).

Family-group name: PACHNODII Péringuey, 1907. Stem: Pachnod-. Note: senior homonym of PACHNODIDAE Steenberg, 1925 (type genus Pachnodus Martens, 1860) in Mollusca.

Pacholenus Schönherr, 1826: 216. Current status: valid genus in CURCULIONIDAE: MOLYTINAE: PACHOLENINI.

Type species: Pacholenus pelliceus Boheman, 1835, by subsequent designation (Vanin and Reichardt 1976: 164). Note: no available species was originally included; the first ones are the two species listed by Schönherr (1835: 101, 102).

Family-group name: PACHOLÉNIDES Lacordaire, 1863. Stem: Pacholen-.

Pachybrachis Chevrolat in Dejean, 1836a: 420. Current status: valid genus in CHRYSOMELIDAE: CRYPTOCEPHALINAE: PACHYBRACHINI.

Type species: Cryptocephalus hieroglyphicus Laicharting, 1781, by subsequent designation (Jacoby 1908: 265).

Family-group name: PACHYBRACHITES Chapuis, 1874. Stem: Pachybrach-.

Pachycarus Solier, 1835a: 660, 666. Current status: valid genus in CARABIDAE: HARPALINAE: HARPALINI: DITOMINA.

Type species: Pachycarus latreillei Solier, 1835, by subsequent designation (E. Desmarest 1851: 106).

Family-group name: PACHYCARINA Stichel, 1923. Stem: Pachycar-.

Pachycera Eschscholtz, 1831: 5, 7. Current status: junior synonym of Hyperops Eschscholtz, 1831 in TENEBRIONIDAE: PIMELIINAE: TENTYRIINI. Note: junior homonym of Pachycera Billberg, 1820 [Hemiptera].

Type species: fixed by Bouchard et al. (2021: 282) according to Article 70.3.2 (ICZN 1999a) as Tenebrio buprestoides Fabricius, 1781, the taxonomic species involved in the misidentifation of Akis laevigata Fabricius, 1801 (sensu Eschscholtz, 1831), as designated originally by monotypy by Eschscholtz (1831).

Family-group name: PACHYCERINA Skopin, 1979. Stem: Pachycer-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

Pachycnema Lepeletier & Audinet-Serville, 1828: 375. Current status: valid genus in SCARABAEIDAE: MELOLONTHINAE: HOPLIINI: LEPITRICHINA. Note: the older name Pachypus Billberg, 1820, a senior objective synonym of this genus-group name, is a nomen oblitum (see Bouchard et al. 2011: 858, Cupello and Ribeiro-Costa 2019: 140).

Type species: Melolontha crassipes Fabricius, 1775, by subsequent designation (Blanchard in Audouin et al. 1843: pl. 44).

Family-group name: PACHYCNÉMIDES Laporte, 1840. Stem: Pachycnem-.

Pachydema Laporte, 1832b: pl. 37. Current status: valid genus in SCARABAEIDAE: MELOLONTHINAE: TANYPROCTINI: TANYPROCTINA.

Type species: Pachydema nigricans Laporte, 1832 (= Melolontha hirticollis Fabricius, 1787), by monotypy.

Family-group name: PACHYDEMIDAE Burmeister, 1855. Stem: Pachydem-. Note: nomen protectum (see A.B.T. Smith 2006: 168).

Pachyderes Guérin-Méneville, 1829a: pl. 12. Current status: valid genus in ELATERIDAE: AGRYPNINAE: OOPHORINI.

Type species: Pachyderes ruficollis Guérin-Méneville, 1829, by monotypy.

Family-group name: PACHYDERINAE Fleutiaux, 1919. Stem: Pachyder-.

Pachydrus Sharp, 1882b: 336, 338. Current status: valid genus in DYTISCIDAE: HYDROPORINAE: PACHYDRINI.

Type species: Pachydrus obesus Sharp, 1882, by subsequent designation (Guignot 1946: 113).

Family-group name: PACHYDRINI F.N. Young, 1980. Stem: Pachydr-.

Pachygastrodes Leleup, 1969: 284. Current status: valid genus in STAPHYLINIDAE: PSELAPHINAE: PSELAPHITAE: PACHYGASTRODINI.

Type species: Pachygastrodes minimipennis Leleup, 1969, by monotypy.

Family-group name: PACHYGASTRODINI Leleup, 1969. Stem: Pachygastrod-.

Pachyglossa Fauvel, 1868: 379. Current status: invalid senior synonym of Pagla Blackwelder, 1952 in STAPHYLINIDAE: ALEOCHARINAE: PAGLINI. Note: new replacement name for Euryglossa Fauvel, 1866; junior homonym of Pachyglossa Hodgson, 1843 [Aves].

Type species: Hoplandria anthracina Fairmaire & Germain, 1861, by subsequent designation (Fenyes 1919: 24) (type fixation for Euryglossa Fauvel).

Family-group name: PACHYGLOSSINI Fenyes, 1919. Stem: Pachygloss-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

Pachymerus Thunberg, 1805: 282. Current status: valid genus in CHRYSOMELIDAE: BRUCHINAE: PACHYMERINI: PACHYMERINA.

Type species: Dermestes bactris Linnaeus, 1763, by monotypy.

Family-group name: PACHYMERINAE Bridwell, 1929. Stem: Pachymer-.

Pachyochthes Reitter, 1897: 248. Current status: junior synonym of Hypodacne J.L. LeConte, 1876 in EUXESTIDAE.

Type species: Pachyochthes edithae Reitter, 1897, by monotypy.

Family-group name: PACHYOCHTHESINAE Reitter, 1911. Stem: Pachyochth-.

Pachypeza Audinet-Serville, 1835a: 42. Current status: valid genus in CERAMBYCIDAE: LAMIINAE: AGAPANTHIINI.

Type species: Saperda pennicornis Germar, 1823, by monotypy.

Family-group name: PACHYPÉZIDES Lacordaire, 1872. Stem: Pachypez-.

Pachyplacus John, 1935: 26. Current status: valid genus in DISCOLOMATIDAE: NOTIOPHYGINAE: PACHYPLACINI.

Type species: Pachyplacus postpressus John, 1935, by monotypy.

Family-group name: PACHYPLACINI John, 1954. Stem: Pachyplac-.

Pachyplectrus J.L. LeConte, 1874b: 53. Current status: valid genus in HYBOSORIDAE: PACHYPLECTRINAE.

Type species: Pachyplectrus laevis J.L. LeConte, 1874, by monotypy.

Family-group name: PACHYPLECTRINAE Ocampo, 2006. Stem: Pachyplectr-.

Pachypterus P.H. Lucas, 1847a: pl. 29. Current status: invalid senior synonym of Neopachypterus Bouchard, Löbl & Merkl, 2007 in TENEBRIONIDAE: BLAPTINAE: OPATRINI: NEOPACHYPTERINA. Note: junior homonym of Pachypterus Swainson, 1839 [Pisces].

Type species: Pachypterus mauritanicus P.H. Lucas, 1847, by monotypy.

Family-group name: PACHYPTÉRATES Mulsant & Rey, 1859. Stem: Pachypter-. Note: family- group name permanently invalid (ICZN 1999a, Article 39).

Pachypus Dejean, 1821: 57. Current status: valid genus in SCARABAEIDAE: MELOLONTHINAE: PACHYPODINI. Note: nomen protectum (see Bouchard et al. 2011: 253).

Type species: Scarabaeus excavatus Fabricius, 1792, by monotypy. Note: the type species has recently been treated as valid (i.e., not as a junior synonym of Scarabaeus candidae Petagna, 1787) by Eberle et al. (2019: 455).

Family-group name: PACHYPODEN Erichson, 1840. Stem: Pachypod-.

Pachyrhinadoretus Ohaus, 1912a: 509. Current status: valid genus in SCARABAEIDAE: RUTELINAE: ADORETINI: PACHYRHINADORETINA.

Type species: Pachyrhinadoretus rugipennis Ohaus, 1912, by subsequent designation (Arrow 1917: 285).

Family-group name: PACHYRHINADORETINA Ohaus, 1912. Stem: Pachyrhinadoret-.

Pachyrhynchus Germar, 1823: 336. Current status: valid genus in CURCULIONIDAE: ENTIMINAE: POLYDRUSITAE: PACHYRHYNCHINI. Note: the senior homonym Pachyrhynchus Wagler, 1822 [Aves] was suppressed for purposes of both the Principle of Priority and Homonymy and Pachyrhynchus Germar, 1823 placed on the Official List of Generic Names in Zoology (ICZN 1970b, as Pachyrhynchus Germar, 1824).

Type species: Pachyrhynchus moniliferus Germar, 1823, by monotypy (see ICZN 1970b). Note: name placed on the Official List of Specific Names in Zoology (ICZN 1970b, as Pachyrhynchus moniliferus Germar, 1824).

Family-group name: PACHYRHYNCHIDES Schönherr, 1826. Stem: Pachyrhynch-.

Pachyrhynchus W. Kirby, 1837: 203. Current status: junior synonym of Ithycerus Schönherr, 1823 in BRENTIDAE: ITHYCERINAE: ITHYCERINI. Note: junior homonym of Pachyrhynchus Germar, 1823 [Coleoptera: CURCULIONIDAE].

Type species: Pachyrhynchus schoenherri W. Kirby, 1837 (= Curculio noveboracensis Forster, 1771), by monotypy.

Family-group name: PACHYRHINCHIDAE W. Kirby, 1837. Stem: Pachyrhynch-. Note: family- group name permanently invalid (ICZN 1999a, Article 39).

Pachyschelus Solier, 1833: 313. Current status: valid genus in BUPRESTIDAE: AGRILINAE: TRACHEINI: PACHYSCHELINA.

Type species: Pachyschelus scutellatus Solier, 1833 (= Trachys terminans Fabricius, 1801), by monotypy.

Family-group name: PACHYSCHELINAE Böving & Craighead, 1931. Stem: Pachyschel-.

Pachysoma W.S. MacLeay, 1821: 507. Current status: valid genus in SCARABAEIDAE: SCARABAEINAE: SCARABAEINI.

Type species: Copris aesculapius G.-A. Olivier, 1790, by subsequent designation (Reiche 1841: 212).

Family-group name: PACHYSOMIDES M.C. Ferreira, 1953. Stem: Pachysomat-.

Pachystola Dejean, 1835: 342. Current status: junior synonym of Lamia Fabricius, 1775 in CERAMBYCIDAE: LAMIINAE: LAMIINI.

Type species: Cerambyx textor Linnaeus, 1758, by monotypy.

Family-group name: PACHYSTOLAEIDAE Gistel, 1848. Stem: Pachystol-.

Pachyta Dejean, 1821: 112. Current status: valid genus in CERAMBYCIDAE: LEPTURINAE: RHAGIINI.

Type species: Leptura octomaculata Fabricius, 1792 (= Leptura quadrimaculata Linnaeus, 1758), by subsequent designation (Westwood 1838c: 41).

Family-group name: PACHYTINI Portevin, 1934. Stem: Pachyt-.

Pachyteles Perty, 1830: 3. Current status: valid genus in CARABIDAE: PAUSSINAE: OZAENINI.

Type species: Pachyteles striola Perty, 1830, by subsequent designation (Hope 1838a: 99).

Family-group name: PACHYTELINI Jeannel, 1946. Stem: Pachytel-.

Pachytrachelus Chaudoir, 1852: 85. Current status: valid genus in CARABIDAE: HARPALINAE: HARPALINI: STENOLOPHINA.

Type species: Pachytrachelus cribriceps Chaudoir, 1852, by monotypy.

Family-group name: PACHYTRACHELI Csiki, 1932. Stem: Pachytrachel-.

Pachytricha Hope, 1841b: 303. Current status: valid genus in SCARABAEIDAE: MELOLONTHINAE: PACHYTRICHINI.

Type species: Pachytricha castanea Hope, 1841, by monotypy.

Family-group name: PACHYTRICHIADAE Burmeister, 1855. Stem: Pachytrich-.

Pachyura Hope, 1834a: 102. Current status: valid genus in BELIDAE: BELINAE: PACHYURITAE: PACHYURINI.

Type species: Pachyura australis Hope, 1834, by monotypy.

Family-group name: PACHYURINI Kuschel, 1959. Stem: Pachyur-.

Pactola Pascoe, 1876: 57. Current status: valid genus in CURCULIONIDAE: CURCULIONINAE: EUGNOMINI: EUGNOMINA.

Type species: Pactola variabilis Pascoe, 1876, by monotypy.

Family-group name: PACTOLINA Voss, 1936. Stem: Pactol-. Note: junior homonym of PACTOLIENS² H. Milne-Edwards, 1837 (type genus Pactolus Leach, 1815) in Crustacea; H. Milne-Edwards’ family-group name is permanently invalid (ICZN 1999a, Article 39) since Pactolus Leach was suppressed under the plenary power for the purposes of the Principle of Priority (ICZN 1966a).

Paederus Fabricius, 1775: 268. Current status: valid genus in STAPHYLINIDAE: PAEDERINAE: PAEDERINI: PAEDERINA.

Type species: Staphylinus riparius Linnaeus, 1758, by subsequent designation (Latreille 1810: 427).

Family-group name: POEDERIDAE Fleming, 1821. Stem: Paeder-.

Paelobius Schönherr, 1808: 27. Current status: junior synonym of Hygrobia Latreille, 1804 in HYGROBIIDAE.

Type species: Dytiscus tardus Herbst, 1779 (= Dytiscus hermanni Fabricius, 1775), by monotypy.

Family-group name: PELOBIINAE Erichson, 1837. Stem: Paelobi-.

Pagla Blackwelder, 1952: 287. Current status: valid genus in STAPHYLINIDAE: ALEOCHARINAE: PAGLINI. Note: new replacement name for Pachyglossa Fauvel, 1868.

Type species: Hoplandria anthracina Fairmaire & Germain, 1861, by subsequent designation (Fenyes 1919: 22) (type fixation for Euryglossa Fauvel for which Pachyglossa Fauvel was a new replacement name).

Family-group name: PAGLINI Newton & Thayer, 1992. Stem: Pagl-.

Pagria Lefèvre, 1884: 92. Current status: valid genus in CHRYSOMELIDAE: EUMOLPINAE: TYPOPHORINI.

Type species: Pagria suturalis Lefèvre, 1884, by subsequent designation (Jacoby 1908: 356).

Family-group name: PAGRIITES Lefèvre, 1884. Stem: Pagri-.

Pagurodactylus Gorham, 1900: 78. Current status: valid genus in MELYRIDAE: MALACHIINAE: PAGURODACTYLINI.

Type species: Pagurodactylus vitticeps Gorham, 1900, by monotypy.

Family-group name: PAGURODACTYLINAE Constantin, 2001. Stem: Pagurodactyl-.

Paipalesomus Schönherr, 1847: 69. Current status: junior synonym of Peribleptus Schönherr, 1843 in CURCULIONIDAE: MOLYTINAE: PAIPALESOMINI.

Type species: Paipalesomus pistriarius Schönherr, 1847 (= Alcides dealbatus Boisduval, 1835), by original designation.

Family-group name: PAIPALESOMINI G.A.K. Marshall, 1932. Stem: Paipalesom-.

Palaechthus C.O. Waterhouse, 1884: 277. Current status: valid genus in CURCULIONIDAE: CYCLOMINAE: LISTRODERINI: PALAECHTHINA.

Type species: Palaechthus glabratus C.O. Waterhouse, 1884, by subsequent designation (Brinck 1948: 51).

Family-group name: PALAECHTINI Brinck, 1948. Stem: Palaechth-.

†Palaeoanoplus Legalov, 2021: 66. Current status: valid genus in CURCULIONIDAE: CURCULIONINAE: †PALAEOANOPLINI. Note: upper Eocene (Baltic amber); combined description of a new family-group taxon and a single new nominal genus (ICZN 1999a, Article 13.5).

Type species: †Palaeoanoplus horridus Legalov, 2021, by original designation.

Family-group name: †PALAEOANOPLINI Legalov, 2021: 65. Stem: Palaeoanopl-.

†Palaeoaxinidium McKay, 1991: 3. Current status: valid genus in CARABIDAE: SCARITINAE: †PALAEOAXINIDIINI. Note: Cretaceous (Botswana).

Type species: †Palaeoaxinidium orapensis MacKay, 1991, by original designation.

Family-group name: †PALAEOAXINIDINI McKay, 1991. Stem: Palaeoaxinidi-.

†Palaeocryptorhynchus Poinar, 2009b: 587. Current status: valid genus in CURCULIONIDAE: MOLYTINAE: †PALAEOCRYPTORHYNCHINI. Note: mid-Cretaceous (Myanmar); Poinar (2009b) used two different spellings, including Palaeocryptorhynus (p. 587); as far as we know no one has acted as First Reviser (ICZN 1999a, Article 24.2.3) and we here select Palaeocryptorhynchus, the spelling in prevailing usage.

Type species: †Palaeocryptorhynchus burmanus Poinar, 2009, by original designation.

Family-group name: †PALEOCRYPTORHYNCHINI Poinar & Legalov in Legalov and Poinar, 2014: 562. Stem: Palaeocryptorhynch-.

†Palaeogyrinus Schlechtendal, 1894: 201. Current status: valid genus in DYTISCIDAE: †PALAEOGYRININAE. Note: upper Oligocene (Germany).

Type species: †Palaeogyrinus strigatus Schlechtendal, 1894, by monotypy.

Family-group name: †PALAEOGYRINIDAE Schlechtendal, 1894. Stem: Palaeogyrin-.

†Palaeomallerus Legalov, 2018b: 75. Current status: valid genus in CURCULIONIDAE: CONODERINAE: †PALAEOMALLERINI. Note: lower Eocene (USA).

Type species: †Palaeomallerus longirostris Legalov, 2018, by original designation.

Family-group name: †PALAEOMALLERINI Legalov, 2018b: 75. Stem: Palaeomaller-.

†Palaeorhamphus Legalov, 2017b: 982. Current status: valid genus in CURCULIONIDAE: CURCULIONINAE: RHAMPHINI: †PALAEORHAMPHINA. Note: upper Eocene (Baltic amber).

Type species: †Palaeorhamphus primitivus Legalov, 2017, by original designation.

Family-group name: †PALAEORHAMPHINA Legalov, 2017b: 981. Stem: Palaeorhamph-.

†Palaeorhopalotria Legalov, 2013b: 64. Current status: valid genus in BELIDAE: OXYCORYNINAE: OXYCORYNITAE: †PALAEORHOPALOTRIINI. Note: upper Eocene (France).

Type species: †Palaeorhopalotria neli Legalov, 2013, by original designation.

Family-group name: †PALAEORHOPALOTRIINI Legalov, 2013b: 63. Stem: Palaeorhopalotri-.

†Palaeotanaos Kirejtshuk, Legalov & Nel, 2016: 1437. Current status: valid genus in BRENTIDAE: APIONINAE: TANAITAE: †PALAEOTANAINI. Note: lower Eocene (France).

Type species: †Palaeotanaos oisensis Kirejtshuk, Legalov & Nel, 2016, by original designation.

Family-group name: †PALAEOTANAINI Legalov, Kirejtshuk & Nel, 2020b: 734. Stem: Palaeotana-.

†Palaeothroscus Iablokoff-Khnzorian, 1962: 83. Current status: junior synonym of Trixagus Kugelann, 1794 in THROSCIDAE. Note: upper Eocene (Baltic amber).

Type species: †Palaeothroscus sosnovskyi Iablokoff-Khnzorian, 1962, by original designation.

Family-group name: †PALAEOTHROSCINI Iablokoff-Khnzorian, 1962. Stem: Palaeothrosc-.

†Palaeotylus Poinar, Vega & Legalov, 2018: 138. Current status: valid genus in POLYPHAGA incertae sedis. Note: mid-Cretaceous (Myanmar).

Type species: †Palaeotylus femoralis Poinar, Vega & Legalov, 2018, by original designation.

Family-group name: †PALAEOTYLINAE Poinar, Vega & Legalov, 2018: 2. Stem: Palaeotyl-.

Palaeoxenus G.H. Horn, 1891: 40. Current status: valid genus in EUCNEMIDAE: PALAEOXENINAE.

Type species: Cryptostoma dohrni G.H. Horn, 1878, by monotypy.

Family-group name: PALAEOXENINAE Muona, 1993. Stem: Palaeoxen-.

Palaestra Laporte, 1840b: 251. Current status: valid genus in MELOIDAE: NEMOGNATHINAE: PALAESTRINI.

Type species: Palaestra rubripennis Laporte, 1840, by monotypy.

Family-group name: PALAESTRINI Bologna, Turco & Pinto, 2013: 394. Stem: Palaestr-.

†Paleocartus Legalov, 2003: 78. Current status: valid genus in NEMONYCHIDAE: †PALEOCARTINAE: †PALEOCARTINI: †PALEOCARTINA. Note: Upper Jurassic (Kazakhstan).

Type species: †Brenthorrhinoides pubescens Gratshev & Zherikhin, 1996, by original designation.

Family-group name: †PALEOCARTINI Legalov, 2003. Stem: Paleocart-.

†Pallichnus Retallack, 1984: 580. Current status: valid genus in †PALLICHNIDAE. Note: Ichnotaxon, Upper Cretaceous/Oligocene (Argentina; South Dakota, USA).

Type species: †Pallichnus dakotensis Retallack, 1984, by original designation.

Family-group name: †PALLICHNIDAE Genise, 2004. Stem: Pallichn-.

Pallodes Erichson, 1843a: 348. Current status: valid genus in NITIDULIDAE: NITIDULINAE: NITIDULINI.

Type species: Strongylus annulifer Laporte, 1840, by subsequent designation (E. Desmarest 1851: 283).

Family-group name: PALLODINI Reitter, 1873: 98. Stem: Pallod-. Note: this name was overlooked in Bouchard et al. (2011), Bouchard and Bousquet (2020a).

Palophagus Kuschel in Kuschel and May, 1990: 704. Current status: valid genus in MEGALOPODIDAE: PALOPHAGINAE: PALOPHAGINI.

Type species: Palophagus bunyae Kuschel, 1990, by original designation.

Family-group name: PALOPHAGINAE Kuschel & May, 1990. Stem: Palophag-.

Palorus Mulsant, 1854: 250. Current status: valid genus in TENEBRIONIDAE: TENEBRIONINAE: PALORINI.

Type species: Hypophlaeus depressus Fabricius, 1790, by monotypy.

Family-group name: PALORINAE E.G. Matthews, 2003. Stem: Palor-.

†Palpattalus Tshernyshev, 2017: 955. Current status: valid genus in MELYRIDAE: MALACHIINAE: †PALPATTALINI. Note: Eocene (Kaliningrad region, Russia).

Type species: †Palpattalus eocenicus Tshernyshev, 2017, by original designation.

Family-group name: †PALPATTALINI Tshernyshev, 2020: 69. Stem: Palpattal-.

Palpomodes Koch, 1952: 223. Current status: valid genus in TENEBRIONIDAE: PIMELIINAE: SEPIDIINI: PALPOMODINA.

Type species: Psammodes physopterus Gebien, 1920, by monotypy.

Family-group name: PALPOMODINA Kamiński & Gearner in Kamiński et al., 2022: 645. Stem: Palpomod-.

Pamborus Latreille in G.-A. Olivier, 1812: 677. Current status: valid genus in CARABIDAE: CARABINAE: PAMBORINI.

Type species: Pamborus alternans Latreille, 1812, by monotypy.

Family-group name: PAMBORIDAE Hope, 1838. Stem: Pambor-.

Panagaeus Latreille, 1802b: 91. Current status: valid genus in CARABIDAE: HARPALINAE: PANAGAEINI: PANAGAEINA.

Type species: Carabus cruxmajor Linnaeus, 1758, by monotypy.

Family-group name: PANAGAEIDES Bonelli, 1810. Stem: Panagae-.

Panaphantus Kiesenwetter in Kraatz, 1858a: 48. Current status: valid genus in STAPHYLINIDAE: PSELAPHINAE: EUPLECTITAE: TRICHONYCHINI: PANAPHANTINA.

Type species: Panaphantus atomus Kiesenwetter, 1858, by monotypy.

Family-group name: PANAPHANTINA Jeannel, 1950. Stem: Panaphant-.

Pandeleteius Schönherr, 1834a: 129. Current status: valid genus in CURCULIONIDAE: ENTIMINAE: POLYDRUSITAE: TANYMECINI: PANDELETEIINA.

Type species: Pandeleteius pauperculus Gyllenhal, 1834 (= Curculio hilaris Herbst, 1797), by original designation.

Family-group name: PANDELETEINI Pierce, 1913. Stem: Pandeletei-.

Panelus Lewis, 1895b: 375. Current status: valid genus in SCARABAEIDAE: SCARABAEINAE: DELTOCHILINI.

Type species: Temnoplectron parvulum C.O. Waterhouse, 1874, by original designation.

Family-group name: PANELINI Arrow, 1931. Stem: Panel-.

Pangaura Gistel, 1856: 366. Current status: junior synonym of Lacon Laporte, 1838 in ELATERIDAE: AGRYPNINAE: AGRYPNINI.

Type species: Elater varius G.-A. Olivier, 1790 (= Elater querceus Herbst, 1784), by subsequent designation (Sánchez-Ruiz 1996: 44).

Family-group name: PANGAURADAE Gistel, 1856. Stem: Pangaur-. Note: nomen oblitum (see Bouchard et al. 2011: 307).

Pangetes Gistel, 1856: 358. Current status: junior synonym of Amara Bonelli, 1810 in CARABIDAE: HARPALINAE: ZABRINI: AMARINA.

Type species: Carabus ovatus Fabricius, 1792, by subsequent designation (Bousquet 2002: 37).

Family-group name: PANGETEIDAE Gistel, 1856. Stem: Panget-.

Paniscus Gistel, 1848a: [5]. Current status: junior synonym of Trichius Fabricius, 1775 in SCARABAEIDAE: CETONIINAE: TRICHIINI. Note: junior homonym of Paniscus Schrank, 1802 [Hymenoptera].

Type species: Scarabaeus fasciatus Linnaeus, 1758, by subsequent designation (Bouchard et al. 2011: 271).

Family-group name: PANISCIDAE Gistel, 1848. Stem: Panisc-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

Pantolia Burmeister, 1842: 567. Current status: valid genus in SCARABAEIDAE: CETONIINAE: STENOTARSIINI: PANTOLIINA.

Type species: Cetonia flavomarginata Gory & Percheron, 1833, by subsequent designation (Sakai and Nagai 1998: 178).

Family-group name: PANTOLIINA Krikken, 1984. Stem: Pantoli-.

Pantomorus Schönherr, 1840a: 942. Current status: valid genus in CURCULIONIDAE: ENTIMINAE: POLYDRUSITAE: NAUPACTINI: NAUPACTINA.

Type species: Pantomorus albosignatus Schönherr, 1840, by original designation.

Family-group name: PANTOMORINA Voss, 1954. Stem: Pantomor-.

Pantopoeus Schönherr, 1842b: 252. Current status: junior synonym of Perperus Schönherr, 1842 in CURCULIONIDAE: ENTIMINAE: ENTIMITAE: BYRSOPAGINI: STRANGALIODINA. Note: First Reviser (ICZN 1999a, Article 24.2.2) for Perperus Schönherr, 1842 versus Pantopoeus Schönherr, 1842 is Lea (1911: 76).

Type species: Pantopoeus cervinus Boheman, 1842, by original designation.

Family-group name: PANTOPÉIDES Lacordaire, 1863. Stem: Pantopoe-.

†Pantostictus Poinar & A.E. Brown, 2009b: 39. Current status: valid genus in HISTERIDAE: ABRAEINAE: †PANTOSTICTINI. Note: mid-Cretaceous (Myanmar).

Type species: †Pantostictus burmanicus Poinar & A.E. Brown, 2009, by original designation.

Family-group name: †PANTOSTICTINI Zhou, Caterino & Ślipiński, 2020: 530. Stem: Pantostict-.

Pantoteles Schönherr, 1845: 59. Current status: valid genus in CURCULIONIDAE: BARIDINAE: PANTOTELINI.

Type species: Pantoteles tenuirostris Boheman, 1845, by original designation.

Family-group name: PANTOTÉLIDES Lacordaire, 1865. Stem: Pantotel-.

Paoligena Pic, 1928b: 42. Current status: valid genus in TENEBRIONIDAE: TENEBRIONINAE: PAOLIGENINI.

Type species: Praogena inhumeralis Pic, 1928, by monotypy.

Family-group name: PAOLIGENINI Ferrer, 2013: 108. Stem: Paoligen-.

Papusus Casey, 1897: 542. Current status: valid genus in STAPHYLINIDAE: SCYDMAENINAE: MASTIGITAE: PAPUSINI.

Type species: Papusus macer Casey, 1897, by monotypy.

Family-group name: PAPUSINI Jałoszyński & Brunke in Jałoszyński et al., 2018: 635. Stem: Papus-.

Parablax O. Schwarz, 1906: 368. Current status: valid genus in ELATERIDAE: PITYOBIINAE: PARABLACINI.

Type species: Metablax trisulcatus O. Schwarz, 1903 (= Parasaphes quinquesulcatus Blackburn, 1900), by subsequent designation (Hyslop 1921: 661).

Family-group name: PARABLACINAE Kundrata, Gunter, H.B. Douglas & Bocák, 2016: 299. Stem: Parablac-.

Paracatops Portevin, 1907: 69. Current status: valid genus in LEIODIDAE: CHOLEVINAE: ANEMADINI: PARACATOPINA.

Type species: Choleva antipoda Kirsch, 1877, by monotypy.

Family-group name: PARACATOPINI Jeannel, 1936. Stem: Paracatop-.

Parachilia Burmeister, 1842: 556. Current status: valid genus in SCARABAEIDAE: CETONIINAE: STENOTARSIINI: PARACHILIINA.

Type species [suggested]: Parachilia melanocala Burmeister, 1842, by subsequent designation (Kraatz 1881: 72). Note: the valid type species fixation is that of Lacordaire (1855: 514), who selected Cetonia bufo Gory & Percheron, 1835, but that nominal species is the type species of the subgenus Chilamblys Kraatz, 1880, not of the nominotypical subgenus; an application to the Commission is necessary to fix Parachilia melanocala Burmeister, 1842 as the type species of Parachilia Burmeister, 1842.

Family-group name: PARACHILIINA Krikken, 1984. Stem: Parachili-.

Parachorius Harold, 1873: 103. Current status: valid genus in SCARABAEIDAE: SCARABAEINAE: PARACHORIINI.

Type species: Parachorius thomsoni Harold, 1873, by monotypy.

Family-group name: PARACHORIINI Tarasov, 2017: 111. Stem: Parachori-.

Paracucujus Sen Gupta & Crowson, 1966: 67. Current status: valid genus in BOGANIIDAE: PARACUCUJINAE.

Type species: Paracucujus rostratus Sen Gupta & Crowson, 1966, by original designation.

Family-group name: PARACUCUJINAE Endrödy-Younga & Crowson, 1986. Stem: Paracucuj-.

†Paradermestes Deng, Ślipiński, Ren & H. Pang, 2017b: 110. Current status: valid genus in DERMESTIDAE: DERMESTINAE: †PARADERMESTINI. Note: Middle/Upper Jurassic (China).

Type species: †Paradermestes jurassicus Deng, Ślipiński, Ren & H. Pang, 2017, by original designation.

Family-group name: †PARADERMESTINI Deng, Ślipiński, Ren & H. Pang, 2017b: 110. Stem: Paradermest-.

Paradoxenusa Bruch, 1937: 354. Current status: valid genus in STAPHYLINIDAE: ALEOCHARINAE: PARADOXENUSINI.

Type species: Paradoxenusa silvestrii Bruch, 1937, by original designation.

Family-group name: PARADOXENUSINI Bruch, 1937. Stem: Paradoxenus-.

Paradrilus Kiesenwetter, 1866: 369. Current status: valid genus in ELATERIDAE: OMALISINAE.

Type species: Paradrilus opacus Kiesenwetter, 1866, by monotypy. Note: combined description of a new genus and a single new species (ICZN 1999a, Article 12.2.6).

Family-group name: PARADRILINAE Kundrata, Baena & Bocák, 2015: 417. Stem: Paradril-.

Paraholopterus Cerda & Cekalović, 1987: 189. Current status: valid genus in CERAMBYCIDAE: CERAMBYCINAE: PARAHOLOPTERINI.

Type species: Paraholopterus nahuelbutensis Cerda & Cekalović, 1987, by original designation.

Family-group name: PARAHOLOPTERINI Martins, 1997. Stem: Paraholopter-.

†Parahygrobia Ponomarenko, 1977: 20. Current status: valid genus in †PARAHYGROBIIDAE. Note: Upper Jurassic (Russia).

Type species: †Parahygrobia natans Ponomarenko, 1977, by original designation.

Family-group name: †PARAHYGROBIIDAE Ponomarenko, 1977. Stem: Parahygrobi-.

Paraleptodema Obenberger, 1936: 106. Current status: junior synonym of Cinyra Laporte & Gory, 1837 in BUPRESTIDAE: CHRYSOCHROINAE: PARALEPTODEMINI: PARALEPTODEMINA.

Type species: Paraleptodema strandi Obenberger, 1936, by monotypy.

Family-group name: PARALEPTODEMINI Cobos, 1975. Stem: Paraleptodem-.

Paralispinus Bernhauer, 1921: 67. Current status: invalid senior synonym of Clavilispinus Bernhauer, 1926 in STAPHYLINIDAE: OSORIINAE: THORACOPHORINI: CLAVILISPININA. Note: replacement name for Ancaeus Fauvel, 1865; junior homonym of Paralispinus Eichelbaum, 1913 [Coleoptera: STAPHYLINIDAE].

Type species: Ancaeus megacephalus Fauvel, 1865, by monotypy (type fixation for Ancaeus Fauvel).

Family-group name: PARALISPINI Blackwelder, 1942. Stem: Paralispin-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

Parallelodemas Faust, 1894a: 306. Current status: valid genus in CURCULIONIDAE: BARIDINAE: BARIDINI: PARALLELODEMINA.

Type species: Parallelodemas perfectum Faust, 1894, by subsequent designation (Morimoto and Yoshihara 1996: 17).

Family-group name: PARALLELODEMINA Legalov, 2018e: 497. Stem: Parallelodem-.

†Paralucanus Nikolajev, 2000b: S329. Current status: valid genus in LUCANIDAE: †PARALUCANINAE. Note: Upper Jurassic (Mongolia).

Type species: †Paralucanus mesozoicus Nikolajev, 2000, by original designation.

Family-group name: †PARALUCANINAE Nikolajev, 2000. Stem: Paralucan-.

Paralycus L.N. Medvedev & Kazantsev, 1992: 55. Current status: junior synonym of Lyropaeus C.O. Waterhouse, 1878 in LYCIDAE: LYROPAEINAE: LYROPAEINI. Note: junior homonym of Paralycus Womersley, 1944 [Acarina].

Type species: Paralycus helvius L.N. Medvedev & Kazantsev, 1992, by monotypy.

Family-group name: PARALYCINAE L.N. Medvedev & Kazantsev, 1992. Stem: Paralyc-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

Paramecolabus Jekel, 1860: 190. Current status: valid genus in ATTELABIDAE: ATTELABINAE: ATTELABINI: PARAMECOLABINA.

Type species: Attelabus discolor Fåhraeus, 1839, by original designation.

Family-group name: PARAMECOLABINA Legalov, 2003. Stem: Paramecolab-.

Paramecosoma Curtis, 1833b: 186. Current status: valid genus in CRYPTOPHAGIDAE: CRYPTOPHAGINAE: CRYPTOPHAGINI.

Type species: Paramecosoma bicolor Curtis, 1833 (= Latridius melanocephalus Herbst, 1793), by monotypy.

Family-group name: PARAMECOSOMINI Reitter, 1874. Stem: Paramecosomat-.

Paramellon C.O. Waterhouse, 1882: iv. Current status: valid genus in TENEBRIONIDAE: PIMELIINAE: COSSYPHODINI: PARAMELLONINA.

Type species: Paramellon sociale C.O. Waterhouse, 1882, by monotypy.

Family-group name: PARAMELLONINAE Andreae, 1961. Stem: Paramellon-.

Parandra Latreille, 1802b: 160. Current status: valid genus in CERAMBYCIDAE: PRIONINAE: PARANDRINI: PARANDRINA.

Type species: Attelabus glaber DeGeer, 1774, by monotypy.

Family-group name: PARANDRIDES Blanchard, 1845. Stem: Parandr-.

†Parandrexis Martynov, 1926: 19. Current status: valid genus in †PARANDREXIDAE: †PARANDREXINAE. Note: Middle-Upper Jurassic (Kazakhstan; China).

Type species: †Parandrexis parvula Martynov, 1926, by original designation.

Family-group name: †PARANDREXIDAE Kirejtshuk, 1994. Stem: Parandrex-.

Paraptochus Seidlitz, 1868: 3, 35. Current status: valid genus in CURCULIONIDAE: ENTIMINAE: CYPHICERITAE: PERITELINI: PERITELINA.

Type species: Paraptochus californicus Seidlitz, 1868 (= Peritelus sellatus Boheman, 1859), by monotypy.

Family-group name: PARAPTOCHI Pierce, 1913. Stem: Paraptoch-.

Parastasia Westwood, 1841d: 204. Current status: valid genus in SCARABAEIDAE: RUTELINAE: RUTELINI: PARASTASIINA.

Type species: Parastasia canaliculata Westwood, 1841, by monotypy.

Family-group name: PARASTASIIDAE Burmeister, 1844. Stem: Parastasi-.

Parasthetops Perkins & J. Balfour-Browne, 1994: 37. Current status: valid genus in HYDRAENIDAE: PROSTHETOPINAE: PARASTHETOPINI.

Type species: Parasthetops nigritus Perkins & J. Balfour-Browne, 1994, by original designation.

Family-group name: PARASTHETOPINI Perkins, 1994. Stem: Parasthetop-.

Parasynaptopsis Legalov, 2003: 377. Current status: valid genus in ATTELABIDAE: ATTELABINAE: EUOPINI.

Type species: Euops chinensis Voss, 1922, by original designation.

Family-group name: PARASYNAPTOPSISINA Legalov, 2007. Stem: Parasynaptopsis- (ICZN 1999a, Article 29.4).

Paratassa Marseul, 1883: 168. Current status: valid genus in BUPRESTIDAE: CHRYSOCHROINAE: PARATASSINI.

Type species: Sphenoptera caroli Marseul, 1883 (= Sphenoptera coraebiformis Fairmaire, 1875), by monotypy.

Family-group name: PARATASSINI Bílý & Volkovitsh, 1996. Stem: Paratass-.

†Parathyrea Alexeev, 1994: 11. Current status: valid genus in BUPRESTIDAE: †PARATHYREINAE. Note: Upper Jurassic (Kazakhstan).

Type species: †Parathyrea jurassica Alexeev, 1994, by original designation.

Family-group name: †PARATHYREINAE Alexeev, 1994. Stem: Parathyre-.

Paratinus Abeille de Perrin, 1891: 220. Current status: junior synonym of Apalochrus Erichson, 1840 in MELYRIDAE: MALACHIINAE: MALACHIINI.

Type species: Apalochrus femoralis Erichson, 1840, by subsequent designation (Evers 1987: 13).

Family-group name: PARATINI Portevin, 1931. Stem: Paratin-.

Paratomapoderus Voss, 1926: 85. Current status: valid genus in ATTELABIDAE: APODERINAE: HOPLAPODERINI: PARATOMAPODERINA.

Type species: Apoderus miniatus Faust, 1882, by original designation.

Family-group name: PARATOMAPODERINA Legalov, 2003. Stem: Paratomapoder-.

Paratrachys Saunders, 1873: 523. Current status: valid genus in BUPRESTIDAE: POLYCESTINAE: PARATRACHEINI.

Type species: Paratrachys hederae Saunders, 1873, by monotypy.

Family-group name: PARATRACHYSAE Cobos, 1980. Stem: Paratrache-.

Parauletanus Legalov, 2007: 31. Current status: valid genus in ATTELABIDAE: RHYNCHITINAE: RHINOCARTITAE: PARAULETANINI.

Type species: Auletanus disparatus Voss, 1922, by original designation.

Family-group name: PARAULETANINA Legalov, 2007. Stem: Parauletan-.

Parecatus Fairmaire, 1900: 245. Current status: junior synonym of Scotinesthes Fairmaire, 1895 in TENEBRIONIDAE: PIMELIINAE: ASIDINI.

Type species: Parecatus plicatulus Fairmaire, 1900, by subsequent designation (Gebien 1937: 741).

Family-group name: PARECATINI Chatanay, 1914. Stem: Parecat-.

Parhydraena Orchymont, 1937: 457. Current status: valid genus in HYDRAENIDAE: HYDRAENINAE: PARHYDRAENINI.

Type species: Hydraena brevipalpis Régimbart, 1906, by original designation.

Family-group name: PARHYDRAENINI Perkins, 1997. Stem: Parhydraen-.

Paristemia Westwood, 1841c: 124. Current status: valid genus in CERAMBYCIDAE: CERAMBYCINAE: TRACHYDERINI: TRACHYDERINA.

Type species: Paristemia platyptera Westwood, 1841, by monotypy.

Family-group name: PARISTÉMIIDES Lacordaire, 1868. Stem: Paristemi-.

Parmena Dejean, 1821: 108. Current status: valid genus in CERAMBYCIDAE: LAMIINAE: PARMENINI.

Type species: Cerambyx fasciatus Villers, 1789 (= Cerambyx balteus Linnaeus, 1767), by subsequent designation (Guérin-Méneville 1826: 187).

Family-group name: PARMÉNAIRES Mulsant, 1839. Stem: Parmen-.

Parmulus Gundlach in Poey, 1854: 323, 336. Current status: junior synonym of Clypastraea Haldeman, 1842 in CORYLOPHIDAE: CORYLOPHINAE: PARMULINI. Note: new replacement name for Clypeaster Dejean, 1821.

Type species: Cossyphus pusillus Gyllenhal, 1810, by monotypy (type fixation for Clypeaster Dejean).

Family-group name: PARMULINI Poey, 1854. Stem: Parmul-.

Parnus Fabricius, 1792a: 245. Current status: junior synonym of Dryops G.-A. Olivier, 1791 in DRYOPIDAE.

Type species: Parnus prolifericornis Fabricius, 1792 (= Dermestes auriculatus Geoffroy, 1785), by subsequent designation (Curtis 1825: pl. 80).

Family-group name: PARNIDEA Leach, 1817. Stem: Parn-.

Paromalus Erichson, 1834: 167. Current status: valid genus in HISTERIDAE: DENDROPHILINAE: PAROMALINI.

Type species: Hister flavicornis Herbst, 1791, by subsequent designation (Westwood 1838c: 22).

Family-group name: PAROMALINI Reitter, 1909. Stem: Paromal-.

Paropsis G.-A. Olivier, 1807: 597. Current status: valid genus in CHRYSOMELIDAE: CHRYSOMELINAE: CHRYSOMELINI.

Type species: Chrysomela australasiae Fabricius, 1801, by subsequent designation (Latreille 1810: 432).

Family-group name: PAROPSINES Motschulsky, 1860. Stem: Paropse-.

Parosorius Bernhauer, 1904: 222. Current status: valid genus in STAPHYLINIDAE: OSORIINAE: OSORIINI: PAROSORIINA.

Type species: Ancaeus foersteri Bernhauer, 1903, by monotypy.

Family-group name: PAROSORII Bernhauer & Schubert, 1911. Stem: Parosori-.

Pasimachus Bonelli, 1813: 476. Current status: valid genus in CARABIDAE: SCARITINAE: PASIMACHINI.

Type species: Scarites depressus Fabricius, 1787, by subsequent designation (Hope 1838a: 94).

Family-group name: PASIMACHIDES Putzeys, 1867. Stem: Pasimach-.

Passalidius Chaudoir, 1863b: 116. Current status: valid genus in CARABIDAE: SCARITINAE: SCARITINI: SCAPTERINA.

Type species: Passalidius afer Chaudoir, 1863 (= Scarites fortipes Boheman, 1860), by subsequent designation (Lorenz 1998: 139).

Family-group name: PASSALIDIINA Csiki, 1927. Stem: Passalidi-.

†Passalopalpus Boucher & Bai in Boucher et al., 2016: 71. Current status: valid genus in †PASSALOPALPIDAE. Note: mid-Cretaceous (Myanmar).

Type species: †Passalopalpus cheni Boucher, Bai & Zhang, 2016, by original designation.

Family-group name: †PASSALOPALPIDAE Boucher & Bai in Boucher et al., 2016: 68. Stem: Passalopalp-.

Passalus Fabricius, 1792b: 240. Current status: valid genus and subgenus in PASSALIDAE: PASSALINAE: PASSALINI.

Type species: Scarabaeus interruptus Linnaeus, 1758, by subsequent designation (Latreille 1810: 429).

Family-group name: PASSALIDA Leach, 1815. Stem: Passal-.

Passandra Dalman in Schönherr, 1817: 146. Current status: valid genus in PASSANDRIDAE: PASSANDRINAE.

Type species: Passandra sexstriata Dalman, 1817, by monotypy.

Family-group name: PASSANDRITES Blanchard, 1845. Stem: Passandr-.

Passandrella Grouvelle, 1916a: 6. Current status: valid genus in PASSANDRIDAE: PASSANDRINAE.

Type species: Passandrella visenda Grouvelle, 1916, by monotypy.

Family-group name: PASSANDRELLINI Grouvelle, 1916. Stem: Passandrell-.

Patrizia Alluaud, 1931: 9. Current status: junior synonym of Agastus Schmidt-Göbel, 1846 in CARABIDAE: HARPALINAE: ZUPHIINI: PATRIZIINA.

Type species: Patrizia zuphioides Alluaud, 1931, by monotypy.

Family-group name: PATRIZIINI Basilewsky, 1953. Stem: Patrizi-.

Patrobus Dejean, 1821: 10. Current status: valid genus in CARABIDAE: PATROBINAE: PATROBINI: PATROBINA.

Type species: here fixed (under ICZN 1999a, Article 70.3.2) as Carabus atrorufus Strøm, 1768, misidentified as Carabus rufipes Fabricius, 1792 (sensu Duftschmid, 1812), in the subsequent designation by Curtis (1827: pl. 192).

Family-group name: PATROBIDAE W. Kirby, 1837. Stem: Patrob-.

Paulusiella Löbl, 2007: 32. Current status: valid genus in ELATERIDAE: HEMIOPINAE: PAULISIELLINI.

Type species: Paulusiella fossulatipennis Mandl, 1974, by original designation.

Family-group name: PAULUSIELLINI Kusy, Motyka & Bocák, 2023: 8. Stem: Paulisiell-.

Paussobrenthus Gestro, 1919: 271. Current status: valid genus in BRENTIDAE: BRENTINAE: BRENTINI: PAUSSOBRENTHINA.

Type species: Paussobrenthus bakeri Gestro, 1919, by monotypy.

Family-group name: PAUSSOBRENTHINI Gestro, 1919. Stem: Paussobrenth-.

Paussus Linnaeus, 1775: 6. Current status: valid genus in CARABIDAE: PAUSSINAE: PAUSSINI: PAUSSINA.

Type species: Paussus microcephalus Linnaeus, 1775, by monotypy.

Family-group name: PAUSSILI Latreille, 1806. Stem: Pauss-.

Paxillus W.S. MacLeay, 1819: 96, 105. Current status: valid genus in PASSALIDAE: PASSALINAE: PASSALINI.

Type species: Paxillus leachii W.S. MacLeay, 1819, by subsequent designation (Gravely 1918: 48).

Family-group name: PAXILLINAE Kuwert, 1891. Stem: Paxill-.

Pectocera Hope, 1842c: 79. Current status: valid genus in ELATERIDAE: DENDROMETRINAE: OXYNOPTERINI.

Type species: Pectocera cantori Hope, 1842, by subsequent designation (Hyslop 1921: 663).

Family-group name: PECTOCERINI Gurjeva, 1974. Stem: Pectocer-.

Pediacus Shuckard, 1839: 150, 185. Current status: valid genus in CUCUJIDAE: PEDIACINAE.

Type species: Cucujus dermestoides Fabricius, 1792, by monotypy.

Family-group name: PEDIACINAE Jin, Zwick, Ślipiński, Marris, Thomas & H. Pang, 2020a: 260. Stem: Pediac-.

Pediculota Ádám, 1987: 156. Current status: valid genus in STAPHYLINIDAE: ALEOCHARINAE: PEDICULOTINI.

Type species: Pediculota hamoriae Ádám, 1987 (= Microdota montandoni Roubal, 1909), by original designation.

Family-group name: PEDICULOTINI Ádám, 1987. Stem: Pediculot-.

Pedilophorus Steffahny, 1842: 35. Current status: valid genus in BYRRHIDAE: BYRRHINAE: PEDILOPHORINI.

Type species: Byrrhus auratus Duftschmid, 1825, by monotypy.

Family-group name: PEDILOPHORINI Casey, 1912. Stem: Pedilophor-.

Pedilus Fischer, 1820: pl. 5. Current status: valid genus in PYROCHROIDAE: PEDILINAE.

Type species: Pedilus fuscus Fischer, 1820, by monotypy.

Family-group name: PÉDILIDES Lacordaire, 1859. Stem: Pedil-.

Pedinus Latreille, 1797: 20. Current status: valid genus in TENEBRIONIDAE: BLAPTINAE: PEDININI: PEDININA.

Type species: Tenebrio femoralis Linnaeus, 1767, by subsequent designation (Stephens 1835b: 159). Note: no species were originally included in this genus by Latreille (1797); the first ones were the two species listed by Latreille (1802b: 175).

Family-group name: PEDINIDEN Eschscholtz, 1829. Stem: Pedin-.

Pegylis Erichson, 1847b: 657. Current status: valid genus in SCARABAEIDAE: MELOLONTHINAE: MELOLONTHINI: PEGYLINA.

Type species: Pegylis morio Erichson, 1847, by monotypy.

Family-group name: PEGYLINI M. Lacroix, 1989. Stem: Pegyl-.

Pelecium W. Kirby, 1819: 377. Current status: valid genus in CARABIDAE: HARPALINAE: PELECIINI.

Type species: Pelecium cyanipes W. Kirby, 1819, by monotypy.

Family-group name: PÉLÉCIDES Chaudoir, 1880. Stem: Peleci-.

Pelecophora Dejean, 1821: 115. Current status: valid genus in RHADALIDAE: RHADALINAE.

Type species: Notoxus illigeri Gyllenhal, 1808, by monotypy.

Family-group name: PELECOPHORINI Majer, 1987. Stem: Pelecophor-.

Pelecotoma Fischer, 1809: 293. Current status: valid genus in RIPIPHORIDAE: PELECOTOMINAE.

Type species: Pelecotoma mosquense Fischer, 1809 (= Ripiphorus fennicus Paykull, 1798), by monotypy.

Family-group name: PELECOTOMINI Guérin-Méneville, 1857. Stem: Pelecotom-.

Pelidnota W.S. MacLeay, 1819: 157. Current status: valid genus in SCARABAEIDAE: RUTELINAE: RUTELINI: RUTELINA. Note: name placed on the Official List of Generic Names in Zoology (ICZN 2003c).

Type species: Scarabaeus punctatus Linnaeus, 1758, by monotypy (see ICZN 2003c). Note: name placed on the Official List of Specific Names in Zoology (ICZN 2003c).

Family-group name: PELIDNOTIDAE Burmeister, 1844. Stem: Pelidnot-.

Peliocypas Schmidt-Göbel, 1846: 33. Current status: valid genus in CARABIDAE: HARPALINAE: LEBIINI: DEMETRIADINA.

Type species: Peliocypas suturalis Schmidt-Göbel, 1846, by subsequent designation (Jeannel 1949b: 991).

Family-group name: PELIOCYPINI Basilewsky, 1984. Stem: Peliocypad-.

Pelmatellus H.W. Bates, 1882: 68. Current status: valid genus in CARABIDAE: HARPALINAE: HARPALINI: PELMATELLINA.

Type species: Pelmatellus nitescens H.W. Bates, 1882, by subsequent designation (Goulet 1974: 84).

Family-group name: PELMATELLINAE H.W. Bates, 1882. Stem: Pelmatell-.

Pelonium Spinola, 1845a: 347. Current status: valid genus in CLERIDAE: KORYNETINAE.

Type species: Enoplium viridipenne W. Kirby, 1818, by subsequent designation (Audinet- Serville 1845: 429).

Family-group name: PELONIINAE Opitz, 2010. Stem: Peloni-.

Pelonomus Erichson, 1847a: 510. Current status: valid genus in DRYOPIDAE.

Type species: Dryops picipes G.-A. Olivier, 1791, by subsequent designation (Lacordaire 1854: 503).

Family-group name: PELONOMINAE Böving & Craighead, 1931. Stem: Pelonom-.

Pelophila Dejean, 1821: 7. Current status: valid genus in CARABIDAE: NEBRIINAE: PELOPHILINI.

Type species: Carabus borealis Paykull, 1790, by monotypy.

Family-group name: PELOPHILINI Kavanaugh, 1996. Stem: Pelophil-.

Pelops Kaup, 1871: 37. Current status: invalid senior synonym of Protomocoelus Zang, 1905 in PASSALIDAE: PASSALINAE: MACROLININI. Note: junior homonym of Pelops Gistel, 1834 [Coleoptera: TENEBRIONIDAE] and Pelops C.L. Koch, 1835 [Acari]; the name Pelops Gistel, 1834 was recently treated as a nomen oblitum by Bousquet and Bouchard (2017: 132).

Type species: Passalus australis Boisduval, 1835, by subsequent designation (Gravely 1918: 107).

Family-group name: PELOPINAE Kuwert, 1896. Stem: Pelop-. Note: senior homonym of PELOPINAE Ewing, 1917 (type genus Pelops C.L. Koch, 1835) in Acari; family-group name permanently invalid (ICZN 1999a, Article 39).

Peloropus Schönherr, 1835: 456. Current status: valid genus in CURCULIONIDAE: CONODERINAE: PELOROPODINI.

Type species: Peloropus ulula Gyllenhal, 1835, by original designation.

Family-group name: PELEROPINI Hustache, 1932. Stem: Peloropod-.

Pelossus J. Thomson, 1864: 222. Current status: valid genus in CERAMBYCIDAE: CERAMBYCINAE: LYGRINI.

Type species: Corethrogaster ruber J. Thomson, 1858, by original designation.

Family-group name: PELOSSINI Tavakilian, 2013: 111, syn. nov. Stem: Peloss-. Note: this family-group name is unnecessary since it was proposed as a new replacement name for LYGRINI Sama, 2008 because of the synonymy of Lygrus Fåhraeus, 1872 with Pelossus J. Thomson, 1864; however, according to Article 40.1 (ICZN 1999a) when a name of a type genus is considered to be a junior synonym, the family-group name is not to be replaced on that account alone.

Peltastica Mannerheim, 1852: 334. Current status: valid genus in DERODONTIDAE: PELTASTICINAE.

Type species: Peltastica tuberculata Mannerheim, 1852, by monotypy.

Family-group name: PELTASTICIDAE J.L. LeConte, 1861. Stem: Peltastic-.

Peltinodes Paulian, 1950: 19. Current status: junior synonym of Holopsis Broun, 1883 in CORYLOPHIDAE: CORYLOPHINAE: PELTINODINI.

Type species: Peltinus gigas Abeille de Perrin, 1894, by monotypy.

Family-group name: PELTINODITAE Paulian, 1950. Stem: Peltinod-.

Peltinus Mulsant & Rey, 1861: 137. Current status: invalid senior synonym of Teplinus Pakaluk, Ślipiński & Lawrence, 1994 in CORYLOPHIDAE: CORYLOPHINAE: TEPLININI. Note: junior homonym of Peltinus Rafinesque, 1815 (unjustified emendation of Peltis Kugelann, 1792) [Coleoptera: PELTIDAE].

Type species: Peltinus velatus Mulsant & Rey, 1861, by monotypy.

Family-group name: PELTININI Paulian, 1950. Stem: Peltin-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

Peltis Kugelann, 1792: 508. Current status: junior synonym of Ostoma Laicharting, 1781 in PELTIDAE. Note: this name has been credited to O.F. Müller (1764: 13) in the literature (e.g., Kolibáč 2007: 366, Kolibáč et al. 2016: 683) but in Opinion 1754 (ICZN 1994a) the Commission ruled that Peltis Geoffroy, 1762 and all other uses of the name prior to the publication of Peltis Kugelann, 1792 are suppressed for the purposes of both the Principle of Priority and the Principle of Homonymy, while Peltis Kugelann, 1792 was placed on the Official List of Generic Names in Zoology (ICZN 1994a); the change of authorship for Peltis from “O.F. Müller, 1764” to “Kugelann, 1792” by the Commission (ICZN 1994a) resulted in the fact that Ostoma Laicharting, 1781 is now the oldest name for this genus and should be used as valid.

Type species: Silpha grossa Linnaeus, 1758, by subsequent designation (Hope 1841a: 150; see ICZN 1994a). Note: name placed on the Official List of Specific Names in Zoology (ICZN 1994a).

Family-group name: PELTIDAE W. Kirby, 1837. Stem: Pelt-.

Peltodytes Régimbart, 1879: 450, 457. Current status: valid genus in HALIPLIDAE.

Type species: Dytiscus caesus Duftschmid, 1805, by subsequent designation (F. Balfour- Browne 1936: 68).

Family-group name: PELTODYTINAE Böving & Craighead, 1931. Stem: Peltodyt-.

Peltonotus Burmeister, 1847: 75. Current status: valid genus in SCARABAEIDAE: DYNASTINAE: CYCLOCEPHALINI.

Type species: Peltonotus morio Burmeister, 1847, by monotypy. Note: Burmeister (1847: 75) mentioned a second species, Melolontha scarabaeina Gyllenhal, 1817 [attributed to Schönherr], but in the Anhang [supplement] in the same publication (p. 521) he placed it in Cyclocephala Dejean, 1821, which is where it is placed currently; therefore, we do not consider Melolontha scarabaeina Gyllenhal, 1817 to be an originally included species.

Family-group name: PELTONOTINI Arrow, 1917. Stem: Peltonot-.

†Peltosyne Ponomarenko, 1977: 105. Current status: valid genus in †PELTOSYNIDAE. Note: Lower Triassic (Kyrgyzstan).

Type species: †Peltosyne triassica Ponomarenko, 1977, by original designation.

Family-group name: †PELTOSYNIDAE Yan, Beutel & Ponomarenko, 2017b: 516. Stem: Peltosyn-.

Peneta Lacordaire, 1859a: 319. Current status: valid genus in TENEBRIONIDAE: PHRENAPATINAE: PENETINI.

Type species: Peneta lebasii Lacordaire, 1859, by subsequent designation (R. Lucas 1920: 492).

Family-group name: PÉNÉTIDES Lacordaire, 1859. Stem: Penet-.

Penia Laporte, 1838: 11. Current status: valid genus in ELATERIDAE: DENDROMETRINAE: DIMINI.

Type species: Elater eschscholtzii Hope, 1831, by monotypy.

Family-group name: PENIINI Dollin, 1990. Stem: Peni-.

Penichrolucanus Deyrolle, 1863: 485. Current status: valid genus in LUCANIDAE: LUCANINAE: FIGULINI.

Type species: Penichrolucanus copricephalus Deyrolle, 1863, by monotypy.

Family-group name: PENICHROLUCANINAE Arrow, 1950. Stem: Penichrolucan-.

Penicillophorus Paulus, 1975: 69. Current status: valid genus in PHENGODIDAE: PENICILLOPHORINAE.

Type species: Penicillophorus ctenotarsus Paulus, 1975, by original designation.

Family-group name: PENICILLOPHORINI Paulus, 1975. Stem: Penicillophor-.

Pentagonica Schmidt-Göbel, 1846: 47. Current status: valid genus in CARABIDAE: HARPALINAE: ODACANTHINI: PENTAGONICINA.

Type species: Pentagonica ruficollis Schmidt-Göbel, 1846, by subsequent designation (Andrewes 1938: 137).

Family-group name: PENTAGONICINAE H.W. Bates, 1873. Stem: Pentagonic-.

Pentaphyllus Dejean, 1821: 68. Current status: valid genus in TENEBRIONIDAE: DIAPERINAE: DIAPERINI: DIAPERINA.

Type species: Mycetophagus testaceus Hellwig, 1792, by monotypy.

Family-group name: PENTAPHYLLAIRES Mulsant, 1854. Stem: Pentaphyll-. Note: senior homonym of PENTAPHYLLINAE Schindewolf, 1942 (type genus Pentaphyllum de Koninck, 1872) in Anthozoa.

Pentaplatarthrus Westwood, 1833: 616. Current status: valid genus in CARABIDAE: PAUSSINAE: PAUSSINI: PENTAPLATARTHRINA.

Type species: Pentaplatarthrus paussoides Westwood, 1833, by original designation.

Family-group name: PENTAPLATARTHRINI Jeannel, 1946. Stem: Pentaplatarthr-.

Pentaria Mulsant, 1856c: 135. Current status: valid genus in SCRAPTIIDAE: ANASPIDINAE: PENTARIINI.

Type species: Pentaria sericaria Mulsant, 1856 (= Anaspis badia Rosenhauer, 1847), by monotypy.

Family-group name: PENTARIINI Franciscolo, 1954. Stem: Pentari-.

Pentarthrum Wollaston, 1854: 129. Current status: valid genus in CURCULIONIDAE: COSSONINAE: PENTARTHRINI.

Type species: Pentarthrum huttoni Wollaston, 1854, by monotypy.

Family-group name: PENTARTHRIDES Lacordaire, 1865. Stem: Pentarthr-.

Penthe Newman, 1838: 373. Current status: valid genus in TETRATOMIDAE: PENTHINAE.

Type species: Helops obliquatus Fabricius, 1798, by subsequent designation (Hope 1841a: 133).

Family-group name: PENTHIDES Lacordaire, 1859. Stem: Penth-.

Pentilia Mulsant, 1850: 502. Current status: valid genus in COCCINELLIDAE: COCCINELLINAE: CRYPTOGNATHINI.

Type species: Pentilia egena Mulsant, 1850, by subsequent designation (Crotch 1874a: 199).

Family-group name: PENTILIINI Casey, 1899. Stem: Pentili-.

Pentodon Hope, 1837: 92. Current status: valid genus in SCARABAEIDAE: DYNASTINAE: PENTODONTINI: PENTODONTINA. Note: name placed on the Official List of Generic Names in Zoology (ICZN 2003c).

Type species: Scarabaeus punctatus Villers, 1789, by original designation (see ICZN 2003c). Note: name ruled not to be invalid by reason of being a junior primary homonym of Scarabaeus punctatus Linnaeus, 1758 and placed on the Official List of Specific Names in Zoology (ICZN 2003c).

Family-group name: PENTODONAIRES Mulsant, 1842. Stem: Pentodont-.

Pericalus W.S. MacLeay, 1825: 15. Current status: valid genus in CARABIDAE: HARPALINAE: LEBIINI: PERICALINA.

Type species: Catascopus cicindeloides W.S. MacLeay, 1825, by monotypy.

Family-group name: PERICALLIDAE Hope, 1838. Stem: Perical-.

Peridinetus Schönherr, 1837: 467. Current status: valid genus in CURCULIONIDAE: BARIDINAE: PERIDINETINI.

Type species: Curculio irroratus Fabricius, 1787, by original designation.

Family-group name: PÉRIDINÉTIDES Lacordaire, 1865. Stem: Peridinet-.

Perieges Schönherr, 1842a: 419. Current status: valid genus in CURCULIONIDAE: BRACHYCERINAE: CRYPTOLARYNGINI.

Type species: Perieges bardus Boheman, 1842, by original designation.

Family-group name: PERIEGINI Legalov, 2003. Stem: Perieg-.

Periglossium Liebke, 1929: 246. Current status: junior synonym of Inna Putzeys, 1861 in CARABIDAE: HARPALINAE: LEBIINI: PERICALINA.

Type species: Periglossium nevermanni Liebke, 1929, by original designation.

Family-group name: PERIGLOSSIINAE Liebke, 1929. Stem: Periglossi-.

Perigona Laporte, 1834c: 151. Current status: valid genus in CARABIDAE: HARPALINAE: PERIGONINI.

Type species: Perigona pallida Laporte, 1834, by monotypy.

Family-group name: PERIGONAE G.H. Horn, 1881. Stem: Perigon-. Note: nomen protectum (see Bouchard et al. 2011: 140).

Perileptus Schaum, 1860: 663. Current status: valid genus in CARABIDAE: TRECHINAE: TRECHINI: TRECHODINA.

Type species: Carabus areolatus Creutzer, 1799, by original designation.

Family-group name: PERILEPTIDES Sloane, 1903. Stem: Perilept-.

Perimylops C. Müller, 1884: 419. Current status: valid genus in PROMECHEILIDAE.

Type species: Perimylops antarcticus C. Müller, 1884, by monotypy. Note: the original spelling antracticus was emended to antarcticus by Gebien (1905: 255); the emended name is in prevailing usage, attributed to the publication of the original spelling, and therefore deemed to be a justified emendation and the correct original spelling (ICZN 1999a, Article 33.2.3.1).

Family-group name: PERIMYLOPIDAE St. George, 1939. Stem: Perimylop-.

Perinthus Casey, 1890a: 192. Current status: valid genus in STAPHYLINIDAE: ALEOCHARINAE: TERMITONANNINI: PERINTHINA.

Type species: Perinthus dudleyanus Casey, 1890, by monotypy.

Family-group name: PERINTHI Bernhauer & Scheerpeltz, 1926. Stem: Perinth-.

Periommatus Chapuis, 1865: 316. Current status: valid genus in CURCULIONIDAE: PLATYPODINAE: TESSEROCERINI: TESSEROCERINA.

Type species: Periommatus longicollis Chapuis, 1865, by monotypy.

Family-group name: PERIOMATINI Schedl, 1939. Stem: Periommat-.

Periptyctus Blackburn, 1895: 234. Current status: valid genus in CORYLOPHIDAE: PERIPTYCTINAE.

Type species: Periptyctus russulus Blackburn, 1895, by monotypy.

Family-group name: PERIPTYCTINAE Ślipiński, Lawrence & Tomaszewska, 2001. Stem: Periptyct-.

Peritelus Germar, 1823: 407. Current status: valid genus in CURCULIONIDAE: ENTIMINAE: CYPHICERITAE: PERITELINI: PERITELINA.

Type species: Peritelus sphaeroides Germar, 1823, by subsequent designation (Schönherr 1826: 193).

Family-group name: PÉRITÉLIDES Lacordaire, 1863. Stem: Peritel-.

†Permocupes Martynov, 1933: 85. Current status: valid genus in †PERMOCUPEDIDAE: †PERMOCUPEDINAE. Note: Permian (Russia).

Type species: †Permocupes semenovi Martynov, 1933, by original designation.

Family-group name: †PERMOCUPIDAE Martynov, 1933. Stem: Permocuped-.

†Permosyne Tillyard, 1924: 431. Current status: valid genus in †PERMOSYNIDAE. Note: Permian (Australia, South Africa, Russia).

Type species: †Permosyne belmontensis Tillyard, 1924, by original designation.

Family-group name: †PERMOSYNIDAE Tillyard, 1924. Stem: Permosyn-.

Perochnoristhus Basilewsky, 1973: 225. Current status: valid genus in CARABIDAE: HARPALINAE: IDIOMORPHINI.

Type species: Perochnoristhus penrithae Basilewsky, 1973, by original designation.

Family-group name: PEROCHNORISTHINAE Basilewsky, 1973. Stem: Perochnoristh-.

Perothops Eschscholtz in Laporte, 1838: table. Current status: valid genus in EUCNEMIDAE: PEROTHOPINAE.

Type species: Perothops cervinus Germar, 1839, by subsequent monotypy (Germar 1839: 197).

Family-group name: PÉROTHOPIDES Lacordaire, 1857. Stem: Perothop-.

Perrhynchites Voss, 1953: 45. Current status: valid genus in ATTELABIDAE: RHYNCHITINAE: RHYNCHITITAE: RHYNCHITINI: PERRHYNCHITINA.

Type species: Rhynchites aereipennis Desbrochers des Loges, 1895, by monotypy.

Family-group name: PERRHYNCHITINA Legalov, 2003. Stem: Perrhynchit-.

Perrotius Fleutiaux, 1938: 134. Current status: valid genus in EUCNEMIDAE: EUCNEMINAE: PERROTIINI.

Type species: Perrotius tenuipes Fleutiaux, 1938, by monotypy.

Family-group name: PERROTINI Muona, 1993. Stem: Perroti-.

Pertinax Harold in Gemminger and Harold, 1868c: 976. Current status: valid subgenus of Passalus Fabricius, 1792 in PASSALIDAE: PASSALINAE: PASSALINI. Note: this name is attributed to Kaup (1869: 20) in the literature; however, it was first made available by Harold in 1868 when is listed several nominal species under this genus name.

Type species: Passalus convexus Dalman, 1817, by subsequent designation (Hincks and Dibb 1935: 38).

Family-group name: PERTINACEAE Kaup, 1871. Stem: Pertinac-.

Perucola Théry, 1925: 122. Current status: valid genus in BUPRESTIDAE: POLYCESTINAE: PERUCOLINI.

Type species: Perucola picturata Théry, 1925, by monotypy.

Family-group name: PERUCOLINI Cobos, 1980. Stem: Perucol-.

Peryphus Dejean, 1821: 17. Current status: valid subgenus of Bembidion Latreille, 1802 in CARABIDAE: TRECHINAE: BEMBIDIINI.

Type species: Carabus littoralis G.-A. Olivier, 1795 (= Bembidion tetracolum Say, 1823), by subsequent designation (Stephens 1835b: 286).

Family-group name: PERYPHIDAE W. Kirby, 1837. Stem: Peryph-.

Petalochilus Schönherr, 1836: 591. Current status: valid genus in CURCULIONIDAE: MOLYTINAE: PETALOCHILINI.

Type species: Petalochilus gemellus Gyllenhal, 1836, by original designation.

Family-group name: PÉTALOCHILIDES Lacordaire, 1863. Stem: Petalochil-.

Petanops Jeannel, 1954c: 99. Current status: junior synonym of Daveyia Lea, 1912 in STAPHYLINIDAE: PSELAPHINAE: PSELAPHITAE: TYRINI: CENTROPHTHALMINA.

Type species: Petanops armstrongi Jeannel, 1954 (= Daveyia mira Lea, 1912), by monotypy.

Family-group name: PETANOPINI Jeannel, 1954. Stem: Petanop-.

Petrejus Harold in Gemminger and Harold, 1868c: 978. Current status: valid genus in PASSALIDAE: PASSALINAE: PASSALINI. Note: this name is attributed to Kaup (1869: 36) in the literature; however, it was first made available by Harold in 1868 when he listed four nominal species under this genus name.

Type species: Passalus mucronatus Burmeister, 1847, by subsequent designation (Brauer 1870: 127).

Family-group name: PETREJINAE Kuwert, 1891. Stem: Petrej-.

Petria Semenov, 1894: 363. Current status: valid genus in TENEBRIONIDAE: ALLECULINAE: CTENIOPODINI.

Type species: Petria tachyptera Semenov, 1894, by subsequent designation (R. Lucas 1920: 496).

Family-group name: PETRIIDAE Semenov, 1894. Stem: Petri-.

Petrognatha Leach, 1819: 495. Current status: valid genus in CERAMBYCIDAE: LAMIINAE: PETROGNATHINI.

Type species: Lamia gigas Fabricius, 1792, by monotypy.

Family-group name: PÉTROGNATHITES Blanchard, 1845. Stem: Petrognath-.

Phacellus Dejean, 1835: 335. Current status: valid genus in CERAMBYCIDAE: LAMIINAE: PHACELLINI.

Type species: Acanthocinus boryi Gory, 1832, by monotypy.

Family-group name: PHACELLIDES Lacordaire, 1872. Stem: Phacell-.

Phaedimus G.R. Waterhouse in Westwood, 1841a: 5. Current status: valid genus in SCARABAEIDAE: CETONIINAE: PHAEDIMINI.

Type species: Mycteristes cumingii G.R. Waterhouse, 1841, by monotypy.

Family-group name: PHAEDIMI Schoch, 1894. Stem: Phaedim-. Note: junior homonym of PHAEDIMIDAE Robineau-Desvoidy, 1863 (type genus Phaedima Robineau-Desvoidy, 1863) in Diptera and PHAEDIMIDAE Thorell, 1890 (type genus Phaedima Thorell, 1881) in Arachnida; Thorell’s name is permanently invalid (based on a preoccupied type genus).

Phaedon Latreille, 1829b: 151. Current status: valid genus in CHRYSOMELIDAE: CHRYSOMELINAE: CHRYSOMELINI.

Type species: Chrysomela armoraciae Linnaeus, 1758, by subsequent designation (Stephens 1835b: 296).

Family-group name: PHAEDONINI Weise, 1915. Stem: Phaedon-.

Phaenithon Schönherr, 1826: 37. Current status: valid genus in ANTHRIBIDAE: ANTHRIBINAE: CORRHECERINI.

Type species: Anthribus costatus Schönherr, 1826 (= Anthribus curvipes Germar, 1823), by original designation.

Family-group name: PHAENITHONINI Pierce, 1930. Stem: Phaenithon-.

Phaenocephalus Wollaston, 1873c: 167. Current status: valid genus in PHALACRIDAE: PHAENOCEPHALINAE.

Type species: Phaenocephalus castaneus Wollaston, 1873, by monotypy.

Family-group name: PHAENOCEPHALIDAE A. Matthews, 1899. Stem: Phaenocephal-.

Phaenocerus Bonvouloir, 1871: 68, 285. Current status: valid genus in EUCNEMIDAE: EUCNEMINAE: PHAENOCERINI. Note: this name was first used by Gemminger in Gemminger and von Harold (1869b: 1469) but it is not available from that date since it was not accompanied by a description or indication (ICZN 1999a: Article 12.1).

Type species: Phaenocerus subclavatus Bonvouloir, 1871, by monotypy.

Family-group name: PHAENOCERINI Muona, 1993. Stem: Phaenocer-.

Phaenognatha Hope, 1842a: 425. Current status: valid genus in SCARABAEIDAE: ACLOPINAE: PHAENOGNATHINI. Note: two original spellings were used, including Phaenognathus (p. 425); subsequently, Hope (1845a: 113) used only Phaenognatha and therefore acted as First Reviser (ICZN 1999a, Article 24.2.4).

Type species: Phaenognatha erichsoni Hope, 1842, by monotypy.

Family-group name: PHAENOGNATHINI Iablokoff-Khnzorian, 1977. Stem: Phaenognath-.

Phaenomeris Hope, 1833: 62. Current status: valid genus in SCARABAEIDAE: PHAENOMERIDINAE. Note: name placed on the Official List of Generic Names in Zoology (ICZN 1962).

Type species: Phaenomeris magnifica Hope, 1833, by monotypy (see ICZN 1962). Note: name placed on the Official List of Specific Names in Zoology (ICZN 1962).

Family-group name: PHAENOMERINI Erichson, 1847. Stem: Phaenomerid- (ICZN 1962).

Phaenomerus Schönherr, 1836: 632. Current status: valid genus in CURCULIONIDAE: CONODERINAE: CAMPYLOSCELINI: PHAENOMERINA. Note: name placed on the Official List of Generic Names in Zoology (ICZN 1962).

Type species: Phaenomerus sundewalli Boheman, 1836, by original designation (see ICZN 1962). Note: name placed on the Official List of Specific Names in Zoology (ICZN 1962).

Family-group name: PHAENOMERINA Faust, 1898. Stem: Phaenomer-.

Phaeochrous Laporte, 1840b: 108. Current status: valid genus in HYBOSORIDAE: HYBOSORINAE.

Type species: Phaeochrous emarginatus Laporte, 1840, by subsequent designation (Paulian 1937: 139).

Family-group name: PHAEOCHROINEN Kolbe, 1912. Stem: Phaeochro-.

Phaeopholus Roelofs, 1873: 180. Current status: valid genus in CURCULIONIDAE: HYPERINAE: HYPERINI: PHAEOPHOLINA. Note: two original spellings were used, including Phoeophilus (p. 193); Alonso-Zarazaga and Lyal (1999: 189) acted as First Revisers (ICZN 1999a, Article 24.2.3) and rejected the spelling Phoeophilus; the alternative original spelling Phaaeopholus was corrected to Phaeopholus in an unnumbered page entitled ‘Errata pour Curculionides recueillis au Japon par M. G. Lewis’ inserted toward the end of the journal volume.

Type species: Phaeopholus ornatus Roelofs, 1874, by monotypy.

Family-group name: PHAEOPHOLINA Legalov, 2011d: 153. Stem: Phaeophol-.

Phalacrus Paykull, 1800: 438. Current status: valid genus in PHALACRIDAE: PHALACRINAE.

Type species: Anisotoma corrusca Panzer, 1796, by subsequent designation (Westwood 1838c: 10).

Family-group name: PHALACRURIDA Leach, 1815. Stem: Phalacr-.

Phalangogonia Burmeister, 1844: 451. Current status: valid genus in SCARABAEIDAE: RUTELINAE: ANOPLOGNATHINI: PHALANGOGONIINA.

Type species: Phalangogonia obesa Burmeister, 1844, by monotypy.

Family-group name: PHALANGOGONIINA Ohaus, 1918. Stem: Phalangogoni-.

Phalepsus Westwood, 1870: 131. Current status: valid genus in STAPHYLINIDAE: PSELAPHINAE: PSELAPHITAE: PHALEPSINI.

Type species: Phalepsus subglobosus Westwood, 1870, by monotypy.

Family-group name: PHALEPSINI Jeannel, 1949. Stem: Phaleps-.

Phaleria Latreille, 1802b: 162. Current status: valid genus in TENEBRIONIDAE: DIAPERINAE: PHALERIINI. Note: name placed on the Official List of Generic Names in Zoology (ICZN 1975).

Type species: Tenebrio cadaverinus Fabricius, 1792, by plenary power (see ICZN 1975). Note: name placed on the Official List of Specific Names in Zoology (ICZN 1975).

Family-group name: PHALÉRIIDES Blanchard, 1845. Stem: Phaleri-.

Phalidura Fischer, 1823b: 265. Current status: junior synonym of Amycterus Schönherr, 1823 in CURCULIONIDAE: CYCLOMINAE: AMYCTERINI.

Type species: Phalidura mirabilis Fischer, 1823 (= Amycterus talpa Schönherr, 1823), by monotypy.

Family-group name: PSALIDURIDAE Pierce, 1914. Stem: Phalidur-.

Phalota Pascoe, 1863b: 559. Current status: valid genus in CERAMBYCIDAE: CERAMBYCINAE: PHALOTINI.

Type species: Phalota tenella Pascoe, 1863, by monotypy.

Family-group name: PHALOTIDES Lacordaire, 1868. Stem: Phalot-.

Phanaeus W.S. MacLeay, 1819: 124. Current status: valid genus in SCARABAEIDAE: SCARABAEINAE: PHANAEINI: PHANAEINA.

Type species: here fixed (under ICZN 1999a, Article 70.3.2) as Phanaeus vindex W.S. MacLeay, 1819, misidentified as Scarabaeus carnifex Linnaeus, 1758 (sensu Linnaeus, 1767), in the subsequent designation by d’Olsoufieff (1924: 23). Note: see Edmonds (1994: 90) for remarks about the misidentification of the type species.

Family-group name: PHANAEIDAE Hope, 1838. Stem: Phanae-.

Phanerochila Fleutiaux, 1896: 314. Current status: valid genus in PTINIDAE: PTILININAE: PHANEROCHILINI.

Type species: Phanerochila boliviensis Fleutiaux, 1896, by monotypy.

Family-group name: PHANEROCHILINI Zahradník in Zahradník and Háva, 2014: 307. Stem: Phanerochil-.

Phanerotoma Solier, 1843: 82, 126. Current status: invalid senior synonym of Ocnodes Fåhraeus, 1870 in TENEBRIONIDAE: PIMELIINAE: SEPIDIINI: MOLURINA. Note: junior homonym of Phanerotoma Wesmael, 1838 [Hymenoptera].

Type species: Phanerotoma elongata Solier, 1843 (= Pimelia laevigata G.-A. Olivier, 1795), by original designation.

Family-group name: PHANEROTOMOIDE C. Koch, 1955. Stem: Phanerotom-. Note: family- group name permanently invalid (ICZN 1999a, Article 39).

Phanerotomea C. Koch, 1958: 58. Current status: junior synonym of Ocnodes Fåhraeus, 1870 in TENEBRIONIDAE: PIMELIINAE: SEPIDIINI: MOLURINA. Note: new replacement name for Phanerotoma Solier, 1843.

Type species: Phanerotoma elongata Solier, 1843 (= Pimelia laevigata G.-A. Olivier, 1795), by original designation (type fixation for Phanerotoma Solier).

Family-group name: PHANEROTOMEINA C. Koch, 1958. Stem: Phanerotome-.

Phantasis J. Thomson, 1860a: 25. Current status: valid genus in CERAMBYCIDAE: LAMIINAE: PHANTASINI.

Type species: Phantasis terribilis J. Thomson, 1860 (= Phrissoma giganteum Guérin- Méneville, 1844), by subsequent designation (J. Thomson 1864: 42).

Family-group name: PHANTASINAE Kolbe, 1897. Stem: Phantas-.

Pharangispa Maulik, 1929: 233. Current status: valid genus in CHRYSOMELIDAE: CASSIDINAE: COELAENOMENODERINI.

Type species: Pharangispa purpureipennis Maulik, 1929, by monotypy.

Family-group name: PHARANGISPINI Uhmann, 1940. Stem: Pharangisp-.

Pharaxonotha Reitter, 1875b: 39, 44. Current status: valid genus in EROTYLIDAE: PHARAXONOTHINAE.

Type species: Pharaxonotha kirschii Reitter, 1875, by monotypy.

Family-group name: PHARAXONOTHINAE Crowson, 1952. Stem: Pharaxonoth-.

Pharochilus Kaup, 1868a: 20. Current status: valid genus in PASSALIDAE: PASSALINAE: MACROLININI.

Type species: Passalus dilatatus Dalman, 1817, by subsequent designation (Gravely 1918: 97).

Family-group name: PHAROCHILINAE Kuwert, 1891. Stem: Pharochil-.

Pharus Mulsant, 1850: 948, 949. Current status: invalid senior synonym of Pharoscymnus Bedel, 1906 in COCCINELLIDAE: COCCINELLINAE: STICHOLOTIDINI. Note: junior homonym of Pharus Konig, 1841 [Mollusca].

Type species: Coccinella sexguttata Gyllenhal, 1808, by monotypy.

Family-group name: PHARINI Casey, 1899. Stem: Phar-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

Phellopsis J.L. LeConte, 1862: 216. Current status: valid genus in ZOPHERIDAE: ZOPHERINAE: PHELLOPSINI.

Type species: Bolitophagus obcordatus W. Kirby, 1837, by subsequent designation (Casey 1907: 470).

Family-group name: PHELLOPSINI Ślipiński & Lawrence, 1999. Stem: Phellops-.

Phengodes Illiger, 1807b: 341. Current status: valid genus in PHENGODIDAE: PHENGODINAE.

Type species: Lampyris plumosa G.-A. Olivier, 1790, by monotypy.

Family-group name: PHENGODINI J.L. LeConte, 1861. Stem: Phengod-.

Pheropsophus Solier, 1834a: 461. Current status: valid genus in CARABIDAE: BRACHININAE: BRACHININI: PHEROPSOPHINA.

Type species [suggested]: Brachinus madagascariensis Dejean, 1831, by subsequent designation (E. Desmarest 1851: 85). Note: the valid type species fixation is that of Hope (1838a: 99), who selected Carabus complanatus Linnaeus, 1767 (sensu Fabricius, 1801) (= Cicindela aequinoctialis Linnaeus, 1763); however, neither the nominal species nor the taxonomic species involved in the type species fixation by Hope (1838a: 99) is currently recognized as a member of the genus Pheropsophus Solier; an application to the Commission is necessary to fix Brachinus madagascariensis Dejean, 1831 as the type species of Pheropsophus Solier, 1834.

Family-group name: PHEROPSOPHINI Jeannel, 1949. Stem: Pheropsoph-.

Phialodes Roelofs, 1874: 137. Current status: valid genus in ATTELABIDAE: ATTELABINAE: ATTELABINI: PHIALODINA.

Type species: Phialodes rufipennis Roelofs, 1874, by subsequent designation (Voss 1925: 264).

Family-group name: PHIALODINA Legalov, 2003. Stem: Phialod-.

Phibalus Gistel, 1856: 384. Current status: valid genus in TENEBRIONIDAE: ALLECULINAE: CTENIOPODINI.

Type species: Chrysomela pubescens Linnaeus, 1767, by monotypy.

Family-group name: PHIBALIDAE Gistel, 1856. Stem: Phibal-.

Philacamatus Bruch, 1933b: 206. Current status: valid genus in STAPHYLINIDAE: ALEOCHARINAE: CREMATOXENINI.

Type species: Philacamatus bosqi Bruch, 1933, by monotypy.

Family-group name: PHILACAMATINI Bruch, 1933. Stem: Philacamat-.

Philanthaxia Deyrolle, 1865: 72. Current status: valid genus in BUPRESTIDAE: BUPRESTINAE: THOMASSETIINI: PHILANTHAXIINA.

Type species: Philanthaxia curta Deyrolle, 1865, by monotypy.

Family-group name: PHILANTHAXIINA Holyński, 1988. Stem: Philanthaxi-.

Phileurus Latreille, 1806: 103. Current status: valid genus in SCARABAEIDAE: DYNASTINAE: PHILEURINI: PHILEURINA.

Type species: Scarabaeus didymus Linnaeus, 1758, by subsequent designation (Latreille 1810: 428).

Family-group name: PHILEURIDAE Burmeister, 1847. Stem: Phileur-.

Philippinauletes Legalov, 2023c: 24. Current status: valid genus in ATTELABIDAE: RHYNCHITINAE: RHINOCARTITAE: PHILIPPINAULETINI. Note: Philippinauletes Legalov (2018c: 795) was published in a work that is nomenclaturally unavailable (ICZN 2012f, Article 8.5.3).

Type species: Philippinauletes rubrauletiformis Legalov, 2023, by original designation. Note: Philippinauletes rubrauletiformis Legalov, 2018 was published in a work that is nomenclaturally unavailable (ICZN 2012f, Article 8.5.3).

Family-group name: PHILIPPINAULETINI Legalov, 2023: 24. Stem: Philippinaulet-. Note: PHILIPPINAULETINI Legalov (2018c: 794) was published in a work that is nomenclaturally unavailable (ICZN 201f2, Article 8.5.3).

Philonthus Stephens, 1829b: 23. Current status: valid genus in STAPHYLINIDAE: STAPHYLININAE: STAPHYLININI: PHILONTHINA.

Type species: Staphylinus politus Linnaeus, 1758, by subsequent designation (Stephens 1835b: 312). Note: the type species usually listed for Philonthus Stephens is Staphylinus splendens Fabricius, 1792 (e.g., Schülke and Smetana 2015: 1034) by subsequent designation of Curtis (1836: pl. 610) but Stephens’ designation is older; both species are currently included in the nominotypical subgenus of Philonthus Stephens, 1829.

Family-group name: PHILONTHIDAE W. Kirby, 1837. Stem: Philonth-.

Philopedon Schönherr, 1826: 97. Current status: valid genus in CURCULIONIDAE: ENTIMINAE: POLYDRUSITAE: CNEORHININI.

Type species: Curculio geminatus Fabricius, 1787 (= Curculio plagiatus Schaller, 1783), by original designation.

Family-group name: PHILOPEDINI Bedel, 1883. Stem: Philoped-.

Philotermes Kraatz, 1857a: 13. Current status: valid genus in STAPHYLINIDAE: ALEOCHARINAE: PHILOTERMITINI.

Type species: Philotermes pilosus Kraatz, 1857, by subsequent designation (Fenyes 1919: 24).

Family-group name: PHILOTERMITINI Seevers, 1957. Stem: Philotermit-.

Philothalpus Kraatz, 1857b: 540. Current status: valid genus in STAPHYLINIDAE: STAPHYLININAE: STAPHYLININI: PHILOTHALPINA.

Type species: Philonthus fervidus Erichson, 1840, by subsequent designation (Herman 2001: 10).

Family-group name: PHILOTHALPINA Chatzimanolis & Brunke in Chani-Posse, Brunke, Chatzimanolis, Schillhammer and Solodovnikov, 2018: 34. Stem: Philothalp-.

Philus Saunders, 1853: 110. Current status: valid genus in VESPERIDAE: PHILINAE.

Type species: Philus inconspicuus Saunders, 1853 (= Stenochorus antennatus Gyllenhal, 1817), by monotypy.

Family-group name: PHILITAE J. Thomson, 1861. Stem: Phil-.

Phlegon Laporte, 1840a: 254. Current status: valid genus in EUCNEMIDAE: PHLEGONINAE.

Type species: Phlegon buqueti Laporte, 1840, by monotypy.

Family-group name: PHLEGONINAE Muona, 1993. Stem: Phlegon-.

Phloeoborus Erichson, 1836: 54. Current status: valid genus in CURCULIONIDAE: SCOLYTINAE: HYLESININI.

Type species: Phloeoborus rudis Erichson, 1836, by subsequent designation (Hopkins 1914: 126).

Family-group name: PHLOEOBORI W.F.H. Blandford, 1893. Stem: Phloeobor-.

Phloeocharis Mannerheim, 1830: 50. Current status: valid genus in STAPHYLINIDAE: PHLOEOCHARINAE.

Type species: Phloeocharis subtilissima Mannerheim, 1830, by monotypy.

Family-group name: PHLOEOCHARINI Erichson, 1839. Stem: Phloeochar-.

Phloeonomus Heer, 1840: 184. Current status: valid genus in STAPHYLINIDAE: OMALIINAE: OMALIINI.

Type species [suggested]: Omalium pusillum Gravenhorst, 1806, by subsequent designation (C.G. Thomson 1859: 51). Note: the valid type species fixation is that of E. Desmarest (1852: 115), who selected Omalium monilicorne Gyllenhal, 1810, but that species is the type species of Xylostiba Ganglbauer, 1895; an application to the Commission is necessary to fix Omalium pusillum Gravenhorst, 1806 as the type species of Phloeonomus Heer, 1840.

Family-group name: PHLOEONOMINI Ádám, 2001. Stem: Phloeonom-.

Phloeophagus Schönherr, 1838: 1047. Current status: valid genus in CURCULIONIDAE: COSSONINAE: RHYNCOLINI: PHLOEOPHAGINA.

Type species: here fixed (under ICZN 1999a, Article 70.3.2) as Phloeophagus turbatus Schönherr, 1845, misidentified as Curculio lignarius Marsham, 1802 (sensu Boheman, 1838), by original designation by Schönherr (1838: 1047).

Family-group name: PHLOEOPHAGINA Voss, 1955. Stem: Phloeophag-.

Phloeophilus Schönherr, 1833: 156. Current status: valid genus in ANTHRIBIDAE: ANTHRIBINAE: PHLOEOPHILINI.

Type species: Phloeophilus agrestis Boheman, 1833, by original designation.

Family-group name: PHLOEOPHILIDES Lacordaire, 1865. Stem: Phloeophil-.

Phloeopora Erichson, 1837: 311. Current status: valid genus in STAPHYLINIDAE: ALEOCHARINAE: OXYPODINI: PHLOEOPORINA.

Type species: Aleochara corticalis Gravenhorst, 1802, by subsequent designation (Westwood 1838c: 19).

Family-group name: PHLOEOPORIDES C.G. Thomson, 1859. Stem: Phloeopor-.

Phloeosinus Chapuis, 1869: 37. Current status: valid genus in CURCULIONIDAE: SCOLYTINAE: PHLOEOSININI. Note: name placed on the Official List of Generic Names in Zoology (ICZN 1981a).

Type species: Hylesinus thujae Perris, 1855 [as thuyae], by subsequent designation (Hopkins 1914: 126; see ICZN 1981a). Note: name placed on the Official List of Specific Names in Zoology (ICZN 1981a).

Family-group name: PHLOEOSINIDES Nüsslin, 1912. Stem: Phloeosin-.

Phloeostichus W. Redtenbacher, 1842: 15. Current status: valid genus in PHLOEOSTICHIDAE.

Type species: Phloeostichus denticollis W. Redtenbacher, 1842, by monotypy.

Family-group name: PHLOEOSTICHINI Reitter, 1911. Stem: Phloeostich-.

Phloeotragus Schönherr, 1823: 1135. Current status: valid genus in ANTHRIBIDAE: ANTHRIBINAE: PHLOEOTRAGINI.

Type species: Anthribus heros Fabricius, 1801, by original designation.

Family-group name: PHLOEOTRAGINAE Kolbe, 1897. Stem: Phloeotrag-.

Phloeotribus Latreille, 1797: 50. Current status: valid genus in CURCULIONIDAE: SCOLYTINAE: PHLOEOTRIBINI. Note: the original spelling was Phloiotribus but Phloeotribus, introduced by Illiger (1804: 108), was ruled to be a justified emendation and Phloeotribus Latreille, 1797 was placed on the Official List of Generic Names in Zoology (ICZN 1979a, as Phloeotribus Latreille, 1796).

Type species: Bostrichus oleae Fabricius, 1792 (= Scolytus scarabaeoides Bernard, 1788), by subsequent monotypy (Latreille 1802b: 204; see ICZN 1979a). Note: Scolytus scarabaeoides Bernard, 1788 was placed on the Official List of Specific Names in Zoology (ICZN 1979a).

Family-group name: PHLOEOTRIBIDAE Chapuis, 1869. Stem: Phloeotrib-.

Phloeotrupes Erichson, 1836: 53. Current status: junior synonym of Phloeoborus Erichson, 1836 in CURCULIONIDAE: SCOLYTINAE: HYLESININI.

Type species: Phloeotrupes grandis Erichson, 1836, by subsequent designation (Hopkins 1914: 127).

Family-group name: PHLOEOTRUPIDES Lacordaire, 1865. Stem: Phloeotrup-.

Phloiophilus Stephens, 1830: 81. Current status: valid genus in PHLOIOPHILIDAE.

Type species: Phloiophilus edwardsii Stephens,1830, by subsequent designation (Stephens 1835b: 314).

Family-group name: PHLOEOPHILINI Kiesenwetter, 1863. Stem: Phloiophil-.

Phlyctaenodes Newman, 1840a: 20. Current status: valid genus in CERAMBYCIDAE: CERAMBYCINAE: PHLYCTAENODINI.

Type species: Phlyctaenodes pustulosa Newman, 1840, by monotypy.

Family-group name: PHLYCTÉNODIDES Lacordaire, 1868. Stem: Phlyctaenod-.

Phobelius Blanchard, 1842c: pl. 14. Current status: valid genus in TENEBRIONIDAE: LAGRIINAE: LAGRIINI: PHOBELIINA. Note: this name is often credited to Blanchard (1845b: 39) in the literature but it was first made available by Blanchard (1842c: pl. 14, fig. 9) when he illustrated the species Phobelius crenatus (see Bouchard et al. 2021: 297).

Type species: Phobelius crenatus Blanchard, 1842, by monotypy.

Family-group name: PHOBELIIDES F. Bates, 1890. Stem: Phobeli-.

Phoberus W.S. MacLeay, 1819: 137. Current status: valid genus in TROGIDAE: TROGINAE.

Type species: Trox horridus Fabricius, 1775, by monotypy.

Family-group name: PHOBERIDAE Gistel, 1848. Stem: Phober-.

Phobetus J.L. LeConte, 1856b: 227. Current status: valid genus in SCARABAEIDAE: MELOLONTHINAE: PHOBETUSINI.

Type species: Phobetus comatus J.L. LeConte, 1856, by monotypy.

Family-group name: PHOBETUSINI Evans & A.B.T. Smith, 2020: 66. Stem: Phobetus- (ICZN 1999a, Article 29.4).

Phodaga J.L. LeConte, 1858a: 76. Current status: valid genus in MELOIDAE: MELOINAE: EUPOMPHINI.

Type species: Phodaga alticeps J.L. LeConte, 1858, by monotypy.

Family-group name: PHODAGAE J.L. LeConte, 1862. Stem: Phodag-.

Phoenicobates Champion, 1914b: 417. Current status: valid genus in CURCULIONIDAE: MOLYTINAE: PHOENICOBATINI.

Type species: Phoenicobates vittatus Champion, 1914, by original designation.

Family-group name: PHOENICOBATINA Champion, 1914. Stem: Phoenicobat-.

Pholeuon Hampe, 1856: 463. Current status: valid genus in LEIODIDAE: CHOLEVINAE: LEPTODIRINI: PHOLEUINA. Note: the original spelling was Poleuon but it was changed to Pholeuon on page [724] of the same journal.

Type species: Pholeuon angusticolle Hampe, 1856, by monotypy.

Family-group name: PHOLEUONES Reitter, 1886. Stem: Pholeu-.

Pholidochlamys Lacordaire, 1865: 473. Current status: valid genus in BRENTIDAE: BRENTINAE: ULOCERINI: PHOLIDOCHLAMYDINA.

Type species: Pholidochlamys madagascariensis Lacordaire, 1865, by monotypy.

Family-group name: PHOLIDOCHLAMYDINAE Damoiseau, 1962. Stem: Pholidochlamyd-.

Pholidotus W.S. MacLeay, 1819: 96, 97. Current status: synonym of Casignetus W.S. MacLeay, 1819 in LUCANIDAE: LUCANINAE: LUCANINI. Note: junior homonym of Pholidotus Brisson, 1762 [Mammalia]; First Reviser (ICZN 1999a, Article 24.2.2) for Pholidotus W.S. MacLeay, 1819 versus Casignetus W.S. MacLeay, 1819 is Westwood (1834: 119).

Type species: Pholidotus lepidosus W.S. MacLeay, 1819 (= Lamprima humboldti Gyllenhal, 1817), by monotypy.

Family-group name: PHOLIDOTINI Kikuta, 1986. Stem: Pholidot-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

Pholidus Mulsant & Rey, 1856: 7. Current status: junior synonym of Euphanias Fairmaire & Laboulbène, 1856 in STAPHYLINIDAE: OXYTELINAE: EUPHANIINI. Note: junior homonym of Pholidus F. Moore, 1854 (unjustified emendation of Phodilus Geoffroy, 1830) [Aves].

Type species: Pholidus insignis Mulsant & Rey, 1856, by monotypy.

Family-group name: PHOLIDINI Acloque, 1895. Stem: Pholid-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

Phoracantha Newman, 1840a: 19. Current status: valid genus in CERAMBYCIDAE: CERAMBYCINAE: PHORACANTHINI.

Type species: Stenocorus semipunctatus Fabricius, 1775, by original designation.

Family-group name: PHORACANTHIDAE Newman, 1840. Stem: Phoracanth-.

Phoroneus Kaup, 1868b: 19. Current status: junior synonym of Passalus Fabricius, 1792 in PASSALIDAE: PASSALINAE: PASSALINI. Note: junior homonym of Phoroneus Stål, 1865 [Hemiptera].

Type species: Passalus quadricollis Eschscholtz, 1829, by subsequent designation (Hincks and Dibb 1935: 38).

Family-group name: PHORONEAE Kaup, 1871. Stem: Phorone-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

†Phoroschizus Bouchard & Bousquet, 2020a: 70. Current status: valid genus in †PHOROSCHIZIDAE. Note: new replacement name for Schizophorus Ponomarenko, 1968; Upper Jurassic (Kazakhstan).

Type species: †Schizophorus crassus Ponomarenko, 1968, by original designation (type fixation for Schizophorus Ponomarenko).

Family-group name: †PHOROSCHIZIDAE Bouchard & Bousquet, 2020a: 70. Stem: Phoroschiz-.

Phosphaenus Laporte, 1833c: 138. Current status: valid genus in LAMPYRIDAE: LAMPYRINAE: LUCIDOTINI: PHOTININA.

Type species: Lampyris hemiptera Geoffroy, 1762, by monotypy.

Family-group name: PHOSPHAENIDES E. Olivier, 1884. Stem: Phosphaen-.

Photinus Laporte, 1833c: 140. Current status: valid genus in LAMPYRIDAE: LAMPYRINAE: LUCIDOTINI: PHOTININA. Note: name placed on the Official List of Generic Names in Zoology (ICZN 2017b).

Type species: Lampyris pallens Fabricius, 1798 [as pullens], by subsequent designation (Paulian 1947: 160; see ICZN 2017b). Note: the spelling of the type species was ruled to be a justified emendation of the original spelling L. pullens by the Commission and Lampyris pallens Fabricius, 1798 was placed on the Official List of Specific Names in Zoology (ICZN 2017b).

Family-group name: PHOTINI J.L. LeConte, 1881. Stem: Photin-. Note: the spelling of the family-group name PHOTININAE Giglio-Tos, 1915 [Mantodea] was ruled to be PHOTINAINAE by the Commission (see ICZN 2017b) to remove the homonymy with J.L. LeConte’s name.

Photuris Dejean, 1833b: 103. Current status: valid genus in LAMPYRIDAE: PHOTURINAE.

Type species: Lampyris versicolor Fabricius, 1798, by subsequent designation (Barber 1951: 11).

Family-group name: PHOTURIDES Lacordaire, 1857. Stem: Photur-.

Phratora Chevrolat in Dejean, 1836a: 405. Current status: valid genus in CHRYSOMELIDAE: CHRYSOMELINAE: CHRYSOMELINI.

Type species: Chrysomela vulgatissima Linnaeus, 1758, by subsequent designation (Motschulsky 1860b: 219). Note: the type species for this genus should be Chrysomela vitellinae Linnaeus, 1758 (ICZN 1999a, Article 68.3) as it was the only taxonomic species cited by Chevrolat in Dejean (1836a: 405), but that species is currently included in the subgenus Phyllodecta W. Kirby, 1837 of the genus Phratora (e.g., Kippenberg 2010: 395); a request should be submitted to the Commission to fix Chrysomela vulgatissima Linnaeus, 1758 as the type species of the genus Phratora Chevrolat, 1836.

Family-group name: PHRATHORINES Motschulsky, 1860. Stem: Phrator-.

Phreatodytes Uéno, 1957: 252. Current status: valid genus in NOTERIDAE: NOTOMICRINAE: PHREATODYTINI.

Type species: Phreatodytes relictus Uéno, 1957, by original designation.

Family-group name: PHREATODYTIDAE Uéno, 1957. Stem: Phreatodyt-.

Phrenapates G.R. Gray in Griffith and Pidgeon, 1831: pls 50, 69. Current status: valid genus in TENEBRIONIDAE: PHRENAPATINAE: PHRENAPATINI.

Type species: Phrenapates bennettii G.R. Gray, 1831, by monotypy.

Family-group name: PHRÉPATIDES Solier, 1834. Stem: Phrenapat-.

Phrissoma Dejean, 1835: 345. Current status: valid genus in CERAMBYCIDAE: LAMIINAE: LAMIINI.

Type species: Lamia crispa Fabricius, 1777, by subsequent designation (E. Desmarest 1860: 324).

Family-group name: PHRISSOMITAE J. Thomson, 1860. Stem: Phrissomat-.

Phrixia Deyrolle, 1865: 66. Current status: valid genus in BUPRESTIDAE: BUPRESTINAE: PHRIXIINI.

Type species: Phrixia filiformis Deyrolle, 1865, by subsequent designation (Cobos 1975: 104).

Family-group name: PHRIXIINI Cobos, 1975. Stem: Phrixi-.

Phrixosoma W.F.H. Blandford, 1896: 143. Current status: valid genus in CURCULIONIDAE: SCOLYTINAE: PHRIXOSOMATINI.

Type species: Phrixosoma rude W.F.H. Blandford, 1897, by subsequent monotypy (W.F.H. Blandford 1897: 148).

Family-group name: PHRIXOSOMINI Wood, 1978. Stem: Phrixosomat-.

Phrydiuchus Gozis, 1885: 129. Current status: valid genus in CURCULIONIDAE: CEUTORHYNCHINAE: CEUTORHYNCHINI.

Type species: Ceutorhynchus topiarius Germar, 1823, by monotypy.

Family-group name: PHRYDIUCHINA Wagner, 1938. Stem: Phrydiuch-.

Phryneta Dejean, 1835: 341. Current status: valid genus in CERAMBYCIDAE: LAMIINAE: PHRYNETINI.

Type species: Lamia marmorea G.-A. Olivier, 1792, by subsequent designation (J. Thomson 1864: 71).

Family-group name: PHRYNETITAE J. Thomson, 1864. Stem: Phrynet-.

Phrynixus Pascoe, 1875: 221. Current status: valid genus in CURCULIONIDAE: MOLYTINAE: PHRYNIXINI: PHRYNIXINA.

Type species: Phrynixus terreus Pascoe, 1875, by monotypy.

Family-group name: PHRYNIXINAE Kuschel, 1964. Stem: Phrynix-.

Phrynocarenum Gebien, 1928: 106. Current status: valid genus in TENEBRIONIDAE: PIMELIINAE: PHRYNOCARENINI.

Type species: Phrynocarenum bruchianum Gebien, 1928 (= Emmallodera strangulate Fairmaire, 1905), by monotypy.

Family-group name: PHRYNOCARENINAE Gebien, 1928. Stem: Phrynocaren-.

Phtegnomus Raffray, 1890a: 104, 107. Current status: valid genus in STAPHYLINIDAE: PSELAPHINAE: EUPLECTITAE: TROGASTRINI: PHTEGNOMINA.

Type species: Phtegnomus oberthuri Raffray, 1890, by monotypy.

Family-group name: PHTEGNOMINI Park, 1951. Stem: Phtegnom-.

Phthorophloeus Rey in Eichhoff, 1883: 128. Current status: junior synonym of Phloeotribus Latreille, 1797 in CURCULIONIDAE: SCOLYTINAE: PHLOEOTRIBINI.

Type species: Phloeophthorus spinulosus Rey, 1883, by monotypy.

Family-group name: PHTHOROPHLOEIDES Nüsslin, 1912. Stem: Phthorophloe-.

Phtora Mulsant, 1854: 228. Current status: invalid senior synonym of Clamoris des Gozis, 1886 in TENEBRIONIDAE: PHRENAPATINAE: PENETINI. Note: junior homonym of Phtora Germar, 1836 [Coleoptera: TENEBRIONIDAE].

Type species: Phtora crenata Mulsant, 1854 (= Clamoris insurgens Gozis, 1886), by monotypy.

Family-group name: PHTHORINI Boddy, 1965. Stem: Phtor-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

Phycocoetes J.L. LeConte in J.L. LeConte and G.H. Horn, 1876: 189. Current status: invalid senior synonym of Thalasselephas Egorov & Korotyaev, 1976 in CURCULIONIDAE: MOLYTINAE: EMPHYASTINI. Note: junior homonym of Phycocoetes Agassiz, 1846 (unjustified emendation of Phucocoetes Jenyns, 1842) [Pisces].

Type species: Phycocoetes testaceus J.L. LeConte, 1876, by monotypy.

Family-group name: PHYCOCOETES J.L. LeConte, 1876. Stem: Phycocoet-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

Phycocus Broun, 1883: 299. Current status: valid genus in SCARABAEIDAE: APHODIINAE: PSAMMODIINI: PHYCOCINA.

Type species: Phycocus graniceps Broun, 1883, by monotypy.

Family-group name: PHYCOCHI Landin, 1960. Stem: Phycoc-.

Phycosecis Pascoe, 1875: 213. Current status: valid genus in PHYCOSECIDAE.

Type species: Phycosecis litoralis Pascoe, 1875, by present designation. Note: Crowson (1964: 313) selected Dermestes limbatus Fabricius, 1781 as the type species for this genus- group name but it is not an originally included nominal species.

Family-group name: PHYCOSECIDAE Crowson, 1952. Stem: Phycosec-.

Phylax Brullé, 1832: 209. Current status: junior synonym of Phylan Sturm, 1826 in TENEBRIONIDAE: BLAPTINAE: DENDARINI: DENDARINA. Note: unnecessary new replacement name for Phylan Sturm, 1826 (see Bouchard and Bousquet 2020a: 100, 101).

Type species: Opatrum gibbum Fabricius, 1775, by subsequent monotypy (Steven 1828: 99) (type fixation for Phylan Sturm).

Family-group name: PHYLACIDES Lacordaire, 1859. Stem: Phylac-.

Phyllarthrius Hope, 1843b: 366. Current status: valid genus in CERAMBYCIDAE: CERAMBYCINAE: PHYLLARTHRIINI.

Type species: Phyllarthrius africanus Hope, 1843, by subsequent designation (J. Thomson 1864: 269).

Family-group name: PHYLLARTHRIINI Lepesme & Breuning, 1956. Stem: Phyllarthri-.

Phyllastolus Gistel, 1856: 372. Current status: junior synonym of Trachyphloeus Germar, 1817 in CURCULIONIDAE: ENTIMINAE: CYPHICERITAE: TRACHYPHLOEINI: TRACHYPHLOEINA.

Type species: Curculio scabriculus Linnaeus, 1771 [as scabricul], by subsequent designation (Alonso-Zarazaga and Lyal 1999: 22, 183).

Family-group name: PHYLLASTOLIDAE Gistel, 1856. Stem: Phyllastol-.

Phyllecthris Dejean, 1836a: 382. Current status: valid genus in CHRYSOMELIDAE: GALERUCINAE: LUPERINI.

Type species: Galeruca dorsalis G.-A. Olivier, 1808, by monotypy.

Family-group name: PHYLLECTHRITES G.H. Horn, 1893. Stem: Phyllecthr-.

Phyllobaenus Dejean, 1833b: 113. Current status: valid genus in CLERIDAE: CLERINAE: CLERITAE: HYDNOCERINI: HYDNOCERINA.

Type species: Clerus humeralis Germar, 1823, by monotypy.

Family-group name: PHYLLOBÉNIDES Lacordaire, 1857. Stem: Phyllobaen-.

Phyllobius Germar, 1823: 447. Current status: valid genus in CURCULIONIDAE: ENTIMINAE: OTIORHYNCHITAE: PHYLLOBIINI. Note: name placed on the Official List of Generic Names in Zoology (ICZN 1981c, as Phyllobius Germar, 1824).

Type species: Curculio pyri Linnaeus, 1758, by subsequent designation (Schönherr 1826: 15; see ICZN 1981c). Note: name placed on the Official List of Specific Names in Zoology (ICZN 1981c).

Family-group name: PHYLLOBIDES Schönherr, 1826. Stem: Phyllobi-.

Phyllobrotica Chevrolat in Dejean, 1836a: 381. Current status: valid genus in CHRYSOMELIDAE: GALERUCINAE: LUPERINI.

Type species: Chrysomela quadrimaculata Linnaeus, 1758, by subsequent designation (C.G. Thomson 1859: 156).

Family-group name: PHYLLOBROTICITES Chapuis, 1875. Stem: Phyllobrotic-.

Phyllocerus Lepeletier & Audinet-Serville, 1825: 116. Current status: valid genus in EUCNEMIDAE: PHYLLOCERINAE: PHYLLOCERINI.

Type species: Phyllocerus flavipennis Lepeletier & Audinet-Serville, 1825, by monotypy.

Family-group name: PHYLLOCERIDAE Reitter, 1905. Stem: Phyllocer-.

Phyllocharis Dalman, 1824: 21. Current status: valid genus in CHRYSOMELIDAE: CHRYSOMELINAE: CHRYSOMELINI.

Type species: Chrysomela cyanipes Fabricius, 1775, by subsequent designation (Hope 1841a: 163).

Family-group name: PHYLLOCHARITES Chapuis, 1874. Stem: Phyllocharit-.

Phyllodecta W. Kirby, 1837: 216. Current status: valid subgenus of Phratora Chevrolat, 1836 in CHRYSOMELIDAE: CHRYSOMELINAE: CHRYSOMELINI.

Type species: Chrysomela vitellinae Linnaeus, 1758, by monotypy. Note: see remark about the type species of Phratora Chevrolat, 1836.

Family-group name: PHYLLODECTAE J.L. LeConte & G.H. Horn, 1883. Stem: Phyllodect-.

Phyllodinarda Wasmann, 1916b: 105. Current status: valid genus in STAPHYLINIDAE: ALEOCHARINAE: PHYLLODINARDINI.

Type species: Phyllodinarda xenocephala Wasmann, 1916, by subsequent designation (Blackwelder 1952: 307).

Family-group name: PHYLLODINARDINI Wasmann, 1916. Stem: Phyllodinard-.

Phyllomanes Gistel, 1848b: xi. Current status: junior synonym of Polydrusus Germar, 1817 in CURCULIONIDAE: ENTIMINAE: POLYDRUSITAE: POLYDRUSINI. Note: unnecessary new replacement name for Polydrosus Schönherr, 1823 (an unjustified emendation of Polydrusus Germar, 1817); junior homonym of Phyllomanes Cabanis, 1847 [Aves].

Type species: Curculio undatus Fabricius, 1781 (= Curculio tereticollis DeGeer, 1775), by subsequent designation (Schönherr 1826: 12; see ICZN 1981c) (type fixation for Polydrusus Germar).

Family-group name: PHYLLOMANISIDAE Gistel, 1848. Stem: Phylloman-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

Phyllonomeus Gistel, 1856: 372. Current status: valid genus in CURCULIONIDAE: LIXINAE: LIXINI.

Type species: Curculio iaceae Fabricius, 1775, by subsequent designation (Alonso-Zarazaga and Lyal 1999: 22, 190).

Family-group name: PHYLLONOMEIDAE Gistel, 1856. Stem: Phyllonome-.

Phyllophorus Hope, 1842c: 73. Current status: junior synonym of Tetralobus Lepeletier & Audinet- Serville, 1828 in ELATERIDAE: TETRALOBINAE: TETRALOBINI. Note: junior homonym of Phyllophorus Grube, 1840 [Echinodermata].

Type species: Elater gigas Fabricius, 1801, by monotypy.

Family-group name: PHYLLOPHORIDAE Hope, 1842. Stem: Phyllophor-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

Phylloplatypus Kato, 1998: 72. Current status: valid genus in CURCULIONIDAE: COSSONINAE: COSSONINI.

Type species: Phylloplatypus pandani Kato, 1998, by original designation.

Family-group name: PHYLLOPLATYPODINI Kato, 1998. Stem: Phylloplatypod-.

Phyllotocidium Blackburn, 1898: 21, 24. Current status: valid genus in SCARABAEIDAE: MELOLONTHINAE: PHYLLOTOCIDIINI.

Type species: Cheiragra macleayi Blackburn, 1892, by original designation.

Family-group name: PHYLLOTOCIDIINI Britton, 1957. Stem: Phyllotocidi-.

Phyllotocus Fischer, 1823b: 255. Current status: valid genus in SCARABAEIDAE: SERICOIDINAE: PHYLLOTOCINI.

Type species: Phyllotocus macleayi Fischer, 1823, by monotypy.

Family-group name: PHYLLOTOCIDAE Burmeister, 1855. Stem: Phyllotoc-.

Phyllotrox Schönherr, 1843: 189. Current status: valid genus in CURCULIONIDAE: CURCULIONINAE: DERELOMINI: PHYLLOTROGINA.

Type species: Phyllotrox semirufus Boheman, 1843, by original designation (as rufus, lapsus).

Family-group name: PHYLLOTROGINA N.M. Franz, 2006. Stem: Phyllotrog-.

Phyllurga Gistel, 1848b: xi, xiii. Current status: junior synonym of Euchlora W.S. MacLeay, 1819 in SCARABAEIDAE: RUTELINAE: ANOMALINI: ANOMALINA. Note: unnecessary new replacement name for Euchlora W.S. MacLeay, 1819.

Type species: Melolontha viridis Fabricius, 1775, by subsequent designation (Hope 1837: 71) (type fixation for Euchlora W.S. MacLeay).

Family-group name: PHYLLURGAEIDAE Gistel, 1848. Stem: Phyllurg-.

Phymaphora Newman, 1838: 389. Current status: valid genus in ENDOMYCHIDAE: LEIESTINAE.

Type species: Phymaphora pulchella Newman, 1838, by monotypy.

Family-group name: PHYMAPHORINA Jakobson, 1915. Stem: Phymaphor-.

Phymasterna Laporte, 1840b: 473. Current status: valid genus in CERAMBYCIDAE: LAMIINAE: PHYMASTERNINI. Note: the senior homonym Phymasterna Dejean, 1835 is available but in a different sense (see Bousquet and Bouchard 2013a: 88); we recommend that an application be submitted to the Commission to suppress Dejean’s name for the Principles of Priority and Homonymy.

Type species: Phymasterna lacteoguttata Laporte, 1840, by monotypy.

Family-group name: PHYMASTERNINI Téocchi, 1989. Stem: Phymastern-.

Phymatolabus Jekel, 1860: 197. Current status: valid genus in ATTELABIDAE: ATTELABINAE: ATTELABINI: PHYMATOLABINA.

Type species: Attelabus dentipennis Gyllenhal, 1839, by original designation.

Family-group name: PHYMATOLABINA Voss, 1925. Stem: Phymatolab-.

Phymatopsinus Voss, 1925: 205. Current status: valid genus in ATTELABIDAE: ATTELABINAE: ATTELABINI: PHYMATOPSININA.

Type species: Attelabus pustula Ancey, 1881, by original designation.

Family-group name: PHYMATOPSININA Legalov, 2003. Stem: Phymatopsin-.

Physea Brullé, 1834: 473. Current status: valid genus in CARABIDAE: PAUSSINAE: OZAENINI. Note: new replacement name for Trachelizus Solier, 1834.

Type species: Trachelizus rufus Solier, 1834 (= Ozaena testudinea Klug, 1834), by monotypy (type fixation for Trachelizus Solier).

Family-group name: PHYSEITAE Jeannel, 1946. Stem: Physe-.

Physocrotaphus Parry, 1849: 180. Current status: valid genus in CARABIDAE: HARPALINAE: PHYSOCROTAPHINI.

Type species: Physocrotaphus ceylonicus Parry, 1849, by monotypy.

Family-group name: PHYSOCROTAPHIDES Chaudoir, 1863. Stem: Physocrotaph-.

Physodactylus Fischer, 1823c: 302. Current status: valid genus in ELATERIDAE: PHYSODACTYLINAE.

Type species: Physodactylus henningii Fischer, 1823, by monotypy.

Family-group name: PHYSODACTYLIDES Lacordaire, 1857. Stem: Physodactyl-.

Physodera Eschscholtz, 1829c: 8. Current status: valid genus in CARABIDAE: HARPALINAE: LEBIINI: PHYSODERINA.

Type species: Physodera dejeanii Eschscholtz, 1829, by monotypy.

Family-group name: PHYSODÉRIDES Chaudoir, 1877. Stem: Physoder-. Note: senior homonym of PHYSODERINAE N.C.E. Miller, 1954 (type genus Physoderes Westwood, 1846) in Hemiptera.

Physogaster Lacordaire, 1830a: 276. Current status: valid genus in TENEBRIONIDAE: PIMELIINAE: PHYSOGASTERINI.

Type species: Physogaster mendocina Lacordaire, 1830, by monotypy.

Family-group name: PHYSOGASTÉRIDES Lacordaire, 1859. Stem: Physogaster-.

Physognathus Solier, 1849: 303. Current status: invalid senior synonym of Solierius Bernhauer, 1921 in STAPHYLINIDAE: SOLIERIINAE. Note: junior homonym of Physognathus Agassiz, 1846 (unjustified emendation of Physignathus Cuvier, 1829) [Reptilia].

Type species: Physognathus obscurus Solier, 1849, by monotypy.

Family-group name: FISOGNATITOS Solier, 1849. Stem: Physognath-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

Physonota Boheman, 1854: 190. Current status: valid genus in CHRYSOMELIDAE: CASSIDINAE: ISCHYROSONYCHINI. Note: this genus was first made available by Chevrolat in Dejean (1836a: 374) in another sense, with Cassida fuscata Klug, 1829 as type species by monotypy; to promote nomenclatural stability a request should be submitted to the Commission to reject the name Physonota Chevrolat, 1836 for the Principles of Priority and Homonymy (see Bousquet and Bouchard 2013a: 129).

Type species: Physonota alutacea Boheman, 1854, by subsequent designation (Hincks 1952: 336).

Family-group name: PHYSONOTITAE Spaeth, 1942. Stem: Physonot-.

Physorhinus Eschscholtz in Laporte, 1838: table. Current status: valid genus in ELATERIDAE: ELATERINAE: PHYSORHININI.

Type species: Elater erythrocephalus Fabricius, 1801, by subsequent designation (Lacordaire 1857: 176). Note: no species were originally included in this genus by Eschscholtz in Laporte (1838); the first ones were the three species described or mentioned by Germar (1840a: 245; 1840b: 439, 440).

Family-group name: PHYSORHINITES Candèze, 1859. Stem: Physorhin-.

Phytobaenus R.F. Sahlberg, 1834: 9. Current status: valid genus in ADERIDAE.

Type species: Phytobaenus amabilis R.F. Sahlberg, 1834, by monotypy.

Family-group name: PHYTOBAENINI Báguena Corella, 1948. Stem: Phytobaen-.

Phytobius Schönherr, 1833: 20. Current status: valid genus in CURCULIONIDAE: CEUTORHYNCHINAE: PHYTOBIINI. Note: new replacement name for Hydaticus Schönherr, 1825, which was placed on the Official Index of Rejected and Invalid Generic Names in Zoology and Phytobius Schönherr, 1833 was placed on the Official List of Generic Names in Zoology (ICZN 2001); the name Phytobius Dejean, 1835 had previously been placed on the Official List of Generic Names in Zoology (ICZN 1989b) but this entry was deleted from the Official List in the more recent ruling (ICZN 2001).

Type species: Rhynchaenus myriophylli Gyllenhal, 1813 (= Curculio leucogaster Marsham, 1802), by original designation (see ICZN 2001) (type fixation for Hydaticus Schönherr). Note: Curculio leucogaster Marsham, 1802 was placed on the Official List of Specific Names in Zoology (ICZN 2001).

Family-group name: PHYTOBIIDAE Gistel, 1848. Stem: Phytobi-.

Phytoecia Dejean, 1835: 351. Current status: valid genus in CERAMBYCIDAE: LAMIINAE: SAPERDINI.

Type species: Cerambyx cylindricus Linnaeus, 1758, by subsequent designation (C.G. Thomson 1859: 153).

Family-group name: PHYTOECIAIRES Mulsant, 1839. Stem: Phytoeci-.

Phytonomus Schönherr, 1823: 1143. Current status: junior synonym of Hypera Germar, 1817 in CURCULIONIDAE: HYPERINAE: HYPERINI: CEPURINA. Note: unnecessary new replacement name for Hypera Germar, 1817.

Type species: Curculio nigrirostris Fabricius, 1775, by subsequent designation (Samouelle 1819: 205) (type fixation for Hypera Germar).

Family-group name: PHYTONOMIDAE Gistel, 1848. Stem: Phytonom-. Note: a number of authors (see Casalini and Colonnelli 2023 and references therein) have treated PHYTONOMINAE and PHYTONOMINI as valid instead of HYPERINAE and HYPERINI; we prefer to conserve usage of HYPERINAE and HYPERINI as valid (ICZN 1999a, Article 40.2, see Bouchard et al. 2011: 619).

Phytoscaphus Schönherr, 1826: 210. Current status: valid genus in CURCULIONIDAE: ENTIMINAE: CYPHICERITAE: CYPHICERINI: PHYTOSCAPHINA.

Type species: Phytoscaphus lixabundus Schönherr, 1826 (= Curculio lanatus Fabricius, 1801), by original designation.

Family-group name: PHYTOSCAPHIDES Lacordaire, 1863. Stem: Phytoscaph-.

Phytosus Curtis, 1838a: pl. 718. Current status: valid genus in STAPHYLINIDAE: ALEOCHARINAE: PHYTOSINI.

Type species: Phytosus spinifer Curtis, 1838, by original designation.

Family-group name: PHYTOSIDES C.G. Thomson, 1867. Stem: Phytos-.

Phytotera Gistel, 1856: 366. Current status: junior synonym of Trachys Fabricius, 1801 in BUPRESTIDAE: AGRILINAE: TRACHEINI: TRACHEINA.

Type species: Buprestis minuta Linnaeus, 1758, by subsequent designation (Bellamy 2003: 11). Note: name placed on the Official List of Specific Names in Zoology (ICZN 2009a).

Family-group name: PHYTOTERADAE Gistel, 1856. Stem: Phytoter-.

Phyxelis Schönherr, 1842b: 122. Current status: valid genus in CURCULIONIDAE: ENTIMINAE: ENTIMITAE: BYRSOPAGINI: PHYXELIDINA.

Type species: Barynotus rigidus Say, 1832, by original designation.

Family-group name: PHYXELES G.H. Horn, 1876. Stem: Phyxelid-.

Piazomias Schönherr, 1840a: 936. Current status: valid genus in CURCULIONIDAE: ENTIMINAE: POLYDRUSITAE: TANYMECINI: PIAZOMIINA.

Type species: Piazomias virescens Boheman, 1840, by original designation.

Family-group name: PIAZOMIINI Reitter, 1913. Stem: Piazomi-.

Piazorhinus Schönherr, 1835: 471. Current status: valid genus in CURCULIONIDAE: CURCULIONINAE: PIAZORHININI.

Type species: Attelabus scutellaris Say, 1826, by original designation.

Family-group name: PIAZORHINIDES Lacordaire, 1863. Stem: Piazorhin-.

Piazurus Schönherr, 1825: 586. Current status: valid genus in CURCULIONIDAE: CONODERINAE: PIAZURINI.

Type species: Poecilma stipitosum Germar, 1823, by original designation.

Family-group name: PIAZURIDES Lacordaire, 1865. Stem: Piazur-.

Picrotus Sharp, 1886: 394. Current status: valid genus in CRYPTOPHAGIDAE: CRYPTOPHAGINAE: PICROTINI.

Type species: Picrotus thoracicus Sharp, 1886, by monotypy.

Family-group name: PICROTINAE Sen Gupta & Crowson, 1971. Stem: Picrot-.

Pidonia Mulsant, 1863: 570. Current status: valid genus in CERAMBYCIDAE: LEPTURINAE: PIDONIINI.

Type species: Leptura lurida Fabricius, 1793, by subsequent designation (Swaine and Hopping 1928: 12).

Family-group name: PIDONIINI Zamoroka, 2022: 308. Stem: Pidoni-.

Piesarthrius Hope, 1834a: 107. Current status: valid genus in CERAMBYCIDAE: CERAMBYCINAE: PIESARTHRIINI.

Type species: Piesarthrius marginellus Hope, 1834, by original designation.

Family-group name: PIESARTHRINI McKeown, 1947. Stem: Piesarthri-.

Piesocorynus Dejean, 1834: 235. Current status: valid genus in ANTHRIBIDAE: ANTHRIBINAE: PIESOCORYNINI.

Type species: Euparius dispar Gyllenhal, 1833, by monotypy.

Family-group name: PIESOCORYNINI B.D. Valentine, 1960. Stem: Piesocoryn-.

Piestus Gravenhorst, 1806: 223. Current status: valid genus in STAPHYLINIDAE: PIESTINAE.

Type species: Piestus sulcatus Gravenhorst, 1806, by monotypy.

Family-group name: PIESTINI Erichson, 1839. Stem: Piest-.

Piezocera Audinet-Serville, 1834b: 92. Current status: valid genus in CERAMBYCIDAE: CERAMBYCINAE: PIEZOCERINI: PIEZOCERINA.

Type species: Piezocera bivittata Audinet-Serville, 1834, by monotypy.

Family-group name: PIÉZOCÉRIDES Lacordaire, 1868. Stem: Piezocer-.

Piezophyllus Hope, 1842c: 76. Current status: valid genus in ELATERIDAE: TETRALOBINAE: PIEZOPHYLLINI.

Type species: Tetralobus robustus Hope, 1842 (= Tetralobus macrocerus Laporte, 1838), by original designation.

Family-group name: PIEZOPHYLLINI Laurent, 1967. Stem: Piezophyll-.

Piezotrachelus Schönherr, 1839: 365. Current status: valid genus in BRENTIDAE: APIONINAE: APIONITAE: PIEZOTRACHELINI.

Type species: Piezotrachelus germarii Gyllenhal, 1839, by original designation.

Family-group name: PIEZOTRACHELINI Voss, 1959. Stem: Piezotrachel-.

Pilinurgus Burmeister, 1842: 658. Current status: valid genus in SCARABAEIDAE: CETONIINAE: CREMASTOCHEILINI: PILINURGINA.

Type species: Cremastocheilus hirtus Gory & Percheron, 1833, by monotypy.

Family-group name: PILINURGINA Krikken, 1984. Stem: Pilinurg-.

Pilipalpus Fairmaire, 1876: 384. Current status: valid genus in PYROCHROIDAE: PILIPALPINAE.

Type species: Pilipalpus dasytoides Fairmaire, 1876, by monotypy.

Family-group name: PILIPALPINI M. Abdullah, 1964. Stem: Pilipalp-.

Pilolabus Jekel, 1860: 196. Current status: valid genus in ATTELABIDAE: ATTELABINAE: PILOLABINI.

Type species: Attelabus klugii Gyllenhal, 1839, by original designation.

Family-group name: PILOLABINI Voss, 1925. Stem: Pilolab-.

Pimelia Fabricius, 1775: 251. Current status: valid genus in TENEBRIONIDAE: PIMELIINAE: PIMELIINI: PIMELIINA.

Type species: Pimelia angulata Fabricius, 1775, by subsequent designation (Hope 1841a: 118).

Family-group name: PIMELIARIAE Latreille, 1802. Stem: Pimeli-.

Pimelopus Erichson, 1842: 159. Current status: valid genus in SCARABAEIDAE: DYNASTINAE: PENTODONTINI: PENTODONTINA.

Type species: Pimelopus porcellus Erichson, 1842, by monotypy.

Family-group name: PIMELOPODEA Burmeister, 1847. Stem: Pimelopod-.

Pimidia Rafinesque, 1815: 113. Current status: junior synonym of Pimelia Fabricius, 1775 in TENEBRIONIDAE: PIMELIINAE: PIMELIINI: PIMELIINA. Note: unnecessary new replacement name for Pimelia Fabricius, 1775.

Type species: Pimelia angulata Fabricius, 1775, by subsequent designation (Hope 1841a: 118) (type fixation for Pimelia Fabricius).

Family-group name: PIMIDINIA Rafinesque, 1815. Stem: Pimidi-.

Pinodytes G.H. Horn, 1881: 248. Current status: junior synonym of Catopocerus Motschulsky, 1869 in LEIODIDAE: CATOPOCERINAE: CATOPOCERINI.

Type species: Catops cryptophagoides Mannerheim, 1852, by monotypy.

Family-group name: PINODYTINI G.H. Horn, 1881. Stem: Pinodyt-.

Pinophilus Gravenhorst, 1802: 201. Current status: valid genus in STAPHYLINIDAE: PAEDERINAE: PINOPHILINI: PINOPHILINA.

Type species: Pinophilus latipes Gravenhorst, 1802, by monotypy.

Family-group name: PINOPHILINIFORMES Nordmann, 1837. Stem: Pinophil-.

Pinotus Erichson, 1847c: 108. Current status: junior synonym of Dichotomius Hope, 1838 in SCARABAEIDAE: SCARABAEINAE: DICHOTOMIINI.

Type species: Pinotus talaus Erichson, 1847, by present designation.

Family-group name: PINOTINAE Kolbe, 1905. Stem: Pinot-.

Piochardia Heyden, 1870: 75. Current status: valid genus in STAPHYLINIDAE: ALEOCHARINAE: ALEOCHARINI: ALEOCHARINA.

Type species: Piochardia lepismiformis Heyden, 1870, by monotypy.

Family-group name: PIOCHARDIAE Fenyes, 1919. Stem: Piochardi-.

Pisenus Casey, 1900: 167, 171. Current status: valid genus in TETRATOMIDAE: PISENINAE.

Type species: Cryptophagus humeralis W. Kirby, 1837, by subsequent designation (Nikitsky 1989: 27).

Family-group name: PISENINI Miyatake, 1960. Stem: Pisen-.

Pissodes Germar, 1817b: 340. Current status: valid genus in CURCULIONIDAE: MOLYTINAE: PISSODINI: PISSODINA.

Type species: Curculio pini Linnaeus, 1758, by subsequent designation (Schönherr 1825: 582).

Family-group name: PISSODISIDAE Gistel, 1848. Stem: Pissod-.

Pithocles J. Thomson, 1864: 291. Current status: junior synonym of Derobrachus Audinet-Serville, 1832 in CERAMBYCIDAE: PRIONINAE: PRIONINI.

Type species: Derobrachus procerus J. Thomson, 1860, by original designation.

Family-group name: PITHOCLITAE J. Thomson, 1864. Stem: Pithocl-.

Pityobius J.L. LeConte, 1853f: 428. Current status: valid genus in ELATERIDAE: PITYOBIINAE: PITYOBIINI.

Type species: Pityobius anguinus J.L. LeConte, 1853, by monotypy.

Family-group name: PITYOBINI Hyslop, 1917. Stem: Pityobi-.

Pityogenes Bedel, 1888: 397. Current status: valid genus in CURCULIONIDAE: SCOLYTINAE: IPINI: IPINA.

Type species: Dermestes chalcographus Linnaeus, 1761 [as calcographus], by original designation. Note: the original spelling of the type species is calcographus but the incorrect subsequent spelling chalcographus is in prevailing usage, attributed to the publication of the original spelling, and therefore deemed to be the correct original spelling (ICZN 1999a, Article 33.3.1).

Family-group name: PITYOGENINA Balachowsky, 1949. Stem: Pityogen-.

Pityophagus Shuckard, 1839: 171. Current status: valid genus in NITIDULIDAE: CRYPTARCHINAE: CRYPTARCHINI.

Type species: Dermestes ferrugineus Linnaeus, 1761, by monotypy.

Family-group name: PITYOPHAGINI Schilsky, 1888. Stem: Pityophag-.

Pityophthorus Eichhoff, 1864: 39, 45. Current status: valid genus in CURCULIONIDAE: SCOLYTINAE: CORTHYLINI: PITYOPHTHORINA.

Type species: Bostrichus lichtensteinii Ratzeburg, 1837, by subsequent designation (Hopkins 1914: 127).

Family-group name: PITYOPHTHORIDAE Eichhoff, 1878. Stem: Pityophthor-.

Placonycha G.H. Horn, 1880a: 111. Current status: junior synonym of Eubrianax Kiesenwetter, 1874 in PSEPHENIDAE: EUBRIANACINAE.

Type species: Dicranopselaphus edwardsii J.L. LeConte, 1874, by monotypy.

Family-group name: PLACONYCHINI G.H. Horn, 1880. Stem: Placonych-.

Placusa Erichson, 1837: 370. Current status: valid genus in STAPHYLINIDAE: ALEOCHARINAE: PLACUSINI.

Type species: Aleochara pumilio Gravenhorst, 1802, by monotypy.

Family-group name: PLACUSATES Mulsant & Rey, 1871. Stem: Placus-.

Plagiarthrina Keys, 1920: 131. Current status: junior synonym of Philhygra Mulsant & Rey, 1873 in STAPHYLINIDAE: ALEOCHARINAE: ATHETINI: ATHETINA.

Type species: Metaxya fordhamiana Keys, 1920 (= Aleochara terminalis Gravenhorst, 1806), by monotypy.

Family-group name: PLAGIARTHRINI Cameron, 1926. Stem: Plagiarthrin-.

Planetes W.S. MacLeay, 1825: 28. Current status: valid genus in CARABIDAE: HARPALINAE: GALERITINI: PLANETINA.

Type species: Planetes bimaculatus W.S. MacLeay, 1825, by monotypy.

Family-group name: PLANETINI Jedlička, 1941. Stem: Planet-.

Planeustomus Jacquelin du Val, 1857b: 58. Current status: valid genus in STAPHYLINIDAE: OXYTELINAE: PLANEUSTOMINI.

Type species: Acrognathus palpalis Erichson, 1839, by monotypy.

Family-group name: PLANEUSTOMITES Jacquelin du Val, 1857. Stem: Planeustom-.

Plastocerus Schaum, 1852: 49. Current status: valid genus in ELATERIDAE: DENDROMETRINAE: PLASTOCERINI.

Type species: Callirhipis angulosus Germar, 1844, by monotypy.

Family-group name: PLASTOCERIDAE Crowson, 1972. Stem: Plastocer-.

Platamodes Ménétriés, 1849b: 233. Current status: valid genus in TENEBRIONIDAE: PIMELIINAE: STENOSINI: PLATAMODINA.

Type species: Platamodes dentipes Ménétriés, 1849, by monotypy.

Family-group name: PLATAMODINA Reitter, 1900. Stem: Platamod-.

Platandria Casey, 1893: 345. Current status: valid genus in STAPHYLINIDAE: ALEOCHARINAE: HOPLANDRIINI: PLATANDRIINA.

Type species: Platandria mormonica Casey, 1893, by monotypy.

Family-group name: PLATANDRIINA Hanley, 2002. Stem: Platandri-.

Platerodrilus Pic, 1921: 13. Current status: valid genus in LYCIDAE: PLATERODRILINAE.

Type species: Platerodrilus sinuatus Pic, 1921, by subsequent designation (N.D. Riley 1923: 124).

Family-group name: PLATERODRILINI Kazantsev, 2004. Stem: Platerodril-.

Plateros Bourgeois, 1879: xix. Current status: valid genus in LYCIDAE: LYCINAE: PLATERODINI.

Type species: Eros brasiliensis P.H. Lucas, 1857, by subsequent designation (Zaragoza Caballero 1999: 2).

Family-group name: PLATERODINAE Kleine, 1929. Stem: Platerod-.

Plateumaris C.G. Thomson, 1859: 154. Current status: valid genus in CHRYSOMELIDAE: DONACIINAE: PLATEUMARINI.

Type species: Donacia nigra Fabricius, 1792 (= Prionus bracatus Scopoli, 1772), by original designation.

Family-group name: PLATEUMARINI Böving, 1922. Stem: Plateumar-.

Platidiolus Chaudoir, 1878b: 77. Current status: valid genus in CARABIDAE: PATROBINAE: PATROBINI: PLATIDIOLINA.

Type species: Platidiolus rufus Chaudoir, 1878, by monotypy.

Family-group name: PLATIDIOLINI Zamotajlov & Lafer, 2001. Stem: Platidiol-.

Platisus Erichson, 1842: 216. Current status: valid genus in CUCUJIDAE: PLATISINAE.

Type species: Platisus obscurus Erichson, 1842, by monotypy.

Family-group name: PLATISINAE Jin, Zwick, Ślipiński, Marris, Thomas & H. Pang, 2020a: 260. Stem: Platis-.

Platyarcha Kirejtshuk, 1987: 78. Current status: valid genus in NITIDULIDAE: CRYPTARCHINAE: PLATYARCHINI.

Type species: Platyarcha pacta Kirejtshuk, 1987, by original designation.

Family-group name: PLATYARCHINI Kirejtshuk, 1998. Stem: Platyarch-.

Platyarthron Guérin-Méneville, 1844a: 230. Current status: valid genus in CERAMBYCIDAE: CERAMBYCINAE: PLATYARTHRINI.

Type species: Platyarthron bilineatum Guérin-Méneville, 1844, by monotypy.

Family-group name: PLATYARTHRINAE H.W. Bates, 1870. Stem: Platyarthr-.

Platyauchenia Sturm, 1843: 358. Current status: valid genus in CHRYSOMELIDAE: CASSIDINAE: ALURNINI.

Type species: Platyauchenia limbata Sturm, 1843 (= Cassida latreillii Laporte, 1840), by monotypy.

Family-group name: PLATYAUCHENIITAE Spaeth, 1929. Stem: Platyaucheni-.

Platycephala Montrouzier, 1861b: 268. Current status: invalid senior synonym of Aralius Kuschel, 1990 in BELIDAE: OXYCORYNINAE: AGLYCYDERITAE: AGLYCYDERINI. Note: junior homonym of Platycephala Fallén, 1820 [Diptera].

Type species: Platycephala olivieri Montrouzier, 1861, by monotypy.

Family-group name: PLATYCEPHALITAE Paulian, 1944. Stem: Platycephal-. Note: family-group name permanently invalid (ICZN 1999a, Article 39); junior homonym of PLATYCEPHALIDAE Swainson, 1839 (type genus Platycephalus Bloch, 1795) in Pisces.

Platyceroides Benesh, 1946: 175. Current status: valid genus in LUCANIDAE: LUCANINAE: PLATYCEROIDINI.

Type species: Platycerus agassii J.L. LeConte, 1861, by original designation.

Family-group name: PLATYCEROIDINI Paulsen & Hawks, 2008. Stem: Platyceroid-.

Platycerus Geoffroy, 1762: 59. Current status: valid genus in LUCANIDAE: LUCANINAE: PLATYCERINI. Note: name placed on the Official List of Generic Names in Zoology (ICZN 1994a).

Type species: Scarabaeus caraboides Linnaeus, 1758, by subsequent designation (Latreille 1810: 429; see ICZN 1994a). Note: name placed on the Official List of Specific Names in Zoology (ICZN 1994a).

Family-group name: PLATYCÉRAIRES Mulsant, 1842. Stem: Platycer-.

Platychile W.S. MacLeay, 1825: 9. Current status: valid genus in CICINDELIDAE: MANTICORINI.

Type species: Manticora pallida Fabricius, 1801, by monotypy.

Family-group name: PLATYCHILIDAE W. Horn, 1893. Stem: Platychil-.

Platycholeus G.H. Horn, 1881: 254. Current status: valid genus in LEIODIDAE: CHOLEVINAE: LEPTODIRINI: PLATYCHOLEINA.

Type species: Ptomaphagus leptinoides Crotch, 1874, by monotypy.

Family-group name: PLATYCHOLEI G.H. Horn, 1881. Stem: Platychole-.

Platycnemus Nordmann, 1837: 6, 135. Current status: junior synonym of Haematodes Laporte, 1835 in STAPHYLINIDAE: STAPHYLININAE: STAPHYLININI: XANTHOPYGINA. Note: name placed on the Official List of Generic Names in Zoology (ICZN 1996c).

Type species: Platycnemus lateritius Nordmann, 1837, by monotypy (see ICZN 1996c). Note: name placed on the Official List of Specific Names in Zoology (ICZN 1996c).

Family-group name: PLATYCNEMIDIFORMES Nordmann, 1837. Stem: Platycnem-.

Platycoelia Dejean, 1833b: 154. Current status: valid genus in SCARABAEIDAE: RUTELINAE: ANOPLOGNATHINI: PLATYCOELIINA.

Type species: Melolontha flavostriata Latreille, 1813, by monotypy.

Family-group name: PLATYCOELIIDAE Burmeister, 1844. Stem: Platycoeli-.

Platycorynus Chevrolat in Dejean, 1836a: 413. Current status: valid genus in CHRYSOMELIDAE: EUMOLPINAE: EUMOLPINI.

Type species: Eumolpus compressicornis Fabricius, 1801, by subsequent designation (Chapuis 1874: 339).

Family-group name: PLATYCORYNI L.N. Medvedev & Rybakova, 1985: 118. Stem: Platycoryn-. Note: this name was overlooked in Bouchard et al. (2011) and Bouchard and Bousquet (2020a); this family-group name was proposed as a new replacement name for CORYNODINI T.A. Marshall, 1865 because of the synonymy and invalidity of its type genus; however, a family-group name is not to be replaced on that account alone (ICZN 1999a, Article 40.1) unless the substitute name was proposed before 1961 and is in prevailing usage (ICZN 1999a, Article 40.2), which is not the case here.

Platycrepidius Candèze, 1859: 159. Current status: valid genus in ELATERIDAE: AGRYPNINAE: PLATYCREPIDIINI. Note: name first proposed as a synonym of Eudactylus Sallé, 1855 and subsequently treated as valid (e.g., Fleutiaux 1934: 181) before 1961 (see ICZN 1999a, Article 11.6).

Type species: Eudactylus wapleri Sallé, 1855, by subsequent designation (Hyslop 1921: 665).

Family-group name: PLATYCREPIDIINI C. Costa & Casari-Chen, 1993. Stem: Platycrepidi-.

Platydascillus Everts, 1909: 6. Current status: valid genus in BYTURIDAE: PLATYDASCILLINAE.

Type species: Platydascillus sumatranus Everts, 1909, by monotypy.

Family-group name: PLATYDASCILLIDAE Pic, 1914. Stem: Platydascill-.

Platydema Laporte & Brullé, 1831: 332, 350. Current status: valid genus in TENEBRIONIDAE: DIAPERINAE: DIAPERINI: DIAPERINA.

Type species: Diaperis violacea Fabricius, 1790, by subsequent designation (Westwood 1838c: 32).

Family-group name: PLATYDEMINAE Reitter, 1917. Stem: Platydem-.

Platygenia W.S. MacLeay, 1819: 151. Current status: valid genus in SCARABAEIDAE: CETONIINAE: PLATYGENIINI.

Type species: Platygenia zairica W.S. MacLeay, 1819 (= Trichius barbatus Afzelius, 1817), by monotypy.

Family-group name: PLATYGENIINI Krikken, 1984. Stem: Platygeni-.

Platygnathus Audinet-Serville, 1832: 150. Current status: valid genus in CERAMBYCIDAE: PRIONINAE: MACROTOMINI: PLATYGNATHINA.

Type species: Prionus octangularis G.-A. Olivier, 1795, by subsequent designation (Laporte 1840b: 404).

Family-group name: PLATYGNATHINA Gilmour, 1954. Stem: Platygnath-.

Platynaspis L. Redtenbacher, 1843: 11, 15. Current status: valid genus in COCCINELLIDAE: COCCINELLINAE: PLATYNASPINI.

Type species: Coccinella bisbipustulata Fabricius, 1792 (= Coccinella luteorubra Goeze, 1777), by monotypy. Note: Goeze’s (1777) work is not consistently binominal (see ICZN 2021a) and therefore an application to the Commission is necessary to retain Coccinella luteorubra Goeze, 1777 as the valid name for the type species.

Family-group name: PLATYNASPIAIRES Mulsant, 1846. Stem: Platynasp-.

Platynectes Régimbart, 1879: 462. Current status: valid genus in DYTISCIDAE: AGABINAE: PLATYNECTINI. Note: name placed on the Official List of Generic Names in Zoology (ICZN 1995a).

Type species: Agabus decemnotatus Aubé, 1838, by subsequent designation (Guignot 1946: 117; see ICZN 1995a). Note: name placed on the Official List of Specific Names in Zoology (ICZN 1995a).

Family-group name: PLATYNECTINI Toussaint & Balke in Toussaint et al., 2017: 504. Stem: Platynect-. Note: family-group name used as valid in recent publications but it is not an available name since it is not accompanied by a description or definition that states in words the characters that are purported to differentiate the taxon (ICZN 1999a, Article 13.1); in order to promote stability, we believe that a request should be submitted to the Commission to accept this family-group name as available from its first date of publication.

Platynoptera Chevrolat, 1834: [18] 1. Current status: valid genus in CLERIDAE: KORYNETINAE.

Type species: Platynoptera lyciformis Chevrolat, 1834, by monotypy.

Family-group name: PLATYNOPTÉROÏDES Spinola, 1844. Stem: Platynopter-.

Platynotus Fabricius, 1801a: 138. Current status: valid genus in TENEBRIONIDAE: BLAPTINAE: PLATYNOTINI: PLATYNOTINA.

Type species: Blaps excavata Fabricius, 1775, by subsequent designation (Hope 1841a: 110).

Family-group name: PLATYNOTAIRES Mulsant & Rey, 1853. Stem: Platynot-.

Platynus Bonelli, 1810: Tab. Syn. Current status: valid genus in CARABIDAE: HARPALINAE: PLATYNINI.

Type species: Carabus angusticollis Fabricius, 1801 (= Carabus assimilis Paykull, 1790), by subsequent monotypy (Germar 1817a: 303). Note: Platynus complanatus Dejean, 1831 is sometimes listed as the type species of this genus (e.g., J. Schmidt 2017: 666) but is not an originally included nominal species in Germar (1817a) and thus cannot be the type species of Platynus Bonelli, 1810 (ICZN 1999a, Article 67.2.2).

Family-group name: PLATYNII Bonelli, 1810. Stem: Platyn-.

Platyomus C.R. Sahlberg, 1823a: 29. Current status: valid genus in CURCULIONIDAE: ENTIMINAE: POLYDRUSITAE: NAUPACTINI: PLATYOMINA. Note: we consider Platyomus used by Schönherr (1823: 1140) a subsequent usage of Platyomus C.R. Sahlberg, 1823.

Type species: Platyomus nodipennis C.R. Sahlberg, 1823, by monotypy.

Family-group name: PLATYOMINA Champion, 1911. Stem: Platyom-.

Platyope Fischer, 1820: pl. 15. Current status: valid genus in TENEBRIONIDAE: PIMELIINAE: PIMELIINI: PIMELIINA.

Type species: Tenebrio leucographus Pallas, 1773, by subsequent designation (Jacquelin du Val 1860b: 261). Note: Pallas (1773) used two different spellings for this species, Tenebrio leucographus (p. 474) and Tenebrio leucogrammus (p. 719); subsequently, Pallas (1781: 54) used T. leucographus as the valid name of the species and therefore acted as First Reviser (ICZN 1999a, Article 24.2.4).

Family-group name: PLATYOPIDAE Semenov, 1893. Stem: Platyop-.

Platypicerus Nunberg, 1953: 46. Current status: junior synonym of Mitosoma Chapuis, 1865 in CURCULIONIDAE: PLATYPODINAE: TESSEROCERINI: TESSEROCERINA.

Type species: Platypicerus hamatus Nunberg, 1953, by monotypy.

Family-group name: PLATYPICERINAE Nunberg, 1953. Stem: Platypicer-.

Platyprosopus Mannerheim, 1830: 36. Current status: valid genus in STAPHYLINIDAE: STAPHYLININAE: PLATYPROSOPINI.

Type species: Platyprosopus elongatus Mannerheim, 1830, by monotypy.

Family-group name: PLATYPROSOPARIA Lynch Arribálzaga, 1884. Stem: Platyprosop-.

Platypsyllus Ritsema, 1869: [23]. Current status: valid genus in LEIODIDAE: PLATYPSYLLINAE. Note: this taxon was also made proposed subsequently the same year by Westwood (1869b: 118).

Type species: Platypsyllus castoris Ritsema, 1869, by monotypy.

Family-group name: PLATYPSYLLIDAE Ritsema, 1869. Stem: Platypsyll-.

Platypus Herbst, 1793: 128. Current status: valid genus in CURCULIONIDAE: PLATYPODINAE: PLATYPODINI.

Type species: Bostrichus cylindrus Fabricius, 1792, by monotypy.

Family-group name: PLATYPODIDAE Shuckard, 1839. Stem: Platypod-.

Platyrhinus Clairville, 1798: 56, 112. Current status: valid genus in ANTHRIBIDAE: ANTHRIBINAE: ZYGAENODINI.

Type species: Platyrhinus costirostris Clairville, 1798 (= Curculio resinosus Scopoli, 1763), by subsequent designation (Pierce 1930: 12).

Family-group name: PLATYRRHINIDAE Bedel, 1882. Stem: Platyrhin-. Note: senior homonym of PLATYRHINIDAE Jordan, 1923 (type genus Platyrhina J. Müller & Henle, 1838) in Pisces.

Platyrhopalus Westwood, 1833: 615, 654. Current status: valid genus in CARABIDAE: PAUSSINAE: PAUSSINI: PAUSSINA.

Type species: Paussus denticornis Donovan, 1804, by original designation.

Family-group name: PLATYRHOPALINI Jeannel, 1946. Stem: Platyrhopal-.

Platyscelis Latreille, 1818: 23. Current status: valid genus in TENEBRIONIDAE: BLAPTINAE: PLATYSCELIDINI. Note: name placed on the Official List of Generic Names in Zoology (ICZN 1993f).

Type species: Tenebrio hypolithus Pallas, 1781, by plenary power (see ICZN 1993f). Note: name placed on the Official List of Specific Names in Zoology (ICZN 1993f).

Family-group name: PLATYSCÉLIDES Lacordaire, 1859. Stem: Platyscelid-.

Platysma Bonelli, 1810: Tab. Syn. Current status: valid genus in CARABIDAE: HARPALINAE: PTEROSTICHINI: PTEROSTICHINA.

Type species: Carabus niger Schaller, 1783, by subsequent designation (Stephens 1835b: 414). Note: No species were originally included in this genus by Bonelli (1810); the first ones were the two species listed by Panzer (1813: 55).

Family-group name: PLATYSMATINI Tschitschérine, 1899. Stem: Platysmat-.

Platysoma Leach, 1817: 77. Current status: valid genus in HISTERIDAE: HISTERINAE: PLATYSOMATINI.

Type species: Hister depressus Fabricius, 1787 (= Hister compressus Herbst, 1783), by subsequent designation (Stephens 1835b: 415).

Family-group name: PLATYSOMINI Bickhardt, 1914. Stem: Platysomat-.

Platyspartus Faust, 1898b: 235, 247. Current status: valid genus in CURCULIONIDAE: ENTIMINAE: OTIORHYNCHITAE: CELEUTHETINI: CELEUTHETINA.

Type species: Piezonotus latiscapus Heller, 1896, by original designation.

Family-group name: PLATYSPARTINA Voss, 1940: 279. Stem: Platyspart-. Note: this name was overlooked in Bouchard et al. (2011) and Bouchard and Bousquet (2020a).

Platysternus Dejean, 1835: 336. Current status: valid genus in CERAMBYCIDAE: LAMIINAE: ANISOCERINI.

Type species: Cerambyx hebraeus Fabricius, 1781, by monotypy.

Family-group name: PLATYSTERNIDES Lacordaire, 1872. Stem: Platystern-. Note: junior homonym of PLATYSTERNIDAE J.E. Gray, 1869 (type genus Platysternon J.E. Gray, 1831) in Reptilia.

Platystomos D.H. Schneider in Fabricius, 1791: 21. Current status: valid genus in ANTHRIBIDAE: ANTHRIBINAE: PLATYSTOMINI.

Type species: Curculio albinus Linnaeus, 1758, by subsequent designation (Pierce 1916: 463).

Family-group name: PLATYSTOMOIDEA Pierce, 1916. Stem: Platystom-.

Platytarsulus Schedl, 1935: 632. Current status: valid genus in CURCULIONIDAE: PLATYPODINAE: TESSEROCERINI: TESSEROCERINA.

Type species: Platytarsulus marshalli Schedl, 1935, by original designation.

Family-group name: PLATYTARSILIDAE Schedl, 1939. Stem: Platytarsul-.

Platyxantha Baly, 1864a: 233. Current status: junior synonym of Taumacera Thunberg, 1814 in CHRYSOMELIDAE: GALERUCINAE: LUPERINI.

Type species: Platyxantha apicalis Baly, 1864, by original designation.

Family-group name: PLATYXANTHITES Chapuis, 1875. Stem: Platyxanth-.

Plaumanniola Costa Lima, 1962: 415. Current status: valid genus in STAPHYLINIDAE: SCYDMAENINAE: SCYDMAENITAE: PLAUMANNIOLINI.

Type species: Plaumanniola sanctaecatharinae Costa Lima, 1962, by monotypy.

Family-group name: PLAUMANNIOLINAE Costa Lima, 1962. Stem: Plaumanniol-.

Plectogaster C.O. Waterhouse, 1881: 429. Current status: valid genus in CERAMBYCIDAE: CERAMBYCINAE: PLECTOGASTERINI.

Type species: Megacoelus pectinicornis H.W. Bates, 1881, by subsequent designation (Quentin and Villiers 1969: 615).

Family-group name: PLECTOGASTERINI Quentin & Villiers, 1969. Stem: Plectogaster-.

Plectris Lepeletier & Audinet-Serville, 1828: 369. Current status: valid genus in SCARABAEIDAE: MELOLONTHINAE: MACRODACTYLINI.

Type species: Plectris tomentosa Lepeletier & Audinet-Serville, 1828, by original designation.

Family-group name: PLECTRIDAE Burmeister, 1855. Stem: Plectr-.

Plectromerus Haldeman, 1847: 43. Current status: valid genus in CERAMBYCIDAE: CERAMBYCINAE: PLECTROMERINI.

Type species [suggested]: Obrium dentatum J.E. LeConte, 1824 (= Callidium dentipes G.-A. Olivier, 1790), by subsequent designation (Linsley 1963: 135). Note: the valid type species fixation is that of J.L. LeConte (1873: 189), who selected Plectromerus concinnatus Haldeman, 1847 (= Curius dentatus Newman, 1840), but that nominal species is the type species of Curius Newman, 1840; although J.L. LeConte (1873: 189) listed Curius dentatus, which is not an originally included nominal species, as type species of Plectromerus, the fact that he listed Plectromerus concinnatus Haldeman, 1847, one of the two originally included species in Plectromerus, at the same time in synonymy with Curius dentatus Newman, he is deemed to have designated Plectromerus concinnatus Haldeman as type species of Plectromerus (Article 69.2.2) (see Bousquet et al. 2009: 53); an application to the Commission is necessary to fix Obrium dentatum J.E. LeConte, 1824 as the type species of Plectromerus Haldeman, 1847.

Family-group name: PLECTROMERINI Nearns & Braham, 2008. Stem: Plectromer-.

Plectrophorus Schönherr, 1826: 106. Current status: invalid senior synonym of Plectrophoroides Wibmer & O’Brien, 1986 in CURCULIONIDAE: ENTIMINAE: POLYDRUSITAE: NAUPACTINI: NAUPACTINA. Note: junior homonym of Plectrophorus Férussac, 1819 [Mollusca].

Type species: Leptocerus lutra Schönherr, 1826, by original designation.

Family-group name: PLECTROPHORINA Voss, 1954. Stem: Plectrophor-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

Plectroscelis Chevrolat in Dejean, 1836a: 393. Current status: junior synonym of Chaetocnema Stephens, 1831 in CHRYSOMELIDAE: GALERUCINAE: ALTICINI.

Type species: Altica aridula Gyllenhal, 1827, by subsequent designation (Monrós and Bechyné 1956: 1134).

Family-group name: PLECTROSCELIDES C.G. Thomson, 1866. Stem: Plectroscelid-.

Plegaderus Erichson, 1834: 203. Current status: valid genus in HISTERIDAE: ABRAEINAE: PLEGADERINI.

Type species: Hister caesus Herbst, 1791, by subsequent designation (Hope 1841a: 105).

Family-group name: PLEGADERINI Portevin, 1929. Stem: Plegader-.

Pleganophorus Hampe, 1855: 97. Current status: valid genus in ENDOMYCHIDAE: PLEGANOPHORINAE.

Type species: Pleganophorus bispinosus Hampe, 1855, by monotypy.

Family-group name: PLÉGANOPHORIDES Jacquelin du Val, 1858. Stem: Pleganophor-.

Pleiarthrocerus Bruch, 1915: 340. Current status: valid genus in CERAMBYCIDAE: CERAMBYCINAE: PLEIARTHROCERINI.

Type species: Pleiarthrocerus opacus Bruch, 1915, by original designation.

Family-group name: PLEIARTHROCERINAE Lane, 1950. Stem: Pleiarthrocer-.

Pleocoma J.L. LeConte, 1856a: 24. Current status: valid genus in PLEOCOMIDAE: PLEOCOMINAE.

Type species: Pleocoma fimbriata J.L. LeConte, 1856, by monotypy.

Family-group name: PLEOCOMINI J.L. LeConte, 1861. Stem: Pleocom-.

Pleonomus Ménétriés, 1849a: 48. Current status: valid genus in ELATERIDAE: DENDROMETRINAE: PLEONOMINI.

Type species: Pleonomus tereticollis Ménétriés, 1849, by subsequent designation (Lacordaire 1857: 222).

Family-group name: PLEONOMINI Semenov-Tjan-Shansky & Pjatakova, 1936. Stem: Pleonom-.

Pleotomus J.L. LeConte, 1861: 184. Current status: valid genus in LAMPYRIDAE: LAMPYRINAE: PLEOTOMINI.

Type species: Pleotomus pallens J.L. LeConte, 1865, by subsequent monotypy (J.L. LeConte 1865: 88).

Family-group name: PLEOTOMINI Summers, 1874. Stem: Pleotom-.

Pleroticus Péringuey, 1896: 521. Current status: valid genus in CARABIDAE: HARPALINAE: CHLAENIINI: CHLAENIINA.

Type species: Vertagus lucidulus Boheman, 1848, by monotypy.

Family-group name: PLEROTICINI Basilewsky, 1950. Stem: Plerotic-.

Plesioclytus Giesbert, 1993: 129. Current status: valid genus in CERAMBYCIDAE: CERAMBYCINAE: PLESIOCLYTINI.

Type species: Plesioclytus relictus Giesbert, 1993, by original designation.

Family-group name: PLESIOCLYTINI Wappes & Skelley, 2015: 2. Stem: Plesioclyt-.

Pleurarius Kaup, 1868b: 1. Current status: valid genus in PASSALIDAE: PASSALINAE: MACROLININI.

Type species: Pleurarius pilipes Kaup, 1868, by monotypy.

Family-group name: PLEURARIINAE Kuwert, 1896. Stem: Pleurari-.

Pleurolabus Jekel, 1860: 197. Current status: valid genus in ATTELABIDAE: ATTELABINAE: ATTELABINI: PLEUROLABINA.

Type species: Rhynchites exaratus Boheman, 1828, by original designation.

Family-group name: PLEUROLABINA Legalov, 2003. Stem: Pleurolab-.

Pleurophorus Mulsant, 1842a: 312. Current status: valid genus in SCARABAEIDAE: APHODIINAE: PSAMMODIINI: PSAMMODIINA.

Type species: Scarabaeus caesus Panzer, 1796, by monotypy. Note: see Branco (2007: 19) for comments about the authorship of this species-group name.

Family-group name: PLEUROPHORATES Mulsant, 1842. Stem: Pleurophor-. Note: senior homonym of PLEUROPHORIDAE Weller, 1886 (type genus Pleurophorus King, 1844) in Mollusca; Weller’s name is permanently invalid (based on a preoccupied type genus).

Pleuropterus Westwood, 1841a: 11. Current status: invalid senior synonym of Heteropaussus J. Thomson, 1860 in CARABIDAE: PAUSSINAE: PAUSSINI: HETEROPAUSSINA. Note: junior homonym of Pleuropterus Burnett, 1829 [Mammalia].

Type species: Cerapterus westermanni Westwood, 1841, by monotypy.

Family-group name: PLEUROPTERINI Wasmann, 1920. Stem: Pleuropter-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

Plinthus Germar, 1817b: 340. Current status: valid genus in CURCULIONIDAE: MOLYTINAE: MOLYTINI: PLINTHINA. Note: name placed on the Official List of Generic Names in Zoology (ICZN 2012d).

Type species: Curculio megerlei Panzer, 1803, by plenary power (see ICZN 2012d). Note: name placed on the Official List of Specific Names in Zoology (ICZN 2012d).

Family-group name: PLINTHIDES Lacordaire, 1863. Stem: Plinth-.

Plocamotrechus Jeannel, 1926: 480, 525. Current status: junior synonym of Pachydesus Motschulsky, 1864 in CARABIDAE: TRECHINAE: TRECHINI: TRECHINA.

Type species: Trechus pallipes Boheman, 1848 (= Plocamotrechus bohemani Jeannel, 1926), by original designation.

Family-group name: PLOCAMOTRECHINA Jeannel, 1960. Stem: Plocamotrech-.

Plocastes Gistel, 1856: 365. Current status: junior synonym of Aesalus Fabricius, 1801 in LUCANIDAE: AESALINAE: AESALINI.

Type species: Lucanus scarabaeoides Panzer, 1795, by monotypy.

Family-group name: PLOCASTEIDAE Gistel, 1856. Stem: Plocast-.

Plochionus Dejean, 1821: 5. Current status: valid genus in CARABIDAE: HARPALINAE: LEBIINI: CALLEIDINA.

Type species: Lebia bonfilsi Audinet-Serville, 1821 (= Carabus pallens Fabricius, 1775), by monotypy.

Family-group name: PLOCHIONIDAE Gozis, 1875. Stem: Plochion-.

Ploeosoma Wollaston, 1854: 147. Current status: valid genus in CERYLONIDAE: CERYLONINAE.

Type species: Ploeosoma ellipticum Wollaston, 1854, by monotypy.

Family-group name: PLEOSOMIDES Fauvel, 1891. Stem: Ploeosomat-.

Plotina Lewis, 1896: 35. Current status: valid genus in COCCINELLIDAE: COCCINELLINAE: PLOTININI.

Type species: Plotina versicolor Lewis, 1896, by original designation.

Family-group name: PLOTININI Miyatake, 1994. Stem: Plotin-.

Plusiotis Burmeister, 1844: 417. Current status: valid genus in SCARABAEIDAE: RUTELINAE: RUTELINI: RUTELINA.

Type species: Pelidnota victorina Hope, 1841, by subsequent designation (Ohaus 1934: 16).

Family-group name: PLUSIOTINA H.W. Bates, 1888. Stem: Plusiotid-.

Pocadius Erichson, 1843a: 318. Current status: valid genus in NITIDULIDAE: NITIDULINAE: NITIDULINI.

Type species: Nitidula ferruginea Fabricius, 1775, by subsequent designation (E. Desmarest 1851: 281).

Family-group name: POCADIINI Seidlitz, 1872 [Gtt.]. Stem: Pocadi-.

Podabrocephalus Pic, 1913: 118. Current status: valid genus in PTILODACTYLIDAE: PTILODACTYLINAE.

Type species: Podabrocephalus sinuaticollis Pic, 1913, by monotypy.

Family-group name: PODABROCEPHALIDAE Pic, 1930. Stem: Podabrocephal-.

Podabrus Dejean, 1833b: 105. Current status: valid genus in CANTHARIDAE: CANTHARINAE: PODABRINI.

Type species: Cantharis alpinus Paykull, 1798, by subsequent designation (Stephens 1835a: 416).

Family-group name: PODABRIDAE Gistel, 1856. Stem: Podabr-.

Podagrica Chevrolat in Dejean, 1836a: 394. Current status: valid genus in CHRYSOMELIDAE: GALERUCINAE: ALTICINI.

Type species: Altica fuscipes Fabricius, 1775, by subsequent designation (Maulik 1926: 273).

Family-group name: PODAGRICINI Bechyné, 1968. Stem: Podagric-.

Podalgus Burmeister, 1847: 90, 117. Current status: valid genus in SCARABAEIDAE: DYNASTINAE: PENTODONTINI: PENTODONTINA. Note: name placed on the Official List of Generic Names in Zoology (ICZN 2004a).

Type species: Podalgus cuniculus Burmeister, 1847, by subsequent designation (Arrow 1908: 341; see ICZN 2004a). Note: name placed on the Official List of Specific Names in Zoology (ICZN 2004a).

Family-group name: PODALGIDAE Burmeister, 1847. Stem: Podalg-.

Podapion C.V. Riley, 1883: 62. Current status: valid genus in BRENTIDAE: APIONINAE: PODAPIITAE.

Type species: Podapion gallicola C.V. Riley, 1883, by monotypy.

Family-group name: PODAPIITAE Wanat, 2001. Stem: Podapi-.

Podolasia Harold, 1869b: 122. Current status: valid genus in SCARABAEIDAE: PODOLASIINAE. Note: new replacement name for Lasiopus J.L. LeConte, 1856.

Type species: Lasiopus ferrugineus J.L. LeConte, 1856, by monotypy (type fixation for Lasiopus J.L. LeConte).

Family-group name: PODOLASIINI Howden, 1997. Stem: Podolasi-.

Podonta Solier, 1835b: 247. Current status: valid genus in TENEBRIONIDAE: ALLECULINAE: CTENIOPODINI.

Type species: Cistela nigrita Fabricius, 1794, by subsequent designation (Cazurro Ruiz 1894: 883).

Family-group name: PODONTINI Portevin, 1934. Stem: Podont-.

Poecilonota Eschscholtz, 1829b: 9. Current status: valid genus in BUPRESTIDAE: CHRYSOCHROINAE: POECILONOTINI: POECILONOTINA. Note: name placed on the Official List of Generic Names in Zoology (ICZN 1996a).

Type species: Buprestis variolosa Paykull, 1799, by plenary power (see ICZN 1996a). Note: name placed on the Official List of Specific Names in Zoology (ICZN 1996a).

Family-group name: POECILONOTINA Jakobson, 1913. Stem: Poecilonot-.

Poecilopeplus Dejean, 1835: 319. Current status: valid genus in CERAMBYCIDAE: CERAMBYCINAE: TRACHYDERINI: TRACHYDERINA.

Type species: Prionus corallifer Sturm, 1826, by monotypy.

Family-group name: POECILOPÉPLIDES Lacordaire, 1868. Stem: Poecilopepl-.

Poecilus Bonelli, 1810: Tab. Syn. Current status: valid genus in CARABIDAE: HARPALINAE: PTEROSTICHINI: PTEROSTICHINA.

Type species: Carabus cupreus Linnaeus, 1758, by subsequent designation (Curtis 1827: pl. 187). Note: no species were originally included in this genus by Bonelli (1810); the first ones were the six species listed by Panzer (1813: 60, 61).

Family-group name: POECILII Bonelli, 1810. Stem: Poecil-.

Poekilosoma Audinet-Serville, 1832: 184. Current status: valid genus in CERAMBYCIDAE: PRIONINAE: ANACOLINI.

Type species: Prionus ornatus Dalman, 1823, by subsequent designation (J. Thomson 1864: 276).

Family-group name: POECILOSOMI H.W. Bates, 1869. Stem: Poekilosomat-.

Pogonocerus Fischer, 1812: 281. Current status: valid genus in PYROCHROIDAE: POGONOCERINAE.

Type species: Pogonocerus thoracicus Fischer, 1812, by monotypy.

Family-group name: POGONOCERINAE Iablokoff-Khnzorian, 1985. Stem: Pogonocer-.

Pogonocherus Dejean, 1821: 107. Current status: valid genus in CERAMBYCIDAE: LAMIINAE: POGONOCHERINI.

Type species: Cerambyx hispidus Linnaeus, 1758, by subsequent designation (Guérin- Méneville 1826: 186).

Family-group name: POGONOCHÉRAIRES Mulsant, 1839. Stem: Pogonocher-.

Pogonopsis Bedel, 1898: 241. Current status: valid genus in CARABIDAE: TRECHINAE: POGONINI.

Type species: Pogonopsis pallida Bedel, 1898, by monotypy.

Family-group name: POGONOPSINI Bedel, 1900. Stem: Pogonopse-.

Pogonostoma Klug, 1835: 382. Current status: valid genus in CICINDELIDAE: CTENOSTOMATINI.

Type species: Pogonostoma chalybaeum Klug, 1835, by subsequent designation (W. Horn 1926: 12).

Family-group name: POGONOSTOMINI Mandl, 1954. Stem: Pogonostomat-.

Pogonus Dejean, 1821: 9. Current status: valid genus in CARABIDAE: TRECHINAE: POGONINI.

Type species: Carabus littoralis Duftschmid, 1812, by monotypy.

Family-group name: POGONIDAE Laporte, 1834. Stem: Pogon-.

Polistichus Bonelli, 1810: Tab. Syn. Current status: valid genus in CARABIDAE: HARPALINAE: ZUPHIINI: ZUPHIINA.

Type species: here fixed (under ICZN 1999a, Article 70.3.2) as Buprestis connexus Geoffroy, 1785, misidentified as Carabus fasciolatus Rossi, 1790 (sensu Fabricius, 1801) in the designation by subsequent monotypy in Latreille (1816: 184).

Family-group name: POLYSTICHINAE H.W. Bates, 1886. Stem: Polistich-.

†Pollostelater Alexeev, 2011: 65. Current status: valid genus in ELATERIDAE: †PROTAGRYPNINAE: †POLLOSTELATERINI. Note: Lower Cretaceous (Russia).

Type species: †Pollostelater baissensis Alexeev, 2011, by original designation.

Family-group name: †POLLOSTELATERINI Alexeev, 2011: 65. Stem: Pollostelater-. Note: this name was overlooked in Bouchard et al. (2011), Bouchard and Bousquet (2020a).

Polpochila Solier, 1849: 217. Current status: valid genus in CARABIDAE: HARPALINAE: HARPALINI: STENOLOPHINA.

Type species: Polpochila parallela Solier, 1849 (= Melanotus chilensis Chaudoir, 1837), by original designation.

Family-group name: POLPOCHILINAE H.W. Bates, 1891. Stem: Polpochil-.

Polyarthron Audinet-Serville, 1832: 189. Current status: valid genus in CERAMBYCIDAE: PRIONINAE: PRIONINI.

Type species: Prionus pectinicornis Fabricius, 1792, by monotypy.

Family-group name: POLYARTHRINI Gounelle, 1911. Stem: Polyarthr-. Note: junior homonym of POLYARTHRIDAE Daday, 1893 (type genus Polyarthra Ehrenberg, 1834) in Rotifera.

Polybothris Dupont, 1833: pl. 77. Current status: valid genus in BUPRESTIDAE: CHRYSOCHROINAE: DICERCINI: DICERCINA. Note: the original spelling was Polybotris but the Commission ruled that Polybothris was the correct original spelling and placed Polybothris Dupont, 1833 on the Official List of Generic Names in Zoology (ICZN 2015a).

Type species: Polybothris croesus Dupont, 1833 (= Buprestis sumptuosa Klug, 1833), by monotypy (see ICZN 2015a). Note: Polybothris croesus Dupont, 1833 was placed on the Official List of Specific Names in Zoology (ICZN 2015a).

Family-group name: POLYBOTHRISIDAE Gistel, 1848. Stem: Polybothrid-.

Polycaon Laporte, 1838: 30. Current status: valid genus in BOSTRICHIDAE: POLYCAONINAE.

Type species: Polycaon chiliensis Laporte, 1838, by monotypy.

Family-group name: POLYCAONINAE Lesne, 1896. Stem: Polycaon-.

Polycatus Heller, 1913: 379. Current status: valid genus in CURCULIONIDAE: ENTIMINAE: OTIORHYNCHITAE: POLYCATINI.

Type species: Polycatus aurofasciatus Heller, 1913, by original designation.

Family-group name: POLYCATINI G.A.K. Marshall, 1956. Stem: Polycat-.

Polycesta Dejean, 1833a: 78. Current status: valid genus in BUPRESTIDAE: POLYCESTINAE: POLYCESTINI: POLYCESTINA.

Type species: Buprestis porcata Fabricius, 1775, by monotypy.

Family-group name: POLYCESTIDES Lacordaire, 1857. Stem: Polycest-.

Polyctesis Marseul, 1865: 264. Current status: valid genus in BUPRESTIDAE: POLYCESTINAE: POLYCTESINI.

Type species: Polyctesis rhois Marseul, 1865, by monotypy.

Family-group name: POLYCTESINI Cobos, 1955. Stem: Polyctes-.

Polycystophorus Gistel, 1856: 385. Current status: junior synonym of Malachius Fabricius, 1775 in MELYRIDAE: MALACHIINAE: MALACHIINI.

Type species: Cantharis aenea Linnaeus, 1758, by subsequent designation (Bouchard et al. 2011: 352).

Family-group name: POLYCYSTOPHORIDAE Gistel, 1856. Stem: Polycystophor-.

Polydrusus Germar, 1817b: 341. Current status: valid genus in CURCULIONIDAE: ENTIMINAE: POLYDRUSITAE: POLYDRUSINI. Note: name placed on the Official List of Generic Names in Zoology (ICZN 1981c).

Type species: Curculio undatus Fabricius, 1781 (= Curculio tereticollis DeGeer, 1775), by subsequent designation (Schönherr 1826: 12; see ICZN 1981c). Note: Curculio undatus Fabricius, 1781 was placed on the Official List of Specific Names in Zoology (ICZN 1981c).

Family-group name: POLYDROSIDES Schönherr, 1823. Stem: Polydrus-.

Polygraphus Erichson, 1836: 57. Current status: valid genus in CURCULIONIDAE: SCOLYTINAE: POLYGRAPHINI.

Type species: Bostrichus pubescens Fabricius, 1792 (= Dermestes poligraphus Linnaeus, 1758), by monotypy.

Family-group name: POLYGRAPHIDAE Chapuis, 1869. Stem: Polygraph-. Note: senior homonym of POLYGRAPHINI Rydon, 1971 (type genus Polygrapha Staudinger, 1887) in Lepidoptera.

Polyhirma Chaudoir, 1850: 44. Current status: senior synonym of Cypholoba Chaudoir, 1850 in CARABIDAE: HARPALINAE: ANTHIINI. Note: First Reviser (ICZN 1999a, Article 24.2.2) for Polyhirma Chaudoir, 1850 versus Cypholoba Chaudoir, 1850 is Péringuey (1896: 339) who selected Polyhirma as the valid name (the accepted synonymy between the two genus-group names found in the literature is given to G. Strohmeyer (1928: 298) who selected Cypholoba); an application to the Commission is necessary to retain Cypholoba Chaudoir, 1850 as the valid name.

Type species: Carabus macilentus G.-A. Olivier, 1795, by subsequent designation (Lorenz 1998: 148).

Family-group name: POLYHIRMI Rousseau, 1905. Stem: Polyhirm-.

Polyphylla Harris, 1841: 30. Current status: valid genus in SCARABAEIDAE: MELOLONTHINAE: MELOLONTHINI: MELOLONTHINA.

Type species: Melolontha variolosa Hentz, 1830, by subsequent designation (R.M. Young 1988: 21).

Family-group name: POLYPHYLLIDAE Burmeister, 1855. Stem: Polyphyll-.

Polypleurus Eschscholtz, 1831: 10, 11. Current status: valid genus in TENEBRIONIDAE: STENOCHIINAE: CNODALONINI.

Type species: Polypleurus geminatus Eschscholtz, 1831, by monotypy.

Family-group name: POLYPLEURI J.L. LeConte, 1862. Stem: Polypleur-.

Polypria Chevrolat, 1874: 330. Current status: valid genus in OEDEMERIDAE: POLYPRIINAE.

Type species: Polypria cruxrufa Chevrolat, 1874, by monotypy.

Family-group name: POLYPRIINAE Lawrence, 2005. Stem: Polypri-.

Polyrhaphis Audinet-Serville, 1835a: 26. Current status: valid genus in CERAMBYCIDAE: LAMIINAE: POLYRHAPHIDINI.

Type species: Lamia horrida Fabricius, 1792 (= Cerambyx spinosus Drury, 1773), by subsequent designation (E. Desmarest 1860: 321).

Family-group name: POLYRHAPHITAE J. Thomson, 1860. Stem: Polyrhaphid-.

Polytus Faust, 1894a: 353. Current status: valid genus in CURCULIONIDAE: DRYOPHTHORINAE: LITOSOMINI: POLYTINA.

Type species: Sitophilus mellerborgii Boheman, 1838, by original designation.

Family-group name: POLYTINI Zimmerman, 1993. Stem: Polyt-.

Pomachilius Eschscholtz, 1829a: 31. Current status: valid genus in ELATERIDAE: ELATERINAE: POMACHILIINI.

Type species: Elater subfasciatus Germar, 1823, by monotypy.

Family-group name: POMACHILIITES Candèze, 1859. Stem: Pomachili-.

Pondonatus John, 1954: 309. Current status: valid genus in DISCOLOMATIDAE: PONDONATINAE.

Type species: Pondonatus turneri John, 1954, by original designation.

Family-group name: PONDONATINAE John, 1954. Stem: Pondonat-.

†Ponomarenkia Yan, Lawrence, Beattie & Beutel, 2017c: 613. Current status: invalid senior synonym of †Ponomarenkium Yan, Lawrence, Beattie & Beutel, 2018 in †PONOMARENKIUMIDAE. Note: junior homonym of Ponomarenkia Perkovsky, 2001 in Coleoptera: AGYRTIDAE. Note: upper Permian (Australia).

Type species: †Ponomarenkia belmonthensis Yan, Lawrence, Beattie & Beutel, 2017, by original designation.

Family-group name: †PONOMARENKIIDAE Yan, Lawrence, Beattie & Beutel, 2017c: 612. Stem: Ponomarenki-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

†Ponomarenkium Yan, Lawrence, Beattie & Beutel, 2018: 110. Current status: valid genus in †PONOMARENKIUMIDAE. Note: upper Permian (Australia).

Type species: †Ponomarenkia belmonthensis Yan, Lawrence, Beattie & Beutel, 2017, by original designation.

Family-group name: †PONOMARENKIUMIDAE Lawrence & Bouchard in Cai et al., 2022: 12, 13 (Supp. Inf.). Stem: Ponomarenkium-.

Poophagus Schönherr, 1837: 590. Current status: valid genus in CURCULIONIDAE: CEUTORHYNCHINAE: CEUTORHYNCHINI.

Type species: Curculio sisymbrii Fabricius, 1777, by original designation.

Family-group name: POOPHAGIDAE Schultze, 1902. Stem: Poophag-.

Popilius Kaup, 1871: 75. Current status: valid genus in PASSALIDAE: PASSALINAE: PROCULINI.

Type species: Passalus marginatus Percheron, 1834, by subsequent designation (Gravely 1918: 26).

Family-group name: POPILIINAE Kuwert, 1896. Stem: Popili-.

Popillia Dejean, 1821: 60. Current status: valid genus in SCARABAEIDAE: RUTELINAE: ANOMALINI: POPILLIINA.

Type species: Trichius bipunctatus Fabricius, 1787, by monotypy.

Family-group name: POPILLIIDAE Ohaus, 1902. Stem: Popilli-.

Poria Mulsant, 1850: 885. Current status: valid genus in COCCINELLIDAE: COCCINELLINAE: COCCIDULINI.

Type species: Poria cyanea Mulsant, 1850, by subsequent designation (Crotch 1874a: 288).

Family-group name: PORIENS Mulsant, 1850. Stem: Pori-.

Porphyraspis Hope, 1841a: 154. Current status: junior synonym of Hemisphaerota Chevrolat, 1836 in CHRYSOMELIDAE: CASSIDINAE: HEMISPHAEROTINI.

Type species: Cassida erythrocera Germar, 1823, by original designation. Note: this species is generally listed as a junior synonym of Imatidium cyaneum Say, 1824, but Germar’s name is older and should be used as the valid name for the species; an application to the Commission is necessary to conserve usage of the commonly used name Hemisphaerota cyanea (Say, 1824).

Family-group name: PORPHYRASPITAE Spaeth, 1929. Stem: Porphyraspid-.

Porphyronota Burmeister, 1842: 622. Current status: valid genus in SCARABAEIDAE: CETONIINAE: DIPLOGNATHINI.

Type species: Cetonia carnifex Fabricius, 1781, by subsequent designation (Holm 1990: 113).

Family-group name: PORPHYRONOTII Péringuey, 1907. Stem: Porphyronot-.

Potamophilus Germar, 1811: 41. Current status: valid genus in ELMIDAE: LARAINAE: POTAMOPHILINI.

Type species: Parnus acuminatus Fabricius, 1792, by monotypy.

Family-group name: POTAMOPHILIDAE Fournel & Géhin, 1845. Stem: Potamophil-. Note: this family-group name was attributed to “Mulsant & Rey, 1872” in Bouchard et al. (2011: 292); however, it was first made available by Fournel and Géhin (1845: 119); the tribe POTAMOPHILINI is currently included in the subfamily LARAINAE J.L. LeConte, 1861 and since POTAMOPHILINI is older than LARAINAE, it should be the valid name for the subfamily; however, usage of LARAINAE J.L. LeConte, 1861 is conserved following Article 35.5 (ICZN 1999a).

Potemnemus J. Thomson, 1864: 81. Current status: valid genus in CERAMBYCIDAE: LAMIINAE: LAMIINI.

Type species: Cerambyx scabrosus G.-A. Olivier, 1790, by original designation.

Family-group name: POTEMNEMINI Aurivillius, 1922. Stem: Potemnem-.

Potergus Bonvouloir, 1871: 64, 110. Current status: valid genus in THROSCIDAE.

Type species: Potergus filiformis Bonvouloir, 1871, by monotypy.

Family-group name: POTERGINI Cobos, 1961. Stem: Poterg-.

†Praelaterium Dolin, 1973: 75. Current status: valid genus in CEROPHYTIDAE. Note: Jurassic (Kyrgyzstan).

Type species: †Praelaterium problematicum Dolin, 1973, by original designation.

Family-group name: †PRAELATERIIDAE Dolin, 1973. Stem: Praelateri-.

†Praemordella Ščegoleva-Barovskaya, 1929: 27. Current status: valid genus in MORDELLIDAE: †PRAEMORDELLINAE. Note: Upper Jurassic (Kazakhstan).

Type species: †Praemordella martynovi Ščegoleva-Barovskaya, 1929, by monotypy.

Family-group name: †PRAEMORDELLINAE Ščegoleva-Barovskaya, 1929. Stem: Praemordell-.

Praeugena Laporte, 1840b: 241. Current status: valid genus in TENEBRIONIDAE: TENEBRIONINAE: PRAEUGENINI. Note: nomen protectum (see Bouchard and Bousquet 2020b: 6).

Type species: Helops marginatus Fabricius, 1792, by subsequent designation (Hope 1841a: 133).

Family-group name: PRAEUGENINA De Moor, 1970. Stem: Praeugen-.

Praocis Eschscholtz, 1829d: 6. Current status: valid genus in TENEBRIONIDAE: PIMELIINAE: PRAOCIINI.

Type species: Praocis rufipes Eschscholtz, 1829, by subsequent designation (Guérin-Méneville 1834b: 8–9).

Family-group name: PRAOCIDAE Eschscholtz, 1829. Stem: Praoci-.

Prasocuris Latreille, 1802b: 224. Current status: valid genus in CHRYSOMELIDAE: CHRYSOMELINAE: CHRYSOMELINI.

Type species: Chrysomela phellandrii Linnaeus, 1758, by monotypy.

Family-group name: PRASOCURISIDAE Gistel, 1848. Stem: Prasocur-.

Prasoidea Weise, 1907: 133. Current status: junior synonym of Odontionopa Chevrolat, 1836 in CHRYSOMELIDAE: EUMOLPINAE: EURYOPINI.

Type species: Colaspis sericea Gyllenhal, 1808, by monotypy.

Family-group name: PRASOIDEINI Clavareau, 1914. Stem: Prasoide-.

Prateus J.L. LeConte, 1862: 238. Current status: valid genus in TENEBRIONIDAE: LAGRIINAE: PRATEINI.

Type species: Prateus fusculus J.L. LeConte, 1862, by original designation.

Family-group name: PRATEINI Aalbu, Kanda, Merkl, Ivie & Johnston, 2023: 161. Stem: Prate-.

Premnobius Eichhoff, 1878: 404. Current status: valid genus in CURCULIONIDAE: SCOLYTINAE: IPINI: PREMNOBIINA.

Type species: Premnobius cavipennis Eichhoff, 1878, by monotypy.

Family-group name: PREMNOBIINI Browne, 1962. Stem: Premnobi-.

Premnotrypes Pierce, 1914: 348. Current status: valid genus in CURCULIONIDAE: ENTIMINAE: ENTIMITAE: PREMNOTRYPINI.

Type species: Premnotrypes solani Pierce, 1914, by original designation.

Family-group name: PREMNOTRYPINI Kuschel, 1956. Stem: Premnotryp-.

Prepusa Chaudoir, 1850: 15. Current status: junior synonym of Eulampra Chaudoir, 1848 in CICINDELIDAE: CICINDELINI: DROMICINA. Note: unnecessary new replacement name for Eulampra Chaudoir, 1848.

Type species: Cicindela miranda Chaudoir, 1843, by monotypy (type fixation for Eulampra Chaudoir).

Family-group name: PREPUSINI W. Horn, 1899. Stem: Prepus-.

Pretilia H.W. Bates, 1866: 302. Current status: valid genus in CERAMBYCIDAE: LAMIINAE: PRETILIINI.

Type species: Pretilia telephoroides H.W. Bates, 1866 (= Saperda tuberculata Fabricius, 1801), by monotypy.

Family-group name: PRETILIINI Martins & Galileo, 1990. Stem: Pretili-.

Priacma J.L. LeConte, 1874c: 87. Current status: valid genus in CUPEDIDAE: PRIACMINAE.

Type species: Cupes serrata J.L. LeConte, 1861, by monotypy.

Family-group name: PRIACMINI Crowson, 1962. Stem: Priacm-.

Priasilpha Broun, 1893b: 1077. Current status: valid genus in PRIASILPHIDAE.

Type species: Priasilpha obscura Broun, 1893, by monotypy.

Family-group name: PRIASILPHINAE Crowson, 1973. Stem: Priasilph-.

Prinobius Mulsant, 1842b: 207. Current status: valid genus in CERAMBYCIDAE: PRIONINAE: MACROTOMINI: MACROTOMINA.

Type species: Prinobius myardi Mulsant, 1842, by monotypy.

Family-group name: PRINOBIINI Vives, 2000. Stem: Prinobi-.

Priocera W. Kirby, 1819: 389. Current status: valid genus in CLERIDAE: CLERINAE: PRIOCERINI.

Type species: Priocera variegata W. Kirby, 1819, by monotypy.

Family-group name: PRIOCERIDAE Laporte, 1838. Stem: Priocer-.

Priolomus Erichson, 1845a: 256. Current status: valid genus in ZOPHERIDAE: COLYDIINAE: SYNCHITINI.

Type species: Priolomus spinicollis Fairmaire, 1869, by subsequent monotypy (Fairmaire 1869: 204).

Family-group name: PRIOLOMINI Dajoz, 1980. Stem: Priolom-.

Prionobrachium Faust, 1894b: 341. Current status: valid genus in CURCULIONIDAE: CURCULIONINAE: PRIONOBRACHIINI.

Type species: Prionobrachium schoenherri Faust, 1894, by monotypy.

Family-group name: PRIONOBRACHIINA Hustache, 1938. Stem: Prionobrachi-.

Prionocerus Perty, 1831: xxxiii. Current status: valid genus in PRIONOCERIDAE.

Type species: Prionocerus coeruleipennis Perty, 1831, by monotypy.

Family-group name: PRIONOCÉRIDES Lacordaire, 1857. Stem: Prionocer-. Note: senior homonym of PRIONOCERINI Savchenko, 1966 (type genus Prionocera Loew, 1844) in Diptera.

Prionomerus Schönherr, 1835: 359. Current status: junior synonym of Odontopus Say, 1832 in CURCULIONIDAE: CURCULIONINAE: CAMAROTINI: PRIONOMERINA.

Type species: Prionomerus carbonarius Gyllenhal, 1835 (= Anthonomus calceatus Say, 1832), by original designation.

Family-group name: PRIONOMÉRIDES Lacordaire, 1863. Stem: Prionomer-.

Prionomma A. White, 1853: 19. Current status: valid genus in CERAMBYCIDAE: PRIONINAE: PRIONINI.

Type species: Prionus orientalis G.-A. Olivier, 1795 (= Prionus atratus Gmelin, 1790 [as attratus]), by monotypy.

Family-group name: PRIONOMMITAE J. Thomson, 1861. Stem: Prionommat-.

Prionus Geoffroy, 1762: 198. Current status: valid genus in CERAMBYCIDAE: PRIONINAE: PRIONINI. Note: name placed on the Official List of Generic Names in Zoology (ICZN 1994a).

Type species: Cerambyx coriarius Linnaeus, 1758, by plenary power (see ICZN 1994a). Note: name placed on the Official List of Specific Names in Zoology (ICZN 1994a).

Family-group name: PRIONII Latreille, 1802. Stem: Prion-.

Prioptera Hope, 1841a: 153, 176. Current status: junior synonym of Basiprionota Chevrolat, 1836 in CHRYSOMELIDAE: CASSIDINAE: BASIPRIONOTINI.

Type species: Cassida octopunctata Fabricius, 1787, by original designation.

Family-group name: PRIOPTERITAE Spaeth, 1914. Stem: Priopter-.

Prioscelis Hope, 1841a: 128. Current status: valid genus in TENEBRIONIDAE: LAGRIINAE: PYCNOCERINI.

Type species: Prioscelis fabricii Hope, 1841, by original designation.

Family-group name: PRIOSCELINA Skopin, 1964. Stem: Prioscelid-.

Prisahypnus Stibick, 1979: 166. Current status: valid genus in ELATERIDAE: HYPNOIDINAE.

Type species: Cryptohypnus frontalis Sharp, 1877, by original designation.

Family-group name: PRISAHYPINI Stibick, 1979. Stem: Prisahypn-.

Pristiptera Dejean, 1833a: 77. Current status: senior synonym of Pelecopselaphus Solier, 1833 in BUPRESTIDAE: CHRYSOCHROINAE: PARALEPTODEMINI: PRISTIPTERINA. Note: this name is a senior subjective synonym of Pelecopselaphus Solier, 1833 and should be used as valid (see Bousquet and Bouchard 2013a: 22); however, the junior name is currently used as valid and an application to the Commission is necessary to conserve the usage of Pelecopselaphus Solier, 1833.

Type species: Buprestis blanda Fabricius, 1781, by monotypy.

Family-group name: PRISTIPTERINA Holyński, 1993. Stem: Pristipter-.

Pristolycus Gorham, 1883b: 407. Current status: valid genus in LAMPYRIDAE: LAMPYRINAE: LUCIDOTINI: LUCIDOTINA.

Type species: Pristolycus sagulatus Gorham, 1883, by monotypy.

Family-group name: PRISTOLYCINI J.R. Winkler, 1953. Stem: Pristolyc-.

Pristonychus Dejean, 1828: 3, 43. Current status: valid subgenus of Laemostenus Bonelli, 1810 in CARABIDAE: HARPALINAE: SPHODRINI: SPHODRINA.

Type species: Carabus terricola Herbst, 1784, by subsequent designation (Westwood 1838c: 2).

Family-group name: PRISTONYCHIDAE Gistel, 1848. Stem: Pristonych-.

†Pristorhynchus Heer, 1847: 190. Current status: valid genus in CURCULIONIDAE: ENTIMINAE incertae sedis. Note: Upper Miocene (Germany).

Type species: †Pristorhynchus ellipticus Heer, 1847, by monotypy.

Family-group name: †PRISTORHYNCHIDEN Heer, 1847. Stem: Pristorhynch-.

Pristosia Motschulsky, 1865: 311. Current status: valid genus in CARABIDAE: HARPALINAE: SPHODRINI: PRISTOSIINA.

Type species: Pristosia picea Motschulsky, 1865, by monotypy.

Family-group name: PRISTOSIAE Lindroth, 1956. Stem: Pristosi-.

†Probelus Arnoldi, 1977: 151. Current status: valid genus in NEMONYCHIDAE: †EOBELINAE: †PROBELINI. Note: Upper Jurassic (Kazakhstan).

Type species: †Probelus curvispinus Arnoldi, 1977, by original designation.

Family-group name: †PROBELINI Legalov, 2009. Stem: Probel-.

Problechilus Eichhoff, 1878: 34, 167. Current status: junior synonym of Gymnochilus Eichhoff, 1868 in CURCULIONIDAE: SCOLYTINAE: HEXACOLINI. Note: unnecessary new replacement name for Gymnochilus Eichhoff, 1868.

Type species: Gymnochilus zonatus Eichhoff, 1868, by monotypy (type fixation for Gymnochilus Eichhoff).

Family-group name: PROBLECHILIDAE Eichhoff, 1878. Stem: Problechil-.

Procerus Dejean, 1821: 5. Current status: valid subgenus of Carabus Linnaeaus, 1758 in CARABIDAE: CARABINAE: CARABINI: CARABINA.

Type species: here fixed (under ICZN 1999a, Article 70.3.2) as Carabus gigas Creutzer, 1799, misidentified as Carabus scabrosus G.-A. Olivier, 1790 (sensu Fabricius, 1801) in the subsequent designation by Hope (1838a: 47).

Family-group name: PROCERIDAE Laporte, 1834. Stem: Procer-.

Procirrus Latreille, 1829a: 436. Current status: valid genus in STAPHYLINIDAE: PAEDERINAE: PINOPHILINI: PROCIRRINA.

Type species: Procirrus lefebvrei Latreille, 1829 [as lefeburi], by monotypy.

Family-group name: PROCIRRI Bernhauer & Schubert, 1912. Stem: Procirr-.

Procletus Péringuey, 1896: 568, 570. Current status: valid genus in CARABIDAE: HARPALINAE: CHLAENIINI: CHLAENIINA.

Type species: Procletus singularis Péringuey,1896, by monotypy.

Family-group name: PROCLETINI Basilewsky, 1950. Stem: Proclet-.

Procrustes Bonelli, 1810: 39, Tab. Syn. Current status: valid genus in CARABIDAE: CARABINAE: CARABINI: CARABINA.

Type species: Carabus coriaceus Linnaeus, 1758, by monotypy.

Family-group name: PROCRUSTOGENICI Roeschke, 1898. Stem: Procrust-.

Proctocera Chevrolat, 1855: 285. Current status: valid genus in CERAMBYCIDAE: LAMIINAE: PROCTOCERINI.

Type species: Proctocera scalaris Chevrolat, 1855, by monotypy.

Family-group name: PROCTOCERINI Aurivillius, 1922. Stem: Proctocer-.

Proctophanes Harold, 1861: 110, 111. Current status: valid genus in SCARABAEIDAE: APHODIINAE: APHODIINI: PROCTOPHANINA.

Type species: Aphodius sculptus Hope, 1848, by monotypy.

Family-group name: PROCTOPHANINI Stebnicka & Howden, 1995. Stem: Proctophan-.

Proculejus Kaup, 1868b: 13. Current status: valid genus in PASSALIDAE: PASSALINAE: PROCULINI.

Type species: Proculejus truquii Kaup, 1868, by subsequent designation (Gravely 1918: 31).

Family-group name: PROCULEJINAE Kuwert, 1896. Stem: Proculej-.

Proculus Kaup, 1868b: 8. Current status: valid genus in PASSALIDAE: PASSALINAE: PROCULINI.

Type species: Passalus goryi Melly, 1833, by subsequent designation (Gravely 1918: 42).

Family-group name: PROCULINAE Kaup, 1868. Stem: Procul-.

†Procurculio Arnoldi, 1977: 157. Current status: valid genus in NEMONYCHIDAE: †ECCOPTARTHRINAE. Note: Upper Jurassic (Kazakhstan).

Type species: †Procurculio fortipes Arnoldi, 1977, by original designation.

Family-group name: †PROCURCULIONINI Arnoldi, 1977. Stem: Procurculion-.

Prodoretus Brenske, 1893: 1. Current status: valid genus in SCARABAEIDAE: RUTELINAE: ADORETINI: PRODORETINA.

Type species: Prodoretus vittatus Brenske, 1893, by subsequent monotypy (Brenske 1893: 10). Note: Prodoretus Brenske, 1893 and Prodoretus vittatus Brenske, 1893 were published in two separate parts (published on different dates) of the same article, hence the use of subsequent monotypy for the type species designation.

Family-group name: PRODORETINA Ohaus, 1912. Stem: Prodoret-.

Proeces Schönherr, 1838: 1080. Current status: valid genus in CURCULIONIDAE: COSSONINAE: COSSONINI. Note: two original spellings were used, including Oeces (p. 1140); Neave (1940: 910) acted as First Reviser (ICZN 1999a, Article 24.2.3) and selected Proeces, the spelling currently in prevailing usage.

Type species: Proeces macer Boheman, 1838, by original designation.

Family-group name: PROECINA Voss, 1956. Stem: Proec-.

Prognathus Blondel, 1827: 413. Current status: junior synonym of Siagonium W. Kirby & Spence, 1815 in STAPHYLINIDAE: PIESTINAE.

Type species: Prognathus rufipennis Blondel, 1827 (= Siagonium quadricorne W. Kirby & Spence, 1815), by monotypy.

Family-group name: PROGNATHITES Blanchard, 1845. Stem: Prognath-.

Proholopterus Monné, 2012: 44. Current status: junior synonym of Holopterus Blanchard, 1851, stat. nov. in CERAMBYCIDAE: CERAMBYCINAE: HOLOPTERINI. Note: unnecessary new replacement name for Holopterus Blanchard, 1851; Holopterus Brehm, 1845 [Aves] is an incorrect subsequent spelling of Hoplopterus Bonaparte, 1831 not in prevailing usage and therefore Holopterus Blanchard, 1851, stat. nov. can be used as the valid name.

Type species: Holopterus chilensis Blanchard, 1851, by monotypy (type fixation for Holopterus Blanchard).

Family-group name: PROHOLOPTERINI Monné, 2012: 44. Stem: Proholopter-.

Prolytta Kaszab, 1959: 110. Current status: valid genus in MELOIDAE: MELOINAE: LYTTINI.

Type species: Lytta lucida Haag-Rutenberg, 1880, by original designation.

Family-group name: PROLYTTINI Selander, 1960. Stem: Prolytt-.

Promecheilus Solier, 1851: 251. Current status: valid genus in PROMECHEILIDAE.

Type species: Promecheilus variegatus Solier, 1851, by monotypy.

Family-group name: PROMÉCHILIDES Lacordaire, 1859. Stem: Promecheil-.

Promecognathus Chaudoir, 1847: 515, 524. Current status: valid genus in CARABIDAE: SCARITINAE: PROMECOGNATHINI.

Type species: Eripus laevissimus Dejean, 1829, by monotypy.

Family-group name: PROMECOGNATHI J.L. LeConte, 1853. Stem: Promecognath-.

Promecops C.R. Sahlberg, 1823a: 30. Current status: valid genus in CURCULIONIDAE: ENTIMINAE: ENTIMITAE: EUDIAGOGINI. Note: we consider Promecops used by Schönherr (1823: 1141) a subsequent use of Promecops C.R. Sahlberg, 1823, not a distinct taxon.

Type species: Promecops nubifer C.R. Sahlberg, 1823, by monotypy. Note: combined description by indication of a new genus and a single new species (ICZN 1999a, Article 12.2.6).

Family-group name: PROMÉCOPIDES Lacordaire, 1863. Stem: Promecop-. Note: this family- group name has priority over EUDIAGOGINI J.L. LeConte, 1874, which is currently use as valid; a request to the Commission is necessary to conserve usage of J.L. LeConte’s name.

Promecotheca Guérin-Méneville, 1840b: 334. Current status: valid genus in CHRYSOMELIDAE: CASSIDINAE: PROMECOTHECINI. Note: as pointed out by Bouchard and Bousquet (2020a: 109), this genus is credited to Blanchard (1853: 312) by several authors including by Gressitt (1957: 279) and Staines (2015: 1), but it was first made available by Guérin-Méneville in 1840 by indication, with the inclusion of at least one available species included under a new genus-group name (ICZN 1999a, Article 12.2.5).

Type species: Promecotheca petelii Guérin-Méneville, 1840, by present designation.

Family-group name: PROMECOTHÉCITES Chapuis, 1875. Stem: Promecothec-.

Prometopia Erichson, 1843a: 279. Current status: valid genus in NITIDULIDAE: PROMETOPIINAE.

Type species: Nitidula sexmaculata Say, 1825, by subsequent designation (R. Lucas 1920: 539).

Family-group name: PROMETOPINAE Böving & Craighead, 1931. Stem: Prometopi-.

Pronomaea Erichson, 1837: 378. Current status: valid genus in STAPHYLINIDAE: ALEOCHARINAE: PRONOMAEINI.

Type species: Pronomaea rostrata Erichson, 1837, by monotypy.

Family-group name: PRONOMÉATES Mulsant & Rey, 1873. Stem: Pronomae-.

Pronoterus Sharp, 1882b: 263. Current status: valid genus in NOTERIDAE: NOTERINAE: NOTERINI.

Type species: Pronoterus punctipennis Sharp, 1882, by monotypy.

Family-group name: PRONOTERINI Nilsson, 2005. Stem: Pronoter-.

Propalticus Sharp, 1879: 88. Current status: valid genus in LAEMOPHLOEIDAE.

Type species: Propalticus oculatus Sharp, 1879, by monotypy.

Family-group name: PROPALTICIDAE Crowson, 1952. Stem: Propaltic-.

Proscarabaeus Schrank, 1781: 225. Current status: junior synonym of Meloe Linnaeus, 1758 in MELOIDAE: MELOINAE: MELOINI.

Type species: Meloe proscarabaeus Linnaeus, 1758, by absolute tautonymy.

Family-group name: PROSCARABAEIDAE Gistel, 1848. Stem: Proscarabae-.

Proscoporhinus Montrouzier, 1861a: 868. Current status: valid genus in ANTHRIBIDAE: ANTHRIBINAE: PROSCOPORHININI.

Type species: Proscoporhinus amyoti Montrouzier, 1861, by monotypy.

Family-group name: PROSCOPORHINIDES Lacordaire, 1865. Stem: Proscoporhin-.

Prosodes Eschscholtz, 1829d: 9. Current status: valid genus in TENEBRIONIDAE: BLAPTINAE: BLAPTINI: PROSODINA.

Type species: Blaps attenuata Fischer, 1820 (= Blaps obtusa Fabricius, 1798), by subsequent designation (Gebien 1937: 846).

Family-group name: PROSODINA Skopin, 1960. Stem: Prosod-.

Prosopocera Blanchard, 1845a: 160, 176. Current status: valid genus in CERAMBYCIDAE: LAMIINAE: PROSOPOCERINI.

Type species: Lamia fronticornis Fabricius, 1781 (= Cerambyx bipunctatus Drury, 1773), by monotypy.

Family-group name: PROSOPOCERITAE J. Thomson, 1864. Stem: Prosopocer-.

Prosopocoilus Hope, 1845b: 30. Current status: valid genus in LUCANIDAE: LUCANINAE: DORCINI.

Type species: Lucanus cavifrons Hope, 1845, by subsequent designation (Arrow 1950: 77).

Family-group name: PROSOPOCOILINI Benesh, 1960. Stem: Prosopocoil-.

Prosopodonta Baly, 1858: 68. Current status: valid genus in CHRYSOMELIDAE: CASSIDINAE: PROSOPODONTINI.

Type species: Prosopodonta limbata Baly, 1858, by original designation.

Family-group name: PROSOPODONTINI Weise, 1910. Stem: Prosopodont-.

Prospheres Saunders, 1868: 7. Current status: valid genus in BUPRESTIDAE: POLYCESTINAE: PROSPHERINI.

Type species: Buprestis aurantiopicta Laporte & Gory, 1838, by monotypy.

Family-group name: PROSPHERESINI Cobos, 1980. Stem: Prospher-.

Prosphodrus Britton, 1959: 106. Current status: valid genus in CARABIDAE: HARPALINAE: PLATYNINI.

Type species: Prosphodrus waltoni Britton, 1959, by monotypy.

Family-group name: PROSPHODRINI J.M. Valentine, 1987. Stem: Prosphodr-.

Prosternon Latreille, 1834: 151. Current status: valid genus in ELATERIDAE: DENDROMETRINAE: PROSTERNINI.

Type species: Elater holosericeus G.-A. Olivier, 1790 (= Elater tessellatus Linnaeus, 1758), by subsequent designation (Westwood 1838c: 26).

Family-group name: PROSTERNIDAE Gistel, 1856. Stem: Prostern-. Note: nomen protectum (see Bouchard et al. 2011: 315).

Prosthetops C.O. Waterhouse, 1879b: 533. Current status: valid genus in HYDRAENIDAE: PROSTHETOPINAE: PROSTHETOPINI.

Type species: Prosthetops capensis C.O. Waterhouse, 1879 (= Ochthebius megacephalus Boheman, 1851), by monotypy.

Family-group name: PROSTHETOPINAE Perkins, 1994. Stem: Prosthetop-.

Prostominia Reitter, 1889f: 315. Current status: valid genus in SALPINGIDAE: PROSTOMINIINAE.

Type species: Prostominia lewisi Reitter, 1889, by monotypy.

Family-group name: PROSTOMININI Grouvelle, 1914. Stem: Prostomini-.

Prostomis Latreille, 1819a: 171. Current status: valid genus in PROSTOMIDAE.

Type species: Trogosita mandibularis Fabricius, 1801, by subsequent monotypy (Lepeletier and Audinet-Serville 1825: 215). Note: Latreille (1819a: 171) listed one species under his genus, “trogosite mandibulaire de Fabricius”; this name being French vernacular is not an available scientific name and cannot be used as the type species.

Family-group name: PROSTOMIDAE C.G. Thomson, 1859. Stem: Prostom-.

†Protactus Heer, 1847: 28. Current status: valid genus in STAPHYLINIDAE: †PROTACTINAE. Note: Upper Miocene (Germany).

Type species: †Protactus erichsonii Heer, 1847, by monotypy.

Family-group name: †PROTACTIDEN Heer, 1847. Stem: Protact-.

†Protagrypnus Dolin, 1973: 75. Current status: valid genus in ELATERIDAE: †PROTAGRYPNINAE: †PROTAGRYPNINI. Note: Jurassic (China; Kyrgkzstan).

Type species: †Protagrypnus exoletus Dolin, 1973, by original designation.

Family-group name: †PROTAGRYPNINI Dolin, 1973. Stem: Protagrypn-.

Protaxis Gahan, 1906: xiv, 93. Current status: valid genus in CERAMBYCIDAE: CERAMBYCINAE: PROTAXINI.

Type species: Protaxis fulvescens Gahan, 1906, by original designation.

Family-group name: PROTAXINI Gahan, 1906. Stem: Protax-.

Proteinus Latreille, 1797: 9. Current status: valid genus in STAPHYLINIDAE: PROTEININAE: PROTEININI. Note: name placed on the Official List of Generic Names in Zoology (ICZN 1969a).

Type species: Dermestes brachypterus Fabricius, 1792, by plenary power (see ICZN 1969a). Note: name placed on the Official List of Specific Names in Zoology (ICZN 1969a).

Family-group name: PROTEININI Erichson, 1839. Stem: Protein-.

Protelater Sharp, 1877c: 482. Current status: valid genus in ELATERIDAE: LISSOMINAE: PROTELATERINI.

Type species: Protelater elongatus Sharp, 1877, by subsequent designation (Hyslop 1921: 667).

Family-group name: PROTELATERIDAE O. Schwarz, 1902. Stem: Protelater-.

Protelmis Grouvelle, 1911: 265. Current status: valid genus in PROTELMIDAE.

Type species: Protelmis limnioides Grouvelle, 1911, by monotypy.

Family-group name: PROTELMINI Jeannel, 1950. Stem: Protelm-.

Proterhinus Sharp, 1878a: 16, 20. Current status: valid genus in BELIDAE: OXYCORYNINAE: AGLYCYDERITAE: AGLYCYDERINI.

Type species: Proterhinus vestitus Sharp, 1878, by subsequent designation (R. Lucas 1920: 541).

Family-group name: PROTERHINIDES Fauvel, 1891. Stem: Proterhin-.

Proterotaphes Kazantsev, 2006: 218. Current status: junior synonym of Plateros Kazantsev, 2004 in LYCIDAE: EROTINAE: PROTEROTAPHINI. Note: unnecessary new replacement name for Proteros Kazantsev, 2004; the name Proteros Holmberg, 1917 [Hymenoptera] was suppressed for both the Principles of Priority and Homonymy and placed on the Index of Rejected and Invalid Generic Names in Zoology (ICZN 2004b).

Type species: Proteros sempiternus Kazantsev, 2004, by original designation (type fixation for Proteros Kazantsev).

Family-group name: PROTEROTAPHINI Kazantsev, 2012b: 399. Stem: Proterotaph-.

Proterus Raffray, 1897a: 231. Current status: valid genus in STAPHYLINIDAE: PSELAPHINAE: GONIACERITAE: PROTERINI.

Type species: Proterus punctatus Raffray, 1897, by monotypy.

Family-group name: PROTERINI Jeannel, 1949. Stem: Proter-.

Proteugnamptus Voss, 1939: 446. Current status: valid genus in ATTELABIDAE: RHYNCHITINAE: RHINOCARTITAE: RHINOCARTINI.

Type species: Proteugnamptus madagassus Voss, 1939, by monotypy.

Family-group name: PROTEUGNAMPTINI Legalov, 2003. Stem: Proteugnampt-.

Prothema Pascoe, 1856: 43. Current status: valid genus in CERAMBYCIDAE: CERAMBYCINAE: PROTHEMINI.

Type species: Prothema signata Pascoe, 1856, by subsequent designation (J. Thomson 1864: 182).

Family-group name: PROTHÉMIDES Lacordaire, 1868. Stem: Prothem-.

Prothydrus Guignot, 1954: 45. Current status: junior synonym of Enhydrus Laporte, 1835 in GYRINIDAE: GYRININAE: DINEUTINI: ENHYDRUSINA. Note: new replacement name for Enhydrus Laporte, 1835; name suppressed for both the Principles of Priority and Homonymy and placed on the Official Index of Rejected and Invalid Generic Names in Zoology (ICZN 1964).

Type species: Gyrinus sulcatus Wiedemann, 1821, by monotypy (type fixation for Enhydrus Laporte).

Family-group name: PROTHYDRINAE Guignot, 1954. Stem: Prothydr-. Note: family-group name permanently invalid (ICZN 1999a, Article 39); name placed on the Official Index of Rejected and Invalid Family-group Names in Zoology (ICZN 1964).

Prothyma Hope, 1838a: 12, 27. Current status: valid genus in CICINDELIDAE: CICINDELINI: DROMICINA.

Type species: Cicindela quadripunctata Fabricius, 1801, by monotypy.

Family-group name: PROTHYMINI W. Horn, 1906. Stem: Prothym-.

Protochthebius Perkins, 1997: 154. Current status: valid genus in HYDRAENIDAE: OCHTHEBIINAE: OCHTHEBIINI.

Type species: Protochthebius satoi Perkins, 1997, by original designation.

Family-group name: PROTOCHTHEBIINA Perkins, 1997. Stem: Protochthebi-.

†Protoclaviger J. Parker & Grimaldi, 2014: 2428. Current status: valid genus in STAPHYLINIDAE: PSELAPHINAE: CLAVIGERITAE: †PROTOCLAVIGERINI. Note: lower Eocene (India).

Type species: †Protoclaviger trichodens J. Parker & Grimaldi, 2014, by monotypy.

Family-group name: †PROTOCLAVIGERINI J. Parker & Grimaldi, 2014: 2428. Stem: Protoclaviger-.

Protocucujus Crowson, 1954: 60. Current status: junior synonym of Ericmodes Reitter, 1878 in PROTOCUCUJIDAE.

Type species: Protocucujus chilensis Crowson, 1954 (= Ericmodes fuscitarsis Reitter, 1878), by original designation.

Family-group name: PROTOCUCUJIDAE Crowson, 1954. Stem: Protocucuj-.

†Protolucanus Nikolajev, 2007b: 18, 213. Current status: valid genus in LUCANIDAE: †PROTOLUCANINAE. Note: Lower Cretaceous (Mongolia).

Type species: †Protolucanus jurassicus Nikolajev, 2007, by original designation.

Family-group name: †PROTOLUCANINAE Nikolajev, 2007. Stem: Protolucan-.

Protomantis Schönherr, 1840a: 721. Current status: valid genus in CURCULIONIDAE: BRACHYCERINAE: BRACHYCERINI.

Type species: Protomantis dregei Gyllenhal, 1840, by original designation.

Family-group name: PROTOMANTIINAE Aurivillius, 1886. Stem: Protomanti-.

Protomeloe M. Abdullah, 1965: 248. Current status: valid genus in MELOIDAE: ELETICINAE: SPASTICINI: PROTOMELOINA.

Type species: Protomeloe argentinensis M. Abdullah, 1965 (= Spastica wagneri Pic, 1915), by original designation.

Family-group name: PROTOMELOINI M. Abdullah, 1965. Stem: Protomelo-.

Protonarthron J. Thomson, 1858b: 180. Current status: valid genus in CERAMBYCIDAE: LAMIINAE: DORCASCHEMATINI.

Type species: Protonarthron diabolicum J. Thomson, 1858, by monotypy.

Family-group name: PROTONARTHRONITAE J. Thomson, 1864. Stem: Protonarthr-.

†Protopassalus Aquino dos Santos, Mattos, Mermudes, Scheffler & Reyes-Castillo, 2021: 4. Current status: valid genus in PASSALIDAE: PASSALINAE: †PROTOPASSALINI. Note: Lower Cretaceous (Brazil).

Type species: †Protopassalus araripensis Aquino dos Santos, Mattos, Mermudes, Scheffler & Reyes-Castillo, 2021, by original designation.

Family-group name: †PROTOPASSALINI Aquino dos Santos, Mattos, Mermudes, Scheffler & Reyes-Castillo, 2021: 4. Stem: Protopassal-.

Protopaussus Gestro, 1892: 706. Current status: valid genus in CARABIDAE: PAUSSINAE: PROTOPAUSSINI.

Type species: Protopaussus feae Gestro, 1892, by monotypy.

Family-group name: PROTOPAUSSINI Gestro, 1892. Stem: Protopauss-.

Protopeltis Crowson, 1964: 287. Current status: valid genus in PROTOPELTIDAE.

Type species: Grynoma viridescens Broun, 1886, by original designation.

Family-group name: PROTOPELTINI Crowson, 1966. Stem: Protopelt-.

Protopselaphus Newton & Thayer, 1995: 227. Current status: valid genus in STAPHYLINIDAE: PROTOPSELAPHINAE.

Type species: Protopselaphus watrousi Newton & Thayer, 1995, by original designation.

Family-group name: PROTOPSELAPHINAE Newton & Thayer, 1995. Stem: Protopselaph-.

†Protorabus Ponomarenko, 1977: 72. Current status: valid genus in CARABIDAE: †PROTORABINAE. Note: Upper Jurassic (Kazakhstan).

Type species: †Protorabus planus Ponomarenko, 1977, by original designation.

Family-group name: †PROTORABINAE Ponomarenko, 1977. Stem: Protorab-.

Protorhopala J. Thomson, 1860a: 91. Current status: valid genus in CERAMBYCIDAE: LAMIINAE: PTEROPLIINI.

Type species: Lamia sexnotata Klug, 1833, by monotypy.

Family-group name: PROTORHOPALITAE J. Thomson, 1864. Stem: Protorhopal-.

†Protoscelis L.N. Medvedev, 1968: 156. Current status: valid genus in CHRYSOMELIDAE: †PROTOSCELIDINAE. Note: Cretaceous, Upper Jurassic (China, Kazakhstan).

Type species: †Protoscelis jurassica L.N. Medvedev, 1968, by original designation.

Family-group name: †PROTOSCELINAE L.N. Medvedev, 1968. Stem: Protoscelid-.

Protosphindus Sen Gupta & Crowson, 1979: 181. Current status: valid genus in SPHINDIDAE: PROTOSPHINDINAE.

Type species: Protosphindus chilensis Sen Gupta & Crowson, 1979, by original designation.

Family-group name: PROTOSPHINDINAE Sen Gupta & Crowson, 1979. Stem: Protosphind-.

Protosternum Sharp, 1890a: 356. Current status: valid genus in HYDROPHILIDAE: SPHAERIDIINAE: PROTOSTERNINI.

Type species: Protosternum atomarium Sharp, 1890, by monotypy.

Family-group name: PROTOSTERNINI Hansen, 1991. Stem: Protostern-.

Protosthetops Perkins in Perkins and J. Balfour-Browne, 1994: 8, in key. Current status: valid genus in HYDRAENIDAE: PROSTHETOPINAE: PROTOSTHETOPINI.

Type species: Prosthetops kenyensis Orchymont, 1948, by original designation.

Family-group name: PROTOSTHETOPINI Perkins, 1994. Stem: Protosthetop-.

Prototrichapion Voss, 1959: 316. Current status: valid genus in BRENTIDAE: APIONINAE: ASPIDAPIITAE: PROTOTRICHAPIINI.

Type species: Apion penicillatum G.A.K. Marshall, 1932, by original designation.

Family-group name: PROTOTRICHAPIINI Wanat, 1995. Stem: Prototrichapi-.

†Prototrox Nikolajev, 2000a: 65. Current status: valid genus in SCARABAEIDAE: †PROTOTROGINAE. Note: Lower Cretaceous (Mongolia; Russia).

Type species: †Prototrox transbaikalicus Nikolajev, 2000, by original designation.

Family-group name: †PROTOTROGINAE Nikolajev, 2000. Stem: Prototrog-.

Proutianus Muona, 1993: 24. Current status: valid genus in EUCNEMIDAE: EUCNEMINAE: PROUTIANINI.

Type species: Phaenocerus americanus G.H. Horn, 1872, by original designation.

Family-group name: PROUTIANINI Muona, 1993. Stem: Proutian-.

Prypnus Schönherr, 1823: 1139. Current status: valid genus in CURCULIONIDAE: ENTIMINAE: POLYDRUSITAE: PRYPNINI.

Type species: Prypnus quinquenodosus Schönherr, 1823, by original designation.

Family-group name: PRYPNIDES Lacordaire, 1863. Stem: Prypn-.

Psalidognathus G.R. Gray in Griffith and Pidgeon, 1831: pls 6, 14. Current status: valid genus in CERAMBYCIDAE: PRIONINAE: PRIONINI.

Type species: Psalidognathus friendii G.R. Gray, 1831, by monotypy.

Family-group name: PSALIDOGNATHITAE J. Thomson, 1861. Stem: Psalidognath-.

Psallidium Herbst, 1795: 342. Current status: valid genus in CURCULIONIDAE: ENTIMINAE: POLYDRUSITAE: PSALLIDIINI.

Type species: Curculio maxillosus Fabricius, 1792, by monotypy.

Family-group name: PSALIDIIDES Lacordaire, 1863. Stem: Psallidi-.

†Psammaegialia Nikolajev, B. Wang & Zhang, 2014: 221. Current status: valid genus in SCARABAEIDAE: APHODIINAE: †PSAMMAEGIALIINI. Note: Lower Cretaceous (China).

Type species: †Psammaegialia abdita Nikolajev, B. Wang & Zhang, 2014, by original designation.

Family-group name: †PSAMMAEGIALIINI Nikolajev, B. Wang & Zhang, 2014: 220. Stem: Psammaegiali-.

Psammobius Heer, 1841: 531. Current status: junior synonym of Psammodius Fallén, 1807 in SCARABAEIDAE: APHODIINAE: PSAMMODIINI: PSAMMODIINA.

Type species: Aphodius sulcicollis Illiger, 1802 (= Scarabaeus asper Fabricius, 1775), by subsequent designation (Tesař 1957: 168).

Family-group name: PSAMMOBIINA Reitter, 1909. Stem: Psammobi-. Note: junior homonym of PSAMMOBIIDAE Fleming, 1828 (type genus Psammobia Lamarck, 1818) in Mollusca.

Psammodius Fallén, 1807: 37. Current status: valid genus in SCARABAEIDAE: APHODIINAE: PSAMMODIINI: PSAMMODIINA.

Type species: Aphodius sulcicollis Illiger, 1802 (= Scarabaeus asper Fabricius, 1775), by subsequent designation (Curtis 1829: pl. 258).

Family-group name: PSAMMODIAIRES Mulsant, 1842. Stem: Psammodi-.

Psammoecus Latreille, 1829b: 135. Current status: valid genus in SILVANIDAE: BRONTINAE: TELEPHANINI.

Type species: Notoxus bipunctatus Fabricius, 1792, by monotypy.

Family-group name: PSAMMOECINI Reitter, 1879. Stem: Psammoec-.

Psathyrus J. Thomson, 1857: 192. Current status: valid genus in CERAMBYCIDAE: CERAMBYCINAE: AUXESINI.

Type species: Psathyrus aeolis J. Thomson, 1857, by monotypy.

Family-group name: PSATHYRINI Quentin, 1954. Stem: Psathyr-.

Psebium Pascoe, 1864b: 289. Current status: valid genus in CERAMBYCIDAE: CERAMBYCINAE: PSEBIINI.

Type species: Psebium brevipenne Pascoe, 1864, by monotypy.

Family-group name: PSÉBIIDES Lacordaire, 1868. Stem: Psebi-.

Pselaphus Herbst, 1791: 106. Current status: valid genus in STAPHYLINIDAE: PSELAPHINAE: PSELAPHITAE: PSELAPHINI.

Type species: Pselaphus heisei Herbst, 1791, by subsequent designation (Guérin-Méneville 1828: 316).

Family-group name: PSELAPHII Latreille, 1802. Stem: Pselaph-.

Pselaptus J.L. LeConte, 1881a: 184. Current status: valid genus in STAPHYLINIDAE: PSELAPHINAE: GONIACERITAE: BRACHYGLUTINI: BRACHYGLUTINA.

Type species: Pselaptus belfragei J.L. LeConte, 1881, by monotypy.

Family-group name: PSELAPTINA Park, 1976. Stem: Pselapt-.

Psenocerus J.L. LeConte, 1852a: 158. Current status: valid genus in CERAMBYCIDAE: LAMIINAE: DESMIPHORINI.

Type species: here fixed (under ICZN 1999a, Article 70.3.2) as Clytus supernotatus Say, 1824, misidentified as Callidium pini G.-A. Olivier, 1795 (sensu J.L. LeConte, 1852) in the original designation by monotypy in J.L. LeConte (1852a: 158).

Family-group name: PSENOCERINI J.L. LeConte, 1873. Stem: Psenocer-.

Psephenoides Gahan, 1914: 189. Current status: valid genus in PSEPHENIDAE: PSEPHENOIDINAE.

Type species: Psephenoides immsi Gahan, 1914, by original designation.

Family-group name: PSEPHENOIDINI Bollow, 1938. Stem: Psephenoid-.

Psephenus Haldeman in Melsheimer, 1853: 34. Current status: valid genus in PSEPHENIDAE: PSEPHENINAE. Note: new replacement name for Eurypalpus J.L. LeConte, 1850.

Type species: Eurypalpus lecontei J.L. LeConte, 1850 (= Fluvicola herricki DeKay, 1844), by monotypy (type fixation for Eurypalpus J.L. LeConte).

Family-group name: PSÉPHÉNIDES Lacordaire, 1854. Stem: Psephen-.

Psepholax A. White in A. White and Doubleday, 1843: 275. Current status: valid genus in CURCULIONIDAE: CRYPTORHYNCHINAE: PSEPHOLACINI.

Type species: Psepholax sulcatus A. White, 1843, by monotypy.

Family-group name: PSÉPHOLACIDES Lacordaire, 1865. Stem: Psepholac-.

Pseudacanthus Kaup, 1868b: 19. Current status: valid genus in PASSALIDAE: PASSALINAE: PROCULINI.

Type species: Passalus mexicanus Truqui, 1857, by subsequent designation (Gravely 1918: 30).

Family-group name: PSEUDACANTHEAE Kaup, 1871. Stem: Pseudacanth-.

Pseudacherusia Kerremans, 1905: 382. Current status: valid genus in BUPRESTIDAE: POLYCESTINAE: TYNDARIDINI: PSEUDACHERUSIINA.

Type species: Pseudacherusia theryi Kerremans, 1905, by monotypy.

Family-group name: PSEUDOACHERUSINI Cobos, 1980. Stem: Pseudacherusi-.

Pseudadoretus Semenov, 1889b: 202. Current status: valid genus in SCARABAEIDAE: RUTELINAE: ADORETINI: ADORETINA.

Type species: Pseudadoretus dilutellus Semenov, 1889, by original designation. Note: Adoretus phthisicus Dohrn, 1882 is sometimes cited as the type species of this genus (e.g., Zorn and A. Bezděk 2016: 320) but Semenov (1889b: 202) clearly designated P. dilutellus [as ditutellus] as type species; two spellings were originally used for the type species, including ditutellus (p. 202); as far as we know no one has acted as First Reviser (ICZN 1999a, Article 24.2.3) and we here select dilutellus as the correct spelling.

Family-group name: PSEUDADORETINA Ohaus, 1912. Stem: Pseudadoret-.

Pseudaplatopterus Kleine, 1940: 179. Current status: valid genus in LYCIDAE: EROTINAE: EROTINI.

Type species: Aplatopterus scheelei Kleine, 1940, by monotypy. Note: just after Kleine’s (1940: 179) description of Pseudaplatopterus, he described a new species under the name “A. Scheeli” in honor of Major Scheele; Kazantsev (2013c: 97) used the spelling “ascheelei” for the type species but we believe the “A.” is either an error for “P.” or stands as an abbreviation for Aplatopterus Reitter, 1911; we think the second option is more likely the correct one.

Family-group name: PSEUDAPLATOPTERINA Kazantsev, 2012b: 399. Stem: Pseudaplatopteri-.

Pseudapotrepus Champion, 1909: 21. Current status: valid genus in CURCULIONIDAE: COSSONINAE: PSEUDAPOTREPINI.

Type species: Pseudapotrepus macrophthalmus Champion, 1909, by original designation.

Family-group name: PSEUDAPOTREPIDES Champion, 1909. Stem: Pseudapotrep-.

Pseudauletes Voss, 1922a: 27, 95. Current status: valid genus in ATTELABIDAE: RHYNCHITINAE: RHYNCHITITAE: AULETINI: PSEUDAULETINA.

Type species: Rhynchites rufiventris Jekel, 1860, by subsequent designation (Legalov 2011e: 96).

Family-group name: PSEUDAULETINA Voss, 1933. Stem: Pseudaulet-.

Pseudeucinetus Heller, 1921: 155. Current status: valid genus in LIMNICHIDAE: THAUMASTODINAE.

Type species: Pseudeucinetus zygops Heller, 1921, by original designation.

Family-group name: PSEUDEUCINETINI Csiki, 1924. Stem: Pseudeucinet-.

Pseudobagous Sharp, 1917: 27. Current status: valid genus in CURCULIONIDAE: BAGOINAE.

Type species: Bagous longulus Gyllenhal, 1836, by original designation.

Family-group name: PSEUDOBAGOINI Sharp, 1917. Stem: Pseudobago-.

Pseudobalaninus Faust, 1889: 97. Current status: valid genus in CURCULIONIDAE: CURCULIONINAE: CURCULIONINI: PSEUDOBALANININA.

Type species: Pseudobalaninus semifasciatus Faust, 1889, by subsequent designation (Heller 1925b: 93).

Family-group name: PSEUDOBALANININA Heller, 1925. Stem: Pseudobalanin-.

†Pseudobrienia Legalov, 2012: 72. Current status: valid genus in †OBRIENIIDAE: †PSEUDOBRIENIINAE. Note: Upper Jurassic (Kazakhstan).

Type species: †Pseudobrienia rasnitsyni Legalov, 2012, by original designation.

Family-group name: †PSEUDOBRIENIINAE Legalov, 2012: 73. Stem: Pseudobrieni-.

Pseudoceocephalus Kleine, 1920: 18. Current status: valid genus in BRENTIDAE: BRENTINAE: TRACHELIZINI: PSEUDOCEOCEPHALINA.

Type species: Brentus depressus Lund, 1802, by original designation.

Family-group name: PSEUDOCEOCEPHALIDAE Kleine, 1922. Stem: Pseudoceocephal-.

Pseudocephalus Newman, 1842b: 353. Current status: valid genus in CERAMBYCIDAE: CERAMBYCINAE: PSEUDOCEPHALINI.

Type species: Pseudocephalus formicides Newman, 1842, by monotypy.

Family-group name: PSEUDOCEPHALINI Aurivillius, 1912. Stem: Pseudocephal-.

Pseudochodaeus Carlson & Ritcher, 1974: 99. Current status: valid genus in OCHODAEIDAE: CHAETOCANTHINAE: PSEUDOCHODAEINI.

Type species: Ochodaeus estriatus Schaeffer, 1906, by original designation.

Family-group name: PSEUDOCHODAEINI Scholtz, 1988. Stem: Pseudochodae-.

Pseudocistela Crotch, 1874b: 108. Current status: valid genus in TENEBRIONIDAE: ALLECULINAE: ALLECULINI: GONODERINA.

Type species: Cistela brevis Say, 1824, by subsequent designation (Novák and Pettersson 2008: 327).

Family-group name: PSEUDOCISTELINI Portevin, 1934. Stem: Pseudocistel-.

Pseudocneorhinus Roelofs, 1873: 177. Current status: valid genus in CURCULIONIDAE: ENTIMINAE: CYPHICERITAE: TRACHYPHLOEINI: PSEUDOCNEORHININA.

Type species: Pseudocneorhinus obesus Roelofs, 1873, by monotypy.

Family-group name: PSEUDOCNEORRHININI Kôno, 1930. Stem: Pseudocneorhin-.

Pseudocolaspis Laporte, 1833a: 19, 23. Current status: valid genus in CHRYSOMELIDAE: EUMOLPINAE: BROMIINI.

Type species: Pseudocolaspis coerulea Laporte, 1833, by subsequent designation (Baly 1878a: 248).

Family-group name: PSEUDOCOLASPITES Chapuis, 1874. Stem: Pseudocolaspid-.

Pseudocrioceris Pic, 1916b: 3. Current status: valid genus in CHRYSOMELIDAE: CRIOCERINAE: PSEUDOCRIOCERINI.

Type species: Brachydactyla annulipes Pic, 1913, by present designation. Note: Heinze (1962: 201) listed Crioceris discoidea Guérin-Méneville, 1844 as the type species of Pseudocrioceris Pic, 1916 but this is not an originally included nominal species.

Family-group name: PSEUDOCRIOCERINI Heinze, 1962. Stem: Pseudocriocer-.

Pseudocycnotrachelus Legalov, 2003: 525. Current status: valid genus in ATTELABIDAE: APODERINAE: APODERINI.

Type species: Apoderus ledyardi Heller, 1915, by original designation.

Family-group name: PSEUDOCYCNOTRACHELINA Legalov, 2003. Stem: Pseudocycnotrachel-.

Pseudocyphus Schaeffer, 1905: 179. Current status: junior synonym of Platyomus C.R. Sahlberg, 1823 in CURCULIONIDAE: ENTIMINAE: POLYDRUSITAE: NAUPACTINI: NAUPACTINA.

Type species: Pseudocyphus flexicaulis Schaeffer, 1905, by monotypy.

Family-group name: PSEUDOCYPHI Pierce, 1913. Stem: Pseudocyph-.

Pseudodorcus Parry, 1870: 94. Current status: valid genus in LUCANIDAE: LUCANINAE: LUCANINI.

Type species: Lucanus hydrophiloides Hope, 1845, by monotypy.

Family-group name: PSEUDODORCINI Benesh, 1960. Stem: Pseudodorc-.

Pseudolampsis G.H. Horn, 1889: 174. Current status: valid genus in CHRYSOMELIDAE: GALERUCINAE: ALTICINI.

Type species: Hypolampsis guttatus J.L. LeConte, 1884, by monotypy.

Family-group name: PSEUDOLAMPSES G.H. Horn, 1889. Stem: Pseudolampse-.

Pseudoleptura J. Thomson, 1861: 142, 148. Current status: junior synonym of Erythrus A. White, 1853 in CERAMBYCIDAE: CERAMBYCINAE: PSEUDOLEPTURINI. Note: unnecessary new replacement name for Erythrus A. White, 1853.

Type species: Erythrus championi A. White, 1853, by original designation.

Family-group name: PSEUDOLEPTURITAE J. Thomson, 1861. Stem: Pseudoleptur-.

Pseudoliodes Portevin, 1926: 78. Current status: junior synonym of Pseudcolenis Reitter, 1885 in LEIODIDAE: LEIODINAE: PSEUDOLIODINI.

Type species: Pseudcolenis grandis Portevin, 1905, by original designation.

Family-group name: PSEUDOLIODINI Portevin, 1926. Stem: Pseudoliod-.

Pseudolotelus Pic, 1901: 67. Current status: valid genus in ADERIDAE.

Type species: Euglenes punctatissimus Reitter, 1885, by subsequent designation (Nardi 2007: 29).

Family-group name: PSEUDOLOTELINA Báguena Corella, 1948. Stem: Pseudolotel-.

Pseudomasoreus Desbrochers des Loges, 1904: 140, 163. Current status: valid genus in CARABIDAE: HARPALINAE: LEBIINI: CYMINDIDINA.

Type species: Cymindis canigoulensis Fairmaire & Laboulbène, 1854, by monotypy.

Family-group name: PSEUDOMASOREINI Jeannel, 1942. Stem: Pseudomasore-.

Pseudomelanactes Mathieu, 1961: 476. Current status: junior synonym of Anthracalaus Fairmaire, 1889 in ELATERIDAE: AGRYPNINAE: PSEUDOMELANACTINI.

Type species: Melanactes agrypnoides Van Dyke, 1932, by original designation.

Family-group name: PSEUDOMELANACTINI Arnett, 1967. Stem: Pseudomelanact-.

Pseudomenes Fleutiaux, 1902: 663. Current status: valid genus in EUCNEMIDAE: PSEUDOMENINAE. Note: new replacement name for Eumenes Bonvouloir, 1871 [cited by Fleutiaux as 1875].

Type species: Eumenes bakewelli Bonvouloir, 1872, by subsequent monotypy (type fixation for Eumenes Bonvouloir).

Family-group name: PSEUDOMENINAE Muona, 1993. Stem: Pseudomen-.

Pseudomesauletes Legalov, 2001b: 53. Current status: valid genus in ATTELABIDAE: RHYNCHITINAE: RHYNCHITITAE: AULETINI: PSEUDOMESAULETINA.

Type species: Auletes uniformis Roelofs, 1875, by original designation.

Family-group name: PSEUDOMESAULETINA Legalov, 2003. Stem: Pseudomesaulet-.

Pseudometrioxena Legalov, 2023b: 39. Current status: valid genus in BELIDAE: OXYCORYNINAE: OXYCORYNITAE: METRIOXENINI: PSEUDOMETRIOXENINA.

Type species: Metrioxena decisa Pascoe, 1885, by original designation.

Family-group name: PSEUDOMETRIOXENINA Legalov, 2023: 39. Stem: Pseudometrioxen-.

Pseudomimus Hartmann, 1904: 416. Current status: valid genus in CURCULIONIDAE: COSSONINAE: RHYNCOLINI: PSEUDOMIMINA.

Type species: Pseudomimus corpulentus Hartmann, 1904, by monotypy.

Family-group name: PSEUDOMIMINI Voss, 1939. Stem: Pseudomim-.

Pseudomorpha W. Kirby, 1823a: 98. Current status: valid genus in CARABIDAE: HARPALINAE: PSEUDOMORPHINI. Note: W. Kirby originally used two different spellings for this taxon, including Heteromorpha (p. 109); subsequently, W. Kirby (1823b: 420) used only Pseudomorpha and therefore acted as First Reviser (ICZN 1999a, Article 24.2.4).

Type species: Pseudomorpha excrucians W. Kirby, 1823, by monotypy.

Family-group name: HETEROMORPHIDAE Hope, 1838. Stem: Pseudomorph-.

Pseudoperinthus Wasmann, 1916a: 194. Current status: valid genus in STAPHYLINIDAE: ALEOCHARINAE: PSEUDOPERINTHINI.

Type species: Pseudoperinthus malayanus Wasmann, 1916, by monotypy.

Family-group name: PSEUDOPERINTHINAE Cameron, 1939. Stem: Pseudoperinth-.

Pseudoperotis Obenberger, 1936: 115. Current status: valid genus in BUPRESTIDAE: CHRYSOCHROINAE: DICERCINI: PSEUDOPEROTINA.

Type species: Oedisterna scabrosula Obenberger, 1936, by original designation.

Family-group name: PSEUDOPEROTINA Tôyama, 1987. Stem: Pseudoperot-.

Pseudophanus J.L. LeConte, 1859a: 84. Current status: junior synonym of Cryptomorpha Wollaston, 1854 in SILVANIDAE: BRONTINAE: TELEPHANINI.

Type species: Pseudophanus signatus J.L. LeConte, 1859 (= Psammoecus desjardinsii Guérin- Méneville, 1844), by monotypy.

Family-group name: PSEUDOPHANINI J.L. LeConte, 1861. Stem: Pseudophan-. Note: junior homonym of PSEUDOPHANINAE Amyot & Audinet-Serville, 1843 (type genus Pseudophana Burmeister, 1835) in Hemiptera.

Pseudophengodes Pic, 1930: 319. Current status: valid genus in PHENGODIDAE: PHENGODINAE.

Type species: Phengodes pulchella Guérin-Méneville, 1843, by subsequent designation (Wittmer 1976b: 425).

Family-group name: PSEUDOPHENGODIDAE Pic, 1930. Stem: Pseudophengod-.

Pseudophrysus Legalov, 2003: 479. Current status: valid genus in ATTELABIDAE: APODERINAE: CLITOSTYLINI: PSEUDOPHRYSINA.

Type species: Cycnotrachelus parvulus Voss, 1928, by original designation.

Family-group name: PSEUDOPHRYSINA Legalov, 2003. Stem: Pseudophrys-.

Pseudoplandria Fenyes, 1921: 30. Current status: valid genus in STAPHYLINIDAE: ALEOCHARINAE: HOPLANDRIINI: PSEUDOPLANDRIINA.

Type species: Pseudoplandria laeta Fenyes, 1921, by original designation.

Family-group name: PSEUDOPLANDRIINA Hanley, 2002. Stem: Pseudoplandri-.

Pseudopsis Newman, 1834: 313. Current status: valid genus in STAPHYLINIDAE: PSEUDOPSINAE.

Type species: Pseudopsis sulcatus Newman, 1834, by monotypy.

Family-group name: PSEUDOPSINI Ganglbauer, 1895. Stem: Pseudops-.

Pseudoryctes Sharp, 1873: 267. Current status: valid genus in SCARABAEIDAE: DYNASTINAE: PENTODONTINI: PSEUDORYCTINA.

Type species: Oryctes mullerianus A. White, 1859, by monotypy.

Family-group name: PSEUDORYCTINA Carne, 1957. Stem: Pseudoryct-.

Pseudotrechus Rosenhauer, 1856: 22. Current status: valid genus in CARABIDAE: HARPALINAE: LEBIINI: PSEUDOTRECHINA.

Type species: Pseudotrechus mutilatus Rosenhauer, 1856, by monotypy.

Family-group name: PSEUDOTRECHINI Basilewsky, 1984. Stem: Pseudotrech-.

Pseudozaena Laporte, 1834b: 55. Current status: valid genus in CARABIDAE: PAUSSINAE: OZAENINI.

Type species: Ozaena megacephala Laporte, 1834 (= Ozaena orientalis Klug, 1834), by monotypy.

Family-group name: PSEUDOZAENINI Sloane, 1905. Stem: Pseudozaen-.

Psilapha Clark, 1865: 377, 389. Current status: valid genus in CHRYSOMELIDAE: GALERUCINAE: ALTICINI.

Type species: Psilapha flava Clark, 1865, by monotypy.

Family-group name: PSILAPHINA Bechyné & Špringlová de Bechyné, 1973. Stem: Psilaph-.

Psilocladus Blanchard, 1845b: 121. Current status: valid genus in LAMPYRIDAE: PSILOCLADINAE.

Type species: Psilocladus miltoderus Blanchard, 1845, by monotypy.

Family-group name: PSILOCLADINA McDermott, 1964. Stem: Psiloclad-.

Psilomorpha Saunders, 1850: 80. Current status: valid genus in CERAMBYCIDAE: CERAMBYCINAE: PSILOMORPHINI.

Type species: Psilomorpha tenuipes Saunders, 1850, by monotypy.

Family-group name: PSILOMORPHIDES Lacordaire, 1868. Stem: Psilomorph-.

Psilonychus Burmeister, 1855: 288. Current status: valid genus in SCARABAEIDAE: MELOLONTHINAE: SCHIZONYCHINI.

Type species: Psilonychus ecklonii Burmeister, 1855, by present designation.

Family-group name: PSILONYCHIDES Péringuey, 1904. Stem: Psilonych-.

Psiloptera Dejean, 1833a: 76. Current status: valid genus in BUPRESTIDAE: CHRYSOCHROINAE: DICERCINI: DICERCINA.

Type species: Buprestis attenuata Fabricius, 1792, by subsequent designation (J. Thomson 1878: 29).

Family-group name: PSILOPTÉRIDES Lacordaire, 1857. Stem: Psilopter-. Note: senior homonym of PSILOPTERINAE Dolgopol de Saez, 1927 (type genus Psilopterus Moreno & Mercerat, 1891) in Aves.

Psoa Herbst, 1797: 214. Current status: valid genus in BOSTRICHIDAE: PSOINAE: PSOINI.

Type species: Psoa viennensis Herbst, 1797, by monotypy.

Family-group name: PSOITAS Blanchard, 1851. Stem: Pso-.

Psydrus J.L. LeConte, 1846: 153. Current status: valid genus in CARABIDAE: PSYDRINAE: PSYDRINI.

Type species: Psydrus piceus J.L. LeConte, 1846, by monotypy.

Family-group name: PSYDRI J.L. LeConte, 1853. Stem: Psydr-.

Psylliodes Latreille, 1829b: 154. Current status: valid genus in CHRYSOMELIDAE: GALERUCINAE: ALTICINI.

Type species: Galeruca affinis Paykull, 1799, by subsequent designation (Blanchard in Audouin et al. 1844: pl. 73). Note: Chrysomela chrysocephala Linnaeus, 1758 is the type species usually listed for this genus (e.g., Konstantinov and Vanderberg 1996: 373; Döberl 2010: 552) by subsequent designation of Maulik (1926: 144) but Blanchard’s designation is earlier; both species are currently included in the nominotypical subgenus of Psylliodes Latreille.

Family-group name: PSYLLIODITES Chapuis, 1875. Stem: Psylliod-.

Psyllobora Chevrolat in Dejean, 1836a: 434. Current status: valid genus in COCCINELLIDAE: COCCINELLINAE: COCCINELLINI.

Type species: Coccinella lineola Fabricius, 1792 (= Psyllobora fabricii Crotch, 1871), by subsequent designation (Timberlake 1943: 41).

Family-group name: PSYLLOBORINI Casey, 1899. Stem: Psyllobor-.

Ptenidium Erichson, 1845a: 34. Current status: valid genus in PTILIIDAE: PTILIINAE: PTENIDIINI. Note: name given precedence over Anisarthria Stephens, 1830 whenever the two are considered synonyms and placed on the Official List of Generic Names in Zoology (ICZN 1984b).

Type species: Scaphidium pusillum Gyllenhal, 1808, by subsequent designation (C.G. Thomson 1859: 63; see ICZN 1984b). Note: given precedence over Dermestes melas Marsham, 1802 whenever the two are considered synonyms and placed on the Official List of Specific Names in Zoology (ICZN 1984b).

Family-group name: PTENIDIINI Flach, 1889. Stem: Ptenidi-.

Pteracantha Newman, 1838: 392. Current status: valid genus in CERAMBYCIDAE: CERAMBYCINAE: TRACHYDERINI: TRACHYDERINA.

Type species: Pteracantha fasciata Newman, 1838, by monotypy.

Family-group name: PTERACANTHITAE J. Thomson, 1864. Stem: Pteracanth-.

Pteracmes Raffray, 1890a: 95, 102. Current status: valid genus in STAPHYLINIDAE: PSELAPHINAE: EUPLECTITAE: TRICHONYCHINI: TRICHONYCHINA.

Type species: Pteracmes schaufussi Raffray, 1890, by monotypy.

Family-group name: PTERACMINI Jeannel, 1962. Stem: Pteracmet-.

Ptericoptus Lacordaire, 1830b: 185. Current status: valid genus in CERAMBYCIDAE: LAMIINAE: APOMECYNINI.

Type species: Ptericoptus dorsalis Audinet-Serville, 1835, by subsequent monotypy (Audinet- Serville 1835a: 61).

Family-group name: PTÉRICOPTIDES Lacordaire, 1872. Stem: Ptericopt-.

Pterobothris Fairmaire & Germain, 1859: 714. Current status: valid genus in BUPRESTIDAE: BUPRESTINAE: PTEROBOTHRINI.

Type species: Pterobothris corrosus Fairmaire & Germain, 1859, by monotypy.

Family-group name: PTEROBOTHRINI Volkovitsh, 2001. Stem: Pterobothr-.

Pterocolus Schönherr in Say, 1831: 5. Current status: valid genus in ATTELABIDAE: PTEROCOLINAE. Note: see Prena (2018: 331) for the authorship of this genus-group name.

Type species: Attelabus ovatus Fabricius, 1801, by monotypy.

Family-group name: PTÉROCOLIDES Lacordaire, 1865. Stem: Pterocol-.

Pterogenius Candèze, 1861: 363. Current status: valid genus in PTEROGENIIDAE.

Type species: Pterogenius nietneri Candèze, 1861, by monotypy.

Family-group name: PTEROGENIIDAE Crowson, 1953. Stem: Pterogeni-.

Pteroloma Gyllenhal, 1827: 418. Current status: valid genus in AGYRTIDAE: PTEROLOMATINAE.

Type species: Harpalus forsstromii Gyllenhal, 1810, by monotypy.

Family-group name: PTEROLOMINA C.G. Thomson, 1862. Stem: Pterolomat-.

Pteronius Blackwelder, 1952: 331. Current status: junior synonym of Proteinus Latreille, 1797 in STAPHYLINIDAE: PROTEININAE: PROTEININI.

Type species: Dermestes brachypterus Fabricius, 1792, by original designation.

Family-group name: PTERONINI Arnett, 1961. Stem: Pteroni-.

Pteroplatus Buquet, 1840b: 287. Current status: valid genus in CERAMBYCIDAE: CERAMBYCINAE: PTEROPLATINI.

Type species: Pteroplatus pulcher Buquet, 1840, by subsequent designation (J. Thomson 1864: 258).

Family-group name: PTEROPLATITAE J. Thomson, 1861. Stem: Pteroplat-.

Pteroplius Lacordaire, 1830b: 183. Current status: valid genus in CERAMBYCIDAE: LAMIINAE: PTEROPLIINI. Note: the original spelling was Pterhoplius, but Pteroplius, an incorrect subsequent spelling, is in prevailing usage, attributed to the publication of the original spelling, and therefore deemed to be the original spelling (ICZN 1999a, Article 33.3.1).

Type species: Pteroplius acuminatus Audinet-Serville, 1835, by subsequent monotypy. Note: Audinet-Serville (1835a: 66) included two species in the genus Pteroplius, one in the section ‘propriè dictus’ and one in the section Rhaphiptera, a new genus-group name; the type species of the genus is the species cited in the first section.

Family-group name: PTEROPLIITAE J. Thomson, 1860. Stem: Pteropli-.

Pterosthetops Perkins in Perkins and J. Balfour-Browne, 1994: 17. Current status: valid genus in HYDRAENIDAE: PROSTHETOPINAE: PROSTHETOPINI.

Type species: Pterosthetops impressus Perkins & J. Balfour-Browne, 1994, by original designation.

Family-group name: PTEROSTHETOPINI Perkins, 1994. Stem: Pterosthetop-.

Pterostichus Bonelli, 1810: Tab. Syn. Current status: valid genus in CARABIDAE: HARPALINAE: PTEROSTICHINI: PTEROSTICHINA.

Type species: Carabus fasciatopunctatus Creutzer, 1799, by subsequent designation (Curtis 1828: pl. 196). Note: no species were originally included in this genus by Bonelli (1810); the first ones were the four species listed by Panzer (1813: 70).

Family-group name: PTEROSTICHII Bonelli, 1810. Stem: Pterostich-.

Pterotarsus Guérin-Méneville, 1829a: pl. 12. Current status: valid genus in ELATERIDAE: THYLACOSTERNINAE.

Type species: Pterotarsus histrio Guérin-Méneville, 1829, by monotypy.

Family-group name: PTEROTARSINI Fleutiaux, 1902. Stem: Pterotars-.

Pterotus J.L. LeConte, 1859a: 86. Current status: valid genus in LAMPYRIDAE: PTEROTINAE.

Type species: Pterotus obscuripennis J.L. LeConte, 1859, by monotypy.

Family-group name: PTEROTINI J.L. LeConte, 1861. Stem: Pterot-.

Ptichopus Kaup, 1868b: 20. Current status: valid genus in PASSALIDAE: PASSALINAE: PASSALINI.

Type species: Passalus angulatus Percheron, 1834, by monotypy.

Family-group name: PTICHOPINAE Kuwert, 1896. Stem: Ptichopod-.

Ptilineurus Reitter, 1902: 24. Current status: valid genus in PTINIDAE: DRYOPHILINAE: PTILINEURINI.

Type species: Ptilinus marmoratus Reitter, 1877, by monotypy.

Family-group name: PTILINEURINI Böving, 1927. Stem: Ptilineur-.

Ptilinus Geoffroy, 1762: 64. Current status: valid genus in PTINIDAE: PTILININAE: PTILININI. Note: name placed on the Official List of Generic Names in Zoology (ICZN 1994a).

Type species: Ptilinus fuscus Geoffroy, 1785, by subsequent monotypy (O.F. Müller 1776: 81), as Ptilinus cylindricus O.F. Müller, 1776, the name of the senior subjective synonym of P. fuscus Geoffroy that has been suppressed for the purposes of both Principles of Priority and Homonymy; see ICZN 1994a. Note: Ptilinus fuscus Geoffroy, 1785 was placed on the Official List of Specific Names in Zoology (ICZN 1994a).

Family-group name: PTILINIDAE Shuckard, 1839. Stem: Ptilin-.

Ptilium Gyllenhal, 1827: 292. Current status: valid genus in PTILIIDAE: PTILIINAE: PTILIINI. Note: name placed on the Official List of Generic Names in Zoology (ICZN 1984b).

Type species: Ptilium caesum Erichson, 1845, by plenary power (see ICZN 1984b). Note: name placed on the Official List of Specific Names in Zoology (ICZN 1984b).

Family-group name: PTILINA Erichson, 1845. Stem: Ptili-.

Ptilodactyla Illiger, 1807b: 342. Current status: valid genus in PTILODACTYLIDAE: PTILODACTYLINAE.

Type species: Ptilodactyla elaterina Illiger, 1807 (= Pyrochroa nitida DeGeer, 1775), by monotypy.

Family-group name: PTILODACTYLIDAE Laporte, 1838. Stem: Ptilodactyl-.

Ptilophorus Dejean, 1834: 218. Current status: valid genus in RIPIPHORIDAE: PTILOPHORINAE.

Type species: Pelecotoma dufourii Latreille, 1818, by monotypy.

Family-group name: PTILOPHORINI Gerstaecker, 1855. Stem: Ptilophor-.

Ptilopterium Gistel, 1848b: x. Current status: junior synonym of Ptilium Gyllenhal, 1827 in PTILIIDAE: PTILIINAE: PTILIINI. Note: unnecessary new replacement name for Ptilium Gyllenhal, 1827.

Type species: Ptilium caesum Erichson, 1845, by plenary power (see ICZN 1984b) (type fixation for Ptilium Gyllenhal).

Family-group name: PTILOPTERIIDAE Gistel, 1848. Stem: Ptilopteri-.

Ptinella Motschulsky, 1845a: 819. Current status: valid genus in PTILIIDAE: PTILIINAE: PTINELLINI. Note: this genus is usually credited to Motschulsky (“1844”/1845a: 819) in the literature, referring to volume 17 (part 4) of the ‘Bulletin de la Société Impériale des Naturalistes de Moscou’, which was issued by 27 January 1845 (Bousquet and Davies 2021: 15); however, it is possible that Ptinella was first made available in the December issue of the ‘Revue Zoologique’ for the year 1844, which was received by the Académie des Sciences of France, also on 27 January 1845 (see Motschulsky, 1844b: 446); Ptinella Motschulsky, 1845 was placed on the Official List of Generic Names in Zoology (ICZN 1985b); we prefer to use the data as it appears on the Official List until the date priority of the two publications can be ascertained.

Type species: Ptilium apterum Guérin-Méneville, 1839, by plenary power (see ICZN 1985b). Note: name placed on the Official List of Specific Names in Zoology (ICZN 1985b).

Family-group name: PTINELLINI Reitter, 1906. Stem: Ptinell-.

Ptinus Linnaeus, 1767: 537, 565. Current status: valid genus in PTINIDAE: PTININAE: PTININI. Note: name placed on the Official List of Generic Names in Zoology (ICZN 1995b).

Type species: Cerambyx fur Linnaeus, 1758, by subsequent designation (Latreille 1810: 427; see ICZN 1995b). Note: name placed on the Official List of Specific Names in Zoology (ICZN 1995b).

Family-group name: PTINIORES Latreille, 1802. Stem: Ptin-.

†Ptisma Kirejtshuk & Azar in Kirejtshuk et al., 2016a: 204. Current status: valid genus in CLAMBIDAE. Note: Lower Cretaceous (Lebanon).

Type species: †Ptisma zasukhae Kirejtshuk & Azar, 2016, by original designation. Note: two original spellings were used in Kirejtshuk et al. (2016a): P. zhasukhai (p. 204) and P. zasukhae (pp. 201–209); Kirejtshuk et al. (2016b: 275) acted as First Revisers (ICZN 1999a, Article 24.2.4) and selected P. zasukhae.

Family-group name: †PTISMIDAE Kirejtshuk & Azar in Kirejtshuk et al., 2016a: 202. Stem: Ptism-.

Ptochus Schönherr, 1826: 187. Current status: valid genus in CURCULIONIDAE: ENTIMINAE: CYPHICERITAE: CYPHICERINI: MYLLOCERINA. Note: name placed on the Official List of Generic Names in Zoology (ICZN 1990d).

Type species: Ptochus porcellus Boheman, 1834, by subsequent designation (Marshall 1916: 259; see ICZN 1990d). Note: name placed on the Official List of Specific Names in Zoology (ICZN 1990d).

Family-group name: PTOCHINI Reitter, 1913. Stem: Ptoch-.

Ptomaphaginus Portevin, 1914: 194. Current status: valid genus in LEIODIDAE: CHOLEVINAE: PTOMAPHAGINI: PTOMAPHAGININA.

Type species: Ptomaphaginus longitarsis Portevin, 1914, by monotypy.

Family-group name: PTOMAPHAGININI Szymczakowski, 1964. Stem: Ptomaphagin-.

Ptomaphagus Hellwig, 1795: 358. Current status: valid genus in LEIODIDAE: CHOLEVINAE: PTOMAPHAGINI: PTOMAPHAGINA.

Type species: Tritoma sericea Fabricius, 1787 (= Silpha subvillosa Goeze, 1777), by monotypy. Note: Goeze’s (1777) work is not consistently binominal (see ICZN 2021a) and therefore an application to the Commission is necessary to retain Silpha subvillosa Goeze, 1777 as the valid name for the type species.

Family-group name: PTOMAPHAGINI Jeannel, 1911. Stem: Ptomaphag-.

Ptosima Dejean, 1833a: 79. Current status: valid genus in BUPRESTIDAE: POLYCESTINAE: PTOSIMINI.

Type species: here fixed (under ICZN 1999a, Article 70.3.2) as Buprestis undecimmaculata Herbst, 1784, misidentified as Buprestis novemmaculata Linnaeus, 1767 (sensu Fabricius, 1775) in the original designation by monotypy in Dejean (1833a: 79).

Family-group name: PTOSIMITES Kerremans, 1903. Stem: Ptosim-.

Ptychoderes Schönherr, 1823: 1135. Current status: valid genus in ANTHRIBIDAE: ANTHRIBINAE: PTYCHODERINI.

Type species: Macrocephalus nebulosus G.-A. Olivier, 1800, by original designation.

Family-group name: PTYCHODERIDAE Jekel, 1855. Stem: Ptychoder-. Note: senior homonym of PTYCHODERIDAE Spengel, 1893 (type genus Ptychodera Eschscholtz, 1825) in Enteropneusta (acorn worms).

Ptychodes Audinet-Serville, 1835a: 74. Current status: valid genus in CERAMBYCIDAE: LAMIINAE: LAMIINI.

Type species: Ptychodes politus Audinet-Serville, 1835, by monotypy.

Family-group name: PTYCHODES J.L. LeConte, 1873. Stem: Ptychod-.

Ptyopteryx Reichardt & C. Costa, 1967: 13. Current status: invalid senior synonym of Iapir Py-Daniel, Fonseca & Barbosa, 1993 in TORRIDINCOLIDAE: TORRIDINCOLINAE. Note: junior homonym of Ptyopteryx Kolenati, 1848 [Trichoptera].

Type species: Ptyopteryx britskii Reichardt & C. Costa, 1967, by original designation.

Family-group name: PTYOPTERINAE M. Abdullah, 1974. Stem: Ptyopteryg-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

Pulicomorpha Mann, 1924: 87. Current status: valid genus in STAPHYLINIDAE: ALEOCHARINAE: CREMATOXENINI.

Type species: Pulicomorpha coecum Mann, 1924, by original designation.

Family-group name: PULICOMORPHINI Mann, 1924. Stem: Pulicomorph-.

Purpuricenus Dejean, 1821: 105. Current status: valid genus in CERAMBYCIDAE: CERAMBYCINAE: TRACHYDERINI: TRACHYDERINA.

Type species: Cerambyx kaehleri Linnaeus, 1758, by subsequent designation (Blanchard in Audouin et al. 1843: pl. 66).

Family-group name: PURPURICENITAE J. Thomson, 1861. Stem: Purpuricen-.

Pycnocerus Westwood, 1842c: 67. Current status: valid genus in TENEBRIONIDAE: LAGRIINAE: PYCNOCERINI. Note: new replacement name for Pachylocerus Hope, 1841.

Type species: Pachylocerus westermanni Hope, 1841, by monotypy (type fixation for Pachylocerus Hope).

Family-group name: PYCNOCÉRIDES Lacordaire, 1859. Stem: Pycnocer-.

Pycnomerus Erichson, 1842: 214. Current status: valid genus in ZOPHERIDAE: ZOPHERINAE: PYCNOMERINI.

Type species: Ips terebrans G.-A. Olivier, 1790, by subsequent designation (Chevrolat 1847b: 644).

Family-group name: PYCNOMERINI Erichson, 1845. Stem: Pycnomer-.

Pygomolpus Bechyné, 1949: 533. Current status: valid genus in CHRYSOMELIDAE: EUMOLPINAE: PYGOMOLPINI.

Type species: Pygomolpus opacus Bechyné, 1949, by monotypy.

Family-group name: PYGOMOLPINI Bechyné, 1949. Stem: Pygomolp-.

Pygostenus Kraatz, 1858b: 361. Current status: valid genus in STAPHYLINIDAE: ALEOCHARINAE: PYGOSTENINI.

Type species: Pygostenus microcerus Kraatz, 1858, by monotypy.

Family-group name: PYGOSTENINI Fauvel, 1899. Stem: Pygosten-.

Pygoxyon Reitter in Leder, 1881: 508. Current status: valid genus in STAPHYLINIDAE: PSELAPHINAE: GONIACERITAE: PYGOXYINI.

Type species: Pygoxyon scydmaeniforme Reitter, 1881, by monotypy.

Family-group name: PYGOXYINI Reitter, 1909. Stem: Pygoxy-.

Pyrestes Pascoe, 1857: 96. Current status: valid genus in CERAMBYCIDAE: CERAMBYCINAE: PYRESTINI.

Type species: Pyrestes haematicus Pascoe, 1857, by subsequent designation (J. Thomson 1864: 159).

Family-group name: PYRESTHIDES Lacordaire, 1868. Stem: Pyrest-.

Pyrochroa Geoffroy, 1762: 338. Current status: valid genus in PYROCHROIDAE: PYROCHROINAE. Note: name placed on the Official List of Generic Names in Zoology (ICZN 1994a).

Type species: Cantharis coccinea Linnaeus, 1761, by subsequent designation (Westwood 1838c: 30; see ICZN 1994a). Note: name placed on the Official List of Specific Names in Zoology (ICZN 1994a).

Family-group name: PYROCHROIDES Latreille, 1806. Stem: Pyrochro-.

Pyrodes Audinet-Serville, 1832: 186. Current status: valid genus in CERAMBYCIDAE: PRIONINAE: MALLASPINI.

Type species: Prionus speciosus G.-A. Olivier, 1795 (= Prionus nitidus Fabricius, 1787), by subsequent designation (E. Desmarest 1860: 308).

Family-group name: PYRODINI Harold, 1879. Stem: Pyrod-.

Pyrophorus Billberg, 1820: 20. Current status: valid genus in ELATERIDAE: AGRYPNINAE: PYROPHORINI: PYROPHORINA.

Type species: Elater noctilucus Linnaeus, 1758, by subsequent designation (J. Meyer 1850: 120).

Family-group name: PYROPHORITES Candèze, 1863. Stem: Pyrophor-.

Pyropus Schönherr, 1836: 641. Current status: valid genus in CURCULIONIDAE: CURCULIONINAE: PYROPINI.

Type species: Pyropus sapphirinus Gyllenhal, 1836, by original designation.

Family-group name: PYROPIDES Lacordaire, 1865. Stem: Pyrop-.

Pyrota Dejean, 1834: 224. Current status: valid genus in MELOIDAE: MELOINAE: PYROTINI.

Type species: Lytta dispar Germar, 1823, by subsequent designation (Aksentjev 1988: 578). Note: the generic assignment of the type species was given as Tetraonyx, which was changed in the corrigenda (Germar, 1823: 623) to Lytta.

Family-group name: PYROTINI MacSwain, 1956. Stem: Pyrot-.

Pytheus Newman, 1840c: 14. Current status: junior synonym of Brachytria Newman, 1840 in CERAMBYCIDAE: CERAMBYCINAE: CERTALLINI.

Type species: Pytheus jugosus Newman, 1840, by monotypy.

Family-group name: PYTHEITAE J. Thomson, 1864. Stem: Pythe-.

Pythiopus C. Koch, 1953: 245. Current status: valid genus in TENEBRIONIDAE: BLAPTINAE: DENDARINI: DENDARINA.

Type species: Pythiopus cornutipectus C. Koch, 1953, by original designation.

Family-group name: PYTHIOPINI C. Koch, 1953. Stem: Pythiop-.

Pytho Latreille, 1797: 23. Current status: valid genus in PYTHIDAE.

Type species: Cucujus coeruleus Fabricius, 1787 (= Tenebrio depressus Linnaeus, 1767), by subsequent designation (Latreille 1810: 429). Note: no species were originally included in this genus by Latreille (1797); the first ones were the three species listed by Fabricius (1801b: 95, 96).

Family-group name: PYTHITES Solier, 1834. Stem: Pyth-.

Pyxidicerus Motschulsky, 1863: 422. Current status: valid genus in STAPHYLINIDAE: PSELAPHINAE: EUPLECTITAE: BYTHINOPLECTINI: PYXIDICERINA.

Type species: Pyxidicerus castaneus Motschulsky, 1863, by monotypy.

Family-group name: PYXIDICERINI Raffray, 1904. Stem: Pyxidicer-.

Quasimus Gozis, 1886: 22. Current status: valid genus in ELATERIDAE: NEGASTRIINAE: QUASIMUSINI: QUASIMUSINA.

Type species: Elater minutissimus Germar, 1822, by monotypy.

Family-group name: QUASIMUSINI Schimmel & Tarnawski, 2009. Stem: Quasimus- (ICZN 1999a, Article 29.4).

Quedius Stephens, 1829b: 22. Current status: valid genus in STAPHYLINIDAE: STAPHYLININAE: STAPHYLININI: QUEDIINA. Note: name placed on the Official List of Generic Names in Zoology (ICZN 1996c).

Type species: Staphylinus levicollis Brullé, 1832, by plenary power (see ICZN 1996c). Note: name placed on the Official List of Specific Names in Zoology (ICZN 1996c).

Family-group name: QUEDIIFORMES Kraatz, 1857. Stem: Quedi-.

Quelaestrygon Smetana, 1999: 241. Current status: valid genus in STAPHYLINIDAE: STAPHYLININAE: STAPHYLININI: QUELAESTRYGONINA.

Type species: Quelaestrygon puetzi Smetana, 1999, by original designation.

Family-group name: QUELAESTRYGONINI Brunke in Brunke et al., 2021: 415. Stem: Quelaestrygon-.

Radama Raffray, 1883: 230. Current status: valid genus in STAPHYLINIDAE: PSELAPHINAE: CLAVIGERITAE: CLAVIGERINI.

Type species: Radama inflatus Raffray, 1883, by subsequent designation (R. Lucas 1920: 564).

Family-group name: RADAMINI Jeannel, 1954. Stem: Radam-.

Raffrayia Reitter, 1882a: 198, 211. Current status: valid genus in STAPHYLINIDAE: PSELAPHINAE: EUPLECTITAE: TRICHONYCHINI: TRICHONYCHINA.

Type species: Trichonyx antennatus Raffray, 1877, by original designation.

Family-group name: RAFFRAYINA Jeannel, 1949. Stem: Raffrayi-.

Ranavala Raffray, 1898a: 206. Current status: valid genus in STAPHYLINIDAE: PSELAPHINAE: EUPLECTITAE: TRICHONYCHINI: TRICHONYCHINA.

Type species: Autoplectus integricollis Raffray, 1887, by subsequent monotypy (Raffray 1898a: 224).

Family-group name: RANAVALINI Jeannel, 1954. Stem: Ranaval-.

Ranolus Blair, 1929: 382. Current status: valid genus in DERMESTIDAE: ORPHILINAE.

Type species: Attagenus cavernicola Blair, 1929, by monotypy. Note: combined description of a new genus-group taxon and a single new species (ICZN 1999a, Article 12.2.6).

Family-group name: RANOLINI Háva in Zahradník and Háva, 2014: 306. Stem: Ranol-.

Raphidopalpa Chevrolat in Dejean, 1836a: 378. Current status: junior synonym of Aulacophora Chevrolat, 1836 in CHRYSOMELIDAE: GALERUCINAE: LUPERINI. Note: First Reviser (ICZN 1999a, Article 24.2.2) for Raphidopalpa Chevrolat, 1836 versus Aulacophora Chevrolat, 1836 is Gestro (1889: 72).

Type species: Crioceris abdominalis Fabricius, 1781, by subsequent designation (Chevrolat 1845: 6).

Family-group name: RHAPHIDOPALPINI Weise, 1902. Stem: Raphidopalp-.

Rathymus Dejean, 1831: 695, 783. Current status: valid genus in CARABIDAE: HARPALINAE: PTEROSTICHINI: PTEROSTICHINA.

Type species: Rathymus carbonarius Dejean, 1831, by monotypy.

Family-group name: RHATHYMINAE H.W. Bates, 1891. Stem: Rathym-.

Raymondionymus Wollaston, 1873b: 531. Current status: valid genus in CURCULIONIDAE: BRACHYCERINAE: RAYMONDIONYMINI. Note: new replacement name for Raymondia Aubé, 1861.

Type species: Raymondia fossor Aubé, 1861, by monotypy (type fixation for Raymondia Aubé).

Family-group name: RAYMONDIONYMINI Reitter, 1913. Stem: Raymondionym-.

Reicheia Saulcy, 1862: 285. Current status: valid genus in CARABIDAE: SCARITINAE: CLIVININI: CLIVININA.

Type species: Reicheia lucifuga Saulcy, 1862, by monotypy.

Family-group name: REICHEINA Jeannel, 1957. Stem: Reichei-.

Reichenbachia Leach, 1826: 451. Current status: valid genus in STAPHYLINIDAE: PSELAPHINAE: GONIACERITAE: BRACHYGLUTINI: BRACHYGLUTINA.

Type species: Bryaxis juncorum Leach, 1817, by subsequent designation (E. Desmarest 1852: 136).

Family-group name: REICHENBACHIINA Jakobson, 1910. Stem: Reichenbachi-.

Rembus W.S. MacLeay, 1825: 16. Current status: invalid senior synonym of Diplocheila Brullé, 1834 in CARABIDAE: HARPALINAE: LICININI: DICAELINA. Note: junior homonym of Rembus Germar, 1823 [Coleoptera: CURCULIONIDAE].

Type species: Carabus politus Fabricius, 1792, by monotypy.

Family-group name: REMBIDAE Gistel, 1848. Stem: Remb-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

Remipedella Semenov-Tjan-Shansky, 1907: 257. Current status: valid genus in TENEBRIONIDAE: BLAPTINAE: BLAPTINI: REMIPEDELLINA.

Type species: Remipedella deserti Semenov-Tjan-Shansky, 1907, by monotypy.

Family-group name: REMIPEDELLINI Semenov-Tjan-Shansky, 1907. Stem: Remipedell-.

Remphan G.R. Waterhouse, 1835: 67. Current status: valid genus in CERAMBYCIDAE: PRIONINAE: MACROTOMINI: REMPHANINA.

Type species: Remphan hopei G.R. Waterhouse, 1835, by monotypy.

Family-group name: REMPHANIDES Lacordaire, 1868. Stem: Remphan-.

Renania Lewis, 1887: 60. Current status: valid genus in EROTYLIDAE: EROTYLINAE: TRITOMINI.

Type species: Renania atrocyanea Lewis, 1887, by monotypy.

Family-group name: RENANIINAE Chûjô, 1941. Stem: Renani-.

Rentonium Crowson, 1966: 121. Current status: valid genus in RENTONIIDAE.

Type species: Rentonium daldiniae Crowson, 1966, by original designation.

Family-group name: RENTONIINAE Crowson, 1966. Stem: Rentoni-.

Repsimus W.S. MacLeay, 1819: 144. Current status: valid genus in SCARABAEIDAE: RUTELINAE: ANOPLOGNATHINI: ANOPLOGNATHINA.

Type species: Anoplognathus dytiscoides W.S. MacLeay, 1819 (= Rutela manicata Swartz, 1817), by subsequent designation (Crotch 1870b: 234).

Family-group name: RHEPSIMIDAE Streubel, 1846. Stem: Repsim-.

†Retromalisus Kazantsev, 2020: 1074. Current status: valid genus in †BERENDTIMIRIDAE: RETROMALISINAE. Note: [geological time not mentioned] (Baltic amber).

Type species: †Retromalisus damzeni Kazantsev, 2020, by original designation.

Family-group name: †RETROMALISINAE Fanti, Vitali & Pankowski, 2023: 135. Stem: Retromalis-.

Reynoldsiella Ray, 1930: 164. Current status: valid genus in MORDELLIDAE: MORDELLINAE: REYNOLDSIELLINI.

Type species: Reynoldsiella parallela Ray, 1930, by original designation.

Family-group name: REYNOLDSIELLINI Franciscolo, 1957. Stem: Reynoldsiell-.

Rhadalus J.L. LeConte, 1852d: 212. Current status: valid genus in RHADALIDAE: RHADALINAE.

Type species: Rhadalus testaceus J.L. LeConte, 1852, by monotypy.

Family-group name: RHADALINI J.L. LeConte, 1861. Stem: Rhadal-.

Rhadinocyba Faust, 1889: 78, 79. Current status: valid genus in BRENTIDAE: APIONINAE: RHADINOCYBITAE: RHADINOCYBINI.

Type species: Rhadinocyba nitidipennis Faust, 1889, by monotypy.

Family-group name: RHADINOCYBINI Alonso-Zarazaga, 1992. Stem: Rhadinocyb-.

Rhadinosomus Schönherr, 1840b: 473. Current status: valid genus in CURCULIONIDAE: CYCLOMINAE: ATERPINI: RHADINOSOMINA. Note: new replacement name for Leptosomus Schönherr, 1823.

Type species: Curculio acuminatus Fabricius, 1775, by original designation (type fixation for Leptosomus Schönherr).

Family-group name: RHADINOSOMIDES Lacordaire, 1863. Stem: Rhadinosom-.

Rhaeboscelis Chevrolat, 1838: 103. Current status: valid genus in BUPRESTIDAE: AGRILINAE: AGRILINI: RHAEBOSCELIDINA. Note: the original spelling was Raeboscelis but the incorrect subsequent spelling Rhaeboscelis is in prevailing usage, attributed to the publication of the original spelling, and therefore deemed to be the correct original spelling (ICZN 1999a, Article 33.3.1).

Type species: Rhaeboscelis purpureus Chevrolat, 1838, by monotypy. Note: Chevrolat (1838: 104) added that “carbonculus de Gory parait devoir rentrer dans cette coupe [it appears that carbonculus of Gory should belong in this genus]” and we consider, from that statement, that carbonculus is doubtfully included in the genus and therefore not an originally included species (ICZN 1999a, Article 67.2.5).

Family-group name: RHAEBOSCELIDI Cobos, 1976. Stem: Rhaeboscelid-.

Rhaebus Fischer, 1823a: pl. 47. Current status: valid genus in CHRYSOMELIDAE: BRUCHINAE: RHAEBINI.

Type species: Rhaebus gebleri Fischer, 1823, by monotypy.

Family-group name: RHAEBITES Blanchard, 1845. Stem: Rhaeb-.

Rhaetulus Westwood, 1871: 353. Current status: valid genus in LUCANIDAE: LUCANINAE: DORCINI.

Type species: Rhaetulus crenatus Westwood, 1871, by monotypy.

Family-group name: RHAETULINAE Miwa, 1931. Stem: Rhaetul-.

Rhagiomorpha Newman, 1840a: 21. Current status: junior synonym of Stenoderus Dejean, 1821 in CERAMBYCIDAE: CERAMBYCINAE: RHAGIOMORPHINI.

Type species: Stenoderus concolor W.S. MacLeay, 1826, by subsequent designation (J. Thomson 1864: 136).

Family-group name: RHAGIOMORPHIDAE Newman, 1841. Stem: Rhagiomorph-.

Rhagium Fabricius, 1775: 182. Current status: valid genus in CERAMBYCIDAE: LEPTURINAE: RHAGIINI.

Type species: Cerambyx inquisitor Linnaeus, 1758, by subsequent designation (Westwood 1838c: 41).

Family-group name: RHAGIADAE W. Kirby, 1837. Stem: Rhagi-.

Rhagocrepis Eschscholtz, 1829c: 5. Current status: junior synonym of Leptotrachelus Latreille, 1829 in CARABIDAE: HARPALINAE: CTENODACTYLINI.

Type species: Rhagocrepis riedelii Eschscholtz, 1829, by monotypy.

Family-group name: RHAGOCREPIDES Chaudoir, 1848. Stem: Rhagocrepid-.

Rhagodera Mannerheim, 1843: 300. Current status: valid genus in ZOPHERIDAE: COLYDIINAE: RHAGODERINI.

Type species: Rhagodera tuberculata Mannerheim, 1843, by monotypy.

Family-group name: RHAGODERINI G.H. Horn, 1878. Stem: Rhagoder-.

Rhagophthalmus Motschulsky, 1853a: 45. Current status: valid genus in RHAGOPHTHALMIDAE.

Type species: Rhagophthalmus scutellatus Motschulsky, 1853, by monotypy.

Family-group name: RHAGOPHTHALMIDAE E. Olivier, 1907. Stem: Rhagophthalm-.

Rhamnusium Latreille, 1829b: 130. Current status: valid genus in CERAMBYCIDAE: LEPTURINAE: RHAMNUSIINI.

Type species: Callidium salicis Fabricius, 1787 (= Cerambyx bicolor Schrank, 1781), by monotypy.

Family-group name: RHAMNUSIINI Sama, 2009. Stem: Rhamnusi-.

Rhamphophorus Poinar & A.E. Brown, 2021: 2. Current status: valid genus in POLYPHAGA incertae sedis.

Type species: Rhamphophorus legalovii Poinar & A.E. Brown, 2021, by original designation.

Family-group name: RHAMPHOPHORINI Poinar & A.E. Brown, 2021: 2. Stem: Rhamphophor-. Note: combined description of a new family-group taxon and a single new nominal genus (ICZN 1999a, Article 13.5).

Rhamphus Clairville, 1798: 57, 104. Current status: valid genus in CURCULIONIDAE: CURCULIONINAE: RHAMPHINI: RHAMPHINA. Note: two original spellings were used, including Ramphus (p. 104); Lepeletier and Audinet-Serville (1825: 284) acted as First Revisers (ICZN 1999a, Article 24.2.3) and selected Rhamphus.

Type species: Rhamphus flavicornis Clairville, 1798 (= Curculio pulicarius Herbst, 1795), by monotypy.

Family-group name: RAMPHORIA Rafinesque, 1815. Stem: Rhamph-.

Rhanis J.L. LeConte, 1853d: 357, 360. Current status: invalid senior synonym of Rhanidea Strohecker, 1953 in ENDOMYCHIDAE: LEIESTINAE. Note: junior homonym of Rhanis C.L. Koch, 1846 [Arachnida].

Type species: Lycoperdina unicolor Ziegler, 1845, by monotypy.

Family-group name: RHANES J.L. LeConte & G.H. Horn, 1883. Stem: Rhanid-. Note: family- group name permanently invalid (ICZN 1999a, Article 39).

Rhaphipodus Audinet-Serville, 1832: 168. Current status: valid genus in CERAMBYCIDAE: PRIONINAE: REMPHANINI.

Type species: Rhaphipodus suturalis Audinet-Serville, 1832, by monotypy.

Family-group name: RHAPHIPODI Lameere, 1912. Stem: Rhaphipod-.

Rhexius J.L. LeConte, 1849: 102. Current status: valid genus in STAPHYLINIDAE: PSELAPHINAE: EUPLECTITAE: TROGASTRINI: RHEXIINA.

Type species: Rhexius insculptus J.L. LeConte, 1849, by monotypy.

Family-group name: RHEXINI Park, 1951. Stem: Rhexi-.

Rhigopsis J.L. LeConte, 1874a: 459. Current status: valid genus in CURCULIONIDAE: ENTIMINAE: ENTIMITAE: BYRSOPAGINI: STRANGALIODINA.

Type species: Rhigopsis effracta J.L. LeConte, 1874, by monotypy.

Family-group name: RHIGOPSINI J.L. LeConte, 1874. Stem: Rhigopse-.

Rhigus Germar, 1823: 438. Current status: valid genus in CURCULIONIDAE: ENTIMINAE: ENTIMITAE: ENTIMINI: RHIGIINA.

Type species: Curculio tribuloides Pallas, 1781, by original designation.

Family-group name: RHIGIINA Legalov, 2020b: 339. Stem: Rhigi- (ICZN 1999a, Article 29.4).

Rhina Latreille, 1802b: 198. Current status: invalid senior synonym of Rhinostomus Rafinesque, 1815 in CURCULIONIDAE: DRYOPHTHORINAE: RHINOSTOMINI. Note: junior homonym of Rhina J.G. Schneider, 1801 [Pisces]; name placed on the Official Index of Rejected and Invalid Generic Names in Zoology (ICZN 1955a).

Type species: Curculio barbicornis Fabricius, 1775, by monotypy. Note: name placed on the Official List of Specific Names in Zoology (ICZN 1955a).

Family-group name: RHINIDAE J.L. LeConte, 1874. Stem: Rhin-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

Rhinaria W. Kirby, 1819: 430. Current status: valid genus in CURCULIONIDAE: CYCLOMINAE: ATERPINI: ATERPINA.

Type species: Rhinaria cristata W. Kirby, 1819, by original designation.

Family-group name: RHINARIDES Jekel, 1865. Stem: Rhinari-.

Rhinastus Schönherr, 1825: 584. Current status: valid genus in CURCULIONIDAE: MOLYTINAE: CHOLINI: RHINASTINA.

Type species: Cholus sternicornis Germar, 1823, by monotypy.

Family-group name: RHINASTINI Vaurie, 1973. Stem: Rhinast-.

Rhinocartus Voss, 1922a: 17, 18. Current status: valid genus in ATTELABIDAE: RHYNCHITINAE: RHINOCARTITAE: RHINOCARTINI.

Type species: Rhinocartus tessmanni Voss, 1922, by monotypy.

Family-group name: RHINOCARTINI Voss, 1931. Stem: Rhinocart-.

Rhinocyllus Germar, 1817b: 341. Current status: valid genus in CURCULIONIDAE: LIXINAE: RHINOCYLLINI.

Type species: Curculio thaumaturgus Rossi, 1794 (= Curculio conicus Frölich, 1792), by monotypy.

Family-group name: RHINOCYLLIDES Lacordaire, 1863. Stem: Rhinocyll-.

Rhinomacer Fabricius, 1781: 199. Current status: name in MYCTERIDAE: MYCTERINAE. Note: name placed on the Official Index of Rejected and Invalid Generic Names in Zoology (see ICZN 2005c).

Type species: Rhinomacer curculioides Fabricius, 1781, by monotypy.

Family-group name: RHINOMACERIDAE Fleming, 1821. Stem: Rhinomacr-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

Rhinoncus Schönherr, 1825: 586. Current status: valid genus in CURCULIONIDAE: CEUTORHYNCHINAE: PHYTOBIINI. Note: name placed on the Official List of Generic Names in Zoology (ICZN 1989b).

Type species: Curculio pericarpius Linnaeus, 1758, by plenary power (see ICZN 1989b). Note: name placed on the Official List of Specific Names in Zoology (ICZN 1989b).

Family-group name: RHINONCIDES C.G. Thomson, 1865. Stem: Rhinonc-.

Rhinophthalmus J. Thomson, 1861: 152. Current status: valid genus in CERAMBYCIDAE: CERAMBYCINAE: URACANTHINI. Note: new replacement name for Stephanops Shuckard, 1838.

Type species: Stephanops nasutus Newman, 1838, by monotypy (type fixation for Stephanops Shuckard).

Family-group name: RHINOPHTHALMITAE J. Thomson, 1861. Stem: Rhinophthalm-.

Rhinorhipus Lawrence, 1988: 4. Current status: valid genus in RHINORHIPIDAE.

Type species: Rhinorhipus tamborinensis Lawrence, 1988, by original designation.

Family-group name: RHINORHIPIDAE Lawrence, 1988. Stem: Rhinorhip-.

Rhinorhynchidius Voss, 1922a: 107. Current status: valid genus in BRENTIDAE: RHINORHYNCHIDIITAE.

Type species: Rhinorhynchidius cylindrirostris Voss, 1922 (= Apion meridionale Lea, 1926), by monotypy.

Family-group name: RHINORHYNCHIDIINAE Zimmerman, 1994. Stem: Rhinorhynchidi-.

Rhinorhynchus Sharp, 1882a: 88. Current status: valid genus in NEMONYCHIDAE: RHINORHYNCHINAE: RHINORHYNCHINI.

Type species: Rhinorhynchus zealandicus Sharp, 1882 (= Rhinomacer rufulus Broun, 1880), by monotypy.

Family-group name: RHINORHYNCHINI Voss, 1922. Stem: Rhinorhynch-.

Rhinoscepsis J.L. LeConte, 1878a: 382. Current status: valid genus in STAPHYLINIDAE: PSELAPHINAE: EUPLECTITAE: METOPIASINI: RHINOSCEPSINA.

Type species: Rhinoscepsis bistriatus J.L. LeConte, 1878, by monotypy.

Family-group name: RHINOSCEPSII Bowman, 1934. Stem: Rhinosceps-.

Rhinosimus Latreille, 1802b: 192. Current status: junior synonym of Salpingus Illiger, 1802 in SALPINGIDAE: SALPINGINAE.

Type species: Curculio planirostris Fabricius, 1787, by monotypy.

Family-group name: RHINOSIMIDAE Hope, 1841. Stem: Rhinosim-.

Rhinostomus Rafinesque, 1815: 115. Current status: valid genus in CURCULIONIDAE: DRYOPHTHORINAE: RHINOSTOMINI. Note: name placed on the Official List of Generic Names in Zoology (ICZN 1955a).

Type species: Curculio barbirostris Fabricius, 1775, by plenary power (see ICZN 1955a). Note: name placed on the Official List of Specific Names in Zoology (ICZN 1955a).

Family-group name: RHINOSTOMINI Kuschel, 1995. Stem: Rhinostom-.

Rhinotragus Germar, 1823: 513. Current status: valid genus in CERAMBYCIDAE: CERAMBYCINAE: RHINOTRAGINI.

Type species: Rhinotragus dorsiger Germar, 1823, by monotypy.

Family-group name: RHINOTRAGITAE J. Thomson, 1861. Stem: Rhinotrag-.

Rhipicera Latreille, 1816: 235. Current status: valid genus in RHIPICERIDAE: RHIPICERINAE.

Type species: Hispa mystacina Fabricius, 1775, by monotypy.

Family-group name: RHIPICERIDES Latreille, 1834. Stem: Rhipicer-.

Rhipidandrus J.L. LeConte, 1862: 236. Current status: valid genus in TENEBRIONIDAE: TENEBRIONINAE: BOLITOPHAGINI.

Type species: Xyletinus flabellicornis Sturm, 1826 (= Melolontha paradoxa Palisot de Beauvois, 1818), by monotypy.

Family-group name: RHIPIDANDRI J.L. LeConte, 1862. Stem: Rhipidandr-.

Rhipidocerus Westwood, 1842d: 70. Current status: valid genus in CERAMBYCIDAE: PRIONINAE: RHIPIDOCERINI.

Type species: Rhipidocerus australasiae Westwood, 1842, by monotypy.

Family-group name: RHIPIDOCERINI Jin, De Keyzer, Ashman, Zwick & Ślipiński, 2023: 507. Stem: Rhipidocer-.

Rhizophagus Herbst, 1793: 18. Current status: valid genus in MONOTOMIDAE: RHIZOPHAGINAE. Note: the original spelling Ryzophagus was changed to Rhizophagus and Rhizophagus Herbst, 1793 placed on the Official List of Generic Names in Zoology (ICZN 1995c).

Type species: Lyctus bipustulatus Fabricius, 1792, by subsequent designation (Westwood 1838c: 13; see ICZN 1995c). Note: name placed on the Official List of Specific Names in Zoology (ICZN 1995c).

Family-group name: RHIZOPHAGI L. Redtenbacher, 1845. Stem: Rhizophag-.

†Rhizophtoma Kirejtshuk & Azar in Kirejtshuk et al., 2009a: 123. Current status: valid genus in MONOTOMIDAE: MONOTOMINAE: †RHIZOPHTOMINI. Note: Lower Cretaceous (Lebanon; Spain).

Type species: †Rhizophtoma elateroides Kirejtshuk & Azar, 2009, by original designation.

Family-group name: †RHIZOPHTOMINAE Kirejtshuk & Azar, 2009. Stem: Rhizophtom-.

Rhizotrogus Berthold, 1827: 362. Current status: valid genus in SCARABAEIDAE: MELOLONTHINAE: RHIZOTROGINI. Note: this name is sometimes credited to Latreille (1825: 371) (e.g., Branco 2007: 20; J. Bezděk 2016: 275) but it was proposed in that work under the French vernacular name Rhizotrogue, not under a scientific name.

Type species: Melolontha aestiva G.-A. Olivier, 1789, by monotypy.

Family-group name: RHIZOTROGIDAE Burmeister, 1855. Stem: Rhizotrog-.

Rhodocanthopus Kaup, 1871: 90. Current status: valid genus in PASSALIDAE: PASSALINAE: PASSALINI.

Type species: Passalus caelatus Erichson, by subsequent designation (Hincks and Dibb 1935: 39)

Family-group name: RHODACANTHOPINAE Kuwert, 1896. Stem: Rhodocanthopod-.

Rhodopina Gressitt, 1951: 439. Current status: valid genus in CERAMBYCIDAE: LAMIINAE: LAMIINI. Note: new replacement name for Rhodopis J. Thomson, 1857.

Type species: Rhodopis pubera J. Thomson, 1857, by monotypy (type fixation for Rhodopis J. Thomson).

Family-group name: RHODOPININI Gressitt, 1951. Stem: Rhodopin-.

Rhodopis J. Thomson, 1857: 174. Current status: invalid senior synonym of Rhodopina Gressit, 1951 in CERAMBYCIDAE: LAMIINAE: LAMIINI. Note: junior homonym of Rhodopis L. Reichenbach, 1854 [Aves].

Type species: Rhodopis pubera J. Thomson, 1857, by monotypy.

Family-group name: RHODOPIDES Lacordaire, 1872. Stem: Rhodopid-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

†Rhombocoleus Rohdendorf, 1961: 432. Current status: valid genus in †RHOMBOCOLEIDAE. Note: Permian (Siberia, Russia).

Type species: †Rhombocoleus andreae Rohdendorf, 1961, by original designation.

Family-group name: †RHOMBOCOLEIDAE Rohdendorf, 1961. Stem: Rhombocole-.

Rhomborhina Hope, 1837: 120. Current status: valid genus in SCARABAEIDAE: CETONIINAE: GOLIATHINI: RHOMBORHININA.

Type species: Goliathus heros Gory & Percheron, 1833, by original designation.

Family-group name: RHOMBORRHINAE Westwood, 1842. Stem: Rhomborh-. Note: this family- group name, issued 1 January 1842, has precedence over CORYPHOCERINA Burmeister, 1842, issued by 28 December 1842.

Rhopalocerus W. Redtenbacher, 1842: 21. Current status: valid genus in ZOPHERIDAE: COLYDIINAE: RHOPALOCERINI. Note: name placed on the Official List of Generic Names in Zoology (ICZN 1986b).

Type species: Rhopalocerus setosus W. Redtenbacher, 1842 (= Monotoma rondanii A. Villa & G.B. Villa, 1833), by monotypy (see ICZN 1986b). Note: Monotoma rondanii A. Villa & G.B. Villa, 1833 was placed on the Official List of Specific Names in Zoology (ICZN 1986b).

Family-group name: RHOPALOCERINI Reitter, 1911. Stem: Rhopalocer-.

Rhopalogastrum Bernhauer, 1912: 68. Current status: valid genus in STAPHYLINIDAE: ALEOCHARINAE: AUTALIINI.

Type species: Rhopalogastrum claviventre Bernhauer, 1912, by monotypy.

Family-group name: RHOPALOGASTRA Fenyes, 1919. Stem: Rhopalogastr-.

Rhopalomelus Boheman, 1848: 165. Current status: valid genus in CARABIDAE: HARPALINAE: CHLAENIINI: CHLAENIINA.

Type species: Rhopalomelus angusticollis Boheman, 1848, by monotypy.

Family-group name: RHOPALOMELINI Alluaud, 1930. Stem: Rhopalomel-.

Rhopalophora Audinet-Serville, 1834b: 100. Current status: valid genus in CERAMBYCIDAE: CERAMBYCINAE: RHOPALOPHORINI.

Type species: Rhopalophora sanguinicollis Audinet-Serville, 1834 (= Callichroma collare Germar, 1823), by monotypy.

Family-group name: RHOPALOPHORITES Blanchard, 1845. Stem: Rhopalophor-.

Rhopalosilpha Arrow, 1929b: 97. Current status: valid genus in DERMESTIDAE: DERMESTINAE: MARIOUTINI.

Type species: Rhopalosilpha wasmanni Arrow, 1929, by monotypy.

Family-group name: RHOPALOSILPHINAE Arrow, 1929. Stem: Rhopalosilph-.

Rhynchaenus Clairville, 1798: 71, 142. Current status: valid genus in CURCULIONIDAE: CURCULIONINAE: RHAMPHINI: RHAMPHINA. Note: two original spellings were used, including Rynchaenus (pp. 56, 57, 71–73); Buchanan (1939: 81) acted as First Reviser (ICZN 1999a, Article 24.2.3) and selected the spelling Rhynchaenus.

Type species: Rhynchaenus xylostei Clairville, 1798 (= Curculio lonicerae Herbst, 1795), by subsequent designation (Crotch 1870a: 50).

Family-group name: RHYNCHENIDAE Blanchard, 1853. Stem: Rhynchaen-.

Rhynchitallus Voss, 1960: 415. Current status: valid genus in ATTELABIDAE: RHYNCHITINAE: RHYNCHITITAE: RHYNCHITINI: RHYNCHITINA.

Type species: Rhynchitallus cyclops Voss, 1960, by original designation.

Family-group name: RHYNCHITALLINA Legalov, 2003. Stem: Rhynchitall-.

Rhynchitapion Wanat, 2021: 59. Current status: valid genus in BRENTIDAE: APIONINAE incertae sedis.

Type species: Rhynchitapion variiforme Wanat, 2021, by original designation.

Family-group name: RHYNCHITAPIINI Wanat, 2021: 56. Stem: Rhynchitapi-.

Rhynchites D.H. Schneider, 1791: 83. Current status: valid genus in ATTELABIDAE: RHYNCHITINAE: RHYNCHITITAE: RHYNCHITINI: RHYNCHITINA.

Type species: Curculio bacchus Linnaeus, 1758, by subsequent designation (Latreille 1810: 430).

Family-group name: RHYNCHITISIDAE Gistel, 1848. Stem: Rhynchit-.

Rhynchitomacer Voss, 1937b: 201. Current status: valid genus in NEMONYCHIDAE: RHINORHYNCHINAE: RHYNCHITOMACRINI.

Type species: Rhynchitomacer flavus Voss, 1937, by original designation.

Family-group name: RHYNCHITOMACERINI May, 1993. Stem: Rhynchitomacr-.

Rhynchitoplesius Voss, 1952: 177. Current status: valid genus in NEMONYCHIDAE: RHINORHYNCHINAE: MECOMACERINI: RHYNCHITOPLESIINA.

Type species: Rhynchitomacer eximius Voss, 1937, by monotypy.

Family-group name: RHYNCHITOPLESIINA Legalov, 2011a: 163. Stem: Rhynchitoplesi-

Rhynchophorus Herbst, 1795: 3, pl. lx. Current status: valid genus in CURCULIONIDAE: DRYOPHTHORINAE: RHYNCHOPHORINI. Note: two original spellings were used, including Rynchophorus (p. 3); Alonso-Zarazaga and Lyal (1999: 64) acted as First Reviser (ICZN 1999a, Article 24.2.3) and selected the spelling Rhynchophorus.

Type species: Curculio palmarum Linnaeus, 1758, by subsequent designation (Schönherr 1826: 23).

Family-group name: RHYNCHOPHORIDES Schönherr, 1833. Stem: Rhynchophor-.

Rhyncogonus Sharp in Blackburn and Sharp, 1885: 176. Current status: valid genus in CURCULIONIDAE: ENTIMINAE: OTIORHYNCHITAE: RHYNCOGONINI.

Type species: Rhyncogonus blackburni Sharp, 1885, by subsequent designation (Zimmerman 1964: 308).

Family-group name: RHYNCOGONIDES Sharp, 1919. Stem: Rhyncogon-.

Rhyncolus Germar, 1817b: 340. Current status: valid genus in CURCULIONIDAE: COSSONINAE: RHYNCOLINI: RHYNCOLINA. Note: name placed on the Official List of Generic Names in Zoology (ICZN 1991b).

Type species: Curculio ater Linnaeus, 1758, by plenary power (see ICZN 1991b). Note: name placed on the Official List of Specific Names in Zoology (ICZN 1991b).

Family-group name: RHYNCHALIDAE Gistel, 1848. Stem: Rhyncol-.

Rhyparosomus Schönherr, 1842a: 200. Current status: valid genus in CURCULIONIDAE: CYCLOMINAE: RHYTHIRRININI.

Type species: Rhyparosomus horridus Boheman, 1842, by original designation.

Family-group name: RHYPAROSOMIDES Lacordaire, 1863. Stem: Rhyparosom-.

Rhyparus Westwood, 1845c: 93. Current status: valid genus in SCARABAEIDAE: APHODIINAE: RHYPARINI. Note: the original spelling was Ryparus but the unjustified emendation Rhyparus, proposed by Agassiz (1846: 328), is in prevailing usage, attributed to the publication of the original spelling, and therefore deemed to be the correct original spelling (ICZN 1999a, Article 33.2.3.1); accepting Agassiz’s emendation as a justified emendation avoids homonymy with Ryparus Spinola, 1845 [Coleoptera: CLERIDAE] (see A.B.T. Smith 2006: 159).

Type species: Rhyparus desjardinsii Westwood, 1845, by monotypy (type fixation for Ryparus Westwood).

Family-group name: RHYPARINA A. Schmidt, 1910. Stem: Rhypar-.

Rhysodes Germar, 1822a: no 1. Current status: valid genus in CARABIDAE: RHYSODINAE: RHYSODINI.

Type species: Rhysodes europaeus Germar, 1822 (= Cucujus sulcatus Fabricius, 1787), by monotypy.

Family-group name: RHYSODITES Laporte, 1840. Stem: Rhysod-.

Rhysopaussus Wasmann, 1896: 616. Current status: valid genus in TENEBRIONIDAE: TENEBRIONINAE: RHYSOPAUSSINI.

Type species: Rhysopaussus dohertyi Wasmann, 1896, by monotypy.

Family-group name: RHYSOPAUSSIDAE Wasmann, 1896. Stem: Rhysopauss-.

Rhyssemus Mulsant, 1842a: 305, 314. Current status: valid genus in SCARABAEIDAE: APHODIINAE: PSAMMODIINI: RHYSSEMINA.

Type species: here fixed (under ICZN 1999a, Article 70.3.2) as Ptinus germanus Linnaeus, 1767, misidentified as Scarabaeus asper Fabricius, 1775 (sensu Mulsant, 1842), in the subsequent designation of C.G. Thomson (1859: 81). Note: see Branco (2007: 20) for additional information regarding the misidentified type species.

Family-group name: RHYSSEMINA Pittino & Mariani, 1986. Stem: Rhyssem-.

Rhythirrinus Schönherr, 1823: 1142. Current status: valid genus in CURCULIONIDAE: CYCLOMINAE: RHYTHIRRININI.

Type species: Curculio inaequalis Fabricius, 1781, by original designation.

Family-group name: RHYTIRHINIDES Lacordaire, 1863. Stem: Rhythirrin-.

Rhyticephalus Chevrolat, 1839: 174. Current status: junior synonym of Ischyromerus Labram & Imhoff, 1838 in BRENTIDAE: BRENTINAE: TRACHELIZINI: RHYTICEPHALINA.

Type species: Rhyticephalus brevicornis Chevrolat, 1839 (= Brenthus madagascariensis Labram & Imhoff, 1838), by subsequent designation (R. Lucas 1920: 574; typification for Rhytidocephalus, an unjustified emendation by Agassiz (1846: 327) of Rhyticephalus Chevrolat).

Family-group name: RHYTICEPHALINI Kleine, 1922. Stem: Rhyticephal-.

Rhytidophloeus Schönherr, 1842a: 290. Current status: valid genus in CURCULIONIDAE: MOLYTINAE: LITHININI: RHYTIDOPHLOEINA.

Type species: Curculio albipes G.-A. Olivier, 1791, by original designation.

Family-group name: RHYTIDOPHLOEINI Voss, 1963. Stem: Rhytidophloe-.

Rhyzobius Stephens, 1829b: 19. Current status: valid genus in COCCINELLIDAE: COCCINELLINAE: COCCIDULINI.

Type species: Nitidula litura Fabricius, 1787, by monotypy.

Family-group name: RHIZOBIINA C.G. Thomson, 1866. Stem: Rhyzobi-.

Riedelinius Legalov, 2003: 362. Current status: valid genus in ATTELABIDAE: ATTELABINAE: EUOPINI.

Type species: Euops armatus Riedel, 1999, by original designation.

Family-group name: RIEDELININA Legalov, 2007. Stem: Riedelin- (ICZN 1999a, Article 29.4).

Ripidius Thunberg, 1806: 4. Current status: valid genus in RIPIPHORIDAE: RIPIDIINAE: RIPIDIINI.

Type species: Ripidius pectinicornis Thunberg, 1806, by monotypy.

Family-group name: RHIPIDIINI Gerstaecker, 1855. Stem: Ripidi-.

Ripiphorus Bosc, 1791: 327. Current status: valid genus in RIPIPHORIDAE: RIPIPHORINAE: RIPIPHORINI. Note: name placed on the Official List of Generic Names in Zoology (ICZN 2020b).

Type species: Ripiphorus subdipterus Fabricius, 1792, by plenary power (ICZN 2020b). Note: no species were originally included in this genus by Bosc (1791); the first ones are the twelve species listed by Fabricius (1792b: 109–112) (see Bousquet and Bouchard 2018: 36–37); name placed on the Official List of Specific Names in Zoology (ICZN 2020b).

Family-group name: RHIPIPHORITES Laporte, 1840. Stem: Ripiphor-.

Rondoniella Kaszab, 1970: 112. Current status: valid genus in TENEBRIONIDAE: PIMELIINAE: CNEMEPLATIINI: RONDONIELLINA.

Type species: Rondoniella costata Kaszab, 1970, by original designation.

Family-group name: RONDONIELLINA Ferrer & Moragues, 2000. Stem: Rondoniell-.

Ropalodontus Mellié, 1847: 109. Current status: valid genus in CIIDAE: CIINAE: OROPHIINI.

Type species: Cis perforatus Gyllenhal, 1813, by monotypy.

Family-group name: RHOPALODONTINI Everts, 1898. Stem: Ropalodont-.

Rosalia Audinet-Serville, 1834a: 561. Current status: valid genus in CERAMBYCIDAE: CERAMBYCINAE: ROSALIINI.

Type species: Cerambyx alpinus Linnaeus, 1758, by monotypy.

Family-group name: ROSALIITES Fairmaire, 1864. Stem: Rosali-.

Rosiroia Bechyné, 1950: 148. Current status: valid genus in CHRYSOMELIDAE: EUMOLPINAE: ROSIROIINI.

Type species: Rosiroia anisotomoides Bechyné, 1950, by monotypy.

Family-group name: ROSIROIINI Bechyné, 1950. Stem: Rosiroi-.

Rostricephalus Fleutiaux, 1918b: 252. Current status: valid genus in ELATERIDAE: LISSOMINAE: PROTELATERINI.

Type species: Rostricephalus vitalisi Fleutiaux, 1918, by monotypy.

Family-group name: ROSTRICEPHALINAE Fleutiaux, 1947. Stem: Rostricephal-.

Rugilus Leach in Samouelle, 1819: 173. Current status: valid genus in STAPHYLINIDAE: PAEDERINAE: LATHROBIINI: STILICINA.

Type species: Staphylinus orbiculatus Paykull, 1789, by original designation.

Family-group name: RUGILINA Hatch, 1957. Stem: Rugil-.

Rupilia Clark, 1864: 260. Current status: valid genus in CHRYSOMELIDAE: GALERUCINAE: GALERUCINI.

Type species: Rupilia ruficollis Clark, 1864, by original designation.

Family-group name: RUPILIITES Chapuis, 1875. Stem: Rupili-.

Rutela Latreille, 1802b: 151. Current status: valid genus in SCARABAEIDAE: RUTELINAE: RUTELINI: RUTELINA.

Type species: Scarabaeus lineola Linnaeus, 1758, by subsequent designation (MacLeay 1819: 72).

Family-group name: RUTELIDAE W.S. MacLeay, 1819. Stem: Rutel-.

Rygmodus A. White, 1846: 11. Current status: valid genus in HYDROPHILIDAE: CYLOMINAE.

Type species: Rygmodus modestus A. White, 1846, by subsequent designation (Knisch 1924: 107).

Family-group name: RYGMODINI Orchymont, 1916. Stem: Rygmod-.

Rypobius J.L. LeConte, 1852e: 141, 142. Current status: valid genus in CORYLOPHIDAE: CORYLOPHINAE: RYPOBIINI.

Type species: Rypobius marinus J.L. LeConte, 1852, by monotypy.

Family-group name: RHYPOBIINI Paulian, 1950. Stem: Rypobi-.

Ryssonotus W.S. MacLeay, 1819: 96, 98. Current status: valid genus in LUCANIDAE: LUCANINAE: RHYSSONOTINI. Note: name also referred to as Rhyssonotus, an unjustified emendation by Agassiz (1846: 329), in recent literature.

Type species: Lucanus nebulosus W. Kirby, 1819, by monotypy. Note: as noted by Reid and Beatson (2016: 3), Lucanus foveolatus Thunberg, 1806, often listed as a senior synonym of Lucanus nebulosus W. Kirby, 1819, is a junior synonym of the North American Lucanus capreolus (Linnaeus, 1763).

Family-group name: RHYSSONOTINI Benesh, 1960. Stem: Ryssonot-.

Sachalinobia Jakobson, 1899: 39. Current status: valid genus in CERAMBYCIDAE: LEPTURINAE: SACHALINOBIINI.

Type species: Sachalinobia retata Jakobson, 1899 (= Brachyta koltzei Heyden, 1887), by original designation.

Family-group name: SACHALINOBIINI Danilevsky, 2010. Stem: Sachalinobi-.

Sacium J.L. LeConte, 1852e: 144. Current status: junior synonym of Clypastraea Haldeman, 1842 in CORYLOPHIDAE: CORYLOPHINAE: PARMULINI. Note: new replacement name for Clypeaster Dejean, 1821.

Type species: Cossyphus pusillus Gyllenhal, 1810, by monotypy (type fixation for Clypeaster Dejean).

Family-group name: SACIINA A. Matthews, 1888. Stem: Saci-.

Sagra Fabricius, 1792b: 51. Current status: valid genus in CHRYSOMELIDAE: SAGRINAE: SAGRINI.

Type species: Tenebrio femoratus Drury, 1773, by subsequent designation (Latreille 1810: 431).

Family-group name: SAGRIDA Leach, 1815. Stem: Sagr-.

Sahlbergius Bernhauer, 1927: 378. Current status: valid genus in STAPHYLINIDAE: ALEOCHARINAE: SAHLBERGIINI.

Type species: Sahlbergius mirabilis Bernhauer, 1927, by monotypy.

Family-group name: SAHLBERGIINI Kistner, 1993. Stem: Sahlbergi-.

Salax Guérin-Méneville, 1834b: 11. Current status: valid genus in TENEBRIONIDAE: PIMELIINAE: TRILOBOCARINI.

Type species: Salax lacordairii Guérin-Méneville, 1834, by monotypy.

Family-group name: SALAXINI Casey, 1907. Stem: Salac-.

Salcedia Fairmaire, 1899: 512. Current status: valid genus in CARABIDAE: SCARITINAE: SALCEDIINI.

Type species: Salcedia perrieri Fairmaire, 1899, by monotypy.

Family-group name: SALCEDIINI Alluaud, 1930. Stem: Salcedi-.

Salpingus Illiger, 1802: 301. Current status: valid genus in SALPINGIDAE: SALPINGINAE.

Type species: Curculio roboris Fabricius, 1787 (= Curculio ruficollis Linnaeus, 1761), by subsequent designation (Crotch 1870b: 215).

Family-group name: SALPINGIDES Leach, 1815. Stem: Salping-.

Sandalus Knoch, 1801: 131. Current status: valid genus in RHIPICERIDAE: SANDALINAE.

Type species: Sandalus niger Knoch, 1801, by subsequent designation (Stephens 1840: 305). Note: the type species usually listed is Sandalus petrophya Knoch, 1801 (e.g., Hájek 2016: 431) by subsequent designation of Guérin-Méneville (1843c [No 2]: 2) but Stephens’ designation is earlier; both species are currently included in Sandalus Knoch, 1801.

Family-group name: SANDALIDAE Jakobson, 1913. Stem: Sandal-.

Sapriniana Strand, 1917: 85. Current status: valid genus in SCARABAEIDAE: AEGIALIINAE: SAPRINIANINI. Note: new replacement name for Saprus Blackburn, 1904.

Type species: Saprus griffithi Blackburn, 1904, by monotypy (type fixation for Saprus Blackburn).

Family-group name: SAPRINIANINI Nikolajev, 2011: 328. Stem: Saprinian-.

Saperda Fabricius, 1775: 184. Current status: valid genus in CERAMBYCIDAE: LAMIINAE: SAPERDINI. Note: name placed on the Official List of Generic Names in Zoology (ICZN 1980).

Type species: Cerambyx carcharias Linnaeus, 1758, by subsequent designation (Guérin- Méneville 1829b: 151). Note: Cerambyx scalaris Linnaeus, 1758 is listed as the type species by subsequent designation in Curtis (1829: pl. 275) on the Official List of Specific Names in Zoology (ICZN 1980); however, as pointed out by Bousquet (2008a: 622), the type species Cerambyx carcharias Linnaeus, 1758 was fixed earlier by Guérin- Méneville (1829b: 151), as recognized by recent authors (e.g., Danilevsky 2020: 473); the Official List of Generic Names in Zoology needs to be updated to reflect the changes for the type species mentioned above.

Family-group name: SAPERDAIRES Mulsant, 1839. Stem: Saperd-.

Saphanus Audinet-Serville, 1834b: 81. Current status: valid genus in CERAMBYCIDAE: SPONDYLIDINAE: SAPHANINI.

Type species: Callidium spinosum Fabricius, 1792 (= Callidium piceum Laicharting, 1784), by monotypy.

Family-group name: SAPHANIDAE Gistel, 1848. Stem: Saphan-.

Saphoglossa Sharp, 1883a: 291. Current status: valid genus in STAPHYLINIDAE: ALEOCHARINAE: HYGRONOMINI: SAPHOGLOSSINA.

Type species: Saphoglossa pictipennis Sharp, 1883, by monotypy.

Family-group name: SAPHOGLOSSAE Bernhauer & Scheerpeltz, 1926. Stem: Saphogloss-.

Saprinus Erichson, 1834: 172. Current status: valid genus in HISTERIDAE: SAPRININAE: SAPRININI.

Type species: Hister nitidulus Fabricius, 1801 (= Hister semistriatus Scriba, 1790), by subsequent designation (Westwood 1838c: 22).

Family-group name: SAPRINITES Blanchard, 1845. Stem: Saprin-.

Saprus Blackburn, 1904: 150, 178. Current status: invalid senior synonym of Sapriniana Strand, 1917 in SCARABAEIDAE: AEGIALIINAE: SAPRINIANINI. Note: junior homonym of Saprus Gistel, 1856 [Coleoptera: SPHAERIUSIDAE].

Type species: Saprus griffithi Blackburn, 1904, by monotypy.

Family-group name: SAPRINI Nikolajev, 2008. Stem: Sapr-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

Sarothrias Grouvelle, 1918: 7. Current status: valid genus in JACOBSONIIDAE.

Type species: Sarothrias eximius Grouvelle, 1918, by monotypy.

Family-group name: SAROTHRIIDAE Crowson, 1955. Stem: Sarothri-.

Sarothrocrepis Chaudoir, 1850: 76. Current status: valid genus in CARABIDAE: HARPALINAE: CYCLOSOMINI.

Type species: Carabus corticalis Fabricius, 1801, by subsequent designation (Lacordaire 1854: 129).

Family-group name: SAROTHROCREPIDAE Chaudoir, 1876. Stem: Sarothrocrepid-.

Sarrotrium Illiger, 1798: 339. Current status: junior synonym of Orthocerus Latreille, 1797 in ZOPHERIDAE: COLYDIINAE: ORTHOCERINI.

Type species: Hispa mutica Linnaeus, 1767 (= Dermestes clavicornis Linnaeus, 1758), by monotypy.

Family-group name: SARROTRIIDES Billberg, 1820. Stem: Sarrotri-. Note: name placed on the Official Index of Rejected and Invalid Family-Group Names in Zoology (ICZN 1995d).

Sawadaeuops Legalov, 2003: 398. Current status: junior synonym of Riedeliops Alonso-Zarazaga & Lyal, 2002 in ATTELABIDAE: ATTELABINAE: EUOPINI.

Type species: Attelabus punctatostriatus Motschulsky, 1860, by original designation.

Family-group name: SAWADAEUOPSINA Legalov, 2007. Stem: Sawadaeuops- (ICZN 1999a, Article 29.4).

†Sayrevilleus Gratshev & Zherikhin, 2000b: 244. Current status: valid genus in ATTELABIDAE: SAYREVILLEINAE: †SAYREVILLEINI. Note: Upper Cretaceous (New Jersey, USA).

Type species: †Sayrevilleus grimaldii Gratshev & Zherikhin, 2000, by original designation.

Family-group name: †SANYREVILLEINA Legalov, 2003. Stem: Sayreville-.

Scalidia Erichson, 1845a: 305. Current status: junior synonym of Catogenus Westwood, 1830 in PASSANDRIDAE: PASSANDRINAE.

Type species: Passandra cylindricollis Lacordaire, 1854, by subsequent monotypy (Lacordaire 1854: 397).

Family-group name: SCALIDIINI Grouvelle, 1916. Stem: Scalidi-.

Scaphicoma Motschulsky, 1863: 435. Current status: valid genus in STAPHYLINIDAE: SCAPHIDIINAE: SCAPHISOMATINI.

Type species: Scaphicoma flavovittata Motschulsky, 1863, by monotypy.

Family-group name: SCAPHICOMITAE Achard, 1924. Stem: Scaphicom-.

Scaphidema L. Redtenbacher, 1848: 591. Current status: valid genus in TENEBRIONIDAE: DIAPERINAE: SCAPHIDEMINI.

Type species: Scaphidium bicolor Fabricius, 1798, by monotypy.

Family-group name: SCAPHIDEMINI Reitter, 1922. Stem: Scaphidem-.

Scaphidium G.-A. Olivier, 1790a: [20] 3. Current status: valid genus in STAPHYLINIDAE: SCAPHIDIINAE: SCAPHIDIINI.

Type species: Scaphidium quadrimaculatum G.-A. Olivier, 1790, by subsequent designation (Latreille 1810: 427).

Family-group name: SCAPHIDILIA Latreille, 1806. Stem: Scaphidi-.

Scaphisoma Leach, 1815a: 89. Current status: valid genus in STAPHYLINIDAE: SCAPHIDIINAE: SCAPHISOMATINI.

Type species: Silpha agaricina Linnaeus, 1758, by monotypy.

Family-group name: SCAPHISOMINI Casey, 1893. Stem: Scaphisomat-.

Scaphium W. Kirby, 1837: 108. Current status: valid genus in STAPHYLINIDAE: SCAPHIDIINAE: SCAPHIINI.

Type species: Scaphium castanipes W. Kirby, 1837, by monotypy.

Family-group name: SCAPHIITAE Achard, 1924. Stem: Scaphi-.

Scaphorhinadoretus Ohaus, 1912a: 426. Current status: valid genus in SCARABAEIDAE: RUTELINAE: ADORETINI: ADORETINA.

Type species: Scaphorhinadoretus bimaculatus Ohaus, 1912, by subsequent monotypy (Ohaus 1912a: 509).

Family-group name: SCAPHORHINADORETINA Ohaus, 1912. Stem: Scaphorhinadoret-.

Scapterus Dejean, 1826: 471. Current status: valid genus in CARABIDAE: SCARITINAE: SCARITINI: SCAPTERINA.

Type species: Scapterus guerini Dejean, 1826, by monotypy.

Family-group name: SCAPTÉRIDES Putzeys, 1867. Stem: Scapter-.

Scarabaeus Linnaeus, 1758: 342, 345. Current status: valid genus in SCARABAEIDAE: SCARABAEINAE: SCARABAEINI. Note: name placed on the Official List of Generic Names in Zoology (ICZN 2014a).

Type species: Scarabaeus sacer Linnaeus, 1758, by subsequent designation (Hope 1837: 22; see ICZN 2014a). Note: name placed on the Official List of Specific Names in Zoology (ICZN 2014a).

Family-group name: SCARABAEÏDES Latreille, 1802. Stem: Scarabae-.

Scarabatermes Howden, 1973: 29. Current status: valid genus in HYBOSORIDAE: CERATOCANTHINAE: SCARABATERMITINI.

Type species: Scarabatermes amazonensis Howden, 1973, by original designation.

Family-group name: SCARABATERMITINI Nikolajev, 1999. Stem: Scarabatermit-.

Scardamyctes Gistel, 1848b: xi. Current status: junior synonym of Magdalis Germar, 1817 in CURCULIONIDAE: MESOPTILIINAE: MAGDALIDINI. Note: unnecessary new replacement name for Thamnophilus Schönherr, 1823 (an unnecessary new replacement name for Magdalis Germar, 1817).

Type species: Curculio violaceus Linnaeus, 1758, by plenary power (see ICZN 1955a) (type fixation for Magdalis Germar).

Family-group name: SCARDAMYCTISIDAE Gistel, 1848. Stem: Scardamyct-. Note: nomen oblitum (see Bouchard et al. 2011: 621).

Scarites Fabricius, 1775: 249. Current status: valid genus in CARABIDAE: SCARITINAE: SCARITINI: SCARITINA.

Type species: Scarites subterraneus Fabricius, 1775, by subsequent designation (Stephens 1840: 352).

Family-group name: SCARITIDES Bonelli, 1810. Stem: Scarit-.

Scatimus Erichson, 1847c: 110. Current status: valid genus in SCARABAEIDAE: SCARABAEINAE: ATEUCHINI: SCATIMINA.

Type species: Scatimus cucullatus Erichson, 1847, by monotypy.

Family-group name: SCATIMINA Vaz-de-Mello, 2008. Stem: Scatim-.

Scatonomus Erichson, 1835: 256. Current status: valid genus in SCARABAEIDAE: SCARABAEINAE: DELTOCHILINI.

Type species: Scatonomus viridis Erichson, 1835, by subsequent designation (Westwood 1847a: 230)

Family-group name: SCATONOMIDES Lacordaire, 1855. Stem: Scatonom-.

Scaurus Fabricius, 1775: 253. Current status: valid genus in TENEBRIONIDAE: TENEBRIONINAE: SCAURINI.

Type species: Scaurus atratus Fabricius, 1775, by monotypy.

Family-group name: SCAURIDES Billberg, 1820. Stem: Scaur-.

Sceleocantha Newman, 1840a: 14. Current status: valid genus in CERAMBYCIDAE: PRIONINAE: SCELEOCANTHINI.

Type species: Prionus pilosicollis Hope, 1834, by subsequent designation (Chevrolat 1848: 411).

Family-group name: SCÉLÉOCANTHIDES Lacordaire, 1868. Stem: Sceleocanth-. Note: this name was treated as unavailable by Bouchard et al. (2011: 456) but was accepted as valid and attributed to Lacordaire, 1868 by Jin et al. (2023: 507).

Scelida Chapuis, 1875: 184. Current status: valid genus in CHRYSOMELIDAE: GALERUCINAE: LUPERINI.

Type species: Scelida elegans Chapuis, 1875, by monotypy.

Family-group name: SCELIDITES Chapuis, 1875. Stem: Scelid-.

Scelodonta Westwood, 1838b: 129. Current status: junior synonym of Heteraspis Chevrolat, 1836 in CHRYSOMELIDAE: EUMOLPINAE: BROMIINI.

Type species: Scelodonta curculionoides Westwood, 1838, by monotypy.

Family-group name: SCELODONTITES Chapuis, 1874. Stem: Scelodont-.

Sceloenopla Chevrolat in Dejean, 1836a: 364. Current status: valid genus in CHRYSOMELIDAE: CASSIDINAE: SCELOENOPLINI.

Type species: Hispa spinipes Fabricius, 1794 (= Hispa maculata G.-A. Olivier, 1792), by monotypy.

Family-group name: SCELOENOPLINI Uhmann, 1930. Stem: Sceloenopl-.

Scelophysa Burmeister, 1844: 168. Current status: valid genus in SCARABAEIDAE: MELOLONTHINAE: HOPLIINI: HOPLIINA.

Type species: Scelophysa pruinosa Burmeister, 1844, by subsequent designation (Dombrow 1999: 179).

Family-group name: SCELOPHYSIDES Péringuey, 1902. Stem: Scelophys-.

Sceptobius Sharp, 1883a: 211. Current status: valid genus in STAPHYLINIDAE: ALEOCHARINAE: SCEPTOBIINI.

Type species: Sceptobius dispar Sharp, 1883, by monotypy.

Family-group name: SCEPTOBIINI Seevers, 1978. Stem: Sceptobi-.

Schedarosus Reitter, 1876b: 42. Current status: junior synonym of Adelina Dejean, 1835 in TENEBRIONIDAE: DIAPERINAE: DIAPERINI: ADELININA.

Type species: Schedarosus cucujiformis Reitter, 1876 (= Pytho pallida Say, 1823), by subsequent designation (Löbl et al. 2008: 42).

Family-group name: SCHEDAROSINI Reitter, 1876. Stem: Schedaros-.

Schedlarius Wood, 1958: 103. Current status: valid genus in CURCULIONIDAE: PLATYPODINAE: SCHEDLARIINI. Note: new replacement name for Chapuisia Dugès, 1886.

Type species: Chapuisia mexicana Dugès, 1886, by monotypy (type fixation for Chapuisia Dugès).

Family-group name: SCHEDLARINI Wood & Bright, 1992. Stem: Schedlari-.

Schematiza Chevrolat in Dejean, 1836a: 377. Current status: valid genus in CHRYSOMELIDAE: GALERUCINAE: GALERUCINI.

Type species: Lycus laevigatus Fabricius, 1801, by subsequent designation (Barber 1947b: 242).

Family-group name: SCHEMATIZITES Chapuis, 1875. Stem: Schematiz-.

Schistodactylus Raffray, 1883: 243. Current status: valid genus in STAPHYLINIDAE: PSELAPHINAE: PSELAPHITAE: SCHISTODACTYLINI.

Type species: Schistodactylus phantasma Raffray, 1883, by monotypy.

Family-group name: SCHISTODACTYLINI Raffray, 1890. Stem: Schistodactyl-.

Schistogenia Kraatz, 1857a: 39. Current status: valid genus in STAPHYLINIDAE: ALEOCHARINAE: ATHETINI: SCHISTOGENIINA.

Type species: Schistogenia crenicollis Kraatz, 1857, by monotypy.

Family-group name: SCHISTOGENIAE Fenyes, 1919. Stem: Schistogeni-.

†Schistomerus Palmer, 1957: 259. Current status: valid genus in DYTISCIDAE: HYDROPORINAE: †SCHISTOMERINI. Note: Miocene (USA).

Type species: †Schistomerus californense Palmer, 1957, by original designation.

Family-group name: †SCHISTOMERINI Palmer, 1957. Stem: Schistomer-.

Schizelythron Kemner, 1925: 110. Current status: valid genus in STAPHYLINIDAE: ALEOCHARINAE: TRICHOPSENIINI.

Type species: Schizelythron javanicum Kemner, 1925, by original designation.

Family-group name: SCHIZELYTHRINAE Kemner, 1925. Stem: Schizelythr-.

†Schizocoleus Rohdendorf, 1961: 438. Current status: valid genus in †SCHIZOCOLEIDAE. Note: Permian/Early Triassic (Russia)

Type species: †Schizocoleus kuznetskiensis Rohdendorf, 1961, by original designation.

Family-group name: †SCHIZOCOLEIDAE Rohdendorf, 1961. Stem: Schizocole-.

Schizogenius Putzeys, 1846: 131. Current status: valid genus in CARABIDAE: SCARITINAE: CLIVININI: SCHIZOGENIINA.

Type species: Schizogenius strigicollis Putzeys, 1846, by subsequent designation (E. Desmarest 1851: 102).

Family-group name: SCHIZOGEINA Dostal, 2017: 115. Stem: Schizogeni-.

Schizognathus Fischer, 1823b: 263. Current status: valid genus in SCARABAEIDAE: RUTELINAE: ANOPLOGNATHINI: SCHIZOGNATHINA.

Type species: Schizognathus macleayi Fischer, 1823, by monotypy.

Family-group name: SCHIZOGNATHINA Ohaus, 1918. Stem: Schizognath-.

Schizonycha Dejean, 1833b: 161. Current status: valid genus in SCARABAEIDAE: MELOLONTHINAE: SCHIZONYCHINI.

Type species: Scarabaeus globator Fabricius, 1781, by subsequent designation (Pope 1960: 68).

Family-group name: SCHIZONYCHIDAE Burmeister, 1855. Stem: Schizonych-.

Schizophilus Bonvouloir, 1871: 73. Current status: valid genus in EUCNEMIDAE: SCHIZOPHILINAE. Note: this name was first used by Gemminger in Gemminger and von Harold (1869b: 1480) but is unavailable from that date since it was not accompanied by a description or indication (ICZN 1999a, Article 12.1).

Type species: Elater subrufa Randall, 1838, by subsequent monotypy (Bonvouloir 1872b: pl. 34, fig. 5). Note: the name on pl. 34 of Bonvouloir’s work is “Schizophilus trilobatus” but it was changed on p. 710 and in the “Corrections” (p. 885) to S. subrufus; the change was proposed at a later date (i.e., 1875) than the plate (1872b) but in one of the parts of the same volume and therefore the spelling must be corrected (ICZN 1999a, Article 32.5.1.1).

Family-group name: SCHIZOPHILINI Muona, 1993. Stem: Schizophil-.

†Schizophorus Ponomarenko, 1968: 130. Current status: invalid senior synonym of †Phoroschizus Bouchard & Bousquet, 2020 in †PHOROSCHIZIDAE. Note: junior homonym of †Schizophorus Balashova, 1953 [Trilobita]; Upper Jurassic (Kazakhstan).

Type species: †Schizophorus crassus Ponomarenko, 1968, by original designation.

Family-group name: †SCHIZOPHORIDAE Ponomarenko, 1968. Stem: Schizophor-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

Schizopus J.L. LeConte, 1858a: 70. Current status: valid genus in SCHIZOPODIDAE: SCHIZOPODINI. Note: name placed on the Official List of Generic names in Zoology (ICZN 1993e).

Type species: Schizopus laetus J.L. LeConte, 1858, by monotypy (see ICZN 1993e). Note: name placed on the Official List of Specific Names in Zoology (ICZN 1993e).

Family-group name: SCHIZOPODIDAE J.L. LeConte, 1859. Stem: Schizopod-.

Schizorhina W. Kirby, 1825: 570. Current status: valid genus in SCARABAEIDAE: CETONIINAE: SCHIZORHININI: SCHIZORHININA.

Type species: Cetonia atropunctata W. Kirby, 1819, by original designation.

Family-group name: SCHIZORRHINIDAE Burmeister, 1842. Stem: Schizorhin-.

Schoenherriella Viana, 1952: 232. Current status: junior synonym of Epipedophyes G.A.K. Marshall, 1946 in CURCULIONIDAE: MOLYTINAE: PETALOCHILINI. Note: new replacement name for Epipedus Schönherr, 1842.

Type species: Epipedus squamifer Boheman, 1842, by original designation (type fixation for Epipedus Schönherr).

Family-group name: SCHOENHERRIELLINAE Viana, 1952. Stem: Schoenherriell-.

Schyzoschelus C. Koch, 1954: 72. Current status: valid genus in TENEBRIONIDAE: BLAPTINAE: PLATYNOTINI: EURYNOTINA.

Type species: Schyzoschelus kaszabi C. Koch, 1954, by original designation.

Family-group name: SCHYZOSCHELINA C. Koch, 1956. Stem: Schyzoschel-.

†Sciabregma Scudder, 1893: 146. Current status: valid genus in CURCULIONIDAE: MOLYTINAE: †SCIABREGMINI. Note: Tertiary (USA).

Type species: †Sciabregma rugosa Scudder, 1893, by monotypy.

Family-group name: †SCIABREGMINI Legalov, Kirejtshuk & Nel, 2020b: 736. Stem: Sciabregm-.

Sciacharis Broun, 1893b: 1062. Current status: valid genus in STAPHYLINIDAE: SCYDMAENINAE: SCYDMAENITAE: STENICHNINI.

Type species: Sciacharis fulva Broun, 1893, by monotypy.

Family-group name: SCIACHARINI Csiki, 1919. Stem: Sciacharit-.

Sciaphilus Schönherr, 1823: 1139. Current status: valid genus in CURCULIONIDAE: ENTIMINAE: CYPHICERITAE: SCIAPHILINI.

Type species: Curculio muricatus Fabricius, 1792 (= Curculio asperatus Bonsdorff, 1785), by original designation.

Family-group name: SCIAPHILINA Sharp, 1891. Stem: Sciaphil-.

Sciaphyes Jeannel, 1910: 7. Current status: valid genus in LEIODIDAE: CHOLEVINAE: SCIAPHYINI.

Type species: Bathyscia sibirica Reitter, 1887, by original designation.

Family-group name: SCIAPHYINI Perreau, 2000. Stem: Sciaphy-.

Sciatrophes Blackburn, 1903: 100. Current status: junior synonym of Baeocera Erichson, 1845 in STAPHYLINIDAE: SCAPHIDIINAE: SCAPHISOMATINI.

Type species: Sciatrophes latens Blackburn, 1903, by monotypy.

Family-group name: SCIATROPHITAE Achard, 1924. Stem: Sciatroph-.

Scirtes Illiger, 1807a: 301. Current status: valid genus in SCIRTIDAE: SCIRTINAE.

Type species: Chrysomela hemisphaerica Linnaeus, 1758, by subsequent monotypy (Leach 1815a: 86).

Family-group name: SCIRTESIDAE Fleming, 1821. Stem: Scirt-.

Scitala Erichson, 1842: 166. Current status: valid genus in SCARABAEIDAE: SERICOIDINAE: SCITALINI.

Type species: Scitala sericans Erichson, 1842, by subsequent designation (Blackburn 1898: 37).

Family-group name: SCITALINI Britton, 1957. Stem: Scital-.

Sclerastes Gistel, 1856: 368. Current status: junior synonym of Ptilinus Geoffroy, 1762 in PTINIDAE: PTILININAE.

Type species: Ptilinus costatus Gyllenhal, 1827 (= Bostrichus fuscus Geoffroy, 1785), by subsequent designation (Bouchard et al. 2011: 340).

Family-group name: SCLERASTEIDAE Gistel, 1856. Stem: Sclerast-.

Sclerocardius Schönherr, 1847: 82. Current status: valid genus in CURCULIONIDAE: MOLYTINAE: ITHYPORINI: SCLEROCARDIINA.

Type species: Sclerocardius bohemani Schönherr, 1847 (= Heteropus africanus Boheman, 1845), by original designation.

Family-group name: SCLÉROCARDIIDES Lacordaire, 1865. Stem: Sclerocardi-.

Scleron Hope, 1841a: 111. Current status: junior synonym of Sclerum Dejean, 1834 in TENEBRIONIDAE: BLAPTINAE: OPATRINI: SCLERINA.

Type species: Opatrum orientale Fabricius, 1775, by original designation.

Family-group name: SCLÉRIDES Lacordaire, 1859. Stem: Scler-.

Scleropterus Schönherr, 1825: 585. Current status: valid genus in CURCULIONIDAE: CEUTORHYNCHINAE: SCLEROPTERINI.

Type species: Cryptorhynchus serratus Germar, 1823, by original designation.

Family-group name: SCLEROPTERIDAE Schultze, 1902. Stem: Scleropter-. Note: junior homonym of SCLEROPTERINAE Saussure, 1877 (type genus Scleropterus de Haan, 1842) in Orthoptera; Saussure’s name is permanently invalid (based on a preoccupied type genus).

Sclerostomus Burmeister, 1847: 410, 423. Current status: valid genus in LUCANIDAE: LUCANINAE: SCLEROSTOMINI. Note: new replacement name for Sclerognathus Hope, 1845.

Type species: Sclerognathus costatus Hope, 1845, by monotypy (type fixation for Sclerognathus Hope).

Family-group name: SCLEROSTOMINI Benesh, 1955. Stem: Sclerostom-.

Scolopterus A. White, 1846: 14. Current status: valid genus in CURCULIONIDAE: CURCULIONINAE: EUGNOMINI: EUGNOMINA.

Type species: Scolopterus tetracanthus A. White, 1846, by subsequent designation (Broun 1880: 473).

Family-group name: SCOLOPTÉRIDES Lacordaire, 1863. Stem: Scolopter-.

Scolytoplatypus C. Schaufuss, 1891b: 31. Current status: valid genus in CURCULIONIDAE: SCOLYTINAE: SCOLYTOPLATYPODINI.

Type species: Scolytoplatypus permirus C. Schaufuss, 1891, by monotypy.

Family-group name: SCOLYTOPLATYPINI W.F.H. Blandford, 1893. Stem: Scolytoplatypod-.

Scolytus Fabricius, 1790: 221. Current status: invalid senior synonym of Omophron Latreille, 1802 in CARABIDAE: OMOPHRONINAE. Note: junior homonym of Scolytus Geoffroy, 1762 [Coleoptera: CURCULIONIDAE].

Type species: Carabus limbatus Fabricius, 1777, by subsequent designation (Latreille 1810: 426) (type fixation for Omophron Latreille).

Family-group name: SCOLYTI Motschulsky, 1850. Stem: Scolyt-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

Scolytus Geoffroy, 1762: 309. Current status: valid genus in CURCULIONIDAE: SCOLYTINAE: SCOLYTINI. Note: name placed on the Official List of Generic Names in Zoology (ICZN 1963c).

Type species: Bostrichus scolytus Fabricius, 1775, by plenary power (see ICZN 1963c). Note: name placed on the Official List of Specific Names in Zoology (ICZN 1963c).

Family-group name: SCOLITARII Latreille, 1804. Stem: Scolyt-.

Scopadus Pascoe, 1857: 100. Current status: valid genus in CERAMBYCIDAE: LAMIINAE: CYRTININI.

Type species: Scopadus ciliatus Pascoe, 1857, by monotypy.

Family-group name: SCOPADINI Villiers, 1980. Stem: Scopad-.

Scopaeus Erichson, 1839b: 29. Current status: valid genus in STAPHYLINIDAE: PAEDERINAE: LATHROBIINI: SCOPAEINA.

Type species: Paederus laevigatus Gyllenhal, 1827, by subsequent designation (Duponchel 1841b: 57). Note: no species were originally included in this genus by Erichson (1839b); the first ones were the nine species listed by Erichson (1840a: 605–609).

Family-group name: SCOPÉATES Mulsant & Rey, 1878. Stem: Scopae-.

Scopodes Erichson, 1842: 123. Current status: valid genus in CARABIDAE: HARPALINAE: ODACANTHINI: PENTAGONICINA.

Type species: Scopodes boops Erichson, 1842, by monotypy.

Family-group name: SCOPODINAE H.W. Bates, 1874. Stem: Scopod-.

Scortizus Westwood, 1834: 119. Current status: valid genus in LUCANIDAE: LUCANINAE: SCLEROSTOMINI.

Type species: Pholidotus irroratus Hope, 1833 (= Lucanus maculatus Klug, 1825), by monotypy.

Family-group name: SCORTIZINI Benesh, 1955. Stem: Scortiz-.

Scotobius Germar, 1823: 135. Current status: valid genus in TENEBRIONIDAE: TENEBRIONINAE: SCOTOBIINI.

Type species: Scotobius pilularius Germar, 1823, by subsequent designation (Lacordaire 1830a: 282).

Family-group name: SCOTOBITES Solier, 1838. Stem: Scotobi-.

Scotocryptus M. Girard, 1874a: 123. Current status: valid genus in LEIODIDAE: LEIODINAE: SCOTOCRYPTINI.

Type species: Scotocryptus meliponae M. Girard, 1874, by monotypy. Note: combined description of a new genus and a single new species (ICZN 1999a, Article 12.2.6); the genus was redescribed and the species subsequently formally described by M. Girard (1875: 574).

Family-group name: SCOTOCRYPTINI Reitter, 1884. Stem: Scotocrypt-.

Scotodipnus Schaum, 1860: 667. Current status: valid genus in CARABIDAE: TRECHINAE: ANILLINI.

Type species: Anillus glaber Baudi di Selve, 1859, by monotypy.

Family-group name: SCOTODIPNINA Jeannel, 1937. Stem: Scotodipn-.

Scotodytes Saulcy, 1865: 18. Current status: valid subgenus of Phloeocharis Mannerheim, 1830 in STAPHYLINIDAE: PHLOEOCHARINAE.

Type species: Scotodytes paradoxus Saulcy, 1865, by monotypy.

Family-group name: SCOTODYTIDAE Saulcy, 1870. Stem: Scotodyt-.

Scotoscymnus Weise, 1901: 458. Current status: junior synonym of Scymnomorphus Weise, 1897 in COCCINELLIDAE: MICROWEISEINAE: MICROWEISEINI. Note: unnecessary new replacement name for Scymnomorphus Weise, 1897.

Type species: Scymnomorphus rotundatus Weise, 1897, by subsequent designation (Pope 1962: 628).

Family-group name: SCOTOSCYMNINAE Duverger, 2003. Stem: Scotoscymn-.

Scraptia Latreille, 1806: 199. Current status: valid genus in SCRAPTIIDAE: SCRAPTIINAE: SCRAPTIINI.

Type species: Melandrya fusca Latreille, 1804 (= Melyris dubia G.-A. Olivier, 1790), by monotypy.

Family-group name: SCRAPTIAEIDAE Gistel, 1848. Stem: Scrapti-.

Scydmaenus Latreille, 1802b: 116. Current status: valid genus in STAPHYLINIDAE: SCYDMAENINAE: SCYDMAENITAE: SCYDMAENINI.

Type species: here fixed (under ICZN 1999a, Article 70.3.2) as Scydmaenus tarsatus P.W.J. Müller & Kunze, 1822, misidentified as Pselaphus hellwigii Herbst, 1791 (sensu Paykull, 1800) in the original designation by monotypy in Latreille (1802b: 116).

Family-group name: SCYDMAENIDES Leach, 1815. Stem: Scydmaen-.

Scymnillus G.H. Horn, 1895a: 110. Current status: junior synonym of Zilus Mulsant, 1850 in COCCINELLIDAE: COCCINELLINAE: COCCIDULINI.

Type species: Scymnillus aterrimus G.H. Horn, 1895, by monotypy.

Family-group name: SCYMNILLINI Casey, 1899. Stem: Scymnill-.

Scymnus Kugelann, 1794: 545. Current status: valid genus in COCCINELLIDAE: COCCINELLINAE: COCCIDULINI.

Type species: Scymnus nigrinus Kugelann, 1794, by subsequent designation (Westwood 1838c: 43). Note: this name is listed as nomen protectum (over Coccinella minima O.F. Müller, 1776) by Kovář (2007: 590) but we were unable to verify that the conditions of Article23.9.1.2 (ICZN 1999a) were met.

Family-group name: SCYMNIAIRES Mulsant, 1846. Stem: Scymn-.

Scythropus Schönherr, 1826: 12, 140. Current status: junior synonym of Pachyrhinus Schönherr, 1823 in CURCULIONIDAE: ENTIMINAE: CYPHICERITAE: SCYTHROPINI. Note: two original spellings were used, including Scytropus (p. 338); subsequently, Schönherr (1840b: 301) used Scythropus and therefore acted as First Reviser (ICZN 1999a, Article 24.2.4).

Type species: Curculio mustela Herbst, 1797 (= Curculio squamulosus Herbst, 1795), by original designation.

Family-group name: SCYTHROPIDES Lacordaire, 1863. Stem: Scythrop-.

Selatosomus Stephens, 1830: 268. Current status: valid genus in ELATERIDAE: DENDROMETRINAE: SELATOSOMINI: SELATOSOMINA.

Type species: Elater aeneus Linnaeus, 1758, by subsequent designation (Curtis 1838a: pl. 694).

Family-group name: SELATOSOMINI Schimmel, Tarnawski, Han & Platia, 2015: 30. Stem: Selatosom-.

†Selengarhynchus Gratshev & Legalov, 2009: 415. Current status: valid genus in NEMONYCHIDAE: †SELENGARHYNCHINAE. Note: Lower Cretaceous (Mongolia).

Type species: †Selengarhynchus ovalis Gratshev & Legalov, 2009, by original designation.

Family-group name: †SELENGARHYNCHINAE Gratshev & Legalov, 2009. Stem: Selengarhynch-.

Selenophorus Dejean, 1829: 4, 80. Current status: valid genus in CARABIDAE: HARPALINAE: HARPALINI: HARPALINA.

Type species: Carabus palliatus Fabricius, 1798, by subsequent designation (Hope 1838a: 84).

Family-group name: SELENOPHORINI Casey, 1914. Stem: Selenophor-.

Selina Motschulsky, 1857c: 110. Current status: valid genus in CARABIDAE: HARPALINAE: LACHNOPHORINI: SELININA.

Type species: Selina westermanni Motschulsky, 1857, by monotypy.

Family-group name: SELININI Jeannel, 1948. Stem: Selin-.

Selvadius Casey, 1899: 134, 137. Current status: valid genus in COCCINELLIDAE: COCCINELLINAE: HYPERASPIDINI: SELVADIINA.

Type species: Selvadius rectus Casey, 1899, by original designation.

Family-group name: SELVADIINI Gordon, 1985. Stem: Selvadi-.

Semicyclus Kaup, 1871: 28. Current status: valid genus in PASSALIDAE: PASSALINAE: SOLENOCYCLINI.

Type species: Semicyclus grayi Kaup, 1871, by monotypy.

Family-group name: SEMICYCLINAE Kuwert, 1891. Stem: Semicycl-.

Semiotus Eschscholtz, 1829a: 31. Current status: valid genus in ELATERIDAE: DENDROMETRINAE: SEMIOTINI.

Type species: Elater furcatus Fabricius, 1792, by subsequent designation (Hyslop 1921: 668).

Family-group name: SEMIOTINA Jakobson, 1913. Stem: Semiot-.

Senodonia Laporte, 1838: 12. Current status: valid genus in ELATERIDAE: LISSOMINAE: PROTELATERINI.

Type species: Semiotus quadricollis Laporte, 1838, by monotypy.

Family-group name: SENODONIINAE Schenkling, 1927. Stem: Senodoni-.

Sepedonastes Gistel, 1856: 382. Current status: junior synonym of Phaleria Latreille, 1802 in TENEBRIONIDAE: DIAPERINAE: PHALERIINI.

Type species: Tenebrio bimaculatus Herbst, 1799 (= Dytiscus bimaculatus Linnaeus, 1767), by subsequent designation (Bouchard et al. 2005: 501).

Family-group name: SEPEDONASTIDAE Gistel, 1856. Stem: Sepedonast-.

Sepedophilus Gistel, 1856: 267, 386. Current status: valid genus in STAPHYLINIDAE: TACHYPORINAE: TACHYPORINI: EUCONOSOMATINA.

Type species: Staphylinus pubescens Paykull, 1790 (= Staphylinus littoreus Linnaeus, 1758), by subsequent designation (Blackwelder 1952: 350). Note: Staphylinus littoreus Linnaeus, 1758 was placed on the Official List of Specific Names in Zoology (ICZN 2003b).

Family-group name: SEPEDOPHILINI Ádám, 2001. Stem: Sepedophil-.

Sepidium Fabricius, 1775: 250. Current status: valid genus in TENEBRIONIDAE: PIMELIINAE: SEPIDIINI: SEPIDIINA.

Type species: Sepidium tricuspidatum Fabricius, 1775, by subsequent designation (Latreille 1810: 429).

Family-group name: SEPIDIAE Eschscholtz, 1829. Stem: Sepidi-.

†Septiventer Bai, Ren, Shih & X.-K. Yang in Bai et al., 2012b: 364. Current status: valid genus in †SEPTIVENTERIDAE. Note: Lower Cretaceous (China).

Type species: †Septiventer quadridentatus Bai, Ren, Shih & X.-K. Yang, 2012, by original designation.

Family-group name: †SEPTIVENTERIDAE Bai, Ren, Shih & X.-K. Yang in Bai et al., 2012b: 364. Stem: Septiventer- (ICZN 1999a, Article 29.4).

Serangium Blackburn, 1889: 187, 209. Current status: valid genus in COCCINELLIDAE: MICROWEISEINAE: SERANGIINI.

Type species: Serangium mysticum Blackburn, 1889, by monotypy.

Family-group name: SERANGIINI Blackwelder, 1945. Stem: Serangi-.

Serica W.S. MacLeay, 1819: 146. Current status: valid genus in SCARABAEIDAE: SERICINAE: SERICINI: SERICINA.

Type species: Scarabaeus brunneus Linnaeus, 1758 [as brunnus], by monotypy.

Family-group name: SERICIDAE W. Kirby, 1837. Stem: Seric-.

Sericoda W. Kirby, 1837: 14. Current status: valid genus in CARABIDAE: HARPALINAE: PLATYNINI.

Type species: Sericoda bembidioides W. Kirby, 1837, by monotypy.

Family-group name: SERICODIADAE W. Kirby, 1837. Stem: Sericod-.

Sericoderus Stephens, 1829a: 158. Current status: valid genus in CORYLOPHIDAE: CORYLOPHINAE: SERICODERINI.

Type species: Sericoderus thoracicus Stephens, 1829 (= Cossyphus lateralis Gyllenhal, 1827), by subsequent monotypy (Stephens 1829a: 188).

Family-group name: SERICODERINA A. Matthews, 1886. Stem: Sericoder-.

Sericoides Guérin-Méneville, 1839b: 301. Current status: valid genus in SCARABAEIDAE: SERICOIDINAE: SERICOIDINI.

Type species: Melolontha glacialis Fabricius, 1775, by subsequent designation (E. Desmarest 1860: 68).

Family-group name: SERICOIDEAE Erichson, 1847. Stem: Sericoid-.

Sericosomus Dejean, 1833a: 96. Current status: junior synonym of Sericus Eschscholtz, 1829 in ELATERIDAE: ELATERINAE: ELATERINI. Note: unnecessary new replacement name for Sericus Eschscholtz, 1829.

Type species: Elater brunneus Linnaeus, 1758, by subsequent designation (Curtis 1838a: pl. 694) (type fixation for Sericus Eschscholtz).

Family-group name: SERICOSOMINA Hyslop, 1917. Stem: Sericosom-.

Sermyla Chapuis, 1875: 224. Current status: invalid senior synonym of Sermylassa Reitter, 1912 in CHRYSOMELIDAE: GALERUCINAE: HYLASPINI. Note: junior homonym of Sermyla Adams, 1854 [Mollusca] and Sermyla Walker, 1854 [Lepidoptera].

Type species: Chrysomela halensis Linnaeus, 1767, by monotypy.

Family-group name: SERMYLITES Chapuis, 1875. Stem: Sermyl-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

Sermylassa Reitter, 1913c: 138. Current status: valid genus in CHRYSOMELIDAE: GALERUCINAE: HYLASPINI. Note: new replacement name for Sermyla Chapuis, 1875.

Type species: Chrysomela halensis Linnaeus, 1767, by monotypy (type fixation for Sermyla Chapuis).

Family-group name: SERMYLASSINI Mroczkowski, 1991. Stem: Sermylass-.

Serraticollis B.E. White, 1942: 17. Current status: junior synonym of Aulacothorax Boheman, 1858 in CHRYSOMELIDAE: GALERUCINAE: SERRATICOLLINI.

Type species: Serraticollis rhois B.E. White, 1942 (= Haltica recticollis J.L. LeConte, 1861), by original designation.

Family-group name: SERRATICOLLINI B.E. White, 1942. Stem: Serraticoll-.

Serropalpus Hellenius, 1786: 310. Current status: valid genus in MELANDRYIDAE: MELANDRYINAE: SERROPALPINI.

Type species: Serropalpus striatus Hellenius, 1786 (= Mordella barbata Schaller, 1783), by subsequent designation (Blanchard in Audouin et al. 1844: pl. 53).

Family-group name: SERROPALPIDES Latreille, 1829. Stem: Serropalp-.

Sertorius Kaup, 1871: 114. Current status: invalid senior synonym of Arrox Zang, 1905 in PASSALIDAE: PASSALINAE: PROCULINI. Note: junior homonym of Sertorius Stål, 1866 [Hemiptera].

Type species: Sertorius agassizi Kaup, 1871, by monotypy.

Family-group name: SERTORINAE Kuwert, 1891. Stem: Sertori-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

Sestyra Pascoe, 1867a: 513. Current status: valid genus in CERAMBYCIDAE: CERAMBYCINAE: SESTYRINI.

Type species: Sestyra cephalotes Pascoe, 1867, by original designation.

Family-group name: SESTYRIDES Lacordaire, 1868. Stem: Sestyr-.

Setapion F. Balfour-Browne, 1944: 71. Current status: valid genus in BRENTIDAE: APIONINAE: SETAPIITAE.

Type species: Setapion quantillum F. Balfour-Browne, 1944, by original designation.

Family-group name: SETAPIITAE Legalov, 2023c: 30. Stem: Setapi-. Note: SETAPIITAE Legalov (2018c: 811) was published in a work that is nomenclaturally unavailable (ICZN 2012f, Article 8.5.3).

Setaria Mulsant & Rey, 1864a: 1. Current status: invalid senior synonym of Setariola Jakobson, 1915 in EROTYLIDAE: PHARAXONOTHINAE. Note: junior homonym of Setaria Viborg, 1795 [Vermes] and Setaria Blyth, 1844 [Aves].

Type species: Setaria sericea Mulsant & Rey, 1864, by monotypy.

Family-group name: SETARIINI Casey, 1900. Stem: Setari-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

Setariola Jakobson, 1915: 941. Current status: valid genus in EROTYLIDAE: PHARAXONOTHINAE. Note: new replacement name for Setaria Mulsant & Rey, 1864.

Type species: Setaria sericea Mulsant & Rey, 1864, by monotypy (type fixation for Setaria Mulsant & Rey).

Family-group name: SETARIOLINAE Crowson, 1952. Stem: Setariol-.

Shirozuella Sasaji, 1967: 24. Current status: valid genus in COCCINELLIDAE: COCCINELLINAE: SHIROZUELLINI.

Type species: Shirozuella mirabilis Sasaji, 1967, by original designation.

Family-group name: SHIROZUELLINI Sasaji, 1967. Stem: Shirozuell-.

Siagona Latreille, 1804a: 141. Current status: valid genus in CARABIDAE: SIAGONINAE: SIAGONINI.

Type species: Cucujus rufipes Fabricius, 1792, by monotypy.

Family-group name: SIAGONES Bonelli, 1813. Stem: Siagon-.

Siamites H. Franz, 1989: 44. Current status: valid genus in STAPHYLINIDAE: SCYDMAENINAE: SCYDMAENITAE: STENICHNINI.

Type species: Siamites loebli H. Franz, 1989, by original designation.

Family-group name: SIAMITINI H. Franz, 1989. Stem: Siamit-.

Sibinia Germar, 1817b: 340. Current status: valid genus in CURCULIONIDAE: CURCULIONINAE: TYCHIINI: TYCHIINA.

Type species: Curculio viscariae Linnaeus, 1761, by subsequent designation (Schönherr 1825: 583).

Family-group name: SIBYNIDAE Marseul, 1863. Stem: Sibini-.

Sibylla J. Thomson, 1858a: 406. Current status: invalid senior synonym of Zehra Özdikmen, 2008 in CERAMBYCIDAE: CERAMBYCINAE: BIMIINI. Note: junior homonym of Sibylla Stål, 1856 [Orthoptera].

Type species: Phoedinus coemeterii J. Thomson, 1856, by monotypy.

Family-group name: SIBYLLINI Cerda, 1973. Stem: Sibyll-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

Siderodactylus Schönherr, 1834a: 125. Current status: junior synonym of Hadromerus Schönherr, 1823 in CURCULIONIDAE: ENTIMINAE: POLYDRUSITAE: TANYMECINI: TANYMECINA.

Type species: Curculio sagittarius G.-A. Olivier, 1807, by original designation.

Family-group name: SIDERODACTYLIDES Jekel, 1865. Stem: Siderodactyl-.

Siettitia Abeille de Perrin, 1904: 226. Current status: valid genus in DYTISCIDAE: HYDROPORINAE: HYDROPORINI: SIETTITIINA.

Type species: Siettitia balsetensis Abeille de Perrin, 1904, by monotypy.

Family-group name: SIETTITIINI Smrž, 1982. Stem: Siettiti-.

Sikhotealinia Lafer, 1996: 392. Current status: valid genus in JURODIDAE.

Type species: Sikhotealinia zhiltzovae Lafer, 1996, by original designation.

Family-group name: SIKHOTEALINIIDAE Lafer, 1996. Stem: Sikhotealini-.

Silis Charpentier, 1825: 194. Current status: valid genus in CANTHARIDAE: SILINAE: SILINI.

Type species: Silis rubricollis Charpentier, 1825 (= Cantharis ruficollis Fabricius, 1775), by subsequent designation (Hope 1841a: 141). Note: Kazantsev (2011: 28) mentioned that the type species was designated by Westwood (1838c) but Westwood cited “Cantharis ruficollis Fab.” as type species, which is not an originally included species, and he did not mention that the species was a synonym of Silis rubricollis Charpentier, one of the two originally included species.

Family-group name: SILIAIRES Mulsant, 1862. Stem: Sil-.

Silluvia Landin, 1950: 5. Current status: valid genus in SCARABAEIDAE: AEGIALIINAE: AEGIALIINI.

Type species: Silluvia elongata Landin, 1950, by original designation.

Family-group name: SILLUVIINAE Landin, 1950. Stem: Silluvi-.

Silpha Linnaeus, 1758: 342, 359. Current status: valid genus in STAPHYLINIDAE: SILPHINAE: SILPHINI.

Type species [suggested]: Silpha obscura Linnaeus, 1758, by subsequent designation (Curtis 1839: pl. 742). Note: the valid type species fixation is that of Latreille (1810: 427) who selected Silpha littoralis Linnaeus, 1758, but that species is the type species of the genus Necrodes Leach, 1815; a subsequent designation based on an originally included nominal species is that of Westwood (1838c: 11), who selected Silpha quadripunctata Linnaeus, 1758, but that species is the type species of the genus Glischrochilus Reitter, 1873; an application to the Commission is necessary to fix Silpha obscura Linnaeus, 1758 as the type species of Silpha Linnaeus, 1758.

Family-group name: SILPHALES Latreille, 1806. Stem: Silph-.

Silphotelus Broun, 1895: 83. Current status: valid genus in STAPHYLINIDAE: PROTEININAE: SILPHOTELINI.

Type species: Silphotelus nitidus Broun, 1895, by monotypy.

Family-group name: SILPHOTELINI Newton & Thayer, 1995. Stem: Silphotel-.

Silusa Erichson, 1837: 377. Current status: valid genus in STAPHYLINIDAE: ALEOCHARINAE: HOMALOTINI: SILUSINA.

Type species: Silusa rubiginosa Erichson, 1837, by monotypy.

Family-group name: SILUSAE Fenyes, 1919. Stem: Silus-.

Silvanus Latreille, 1804a: 158. Current status: valid genus in SILVANIDAE: SILVANINAE.

Type species: Ips unidentata G.-A. Olivier, 1790, by monotypy.

Family-group name: SYLVANIDAE W. Kirby, 1837. Stem: Silvan-.

Simo Dejean, 1821: 92. Current status: valid genus in CURCULIONIDAE: ENTIMINAE: CYPHICERITAE: PERITELINI: SIMONINA.

Type species: Curculio hirticornis Herbst, 1795, by monotypy.

Family-group name: SIMOINI Pierce, 1913. Stem: Simon-.

Simous Chaudoir, 1882: 373. Current status: valid genus in CARABIDAE: HARPALINAE: OODINI.

Type species: Oodes nigriceps Wiedemann, 1821, by subsequent designation (Andrewes 1938: 138).

Family-group name: SIMOINI Basilewsky, 1953. Stem: Simo-.

Simplocaria Stephens, 1829b: 9. Current status: valid genus in BYRRHIDAE: BYRRHINAE: SIMPLOCARIINI. Note: name placed on the Official List of Generic Names in Zoology (ICZN 1985c).

Type species: Byrrhus semistriatus Fabricius, 1794, by subsequent designation (Curtis 1830: pl. 335; see ICZN 1985c where the same type species was recorded on the Official List as subsequently designated by Jacquelin du Val (1859b: 267)). Note: name placed on the Official List of Specific Names in Zoology (ICZN 1985c).

Family-group name: SIMPLOCARIATES Mulsant & Rey, 1869. Stem: Simplocari-.

Simus Raffray, 1882: 6. Current status: invalid senior synonym of Ipsimus Reitter, 1885 in STAPHYLINIDAE: PSELAPHINAE: GONIACERITAE: GONIACERINI. Note: junior homonym of Simus Bonaparte, 1838 [Reptilia].

Type species: Simus fracticornis Raffray, 1882, by monotypy.

Family-group name: SIMINI L.W. Schaufuss, 1890. Stem: Sim-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

Sinanarchusa Pace, 2013b: 23. Current status: valid genus in STAPHYLINIDAE: ALEOCHARINAE: HIMALUSINI.

Type species: Sinanarchusa daxuensis Pace, 2013, by original designation.

Family-group name: SINANARCHUSINI Pace, 2013b: 22. Stem: Sinanarchus-.

Singhikalia Kapur, 1963: 16. Current status: valid genus in COCCINELLIDAE: COCCINELLINAE: COCCINELLINI.

Type species: Singhikalia ornata Kapur, 1963, by original designation.

Family-group name: SINGHIKALIINI Miyatake, 1972. Stem: Singhikali-.

Singiliomimus Péringuey, 1896: 243. Current status: valid genus in CARABIDAE: HARPALINAE: LEBIINI: DROMIUSINA.

Type species: Singiliomimus posticalis Péringuey, 1896, by subsequent designation (Lorenz 1998: 161).

Family-group name: SINGILIOMIMINA Basilewsky, 1984. Stem: Singiliomim-.

Singilis Rambur, 1837: 25. Current status: valid genus in CARABIDAE: HARPALINAE: LEBIINI: DROMIUSINA.

Type species: Singilis bicolor Rambur, 1837, by subsequent designation (Antoine 1963: 551).

Family-group name: SINGILINI Jeannel, 1949. Stem: Singil-.

†Sinisilvana Hong, 2002: 134. Current status: valid genus in †SINISILVANIDAE. Note: Eocene (China).

Type species: †Sinisilvana fushunensis Hong, 2002, by original designation.

Family-group name: †SINISILVANIDAE Hong, 2002. Stem: Sinisilvan-.

Sinodendron Hellwig in Fabricius, 1791: 18. Current status: valid genus in LUCANIDAE: SYNDESINAE: SINODENDRINI. Note: this name was first proposed as a synonym of Ligniperda Fabricius, 1791 and, since it was used as valid before 1961 (e.g., Latreille 1804a: 149), it therefore dates from its proposal in synonymy (ICZN 1999a, Article 11.6.1), as opposed to being credited to Hellwig (1792: 391) as in recent literature (e.g., Bartolozzi et al. 2016: 60).

Type species: Scarabaeus cylindricus Linnaeus, 1758, by monotypy.

Family-group name: SINODENDRIENS Mulsant, 1842. Stem: Sinodendr-.

†Sinodryopites Hong, 2002: 110. Current status: valid genus in SCIRTIDAE incertae sedis. Note: Eocene (China).

Type species: †Sinodryopites ovalifemorales Hong, 2002, by original designation.

Family-group name: †SINODRYOPITIDAE Hong, 2002. Stem: Sinodryopit-.

Sinopyrophorus Bi & Li in Bi et al., 2019: 83. Current status: valid genus in SINOPYROPHORIDAE.

Type species: Sinopyrophorus schimmeli Bi & X.-Y. Li, 2019, by original designation.

Family-group name: SINOPYROPHORINAE Bi & X.-Y. Li in Bi et al., 2019: 89. Stem: Sinopyrophor-.

Sinoxylon Duftschmid, 1825: 85. Current status: valid genus in BOSTRICHIDAE: BOSTRICHINAE: SINOXYLINI.

Type species: Dermestes muricatus Linnaeus, 1767 nomen oblitum (= Bostrichus sexdentatus G.-A. Olivier, 1790 nomen protectum), by monotypy. Note: see Ivie (2010: 40) regarding the identity of the type species.

Family-group name: SINOXYLIDAE Marseul, 1857. Stem: Sinoxyl-.

Sinozolus Deuve, 1997: 32. Current status: valid genus in CARABIDAE: TRECHINAE: SINOZOLINI: SINOZOLINA.

Type species: Sinozolus yuae Deuve, 1997, by monotypy. Note: combined description of a new genus and a single new species (ICZN 1999a, Article 13.4).

Family-group name: SINOZOLINI Deuve, 1997. Stem: Sinozol-.

Sintor Schönherr, 1839: 148. Current status: valid genus in ANTHRIBIDAE: ANTHRIBINAE: SINTORINI.

Type species: Sintor quadrilineatus Fåhraeus, 1839, by original designation.

Family-group name: SINTORIDES Lacordaire, 1865. Stem: Sintor-.

Sipalia Mulsant & Rey, 1853b: 32. Current status: invalid senior synonym of Cyllopisalia Pace, 1982 in STAPHYLINIDAE: ALEOCHARINAE: HOMALOTINI: BOLITOCHARINA. Note: name suppressed for the purposes of the Principle of Priority and placed on the Official Index of Rejected and Invalid Generic Names in Zoology (ICZN 2005d).

Type species: Homalota difformis Mulsant & Rey, 1853, by subsequent designation (Fauvel 1902: 40). Note: name placed on the Official List of Specific Names in Zoology (ICZN 2005d).

Family-group name: SIPALIAE Casey, 1910. Stem: Sipali-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

Sipalinus G.A.K. Marshall, 1943: 119. Current status: valid genus in CURCULIONIDAE: DRYOPHTHORINAE: ORTHOGNATHINI. Note: new replacement name for Sipalus Schönherr, 1825.

Type species: Calandra granulata Fabricius, 1801 (= Curculio gigas Fabricius, 1775), by original designation (type fixation for Sipalus Schönherr).

Family-group name: SIPALININAE Zimmerman, 1993. Stem: Sipalin-.

Sipalomimus Voss, 1958: 125. Current status: valid genus in CURCULIONIDAE: DRYOPHTHORINAE: SIPALOMIMINI.

Type species: Sipalomimus singularis Voss, 1958, by original designation.

Family-group name: SIPALOMIMINI Legalov, 2020a: 325. Stem: Sipalomim-.

Sipalus Schönherr, 1825: 587. Current status: invalid senior synonym of Sipalinus G.A.K. Marshall, 1943 in CURCULIONIDAE: DRYOPHTHORINAE: ORTHOGNATHINI. Note: junior homonym of Sipalus Fischer, 1813 [Mammalia].

Type species: Calandra granulata Fabricius, 1801 (= Curculio gigas Fabricius, 1775), by original designation.

Family-group name: SIPALIDES Lacordaire, 1865. Stem: Sipal-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

Sisyphus Latreille, 1806: 79. Current status: valid genus in SCARABAEIDAE: SCARABAEINAE: SISYPHINI. Note: the original spelling was Sisyphe but Sisyphus, an incorrect subsequent spelling, is in prevailing usage, attributed to the publication of the original spelling, and therefore deemed to be the correct original spelling (ICZN 1999a, Article 33.3.1).

Type species: Scarabaeus schaefferi Linnaeus, 1758, by monotypy.

Family-group name: SISYPHAIRES Mulsant, 1842. Stem: Sisyph-.

Sitaris Latreille, 1802b: 187. Current status: valid genus in MELOIDAE: NEMOGNATHINAE: NEMOGNATHINI: SITARINA.

Type species: Necydalis humeralis Fabricius, 1775 (= Necydalis muralis Forster, 1771), by monotypy. Note: Fabricius’s specific name was spelled humuralis but the spelling humeralis, an incorrect subsequent spelling, is in prevailing usage, attributed to the publication of the original spelling, and therefore deemed to be the correct original spelling (ICZN 1999a, Article 33.3.1).

Family-group name: SITARATES Mulsant, 1857. Stem: Sitar-.

Sitona Germar, 1817b: 341. Current status: valid genus in CURCULIONIDAE: ENTIMINAE: POLYDRUSITAE: SITONINI.

Type species: Curculio lineatus Linnaeus, 1758, by subsequent designation (Schönherr 1823: 1141).

Family-group name: SITONISIDAE Gistel, 1848. Stem: Siton-.

Sitophilus Schönherr, 1838: 967. Current status: valid genus in CURCULIONIDAE: DRYOPHTHORINAE: LITOSOMINI: SITOPHILINA. Note: name placed on the Official List of Generic Names in Zoology (ICZN 1959b).

Type species: Curculio oryzae Linnaeus, 1763, by original designation (see ICZN 1959b). Note: although the species was originally spelled oryza, the emended spelling oryzae was validated by the Commission and the name placed on the Official List of Specific Names in Zoology (ICZN 1959b).

Family-group name: SCASITOPHILIPHIDILIA Csiki, 1936. Stem: Sitophil-.

Skatitoxenus Kistner & Pasteels, 1969: 1190. Current status: valid genus in STAPHYLINIDAE: ALEOCHARINAE: SKATITOXENINI.

Type species: Skatitoxenus coatoni Kistner & Pasteels, 1969, by original designation.

Family-group name: SKATITOXENINI Kistner & Pasteels, 1969. Stem: Skatitoxen-.

Slipinskia Bocák & Bocáková, 1992: 257. Current status: junior synonym of Flagrax Kasantsev, 1992 in LYCIDAE: EROTINAE: SLIPINSKIINI.

Type species: Stadenus aethiops Kleine, 1933, by original designation.

Family-group name: SLIPINSKIININA Bocák & Bocáková, 1992. Stem: Slipinski-.

Sloanoglymmius R.T. Bell & J.R. Bell, 1991: 185. Current status: valid genus in CARABIDAE: RHYSODINAE: SLOANOGLYMMIINI.

Type species: Rhysodes planatus Lea, 1904, by original designation.

Family-group name: SLOANOGLYMMIINA R.T. Bell & J.R. Bell, 1991. Stem: Sloanoglymmi-.

†Slonik Zherikhin, 1977b: 180. Current status: valid genus in NEMONYCHIDAE: †SLONIKINAE. Note: Lower Cretaceous (Kazakhstan).

Type species: †Slonik sibiricus Zherikhin, 1977, by original designation.

Family-group name: †SLONIKINAE Zherikhin, 1977. Stem: Slonik-.

Smicrips J.L. LeConte, 1878a: 399. Current status: valid genus in SMICRIPIDAE.

Type species: Smicrips palmicola J.L. LeConte, 1878 (= Tisiphone hypocoproides Reitter, 1876), by monotypy.

Family-group name: SMICRIPINI G.H. Horn, 1880. Stem: Smicrip-.

Smicronyx Schönherr, 1843: 313. Current status: valid genus in CURCULIONIDAE: CURCULIONINAE: SMICRONYCHINI. Note: new replacement name for Micronyx Schönherr, 1835.

Type species: Micronyx reichii Gyllenhal, 1835, by original designation (type fixation for Micronyx Schönherr).

Family-group name: SMICRONYCINA Seidlitz, 1891. Stem: Smicronych-. Note: nomen protectum (see Bouchard et al. 2011: 584).

Smodicum Haldeman, 1847: 38. Current status: valid genus in CERAMBYCIDAE: CERAMBYCINAE: SMODICINI.

Type species: Callidium cucujiforme Say, 1826, by monotypy.

Family-group name: SMODICIDES Lacordaire, 1868. Stem: Smodic-.

Sobarus Harold, 1879: 164. Current status: valid genus in CERAMBYCIDAE: PRIONINAE: ANACOLINI. Note: junior homonym of Sobarus Loew, 1855 [Diptera]; since no new replacement name has been proposed for Sobarus Harold, we here propose Basorus Bouchard & Bousquet, nomen novum.

Type species: Sobarus poggei Harold, 1879, by monotypy.

Family-group name: SOBARINI Lameere, 1913. Stem: Sobar-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

Sogda Lopatin, 1961: 121. Current status: valid genus in LEIODIDAE: LEIODINAE: SOGDINI.

Type species: Sogda pavlovskii Lopatin, 1961, by original designation.

Family-group name: SOGDIIDAE Lopatin, 1961. Stem: Sogd-.

Soleniscus Bonvouloir, 1871: 76 [in key]. Current status: invalid senior synonym of Cussolenis Fleutiaux, 1918 in ELATERIDAE: THYLACOSTERNINAE. Note: junior homonym of Soleniscus Meek & Worthen, 1860 [Mollusca]; this name was first used by Gemminger in Gemminger and von Harold (1869b: 1486) but is unavailable from that date as the species associated with the name is a nomen nudum; the plates of Bonvouloir’s work on EUCNEMIDAE referred to by Gemminger were not engraved until 1870 as listed at the top of the plates, and the part including Soleniscus was not published until 1871.

Type species: Soleniscus mutabilis Bonvouloir, 1875, by subsequent monotypy (de Bonvouloir 1875: 828).

Family-group name: SOLENISCINAE Lameere, 1900. Stem: Solenisc-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

Solenocyclus Kaup, 1868a: 10. Current status: valid genus in PASSALIDAE: PASSALINAE: SOLENOCYCLINI.

Type species: Passalus exaratus Klug, 1833, by monotypy.

Family-group name: SOLENOCYCLEAE Kaup, 1871. Stem: Solenocycl-.

Solenogenys Westwood, 1860: 170. Current status: valid genus in CARABIDAE: SCARITINAE: SALCEDIINI: SOLENOGENYINA. Note: this taxon is a junior synonym of Aulacinia J. Thomson, 1857 and an application to the Commission is necessary to preserve Solenogenys as a valid taxon.

Type species: Solenogenys foeda Westwood, 1860 (= Aulacinia rhysodoides J. Thompson, 1857), by monotypy.

Family-group name: SOLENOGENYINA R.T. Bell, 1998. Stem: Solenogeny-.

Solenoptera Audinet-Serville, 1832: 183. Current status: valid genus in CERAMBYCIDAE: PRIONINAE: SOLENOPTERINI.

Type species: Cerambyx thomae Linnaeus, 1767, by subsequent designation (Cazurro Ruiz 1896: 430).

Family-group name: SOLÉNOPTÉRIDES Lacordaire, 1868. Stem: Solenopter-.

Solierius Bernhauer, 1921: 68. Current status: valid genus in STAPHYLINIDAE: SOLIERIINAE. Note: new replacement name for Physognathus Solier 1849 (junior homonym of Physognathus Agassiz 1846, an emendation of Physignathus Cuvier, 1829).

Type species: Physognathus obscurus Solier, 1849, by monotypy (type fixation for Physognathus Solier).

Family-group name: SOLIERIINAE Newton & Thayer, 1992. Stem: Solieri-.

Somatipion L.W. Schaufuss, 1877b: 457. Current status: valid genus in STAPHYLINIDAE: PSELAPHINAE: PSELAPHITAE: TYRINI: SOMATIPIONINA.

Type species: Somatipion globulifer L.W. Schaufuss, 1877 (= Faronus punctatus King, 1866), by monotypy.

Family-group name: SOMATIPIONINI Jeannel, 1949. Stem: Somatipion-.

Somatodes Schönherr, 1823: 1139. Current status: junior synonym of Pachyrhynchus Germar, 1823 in CURCULIONIDAE: ENTIMINAE: POLYDRUSITAE: PACHYRHYNCHINI. Note: name suppressed for the Principles of Priority and Homonymy and placed on the Official Index of Rejected and Invalid Generic Names in Zoology (see ICZN 1994c).

Type species: Somatodes sanctus Schönherr, 1823, by original designation. Note: name suppressed for the Principle of Priority and placed on the Official Index of Rejected and Invalid Specific Names in Zoology (see ICZN 1994c).

Family-group name: SOMATODIDES Schönherr, 1823. Stem: Somatod-. Note: family-group name permanently invalid (ICZN 1999a, Article 39); name placed on the Official Index of Rejected and Invalid Family-group Names in Zoology (ICZN 1994c).

Somatodes Schönherr, 1840a: 800. Current status: valid genus in CURCULIONIDAE: CYCLOMINAE: CYCLOMINI. Note: the senior homonym Somatodes Schönherr, 1823, and all uses of Somatodes prior to the publication of Somatodes Schönherr, 1840, was suppressed for the purposes of both the Principle of Priority and Homonymy and placed on the Official Index of Rejected and Invalid Generic Names in Zoology while Somatodes Schönherr, 1840 was placed on the Official List of Generic Names in Zoology (ICZN 1994c).

Type species: Somatodes misumenus Gyllenhal, 1840, by original designation (see ICZN 1994c). Note: name placed on the Official List of Specific Names in Zoology (ICZN 1994c, as type species fixed by monotypy).

Family-group name: SOMATODIDES Lacordaire, 1863. Stem: Somatod-. Note: junior homonym of SOMATODINI Schönherr, 1823 (type genus Somatodes Schönherr, 1823) in Coleoptera: CURCULIONIDAE; Schönherr’s name is permanently invalid (based on a suppressed type genus).

Somotrichus Seidlitz, 1887: 7 [Gatt.]. Current status: valid genus in CARABIDAE: HARPALINAE: LEBIINI: PERICALINA.

Type species: Carabus elevatus Fabricius, 1792 [p. 162] (= Lebia unifasciata Dejean, 1831), by monotypy.

Family-group name: SOMOTRICHINI Mateu, 1963. Stem: Somotrich-.

Sonnetius Casey, 1922: 321. Current status: junior synonym of Barymerus Lacordaire, 1865 in CURCULIONIDAE: BARIDINAE: MADARINI: BARYMERINA.

Type species: Sonnetius binarius Casey, 1922, by present designation.

Family-group name: SONNETIINI Casey, 1922. Stem: Sonneti-.

Sophrorhinus Rouzet, 1855: 80. Current status: valid genus in CURCULIONIDAE: CRYPTORHYNCHINAE: SOPHRORHININI.

Type species: Sophrorhinus duvernoyi Rouzet, 1855, by monotypy.

Family-group name: SOPHRORHINIDES Lacordaire, 1865. Stem: Sophrorhin-.

Sosylopsis Grouvelle, 1910a: 269. Current status: valid genus in TEREDIDAE: TEREDINAE: SOSYLOPSINI.

Type species: Sosylopsis geayi Grouvelle, 1910, by monotypy.

Family-group name: SOSYLOPSINI Dajoz, 1980. Stem: Sosylops-.

Spalacopsis Newman, 1842a: 305. Current status: valid genus in CERAMBYCIDAE: LAMIINAE: AGAPANTHIINI.

Type species: Spalacopsis stellio Newman, 1842 (= Hippopsis filum Klug, 1829), by subsequent designation (J. Thomson 1864: 95).

Family-group name: SPALACOPSIDES Lacordaire, 1872. Stem: Spalacopse-.

Spanglerogyrus Folkerts, 1979: 2. Current status: valid genus in GYRINIDAE: SPANGLEROGYRINAE.

Type species: Spanglerogyrus albiventris Folkerts, 1979, by original designation.

Family-group name: SPANGLEROGYRINAE Folkerts, 1979. Stem: Spanglerogyr-.

Spaniophaenus Reitter, 1875b: 4, 8. Current status: valid genus in CRYPTOPHAGIDAE: CRYPTOPHAGINAE: CRYPTOPHAGINI.

Type species: Cryptophagus amplicollis Brisout de Barneville, 1866 (= Cryptophagus lapidarius Fairmaire, 1862), by subsequent designation (Otero and Diaz Pazos 1995: 183). Note: Otero and Diaz Pazos (1995: 183) designated Cryptophagus lapidarius as type species, which is not an originally included species; however, as they listed it in synonymy with Cryptophagus amplicollis, one of the two originally included species, they are deemed to have designated the latter species as type species (ICZN 1999a, Article 69.2.2); Cryptophagus laticollis L. Miller, 1858 (= Catopochrotides termitophilus Kieseritzky & Reikhardt, 1936) is often listed as type species of Spaniophaenus, by subsequent designation of Leschen (1996: 601) but Otero and Diaz Pazos’ designation is earlier; both species are currently included in the genus Spaniophaenus Reitter.

Family-group name: SPANIOPHAENI Casey, 1900. Stem: Spaniophaen-.

Sparedrus Dejean, 1821: 72. Current status: valid genus in OEDEMERIDAE: CALOPODINAE.

Type species: Calopus testaceus Andersch, 1797, by monotypy.

Family-group name: SPAREDRIIDAE Gistel, 1848. Stem: Sparedr-. Note: nomen oblitum (see Bouchard et al. 2011: 435).

Sparostes Putzeys, 1867: 27. Current status: valid genus in CARABIDAE: SCARITINAE: CLIVININI: SPAROSTESINA.

Type species: Sparostes brevicollis Putzeys, 1867, by subsequent designation (Andrewes 1929: 344).

Family-group name: SPAROSTESINA Dostal, 2017: 117. Stem: Sparostes-.

Sparrmannia Laporte, 1840b: 132. Current status: valid genus in SCARABAEIDAE: MELOLONTHINAE: TANYPROCTINI: TANYPROCTINA. Note: nomen protectum (see Bouchard et al. 2011: 255); the original spelling was Sparmannia but the unjustified emendation Sparrmannia, introduced by von Dalla Torre (1913: 291), is in prevailing usage, attributed to Laporte (1840b), and therefore is a justified emendation and the correct original spelling (ICZN 1999a, Article 33.2.3.1).

Type species: Melolontha alopex Fabricius, 1787, by subsequent designation (Evans 1989: 14).

Family-group name: SPARRMANNINI Péringuey, 1904. Stem: Sparrmanni-.

Spastica Guérin-Méneville, 1844a: 137. Current status: valid genus in MELOIDAE: ELETICINAE: SPASTICINI: SPASTICINA. Note: this name was first proposed as a synonym of Gnathium W. Kirby, 1819 and treated as an available name before 1961 (e.g., Lacordaire 1859b: 679), Spastica was therefore made available from its first publication as a synonym (ICZN 1999a, Article 11.6.1).

Type species: Gnathium flavicolle Guérin-Méneville, 1834, by monotypy.

Family-group name: SPASTICINA Kaszab, 1959. Stem: Spastic-.

Spelaeobates J. Müller, 1901: 16, 28. Current status: valid genus in LEIODIDAE: CHOLEVINAE: LEPTODIRINI: SPELAEOBATINA.

Type species: Spelaeobates novaki J. Müller, 1901, by subsequent designation (Jeannel 1910: 25).

Family-group name: SPELAEOBATINA Guéorguiev, 1974. Stem: Spelaeobat-.

Speleobama Park, 1951: 52. Current status: valid genus in STAPHYLINIDAE: PSELAPHINAE: GONIACERITAE: SPELEOBAMINI.

Type species: Speleobama vana Park, 1951, by original designation.

Family-group name: SPELEOBAMINI Park, 1951. Stem: Speleobam-.

Spercheus Kugelann in Illiger, 1798: 241. Current status: valid genus in SPERCHEIDAE.

Type species: Dytiscus emarginatus Schaller, 1783, by subsequent designation (Latreille 1810: 428).

Family-group name: SPERCHEINI Erichson, 1837. Stem: Sperche-.

Sperchopsis J.L. LeConte, 1861: 47. Current status: valid genus in HYDROPHILIDAE: HYDROPHILINAE: HYDROBIUSINI.

Type species: Spercheus tesselatus Ziegler, 1844, by original designation.

Family-group name: SPERCHOPSINI Hansen, 1991. Stem: Sperchops-.

Spermophagus Schönherr, 1833: 102. Current status: valid genus in CHRYSOMELIDAE: BRUCHINAE: AMBLYCERINI: SPERMOPHAGINA.

Type species: Spermophagus titivilitius Boheman, 1833, by original designation.

Family-group name: SPERMOPHAGINI Borowiec, 1987. Stem: Spermophag-.

Sphadasmus Schönherr, 1844: 290. Current status: valid genus in CURCULIONIDAE: CONODERINAE: SPHADASMINI. Note: new replacement name for Cyrtomon Schönherr, 1836.

Type species: Cyrtomon camelus Gyllenhal, 1836, by original designation (type fixation for Cyrtomon Schönherr).

Family-group name: SPHADASMIDES Lacordaire, 1865. Stem: Sphadasm-.

Sphaenelater O. Schwarz, 1902: 365. Current status: valid genus in ELATERIDAE: LISSOMINAE: PROTELATERINI.

Type species: Sphaenelater nigricornis O. Schwarz, 1902, by monotypy.

Family-group name: SPHAENELATERINI Stibick, 1979. Stem: Sphaenelater-.

Sphaenothecus Dupont, 1836: pls 141–164 [p. 2]. Current status: valid genus in CERAMBYCIDAE: CERAMBYCINAE: TRACHYDERINI: TRACHYDERINA.

Type species: Sphaenothecus tomentosus Dupont, 1838 (= Sphaenothecus trilineatus Dupont, 1838), by subsequent designation (J. Thomson 1864: 205). Note: no species were originally included in this genus by Dupont (1836); the first ones were the four species listed by Dupont (1838: 56–59).

Family-group name: SPHAENOTHECITAE J. Thomson, 1861. Stem: Sphaenothec-.

Sphaeratrix Gistel, 1848b: 190. Current status: valid genus in CHRYSOMELIDAE: CHRYSOMELINAE: CHRYSOMELINI.

Type species: Sphaeratrix latifrons Gistel, 1848, by monotypy.

Family-group name: SPHAERATRIXINI Daccordi, 1981. Stem: Sphaeratrix-.

Sphaericus Wollaston, 1854: 263. Current status: valid genus in PTINIDAE: PTININAE: SPHAERICINI.

Type species: Ptinus dawsoni Wollaston, 1854, by subsequent designation (Bellés 1991: 24).

Family-group name: SPHAERICINI Portevin, 1931. Stem: Sphaeric-.

Sphaeridium Fabricius, 1775: 66. Current status: valid genus in HYDROPHILIDAE: SPHAERIDIINAE: SPHAERIDIINI.

Type species: Dermestes scarabaeoides Linnaeus, 1758, by subsequent designation (Latreille 1810: 428).

Family-group name: SPHAERIDIOTA Latreille, 1802. Stem: Sphaeridi-. Note: senior homonym of SPHAERIDIINI Richards, 1968 (type genus Sphaeridia Linnaniemi, 1912) in Collembola.

Sphaerion Audinet-Serville, 1834b: 68. Current status: valid genus in CERAMBYCIDAE: CERAMBYCINAE: ELAPHIDIINI.

Type species: Elaphidion cyanipennis Audinet-Serville, 1834, by original designation.

Family-group name: SPHÉRIONIDES Lacordaire, 1868. Stem: Sphaeri-. Note: junior homonym of SPHAERIIDAE Deshayes, 1855 (type genus Sphaerium Scopoli, 1877) in Mollusca.

Sphaeriopoeus Kuschel, 2003a: 59. Current status: valid genus in CURCULIONIDAE: CURCULIONINAE: SPHAERIOPOEINI.

Type species: Sphaeriopoeus faber Kuschel, 2003, by original designation.

Family-group name: SPHAERIOPOEINI Kuschel, 2003. Stem: Sphaeriopoe-.

Sphaerites Duftschmid, 1805: 205. Current status: valid genus in SPHAERITIDAE.

Type species: Hister glabratus Fabricius, 1792, by monotypy.

Family-group name: SPHAERITIDAE Shuckard, 1839. Stem: Sphaerit-.

Sphaerius Waltl, 1838: 272. Current status: valid genus in SPHAERIUSIDAE. Note: name suppressed for the purposes of both the Principle of Priority and Homonymy and placed on the Official Index of Rejected and Invalid Generic Names in Zoology (ICZN 1985d) but subsequently placed on the Official List of Generic Names in Zoology (ICZN 2000).

Type species: Sphaerius acaroides Waltl, 1838, by monotypy (see ICZN 2000). Note: name placed on the Official List of Specific Names in Zoology (ICZN 2000).

Family-group name: SPHAERINA Erichson, 1845. Stem: Sphaerius- (ICZN 2000).

Sphaerocharis Lacordaire, 1848: 634. Current status: valid genus in CHRYSOMELIDAE: LAMPROSOMATINAE: SPHAEROCHARINI.

Type species: Lamprosoma marginicolle Guérin-Méneville, 1844, by subsequent designation (Achard 1914b: 21).

Family-group name: SPHOEROCHARIDES Chapuis, 1874. Stem: Sphaerochar-.

Sphaeronum Sharp, 1876b: 224. Current status: valid genus in STAPHYLINIDAE: PAEDERINAE: LATHROBIINI: SPHAERONINA.

Type species: Sphaeronum pallidum Sharp, 1876, by subsequent designation (Casey 1905: 55).

Family-group name: SPHAERONIA Casey, 1905. Stem: Sphaeron-.

Sphaeronychus Dejean, 1836a: 383. Current status: valid genus in CHRYSOMELIDAE: GALERUCINAE: ALTICINI. Note: the original spelling was Sphraeronychus but Sphaeronychus, an incorrect subsequent spelling, is prevailing usage, attributed to the publication of the original spelling, and therefore deemed to be the correct original spelling (ICZN 1999a, Article 33.3.1).

Type species: Altica melanura G.-A. Olivier, 1808, by monotypy.

Family-group name: SPHAERONYCHINI Bechyné & Špringlová de Bechyné, 1960. Stem: Sphaeronych-.

Sphaerosoma Stephens, 1832a: 391. Current status: valid genus in ALEXIIDAE. Note: this genus-group name is often credited to “Samouelle, 1819” in the literature; however, we use the corrected name and author proposed by Löbl and Smetana (2011: 31).

Type species: Sphaerosoma quercus Stephens, 1832 (= Tritoma pilosum Panzer, 1793), by monotypy. Note: we use the data published by Löbl and Smetana (2011: 31) regarding the author and year of the type species as well as its valid name.

Family-group name: SPHAEROSOMINAE Ganglbauer, 1899. Stem: Sphaerosomat-.

Sphaerotrypes W.F.H. Blandford, 1894: 61. Current status: valid genus in CURCULIONIDAE: SCOLYTINAE: DIAMERINI.

Type species: Sphaerotrypes pila W.F.H. Blandford, 1894, by subsequent designation (Hopkins 1914: 129).

Family-group name: SPHAEROTRYPINI Murayama, 1963. Stem: Sphaerotryp-.

Sphallotrichus Fragoso, 1982: 145. Current status: valid genus in CERAMBYCIDAE: CERAMBYCINAE: CERAMBYCINI: SPHALLOTRICHINA.

Type species: Cerambyx setosus Germar, 1823, by original designation.

Family-group name: SPHALLOTRICHINA Martins & Monné, 2005. Stem: Sphallotrich-.

Sphenocorynes Schönherr, 1838: 866. Current status: valid genus in CURCULIONIDAE: DRYOPHTHORINAE: SPHENOPHORINI: SPHENOCORYNINA.

Type species: Curculio quadripunctatus Weber, 1801 (= Rhynchophorus cinereus Illiger, 1800), by original designation.

Family-group name: SPHÉNOCORYNIDES Lacordaire, 1865. Stem: Sphenocoryn-.

Sphenophorus Schönherr, 1838: 874. Current status: valid genus in CURCULIONIDAE: DRYOPHTHORINAE: SPHENOPHORINI: SPHENOPHORINA. Note: name placed on the Official List of Generic Names in Zoology (ICZN 1959b).

Type species: Curculio abbreviatus Fabricius, 1787, by original designation (see ICZN 1959b). Note: name validated by the Commission (i.e., not a primary homonym of Curculio abbreviates Linnaeus, 1758) and placed on the Official List of Specific Names in Zoology (ICZN 1959b).

Family-group name: SPHÉNOPHORIDES Lacordaire, 1865. Stem: Sphenophor-.

Sphenoptera Dejean, 1833a: 81. Current status: valid genus in BUPRESTIDAE: CHRYSOCHROINAE: SPHENOPTERINI.

Type species: Buprestis antiqua Illiger, 1803, by subsequent designation (Volkovitsh and Kalashian 2002: 166).

Family-group name: SPHÉNOPTÉRIDES Lacordaire, 1857. Stem: Sphenopter-.

Sphinctovalgus Kolbe, 1904: 51. Current status: valid genus in SCARABAEIDAE: CETONIINAE: VALGINI: VALGINA.

Type species: Sphinctovalgus conradti Kolbe, 1904, by monotypy.

Family-group name: SPHINCTOVALGINAE Kolbe, 1904. Stem: Sphinctovalg-.

Sphindiphorus Sen Gupta & Crowson, 1979: 184. Current status: valid genus in SPHINDIDAE: SPHINDIPHORINAE.

Type species: Sphindiphorus natalensis Sen Gupta & Crowson, 1979, by original designation.

Family-group name: SPHINDIPHORINI Sen Gupta & Crowson, 1979. Stem: Sphindiphor-.

Sphindocis Fall, 1917: 170. Current status: valid genus in TETRATOMIDAE: SPHINDOCIINAE.

Type species: Sphindocis denticollis Fall, 1917, by monotypy.

Family-group name: SPHINDOCIINAE Lawrence, 1974. Stem: Sphindoci-.

Sphindus Dejean, 1821: 102. Current status: valid genus in SPHINDIDAE: SPHINDINAE. name placed (as Sphindus Megerle in Dejean, 1821) on the Official List of Generic Names in Zoology (ICZN 1997).

Type species: Nitidula dubia Gyllenhal, 1808, by monotypy (see ICZN 1997). Note: name placed on the Official List of Specific Names in Zoology (ICZN 1997).

Family-group name: SPHINDIDES Jacquelin du Val, 1860. Stem: Sphind-.

Sphingnotus Perroud, 1855b: 410. Current status: valid genus in CERAMBYCIDAE: LAMIINAE: TMESISTERNINI.

Type species: Tmesisternus mirabilis Boisduval, 1835, by subsequent designation (J. Thomson 1864: 31).

Family-group name: SPINGNOTHITAE J. Thomson, 1864. Stem: Sphingnot-.

Sphodrosomus Perroud & Montrouzier, 1865: 58. Current status: valid genus in CARABIDAE: HARPALINAE: PTEROSTICHINI: ABACOMORPHINA.

Type species: Sphodrosomus seisseti Perroud & Montrouzier, 1865, by monotypy.

Family-group name: SPHODROSOMINI Tschitschérine, 1902. Stem: Sphodrosom-.

Sphodrus Clairville, 1806: 84. Current status: valid genus in CARABIDAE: HARPALINAE: SPHODRINI: SPHODRINA.

Type species: Carabus planus Fabricius, 1792 (= Carabus leucophthalmus Linnaeus, 1758), by monotypy.

Family-group name: SPHODRIDAE Laporte, 1834. Stem: Sphodr-.

Sphoerodes Chaudoir, 1883: 519. Current status: valid genus in CARABIDAE: HARPALINAE: OODINI.

Type species: Oodes striatus Dejean, 1831, by monotypy.

Family-group name: SPHAERODINI Jeannel, 1949. Stem: Sphoerod-.

Sphondylia Weise, 1902: 120. Current status: valid genus in MEGALOPODIDAE: MEGALOPODINAE: SPHONDYLIINI.

Type species: Sphondylia magnicollis Weise, 1902, by subsequent designation (Jacoby and Clavareau 1905: 7).

Family-group name: SPHONDYLIINI Rodríguez-Mirón in Rodríguez-Mirón et al., 2021: 694. Stem: Sphondyli-.

Sphuridaethes Pace, 1988: 980. Current status: valid genus in STAPHYLINIDAE: ALEOCHARINAE: COROTOCINI: SPHURIDAETHINA.

Type species: Sphuridaethes loebli Pace, 1988, by original designation.

Family-group name: SPHURIDAETHINA Pace, 1988. Stem: Sphuridaeth-.

Spilophora Boheman, 1850: 107. Current status: valid genus in CHRYSOMELIDAE: CASSIDINAE: SPILOPHORINI.

Type species: Calyptocephala trigemina Guérin-Méneville, 1844, by subsequent designation (Monrós and Viana 1951: 389).

Family-group name: SPILOPHORITES Chapuis, 1875. Stem: Spilophor-. Note: senior homonym of SPILOPHORINA Krikken, 1984 (type genus Spilophorus Westwood, 1848) in Coleoptera: SCARABAEIDAE.

Spilophorus Schaum, 1848: 61. Current status: valid genus in SCARABAEIDAE: CETONIINAE: CREMASTOCHEILINI: SPILOPHORINA.

Type species: Cremastocheilus maculatus Gory & Percheron, 1833, by subsequent designation (Kraatz 1899: 63).

Family-group name: SPILOPHORINA Krikken, 1984. Stem: Spilophor-. Note: junior homonym of SPILOPHORINI Chapuis, 1875 (type genus Spilophora Boheman, 1850) in Coleoptera: CHRYSOMELIDAE.

Spilopyra Baly, 1860: 24. Current status: valid genus in CHRYSOMELIDAE: SPILOPYRINAE.

Type species: Spilopyra sumptuosa Baly, 1860, by original designation.

Family-group name: SPILOPYRITES Chapuis, 1874. Stem: Spilopyr-.

Spintheria J. Thomson, 1861: 357. Current status: valid genus in CERAMBYCIDAE: CERAMBYCINAE: SPINTHERIINI.

Type species: Tmesisternus gratiosus Pascoe, 1856, by monotypy.

Family-group name: SPINTHÉRIIDES Lacordaire, 1869. Stem: Spintheri-.

Spodochlamys Burmeister, 1855: 528. Current status: valid genus in SCARABAEIDAE: RUTELINAE: ANATISTINI.

Type species: Spodochlamys caesarea Burmeister, 1855, by monotypy.

Family-group name: SPODOCHLAMYDINI Ohaus, 1918. Stem: Spodochlamyd-.

Spondylis Fabricius, 1775: 159. Current status: valid genus in CERAMBYCIDAE: SPONDYLIDINAE: SPONDYLIDINI.

Type species: Attelabus buprestoides Linnaeus, 1758, by subsequent designation (Latreille 1810: 431).

Family-group name: SPONDYLII Audinet-Serville, 1832. Stem: Spondylid-.

Spongocerus W.F.H. Blandford, 1893: 431. Current status: junior synonym of Scolytoplatypus C. Schaufuss, 1891 in CURCULIONIDAE: SCOLYTINAE: SCOLYTOPLATYPODINI.

Type species: Scolytoplatypus tycon W.F.H. Blandford, 1893, by subsequent designation (Hopkins 1914: 129).

Family-group name: SPONGOCERINAE Hagedorn, 1909. Stem: Spongocer-.

Spurius Kaup, 1871: 75. Current status: valid genus in PASSALIDAE: PASSALINAE: PROCULINI.

Type species: Passalus bicornis Truqui, 1857, by monotypy.

Family-group name: SPURIINAE Kuwert, 1896. Stem: Spuri-.

Stagobius Schiødte, 1848: 78. Current status: junior synonym of Leptodirus F.J. Schmidt, 1832 in LEIODIDAE: CHOLEVINAE: LEPTODIRINI: LEPTODIRINA.

Type species: Stagobius troglodytes Schiødte, 1848 (= Leptodirus hochenwartii F.J. Schmidt, 1832), by monotypy.

Family-group name: STAGOBIINAE Schiødte, 1849. Stem: Stagobi-.

Staminodeus N.M. Franz, 2002: 413. Current status: valid genus in CURCULIONIDAE: CURCULIONINAE: DERELOMINI: STAMINODEINA.

Type species: Staminodeus vectoris N.M. Franz, 2002, by original designation.

Family-group name: STAMINODEINA N.M. Franz, 2006. Stem: Staminode-.

Stanus Ślipiński, Tomaszewska & Lawrence, 2009: 423. Current status: valid genus in CORYLOPHIDAE: CORYLOPHINAE: STANINI.

Type species: Stanus bowesteadi Ślipiński, Tomaszewska & Lawrence, 2009, by original designation.

Family-group name: STANINI Robertson, Ślipiński & McHugh in Robertson et al., 2012: 229. Stem: Stan-.

Staphylinus Linnaeus, 1758: 342, 421. Current status: valid genus in STAPHYLINIDAE: STAPHYLININAE: STAPHYLININI: STAPHYLININA. Note: name placed on the Official List of Generic Names in Zoology (ICZN 1959a).

Type species: Staphylinus erythropterus Linnaeus, 1758, by plenary power (see ICZN 1959a). Note: the species name was originally spelled erytropterus but it was modified to erythropterus by the Commission and placed on the Official List of Specific Names in Zoology (ICZN 1959a).

Family-group name: STAPHYLINIAE Latreille, 1802. Stem: Staphylin-.

Statira Lepeletier & Audinet-Serville, 1828: 479. Current status: valid genus in TENEBRIONIDAE: LAGRIINAE: LAGRIINI: STATIRINA.

Type species: Statira agroides Lepeletier & Audinet-Serville, 1828, by subsequent designation (Blanchard in Audouin et al. 1844: pl. 53bis).

Family-group name: STATYRITES Blanchard, 1845. Stem: Statir-.

Steatoderus Dejean, 1833a: 94. Current status: junior synonym of Elater Linnaeus, 1758 in ELATERIDAE: ELATERINAE: ELATERINI.

Type species: Elater ferrugineus Linnaeus, 1758, by monotypy.

Family-group name: STEATODERIDAE Gistel, 1848. Stem: Steatoder-.

Stenaesthetus Sharp, 1874a: 79. Current status: valid genus in STAPHYLINIDAE: EUAESTHETINAE: STENAESTHETINI.

Type species: Stenaesthetus sunioides Sharp, 1874, by monotypy.

Family-group name: STENAESTHETINI Bernhauer & Schubert, 1911. Stem: Stenaesthet-.

Stenalia Mulsant, 1856c: 83. Current status: valid genus in MORDELLIDAE: MORDELLINAE: STENALIINI.

Type species: Mordella testacea Fabricius, 1787, by subsequent designation (J.L. LeConte 1859b: 18).

Family-group name: STENALIINI Franciscolo, 1955. Stem: Stenali-.

Stenapion Wagner, 1912b: 20. Current status: valid genus in BRENTIDAE: APIONINAE: APIONITAE: APIONINI: STENAPIINA.

Type species: Apion constricticolle Sharp, 1890, by subsequent designation (Kissinger 1968: 29).

Family-group name: STENAPIINA Poinar & Legalov, 2015: 154. Stem: Stenapi-.

Stenaspidius Westwood, 1848b: 366. Current status: valid genus in BOLBOCERATIDAE: BOLBOCERATINAE: STENASPIDIINI.

Type species: Bolboceras nigricornis Westwood, 1848, by monotypy.

Family-group name: STENASPIDIINI Nikolajev, 2003. Stem: Stenaspidi-.

Stenaspis Audinet-Serville, 1834b: 51. Current status: valid genus in CERAMBYCIDAE: CERAMBYCINAE: TRACHYDERINI: TRACHYDERINA.

Type species: Stenaspis verticalis Audinet-Serville, 1834, by monotypy.

Family-group name: STÉNASPIDES Lacordaire, 1868. Stem: Stenaspid-.

Stenelmis Dufour, 1835: 158. Current status: valid genus in ELMIDAE: ELMINAE: ELMINI: STENELMINA.

Type species: Limnius canaliculatus Gyllenhal, 1808, by subsequent designation (C.G. Thomson 1859: 21).

Family-group name: STENELMISATES Mulsant & Rey, 1872. Stem: Stenelm-.

Stenhomalus A. White, 1855: 243. Current status: valid genus in CERAMBYCIDAE: CERAMBYCINAE: STENHOMALINI.

Type species: Stenhomalus fenestratus A. White, 1855, by monotypy.

Family-group name: STENHOMALINI Miroshnikov, 1989. Stem: Stenhomal-.

Stenichnus C.G. Thomson, 1859: 61. Current status: valid genus in STAPHYLINIDAE: SCYDMAENINAE: SCYDMAENITAE: STENICHNINI.

Type species: Scydmaenus exilis Erichson, 1837 (= Scydmaenus bicolor Denny, 1825), by original designation.

Family-group name: STENICHNINI Fauvel, 1885. Stem: Stenichn-.

Stenobia Lacordaire, 1872: 442. Current status: valid genus in CERAMBYCIDAE: LAMIINAE: STENOBIINI.

Type species: Stenobia pradieri Lacordaire, 1872, by monotypy.

Family-group name: STENOBIINI Breuning, 1950. Stem: Stenobi-.

Stenocerus Schönherr, 1826: 39. Current status: valid genus in ANTHRIBIDAE: ANTHRIBINAE: STENOCERINI.

Type species: Anthribus fulvitarsis Germar, 1823, by original designation.

Family-group name: STENOCERINARUM Kolbe, 1895. Stem: Stenocer-.

Stenochia W. Kirby, 1819: 423. Current status: junior synonym of Strongylium W. Kirby, 1819 in TENEBRIONIDAE: STENOCHIINAE: STENOCHIINI.

Type species: Stenochia rufipes W. Kirby, 1819, by subsequent designation (Hope 1841a: 133).

Family-group name: STENOCHIADAE W. Kirby, 1837. Stem: Stenochi-.

Stenocorus Geoffroy, 1762: 221. Current status: valid genus in CERAMBYCIDAE: LEPTURINAE: STENOCORINI. Note: name placed on the Official List of Generic Names in Zoology (ICZN 1994a).

Type species: Leptura meridiana Linnaeus, 1758, by plenary power (see ICZN 1994a). Note: name placed on the Official List of Specific Names in Zoology (ICZN 1994a).

Family-group name: STENOCORITAE J. Thomson, 1861. Stem: Stenocor-.

Stenocorynus Schönherr, 1823: 1142. Current status: valid genus in CURCULIONIDAE: ENTIMINAE: ENTIMITAE: BYRSOPAGINI: STRANGALIODINA.

Type species: Curculio crenulatus Fabricius, 1775, by original designation.

Family-group name: STENOCORYNINI McKeown, 1939. Stem: Stenocoryn-.

Stenocyphon Lawrence, 2001: 357. Current status: valid genus in SCIRTIDAE: STENOCYPHONINAE.

Type species: Stenocyphon sasajii Lawrence, 2001, by original designation.

Family-group name: STENOCYPHONINAE Lawrence & Yoshitomi, 2007. Stem: Stenocyphon-.

Stenodera Eschscholtz, 1818: 469. Current status: valid genus in MELOIDAE: NEMOGNATHINAE: STENODERINI. Note: Stenodera sexpunctata Eschscholtz, 1818 was placed on the Official List of Generic Names in Zoology (ICZN 2020a).

Type species: Stenodera sexpunctata Eschscholtz, 1818 (= Meloe caucasica Pallas, 1782), by monotypy (see ICZN 2020a). Note: name placed on the Official List of Specific Names in Zoology (ICZN 2020a).

Family-group name: STENODERINI Selander, 1991. Stem: Stenoder-.

Stenoderus Dejean, 1821: 112. Current status: valid genus in CERAMBYCIDAE: CERAMBYCINAE: RHAGIOMORPHINI. Note: name placed on the Official List of Generic Names in Zoology (ICZN 2020a).

Type species: Cerambyx abbreviatus Fabricius, 1801 (= Stenocorus suturalis G.-A. Olivier, 1800), by monotypy (see ICZN 2020a). Note: Cerambyx abbreviatus Fabricius, 1801 placed on the Official List of Specific Names in Zoology (ICZN 2020a).

Family-group name: STENODERINAE Pascoe, 1867. Stem: Stenoderus- (see ICZN 2020a).

Stenodontes Audinet-Serville, 1832: 173. Current status: valid genus in CERAMBYCIDAE: PRIONINAE: MACROTOMINI: MALLODONINA.

Type species: Prionus exsertus G.-A. Olivier, 1795, by subsequent designation (E. Desmarest 1860: 307).

Family-group name: STENODONTINI Lameere, 1903. Stem: Stenodont-. Note: homonym of STENODONTINA Schmiedeknecht, 1903 (type genus Stenodontus Berthoumieu, 1897) in Hymenoptera.

Stenolamus Gebien, 1920: 107. Current status: valid genus in TENEBRIONIDAE: BLAPTINAE: OPATRINI: STENOLAMINA.

Type species: Stenolamus sulciceps Gebien, 1920, by subsequent designation (Gebien 1938b: 397).

Family-group name: STENOLAMINA C. Koch, 1956. Stem: Stenolam-.

Stenolophus Dejean, 1821: 15. Current status: valid genus in CARABIDAE: HARPALINAE: HARPALINI: STENOLOPHINA.

Type species: here fixed (under ICZN 1999a, Article 70.3.2) as Carabus teutonus Schrank, 1781, misidentified as Carabus vaporariorum Linnaeus, 1758 (sensu Fabricius, 1787) in the subsequent designation by Westwood (1838c: 5).

Family-group name: STENOLOPHIDAE W. Kirby, 1837. Stem: Stenoloph-.

Stenomela Erichson, 1847c: 159. Current status: valid genus in CHRYSOMELIDAE: SPILOPYRINAE.

Type species: Stenomela pallida Erichson, 1847, by monotypy.

Family-group name: STENOMÈLITES Chapuis, 1874. Stem: Stenomel-.

Stenomorphus Dejean, 1831: 696. Current status: valid genus in CARABIDAE: HARPALINAE: HARPALINI: HARPALINA.

Type species: Stenomorphus angustatus Dejean, 1831, by monotypy.

Family-group name: STENOMORPHIDAE Laporte, 1834. Stem: Stenomorph-.

Stenopelmus Schönherr, 1835: 468. Current status: valid genus in CURCULIONIDAE: BRACHYCERINAE: TANYSPHYRINI.

Type species: Stenopelmus rufinasus Gyllenhal, 1835, by original designation.

Family-group name: STENOPELMI J.L. LeConte, 1876. Stem: Stenopelm-.

Stenophanes Solsky, 1876: 294. Current status: valid genus in TENEBRIONIDAE: STENOCHIINAE: CNODALONINI.

Type species: Hedyphanes mesostenus Solsky, 1871, by original designation.

Family-group name: STENOPHANINI Reitter, 1922. Stem: Stenophan-.

Stenopodius G.H. Horn, 1884: 301. Current status: valid genus in CHRYSOMELIDAE: CASSIDINAE: CHALEPINI.

Type species: Stenopodius flavidus G.H. Horn, 1884, by monotypy.

Family-group name: STENOPODIIDES G.H. Horn, 1884. Stem: Stenopodi-. Note: nomen oblitum (see Bouchard and Bousquet 2020a: 108).

Stenopterus Illiger, 1804: 120. Current status: valid genus in CERAMBYCIDAE: CERAMBYCINAE: STENOPTERINI.

Type species: Necydalis rufa Linnaeus, 1767, by monotypy.

Family-group name: STENOPTERIDAE Gistel, 1848. Stem: Stenopter-.

Stenorhis Jordan, 1928: 167. Current status: valid genus in ANTHRIBIDAE: CHORAGINAE: STENORHININI.

Type species: Stenorhis ampedus Jordan, 1928, by original designation.

Family-group name: STENORHININI B.D. Valentine in Trýzna and B.D. Valentine, 2011: 65. Stem: Stenorhin-.

Stenoria Mulsant, 1857a: 186. Current status: valid genus in MELOIDAE: NEMOGNATHINAE: NEMOGNATHINI: SITARINA.

Type species: Sitaris apicalis Latreille, 1804, by monotypy.

Family-group name: STENORIIDES J.B. Parker & Böving, 1924. Stem: Stenori-.

Stenoscelis Wollaston, 1861b: 141. Current status: valid genus in CURCULIONIDAE: COSSONINAE: RHYNCOLINI: STENOSCELIDINA.

Type species: Stenoscelis hylastoides Wollaston, 1861, by monotypy.

Family-group name: STENOSCELIDES Wollaston, 1877. Stem: Stenoscelid-.

Stenosis Herbst, 1799: 160. Current status: valid genus in TENEBRIONIDAE: PIMELIINAE: STENOSINI: STENOSINA.

Type species: fixed by Bouchard et al. (2021: 351) according to Article 70.3.2 (ICZN 1999a) as Tagenia intermedia Solier, 1838, the taxonomic species involved in the misidentification of Pimelia angustata Fabricius, 1775 (sensu Herbst, 1799), as designated subsequently by Lacordaire (1859a: 102).

Family-group name: STENOSIDAE Schaum, 1859. Stem: Stenos-.

Stenosphenus Haldeman, 1847: 39. Current status: valid genus in CERAMBYCIDAE: CERAMBYCINAE: ELAPHIDIINI.

Type species: Callidium notatum G.-A. Olivier, 1800, by monotypy.

Family-group name: STENOSPHENINI J.L. LeConte, 1873. Stem: Stenosphen-.

Stenostoma Latreille, 1810: 217. Current status: valid genus in OEDEMERIDAE: OEDEMERINAE: STENOSTOMATINI.

Type species: Leptura rostrata Fabricius, 1787, by monotypy.

Family-group name: STÉNOSTOMATES Mulsant, 1858. Stem: Stenostomat-.

Stenotarsia Burmeister, 1842: 590. Current status: valid genus in SCARABAEIDAE: CETONIINAE: STENOTARSIINI: STENOTARSIINA.

Type species: Cetonia velutina Gory & Percheron, 1835, by subsequent designation (Sakai and Nagai 1998: 179). Note: this is a junior primary homonym of Cetonia velutina G.-A. Olivier, 1789 [South Africa], which was also treated as a member of Cetonia by Gory and Percheron (1833: 61); Burmeister (1842: 368) placed Cetonia velutina G.-A. Olivier, 1789 in his new genus Gametis, and it was subsequently treated by Boheman (1857: 19) as a synonym of Gametis subfasciatus (Swederus, 1787) [now Gametoides subfasciata (Swederus, 1787)]; the primary homonym Cetonia velutina Gory & Percheron, 1835 [Madagascar] should be considered permanently invalid unless conditions outlined in Article 57.2 (ICZN 1999a) are met.

Family-group name: STENOTARSIDEN Kraatz, 1880. Stem: Stenotarsi-.

Stenotarsus Perty, 1832: 112. Current status: valid genus in ENDOMYCHIDAE: ENDOMYCHINAE.

Type species: Stenotarsus brevicollis Perty, 1832, by monotypy.

Family-group name: STÉNOTARSITES Chapuis, 1876. Stem: Stenotars-.

Stenotrachelus Berthold, 1827: 371. Current status: valid genus in STENOTRACHELIDAE: STENOTRACHELINAE.

Type species: Dryops aeneus Paykull, 1799, by subsequent monotypy (Lepeletier and Audinet- Serville 1828: 488). Note: although Stenotrachelus aeneus (Paykull, 1799) is used as a valid name in recent literature (e.g., Löbl 2020: 499), Paykull (1799: 152) clearly refers to Dryops aenea (Fabricius 1792b: 75), who in turn refers to his Lagria aenea (Fabricius 1787: 94); more research is needed to confirm if Dryops aeneus Paykull, 1799 should indeed be used as the correct type species for this genus-group name or if a misidentification was involved.

Family-group name: STENOTRACHELIDAE C.G. Thomson, 1859. Stem: Stenotrachel-.

Stenus Latreille, 1797: 77. Current status: valid genus in STAPHYLINIDAE: STENINAE.

Type species: Staphylinus juno Paykull, 1789, by subsequent monotypy (Paykull 1800: 433).

Family-group name: STENIDAE W.S. MacLeay, 1825. Stem: Sten-.

Stephanocephalus Kaup, 1868b: 19. Current status: valid genus in PASSALIDAE: PASSALINAE: SOLENOCYCLINI.

Type species: Passalus hostilis Percheron, 1841, by monotypy.

Family-group name: STEPHANOCEPHALEAE Kaup, 1871. Stem: Stephanocephal-.

Stephanorhynchus A. White, 1846: 17. Current status: valid genus in CURCULIONIDAE: CURCULIONINAE: EUGNOMINI: EUGNOMINA.

Type species: Stephanorhynchus curvipes A. White, 1846, by monotypy.

Family-group name: STEPHANORHYNCHINA Voss, 1936. Stem: Stephanorhynch-.

Stephanorrhina Burmeister, 1842: 208. Current status: valid genus in SCARABAEIDAE: CETONIINAE: GOLIATHINI: RHOMBORHININA.

Type species: Cetonia guttata G.-A. Olivier, 1789, by monotypy.

Family-group name: STEPHANORRHININA Schenkling, 1921. Stem: Stephanorrhin-.

Stereocorynes Wollaston, 1873b: 500, 588. Current status: valid genus in CURCULIONIDAE: COSSONINAE: RHYNCOLINI: STENOSCELIDINA.

Type species: Cossonus truncorum Germar, 1823, by monotypy.

Family-group name: STEREOCORYNINA Voss, 1955. Stem: Stereocoryn-.

Stereodermus Lacordaire, 1865: 419. Current status: valid genus in BRENTIDAE: BRENTINAE: TRACHELIZINI: STEREODERMINA.

Type species: Arrenodes pygmaeus Schönherr, 1833, by original designation.

Family-group name: STEREODERMINA Sharp, 1895. Stem: Stereoderm-.

Stereolus Rafinesque, 1815: 112. Current status: junior synonym of Trixagus Kugelann, 1794 in THROSCIDAE. Note: unnecessary new replacement name for Throscus Latreille, 1797.

Type species: Elater dermestoides Linnaeus, 1767, by subsequent designation (Curtis 1827: pl. 163) (type fixation for Throscus Latreille).

Family-group name: STEREOLIA Rafinesque, 1815. Stem: Stereol-. Note: nomen oblitum (see Bouchard et al. 2011: 306).

Stereomera Arrow, 1905: 534. Current status: valid genus in SCARABAEIDAE: APHODIINAE: STEREOMERINI.

Type species: Stereomera pusilla Arrow, 1905, by monotypy.

Family-group name: STEREOMERINI Howden & Storey, 1992. Stem: Stereomer-.

Sternacanthus Audinet-Serville, 1832: 172. Current status: valid genus in CERAMBYCIDAE: CERAMBYCINAE: TRACHYDERINI: TRACHYDERINA.

Type species: Prionus undatus G.-A. Olivier, 1795, by monotypy.

Family-group name: STERNACANTHITAE J. Thomson, 1864. Stem: Sternacanth-.

Sternechosomus Voss, 1958: 46. Current status: valid genus in CURCULIONIDAE: MOLYTINAE: METATYGINI.

Type species: Sternechosomus octotuberculatus Voss, 1958, by original designation.

Family-group name: STERNECHOSOMINI Voss, 1958. Stem: Sternechosom-.

Sternechus Schönherr, 1826: 251. Current status: valid genus in CURCULIONIDAE: MOLYTINAE: STERNECHINI.

Type species: Orobitis trachyptomus Germar, 1823, by original designation.

Family-group name: STERNÉCHIDES Lacordaire, 1863. Stem: Sternech-.

Sternocera Eschscholtz, 1829b: 8. Current status: valid genus in BUPRESTIDAE: JULODINAE. Note: this name is a nomen protectum and its senior synonym Sternoxus Billberg, 1820 is a nomen oblitum (Bílý 2001: 148).

Type species: Buprestis chrysis Fabricius, 1775, by subsequent designation (E. Desmarest 1860: 39).

Family-group name: STERNOCERINI Csiki, 1904. Stem: Sternocer-.

Sternodea Reitter, 1875b: 78. Current status: valid genus in CRYPTOPHAGIDAE: CRYPTOPHAGINAE: CAENOSCELINI.

Type species: Sternodea baudii Reitter, 1875, by monotypy.

Family-group name: STERNODEINI Casey, 1900. Stem: Sternode-.

Sternopriscus Sharp, 1880: cxlviii. Current status: valid genus in DYTISCIDAE: HYDROPORINAE: HYDROPORINI: STERNOPRISCINA.

Type species: Hydroporus multimaculatus Clark, 1862, by monotypy.

Family-group name: STERNOPRISCINI Branden, 1884. Stem: Sternoprisc-.

Sternotomis Percheron in Guérin-Méneville and Percheron, 1836: pl. 16. Current status: valid genus in CERAMBYCIDAE: LAMIINAE: STERNOTOMINI.

Type species: Sternotomis aper Percheron, 1836 (= Lamia ducalis Klug, 1835), by subsequent designation (J. Thomson 1868: 176). Note: J. Thomson (1868: 176) selected Lamia ducalis Klug, 1835 as type species, which was not an originally included nominal species, but since he listed it in synonymy with Sternotomis aper Percheron, 1836, an originally included nominal species, he is deemed to have designated the latter species as type species (ICZN 1999a, Article 69.2.2).

Family-group name: STERNOTOMITAE J. Thomson, 1860. Stem: Sternotom-.

Steropes Steven, 1806: 166. Current status: valid genus in ANTHICIDAE: STEROPINAE.

Type species: Steropes caspius Stevens, 1806, by monotypy.

Family-group name: STÉROPITES Jacquelin du Val, 1863. Stem: Sterop-. Note: junior homonym of STEROPINAE Dana, 1854 (type genus Sterope Goodsir, 1845) in Crustacea.

Stethaspis Hope, 1837: 71, 104. Current status: valid genus in SCARABAEIDAE: MELOLONTHINAE: COLYMBOMORPHINI. Note: name placed on the Official List of Generic Names in Zoology (ICZN 1983b).

Type species: Melolontha suturalis Fabricius, 1775, by plenary power (see ICZN 1983b). Note: name placed on the Official List of Specific Names in Zoology (ICZN 1983b).

Family-group name: STETHASPIDIDAE Burmeister, 1855. Stem: Stethaspid-.

Stethodesma Hope in Bainbridge, 1841: 482. Current status: valid genus in SCARABAEIDAE: CETONIINAE: GYMNETINI: GYMNETINA.

Type species: Stethodesma strachani Hope, 1841, by monotypy.

Family-group name: STETHODESMAE Schoch, 1894. Stem: Stethodesmat-.

Stethorus Weise, 1885a: 65. Current status: valid genus in COCCINELLIDAE: COCCINELLINAE: COCCIDULINI.

Type species: Coccinella minima Rossi, 1794 (= Scymnus pusillus Herbst, 1797), by subsequent designation (Korschefsky 1931: 111). Note: Korschefsky (1931: 111) selected Stethorus punctillum Weise, 1891 as type species, not an originally included species, but since he listed it in synonymy with Coccinella minima Rossi, 1794, an originally included species, he is deemed to have designated the latter species as type species (ICZN 1999a, Article 69.2.2).

Family-group name: STETHORINI Dobzhansky, 1924. Stem: Stethor-.

Sthereus Motschulsky, 1845c: 373. Current status: valid genus in CURCULIONIDAE: MOLYTINAE: STHEREINI.

Type species: Sthereus quadrituberculatus Motschulsky, 1845, by subsequent designation (Buchanan 1936: 178).

Family-group name: STHEREINI Hatch, 1971. Stem: Sthere-.

Sticholotis Crotch, 1874a: 200. Current status: valid genus in COCCINELLIDAE: COCCINELLINAE: STICHOLOTIDINI.

Type species: Sticholotis substriatus Crotch, 1874, by original designation.

Family-group name: STICHOLOTINI Weise, 1901. Stem: Sticholotid-.

Stictocema Jacoby, 1906: 33. Current status: valid genus in CHRYSOMELIDAE: GALERUCINAE: METACYCLINI.

Type species: Stictocema fasciata Jacoby, 1906, by subsequent designation (Wilcox 1971: 169).

Family-group name: STICTOCEMITES Laboissière, 1922. Stem: Stictocem-.

Stictocranius J.L. LeConte, 1867: 374. Current status: valid genus in STAPHYLINIDAE: EUAESTHETINAE: STICTOCRANIINI.

Type species: Stictocranius puncticeps J.L. LeConte, 1867, by monotypy.

Family-group name: STICTOCRANIINI Jakobson, 1914. Stem: Stictocrani-.

Stigmatrachelus Schönherr, 1840b: 123. Current status: valid genus in CURCULIONIDAE: ENTIMINAE: POLYDRUSITAE: DERMATODINI.

Type species: Curculio cinctus G.-A. Olivier, 1807, by original designation.

Family-group name: STIGMATRACHELINI Richard, 1983. Stem: Stigmatrachel-.

Stigmodera Eschscholtz, 1829b: 9. Current status: valid genus in BUPRESTIDAE: BUPRESTINAE: STIGMODERINI.

Type species: Buprestis macularia Donovan, 1805, by subsequent designation (Solier 1833: 293).

Family-group name: STIGMODÉRIDES Lacordaire, 1857. Stem: Stigmoder-.

Stilbus Seidlitz, 1872: 35 [Gatt.]. Current status: valid genus in PHALACRIDAE: PHALACRINAE. Note: new replacement name for Olistherus Seidlitz, 1872.

Type species: Anisotoma testacea Panzer, 1797, by subsequent designation (Guillebeau 1894: 283) (type fixation for Eustilbus Sharp, 1888, an unnecessary new replacement name for Stilbus Seidlitz).

Family-group name: STILBINI Jakobson, 1915. Stem: Stilb-.

Stilicopsis Sachse, 1852: 144. Current status: valid genus in STAPHYLINIDAE: PAEDERINAE: LATHROBIINI: STILICOPSINA.

Type species: Stilicopsis paradoxa Sachse, 1852, by monotypy.

Family-group name: STILICOPSES Casey, 1905. Stem: Stilicops-.

Stilicus Berthold, 1827: 331. Current status: junior synonym of Rugilus Leach, 1819 in STAPHYLINIDAE: PAEDERINAE: LATHROBIINI: STILICINA.

Type species: Staphylinus orbiculatus Paykull, 1789, by subsequent designation (Blanchard in Audouin et al. 1842: pl. 27).

Family-group name: STILICI Casey, 1905. Stem: Stilic-.

Stilipalpus Jeannel, 1951b: 73. Current status: valid genus in STAPHYLINIDAE: PSELAPHINAE: BATRISITAE: BATRISINI: STILIPALPINA.

Type species: Stilipalpus clavipes Jeannel, 1951, by original designation.

Family-group name: STILIPALPINA Jeannel, 1954. Stem: Stilipalp-.

Stilpnonotus G.R. Gray in Griffith and Pidgeon, 1831: pls 48, 61. Current status: valid genus in MYCTERIDAE: EURYPINAE.

Type species: Stilpnonotus eurypiformis G.R. Gray, 1831, by monotypy.

Family-group name: STILPNONOTINAE Borchmann, 1936. Stem: Stilpnonot-.

Stizopus Erichson, 1843b: 245. Current status: valid genus in TENEBRIONIDAE: BLAPTINAE: OPATRINI: STIZOPODINA.

Type species: Stizopus laticollis Erichson, 1843, by monotypy.

Family-group name: STIZOPIDES Lacordaire, 1859. Stem: Stizopod-.

Stolas Billberg, 1820: 58. Current status: valid genus in CHRYSOMELIDAE: CASSIDINAE: MESOMPHALIINI.

Type species: Cassida discors Fabricius, 1801 (= Cassida discoides Linnaeus, 1758), by subsequent designation (Barber and Bridwell 1940: 9).

Family-group name: STOLAINI Hincks, 1952. Stem: Stolad-.

Stolius Lewis, 1895a: 433. Current status: valid genus in STENOTRACHELIDAE: STOLIINAE.

Type species: Stolius vagepictus Lewis, 1895, by monotypy.

Family-group name: STOLIINAE Nikitsky, 1985. Stem: Stoli-.

Stomis Clairville, 1806: 46. Current status: valid genus in CARABIDAE: HARPALINAE: PTEROSTICHINI: PTEROSTICHINA.

Type species: Carabus pumicatus Panzer, 1795, by monotypy.

Family-group name: STOMIDES Gené, 1839. Stem: Stomid-.

Storeus Schönherr, 1843: 293. Current status: valid genus in CURCULIONIDAE: CURCULIONINAE: STOREINI.

Type species: Storeus variegatus Boheman, 1843, by original designation.

Family-group name: STORÉIDES Lacordaire, 1863. Stem: Store-.

Storthodontus Chaudoir, 1855: 23. Current status: valid genus in CARABIDAE: SCARITINAE: SCARITINI: STORTHODONTINA. Note: Chaudoir originally used two different spellings, including Horthodontus (p. 4); Chaudoir (1879: 128, 176) used Storthodontus and therefore acted as First Reviser (ICZN 1999a, Article 24.2.4).

Type species: Storthodontus nimrod Chaudoir, 1855, by subsequent designation (Motschulsky 1857b: 94).

Family-group name: STORTHODONTINI Jeannel, 1946. Stem: Storthodont-.

Strangaliodes Schönherr, 1842a: 219. Current status: valid genus in CURCULIONIDAE: ENTIMINAE: ENTIMITAE: BYRSOPAGINI: STRANGALIODINA.

Type species: Strangaliodes albosquamosus Boheman, 1842, by original designation.

Family-group name: STRANGALIODIDES Lacordaire, 1863. Stem: Strangaliod-.

Strategus W. Kirby in W. Kirby and Spence, 1828b: 349, 644. Current status: valid genus in SCARABAEIDAE: DYNASTINAE: ORYCTINI.

Type species: Scarabaeus aloeus Linnaeus, 1758, by subsequent designation (Hope 1837: 87).

Family-group name: STRATEGIDAE Burmeister, 1847. Stem: Strateg-.

Streptocerus Fairmaire, 1850: 53. Current status: valid genus in LUCANIDAE: LAMPRIMINAE: STREPTOCERINI.

Type species: Streptocerus speciosus Fairmaire, 1850, by monotypy.

Family-group name: STREPTOCERINI Kikuta, 1986. Stem: Streptocer-.

Striatoquasimus Schimmel & Tarnawski, 2009: 28, 105. Current status: valid genus in ELATERIDAE: NEGASTRIINAE: QUASIMUSINI: STRIATOQUASINA.

Type species: Striatoquasimus dolini Schimmel & Tarnawski, 2009, by original designation.

Family-group name: STRIATOQUASINA Schimmel & Tarnawski, 2009. Stem: Striatoquas- (ICZN 1999a, Article 29.4).

Strigota Casey, 1910: 176. Current status: valid genus in STAPHYLINIDAE: ALEOCHARINAE: ATHETINI: ATHETINA.

Type species: Strigota oppidana Casey, 1910 (= Homalota ambigua Erichson, 1839), by original designation (according to Casey’s first species rule, stipulated on page 90).

Family-group name: STRIGOTAE Casey, 1910. Stem: Strigot-.

Strombophorus Hagedorn, 1909: 740. Current status: valid genus in CURCULIONIDAE: SCOLYTINAE: DIAMERINI.

Type species: Strombophorus crenatus Hagedorn, 1909, by subsequent designation (Hopkins 1914: 130).

Family-group name: STROMBOPHORINI Schedl, 1959. Stem: Strombophor-.

Stromboscerus Schönherr, 1838: 814. Current status: valid genus in CURCULIONIDAE: DRYOPHTHORINAE: STROMBOSCERINI.

Type species: Stromboscerus schuppeli Gyllenhal, 1838, by original designation.

Family-group name: STROMBOSCÉRIDES Lacordaire, 1865. Stem: Stromboscer-.

Strongylium W. Kirby, 1819: 417. Current status: valid genus in TENEBRIONIDAE: STENOCHIINAE: STENOCHIINI. Note: nomen protectum (Bouchard et al. 2005: 501).

Type species: Strongylium chalconatum W. Kirby, 1819, by monotypy.

Family-group name: STRONGYLIIDES Lacordaire, 1859. Stem: Strongyli-.

Strongylopterus Schönherr, 1837: 473. Current status: valid genus in CURCULIONIDAE: CRYPTORHYNCHINAE: PSEPHOLACINI.

Type species: Strongylopterus ovatus Boheman, 1837, by original designation.

Family-group name: STRONGYLOPTÉRIDES Lacordaire, 1865. Stem: Strongylopter-.

Strongylurus Hope, 1834a: 107. Current status: valid genus in CERAMBYCIDAE: CERAMBYCINAE: STRONGYLURINI.

Type species: Strongylurus scutellatus Hope, 1834, by monotypy.

Family-group name: STRONGYLURIDES Lacordaire, 1868. Stem: Strongylur-.

Strongylus Herbst, 1792: 179. Current status: invalid senior synonym of Cyllodes Erichson, 1843 in NITIDULIDAE: NITIDULINAE: CYLLODINI. Note: junior homonym of Strongylus O.F. Müller, 1780 [Nematoda].

Type species [suggested]: Strongylus ater Herbst, 1792, by subsequent designation (Jelínek and Audisio 2003: 166). Note: the valid type species fixation is that of Curtis (1831: pl. 339), who selected Nitidula strigata Fabricius, 1787, but that is the type species of Cryptarcha Shuckard, 1840; an application to the Commission is needed to retain Strongylus ater Herbst, 1792 as the type species of Strongylus Herbst, 1792.

Family-group name: STRONGYLINAE Erichson, 1843. Stem: Strongyl-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

Strophosoma Billberg, 1820: 44. Current status: valid genus in CURCULIONIDAE: ENTIMINAE: POLYDRUSITAE: BRACHYDERINI. Note: we consider the name Strophosomum Gistel, 1856 as an incorrect subsequent spelling of Strophosoma Billberg, notwithstanding the status given by Alonso-Zarazaga and Lyal (1999: 147), who consider the name an unnecessary new replacement name for Billberg’s name.

Type species: Curculio coryli Fabricius, 1775 (= Curculio melanogrammus Forster, 1771), by subsequent designation (Pierce 1913: 411).

Family-group name: STROPHOSOMIDAE Gistel, 1848. Stem: Strophosomat-.

Styanax Pascoe, 1871b: 164. Current status: valid genus in CURCULIONIDAE: MOLYTINAE: STYANACINI.

Type species: Styanax carbonarius Pascoe, 1871, by monotypy.

Family-group name: STYANACINAE Chûjô & Voss, 1960. Stem: Styanac-.

Stylopodus Benick, 1917: 190. Current status: junior synonym of Megalopinus Eichelbaum, 1915 in STAPHYLINIDAE: MEGALOPSIDIINAE.

Type species: Megalops cephalotes Erichson, 1840, by original designation.

Family-group name: STYLOPODINAE Blackwelder, 1943. Stem: Stylopod-.

Stylosomus Suffrian, 1847: 8. Current status: valid genus in CHRYSOMELIDAE: CRYPTOCEPHALINAE: CRYPTOCEPHALINI: STYLOSOMINA.

Type species [suggested]: Cryptocephalus tamaricis Herrich-Schäffer, 1838, by subsequent designation (Gressitt and Kimoto 1961: 109). Note: Suffrian (1847: 8) originally included a single nominal species (p. 10) in his new genus, Cryptocephalus minutissimus Germar, 1823, but that species is currently placed in the subgenus Stylomicrus Petitpierre & Alonso-Zarazaga, 2000, not the nominotypical subgenus; subsequently, Suffrian (1848: 146) described the genus in length and included three species, including C. tamaricis; an application to the Commission is necessary to fix Cryptocephalus tamaricis Herrich- Schäffer, 1838 as the type species of Stylosomus Suffrian, 1847.

Family-group name: STYLOSOMITES Chapuis, 1874. Stem: Stylosom-.

Styphlomerus Chaudoir in Putzeys, 1875: iv. Current status: valid genus in CARABIDAE: BRACHININAE: BRACHININI: APTININA.

Type species: Brachinus quadrimaculatus Dejean, 1831, by subsequent designation (Andrewes 1938: 138).

Family-group name: STYPHLOMERINI Habu, 1984. Stem: Styphlomer-.

Styphlus Schönherr, 1826: 258. Current status: valid genus in CURCULIONIDAE: CURCULIONINAE: STYPHLINI.

Type species: Styphlus penicillus Schönherr, 1826, by original designation.

Family-group name: STYPHLIDAE Jekel, 1861. Stem: Styphl-.

Subcoccinella Agassiz, 1846: 156. Current status: valid genus in COCCINELLIDAE: COCCINELLINAE: EPILACHNINI. Note: this name was first introduced under the vernacular name “Subcoccinelle” by Huber (1842: 376) and therefore it is not available from that publication; this genus-group name has been treated as a junior objective synonym of Lasia Hope, 1841 (e.g., Tomaszewska and Szawaryn 2016: 68), which is a junior homonym of Lasia Wiedemann, 1824 [Diptera]; however, we could not find evidence that Agassiz (1846: 156) proposed Subcoccinella as a new replacement name for Lasia Hope.

Type species: Coccinella globosa D.H. Schneider, 1792 (= Coccinella vigintiquatuorpunctata Linnaeus, 1758), by original designation (type fixation for Lasia Hope).

Family-group name: SUBCOCCINELLINI Jakobson, 1915. Stem: Subcoccinell-.

Subprotelater Fleutiaux, 1916: 387. Current status: valid genus in ELATERIDAE: SUBPROTELATERINAE.

Type species: Subprotelater bakeri Fleutiaux, 1916, by monotypy.

Family-group name: SUBPROTELATERINAE Fleutiaux, 1920. Stem: Subprotelater-.

Sueus Murayama, 1951: 1. Current status: valid genus in CURCULIONIDAE: SCOLYTINAE: HYORRHYNCHINI.

Type species: Sueus sphaerotrypoides Murayama, 1951 (= Hyorrhynchus niisimai Eggers, 1926), by original designation.

Family-group name: SUEINAE Murayama, 1959. Stem: Sue-.

Sugimotoa Habu, 1975: 77. Current status: valid genus in CARABIDAE: HARPALINAE: LEBIINI: SUGIMOTOINA.

Type species: Sugimotoa parallela Habu, 1975 (= Pediomorphus anthracoides Straneo, 1967), by original designation.

Family-group name: SUGIMOTOINA Habu, 1975. Stem: Sugimoto-.

Sukunahikona Kamiya, 1960: 22. Current status: junior synonym of Scymnomorphus Weise, 1897 in COCCINELLIDAE: MICROWEISEINAE: SUKUNAHIKONINI.

Type species: Sukunahikona japonica Kamiya, 1960 (= Alexia japonica Reitter, 1889), by original designation.

Family-group name: SUKUNAHIKONINI Kamiya, 1960. Stem: Sukunahikon-.

Sumnius Weise, 1892: 29. Current status: valid genus in COCCINELLIDAE: COCCINELLINAE: COCCIDULINI.

Type species: Sumnius cardoni Weise, 1892, by subsequent designation (Korschefsky 1931: 94).

Family-group name: SUMNIINI Hoang, 1982. Stem: Sumni-.

Suniops Voss, 1929a: 362. Current status: valid genus in ATTELABIDAE: ATTELABINAE: EUOPINI. Note: new replacement name for Synechops Voss, 1924.

Type species: Euops semicuprea Voss, 1924, by subsequent designation (Legalov 2003: 365) (type fixation for Synechops Voss).

Family-group name: SUNIOPSINA Legalov, 2003. Stem: Suniops- (ICZN 1999a, Article 29.4).

Suphis Aubé, 1836: 42. Current status: valid genus in NOTERIDAE: NOTERINAE: NOTERINI.

Type species: Suphis cimicoides Aubé, 1837, by subsequent monotypy (Aubé 1837: 207).

Family-group name: SUPHISINI Sharp, 1880. Stem: Suphid-.

Svetlanaebyctiscus Legalov, 2001a: 341. Current status: valid genus in ATTELABIDAE: RHYNCHITINAE: RHYNCHITITAE: BYCTISCINI: SVETLANAEBYCTISCINA.

Type species: Coenorhinus vitis Ter-Minassian, 1959, by original designation.

Family-group name: SVETLANAEBYCTISCINA Legalov, 2003. Stem: Svetlanaebyctisc-.

Sybaris Stephens, 1832b: 70. Current status: valid genus in MELOIDAE: MELOINAE: LYTTINI.

Type species: Sybaris immunis Stephens, 1832, by monotypy.

Family-group name: SYBARIDES Wellman, 1910. Stem: Sybare-.

Sydax Lacordaire, 1868: 335. Current status: valid genus in CERAMBYCIDAE: CERAMBYCINAE: SYDACINI.

Type species: Sydax stramineus Lacordaire, 1868, by original designation.

Family-group name: SYDACINI Martins in Martins et al., 2014: 381. Stem: Sydac-.

Syllitus Pascoe, 1859a: 24. Current status: valid genus in CERAMBYCIDAE: CERAMBYCINAE: RHAGIOMORPHINI. Note: name placed on the Official List of Generic Names in Zoology (ICZN 2020a).

Type species: Stenoderus grammicus Newman, 1840, by subsequent designation (McKeown 1947: 73; see ICZN 2020a). Note: name placed on the Official List of Specific Names in Zoology (ICZN 2020a).

Family-group name: SYLLITAE J. Thomson, 1864. Stem: Syllit-.

Symbiotes L. Redtenbacher, 1847b: 198. Current status: valid genus in ANAMORPHIDAE.

Type species: Symbiotes latus L. Redtenbacher, 1847, by monotypy.

Family-group name: SYMBIOTINAE Joy, 1932. Stem: Symbiot-.

Symmathetes Schönherr, 1847: 31. Current status: junior synonym of Pantomorus Schönherr, 1840 in CURCULIONIDAE: ENTIMINAE: POLYDRUSITAE: NAUPACTINI: NAUPACTINA.

Type species: Symmathetes kollari Schönherr, 1847, by original designation.

Family-group name: SYMMATHETES J.L. LeConte, 1874. Stem: Symmathet-.

Symmerus Chapuis, 1865: 42, 319. Current status: invalid senior synonym of Chaetastus Nunberg, 1953 in CURCULIONIDAE: PLATYPODINAE: TESSEROCERINI: TESSEROCERINA. Note: junior homonym of Symmerus Walker, 1848 [Diptera].

Type species: Symmerus tuberculatus Chapuis, 1865, by monotypy.

Family-group name: SYMMERARIAE H. Strohmeyer, 1914. Stem: Symmer-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

Symmixus Bernhauer, 1915b: 56. Current status: valid genus in STAPHYLINIDAE: TACHYPORINAE: TACHYPORINI: TACHYPORINA.

Type species: Symmixus sikkimensis Bernhauer, 1915, by monotypy.

Family-group name: SYMMIXINI Bernhauer, 1915. Stem: Symmix-.

Sympiezopus Schönherr, 1837: 707. Current status: valid genus in CURCULIONIDAE: CONODERINAE: CORYSSOPODINI.

Type species: Sympiezopus aciculatus Schönherr, 1837, by original designation.

Family-group name: SYMPIEZOPINORUM Faust, 1886. Stem: Sympiezopod-.

Sympiezoscelus G.R. Waterhouse, 1853: 203. Current status: valid genus in CURCULIONIDAE: CRYPTORHYNCHINAE: PSEPHOLACINI.

Type species: Sympiezoscelus spencii G.R. Waterhouse, 1853, by monotypy.

Family-group name: SYMPIÉZOSCÉLIDES Lacordaire, 1865. Stem: Sympiezoscel-.

Sympolemon Wasmann, 1900b: 262. Current status: valid genus in STAPHYLINIDAE: ALEOCHARINAE: PYGOSTENINI.

Type species: Sympolemon anommatis Wasmann, 1900, by monotypy.

Family-group name: SYMPOLEMONINI Fenyes, 1919. Stem: Sympolemont-.

Synapion Schilsky, 1902: no 13. Current status: valid genus in BRENTIDAE: APIONINAE: APIONITAE: APIONINI: SYNAPIINA.

Type species: Apion ebeninum W. Kirby, 1808, by original designation.

Family-group name: SYNAPIINA Alonso-Zarazaga, 1990. Stem: Synapi-.

Synaptonyx G.R. Waterhouse, 1853: 187. Current status: valid genus in CURCULIONIDAE: ENTIMINAE: ENTIMITAE: BYRSOPAGINI: STRANGALIODINA.

Type species: Synaptonyx ovatus G.R. Waterhouse, 1853, by monotypy.

Family-group name: SYNAPTONYCIDES Lacordaire, 1863. Stem: Synaptonych-.

Synaptops Jekel, 1860: 221. Current status: valid genus in ATTELABIDAE: ATTELABINAE: EUOPINI.

Type species: Euops nietneri Jekel, 1860 (= Rhynchites suffundens Walker, 1859), by original designation.

Family-group name: SYNAPTOPSINA Legalov, 2003. Stem: Synaptops- (ICZN 1999a, Article 29.4).

Synaptus Eschscholtz, 1829a: 32. Current status: valid genus in ELATERIDAE: ELATERINAE: SYNAPTINI.

Type species: Elater filiformis Fabricius, 1781, by subsequent designation (Lacordaire 1857: 215).

Family-group name: SYNAPTIDAE Gistel, 1856. Stem: Synapt-.

Syncalypta Dillwyn, 1829: 25. Current status: junior synonym of Chaetophora W. Kirby, 1823 in BYRRHIDAE: SYNCALYPTINAE: SYNCALYPTINI.

Type species: Byrrhus arenarius Sturm, 1807 (= Byrrhus spinosus Rossi, 1794), by subsequent designation (Westwood 1838c: 21).

Family-group name: SYNCALYPTAIRES Mulsant & Rey, 1869. Stem: Syncalypt-.

Synchita Hellwig, 1792: 401. Current status: valid genus in ZOPHERIDAE: COLYDIINAE: SYNCHITINI.

Type species: Synchita juglandis Hellwig, 1792 (= Elophorus humeralis Fabricius, 1792), by subsequent designation (Blanchard in Audouin et al. 1843: pl. 62).

Family-group name: SYNCHITAE L. Redtenbacher, 1845. Stem: Synchit-.

Synchroa Newman, 1838: 378. Current status: valid genus in SYNCHROIDAE.

Type species: Synchroa punctata Newman, 1838, by monotypy.

Family-group name: SYNCHROÏDES Lacordaire, 1859. Stem: Synchro-.

Syncosmetus Sharp, 1891: 6. Current status: valid genus in CIIDAE: XYLOGRAPHELLINAE: SYNCOSMETINI.

Type species: Syncosmetus japonicus Sharp, 1891, by monotypy.

Family-group name: SYNCOSMETINA Lopes-Andrade, 2008. Stem: Syncosmet-.

Syndesus W.S. MacLeay, 1819: 96, 104. Current status: valid genus in LUCANIDAE: SYNDESINAE: SYNDESINI.

Type species: Sinodendron cornutum Fabricius, 1801, by monotypy.

Family-group name: SYNDESIDAE W.S. MacLeay, 1819. Stem: Syndes-.

Syndicus Motschulsky, 1852: 502. Current status: valid genus in STAPHYLINIDAE: SCYDMAENINAE: SCYDMAENITAE: STENICHNINI.

Type species: Syndicus pilicornis Motschulsky, 1852, by monotypy.

Family-group name: SYNDICINI Csiki, 1919. Stem: Syndic-.

Synechocera Deyrolle, 1865: 115. Current status: valid genus in BUPRESTIDAE: AGRILINAE: CORAEBINI: SYNECHOCERINA.

Type species: Buprestis deplana Fabricius, 1801, by monotypy.

Family-group name: SYNECHOCERINA Majer, 2000. Stem: Synechocer-.

Synercticus Newman, 1842c: 403. Current status: valid genus in BORIDAE: SYNERCTICINAE.

Type species: Synercticus heteromerus Newman, 1842, by monotypy.

Family-group name: SYNERCTICINAE Lawrence & Pollock, 1994. Stem: Synerctic-.

Syneta Dejean, 1835: 359. Current status: valid genus in CHRYSOMELIDAE: SYNETINAE.

Type species: Crioceris betulae Fabricius, 1792, by monotypy.

Family-group name: SYNETAE J.L. LeConte & G.H. Horn, 1883. Stem: Synet-.

Synirmus Bedel, 1883: 61. Current status: junior synonym of Tropiphorus Schönherr, 1842 in CURCULIONIDAE: ENTIMINAE: ENTIMITAE: BYRSOPAGINI: TROPIPHORINA. Note: unnecessary new replacement name for Tropiphorus Schönherr, 1842.

Type species: Curculio mercurialis Fabricius, 1801 (= Curculio elevatus Herbst, 1795), by original designation (see ICZN 1988c) (type fixation for Tropiphorus Schönherr).

Family-group name: SYNIRMINI Bedel, 1883. Stem: Synirm-.

Synochodaeus Kolbe, 1907: 27. Current status: valid genus in OCHODAEIDAE: CHAETOCANTHINAE: SYNOCHODAEINI.

Type species: Synochodaeus modestus Kolbe, 1907, by monotypy.

Family-group name: SYNOCHODAEINI Scholtz, 1988. Stem: Synochodae-.

Synoditulus Reichensperger, 1924: 212. Current status: valid genus in HISTERIDAE: HAETERIINAE: SYNODITULINI.

Type species: Synodites separatus Reichensperger, 1924, by monotypy.

Family-group name: SYNODITULINI Tishechkin, 2007. Stem: Synoditul-.

Synonycha Chevrolat in Dejean, 1836a: 436. Current status: valid genus in COCCINELLIDAE: COCCINELLINAE: COCCINELLINI.

Type species: Coccinella versicolor Fabricius, 1792 (= Coccinella grandis Thunberg, 1781), by monotypy.

Family-group name: SYNONYCHINI Weise, 1885. Stem: Synonych-.

Synophthalmus Lacordaire, 1863: 516. Current status: junior synonym of Phytophilus Schönherr, 1835 stat. nov. in CURCULIONIDAE: CONODERINAE: CORYSSOMERINI. Note: unnecessary new replacement name for Phytophilus Schönherr, 1835 since Phytophilus, as used by Guérin-Méneville (1838: 99), is an incorrect subsequent spelling for Phitophilus Guérin- Méneville, 1831 in TENEBRIONIDAE (see Bouchard et al. 2021: 538).

Type species: Phytophilus cruciferus Gyllenhal, 1835, by original designation (type fixation for Phytophilus Schönherr).

Family-group name: SYNOPHTHALMIDES Lacordaire, 1863. Stem: Synophthalm-.

Syntelia Westwood, 1864c: 11. Current status: valid genus in SYNTELIIDAE.

Type species: Syntelia indica Westwood, 1864, by subsequent designation (Yuasa 1930: 256).

Family-group name: SYNTELIIDAE Lewis, 1882. Stem: Synteli-.

Syntomium Curtis, 1828: pl. 228. Current status: valid genus in STAPHYLINIDAE: OXYTELINAE: SYNTOMIINI.

Type species: Syntomium nigroaeneum Curtis, 1828 (= Omalium aeneum P.W.J. Müller, 1821), by original designation.

Family-group name: SYNTOMIINAE Böving & Craighead, 1931. Stem: Syntomi-.

Synuchus Gyllenhal, 1810: 77. Current status: valid genus in CARABIDAE: HARPALINAE: SPHODRINI: SYNUCHINA.

Type species: Carabus vivalis Illiger, 1798, by monotypy.

Family-group name: SYNUCHI Lindroth, 1956. Stem: Synuch-.

Syphorbus Pascoe, 1881: 92. Current status: valid genus in CURCULIONIDAE: MOLYTINAE: HYLOBIINI: HYLOBIINA.

Type species: Syphorbus turgidus Pascoe, 1881, by monotypy.

Family-group name: SYPHORBINA G.A.K. Marshall, 1932. Stem: Syphorb-.

Sysolus Grouvelle, 1908: 429. Current status: valid genus in TEREDIDAE: TEREDINAE: SYSOLINI.

Type species: Sysolus antennatus Grouvelle, 1908, by monotypy.

Family-group name: SYSOLINI Ślipiński & Pal, 1985. Stem: Sysol-.

Systellopus Sharp, 1877a: 315. Current status: valid genus in SCARABAEIDAE: MELOLONTHINAE: SYSTELLOPINI.

Type species: Systellopus obtusus Sharp, 1877, by subsequent designation (Allsopp 1990: 204).

Family-group name: SYSTELLOPIDES Sharp, 1877. Stem: Systellop-.

Systena Chevrolat in Dejean, 1836a: 390. Current status: valid genus in CHRYSOMELIDAE: GALERUCINAE: ALTICINI.

Type species: Galleruca frontalis Fabricius, 1801, by subsequent designation (Monrós and Bechyné 1956: 1133).

Family-group name: SYSTENAE G.H. Horn, 1889. Stem: Systen-.

Systenocerus Weise in Heyden et al., 1883: 93. Current status: junior synonym of Platycerus Geoffroy, 1762 in LUCANIDAE: LUCANINAE: PLATYCERINI.

Type species: Scarabaeus caraboides Linnaeus, 1758, by subsequent designation (Didier and Séguy 1953: 169).

Family-group name: SYSTENOCERINI Portevin, 1931. Stem: Systenocer-.

Systolosoma Solier, 1849: 241. Current status: valid genus in TRACHYPACHIDAE: TRACHYPACHINAE.

Type species: Systolosoma breve Solier, 1849, by original designation.

Family-group name: SYSTOLOSOMINI Erwin, 1991. Stem: Systolosomat-.

Syzetoninus Blackburn, 1891: 339. Current status: valid genus in ADERIDAE.

Type species: Syzetoninus mundus Blackburn, 1891, by subsequent designation (Nardi 2007: 30).

Family-group name: SYZETONININI Báguena Corella, 1948. Stem: Syzetonin-.

Tachinus Gravenhorst, 1802: 134. Current status: valid genus in STAPHYLINIDAE: TACHYPORINAE: TACHINUSINI. Note: name placed on the Official List of Generic Names in Zoology (ICZN 1993g).

Type species: Staphylinus rufipes Linnaeus, 1758, by subsequent designation (Latreille 1810: 427; see ICZN 1993g). Note: name placed on the Official List of Specific Names in Zoology (ICZN 1993g).

Family-group name: TACHINIDAE Fleming, 1821. Stem: Tachinus- (ICZN 1993g).

Tachygonus Guérin-Méneville, 1833a: pl. 38. Current status: valid genus in CURCULIONIDAE: CURCULIONINAE: RHAMPHINI: TACHYGONINA. Note: the name Tachygonus was proposed simultaneously by both Schönherr (1833: 6, 311) and Guérin-Méneville (1833a: pl. 38) in June 1833 and recorded on 6 July 1833 in the ‘Bibliographie de la France’ (see Bousquet 2016: 234, 484); Alonso-Zarazaga and Lyal (1999: 82) acted as First Revisers (ICZN 1999a, Article 24.2.2) and selected Guérin-Méneville’s name to have precedence; we consider Tachygonus used by Schönherr, 1833 a subsequent usage of Tachygonus Guérin-Méneville, 1833.

Type species: Tachygonus horridus Guérin-Méneville, 1833, by monotypy.

Family-group name: TACHYGONIDES Lacordaire, 1865. Stem: Tachygon-.

Tachyoryctidium Jeannel & Paulian, 1945: 58. Current status: junior synonym of Euxestoxenus Arrow, 1925 in EUXESTIDAE.

Type species: Tachyoryctidium chappuisi Jeannel & Paulian, 1945, by original designation.

Family-group name: TACHYORYCTIDIINI Jeannel & Paulian, 1945. Stem: Tachyoryctidi-.

Tachyporus Gravenhorst, 1802: 124. Current status: valid genus in STAPHYLINIDAE: TACHYPORINAE: TACHYPORINI: TACHYPORINA. Note: name placed on the Official List of Generic Names in Zoology (ICZN 1993g).

Type species: Staphylinus chrysomelinus Linnaeus, 1758, by subsequent designation (Latreille 1810: 427; see ICZN 1993g). Note: name placed on the Official List of Specific Names in Zoology (ICZN 1993g).

Family-group name: TACHYPORIDAE W.S. MacLeay, 1825. Stem: Tachypor-.

Tachypterellus Fall & Cockerell, 1907: 214. Current status: valid subgenus of Anthonomus Germar, 1817 in CURCULIONIDAE: CURCULIONINAE: ANTHONOMINI. Note: new replacement name for Tachypterus Dietz, 1891.

Type species: Anthonomus quadrigibbus Say, 1832, by original designation (type fixation for Tachypterus Dietz).

Family-group name: TACHYPTERELLINA Voss, 1944. Stem: Tachypterell-.

Tachys Dejean, 1821: 16. Current status: valid genus in CARABIDAE: TRECHINAE: TACHYINI: TACHYINA. Note: name placed on the Official List of Generic Names in Zoology (ICZN 1990c).

Type species: Tachys scutellaris Stephens, 1828, by plenary power (see ICZN 1990c). Note: name placed on the Official List of Specific Names in Zoology (ICZN 1990c).

Family-group name: TACHYAIRES Motschulsky, 1862. Stem: Tachy-.

Tachyusa Erichson, 1837: 307. Current status: valid genus in STAPHYLINIDAE: ALEOCHARINAE: TACHYUSINI. Note: name placed on the Official List of Generic Names in Zoology (ICZN 1961b).

Type species: Tachyusa constricta Erichson, 1837, by plenary power (see ICZN 1961b). Note: name placed on the Official List of Specific Names in Zoology (ICZN 1961b).

Family-group name: TACHYUSIDES C.G. Thomson, 1859. Stem: Tachyus-.

Tadius Pascoe, 1885: 253. Current status: valid genus in CURCULIONIDAE: BRACHYCERINAE: ERIRHININI.

Type species: Tadius erirhinoides Pascoe, 1885, by monotypy.

Family-group name: TADIINI Zimmerman, 1993. Stem: Tadi-.

Taeniocerus W.F.H. Blandford, 1893: 437. Current status: junior synonym of Scolytoplatypus C. Schaufuss, 1891 in CURCULIONIDAE: SCOLYTINAE: SCOLYTOPLATYPODINI. Note: junior homonym of Taeniocerus Kaup, 1871 [Coleoptera: PASSALIDAE].

Type species: Scolytoplatypus mikado W.F.H. Blandford, 1893, by subsequent designation (Hopkins 1914: 130).

Family-group name: TAENIOCERINI W.F.H. Blandford, 1893. Stem: Taeniocer-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

Taeniodera Burmeister, 1842: 325. Current status: valid genus in SCARABAEIDAE: CETONIINAE: TAENIODERINI: TAENIODERINA.

Type species: Macronota monacha Gory & Percheron, 1834, by subsequent designation (Cazurro Ruiz 1897a: 585).

Family-group name: TAENIODERINA Mikšić, 1976. Stem: Taenioder-.

Taenioncus Kirejtshuk, 1984: 192. Current status: valid genus in NITIDULIDAE: EPURAEINAE: TAENIONCINI.

Type species: Carpophilus cylindricus Murray, 1864, by original designation.

Family-group name: TAENIONCINI Kirejtshuk, 1998. Stem: Taenionc-.

Taeniotes Audinet-Serville, 1835a: 90. Current status: valid genus in CERAMBYCIDAE: LAMIINAE: LAMIINI.

Type species: Cerambyx pulverulentus G.-A. Olivier, 1790, by subsequent designation (E. Desmarest 1860: 324).

Family-group name: TAENIOTITAE J. Thomson, 1864. Stem: Taeniot-.

Tagenia Latreille, 1802b: 170. Current status: junior synonym of Stenosis Herbst, 1799 in TENEBRIONIDAE: PIMELIINAE: STENOSINI: STENOSINA.

Type species: Tenebrio filiformis Fabricius, 1792, by monotypy.

Family-group name: TAGÉNITES Solier, 1834. Stem: Tageni-.

†Taimyraltica Nadein in Nadein and Perkovsky, 2018: 98. Current status: valid genus in CHRYSOMELIDAE: GALERUCINAE: †TAIMYRALTICINI. Note: Upper Cretaceous (Russia).

Type species: †Taimyraltica calcarata Nadein, 2018, by original designation.

Family-group name: †TAIMYRALTICINI Nadein in Nadein and Perkovsky 2017: 98. Stem: Taimyraltic-.

Tainophthalmus Desbrochers des Loges, 1873b: 426. Current status: valid genus in CURCULIONIDAE: ENTIMINAE: POLYDRUSITAE: TANYMECINI: TAINOPHTHALMINA.

Type species: Tainophthalmus crotchi Desbrochers des Loges, 1873, by monotypy.

Family-group name: TAINOPHTHALMIDAE Desbrochers des Loges, 1873. Stem: Tainophthalm-.

Talanus Jacquelin du Val, 1857a: 156. Current status: valid genus in TENEBRIONIDAE: STENOCHIINAE: TALANINI.

Type species: Talanus cribrarius Jacquelin du Val, 1857, by monotypy.

Family-group name: TALANIDES Champion, 1887. Stem: Talan-.

†Taldycupes Rohdendorf, 1961: 421. Current status: valid genus in †PERMOCUPEDIDAE: †TALDYCUPEDINAE. Note: Permian (Russia).

Type species: †Taldycupes khalfini Rohdendorf, 1961, by original designation.

Family-group name: †TALDYCUPIDAE Rohdendorf, 1961. Stem: Taldycuped-.

Tamotus L.W. Schaufuss, 1872: 248. Current status: valid genus in STAPHYLINIDAE: EUAESTHETINAE: EUAESTHETINI.

Type species: Tamotus femoratus L.W. Schaufuss, 1874, by subsequent monotypy (Schaufuss 1874: 289).

Family-group name: TAMOTINI Coiffait, 1984. Stem: Tamot-.

Tamulus Gistel, 1848b: ix, 127. Current status: junior synonym of Malachius Fabricius, 1775 in MELYRIDAE: MALACHIINAE: MALACHIINI. Note: unnecessary new replacement name for Malachius Fabricius, 1775.

Type species: Cantharis aenea Linnaeus, 1758, by subsequent designation (Latreille 1810: 426) (type fixation for Malachius Fabricius).

Family-group name: TAMULIDAE Gistel, 1848. Stem: Tamul-.

Tanaos Schönherr, 1826: 63. Current status: valid genus in BRENTIDAE: APIONINAE: TANAITAE: TANAINI.

Type species: Apion sanguineum Thunberg, 1815, by original designation.

Family-group name: TANAONIDES Schönherr, 1839. Stem: Tana-.

Tanygnathinus Reitter, 1909: 164. Current status: junior synonym of Atanygnathus Jakobson, 1909 in STAPHYLINIDAE: STAPHYLININAE: STAPHYLININI: TANYGNATHININA. Note: replacement name for Tanygnathus Erichson, 1839.

Type species: Tanygnathus terminalis Erichson, 1839, by monotypy (type fixation for Tanygnathus Erichson).

Family-group name: TANYGNATHININI Reitter, 1909. Stem: Tanygnathin-.

Tanygnathus Erichson, 1839a: 417. Current status: invalid senior synonym of Atanygnathus Jakobson, 1909 in STAPHYLINIDAE: STAPHYLININAE: STAPHYLININI: TANYGNATHININA. Note: junior homonym of Tanygnathus Wagler, 1832 [Aves].

Type species: Tanygnathus terminalis Erichson, 1839, by monotypy.

Family-group name: TANYGNATHINI Casey, 1915. Stem: Tanygnath-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

Tanymecus Germar, 1817b: 341. Current status: valid genus in CURCULIONIDAE: ENTIMINAE: POLYDRUSITAE: TANYMECINI: TANYMECINA.

Type species: Curculio palliatus Fabricius, 1787, by subsequent designation (Schönherr 1823: 1144).

Family-group name: TANYMÉCIDES Lacordaire, 1863. Stem: Tanymec-.

Tanypleurus Raffray, 1890a: 128, 130. Current status: invalid senior synonym of Natypleurus Newton & Thayer, 1992 in STAPHYLINIDAE: PSELAPHINAE: GONIACERITAE: INIOCYPHINI: NATYPLEURINA. Note: junior homonym of Tanypleurus Steenstrup & Lütken, 1861 [Crustacea].

Type species: Tanypleurus malaianus Raffray, 1890, by subsequent monotypy (Raffray 1890b: 206).

Family-group name: TANYPLEURINI Jeannel, 1949. Stem: Tanypleur-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

Tanyproctus Ménétriés, 1832: 185. Current status: valid genus in SCARABAEIDAE: MELOLONTHINAE: TANYPROCTINI: TANYPROCTINA.

Type species: Tanyproctus persicus Ménétriés, 1832, by monotypy.

Family-group name: TANYPROCTINI Erichson, 1847. Stem: Tanyproct-.

Tanyrhynchus Schönherr, 1826: 212. Current status: valid genus in CURCULIONIDAE: ENTIMINAE: ENTIMITAE: TANYRHYNCHINI.

Type species: Tanyrhynchus terranus Schönherr, 1826 (= Curculio strigirostris Sparrman, 1785), by original designation.

Family-group name: TANYRHYNCHIDES Schönherr, 1826. Stem: Tanyrhynch-.

Tanysphyrus Germar, 1817b: 340. Current status: valid genus in CURCULIONIDAE: BRACHYCERINAE: TANYSPHYRINI.

Type species: Curculio lemnae Fabricius, 1792, by monotypy.

Family-group name: TANYSPHYRIDAE Gistel, 1848. Stem: Tanysphyr-.

Tapeina Lepeletier & Audinet-Serville, 1828: 545. Current status: valid genus in CERAMBYCIDAE: LAMIINAE: TAPEININI.

Type species: Tapeina coronata Lepeletier & Audinet-Serville, 1828, by subsequent designation (Chevrolat 1849a: 344).

Family-group name: TAPEINITES J. Thomson, 1857. Stem: Tapein-.

Taphes C.O. Waterhouse, 1878: 102. Current status: valid genus in LYCIDAE: EROTINAE: TAPHINI.

Type species: Taphes brevicollis C.O. Waterhouse, 1878, by original designation.

Family-group name: TAPHININA Bocák & Bocáková, 1990. Stem: Taph-.

Taphroderes Schönherr, 1823: 1137. Current status: valid genus in BRENTIDAE: BRENTINAE: TAPHRODERINI.

Type species: Brentus foveatus Lund, 1802, by original designation.

Family-group name: TAPHRODÉRIDES Lacordaire, 1865. Stem: Taphroder-.

Taphrorychus Eichhoff, 1878: 204. Current status: valid genus in CURCULIONIDAE: SCOLYTINAE: DRYOCOETINI.

Type species: Bostrichus bicolor Herbst, 1793, by subsequent designation (Hopkins 1914: 130).

Family-group name: TAPHRORYCHINI Reitter, 1913. Stem: Taphrorych-.

Tapinotus Schönherr, 1826: 21, 292. Current status: valid genus in CURCULIONIDAE: CEUTORHYNCHINAE: CEUTORHYNCHINI. Note: the spelling Tapeinotus (p. 21) was corrected to Tapinotus in the corrigenda [unpaginated page].

Type species: Tapinotus ephippiger Schönherr, 1826 (= Attelabus sellatus Fabricius, 1794), by original designation.

Family-group name: TAPINOTINAE Joy, 1932. Stem: Tapinot-.

Tapiromimus Wollaston, 1877: 149. Current status: valid genus in CURCULIONIDAE: COSSONINAE: TAPIROMIMINI.

Type species: Tapiromimus gibbirostris Wollaston, 1877, by monotypy.

Family-group name: TAPIROMIMINI Voss, 1972. Stem: Tapiromim-.

Tarphius Erichson, 1845a: 256. Current status: valid genus in ZOPHERIDAE: COLYDIINAE: SYNCHITINI.

Type species: Tarphius gibbulus Erichson, 1845, by monotypy.

Family-group name: TARPHIINAE Sharp, 1894. Stem: Tarphi-.

Tarquinius Kuwert, 1891: 164. Current status: valid genus in PASSALIDAE: PASSALINAE: MACROLININI.

Type species: Tarquinius paradoxus Kuwert, 1891, by monotypy.

Family-group name: TARQUININAE Kuwert, 1891. Stem: Tarquini-.

Tarsostenus Spinola, 1845a: 287. Current status: valid genus in CLERIDAE: KORYNETINAE.

Type species: Clerus univittatus Rossi, 1792, by monotypy.

Family-group name: TARSOSTÉNITES Jacquelin du Val, 1860. Stem: Tarsosten-.

Tarus Clairville, 1806: 94. Current status: junior synonym of Cymindis Latreille, 1805 in CARABIDAE: HARPALINAE: LEBIINI: CYMINDIDINA.

Type species: Buprestis humeralis Geoffroy, 1785, by subsequent designation (Curtis 1828: pl. 235).

Family-group name: TARIDAE Gistel, 1848. Stem: Tar-.

Tasmosalpingus Lea, 1919a: 743. Current status: valid genus in TASMOSALPINGIDAE.

Type species: Tasmosalpingus quadrispilotus Lea, 1919, by original designation.

Family-group name: TASMOSALPINGINAE Lawrence & Britton, 1991. Stem: Tasmosalping-.

Taurocerastes R.A. Philippi, 1865: 115. Current status: valid genus in GEOTRUPIDAE: TAUROCERASTINAE.

Type species: Taurocerastes patagonicus R.A. Philippi, 1865, by monotypy.

Family-group name: TAUROCERASTIDAE Germain, 1897. Stem: Taurocerast-.

Tauroma Hope, 1839: 97. Current status: junior synonym of Omocerus Chevrolat, 1835 in CHRYSOMELIDAE: CASSIDINAE: OMOCERINI.

Type species: Cassida taurus Fabricius, 1787, by original designation.

Family-group name: TAUROMITAE Spaeth, 1923. Stem: Taurom-.

Taxicera Mulsant & Rey, 1873: 188. Current status: valid genus in STAPHYLINIDAE: ALEOCHARINAE: TAXICERINI.

Type species: Taxicera perfoliata Mulsant & Rey, 1873, by subsequent designation (Blackwelder 1952: 374).

Family-group name: TAXICERINA Lohse, 1989. Stem: Taxicer-.

Techmessa F. Bates, 1874: 113. Current status: valid genus in PYROCHROIDAE: PILIPALPINAE.

Type species: Techmessa concolor F. Bates, 1874, by subsequent designation (Watt 1987: 124).

Family-group name: TECHMESSINAE Paulus, 1972. Stem: Techmess-.

Tefflus Leach, 1819: 494. Current status: valid genus in CARABIDAE: HARPALINAE: PANAGAEINI: TEFFLINA.

Type species: Carabus meyerlei Fabricius, 1801, by monotypy.

Family-group name: TEFFLINI Basilewsky, 1946. Stem: Teffl-.

Tegrodera J.L. LeConte, 1851a: 159. Current status: valid genus in MELOIDAE: MELOINAE: EUPOMPHINI.

Type species: Tegrodera erosa J.L. LeConte, 1851, by monotypy.

Family-group name: TEGRODERINI L.S. Dillon, 1952. Stem: Tegroder-.

Telacis Poey, 1854: 322. Current status: junior synonym of Cteniopus Solier, 1835 in TENEBRIONIDAE: ALLECULINAE: CTENIOPODINI. Note: new replacement name for Cistela Fabricius, 1775.

Type species: Chrysomela sulphurea Linnaeus, 1758, by subsequent designation (Latreille 1810: 429) (type fixation for Cistela Fabricius).

Family-group name: TELACIANAE Poey, 1854. Stem: Telac-.

Teledapus Pascoe, 1871c: 268. Current status: valid genus in CERAMBYCIDAE: LEPTURINAE: TELEDAPINI.

Type species: Teledapus dorcadioides Pascoe, 1871, by monotypy.

Family-group name: TELEDAPINAE Pascoe, 1871. Stem: Teledap-.

Telegeusis G.H. Horn, 1895b: 242. Current status: valid genus in OMETHIDAE: TELEGEUSINAE.

Type species: Telegeusis debilis G.H. Horn, 1895, by monotypy.

Family-group name: TELEGEUSIDAE Leng, 1920. Stem: Telegeus-.

Telephanus Erichson, 1846: 329. Current status: valid genus in SILVANIDAE: BRONTINAE: TELEPHANINI.

Type species: Telephanus atricapillus Erichson, 1846, by monotypy.

Family-group name: TELEPHANIDAE J.L. LeConte, 1861. Stem: Telephan-.

Telephorus Schäffer, 1766: pl. cxxiii. Current status: junior synonym of Cantharis Linnaeus, 1758 in CANTHARIDAE: CANTHARINAE: CANTHARINI. Note: this name was first made available by Schäffer (1764) under the spelling Thelophorus (see Bousquet 2016: 469); the spelling Telephorus, which is in prevailing usage, is an incorrect subsequent spelling of Thelephorus but since Telephorus is not attributed to the publication of the original spelling, Article 33.3.1 (ICZN 1999a) cannot be used; an application to the Commission is necessary to conserve the currently-used spelling Telephorus.

Type species: Cantharis fusca Linnaeus, 1758, by subsequent designation (Latreille 1810: 426).

Family-group name: TELEPHORIDES Leach, 1815. Stem: Telephor-.

Telmatophilus Heer, 1841: 417. Current status: valid genus in CRYPTOPHAGIDAE: CRYPTOPHAGINAE: CRYPTOPHAGINI.

Type species: Ips caricis G.-A. Olivier, 1790, by subsequent designation (Chevrolat 1849a: 445). Note: Cryptophagus typhae Fallén, 1802 is often listed as type species of Telmatophilus, by subsequent designation of Leschen (1996: 603) but Chevrolat’s designation is earlier; both species are currently included in the genus Telmatophilus Heer.

Family-group name: TELMATOPHILIDES Jacquelin du Val, 1859. Stem: Telmatophil-.

Telochilus Krikken, 1975: 21. Current status: valid genus in SCARABAEIDAE: CETONIINAE: CREMASTOCHEILINI: TELOCHILINA.

Type species: Telochilus freyi Krikken, 1975, by original designation.

Family-group name: TELOCHILINA Krikken, 1984. Stem: Telochil-.

Telsimia Casey, 1899: 165. Current status: valid genus in COCCINELLIDAE: COCCINELLINAE: TELSIMIINI.

Type species: Telsimia tetrasticta Casey, 1899, by subsequent designation (Chapin 1926: 130).

Family-group name: TELSIMIINI Casey, 1899. Stem: Telsimi-.

Temnaspis Lacordaire, 1845: 716. Current status: valid genus in MEGALOPODIDAE: MEGALOPODINAE: MEGALOPODINI: TEMNASPIDINA.

Type species: Megalopus javanus Guérin-Méneville, 1844, by subsequent designation (Jacoby and Clavareau 1905: 12).

Family-group name: TEMNASPIDINA Rodríguez-Mirón in Rodríguez-Mirón et al., 2021: 695. Stem: Temnaspid-.

Temnocerus Thunberg, 1815: 110. Current status: valid genus in ATTELABIDAE: RHYNCHITINAE: RHYNCHITITAE: RHYNCHITINI: TEMNOCERINA.

Type species: Attelabus planirostris Fabricius, 1801 (= Curculio nanus Paykull, 1792), by subsequent designation (Crotch 1870b: 227).

Family-group name: TEMNOCERINA Legalov, 2003. Stem: Temnocer-.

Temnoscheila Westwood, 1830b: 231. Current status: valid genus in TROGOSSITIDAE: TROGOSSITINAE: TROGOSSITINI.

Type species: Trogossita caerulea G.-A. Olivier, 1790, by subsequent designation (E. Desmarest 1851: 275).

Family-group name: TEMNOCHILINI Léveillé, 1888. Stem: Temnoscheil-.

Temnostega Enderlein, 1905: 719. Current status: valid genus in CARABIDAE: TRECHINAE: TRECHINI: TRECHINA.

Type species: Temnostega antarctica Enderlein, 1905, by monotypy.

Family-group name: TEMNOSTEGINI Enderlein, 1909. Stem: Temnosteg-.

Tenebrio Linnaeus, 1758: 417. Current status: valid genus in TENEBRIONIDAE: TENEBRIONINAE: TENEBRIONINI.

Type species: Tenebrio molitor Linnaeus, 1758, by subsequent designation (Latreille 1810: 429).

Family-group name: TENEBRIONITES Latreille, 1802. Stem: Tenebrion-.

Tenebroides Piller & Mitterpacher, 1783: 87. Current status: valid genus in TROGOSSITIDAE: TROGOSSITINAE: TROGOSSITINI.

Type species: Tenebroides complanatus Piller & Mitterpacher, 1783 (= Tenebrio mauritanicus Linnaeus, 1758), by monotypy.

Family-group name: TENEBRIOIDINI Ganglbauer, 1899. Stem: Tenebroid-.

Tentyria Latreille, 1802b: 170. Current status: valid genus in TENEBRIONIDAE: PIMELIINAE: TENTYRIINI. Note: name placed on the Official List of Generic Names in Zoology (ICZN 2010b).

Type species: Tentyria ligurica Solier, 1835, by plenary power (see ICZN 2010b). Note: name placed on the Official List of Specific Names in Zoology (ICZN 2010b).

Family-group name: TENTYRIDAE Eschscholtz, 1831. Stem: Tentyri-.

Tephraea Burmeister, 1842: 419. Current status: valid genus in SCARABAEIDAE: CETONIINAE: CETONIINI: CETONIINA.

Type species: Cetonia pulverulenta Gory & Percheron, 1834, by subsequent designation (Kraatz 1882a: 67).

Family-group name: TEPHRAEIDES Schenkling, 1921. Stem: Tephrae-.

Teplinus Pakaluk, Ślipiński & Lawrence, 1994: 228. Current status: valid genus in CORYLOPHIDAE: CORYLOPHINAE: TEPLININI. Note: new replacement name for Peltinus Mulsant & Rey, 1861.

Type species: Peltinus velatus Mulsant & Rey, 1861, by monotypy (type fixation for Peltinus Mulsant & Rey).

Family-group name: TEPLININI Pakaluk, Ślipiński & Lawrence, 1994. Stem: Teplin-.

Terambus Gistel, 1848b: 130. Current status: junior synonym of Aromia Audinet-Serville, 1834 in CERAMBYCIDAE: CERAMBYCINAE: CALLICHROMATINI.

Type species: Cerambyx moschatus Linnaeus, 1758, by subsequent designation (Bousquet et al. 2009: 42).

Family-group name: TERAMBIDAE Gistel, 1848. Stem: Teramb-.

Teredus Dejean, 1835: 313. Current status: valid genus in TEREDIDAE: TEREDINAE: TEREDINI.

Type species: Lyctus nitidus Fabricius, 1792 (= Ips cylindrica G.-A. Olivier, 1790), by monotypy.

Family-group name: TEREDINI Seidlitz, 1888. Stem: Tered-.

Tereticus C.O. Waterhouse, 1879c: 534. Current status: valid genus in CERAMBYCIDAE: PRIONINAE: TERETICINI.

Type species: Tereticus pectinicornis C.O. Waterhouse, 1879, by monotypy.

Family-group name: TERETICI Lameere, 1913. Stem: Teretic-.

Teretrius Erichson, 1834: 201. Current status: valid genus in HISTERIDAE: ABRAEINAE: TERETRIINI.

Type species: Hister picipes Fabricius, 1792 (= Teretrius fabricii Mazur, 1972), by monotypy.

Family-group name: TERETRIINAE Bickhardt, 1914. Stem: Teretri-.

Termitella Wasmann, 1911: 170. Current status: valid genus in STAPHYLINIDAE: ALEOCHARINAE: COROTOCINI: TERMITOGASTRINA.

Type species: Termitella lujae Wasmann, 1911, by monotypy.

Family-group name: TERMITELLICI Jacobson, Kistner & Pasteels, 1986. Stem: Termitell-.

Termitochara Wasmann, 1893b: 247. Current status: valid genus in STAPHYLINIDAE: ALEOCHARINAE: COROTOCINI: TERMITOCHARINA.

Type species: Termitochara kraatzi Wasmann, 1893, by monotypy.

Family-group name: TERMITOCHARINA Seevers, 1957. Stem: Termitochar-.

Termitocupidus Jacobson, Kistner & Pasteels, 1986: 36. Current status: valid genus in STAPHYLINIDAE: ALEOCHARINAE: COROTOCINI: TERMITOCUPIDINA.

Type species: Termitocupidus judithae Jacobson, Kistner & Pasteels, 1986, by original designation.

Family-group name: TERMITOCUPIDINA Jacobson, Kistner & Pasteels, 1986. Stem: Termitocupid-.

Termitoderus Mateu, 1966: 718. Current status: valid genus in SCARABAEIDAE: APHODIINAE: TERMITODERINI.

Type species: Termitoderus grassei Mateu, 1966, by monotypy.

Family-group name: TERMITODERINI Tangelder & Krikken, 1982. Stem: Termitoder-.

Termitodiscus Wasmann, 1899: 147. Current status: valid genus in STAPHYLINIDAE: ALEOCHARINAE: TERMITODISCINI.

Type species: Termitodiscus heimi Wasmann, 1899, by monotypy.

Family-group name: TERMITODISCINI Wasmann, 1904. Stem: Termitodisc-.

Termitogaster Casey, 1889: 384. Current status: valid genus in STAPHYLINIDAE: ALEOCHARINAE: COROTOCINI: TERMITOGASTRINA.

Type species: Termitogaster insolens Casey, 1889, by monotypy.

Family-group name: TERMITOGASTRI Bernhauer & Scheerpeltz, 1926. Stem: Termitogastr-.

Termitohospes Seevers, 1941: 333. Current status: valid genus in STAPHYLINIDAE: ALEOCHARINAE: TERMITOHOSPITINI: TERMITOHOSPITINA.

Type species: Termitohospes miricorniger Seevers, 1941, by original designation.

Family-group name: TERMITOHOSPINI Seevers, 1941. Stem: Termitohospit-.

Termitoiceus Silvestri, 1901: 5. Current status: valid genus in STAPHYLINIDAE: ALEOCHARINAE: COROTOCINI: TERMITOICEINA.

Type species: Termitoiceus anastrephoproctus Silvestri, 1901, by original designation.

Family-group name: TERMITOICEINA Jacobson, Kistner & Pasteels, 1986. Stem: Termitoice-.

Termitomimus Trägårdh, 1907: 173. Current status: valid genus in STAPHYLINIDAE: ALEOCHARINAE: COROTOCINI: COROTOCINA.

Type species: Termitomimus entendveniensis Trägårdh, 1907, by monotypy.

Family-group name: TERMITOMIMINI Fenyes, 1921. Stem: Termitomim-.

Termitonannus Wasmann, 1902: 2. Current status: valid genus in STAPHYLINIDAE: ALEOCHARINAE: TERMITONANNINI: TERMITONANNINA.

Type species: Termitonannus schmalzi Wasmann, 1902, by subsequent designation (Fenyes 1919: 25).

Family-group name: TERMITONANNINI Fenyes, 1919. Stem: Termitonann-.

Termitonda Seevers, 1957: 238. Current status: valid genus in STAPHYLINIDAE: ALEOCHARINAE: TERMITOPAEDIINI.

Type species: Termitonda liberiae Seevers, 1957, by original designation.

Family-group name: TERMITONDINA Seevers, 1957. Stem: Termitond-.

Termitopaedia Wasmann, 1911: 114. Current status: valid genus in STAPHYLINIDAE: ALEOCHARINAE: TERMITOPAEDIINI.

Type species: Termitopaedia kohli Wasmann, 1911, by monotypy.

Family-group name: TERMITOPAEDIINI Seevers, 1957. Stem: Termitopaedi-.

Termitopithus Seevers, 1957: 80. Current status: valid genus in STAPHYLINIDAE: ALEOCHARINAE: COROTOCINI: TERMITOPITHINA.

Type species: Termitopithus crassiusculus Seevers, 1957, by original designation.

Family-group name: TERMITOPITHINA Jacobson, Kistner & Pasteels, 1986. Stem: Termitopith-.

Termitopsenius Wasmann, 1902: 4. Current status: valid genus in STAPHYLINIDAE: ALEOCHARINAE: TRICHOPSENIINI.

Type species: Termitopsenius limulus Wasmann, 1902, by monotypy.

Family-group name: TERMITOPSENINI Wasmann, 1916. Stem: Termitopseni-.

Termitoptochus Silvestri, 1910: 37. Current status: valid genus in STAPHYLINIDAE: ALEOCHARINAE: COROTOCINI: TERMITOPTOCHINA.

Type species: Termitoptochus indicus Silvestri, 1910, by monotypy.

Family-group name: TERMITOPTOCHINI Fenyes, 1921. Stem: Termitoptoch-.

Termitospectrum Mann, 1926: 154. Current status: valid genus in STAPHYLINIDAE: ALEOCHARINAE: TERMITUSINI: TERMITOSPECTRINA.

Type species: Termitospectrum thoracicum Mann, 1926, by original designation.

Family-group name: TERMITOSPECTRINA Seevers, 1957. Stem: Termitospectr-.

Termitotelus Wasmann, 1908: 444. Current status: valid genus in STAPHYLINIDAE: ALEOCHARINAE: ATHETINI: TERMITOTELINA.

Type species: Termitotelus schultzei Wasmann, 1908, by monotypy.

Family-group name: TERMITOTELINA Kistner, 1970. Stem: Termitotel-.

Termitotrox Reichensperger, 1915: 16. Current status: valid genus in SCARABAEIDAE: TERMITOTROGINAE.

Type species: Termitotrox consobrinus Reichenbacher, 1915, by monotypy.

Family-group name: TERMITOTROGINI Wasmann, 1918. Stem: Termitotrog-.

Termitozyras Seevers, 1957: 242. Current status: valid genus in STAPHYLINIDAE: ALEOCHARINAE: LOMECHUSINI: TERMITOZYRINA.

Type species: Termitozyras adamsoni Seevers, 1957, by original designation.

Family-group name: TERMITOZYRINA Seevers, 1957. Stem: Termitozyr-.

Termitusa Wasmann, 1905: 199. Current status: valid genus in STAPHYLINIDAE: ALEOCHARINAE: TERMITUSINI: TERMITUSINA.

Type species: Termitusa sjoestedti Wasmann, 1905, by monotypy.

Family-group name: TERMITUSAE Fenyes, 1919. Stem: Termitus-.

Tessaromma Newman, 1840a: 20. Current status: valid genus in CERAMBYCIDAE: CERAMBYCINAE: TESSAROMMATINI.

Type species: Tessaromma undatum Newman, 1840, by monotypy.

Family-group name: TESSAROMMIDES Lacordaire, 1868. Stem: Tessarommat-.

Tesserocerus Saunders, 1837: 155, 157. Current status: valid genus in CURCULIONIDAE: PLATYPODINAE: TESSEROCERINI: TESSEROCERINA.

Type species: Platypus insignis Saunders, 1837, by monotypy.

Family-group name: TESSEROCERINAE H. Strohmeyer, 1914. Stem: Tesserocer-.

Tetrabrachys Kapur, 1948: 319. Current status: valid genus in COCCINELLIDAE: COCCINELLINAE: COCCIDULINI. Note: new replacement name for Lithophilus Frölich, 1799.

Type species: Lithophilus ruficollis Frölich, 1799 (= Tritoma connata Panzer, 1796), by monotypy (type fixation for Lithophilus Frölich).

Family-group name: TETRABRACHINAE Kapur, 1948. Stem: Tetrabrach-.

Tetracha Hope, 1838a: 6, 7. Current status: valid genus in CICINDELIDAE: MEGACEPHALINI. Note: nomen protectum (see Bousquet 2002: 23).

Type species: Cicindela carolina Linnaeus, 1763, by original designation.

Family-group name: TETRACHAE Leng & Mutchler, 1916. Stem: Tetrach-.

Tetradelus Fauvel, 1904: 90. Current status: valid genus in STAPHYLINIDAE: OMALIINAE: OMALIINI.

Type species: Tetradelus trigonuroides Fauvel, 1904, by monotypy.

Family-group name: TETRADELI Fauvel, 1904. Stem: Tetradel-.

Tetragonoderus Dejean, 1829: 5, 485. Current status: valid genus in CARABIDAE: HARPALINAE: CYCLOSOMINI. Note: nomen protectum (see Bousquet 2002: 50).

Type species: Carabus quadrum Fabricius, 1792, by subsequent designation (Hope 1838a: 89).

Family-group name: TÉTRAGONODÉRIDES Chaudoir, 1871. Stem: Tetragonoder-.

Tetralobus Lepeletier & Audinet-Serville, 1828: 594. Current status: valid genus in ELATERIDAE: TETRALOBINAE: TETRALOBINI.

Type species: Elater flabellicornis Linnaeus, 1767, by subsequent designation (Lacordaire 1857: 164).

Family-group name: TETRALOBITES Laporte, 1840. Stem: Tetralob-.

†Tetrameropsis Kirejtshuk & Azar, 2008: 37. Current status: valid genus in †TETRAMEROPSIDAE. Note: Lower Cretaceous (Lebanon).

Type species: †Tetrameropsis mesozoica Kirejtshuk & Azar, 2008, by original designation.

Family-group name: †TETRAMEROPSINAE Kirejtshuk & Azar, 2008. Stem: Tetramerops- (ICZN 1999a, Article 29.4).

Tetraonyx Latreille, 1809b: 204, 237. Current status: valid genus in MELOIDAE: TETRAONYCINAE.

Type species: Tetraonyx octomaculatum Latreille, 1809, by monotypy.

Family-group name: TETRAONYCIDAE Böving & Craighead, 1931. Stem: Tetraonyc-.

Tetraopes Schönherr, 1817: 401. Current status: valid genus in CERAMBYCIDAE: LAMIINAE: TETRAOPINI.

Type species: Lamia tornator Fabricius, 1775 (= Cerambyx tetrophthalmus Forster, 1771), by subsequent designation (Drapiez 1845: 374).

Family-group name: TETRAOPESITAE J. Thomson, 1860. Stem: Tetraop-.

Tetraphalerus C.O. Waterhouse, 1901: 520. Current status: valid genus in OMMATIDAE: TETRAPHALERINAE.

Type species: Tetraphalerus wagneri C.O. Waterhouse, 1901, by monotypy.

Family-group name: TETRAPHALERINI Crowson, 1962. Stem: Tetraphaler-.

Tetratoma Fabricius, 1790: 217. Current status: valid genus in TETRATOMIDAE: TETRATOMINAE: TETRATOMINI.

Type species: Tetratoma fungorum Fabricius, 1790, by subsequent designation (Latreille 1810: 429).

Family-group name: TETRATOMAEDES Billberg, 1820. Stem: Tetratom-.

Tetraulax Jordan, 1903: 182. Current status: valid genus in CERAMBYCIDAE: LAMIINAE: TETRAULAXINI.

Type species: Tetraulax lateralis Jordan, 1903, by original designation.

Family-group name: TETRAULAXINI Breuning & Téocchi, 1977. Stem: Tetraulax-.

Tetrigus Candèze, 1857: 254. Current status: valid genus in ELATERIDAE: AGRYPNINAE: HEMIRHIPINI: LUDIOCTENINA.

Type species: Tetrigus parallelus Candèze, 1857, by subsequent designation (Hyslop 1921: 671).

Family-group name: TETRIGUSINA Schimmel & Tarnawski, 2012: 116. Stem: Tetrigus- (ICZN 1999a, Article 29.4).

Tetropium W. Kirby, 1837: 174. Current status: valid genus in CERAMBYCIDAE: SPONDYLIDINAE: TETROPIINI. Note: name placed on the Official List of Generic Names in Zoology (ICZN 1988b).

Type species: Callidium cinnamopterum W. Kirby, 1837, by subsequent designation (J. Thomson 1864: 266; see ICZN 1988b). Note: name placed on the Official List of Specific Names in Zoology (ICZN 1988b).

Family-group name: TETROPIINA Seidlitz, 1891. Stem: Tetropi-.

Tetrops Stephens, 1829b: 16. Current status: valid genus in CERAMBYCIDAE: LAMIINAE: TETROPINI. Note: this name was first made available by W. Kirby in W. Kirby and Spence (1826: 498) citing only Lamia tornator Fabricius, 1775 (= Cerambyx tetrophthalmus Forster, 1771) in it, but currently that species is placed in the genus Tetraopes Dalman, 1817; as advocated by Vives and Alonso- Zarazaga (2000: 660–661) and Bousquet et al. (2009: 38–39), a request should be submitted to the Commission to suppress the name Tetrops W. Kirby, 1826 for the purpose of the Principles of Priority and Homonymy.

Type species: Leptura praeusta Linnaeus, 1758, by monotypy.

Family-group name: TETROPINI Portevin, 1927. Stem: Tetrop-.

Thalasselephas Egorov & Korotyaev, 1976: 47. Current status: valid genus in CURCULIONIDAE: MOLYTINAE: EMPHYASTINI.

Type species: Thalasselephas major Egorov & Korotyaev, 1976, by original designation.

Family-group name: THALASSELEPHANTINI Alonso-Zarazaga & Lyal, 1999. Stem: Thalasselephant-.

Thalassophilus Wollaston, 1854: xx, 71. Current status: valid genus in CARABIDAE: TRECHINAE: TRECHINI: TRECHODINA.

Type species: Thalassophilus whitei Wollaston, 1854, by monotypy.

Family-group name: THALASSOPHILI Csiki, 1928. Stem: Thalassophil-.

Thalia Hope, 1838a: 70. Current status: junior synonym of Poecilus Bonelli, 1810 in CARABIDAE: HARPALINAE: PTEROSTICHINI: PTEROSTICHINA. Note: unnecessary new replacement name for Feronia Latreille, 1816; junior homonym of Thalia Bruguière, 1791 [Mollusca] and Thalia Blumenbach, 1827 [Tunicata].

Type species: Carabus cupreus Linnaeus, 1758, by subsequent designation (Blanchard in Audouin et al. 1842: pl. 22) (type fixation for Feronia Latreille).

Family-group name: THALIADAE Hope, 1838. Stem: Thali-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

Thaliabaris Bondar, 1943: 131. Current status: valid genus in CURCULIONIDAE: BARIDINAE: MADARINI: MADARINA.

Type species: Thaliabaris gica Bondar, 1943, by original designation.

Family-group name: THALIABARINI Bondar, 1943. Stem: Thaliabarid-.

Thallisella Crotch, 1876: 402. Current status: valid genus in EROTYLIDAE: LANGURIINAE: THALLISELLINI.

Type species: Thallisella peruviana Crotch, 1876, by original designation.

Family-group name: THALLISELLINI Sen Gupta, 1968. Stem: Thallisell-.

Thalycra Erichson, 1843a: 305. Current status: valid genus in NITIDULIDAE: NITIDULINAE: NITIDULINI.

Type species: Strongylus sericeus Sturm, 1839 (= Nitidula fervida G.-A. Olivier, 1790), by monotypy.

Family-group name: THALYCRINA C.G. Thomson, 1859. Stem: Thalycr-.

Thamiaraea C.G. Thomson, 1858: 35. Current status: valid genus in STAPHYLINIDAE: ALEOCHARINAE: ATHETINI: THAMIARAEINA.

Type species: Aleochara cinnamomea Gravenhorst, 1802, by monotypy.

Family-group name: THAMIARAEINI Fenyes, 1921. Stem: Thamiarae-.

Thamnophilus Schönherr, 1823: 1136. Current status: junior synonym of Magdalis Germar, 1817 in CURCULIONIDAE: MESOPTILIINAE: MAGDALIDINI. Note: unnecessary new replacement name for Magdalis Germar, 1817; junior homonym of Thamnophilus Vieillot, 1816 [Aves]; name placed on the Official Index of Rejected and Invalid Names in Zoology (see ICZN 1955a).

Type species: Curculio violaceus Linnaeus, 1758, by plenary power (see ICZN 1955a) (type fixation for Magdalis Germar).

Family-group name: THAMNOPHILIDES Schönherr, 1823. Stem: Thamnophil-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

Thamnurgus Eichhoff, 1864: 40, 45. Current status: valid genus in CURCULIONIDAE: SCOLYTINAE: DRYOCOETINI.

Type species: Bostrichus euphorbiae Küster, 1845, by subsequent designation (Cazurro Ruiz 1897a: 186).

Family-group name: THAMNURGINAE Nüsslin, 1911. Stem: Thamnurg-.

Thanasimus Latreille, 1805: 270. Current status: valid genus in CLERIDAE: CLERINAE: CLERITAE: CLERINI.

Type species: Attelabus formicarius Linnaeus, 1758, by subsequent designation (Latreille 1810: 427).

Family-group name: THANASIMIIDAE Gistel, 1848. Stem: Thanasim-.

Thaneroclerus Lefebvre, 1838b: xiii. Current status: valid genus in THANEROCLERIDAE: THANEROCLERINAE: THANEROCLERINI.

Type species: Clerus buqueti Lefebvre, 1835 [as buquet], by monotypy.

Family-group name: THANEROCLERINAE Chapin, 1924. Stem: Thanerocler-.

Thaumaphrastus Blaisdell, 1927: 123. Current status: junior synonym of Thorictodes Reitter, 1875 in DERMESTIDAE: DERMESTINAE: THORICTINI.

Type species: Thaumaphrastus karanisensis Blaisdell, 1927 (= Thorictodes heydeni Reitter, 1875), by monotypy.

Family-group name: THAUMAPHRASTINAE W.H. Anderson, 1949. Stem: Thaumaphrast-.

Thaumastocephalus Poggi, Nonveiller, Colla, Pavićević & Rada, 2001: 3. Current status: valid genus in STAPHYLINIDAE: PSELAPHINAE: BATRISITAE: THAUMASTOCEPHALINI.

Type species: Thaumastocephalus folliculipalpus Poggi, Nonveiller, Colla, Pavićević & Rada, 2001, by original designation.

Family-group name: THAUMASTOCEPHALINI Poggi, Nonveiller, Colla, Pavićević & Rada, 2001. Stem: Thaumastocephal-.

Thaumastodus Champion, 1924: 25. Current status: junior synonym of Pseudeucinetus Heller, 1921 in LIMNICHIDAE: THAUMASTODINAE.

Type species: Thaumastodus fusiformis Champion, 1924 (= Pseudeucinetus zygops Heller, 1921), by original designation.

Family-group name: THAUMASTODINAE Champion, 1924. Stem: Thaumastod-.

Thaumasus Reiche, 1853: xxiii. Current status: valid genus in CERAMBYCIDAE: CERAMBYCINAE: TORNEUTINI.

Type species: Ips gigas G.-A. Olivier, 1792, by monotypy.

Family-group name: THAUMASIDAE J. Thomson, 1864. Stem: Thaumas-.

†Thayeralinus Solodovnikov & Yue in Solodovnikov et al., 2013: 383. Current status: valid genus in STAPHYLINIDAE: STAPHYLININAE: †THAYERALININI. Note: Lower Cretaceous (China).

Type species: †Thayeralinus fieldi Solodovnikov & Yue, 2013, by original designation.

Family-group name: †THAYERALININI Solodovnikov & Yue in Solodovnikov et al., 2013: 383. Stem: Thayeralin-.

Thecesternus Say, 1831: 8. Current status: valid genus in CURCULIONIDAE: CURCULIONINAE: THECESTERNINI.

Type species: Brachycerus humeralis Say, 1827, by monotypy.

Family-group name: THÉCESTERNIDES Lacordaire, 1863. Stem: Thecestern-.

Thecturota Casey, 1893: 357. Current status: valid genus in STAPHYLINIDAE: ALEOCHARINAE: HOMALOTINI: HOMALOTINA.

Type species: Thecturota tenuissima Casey, 1893, by subsequent designation (Fenyes 1919: 25).

Family-group name: THECTUROTAE Fenyes, 1919. Stem: Thecturot-.

Theocerus Raffray, 1897a: 280. Current status: valid genus in STAPHYLINIDAE: PSELAPHINAE: CLAVIGERITAE: CLAVIGERINI.

Type species: Theocerus crenulatus Raffray, 1897, by monotypy.

Family-group name: THEOCERINI Jeannel, 1954. Stem: Theocer-.

Theocris J. Thomson, 1858b: 193. Current status: valid genus in CERAMBYCIDAE: LAMIINAE: THEOCRINI.

Type species: Theocris saga J. Thomson, 1858, by monotypy.

Family-group name: THÉOCRIDES Lacordaire, 1872. Stem: Theocr-.

Theone Gistel, 1857: 585. Current status: valid genus in CHRYSOMELIDAE: GALERUCINAE: GALERUCINI.

Type species: Galleruca silphoides Dalman, 1823, by original designation.

Family-group name: THEONINA Laboissière, 1934. Stem: Theon-.

Therates Latreille, 1816: 179. Current status: valid genus in CICINDELIDAE: CICINDELINI: THERATINA.

Type species: Cicindela labiata Weber, 1801, by monotypy. Note: this species was described in 1801 by both Weber (p. 44) and Fabricius (p. 232); bibliographic research suggests that Weber’s book (issued by March 1801, see Evenhuis 1997b: 809) was issued before Fabricius first volume of his ‘Systema Eleutheratorum’ (issued by 26 April 1801, see Bousquet 2017a: 108; see also Chapin 1946: 79).

Family-group name: THERATIDAE W. Horn, 1893. Stem: Therat-.

Thermonectus Dejean, 1833a: 53. Current status: valid genus in DYTISCIDAE: DYTISCINAE: ACILIINI. Note: the original spelling was Thermonetus but Thermonectus, an incorrect subsequent spelling, is in prevailing usage, attributed to the publication of the original spelling, and therefore deemed to be the correct original spelling (ICZN 1999a, Article 33.3.1).

Type species: Dytiscus circumscriptus Latreille, 1809, by monotypy.

Family-group name: THERMONECTINI Sharp, 1880. Stem: Thermonect-.

Thilmanus Baudi di Selve, 1872: 97. Current status: valid genus in ELATERIDAE: OMALISINAE. Note: this name is credited to Gemminger in Gemminger and Harold (1869c: 1683) by the vast majority of authors in the literature but there is no description nor indication that would make the name available from that work.

Type species: Thilmanus obscurus Baudi di Selve, 1872, by monotypy. Note: Baudi di Selve (1874: clxxxvii) noted that the word “obscurus” was erroneously left out (at the time of the impression of the work) before the description, which started with the word “fuscus” and that subsequent authors have incorrectly interpreted that the name of the species was Thilmanus fuscus.

Family-group name: THILMANINAE Kazantsev, 2004. Stem: Thilman-.

Thinobatis Eschscholtz, 1831: 5, 8. Current status: valid genus in TENEBRIONIDAE: PIMELIINAE: THINOBATINI.

Type species: Thinobatis ferruginea Eschscholtz, 1831, by monotypy.

Family-group name: THINOBATIDES Lacordaire, 1859. Stem: Thinobat-.

Thinobius Kiesenwetter, 1844: 355. Current status: valid genus in STAPHYLINIDAE: OXYTELINAE: THINOBIINI.

Type species: Thinobius ciliatus Kiesenwetter, 1844, by monotypy.

Family-group name: THINOBIIDES J.R. Sahlberg, 1876. Stem: Thinobi-.

Thinopinus J.L. LeConte, 1852d: 215. Current status: valid genus in STAPHYLINIDAE: STAPHYLININAE: STAPHYLININI: STAPHYLININA.

Type species: Thinopinus pictus J.L. LeConte, 1852, by monotypy.

Family-group name: THINOPININAE Böving & Craighead, 1931. Stem: Thinopin-.

Thinorycter Semenov-Tjan-Shansky & Reikhardt, 1925: 83. Current status: valid genus in SCARABAEIDAE: DYNAMOPODINAE: THINORYCTERINI.

Type species: Thinorycter chlamydatus Semenov-Tjan-Shansky & Reikhardt, 1925, by original designation.

Family-group name: THINORYCTERINA Semenov-Tjan-Shansky & Reikhardt, 1925. Stem: Thinorycter-.

Thione Sharp, 1899: 544. Current status: valid genus in MONOTOMIDAE: MONOTOMINAE: THIONINI.

Type species: Thione cephalotes Sharp, 1899, by subsequent designation (Sen Gupta 1988: 51).

Family-group name: THIONINAE Crowson, 1952. Stem: Thion-.

Thisias Champion, 1889: 102. Current status: valid genus in MYCTERIDAE: EURYPINAE.

Type species: Thisias marmoratus Champion, 1889, by monotypy.

Family-group name: THISIINA Seidlitz, 1917. Stem: Thisi-.

Thliboclivina Kult, 1959: 175 [in key]. Current status: valid genus in CARABIDAE: SCARITINAE: CLIVININI: ARDISTOMINA.

Type species: Clivina microphthalma Burgeon, 1937, by original designation.

Family-group name: THLIBOCLIVININA Balkenohl, 2022: 107. Stem: Thliboclivin-.

Thomassetia Théry, 1929: 183. Current status: valid genus in BUPRESTIDAE: BUPRESTINAE: THOMASSETIINI: THOMASSETIINA.

Type species: Philanthaxia natalensis Théry, 1929, by monotypy.

Family-group name: THOMASSETIINI Bellamy, 1987. Stem: Thomasseti-.

Thonalmus Bourgeois, 1884: 376. Current status: valid genus in LYCIDAE: LYCINAE: THONALMINI.

Type species: Cantharis bicolor Linnaeus, 1763, by subsequent designation (Bourgeois 1891: 343).

Family-group name: THONALMINI Kleine, 1933. Stem: Thonalm-.

Thoracophorus Motschulsky, 1837: 98. Current status: valid genus in STAPHYLINIDAE: OSORIINAE: THORACOPHORINI: THORACOPHORINA. Note: the original spelling was Thoraxophorus but Thoracophorus, an unjustified emendation by Gemminger and von Harold (1868b: 677), is in prevailing usage, attributed to the original author and date, and therefore is deemed to be a justified emendation (ICZN 1999a, Article 33.2.3.1).

Type species: Thoracophorus corticinus Motschulsky, 1837, by monotypy.

Family-group name: THORACOPHORINAE Reitter, 1909. Stem: Thoracophor-.

Thorictosoma Lea, 1919b: 257. Current status: valid genus in TENEBRIONIDAE: PIMELIINAE: CNEMEPLATIINI: THORICTOSOMATINA.

Type species: Thorictosoma ectatommae Lea, 1919, by original designation.

Family-group name: THORICTOSOMATINA Watt, 1992. Stem: Thorictosomat-.

Thorictus Germar, 1834: No 15. Current status: valid genus in DERMESTIDAE: DERMESTINAE: THORICTINI.

Type species: Thorictus castaneus Germar, 1834, by monotypy.

Family-group name: THORICTIDES Erichson, 1846. Stem: Thorict-. Note: the authorship of this family-group name was credited to “Agassiz” in Bouchard et al. (2011: 334) but here we follow Bouchard et al. (2021: 9) in crediting new nomenclatural acts in the fascicle on Coleoptera of Agassiz’ ‘Nomenclator Zoologicus’ to the beetle specialist W.F. Erichson, whose contribution to the publication was acknowledged on the fascicle’s title page.

Thranius Pascoe, 1859a: 22. Current status: valid genus in CERAMBYCIDAE: CERAMBYCINAE: THRANIINI.

Type species: Thranius bimaculatus Pascoe, 1859, by subsequent designation (J. Thomson 1864: 162).

Family-group name: THRANIINI Gahan, 1906. Stem: Thrani-.

Thrincopyge J.L. LeConte, 1858b: 17. Current status: valid genus in BUPRESTIDAE: POLYCESTINAE: THRINCOPYGINI.

Type species: Buprestis ambiens J.L. LeConte, 1854, by original designation.

Family-group name: THRINCOPYGINI J.L. LeConte, 1861. Stem: Thrincopyg-.

Thrombosternus G.A.K. Marshall, 1955b: 270. Current status: valid genus in CURCULIONIDAE: MOLYTINAE: CYCLOTERINI: THROMBOSTERNINA.

Type species: Thrombosternus cucullatus G.A.K. Marshall, 1955, by original designation.

Family-group name: THROMBOSTERNINA Voss, 1965. Stem: Thrombostern-.

†Throscogenius Iablokoff-Khnzorian, 1962: 81. Current status: valid genus in EUCNEMIDAE: MACRAULACINAE: †THROSCOGENIINI. Note: Eocene (Lithuania).

Type species: †Throscogenius takhtajani Iablokoff-Khnzorian, 1962, by original designation.

Family-group name: †THROSCOGENIINAE Iablokoff-Khnzorian, 1962. Stem: Throscogeni-.

Throscus Latreille, 1797: 42. Current status: junior synonym of Trixagus Kugelann, 1794 in THROSCIDAE.

Type species: Elater dermestoides Linnaeus, 1767, by subsequent designation (Curtis 1827: pl. 163). Note: no species were originally included in this genus by Latreille (1797); the first ones were the two species listed by Latreille (1802b: 100).

Family-group name: THROSCITES Laporte, 1840. Stem: Throsc-. Note: nomen protectum (see Bouchard et al. 2011: 306).

Thryptocerus Chaudoir, 1878b: 74. Current status: valid genus in CARABIDAE: HARPALINAE: OODINI.

Type species: Thryptocerus politus Chaudoir, 1878, by monotypy.

Family-group name: THRYPTOCERINI Jeannel, 1949. Stem: Thryptocer-.

Thylacites Germar, 1817b: 341. Current status: invalid senior synonym of Brachyderes Schönherr, 1823 in CURCULIONIDAE: ENTIMINAE: POLYDRUSITAE: BRACHYDERINI. Note: name suppressed for the Principle of Priority and placed on the Official Index of Rejected and Invalid Names in Zoology (ICZN 1987a).

Type species: Curculio incanus Linnaeus, 1758, by subsequent designation (Samouelle 1819: 205). Note: name placed on the Official List of Specific Names in Zoology (ICZN 1987a).

Family-group name: THYLACITIDAE W. Kirby, 1837. Stem: Thylacit-. Note: family-group name permanently invalid (ICZN 1999a, Article 39); name placed on the Official List of Rejected and Invalid Family-Group Names in Zoology (ICZN 1987a).

Thylacosternus Gemminger in Gemminger and Harold, 1869b: 1486. Current status: valid genus in ELATERIDAE: THYLACOSTERNINAE. Note: this name is credited to Bonvouloir (1871: 76) in the literature but it was made available by Gemminger in 1869 when he associated an available species with it; the other ten species listed by Gemminger and von Harold (1869b: 1486, 1487) are nomina nuda.

Type species: Pterotarsus walckenaerii Guérin-Méneville, 1843, by monotypy.

Family-group name: THYLACOSTERNINAE Fleutiaux, 1920. Stem: Thylacostern-.

Thylodrias Motschulsky, 1839: 75. Current status: valid genus in DERMESTIDAE: TRINODINAE: THYLODRIINI.

Type species: Thylodrias contractus Motschulsky, 1839, by monotypy.

Family-group name: THELYDRIINI Semenov-Tjan-Shansky, 1909. Stem: Thylodri-.

Thymalus Latreille, 1802b: 133. Current status: valid genus in THYMALIDAE: THYMALINAE.

Type species: Cassida brunnea Thunberg, 1787 (= Cassida limbata Fabricius, 1787), by monotypy.

Family-group name: THYMALINI Léveillé, 1888. Stem: Thymal-.

Thyreopterus Dejean, 1831: 279, 445. Current status: valid genus in CARABIDAE: HARPALINAE: LEBIINI: PERICALINA.

Type species: Thyreopterus flavosignatus Dejean, 1831, by subsequent designation (Hope 1838a: 105).

Family-group name: THYRÉOPTÉRIDES Chaudoir, 1869. Stem: Thyreopter-.

Thyrsia Dalman, 1819: 117. Current status: valid genus in CERAMBYCIDAE: CERAMBYCINAE: THYRSIINI.

Type species: Thyrsia lateralis Dalman, 1819, by monotypy.

Family-group name: THYRSIINI Marinoni & Napp, 1984. Stem: Thyrsi-.

Thysanotus Chaudoir, 1848: 123. Current status: valid genus in CARABIDAE: HARPALINAE: LEBIINI: PERICALINA.

Type species: Eurydera anchomenoides Laporte & Gory, 1835, by monotypy.

Family-group name: THYSANOTINI Jeannel, 1949. Stem: Thysanot-.

Thysdarius Fairmaire, 1904b: 117. Current status: valid genus in STAPHYLINIDAE: PSELAPHINAE: CLAVIGERITAE: CLAVIGERINI. Note: new replacement name for Thysdrus Fairmaire, 1899.

Type species: Thysdrus perrieri Fairmaire, 1899, by monotypy (type fixation for Thysdrus Fairmaire).

Family-group name: THYSDARIINI Jeannel, 1954. Stem: Thysdari-.

Tibionema Solier, 1851: 30. Current status: valid genus in ELATERIDAE: PITYOBIINAE: *TIBIONEMINI.

Type species: Tibionema rufiventre Solier, 1851, by original designation.

Family-group name: *TIBIONEMINI Motyka, Kusy, Arias-Bohart, Bybee & Bocák, 2023a: 7. Stem: Tibionem-.

Tichonia Semenov, 1904: 201. Current status: junior synonym of Microcephalus Dejean, 1828 in CARABIDAE: HARPALINAE: PTEROSTICHINI: MICROCEPHALINA. Note: new replacement name for Caletor Tschitschérine 1903, an unnecessary new replacement name for Microcephalus Dejean, 1828; junior homonym of Tichonia Hübner, 1825 [Lepidoptera].

Type species: Microcephalus depressicollis Dejean, 1828, by monotypy (type fixation for Microcephalus Dejean).

Family-group name: TICHONIINI Semenov, 1904. Stem: Tichoni-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

Tichonilla Strand, 1942: 393. Current status: junior synonym of Microcephalus Dejean, 1828 in CARABIDAE: HARPALINAE: PTEROSTICHINI: MICROCEPHALINA. Note: unnecessary new replacement name for Tichonia Semenov, 1904 (an unnecessary new replacement name for Microcephalus Dejean, 1828).

Type species: Microcephalus depressicollis Dejean, 1828, by monotypy (type fixation for Microcephalus Dejean).

Family-group name: TICHONILLINA Emden, 1958. Stem: Tichonill-.

Tillomorpha Blanchard, 1851: 482. Current status: valid genus in CERAMBYCIDAE: CERAMBYCINAE: TILLOMORPHINI.

Type species: Tillomorpha lineoligera Blanchard, 1851, by monotypy.

Family-group name: TILLOMORPHIDES Lacordaire, 1868. Stem: Tillomorph-.

Tillus G.-A. Olivier, 1790a: [22] 3. Current status: valid genus in CLERIDAE: TILLINAE.

Type species: Chrysomela elongata Linnaeus, 1758, by subsequent designation (Latreille 1810: 427). Note: name placed on the Official List of Specific Names in Zoology (ICZN 2011c).

Family-group name: TILLII Fischer, 1813. Stem: Till-.

Timarcha Samouelle, 1819: 213. Current status: valid genus in CHRYSOMELIDAE: CHRYSOMELINAE: TIMARCHINI. Note: this genus is often credited to Latreille (1829b: 150) (e.g., Gómez- Zurita and Kippenberg 2010: 437) but it was made available earlier by Samouelle in 1819 when he associated two available nominal species, Chrysomela tenebricosa Fabricius, 1775 (on page 213) and Chrysomela coriaria Laicharting, 1781 (on page 445), with the name.

Type species: Chrysomela tenebricosa Fabricius, 1775, by subsequent designation (Chevrolat 1843a: 655). Note: the type species often listed for the genus Timarcha, Tenebrio laevigatus Linnaeus, 1767 (e.g., Gómez-Zurita and Kippenberg 2010: 437), is not a species originally included species in Timarcha; Chrysomela tenebricosa Fabricius, 1775 was given precedence over Tenebrio laevigatus Linnaeus, 1767 and placed on the Official List of Specific Names in Zoology (ICZN 2021b).

Family-group name: TIMARCHAEINES Motschulsky, 1860. Stem: Timarch-.

†Timarchopsis Brauer, J. Redtenbacher & Ganglbauer, 1889: 17. Current status: valid genus in †COPTOCLAVIDAE: †TIMARCHOPSINAE. Note: Jurassic (Russia).

Type species: †Timarchopsis czekanowskii Brauer, J. Redtenbacher & Ganglbauer, 1889, by monotypy.

Family-group name: †TIMARCHOPSINAE B. Wang, Ponomarenko & Zhang, 2010. Stem: Timarchops- (ICZN 1999a, Article 29.4).

Timeparthenus Silvestri, 1901: 10. Current status: valid genus in STAPHYLINIDAE: ALEOCHARINAE: COROTOCINI: TIMEPARTHENINA.

Type species: Timeparthenus regius Silvestri, 1901, by original designation.

Family-group name: TIMEPARTHENINI Fenyes, 1921. Stem: Timeparthen-.

Timola Pascoe, 1886: 331. Current status: valid genus in CURCULIONIDAE: CURCULIONINAE: CURCULIONINI: TIMOLINA.

Type species: Timola suturalis Pascoe, 1886 (= Balaninus suturalis Gyllenhal, 1835), by monotypy.

Family-group name: TIMOLINA Heller, 1925. Stem: Timol-.

Tiracerus Besuchet, 1986: 262. Current status: valid genus in STAPHYLINIDAE: PSELAPHINAE: CLAVIGERITAE: TIRACERINI.

Type species: Articerus curvicornis Westwood, 1856, by original designation.

Family-group name: TIRACERINI Besuchet, 1986. Stem: Tiracer-.

Tisiphone Reitter, 1876a: 297. Current status: invalid senior synonym of Smicrips J.L. LeConte, 1878 in SMICRIPIDAE. Note: junior homonym of Tisiphone Hübner, 1819 [Lepidoptera].

Type species: Tisiphone hypocoproides Reitter, 1876, by subsequent designation (Casey 1916: 104)

Family-group name: TISIPHONINAE Sharp, 1900. Stem: Tisiphon-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

Titaena Erichson, 1842: 178. Current status: valid genus in TENEBRIONIDAE: TENEBRIONINAE: TITAENINI.

Type species: Titaena columbina Erichson, 1842, by subsequent designation (Chevrolat 1849a: 594).

Family-group name: TITAENINI Fauvel, 1905. Stem: Titaen-.

Titanus Audinet-Serville, 1832: 133. Current status: valid genus in CERAMBYCIDAE: PRIONINAE: PRIONINI.

Type species: Cerambyx giganteus Linnaeus, 1771, by monotypy.

Family-group name: TITANITAE J. Thomson, 1864. Stem: Titan-.

Tmesiphorus J.L. LeConte, 1849: 75. Current status: valid genus in STAPHYLINIDAE: PSELAPHINAE: PSELAPHITAE: TMESIPHORINI.

Type species: Pselaphus carinatus Say, 1824, by subsequent designation (Bowman 1934: 144).

Family-group name: TMESIPHORINI Jeannel, 1949. Stem: Tmesiphor-.

Tmesisternus Latreille, 1829b: 121. Current status: valid genus in CERAMBYCIDAE: LAMIINAE: TMESISTERNINI.

Type species: Tmesisternus bizonulatus Guérin-Méneville, 1831, by subsequent monotypy (Guérin-Méneville 1831b: pl. 45). Note: this species is incorrectly listed in the literature as a junior synonym of Tmesisternus trivittatus Guérin-Méneville, 1831, which is available from plate 7 (issued 13 October 1831, see Bousquet 2016: 235) of the ‘Voyage autour du monde...La Coquille’; T. bizonulatus is available from plate 45 (issue by March 1831, see Bousquet 2016: 234) of the ‘Iconographie du règne animal’.

Family-group name: TMESISTERNITAE Blanchard, 1853. Stem: Tmesistern-.

Tmesorrhina Westwood, 1842b: 71. Current status: valid genus in SCARABAEIDAE: CETONIINAE: GOLIATHINI: RHOMBORHININA.

Type species: Gnathocera amabilis Bainbridge, 1840, by subsequent designation (Sakai and Nagai 1998: 230).

Family-group name: TMESORRHINAE Schoch, 1894. Stem: Tmesorrhin-.

Tolyphus Erichson, 1845a: 108. Current status: valid genus in PHALACRIDAE: PHALACRINAE.

Type species: Phalacrus granulatus Guérin-Méneville, 1834, by monotypy.

Family-group name: TOLYPHINI Guillebeau, 1892. Stem: Tolyph-.

Tomicus Latreille, 1802b: 203. Current status: valid genus in CURCULIONIDAE: SCOLYTINAE: HYLURGINI. Note: name placed on the Official List of Generic Names in Zoology (ICZN 1963a).

Type species: Dermestes piniperda Linnaeus, 1758, by monotypy (see ICZN 1963a). Note: name placed on the Official List of Specific Names in Zoology (ICZN 1963a).

Family-group name: TOMICINI Wood, 1978. Stem: Tomic-.

Tomoderus LaFerté-Sénectère in Guérin-Méneville, 1849: [no 26] 1. Current status: valid genus in ANTHICIDAE: TOMODERINAE.

Type species: Tomoderus cruciatus LaFerté-Sénectère, 1849, by subsequent designation (Bonadona 1958: 28).

Family-group name: TOMODERINI Bonadona, 1961. Stem: Tomoder-.

Tomorhinus Jekel, 1856: 8. Current status: valid genus in CURCULIONIDAE: ENTIMINAE: ENTIMITAE: LORDOPINI.

Type species: Hypsonotus designatus Fåhraeus, 1842, by monotypy.

Family-group name: TOMORHINI Jekel, 1856. Stem: Tomorhin-.

Tomyris Chapuis, 1874: 265. Current status: invalid senior synonym of Eboo Reid, 1993 in CHRYSOMELIDAE: EUMOLPINAE: BROMIINI. Note: junior homonym of Tomyris Eichwald, 1831 [Reptilia].

Type species: Tomyris pulchella Chapuis, 1874, by monotypy.

Family-group name: TOMYRITES Chapuis, 1874. Stem: Tomyri-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

Tonerus K.B. Miller, 2009: 201. Current status: valid genus in NOTERIDAE: NOTERINAE: NOTERINI.

Type species: Tonerus wheeleri K.B. Miller, 2009, by original designation.

Family-group name: TONERINI K.B. Miller, 2009. Stem: Toner-.

Tonesia Casey, 1922: 337. Current status: valid genus in CURCULIONIDAE: BARIDINAE: EUTOXINI: TONESIINA.

Type species: Tonesia glabra Casey, 1922, by original designation.

Family-group name: TONESIINA Alonso-Zarazaga & Lyal, 1999. Stem: Tonesi-.

Tophoderes Dejean, 1834: 236. Current status: valid genus in ANTHRIBIDAE: ANTHRIBINAE: TOPHODERINI.

Type species: Anthribus frenatus Klug, 1833, by monotypy.

Family-group name: TOPHODÉRIDES Lacordaire, 1865. Stem: Tophoder-.

Toramus Grouvelle, 1916b: 26. Current status: valid genus in EROTYLIDAE: CRYPTOPHILINAE: TORAMINI. Note: new replacement name for Tomarus J.L. LeConte, 1861.

Type species: Tomarus pulchellus J.L. LeConte, 1863, by subsequent monotypy (J.L. LeConte 1863b: 71) (type fixation for Tomarus J.L. LeConte).

Family-group name: TORAMINAE Sen Gupta, 1967. Stem: Toram-.

Torcus Casey, 1922: 315. Current status: valid genus in CURCULIONIDAE: BARIDINAE: APOSTASIMERINI: TORCINA.

Type species: Torcus fasciatus Casey, 1922, by original designation.

Family-group name: TORCOCINA Bondar, 1943. Stem: Torc-.

Tormissus Broun, 1893b: 1021. Current status: valid genus in HYDROPHILIDAE: CYLOMINAE.

Type species: Tormissus magnulus Broun, 1893 (= Hydrostygnus linsi Sharp, 1884), by subsequent designation (Knisch 1924: 107).

Family-group name: TORMISSINI Hansen, 1991. Stem: Tormiss-.

Torneuma Wollaston, 1860: 453. Current status: valid genus in CURCULIONIDAE: CRYPTORHYNCHINAE: TORNEUMATINI.

Type species: Torneuma caecum Wollaston, 1860, by monotypy.

Family-group name: TORNEUMATINI Bedel, 1884. Stem: Torneumat-.

Torneutes Reich, 1838: 9. Current status: valid genus in CERAMBYCIDAE: CERAMBYCINAE: TORNEUTINI.

Type species: Torneutes pallidipennis Reich, 1838, by monotypy.

Family-group name: TORNEUTITAE J. Thomson, 1861. Stem: Torneut-.

Torrentomus Bierig, 1934: 213. Current status: junior synonym of Thinobius Kiesenwetter, 1844 in STAPHYLINIDAE: OXYTELINAE: THINOBIINI.

Type species: Torrentomus torrei Bierig, 1934, by original designation.

Family-group name: TORRENTOMI Bierig, 1934. Stem: Torrentom-.

Torridincola Steffan, 1964: 193. Current status: valid genus in TORRIDINCOLIDAE: TORRIDINCOLINAE.

Type species: Torridincola rhodesica Steffan, 1964, by original designation.

Family-group name: TORRIDINCOLIDAE Steffan, 1964. Stem: Torridincol-.

Tostegoptera Blanchard in H. Milne-Edwards et al., 1851: 149. Current status: valid genus in SCARABAEIDAE: MELOLONTHINAE: RHIZOTROGINI.

Type species: Melolontha lanceolata Say, 1824, by monotypy.

Family-group name: TOSTEGOPTERAE J.L. LeConte, 1861. Stem: Tostegopter-.

Toxicum Latreille, 1802b: 174. Current status: valid genus in TENEBRIONIDAE: TENEBRIONINAE: TOXICINI: TOXICINA.

Type species: Toxicum richesianum Latreille, 1802, by monotypy. Note: combined description of a new genus and a single new species (ICZN 1999a, Article 12.2.6).

Family-group name: TOXICIDEN Oken, 1843. Stem: Toxic-.

Toxidium J.L. LeConte, 1860b: 322, 324. Current status: valid genus in STAPHYLINIDAE: SCAPHIDIINAE: SCAPHISOMATINI.

Type species: Toxidium gammaroides J.L. LeConte, 1860, by monotypy.

Family-group name: TOXIDIINI Achard, 1924. Stem: Toxidi-.

Toxoderus Fauvel, 1900b: 189. Current status: junior synonym of Homalotrichus Solier, 1849 in STAPHYLINIDAE: OXYTELINAE: COPROPHILINI. Note: new replacement name for Sharpia Fauvel, 1878.

Type species: Sharpia banksi Fauvel, 1878, by monotypy (type fixation for Sharpia Fauvel).

Family-group name: TOXODERI Bernhauer & Schubert, 1911. Stem: Toxoder-. Note: junior homonym of TOXODERINI Saussure, 1869 (type genus Toxodera Audinet-Serville, 1837) in Mantodea.

Toxognathus Fairmaire, 1878: 271. Current status: valid genus in ELATERIDAE: PHYSODACTYLINAE.

Type species: Toxognathus costulatus Fairmaire, 1878, by monotypy.

Family-group name: TOXOGNATHINAE Fleutiaux, 1941. Stem: Toxognath-.

†Toxorhynchus Scudder, 1893: 26. Current status: valid genus in BRENTIDAE: APIONINAE: APIONITAE: APIONINI: TOXORHYNCHINA. Note: Oligocene (Colorado, USA).

Type species: †Toxorhynchus minusculus Scudder, 1893, by original designation.

Family-group name: †TOXORHYNCHINI Scudder, 1893. Stem: Toxorhynch-.

Toxoscelus Deyrolle, 1865: 127. Current status: valid genus in BUPRESTIDAE: AGRILINAE: CORAEBINI: TOXOSCELINA.

Type species: Toxoscelus undatus Deyrolle, 1865, by subsequent designation (Descarpentries and Villiers 1967: 490).

Family-group name: TOXOSCELINA Majer, 2000. Stem: Toxoscel-.

Toxotus Dejean, 1821: 112. Current status: junior synonym of Stenocorus Geoffroy, 1762 in CERAMBYCIDAE: LEPTURINAE: STENOCORINI.

Type species: Leptura meridiana Linnaeus, 1758, by subsequent designation (Westwood 1838c: 41).

Family-group name: TOXOTI J.L. LeConte & G.H. Horn, 1883. Stem: Toxot-. Note: junior homonym of TOXOTIDAE Günther, 1860 (type genus Toxotes Cloquet, 1816) in Pisces.

Trabisus Raffray, 1890a: 110. Current status: junior synonym of Atheropterus Raffray, 1882 in STAPHYLINIDAE: PSELAPHINAE: BATRISITAE: BATRISINI: BATRISINA.

Type species: Batrisus zanzibaricus Raffray, 1877, by subsequent designation (Jeannel 1949a: 117).

Family-group name: TRABISINA Jeannel, 1949. Stem: Trabis-.

Trachelizus Dejean, 1834: 243. Current status: valid genus in BRENTIDAE: BRENTINAE: TRACHELIZINI: TRACHELIZINA. Note: this name was first made available by Gyllenhal in Schönherr (1833: 334) but in a different sense; an application to the Commission in needed to suppress Trachelizus Gyllenhal, 1833 (see Alonso-Zarazaga and Lyal 1999: 56).

Type species: Brentus bisulcatus Lund, 1802, by subsequent designation (Schönherr 1840a: 490).

Family-group name: TRACHÉLIZIDES Lacordaire, 1865. Stem: Tracheliz-.

Trachelonia Gistel, 1850: 75. Current status: junior synonym of Collyris Fabricius, 1801 in CICINDELIDAE: COLLYRIDINI: COLLYRIDINA. Note: unnecessary new replacement name for Collyris Fabricius, 1801.

Type species: Cicindela longicollis Fabricius, 1787, by subsequent designation (Latreille 1810: 425) (type fixation for Collyris Fabricius).

Family-group name: TRACHELONIADAE Gistel, 1850. Stem: Tracheloni-.

Trachelophorus Jekel, 1860: 158. Current status: valid genus in ATTELABIDAE: APODERINAE: TRACHELOPHORINI.

Type species: Apoderus giraffa Jekel, 1860, by subsequent designation (Voss 1925: 5).

Family-group name: TRACHELOPHORINI Voss, 1926. Stem: Trachelophor-.

Trachelostenus Solier, 1851: 255. Current status: valid genus in TENEBRIONIDAE: TENEBRIONINAE: TRACHELOSTENINI.

Type species: Trachelostenus inaequalis Solier, 1851, by monotypy.

Family-group name: TRACHÉLOSTÉNIDES Lacordaire, 1859. Stem: Trachelosten-.

Trachodes Germar, 1823: 325. Current status: valid genus in CURCULIONIDAE: MOLYTINAE: TRACHODINI.

Type species: Curculio squamifer Paykull, 1800 (= Curculio hispidus Linnaeus, 1758), by subsequent designation (Schönherr 1825: 584).

Family-group name: TRACHODISIDAE Gistel, 1848. Stem: Trachod-.

Trachyderes Dalman in Schönherr, 1817: 364. Current status: valid genus in CERAMBYCIDAE: CERAMBYCINAE: TRACHYDERINI: TRACHYDERINA.

Type species: Cerambyx succinctus Linnaeus, 1758, by subsequent designation (Blanchard in Audouin et al. 1843: pl. 65).

Family-group name: TRACHYDÉRIDES Dupont, 1836. Stem: Trachyder-.

Trachykele Marseul, 1865: 149. Current status: valid genus in BUPRESTIDAE: BUPRESTINAE: BUPRESTINI: TRACHYKELINA.

Type species: Trachykele blondeli Marseul, 1865, by monotypy.

Family-group name: TRACHYKELINA Holyński, 1988. Stem: Trachykel-.

Trachynotus Latreille, 1828a: 579. Current status: valid genus in TENEBRIONIDAE: PIMELIINAE: SEPIDIINI: TRACHYNOTINA.

Type species: Sepidium vittatum Fabricius, 1781, by subsequent designation (Kamiński et al. 2019: 93).

Family-group name: TRACHYNOTINA C. Koch, 1955. Stem: Trachynot-. Note: junior homonym of TRACHYNOTIDAE Schrammen, 1924 (type genus Trachynotus Schrammen, 1924) in Porifera and TRACHYNOTIDAE Förster, 1869 (type genus Trachynotus Gravenhorst, 1829) in Hymenoptera; both Schrammen and Förster names are permanently invalid (based on a preoccupied type genera).

Trachypachus Motschulsky, 1844a: 86. Current status: valid genus in TRACHYPACHIDAE: TRACHYPACHINAE.

Type species: Blethisa zetterstedtii Gyllenhal, 1827, by subsequent designation (C.G. Thomson 1859: 3).

Family-group name: TRACHYPACHINI C.G. Thomson, 1857. Stem: Trachypach-.

Trachyphilus Faust, 1887: 164. Current status: valid genus in CURCULIONIDAE: ENTIMINAE: CYPHICERITAE: TRACHYPHLOEINI: TRACHYPHILINA.

Type species: Trachyphilus saluber Faust, 1887, by monotypy.

Family-group name: TRACHYPHILINA Voss, 1948. Stem: Trachyphil-.

Trachyphloeus Germar, 1817b: 341. Current status: valid genus in CURCULIONIDAE: ENTIMINAE: CYPHICERITAE: TRACHYPHLOEINI: TRACHYPHLOEINA.

Type species: Curculio scabriculus Linnaeus, 1771 [as scabricul], by monotypy.

Family-group name: TRACHYPHLOEIDAE Gistel, 1848. Stem: Trachyphloe-.

Trachypholis Erichson, 1845a: 257. Current status: valid genus in ZOPHERIDAE: COLYDIINAE: SYNCHITINI.

Type species: Opatrum hispidum Weber, 1801, by monotypy.

Family-group name: TRACHYPHOLINI Grouvelle, 1911. Stem: Trachypholid-.

Trachys Fabricius, 1801a: xviii. Current status: valid genus in BUPRESTIDAE: AGRILINAE: TRACHEINI: TRACHEINA. Note: name placed on the Official List of Generic Names in Zoology (ICZN 2009a).

Type species: Buprestis minuta Linnaeus, 1758, by subsequent designation (Westwood 1838c: 25; see ICZN 2009a). Note: no species were originally included in this genus by Fabricius (1801a); the first ones were the 11 species listed by Fabricius (1801b: 218– 220); name placed on the Official List of Specific Names in Zoology (ICZN 2009a).

Family-group name: TRACHISIDAE Laporte, 1835. Stem: Trache-.

Trachyscelis Latreille, 1809a: 379. Current status: valid genus in TENEBRIONIDAE: DIAPERINAE: TRACHYSCELINI.

Type species: Trachyscelis aphodioides Latreille, 1809, by monotypy.

Family-group name: TRACHYSCÉLIDES Blanchard, 1845. Stem: Trachyscel-.

Tragocephala Dejean, 1835: 342. Current status: valid genus in CERAMBYCIDAE: LAMIINAE: TRAGOCEPHALINI.

Type species: Lamia formosa G.-A. Olivier, 1792, by subsequent designation (Chevrolat 1849a: 632).

Family-group name: TRAGOCEPHALITES J. Thomson, 1857. Stem: Tragocephal-.

Tragocerus Latreille, 1829b: 121. Current status: valid genus in CERAMBYCIDAE: CERAMBYCINAE: TRAGOCERINI.

Type species: Prionus bidentatus Donovan, 1805, by subsequent monotypy (Guérin-Méneville 1831b: pl. 45).

Family-group name: TRAGOCERINAE Pascoe, 1867. Stem: Tragocer-.

Tragosoma Audinet-Serville, 1832: 159. Current status: valid genus in CERAMBYCIDAE: PRIONINAE: MEROSCELISINI.

Type species: Cerambyx depsarius Linnaeus, 1767, by monotypy.

Family-group name: TRAGOSOMITAE J. Thomson, 1864. Stem: Tragosomat-.

Tranes Schönherr, 1843: 129. Current status: valid genus in CURCULIONIDAE: MOLYTINAE: TRANESINI.

Type species: Tranes vigorsii Boheman, 1843, by original designation.

Family-group name: TRANESINI Legalov, 2018d: 345. Stem: Tranes-.

Trechicus J.L. LeConte, 1853e: 384, 386. Current status: valid subgenus of Perigona Laporte, 1834 in CARABIDAE: HARPALINAE: PERIGONINI.

Type species: Trechicus umbripennis J.L. LeConte, 1853 (= Bembidium nigriceps Dejean, 1831), by subsequent designation (Jeannel 1926: 247).

Family-group name: TRECHICHINAE H.W. Bates, 1873. Stem: Trechic-. Note: nomen oblitum (see Bouchard et al. 2011: 140).

Trechodes Blackburn, 1901: 119. Current status: valid genus in CARABIDAE: TRECHINAE: TRECHINI: TRECHODINA.

Type species: Bembidium secalioides Blackburn, 1891, by subsequent designation (Jeannel 1926: 484).

Family-group name: TRECHODINI Jeannel, 1926. Stem: Trechod-. Note: the recent placement of Perileptus Schaum, 1860 within the valid subtribe TRECHODINA by Maddison et al. (2019: 173) creates a priority problem as PERILEPTINA Sloane, 1903 is the oldest available family-group name for this subtribe.

Trechus Clairville, 1806: 22. Current status: valid genus in CARABIDAE: TRECHINAE: TRECHINI: TRECHINA.

Type species [suggested]: Carabus quadristriatus Schrank, 1781, misidentified as Carabus rubens Fabricius, 1792 (sensu Clairville, 1806) in the subsequent designation by Blanchard in Audouin et al. (1841: pl. 25). Note: the valid type species fixation is that of Latreille (1810: 426), who selected Carabus meridianus Linnaeus, 1761, but this is currently the type species of the genus Acupalpus Latreille, 1829; a new application to the Commission is necessary to fix Carabus quadristriatus Schrank, 1781 as the type species of Trechus Clairville, 1806 since a previous request was postponed (ICZN 1950: 447).

Family-group name: TRECHII Bonelli, 1810. Stem: Trech-.

Tretothorax Lea, 1910: 211. Current status: valid genus in SALPINGIDAE: DACODERINAE.

Type species: Tretothorax cleistostoma Lea, 1910, by monotypy.

Family-group name: TRETOTHORACIDAE Lea, 1910. Stem: Tretothorac-.

Triacrus Nordmann, 1837: 5, 19. Current status: valid genus in STAPHYLINIDAE: STAPHYLININAE: STAPHYLININI: XANTHOPYGINA.

Type species: Triacrus dilatus Nordmann, 1837, by monotypy.

Family-group name: TRIACRI Bernhauer, 1931. Stem: Triacr-.

†Triadocupes Ponomarenko, 1966: 51. Current status: valid genus in †TRIADOCUPEDIDAE. Note: Upper Triassic (Kyrgyzstan).

Type species: †Triadocupes ferghanensis Ponomarenko, 1966, by original designation.

Family-group name: †TRIADOCUPEDINAE Ponomarenko, 1966. Stem: Triadocuped-.

†Triamyxa Qvarnström, Fikáček, Wernström, Huld, Beutel, Arriaga-Varela, Ahlberg & Niedźwiedzki, 2021: 3378. Current status: valid genus in HYDROSCAPHIDAE: †TRIAMYXINAE. Note: Triassic (Poland).

Type species: †Triamyxa coprolithica Qvarnström et al., 2021, by original designation.

Family-group name: †TRIAMYXIDAE Qvarnström, Fikáček, Wernström, Huld, Beutel, Arriaga-Varela, Ahlberg & Niedźwiedzki, 2021: 3376. Stem: Triamyx-.

†Triaplus Ponomarenko, 1977: 17. Current status: valid genus in †TRIAPLIDAE. Note: Permian/Triassic (Kyrgyzstan; Russia).

Type species: †Triaplus macroplatus Ponomarenko, 1977, by original designation.

Family-group name: †TRIAPLIDAE Ponomarenko, 1977. Stem: Triapl-.

Triarthron Märkel, 1840: 141. Current status: valid genus in LEIODIDAE: LEIODINAE: SOGDINI.

Type species: Triarthron maerkelii W.L.E. Schmidt, 1840, by monotypy.

Family-group name: TRIARTHRINI Jeannel, 1962. Stem: Triarthr-. Note: junior homonym of TRIARTHRIDAE Ulrich, 1930 (type genus Triarthrus Green, 1832) in Trilobita.

Tribalus Erichson, 1834: 164. Current status: valid genus in HISTERIDAE: TRIBALINAE.

Type species: Hister capensis Paykull, 1811, by subsequent designation (Hope 1841a: 105).

Family-group name: TRIBALINI Bickhardt, 1914. Stem: Tribal-.

Tribax Fischer, 1817: 463. Current status: valid subgenus of Carabus Linnaeus, 1758 in CARABIDAE: CARABINAE: CARABINI: CARABINA. Note: this name was originally proposed as Tribacis, which is the genitive of Tribax; according to Article 11.8.1 (ICZN 1999a) the name, if it meets the other requirements of availability, is available but must corrected to the nominative singular.

Type species: Carabus puschkini Adams, 1817, by subsequent designation (Breuning 1934: 1033).

Family-group name: TRIBACOGENICI Csiki, 1927. Stem: Tribac-.

Tribolium W.S. MacLeay, 1825: 47. Current status: valid genus in TENEBRIONIDAE: TENEBRIONINAE: TRIBOLIINI. Note: name placed on the Official List of Generic Names in Zoology (ICZN 1988e).

Type species: Colydium castaneum Herbst, 1797, by monotypy (see ICZN 1988e). Note: name placed on the Official List of Specific Names in Zoology (ICZN 1988e).

Family-group name: TRIBOLIIDAE Gistel, 1848. Stem: Triboli-.

Trichalus C.O. Waterhouse, 1877: 82. Current status: valid genus in LYCIDAE: METRIORRHYNCHINAE: METRIORRHYNCHINI: METRIORRHYNCHINA.

Type species: Trichalus flavopictus C.O. Waterhouse, 1877, by subsequent designation (C.O. Waterhouse 1878: 103).

Family-group name: TRICHALINAE Kleine, 1929. Stem: Trichal-.

Trichapion Wagner, 1912a: 99, 116. Current status: valid genus in BRENTIDAE: APIONINAE: APIONITAE: APIONINI: SYNAPIINA.

Type species: Apion aurichalceum Wagner, 1912, by subsequent designation (Kissinger 1959: 248).

Family-group name: TRICHAPIINA Alonso-Zarazaga, 1990. Stem: Trichapi-.

Trichelodes Carter, 1935: 189. Current status: valid genus in DERMESTIDAE: TRINODINAE: TRICHELODINI.

Type species: Trichelodes delicatula Carter, 1935, by monotypy.

Family-group name: TRICHELODINI Peacock, 1978. Stem: Trichelod-.

Trichinorhipis Barr, 1948: 70. Current status: valid genus in BUPRESTIDAE: BUPRESTINAE: XENORHIPIDINI: TRICHINORHIPIDINA.

Type species: Trichinorhipis knulli Barr, 1948, by original designation.

Family-group name: TRICHINORHIPIDINA Bellamy, 2006. Stem: Trichinorhipid-.

Trichis Klug, 1832: pl. xxi. Current status: valid genus in CARABIDAE: HARPALINAE: LEBIINI: TRICHINA.

Type species: Trichis pallida Klug, 1832, by subsequent designation (Andrewes 1938: 138).

Family-group name: TRICHINI Basilewsky, 1984. Stem: Trich-.

Trichius Fabricius, 1775: 40. Current status: valid genus in SCARABAEIDAE: CETONIINAE: TRICHIINI. Note: name placed on the Official List of Generic Names in Zoology (ICZN 2004d).

Type species: Scarabaeus fasciatus Linnaeus, 1758, by subsequent designation (Latreille 1810: 428; see ICZN 2004d). Note: name placed on the Official List of Specific Names in Zoology (ICZN 2004d).

Family-group name: TRICHIADAE Fleming, 1821. Stem: Trichi-.

Trichochrysea Baly, 1861: 195. Current status: valid genus in CHRYSOMELIDAE: EUMOLPINAE: BROMIINI.

Type species: Trichochrysea mouhoti Baly, 1861, by original designation.

Family-group name: TRICHOCHRYSEINI Clavareau, 1914. Stem: Trichochryse-.

Trichodes Herbst, 1792: 154. Current status: valid genus in CLERIDAE: CLERINAE: OPILONITAE: DIEROPSEINI. Note: name placed on the Official List of Generic Names in Zoology (ICZN 1984a).

Type species: Attelabus apiarius Linnaeus, 1758, by subsequent designation (Hope 1841a: 137; see ICZN 1984a). Note: name placed on the Official List of Specific Names in Zoology (ICZN 1984a).

Family-group name: TRICHODIDAE Streubel, 1839. Stem: Trichod-. Note: as pointed out by Bouchard and Bousquet (2020a: 92), TRICHODIDAE Streubel, 1839 is an available family- group name and has priority over DIEROPSEINI J.R. Winkler, 1964 based on the recent classification of Bartlett (2021), an application to the Commission is needed to conserve DIEROPSEINI J.R. Winkler, 1964 as the valid name for this tribe; TRICHODINI Streubel, 1839 is a senior homonym of TRICHODINA van der Hoeven, 1849 (type genus Trichoda O.F. Müller, 1773) in Protozoa: Ciliophora.

Trichodocerus Chevrolat, 1879: 123. Current status: valid genus in CURCULIONIDAE: CONODERINAE: TRICHODOCERINI.

Type species: Trichodocerus spinolae Chevrolat, 1879, by subsequent designation (Champion 1906: 713).

Family-group name: TRICHODOCERIDES Champion, 1906. Stem: Trichodocer-.

Trichomesia Pascoe, 1859a: 18. Current status: valid genus in CERAMBYCIDAE: CERAMBYCINAE: TRICHOMESIINI.

Type species: Trichomesia newmani Pascoe, 1859, by monotypy.

Family-group name: TRICHOMESIINI Aurivillius, 1912. Stem: Trichomesi-.

Trichonyx Chaudoir, 1845: 164. Current status: valid genus in STAPHYLINIDAE: PSELAPHINAE: EUPLECTITAE: TRICHONYCHINI: TRICHONYCHINA.

Type species: Pselaphus sulcicollis H.G.L. Reichenbach, 1816, by monotypy.

Family-group name: TRICHONYIDES Reitter, 1882. Stem: Trichonych-.

Trichophya Mannerheim, 1830: 73. Current status: valid genus in STAPHYLINIDAE: TRICHOPHYINAE.

Type species: Aleochara pilicornis Gyllenhal, 1810, by monotypy.

Family-group name: TRICHOPHYINI C.G. Thomson, 1858. Stem: Trichophy-.

Trichoplus Burmeister, 1842: 660. Current status: valid genus in SCARABAEIDAE: CETONIINAE: CREMASTOCHEILINI: TRICHOPLINA.

Type species: Cremastocheilus laevis Gory & Percheron, 1833, by monotypy.

Family-group name: TRICHOPLINA Krikken, 1984. Stem: Trichopl-.

Trichopselaphus Chaudoir, 1843: 399. Current status: valid genus in CARABIDAE: HARPALINAE: HARPALINI: HARPALINA.

Type species: Trichopselaphus subiridescens Chaudoir, 1843, by monotypy.

Family-group name: TRICHOPSELAPHINI Tschitschérine, 1900. Stem: Trichopselaph-.

Trichopsenius G.H. Horn, 1877: 88. Current status: valid genus in STAPHYLINIDAE: ALEOCHARINAE: TRICHOPSENIINI.

Type species: Hypocyptus depressus J.L. LeConte, 1863, by monotypy.

Family-group name: TRICHOPSENII J.L. LeConte & G.H. Horn, 1883. Stem: Trichopseni-.

Trichopteryx W. Kirby in W. Kirby and Spence, 1826: 41. Current status: invalid senior synonym of Acrotrichis Motschulsky, 1848 in PTILIIDAE: PTILIINAE: ACROTRICHINI. Note: junior homonym of Trichopteryx Hübner, 1825 [Lepidoptera].

Type species: Dermestes atomarius DeGeer, 1774, by monotypy.

Family-group name: TRICHOPTERYGIA Erichson, 1845. Stem: Trichopteryg-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

Trichotichnus Morawitz, 1863: 63. Current status: valid genus in CARABIDAE: HARPALINAE: HARPALINI: HARPALINA. Note: nomen protectum (see Bousquet 2008b: 328).

Type species: Trichoptichnus longitarsis Morawitz, 1863, by monotypy.

Family-group name: TRICHOTICHNINI Jeannel, 1942. Stem: Trichotichn-.

†Tricoleus Ponomarenko, 1969a: 139. Current status: valid genus in †TRICOLEIDAE. Note: Jurassic (Kazakhstan).

Type species: †Tricoleus punctatus Ponomarenko, 1969, by original designation.

Family-group name: †TRICOLEIDAE Ponomarenko, 1969. Stem: Tricole-.

Tricondyla Latreille in Latreille and Dejean, 1822: 65. Current status: valid genus in CICINDELIDAE: COLLYRIDINI: TRICONDYLINA.

Type species: Cicindela aptera G.-A. Olivier, 1791, by monotypy.

Family-group name: TRICONDYLINA Naviaux, 1991. Stem: Tricondyl-.

Tricorynus G.R. Waterhouse, 1849: lxviii. Current status: valid genus in PTINIDAE: MESOCOELOPODINAE: TRICORYNINI.

Type species: Tricorynus zeae G.R. Waterhouse, 1849, by monotypy.

Family-group name: TRICORYNINAE R.E. White, 1971. Stem: Tricoryn-.

Tricrania J.L. LeConte, 1860a: 320. Current status: valid genus in MELOIDAE: NEMOGNATHINAE: NEMOGNATHINI: NEMOGNATHINA.

Type species: Horia sanguinipennis Say, 1824, by subsequent designation (Wellman 1910b: 219).

Family-group name: TRICRANIIDES Wellman, 1910. Stem: Tricrani-.

Trictenotoma G.R. Gray in Griffith and Pidgeon, 1831: 534, pl. 5. Current status: valid genus in TRICTENOTOMIDAE.

Type species: Trictenotoma childreni G.R. Gray, 1831, by monotypy.

Family-group name: TRICTÉNOTOMIDES Blanchard, 1845. Stem: Trictenotom-.

Trientoma Solier, 1835c: 253, 256. Current status: valid genus in TENEBRIONIDAE: PIMELIINAE: EDROTINI.

Type species: Trientoma varvasi Solier, 1835, by monotypy.

Family-group name: TRIENTOMINI Casey, 1907. Stem: Trientom-.

Trigonarthron Boppe, 1913: 403. Current status: valid genus in CERAMBYCIDAE: CERAMBYCINAE: TRIGONARTHRINI.

Type species: Trigonarthron cinnabarinum Boppe, 1913, by monotypy.

Family-group name: TRIGONARTHRINI Villiers, 1984. Stem: Trigonarthr-.

Trigonobregma Scheerpeltz, 1944: 170. Current status: valid genus in STAPHYLINIDAE: OXYTELINAE incertae sedis.

Type species: Thinobius ocularis Fauvel, 1902, by monotypy.

Family-group name: TRIGONOBREGMINI Scheerpeltz, 1944. Stem: Trigonobregmat-.

Trigonocolus Lacordaire, 1863: 593. Current status: valid genus in CURCULIONIDAE: MOLYTINAE: TRIGONOCOLINI. Note: new replacement name for Megarhinus Schönherr, 1835.

Type species: Megarhinus firmus Gyllenhal, 1835, by original designation (type fixation for Megarhinus Schönherr).

Family-group name: TRIGONOCOLIDES Lacordaire, 1863. Stem: Trigonocol-.

Trigonodactyla Dejean, 1831: 278, 288. Current status: junior synonym of Hexagonia W. Kirby, 1825 in CARABIDAE: HARPALINAE: HEXAGONIINI.

Type species: Trigonodactyla terminata Dejean, 1831 (= Hexagonia terminalis Gemminger & Harold, 1868), by subsequent designation (Hope 1838a: 104).

Family-group name: TRIGONODACTYLIENS Brullé, 1834. Stem: Trigonodactyl-.

Trigonogenium Harold, 1869b: 123. Current status: valid genus in BUPRESTIDAE: BUPRESTINAE: TRIGONOGENIINI. Note: new replacement name for Trigonophorus Solier, 1849.

Type species: Trigonophorus angulosus Solier, 1849, by monotypy (type fixation for Trigonophorus Solier).

Family-group name: TRIGONOGENIINI Cobos, 1956. Stem: Trigonogeni-.

Trigonognatha Motschulsky, 1857a: 25. Current status: valid subgenus of Myas Sturm, 1826 in CARABIDAE: HARPALINAE: PTEROSTICHINI: PTEROSTICHINA.

Type species: Trigonognatha cuprescens Motschulsky, 1857, by monotypy.

Family-group name: TRIGONOGNATHIDES Tschitschérine, 1898. Stem: Trigonognath-.

Trigonoptera Perroud, 1855a: 336. Current status: valid genus in CERAMBYCIDAE: LAMIINAE: TMESISTERNINI.

Type species: Trigonoptera maculata Perroud, 1855, by monotypy.

Family-group name: TRIGONOPTERINI Aurivillius, 1922. Stem: Trigonopter-.

Trigonopus Mulsant & Rey, 1853d: 105. Current status: valid genus in TENEBRIONIDAE: BLAPTINAE: PLATYNOTINI: PLATYNOTINA.

Type species: Trigonopus capicola Mulsant & Rey, 1853, by subsequent designation (Cazurro Ruiz 1897b: 539).

Family-group name: TRIGONOPAIRES Mulsant & Rey, 1853. Stem: Trigonopod-.

Trigonorhinus Wollaston, 1861a: 102. Current status: valid genus in ANTHRIBIDAE: ANTHRIBINAE: ANTHRIBINI.

Type species: Trigonorhinus pardalis Wollaston, 1861 (= Brachytarsus areolatus Boheman, 1845), by monotypy.

Family-group name: TRIGONORHININI B.D. Valentine, 1999. Stem: Trigonorhin-.

Trigonoscuta Motschulsky, 1853b: 79. Current status: valid genus in CURCULIONIDAE: ENTIMINAE: POLYDRUSITAE: PSALLIDIINI.

Type species: Trigonoscuta pilosa Motschulsky, 1853, by monotypy.

Family-group name: TRIGONOSCUTAE J.L. LeConte, 1874. Stem: Trigonoscut-.

Trigonostomus Brenske, 1893: 1. Current status: valid genus in SCARABAEIDAE: RUTELINAE: ADORETINI: TRIGONOSTOMUSINA. Note: this name was deemed available from Brenske, 1893, not invalid by reason of being an incorrect subsequent spelling of Trigonostomum Burmeister, 1844 and placed on the Official List of Generic Names in Zoology (ICZN 2009b).

Type species: Trigonostomus mucoreum Burmeister, 1844, by plenary power (see ICZN 2009b). Note: name placed on the Official List of Specific Names in Zoology (ICZN 2009b).

Family-group name: TRIGONOSTOMINA Ohaus, 1912. Stem: Trigonostomus- (ICZN 2009b).

Trigonotoma Dejean, 1828: 4, 182. Current status: valid genus in CARABIDAE: HARPALINAE: PTEROSTICHINI: PTEROSTICHINA.

Type species: here fixed (under ICZN 1999a, Article 70.3.2) as Trigonotoma indica Brullé, 1834, misidentified as Omaseus viridicollis W.S. MacLeay, 1825 (sensu Dejean, 1825) in the subsequent designation by Hope (1838a: 111).

Family-group name: TROGONOTOMIDAE Laporte, 1834. Stem: Trigonotom-.

Trigonurus Mulsant, 1848: 515. Current status: valid genus in STAPHYLINIDAE: TRIGONURINAE.

Type species: Trigonurus mellyi Mulsant, 1848, by monotypy.

Family-group name: TRIGONURIDES Reiche, 1866. Stem: Trigonur-.

Trilobitideus Raffray, 1898b: 351. Current status: valid genus in STAPHYLINIDAE: ALEOCHARINAE: TRILOBITIDEINI.

Type species: Trilobitideus mirabilis Raffray, 1898, by monotypy.

Family-group name: TRILOBITIDEIDAE Fauvel, 1899. Stem: Trilobitide-.

Trilobocara Solier, 1851: 129. Current status: valid genus in TENEBRIONIDAE: PIMELIINAE: TRILOBOCARINI.

Type species: Trilobocara ciliatum Solier, 1851, by original designation.

Family-group name: TRIBOLOCARIDES Lacordaire, 1859. Stem: Trilobocar-.

Trimerus Chaudoir, 1878b: 89. Current status: invalid senior synonym of Amorphomerus Sloane, 1923 in CARABIDAE: HARPALINAE: AMORPHOMERINI. Note: junior homonym of Trimerus Green, 1832 [Trilobita].

Type species: Trimerus raffrayi Chaudoir, 1878, by monotypy.

Family-group name: TRIMERINAE Alluaud, 1922. Stem: Trimer-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

Trimiodytes Raffray, 1897b: 52. Current status: valid genus in STAPHYLINIDAE: PSELAPHINAE: EUPLECTITAE: TRICHONYCHINI: TRICHONYCHINA.

Type species: Trimiodytes palustris Raffray, 1897, by monotypy.

Family-group name: TRIMIODYTINA Jeannel, 1964. Stem: Trimiodyt-.

Trimium Aubé, 1833: 44. Current status: valid genus in STAPHYLINIDAE: PSELAPHINAE: EUPLECTITAE: TRICHONYCHINI: TRIMIINA.

Type species: Pselaphus brevicornis H.G.L. Reichenbach, 1816, by monotypy.

Family-group name: TRIMIINI Brendel & Wickham, 1890. Stem: Trimi-.

Trimytis J.L. Leconte, 1851a: 141. Current status: valid genus in TENEBRIONIDAE: PIMELIINAE: EDROTINI.

Type species: Trimytis pruinosa J.L. LeConte, 1851, by monotypy.

Family-group name: TRIMYTINI Casey, 1907. Stem: Trimytid-.

Trinodes Dejean, 1821: 47. Current status: valid genus in DERMESTIDAE: TRINODINAE: TRINODINI.

Type species: Nitidula hirta Fabricius, 1781, by monotypy.

Family-group name: TRINODINI Casey, 1900. Stem: Trinod-.

Trinoparvus Háva, 2004: 152, 155. Current status: valid genus in DERMESTIDAE: TRINODINAE: TRINODINI.

Type species: Trinoparvus villosus Háva, 2004, by original designation.

Family-group name: TRINOPARVINI Háva, 2010. Stem: Trinoparv-.

Triodonyx Saylor, 1942: 158. Current status: valid genus in SCARABAEIDAE: MELOLONTHINAE: RHIZOTROGINI.

Type species: Phyllophaga gigantissima Saylor, 1935, by original designation.

Family-group name: TRIODONINA Rivera-Gasperín & Morón, 2013: 816. Stem: Triodon-.

Triorophus J.L. LeConte, 1851a: 141. Current status: valid genus in TENEBRIONIDAE: PIMELIINAE: EDROTINI.

Type species: Triorophus laevis J.L. LeConte, 1851, by subsequent designation (Casey 1907: 432).

Family-group name: TRIOROPHI J.L. LeConte & G.H. Horn, 1883. Stem: Trioroph-.

Triphyllus Dejean, 1821: 102. Current status: valid genus in MYCETOPHAGIDAE: MYCETOPHAGINAE: MYCETOPHAGINI.

Type species: Mycetophagus punctatus Fabricius, 1792 (= Nitidula bicolor Fabricius, 1777), by subsequent designation (Westwood 1838c: 14).

Family-group name: TRIPHYLLIDAE Crotch, 1873. Stem: Triphyll-.

Triplax Herbst, 1793: 146. Current status: valid genus in EROTYLIDAE: EROTYLINAE: TRITOMINI.

Type species: Silpha russica Linnaeus, 1758, by subsequent designation (Curtis 1838a: pl. 706).

Family-group name: TRIPLACINAE Streubel, 1839. Stem: Triplac-.

Trisignis Park & Schuster, 1955: 1. Current status: valid genus in STAPHYLINIDAE: PSELAPHINAE: EUPLECTITAE: TRICHONYCHINI: PANAPHANTINA.

Type species: Trisignis marshi Park & Schuster, 1955, by original designation.

Family-group name: TRISIGNINA Park & Schuster, 1955. Stem: Trisign-.

Tristaria Reitter, 1878a: 320. Current status: valid genus in BOSTRICHIDAE: LYCTINAE: TROGOXYLINI.

Type species: Tristaria grouvellei Reitter, 1878, by subsequent designation (Lesne 1911: 206).

Family-group name: TRISTARIINI Lesne, 1921. Stem: Tristari-.

†Tritarsus Hong, 2002: 105. Current status: valid genus in †TRITARSUSIDAE. Note: Eocene (China).

Type species: †Tritarsus latus Hong, 2002, by original designation.

Family-group name: †TRITARSUSIDAE Hong, 2002. Stem: Tritarsus- (ICZN 1999a, Article 29.4).

Tritoma Fabricius, 1775: 68. Current status: valid genus in EROTYLIDAE: EROTYLINAE: TRITOMINI. Note: the senior homonym Tritoma Geoffroy, 1762 (and all other uses of the name prior to the publication of Tritoma Fabricius, 1775) was suppressed for both Principles of Priority and Homonymy and placed on the Official Index of Rejected and Invalid Generic Names in Zoology while Tritoma Fabricius, 1775 was placed on the Official List of Generic Names in Zoology (ICZN 1994a).

Type species: Tritoma bipustulata Fabricius, 1775, by subsequent designation (Latreille 1810: 432; see ICZN 1994a). Note: name placed on the Official List of Specific Names in Zoology (ICZN 1994a).

Family-group name: TRITOMIDAE Curtis, 1834. Stem: Tritom-.

Tritoma Geoffroy, 1762: 335. Current status: invalid senior synonym of Mycetophagus Fabricius, 1792 in MYCETOPHAGIDAE: MYCETOPHAGINAE: MYCETOPHAGINI. Note: name suppressed for both Principles of Priority and Homonymy and placed on the Official Index of Rejected and Invalid Names in Zoology (see ICZN 1994a).

Type species: to be determined.

Family-group name: TRITOMIDAE Crotch, 1873. Stem: Tritom-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

Trixagus Kugelann, 1794: 534. Current status: valid genus in THROSCIDAE.

Type species: Dermestes adstrictor Herbst, 1792 (= Elater dermestoides Linnaeus, 1767), by subsequent designation (Barber 1942: 8, 32).

Family-group name: TRIXAGIDAE Gistel, 1848. Stem: Trixag-.

Trochalus Laporte, 1832c: pl. 44. Current status: valid genus in SCARABAEIDAE: SERICINAE: SERICINI: TROCHALINA.

Type species: Trochalus rotundatus Laporte, 1832, by monotypy.

Family-group name: TROCHALINAE Brenske, 1898. Stem: Trochal-.

Trochoideus Westwood, 1833: 673. Current status: valid genus in ENDOMYCHIDAE: PLEGANOPHORINAE.

Type species: Paussus cruciatus Dalman, 1826, by monotypy.

Family-group name: TROCHOIDÉITES Chapuis, 1876. Stem: Trochoide-.

Trogaster Sharp, 1874b: 83. Current status: valid genus in STAPHYLINIDAE: PSELAPHINAE: EUPLECTITAE: TROGASTRINI: TROGASTRINA.

Type species: Trogaster aberrans Sharp, 1874, by monotypy.

Family-group name: TROGASTERINI Brendel & Wickham, 1890. Stem: Trogastr-.

Trogloderus J.L. LeConte, 1879: 2. Current status: valid genus in TENEBRIONIDAE: BLAPTINAE: AMPHIDORINI.

Type species: Trogloderus costatus J.L. LeConte, 1879, by monotypy.

Family-group name: TROGLODERINA La Rivers, 1948. Stem: Trogloder-.

Troglops Erichson, 1840b: 125. Current status: valid genus in MELYRIDAE: MALACHIINAE: MALACHIINI.

Type species: Cantharis albicans Linnaeus, 1767, by subsequent designation (Chevrolat 1849a: 700).

Family-group name: TROGLOPATES Mulsant & Rey, 1867. Stem: Troglop-.

Trogocryptus Sharp, 1900: 602. Current status: valid genus in SALPINGIDAE: PROSTOMINIINAE.

Type species: Trogocryptus nigripectus Sharp, 1900, by monotypy.

Family-group name: TROGOCRYPTINAE Lawrence, 1991. Stem: Trogocrypt-.

Trogoderma Dejean, 1821: 46. Current status: valid genus in DERMESTIDAE: MEGATOMINAE: MEGATOMINI: TROGODERMINA.

Type species: Anthrenus elongatulus Fabricius, 1801 (= Anthrenus glaber Herbst, 1783), by subsequent designation (Hope 1841a: 143).

Family-group name: TROGODERMATES Mulsant & Rey, 1867. Stem: Trogoderm-. Note: this name was treated as unavailable by Bouchard et al. (2011: 335); however, it has been used in latinized form and generally accepted since (e.g., Háva 2015: xxii, Zhou et al. 2022: 18).

Trogodes Boheman, 1857: 54. Current status: valid genus in SCARABAEIDAE: CETONIINAE: CREMASTOCHEILINI: TROGODINA. Note: several authors credit the authorship of this name to Westwood since the name Trogodes in the original description is followed by “(Westw.)”; however, the signature at the end of the combined description of the genus and species is “BHN” [Boheman] and we believe that Boheman should be credited with both the genus-group and species-group names (see Bouchard and Bousquet 2020a: 83, incorrectly credited to “Boheman, 1851” instead of “Boheman, 1857”).

Type species: Trogodes rotundicollis Boheman, 1857, by monotypy.

Family-group name: TROGODINA Krikken, 1984. Stem: Trogod-.

Trogoparvus Háva, 2002: 27. Current status: valid genus in DERMESTIDAE: TROGOPARVINAE.

Type species: Trogoparvus sumatrensis Háva, 2002, by original designation.

Family-group name: TROGOPARVINAE Zhou, Nicholls, Liu, Hartley, Szito, Ślipiński & Zwick, 2022: 14. Stem: Trogoparv-.

Trogophloeus Mannerheim, 1830: 49. Current status: valid subgenus of Carpelimus Leach, 1819 in STAPHYLINIDAE: OXYTELINAE: OXYTELINI.

Type species: Oxytelus corticinus Gravenhorst, 1806, by monotypy.

Family-group name: TROGOPHLÉAIRES Mulsant & Rey, 1878. Stem: Trogophloe-.

Trogossita G.-A. Olivier, 1790a: [19] 5. Current status: junior synonym of Tenebroides Piller & Mitterpacher, 1783 in TROGOSSITIDAE: TROGOSSITINAE: TROGOSSITINI.

Type species: Tenebrio mauritanicus Linnaeus, 1758, by subsequent designation (Partington 1837: 816).

Family-group name: TROGOSSITARII Latreille, 1802. Stem: Trogossit-.

Trogoxylon J.L. LeConte, 1862: 209. Current status: valid genus in BOSTRICHIDAE: LYCTINAE: TROGOXYLINI.

Type species: Xylotrogus parallelipipedus Melsheimer, 1844, by original designation.

Family-group name: TROGOXYLINI Lesne, 1921. Stem: Trogoxyl-.

Tropicus M.F. Pacheco, 1964: 102. Current status: valid genus in HETEROCERIDAE: HETEROCERINAE: TROPICINI.

Type species: Heterocerus pusillus Say, 1823, by original designation.

Family-group name: TROPICINI M.F. Pacheco, 1964. Stem: Tropic-.

Tropideres Schönherr, 1823: 1135. Current status: valid genus in ANTHRIBIDAE: ANTHRIBINAE: TROPIDERINI.

Type species: Curculio albirostris Schaller, 1783, by original designation.

Family-group name: TROPIDÉRIDES Lacordaire, 1865. Stem: Tropider-.

Tropidion J. Thomson, 1867: 138. Current status: valid genus in CERAMBYCIDAE: CERAMBYCINAE: TROPIDINI: TROPIDINA.

Type species: Ibidion flavipes J. Thomson, 1867, by subsequent designation (Martins 1968: 355).

Family-group name: TROPIDINA Martins & Galileo, 2007. Stem: Tropid-.

Tropidosoma Perty, 1830: 15. Current status: junior synonym of Allocerus Lacordaire, 1830 in CERAMBYCIDAE: CERAMBYCINAE: TRACHYDERINI: TRACHYDERINA.

Type species: Prionus spencei W. Kirby, 1818, by monotypy.

Family-group name: TROPIDOSOMITAE J. Thomson, 1864. Stem: Tropidosomat-.

Tropiphorus Schönherr, 1842a: 257. Current status: valid genus in CURCULIONIDAE: ENTIMINAE: ENTIMITAE: BYRSOPAGINI: TROPIPHORINA. Note: name placed on the Official List of Generic Names in Zoology (ICZN 1988c).

Type species: Curculio mercurialis Fabricius, 1801 (= Curculio elevatus Herbst, 1795), by original designation (see ICZN 1988c). Note: Curculio mercurialis Fabricius, 1801 was placed on the Official List of Specific Names in Zoology (ICZN 1988c).

Family-group name: TROPIPHORIDAE Marseul, 1863. Stem: Tropiphor-.

Tropocalymma J. Thomson, 1864: 138. Current status: valid genus in CERAMBYCIDAE: CERAMBYCINAE: TROPOCALYMMATINI.

Type species: Tropis dimidiata Newman, 1841, by original designation.

Family-group name: TROPOCALYMMIDES Lacordaire, 1868. Stem: Tropocalymmat-.

Tropopterus Solier, 1849: 211. Current status: valid genus in CARABIDAE: PSYDRINAE: MORIOMORPHINI: TROPOPTERINA.

Type species: Tropopterus giraudyi Solier, 1849, by subsequent designation (Cazurro Ruiz 1897b: 641).

Family-group name: TROPOPTERIDES Sloane, 1898. Stem: Tropopter-.

Trotommidea Reitter, 1883b: 307. Current status: valid genus in SCRAPTIIDAE: SCRAPTIINAE: SCRAPTIINI.

Type species: Trotommidea salonae Reitter, 1883, by monotypy.

Family-group name: TROTOMMIDEINI Pic, 1903. Stem: Trotommide-.

Trox Fabricius, 1775: 31. Current status: valid genus in TROGIDAE: TROGINAE.

Type species: Scarabaeus sabulosus Linnaeus, 1758, by subsequent designation (Latreille 1810: 428).

Family-group name: TROGIDAE W.S. MacLeay, 1819. Stem: Trog-.

Trypanaeus Eschscholtz, 1829b: 11. Current status: valid genus in HISTERIDAE: ABRAEINAE: TRYPANAEINI.

Type species [suggested]: Bostrichus thoracicus Fabricius, 1801, by subsequent designation (Bickhardt 1916: 43). Note: the valid type species fixation is that of Hope (1841a: 105), who selected Bostrichus proboscideus Fabricius, 1801, but that species is the type species of the genus Coptotrophis Lewis, 1902; an application to the Commission is necessary to fix Bostrichus thoracicus Fabricius, 1801 as the type species of Trypanaeus Eschscholtz, 1829.

Family-group name: TRYPANÉENS Marseul, 1857. Stem: Trypanae-.

Trypanidius Blanchard, 1842c: pl. 22. Current status: valid genus in CERAMBYCIDAE: LAMIINAE: ACANTHOCININI.

Type species: Trypanidius andicola Blanchard, 1842, by monotypy.

Family-group name: TRYPANIDIITAE J. Thomson, 1860. Stem: Trypanidi-.

Trypetes Schönherr, 1836: 595. Current status: valid genus in CURCULIONIDAE: MOLYTINAE: TRYPETIDINI. Note: name placed on the Official List of Generic Names in Zoology (ICZN 1974a).

Type species: Trypetes rhinoides Gyllenhal, 1836, by original designation (see ICZN 1974a). Note: name placed on the Official List of Specific Names in Zoology (ICZN 1974a).

Family-group name: TRYPÉTIDES Lacordaire, 1865. Stem: Trypetid- (ICZN 1974a).

Trypeticus Marseul, 1864: 281. Current status: valid genus in HISTERIDAE: ABRAEINAE: TRYPETICINI.

Type species: Trypanaeus gilolous Marseul, 1864, by subsequent designation (Lewis 1897: 194).

Family-group name: TRYPETICINAE Bickhardt, 1913. Stem: Trypetic-.

Trypodendron Stephens, 1830: 353. Current status: valid genus in CURCULIONIDAE: SCOLYTINAE: XYLOTERINI.

Type species: Dermestes domesticus Linnaeus, 1758, by subsequent designation (Westwood 1838c: 39).

Family-group name: TRYPODENDRINA Nunberg, 1954. Stem: Trypodendr-.

Trypophloeus Fairmaire, 1864: 105. Current status: valid genus in CURCULIONIDAE: SCOLYTINAE: TRYPOPHLOEINI.

Type species: Bostrichus binodulus Ratzeburg, 1837, by monotypy.

Family-group name: TRYPOPHLOEINAE Nüsslin, 1911. Stem: Trypophloe-.

†Tshekardocoleus Rohdendorf, 1944: 252. Current status: valid genus in †TSHEKARDOCOLEIDAE. Note: Permian (Russia).

Type species: †Tshekardocoleus magnus Rohdendorf, 1944, by original designation.

Family-group name: †TSHEKARDOCOLEIDAE Rohdendorf, 1944. Stem: Tshekardocole-.

Tychaeus Fischer, 1823b: 266. Current status: junior synonym of Nemorhinus Schönherr, 1823 in BRENTIDAE: BRENTINAE: BRENTINI: TYCHAEINA.

Type species: Brentus curvidens Lund, 1802 (= Brentus myrmecophaga Herbst, 1797), by subsequent designation (R. Lucas 1920: 661).

Family-group name: TYCHAEIDAE Schoenfeldt, 1908. Stem: Tychae-.

Tychius Germar, 1817b: 340. Current status: valid genus in CURCULIONIDAE: CURCULIONINAE: TYCHIINI: TYCHIINA.

Type species: Curculio quinquepunctatus Linnaeus, 1758, by subsequent designation (Schönherr 1825: 583).

Family-group name: TYCHIIDAE Gistel, 1848. Stem: Tychi-.

Tychus Leach, 1817: 81, 84. Current status: valid genus in STAPHYLINIDAE: PSELAPHINAE: GONIACERITAE: TYCHINI.

Type species: Pselaphus niger Paykull, 1800, by monotypy.

Family-group name: TYCHINI Raffray, 1904. Stem: Tych-. Note: junior homonym of TYCHINAE Dana, 1851 (type genus Tyche T. Bell, 1835) in Crustacea.

Tydessa Peacock, 1982: 362. Current status: valid genus in PYROCHROIDAE: TYDESSINAE.

Type species: Dasytes constrictus Lewis, 1895 (= Dasytes lewisi Pic, 1937), by original designation.

Family-group name: TYDESSINI Nikitsky, 1986. Stem: Tydess-.

Tylauchenia Burmeister, 1872: 377. Current status: valid genus in BUPRESTIDAE: POLYCESTINAE: TYNDARIDINI: TYLAUCHENIINA. Note: this name is considered an invalid junior synonym of Ocypetes Saunders, 1871 by some authors but Saunders’ name is a junior homonym of Ocypetes Risso, 1826 [Arachnida], Ocypetes Wagler, 1829 [Aves] and Ocypetes Lesson, 1842 [Mammalia].

Type species: Buprestis crassicollis Laporte & Gory, 1838, by monotypy.

Family-group name: TYLACHENIAE Cobos, 1959. Stem: Tylaucheni-.

Tylodes C.R. Sahlberg, 1823c: 49. Current status: valid genus in CURCULIONIDAE: CRYPTORHYNCHINAE: CRYPTORHYNCHINI: TYLODINA.

Type species: Tylodes armadillo C.R. Sahlberg, 1823, by monotypy.

Family-group name: TYLODIDES Lacordaire, 1865. Stem: Tylod-.

Tylonotus Schaum, 1863a: 74. Current status: invalid senior synonym of Nototylus Gemminger & Harold, 1868 in CARABIDAE: NOTOTYLINAE. Note: junior homonym of Tylonotus Haldeman, 1847 [Coleoptera: CERAMBYCIDAE].

Type species: Tylonotus fryi Schaum, 1863, by monotypy.

Family-group name: TYLONOTINI Schaum, 1863. Stem: Tylonot-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

Tylosis J.L. LeConte, 1850c: 9. Current status: valid genus in CERAMBYCIDAE: CERAMBYCINAE: TRACHYDERINI: TRACHYDERINA.

Type species: Tylosis maculatus J.L. LeConte, 1850, by subsequent designation (J. Thomson 1864: 200).

Family-group name: TYLOSITAE J. Thomson, 1861. Stem: Tylose-.

Tyndaris J. Thomson, 1857: 168. Current status: valid genus in BUPRESTIDAE: POLYCESTINAE: TYNDARIDINI: TYNDARIDINA.

Type species: Ptosima planata Laporte & Gory, 1838, by monotypy.

Family-group name: TYNDARINI Cobos, 1955. Stem: Tyndarid-.

Typhaea Stephens, 1829b: 8. Current status: valid genus in MYCETOPHAGIDAE: MYCETOPHAGINAE: TYPHAEINI.

Type species: Mycetophagus testaceus Fabricius, 1792 (= Dermestes stercoreus Linnaeus, 1758), by subsequent designation (Westwood 1838c: 14).

Family-group name: TYPHAEINA C.G. Thomson, 1863. Stem: Typhae-.

Typhlocharis Dieck, 1869: 6. Current status: valid genus in CARABIDAE: TRECHINAE: ANILLINI.

Type species: Typhlocharis silvanoides Dieck, 1869, by monotypy.

Family-group name: TYPHLOCHARINA Jeanne, 1973. Stem: Typhlocharit-.

Typhlorhinus Kuschel, 1954b: 287. Current status: junior synonym of Hapactorrhynchus Richard, 1953 in CURCULIONIDAE: ENTIMINAE: OTIORHYNCHITAE: TYPHLORHININI.

Type species: Typhlorhinus jeanneli Kuschel, 1954 (= Hapactorrhynchus vadoni Richard, 1953), by original designation.

Family-group name: TYPHLORHININI Kuschel, 1954. Stem: Typhlorhin-.

Typhlusechus Linell, 1897: 154. Current status: valid genus in TENEBRIONIDAE: PIMELIINAE: STENOSINI: TYPHLUSECHIINA.

Type species: Typhlusechus singularis Linell, 1897, by original designation.

Family-group name: TYPHLUSECHINI Casey, 1907. Stem: Typhlusech-.

Typhocesis Pascoe, 1863b: 561. Current status: valid genus in CERAMBYCIDAE: CERAMBYCINAE: TYPHOCESINI.

Type species: Typhocesis macleayi Pascoe, 1863, by monotypy.

Family-group name: TYPHOCÉSIDES Lacordaire, 1868. Stem: Typhoces-.

Typoderus G.A.K. Marshall, 1953: 104. Current status: valid genus in CURCULIONIDAE: MOLYTINAE: MOLYTINI: TYPODERINA.

Type species: Typoderus machadoi G.A.K. Marshall, 1953, by original designation.

Family-group name: TYPODERINA Voss, 1965. Stem: Typoder-.

Typophorus Chevrolat in Dejean, 1836a: 412. Current status: valid genus in CHRYSOMELIDAE: EUMOLPINAE: TYPOPHORINI.

Type species: Eumolpus nigritus Fabricius, 1801, by subsequent designation (Monrós and Bechyné 1956: 1127).

Family-group name: TYPOPHORINAE Baly, 1865. Stem: Typophor-.

Tyrus Aubé, 1833: 15. Current status: valid genus in STAPHYLINIDAE: PSELAPHINAE: PSELAPHITAE: TYRINI: TYRINA.

Type species: Pselaphus mucronatus Panzer, 1802, by monotypy.

Family-group name: TYRIDES Reitter, 1882. Stem: Tyr-.

Tytthaspis Crotch, 1874a: 181. Current status: valid genus in COCCINELLIDAE: COCCINELLINAE: COCCINELLINI.

Type species: Coccinella sedecimpunctata Linnaeus, 1761, by original designation.

Family-group name: TYTTHASPIDES Crotch, 1874. Stem: Tytthaspid-.

Tytthonyx J.L. LeConte, 1851b: 347. Current status: valid genus in CANTHARIDAE: MALTHININAE: TYTTHONYXINI.

Type species: Lampyris erythrocephala Fabricius, 1801, by monotypy.

Family-group name: TYTTHONYINI Arnett, 1962. Stem: Tytthonyx-.

Uleiota Latreille, 1797: 46. Current status: valid genus in SILVANIDAE: BRONTINAE: BRONTINI.

Type species: Cucujus flavipes Fabricius, 1779 (= Cerambyx planatus Linnaeus, 1761), by subsequent monotypy (Latreille 1802b: 210).

Family-group name: ULEIOTAEIDAE Gistel, 1848. Stem: Uleiot-.

Ulocerus Schönherr, 1823: 1137. Current status: valid genus in BRENTIDAE: BRENTINAE: ULOCERINI: ULOCERINA.

Type species: Ulocerus lacerosus Schönherr, 1823, by original designation.

Family-group name: ULOCERIDES Schönherr, 1823. Stem: Ulocer-.

Ulodes Erichson, 1842: 180. Current status: valid genus in ULODIDAE.

Type species: Ulodes verrucosus Erichson, 1842, by monotypy.

Family-group name: ULODINAE Pascoe, 1869. Stem: Ulod-.

Uloma Dejean, 1821: 67. Current status: valid genus in TENEBRIONIDAE: TENEBRIONINAE: ULOMINI. Note: name placed on the Official List of Generic Names in Zoology (ICZN 1975).

Type species: Tenebrio culinaris Linnaeus, 1758, by plenary power (see ICZN 1975). Note: name placed on the Official List of Specific Names in Zoology (ICZN 1975).

Family-group name: ULOMITES Blanchard, 1845. Stem: Ulom-.

Ulomascus Fairmaire, 1848: 173. Current status: valid genus in CURCULIONIDAE: CURCULIONINAE: ULOMASCINI.

Type species: Ulomascus caviventris Fairmaire, 1848, by monotypy.

Family-group name: ULOMASCIDES Lacordaire, 1865. Stem: Ulomasc-.

†Ulyana Zherikhin, 1993: 27. Current status: valid genus in ANTHRIBIDAE: †ULYANINAE. Note: Cretaceous (Mongolia; Russia).

Type species: †Ulyana nobilis Zherikhin, 1993, by original designation.

Family-group name: †ULYANIDAE Zherikhin, 1993. Stem: Ulyan-.

†Ulyanisca Gratshev, 1998: 45. Current status: valid genus in BRENTIDAE: ITHYCERINAE: †ULYANISCINI. Note: Lower Cretaceous (Mongolia).

Type species: †Ulyanisca dentipes Gratschev, 1998, by original designation.

Family-group name: †ULYANISCINI Legalov, 2009. Stem: Ulyanisc-.

Undulifer Gemminger in Gemminger and Harold, 1868c: 974. Current status: valid genus in PASSALIDAE: PASSALINAE: PROCULINI. Note: this name was previously attributed to Kaup (1869: 6) in the literature; however, it was first made available by Gemminger in 1868 when he associated one nominal species under this name.

Type species: Passalus incisus Truqui, 1857, by monotypy.

Family-group name: UNDULIFERINAE Kuwert, 1891. Stem: Undulifer-.

Unxia J. Thomson, 1861: 234, 252. Current status: valid genus in CERAMBYCIDAE: CERAMBYCINAE: UNXIINI.

Type species: Unxia insignis J. Thomson, 1861 (= Cosmisoma insigne Guérin-Méneville, 1844), by monotypy.

Family-group name: UNXIINI Napp, 2007. Stem: Unxi-.

Upinella Mulsant, 1857b: 17. Current status: valid genus in TENEBRIONIDAE: ALLECULINAE: ALLECULINI: ALLECULINA.

Type species: Allecula aterrima Rosenhauer, 1847, by monotypy.

Family-group name: UPINELLAE J.L. LeConte, 1866. Stem: Upinell-.

Upis Fabricius, 1792b: 515. Current status: valid genus in TENEBRIONIDAE: STENOCHIINAE: CNODALONINI.

Type species: Attelabus ceramboides Linnaeus, 1758, by monotypy.

Family-group name: UPIDAE C.G. Thomson, 1859. Stem: Upid-.

Uptona G.S. Medvedev & Lawrence, 1986: 582. Current status: valid genus in TENEBRIONIDAE: DIAPERINAE: HYOCIINI: UPTONINA.

Type species: Uptona pallida G.S. Medvedev & Lawrence, 1986, by original designation.

Family-group name: UPTONINA G.S. Medvedev & Lawrence, 1986. Stem: Upton-.

Uracanthus Hope, 1833: 64. Current status: valid genus in CERAMBYCIDAE: CERAMBYCINAE: URACANTHINI.

Type species: Uracanthus triangularis Hope, 1833, by monotypy.

Family-group name: URACANTHITAE Blanchard, 1853. Stem: Uracanth-.

†Uralocoleus Zalesskiy, 1947: 858. Current status: valid genus in †TSHEKARDOCOLEIDAE. Note: Permian (Russia).

Type species: †Uralocoleus splendens Zalesskiy, 1947, by original designation.

Family-group name: †URALOCOLEIDAE Zalesskiy, 1947. Stem: Uralocole-.

Urodon Schönherr, 1823: 1134. Current status: junior synonym of Bruchela Dejean, 1821 in ANTHRIBIDAE: URODONTINAE. Note: unnecessary new replacement name for Bruchela Dejean, 1821.

Type species: Anthribus sericeus Fabricius, 1801 (= Bruchus rufipes G.-A. Olivier, 1790), by original designation.

Family-group name: URODONTIDES C.G. Thomson, 1859. Stem: Urodont-.

Uroplata Chevrolat in Dejean, 1836a: 365. Current status: valid genus in CHRYSOMELIDAE: CASSIDINAE: UROPLATINI. Note: name placed on the Official List of Generic Names in Zoology (ICZN 1985f).

Type species: Hispa mucronata G.-A. Olivier, 1808, by subsequent designation (R.E. White 1981: 714; see ICZN 1985f). Note: name placed on the Official List of Specific Names in Zoology (ICZN 1985f).

Family-group name: UROPLATINI Weise, 1910. Stem: Uroplat-.

Uroptera Berthold, 1827: 383. Current status: valid genus in BRENTIDAE: BRENTINAE: TRACHELIZINI: PSEUDOCEOCEPHALINA.

Type species: here fixed (under ICZN 1999a, Article 70.3.2) as Ceocephalus appendiculatus Boheman, 1833, misidentified as Brentus caudatus Herbst, 1797 (sensu Latreille, 1828) in the original designation by subsequent monotypy in Latreille (1828b: 595, as Uropterus).

Family-group name: UROPTERINI Jakobson, 1911. Stem: Uropter-.

Usechus Motschulsky, 1845b: 79. Current status: valid genus in ZOPHERIDAE: ZOPHERINAE: USECHINI.

Type species: Usechus lacerta Motschulsky, 1845, by monotypy.

Family-group name: USECHINI G.H. Horn, 1867. Stem: Usech-.

Vacronus Casey, 1907: 501, 508. Current status: junior synonym of Alaephus G.H. Horn, 1870 in TENEBRIONIDAE: PIMELIINAE: VACRONINI.

Type species: Vacronus tenuicornis Casey, 1907, by original designation.

Family-group name: VACRONINAE Gebien, 1910. Stem: Vacron-.

Vadonaxia Descarpentries, 1970: 189. Current status: valid genus in BUPRESTIDAE: CHRYSOCHROINAE: VADONAXIINI.

Type species: Vadonaxia peyrierasi Descarpentries, 1970, by original designation.

Family-group name: VADONAXIINI Descarpentries, 1970. Stem: Vadonaxi-.

Valda Casey, 1893: 493. Current status: valid genus in STAPHYLINIDAE: PSELAPHINAE: GONIACERITAE: VALDINI.

Type species: Valda frontalis Casey, 1893, by monotypy.

Family-group name: VALDIINI Park, 1953. Stem: Vald-.

Valenfriesia Alonso-Zarazaga & Lyal, 1999: 38. Current status: valid genus in ANTHRIBIDAE: CHORAGINAE: VALENFRIESIINI. Note: new replacement name for Notioxenus Wollaston, 1861.

Type species: Notioxenus bewickii Wollaston, 1861, by subsequent designation (Jordan 1907: 380) (type fixation for Notioxenus Wollaston).

Family-group name: VALENFRIESIINI Alonso-Zarazaga & Lyal, 1999. Stem: Valenfriesi-.

Valgus Scriba, 1790: 66. Current status: valid genus in SCARABAEIDAE: CETONIINAE: VALGINI.

Type species: Scarabaeus hemipterus Linnaeaus, 1758, by monotypy.

Family-group name: VALGUAIRES Mulsant, 1842. Stem: Valg-.

Vansonium C. Koch, 1950b: 354. Current status: valid genus in TENEBRIONIDAE: PIMELIINAE: CRYPTOCHILINI: VANSONIINA.

Type species: Vansonium bushmanicum C. Koch, 1950, by original designation.

Family-group name: VANSONINI C. Koch, 1955. Stem: Vansoni-.

Vatellus Aubé, 1837: 218, 221. Current status: valid genus in DYTISCIDAE: HYDROPORINAE: VATELLINI. Note: the older name Leucorea Laporte, 1835, which was later replaced with Vatellus by Aubé (1837), was suppressed for the Principle of Priority but not the Principle of Homonymy and placed on the Official Index of Rejected and Invalid Names in Zoology while Vatellus Aubé, 1837 was placed on the Official List of Generic Names in Zoology (ICZN 1992).

Type species: Hydroporus tarsatus Laporte, 1835, by monotypy (see ICZN 1992). Note: name placed on the Official List of Specific Names in Zoology (ICZN 1992).

Family-group name: VATELLINI Sharp, 1880. Stem: Vatell-.

Vatesus Sharp, 1876a: 201. Current status: valid genus in STAPHYLINIDAE: TACHYPORINAE: VATESINI.

Type species: Vatesus latitans Sharp, 1876, by monotypy.

Family-group name: VATESINI Seevers, 1958. Stem: Vates-.

Vatinius Harold in Gemminger and Harold, 1868c: 978. Current status: junior synonym of Passalus Fabricius, 1792 in PASSALIDAE: PASSALINAE: PASSALINI. Note: junior homonym of Vatinius Stål, 1865 [Hemiptera]; this name is attributed to Kaup (1869: 35) in the literature; however, it was first made available by Harold in 1868 when he associated several nominal species with this name.

Type species: Passalus toriferus Eschscholtz, 1828 [as torifer], by subsequent designation (Hincks and Dibb 1935: 38).

Family-group name: VATINIINAE Kuwert, 1896. Stem: Vatini-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

Vellejus Kaup, 1871: 35. Current status: junior synonym of Labienus Kaup, 1871 in PASSALIDAE: PASSALINAE: MACROLININI. Note: junior homonym of Vellejus Stål, 1865 [Hemiptera].

Type species: Passalus compergus Boisduval, 1835, by subsequent designation (Hincks and Dibb 1935: 93).

Family-group name: VELLEJINAE Kuwert, 1891. Stem: Vellej-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

Velora J. Thomson, 1864: 56. Current status: valid genus in CERAMBYCIDAE: LAMIINAE: DESMIPHORINI.

Type species: Velora australis J. Thomson, 1864, by original designation.

Family-group name: VELORINI Aurivillius, 1917. Stem: Velor-.

Verania Mulsant, 1850: 358. Current status: junior synonym of Micraspis Chevrolat, 1836 in COCCINELLIDAE: COCCINELLINAE: COCCINELLINI. Note: junior homonym of Verania Krohn, 1846 [Mollusca].

Type species: Coccinella comma Thunberg, 1781, by subsequent designation (Crotch 1874a: 175).

Family-group name: VERANIINI F.A. Schilder & M. Schilder, 1928. Stem: Verani-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

Vesperella Dayrem, 1933: 89. Current status: valid genus in CERAMBYCIDAE: CERAMBYCINAE: VESPERELLINI.

Type species: Vesperella pallida Dayrem, 1933, by monotypy.

Family-group name: VESPERELLINI Sama, 2008. Stem: Vesperell-.

Vesperoctenus H.W. Bates, 1891: 159. Current status: valid genus in CERAMBYCIDAE: PRIONINAE: VESPEROCTENINI.

Type species: Vesperoctenus florhi H.W. Bates, 1891, by monotypy.

Family-group name: VESPEROCTENINI Vives, 2005. Stem: Vesperocten-.

Vesperus Dejean, 1821: 111. Current status: valid genus in VESPERIDAE: VESPERINAE.

Type species: Stenocorus strepens Fabricius, 1792, by subsequent designation (Chevrolat 1849b: 216).

Family-group name: VESPÉRAIRES Mulsant, 1839. Stem: Vesper-.

Vesta Laporte, 1833c: 126, 132. Current status: valid genus in LAMPYRIDAE incertae sedis.

Type species: Vesta chevrolatii Laporte, 1833, by monotypy.

Family-group name: VESTINA McDermott, 1964. Stem: Vest-.

†Vetuproteinus Cai, Newton & Thayer in Cai et al., 2016: 2. Current status: valid genus in STAPHYLINIDAE: PROTEININAE: †VETUPROTEININI. Note: mid-Cretaceous (Myanmar).

Type species: †Vetuproteinus cretaceus Cai, Newton & Thayer, 2016, by original designation.

Family-group name: †VETUPROTEININI Cai, Newton & Thayer in Cai et al., 2016: 2. Stem: Vetuprotein-.

Veturius Kaup, 1871: 110. Current status: valid genus in PASSALIDAE: PASSALINAE: PROCULINI.

Type species: Passalus heydenii Kaup, 1868, by subsequent designation (Gravely 1918: 35).

Family-group name: VETURINAE Kuwert, 1891. Stem: Veturi-.

Vikhrevia Kazantsev, 2013d: 248. Current status: valid genus in LYCIDAE incertae sedis.

Type species: Vikhrevia ludificator Kazantsev, 2013, by original designation.

Family-group name: VIKHREVINI Kazantsev, 2013d: 247. Stem: Vikhrev- (ICZN 1999a, Article 29.4).

Vindex Kaup, 1871: 78. Current status: valid genus in PASSALIDAE: PASSALINAE: PROCULINI.

Type species: Passalus agnoscendus Percheron, 1841, by monotypy.

Family-group name: VINDICINAE Kuwert, 1896. Stem: Vindic-.

Viticis Lea, 1930: 463. Current status: valid genus in CURCULIONIDAE: CURCULIONINAE: VITICIINI.

Type species: Viticis bidentatus Lea, 1930, by original designation.

Family-group name: VITICIINAE Morimoto, 1983. Stem: Vitici-.

Viticlerus Miyatake, 1977: 105. Current status: valid genus in THANEROCLERIDAE: THANEROCLERINAE: THANEROCLERINI.

Type species: Viticlerus formicinus Miyatake, 1977, by original designation.

Family-group name: VITICLERINI J.R. Winkler, 1982. Stem: Viticler-.

Vossicartus Legalov, 2003: 79. Current status: valid genus in ATTELABIDAE: SAYREVILLEINAE: VOSSICARTINI.

Type species: Rhinocartus bruncki Voss, 1974, by original designation.

Family-group name: VOSSICARTINI Legalov, 2003. Stem: Vossicart-.

Vrilletta J.L. LeConte, 1874b: 64. Current status: valid genus in PTINIDAE: XYLETININAE: XYLETININI.

Type species: Vrilletta murrayi J.L. LeConte, 1874, by subsequent designation (R.E. White 1974: 453). Note: R.E. White (1974: 453) listed R. Lucas (1920: 668) as the author who selected the type species; however, R. Lucas listed two species and this is considered a selection made in an ambiguous manner and cannot be considered as a designation under the Code (ICZN 1999a, Article 67.5.3).

Family-group name: VRILETTINI Böving, 1927. Stem: Vrillett-.

†Wabbel Alekseev, 2017: 30. Current status: valid genus in †WABBELIDAE. Note: mid-Eocene (Kaliningrad region, Russia).

Type species: †Wabbel cerebricavus Alekseev, 2017, by original designation.

Family-group name: †WABBELIDAE Alekseev, 2017: 34. Stem: Wabbel-.

†Waidelotus Bukejs, Alekseev & Pollock, 2019: 264. Current status: valid genus in PYROCHROIDAE: †WAIDELOTINAE. Note: mid-Eocene (Kaliningrad region, Russia).

Type species: †Waidelotus hoffeinsorum Bukejs, Alekseev & Pollock, 2019, by original designation.

Family-group name: †WAIDELOTINAE Bukejs, Alekseev & Pollock, 2019: 262. Stem: Waidelot-.

Warwickia A.B.T. Smith & Evans, 2005: 42. Current status: valid genus in SCARABAEIDAE: MELOLONTHINAE: WARWICKIINI. Note: new replacement name for Benedictia Sanderson, 1939.

Type species: Benedictia pilosa Sanderson, 1939, by original designation (type fixation for Benedictia Sanderson).

Family-group name: WARWICKIINI Evans & A.B.T. Smith, 2020: 68. Stem: Warwicki-.

Webbia Hopkins, 1915: 222. Current status: valid genus in CURCULIONIDAE: SCOLYTINAE: XYLEBORINI.

Type species: Webbia dipterocarpi Hopkins, 1915, by original designation.

Family-group name: WEBBINAE Hopkins, 1915. Stem: Webbi-.

Wittmeraltica Bechyné, 1956: 1011. Current status: valid genus in CHRYSOMELIDAE: GALERUCINAE: ALTICINI.

Type species: Wittmeraltica muriensis Bechyné, 1956, by monotypy.

Family-group name: WITTMERALTICINA Bechyné, 1968. Stem: Wittmeraltic-.

Wittmeroquasimus Dolin, 1993: 195. Current status: valid genus in ELATERIDAE: NEGASTRIINAE: QUASIMUSINI: WITTMEROQUASINA.

Type species: Quasimus ocellatus Dolin, 1993, by original designation.

Family-group name: WITTMEROQUASINA Schimmel & Tarnawski, 2009. Stem: Wittmeroquas- (ICZN 1999a, Article 29.4).

Wooldridgeus Spangler, 1999: 181. Current status: junior synonym of Platypelochares Champion, 1923 in LIMNICHIDAE: LIMNICHINAE: WOOLDRIDGEINI.

Type species: Wooldridgeus perforatus Spangler, 1999, by original designation.

Family-group name: WOOLDRIDGEINI Spangler, 1999. Stem: Wooldridge-.

Wow Jałoszyński, Maruyama & Klimaszewski, 2023: 575. Current status: valid genus in STAPHYLINIDAE: ALEOCHARINAE: WOWINI.

Type species: Wow assingi Jałoszyński, Maruyama & Klimaszewski, 2023, by original designation.

Family-group name: WOWINI Jałoszyński, Maruyama & Klimaszewski, 2023: 574. Stem: Wow-.

Xantholinus Dejean, 1821: 23. Current status: valid genus in STAPHYLINIDAE: STAPHYLININAE: XANTHOLININI. Note: name placed on the Official List of Generic Names in Zoology (ICZN 1983c).

Type species: Staphylinus linearis G.-A. Olivier, 1795, by plenary power (see ICZN 1983c). Note: name placed on the Official List of Specific Names in Zoology (ICZN 1983c, as Staphylinus linearis G.-A. Olivier, 1794).

Family-group name: XANTHOLININI Erichson, 1839. Stem: Xantholin-. Note: family-group name given precedence over AGRODINI Nordmann, 1837 and GYROHYPNINI Kirby, 1837 and placed on the Official List of Family-Group Names in Zoology (ICZN 1996c).

Xanthopygus Kraatz, 1857b: 539. Current status: valid genus in STAPHYLINIDAE: STAPHYLININAE: STAPHYLININI: XANTHOPYGINA.

Type species: Staphylinus xanthopygus Nordmann, 1837, by absolute tautonymy.

Family-group name: XANTHOPYGINA Sharp, 1884. Stem: Xanthopyg-.

Xenaroswelliana Erwin, 2007: 563. Current status: valid genus in CARABIDAE: HARPALINAE: XENAROSWELLIANINI.

Type species: Xenaroswelliana deltaquadrant Erwin, 2007, by original designation.

Family-group name: XENAROSWELLANINI Erwin, 2007. Stem: Xenaroswellian-.

Xenaster Bierig, 1939: 179. Current status: invalid senior synonym of Xenasterides Newton, 2017 in STAPHYLINIDAE: PAEDERINAE: LATHROBIINI: STILICOPSINA. Note: junior homonym of Xenaster Simonwitsch, 1871 [Echinodermata].

Type species: Xenaster plaumanni Bierig, 1939, by original designation.

Family-group name: XENASTERES Bierig, 1939. Stem: Xenaster-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

Xenicotela H.W. Bates, 1884: 242. Current status: valid genus in CERAMBYCIDAE: LAMIINAE: XENICOTELINI.

Type species: Xenicotela fuscula H.W. Bates, 1884 (= Monohammus pardalinus H.W. Bates, 1884), by monotypy.

Family-group name: XENICOTELINI Matsushita, 1933. Stem: Xenicotel-.

Xenobythus Peyerimhoff, 1901: 203. Current status: valid genus in STAPHYLINIDAE: PSELAPHINAE: GONIACERITAE: BYTHININI: XENOBYTHINA.

Type species: Xenobythus serullazi Peyerimhoff, 1901, by monotypy.

Family-group name: XENOBYTHINA Jeannel, 1950. Stem: Xenobyth-.

Xenocephalus Wasmann, 1887: 411. Current status: junior synonym of Vatesus Sharp, 1876 in STAPHYLINIDAE: TACHYPORINAE: VATESINI. Note: junior homonym of Xenocephalus Kaup, 1858 [Pisces].

Type species: Xenocephalus clypeatus Wasmann, 1887, by monotypy.

Family-group name: XENOCEPHALINI Wasmann, 1887. Stem: Xenocephal-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

Xenocerus Schönherr, 1833: 3, 117. Current status: valid genus in ANTHRIBIDAE: ANTHRIBINAE: XENOCERINI.

Type species: Xenocerus saperdoides Gyllenhal, 1833, by original designation.

Family-group name: XÉNOCERIDES Lacordaire, 1865. Stem: Xenocer-.

Xenodusa Wasmann, 1894: 205. Current status: valid genus in STAPHYLINIDAE: ALEOCHARINAE: LOMECHUSINI: LOMECHUSINA.

Type species: Atemeles cava J.L. LeConte, 1863, by subsequent designation (Fenyes 1919: 26).

Family-group name: XENODUSAE Seevers, 1978. Stem: Xenodus-.

Xenofrea H.W. Bates, 1885: 373. Current status: valid genus in CERAMBYCIDAE: LAMIINAE: XENOFREINI.

Type species: Xenofrea areolata H.W. Bates, 1885, by subsequent designation (Marinoni 1977: 50).

Family-group name: XENOFREINI Aurivillius, 1923. Stem: Xenofre-.

Xenolea J. Thomson, 1864: 91. Current status: valid genus in CERAMBYCIDAE: LAMIINAE: XENOLEINI.

Type species: Xenolea collaris J. Thomson, 1864, by original designation.

Family-group name: XÉNOLÉIDES Lacordaire, 1872. Stem: Xenole-.

Xenomycetes G.H. Horn, 1880b: 141. Current status: valid genus in ENDOMYCHIDAE: XENOMYCETINAE.

Type species: Xenomycetes morrisoni G.H. Horn, 1880, by monotypy.

Family-group name: XENOMYCETINAE Strohecker, 1962. Stem: Xenomycet-.

Xenopsis Saunders, 1867: 514. Current status: valid genus in BUPRESTIDAE: POLYCESTINAE: POLYCESTINI: XENOPSINA.

Type species: Xenopsis laevis Saunders, 1867, by monotypy.

Family-group name: XENOPSINA Volkovitsh, 2008. Stem: Xenops- (ICZN 1999a, Article 29.4).

Xenorchestes Wollaston, 1854: 417. Current status: valid genus in ANTHRIBIDAE: CHORAGINAE: XENORCHESTINI.

Type species: Xenorchestes saltitans Wollaston, 1854, by monotypy.

Family-group name: XÉNORCHESTIDES Lacordaire, 1865. Stem: Xenorchest-.

Xenorhipis J.L. LeConte, 1867: 384. Current status: valid genus in BUPRESTIDAE: BUPRESTINAE: XENORHIPIDINI: XENORHIPIDINA.

Type species: Xenorhipis brendeli J.L. LeConte, 1867, by monotypy.

Family-group name: XENORHIPINI Cobos, 1986. Stem: Xenorhipid-.

Xenoscelinus Grouvelle, 1910b: 143. Current status: junior synonym of Cathartocryptus Sharp, 1886 in EROTYLIDAE: PHARAXONOTHINAE.

Type species: Xenoscelinus malaicus Grouvelle, 1910, by monotypy.

Family-group name: XENOSCELININI Sen Gupta & Crowson, 1971. Stem: Xenoscelin-.

Xenoscelis Wollaston, 1864: 132. Current status: valid genus in EROTYLIDAE: XENOSCELINAE. Note: new replacement name for Pristoscelis Wollaston, 1862.

Type species: Pristoscelis deplanatus Wollaston, 1862, by monotypy (type fixation for Pristoscelis Wollaston).

Family-group name: XENOSCELINI Ganglbauer, 1899. Stem: Xenoscel-.

Xenospasta Selander, 1966: 467. Current status: valid genus in MELOIDAE: ELETICINAE: SPASTICINI: XENOSPASTINA.

Type species: Xenospasta flava Selander, 1966, by original designation.

Family-group name: XENOSPASTINA Selander, 1966. Stem: Xenospast-.

Xenostanus Y.-D. Li, Szawaryn & Cai in Y.-D. Li et al., 2022: 413. Current status: valid genus in CORYLOPHIDAE: CORYLOPHINAE: XENOSTANINI.

Type species: Xenostanus jiangkuni Y.-D. Li, Szawaryn & Cai, 2022, by original designation.

Family-group name: XENOSTANINI Y.-D. Li, Szawaryn & Cai in Y.-D. Li et al., 2022: 413. Stem: Xenostan-.

Xerasiborus Jordal, 2021: 102. Current status: valid genus in CURCULIONIDAE: SCOLYTINAE: HYPOBORINI: XERASIBORINA.

Type species: Xerasiborus euphorbiae Jordal, 2021, by original designation.

Family-group name: XERASIBORINA Jordal, 2021: 102. Stem: Xerasibor-.

Xerobia Gistel, 1856: 373. Current status: junior synonym of Cleonis Dejean, 1821 in CURCULIONIDAE: LIXINAE: CLEONINI.

Type species: Curculio sulcirostris Linnaeus, 1758, by subsequent designation (Alonso- Zarazaga and Lyal 1999: 22, 191).

Family-group name: XEROBIADAE Gistel, 1856. Stem: Xerobi-.

Xestobium Motschulsky, 1845b: 35. Current status: valid genus in PTINIDAE: ERNOBIINAE: XESTOBIINI.

Type species: Dermestes tessellatus Fabricius, 1775 (= Ptinus rufovillosus DeGeer, 1774), by original designation.

Family-group name: XESTOBIINI Böving, 1927. Stem: Xestobi-.

Xiphaspis G.A.K. Marshall, 1920: 393. Current status: valid genus in CURCULIONIDAE: XIPHASPIDINAE.

Type species: Xiphaspis longiclavis G.A.K. Marshall, 1920, by original designation.

Family-group name: XIPHASPIDINAE G.A.K. Marshall, 1920. Stem: Xiphaspid-.

Xiphoscelis Burmeister, 1842: 613. Current status: valid genus in SCARABAEIDAE: CETONIINAE: XIPHOSCELIDINI.

Type species: Xiphoscelis schuckardi Burmeister, 1842, by subsequent designation (Lacordaire 1855: 495). Note: Lacordaire (1855: 495) designated Cetonia gariepena Gory & Percheron, 1833 as type species of Xiphoscelis, a nominal species not originally included, but since he placed that species in synonymy with Xiphoscelis schuckardi Burmeister, a species originally included, he is deemed to have designated the latter species as type species (ICZN 1999a, Article 69.2.2).

Family-group name: XIPHOSCELIDEAE Burmeister, 1842. Stem: Xiphoscelid-.

Xixuthrus J. Thomson, 1864: 296. Current status: valid genus in CERAMBYCIDAE: PRIONINAE: MACROTOMINI: XIXUTHRINA.

Type species: Macrotoma microcerus A. White, 1853, by original designation.

Family-group name: XIXUTHRI Lameere, 1912. Stem: Xixuthr-.

Xylariophilus Pal & Lawrence, 1986: 185. Current status: valid genus in TEREDIDAE: XYLARIOPHILINAE.

Type species: Xylariophilus honoratus Pal & Lawrence, 1986, by original designation. Note: Xylariophilus comatus Pal & Lawrence, 1986 is sometimes mentioned as the type species (e.g., Ślipiński 2007: 551).

Family-group name: XYLARIOPHILINAE Pal & Lawrence, 1986. Stem: Xylariophil-.

Xyleborips Reitter, 1913b: 111. Current status: junior synonym of Monarthrum Kirsch, 1866 in CURCULIONIDAE: SCOLYTINAE: CORTHYLINI: CORTHYLINA.

Type species: Xyloborus meuseli Reitter, 1905, by monotypy.

Family-group name: XYLEBORIPINA Reitter, 1913. Stem: Xyleborip-.

Xyleborus Eichhoff, 1864: 37, 45. Current status: valid genus in CURCULIONIDAE: SCOLYTINAE: XYLEBORINI. Note: the senior homonym Xyleborus Bowdich, 1825 [Coleoptera: BUPRESTIDAE], and all other uses of the name Xyleborus prior to Xyleborus Eichhoff, 1864, were suppressed for the purposes of the Principle of Priority and the Principle of Homonymy by the Commission and Xyleborus Eichhoff, 1864 was placed on the Official List of Generic Names in Zoology (ICZN 1968a).

Type species: Bostrichus monographus Fabricius, 1792, by subsequent designation (Lacordaire 1865: 381; see ICZN 1968a). Note: name placed on the Official List of Specific Names in Zoology (ICZN 1968a).

Family-group name: XYLEBORI J.L. LeConte, 1876. Stem: Xylebor-.

Xylechinus Chapuis, 1869: 36. Current status: valid genus in CURCULIONIDAE: SCOLYTINAE: HYLURGINI.

Type species: Hylesinus pilosus Ratzeburg, 1837, by monotypy.

Family-group name: XYLECHINIDES Nüsslin, 1912. Stem: Xylechin-.

Xyletinus Latreille, 1809a: 376. Current status: valid genus in PTINIDAE: XYLETININAE: XYLETININI. Note: name placed on the Official List of Generic Names in Zoology (ICZN 1971).

Type species: Ptilinus ater Panzer, 1796, by plenary power (ICZN 1971, as Ptilinus ater Creutzer, 1796). Note: Panzer (1796) listed Creutzer after the species’ name on the plate; however, Panzer was responsible for the work and he is the one that satisfied the criteria of availability (ICZN 1999a, Article 50.1).

Family-group name: XYLETINIDAE Gistel, 1848. Stem: Xyletin-.

Xylinades Latreille, 1828b: 590. Current status: junior synonym of Xylinada Berthold, 1827 in ANTHRIBIDAE: ANTHRIBINAE: XYLINADINI.

Type species: Xylinades westermanni Schönherr, 1833, by subsequent designation (Schönherr 1833: 177). Note: no species were originally included in this genus by Latreille (1828b); the first ones were the six species listed by Schönherr (1833: 178–181).

Family-group name: XYLINADIDES Lacordaire, 1865. Stem: Xylinad-.

Xylita Paykull, 1798: 249. Current status: valid genus in MELANDRYIDAE: MELANDRYINAE: XYLITINI.

Type species: Elater buprestoides Fabricius, 1792 (= Serropalpus laevigatus Hellenius, 1786), by subsequent designation (Mulsant 1856b: 86). Note: Mulsant (1856b: 86) designated Serropalpus laevigatus Hellenius, 1786 as type species, a species not originally included in Xylita, but since he listed the species in synonymy with Elater buprestoides Fabricius, 1792 at the same time, a species originally included in Xylita, he is deemed to have selected the latter species (ICZN 1999a, Article 69.2.2).

Family-group name: XYLITINA C.G. Thomson, 1864. Stem: Xylit-.

Xylobius Latreille, 1834: 124. Current status: junior synonym of Xylophilus Mannerheim, 1823 in EUCNEMIDAE: MELASINAE: XYLOBIINI. Note: unnecessary new replacement name for Xylophilus Mannerheim, 1823.

Type species: Elater alni Fabricius, 1801 (= Elater corticalis Paykull, 1800), by subsequent designation (Guérin-Méneville 1843b: 175).

Family-group name: XYLOBIINI Reitter, 1911. Stem: Xylobi-.

Xyloctonus Eichhoff, 1872: 134. Current status: valid genus in CURCULIONIDAE: SCOLYTINAE: XYLOCTONINI: XYLOCTONINA.

Type species: Xyloctonus scolytoides Eichhoff, 1872, by monotypy.

Family-group name: XYLOCTONIDAE Eichhoff, 1878. Stem: Xylocton-.

Xylographella Miyatake, 1985: 279. Current status: valid genus in CIIDAE: CIINAE: XYLOGRAPHELLINAE: XYLOGRAPHELLINI.

Type species: Xylographella punctata Miyatake, 1985, by monotypy.

Family-group name: XYLOGRAPHELLINI Kawanabe & Miyatake, 1996. Stem: Xylographell-.

Xylonichus Boisduval, 1835: 186. Current status: valid genus in SCARABAEIDAE: MELOLONTHINAE: COLYMBOMORPHINI.

Type species: Xylonichus eucalypti Boisduval, 1835, by monotypy.

Family-group name: XYLONYCHINI Britton, 1957. Stem: Xylonich-.

Xylopertha Guérin-Méneville, 1845: xvii. Current status: valid genus in BOSTRICHIDAE: BOSTRICHINAE: XYLOPERTHINI.

Type species: Apate sinuata Fabricius, 1792 (= Bostrichus retusus G.-A. Olivier, 1790), by subsequent designation (E. Desmarest 1860: 258).

Family-group name: XYLOPERTHINI Lesne, 1921. Stem: Xyloperth-.

Xylophilus Latreille, 1828b: 589. Current status: invalid senior synonym of Euglenes Westwood, 1830 in ADERIDAE. Note: Latreille (1825: 383) used the vernacular name “Xylophile” for this genus, the name is therefore unavailable from that work; junior homonym of Xylophilus Mannerheim, 1823 [Coleoptera: EUCNEMIDAE].

Type species: Anthicus oculatus Paykull, 1798, by subsequent designation (Curtis 1830: pl. 299). Note: the type species often given in the literature is Notoxus populneus Creutzer, 1796 (e.g., Nardi 2007: 23) by subsequent designation of Westwood (1830a: 57, 61); however, we failed to find any indication in Westwood’s work that he designated a type species for Xylophilus Latreille, 1828. The valid type species fixation is that of Curtis (1830: pl. 299) and therefore Xylophilus Latreille, 1828 is a senior synonym of Euglenes Westwood, 1830, not of Aderus Stephens, 1829, as is sometimes recognized (e.g., Nardi 2020: 626).

Family-group name: XYLOPHILIDAE Shuckard, 1839. Stem: Xylophil-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

Xylorhiza Dejean, 1835: 344. Current status: valid genus in CERAMBYCIDAE: LAMIINAE: XYLORHIZINI.

Type species: Lamia adusta Wiedemann, 1819, by monotypy.

Family-group name: XYLORHIZIDES Lacordaire, 1872. Stem: Xylorhiz-.

Xylosteus Frivaldszky von Frivald, 1837: 180. Current status: valid genus in CERAMBYCIDAE: LEPTURINAE: XYLOSTEINI.

Type species: Xylosteus spinolae Frivaldszky von Frivald, 1837, by monotypy.

Family-group name: XYLOSTEINA Reitter, 1913. Stem: Xyloste-.

Xyloterus Erichson, 1836: 60. Current status: junior synonym of Trypodendron Stephens, 1830 in CURCULIONIDAE: SCOLYTINAE: XYLOTERINI.

Type species: Bostrichus lineatus G.-A. Olivier, 1795, by subsequent designation (C.G. Thomson 1859: 146).

Family-group name: XYLOTERI J.L. LeConte, 1876. Stem: Xyloter-.

Xylotrogus Stephens, 1830: 116. Current status: valid genus in BOSTRICHIDAE: LYCTINAE: LYCTINI.

Type species: Xylotrogus brunneus Stephens, 1830, by monotypy (see ICZN 1994d).

Family-group name: XYLOTROGIDAE Schönfeldt, 1887. Stem: Xylotrog-.

Xylotrupes Hope, 1837: 15, 19, 29, 30. Current status: valid genus in SCARABAEIDAE: DYNASTINAE: DYNASTINI: XYLOTRUPINA.

Type species [suggested]: Scarabaeus gideon Linnaeus, 1767, by subsequent designation (E. Desmarest 1860: 92). Note: the valid type species fixation is that of Shuckard in Swainson and Shuckard (1840: 216), who selected Scarabaeus centaurus Fabricius, 1775, but that species is currently included in the genus Augosoma Burmeister, 1841; an application to the Commission is necessary to fix Scarabaeus gideon Linnaeus, 1767 as the type species of Xylotrupes Hope, 1837.

Family-group name: XYLOTRUPIDAE Hope, 1838. Stem: Xylotrup-.

Xyroscelis Saunders, 1868: 53. Current status: valid genus in BUPRESTIDAE: POLYCESTINAE: XYROSCELIDINI.

Type species: Amorphosoma crocatum Gory & Laporte, 1839, by monotypy.

Family-group name: XIROSCELINI Cobos, 1955. Stem: Xyroscelid-.

Xystosomus Schaum, 1863b: 89. Current status: valid genus in CARABIDAE: TRECHINAE: TACHYINI: XYSTOSOMINA.

Type species: Tachys inflatus Schaum, 1859, by subsequent designation (Erwin 1973: 4).

Family-group name: XYSTOSOMINA Erwin, 1994. Stem: Xystosom-.

Xystrocera Audinet-Serville, 1834b: 69. Current status: valid genus in CERAMBYCIDAE: CERAMBYCINAE: XYSTROCERINI.

Type species: Cerambyx globosus G.-A. Olivier, 1795, by subsequent designation (J. Thomson 1864: 247).

Family-group name: XYSTROCÉRITES Blanchard, 1845. Stem: Xystrocer-.

Xystropus Solier, 1835b: 241. Current status: valid genus in TENEBRIONIDAE: ALLECULINAE: ALLECULINI: XYSTROPODINA.

Type species: Helops pilosus Solier, 1835, by monotypy. Note: see Bousquet et al. (2015: 139) for comments regarding the species originally included in this genus.

Family-group name: XYSTROPIDES Solier, 1835. Stem: Xystropod-.

Yanga Muona, 1993: 25. Current status: invalid senior synonym of Muonaja Özdikmen, 2008 in EUCNEMIDAE: EUCNEMINAE: MUONAJINI. Note: junior homonym of Yanga Distant, 1904 [Hemiptera].

Type species: Yanga yonde Muona, 1993, by original designation.

Family-group name: YANGINI Muona, 1993. Stem: Yang-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

†Yixianscarabaeus Nikolajev, 2015b: 299. Current status: valid genus in SCARABAEIDAE: EREMAZINAE: †YIXIANSCARABAEINI. Note: Lower Cretaceous (China).

Type species: †Yixianscarabaeus sulcatus Nikolajev, 2015, by original designation.

Family-group name: †YIXIANSCARABAEINAE Nikolajev, 2015b: 298. Stem: Yixianscarabae-.

Zabrus Clairville, 1806: 80, 81. Current status: valid genus in CARABIDAE: HARPALINAE: ZABRINI: ZABRINA.

Type species: Carabus gibbus Fabricius, 1794 (= Carabus tenebrioides Goeze, 1777), by monotypy. Note: Goeze’s (1777) work is not consistently binominal (see ICZN 2021a) and therefore an application to the Commission is necessary to retain Carabus tenebrioides Goeze, 1777 as the valid name for the type species.

Family-group name: ZABRIDES Bonelli, 1810. Stem: Zabr-.

Zacotus J.L. LeConte, 1869a: 373. Current status: valid genus in CARABIDAE: BROSCINAE: BROSCINI: BROSCINA.

Type species: Zacotus matthewsii J.L. LeConte, 1869, by monotypy.

Family-group name: ZACOTINI G.H. Horn, 1881. Stem: Zacot-.

Zadenos Laporte, 1840b: 210. Current status: valid subgenus of Selenepistoma Dejean, 1834 in TENEBRIONIDAE: BLAPTINAE: DENDARINI: MELAMBIINA.

Type species: Opatrum longipalpe Wiedemann, 1823, by monotypy.

Family-group name: ZADENINA C. Koch, 1956. Stem: Zaden-.

Zaplous J.L. LeConte, 1878a: 415. Current status: valid genus in CERAMBYCIDAE: LAMIINAE: POGONOCHERINI.

Type species: Zaplous hubbardi J.L. LeConte, 1878 (= Ecyrus annulatus Chevrolat, 1862), by monotypy.

Family-group name: ZAPLOI J.L. LeConte & G.H. Horn, 1883. Stem: Zaplo-.

Zarax Pascoe, 1867b: 410. Current status: junior synonym of Eurypoda Saunders, 1853 in CERAMBYCIDAE: PRIONINAE: EURYPODINI.

Type species: Zarax eurypodioides Pascoe, 1867 (= Eurypoda nigrita J. Thomson, 1865), by monotypy.

Family-group name: ZARACIDES Lacordaire, 1868. Stem: Zarac-.

Zarudniola Semenov-Tjan-Shansky & Martynov in Martynov, 1925: 74. Current status: junior synonym of Karumia Escalera, 1913 in DASCILLIDAE: KARUMIINAE: KARUMIINI.

Type species: Zarudniola staphylinus Semenov-Tjan-Shansky & Martynov, 1925, by monotypy.

Family-group name: ZARUDNIOLIDAE Semenov-Tjan-Shansky & Martynov, 1925. Stem: Zarudniol-.

Zelliboria Lane, 1951: 5. Current status: valid genus in CERAMBYCIDAE: CERAMBYCINAE: PIEZOCERINI: PIEZOCERINA.

Type species: Rhagium daedaleum Perty, 1832, by original designation.

Family-group name: ZELLIBORIINAE Lane, 1951. Stem: Zellibori-.

Zelma Andrewes, 1920: 451. Current status: junior synonym of Salcedia Fairmaire, 1899 in CARABIDAE: SCARITINAE: SALCEDIINI: SALCEDIINA.

Type species: Zelma miranda Andrewes, 1920, by monotypy.

Family-group name: ZELMIDES Andrewes, 1929. Stem: Zelm-.

Zenoa Say, 1835: 153. Current status: valid genus in CALLIRHIPIDAE.

Type species: Sandalus brunneus Say, 1835 (= Melasis picea Palisot de Beauvois 1805), by monotypy.

Family-group name: ZENOINI J.L. LeConte, 1866. Stem: Zeno-.

Zenodosus Wolcott, 1910: 321. Current status: valid genus in THANEROCLERIDAE: ZENODOSINAE.

Type species: Clerus sanguineus Say, 1835, by original designation.

Family-group name: ZENODOSINI Kolibáč, 1992. Stem: Zenodos-.

Zethopsus Reitter, 1880: 85. Current status: valid genus in STAPHYLINIDAE: PSELAPHINAE: EUPLECTITAE: BYTHINOPLECTINI: BYTHINOPLECTINA. Note: new replacement name for Zethus L.W. Schaufuss, 1872.

Type species: Zethus opacus L.W. Schaufuss, 1877, by subsequent monotypy (L.W. Schaufuss 1877a: 12) (type fixation for Zethus L.W. Schaufuss).

Family-group name: ZETHOPSINA Jeannel, 1952. Stem: Zethops-.

Zethus L.W. Schaufuss, 1872: 246. Current status: invalid senior synonym of Zethopsus Reitter, 1880 in STAPHYLINIDAE: PSELAPHINAE: EUPLECTITAE: BYTHINOPLECTINI: BYTHINOPLECTINA. Note: junior homonym of Zethus Fabricius, 1805 [Hymenoptera].

Type species: Zethus opacus L.W. Schaufuss, 1877, by subsequent monotypy (L.W. Schaufuss 1877a: 12).

Family-group name: ZETHINI L.W. Schaufuss, 1890. Stem: Zeth-. Note: family-group name permanently invalid (ICZN 1999a, Article 39).

Zeugophora Kunze, 1818: 71. Current status: valid genus in MEGALOPODIDAE: ZEUGOPHORINAE. Note: name placed on the Official List of Generic Names in Zoology (ICZN 1986a).

Type species: Crioceris subspinosa Fabricius, 1781, by subsequent designation (Westwood 1838c: 42; see ICZN 1986a). Note: name placed on the Official List of Specific Names in Zoology (ICZN 1986a).

Family-group name: ZEUGOPHORINAE Böving & Craighead, 1931. Stem: Zeugophor-.

Zherichinixena Legalov, 2009b: 309, 310. Current status: valid genus in BELIDAE: OXYCORYNINAE: OXYCORYNITAE: METRIOXENINI: ZHERICHINIXENINA.

Type species: Zherichinixena nigra Legalov, 2009, by original designation.

Family-group name: ZHERICHINIXENINA Legalov, 2009. Stem: Zherichinixen-.

Zilora Mulsant, 1856b: 84. Current status: valid genus in MELANDRYIDAE: MELANDRYINAE: ZILORINI.

Type species: here fixed (under ICZN 1999a, Article 70.3.2) as Parnus obscurus Fabricius, 1794, misidentified as Xylita ferruginea Paykull, 1798 (sensu Mulsant, 1856) in the original designation by monotypy in Mulsant (1856b: 84).

Family-group name: ZILORINI Desbrochers des Loges, 1900. Stem: Zilor-.

Zilus Mulsant, 1850: 958. Current status: valid genus in COCCINELLIDAE: COCCINELLINAE: COCCIDULINI.

Type species: Scymnus fulvipes Mulsant, 1850, by monotypy.

Family-group name: ZILINI Gordon, 1985. Stem: Zil-.

Zolinopatrobus Deuve & Tian, 2001: 418. Current status: junior synonym of Lissopogonus Andrewes, 1923 in CARABIDAE: PATROBINAE: LISSOPOGONINI.

Type species: Zolinopatrobus nanlingensis Deuve & Tian, 2001, by original designation.

Family-group name: ZOLINOPATROBINA Deuve & Tian, 2001. Stem: Zolinopatrob-.

Zimmiellus Kuschel in Kuschel & Leschen, 2011: 598. Current status: valid genus in NEMONYCHIDAE: RHINORHYNCHINAE: ZIMMIELLINI.

Type species: Zimmiellus fronto Kuschel, 2011, by original designation.

Family-group name: ZIMMIELLINI Legalov, 2023c: 21. Stem: Zimmiell-. Note: ZIMMIELLINI (Legalov, 2017a: 71) was published in a work that is nomenclaturally unavailable (ICZN 2012f, Article 8.5.3).

Zolodinus Blanchard, 1853: 159. Current status: valid genus in TENEBRIONIDAE: ZOLODININAE.

Type species: Zolodinus zelandicus Blanchard, 1853, by monotypy.

Family-group name: ZOLODININAE Watt, 1975. Stem: Zolodin-.

Zoltanzonitis Bologna & Pinto, 2018: 438. Current status: valid genus in MELOIDAE: NEMOGNATHINAE: ZOLTANZONITINI.

Type species: Zonitis natala Beauregard, 1890, by original designation.

Family-group name: ZOLTANZONITINI Riccieri, Capogna, Pinto & Bologna, 2023: 109. Stem: Zoltanzonit- (ICZN 1999a, Article 29.4).

Zolus Sharp, 1886: 371. Current status: junior synonym of Oopterus Guérin-Méneville, 1841 in CARABIDAE: TRECHINAE: ZOLINI.

Type species: Zolus helmsi Sharp, 1886, by monotypy.

Family-group name: ZOLINI Sharp, 1886. Stem: Zol-.

Zonabris Harold, 1879: 134. Current status: junior synonym of Mylabris Fabricius, 1775 in MELOIDAE: MELOINAE: MYLABRINI. Note: new replacement name for Mylabris Fabricius, 1775.

Type species: Meloe cichorii Linnaeus, 1758, by subsequent designation (Latreille 1810: 430) (see nomenclatural problem with typification of Mylabris Fabricius).

Family-group name: ZONABRINI Wellman, 1908. Stem: Zonabr-.

Zonitis Fabricius, 1775: 126. Current status: valid genus in MELOIDAE: NEMOGNATHINAE: NEMOGNATHINI: ZONITIDINA. Note: name placed on the Official List of Generic Names in Zoology (ICZN 1999c).

Type species: Zonitis flava Fabriciuis, 1775, by subsequent designation (Selander 1987: 341; see ICZN 1999c). name placed on the Official List of Specific Names in Zoology (ICZN 1999c).

Family-group name: ZONITAIRES Mulsant, 1857. Stem: Zonitid- (ICZN 1999c).

Zopheromimus Alexeev & Nabozhenko, 2023: 438. Current status: valid genus in ZOPHERIDAE: ZOPHERINAE: ZOPHEROMIMINI.

Type species: Zopheromimus auriborussiensis Alexeev & Nabozhenko, 2023, by original designation. Note: combined description of a new genus and a single new species (ICZN 1999a, Article 13.4).

Family-group name: ZOPHEROMIMINI Alexeev & Nabozhenko, 2023: 437. Stem: Zopheromim-.

Zopherosis A. White, 1859: 121. Current status: valid genus in ZOPHERIDAE: ZOPHERINAE: ZOPHERINI.

Type species: Zopherosis georgii A. White, 1859, by monotypy.

Family-group name: ZOPHEROSINI Casey, 1907. Stem: Zopherose-.

Zopherus G.R. Gray in Griffith and Pidgeon, 1831: pls 50, 69. Current status: valid genus in ZOPHERIDAE: ZOPHERINAE: ZOPHERINI.

Type species: Zopherus mexicanus G.R. Gray, 1831, by monotypy. Note: a second species in this genus, Zopherus chilensis G.R. Gray, 1832, was made available in the second volume of this work published one year later (Griffith and Pidgeon 1832: 796, pl. 124).

Family-group name: ZOPHÉRITES Solier, 1834. Stem: Zopher-.

Zophosis Latreille, 1802b: 167. Current status: valid genus in TENEBRIONIDAE: PIMELIINAE: ZOPHOSINI.

Type species: Erodius testudinarius Fabricius, 1781, by monotypy.

Family-group name: ZOPHOSITES Solier, 1834. Stem: Zophos-.

Zuphium Latreille, 1805: 198, 227. Current status: valid genus in CARABIDAE: HARPALINAE: ZUPHIINI.

Type species: Carabus olens Rossi, 1790, by subsequent designation (Latreille 1810: 426).

Family-group name: ZUPHIETAE Bonelli, 1810. Stem: Zuphi-.

Zygaenodes Pascoe, 1859b: 328. Current status: junior synonym of Exechesops Schönherr, 1847 in ANTHRIBIDAE: ANTHRIBINAE: ZYGAENODINI.

Type species: Zygaenodes wollastoni Pascoe, 1859, by monotypy.

Family-group name: ZYGÉNODIDES Lacordaire, 1865. Stem: Zygaenod-.

Zygia Fabricius, 1775: 126. Current status: junior synonym of Melyris Fabricius, 1775 in MELYRIDAE: MELYRINAE: MELYRINI. Note: the First Reviser (ICZN 1999a, Article 24.2.2) for Melyris Fabricius, 1775 versus Zygia Fabricius, 1775 is Latreille (1816: 243).

Type species: Zygia oblonga Fabricius, 1775, by monotypy.

Family-group name: ZYGIIDAE Jakobson, 1911. Stem: Zygi-.

Zygobaris J.L. LeConte in J.L. LeConte and G.H. Horn, 1876: 301, 317. Current status: valid genus in CURCULIONIDAE: BARIDINAE: APOSTASIMERINI: DIORYMERINA.

Type species: Zygobaris nitens J.L. LeConte, 1876, by subsequent designation (Champion 1908: 330).

Family-group name: ZYGOBARI Pierce, 1907. Stem: Zygobarid-.

Zygocera Erichson, 1842: 223. Current status: valid genus in CERAMBYCIDAE: LAMIINAE: ZYGOCERINI. Note: this name was first used and made available by Dejean (1835: 344) but in a different taxonomic sense; we believe the best avenue is to apply to the Commission to suppress Dejean’s name for the purpose of the Principles of Priority and Homonymy (see Bousquet and Bouchard 2013a: 95).

Type species: Zygocera canosa Erichson, 1842, by monotypy.

Family-group name: ZYGOCERITAE J. Thomson, 1864. Stem: Zygocer-.

Zygogramma Chevrolat in Dejean, 1836a: 398. Current status: valid genus in CHRYSOMELIDAE: CHRYSOMELINAE: CHRYSOMELINI.

Type species: Chrysomela pulcra Fabricius, 1792 (= Chrysomela suturalis Fabricius, 1775), by subsequent designation (Motschulsky 1860b: 181).

Family-group name: ZYGOGRAMMINI Weise, 1915. Stem: Zygogrammat-.

Zygops Schönherr, 1825: 586. Current status: valid genus in CURCULIONIDAE: CONODERINAE: ZYGOPINI. Note: name placed on the Official List of Generic Names in Zoology (ICZN 1987e).

Type species: Poecilma wiedii Germar, 1823, by original designation (see ICZN 1987e). Note: placed on the Official List of Specific Names in Zoology (ICZN 1987e, as Poecilma wiedii Germar, 1824).

Family-group name: ZYGOPIDES Lacordaire, 1865. Stem: Zygop-.

Zyras Stephens, 1835a: 430. Current status: valid genus in STAPHYLINIDAE: ALEOCHARINAE: LOMECHUSINI: MYRMEDONIINA. Note: name placed on the Official List of Generic Names in Zoology (ICZN 1961a).

Type species: Aleochara haworthi Stephens, 1832, by monotypy (see ICZN 1961a). Note: name placed on the Official List of Specific Names in Zoology (ICZN 1961a).

Family-group name: ZYRINI Bradley, 1930. Stem: Zyr-.